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Silva Fennica 1926-1997
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Acta Forestalia Fennica
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Articles containing the keyword 'EC'.

Category: Commentary

article id 461, category Commentary
Sari Palmroth. (2009). Boreal forest and climate change – from processes and transport to trees, ecosystems and atmosphere. Silva Fennica vol. 43 no. 4 article id 461. https://doi.org/10.14214/sf.461
Book review: Hari, P. & Kulmala, L. (eds.). 2008. Boreal Forest and Climate Change. Advances in Global Change Research 34. Springer. 582 p. ISBN 978-1-4020-8717-2.
  • Palmroth, Duke University, Nicholas School of the Environment, Box 90328 Duke University, Durham, USA ORCID ID:E-mail: sari.palmroth@duke.edu (email)
article id 547, category Commentary
Timo Kuuluvainen. (2002). Introduction. Disturbance dynamics in boreal forests: defining the ecological basis of restoration and management of biodiversity. Silva Fennica vol. 36 no. 1 article id 547. https://doi.org/10.14214/sf.547
  • Kuuluvainen, Department of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland ORCID ID:E-mail: timo.kuuluvainen@helsinki.fi (email)

Category: Research article

article id 10242, category Research article
Shisheng Long, Siqi Zeng, Falin Liu, Guangxing Wang. (2020). Influence of slope, aspect and competition index on the height-diameter relationship of Cyclobalanopsis glauca trees for improving prediction of height in mixed forests. Silva Fennica vol. 54 no. 1 article id 10242. https://doi.org/10.14214/sf.10242
Highlights: In this study, the effects of slope, aspect and competition index (CI) on the H-DBH relationship were explored and an improved CI was developed and included to improve predictions of Cyclobalanopsis glauca tree height; It was found that the effects were statistically significant and considering slope, aspect and CI for developing the H-DBH models significantly increased the H prediction accuracy.

Diameter at breast height (DBH) and height (H) of trees are two important variables used in forest management plans. However, collecting the measurements of H is time-consuming and costly. Instead, the H-DBH relationship is modeled and used to estimate H. But, ignoring the effects of slope, aspect and tree competition on the H-DBH relationship often impedes the improvement of H predictions. In this study, to improve predictions of Cyclobalanopsis glauca (Thunb.) Oerst. tree H in mixed forests, we compared eleven H-DBH models and examined the influence of slope and aspect on the H-DBH relationship using 426 trees. We then improved Hegyi competition index and explored its effect on the H predictions by including it in the selected models. Results showed 1) There were statistically significant effects of slope and aspect on the H-DBH relationship; 2) The log transformation and exponential model performed best for sunny- and shady-steep, respectively, and the Gompertz’s model was optimal for both sunny- and shady-gentle; 3) Compared with the whole dataset, the division of the data into the slope and aspect sub-datasets significantly reduced the RMSE of H predictions; 4) Compared with the selected models without competition index, adding the original Hegyi and improved Hegyi_I into the models improved the H predictions but only the models containing the improved Hegyi_I significantly increased the prediction accuracy at the significant level of 0.1. This study implied that modeling the H-DBH relationship under different slopes and aspects and including the improved Hegyi_I provided the great potential to improve the H predictions.

  • Long, Faculty of Forestry, Central South University of Forestry and Technology, Changsha, Hunan 410004, China ORCID ID:E-mail: shisheng3604@21cn.com
  • Zeng, Faculty of Forestry, Central South University of Forestry and Technology, Changsha, Hunan 410004, China ORCID ID:E-mail: zengsiqi@21cn.com
  • Liu, Faculty of Forestry, Central South University of Forestry and Technology, Changsha, Hunan 410004, China ORCID ID:E-mail: liufl680@126.com
  • Wang, Research Center of Forestry Remote Sensing & Information Engineering, Central South University of Forestry and Technology, Changsha 410004, China; Department of Geography and Environmental Resources, Southern Illinois University, Carbondale, IL 62901, USA ORCID ID:E-mail: gxwang@siu.edu (email)
article id 10242, category Research article
Shisheng Long, Siqi Zeng, Falin Liu, Guangxing Wang. (2020). Influence of slope, aspect and competition index on the height-diameter relationship of Cyclobalanopsis glauca trees for improving prediction of height in mixed forests. Silva Fennica vol. 54 no. 1 article id 10242. https://doi.org/10.14214/sf.10242
Highlights: In this study, the effects of slope, aspect and competition index (CI) on the H-DBH relationship were explored and an improved CI was developed and included to improve predictions of Cyclobalanopsis glauca tree height; It was found that the effects were statistically significant and considering slope, aspect and CI for developing the H-DBH models significantly increased the H prediction accuracy.

Diameter at breast height (DBH) and height (H) of trees are two important variables used in forest management plans. However, collecting the measurements of H is time-consuming and costly. Instead, the H-DBH relationship is modeled and used to estimate H. But, ignoring the effects of slope, aspect and tree competition on the H-DBH relationship often impedes the improvement of H predictions. In this study, to improve predictions of Cyclobalanopsis glauca (Thunb.) Oerst. tree H in mixed forests, we compared eleven H-DBH models and examined the influence of slope and aspect on the H-DBH relationship using 426 trees. We then improved Hegyi competition index and explored its effect on the H predictions by including it in the selected models. Results showed 1) There were statistically significant effects of slope and aspect on the H-DBH relationship; 2) The log transformation and exponential model performed best for sunny- and shady-steep, respectively, and the Gompertz’s model was optimal for both sunny- and shady-gentle; 3) Compared with the whole dataset, the division of the data into the slope and aspect sub-datasets significantly reduced the RMSE of H predictions; 4) Compared with the selected models without competition index, adding the original Hegyi and improved Hegyi_I into the models improved the H predictions but only the models containing the improved Hegyi_I significantly increased the prediction accuracy at the significant level of 0.1. This study implied that modeling the H-DBH relationship under different slopes and aspects and including the improved Hegyi_I provided the great potential to improve the H predictions.

  • Long, Faculty of Forestry, Central South University of Forestry and Technology, Changsha, Hunan 410004, China ORCID ID:E-mail: shisheng3604@21cn.com
  • Zeng, Faculty of Forestry, Central South University of Forestry and Technology, Changsha, Hunan 410004, China ORCID ID:E-mail: zengsiqi@21cn.com
  • Liu, Faculty of Forestry, Central South University of Forestry and Technology, Changsha, Hunan 410004, China ORCID ID:E-mail: liufl680@126.com
  • Wang, Research Center of Forestry Remote Sensing & Information Engineering, Central South University of Forestry and Technology, Changsha 410004, China; Department of Geography and Environmental Resources, Southern Illinois University, Carbondale, IL 62901, USA ORCID ID:E-mail: gxwang@siu.edu (email)
article id 10084, category Research article
Mihails Čugunovs, Eeva-Stiina Tuittila, Jari Kouki. (2020). Proximity to charred logs in burned forests likely affects decomposition processes in the soil. Silva Fennica vol. 54 no. 1 article id 10084. https://doi.org/10.14214/sf.10084
Highlights: Standardised organic substrate decomposition was tentatively observed to be faster adjacent to non-charred downed logs than away from the logs or adjacent to charred logs; A spatial linkage was observed between non-charred logs and decomposition in the soil in burned boreal forests; Proximity to a charred log may provide a micro-environment where decomposition rates differ from the surrounding forest soil.

We studied the spatial decomposition rates of standardised organic substrates in soils (burned boreal pine-dominated sub-xeric forests in eastern Finland), with respect to charred and non-charred coarse woody debris (CWD). Decomposition rates of rooibos plant litter inside teabags (C:N = 42.870 ± 1.841) and pressed-sheet Nordic hardwood pulp (consisting of mainly alpha-cellulose) were measured at 0.2 m distance from 20 charred (LC0.2) and 40 non-charred logs (LNC0.2). We also measured decomposition at 60 plots located 3–10 m away from downed logs (L3,10). The rooibos decomposition rate constant ‘k’ was 8.4% greater at the LNC0.2 logs than at the L3,10 or LC0.2 logs. Cellulose decomposed more completely in 1 micron mesh bags at LNC0.2 (44% of buried bags had leftover material) than at LC0.2 (76%) or L3,10 (70%). Decomposition of cellulose material was rapid but varied greatly between sampling plots. Our results indicate that decomposition of the standardised organic matter was more rapid close to CWD pieces than further away. However, only the plots located near non-charred logs (LNC0.2) exhibited high decomposition rates, with no corresponding increase observed at the charred logs (LC0.2). This suggests a possible noteworthy indirect effect of forest burning on soil organic matter (SOM) decomposition rates close to charred CWD after forest fires. We urge for more studies on this tentative observation as it may affect the estimates on how fires affect carbon cycling in forests.

  • Čugunovs, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: mihails.cugunovs@gmail.com (email)
  • Tuittila, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID: https://orcid.org/0000-0001-8861-3167 E-mail: eeva-stiina.tuittila@uef.fi
  • Kouki, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID: https://orcid.org/0000-0003-2624-8592 E-mail: jari.kouki@uef.fi
article id 10151, category Research article
Jyri Hietala, Riitta Hänninen, Matleena Kniivilä, Anne Toppinen. (2019). Networks in international opportunity recognition among Finnish wood product industry SMEs. Silva Fennica vol. 53 no. 4 article id 10151. https://doi.org/10.14214/sf.10151
Highlights: In line with earlier literature, we found the networks in our study to positively impact international opportunity recognition; Despite the reliance on various network forms and levels, a strategic stance towards opportunity recognition can be characterized as being more reactive than proactive; Institutional networks represented a more systematic way of recognizing international opportunities among case companies.

Bioeconomy development will create new opportunities for firms operating in the international wood products markets, and identifying and exploiting these opportunities is emphasized as a key concept to achieving business success. Our study will attempt to address a gap in the literature on sawmill industry business development from the viewpoint of international opportunity recognition. The aim of our study is to provide a holistic description on how small and medium-sized enterprises (SMEs) in the wood products industry recognize and exploit international business opportunities, and how they utilize network perspectives in this context. The subject was examined through Finnish wood product industry SMEs by interviewing 11 managers and industry representatives. The results suggest that SMEs recognize international opportunities reactively per se. Social networks formed in professional forums were an important information channel for identifying international opportunities. Through vertical business networks, such as sales agents, firms have been able to increase their international market presence and free their own resources for other important activities. Horizontal dyadic business networks were seen to facilitate new international opportunities through cooperation, while excessive reliance on vertical networks raised concerns and seemed not to be effective in international opportunity recognition. Institutional networks formed a systematic way of recognizing international opportunities, but more so at the initial market entry stage.

  • Hietala, United Bankers, Aleksanterinkatu 21 A, FI-00100 Helsinki, Finland ORCID ID:E-mail: jyri.hietala@unitedbankers.fi (email)
  • Hänninen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID ID:E-mail: riitta.hanninen@luke.fi
  • Kniivilä, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID ID:E-mail: matleena.kniivila@luke.fi
  • Toppinen, University of Helsinki, Helsinki Institute of Sustainability Science, Latokartanonkaari 7, P.O. 27, FI-00014 University of Helsinki, Finland ORCID ID:E-mail: anne.toppinen@helsinki.fi
article id 10179, category Research article
Lauri Korhonen, Jaakko Repola, Tomi Karjalainen, Petteri Packalen, Matti Maltamo. (2019). Transferability and calibration of airborne laser scanning based mixed-effects models to estimate the attributes of sawlog-sized Scots pines. Silva Fennica vol. 53 no. 3 article id 10179. https://doi.org/10.14214/sf.10179
Highlights: Attributes of individual sawlog-sized pines estimated by transferring ALS-based models between sites; Mixed effects models were more accurate than k-NN imputation tested earlier; Calibration with a small number of field measured trees improved the accuracy.

Airborne laser scanning (ALS) data is nowadays often available for forest inventory purposes, but adequate field data for constructing new forest attribute models for each area may be lacking. Thus there is a need to study the transferability of existing ALS-based models among different inventory areas. The objective of our study was to apply ALS-based mixed models to estimate the diameter, height and crown base height of individual sawlog sized Scots pines (Pinus sylvestris L.) at three different inventory sites in eastern Finland. Different ALS sensors and acquisition parameters were used at each site. Multivariate mixed-effects models were fitted at one site and the models were validated at two independent test sites. Validation was carried out by applying the fixed parts of the mixed models as such, and by calibrating them using 1–3 sample trees per plot. The results showed that the relative RMSEs of the predictions were 1.2–6.5 percent points larger at the test sites compared to the training site. Systematic errors of 2.4–6.2 percent points also emerged at the test sites. However, both the RMSEs and the systematic errors decreased with calibration. The results showed that mixed-effects models of individual tree attributes can be successfully transferred and calibrated to other ALS inventory areas in a level of accuracy that appears suitable for practical applications.

  • Korhonen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID: http://orcid.org/0000-0002-9352-0114 E-mail: lauri.korhonen@uef.fi (email)
  • Repola, Natural Resources Institute of Finland (Luke), Natural resources, Eteläranta 55, FI-96300 Rovaniemi, Finland ORCID ID:E-mail: jaakko.repola@luke.fi
  • Karjalainen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: tomikar@uef.fi
  • Packalen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: petteri.packalen@uef.fi
  • Maltamo, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: matti.maltamo@uef.fi
article id 10074, category Research article
Sebastian Kühle, Alfred Teischinger, Manfred Gronalt. (2019). Optimal location of laminated beech production plants within the solid hardwood supply network in Austria. Silva Fennica vol. 53 no. 3 article id 10074. https://doi.org/10.14214/sf.10074
Highlights: This paper provides data to the solid hardwood business and develops a mixed integer linear program model to design a laminated beech wood supply network; It covers the strategic decision where to locate a new production facility within the existing supply network with the lowest supply network cost; Sufficient sawn wood suppliers and potential facility locations are provided.

Due to changes in forest management in various European countries, hardwood forest areas and amounts will increase. Sustainable and individual utilization concepts have to be developed for the upcoming available resource. Studies conclude that there is low potential for hardwoods in the traditional appearance market thus the application areas have to be extended to new structural innovative products. This paper examines the extension to a future laminated beech wood supply network which would be a combination of already existing and new production facilities. For a better future use of hardwood raw materials it is necessary to consider the entire supply chain. This also better shows a total hardwood value chain. Therefore, this paper provides data to the solid hardwood business and develops a mixed integer linear programming to design a laminated beech wood supply network. The model is applied to Austria as the sample region. It covers the important strategic decisions where to locate a downstream facility within the existing production network with the lowest supply network cost. Fourteen scenarios are developed to examine various future network configurations. Results about optimal material flows and used sawmills as well as downstream production facilities are presented in form of material and financial performances. Two optimal laminated beech production locations are determined by the calculated scenarios results, and the impact of a new sawmill is analyzed which is focused on beech.

  • Kühle, BOKU - University of Natural Resources and Life Sciences, Vienna, Department of Material Science and Process Engineering, and Renewable Institute of Wood Technology Materials, Konrad-Lorenz-Straße 24, 3430 Tulln, Austria ORCID ID:E-mail: skuehle@boku.ac.at (email)
  • Teischinger, BOKU - University of Natural Resources and Life Sciences, Vienna, Department of Material Science and Process Engineering, and Renewable Institute of Wood Technology Materials, Konrad-Lorenz-Straße 24, 3430 Tulln, Austria ORCID ID:E-mail: alfred.teischinger@boku.ac.at
  • Gronalt, BOKU - University of Natural Resources and Life Sciences, Vienna, Department of Economics and Social Sciences, Institute of Production and Logistics, Feistmantelstraße 4, 1180 Vienna, Austria ORCID ID:E-mail: manfred.gronalt@boku.ac.at
article id 10043, category Research article
Outi Manninen, Rainer Peltola. (2019). Continuous picking may increase bilberry yields. Silva Fennica vol. 53 no. 3 article id 10043. https://doi.org/10.14214/sf.10043
Highlights: Bilberry fruit production and fruit set increased under continuous picking by rake in three-year study; Bilberry flower number and fruit mass were not affected by picking; Bilberry compensated for biomass loss; The highest relative deciduous species abundance was found in the picking treatment plots at the end of the experiment.

Accumulated knowledge about the health benefits of bilberry (Vaccinium myrtillus L.) has increased the demand and utilization of wild bilberries. Intensive berry picking by metal rakes is believed to cause damage in bilberry stands in areas under continuous picking pressure, and hence expected to hamper the production of berries in forthcoming years. We conducted an experiment to examine the effect of continuous bilberry picking by metal rake on the number of bilberry flowers and fruits, fruit mass, compensation for biomass loss after picking, and plant functional type abundance in the understorey in northern Finland. Bilberry lost less than 0.5% of its biomass annually during the three-year study period due to rake harvesting. The number of flowers was not significantly affected by damage caused by picking, while both fruit production and fruit set increased without any indication of reduced fruit mass, and biomass loss was fully compensated. Moreover, the relative abundance of plant functional types was not affected by picking during the study. We suggest that the low intensity and timing of damage act as a buffer against the adverse effects of picking on bilberry fruit production. On the basis of this study, it is reasonable to anticipate that there are no indications that current intensive berry picking would not be on a sustainable level.

  • Manninen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Paavo Havaksentie 3, FI-90014 University of Oulu, Finland ORCID ID:E-mail: outi.manninen@luke.fi (email)
  • Peltola, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Ounasjoentie 6, FI-96200 Rovaniemi, Finland ORCID ID:E-mail: rainer.peltola@luke.fi
article id 10088, category Research article
Antti-Jussi Lindroos, Kira Ryhti, Tomi Kaakkurivaara, Jori Uusitalo, Heljä-Sisko Helmisaari. (2019). Leaching of heavy metals and barium from forest roads reinforced with fly ash. Silva Fennica vol. 53 no. 2 article id 10088. https://doi.org/10.14214/sf.10088
Highlights: Heavy metal concentrations were generally low in percolation and ditch water samples of ash roads, but elevated concentrations were found in some parts of ash roads; Risk for heavy metal leaching is negligible if road parts producing high concentrations are rare.

The aim of this study was to determine the effect of leaching of heavy metals (Cr, As, Cd, Cu, Ni, Pb, Zn, Co, Mo) and earth-alkaline metal, barium (Ba), on the percolation and ditch water quality from the forest roads that contained ash in the road structures. Water quality was studied in the immediate vicinity below the ash layers as well as deeper in the road structure. Water quality was also determined in the drainage water in ditches that crossed the forest roads. A mixture of wood and peat based fly ash was used in the road structures. The treatments were: 1) no ash, 2) a 15 cm layer of ash/gravel mixture, 3) a 20 cm layer of ash/gravel mixture, 4) a 25 cm layer of ash, and 5) a 50 cm layer of ash. Large variation in the concentrations of Cr, As, Cu, Ni, Pb, Mo and Ba in the percolation water, even within the same treatment, caused difficulties to generalize the results. The concentrations of Cr, As, Ni, Pb, Mo and Ba in water samples were high in some treatment plot lysimeters containing ash compared to the control (no ash). On the other hand, many lysimeters had low and similar concentrations in water samples in the treatment plots containing ash compared to concentrations in the control plots. The ash in the roads did not affect the concentrations in the ditches. The leaching is uneven and seems to take place only from some parts of the ash layer. Risk for leaching is minimal if such parts are not widely spread.

  • Lindroos, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID ID:E-mail: antti.lindroos@luke.fi (email)
  • Ryhti, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID ID:E-mail: kira.ryhti@helsinki.fi
  • Kaakkurivaara, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID ID:E-mail: tomi.kaakkurivaara@gmail.com
  • Uusitalo, Natural Resources Institute Finland (Luke), Korkeakoulunkatu 7, FI-33720 Tampere, Finland ORCID ID:E-mail: jori.uusitalo@luke.fi
  • Helmisaari, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID ID:E-mail: helja-sisko.helmisaari@helsinki.fi
article id 10068, category Research article
Lari Melander, Risto Ritala, Markus Strandström. (2019). Classifying soil stoniness based on the excavator boom vibration data in mounding operations. Silva Fennica vol. 53 no. 2 article id 10068. https://doi.org/10.14214/sf.10068
Highlights: An excavator was equipped with an inertial measurement unit for taking automatic measurements of soil stoniness during mounding work; Supervised machine-learning classifiers were trained utilizing both the automatically measured data and manual stoniness measurements; The class prediction for the soil stoniness achieved an accuracy of 70% when assigned to constant grid cells.

The stoniness index of forest soil describes the stone content in the upper soil layer at depths of 20–30 centimeters. This index is not available in any existing map databases, and traditional measurements for the stoniness of the soil have always necessitated laborious soil-penetration methods. Knowledge of the stone content of a forest site could be of use in a variety of forestry operations. This paper presents a novel approach to obtaining automatic measurements of soil stoniness during an excavator-based mounding operation. The excavator was equipped with only a low-cost inertial measurement unit and a satellite navigation receiver. Using the data from these sensors and manually conducted soil stoniness measurements, supervised machine learning methods were utilized to build a model that is capable of predicting the stoniness class of a given mounding location. This study compares different classifiers and feature selection methods to find the most promising solution for this learning problem. The discussion includes a proposition for a meaningful measurement resolution of the soil’s stoniness, and a practical method for evaluating the variability of the stone content of the soil. The results indicate that it is possible to predict the soil stoniness class with 70% accuracy using only the inertial and location measurements.

  • Melander, Automation Technology and Mechanical Engineering, Tampere University, FI-33014 Tampere University, Finland ORCID ID: http://orcid.org/0000-0003-3662-5187 E-mail: lari.melander@tuni.fi (email)
  • Ritala, Automation Technology and Mechanical Engineering, Tampere University, FI-33014 Tampere University, Finland ORCID ID: http://orcid.org/0000-0003-0721-9948 E-mail: risto.ritala@tuni.fi
  • Strandström, Metsäteho Oy, Vernissakatu 1, FI-01300 Vantaa, Finland ORCID ID:E-mail: markus.strandstrom@metsateho.fi
article id 10048, category Research article
Urszula Zajączkowska, Karina Kaczmarczyk, Janusz Liana. (2019). Birch sap exudation: influence of tree position in a forest stand on birch sap production, trunk wood anatomy and radial bending strength. Silva Fennica vol. 53 no. 2 article id 10048. https://doi.org/10.14214/sf.10048
Highlights: Birch trees along the forest edge exude more xylem sap but less concentrated than the trees from the interior; Radial bending strength of wood in birch trunk is higher in the trees from forest edge; Trees exhibit higher bending strength in western side of the trunk, where the number of vessels and the wood potential conductivity index are smaller.

It is commonly accepted that the period of early-spring xylem sap exudation marks a stage during which a positive pressure builds inside the tree trunks. This state changes when leaves appear, initiating water transport within the trunk. It is unknown, however, how the wood anatomical structure and its mechanical resistance influences the sap. We present the results of research on the relationship between exudation of sap from Betula pendula Roth trees from the interior of a forest stand and from its edge, and the anatomical structure of the trunk wood and its bending strength. During the period between March 21 and April 18, we performed five sets of measurements of sap exudation from trees at the edge of the stand and from the forest interior. The resulting radial wood samples were tested for bending strength using a fractometer. We tested the sap for electrolytic conductivity and sugars content. For the anatomical analysis of the wood, we determined the number of vessels per 1 mm2, average vessel lumen area and potential conductivity index. We found that the trees along the edge of the stand exude more sap, but it is less concentrated than the sap from the trees from the interior. Bending strength perpendicular to wood fibres is higher in the trees from the stand edge and in the western side of the trunk, where the number of vessels per 1 mm2 and conductivity index are smaller. Seemingly, this is the result of western winds, which are dominant in Poland.

  • Zajączkowska, Department of Forest Botany, Faculty of Forestry, Warsaw University of Life Sciences, 159 Nowoursynowska St., 02-776 Warsaw, Poland ORCID ID:E-mail: urszula.zajaczkowska@wl.sggw.pl (email)
  • Kaczmarczyk, Department of Forest Botany, Faculty of Forestry, Warsaw University of Life Sciences, 159 Nowoursynowska St., 02-776 Warsaw, Poland ORCID ID:E-mail: karina.kaczmarczyk@wl.sggw.pl
  • Liana, Department of Forest Botany, Faculty of Forestry, Warsaw University of Life Sciences, 159 Nowoursynowska St., 02-776 Warsaw, Poland ORCID ID:E-mail: janusz.liana@wl.sggw.pl
article id 10052, category Research article
Pentti Niemistö, Harri Kilpeläinen, Henrik Heräjärvi. (2019). Effect of pruning season and tool on knot occlusion and stem discolouration in Betula pendula – situation five years after pruning. Silva Fennica vol. 53 no. 1 article id 10052. https://doi.org/10.14214/sf.10052
Highlights: The occlusion was the fastest in the case of small living branches of fast growing trees pruned in springtime; Occlusion was quicker after saw pruning than after secateurs pruning, due to shorter knot stubs; Branches that were pruned in living state occluded faster than the ones pruned as dead; Dead branches hit down with a stick occluded slowly.

This paper investigates and models the effects of pruning season and tool on wound occlusion with varying tree and branch characteristics of silver birch (Betula pendula Roth) stems at the pruning height of 0−4 metres. Dates of eight secateurs prunings, three saw prunings and two sticks prunings as well as unpruned control were tested in permanent plots on four sites. Knot occlusion and discolouration in stemwood were measured from about 1600 studied knots of 112 sample trees felled five to six years after pruning in 2010. Knot occlusion rate was modelled according to pruning tool, date, tree growth, and branch characteristics. The occlusion was the fastest in trees pruned in spring or early summer, and the slowest in trees pruned in autumn. Stubs of living branches occluded faster than the dead ones with the same diameter. Saw pruning resulted in clearly better occlusion rates than secateurs pruning, caused by the shorter knot stubs after saw pruning. Hitting dead branches away with a stick resulted in the worst occlusion status. The colour defects spread more often upward from the knot than downward. Discolouration in stemwood was detected more frequently near to the pruned branches than the unpruned ones, and more widely near to the stubs of dead branches than the living ones. Most saw and secateurs pruned branches were completely occluded during the experiment, so these prunings were suitable for all branches under 20 mm in diameter, and for living branches even up to 30 mm in fast-growing trees.

  • Niemistö, Natural Resources Institute Finland (Luke), Natural resources, Kampusranta 9 C, FI-60320 Seinäjoki, Finland ORCID ID:E-mail: pentti.niemisto@luke.fi (email)
  • Kilpeläinen, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6 B, FI-80100 Joensuu, Finland ORCID ID:E-mail: harri.kilpelainen@luke.fi
  • Heräjärvi, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6 B, FI-80100 Joensuu, Finland ORCID ID:E-mail: henrik.herajarvi@luke.fi
article id 9918, category Research article
Ari Nikula, Vesa Nivala, Juho Matala, Kari Heliövaara. (2019). Modelling the effect of habitat composition and roads on the occurrence and number of moose damage at multiple scales. Silva Fennica vol. 53 no. 1 article id 9918. https://doi.org/10.14214/sf.9918
Highlights: The occurrence and number of moose damage were modelled with a zero-inflated count model; An admixture of mature forests within plantations increased the number of damage; Vicinity of inhabited areas and roads reduced damage; Plantations in landscapes with a large amount of pine-dominated thinning forests had less damage in Lapland; Damage risk assessment should include characteristics specific to each region.

We modelled the effect of habitat composition and roads on the number and occurrence of moose (Alces alces L.) damage in Ostrobothnia and Lapland using a zero-inflated count model. Models were developed for 1 km2, 25 km2 and 100 km2 landscapes consisting of equilateral rectangular grid cells. Count models predict the number of damage, i.e. the number of plantations and zero models the probability of a landscape being without damage for a given habitat composition. The number of moose damage in neighboring grid cells was a significant predictor in all models. The proportion of mature forest was the most frequent significant variable, and an increasing admixture of mature forests among plantations increased the number and occurrence of damage. The amount of all types of plantations was the second most common significant variable predicting increasing damage along with increasing amount of plantations. An increase in thinning forests as an admixture also increased damage in 1 km2 landscapes in both areas, whereas an increase in pine-dominated thinning forests in Lapland reduced the number of damage in 25 km2 landscapes. An increasing amount of inhabited areas in Ostrobothnia and the length of connecting roads in Lapland reduced the number of damage in 1 and 25 km2 landscapes. Differences in model variables between areas suggest that models of moose damage risk should be adjusted according to characteristics that are specific to the study area.

  • Nikula, Natural Resources Institute Finland (Luke), Bioeconomy and Environment, Ounasjoentie 6, FI-96200 Rovaniemi, Finland ORCID ID:E-mail: ari.nikula@luke.fi (email)
  • Nivala, Natural Resources Institute Finland (Luke), Bioeconomy and Environment, Ounasjoentie 6, FI-96200 Rovaniemi, Finland ORCID ID:E-mail: vesa.nivala@luke.fi
  • Matala, Natural Resources Institute Finland (Luke), Natural resources, Yliopistokatu 6, FI-80100 Joensuu, Finland ORCID ID:E-mail: juho.matala@luke.fi
  • Heliövaara, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID ID:E-mail: kari.heliovaara@helsinki.fi
article id 10036, category Research article
Karin Hjelm, Lars Rytter. (2018). The demand of hybrid aspen (Populus tremula × P. tremuloides) on site conditions for a successful establishment on forest land. Silva Fennica vol. 52 no. 5 article id 10036. https://doi.org/10.14214/sf.10036
Highlights: Low pH (below 3.5) reduced growth, but not survival, in a greenhouse study; Site preparation methods did not affect survival in field, but differences were found for growth; Mounding had generally the best effect on growth; Clonal differences were found that could be useful for improving establishment and growth.

Hybrid aspen (Populus tremula L. × P. tremuloides Michx.) is a deciduous tree species suitable for producing large amounts of renewable biomass during short rotations. Its potential under North European conditions could be largely extended if not only agricultural land but also forest land was used for cultivation. Unfortunately, the knowledge of appropriate forest site conditions and effects of site preparation methods on hybrid aspen establishment is limited. In this paper, two studies that explore these questions are presented. In the first study, the sensitivity to acid soils was tested under greenhouse conditions in two type of soils: a) peat soil limed to certain pH levels (3.4–5.7) and b) collected forest soils where pH varied from 3.9 to 5.3. The lowest pH level resulted in reduced growth, elsewhere no significant differences were found. The second study was applied in the field and investigated the effect of four site preparation methods on survival and growth. The methods were: 1) control with no site preparation, 2) patch scarification, 3) mounding and 4) soil inversion. While no differences were found for survival, mounding was generally the method with the highest growth and patch scarification was least successful. The result was probably an effect of good soil aeration and less competition from vegetation after mounding. The field study also revealed clonal differences in growth performance, which stresses the importance of clone selection prior to planting. The results of these studies indicate that hybrid aspen is less sensitive to variation in pH and site preparation methods compared with other poplar species, as have been found in similar studies.

  • Hjelm, Skogforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden ORCID ID:E-mail: karin.hjelm@skogforsk.se (email)
  • Rytter, Skogforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden ORCID ID: http://orcid.org/0000-0001-6183-4832 E-mail: lars.rytter@skogforsk.se
article id 10001, category Research article
Karoliina Hämäläinen, Teemu Tahvanainen, Kaisa Junninen. (2018). Characteristics of boreal and hemiboreal herb-rich forests as habitats for polypore fungi. Silva Fennica vol. 52 no. 5 article id 10001. https://doi.org/10.14214/sf.10001
Highlights: Polypore species richness and diversity were affected positively by dead-wood diversity, and negatively by increasing latitude; Red-listed species responded only to the abundance of large-diameter dead wood; Main factor determining composition of polypore assemblages was host-tree species; High proportion of deciduous dead-wood in herb-rich forests provides complementary effect on polypore assemblages in boreal forest landscapes.

Herb-rich forests are often considered biodiversity hotspots in the boreal zone but their fungal assemblages, particularly those of wood-decaying fungi, remain poorly known. We studied herb-rich forests as habitats for polypores, a distinct group of wood-decaying fungi, and assessed the importance of tree- and stand-scale variables for polypore species richness, abundance, and diversity, including red-listed species. The data include 71 herb-rich forest stands in Finland and 4797 dead wood items, on which we made 2832 observations of 101 polypore species. Dead-wood diversity was the most important variable explaining polypore species richness and diversity, whereas increasing latitude had a negative effect. Red-listed species showed a positive response to the abundance of large-diameter dead wood, which, especially birch, supported also high general abundance of polypores. The composition of polypore assemblages reflected their host-tree species. The red-listed species did not show explicit patterns in the ordination space. Compared to old-growth spruce forests, herb-rich forests seem to host lower polypore species richness and less red-listed species. However, because of high proportion of deciduous trees in the dead wood profile, herb-rich forests have a clear complementary effect on polypore assemblages in boreal forest landscapes.

  • Hämäläinen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: karoham@uef.fi (email)
  • Tahvanainen, Department of Environmental and Biological Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: teemu.tahvanainen@uef.fi
  • Junninen, Metsähallitus Parks & Wildlife Finland, c/o UEF/Borealis, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: kaisa.junninen@metsa.fi
article id 10001, category Research article
Karoliina Hämäläinen, Teemu Tahvanainen, Kaisa Junninen. (2018). Characteristics of boreal and hemiboreal herb-rich forests as habitats for polypore fungi. Silva Fennica vol. 52 no. 5 article id 10001. https://doi.org/10.14214/sf.10001
Highlights: Polypore species richness and diversity were affected positively by dead-wood diversity, and negatively by increasing latitude; Red-listed species responded only to the abundance of large-diameter dead wood; Main factor determining composition of polypore assemblages was host-tree species; High proportion of deciduous dead-wood in herb-rich forests provides complementary effect on polypore assemblages in boreal forest landscapes.

Herb-rich forests are often considered biodiversity hotspots in the boreal zone but their fungal assemblages, particularly those of wood-decaying fungi, remain poorly known. We studied herb-rich forests as habitats for polypores, a distinct group of wood-decaying fungi, and assessed the importance of tree- and stand-scale variables for polypore species richness, abundance, and diversity, including red-listed species. The data include 71 herb-rich forest stands in Finland and 4797 dead wood items, on which we made 2832 observations of 101 polypore species. Dead-wood diversity was the most important variable explaining polypore species richness and diversity, whereas increasing latitude had a negative effect. Red-listed species showed a positive response to the abundance of large-diameter dead wood, which, especially birch, supported also high general abundance of polypores. The composition of polypore assemblages reflected their host-tree species. The red-listed species did not show explicit patterns in the ordination space. Compared to old-growth spruce forests, herb-rich forests seem to host lower polypore species richness and less red-listed species. However, because of high proportion of deciduous trees in the dead wood profile, herb-rich forests have a clear complementary effect on polypore assemblages in boreal forest landscapes.

  • Hämäläinen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: karoham@uef.fi (email)
  • Tahvanainen, Department of Environmental and Biological Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: teemu.tahvanainen@uef.fi
  • Junninen, Metsähallitus Parks & Wildlife Finland, c/o UEF/Borealis, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: kaisa.junninen@metsa.fi
article id 10001, category Research article
Karoliina Hämäläinen, Teemu Tahvanainen, Kaisa Junninen. (2018). Characteristics of boreal and hemiboreal herb-rich forests as habitats for polypore fungi. Silva Fennica vol. 52 no. 5 article id 10001. https://doi.org/10.14214/sf.10001
Highlights: Polypore species richness and diversity were affected positively by dead-wood diversity, and negatively by increasing latitude; Red-listed species responded only to the abundance of large-diameter dead wood; Main factor determining composition of polypore assemblages was host-tree species; High proportion of deciduous dead-wood in herb-rich forests provides complementary effect on polypore assemblages in boreal forest landscapes.

Herb-rich forests are often considered biodiversity hotspots in the boreal zone but their fungal assemblages, particularly those of wood-decaying fungi, remain poorly known. We studied herb-rich forests as habitats for polypores, a distinct group of wood-decaying fungi, and assessed the importance of tree- and stand-scale variables for polypore species richness, abundance, and diversity, including red-listed species. The data include 71 herb-rich forest stands in Finland and 4797 dead wood items, on which we made 2832 observations of 101 polypore species. Dead-wood diversity was the most important variable explaining polypore species richness and diversity, whereas increasing latitude had a negative effect. Red-listed species showed a positive response to the abundance of large-diameter dead wood, which, especially birch, supported also high general abundance of polypores. The composition of polypore assemblages reflected their host-tree species. The red-listed species did not show explicit patterns in the ordination space. Compared to old-growth spruce forests, herb-rich forests seem to host lower polypore species richness and less red-listed species. However, because of high proportion of deciduous trees in the dead wood profile, herb-rich forests have a clear complementary effect on polypore assemblages in boreal forest landscapes.

  • Hämäläinen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: karoham@uef.fi (email)
  • Tahvanainen, Department of Environmental and Biological Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: teemu.tahvanainen@uef.fi
  • Junninen, Metsähallitus Parks & Wildlife Finland, c/o UEF/Borealis, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: kaisa.junninen@metsa.fi
article id 10013, category Research article
Hardo Becker, Jürgen Aosaar, Mats Varik, Gunnar Morozov, Kristiina Aun, Ülo Mander, Kaido Soosaar, Veiko Uri. (2018). Annual net nitrogen mineralization and litter flux in well-drained downy birch, Norway spruce and Scots pine forest ecosystems. Silva Fennica vol. 52 no. 4 article id 10013. https://doi.org/10.14214/sf.10013
Highlights: The net nitrogen mineralization (NNM) flux in drained peat soils depends largely on the C/N ratio and tree species; The soil NNM process is affected by trees through organic litter input into soil; Pine stand in low-fertility drained transitional bog is dominated by net ammonification; Birch and spruce stands on the fertile drained peat soil with higher pH and N content are dominated by net nitrification.

The main aim of the current study was to estimate the annual net nitrogen mineralization (NNM) flux in stands of different tree species growing on drained peatlands, as well as to clarify the effect of tree species, soil properties and litter on annual NNM dynamics. Three study sites were set up in May 2014: a downy birch (Betula pubescens Ehrh.) stand and a Norway spruce (Picea abies (L.) Karst.) stand in Oxalis full-drained swamp (ODS) and a Scots pine (Pinus sylvestris L.) stand in Myrtillus full-drained swamp (MDS). The NNM flux was estimated using the in situ method with incubated polyethylene bags. The highest value of NNM was found in stands that were growing on fertile ODS: 127.5 kg N ha–1 yr–1 and 87.7 kg N ha–1 yr–1, in the downy birch stand and in the Norway spruce stand, respectively. A significantly lower annual NNM flux (11.8 kg N ha–1 yr–1) occurred in the Scots pine stand growing in MDS. Nitrification was highest at fertile ODS sites and ammonification was the highest at the low fertility MDS site. For all study sites, positive correlation was found between soil temperature and NNM intensity. The difference in annual NNM between the downy birch stand and the Norway spruce stand growing on similar drained fertile peatlands was due to litter quality. The annual N input into the soil through leaf litter was the highest at the downy birch site where also the C/N ratio of litter was the lowest. The second highest N input into the soil was found in the spruce stand and the lowest in the pine stand.

  • Becker, Chair of Silviculture and Forest Ecology, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Friedrich Reinhold Kreutzwaldi 1, 51014 Tartu, Estonia ORCID ID:E-mail: hardo.becker@emu.ee (email)
  • Aosaar, Chair of Silviculture and Forest Ecology, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Friedrich Reinhold Kreutzwaldi 1, 51014 Tartu, Estonia ORCID ID:E-mail: jyrgen.aosaar@emu.ee
  • Varik, Chair of Silviculture and Forest Ecology, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Friedrich Reinhold Kreutzwaldi 1, 51014 Tartu, Estonia ORCID ID:E-mail: mats.varik@emu.ee
  • Morozov, Chair of Silviculture and Forest Ecology, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Friedrich Reinhold Kreutzwaldi 1, 51014 Tartu, Estonia ORCID ID:E-mail: gunnar.morozov@emu.ee
  • Aun, Chair of Silviculture and Forest Ecology, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Friedrich Reinhold Kreutzwaldi 1, 51014 Tartu, Estonia ORCID ID:E-mail: kristiina.aun@emu.ee
  • Mander, Department of Geography, Institute of Ecology & Earth Sciences, University of Tartu, Ülikooli 18, 50090 Tartu, Estonia ORCID ID:E-mail: ulo.mander@ut.ee
  • Soosaar, Department of Geography, Institute of Ecology & Earth Sciences, University of Tartu, Ülikooli 18, 50090 Tartu, Estonia ORCID ID:E-mail: kaido.soosaar@ut.ee
  • Uri, Chair of Silviculture and Forest Ecology, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Friedrich Reinhold Kreutzwaldi 1, 51014 Tartu, Estonia ORCID ID:E-mail: veiko.uri@emu.ee
article id 9972, category Research article
Jukka Malinen, Harri Kilpeläinen, Erkki Verkasalo. (2018). Validating the predicted saw log and pulpwood proportions and gross value of Scots pine and Norway spruce harvest at stand level by Most Similar Neighbour analyses and a stem quality database. Silva Fennica vol. 52 no. 4 article id 9972. https://doi.org/10.14214/sf.9972
Highlights: Non-parametric prediction together with external stem quality database provides predictions usable for pre-harvest assessment at a stand level; The prediction of Norway spruce assortment recovery and value proved to be more accurate than the predictions for Scots pine; RMSE and bias of unit prices were 3.50 € m–3 and 0.58 € m–3 for pine and 2.60 € m–3 and 0.35 € m–3 for spruce.

Detailed pre-harvest information about the volumes and properties of growing stocks is needed for increased precision in wood procurement planning for just-in-time wood deliveries by cut-to-length (CTL) harvesters. In the study, the non-parametric Most Similar Neighbour (MSN) methodology was evaluated for predicting external quality of Scots pine and Norway spruce, expressed as stem sections fulfilling the saw log dimension and quality requirements of Finnish forest industry, as they affect the recovery of timber assortments and the value of a pre-harvest stand. Effects of external tree quality were evaluated using saw log recovery and saw log reduction caused by stem defects, as well as total timber value (€) and average unit value (€ m–3) in a stand. Root mean square error (RMSE) of saw log recovery and reduction were 9.12 percentile points (pp) for Scots pine and 6.38 pp for Norway spruce stands. In the unit value considerations, the predictions compared with measurements resulted in the RMSE of 3.50 € m–3 and the bias of 0.58 € m–3 in Scots pine stands and 2.60 € m–3, and 0.35 € m–3 in Norway spruce stands, respectively. The presented MSN based approach together with the utilization of the external stem quality database included in the ARVO software could provide dimension and external quality predictions usable for pre-harvest assessment of timber stock at a stand level. This prediction methodology is usable especially in analyses where timber assortment recoveries, values and unit prices are compared when different bucking objectives are used.

  • Malinen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: jukka.malinen@uef.fi (email)
  • Kilpeläinen, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6, FI-80100 Joensuu, Finland ORCID ID:E-mail: harri.kilpelainen@luke.fi
  • Verkasalo, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6, FI-80100 Joensuu, Finland ORCID ID:E-mail: erkki.verkasalo@luke.fi
article id 9972, category Research article
Jukka Malinen, Harri Kilpeläinen, Erkki Verkasalo. (2018). Validating the predicted saw log and pulpwood proportions and gross value of Scots pine and Norway spruce harvest at stand level by Most Similar Neighbour analyses and a stem quality database. Silva Fennica vol. 52 no. 4 article id 9972. https://doi.org/10.14214/sf.9972
Highlights: Non-parametric prediction together with external stem quality database provides predictions usable for pre-harvest assessment at a stand level; The prediction of Norway spruce assortment recovery and value proved to be more accurate than the predictions for Scots pine; RMSE and bias of unit prices were 3.50 € m–3 and 0.58 € m–3 for pine and 2.60 € m–3 and 0.35 € m–3 for spruce.

Detailed pre-harvest information about the volumes and properties of growing stocks is needed for increased precision in wood procurement planning for just-in-time wood deliveries by cut-to-length (CTL) harvesters. In the study, the non-parametric Most Similar Neighbour (MSN) methodology was evaluated for predicting external quality of Scots pine and Norway spruce, expressed as stem sections fulfilling the saw log dimension and quality requirements of Finnish forest industry, as they affect the recovery of timber assortments and the value of a pre-harvest stand. Effects of external tree quality were evaluated using saw log recovery and saw log reduction caused by stem defects, as well as total timber value (€) and average unit value (€ m–3) in a stand. Root mean square error (RMSE) of saw log recovery and reduction were 9.12 percentile points (pp) for Scots pine and 6.38 pp for Norway spruce stands. In the unit value considerations, the predictions compared with measurements resulted in the RMSE of 3.50 € m–3 and the bias of 0.58 € m–3 in Scots pine stands and 2.60 € m–3, and 0.35 € m–3 in Norway spruce stands, respectively. The presented MSN based approach together with the utilization of the external stem quality database included in the ARVO software could provide dimension and external quality predictions usable for pre-harvest assessment of timber stock at a stand level. This prediction methodology is usable especially in analyses where timber assortment recoveries, values and unit prices are compared when different bucking objectives are used.

  • Malinen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: jukka.malinen@uef.fi (email)
  • Kilpeläinen, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6, FI-80100 Joensuu, Finland ORCID ID:E-mail: harri.kilpelainen@luke.fi
  • Verkasalo, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6, FI-80100 Joensuu, Finland ORCID ID:E-mail: erkki.verkasalo@luke.fi
article id 10006, category Research article
Matti Maltamo, Tomi Karjalainen, Jaakko Repola, Jari Vauhkonen. (2018). Incorporating tree- and stand-level information on crown base height into multivariate forest management inventories based on airborne laser scanning. Silva Fennica vol. 52 no. 3 article id 10006. https://doi.org/10.14214/sf.10006
Highlights: The most accurate tree-level alternative is to include crown base height (CBH) to nearest neighbour imputation; Also mixed-effects models can be applied to predict CBH using tree attributes and airborne laser scanning (ALS) metrics; CBH prediction can be included with an accuracy of 1–1.5 m to forest management inventory applications.

This study examines the alternatives to include crown base height (CBH) predictions in operational forest inventories based on airborne laser scanning (ALS) data. We studied 265 field sample plots in a strongly pine-dominated area in northeastern Finland. The CBH prediction alternatives used area-based metrics of sparse ALS data to produce this attribute by means of: 1) Tree-level imputation based on the k-nearest neighbor (k-nn) method and full field-measured tree lists including CBH observations as reference data; 2) Tree-level mixed-effects model (LME) prediction based on tree diameter (DBH) and height and ALS metrics as predictors of the models; 3) Plot-level prediction based on analyzing the computational geometry and topology of the ALS point clouds; and 4) Plot-level regression analysis using average CBH observations of the plots for model fitting. The results showed that all of the methods predicted CBH with an accuracy of 1–1.5 m. The plot-level regression model was the most accurate alternative, although alternatives producing tree-level information may be more interesting for inventories aiming at forest management planning. For this purpose, k-nn approach is promising and it only requires that field measurements of CBH is added to the tree lists used as reference data. Alternatively, the LME-approach produced good results especially in the case of dominant trees.

  • Maltamo, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: matti.maltamo@uef.fi (email)
  • Karjalainen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: tomimkarjalainen@gmail.com
  • Repola, Natural Resources Institute of Finland (Luke), Natural resources, Eteläranta 55, FI-96300 Rovaniemi, Finland ORCID ID:E-mail: jaakko.repola@luke.fi
  • Vauhkonen, Natural Resources Institute of Finland (Luke), Bioeconomy and environment, Yliopistokatu 6, 80100 Joensuu, Finland ORCID ID:E-mail: jari.vauhkonen@luke.fi
article id 9983, category Research article
Mika Rekola, Jaakko Nippala, Päivi Tynjälä, Anne Virtanen. (2018). Modelling competences and anticipating the future competence needs in the forest sector. Silva Fennica vol. 52 no. 3 article id 9983. https://doi.org/10.14214/sf.9983
Highlights: The most frequently used practices of modelling competences in Finnish forest sector organisations were superior-subordinate review discussions and quantitative surveys; Competence modelling was used for several human resources functions but surprisingly not for hiring and compensation; The experts interviewed underlined the need for generic competences in the future, especially they highlighted the importance of information processing and personal self-management skills.

This explorative study examined practices of competence modelling in the forest sector organisations and how organisations anticipate changes in competence needs in the future. Semi-structured in-depth interviews (n=10) were conducted amongst forest sector experts in Finland and data was analysed by thematic analysis. The findings showed that the practices of modelling competences were diverse, most frequently used ones being superior-subordinate review discussions and quantitative competence surveys. In addition to these formal systems, informal modelling, especially on the team level and in smaller companies was also frequent. Organisations used competence modelling for several human resources functions, such as appraisal, motivation and promotion of employees. Surprisingly hiring and compensation functions were not mentioned. Perceptions related to competence modelling were generally speaking positive. The most important challenges were the lack of further actions and sometimes the extraordinary burden to the employees. When anticipating the future, the experts interviewed mentioned several commonly recognised trends, e.g., development of information technology, fragmentation of working life and structural changes in labour markets. All these require more generic competences related to information processing and personal self-management, especially respondents highlighted the importance of self-awareness skills. It is concluded that several useful practices for competence modelling already exist and that present study provides a basis for further quantitative further study.

  • Rekola, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID ID:E-mail: mika.rekola@helsinki.fi (email)
  • Nippala, Suomen Partiolaiset – Finlands Scouter ry, Töölönkatu 55, FI-00250 Helsinki, Finland ORCID ID:E-mail: jaakko.nippala@partio.fi
  • Tynjälä, Finnish Institute for Educational Research, P.O. Box 35, FI-40014 University of Jyväskylä, Finland ORCID ID:E-mail: paivi.a.tynjala@jyu.fi
  • Virtanen, Finnish Institute for Educational Research, P.O. Box 35, FI-40014 University of Jyväskylä, Finland ORCID ID:E-mail: anne.virtanen@jyu.fi
article id 9947, category Research article
Eric R. Labelle, Linus Huß. (2018). Creation of value through a harvester on-board bucking optimization system operated in a spruce stand. Silva Fennica vol. 52 no. 3 article id 9947. https://doi.org/10.14214/sf.9947
Highlights: Use of harvester on-board computer bucking optimization remains highly under-utilized in German forestry; Revenue per tree and harvesting productivity were both statistically higher with automatic bucking as compared to quality bucking during a thinning operation in a spruce dominated stand.

Tree bucking, defined as the process in which a stem is segmented into shorter logs of varying lengths, has a significant effect on the value adding potential of a forest enterprise. Because of its importance in terms of correct product and length combinations, improper bucking can lead to financial losses. In this study, two treatments (OFF: quality bucking performed by the operator while using hot keys and ON: automatic bucking using the optimized suggestions from the harvester on-board computer; OBC) were tested in a Norway spruce (Picea abies [L.] Karst.) dominated stand located in Germany. Both treatments had the aim to maximize the value of a stem. The research took place in an 80-year old spruce and beech stand under a regenerative cutting. Fully-mechanized harvesting was performed with an 8-wheel Ponsse Bear single-grip harvester equipped with a H8 harvesting head. Results indicated that the product recovery of the two treatments differed by 4% in undamaged trees (no broken tree-tops or stems) to the benefit of manual bucking. However, the revenue of trees subjected to optimized bucking were up to 4% higher (in average 3%) than those of the manual bucking once expressed on a per cubic meter basis. Moreover, the harvesting productivity of the ON treatment was at the maximum 17% higher compared to the OFF treatment. Based on the results from this case study, the use of an optimization software in Norway spruce dominated stands with the aim to maximize the value of single stems showed promising results.

  • Labelle, Assistant Professorship of Forest Operations, Department of Ecology and Ecosystem Management, Technical University of Munich, Hans-Carl-von-Carlowitz-Platz 2, D-85354 Freising, Germany ORCID ID:E-mail: eric.labelle@tum.de (email)
  • Huß, Assistant Professorship of Forest Operations, Department of Ecology and Ecosystem Management, Technical University of Munich, Hans-Carl-von-Carlowitz-Platz 2, D-85354 Freising, Germany ORCID ID:E-mail: linus.huss@gmx.de
article id 9940, category Research article
Esteban Ceriani-Nakamurakare, Sergio Ramos, Carolina A. Robles, María V. Novas, María F. D´Jonsiles, Paola Gonzalez-Audino, Cecilia Carmarán. (2018). Metagenomic approach of associated fungi with Megaplatypus mutatus (Coleoptera: Platypodinae). Silva Fennica vol. 52 no. 3 article id 9940. https://doi.org/10.14214/sf.9940
Highlights: There were no significant effects of host plant and location on fungal richness; Two fungal species, belonging to Fusarium and Candida genera, were present in all the studied associations; Results suggest that host plant identity would not be crucial to determine the composition of fungal communities associated to Megaplatypus mutatus.

Megaplatypus mutatus is a major forest pest in Argentina and an emerging pest in Europe. In this study the multitrophic interactions between M. mutatus and associated fungi were assessed with a metagenomics approach (454-pyrosequencing). A total of 270 collection points from insect galleries from three locations in Argentina were pooled for pyrosequencing analyses. Two hosts, Populus deltoides and Casuarina cunninghamiana, were independently evaluated to characterize the fungal communities associated to M. mutatus; compare the culture-independent approach with previous culturing studies, in terms of data recovery related to the fungal community composition, and test the specificity of the fungal communities amongst locations and hosts. A Generalized Linear Mixed Model was performed to compare the fungal richness in each dataset, which showed no significant differences between taxa richness amongst locations. Principal Coordinates Analyses showed a separation between fungal communities within the same host, suggesting that host identity would not be crucial to determine the specificity in fungal communities. Candida insectalens and one Fusarium species, present in all hosts and locations, achieved 37.6% of the total relative frequency per taxa. These results complement the data from culturing methods previously reported, thus improving the accuracy and understanding of the fungal assemblages associated to M. mutatus.

  • Ceriani-Nakamurakare, Universidad de Buenos Aires. Facultad de Ciencias Exactas y Naturales. Depto. Biodiversidad y Biología Experimental. Buenos Aires, Argentina; CONICET- Universidad de Buenos Aires. Instituto de Micología y Botánica (INMIBO). Buenos Aires, (C1428EHA) Argentina ORCID ID:E-mail: cerianinaka@gmail.com
  • Ramos, Instituto Nacional de Tecnología Agropecuaria. Estación Experimental Agropecuaria Concordia. Entre Ríos, (E3200) Argentina ORCID ID:E-mail: ramos.sergio@inta.gob.ar
  • Robles, Universidad de Buenos Aires. Facultad de Ciencias Exactas y Naturales. Depto. Biodiversidad y Biología Experimental. Buenos Aires, Argentina; CONICET- Universidad de Buenos Aires. Instituto de Micología y Botánica (INMIBO). Buenos Aires, (C1428EHA) Argentina ORCID ID:E-mail: carorobles@bg.fcen.uba.ar
  • Novas, Universidad de Buenos Aires. Facultad de Ciencias Exactas y Naturales. Depto. Biodiversidad y Biología Experimental. Buenos Aires, Argentina; CONICET- Universidad de Buenos Aires. Instituto de Micología y Botánica (INMIBO). Buenos Aires, (C1428EHA) Argentina ORCID ID:E-mail: vicnovas@bg.fcen.uba.ar
  • D´Jonsiles, Universidad de Buenos Aires. Facultad de Ciencias Exactas y Naturales. Depto. Biodiversidad y Biología Experimental. Buenos Aires, Argentina; CONICET- Universidad de Buenos Aires. Instituto de Micología y Botánica (INMIBO). Buenos Aires, (C1428EHA) Argentina ORCID ID:E-mail: lalijonsi@gmail.com
  • Gonzalez-Audino, Centro de Investigaciones de Plagas e Insecticidas (CITEFA-CONICET). Buenos Aires, (B1603ALO) Argentina ORCID ID:E-mail: pgonzalezaudino@citedef.gob.ar
  • Carmarán, Centro de Investigaciones de Plagas e Insecticidas (CITEFA-CONICET). Buenos Aires, (B1603ALO) Argentina ORCID ID:E-mail: carmaran@bg.fcen.uba.ar (email)
article id 9927, category Research article
Eva Dařenová, Richard A. Crabbe, Robert Knott, Barbora Uherková, Jan Kadavý. (2018). Effect of coppicing, thinning and throughfall reduction on soil water content and soil CO2 efflux in a sessile oak forest. Silva Fennica vol. 52 no. 2 article id 9927. https://doi.org/10.14214/sf.9927
Highlights: The coppice stand reached similar soil CO2 efflux rates eight years after harvest as the original stand; Thinning increased SWC but did not affect soil CO2 efflux; 30% throughfall reduction decreased soil CO2 efflux by 50.7%.

In this study we determined the effect of transformation of a mature sessile oak forest stand into a coppiced forest, and of thinning and throughfall reduction in a coppice stand on soil water content (SWC) and soil CO2 efflux. The precipitation reduction was induced by installing parallel drainage channels in both unthinned and thinned coppice stands. The driving factor for temporal dynamics of soil CO2 efflux in all plots was soil temperature. The other factor was soil water content but only up to about 15%. Above this threshold, there was no more effect on CO2 efflux. We found no clear difference in SWC or soil CO2 efflux between the mature and coppiced stand eight years after harvesting. On the other hand, thinning of the coppice stand resulted in increase in SWC up to 22% in proportion, which we assume to be a result of increased gap fraction of the canopy. However, no effect on soil CO2 efflux was observed two years after the thinning. Installation of the drainage channels in two plots covering 30% of the ground area resulted in decrease in SWC up to a proportional 30.5% and thus contributed up to 50.7% reduction in soil CO2 efflux.

  • Dařenová, Global Change Research Institute CAS, v.v.i., Belidla 4a, 603 00 Brno, Czech Republic ORCID ID:E-mail: darenova.e@czechglobe.cz (email)
  • Crabbe, Global Change Research Institute CAS, v.v.i., Belidla 4a, 603 00 Brno, Czech Republic ORCID ID:E-mail: crabbe.r@czechglobe.cz
  • Knott, Mendel University in Brno, Zemedelska 3, 613 00 Brno, Czech Republic ORCID ID:E-mail: robert.knott@mendelu.cz
  • Uherková, Mendel University in Brno, Zemedelska 3, 613 00 Brno, Czech Republic ORCID ID:E-mail: xfedorov@node.mendelu.cz
  • Kadavý, Mendel University in Brno, Zemedelska 3, 613 00 Brno, Czech Republic ORCID ID:E-mail: jan.kadavy@mendelu.cz
article id 9914, category Research article
Jun Tanabe, Futoshi Ishiguri, Akira Tamura, Yuya Takashima, Jyunichi Ohshima, Kazuya Iizuka, Shinso Yokota. (2018). Within-tree radial and among-family variations in wood density, microfibril angle, and mechanical properties in Picea glehnii. Silva Fennica vol. 52 no. 2 article id 9914. https://doi.org/10.14214/sf.9914
Highlights: The modulus of elasticity was affected by both microfibril angle and wood density, whereas the modulus of rupture was mainly affected by wood density in Picea glehnii; A larger degree of among-family variation in wood properties was detected in juvenile wood than in mature wood, indicating that genetic improvements in the mechanical properties may be more effective for juvenile wood.

Genetic improvements in the mechanical properties of wood are important in forestry species used for lumber, such as Picea. The within-tree radial and among-family variations for the modulus of elasticity (MOE), modulus of rupture (MOR), and their related traits [i.e., microfibril angle (MFA) of the S2 layer in latewood tracheid and air-dry density (AD)] were evaluated in nine open-pollinated families of Picea glehnii (F. Schmidt) Mast. The radial variation in MOR was mainly affected by AD, whereas MOE was affected by MFA and AD. Higher F-values obtained by analysis of variance and coefficient of variation were observed for all properties at the 6th–15th annual ring, except for AD at the 6th–10th annual ring. This result suggests that the contribution of genetic effect is larger in these highly variable regions. In addition, positive correlation coefficients were obtained between wood properties at the 6th–15th annual ring and mean values of these properties. Therefore, genetic improvements for MOE, MOR, and their related traits in P. glehnii is likely to be more effective in juvenile wood, specifically at the 6th–15th annual ring from the pith.

  • Tanabe, Faculty of Education, Chiba University, Chiba, 263-8522, Japan ORCID ID:E-mail: tanabe_j@chiba-u.jp (email)
  • Ishiguri, Faculty of Agriculture, Utsunomiya University, Utsunomiya 321-8505, Japan ORCID ID:E-mail: ishiguri@cc.utsunomiya-u.ac.jp
  • Tamura, Forest Tree Breeding Center, Forestry and Forest Products Research Institute, Hitachi, 319-1301, Japan ORCID ID:E-mail: akirat@affrc.go.jp
  • Takashima, Forest Tree Breeding Center, Forestry and Forest Products Research Institute, Hitachi, 319-1301, Japan ORCID ID:E-mail: ytakashima@ffpri.affrc.go.jp
  • Ohshima, Faculty of Agriculture, Utsunomiya University, Utsunomiya 321-8505, Japan ORCID ID:E-mail: joshima@cc.utsunomiya-u.ac.jp
  • Iizuka, Faculty of Agriculture, Utsunomiya University, Utsunomiya 321-8505, Japan ORCID ID:E-mail: kiizuka@cc.utsunomiya-u.ac.jp
  • Yokota, Faculty of Agriculture, Utsunomiya University, Utsunomiya 321-8505, Japan ORCID ID:E-mail: yokotas@cc.utsunomiya-u.ac.jp
article id 7816, category Research article
Pentti Niemistö, Harri Kilpeläinen, Eero Poutiainen. (2018). Effect of first thinning type and age on growth, stem quality and financial performance of a Scots pine stand in Finland. Silva Fennica vol. 52 no. 2 article id 7816. https://doi.org/10.14214/sf.7816
Highlights: Early commercial first thinning at the top height of 11 m with tree selection targeting high quality of remaining stems was more profitable in a Scots pine stand than thinning from below at similar time or delayed thinning 10 years later; After early first thinning only one intermediate thinning was needed, but a late thinning at age of 60 years was feasible in connection with increased amount of high quality butt logs after artificial pruning; The optimal rotation period for a Scots pine stand using a lower interest rate of 1% was 80–85 years depending on the thinning pattern. With a higher rate of 4%, the optimal rotation took only 60 years.

The objective of the study was to ascertain the effects of tree selection (thinning from below, from above and according to stem quality) and timing of first commercial thinning (early and delayed) on the growth, yield and quality of trees in a Scots pine (Pinus sylvestris L.) stand. A long-term field experiment (25 years) was measured in 5-year periods and the further development was simulated with growth and yield models to final cuttings using alternative rotation periods of 55–85 years. The measurements included also the exact location and type of technical defects detected on all trees in the experimental plots. The measured volume increment per unit area during the study period, 25 years after the early thinning stage was the lowest in the plots thinned from below, and the highest in the plots thinned from above or in the delayed thinning plots. However, the largest volume of saw logs during the whole rotation of 80 years was yielded after early first thinning according to the quality. The largest volume of very high-quality butt logs was produced by pruning connected with early thinning from above, and a smaller volume after early thinning according to stem quality but no after thinning from below or delayed first thinning. Without pruning an early quality thinning with one intermediate thinning was the most profitable thinning treatment in the Scots pine stand regardless the rotation length or the interest rate used. By interest rates of 1% and 2%, the optimal rotations were 80–85 years and 70 years respectively. A late thinning at the age of 60 year with long rotation was profitable only for the pruned pine stands with a low interest rate.

  • Niemistö, Natural Resources Institute Finland (Luke), Natural resources, Kampusranta 9 C, FI-60320 Seinäjoki, Finland ORCID ID:E-mail: pentti.niemisto@luke.fi (email)
  • Kilpeläinen, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6, FI-80100 Joensuu, Finland ORCID ID:E-mail: harri.kilpelainen@luke.fi
  • Poutiainen, Oulu ORCID ID:E-mail: eero1.poutiainen@dnainternet.net
article id 7738, category Research article
Samuel Egbäck, Bo Karlsson, Karl-Anders Högberg, Kenneth Nyström, Mateusz Liziniewicz, Urban Nilsson. (2018). Effects of phenotypic selection on height-diameter ratio of Norway spruce and Scots pine in Sweden. Silva Fennica vol. 52 no. 2 article id 7738. https://doi.org/10.14214/sf.7738
Highlights: Swedish plus-tree selection promoted less slender Norway spruce trees and more slender Scots pine trees compared to neighboring trees; Similar results were also found for progeny trials which indicated that genetics played a prominent role in phenotypic appearance.

Genetically improved Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) are extensively used in operational Swedish forestry plantations. However, relatively little is known about the stem slenderness (height-diameter ratio) of genetically improved material. Thus, in this study we investigated effects of plus-tree selection on stem slenderness of Norway spruce and Scots pine in Sweden by evaluating both the plus-tree selection and a large number of progeny trials. Species-specific models for predicting the height-diameter ratio were estimated using regression and mixed model approach. Our results show that phenotypic plus-tree selection promoted less slender Norway spruce trees and more slender Scots pine trees compared to neighboring trees. Similar results were also found for the progeny trials which indicated that genetics played a prominent role in the phenotypic appearance. Compared to the progeny of neighboring trees, Norway spruce plus-tree progenies had a 5.3% lower height-diameter ratio, while Scots pine plus-tree progenies had a 1.5% greater height-diameter ratio. The narrow sense heritability for height-diameter ratio was 0.19 for Norway spruce and 0.11 for Scots pine, indicating that it is possible to modify the height-diameter ratio by breeding. Correlation coefficients between breeding values for height-diameter ratio and diameter were negative for Scots pine (–0.71) and Norway spruce (–0.85), indicating that selection for diameter only would result in less slender stems of both species. Similar correlations were also found between breeding values for height-diameter ratio and height of Scots pine (–0.34) and Norway spruce (–0.74).

  • Egbäck, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, 230 53 Alnarp, Sweden ORCID ID:E-mail: samuel.egback@slu.se (email)
  • Karlsson, Skogforsk, Ekebo, 268 90 Svalöv, Sweden ORCID ID:E-mail: bo.karlsson@skogforsk.se
  • Högberg, Skogforsk, Ekebo, 268 90 Svalöv, Sweden ORCID ID:E-mail: karl-anders.hogberg@skogforsk.se
  • Nyström, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd, 901 83 Umeå, Sweden ORCID ID:E-mail: kenneth.nystrom@slu.se
  • Liziniewicz, Skogforsk, Ekebo, 268 90 Svalöv, Sweden ORCID ID:E-mail: Mateusz.Liziniewicz@skogforsk.se
  • Nilsson, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, 230 53 Alnarp, Sweden ORCID ID:E-mail: urban.nilsson@slu.se
article id 7791, category Research article
Tadeusz B. Splawinski, Sylvie Gauthier, Nicole J. Fenton, Daniel Houle, Yves Bergeron. (2018). The colonization of young fire initiated stands by the crustose lichen Trapeliopsis granulosa and its potential effect on conifer establishment and stand succession. Silva Fennica vol. 52 no. 1 article id 7791. https://doi.org/10.14214/sf.7791
Highlights: T. granulosa is a poor seedbed for jack pine establishment; The presence of extensive T. granulosa cover can limit ongoing tree recruitment, thereby maintaining open lichen woodland; Dry open conditions favor the establishment of T. granulosa; Stands with significant T. granulosa cover may be good candidates for afforestation initiatives due to lower evaporation potential and decreased water stress.

The resilience of closed-crown coniferous stands within the boreal forest of North America is highly dependent on successful re-establishment of tree species following fire. A shift from closed-crown forest to open lichen woodland is possible following poor natural regeneration during the initial establishment phase, followed by the development of extensive lichen cover, which may hinder ongoing recruitment. We examined the development of the crustose lichen Trapeliopsis granulosa (Hoffm.) 18 to 21 years following fire within six sites in the boreal forest of northwestern Quebec, and explored its potential to affect ongoing recruitment during early successional stages of stand development. Germination and survivorship trials were conducted within the laboratory to determine the establishment rate of Pinus banksiana Lamb. (jack pine) on T. granulosa, mineral soil, and burnt duff under two separate watering frequencies (observed and drought). Survival and establishment rates of jack pine were highest on burnt duff, and poor on both T. granulosa and mineral soil. Under the drought treatment, no seedlings survived on any substrates. In the field, T. granulosa cover had a positive relationship with mineral soil cover, and negative relationships with duff cover, ericaceous shrub cover, organic layer depth, other lichen cover, and Sphagnum moss cover. No discernable relationship was found between T. granulosa and tree density, rock cover, dead wood cover or other moss cover. The development of extensive T. granulosa cover in fire-initiated stands can impede ongoing recruitment of conifer species due to its poor seedbed quality, thereby maintaining open forests.

  • Splawinski, Institut de recherche sur les forêts, Université du Québec en Abitibi-Témiscamingue, 445, boul. de l’Université, Rouyn-Noranda, QC, J9X 5E4, Canada ORCID ID:E-mail: tsplawinski@gmail.com (email)
  • Gauthier, Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, 1055 rue du PEPS, P.O. Box 10380, Stn Sainte Foy, QC, G1V 4C7, Canada ORCID ID:E-mail: sylvie.gauthier@rncan-nrcan.gc.ca
  • Fenton, Institut de recherche sur les forêts (IRF), Université du Québec en Abitibi-Témiscamingue, 445 boul. de l’Université, Rouyn-Noranda, QC, J9X 5E4, Canada ORCID ID:E-mail: nicole.fenton@uqat.ca
  • Houle, Ministère des Forêts, de la Faune et des Parcs, Direction de la recherché forestière, Québec, QC, G1P 3W8, Canada; Ouranos Climate Change Consortium, Montréal, QC, H3A 1B9, Canada ORCID ID:E-mail: daniel.houle@mffp.gouv.qc.ca
  • Bergeron, Centre d’étude sur la forêt and Chaire industrielle en aménagement forestier durable, Université du Québec à Montréal, CP 8888 Succursale A, Montréal, QC, H3C 3P8, Canada ORCID ID:E-mail: bergeron.yves@uqam.ca
article id 7783, category Research article
Markku T. Lehtinen, Pertti Pulkkinen. (2017). Effects of Scots pine paternal genotypes of two contiguous seed orchards on the budset and frost hardening of first-year progeny. Silva Fennica vol. 51 no. 5 article id 7783. https://doi.org/10.14214/sf.7783
Highlights: This environmentally controlled study on Scots pine demonstrated the effect of the paternal genotype on the budset and frost hardening of the progeny; With the applied study design, no significant indication of an environmental influence on the effect of the Scots pine paternal genotype was obtained.

In Scots pine (Pinus sylvestris L.), it has been shown that the parental conditions have a role in the phenological variation among first-year seedlings. For this reason, it is argued that they should be comprehensively controlled before estimating the parental genotype effects. This controlled-cross study examined the effects of a set of fathers of Scots pines on the timing of budset and autumn frost hardening of first-year seedlings. The paternal genotypes had either a northern or southern provenance, but had spent a period of over 25 years as grafts in a shared climatic environment in two closely located southern orchards. Pollen applied in the crosses was collected from these orchards in one year and all the maternal genotypes were pollinated in only one seed orchard. The results of freeze tests and budset observations of the consequent progeny were analysed and additionally compared with results obtained using seedlings from seed lots of natural forests in order to estimate the ability of northern paternal genotypes to maintain a northern effect under southern conditions. This environmentally controlled study demonstrated a significant effect of the paternal genotype on the budset and autumn frost hardening of first-year seedling of Scots pine. With the applied study design, no significant indication of an environmental influence on the effect of the paternal genotype was obtained. The accuracy of the observations is discussed. It is concluded that the results suggest a minor role of mutability in the effects of Scots pine paternal genotypes.

  • Lehtinen, University of Helsinki, Department of Agricultural Sciences, Latokartanonkaari 5 and 7, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID ID:E-mail: markku.t.lehtinen@helsinki.fi (email)
  • Pulkkinen, Natural Resources Institute Finland (Luke), Green Technology, Haapastensyrjäntie 34, FI-12600 Läyliäinen, Finland ORCID ID:E-mail: pertti.pulkkinen@luke.fi
article id 7721, category Research article
Sakari Tuominen, Andras Balazs, Eija Honkavaara, Ilkka Pölönen, Heikki Saari, Teemu Hakala, Niko Viljanen. (2017). Hyperspectral UAV-imagery and photogrammetric canopy height model in estimating forest stand variables. Silva Fennica vol. 51 no. 5 article id 7721. https://doi.org/10.14214/sf.7721
Highlights: Hyperspectral imagery and photogrammetric 3D point cloud based on RGB imagery were acquired under weather conditions changing from cloudy to sunny; Calibration of hyperspectral imagery was required for compensating the effect of varying weather conditions; The combination of hyperspectral imagery and photogrammetric point cloud data resulted in accurate forest estimates, especially for volumes per tree species.

Remote sensing using unmanned aerial vehicle (UAV) -borne sensors is currently a highly interesting approach for the estimation of forest characteristics. 3D remote sensing data from airborne laser scanning or digital stereo photogrammetry enable highly accurate estimation of forest variables related to the volume of growing stock and dimension of the trees, whereas recognition of tree species dominance and proportion of different tree species has been a major complication in remote sensing-based estimation of stand variables. In this study the use of UAV-borne hyperspectral imagery was examined in combination with a high-resolution photogrammetric canopy height model in estimating forest variables of 298 sample plots. Data were captured from eleven separate test sites under weather conditions varying from sunny to cloudy and partially cloudy. Both calibrated hyperspectral reflectance images and uncalibrated imagery were tested in combination with a canopy height model based on RGB camera imagery using the k-nearest neighbour estimation method. The results indicate that this data combination allows accurate estimation of stand volume, mean height and diameter: the best relative RMSE values for those variables were 22.7%, 7.4% and 14.7%, respectively. In estimating volume and dimension-related variables, the use of a calibrated image mosaic did not bring significant improvement in the results. In estimating the volumes of individual tree species, the use of calibrated hyperspectral imagery generally brought marked improvement in the estimation accuracy; the best relative RMSE values for the volumes for pine, spruce, larch and broadleaved trees were 34.5%, 57.2%, 45.7% and 42.0%, respectively.

  • Tuominen, Natural Resources Institute Finland (Luke), Economics and society, P.O. Box 2, FI-00791 Helsinki, Finland ORCID ID: http://orcid.org/0000-0001-5429-3433 E-mail: sakari.tuominen@luke.fi (email)
  • Balazs, Natural Resources Institute Finland (Luke), Economics and society, P.O. Box 2, FI-00791 Helsinki, Finland ORCID ID:E-mail: andras.balazs@luke.fi
  • Honkavaara, Finnish Geospatial Research Institute, National Land Survey of Finland, Geodeetinrinne 2, FI-02430 Masala, Finland ORCID ID:E-mail: eija.honkavaara@nls.fi
  • Pölönen, University of Jyväskylä, Faculty of Information Technology, P.O. Box 35, FI-40014 Jyväskylä, Finland ORCID ID:E-mail: ilkka.polonen@jyu.fi
  • Saari, VTT Microelectronics, P.O. Box 1000, FI-02044 VTT, Finland ORCID ID:E-mail: heikki.saari@vtt.fi
  • Hakala, Finnish Geospatial Research Institute, National Land Survey of Finland, Geodeetinrinne 2, FI-02430 Masala, Finland ORCID ID:E-mail: teemu.hakala@nls.fi
  • Viljanen, Finnish Geospatial Research Institute, National Land Survey of Finland, Geodeetinrinne 2, FI-02430 Masala, Finland ORCID ID:E-mail: niko.viljanen@nls.fi
article id 7693, category Research article
Chunyu Zhu, Jiaojun Zhu, Xiao Zheng, Deliang Lu, Xiufen Li. (2017). Comparison of gap formation and distribution pattern induced by wind/snowstorm and flood in a temperate secondary forest ecosystem, Northeast China. Silva Fennica vol. 51 no. 5 article id 7693. https://doi.org/10.14214/sf.7693
Highlights: The canopy gaps induced by wind/snowstorm were aggregated in steep slope and high altitude areas, while the gaps formed by flood were gathered in steep slope and low altitude areas; The wind/snowstorm mainly driven the formation of medium gaps, while the flood mainly promoted the percentage of small gaps and vacant lands.

Canopy gap is the driving force of forest succession. Due to the uncontrollability, however, the influences of natural disturbances on gap formation and gap distribution pattern have been rarely understood in temperate secondary forest ecosystems. We monitored the gap formation and gap distribution pattern using high-resolution remote sensing images before and after two disturbances (wind/snowstorm in 2003 and flood in 2013). The results showed that after wind/snowstorm, the gap nearest neighbor index (GNNI) decreased, the vacant land area did not obviously change while the gap fraction and gaps density (especially medium size) increased. After the flood, GNNI decreased, the number of small gaps increased but larger gaps were in many cases extended to vacant land areas leading to a smaller total number of medium and large gaps but considerable increase in vacant land area. We also found that the gap densities increased with slope and altitude for wind/snowstorm-formed gaps, but they increased with increasing slope and decreasing altitude for flood-formed gaps. These results indicated that gaps were aggregated in steep slope and high altitude areas after wind/snowstorm, but in steep slope and low altitude areas after the flood. Medium gaps were mainly created by the wind/snowstorm due to the individual-level death of dominant tree with the continuous fall of surrounding trees. While, vacant lands were obviously created during the flood because of integral sweeping. Besides, smaller trees were easily damaged by runoff of flood, which induced small gaps. In summary, forest managers may pay more attention to use gaps to accelerate forest succession after wind/snowstorms and to restore vegetation in vacant lands after floods.

  • Zhu, ORCID ID:E-mail: Chunyuzhu123@126.com
  • Zhu, ORCID ID:E-mail: jiaojunzhu@iae.ac.cn (email)
  • Zheng, ORCID ID:E-mail: xiaozheng@iae.ac.cn
  • Lu, ORCID ID:E-mail: delianglu14@hotmail.com
  • Li, ORCID ID:E-mail: delianglu14@hotmail.com
article id 2018, category Research article
Sima Mohtashami, Lars Eliasson, Gunnar Jansson, Johan Sonesson. (2017). Influence of soil type, cartographic depth-to-water, road reinforcement and traffic intensity on rut formation in logging operations: a survey study in Sweden. Silva Fennica vol. 51 no. 5 article id 2018. https://doi.org/10.14214/sf.2018
Highlights: Soil type and traffic intensity had significant effects on rut formation; Further studies are required to identify all factors affecting rut formation, especially on soils with medium bearing capacity; The cartographic depth-to-water index (DTW) alone did not predict rut formation, but used in combination with other information, e.g. soil type, could be an interesting tool for delineating soil areas that are potentially vulnerable to rut formation in logging operations.

Rut formation caused by logging operations has been recognised as a challenge for Swedish forestry. Frequent traffic with heavy machines on extraction roads, together with a warmer climate, is one of the factors that increases the risk of rut formation in forests. One possible way to control this impact of logging operations is to design and apply decision support tools that enable operators to take sensitive areas into account when planning extraction roads. In this study, 16 different logging sites in south-eastern Sweden were surveyed after clear-cut. Information was collected about extraction roads (i.e. traffic intensity, whether the roads had been reinforced with slash) and ruts. Digital maps such as cartographic depth-to-water (DTW) index and soil type were also examined for any connection to rut positions. Soil type and traffic intensity were found to be significant factors in rut formation, while DTW and slash reinforcement were not. However, the DTW map combined with other information, such as soil type, could contribute to decision support tools that improve planning of extraction roads.

  • Mohtashami, The forestry research institute of Sweden, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail: sima.mohtashami@skogforsk.se (email)
  • Eliasson, The forestry research institute of Sweden, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID: http://orcid.org/0000-0002-2038-9864 E-mail: lars.eliasson@skogforsk.se
  • Jansson, The forestry research institute of Sweden, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID: http://orcid.org/0000-0002-3018-9161 E-mail: gunnar.jansson@skogforsk.se
  • Sonesson, The forestry research institute of Sweden, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID: http://orcid.org/0000-0002-2018-7496 E-mail: johan.sonesson@skogforsk.se
article id 1740, category Research article
Ram P. Sharma, Zdeněk Vacek, Stanislav Vacek. (2017). Modelling tree crown-to-bole diameter ratio for Norway spruce and European beech. Silva Fennica vol. 51 no. 5 article id 1740. https://doi.org/10.14214/sf.1740
Highlights: Modelled crown-to-bole diameter ratio (CDBDR) using tree and stand-level predictors, and sample plot random effects; Spatially explicit mixed-effects model described the largest part of CDBDR variation with no significant trend in the residuals; The CDBDR increased with increasing stand development stage and site quality, but decreased with decreasing proportion of the species of interest, and increasing competition.

Crown dimensions are correlated to growth of other parts of a tree and often used as predictors in growth models. The crown-to-bole diameter ratio (CDBDR), which is a ratio of maximum crown width to diameter at breast height (DBH), was modelled using data from permanent sample plots located on Norway spruce (Picea abies (L.) Karst.) and European beech (Fagus sylvatica L.) stands in different parts of the Czech Republic. Among various tree and stand-level measures evaluated, DBH, height to crown base (HCB), dominant height (HDOM), basal area of trees larger in diameter than a subject tree (BAL), basal area proportion of the species of interest (BAPOR), and Hegyi’s competition index (CI) were found to be significant predictors in the CDBDR model. Random effects were included using the mixed-effects modelling to describe sample plot-level variation. For each species, the mixed-effects model described a larger part of the variation of the CDBDR than nonlinear ordinary least squares model with no trend in the residuals. The spatially explicit mixed-effects model showed more attractive fit statistics [conditional R2 ≈ 0.73 (spruce), 0.78 (beech)] than its spatially inexplicit counterpart [conditional R2 ≈ 0.71 (spruce), 0.76 (beech)]. The model showed that CDBDR increased with increasing HDOM – a measure that combines the stand development stage and site quality – but decreased with increasing HCB and competition (increasing BAL and CI), and decreasing proportions of the species of interest (increasing BAPOR). For both species, the spatially explicit mixed-effects model should be a preferred choice for a precise prediction of the CDBDR. The CDBDR model will have various management implications such as determination of spacing, stand basal area, stocking, and planning of appropriate species mixture.

  • Sharma, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Kamýcká 129, 16521, Praha 6 – Suchdol, Czech Republic ORCID ID:E-mail: sharmar@fld.czu.cz (email)
  • Vacek, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Kamýcká 129, 16521, Praha 6 – Suchdol, Czech Republic ORCID ID:E-mail: vacekz@fld.czu.cz
  • Vacek, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Kamýcká 129, 16521, Praha 6 – Suchdol, Czech Republic ORCID ID:E-mail: vacekstanislav@fld.czu.cz
article id 1740, category Research article
Ram P. Sharma, Zdeněk Vacek, Stanislav Vacek. (2017). Modelling tree crown-to-bole diameter ratio for Norway spruce and European beech. Silva Fennica vol. 51 no. 5 article id 1740. https://doi.org/10.14214/sf.1740
Highlights: Modelled crown-to-bole diameter ratio (CDBDR) using tree and stand-level predictors, and sample plot random effects; Spatially explicit mixed-effects model described the largest part of CDBDR variation with no significant trend in the residuals; The CDBDR increased with increasing stand development stage and site quality, but decreased with decreasing proportion of the species of interest, and increasing competition.

Crown dimensions are correlated to growth of other parts of a tree and often used as predictors in growth models. The crown-to-bole diameter ratio (CDBDR), which is a ratio of maximum crown width to diameter at breast height (DBH), was modelled using data from permanent sample plots located on Norway spruce (Picea abies (L.) Karst.) and European beech (Fagus sylvatica L.) stands in different parts of the Czech Republic. Among various tree and stand-level measures evaluated, DBH, height to crown base (HCB), dominant height (HDOM), basal area of trees larger in diameter than a subject tree (BAL), basal area proportion of the species of interest (BAPOR), and Hegyi’s competition index (CI) were found to be significant predictors in the CDBDR model. Random effects were included using the mixed-effects modelling to describe sample plot-level variation. For each species, the mixed-effects model described a larger part of the variation of the CDBDR than nonlinear ordinary least squares model with no trend in the residuals. The spatially explicit mixed-effects model showed more attractive fit statistics [conditional R2 ≈ 0.73 (spruce), 0.78 (beech)] than its spatially inexplicit counterpart [conditional R2 ≈ 0.71 (spruce), 0.76 (beech)]. The model showed that CDBDR increased with increasing HDOM – a measure that combines the stand development stage and site quality – but decreased with increasing HCB and competition (increasing BAL and CI), and decreasing proportions of the species of interest (increasing BAPOR). For both species, the spatially explicit mixed-effects model should be a preferred choice for a precise prediction of the CDBDR. The CDBDR model will have various management implications such as determination of spacing, stand basal area, stocking, and planning of appropriate species mixture.

  • Sharma, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Kamýcká 129, 16521, Praha 6 – Suchdol, Czech Republic ORCID ID:E-mail: sharmar@fld.czu.cz (email)
  • Vacek, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Kamýcká 129, 16521, Praha 6 – Suchdol, Czech Republic ORCID ID:E-mail: vacekz@fld.czu.cz
  • Vacek, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Kamýcká 129, 16521, Praha 6 – Suchdol, Czech Republic ORCID ID:E-mail: vacekstanislav@fld.czu.cz
article id 1740, category Research article
Ram P. Sharma, Zdeněk Vacek, Stanislav Vacek. (2017). Modelling tree crown-to-bole diameter ratio for Norway spruce and European beech. Silva Fennica vol. 51 no. 5 article id 1740. https://doi.org/10.14214/sf.1740
Highlights: Modelled crown-to-bole diameter ratio (CDBDR) using tree and stand-level predictors, and sample plot random effects; Spatially explicit mixed-effects model described the largest part of CDBDR variation with no significant trend in the residuals; The CDBDR increased with increasing stand development stage and site quality, but decreased with decreasing proportion of the species of interest, and increasing competition.

Crown dimensions are correlated to growth of other parts of a tree and often used as predictors in growth models. The crown-to-bole diameter ratio (CDBDR), which is a ratio of maximum crown width to diameter at breast height (DBH), was modelled using data from permanent sample plots located on Norway spruce (Picea abies (L.) Karst.) and European beech (Fagus sylvatica L.) stands in different parts of the Czech Republic. Among various tree and stand-level measures evaluated, DBH, height to crown base (HCB), dominant height (HDOM), basal area of trees larger in diameter than a subject tree (BAL), basal area proportion of the species of interest (BAPOR), and Hegyi’s competition index (CI) were found to be significant predictors in the CDBDR model. Random effects were included using the mixed-effects modelling to describe sample plot-level variation. For each species, the mixed-effects model described a larger part of the variation of the CDBDR than nonlinear ordinary least squares model with no trend in the residuals. The spatially explicit mixed-effects model showed more attractive fit statistics [conditional R2 ≈ 0.73 (spruce), 0.78 (beech)] than its spatially inexplicit counterpart [conditional R2 ≈ 0.71 (spruce), 0.76 (beech)]. The model showed that CDBDR increased with increasing HDOM – a measure that combines the stand development stage and site quality – but decreased with increasing HCB and competition (increasing BAL and CI), and decreasing proportions of the species of interest (increasing BAPOR). For both species, the spatially explicit mixed-effects model should be a preferred choice for a precise prediction of the CDBDR. The CDBDR model will have various management implications such as determination of spacing, stand basal area, stocking, and planning of appropriate species mixture.

  • Sharma, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Kamýcká 129, 16521, Praha 6 – Suchdol, Czech Republic ORCID ID:E-mail: sharmar@fld.czu.cz (email)
  • Vacek, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Kamýcká 129, 16521, Praha 6 – Suchdol, Czech Republic ORCID ID:E-mail: vacekz@fld.czu.cz
  • Vacek, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Kamýcká 129, 16521, Praha 6 – Suchdol, Czech Republic ORCID ID:E-mail: vacekstanislav@fld.czu.cz
article id 1740, category Research article
Ram P. Sharma, Zdeněk Vacek, Stanislav Vacek. (2017). Modelling tree crown-to-bole diameter ratio for Norway spruce and European beech. Silva Fennica vol. 51 no. 5 article id 1740. https://doi.org/10.14214/sf.1740
Highlights: Modelled crown-to-bole diameter ratio (CDBDR) using tree and stand-level predictors, and sample plot random effects; Spatially explicit mixed-effects model described the largest part of CDBDR variation with no significant trend in the residuals; The CDBDR increased with increasing stand development stage and site quality, but decreased with decreasing proportion of the species of interest, and increasing competition.

Crown dimensions are correlated to growth of other parts of a tree and often used as predictors in growth models. The crown-to-bole diameter ratio (CDBDR), which is a ratio of maximum crown width to diameter at breast height (DBH), was modelled using data from permanent sample plots located on Norway spruce (Picea abies (L.) Karst.) and European beech (Fagus sylvatica L.) stands in different parts of the Czech Republic. Among various tree and stand-level measures evaluated, DBH, height to crown base (HCB), dominant height (HDOM), basal area of trees larger in diameter than a subject tree (BAL), basal area proportion of the species of interest (BAPOR), and Hegyi’s competition index (CI) were found to be significant predictors in the CDBDR model. Random effects were included using the mixed-effects modelling to describe sample plot-level variation. For each species, the mixed-effects model described a larger part of the variation of the CDBDR than nonlinear ordinary least squares model with no trend in the residuals. The spatially explicit mixed-effects model showed more attractive fit statistics [conditional R2 ≈ 0.73 (spruce), 0.78 (beech)] than its spatially inexplicit counterpart [conditional R2 ≈ 0.71 (spruce), 0.76 (beech)]. The model showed that CDBDR increased with increasing HDOM – a measure that combines the stand development stage and site quality – but decreased with increasing HCB and competition (increasing BAL and CI), and decreasing proportions of the species of interest (increasing BAPOR). For both species, the spatially explicit mixed-effects model should be a preferred choice for a precise prediction of the CDBDR. The CDBDR model will have various management implications such as determination of spacing, stand basal area, stocking, and planning of appropriate species mixture.

  • Sharma, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Kamýcká 129, 16521, Praha 6 – Suchdol, Czech Republic ORCID ID:E-mail: sharmar@fld.czu.cz (email)
  • Vacek, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Kamýcká 129, 16521, Praha 6 – Suchdol, Czech Republic ORCID ID:E-mail: vacekz@fld.czu.cz
  • Vacek, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Kamýcká 129, 16521, Praha 6 – Suchdol, Czech Republic ORCID ID:E-mail: vacekstanislav@fld.czu.cz
article id 7753, category Research article
Aarne Hovi, Pekka Raitio, Miina Rautiainen. (2017). A spectral analysis of 25 boreal tree species. Silva Fennica vol. 51 no. 4 article id 7753. https://doi.org/10.14214/sf.7753
Highlights: An extensive spectral library containing leaf and needle reflectance and transmittance spectra was collected; The spectra openly available in SPECCHIO Spectral Information System; Effects of tree species, leaf/needle side, canopy position, and needle age on spectra were quantified; Seasonal variations were measured for four species; Spectra analysis highlights the importance of shortwave-infrared region in separating tree species.

Spectral libraries have a fundamental role in the development of interpretation methods for airborne and satellite-borne remote sensing data. This paper presents to-date the largest spectral measurement campaign of boreal tree species. Reflectance and transmittance spectra of over 600 leaf and needle samples from 25 species were measured in the Helsinki area (Finland) using integrating sphere systems attached to an ASD FieldSpec 4 spectroradiometer. Factors influencing the spectra and red edge inflection point (REIP) were quantified using one-way analysis of variance. Tree species differed most in the shortwave-infrared (1500–2500 nm) and least in the visible (400–700 nm) wavelength region. Species belonging to same genera showed similar spectral characteristics. Upper (adaxial) and lower (abaxial) leaf sides differed most in the visible region. Canopy position (sunlit/shaded) had a minor role in explaining spectral variation. For evergreen conifers, current and previous year needles differed in their spectra, current-year needles resembling those of broadleaved and deciduous conifers. Two broadleaved species were monitored throughout the growing season (May–October), and two conifers were measured twice during summer (June, September). Rapid changes were observed in the spectra in early spring and late autumn, whereas seasonal variations during summer months were relatively small for both broadleaved and coniferous species. Based on our results, shortwave-infrared seems promising in separating tree species, although it is to-date least studied. The spectral library reported here (Version 1.0) is publicly available through the SPECCHIO Spectral Information System.

  • Hovi, Aalto University, Department of Built Environment, P.O. Box 14100, FI-00076 Aalto, Finland ORCID ID:E-mail: aarne.hovi@aalto.fi (email)
  • Raitio, Aalto University, Department of Built Environment, P.O. Box 14100, FI-00076 Aalto, Finland ORCID ID:E-mail: pekka.raitio@aalto.fi
  • Rautiainen, Aalto University, Department of Built Environment, P.O. Box 14100, FI-00076 Aalto, Finland; Aalto University, Department of Electronics and Nanoengineering, P.O. Box 15500, FI-00076 Aalto, Finland ORCID ID:E-mail: miina.a.rautiainen@aalto.fi
article id 7753, category Research article
Aarne Hovi, Pekka Raitio, Miina Rautiainen. (2017). A spectral analysis of 25 boreal tree species. Silva Fennica vol. 51 no. 4 article id 7753. https://doi.org/10.14214/sf.7753
Highlights: An extensive spectral library containing leaf and needle reflectance and transmittance spectra was collected; The spectra openly available in SPECCHIO Spectral Information System; Effects of tree species, leaf/needle side, canopy position, and needle age on spectra were quantified; Seasonal variations were measured for four species; Spectra analysis highlights the importance of shortwave-infrared region in separating tree species.

Spectral libraries have a fundamental role in the development of interpretation methods for airborne and satellite-borne remote sensing data. This paper presents to-date the largest spectral measurement campaign of boreal tree species. Reflectance and transmittance spectra of over 600 leaf and needle samples from 25 species were measured in the Helsinki area (Finland) using integrating sphere systems attached to an ASD FieldSpec 4 spectroradiometer. Factors influencing the spectra and red edge inflection point (REIP) were quantified using one-way analysis of variance. Tree species differed most in the shortwave-infrared (1500–2500 nm) and least in the visible (400–700 nm) wavelength region. Species belonging to same genera showed similar spectral characteristics. Upper (adaxial) and lower (abaxial) leaf sides differed most in the visible region. Canopy position (sunlit/shaded) had a minor role in explaining spectral variation. For evergreen conifers, current and previous year needles differed in their spectra, current-year needles resembling those of broadleaved and deciduous conifers. Two broadleaved species were monitored throughout the growing season (May–October), and two conifers were measured twice during summer (June, September). Rapid changes were observed in the spectra in early spring and late autumn, whereas seasonal variations during summer months were relatively small for both broadleaved and coniferous species. Based on our results, shortwave-infrared seems promising in separating tree species, although it is to-date least studied. The spectral library reported here (Version 1.0) is publicly available through the SPECCHIO Spectral Information System.

  • Hovi, Aalto University, Department of Built Environment, P.O. Box 14100, FI-00076 Aalto, Finland ORCID ID:E-mail: aarne.hovi@aalto.fi (email)
  • Raitio, Aalto University, Department of Built Environment, P.O. Box 14100, FI-00076 Aalto, Finland ORCID ID:E-mail: pekka.raitio@aalto.fi
  • Rautiainen, Aalto University, Department of Built Environment, P.O. Box 14100, FI-00076 Aalto, Finland; Aalto University, Department of Electronics and Nanoengineering, P.O. Box 15500, FI-00076 Aalto, Finland ORCID ID:E-mail: miina.a.rautiainen@aalto.fi
article id 7728, category Research article
Liam Donnelly, Sven-Olof Lundqvist, Conor O’Reilly. (2017). Inter- and intra-annual wood property variation in juvenile wood between six Sitka spruce clones. Silva Fennica vol. 51 no. 4 article id 7728. https://doi.org/10.14214/sf.7728
Highlights: Wood property differences resulted primarily from variation in the proportions of early- and latewood in each annual ring; Width of early- and latewood bands in each ring was found to be a more important determinant of juvenile wood quality than the characteristics of the cells within each band; Wood properties differed greatly between clones, suggesting that there is potential to improve juvenile wood properties through selective breeding.

Increased growth rates have reduced rotation lengths, increasing the proportion of juvenile wood relative to mature wood, which may negatively affect mechanical performance of sawn timber. However, there is limited information available on the potential impact of breeding for vigour on juvenile wood in Sitka spruce (Picea sitchensis (Bong.) Carrière). In this study, the relationship between vigour (based on total height) and wood properties was investigated in six-year-old Sitka spruce clones grown in two replicated field trials in Ireland. Six clones were evaluated, two clones from each of three vigour (high, intermediate and low) classes. Discs were cut from the base of one ramet per replication for each clone to assess wood quality attributes. Radial tracheid width was significantly and positively correlated with ring width and height, and was negatively correlated with density. The wood of the most vigorous clone had significantly larger ring width with thinner cell walls and wider tracheids than all clones in the two other vigour classes, resulting in lower mean wood density. Latewood properties for all wood attributes measured differed significantly between the two sites. Wood property differences resulted primarily from variation in the proportions of early- and latewood in each annual ring. Additionally, the width of early- and latewood bands in each ring was found to be a more important determinant of juvenile wood quality than the characteristics of the cells within each band. Wood properties differed greatly between clones, suggesting that there is potential to improve juvenile wood properties through selective breeding.

  • Donnelly, UCD Forestry, University College Dublin, Belfield, Dublin 4, Ireland ORCID ID:E-mail: liam.donnelly@ucdconnect.ie (email)
  • Lundqvist, Innventia Ab, Drottning Kristinas väg 61, SE-114 86 Stockholm, Sweden ORCID ID:E-mail: svenolof.lundqvist@innventia.com
  • O’Reilly, UCD Forestry, University College Dublin, Belfield, Dublin 4, Ireland ORCID ID:E-mail: conor.oreilly@ucd.ie
article id 6991, category Research article
Isabel León, Juan José García, Manuel Fernández, Javier Vázquez-Piqué, Raúl Tapias. (2017). Differences in root growth of Quercus ilex and Quercus suber seedlings infected with Phytophthora cinnamomi. Silva Fennica vol. 51 no. 4 article id 6991. https://doi.org/10.14214/sf.6991
Highlights: Root growth of two Quercus sp. differs significantly after infection with Phytophthora cinnamomi; We observed a marked decrease in the growth of new roots in Quercus ilex with increasing inoculum level; Roots were longer but thinner with a moderate inoculum level in Quercus suber.

In the southwest of the Iberian Peninsula, Phytophthora cinnamomi Rands is causing irreversible damage to populations of the two most common species of Quercus, the holm oak (Quercus ilex L.) and the cork oak (Quercus suber L.). Although the symptoms are similar in the two species, the mortality rates are different. We found significant differences in the post-infection growth of the root system as a function of tree species, as well as initial plant size, and inoculum level. We observed a marked decrease in the growth of new roots in Q. ilex with increasing inoculum level, while in Q. suber, we found longer but thinner roots with a moderate inoculum level. In both species, we observed a worsening in the water status of the plants from the lowest inoculum level.

  • León, University of Huelva, Agroforestry department, Calle Dr. Cantero Cuadrado, 6, 21004 Huelva, Spain ORCID ID:E-mail: isabel.leon@dcaf.uhu.es
  • García, University of Huelva, Agroforestry department, Calle Dr. Cantero Cuadrado, 6, 21004 Huelva, Spain ORCID ID:E-mail: juanjose.garcia@dcaf.uhu.es
  • Fernández, University of Huelva, Agroforestry department, Calle Dr. Cantero Cuadrado, 6, 21004 Huelva, Spain ORCID ID:E-mail: nonoe@uhu.es
  • Vázquez-Piqué, University of Huelva, Agroforestry department, Calle Dr. Cantero Cuadrado, 6, 21004 Huelva, Spain ORCID ID:E-mail: jpique@dcaf.uhu.es
  • Tapias, University of Huelva, Agroforestry department, Calle Dr. Cantero Cuadrado, 6, 21004 Huelva, Spain ORCID ID:E-mail: rtapias@uhu.es (email)
article id 6991, category Research article
Isabel León, Juan José García, Manuel Fernández, Javier Vázquez-Piqué, Raúl Tapias. (2017). Differences in root growth of Quercus ilex and Quercus suber seedlings infected with Phytophthora cinnamomi. Silva Fennica vol. 51 no. 4 article id 6991. https://doi.org/10.14214/sf.6991
Highlights: Root growth of two Quercus sp. differs significantly after infection with Phytophthora cinnamomi; We observed a marked decrease in the growth of new roots in Quercus ilex with increasing inoculum level; Roots were longer but thinner with a moderate inoculum level in Quercus suber.

In the southwest of the Iberian Peninsula, Phytophthora cinnamomi Rands is causing irreversible damage to populations of the two most common species of Quercus, the holm oak (Quercus ilex L.) and the cork oak (Quercus suber L.). Although the symptoms are similar in the two species, the mortality rates are different. We found significant differences in the post-infection growth of the root system as a function of tree species, as well as initial plant size, and inoculum level. We observed a marked decrease in the growth of new roots in Q. ilex with increasing inoculum level, while in Q. suber, we found longer but thinner roots with a moderate inoculum level. In both species, we observed a worsening in the water status of the plants from the lowest inoculum level.

  • León, University of Huelva, Agroforestry department, Calle Dr. Cantero Cuadrado, 6, 21004 Huelva, Spain ORCID ID:E-mail: isabel.leon@dcaf.uhu.es
  • García, University of Huelva, Agroforestry department, Calle Dr. Cantero Cuadrado, 6, 21004 Huelva, Spain ORCID ID:E-mail: juanjose.garcia@dcaf.uhu.es
  • Fernández, University of Huelva, Agroforestry department, Calle Dr. Cantero Cuadrado, 6, 21004 Huelva, Spain ORCID ID:E-mail: nonoe@uhu.es
  • Vázquez-Piqué, University of Huelva, Agroforestry department, Calle Dr. Cantero Cuadrado, 6, 21004 Huelva, Spain ORCID ID:E-mail: jpique@dcaf.uhu.es
  • Tapias, University of Huelva, Agroforestry department, Calle Dr. Cantero Cuadrado, 6, 21004 Huelva, Spain ORCID ID:E-mail: rtapias@uhu.es (email)
article id 6977, category Research article
Anett Schibalski, Aleksi Lehtonen, Thomas Hickler, Boris Schröder. (2017). Identifying important topics for model refinement in a widely used process-based model informed by correlative model analyses in a boreal forest. Silva Fennica vol. 51 no. 4 article id 6977. https://doi.org/10.14214/sf.6977
Highlights: Continental-scale model parameterization of widely used LPJ-GUESS experiences problems when applied on the regional level; Competition, disturbances and soil conditions are crucial for explaining treeline position in Finland, besides climatic limitation; Picea abies is overly dominant in LPJ-GUESS model, as key competitive mechanisms are not implemented in sufficient detail.

Models attempting to predict treeline shifts in changing climates must include the relevant ecological processes in sufficient detail. A previous correlative model study has pointed to nutrients, competition, and temperature as the most important factors shaping the treelines of Pinus sylvestris L., Picea abies (L.) H. Karst. and Betula pubescens Ehrh. in Finnish Lapland. Here, we applied a widely used process-based dynamic vegetation model (LPJ-GUESS) to (i) test its capability to simulate observed spatial and temporal patterns of the main tree species in Finnish Lapland, and (ii) to explore the model representation of important processes in order to guide further model development. A European parameterization of LPJ-GUESS overestimated especially P. abies biomass and the species’ northern range limit. We identified implemented processes to adjust (competition, disturbance) and crucial processes in boreal forests to include (nutrient limitation, forest management) which account for the model’s failure to (edaphically) restrict P. abies in Finnish Lapland and the resulting species imbalance. Key competitive mechanisms are shade and drought tolerance, nutrient limitation, fire resistance, and susceptibility to disturbances (storm, herbivory) which we discussed with respect to boreal ecology and promising model developments to provide a starting point for future model development.

  • Schibalski, Institute of Geoecology, Technische Universität Braunschweig, Langer Kamp 19c, D-38106 Braunschweig, Germany ORCID ID:E-mail: a.schibalski@tu-braunschweig.de (email)
  • Lehtonen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 2, FI-00791 Helsinki, Finland ORCID ID:E-mail: aleksi.lehtonen@luke.fi
  • Hickler, Biodiversity and Climate Research Centre (BiK-F), Senckenberganlage 25, D-60325 Frankfurt am Main, Germany; Department of Physical Geography, Goethe University, Altenhöferallee 1, D-60438 Frankfurt am Main, Germany ORCID ID:E-mail: thomas.hickler@senckenberg.de
  • Schröder, Institute of Geoecology, Technische Universität Braunschweig, Langer Kamp 19c, D-38106 Braunschweig, Germany; Berlin-Brandenburg Institute of Advanced Biodiversity Research BBIB, Altensteinstr. 6, D-14195 Berlin, Germany ORCID ID:E-mail: boris.schroeder@tu-bs.de
article id 1691, category Research article
Enéas Ricardo Konzen, Raquel Peron, Márcio Akira Ito, Gilvano Ebling Brondani, Siu Mui Tsai. (2017). Molecular identification of bamboo genera and species based on RAPD-RFLP markers. Silva Fennica vol. 51 no. 4 article id 1691. https://doi.org/10.14214/sf.1691
Highlights: We investigated the potential of RAPD-RFLP (Random Amplified Polymorphic DNA, Restriction Fragment Length Polymorphism) in detecting genetic relationships among bamboo genera and species; RFLP profiles resolved in acrylamide gels revealed high number of markers, which accurately differentiated species and genera, based on cophenetic correlation coefficients; We recommend RAPD-RFLP for analyses of genetic diversity and divergence among bamboo genera, species and varieties.

Bamboo species have a very significant ecological and economic impact. Determining morphological and genetic differences among bamboo genera and species are crucial to explore desirable traits for breeding purposes. Several advances have been made in the taxonomy of bamboos by using molecular fingerprinting tools and next generation sequencing technologies. Nevertheless, classical molecular markers such as RAPD (Random Amplified Polymorphic DNA), AFLP (Amplified Fragment Length Polymorphism) and ISSR (Inter Simple Sequence Repeats) also provide an accurate discrimination among genera and species. Moreover, the RAPD-RFLP (Random Amplified Polymorphic DNA, Restriction Fragment Length Polymorphism) method, in which amplification products from RAPD are digested with restriction enzymes, is a reliable, fast and cost-effective method for fingerprinting. RAPD-RFLP has been scarcely used in the literature and no report regarding bamboo taxonomy is available with this method. Here we explored the molecular (RAPD, RAPD-RFLP) variation among genera (Bambusa, Dendrocalamus, Guadua and Phyllostachys) and species of bamboo cultivated in Brazil. Both molecular markers allowed clear distinction among the genera studied. Moreover, high cophenetic correlation values in UPGMA clusters indicated their potential for discriminating bamboo species. The digestion of RAPD products (RFLP) resulted in high number of polymorphic bands and produced very characteristic profiles for each genus with three enzyme combinations (HindIII/HaeIII, HinfI/RsaI, and single digestion with MspI). We recommend RAPD-RFLP as a reproducible and informative method for screening differences among genera, species and varieties of bamboos. Providing a cost-effective and accurate method for species identification and characterization is straightforward for bamboo conservation, management and breeding.

  • Konzen, Cell and Molecular Biology Laboratory, Center for Nuclear Energy in Agriculture (CENA), University of Sao Paulo (USP), Centenário Av., 303, Piracicaba, SP, P.O. Box 96, Brazil; Forest Sciences, Federal University of Lavras (UFLA), Lavras, MG, P.O. Box 3037, Brazil ORCID ID:E-mail: erkonzen@gmail.com (email)
  • Peron, Cell and Molecular Biology Laboratory, Center for Nuclear Energy in Agriculture (CENA), University of Sao Paulo (USP), Centenário Av., 303, Piracicaba, SP, P.O. Box 96, Brazil ORCID ID:E-mail: rperon@purdue.edu
  • Ito, Embrapa Western Agriculture, Brazilian Agricultural Research Corporation (EMBRAPA), BR 163 Rd., km 253, Dourados, MS, P.O. Box 449, Brazil ORCID ID:E-mail: marcio.ito@embrapa.br
  • Brondani, Forest Sciences, Federal University of Lavras (UFLA), Lavras, MG, P.O. Box 3037, Brazil ORCID ID:E-mail: gebrondani@gmail.com
  • Tsai, Cell and Molecular Biology Laboratory, Center for Nuclear Energy in Agriculture (CENA), University of Sao Paulo (USP), Centenário Av., 303, Piracicaba, SP, P.O. Box 96, Brazil ORCID ID:E-mail: tsai@cena.usp.br
article id 1691, category Research article
Enéas Ricardo Konzen, Raquel Peron, Márcio Akira Ito, Gilvano Ebling Brondani, Siu Mui Tsai. (2017). Molecular identification of bamboo genera and species based on RAPD-RFLP markers. Silva Fennica vol. 51 no. 4 article id 1691. https://doi.org/10.14214/sf.1691
Highlights: We investigated the potential of RAPD-RFLP (Random Amplified Polymorphic DNA, Restriction Fragment Length Polymorphism) in detecting genetic relationships among bamboo genera and species; RFLP profiles resolved in acrylamide gels revealed high number of markers, which accurately differentiated species and genera, based on cophenetic correlation coefficients; We recommend RAPD-RFLP for analyses of genetic diversity and divergence among bamboo genera, species and varieties.

Bamboo species have a very significant ecological and economic impact. Determining morphological and genetic differences among bamboo genera and species are crucial to explore desirable traits for breeding purposes. Several advances have been made in the taxonomy of bamboos by using molecular fingerprinting tools and next generation sequencing technologies. Nevertheless, classical molecular markers such as RAPD (Random Amplified Polymorphic DNA), AFLP (Amplified Fragment Length Polymorphism) and ISSR (Inter Simple Sequence Repeats) also provide an accurate discrimination among genera and species. Moreover, the RAPD-RFLP (Random Amplified Polymorphic DNA, Restriction Fragment Length Polymorphism) method, in which amplification products from RAPD are digested with restriction enzymes, is a reliable, fast and cost-effective method for fingerprinting. RAPD-RFLP has been scarcely used in the literature and no report regarding bamboo taxonomy is available with this method. Here we explored the molecular (RAPD, RAPD-RFLP) variation among genera (Bambusa, Dendrocalamus, Guadua and Phyllostachys) and species of bamboo cultivated in Brazil. Both molecular markers allowed clear distinction among the genera studied. Moreover, high cophenetic correlation values in UPGMA clusters indicated their potential for discriminating bamboo species. The digestion of RAPD products (RFLP) resulted in high number of polymorphic bands and produced very characteristic profiles for each genus with three enzyme combinations (HindIII/HaeIII, HinfI/RsaI, and single digestion with MspI). We recommend RAPD-RFLP as a reproducible and informative method for screening differences among genera, species and varieties of bamboos. Providing a cost-effective and accurate method for species identification and characterization is straightforward for bamboo conservation, management and breeding.

  • Konzen, Cell and Molecular Biology Laboratory, Center for Nuclear Energy in Agriculture (CENA), University of Sao Paulo (USP), Centenário Av., 303, Piracicaba, SP, P.O. Box 96, Brazil; Forest Sciences, Federal University of Lavras (UFLA), Lavras, MG, P.O. Box 3037, Brazil ORCID ID:E-mail: erkonzen@gmail.com (email)
  • Peron, Cell and Molecular Biology Laboratory, Center for Nuclear Energy in Agriculture (CENA), University of Sao Paulo (USP), Centenário Av., 303, Piracicaba, SP, P.O. Box 96, Brazil ORCID ID:E-mail: rperon@purdue.edu
  • Ito, Embrapa Western Agriculture, Brazilian Agricultural Research Corporation (EMBRAPA), BR 163 Rd., km 253, Dourados, MS, P.O. Box 449, Brazil ORCID ID:E-mail: marcio.ito@embrapa.br
  • Brondani, Forest Sciences, Federal University of Lavras (UFLA), Lavras, MG, P.O. Box 3037, Brazil ORCID ID:E-mail: gebrondani@gmail.com
  • Tsai, Cell and Molecular Biology Laboratory, Center for Nuclear Energy in Agriculture (CENA), University of Sao Paulo (USP), Centenário Av., 303, Piracicaba, SP, P.O. Box 96, Brazil ORCID ID:E-mail: tsai@cena.usp.br
article id 1631, category Research article
Jonas Koala, Louis Sawadogo, Patrice Savadogo, Ermias Aynekulu, Janne Heiskanen, Mohammed Saïd. (2017). Allometric equations for below-ground biomass of four key woody species in West African savanna-woodlands. Silva Fennica vol. 51 no. 3 article id 1631. https://doi.org/10.14214/sf.1631
Highlights: Species-specific equations for belowground biomass (BGB) predicted biomass with less bias than generic equations; All the generic equations underestimated BGB; For accurate estimation of BGB in savanna-woodlands, species-specific equations are needed for more species.

Accurate estimates of both above-ground biomass (AGB) and below-ground biomass (BGB) are essential for estimating carbon (C) balances at various geographical scales and formulating effective climate change mitigation programs. However, estimating BGB is challenging, particularly for forest ecosystems, so robust allometric equations are needed. To obtain such equations for savanna-woodlands of the West African north sudanian zone, we selected four common native woody species (Anogeissus leiocarpa (DC.) Guill. & Perr., Detarium microcarpum Guill. & Perr., Piliostigma thonningii (Schumach.) Milne-Redh. and Vitellaria paradoxa C.F. Gaertn.). At two sites in Burkina Faso, we determined the BGB of 30 trees of each of these species by excavation, and measured various above-ground dimensional variables. The root:shoot ratio varied widely among the species, from 0.1 to 3.4. Depending on the species, allometric equations based on stem basal area at 20 cm height, basal area at breast height and tree height explained 50–95% of the variation in BGB. The best generic equation we obtained, based on basal area at 20 cm, explained 60% of the variation in BGB across the species. Three previously published generic allometric equations underestimated BGB by 8 to 63%. The presented equations should significantly improve the accuracy of BGB estimates in savanna-woodlands and help avoid costly needs to excavate root systems.

  • Koala, Centre National de Recherche Scientifique et Technologique (CNRST), Institut de l’Environnement et de Recherches Agricoles (INERA), Département Productions Forestières, 03 BP 7047, Ouagadougou 03, Burkina Faso ORCID ID:E-mail: ezeyamb@yahoo.fr (email)
  • Sawadogo, Centre National de Recherche Scientifique et Technologique (CNRST), Institut de l’Environnement et de Recherches Agricoles (INERA), Département Productions Forestières, 03 BP 7047, Ouagadougou 03, Burkina Faso ORCID ID:E-mail: sawadogo_ls@hotmail.com
  • Savadogo, World Agroforestry Centre & International Crop Research Institute for the Semi-Arid Tropics (ICRAF-ICRISAT), West and Central Africa Region-Sahel Node, BP 12404, Niamey, Niger ORCID ID:E-mail: savadogo.patrice@gmail.com
  • Aynekulu, World Agroforestry Centre (ICRAF), United Nations Avenue, P.O. Box 30677-00100, Nairobi, Kenya ORCID ID:E-mail: e.betemariam@cgiar.org
  • Heiskanen, University of Helsinki, Department of Geosciences and Geography, P.O. Box 68, FI-00014 University of Helsinki, Finland ORCID ID:E-mail: janne.heiskanen@helsinki.fi
  • Saïd, International Livestock Research Institute (ILRI). P.O. Box 30709, Nairobi, Kenya ORCID ID:E-mail: m.said@cgiar.org
article id 5643, category Research article
Katarzyna A. Jadwiszczak, Stanisław Kłosowski, Iwona Zalewska, Agata Banaszek, Agnieszka Chrzanowska. (2017). Genetic diversity and sexual reproduction in relict populations of Betula nana. Silva Fennica vol. 51 no. 2 article id 5643. https://doi.org/10.14214/sf.5643
Highlights: Genetic diversity parameters and meiotic recombination frequencies in the relict populations were comparable to those from widespread localities; Contribution of seeds without ovule was very high; Fully developed seeds germinated better in central populations; Significant differences of groundwater parameters were observed between relict and central populations.

In the present study, the impact of geographical isolation and habitat conditions on genetic diversity and sexual reproduction was tested in four relict populations of dwarf birch Betula nana L. in Poland and Belarus. Amplified fragment length polymorphism (AFLP) method revealed that the endangered central European stands were not genetically extirpated compared with the widespread localities from Finland and Russia, which can result from infrequent outcrossing events in long-living clonal populations. However, genetic clustering methods indicated significant differentiation of the Polish populations because of their small sizes and long-term geographical isolation. Considerable numbers of empty seeds were observed in both relict and central locations, although fully developed seeds germinated better in widespread populations. Analysis of groundwater chemical parameters indicated that two relict populations were significantly different from the remaining samples with respect to pH, electrical conductivity and concentrations of phosphorus ions, which can also influence the efficiency of sexual reproduction. In the light of results obtained it seems that endangered B. nana localities are relatively stable.

  • Jadwiszczak, Institute of Biology, University of Białystok, Ciołkowskiego 1J, 15-245 Białystok, Poland ORCID ID: http://orcid.org/0000-0002-9345-8891 E-mail: jadwiszczak2010@gmail.com (email)
  • Kłosowski, Department of Environment Protection and Modelling, The Jan Kochanowski University, Świętokrzyska 15, 25-406 Kielce, Poland ORCID ID:E-mail: stanislaw.klosowski@ujk.kielce.pl
  • Zalewska, Faculty of Pharmacy, Medical University of Białystok, Mickiewicza 2a, 15-222 Białystok, Poland ORCID ID:E-mail: iwonazalewska1988@gmail.com
  • Banaszek, Institute of Biology, University of Białystok, Ciołkowskiego 1J, 15-245 Białystok, Poland ORCID ID:E-mail: banaszek@uwb.edu.pl
  • Chrzanowska, Institute of Biology, University of Białystok, Ciołkowskiego 1J, 15-245 Białystok, Poland ORCID ID:E-mail: maga.chrzanowska@gmail.com
article id 1781, category Research article
Petr Čermák, Michal Rybníček, Tomáš Žid, Kjell Andreassen, Isabella Børja, Tomáš Kolář. (2017). Impact of climate change on growth dynamics of Norway spruce in south-eastern Norway. Silva Fennica vol. 51 no. 2 article id 1781. https://doi.org/10.14214/sf.1781
Highlights: Correlations between tree-ring width and climate parameters showed temporal instability in their relationship during the period 1915–2012; A statistically significant positive correlation of April–May precipitation on tree-ring growth was identified since the mid-1970s; The concomitant temperature increase may have contributed to the changes of growth dynamics.

The ongoing climate change may have a distinct effect on Norway spruce growth, one of the most important tree species in European forest management. Therefore, the understanding and assessment of climate-growth relationship can help to reveal relevant patterns in temporal variability that may result in lower tree vitality and decline. The main objective of our study was to evaluate the long-term climate-growth variability of Norway spruce in south-eastern Norway, at the northern edge of the temperate zone. We sampled in total 270 dominant and co-dominant trees from 18 plots in south-eastern Norway. We analysed stem cores and evaluated crown condition parameters to assess the retrospective tree growth and vitality. Despite considerable differences in the crown parameters, high similarity among tree-ring width (TRW) series allowed compiling the regional tree-ring width chronology. Correlations between TRW and climate parameters showed temporal instability in their relationship during the period 1915–2012. While we did not detect any significant relationships between TRW and climate parameters in the first half of the study period (1915–1963), a significant correlation between TRW and spring precipitation was observed for the period 1964–2012. This shift appeared concurrent with temperatures reaching above-average values compared to the average of the climate normal period 1961–1990.

  • Čermák, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědělská 3, 613 00 Brno, Czech Republic ORCID ID:E-mail: cermacek@mendelu.cz (email)
  • Rybníček, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědělská 3, 613 00 Brno, Czech Republic; Global Change Research Institute, The Czech Academy of Sciences, Bělidla 986/4a, 603 00 Brno, Czech Republic ORCID ID:E-mail: michalryb@post.cz
  • Žid, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědělská 3, 613 00 Brno, Czech Republic ORCID ID:E-mail: tom.z@centrum.cz
  • Andreassen, Norwegian Institute of Bioeconomy Research (NIBIO), P.O. Box 115, NO-1431 Ås, Norway ORCID ID:E-mail: Kjell.Andressen@nibio.no
  • Børja, Norwegian Institute of Bioeconomy Research (NIBIO), P.O. Box 115, NO-1431 Ås, Norway ORCID ID:E-mail: Isabella.Borja@nibio.no
  • Kolář, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědělská 3, 613 00 Brno, Czech Republic; Global Change Research Institute, The Czech Academy of Sciences, Bělidla 986/4a, 603 00 Brno, Czech Republic ORCID ID:E-mail: koldatom@gmail.com
article id 1781, category Research article
Petr Čermák, Michal Rybníček, Tomáš Žid, Kjell Andreassen, Isabella Børja, Tomáš Kolář. (2017). Impact of climate change on growth dynamics of Norway spruce in south-eastern Norway. Silva Fennica vol. 51 no. 2 article id 1781. https://doi.org/10.14214/sf.1781
Highlights: Correlations between tree-ring width and climate parameters showed temporal instability in their relationship during the period 1915–2012; A statistically significant positive correlation of April–May precipitation on tree-ring growth was identified since the mid-1970s; The concomitant temperature increase may have contributed to the changes of growth dynamics.

The ongoing climate change may have a distinct effect on Norway spruce growth, one of the most important tree species in European forest management. Therefore, the understanding and assessment of climate-growth relationship can help to reveal relevant patterns in temporal variability that may result in lower tree vitality and decline. The main objective of our study was to evaluate the long-term climate-growth variability of Norway spruce in south-eastern Norway, at the northern edge of the temperate zone. We sampled in total 270 dominant and co-dominant trees from 18 plots in south-eastern Norway. We analysed stem cores and evaluated crown condition parameters to assess the retrospective tree growth and vitality. Despite considerable differences in the crown parameters, high similarity among tree-ring width (TRW) series allowed compiling the regional tree-ring width chronology. Correlations between TRW and climate parameters showed temporal instability in their relationship during the period 1915–2012. While we did not detect any significant relationships between TRW and climate parameters in the first half of the study period (1915–1963), a significant correlation between TRW and spring precipitation was observed for the period 1964–2012. This shift appeared concurrent with temperatures reaching above-average values compared to the average of the climate normal period 1961–1990.

  • Čermák, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědělská 3, 613 00 Brno, Czech Republic ORCID ID:E-mail: cermacek@mendelu.cz (email)
  • Rybníček, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědělská 3, 613 00 Brno, Czech Republic; Global Change Research Institute, The Czech Academy of Sciences, Bělidla 986/4a, 603 00 Brno, Czech Republic ORCID ID:E-mail: michalryb@post.cz
  • Žid, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědělská 3, 613 00 Brno, Czech Republic ORCID ID:E-mail: tom.z@centrum.cz
  • Andreassen, Norwegian Institute of Bioeconomy Research (NIBIO), P.O. Box 115, NO-1431 Ås, Norway ORCID ID:E-mail: Kjell.Andressen@nibio.no
  • Børja, Norwegian Institute of Bioeconomy Research (NIBIO), P.O. Box 115, NO-1431 Ås, Norway ORCID ID:E-mail: Isabella.Borja@nibio.no
  • Kolář, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědělská 3, 613 00 Brno, Czech Republic; Global Change Research Institute, The Czech Academy of Sciences, Bělidla 986/4a, 603 00 Brno, Czech Republic ORCID ID:E-mail: koldatom@gmail.com
article id 1713, category Research article
Lars-Göran Stener, Lars Rytter, Gunnar Jansson. (2017). Effects of pruning on wood properties of planted silver birch in southern Sweden. Silva Fennica vol. 51 no. 2 article id 1713. https://doi.org/10.14214/sf.1713
Highlights: Pruning silver birch trees increased the production of defect-free wood outside the knots; Most wood defects were found inside the knots; Pruned birch trees provide butt logs with higher value than unpruned trees.

Pruning was performed at midsummer in two genetically homogenous and managed planted silver birch stands in southern Sweden – one aged 9 and one aged 10 years. Wood defects were analysed 10 years thereafter, using the five uppermost twigs of the stems up to a height of 30 dm. The number of trees examined at each site was around 70, of which half were pruned. The main findings were that: a) compared to unpruned trees, pruned trees produced more defect-free wood outside the knots; b) most wood defects were found inside the knots; and c) wood defects like rot and bark ingrowth were similar for pruned and unpruned trees, while discolouration was marginally higher for pruned trees inside knots but similar outside knots. Overall, the results confirm previous findings that pruned birch trees will provide butt logs with higher value than unpruned trees.

  • Stener, The Forestry Research Institute of Sweden, Ekebo 2250, 268 90 Svalöv, Sweden ORCID ID:E-mail: lars-goran.stener@skogforsk.se (email)
  • Rytter, The Forestry Research Institute of Sweden, Ekebo 2250, 268 90 Svalöv, Sweden ORCID ID:E-mail: lars.rytter@skogforsk.se
  • Jansson, The Forestry Research Institute of Sweden, Uppsala Science Park, 751 83 Uppsala, Sweden ORCID ID:E-mail: gunnar.jansson@skogforsk.se
article id 1778, category Research article
Adriano Mazziotta, Dmitry Podkopaev, María Triviño, Kaisa Miettinen, Tähti Pohjanmies, Mikko Mönkkönen. (2017). Quantifying and resolving conservation conflicts in forest landscapes via multiobjective optimization. Silva Fennica vol. 51 no. 1 article id 1778. https://doi.org/10.14214/sf.1778
Highlights: We introduce a compatibility index quantifying how targeting a management objective in the forest landscape affects another objective; To resolve conflicts we find compromise solutions minimizing the maximum deterioration among objectives; We apply our approach for a case study of forest management for biodiversity conservation and development; Multiple use management and careful planning can reduce biodiversity conflicts in forest ecosystems.

Environmental planning for of the maintenance of different conservation objectives should take into account multiple contrasting criteria based on alternative uses of the landscape. We develop new concepts and approaches to describe and measure conflicts among conservation objectives and for resolving them via multiobjective optimization. To measure conflicts we introduce a compatibility index that quantifies how much targeting a certain conservation objective affects the capacity of the landscape for providing another objective. To resolve such conflicts we find compromise solutions defined in terms of minimax regret, i.e. minimizing the maximum percentage of deterioration among conservation objectives. Finally, we apply our approach for a case study of management for biodiversity conservation and development in a forest landscape. We study conflicts between six different forest species, and we identify management solutions for simultaneously maintaining multiple species’ habitat while obtaining timber harvest revenues. We employ the method for resolving conflicts at a large landscape level across a long 50-years forest planning horizon. Our multiobjective approach can be an instrument for guiding hard choices in the conservation-development nexus with a perspective of developing decision support tools for land use planning. In our case study multiple use management and careful landscape level planning using our approach can reduce conflicts among biodiversity objectives and offer room for synergies in forest ecosystems.

  • Mazziotta, University of Jyväskylä, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland; Center for Macroecology Evolution and Climate, University of Copenhagen, Universitetsparken 15, DK-2100 Copenhagen, Denmark; Stockholm Resilience Centre, Stockholm University, Kräftriket 2b, 11429 Stockholm, Sweden ORCID ID: http://orcid.org/0000-0003-2088-3798 E-mail: a_mazziotta@hotmail.com (email)
  • Podkopaev, University of Jyväskylä, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland; Systems Research Institute, Polish Academy of Sciences, Newelska 6, 01-447 Warsaw, Poland ORCID ID:E-mail: dmitry.podkopaev@ibspan.waw.pl
  • Triviño, University of Jyväskylä, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland ORCID ID:E-mail: maria.trivino@jyu.fi
  • Miettinen, University of Jyväskylä, Faculty of Information Technology, P.O. Box 35, FI-40014 University of Jyväskylä, Finland ORCID ID:E-mail: kaisa.miettinen@jyu.fi
  • Pohjanmies, University of Jyväskylä, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland ORCID ID:E-mail: tahti.t.pohjanmies@jyu.fi
  • Mönkkönen, University of Jyväskylä, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland ORCID ID:E-mail: mikko.monkkonen@jyu.fi
article id 1778, category Research article
Adriano Mazziotta, Dmitry Podkopaev, María Triviño, Kaisa Miettinen, Tähti Pohjanmies, Mikko Mönkkönen. (2017). Quantifying and resolving conservation conflicts in forest landscapes via multiobjective optimization. Silva Fennica vol. 51 no. 1 article id 1778. https://doi.org/10.14214/sf.1778
Highlights: We introduce a compatibility index quantifying how targeting a management objective in the forest landscape affects another objective; To resolve conflicts we find compromise solutions minimizing the maximum deterioration among objectives; We apply our approach for a case study of forest management for biodiversity conservation and development; Multiple use management and careful planning can reduce biodiversity conflicts in forest ecosystems.

Environmental planning for of the maintenance of different conservation objectives should take into account multiple contrasting criteria based on alternative uses of the landscape. We develop new concepts and approaches to describe and measure conflicts among conservation objectives and for resolving them via multiobjective optimization. To measure conflicts we introduce a compatibility index that quantifies how much targeting a certain conservation objective affects the capacity of the landscape for providing another objective. To resolve such conflicts we find compromise solutions defined in terms of minimax regret, i.e. minimizing the maximum percentage of deterioration among conservation objectives. Finally, we apply our approach for a case study of management for biodiversity conservation and development in a forest landscape. We study conflicts between six different forest species, and we identify management solutions for simultaneously maintaining multiple species’ habitat while obtaining timber harvest revenues. We employ the method for resolving conflicts at a large landscape level across a long 50-years forest planning horizon. Our multiobjective approach can be an instrument for guiding hard choices in the conservation-development nexus with a perspective of developing decision support tools for land use planning. In our case study multiple use management and careful landscape level planning using our approach can reduce conflicts among biodiversity objectives and offer room for synergies in forest ecosystems.

  • Mazziotta, University of Jyväskylä, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland; Center for Macroecology Evolution and Climate, University of Copenhagen, Universitetsparken 15, DK-2100 Copenhagen, Denmark; Stockholm Resilience Centre, Stockholm University, Kräftriket 2b, 11429 Stockholm, Sweden ORCID ID: http://orcid.org/0000-0003-2088-3798 E-mail: a_mazziotta@hotmail.com (email)
  • Podkopaev, University of Jyväskylä, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland; Systems Research Institute, Polish Academy of Sciences, Newelska 6, 01-447 Warsaw, Poland ORCID ID:E-mail: dmitry.podkopaev@ibspan.waw.pl
  • Triviño, University of Jyväskylä, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland ORCID ID:E-mail: maria.trivino@jyu.fi
  • Miettinen, University of Jyväskylä, Faculty of Information Technology, P.O. Box 35, FI-40014 University of Jyväskylä, Finland ORCID ID:E-mail: kaisa.miettinen@jyu.fi
  • Pohjanmies, University of Jyväskylä, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland ORCID ID:E-mail: tahti.t.pohjanmies@jyu.fi
  • Mönkkönen, University of Jyväskylä, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland ORCID ID:E-mail: mikko.monkkonen@jyu.fi
article id 1709, category Research article
Guoping Chen, Cong Shi, Shanshan Cheng, Tiejian Zhao, Guoquan Liu, Fuchen Shi. (2017). The structure and soil characteristics of a Pinus tabuliformis planted forest after 60 years of natural development in North China. Silva Fennica vol. 51 no. 1 article id 1709. https://doi.org/10.14214/sf.1709
Highlights: Increasing proportions of broadleaf tree species was shown to affect nutrient content of the forest floor and soil, and the soil microbial community in the process of natural development of Pinus tabuliformis planted forest. In this regard, this study can act as a reference for management of the near-natural transformation of P. tabuliformis planted forests and for the choice of the tree species used.

This study evaluated the transformation of a Pinus tabuliformis Carrière forest into a near-natural forest after 60 years of natural development. The structure and soil characteristics of P. tabuliformis planted forest, the near-natural forest (coniferous-broadleaved P. tabuliformis mixed forest), and secondary forest (Quercus mongolica Fisch. ex Ledeb. forest) were compared. Tree, shrub and herb species diversity of the mixed and Q. mongolica forests was higher than that of the planted P. tabuliformis forest. Examination of soil characteristics revealed that compared to the pure pine forest, nitrogen (N) and phosphorus (P) concentrations of the mixed and Q. mongolica forests increased in the forest floor and soil, but total carbon (C) concentration decreased in the forest floor, countered by increases in the soil. Furthermore, soil cation exchange capacity (CEC) and pH in the P. tabuliformis forest increased when deciduous broadleaved species were present. Total microbial biomass and bacterial biomass in the soils were greatest in the Q. mongolica forest, followed by the mixed, and then the P. tabuliformis forests. However, fungal biomass did not significantly differ among the three forests. Overall, the findings of this study suggest that different forest types can affect soil microbial biomass and community structure. Meanwhile, the natural development is recommended as a potential management alternative to near-natural transformation of a P. tabuliformis planted forest.

  • Chen, Department of Plant Biology & Ecology, College of Life Sciences, Nankai University, Weijin Road 94, Tianjin 300071, P.R. China ORCID ID:E-mail: guopingchern@mail.nankai.edu.cn
  • Shi, Graduate School of Agriculture, Hokkaido University, Sapporo, Hokkaido 060-8689, Japan ORCID ID:E-mail: cshi1@for.agr.hokudai.ac.jp
  • Cheng, School of Environment and Energy, Shenzhen Graduate School of Peking University, Shenzhen 518055, China ORCID ID:E-mail: 1401213932@sz.pku.edu.cn
  • Zhao, Baxian Mountain National Nature Reserve, Tianjin 301900, China Received 29 September 2016 Revised ORCID ID:E-mail: zhaotiejiann456@sina.com
  • Liu, Baxian Mountain National Nature Reserve, Tianjin 301900, China Received 29 September 2016 Revised ORCID ID:E-mail: liuguoquan01@163.com
  • Shi, Department of Plant Biology & Ecology, College of Life Sciences, Nankai University, Weijin Road 94, Tianjin 300071, P.R. China ORCID ID:E-mail: fcshi@nankai.edu.cn (email)
article id 1709, category Research article
Guoping Chen, Cong Shi, Shanshan Cheng, Tiejian Zhao, Guoquan Liu, Fuchen Shi. (2017). The structure and soil characteristics of a Pinus tabuliformis planted forest after 60 years of natural development in North China. Silva Fennica vol. 51 no. 1 article id 1709. https://doi.org/10.14214/sf.1709
Highlights: Increasing proportions of broadleaf tree species was shown to affect nutrient content of the forest floor and soil, and the soil microbial community in the process of natural development of Pinus tabuliformis planted forest. In this regard, this study can act as a reference for management of the near-natural transformation of P. tabuliformis planted forests and for the choice of the tree species used.

This study evaluated the transformation of a Pinus tabuliformis Carrière forest into a near-natural forest after 60 years of natural development. The structure and soil characteristics of P. tabuliformis planted forest, the near-natural forest (coniferous-broadleaved P. tabuliformis mixed forest), and secondary forest (Quercus mongolica Fisch. ex Ledeb. forest) were compared. Tree, shrub and herb species diversity of the mixed and Q. mongolica forests was higher than that of the planted P. tabuliformis forest. Examination of soil characteristics revealed that compared to the pure pine forest, nitrogen (N) and phosphorus (P) concentrations of the mixed and Q. mongolica forests increased in the forest floor and soil, but total carbon (C) concentration decreased in the forest floor, countered by increases in the soil. Furthermore, soil cation exchange capacity (CEC) and pH in the P. tabuliformis forest increased when deciduous broadleaved species were present. Total microbial biomass and bacterial biomass in the soils were greatest in the Q. mongolica forest, followed by the mixed, and then the P. tabuliformis forests. However, fungal biomass did not significantly differ among the three forests. Overall, the findings of this study suggest that different forest types can affect soil microbial biomass and community structure. Meanwhile, the natural development is recommended as a potential management alternative to near-natural transformation of a P. tabuliformis planted forest.

  • Chen, Department of Plant Biology & Ecology, College of Life Sciences, Nankai University, Weijin Road 94, Tianjin 300071, P.R. China ORCID ID:E-mail: guopingchern@mail.nankai.edu.cn
  • Shi, Graduate School of Agriculture, Hokkaido University, Sapporo, Hokkaido 060-8689, Japan ORCID ID:E-mail: cshi1@for.agr.hokudai.ac.jp
  • Cheng, School of Environment and Energy, Shenzhen Graduate School of Peking University, Shenzhen 518055, China ORCID ID:E-mail: 1401213932@sz.pku.edu.cn
  • Zhao, Baxian Mountain National Nature Reserve, Tianjin 301900, China Received 29 September 2016 Revised ORCID ID:E-mail: zhaotiejiann456@sina.com
  • Liu, Baxian Mountain National Nature Reserve, Tianjin 301900, China Received 29 September 2016 Revised ORCID ID:E-mail: liuguoquan01@163.com
  • Shi, Department of Plant Biology & Ecology, College of Life Sciences, Nankai University, Weijin Road 94, Tianjin 300071, P.R. China ORCID ID:E-mail: fcshi@nankai.edu.cn (email)
article id 1692, category Research article
Aleksey Fedorkov, Ludmila Gutiy. (2017). Performance of lodgepole pine and Scots pine in field trials located in north-west Russia. Silva Fennica vol. 51 no. 1 article id 1692. https://doi.org/10.14214/sf.1692
Highlights: Stem volume was bigger for lodgepole pine in comparison to local Scots pine except for the southernmost origin; The proportion of stems with no defects was lower for all lodgepole pine seed sources than for local Scots pine; Lodgepole pine stem growth traits were significantly related to latitude of seed origin.

Mortality, stem growth and quality of lodgepole pine (Pinus contorta var. latifolia) originating from the six Swedish seed orchards and local Scots pine (Pinus sylvestris L.) were estimated in four field trials established in the Komi Republic (north-west Russia). A randomized row-plot design with 6–12 replicates of each entry was used. The tree mortality was slightly higher for Scots pine than that for lodgepole pine, except for the lodgepole pine seed sources of the southern origins with lower survival. Scots pine stem quality was better than that of lodgepole pine, but the lodgepole pine stem growth was faster except the seed source of the southernmost origin. The lodgepole pine seed sources of northern origins had better stem growth (height, diameter at breast height and volume), while the effect of latitude on the quality traits was insignificant.

  • Fedorkov, Institute of Biology, Komi Science Center, Russian Academy of Sciences, 28 Kommunisticheskaya st., Syktyvkar 167982, Russia ORCID ID: http://orcid.org/0000-0001-7800-7534 E-mail: fedorkov@ib.komisc.ru (email)
  • Gutiy, Syktyvkar Forest Institute (branch), Saint-Petersburg State Forest Technical University, 39 Lenin st., Syktyvkar 167000, Russia ORCID ID:E-mail: lguti@mail.ru
article id 1566, category Research article
Valdir Marcos Stefenon, Jordana Caroline Nagel, Igor Poletto. (2016). Evidences of genetic bottleneck and fitness decline in Luehea divaricata populations from southern Brazil. Silva Fennica vol. 50 no. 5 article id 1566. https://doi.org/10.14214/sf.1566
Highlights: Signatures of genetic bottlenecks and reduction of populations’ fitness were observed in populations of Luehea divaricata in southern Brazil; Lower levels of observed heterozygosity are correlated with populations’ fitness, decreasing germination capacity and increasing the proportion of anomalous germinated plantlets; Promoting the connection among populations is proposed as a key strategy towards conservation of L. divaricata genetic resources in its southernmost distribution range.

Extant populations growing in regions that were refugia during the last glacial period are expected to show higher genetic diversity than populations that moved from these refugia into new areas in higher latitudes. Such new populations likely faced harsher climatic conditions, being established with reduced population size and experiencing the effects of genetic bottlenecks. In this study we employed data from nuclear SSR markers for detecting molecular signatures of genetic bottlenecks, and germination experiments to evaluate reduction of populations’ fitness in natural populations of Luehea divaricata Mart. et Zucc., growing in the southern range of the species distribution (around 30°S latitude). Signatures of genetic bottlenecks and reduction of populations’ fitness were observed in all populations. Lower levels of observed heterozygosity are correlated with populations’ fitness, decreasing germination capacity and increasing the proportion of anomalous germinated plantlets. Promoting the connection among populations is proposed as a key strategy towards conservation of L. divaricata genetic resources in its southernmost distribution range. The offspring from crosses among populations would significantly increase the observed heterozygosity and fitness of multiple populations.

  • Stefenon, Nucleus of Genomics and Molecular Ecology, Interdisciplinary Center of Biotechnological Research, Universidade Federal do Pampa, BR290 km, 97300-000, São Gabriel, RS, Brazil ORCID ID:E-mail: valdirstefenon@unipampa.edu.br (email)
  • Nagel, Nucleus of Genomics and Molecular Ecology, Interdisciplinary Center of Biotechnological Research, Universidade Federal do Pampa, BR290 km, 97300-000, São Gabriel, RS, Brazil ORCID ID:E-mail: jordana.nagel@yahoo.com.br
  • Poletto, Laboratory of Plant Protection, Universidade Federal do Pampa, BR290 km, 97300-000, São Gabriel, RS, Brazil ORCID ID:E-mail: igorpoletto@unipampa.edu.br
article id 1687, category Research article
Hannu Hökkä, Jori Uusitalo, Harri Lindeman, Jari Ala-Ilomäki. (2016). Performance of weather parameters in predicting growing season water table depth variations on drained forested peatlands – a case study from southern Finland. Silva Fennica vol. 50 no. 4 article id 1687. https://doi.org/10.14214/sf.1687
Highlights: Four-week precipitation and evapotranspiration explain much of drained peatland water table depth variation during a growing season.

The amount of water in peat soil is one factor affecting its bearing capacity, which is a crucial aspect in planning peatland timber harvesting operations. We studied the influence of weather variables on the variation of drained peatland growing season water conditions, here the ground water table depth (WTD). WTD was manually monitored four times in 2014 and three times in 2015 in 10–30 sample plots located in four drained peatland forests in south-western Finland. For each peatland, precipitation and evapotranspiration were calculated from the records of the nearest Finnish Meteorological Institute field stations covering periods from one day to four weeks preceding the WTD monitoring date. A mixed linear model was constructed to investigate the impact of the weather parameters on WTD. Precipitation of the previous four–week period was the most important explanatory variable. The four-week evapotranspiration amount was interacting with the Julian day showing a greater effect in late summer. Other variables influencing WTD were stand volume within the three-metre radius sample plot and distance from nearest ditch. Our results show the potential of weather parameters, specifically that of the previous four-week precipitation and evapotranspiration, for predicting drained peatland water table depth variation and subsequently, the possibility to develop a more general empirical model to assist planning of harvesting operations on drained peatlands.

  • Hökkä, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Paavo Havaksen tie 3, FI-90014 Oulun yliopisto, Finland ORCID ID:E-mail: hannu.hokka@luke.fi (email)
  • Uusitalo, Natural Resources Institute Finland (Luke), Green Technology, Kaironiementie 15, FI-39700 Parkano, Finland ORCID ID:E-mail: jori.uusitalo@luke.fi
  • Lindeman, Natural Resources Institute Finland (Luke), Green Technology, Kaironiementie 15, FI-39700 Parkano, Finland ORCID ID:E-mail: harri.lindeman@luke.fi
  • Ala-Ilomäki, Natural Resources Institute Finland (Luke), Green Technology, Jokiniemenkuja 1, FI-01370 Vantaa, Finland ORCID ID:E-mail: jari.ala-ilomaki@luke.fi
article id 1682, category Research article
Jiří Korecký, Yousry A. El-Kassaby. (2016). Pollination dynamics variation in a Douglas-fir seed orchard as revealed by microsatellite analysis. Silva Fennica vol. 50 no. 4 article id 1682. https://doi.org/10.14214/sf.1682
Highlights: Important characteristics such as parental reproductive success, pollen contamination, and selfing rate in the second generation Douglas-fir seed orchard have been estimated; Since this research is a part of a multi-year study, outputs were compared to those from two other years; Results are in line and differences in pollination dynamics across years are attributable to the various crop management practices.

As part of a multi-year monitoring study of pollination dynamics in a second generation Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) seed orchard, we estimated with the aid of eight microsatellite markers three important reproductive biology characteristics affecting the genetic worth and diversity of seed crops; namely parental reproductive success, pollen contamination, and selfing rate. The obtained results were compared to those from two previous years to gauge appropriate seed crop management practices and ultimately allow approximate generalization of seed crop genetic quality. We determined that 80% of parental gametes were produced by 52% of the parents, 13% of paternal gametes resulted from pollen contamination (i.e., gene flow), and 12% of the seed were the product of selfing. The obtained results were in line with those observed for 2005 and 2009 where 80% of gametes being produced by 37–48% of the parents, 10–18% pollen contamination, and 15–17% selfing rate. The observed reproductive biology parameters differences are attributable to the various crop management practices implemented (i.e., bloom delay and supplemental-mass-pollination) across years and calls for justification due to the observed minimal differences on seed crops genetic quality.

  • Korecký, Department of Genetics and Physiology of Forest Trees, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Praha 6, 165 21, Czech Republic ORCID ID: http://orcid.org/0000-0001-7859-1750 E-mail: korecky@fld.czu.cz (email)
  • El-Kassaby, Department of Forest and Conservation Sciences, Faculty of Forestry, University of British Columbia, Vancouver, BC, V6T 1Z4, Canada ORCID ID: http://orcid.org/0000-0002-4887-8977 E-mail: y.el-kassaby@ubc.ca
article id 1663, category Research article
Marta Kempf, Monika Konnert. (2016). Distribution of genetic diversity in Fagus sylvatica at the north-eastern edge of the natural range. Silva Fennica vol. 50 no. 4 article id 1663. https://doi.org/10.14214/sf.1663
Highlights: European beech at the north-eastern edge of its natural range in Poland have a high level of genetic variation, similar to the populations from Central Europe; The differences between the beech provenances from the two centres in Poland, which were previously identified on the basis of pollen analyses and phenotypic traits, have now been genetically confirmed.

An understanding of the genetic variation of the beech, especially at the edge of its natural distribution, is essential because of the change in natural distribution of the species resulting from changing climatic conditions. The main aim of the study was to determine the level of genetic diversity of European beech at the north-eastern edge of its natural range. The other aim was to check the genetic variation of beech from the two centres, the north and the south of Poland, which were identified in previous findings based on pollen analyses and phenotypic traits. The research material was the progeny of twelve beech provenances. The genetic structure of the populations was determined by ten highly variable microsatellite DNA loci. The results confirmed the high genetic diversity of beech at the north-eastern edge of its natural distribution, which infers the probability of their good adaptation to the changing climate and an extension of the range. Genetic analyses confirmed the existence of two genetic centres for beech in Poland. The populations from south-eastern Poland had a slightly higher diversity than the populations from the north-western area, which may indicate that the colonisation of Poland occurred by two routes. The results are important for creating the borders of the provenance regions and for limiting the transfer of seeds and seedlings. The choice of forest reproductive material, based on the knowledge of genetic diversity, is very important for the stability of future forests.

  • Kempf, Department of Genetics and Forest Tree Breeding, Faculty of Forestry, University of Agriculture in Krakow, Al. 29-listopada 46, 31–425 Krakow ORCID ID:E-mail: m.kempf@ur.krakow.pl (email)
  • Konnert, Bavarian Office for Forest Seeding and Planting, Forstamtsplatz 1, 83317 Teisendorf, Germany ORCID ID:E-mail: monika.konnert@asp.bayern.de
article id 1611, category Research article
Ilpo Ervasti. (2016). Wood fiber contents of different materials in the paper industry material chain expressed in roundwood equivalents (RWEs). Silva Fennica vol. 50 no. 4 article id 1611. https://doi.org/10.14214/sf.1611
Highlights: The paper industry material recycling chain consists of different material streams; Consumed paper products form a huge fiber source for industry. However, a reliable measuring system should be developed to quantify these fiber volumes; Converting factors are introduced to quantify the wood fiber contents of different paper industry-related materials of the chain, namely, wood pulp, paper and recovered paper; By using these conversion factors it is possible to convert for example global recovered paper utilization volumes into wood fiber volumes expressed in roundwood equivalents (RWEs).

At present, there are no means for reliably comparing the wood fiber contents of different material streams within the paper industry material chain with each other. The aim of this article is to introduce conversion factors that make it possible to quantify the volume of wood expressed in roundwood equivalent (RWE) values for different paper industry-related materials in the material chain. These conversion factors apply to wood pulp, paper, and recovered paper. European data are used in quantifying the paper industry material streams and calculating the RWE conversion factors. The introduced conversion factors can be used to estimate RWE volumes at a global scale. With assumption that paper recycling did not occur and that paper production volume remained unchanged, an additional volume of 666 million m3 RWEs would be required globally per annum to produce 167 million tons of virgin wood pulp to replace 222 million tons of recovered paper utilized by the paper industry in 2010. This volume is approximately the same as 1.6 times the total removal of wood in Europe (EU27), or the total annual removal of wood in the USA, Canada, and Brazil combined.

  • Ervasti, Aalto University, Industrial Engineering and Management, Maarintie 8, 02150 Espoo, Finland ORCID ID:E-mail: ilervasti@gmail.com (email)
article id 1611, category Research article
Ilpo Ervasti. (2016). Wood fiber contents of different materials in the paper industry material chain expressed in roundwood equivalents (RWEs). Silva Fennica vol. 50 no. 4 article id 1611. https://doi.org/10.14214/sf.1611
Highlights: The paper industry material recycling chain consists of different material streams; Consumed paper products form a huge fiber source for industry. However, a reliable measuring system should be developed to quantify these fiber volumes; Converting factors are introduced to quantify the wood fiber contents of different paper industry-related materials of the chain, namely, wood pulp, paper and recovered paper; By using these conversion factors it is possible to convert for example global recovered paper utilization volumes into wood fiber volumes expressed in roundwood equivalents (RWEs).

At present, there are no means for reliably comparing the wood fiber contents of different material streams within the paper industry material chain with each other. The aim of this article is to introduce conversion factors that make it possible to quantify the volume of wood expressed in roundwood equivalent (RWE) values for different paper industry-related materials in the material chain. These conversion factors apply to wood pulp, paper, and recovered paper. European data are used in quantifying the paper industry material streams and calculating the RWE conversion factors. The introduced conversion factors can be used to estimate RWE volumes at a global scale. With assumption that paper recycling did not occur and that paper production volume remained unchanged, an additional volume of 666 million m3 RWEs would be required globally per annum to produce 167 million tons of virgin wood pulp to replace 222 million tons of recovered paper utilized by the paper industry in 2010. This volume is approximately the same as 1.6 times the total removal of wood in Europe (EU27), or the total annual removal of wood in the USA, Canada, and Brazil combined.

  • Ervasti, Aalto University, Industrial Engineering and Management, Maarintie 8, 02150 Espoo, Finland ORCID ID:E-mail: ilervasti@gmail.com (email)
article id 1611, category Research article
Ilpo Ervasti. (2016). Wood fiber contents of different materials in the paper industry material chain expressed in roundwood equivalents (RWEs). Silva Fennica vol. 50 no. 4 article id 1611. https://doi.org/10.14214/sf.1611
Highlights: The paper industry material recycling chain consists of different material streams; Consumed paper products form a huge fiber source for industry. However, a reliable measuring system should be developed to quantify these fiber volumes; Converting factors are introduced to quantify the wood fiber contents of different paper industry-related materials of the chain, namely, wood pulp, paper and recovered paper; By using these conversion factors it is possible to convert for example global recovered paper utilization volumes into wood fiber volumes expressed in roundwood equivalents (RWEs).

At present, there are no means for reliably comparing the wood fiber contents of different material streams within the paper industry material chain with each other. The aim of this article is to introduce conversion factors that make it possible to quantify the volume of wood expressed in roundwood equivalent (RWE) values for different paper industry-related materials in the material chain. These conversion factors apply to wood pulp, paper, and recovered paper. European data are used in quantifying the paper industry material streams and calculating the RWE conversion factors. The introduced conversion factors can be used to estimate RWE volumes at a global scale. With assumption that paper recycling did not occur and that paper production volume remained unchanged, an additional volume of 666 million m3 RWEs would be required globally per annum to produce 167 million tons of virgin wood pulp to replace 222 million tons of recovered paper utilized by the paper industry in 2010. This volume is approximately the same as 1.6 times the total removal of wood in Europe (EU27), or the total annual removal of wood in the USA, Canada, and Brazil combined.

  • Ervasti, Aalto University, Industrial Engineering and Management, Maarintie 8, 02150 Espoo, Finland ORCID ID:E-mail: ilervasti@gmail.com (email)
article id 1567, category Research article
Eetu Kotivuori, Lauri Korhonen, Petteri Packalen. (2016). Nationwide airborne laser scanning based models for volume, biomass and dominant height in Finland. Silva Fennica vol. 50 no. 4 article id 1567. https://doi.org/10.14214/sf.1567
Highlights: Pooled data from nine inventory projects in Finland were used to create nationwide laser-based regression models for dominant height, volume and biomass; Volume and biomass models provided regionally different means than real means, but for dominant height the mean difference was small; The accuracy of general volume predictions was nevertheless comparable to relascope-based field inventory by compartments.

The aim of this study was to examine how well stem volume, above-ground biomass and dominant height can be predicted using nationwide airborne laser scanning (ALS) based regression models. The study material consisted of nine practical ALS inventory projects taken from different parts of Finland. We used field sample plots and airborne laser scanning data to create nationwide and regional models for each response variable. The final models had one or two ALS predictors, which were chosen based on the root mean square error (RMSE), and cross-validated. Finally, we tested how much predictions would improve if the nationwide models were calibrated with a small number of regional sample plots. Although forest structures differ among different parts of Finland, the nationwide volume and biomass models performed quite well (leave-inventory-area-out RMSE 22.3% to 33.8%, mean difference [MD] –13.8% to 18.7%) compared with regional models (leave-plot-out RMSE 20.2% to 26.8%). However, the nationwide dominant height model (RMSE 5.4% to 7.7%, MD –2.0% to 2.8%, with the exception of the Tornio region – RMSE 11.4%, MD –9.1%) performed nearly as well as the regional models (RMSE 5.2% to 6.7%). The results show that the nationwide volume and biomass models provided different means than real means at regional level, because forest structure and ALS device have a considerable effect on the predictions. Large MDs appeared especially in northern Finland. Local calibration decreased the MD and RMSE of volume and biomass models. However, the nationwide dominant height model did not benefit much from calibration.

  • Kotivuori, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: eetu.kotivuori@uef.fi (email)
  • Korhonen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: lauri.korhonen@uef.fi
  • Packalen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: petteri.packalen@uef.fi
article id 1568, category Research article
Jouni Siipilehto, Harri Lindeman, Mikko Vastaranta, Xiaowei Yu, Jori Uusitalo. (2016). Reliability of the predicted stand structure for clear-cut stands using optional methods: airborne laser scanning-based methods, smartphone-based forest inventory application Trestima and pre-harvest measurement tool EMO. Silva Fennica vol. 50 no. 3 article id 1568. https://doi.org/10.14214/sf.1568
Highlights: An airborne laser scanning grid-based approach for determining stand structure enabled bi- or multimodal predicted distributions that fitted well to the ground-truth harvester data; EMO and Trestima applications needed stand-specific inventory for sample measurements or sample photos, respectively, and at their best, provided superior accuracy for predicting certain stand characteristics.

Accurate timber assortment information is required before cuttings to optimize wood allocation and logging activities. Timber assortments can be derived from diameter-height distribution that is most often predicted from the stand characteristics provided by forest inventory. The aim of this study was to assess and compare the accuracy of three different pre-harvest inventory methods in predicting the structure of mainly Scots pine-dominated, clear-cut stands. The investigated methods were an area-based approach (ABA) based on airborne laser scanning data, the smartphone-based forest inventory Trestima app and the more conventional pre-harvest inventory method called EMO. The estimates of diameter-height distributions based on each method were compared to accurate tree taper data measured and registered by the harvester’s measurement systems during the final cut. According to our results, grid-level ABA and Trestima were generally the most accurate methods for predicting diameter-height distribution. ABA provides predictions for systematic 16 m × 16 m grids from which stand-wise characteristics are aggregated. In order to enable multimodal stand-wise distributions, distributions must be predicted for each grid cell and then aggregated for the stand level, instead of predicting a distribution from the aggregated stand-level characteristics. Trestima required a sufficient sample for reliable results. EMO provided accurate results for the dominating Scots pine but, it could not capture minor admixtures. ABA seemed rather trustworthy in predicting stand characteristics and diameter distribution of standing trees prior to harvesting. Therefore, if up-to-date ABA information is available, only limited benefits can be obtained from stand-specific inventory using Trestima or EMO in mature pine or spruce-dominated forests.

  • Siipilehto, Natural Research Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 18, FI-01301 Vantaa, Finland ORCID ID:E-mail: jouni.siipilehto@luke.fi (email)
  • Lindeman,  Natural Research Institute Finland, Green Technology, Kaironiementie 15, 39700 Parkano ORCID ID:E-mail: harri.lindeman@luke.fi
  • Vastaranta, University of Helsinki, Department of Forest Sciences, P.O. Box 62 (Viikinkaari 11), FI-00014 University of Helsinki ORCID ID:E-mail: mikko.vastaranta@helsinki.fi
  • Yu, Finnish Geospatial Research Institute (FGI), Department of Remote Sensing and Photogrammetry, National Land Survey of Finland, P.O. Box 15 (Geodeetinrinne 2), FI-02431, Masala, Finland ORCID ID:E-mail: xiaowei.yu@maanmittauslaitos.fi
  • Uusitalo,  Natural Research Institute Finland, Green Technology, Kaironiementie 15, 39700 Parkano ORCID ID:E-mail: jori.uusitalo@luke.fi
article id 1546, category Research article
Jussi Manner, Lauri Palmroth, Tomas Nordfjell, Ola Lindroos. (2016). Load level forwarding work element analysis based on automatic follow-up data. Silva Fennica vol. 50 no. 3 article id 1546. https://doi.org/10.14214/sf.1546
Highlights: Recent developments in on-board technology enables automatic collection of follow-up data on forwarder work; Time consumption per load was more strongly associated with Loading drive distance than with extraction distance, indicating that the relevance of extraction distance as a main indicator of forwarding productivity should be re-considered; Data, within variables, were positively skewed with a few exceptions with normal distributions.

Recent developments in on-board technology have enabled automatic collection of follow-up data on forwarder work. The objective of this study was to exploit this possibility to obtain highly representative information on time consumption of specific work elements (including overlapping crane work and driving), with one load as unit of observation, for large forwarders in final felling operations. The data used were collected by the John Deere TimberLink system as nine operators forwarded 8868 loads, in total, at sites in mid-Sweden. Load-sizes were not available. For the average and median extraction distances (219 and 174 m, respectively), Loading, Unloading, Driving empty, Driving loaded and Other time effective work (PM) accounted for ca. 45, 19, 8.5, 7.5 and 14% of total forwarding time consumption, respectively. The average and median total time consumptions were 45.8 and 42.1 minutes/load, respectively. The developed models explained large proportions of the variation of time consumption for the work elements Driving empty and Driving loaded, but minor proportions for the work elements Loading and Unloading. Based on the means, the crane was used during 74.8% of Loading PM time, the driving speed was nonzero during 31.9% of the Loading PM time, and Simultaneous crane work and driving occurred during 6.7% of the Loading PM time. Time consumption per load was more strongly associated with Loading drive distance than with extraction distance, indicating that the relevance of extraction distance as a main indicator of forwarding productivity should be re-considered.

  • Manner, The Forestry Research Institute of Sweden (Skogforsk), Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail: jussi.manner@skogforsk.se (email)
  • Palmroth, John Deere Forestry, P.O. Box 472, FI-33101 Tampere, Finland ORCID ID:E-mail: PalmrothLauri@JohnDeere.com
  • Nordfjell, Swedish University of Agricultural Sciences, Department of Forest Biomaterials and Technology, SE-901 83 Umeå, Sweden ORCID ID:E-mail: tomas.nordfjell@slu.se
  • Lindroos, Swedish University of Agricultural Sciences, Department of Forest Biomaterials and Technology, SE-901 83 Umeå, Sweden ORCID ID:E-mail: ola.lindroos@slu.se
article id 1514, category Research article
Heidi Hallongren, Ville Kankaanhuhta, Mikael Kukkonen. (2016). Cleaning Scots pine seedling stands with mechanical uprooters – a work quality comparison of two related devices. Silva Fennica vol. 50 no. 3 article id 1514. https://doi.org/10.14214/sf.1514
Highlights: The productivity of the narrower modified device was significantly better than the wider original device; Work quality did not differ significantly between devices when stand characteristics, regeneration success and pre-existing damage were taken into account; Results indicate that mechanical uprooting devices may be further developed to a cost-effective alternative to motor-manual techniques for the early cleaning of direct seeded commercial Scots pine stands.

Commercial forests require early cleaning to ensure the unhindered and uniform growth of crop trees. In order to be cost effective, non-crop vegetation should be uprooted to prevent their recovery. Performing this work manually is a labour-intensive task but it can be done mechanically. We evaluated the efficiency of two uprooting devices in direct seeded Scots pine (Pinus sylvestris L.) stands ca. 1 m tall. Productivity and quality of the uprooting work was investigated across eight stands and ca. 160 sample plots in northern Karelia, eastern Finland. Time consumption of the uprooters was analyzed through a linear regression model and the work quality through a multilevel multivariate model in terms of the number of individual Scots pine seedlings, processing units (i.e., a bunch of seedlings to be harvested in the future) and broadleaves. The productivity of the narrower modified device was significantly better in terms of time consumption than the wider original device. Work quality did not differ significantly between devices when stand characteristics, regeneration success and pre-existing damage were taken into account. Results indicate that mechanical uprooting devices may be further developed to a cost-effective alternative to motor-manual techniques for the early cleaning of direct seeded commercial Scots pine stands.

  • Hallongren, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: heidi.hallongren@luke.fi (email)
  • Kankaanhuhta, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: ville.kankaanhuhta@luke.fi
  • Kukkonen, University of Eastern Finland, School of Forest Sciences. P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: mikaelkukkonen@hotmail.com
article id 1492, category Research article
Li-Bin Liu, Yang-Yang Wu, Gang Hu, Zhong-Hua Zhang, An-Yun Cheng, Shi-Jie Wang, Jian Ni. (2016). Biomass of karst evergreen and deciduous broad-leaved mixed forest in central Guizhou province, southwestern China: a comprehensive inventory of a 2 ha plot. Silva Fennica vol. 50 no. 3 article id 1492. https://doi.org/10.14214/sf.1492
Highlights: Comprehensive inventory of the karst secondary forest based on a 2 ha large plot enhanced the reliability of biomass estimates; The biomass was 158.1 Mg ha−1, and the five dominant tree species accounted for 92.4% of aboveground tree biomass; The estimated necromass of woody debris and litter in the karst secondary forest was 17.6 Mg ha−1.

The biomass of a secondary evergreen and deciduous broad-leaved mixed forest was comprehensively inventoried in a permanent 2 ha plot in southwestern China. Biomass models, sub-sampling, soil pit method, and published data were utilized to determine the biomass of all components. Results showed that the total biomass of the forest was 158.1 Mg ha−1; the total biomass included the major aboveground (137.7 Mg ha−1) and belowground (20.3 Mg ha−1) biomass components of vascular plants as well as the minor biomass components of bryophytes (0.078 Mg ha−1) and lichens (0.043 Mg ha−1). The necromass was 17.6 Mg ha−1 and included woody debris (9.0 Mg ha−1) and litter (8.6 Mg ha−1). The spatial pattern of the aboveground biomass was determined by the spatial distribution of dominant trees with large diameter, tall height, and dense wood. The belowground biomass differed in terms of root diameter and decreased with increasing soil depth. The belowground biomass in each soil pit in local habitats was not related to the spatial distribution of woody plants and soil pit depth. The karst forest presented lower biomass compared than the nonkarst forests in the subtropical zone. Biomass carbon in the karst terrains would increase substantially if degraded karst vegetation could be successfully restored to the forest. Comprehensive site-based biomass inventory of karst vegetation will contribute not only to provide data for benchmarking global and regional vegetation and carbon models but also for regional carbon inventory and vegetation restoration.

  • Liu, State Key Laboratory of Environmental Geochemistry, Institute of Geochemistry, Chinese Academy of Sciences, Lincheng West Road 99, 550081 Guiyang, China; Puding Karst Ecosystem Research Station, Chinese Academy of Sciences, 562100 Puding, China; University of Chinese Academy of Sciences, 100049 Beijing, China ORCID ID:E-mail: liulibin@mail.gyig.ac.cn
  • Wu, State Key Laboratory of Environmental Geochemistry, Institute of Geochemistry, Chinese Academy of Sciences, Lincheng West Road 99, 550081 Guiyang, China; Puding Karst Ecosystem Research Station, Chinese Academy of Sciences, 562100 Puding, China; University of Chinese Academy of Sciences, 100049 Beijing, China ORCID ID:E-mail: wuyang2468@hotmail.com
  • Hu, School of Chemistry and Life Science, Guangxi Teachers Education University, 530001 Nanning, China; Key Laboratory of Beibu Gulf Environment Change and Resources Utilization of Ministry of Education, Guangxi Teachers Education University, 530001 Nanning, China ORCID ID:E-mail: ahhugang@gmail.com
  • Zhang, School of Chemistry and Life Science, Guangxi Teachers Education University, 530001 Nanning, China; Key Laboratory of Beibu Gulf Environment Change and Resources Utilization of Ministry of Education, Guangxi Teachers Education University, 530001 Nanning, China ORCID ID:E-mail: gxtczzh@gmail.com
  • Cheng, State Key Laboratory of Environmental Geochemistry, Institute of Geochemistry, Chinese Academy of Sciences, Lincheng West Road 99, 550081 Guiyang, China; Puding Karst Ecosystem Research Station, Chinese Academy of Sciences, 562100 Puding, China ORCID ID:E-mail: chenganyun@vip.skleg.cn
  • Wang, State Key Laboratory of Environmental Geochemistry, Institute of Geochemistry, Chinese Academy of Sciences, Lincheng West Road 99, 550081 Guiyang, China; Puding Karst Ecosystem Research Station, Chinese Academy of Sciences, 562100 Puding, China ORCID ID:E-mail: wangshijie@vip.skleg.cn
  • Ni, State Key Laboratory of Environmental Geochemistry, Institute of Geochemistry, Chinese Academy of Sciences, Lincheng West Road 99, 550081 Guiyang, China; Puding Karst Ecosystem Research Station, Chinese Academy of Sciences, 562100 Puding, China ORCID ID:E-mail: nijian@vip.skleg.cn (email)
article id 1492, category Research article
Li-Bin Liu, Yang-Yang Wu, Gang Hu, Zhong-Hua Zhang, An-Yun Cheng, Shi-Jie Wang, Jian Ni. (2016). Biomass of karst evergreen and deciduous broad-leaved mixed forest in central Guizhou province, southwestern China: a comprehensive inventory of a 2 ha plot. Silva Fennica vol. 50 no. 3 article id 1492. https://doi.org/10.14214/sf.1492
Highlights: Comprehensive inventory of the karst secondary forest based on a 2 ha large plot enhanced the reliability of biomass estimates; The biomass was 158.1 Mg ha−1, and the five dominant tree species accounted for 92.4% of aboveground tree biomass; The estimated necromass of woody debris and litter in the karst secondary forest was 17.6 Mg ha−1.

The biomass of a secondary evergreen and deciduous broad-leaved mixed forest was comprehensively inventoried in a permanent 2 ha plot in southwestern China. Biomass models, sub-sampling, soil pit method, and published data were utilized to determine the biomass of all components. Results showed that the total biomass of the forest was 158.1 Mg ha−1; the total biomass included the major aboveground (137.7 Mg ha−1) and belowground (20.3 Mg ha−1) biomass components of vascular plants as well as the minor biomass components of bryophytes (0.078 Mg ha−1) and lichens (0.043 Mg ha−1). The necromass was 17.6 Mg ha−1 and included woody debris (9.0 Mg ha−1) and litter (8.6 Mg ha−1). The spatial pattern of the aboveground biomass was determined by the spatial distribution of dominant trees with large diameter, tall height, and dense wood. The belowground biomass differed in terms of root diameter and decreased with increasing soil depth. The belowground biomass in each soil pit in local habitats was not related to the spatial distribution of woody plants and soil pit depth. The karst forest presented lower biomass compared than the nonkarst forests in the subtropical zone. Biomass carbon in the karst terrains would increase substantially if degraded karst vegetation could be successfully restored to the forest. Comprehensive site-based biomass inventory of karst vegetation will contribute not only to provide data for benchmarking global and regional vegetation and carbon models but also for regional carbon inventory and vegetation restoration.

  • Liu, State Key Laboratory of Environmental Geochemistry, Institute of Geochemistry, Chinese Academy of Sciences, Lincheng West Road 99, 550081 Guiyang, China; Puding Karst Ecosystem Research Station, Chinese Academy of Sciences, 562100 Puding, China; University of Chinese Academy of Sciences, 100049 Beijing, China ORCID ID:E-mail: liulibin@mail.gyig.ac.cn
  • Wu, State Key Laboratory of Environmental Geochemistry, Institute of Geochemistry, Chinese Academy of Sciences, Lincheng West Road 99, 550081 Guiyang, China; Puding Karst Ecosystem Research Station, Chinese Academy of Sciences, 562100 Puding, China; University of Chinese Academy of Sciences, 100049 Beijing, China ORCID ID:E-mail: wuyang2468@hotmail.com
  • Hu, School of Chemistry and Life Science, Guangxi Teachers Education University, 530001 Nanning, China; Key Laboratory of Beibu Gulf Environment Change and Resources Utilization of Ministry of Education, Guangxi Teachers Education University, 530001 Nanning, China ORCID ID:E-mail: ahhugang@gmail.com
  • Zhang, School of Chemistry and Life Science, Guangxi Teachers Education University, 530001 Nanning, China; Key Laboratory of Beibu Gulf Environment Change and Resources Utilization of Ministry of Education, Guangxi Teachers Education University, 530001 Nanning, China ORCID ID:E-mail: gxtczzh@gmail.com
  • Cheng, State Key Laboratory of Environmental Geochemistry, Institute of Geochemistry, Chinese Academy of Sciences, Lincheng West Road 99, 550081 Guiyang, China; Puding Karst Ecosystem Research Station, Chinese Academy of Sciences, 562100 Puding, China ORCID ID:E-mail: chenganyun@vip.skleg.cn
  • Wang, State Key Laboratory of Environmental Geochemistry, Institute of Geochemistry, Chinese Academy of Sciences, Lincheng West Road 99, 550081 Guiyang, China; Puding Karst Ecosystem Research Station, Chinese Academy of Sciences, 562100 Puding, China ORCID ID:E-mail: wangshijie@vip.skleg.cn
  • Ni, State Key Laboratory of Environmental Geochemistry, Institute of Geochemistry, Chinese Academy of Sciences, Lincheng West Road 99, 550081 Guiyang, China; Puding Karst Ecosystem Research Station, Chinese Academy of Sciences, 562100 Puding, China ORCID ID:E-mail: nijian@vip.skleg.cn (email)
article id 1441, category Research article
Dorota Zawadzka, Stanisław Drozdowski, Grzegorz Zawadzki, Jerzy Zawadzki. (2016). The availability of cavity trees along an age gradient in fresh pine forests. Silva Fennica vol. 50 no. 3 article id 1441. https://doi.org/10.14214/sf.1441
Highlights: The density of cavity trees in pine-dominated, managed forests varied in relation to stand age and was highest in stands older than 130 years of age; Cavities excavated by woodpeckers dominated among all cavities; The number of trees with cavities appears insufficient to ensure the effective protection of bird diversity in managed stands of Augustów Forest.

Given their importance as a resource for many forest organisms, tree cavities were inventoried in the managed pine forests of north-east Poland, in relation to the: 70–100, 101–130 and >130 year age-classes within the clear-cutting system. The densities at which cavities were present was found to depend on forest age, given that stands 70–100 years old were characterised by an average density of 0.62 trees ha–1, while forests older than 130 years reported 3.28 trees ha–1. Stands aged 70–100 years differed from those aged 130+ in having just 0.27 trees ha–1 of cavity trees, as compared with 2.91 trees ha–1. The total volume of cavity trees in stands up to 100 years old was 0.37 m3 ha–1 on average, as compared with 5.42 m3 ha–1 in stands over 130 years old. The cavities created by woodpeckers constituted 76% of all of those found, and included 53% excavated by great spotted woodpeckers (Dendrocopos major L.) and 23% by black woodpeckers (Dryocopus martius L.) The proportion of cavities excavated by D. major was highest in the youngest age class of stands. There, cavities made by D. martius constituted only 8% of the total, as compared with 31% in the oldest stands. The abundance of cavity trees thus differed along an age gradient, though in any event the availability of cavity trees appears to be too limited to provide for the needs of hole-nesting birds. Forest managers must thus take more account than hitherto of the need to protect cavity trees.

  • Zawadzka, Institute of Forest Science, University of Łódź, Branch in Tomaszów Mazowiecki, Konstytucji 3 Maja 65/67, 97-200 Tomaszów Mazowiecki, Poland ORCID ID:E-mail: dorota_zaw@wp.pl
  • Drozdowski, Department of Silviculture, Warsaw University of Life Sciences - SGGW, Nowoursynowska 159, 02-776 Warsaw, Poland ORCID ID:E-mail: stanislaw_drozdowski@sggw.pl (email)
  • Zawadzki, Eagle Conservation Committee, Okółek 14, 16-506 Giby, Poland ORCID ID:E-mail: grzesiekgfz@op.pl
  • Zawadzki, Eagle Conservation Committee, Okółek 14, 16-506 Giby, Poland ORCID ID:E-mail: jerzy_zaw@wp.pl
article id 1495, category Research article
Per-Ola Olsson, Tuula Kantola, Päivi Lyytikäinen-Saarenmaa, Anna Maria Jönsson, Lars Eklundh. (2016). Development of a method for monitoring of insect induced forest defoliation – limitation of MODIS data in Fennoscandian forest landscapes. Silva Fennica vol. 50 no. 2 article id 1495. https://doi.org/10.14214/sf.1495
Highlights: We developed and tested a method to monitor insect induced defoliation in forests based on coarse-resolution satellite data (MODIS); MODIS data may fail to detect defoliation in fragmented landscapes, especially if defoliation history is long. More homogenous forests results in higher detection accuracies; The method may be applied to future coarse and medium-resolution satellite data with high temporal resolution.

We investigated if coarse-resolution satellite data from the MODIS sensor can be used for regional monitoring of insect disturbances in Fennoscandia. A damage detection method based on z-scores of seasonal maximums of the 2-band Enhanced Vegetation Index (EVI2) was developed. Time-series smoothing was applied and Receiver Operating Characteristics graphs were used for optimisation. The method was developed in fragmented and heavily managed forests in eastern Finland dominated by Scots pine (Pinus sylvestris L.) (pinaceae) and with defoliation of European pine sawfly (Neodiprion sertifer Geoffr.) (Hymenoptera: Diprionidae) and common pine sawfly (Diprion pini L.) (Hymenoptera: Diprionidae). The method was also applied to subalpine mountain birch (Betula pubescens ssp. Czerepanovii N.I. Orlova) forests in northern Sweden, infested by autumnal moth (Epirrita autumnata Borkhausen) and winter moth (Operophtera brumata L.). In Finland, detection accuracies were fairly low with 50% of the damaged stands detected, and a misclassification of healthy stands of 22%. In areas with long outbreak histories the method resulted in extensive misclassification. In northern Sweden accuracies were higher, with 75% of the damage detected and a misclassification of healthy samples of 19%. Our results indicate that MODIS data may fail to detect damage in fragmented forests, particularly when the damage history is long. Therefore, regional studies based on these data may underestimate defoliation. However, the method yielded accurate results in homogeneous forest ecosystems and when long-enough periods without damage could be identified. Furthermore, the method is likely to be useful for insect disturbance detection using future medium-resolution data, e.g. from Sentinel‑2.

  • Olsson, Lund University, Department of Physical Geography and Ecosystem Science, Sölvegatan 12, S-223 62 Lund, Sweden ORCID ID:E-mail: per-ola.olsson@nateko.lu.se (email)
  • Kantola, Texas A & M University, Knowledge Engineering Laboratory, Department of Entomology, College Station, TX, USA ORCID ID:E-mail: tuula.kantola@helsinki.fi
  • Lyytikäinen-Saarenmaa, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID ID:E-mail: paivi.lyytikainen-saarenmaa@helsinki.fi
  • Jönsson, Lund University, Department of Physical Geography and Ecosystem Science, Sölvegatan 12, S-223 62 Lund, Sweden ORCID ID:E-mail: anna_maria.jonsson@nateko.lu.se
  • Eklundh, Lund University, Department of Physical Geography and Ecosystem Science, Sölvegatan 12, S-223 62 Lund, Sweden ORCID ID:E-mail: lars.eklundh@nateko.lu.se
article id 1462, category Research article
Pekka Punttila, Olli Autio, Janne S. Kotiaho, D. Johan Kotze, Olli J. Loukola, Norbertas Noreika, Anna Vuori, Kari Vepsäläinen. (2016). The effects of drainage and restoration of pine mires on habitat structure, vegetation and ants. Silva Fennica vol. 50 no. 2 article id 1462. https://doi.org/10.14214/sf.1462
Highlights: Mire drainage shifted floristic composition and ant assemblages towards forest communities; Raising the water-table level by ditch filling and the thinning of trees affected mire communities positively already 1–3 years after the start of restoration; The extent of tree cover, the coverage of Sphagnum mosses and the water-table level were major determinants of ant assemblage structure.

Habitat loss and degradation are the main threats to biodiversity worldwide. For example, nearly 80% of peatlands in southern Finland have been drained. There is thus a need to safeguard the remaining pristine mires and to restore degraded ones. Ants play a pivotal role in many ecosystems and like many keystone plant species, shape ecosystem conditions for other biota. The effects of mire restoration and subsequent vegetation succession on ants, however, are poorly understood. We inventoried tree stands, vegetation, water-table level, and ants (with pitfall traps) in nine mires in southern Finland to explore differences in habitats, vegetation and ant assemblages among pristine, drained (30–40 years ago) and recently restored (1–3 years ago) pine mires. We expected that restoring the water-table level by ditch filling and reconstructing sparse tree stands by cuttings will recover mire vegetation and ants. We found predictable responses in habitat structure, floristic composition and ant assemblage structure both to drainage and restoration. However, for mire-specialist ants the results were variable and longer-term monitoring is needed to confirm the success of restoration since these social insects establish perennial colonies with long colony cycles. We conclude that restoring the water-table level and tree stand structure seem to recover the characteristic vegetation and ant assemblages in the short term. This recovery was likely enhanced because drained mires still had both acrotelm and catotelm, and connectedness was still reasonable for mire organisms to recolonize the restored mires either from local refugia or from populations of nearby mires.

  • Punttila, Finnish Environment Institute, P.O. Box 140, FI-00251 Helsinki, Finland ORCID ID:E-mail: pekka.punttila@ymparisto.fi (email)
  • Autio, Centre for Economic Development, Transport and the Environment in South Ostrobothnia, P.O. Box 252, FI-65101 Vaasa, Finland ORCID ID:E-mail: olli.autio@ely-keskus.fi
  • Kotiaho, University of Jyväskylä, Department of Biology & Environmental Sciences, P.O. Box 35, FI-40014 Jyväskylä, Finland ORCID ID:E-mail: janne.kotiaho@jyu.fi
  • Kotze, University of Helsinki, Department of Environmental Sciences, P.O. Box 65, FI-00014, University of Helsinki, Finland ORCID ID:E-mail: johan.kotze@helsinki.fi
  • Loukola, University of Oulu, Department of Biology, P.O. Box 3000, FI-90014 Oulu, Finland ORCID ID:E-mail: olli.loukola@gmail.com
  • Noreika, University of Helsinki, Department of Environmental Sciences, P.O. Box 65, FI-00014, University of Helsinki, Finland; University of Helsinki, Department of Biosciences, P.O. Box 65, FI-00014 University of Helsinki, Finland ORCID ID:E-mail: norbertas.noreika@gmail.com
  • Vuori, University of Jyväskylä, Department of Biology & Environmental Sciences, P.O. Box 35, FI-40014 Jyväskylä, Finland ORCID ID:E-mail: anna@kureniemi.fi
  • Vepsäläinen, University of Helsinki, Department of Biosciences, P.O. Box 65, FI-00014 University of Helsinki, Finland ORCID ID:E-mail: kari.vepsalainen@helsinki.fi
article id 1448, category Research article
Andrew McEwan, Natascia Magagnotti, Raffaele Spinelli. (2016). The effects of number of stems per stool on cutting productivity in coppiced Eucalyptus plantations. Silva Fennica vol. 50 no. 2 article id 1448. https://doi.org/10.14214/sf.1448
Highlights: Double- and single stem coppice stools were harvested mechanically; Stem size had the strongest impact on productivity; An experienced operator performed equally well with both stool treatments; Cost was ~10% higher with double stems for the less experienced operator; Operator experience may play a major role when cutting coppice stands.

A time study was conducted to determine whether stem crowding had any impact on harvester productivity in Eucalyptus grandis stands. This represents an important element when trying to balance the advantages and disadvantages of coppice management in fast growing plantations designated for mechanized harvesting (i.e. machine felling, delimbing, debarking and cross-cutting). The study material consisted of 446 coppice stems, half of which grew as single stems per stool and half as double stems per stool as a result of different coppice reduction strategies. The dataset was balanced and randomized, with both subsets replicating exactly the same stem size distribution and the single and double stems alternating randomly. Harvester productivity ranged between 6 and 50 m3 under bark per productive machine hour, following the variation of tree diameter from 10 to 40 cm at breast height (1.37 m according to South African standards). Regression analysis indicated that both tree size and stem crowding (e.g. one or two stems per stool) had a significant effect on harvester productivity, which increased with stem size and decreased with stem crowding. However, operator experience may overcome the effect of stem crowding, which was not significant when the harvester was manned by a highly experienced operator. In any case, the effect of stem size was much greater than that of stem crowding, which resulted in a cost difference of less than 10%. However, this figure excludes the possible effects of stem crowding on volume recovery and stem development, which should be addressed in the future.

  • McEwan, Nelson Mandela Metropolitan University – George Campus, Saasveld, 6529, George, South Africa ORCID ID:E-mail: Andrew.McEwan@nmmu.ac.za
  • Magagnotti, CNR IVALSA, Via Madonna del Piano 10, I-50019 Sesto Fiorentino (FI), Italy ORCID ID:E-mail: magagnotti@ivalsa.cnr.it
  • Spinelli, CNR IVALSA, Via Madonna del Piano 10, I-50019 Sesto Fiorentino (FI), Italy ORCID ID:E-mail: spinelli@ivalsa.cnr.it (email)
article id 1414, category Research article
Rami Saad, Jörgen Wallerman, Johan Holmgren, Tomas Lämås. (2016). Local pivotal method sampling design combined with micro stands utilizing airborne laser scanning data in a long term forest management planning setting. Silva Fennica vol. 50 no. 2 article id 1414. https://doi.org/10.14214/sf.1414
Highlights: Most similar neighbor imputation was used to estimate forest variables using airborne laser scanning data as auxiliary data; For selecting field reference plots the local pivotal method (LPM) was compared to systematic sampling design; The LPM sampling design combined with a micro stand approach showed potential for improvement and has the potential to be a competitive method when considering cost efficiency.

A new sampling design, the local pivotal method (LPM), was combined with the micro stand approach and compared with the traditional systematic sampling design for estimation of forest stand variables. The LPM uses the distance between units in an auxiliary space – in this case airborne laser scanning (ALS) data – to obtain a well-spread sample. Two sets of reference plots were acquired by the two sampling designs and used for imputing data to evaluation plots. The first set of reference plots, acquired by LPM, made up four imputation alternatives (varying number of reference plots) and the second set of reference plots, acquired by systematic sampling design, made up two alternatives (varying plot radius). The forest variables in these alternatives were estimated using the nonparametric method of most similar neighbor imputation, with the ALS data used as auxiliary data. The relative root mean square error (RelRMSE), stem diameter distribution error index and suboptimal loss were calculated for each alternative, but the results showed that neither sampling design, i.e. LPM vs. systematic, offered clear advantages over the other. It is likely that the obtained results were a consequence of the small evaluation dataset used in the study (n = 30). Nevertheless, the LPM sampling design combined with the micro stand approach showed potential for improvement and might be a competitive method when considering the cost efficiency.

  • Saad, Swedish University of Agricultural Sciences (SLU), Department of Forest Resource Management, Skogsmarksgränd, SE-901 83 Umeå, Sweden ORCID ID:E-mail: rami.saad@slu.se (email)
  • Wallerman, Swedish University of Agricultural Sciences (SLU), Department of Forest Resource Management, Skogsmarksgränd, SE-901 83 Umeå, Sweden ORCID ID:E-mail: jorgen.wallerman@slu.se
  • Holmgren, Swedish University of Agricultural Sciences (SLU), Department of Forest Resource Management, Skogsmarksgränd, SE-901 83 Umeå, Sweden ORCID ID:E-mail: johan.holmgren@slu.se
  • Lämås, Swedish University of Agricultural Sciences (SLU), Department of Forest Resource Management, Skogsmarksgränd, SE-901 83 Umeå, Sweden ORCID ID:E-mail: tomas.lamas@slu.se
article id 1410, category Research article
Seppo Nevalainen, Juho Matala, Kari T. Korhonen, Antti Ihalainen, Ari Nikula. (2016). Moose damage in National Forest Inventories (1986–2008) in Finland. Silva Fennica vol. 50 no. 2 article id 1410. https://doi.org/10.14214/sf.1410
Highlights: Almost 100 000 stands were studied; The proportion of damage doubled during the study period; Tree species mixture had a clear effect on the damage frequency; The damage was more common in mineral soils than in peatlands, in artificially than in naturally regenerated stands and in stands that needed thinning or clearing or in which soil preparation was used.

The occurrence of moose damage was studied using data from three National Forest Inventories (NFIs) accomplished between 1986 and 2008 in Finland. The combined data included a total of 97 390 young stands. The proportion of moose damage increased from 3.6% to 8.6% between the 8th NFI (1986–1994) and the 10th NFI (2004–2008). The majority (75%) of the damage occurred in Scots pine-dominated stands. The proportion of damage was higher in aspen-dominated stands than in stands dominated by any other tree species. The tree species mixture also had a clear effect on the occurrence of damage. Pure Scots pine stands had less damage than mixed Scots pine stands, and moose damage decreased linearly with the increasing proportion of Scots pine. Stands on mineral soil had more frequent moose damage than stands on peatlands. The fertility class of the site had no straightforward effect on the damage frequency. Artificially regenerated stands had more damage than naturally regenerated stands. Accomplished soil preparation measures and the need for thinning or clearing operations increased moose damage. High proportions of moose damage in young stands were found around the country. In the 10th NFI, the largest concentration of damage was found in southwestern Finland. Our study shows the temporal and spatial changes in the occurrence of moose damage and pinpoints some important silvicultural factors affecting the relative risk of young stands over a large geographical area.

  • Nevalainen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail: seppo.nevalainen@luke.fi (email)
  • Matala, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail: juho.matala@luke.fi
  • Korhonen, Natural Resources Institute Finland (Luke), Economics and society, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail: kari.t.korhonen@luke.fi
  • Ihalainen, Natural Resources Institute Finland (Luke), Economics and society, P.O. Box 18, FI-01301 Vantaa, Finland ORCID ID:E-mail: antti.ihalainen@luke.fi
  • Nikula, Natural Resources Institute Finland (Luke), Economics and society, P.O. Box 16, FI-96301 Rovaniemi, Finland ORCID ID:E-mail: ari.nikula@luke.fi
article id 1323, category Research article
Tiina Laine, Kalle Kärhä, Antti Hynönen. (2016). A survey of the Finnish mechanized tree-planting industry in 2013 and its success factors. Silva Fennica vol. 50 no. 2 article id 1323. https://doi.org/10.14214/sf.1323
Highlights: In 2013, 31 planting machines were operated by 22 businesses and planted 4.7 million seedlings on 2663 hectares in Finland; Critical success factors included expertise of planting machine operators, high quality planting, adequate amount of work, stoniness, and removal of slash; Growth of the industry will depend on improved cost-efficiency, appropriate worksites, marketing, development of planting machines.

The aim of the study was to update the information pertaining to mechanized tree-planting activity in Finland in 2013 and its success factors. All businesses providing a mechanized tree-planting service were interviewed and asked to describe their equipment and activities, identify critical success factors (CSFs), and suggest areas for improvement. In 2013, 31 planting machines (18 Bracke P11.a, 11 M-Planter and 2 Risutec) operated by 22 businesses planted approximately 4.7 million seedlings on 2663 hectares. CSFs included expertise of planting machine operators, high quality planting, adequate amount of work, stoniness, and removal of slash. Based on the survey, some recommendations for improving mechanized planting work can be made. Firstly, improving the cost-efficiency of mechanized planting is necessary to enhance businesses’ profitability. Secondly, worksite selection is crucial as stoniness, stumps and slash debris diminish productivity. Lastly, the popularity of mechanized planting in the future will benefit from more marketing. Many businesses were unwilling to increase the area of service, invest in new equipment, or increase the volume of planting work but they believed that mechanized planting will become more popular in the near future.

  • Laine, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: tiina.laine@luke.fi (email)
  • Kärhä, Stora Enso Wood Supply Finland, P.O. Box 309, FI-00101 Helsinki, Finland ORCID ID:E-mail: kalle.karha@storaenso.com
  • Hynönen, University of Eastern Finland, Faculty of Science and Forestry, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: hynonena@gmail.com
article id 1406, category Research article
Tatiana V. Stankova. (2016). A dynamic whole-stand growth model, derived from allometric relationships. Silva Fennica vol. 50 no. 1 article id 1406. https://doi.org/10.14214/sf.1406
Highlights: A dynamic whole-stand model was derived from simple allometries and biological rationale; The state-space modelling approach was applied, suggesting several novelties to overcome scarcity of longitudinal data; The model consists of a three-dimensional state vector, defined by dominant stand height, stand density and mean stem volume, and three transition functions; It was tested with data from Scots pine plantations.

Growth and yield tables are constrained by a standard production regime and the stand-level dynamic models are an attractive alternative for the even-aged monospecific stands. The objective of this study is to derive a parsimonious and biologically sound whole-stand dynamic growth model and to validate its consistency and relevance for prediction of stand growth and yield. The state-space modelling approach was employed, introducing several novelties in comparison with its current usage. The model consists of a three-dimensional state vector, defined by dominant stand height, number of trees per hectare and mean stem volume, and three transition functions. A site index model was incorporated for height growth projection and transition functions for stand density and mean stem volume with respect to dominant height increase were derived from simple allometries and biological rationale. The model was examined with data from 79 permanent sample plots in Scots pine plantations. Nonlinear Seemingly Unrelated Regression was used to account for cross-equation correlations, and the Base-Height-Invariant dummy variable method was employed to estimate dynamic-form equations. Model consistency was validated in terms of accuracy of predictions and applicability to both thinned and unthinned stands. The new dynamic growth model is a parsimonious biometric whole-stand model that simulates the expected stand development for a broad spectrum of potential management alternatives and can be incorporated in a computer program for further use. It may be especially useful for application when a scarcity of longitudinal data prevents the use of more complicated modelling approaches.

  • Stankova, Forest Research Institute of Bulgarian Academy of Sciences, Department of Forest Genetics, Physiology and Plantations, 132 “St. Kliment Ohridski” Blvd., 1756 Sofia, Bulgaria ORCID ID: http://orcid.org/0000-0002-9932-7286 E-mail: tatianastankova@yahoo.com (email)
article id 1341, category Research article
Přemysl Humplík, Petr Čermák, Tomáš Žid. (2016). Electrical impedance tomography for decay diagnostics of Norway spruce (Picea abies): possibilities and opportunities. Silva Fennica vol. 50 no. 1 article id 1341. https://doi.org/10.14214/sf.1341
Highlights: Statistical parameters of EIT datasets with values of electrical resistance of heartwood are possible to employ in refining heartwood rot diagnostics; Sapwood proportion is decreasing as the proportion of decay on the radial cut expands; Using EIT datasets and sapwood proportion, trees with rot can be split into two groups as per proportion of decay: [< 35%] and [> 35%].

The paper aimed at testing the potential of refining tree rot diagnostics carried out by means of electrical impedance tomography (EIT). Examined was the use of EIT datasets with electrical resistance values and sapwood proportion determined on the basis of tomograms. Making use of datasets with resistance values in EIT rot diagnostics is not a default method, although datasets stay unaffected by a fixed colour scale and subsequent subjective evaluation unlike tomograms. Tomography measurement was carried out for 27 individuals of Norway spruce (Picea abies [L.] Karst.) in two stands north-east of Brno, Czech Republic. Once felled down, radial cut-outs were sampled at the measurement site and used for localising rot and determining the extent of the area of decay. The results were subsequently compared with tomograms. EIT datasets containing values of electrical resistance found by measuring were statistically processed and compared with the extent of rot area identified within the cuts. Sapwood proportion values were also detected using the tomograms. The baseline assumption that sapwood proportion decreases as the rot area in the radial cut expands was confirmed. In trees with rot percentage to 35% approximately, sapwood proportion was exceeding 30% except one tree. In trees with rot percentage exceeding 35%, sapwood proportion was below 30%. On the basis of interpreted datasets, the trees can be split into three characteristic groups that correspond to the occurrence, extent and nature of the rot.

  • Humplík, Mendel University in Brno, Faculty of Forestry and Wood Technology, Zemědělská 1665/3, 613 00 Brno, Czech Republic ORCID ID:E-mail: premysl.humplik@mendelu.cz (email)
  • Čermák, Mendel University in Brno, Faculty of Forestry and Wood Technology, Zemědělská 1665/3, 613 00 Brno, Czech Republic ORCID ID:E-mail: petr.cermak@mendelu.cz
  • Žid, Mendel University in Brno, Faculty of Forestry and Wood Technology, Zemědělská 1665/3, 613 00 Brno, Czech Republic ORCID ID:E-mail: tomas.zid@mendelu.cz
article id 1403, category Research article
Kristina Mjöfors, Monika Strömgren, Hans-Örjan Nohrstedt, Annemieke Ingrid Gärdenäs. (2015). Impact of site-preparation on soil-surface CO2 fluxes and litter decomposition in a clear-cut in Sweden. Silva Fennica vol. 49 no. 5 article id 1403. https://doi.org/10.14214/sf.1403
Highlights: Disturbances of the soil did not lead to higher CO2 emissions from the soil; Heavy mixing of the soil lead to lower CO2 emissions from the soil; Buried needles and coarse roots decomposed faster than those on the surface; Abundance of δ15N decreased in needles and roots after site preparation.

Boreal forest soil contains significant amounts of organic carbon. Soil disturbance, caused for example by site preparation or stump extraction, may increase decomposition and thus lead to higher CO2 emissions, contributing to global warming. The aim of this study was to quantify responses of soil-surface CO2 fluxes (Rs) and litter (needle and root) decomposition rates following various kinds of soil disturbance commonly caused by mechanical site preparation and stump harvest. For this purpose four treatments were applied in a clear-cut site in central Sweden: i) removal of the humus layer and top 2 cm of mineral soil, ii) placement of a humus layer and 2 cm of mineral soil upside down on top of undisturbed soil, forming a double humus layer buried under mineral soil, iii) heavy mixing of the humus layer and mineral soil, and iv) no disturbance (control). Rs measurements were acquired with a portable respiration system during two growing seasons. To assess the treatments’ effects on litter decomposition rates, needles or coarse roots (Ø = 6 mm) were incubated in litterbags at positions they would be located after the treatments (buried, or on top of the soil). The results indicate that site preparation-simulating treatments have no effect or may significantly reduce, rather than increase, CO2 emissions during the following two years. They also show that buried litter decomposes more rapidly than litter on the surface, but in other respects the treatments have little effect on litter decomposition rates.

  • Mjöfors, Swedish University of Agricultural Sciences (SLU), Department of Soil and Environment, P.O. Box 7014, 150 07 Uppsala, Sweden ORCID ID:E-mail: kristina.mjofors@slu.se (email)
  • Strömgren, Swedish University of Agricultural Sciences (SLU), Department of Soil and Environment, P.O. Box 7014, 150 07 Uppsala, Sweden ORCID ID:E-mail: Monika.stromgren@slu.se
  • Nohrstedt, Swedish University of Agricultural Sciences (SLU), Department of Soil and Environment, P.O. Box 7014, 150 07 Uppsala, Sweden ORCID ID:E-mail: Hans-orjan.nohrstedt@slu.se
  • Gärdenäs, Swedish University of Agricultural Sciences (SLU), Department of Soil and Environment, P.O. Box 7014, 150 07 Uppsala, Sweden ORCID ID:E-mail: Annemieke.gardenas@slu.se
article id 1403, category Research article
Kristina Mjöfors, Monika Strömgren, Hans-Örjan Nohrstedt, Annemieke Ingrid Gärdenäs. (2015). Impact of site-preparation on soil-surface CO2 fluxes and litter decomposition in a clear-cut in Sweden. Silva Fennica vol. 49 no. 5 article id 1403. https://doi.org/10.14214/sf.1403
Highlights: Disturbances of the soil did not lead to higher CO2 emissions from the soil; Heavy mixing of the soil lead to lower CO2 emissions from the soil; Buried needles and coarse roots decomposed faster than those on the surface; Abundance of δ15N decreased in needles and roots after site preparation.

Boreal forest soil contains significant amounts of organic carbon. Soil disturbance, caused for example by site preparation or stump extraction, may increase decomposition and thus lead to higher CO2 emissions, contributing to global warming. The aim of this study was to quantify responses of soil-surface CO2 fluxes (Rs) and litter (needle and root) decomposition rates following various kinds of soil disturbance commonly caused by mechanical site preparation and stump harvest. For this purpose four treatments were applied in a clear-cut site in central Sweden: i) removal of the humus layer and top 2 cm of mineral soil, ii) placement of a humus layer and 2 cm of mineral soil upside down on top of undisturbed soil, forming a double humus layer buried under mineral soil, iii) heavy mixing of the humus layer and mineral soil, and iv) no disturbance (control). Rs measurements were acquired with a portable respiration system during two growing seasons. To assess the treatments’ effects on litter decomposition rates, needles or coarse roots (Ø = 6 mm) were incubated in litterbags at positions they would be located after the treatments (buried, or on top of the soil). The results indicate that site preparation-simulating treatments have no effect or may significantly reduce, rather than increase, CO2 emissions during the following two years. They also show that buried litter decomposes more rapidly than litter on the surface, but in other respects the treatments have little effect on litter decomposition rates.

  • Mjöfors, Swedish University of Agricultural Sciences (SLU), Department of Soil and Environment, P.O. Box 7014, 150 07 Uppsala, Sweden ORCID ID:E-mail: kristina.mjofors@slu.se (email)
  • Strömgren, Swedish University of Agricultural Sciences (SLU), Department of Soil and Environment, P.O. Box 7014, 150 07 Uppsala, Sweden ORCID ID:E-mail: Monika.stromgren@slu.se
  • Nohrstedt, Swedish University of Agricultural Sciences (SLU), Department of Soil and Environment, P.O. Box 7014, 150 07 Uppsala, Sweden ORCID ID:E-mail: Hans-orjan.nohrstedt@slu.se
  • Gärdenäs, Swedish University of Agricultural Sciences (SLU), Department of Soil and Environment, P.O. Box 7014, 150 07 Uppsala, Sweden ORCID ID:E-mail: Annemieke.gardenas@slu.se
article id 1395, category Research article
Joseph Buongiorno. (2015). Income and time dependence of forest product demand elasticities and implications for forecasting. Silva Fennica vol. 49 no. 5 article id 1395. https://doi.org/10.14214/sf.1395
Highlights: Elasticities of demand with gross domestic product and prices were stable over time and income level for sawnwood and particleboard only; Other product elasticities differed with income and time, leading in conjunction with a sector model to higher projected world demand and prices than obtained by ignoring differences between countries and over time.

In view of improving multi-country forest sector models, this study investigated to what extent the price and income elasticity of demand for forest products had changed in the past two decades, and how much they depended on the countries income level. For each of seven major product groups annual observations were divided between high-income (top 20% in gross domestic product per capita) and low-income, and between recent (2004–2013) and older (1992–2003) observations. The results indicated that for sawnwood and particleboard the data could be pooled across all countries and years. For the other commodity groups (veneer & plywood, fiberboard, newsprint, printing & writing paper, other paper & paperboard), there were statistically significant differences in gross domestic product or price elasticity between high and low-income levels or old and recent observations. Efficient elasticities were obtained by pooling the maximum number of observations while respecting the statistically significant differences. The resulting GDP elasticities were the same, or very close, across income levels for all products. The price elasticities differed by income level only for newsprint and for veneer and plywood. International forest sector projections to 2065 obtained with these elasticities compared with those based on pooling all data across time and income levels gave less than 3% difference for world consumption of sawnwood, particleboard, fiberboard, and newsprint, but 19% higher consumption for veneer and plywood, 31% for printing and writing paper, and 18% for other paper and paperboard. The world price was 1% to 11% higher for end products and 3% to 22% higher for raw materials and intermediate products.

  • Buongiorno, University of Wisconsin-Madison, Department of Forest and Wildlife Ecology, 1630 Linden Drive, Madison, WI 53706, USA ORCID ID:E-mail: jbuongio@wisc.edu (email)
article id 1342, category Research article
Blas Mola-Yudego, Gianni Picchi, Dominik Röser, Raffaele Spinelli. (2015). Assessing chipper productivity and operator effects in forest biomass operations. Silva Fennica vol. 49 no. 5 article id 1342. https://doi.org/10.14214/sf.1342
Highlights: A model is constructed to assess the productivity in chipping of wood biomass at roadside; The data includes 172 trials and 67 operators in Italy; The operator effect was included in a mixed model approach; The R2 were 0.76 (fixed part) and 0.88 (incl. operator effects).

The present research focuses on the productivity of energy wood chipping operations at several sites in Italy. The aim was to assess the productivity and specifically the effect attributed to the operator in the chipping of wood biomass. The research included 172 trials involving 67 operators across the country that were analysed using a mixed model approach, in order to assess productivity, and to isolate the operator effect from other potential variables. The model was constructed using different predictors aiming to explain the variability due to the machines and the raw-materials. The final model included the average piece weight of raw material chipped as well as the power of the machine. The coefficients of determination (R2) were 0.76 for the fixed part of the model, and 0.88 when the effects due to the operators were included. The operators’ performance compared to their peers was established, and it was compared to a subjective classification based on the operator’s previous experience. The results of this study can help to the planning and logistics of raw material supply for bioenergy, as well as to a more effective training of future forest operators.

  • Mola-Yudego, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland; NIBIO Norwegian Institute of Bioeconomy Research, P.O. Box 115, 1431 Ås, Norway ORCID ID: http://orcid.org/0000-0003-0286-0170 E-mail: blas.mola@uef.fi (email)
  • Picchi, CNR IVALSA, Via Madonna del Piano 10, I-50019 Sesto Fiorentino, Italy ORCID ID:E-mail: picchi@ivalsa.cnr.it
  • Röser, Forest Feedstocks Group, FPInnovations, Vancouver, British Columbia, Canada ORCID ID:E-mail: dominik.roser@fpinnovations.ca
  • Spinelli, CNR IVALSA, Via Madonna del Piano 10, I-50019 Sesto Fiorentino, Italy ORCID ID:E-mail: spinelli@ivalsa.cnr.it
article id 1340, category Research article
Mostafa Farhadi, Mulualem Tigabu, Per Christer Odén. (2015). Near Infrared Spectroscopy as non-destructive method for sorting viable, petrified and empty seeds of Larix sibirica. Silva Fennica vol. 49 no. 5 article id 1340. https://doi.org/10.14214/sf.1340
Highlights: Near Infrared spectroscopy discriminates filled-viable, empty and petrified seeds of Larix sibirica with 98%, 82% and 87% accuracy, respectively based on spectral differences attributed to moisture and storage reserves; The classification accuracy reached 100% when sorting seeds into viable and non-viable class; The results demonstrate that NIR spectroscopy has great potential as non-destructive sorting technique to upgrade seed lot quality.

Larix sibirica Ledeb. is one of the promising timber species for planting in the boreal ecosystem; but poor seed lot quality is the major hurdle for production of sufficient quantity of planting stocks. Here, we evaluated the potential of Near Infrared (NIR) Spectroscopy for sorting viable and non-viable seeds, as the conventional sorting technique is inefficient. NIR reflectance spectra were collected from single seeds, and discriminant models were developed with Orthogonal Projections to Latent Structure – Discriminant Analysis (OPLS-DA). The computed model predicted the class membership of filled-viable, empty and petrified seeds in the test set with 98%, 82% and 87% accuracy, respectively. When two-class OPLS-DA model was fitted to discriminate viable and non-viable (empty and petrified seeds combined), the predicted class membership of test set samples was 100% for both classes. The origins of spectral differences between non-viable (petrified and empty) and viable seeds were attributed to differences in seed moisture content and storage reserves. In conclusion, the result provides evidence that NIR spectroscopy is a powerful non-destructive method for sorting non-viable seeds of Larix sibirica; thus efforts should be made to develop on-line sorting system for large-scale seed handling.

  • Farhadi, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53 Alnarp, Sweden ORCID ID:E-mail: mostafa.farhadi@slu.se
  • Tigabu, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53 Alnarp, Sweden ORCID ID:E-mail: mulualem.tigabu@slu.se (email)
  • Odén, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53 Alnarp, Sweden ORCID ID:E-mail: per.oden@slu.se
article id 1334, category Research article
Abolfazl Daneshvar, Mulualem Tigabu, Asaddollah Karimidoost, Per Christer Oden. (2015). Single seed Near Infrared Spectroscopy discriminates viable and non-viable seeds of Juniperus polycarpos. Silva Fennica vol. 49 no. 5 article id 1334. https://doi.org/10.14214/sf.1334
Highlights: Near Infrared (NIR) Spectroscopy discriminates viable and non-viable (empty, insect-attacked and shriveled) seeds of J. polycarpos with 98% and 100% accuracy, respectively; The origins of spectral differences between non-viable and viable seeds were attributed to differences in seed coat chemical composition and storage reserves; The results demonstrate that NIR spectroscopy has great potential as seed sorting technology to ensure precision sowing.

A large quantity of non-viable (empty, insect-attacked and shriveled) seeds of Juniperus polycarpos (K. Koch) is often encountered during seed collection, which should be removed from the seed lots to ensure precision sowing in the nursery or out in the field. The aims of this study were to evaluate different modelling approaches and to examine the sensitivity of the change in detection system (Silicon-detector in the shorter vis-a-vis InGsAs-detector in the longer NIR regions) for discriminating non-viable seeds from viable seeds by Near Infrared (NIR) spectroscopy. NIR reflectance spectra were collected from single seeds, and discriminant models were developed by Partial Least Squares – Discriminant Analysis (PLS-DA) and Orthogonal Projection to Latent Structures – Discriminant Analysis (OPLS-DA) using the entire or selected NIR regions. Both modelling approaches resulted in 98% and 100% classification accuracy for viable and non-viable seeds in the test set, respectively. However, OPLS-DA models were superb in terms of model parsimony and information quality. Modelling in the shorter and longer wavelength region also resulted in similar classification accuracy, suggesting that prediction of class membership is insensitive to change in the detection system. The origins of spectral differences between non-viable and viable seeds were attributed to differences in seed coat chemical composition, mainly terpenoids that were dominant in non-viable seeds and storage reserves in viable seeds. In conclusion, the results demonstrate that NIR spectroscopy has great potential as seed sorting technology to upgrade seed lot quality that ensures precision sowing.

  • Daneshvar, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53, Alnarp, Sweden; (permanent address) Department of Natural Resources, Gonbad Kavous University, Shahid Fallahi Street, P.O. Box 163, Gonbad, Iran ORCID ID:E-mail: abolfazl.daneshvar@slu.se
  • Tigabu, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53, Alnarp, Sweden ORCID ID:E-mail: mulualem.tigabu@slu.se (email)
  • Karimidoost, Agriculture and Natural Resources Research Center of Golestan Province, Beheshti Ave. P.O. Box 4915677555, Gorgan, Iran ORCID ID:E-mail: karimidoost@yahoo.com
  • Oden, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53, Alnarp, Sweden ORCID ID:E-mail: per.oden@slu.se
article id 1312, category Research article
Simon Karl Nils Berg, Tomas Nordfjell, Dan Bergström. (2015). Effect of stump size and timing of stump harvesting on ground disturbance and root breakage diameter. Silva Fennica vol. 49 no. 5 article id 1312. https://doi.org/10.14214/sf.1312
Highlights: The ground disturbance and root breakage diameter during conventional stump harvesting on mineral soil were quantified; A function for estimating the disturbed area based on stump size was constructed; Many fine roots were found to be harvested; The total ground disturbance at the site after stump harvesting was similar to that caused by soil scarification.

Stump wood is a possible alternative to fossil fuel. Its harvesting, however, disturbs the ground and this has not yet been quantified at stump level. Such disturbance is likely to be dependent on stump size, type of soil and timing of stump harvesting. Therefore, we measured ground disturbance and root breakage diameter at two Norway spruce sites with sandy glacial till soil. The sites were harvested with a fork type head, 6 and 18 months after clear cutting. Measurements were made within 2 weeks of harvest. No difference was found between the two sites. The mean area of disturbed ground was 6.06 (std 3.14) m2 per stump and increased exponentially with stump size. A regression function modelling the relationship was constructed. Unexpectedly, many fine roots where extracted in the harvest. The arithmetic and basal area weighted mean root breakage diameter was 4.6 (std 2.2) and 29.5 (std 17.9) mm, respectively. There seems to be a limited increase in root breakage diameter with increased stump size. The small root breakage diameter is associated with reduced fuel quality and greater nutrient removal. It appears that much of the ground disturbance is associated with the creation of ruts rather than stump harvest per se. Stump harvesting disturbs a larger percentage of the area of a harvested site than mounding. Postponing stump harvest by one year did not decrease the ground disturbance or increase the root breakage diameter. To achieve less disturbance and larger root breakage diameter, probably new stump harvesting technology is required.

  • Berg, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden ORCID ID:E-mail: phd.simon.berg@gmail.com (email)
  • Nordfjell, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden ORCID ID:E-mail: tomas.nordfjell@slu.se
  • Bergström, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden ORCID ID:E-mail: dan.bergstrom@slu.se
article id 1310, category Research article
Deliang Lu, Jiaojun Zhu, Yirong Sun, Lile Hu, Guangqi Zhang. (2015). Gap closure process by lateral extension growth of canopy trees and its effect on woody species regeneration in a temperate secondary forest, Northeast China. Silva Fennica vol. 49 no. 5 article id 1310. https://doi.org/10.14214/sf.1310
Highlights: Gap closure process by lateral extension growth can be described by quadratic functions; Large gaps (514–621 m2) had higher closure rates but lower closure percentages compared with middle (174–321 m2) and small gaps (68–125 m2); Gaps promoted woody species regeneration in early stage; Large and middle gaps would provide opportunities for filling regeneration, but regeneration in small gaps may eventually fail.

Gap formation and its effects on regeneration have been reported as being important in forest development, but seldom studies concentrated on the gap closure process by lateral extension growth of canopy trees surrounding gaps. We monitored the closure process of 12 artificial gaps for 7 years with three size classes: small (from 68 m2 to 125 m2), middle (from 174 m2 to 321 m2), and large (from 514 m2 to 621 m2); and investigated the regeneration twice in a temperate secondary forest, Northeast China. The closure process can be described through quadratic functions, which showed the closure rates slowed down with gap ages. Large gaps had a higher closure rate (39 m2 a–1) than middle gaps (25 m2 a–1) and small gaps (11 m2 a–1). According to the quadratic equations, the lateral growth could last 11, 13 and 16 years for small, middle and large gaps with a remaining size of 12, 69 and 223 m2, respectively. As expected, regeneration exhibited the highest seedling density and volume in large gaps. There was no significant difference in regeneration density between middle gaps, small gaps and forest understory in the final investigation; but the volume of regenerated woody species increased significantly from small gaps to large gaps compared with forest understory. These results may provide references on the choice of appropriate gap sizes to promote the regeneration in temperate secondary forests.

  • Lu, State Key Laboratory of Forest and Soil Ecology, Qingyuan Forest CERN, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110016, China;  University of Chinese Academy of Sciences, Beijing 100049, China ORCID ID:E-mail: delianglu14@hotmail.com
  • Zhu, State Key Laboratory of Forest and Soil Ecology, Qingyuan Forest CERN, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110016, China ORCID ID:E-mail: jiaojunzhu@iae.ac.cn (email)
  • Sun, State Key Laboratory of Forest and Soil Ecology, Qingyuan Forest CERN, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110016, China ORCID ID:E-mail: yirongsun@iae.ac.cn
  • Hu,  Chinese Research Academy of Environmental Sciences, Beijing 100012, China ORCID ID:E-mail: lilehu@gmail.com
  • Zhang, State Key Laboratory of Forest and Soil Ecology, Qingyuan Forest CERN, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110016, China;  University of Chinese Academy of Sciences, Beijing 100049, China ORCID ID:E-mail: zgq04713@163.com
article id 1391, category Research article
Roberts Matisons, Jānis Jansons, Juris Katrevičs, Āris Jansons. (2015). Relation of tree-ring width and earlywood vessel size of alien Quercus rubra L. with climatic factors in Latvia. Silva Fennica vol. 49 no. 4 article id 1391. https://doi.org/10.14214/sf.1391
Highlights: Climate-growth relationships of red oak from three sites in Latvia were studied; Tree-ring width was mainly affected by temperature and precipitation in late summer; Vessel size was correlated with temperature parameters in autumn–spring; Sets of climatic factors significant for growth of red oak differed between sites; Changes in climate-growth relationships occurred during 20th century.

The effect of climatic factors on wood anatomy of the alien red oak (Quercus rubra L.) growing in three experimental plantations in Latvia was assessed by classical dendrochronological techniques. Two tree-ring proxies – tree-ring width (TRW) and mean area of earlywood vessel lumen (VLA) – were studied on 33 trees. Annual variation of TRW amongst trees was similar (mean r = 0.46), but there was more individuality in VLA (mean r = 0.26); nevertheless, chronologies of both proxies had rather synchronous variation amongst the sites. Annual variation of TRW was affected by factors related to water deficit in late summer, as suggested by the negative effect of temperature and positive effect of precipitation that have intensified during the 20th century, likely due to warming. Although weather conditions during the dormant period did not directly affect TRW, temperature during the autumn-spring period has been the main climatic determinant of VLA likely via influence on overwintering and hence vigour of tree. This suggests that conductive properties of wood and hence the susceptibility to water deficit have been affected by weather conditions before the formation of tree rings. During the 20th century, sensitivity of VLA has shifted from temperature in winter to temperature in autumn likely due to climate change. Still, the positive effect of these factors suggests that warming of climate would increase VLA and hence the risk of embolism and xylem disfunction. Therefore, the importance of availability of water for growth of red oak in Latvia is increasing.

  • Matisons, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 ORCID ID:E-mail: robism@inbox.lv (email)
  • Jansons, Latvian Forest Competence Centre, Dzērbenes str. 27, Riga, Latvia, LV 1006 ORCID ID:E-mail: janis.jansons@silava.lv
  • Katrevičs, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 ORCID ID:E-mail: juris.katrevics@silava.lv
  • Jansons, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 ORCID ID:E-mail: aris.jansons@silava.lv
article id 1354, category Research article
Johannes Edvardsson, Anton Hansson. (2015). Multiannual hydrological responses in Scots pine radial growth within raised bogs in southern Sweden. Silva Fennica vol. 49 no. 4 article id 1354. https://doi.org/10.14214/sf.1354
Highlights: Annual growth of Scots pine (Pinus sylvestris L.) at Boreal raised bogs was found to reflect a synthesis of climate controlled moisture variability over the preceding one to four year period; Excessive soil moisture is a growth limiting factor for trees at raised bogs; River discharge data reflect hydrological conditions in peat bogs better than precipitation data.

To explore interactions between climate and peatland hydrology, Scots pine (Pinus sylvestris L.) growing at four raised bogs in southern Sweden were subject to a dendroclimatological study. Radial tree growth reflecting climate and water table fluctuations over multiannual periods was detected as significant negative correlations between tree-ring width (TRW) chronologies and the preceding one to four years total precipitation or river discharge. Systematically stronger negative correlations were obtained when river discharge instead of precipitation was compared to radial tree growth. This indicates that river discharge reflect moisture variability of peat bogs better than what precipitation data does. Meanwhile, monthly precipitation and radial tree growth did not show any clear correlation, whereas spring and early summer temperatures had a positive influence on the tree growth. Our study shows that growth variability of bog pines in the Boreal zone reflect hydrological responses related to a synthesis of climate controlled moisture variability over several year periods.

  • Edvardsson, Dendrolab.ch, Institute of Geological Sciences, University of Bern, Baltzerstrasse 1+3, CH-3012 Bern, Switzerland & Quaternary Sciences, Department of Geology, Lund University, Sölvegatan 12, SE-223 62 Lund, Sweden ORCID ID: http://orcid.org/0000-0001-5163-1599 E-mail: johannes.edvardsson@dendrolab.ch (email)
  • Hansson, Quaternary Sciences, Department of Geology, Lund University, Sölvegatan 12, SE-223 62 Lund, Sweden ORCID ID:E-mail: anton.hansson@geol.lu.se
article id 1283, category Research article
Ivana Bjedov, Dragica Obratov–Petković, Danijela Mišić, Branislav Šiler, Jelena M Aleksic. (2015). Genetic patterns in range-edge populations of Vaccinium species from the central Balkans: implications on conservation prospects and sustainable usage. Silva Fennica vol. 49 no. 4 article id 1283. https://doi.org/10.14214/sf.1283
Highlights: We studied fragmentary distributed range-edge populations of Vaccinium myrtillus, Vaccinium uliginosum and Vaccinium vitis-idaea from the Balkans using RAPDs; Low genetic diversities and high genetic differentiation were found in all species; The prevalence of clonal individuals was not observed; Past interspecific hybridization among V. vitis-idaea and the other two species was detected; Guidelines for conservation and sustainable usage were provided.

Vaccinium myrtillus L., Vaccinium uliginosum L. and Vaccinium vitis-idaea L. are perennial, cold-adapted clonal shrubs distributed throughout Europe, northern Asia and North America. Due to their usage in food (berries) and pharmaceutical industry (berries and leaves), their natural populations are exposed to anthropogenic and other impacts that affect their genetic make-up. We analyzed 14 fragmentary distributed and small-sized peripheral populations of these species from the Balkans, which represents the southeastern-European marginal area of their wide European distributions, using RAPD molecular markers. The contemporary genetic patterns in all three species within the Balkans were generally similar, and in comparison to previous reports on populations of these species found in northward Europe, where they have a more continuous distribution, the levels of genetic diversity were more or less halved, genetic differentiation was several times higher, gene flow exceptionally low, and the expected prevalence of clonal individuals was lacking. The population dynamics of all three species within the Balkans was complex and distinct, and was characterized by a past admixture of individuals from discrete populations of the same species and interspecific hybridisation not only between V. myrtillus and V. vitis-idaea but also between V. uliginosum and V. vitis-idaea, the latter not being reported to date. Conservation measures suitable for preservation of presumably genetically distinct portions of the Balkans’ gene pools of studied species have been suggested, while the utility of interspecific hybrids in breeding programs and/ or in food/pharmaceutical industry is yet to be assessed. 

  • Bjedov, University of Belgrade, Faculty of Forestry, Kneza Višeslava 1, 11000 Belgrade, Serbia ORCID ID:E-mail: ivana.bjedov@sfb.bg.ac.rs
  • Obratov–Petković, University of Belgrade, Faculty of Forestry, Kneza Višeslava 1, 11000 Belgrade, Serbia ORCID ID:E-mail: dragica.obratov-petkovic@sfb.bg.ac.rs
  • Mišić, University of Belgrade, Institute for Biological Research “Siniša Stanković”, Boulevard Despota Stefana 142, 11000 Belgrade, Serbia ORCID ID:E-mail: dmisic@ibiss.bg.ac.rs
  • Šiler, University of Belgrade, Institute for Biological Research “Siniša Stanković”, Boulevard Despota Stefana 142, 11000 Belgrade, Serbia ORCID ID:E-mail: branislav.siler@ibiss.bg.ac.rs
  • Aleksic, University of Belgrade, Institute of Molecular Genetics and Genetic Engineering, Vojvode Stepe 444a, P.O. Box 23, 11010 Belgrade, Serbia ORCID ID:E-mail: aleksic_jelena@yahoo.com.au (email)
article id 1236, category Research article
Monika Litkowiec, Beata P. Plitta-Michalak, Andrzej Lewandowski, Grzegorz Iszkuło. (2015). Homogenous genetic structure in populations of Taxus baccata with varied proportions of male and female individuals. Silva Fennica vol. 49 no. 4 article id 1236. https://doi.org/10.14214/sf.1236
Highlights: Polish populations of Taxus baccata showed a high level of genetic diversity within populations and moderate genetic differentiation between them after nSSR marker testing; No significant differences in the genetic variation between T. baccata male and female individuals were observed, and microsatellite loci neutrality was verified; Determining the sex ratio in T. baccata populations is not essential to develop a clear understanding of genetic differentiation and diversity within and between populations of this species.

English yew (Taxus baccata L.) is a strictly outcrossing and dioecious species whose populations are small and isolated. It is known that sex ratios may vary in natural populations due to local environmental conditions or stochastic events. However, unbalanced sex ratios may have negative impacts on genetic diversity through enhanced genetic drift and inbreeding. The present study represents one of the first attempts to compare the genetic variation at microsatellite loci within and between populations with different gender proportions. Our results indicated that there were no significant correlations between sex ratio and the extent of genetic variation in different populations. All populations exhibited high levels of genetic diversity. Additionally, the genetic structure was characterized separately in male and female individuals. Statistical analyses of the set estimators describing the genetic structure of male and female individuals of T. baccata revealed no significant differences between the two groups. Molecular analysis verified that microsatellite nuclear loci neutrality developed for T. baccata, as there were no significant differences in the genetic variation between males and females and no evidence for any outlier loci using coalescent and hierarchical Bayesian simulations. The results demonstrate that ignoring biased sex ratios in T. baccata populations had no effect on the assessment of genetic differentiation and genetic diversity within and between populations of this species. These results are discussed with regards to the practical application of molecular markers in conservation programs.

  • Litkowiec, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland ORCID ID:E-mail: mlit@man.poznan.pl (email)
  • Plitta-Michalak, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland ORCID ID:E-mail: beata-plitta@wp.pl
  • Lewandowski, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland ORCID ID:E-mail: alew@man.poznan.pl
  • Iszkuło, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland & University of Zielona Góra, Faculty of Biological Sciences, Prof. Z. Szafrana 1, 65-516 Zielona Góra, Poland ORCID ID:E-mail: iszkulo@man.poznan.pl
article id 1337, category Research article
Leszek Bujoczek, Małgorzata Bujoczek, Jan Banaś, Stanisław Zięba. (2015). Spruce regeneration on woody microsites in a subalpine forest in the western Carpathians. Silva Fennica vol. 49 no. 3 article id 1337. https://doi.org/10.14214/sf.1337
Highlights: The occurrence probability of Picea abies seedlings on fallen deadwood was found to increase with diameter and decay stage of deadwood and with the volume of living trees, and to decrease with the density of living trees, sapling density, and land slope. It was also higher on stumps with greater diameter and in plots with higher sapling density, but decreased with increasing stump height.

The density of Picea abies [L.] Karst. regeneration on different microsites, the quantity and quality of woody microsites, and seedling occurrence probability on stumps and fallen deadwood were studied in a subalpine forest that has been under protection for approximately 30–40 years (Gorce Mountains in the western Carpathians). Thirty percent of seedlings and 29% of saplings grew on stumps and fallen deadwood, while the remaining regeneration occurred on soil surface and mounds created by uprooted trees. The occurrence probability of Picea seedlings on fallen deadwood increased with deadwood diameter and decay stage and with the volume of living trees, and decreased with increased density of living trees, sapling density, and land slope. Furthermore, seedlings were more likely to grow on stumps with a greater diameter and in plots with higher sapling density, but less likely to grow on higher stumps. Stumps and fallen deadwood covered about 4% of the forest floor, but the material that is most important for promoting regeneration (strongly decomposed logs and those of a diameter exceeding 30 cm) took up only about 22 m2 ha-1. We have concluded that in a subalpine forest that has been protected for 30–40 years regeneration processes take place mostly on soil surface and stumps. The role of fallen deadwood increases over time as a greater number of suitable logs (in terms of size and decay stage) become available.

  • Bujoczek, University of Agriculture in Krakow, ORCID ID:E-mail: lbujoczek@gmail.com (email)
  • Bujoczek, University of Agriculture in Krakow, ORCID ID:E-mail: bujoczek.m@gmail.com
  • Banaś, University of Agriculture in Krakow, ORCID ID:E-mail: rlbanas@cyf-kr.edu.pl
  • Zięba, University of Agriculture in Krakow, ORCID ID:E-mail: rlzieba@cyf-kr.edu.pl
article id 1267, category Research article
Caroline Mary Adrianne Franklin, Karen A Harper, Liam Kyte Murphy. (2015). Structural dynamics at boreal forest edges created by a spruce budworm outbreak. Silva Fennica vol. 49 no. 3 article id 1267. https://doi.org/10.14214/sf.1267
Highlights: Insect outbreak edges were 10 m wide with different canopy cover, stem density and tree structural diversity than adjacent ecosystems; Although edge influence on forest structure was weak, forest influence was stronger and extended further, creating an edge zone skewed towards the disturbed area; After thirty years, high-contrast and structurally-diverse transition zones persist on the landscape.
Natural disturbances such as insect outbreaks create boundaries that influence vegetation patterns and ecological processes.  To better understand the effects of natural edge creation on relatively intact forests and adjacent disturbed areas, we investigated forest structure on both sides of 30 year-old forest edges created by a spruce budworm (Choristoneura fumiferana Clemens) outbreak in the boreal forest of Cape Breton Highlands National Park, Canada.  Our objectives were: 1) to determine edge influence (compared to interior forest) and forest influence (compared to disturbed areas) on vegetation structure, and 2) to gain insight into the structural development of the edges.  Canopy cover, tree density, radial growth and deadwood were sampled in 5 m x 20 m plots along 120 m transects across six edges.  Randomization tests were used to estimate the magnitude and distance of edge and forest influence.  Narrow transition zones approximately 10 m wide characterized the spruce budworm-induced edges.  Edge influence did not extend into the forest; however, forest influence on structure was detected up to 40 m from the edge into the disturbed area.  We found evidence of the insect outbreak in the form of reduced radial growth during the disturbance across the entire disturbed area-forest gradient, which indicates that spruce budworm activity may not have ceased directly at the edge.  Tree mortality caused by the insect outbreak resulted in snags, many of which have transformed into logs since the outbreak collapsed.  Spruce budworm outbreak-induced forest edges are narrow but dynamic boundaries separating two distinct vegetation communities in the boreal landscape.
  • Franklin, Department of Renewable Resources, University of Alberta, 751 General Services Building, Edmonton, Alberta, T6G 2H1, Canada ORCID ID:E-mail: cfrankli@ualberta.ca (email)
  • Harper, School for Resource and Environmental Studies, Dalhousie University, Suite 5010, 6100 University Ave., Halifax, Nova Scotia, B3H 3J5, Canada ORCID ID:E-mail: Karen.Harper@dal.ca
  • Murphy, Department of Environmental Science, Saint Mary’s University, 923 Robie St., Halifax, Nova Scotia, B3H 3C3, Canada ORCID ID:E-mail: liamkmurphy@gmail.com
article id 1262, category Research article
Āris Jansons, Roberts Matisons, Māra Zadiņa, Linards Sisenis, Jānis Jansons. (2015). The effect of climatic factors on height increment of Scots pine in sites differing by continentality in Latvia. Silva Fennica vol. 49 no. 3 article id 1262. https://doi.org/10.14214/sf.1262
Highlights: Height increment-climate relationships of Scots pine were assessed using dendrochronological techniques; Annual height increment was significantly affected by climatic factors; In western Latvia, temperature in preceding summer mainly affected height increment; In eastern Latvia height increment was affected by previous autumn temperature; During the 20th century, the effect of climatic factors has altered likely dues to climate change.
Height growth of trees is a crucial parameter that influences the composition and productivity of forest stands and quality of timber; however, the relationships between annual height increment (HI) and climatic factors have been poorly studied. In this study, the effect of monthly mean temperature and precipitation sums on the HI of Scots pine in two sites in Latvia have been determined using dendrochronological techniques. Correlation and response function analyses were conducted for entire chronologies of HI and for 50-year intervals within them. Climatic factors significantly affected the HI of Scots pine; however, not only did the suite of significant factors differ between the sites, but the influence of these factors changed during the 20th century. In the site in western Latvia where climate is milder, temperature in the preceding summer was the main climatic determinant of HI. The effect of temperature in the dormant period and spring was significant during the first part of the 20th century, while the effect of temperature in the previous September and November has become significant since the second half of the 20th century. In the site in eastern Latvia where summers are hotter, HI has been affected by both temperature and water deficit related factors in the summer. However, since the later part of the 20th century, the effect of temperature in the previous October has intensified and become the main climatic determinant of HI.
  • Jansons,  LSFRI “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 ORCID ID:E-mail: aris.jansons@silava.lv
  • Matisons,  LSFRI “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 ORCID ID:E-mail: robism@inbox.lv (email)
  • Zadiņa,  LSFRI “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 ORCID ID:E-mail: mara.zadina@silava.lv
  • Sisenis, LUA Forestry Faculty, Akadēmijas str. 11, Jelgava, Latvia ORCID ID:E-mail: linards.sisenis@llu.lv
  • Jansons, Forest Competence Centre, Dzērbenes str. 27, Riga, Latvia, LV1006 ORCID ID:E-mail: janis.jansons@silava.lv
article id 1262, category Research article
Āris Jansons, Roberts Matisons, Māra Zadiņa, Linards Sisenis, Jānis Jansons. (2015). The effect of climatic factors on height increment of Scots pine in sites differing by continentality in Latvia. Silva Fennica vol. 49 no. 3 article id 1262. https://doi.org/10.14214/sf.1262
Highlights: Height increment-climate relationships of Scots pine were assessed using dendrochronological techniques; Annual height increment was significantly affected by climatic factors; In western Latvia, temperature in preceding summer mainly affected height increment; In eastern Latvia height increment was affected by previous autumn temperature; During the 20th century, the effect of climatic factors has altered likely dues to climate change.
Height growth of trees is a crucial parameter that influences the composition and productivity of forest stands and quality of timber; however, the relationships between annual height increment (HI) and climatic factors have been poorly studied. In this study, the effect of monthly mean temperature and precipitation sums on the HI of Scots pine in two sites in Latvia have been determined using dendrochronological techniques. Correlation and response function analyses were conducted for entire chronologies of HI and for 50-year intervals within them. Climatic factors significantly affected the HI of Scots pine; however, not only did the suite of significant factors differ between the sites, but the influence of these factors changed during the 20th century. In the site in western Latvia where climate is milder, temperature in the preceding summer was the main climatic determinant of HI. The effect of temperature in the dormant period and spring was significant during the first part of the 20th century, while the effect of temperature in the previous September and November has become significant since the second half of the 20th century. In the site in eastern Latvia where summers are hotter, HI has been affected by both temperature and water deficit related factors in the summer. However, since the later part of the 20th century, the effect of temperature in the previous October has intensified and become the main climatic determinant of HI.
  • Jansons,  LSFRI “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 ORCID ID:E-mail: aris.jansons@silava.lv
  • Matisons,  LSFRI “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 ORCID ID:E-mail: robism@inbox.lv (email)
  • Zadiņa,  LSFRI “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 ORCID ID:E-mail: mara.zadina@silava.lv
  • Sisenis, LUA Forestry Faculty, Akadēmijas str. 11, Jelgava, Latvia ORCID ID:E-mail: linards.sisenis@llu.lv
  • Jansons, Forest Competence Centre, Dzērbenes str. 27, Riga, Latvia, LV1006 ORCID ID:E-mail: janis.jansons@silava.lv
article id 1239, category Research article
Tomi Kaakkurivaara, Nuutti Vuorimies, Pauli Kolisoja, Jori Uusitalo. (2015). Applicability of portable tools in assessing the bearing capacity of forest roads. Silva Fennica vol. 49 no. 2 article id 1239. https://doi.org/10.14214/sf.1239
Highlights: The dynamic cone penetrometer (DCP) and light falling weight deflectometer (LFWD) are useful tools for measuring bearing capacity; The measurement results are not same as with the falling weight deflectometer (FWD), but comparable.
Forest roads provide access to logging sites and enable transportation of timber from forest to mills. Efficient forest management and forest industry are impossible without a proper forest road network. The bearing capacity of forest roads varies significantly by weather conditions and seasons since they are generally made of poor materials and the constructed layers may be mixed with subgrade. A bearing capacity assessment is valuable information when trafficability is uncertain and rutting is obvious. In this study, bearing capacity measurements were carried out using the light falling weight deflectometer (LFWD), the dynamic cone penetrometer (DCP) and the conventional falling weight deflectometer (FWD). The aim was to compare their measurement results in relation to road characteristics and moisture conditions. Data were collected from 35 test road sections in four consecutive springs and during one summer. The test road sections had measurement points both on the wheel path and the centre line. The data show logical correlations between measured quantities, and the study presents reliable regression models between measuring devices. The results indicate that light portable tools, the DCP and the LFWD, can in most cases be used instead of the expensive falling weight deflectometer on forest roads.
  • Kaakkurivaara, Natural Resources Institute Finland, Green technology, Kaironiementie 15, FI-39700 Parkano, Finland ORCID ID:E-mail: tomi.kaakkurivaara@gmail.com (email)
  • Vuorimies, Tampere University of Technology, P.O.Box 600, FI-33101 Tampere, Finland ORCID ID:E-mail: nuutti.vuorimies@tut.fi
  • Kolisoja, Tampere University of Technology, P.O.Box 600, FI-33101 Tampere, Finland ORCID ID:E-mail: pauli.kolisoja@tut.fi
  • Uusitalo, Natural Resources Institute Finland, Green technology, Kaironiementie 15, FI-39700 Parkano, Finland ORCID ID:E-mail: jori.uusitalo@luke.fi
article id 1239, category Research article
Tomi Kaakkurivaara, Nuutti Vuorimies, Pauli Kolisoja, Jori Uusitalo. (2015). Applicability of portable tools in assessing the bearing capacity of forest roads. Silva Fennica vol. 49 no. 2 article id 1239. https://doi.org/10.14214/sf.1239
Highlights: The dynamic cone penetrometer (DCP) and light falling weight deflectometer (LFWD) are useful tools for measuring bearing capacity; The measurement results are not same as with the falling weight deflectometer (FWD), but comparable.
Forest roads provide access to logging sites and enable transportation of timber from forest to mills. Efficient forest management and forest industry are impossible without a proper forest road network. The bearing capacity of forest roads varies significantly by weather conditions and seasons since they are generally made of poor materials and the constructed layers may be mixed with subgrade. A bearing capacity assessment is valuable information when trafficability is uncertain and rutting is obvious. In this study, bearing capacity measurements were carried out using the light falling weight deflectometer (LFWD), the dynamic cone penetrometer (DCP) and the conventional falling weight deflectometer (FWD). The aim was to compare their measurement results in relation to road characteristics and moisture conditions. Data were collected from 35 test road sections in four consecutive springs and during one summer. The test road sections had measurement points both on the wheel path and the centre line. The data show logical correlations between measured quantities, and the study presents reliable regression models between measuring devices. The results indicate that light portable tools, the DCP and the LFWD, can in most cases be used instead of the expensive falling weight deflectometer on forest roads.
  • Kaakkurivaara, Natural Resources Institute Finland, Green technology, Kaironiementie 15, FI-39700 Parkano, Finland ORCID ID:E-mail: tomi.kaakkurivaara@gmail.com (email)
  • Vuorimies, Tampere University of Technology, P.O.Box 600, FI-33101 Tampere, Finland ORCID ID:E-mail: nuutti.vuorimies@tut.fi
  • Kolisoja, Tampere University of Technology, P.O.Box 600, FI-33101 Tampere, Finland ORCID ID:E-mail: pauli.kolisoja@tut.fi
  • Uusitalo, Natural Resources Institute Finland, Green technology, Kaironiementie 15, FI-39700 Parkano, Finland ORCID ID:E-mail: jori.uusitalo@luke.fi
article id 1218, category Research article
Mikko Niemi, Mikko Vastaranta, Jussi Peuhkurinen, Markus Holopainen. (2015). Forest inventory attribute prediction using airborne laser scanning in low-productive forestry-drained boreal peatlands. Silva Fennica vol. 49 no. 2 article id 1218. https://doi.org/10.14214/sf.1218
Highlights: Following current forest inventory practises, stem volume was predicted in low-productive drained peatlands (LPDPs) with a root mean square error (RMSE) of 13.7 m3 ha–1; When 30 reference plots measured from LPDPs were added to the prediction, RMSE was decreased to 10.0 m3 ha–1; Additional reference plots from LPDPs did not affect the forest inventory attribute predictions in productive forests.
Nearly 30% of Finland’s land area is covered by peatlands. In Northern parts of the country there is a significant amount of low-productive drained peatlands (LPDPs) where the average annual stem volume growth is less than 1 m3 ha–1. The re-use of LPDPs has been considered thoroughly since Finnish forest legislation was updated and the forest regeneration prerequisite was removed from LPDPs in January 2014. Currently, forestry is one of the re-use alternatives, thus detailed forest resource information is required for allocating activities. However, current forest inventory practices have not been evaluated for sparse growing stocks (e.g., LPDPs). The purpose of our study was to evaluate the suitability of airborne laser scanning (ALS) for mapping forest inventory attributes in LPDPs. We used ALS data with a density of 0.8 pulses per m2, 558 field-measured reference plots (500 from productive forests and 58 from LPDPs) and k nearest neighbour (k-NN) estimation. Our main aim was to study the sensitivity of predictions to the number of LPDP reference plots used in the k-NN estimation. When the reference data consisted of 500 plots from productive forest stands, the root mean square errors (RMSEs) for the prediction accuracy of Lorey’s height, basal area and stem volume were 1.4 m, 2.7 m2 ha–1 and 13.7 m3 ha–1 in LPDPs, respectively. When 30 additional reference plots were allocated to LPDPs, the respective RMSEs were 1.1 m, 1.7 m2 ha–1 and 10.0 m3 ha–1. Additional reference plot allocation did not affect the predictions in productive forest stands.
  • Niemi, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014, Finland & Centre of Excellence in Laser Scanning Research, Finnish Geospatial Research Institute FGI, Geodeetinrinne 2, FI-02430, Finland ORCID ID:E-mail: mikko.t.niemi@helsinki.fi (email)
  • Vastaranta, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014, Finland & Centre of Excellence in Laser Scanning Research, Finnish Geospatial Research Institute FGI, Geodeetinrinne 2, FI-02430, Finland ORCID ID:E-mail: mikko.vastaranta@helsinki.fi
  • Peuhkurinen, Arbonaut Oy Ltd., Latokartanontie 7 A, FI-00700, Finland ORCID ID:E-mail: jussi.peuhkurinen@arbonaut.com
  • Holopainen, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014, Finland & Centre of Excellence in Laser Scanning Research, Finnish Geospatial Research Institute FGI, Geodeetinrinne 2, FI-02430, Finland ORCID ID:E-mail: markus.holopainen@helsinki.fi
article id 1220, category Research article
Elisabeth Düthorn, Lea Schneider, Oliver Konter, Philipp Schön, Mauri Timonen, Jan Esper. (2015). On the hidden significance of differing micro-sites on tree-ring based climate reconstructions. Silva Fennica vol. 49 no. 1 article id 1220. https://doi.org/10.14214/sf.1220
Highlights: Pines and spruces show growth level differences in wet and dry micro-sites with higher growth rates in the dry sites; Spruces show a robust climate-growth relationship with June-July temperatures; Application of collective detrending methods can bias long-term trends in climate reconstructions, if relict and recent samples originate from different micro-sites.
Tree-ring chronologies are commonly extended back in time by combining samples from living trees with relict material preserved in man-made structures or natural archives (e.g. lakes). Although spatially close, these natural archives and living-tree-sites often comprise different micro-climates. Inhomogeneous growth conditions among these habitats, which may yield offsets in growth-rates, require caution in data processing. Here we assess species-specific growth dynamics in two micro-habitats and their potential effects on long chronologies by combining tree-ring data from different living-tree-sites with an “artificial” subfossil dataset. Well replicated (n > 80) Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) chronologies from northern Fennoscandia, sampled directly at the lakeshore (wet) and several meters beyond the lakeshore (dry) reveal high coherence of the variance between micro-sites (rspruce = 0.59, rpine = 0.68). Significant differences of the Regional Curves (RC) indicate faster growth of both species at the drier site though. Growth differences are more pronounced between the spruce micro-sites. The combination of recent dry and wet spruce data with artificial relict data results in two long chronologies covering the last 800 years with substantially different trends, although they consist of the same relict material and the micro-site chronologies correlate significantly over the past two centuries. The combination of spruce samples from dry inland micro-sites with subfossil samples originating from the wet lake shore can result in an underestimation of past temperatures prior to the 19th century. Such effects, hidden in the composition of long chronologies (living trees + subfossil samples) can bias long-term trends in climate reconstructions.
  • Düthorn, Department of Geography, Johannes Gutenberg University, Becherweg 21, 55099 Mainz, Germany ORCID ID:E-mail: duethorn@uni-mainz.de (email)
  • Schneider, Department of Geography, Johannes Gutenberg University, Becherweg 21, 55099 Mainz, Germany ORCID ID:E-mail: l.schneider@geo.uni-mainz.de
  • Konter, Department of Geography, Johannes Gutenberg University, Becherweg 21, 55099 Mainz, Germany ORCID ID:E-mail: O.Konter@geo.uni-mainz.de
  • Schön, Department of Geography, Johannes Gutenberg University, Becherweg 21, 55099 Mainz, Germany ORCID ID:E-mail: philipp.schoen@gmx.de
  • Timonen, Natural Resources Institute Finland (Luke), Natural resources and bioproduction, FI-96301 Rovaniemi, Finland ORCID ID:E-mail: mauri.timonen@metla.fi
  • Esper, Department of Geography, Johannes Gutenberg University, Becherweg 21, 55099 Mainz, Germany ORCID ID:E-mail: J.Esper@geo.uni-mainz.de
article id 1219, category Research article
Thomas P. Sullivan, Druscilla S. Sullivan. (2014). Diversifying clearcuts with green-tree retention and woody debris structures: conservation of mammals across forest ecological zones. Silva Fennica vol. 48 no. 5 article id 1219. https://doi.org/10.14214/sf.1219
Highlights: Species diversity of small mammals increased with structural complexity left on clearcut sites; Productivity of red-backed vole populations was higher in sites with green-tree retention (GTR) and windrows of woody debris; GTR and windrows may provide additive effect for providing habitat to conserve mammals on clearcuts.
We tested the hypotheses (H) that on newly clearcut-harvested sites, (H1) abundance and species diversity of the forest-floor small mammal community, and (H2) abundance, reproduction, and recruitment of red-backed voles (Myodes gapperi Vigors), would increase with higher levels of structural retention via green-tree retention (GTR) and woody debris (dispersed and constructed into windrows). Study areas were located in three forest ecological zones in southern British Columbia, Canada. For H1, mean total abundance did generally increase with the gradient of retained habitat structure. Mean species richness and diversity were similar among treatment sites but did show an increasing gradient with structural compexity. For H2, mean abundance, reproduction, and recruitment of M. gapperi were higher in GTR and windrow sites than those without retained structures. There was a positive relationship between mean abundance of M. gapperi and total volume of woody debris across treatments. This study is the first investigation of the responses of forest-floor small mammals to an increasing gradient of retained habitat structure via GTR and woody debris on clearcuts. Our assessment of a combination of these two interventions suggested a potentially strong additive effect that could be cautiously extrapolated across three forest ecological zones. With the advent of low levels of GTR on clearcuts, woody debris structures should help provide some habitat to conserve forest mammals on harvest openings.
  • Sullivan, Department of Forest and Conservation Sciences, Faculty of Forestry, University of BC, 2424 Main Mall, Vancouver, BC, Canada V6T 1Z4 ORCID ID:E-mail: tom.sullivan@ubc.ca (email)
  • Sullivan, Applied Mammal Research Institute, 11010 Mitchell Avenue, Summerland, BC, Canada V0H 1Z8 ORCID ID:E-mail: dru.sullivan@appliedmammal.com
article id 1211, category Research article
Xiao Chen, Deborah Page-Dumroese, Ruiheng Lv, Weiwei Wang, Guolei Li, Yong Liu. (2014). Interaction of initial litter quality and thinning intensity on litter decomposition rate, nitrogen accumulation and release in a pine plantation. Silva Fennica vol. 48 no. 4 article id 1211. https://doi.org/10.14214/sf.1211
Highlights: Litter quality and thinning showed an interaction on one year litter decomposition rates, N accumulation, and net N release; N accumulated until the underlying critical acid-unhydrolyzable residue to nitrogen ratio (approximately 57–69) was met; Increased N concentration in litter and thinning intensity induced rapid litter decomposition and N cycling in coniferous plantation with a slow decomposition rate.
Thinning alters litter quality and microclimate under forests. Both of these two changes after thinning induce alterations of litter decomposition rates and nutrient cycling. However, a possible interaction between these two changes remains unclear. We placed two types of litter (LN, low N concentration litter; HN, high N concentration litter) in a Chinese pine (Pinus tabulaeformis Carrière) plantation under four thinning treatments to test the impacts of litter quality, thinning or their combination on decomposition rate and N cycling. In our study, N was accumulated to approach an underlying critical acid-unhydrolyzable residue to nitrogen ratio (approximately 57–69) in litter. Moreover, an interaction between litter quality and thinning on decomposition rates, N accumulation and net release did exist. On one hand, one year decomposition rate of LN was elevated after thinning while that of HN remained the same or even lower (under light thinning); N accumulation of LN declined with light thinning and was restored with the increase of thinning intensity whereas that of HN did not decline with thinning and increased under heavy thinning; Net N release from LN was only found in light and heavy thinning while that from HN was found in all treatments, moreover net N release from LN and HN were both elevated under heavy thinning. On the other hand, HN decomposed faster, accumulated less and released more N than LN did under all treatments. Generally, high N concentration in litter and high-intensity thinning can lead to rapid litter decomposition and N cycling in coniferous plantations.
  • Chen, Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing, 100083, China ORCID ID:E-mail: chenxiao_0123@126.com
  • Page-Dumroese, U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, 1221 South Main Street, Moscow, ID 83843, USA ORCID ID:E-mail: ddumroese@fs.fed.us
  • Lv, College of Plant Science and Technology, Tarim University, Alar Xinjiang, 843300, China ORCID ID:E-mail: lvrh514723@126.com
  • Wang, Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing, 100083, China ORCID ID:E-mail: fuyuerdejia@126.com
  • Li, Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing, 100083, China ORCID ID:E-mail: glli226@163.com
  • Liu, Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing, 100083, China ORCID ID:E-mail: lyong@bjfu.edu.cn (email)
article id 1207, category Research article
Olli-Pekka Tikkanen, Irina A. Chernyakova. (2014). Past human population history affects current forest landscape structure of Vodlozero National Park, Northwest Russia. Silva Fennica vol. 48 no. 4 article id 1207. https://doi.org/10.14214/sf.1207
Highlights: In large scale (0–20 km), the proportion of spruce in forest stands was positively affected by distance from old villages. This relationship was non-linear with a threshold distance of 15 km; In small scale (0–5 km), old villages affected tree species composition and age structure of forests. Effect on age structure was the strongest on stands growing on mineral soils.
The information about location and size of past human settlements can give new insights into the analysis of landscape structures. Vodlozero National Park (NP) is one the largest strictly protected areas in Northwestern Europe. We mapped the location of historic villages, which were abandoned in 1958, and studied the effect of past human activity in the forest landscape in two different scales using forest survey data. We assessed the possible change in age, volume and tree species composition from the edge of open fields up to the distance of 5 km from villages. We made a larger landscape analysis using a grid of forest stands covering the whole northern part of the NP. The past human activity was clearly visible in the present forest landscape. Distance from villages affected age, volume and tree species composition of the forest stands. This effect was the strongest within the first two kilometers from the villages. At the level of whole northern NP, the proportion of spruce markedly increased after approximately 15 km from the nearest old village. The changes in the forests surrounding the villages were most likely the result of the intensive use of wood for different commodities needed in households and farming, in addition to short rotation slash and burn agriculture. If the occurrence of forest fires was more frequent closer to villages than in more remote areas, it can well explain the observed pattern in the abundance of spruce in the larger landscape that is less tolerant to fire than pine.
  • Tikkanen, Department of Biology, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland (Current: School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland) & Interdisciplinary Research and Educational Center of Cross-border Communication CARELICA, Institute of History, Political and Social Sciences, Petrozavodsk State University, 33 Lenin Prospectus, 185910 Petrozavodsk, Republic of Karelia, Russia ORCID ID:E-mail: Olli-Pekka.Tikkanen@uef.fi (email)
  • Chernyakova, Interdisciplinary Research and Educational Center of Cross-border Communication CARELICA, Institute of History, Political and Social Sciences, Petrozavodsk State University, 33 Lenin Prospectus, 185910 Petrozavodsk, Republic of Karelia, Russia ORCID ID:E-mail: irina.chernyakova@onego.ru
article id 1161, category Research article
Tiina Laine, Veli-Matti Saarinen. (2014). Comparative study of the Risutec Automatic Plant Container (APC) and Bracke planting devices. Silva Fennica vol. 48 no. 3 article id 1161. https://doi.org/10.14214/sf.1161
Highlights: As currently designed, the prototype Risutec APC fitted with an automatic feeding system offers no significant advantage over the Bracke planting device in terms of planting productivity or quality; Cost estimates suggest that an idealized automated feeding system could increase productivity and decrease unit costs.
The productivity of mechanized planting could be increased by minimizing the time spent manually reloading seedling cassettes. This study compared the work-time distribution, productivity and quality of the prototype Risutec APC fitted with an automatic feeding system and the commonly-used and manually-loaded Bracke P11.a. An approach of comparative time study was employed that compared performance of two operators using both machines in four sites where slash and stumps had been removed. Operating costs were estimated and compared for these two machines and an idealized machine with automatic feeding system (referred as AUT). AUT was assumed to be similar to the Bracke planting machine with the only difference being in automatic feeding. Productivity of the Risutec APC (196 seedlings per productive work hour [pl PWh0–1]) was lower than that of Bracke (244 pl PWh0–1), making the unit cost 35.7% higher. A large portion (17.6%) of the productive work time of Risutec APC was interrupted by malfunctions, so it cannot be considered robust and reliable yet. Quality of the planting work was reasonable for both machines. The results suggest that an idealized AUT could increase planting capacity (hectares per year [ha yr–1]) by 15.4% and lower the unit cost (Euro per seedling [€ pl–1]) by 4.7% compared to today’s machines. The importance of an automated feeding system increases with planting efficiency because relatively more time is spent reloading seedlings. Proper automatic feeding system could offer a cost-effective solution and could enhance productivity, but the Risutec APC has yet to meet the technical and economic standards required to be competitive.
  • Laine, Finnish Forest Research Institute (METLA), Suonenjoki Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: tiina.laine@metla.fi (email)
  • Saarinen, Finnish Forest Research Institute (METLA), Suonenjoki Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: veli-matti.saarinen@metla.fi
article id 1087, category Research article
Ilkka Korpela, Lauri Mehtätalo, Lauri Markelin, Anne Seppänen, Annika Kangas. (2014). Tree species identification in aerial image data using directional reflectance signatures. Silva Fennica vol. 48 no. 3 article id 1087. https://doi.org/10.14214/sf.1087
Highlights: Multispectral reflectance data showed a strong and spectrally correlated tree effect; There was no gain in species classification from using species-specific differences of directional reflectance in real data and only a marginal improvement in simulated data; The directional signatures extracted in multiple images are obscured by the intrinsic within-species variation, correlated observations and inherent reflectance calibration errors.
Tree species identification using optical remote sensing is challenging. Modern digital photogrammetric cameras enable radiometrically quantitative remote sensing and the estimation of reflectance images, in which the observations depend largely on the reflectance properties of targets. Previous research has shown that there are species-specific differences in how the brightness observed changes when the viewing direction in an aerial image is altered. We investigated if accounting for such directional signatures enhances species classification, using atmospherically corrected, real and simulated multispectral Leica ADS40 line-camera data. Canopy in direct and diffuse illumination were differentiated and species-specific variance-covariance structures were analyzed in real reflectance data, using mixed-effects modeling. Species classification simulations aimed at elucidating the level of accuracy that can be achieved by using images of different quality, number and view-illumination geometry. In real data, a substantial variance component was explained by tree effect, which demonstrates that observations from a tree correlate between observation geometries as well as spectrally. Near-infrared band showed the strongest tree effect, while the directionality was weak in that band. The gain from directional signatures was insignificant in real data, while simulations showed a potential gain of 1–3 percentage points in species classification accuracy. The quality of reflectance calibration was found to be important as well as the image acquisition geometry. We conclude that increasing the number of image observations cancels out random observation noise and reflectance calibration errors, but fails to eliminate the tree effect and systematic calibration inaccuracy. Directional reflectance constitutes a marginal improvement in tree species classification.
  • Korpela, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014, Finland ORCID ID:E-mail: ilkka.korpela@helsinki.fi (email)
  • Mehtätalo, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: lauri.mehtatalo@uef.fi
  • Markelin, Department of Remote Sensing and Photogrammetry, Finnish Geodetic Institute, P.O. Box 15, FI-02431 Masala, Finland ORCID ID:E-mail: lauri.markelin@fgi.fi
  • Seppänen, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: anne.seppanen@arbonaut.com
  • Kangas, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014, Finland ORCID ID:E-mail: annika.kangas@helsinki.fi
article id 1087, category Research article
Ilkka Korpela, Lauri Mehtätalo, Lauri Markelin, Anne Seppänen, Annika Kangas. (2014). Tree species identification in aerial image data using directional reflectance signatures. Silva Fennica vol. 48 no. 3 article id 1087. https://doi.org/10.14214/sf.1087
Highlights: Multispectral reflectance data showed a strong and spectrally correlated tree effect; There was no gain in species classification from using species-specific differences of directional reflectance in real data and only a marginal improvement in simulated data; The directional signatures extracted in multiple images are obscured by the intrinsic within-species variation, correlated observations and inherent reflectance calibration errors.
Tree species identification using optical remote sensing is challenging. Modern digital photogrammetric cameras enable radiometrically quantitative remote sensing and the estimation of reflectance images, in which the observations depend largely on the reflectance properties of targets. Previous research has shown that there are species-specific differences in how the brightness observed changes when the viewing direction in an aerial image is altered. We investigated if accounting for such directional signatures enhances species classification, using atmospherically corrected, real and simulated multispectral Leica ADS40 line-camera data. Canopy in direct and diffuse illumination were differentiated and species-specific variance-covariance structures were analyzed in real reflectance data, using mixed-effects modeling. Species classification simulations aimed at elucidating the level of accuracy that can be achieved by using images of different quality, number and view-illumination geometry. In real data, a substantial variance component was explained by tree effect, which demonstrates that observations from a tree correlate between observation geometries as well as spectrally. Near-infrared band showed the strongest tree effect, while the directionality was weak in that band. The gain from directional signatures was insignificant in real data, while simulations showed a potential gain of 1–3 percentage points in species classification accuracy. The quality of reflectance calibration was found to be important as well as the image acquisition geometry. We conclude that increasing the number of image observations cancels out random observation noise and reflectance calibration errors, but fails to eliminate the tree effect and systematic calibration inaccuracy. Directional reflectance constitutes a marginal improvement in tree species classification.
  • Korpela, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014, Finland ORCID ID:E-mail: ilkka.korpela@helsinki.fi (email)
  • Mehtätalo, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: lauri.mehtatalo@uef.fi
  • Markelin, Department of Remote Sensing and Photogrammetry, Finnish Geodetic Institute, P.O. Box 15, FI-02431 Masala, Finland ORCID ID:E-mail: lauri.markelin@fgi.fi
  • Seppänen, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: anne.seppanen@arbonaut.com
  • Kangas, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014, Finland ORCID ID:E-mail: annika.kangas@helsinki.fi
article id 980, category Research article
Atte Komonen, Panu Halme, Mari Jäntti, Tuuli Koskela, Janne S. Kotiaho, Tero Toivanen. (2014). Created substrates do not fully mimic natural substrates in restoration: the occurrence of polypores on spruce logs. Silva Fennica vol. 48 no. 1 article id 980. https://doi.org/10.14214/sf.980
Highlights: Polypore communities were more homogeneous among created than among natural logs; The old-growth forest indicator Phellinus ferrugineofuscus occurred frequently on natural logs, but occupied only a few created logs; Results show that created logs do not fully mimic natural logs.
Many protected areas have been under intensive forest management prior to protection and thus lack natural ecosystem structures and dynamics. Dead wood is a key structure in forests harboring hundreds of threatened species. We investigated the ecological success of dead wood creation as a boreal forest restoration measure. We analysed whether the polypore communities of chain-saw felled and girdled (subsequently fallen) Norway spruce (Picea abies (L.) H. Karst.) logs differ from naturally formed spruce logs of similar decay stage and size. The study was conducted in Leivonmäki National Park in central Finland 8 years after the restoration measures. The average number of polypore species was highest on the chain-saw felled logs and most of the common polypore species were most frequent on this substrate. However, among the natural logs, number of species increased more steeply with increasing number of logs, suggesting greater variation in community composition on this substrate. The old-growth forest indicator Phellinus ferrugineofuscus occurred frequently on natural logs, occupied a few girdled logs but was absent from chain-saw felled logs. Our results show that from the polypore perspective created logs do not fully mimic natural logs, suggesting that creating substrates for species may pose a challenge for restoration.
  • Komonen, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland ORCID ID:E-mail: atte.komonen@jyu.fi (email)
  • Halme, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland ORCID ID:E-mail: panu.halme@jyu.fi
  • Jäntti, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland ORCID ID:E-mail: mari.j.jantti@student.jyu.fi
  • Koskela, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland ORCID ID:E-mail: tuuli.e.koskela@student.jyu.fi
  • Kotiaho, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland ORCID ID:E-mail: janne.kotiaho@jyu.fi
  • Toivanen, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland; Current: Birdlife Finland, Annankatu 29 A 16, FI-00100 Helsinki, Finland ORCID ID:E-mail: tero.toivanen@birdlife.fi
article id 980, category Research article
Atte Komonen, Panu Halme, Mari Jäntti, Tuuli Koskela, Janne S. Kotiaho, Tero Toivanen. (2014). Created substrates do not fully mimic natural substrates in restoration: the occurrence of polypores on spruce logs. Silva Fennica vol. 48 no. 1 article id 980. https://doi.org/10.14214/sf.980
Highlights: Polypore communities were more homogeneous among created than among natural logs; The old-growth forest indicator Phellinus ferrugineofuscus occurred frequently on natural logs, but occupied only a few created logs; Results show that created logs do not fully mimic natural logs.
Many protected areas have been under intensive forest management prior to protection and thus lack natural ecosystem structures and dynamics. Dead wood is a key structure in forests harboring hundreds of threatened species. We investigated the ecological success of dead wood creation as a boreal forest restoration measure. We analysed whether the polypore communities of chain-saw felled and girdled (subsequently fallen) Norway spruce (Picea abies (L.) H. Karst.) logs differ from naturally formed spruce logs of similar decay stage and size. The study was conducted in Leivonmäki National Park in central Finland 8 years after the restoration measures. The average number of polypore species was highest on the chain-saw felled logs and most of the common polypore species were most frequent on this substrate. However, among the natural logs, number of species increased more steeply with increasing number of logs, suggesting greater variation in community composition on this substrate. The old-growth forest indicator Phellinus ferrugineofuscus occurred frequently on natural logs, occupied a few girdled logs but was absent from chain-saw felled logs. Our results show that from the polypore perspective created logs do not fully mimic natural logs, suggesting that creating substrates for species may pose a challenge for restoration.
  • Komonen, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland ORCID ID:E-mail: atte.komonen@jyu.fi (email)
  • Halme, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland ORCID ID:E-mail: panu.halme@jyu.fi
  • Jäntti, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland ORCID ID:E-mail: mari.j.jantti@student.jyu.fi
  • Koskela, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland ORCID ID:E-mail: tuuli.e.koskela@student.jyu.fi
  • Kotiaho, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland ORCID ID:E-mail: janne.kotiaho@jyu.fi
  • Toivanen, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland; Current: Birdlife Finland, Annankatu 29 A 16, FI-00100 Helsinki, Finland ORCID ID:E-mail: tero.toivanen@birdlife.fi
article id 980, category Research article
Atte Komonen, Panu Halme, Mari Jäntti, Tuuli Koskela, Janne S. Kotiaho, Tero Toivanen. (2014). Created substrates do not fully mimic natural substrates in restoration: the occurrence of polypores on spruce logs. Silva Fennica vol. 48 no. 1 article id 980. https://doi.org/10.14214/sf.980
Highlights: Polypore communities were more homogeneous among created than among natural logs; The old-growth forest indicator Phellinus ferrugineofuscus occurred frequently on natural logs, but occupied only a few created logs; Results show that created logs do not fully mimic natural logs.
Many protected areas have been under intensive forest management prior to protection and thus lack natural ecosystem structures and dynamics. Dead wood is a key structure in forests harboring hundreds of threatened species. We investigated the ecological success of dead wood creation as a boreal forest restoration measure. We analysed whether the polypore communities of chain-saw felled and girdled (subsequently fallen) Norway spruce (Picea abies (L.) H. Karst.) logs differ from naturally formed spruce logs of similar decay stage and size. The study was conducted in Leivonmäki National Park in central Finland 8 years after the restoration measures. The average number of polypore species was highest on the chain-saw felled logs and most of the common polypore species were most frequent on this substrate. However, among the natural logs, number of species increased more steeply with increasing number of logs, suggesting greater variation in community composition on this substrate. The old-growth forest indicator Phellinus ferrugineofuscus occurred frequently on natural logs, occupied a few girdled logs but was absent from chain-saw felled logs. Our results show that from the polypore perspective created logs do not fully mimic natural logs, suggesting that creating substrates for species may pose a challenge for restoration.
  • Komonen, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland ORCID ID:E-mail: atte.komonen@jyu.fi (email)
  • Halme, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland ORCID ID:E-mail: panu.halme@jyu.fi
  • Jäntti, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland ORCID ID:E-mail: mari.j.jantti@student.jyu.fi
  • Koskela, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland ORCID ID:E-mail: tuuli.e.koskela@student.jyu.fi
  • Kotiaho, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland ORCID ID:E-mail: janne.kotiaho@jyu.fi
  • Toivanen, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland; Current: Birdlife Finland, Annankatu 29 A 16, FI-00100 Helsinki, Finland ORCID ID:E-mail: tero.toivanen@birdlife.fi
article id 949, category Research article
Ken Olaf Storaunet, Jørund Rolstad, Erlend Rolstad. (2014). Effects of logging on the threatened epiphytic lichen Usnea longissima: an experimental approach. Silva Fennica vol. 48 no. 1 article id 949. https://doi.org/10.14214/sf.949
Highlights: A re-inventory of the threatened lichen Usnea longissima in ten Norway spruce forest stands where experimental selective loggings had been conducted 5 to 8 years before revealed that the number of lichen thalli had increased with 34%; The number of thalli increased more where the forest was open whether or not the low tree density was caused by the loggings.
Usnea longissima Ach. is a circumboreal epiphytic lichen draping tree canopies in moist coastal and mountainous forests. It is extinct from many European and North-American localities, presumably due to industrial forestry and air pollution, but still has a stronghold in parts of Scandinavia and U.S. and Canadian Pacific Northwest. In 2005/06 we used a comparative and retrospective approach to evaluate how present and historic tree and stand characteristics influenced the occurrence and abundance of the lichen (Storaunet et al. 2008). In 2012, we re-inventoried ten Norway spruce forest stands with 401 U. longissima-bearing trees and recorded changes in the number of U. longissima thalli. Seven of the stands had been experimentally, selectively logged 5–8 years before, where the lichen-bearing trees had been marked in the field and were avoided during the logging operation. Total number of lichen-bearing trees decreased slightly (2.9%), whereas the number of thalli had increased with 34%. Number of thalli increased more where the forest was open (low basal area, m2ha-1) whether or not the low tree density was caused by the logging events. At high tree densities the change in number of thalli was negligible. We suggest that selective logging, securing lichen-bearing trees, may be a viable management option to keep tree density from becoming too dense, thereby enhancing growth and establishment of U. longissima.
  • Storaunet, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway ORCID ID:E-mail: stk@skogoglandskap.no (email)
  • Rolstad, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway ORCID ID:E-mail: roj@skogoglandskap.no
  • Rolstad, Skogfaglig Rådgivning, Holmsida 126, NO-1488 Hakadal, Norway ORCID ID:E-mail: roe@skogoglandskap.no
article id 1071, category Research article
Ursula Kretschmer, Nadeschda Kirchner, Christopher Morhart, Heinrich Spiecker. (2013). A new approach to assessing tree stem quality characteristics using terrestrial laser scans. Silva Fennica vol. 47 no. 5 article id 1071. https://doi.org/10.14214/sf.1071
Highlights: Minimal deviations of the bark surface can be detected and visualized based on terrestrial laser scan data; Additionally the geometrical properties of bark scars and branched knots can be assessed; Two methods using two different approaches are presented: (1) a method using intensity data and (2) a method using bark surface models.
This paper presents an approach to assess and measure bark characteristics as indicators of wood quality using terrestrial laser scan data. In addition to the detection and measurement by use of the intensity information of the scan data a new approach was established. Bark surface models are calculated for each tree. They offer the representation of the bark as a height model. The reference is the tree stem approximated by a chain of cylinders. Minimal deviations of the bark surface can be detected and visualized and the geometrical properties of bark scars and branched knots can be assessed. Results of the measurement of 18 scars are presented using the two approaches: (1) a method using intensity data or (2) using bark surface models. The selection of the adequate approach depends on the stem characteristics. In a next step, methods for automatic measurement of bark scars will be developed.
  • Kretschmer, Chair of Forest Growth, Albert-Ludwigs-University Freiburg, Tennenbacher Str. 4, 79106 Freiburg, Germany ORCID ID:E-mail: ursula.kretschmer@iww.uni-freiburg.de (email)
  • Kirchner, VOLKE Consulting Engineers GmbH, Schätzweg 7-9, 80935 München, Germany ORCID ID:E-mail: nadeschda.kirchner@volke.muc.de
  • Morhart, Chair of Forest Growth, Albert-Ludwigs-University Freiburg, Tennenbacher Str. 4, 79106 Freiburg, Germany ORCID ID:E-mail: christopher.morhart@iww.uni-freiburg.de
  • Spiecker, Chair of Forest Growth, Albert-Ludwigs-University Freiburg, Tennenbacher Str. 4, 79106 Freiburg, Germany ORCID ID:E-mail: instww@uni-freiburg.de
article id 937, category Research article
Rene Zamora-Cristales, Kevin Boston, John Sessions, Glen Murphy. (2013). Stochastic simulation and optimization of mobile chipping economics in processing and transport of forest biomass from residues. Silva Fennica vol. 47 no. 5 article id 937. https://doi.org/10.14214/sf.937
Highlights: A stochastic simulation model is proposed to analyze forest biomass operations; The cost of chipper and truck waiting times was estimated in forest biomass recovery operations; The economic effect of truck-machine interactions under uncertainty was analyzed; Road characteristics and processing location have an economic impact in truck and chipper waiting times
We analyzed the economics of mobile chipping and transport of biomass from forest residues for energy purposes under uncertainty. A discrete-event simulation model was developed and utilized to quantify the impacts of controllable and environmental variables on productivity in order to determine the most cost effective transportation options under steep terrain conditions. Truck-chipper interactions were analyzed to show their effect on truck and chipper standing time. A costing model was developed to account for operating and standing time cost (for the chipper and trucks). The model used information from time studies of each activity in the productive cycle and spatial-temporal information obtained from geographic information system (GIS) devices, and tracking analysis of machine and truck movements. The model was validated in field operations, and proved to be accurate in providing the expected productivity. A cost distribution was elaborated to support operational decisions of forest managers, landowners and risk-averse contractors. Different scenarios were developed to illustrate the economic effects due to changes in road characteristics such as in-highway transport distance, in-forest internal road distance and pile to trailer chipper traveling distances.
  • Zamora-Cristales, Department of Forest Engineering, Resources, and Management, College of Forestry, Oregon State University, 280 Peavy Hall, Corvallis, OR 97331, USA ORCID ID:E-mail: rene.zamora@oregonstate.edu (email)
  • Boston, Department of Forest Engineering, Resources, and Management, College of Forestry, Oregon State University, 280 Peavy Hall, Corvallis, OR 97331, USA ORCID ID:E-mail: kevin.boston@oregonstate.edu
  • Sessions, Department of Forest Engineering, Resources, and Management, College of Forestry, Oregon State University, 280 Peavy Hall, Corvallis, OR 97331, USA ORCID ID:E-mail: john.sessions@oregonstate.edu
  • Murphy, Waiariki Institute of Technology, Rotorua, New Zealand ORCID ID:E-mail: glen.murphy@waiariki.ac.nz
article id 937, category Research article
Rene Zamora-Cristales, Kevin Boston, John Sessions, Glen Murphy. (2013). Stochastic simulation and optimization of mobile chipping economics in processing and transport of forest biomass from residues. Silva Fennica vol. 47 no. 5 article id 937. https://doi.org/10.14214/sf.937
Highlights: A stochastic simulation model is proposed to analyze forest biomass operations; The cost of chipper and truck waiting times was estimated in forest biomass recovery operations; The economic effect of truck-machine interactions under uncertainty was analyzed; Road characteristics and processing location have an economic impact in truck and chipper waiting times
We analyzed the economics of mobile chipping and transport of biomass from forest residues for energy purposes under uncertainty. A discrete-event simulation model was developed and utilized to quantify the impacts of controllable and environmental variables on productivity in order to determine the most cost effective transportation options under steep terrain conditions. Truck-chipper interactions were analyzed to show their effect on truck and chipper standing time. A costing model was developed to account for operating and standing time cost (for the chipper and trucks). The model used information from time studies of each activity in the productive cycle and spatial-temporal information obtained from geographic information system (GIS) devices, and tracking analysis of machine and truck movements. The model was validated in field operations, and proved to be accurate in providing the expected productivity. A cost distribution was elaborated to support operational decisions of forest managers, landowners and risk-averse contractors. Different scenarios were developed to illustrate the economic effects due to changes in road characteristics such as in-highway transport distance, in-forest internal road distance and pile to trailer chipper traveling distances.
  • Zamora-Cristales, Department of Forest Engineering, Resources, and Management, College of Forestry, Oregon State University, 280 Peavy Hall, Corvallis, OR 97331, USA ORCID ID:E-mail: rene.zamora@oregonstate.edu (email)
  • Boston, Department of Forest Engineering, Resources, and Management, College of Forestry, Oregon State University, 280 Peavy Hall, Corvallis, OR 97331, USA ORCID ID:E-mail: kevin.boston@oregonstate.edu
  • Sessions, Department of Forest Engineering, Resources, and Management, College of Forestry, Oregon State University, 280 Peavy Hall, Corvallis, OR 97331, USA ORCID ID:E-mail: john.sessions@oregonstate.edu
  • Murphy, Waiariki Institute of Technology, Rotorua, New Zealand ORCID ID:E-mail: glen.murphy@waiariki.ac.nz
article id 1057, category Research article
Jouni Siipilehto, Lauri Mehtätalo. (2013). Parameter recovery vs. parameter prediction for the Weibull distribution validated for Scots pine stands in Finland. Silva Fennica vol. 47 no. 4 article id 1057. https://doi.org/10.14214/sf.1057
Highlights: A parameter recovery method (PRM) was developed for forest stand inventories and compared with previously developed parameter prediction methods (PPM) in Finland; PRM for the 2-parameter Weibull function provided compatibility for the main stand characteristics: stem number, basal area and one of the four optional mean characteristics; PRM provided comparable and at its best, superior accuracy in volume characteristics compared with PPM.
The moment-based parameter recovery method (PRM) has not been applied in Finland since the 1930s, even after a continuation of forest stand structure modelling in the 1980s. This paper presents a general overview of PRM and some useful applications. Applied PRM provided compatibility for the included stand characteristics of stem number (N) and basal area (G) with either mean (D), basal area-weighted mean (DG), median (DM) or basal area-median (DGM) diameter at breast height (dbh). A two-parameter Weibull function was used to describe the dbh-frequency distribution of Scots pine stands in Finland. In the validation, PRM was compared with existing parameter prediction models (PPMs). In addition, existing models for stand characteristics were used for the prediction of unknown characteristics. Validation consisted of examining the performance of the predicted distributions with respect to variation in stand density and accuracy of the localised distributions, as well as accuracy in terms of bias and the RMSE in stand characteristics in the independent test data set. The validation data consisted of 467 randomly selected stands from the National Forest Inventory based plots. PRM demonstrated excellent accuracy if G and N were both known. At its best, PRM provided accuracy that was superior to any existing model in Finland – especially in young stands (mean height < 9 m), where the RMSE in total and pulp wood volumes, 3.6 and 5.7%, respectively, was reduced by one-half of the values obtained using the best performing existing PPM (8.7–11.3%). The unweighted Weibull distribution solved by PRM was found to be competitive with weighted existing PPMs for advanced stands. Therefore, using PRM, the need for a basal area weighted distribution proved unnecessary, contrary to common belief. Models for G and N were shown to be unreliable and need to be improved to obtain more reliable distributions using PRM.
  • Siipilehto, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland ORCID ID:E-mail: jouni.siipilehto@metla.fi (email)
  • Mehtätalo, University of Eastern Finland, School of Computing, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: lauri.mehtatalo@uef.fi
article id 1046, category Research article
Eva-Maria Nordström, Hampus Holmström, Karin Öhman. (2013). Evaluating continuous cover forestry based on the forest owner’s objectives by combining scenario analysis and multiple criteria decision analysis. Silva Fennica vol. 47 no. 4 article id 1046. https://doi.org/10.14214/sf.1046
Highlights: Scenario analysis and multiple criteria decision analysis were combined to evaluate alternative forest management strategies for Linköping municipality, Sweden; Continuous cover forestry (CCF) promoted ecological and social objectives better than even-aged forestry but was worse for economic objectives; Ecological and social objectives were important to the municipality and thus, in summary, CCF seemed to be a suitable strategy.
Forests are increasingly managed both to provide a sustainable yield of timber and for supplying a range of ecosystem services in line with the concept of sustainable forest management. Several incommensurable interests must then be considered, and it is necessary to strike a balance between different objectives. In evaluation of trade-offs to be made, both objective factors and subjective values need to be taken into account. In recent years, continuous cover forestry (CCF) has been put forward as an alternative to even-aged forestry. The aim of this study was to use scenario analysis in combination with multi criteria decision analysis (MCDA) to evaluate whether CCF is a suitable strategy based on the decision makers’ objectives and preferences for sustainable forest management in a specific landscape. This approach was applied to a planning case on the forest estate of the Linköping municipality in southwestern Sweden. The scenario analyses provided insights into relevant quantitative factors, while the MCDA evaluation helped in clarifying the objectives of the forest management and in assessing the relative importance of various objectives. The scenario analyses showed that in this case CCF is a good management strategy in ecological and social terms but yields worse economic outcomes than conventional even-aged forestry. In the Linköping case, there was a relatively strong emphasis on ecological and social aspects and thus, in summary, CCF seemed to be the most suitable option.
  • Nordström, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd, SE-901 83 Umeå, Sweden ORCID ID:E-mail: eva-maria.nordstrom@slu.se (email)
  • Holmström, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd, SE-901 83 Umeå, Sweden ORCID ID:E-mail: hampus.holmstrom@slu.se
  • Öhman, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd, SE-901 83 Umeå, Sweden ORCID ID:E-mail: karin.ohman@slu.se
article id 1009, category Research article
Inger Sundheim Fløistad, Aksel Granhus. (2013). Timing and duration of short-day treatment influence morphology and second bud flush in Picea abies seedlings. Silva Fennica vol. 47 no. 3 article id 1009. https://doi.org/10.14214/sf.1009
Highlights: The duration of short-day treatment, calculated as number of days, influenced the root collar diameter growth more than the timing of the treatment; If short-day treatment starts early in summer, a longer duration of the treatment is recommended to avoid second bud flush.
A slower reaction of diameter growth cessation compared to that of height growth in response to short day (SD) treatment is well documented in Picea abies (L.) Karst. seedlings, suggesting that the height/diameter ratio of seedlings could be controlled through appropriate timing and/or duration of SD treatment is forest nurseries. Here, we applied specific combinations of timing (starting date 20 and 27 June, 4 or 11 July) and duration (7, 10, 14 or 17 days) of SD treatment to assess the possibility of obtaining more sturdy seedlings. We observed a rapid and uniform height growth cessation following SD treatment compared with the delayed cessation of diameter growth. Height growth responded significantly only to starting date of SD treatment, resulting in taller seedlings for later starting dates. Diameter growth responded to the duration of SD treatment, with significantly less diameter growth in seedlings exposed to 14 or 17 days of SD treatment than in seedlings exposed to 7 or 10 days of SD treatment. Also starting date influenced diameter growth, resulting in significantly more diameter growth with the earliest starting date compared with the two latest starting dates of the SD treatment.  A second bud flush occurred only in seedlings exposed to SD treatment starting on 20 or 27 of June and only following 7-14 days of duration. This implies a need of longer duration if the SD treatment starts early. This will be at the expense of sustained diameter growth, thus compromising the objective of obtaining more sturdy seedlings.
  • Fløistad, Norwegian Institute for Agricultural and Environmental Research, Høgskolevn 7, N-1430 Ås, Norway & Norwegian Forest and Landscape Institute, P.O. Box 115, N-1431 Ås, Norway ORCID ID:E-mail: isf@skogoglandskap.no (email)
  • Granhus, Norwegian Forest and Landscape Institute, P.O. Box 115, N-1431 Ås, Norway ORCID ID:E-mail: aksel.granhus@skogoglandskap.no
article id 973, category Research article
Sergio Rossi, Hubert Morin, François Gionest, Danielle Laprise. (2013). Spatially explicit structure of natural stands dominated by black spruce. Silva Fennica vol. 47 no. 3 article id 973. https://doi.org/10.14214/sf.973
Black spruce [Picea mariana (Mill.) BSP] regeneration emerges in clusters near the pre-existing boles within a few years after the passage of fire. This paper tested the hypothesis that black spruce forests still maintain the spatial structure deriving from postfire stand initiation. Trees and saplings were monitored during 2000-2007 and the horizontal and vertical structure of the stands were investigated on four permanent plots in the boreal forest of Quebec, Canada. Plots showed 1300-2150 trees ha-1, and were composed of trees with homogeneous sizes and a very small proportion of saplings. These characteristics identify single cohorts generated by complete, or almost-complete, stand replacement by fire. Ripley’s L(r) functions showed that the spatial pattern of trees and saplings ranged from random to aggregated, thus demonstrating that the clustering distribution of the individuals in black spruce forests can be maintained even after 80-120 years from stand initiation. These findings could results from incomplete self-thinning or from an environment with heterogeneous distribution of resources. The practices of ecosystem management recently developed in Eastern Canada should take into account both the horizontal and vertical structure to better modulate the competition among individuals during partial harvesting.
  • Rossi, Département des Sciences Fondamentales, Université du Québec à Chicoutimi, 555 boulevard de l’Université, Chicoutimi (QC), G7H2B1, Canada ORCID ID:E-mail: sergio.rossi@uqac.ca (email)
  • Morin, Département des Sciences Fondamentales, Université du Québec à Chicoutimi, 555 boulevard de l’Université, Chicoutimi (QC), G7H2B1, Canada ORCID ID:E-mail: hubert_morin@uqac.ca
  • Gionest, Département des Sciences Fondamentales, Université du Québec à Chicoutimi, 555 boulevard de l’Université, Chicoutimi (QC), G7H2B1, Canada ORCID ID:E-mail: francois_gionest@uqac.ca
  • Laprise, Département des Sciences Fondamentales, Université du Québec à Chicoutimi, 555 boulevard de l’Université, Chicoutimi (QC), G7H2B1, Canada ORCID ID:E-mail: danielle_laprise@uqac.ca
article id 958, category Research article
Back Tomas Ersson, Linus Jundén, Urban Bergsten, Martin Servin. (2013). Simulated productivity of one- and two-armed tree planting machines. Silva Fennica vol. 47 no. 2 article id 958. https://doi.org/10.14214/sf.958
Highlights: Using discrete-event simulation and detailed terrain and machine models, the productivities of excavator-based one- and two-armed tree planting machines were simulated; The machines’ arms were equipped with one-and two-headed planting devices; Two planting heads per arm rather than two arms per base machine is better for increasing the productivity of intermittently advancing planting machines on Nordic clearcuts.
To increase mechanized planting, planting machine productivity must increase in order to improve cost-efficiency. To determine if excavators with two crane arms could potentially help to increase planting machine productivity under Nordic clearcut conditions, we modelled one-armed and semi-automated two-armed excavators with one- and two-headed planting devices. Using a recently developed tool for discrete-event simulation, these machine models then mounded and planted seedlings on terrain models with moraine soil having various frequencies of obstacles (stumps, roots and stones). Compared to if the two heads were mounted pairwise on only one arm, the results showed that productivity did not increase if two planting heads were attached individually to two separate crane arms. But productivity did increase if the planting machine had four planting heads mounted pairwise on two separate arms. However, despite assuming automated mounding and crane motion between planting spots, the two-armed, four-headed model never achieved high enough productivity levels to make it more cost-efficient than one-armed machines. The simulations illustrate that our terrain models generate realistic root architecture and boulder content distributions in moraine soil, while our machine models functionally describe mechanized planting work. Based on our assumptions, we conclude that further development work on two-armed excavator-based planting machines for Nordic clearcut conditions is not warranted. Our simulations reveal that increasing the number of planting heads per crane arm rather than number of crane arms per base machine offers the greatest potential to raise the productivity of intermittently advancing planting machines.
  • Ersson,  Swedish University of Agricultural Sciences, Department of Forest Biomaterials and Technology, SE-901 83 Umeå, Sweden ORCID ID:E-mail: back.tomas.ersson@slu.se (email)
  • Jundén,  UMIT Research Lab, Umeå University, SE-901 87 Umeå, Sweden ORCID ID:E-mail: linus.junden@gmail.com
  • Bergsten,  Swedish University of Agricultural Sciences, Department of Forest Biomaterials and Technology, SE-901 83 Umeå, Sweden ORCID ID:E-mail: urban.bergsten@slu.se
  • Servin,  UMIT Research Lab, Umeå University, SE-901 87 Umeå, Sweden ORCID ID:E-mail: martin.servin@physics.umu.se
article id 954, category Research article
Anna-Maria Eriksson, Jörgen Olsson, Bengt Gunnar Jonsson, Sara Toivanen, Mattias Edman. (2013). Effects of restoration fire on dead wood heterogeneity and availability in three Pinus sylvestris forests in Sweden. Silva Fennica vol. 47 no. 2 article id 954. https://doi.org/10.14214/sf.954
Restoration fires are increasingly used as a conservation tool in Sweden to recreate forests with characteristics of previous forests that were periodically disturbed by fires and promote fire-dependent species. Restoration fires can result in large inputs of fresh dead wood, but there are risks of losing some of the existing, pre-fire dead wood. To assess these counteracting effects we studied the heterogeneity and availability of dead wood before and after three restoration fires in boreal Scots pine forests. Specifically, we studied volumes of stumps, high stumps, snags and logs. The fires decreased the total volume of pre-fire dead wood (23-41%) and consumed logs in late decay stages (26-54%) to a higher extent than logs in earlier stages. The input of new fresh dead wood after the fires exceeded losses of pre-fire dead wood and resulted in a net increase of dead wood in all three sites. The added dead wood consisted of fresh snags killed by the fires. Fire also affected log characteristics: reducing their vegetation coverage (60-98%), decreasing their ground contact (4-50%) and increasing their surface area of charred wood (>50%). Such changes have important consequences for the micro environmental conditions inside logs, but have been rarely studied in relation to restoration fires. Our results show that restoration fire causes changes in dead wood availability and characteristics of logs. The results imply that ideally stands with low abundance of rare and heavily decayed wood substrates should be burned to optimize dead wood values. Alternatively, management practices should include protection of these substrates during restoration fires.
  • Eriksson, Department of Natural Sciences, Mid Sweden University, SE-851 70, Sundsvall, Sweden ORCID ID:E-mail: anna-maria.eriksson@miun.se (email)
  • Olsson, Department of Ecology and Environmental Science, Umeå University, SE-901 87, Umeå, Sweden ORCID ID:E-mail: jorgen.m.olsson@slu.se
  • Jonsson, Department of Natural Sciences, Mid Sweden University, SE-851 70, Sundsvall, Sweden ORCID ID:E-mail: bengt-gunnar.jonsson@miun.se
  • Toivanen, Department of Natural Sciences, Mid Sweden University, SE-851 70, Sundsvall, Sweden ORCID ID:E-mail: sara.toivanen@lansstyrelsen.se
  • Edman, Department of Natural Sciences, Mid Sweden University, SE-851 70, Sundsvall, Sweden ORCID ID:E-mail: mattias.edman@miun.se
article id 952, category Research article
Lauri Korhonen, Inka Pippuri, Petteri Packalén, Ville Heikkinen, Matti Maltamo, Juho Heikkilä. (2013). Detection of the need for seedling stand tending using high-resolution remote sensing data. Silva Fennica vol. 47 no. 2 article id 952. https://doi.org/10.14214/sf.952
Seedling stands are problematic in airborne laser scanning (ALS) based stand level forest management inventories, as the stem density and species proportions are difficult to estimate accurately using only remotely sensed data. Thus the seedling stands must still be checked in the field, which results in an increase in costs. In this study we tested an approach where ALS data and aerial images are used to directly classify the seedling stands into two categories: those that involve tending within the next five years and those which involve no tending. Standard ALS-based height and density features, together with texture and spectral features calculated from aerial images, were used as inputs to two classifiers: logistic regression and the support vector machine (SVM). The classifiers were trained using 208 seedling plots whose tending need was estimated by a local forestry expert. The classification was validated on 68 separate seedling stands. In the training data, the logistic model’s kappa coefficient was 0.55 and overall accuracy (OA) 77%. The SVM did slightly better with a kappa = 0.71 and an OA = 86%. In the stand level validation data, the performance decreased for both the logistic model (kappa = 0.38, OA = 71%) and the SVM (kappa = 0.37, OA = 72%). Thus our approach cannot totally replace the field checks. However, in considering the stands where the logistic model predictions had high reliability, the number of misclassifications reduced drastically. The SVM however, was not as good at recognizing reliable cases.
  • Korhonen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: lauri.korhonen@uef.fi (email)
  • Pippuri, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: inka.pippuri@uef.fi
  • Packalén, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: petteri.packalen@uef.fi
  • Heikkinen, School of Computing, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: ville.heikkinen@uef.fi
  • Maltamo, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: matti.maltamo@uef.fi
  • Heikkilä, Finnish Forest Centre, Public Services, Maistraatinportti 4 A, FI-00240 Helsinki, Finland ORCID ID:E-mail: juho.heikkila@metsakeskus.fi
article id 935, category Research article
Loice M. A. Omoro, Mike Starr, Petri K. E. Pellikka. (2013). Tree biomass and soil carbon stocks in indigenous forests in comparison to plantations of exotic species in the Taita Hills of Kenya. Silva Fennica vol. 47 no. 2 article id 935. https://doi.org/10.14214/sf.935
Carbon (C) densities of the tree biomass and soil (0–50 cm) in indigenous forest and plantations of eucalyptus, cypress and pine in the Taita Hills, Kenya were determined and compared. The cypress and pine plantations were about 30-years-old and eucalyptus plantations about 50-years-old. Biomass C densities were estimated from breast height diameter and wood density using allometric functions developed for tropical species and an assumed C content of 50%. Belowground biomass C densities were estimated using root:shoot biomass ratios. Soil organic C (SOC) densities were calculated from measured organic carbon contents (0–20 and 20–50 cm layers) and modelled bulk density values. Mean total biomass C and SOC densities for indigenous forest were greater than those of the plantations, and the difference was significant (p < 0.05) in the cases of cypress and pine biomass and pine SOC. The correlation between biomass C and SOC densities was nearly significant in the case of indigenous forest, but negative. Biomass C densities were not significantly correlated with mean annual precipitation, mean annual temperature or potential evapotranspiration, but pine biomass C densities were significantly correlated to actual evapotranspiration. SOC densities were more strongly correlated to mean annual precipitation than biomass C densities, but only significantly so in the case of pine. Neither biomass C nor SOC densities were correlated to plant available water capacity of the soil. Indigenous forest SOC densities were significantly correlated to soil clay contents, but negatively. Indigenous forests sequester more C in biomass and soil than do 30 to 50-year-old plantations of exotics, but it remains unclear if this is an intrinsic difference between indigenous forest and plantations of exotics or because of insufficient time for SOC levels in plantations to recover after clearance of original indigenous forest.
  • Omoro, Viikki Tropical Resources Institute, Department of Forest Sciences, P.O. Box 27 (Latokartanonkaari 7), FI-00014 University of Helsinki, Finland ORCID ID:E-mail: loice.omoro@helsinki.fi
  • Starr, Department of Forest Sciences, P. O. Box 27 (Latokartanonkaari 7), FI-00014 University of Helsinki, Finland ORCID ID:E-mail: mike.starr@helsinki.fi (email)
  • Pellikka, Department of Geosciences and Geography, P. O. Box 64 (Gustaf Hällströminkatu 2), FI-00014 University of Helsinki, Finland ORCID ID:E-mail: petri.pellikka@helsinki.fi
article id 901, category Research article
Luis A. Apiolaza, Rosa M. Alzamora. (2013). Building deployment portfolios for genotypes under performance instability. Silva Fennica vol. 47 no. 1 article id 901. https://doi.org/10.14214/sf.901
We used portfolio theory to analyze the tradeoffs between returns and performance instability of deployment units for Pinus radiata D. Don. We considered three groups of 34 trees each grown to produce appearance lumber, structural lumber, or both. Risk was based on the variability of tree returns in scenarios of changing volume, wood stiffness and presence of resin defects due to genotype by environment interaction inducing both changes of scale and differential tree response to environmental scenarios. The return of structural trees was highly variable with a mean of 3.11 NZ $/stem/year, followed by appearance-structural trees (3.48 NZ $/stem/year). In contrast, appearance trees had the lowest returns (1.99 NZ $/stem/year) and variability. The portfolio model selected structural trees in high-risk scenarios, but selection was apportioned between structural and appearance-structural trees as the risk decreased. The model selected only appearance trees for high-risk aversion. The analysis also considered silvicultural regimes, where the appearance-structural regime was selected under high variability. As risk decreased the appearance grades regime was also selected. The structural regime was rarely selected due to the variability of stiffness between trees. Using genotypes improved for stiffness could increase the expected value and reduce variability for structural purposes, making the structural regime more appealing.
  • Apiolaza, School of Forestry, University of Canterbury, Private Bag 4800, 8042 Canterbury, New Zealand ORCID ID:E-mail: luis.apiolaza@canterbury.ac.nz (email)
  • Alzamora, Instituto de Manejo Forestal, Universidad Austral, Valdivia, Chile ORCID ID:E-mail: ralzamor@uach.cl
article id 894, category Research article
Heimo Karppinen, Mika Korhonen. (2013). Do forest owners share the public’s values? An application of Schwartz’s value theory. Silva Fennica vol. 47 no. 1 article id 894. https://doi.org/10.14214/sf.894
The purpose of this study is to investigate the differences in value priorities between Finnish forest owners and the general public. A conclusion is drawn whether and to what extent value changes in society are reflected in forest owners’ values and objectives, and, finally, in their actual forestry behavior. In addition, the study highlights the differences in value priorities among forest owners in various demographic groups. The data set used in this study was based on a nationwide mail survey on Finnish non-industrial private forest owners conducted in 2009 and consisting of 2116 observations of forest owners. Schwartz’s value theory was a good fit for testing the value priorities of forest owners. The three most important values were benevolence, security and conformity, both among the forest owners and the public. Tradition was ranked the fourth most important value by the forest owners, but very low by the public. The forest owners ranked universalism slightly lower than the public in general. This difference was clearly greater when the female forest owners were compared to women in the whole population. The probability of a forest owner belonging to the Softies (high emphasis on universalism and benevolence) increased with age and was higher for the female owners and the owners with recreational or multiple objectives compared to the indifferent owners. The multiobjective owners and recreationists had relatively similar value profiles. The previous literature suggests that multiobjective owners are the most active forest owner group and that recreationists and indifferent owners are the most passive groups in their timber supply behavior. The relationship between values and forestry behavior thus remains ambiguous.
  • Karppinen, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID ID:E-mail: heimo.karppinen@helsinki.fi (email)
  • Korhonen, Kämnerintie 7e 41, FI-00750 Helsinki, Finland ORCID ID:E-mail: mika.korhonen@gmx.com
article id 924, category Research article
Minna Pulkkinen. (2012). On non-circularity of tree stem cross-sections: effect of diameter selection on cross-section area estimation, Bitterlich sampling and stem volume estimation in Scots pine. Silva Fennica vol. 46 no. 5B article id 924. https://doi.org/10.14214/sf.924
In the common methods of forest mensuration, including stem volume models and Bitterlich sampling, stem cross-sections are assumed to be circular. In nature this assumption is never exactly fulfilled. Errors due to non-circularity have been presumed to be small and unimportant but studied little: theoretical and empirical studies exist on cross-section area estimation, but errors in stem volume estimation have not been investigated at all, and errors in Bitterlich sampling are theoretically known only for stand basal area estimation. In the theoretical part of this study, we developed methods for quantifying the systematic and sampling errors that 22 common ways of selecting diameter within non-circular cross-sections induce (i) in area estimates by the circle area formula, (ii) in stand total estimates by Bitterlich sampling, and (iii) in stem volume estimates by a volume equation, by a cubic-spline-interpolated stem curve, and by a generalised volume estimator. In the empirical part, based on the digital images of 709 discs taken at 6–10 heights in 81 Scots pine stems from different parts of Finland, we investigated the variation in cross-section shape, and demonstrated the magnitude of the errors presented in the theoretical part. We found that non-circularity causes systematic overestimation of area and volume, and inflicts potentially systematic error on stand total estimates by Bitterlich sampling. In our data these effects were small, but the finding is not generalisable due the skewed size distribution and poor geographical representativeness of the data. We recommend using diameter derived from girth for both tree and stand level estimation, as it involves no sampling error and produces clearly the most stable systematic errors.
  • Pulkkinen, ORCID ID:E-mail: minna.pulkkinen@iki.fi (email)
article id 919, category Research article
Karri Uotila, Juho Rantala, Timo Saksa. (2012). Estimating the need for early cleaning in Norway spruce plantations in Finland. Silva Fennica vol. 46 no. 5 article id 919. https://doi.org/10.14214/sf.919
Effective management of Norway spruce (Picea abies (L.) Karst.) plantations requires detailed information on stand development, which is costly to measure. However, estimating the need for early stand management from site attributes that persists stabile after ones measured, may provide an inexpensive alternative. This study compared hardwood competition in spruce plantations of varying ages and tested the usability of this information in estimating the need for early cleaning. The data included 197 spruce plantations (4–7 years old) inventoried in southern Finland in 2007. The level (Low, Substantial, High) of need for early cleaning was subjectively determined by contrasting location and size of competing hardwoods to a conifer crop tree. Then the stage of the need for early cleaning was modelled according to site and stand attributes. Nearly 60% of the conifer crop trees in the plantations were subjectively judged to require early cleaning (Substantial 37.2%, High 21.2%), but only 10 per cent of the evaluated area was cleaned. Need for cleaning was intense on peatlands or damp soils, whereas it was mild on unprepared soils or cleaned sites. Traditional site characteristics used in forest management planning can be useful for recognising the peripheral cases, where need for cleaning is probably high or low. However, on a typical mineral soil plantation (uncleaned, soil prepared) the model indicates the differences in the need for early cleaning weakly. The need for early cleaning was already high in 4-year-old plantations, why stand age did not have significant effect on development of the need. Thus, the timing of an operation can not be predicted with the model. Nonetheless, early cleaning very likely opens growth space of crop trees in a 4–7-year-old spruce plantation. Therefore, from an aspect of crop growth, an uncleaned Norway spruce plantation in this age group is quite consistently worth cleaning.
  • Uotila, Finnish Forest Research Institute, Suonenjoki Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: karri.uotila@metla.fi (email)
  • Rantala, Metsä Group, Tampere, Finland ORCID ID:E-mail: juho.rantala@metsagroup.com
  • Saksa, Finnish Forest Research Institute, Suonenjoki Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: timo.saksa@metla.fi
article id 917, category Research article
Anna Gunulf, Rebecka Mc Carthy, Jonas Rönnberg. (2012). Control efficacy of stump treatment and influence of stump height on natural spore infection by Heterobasidion spp. of precommercial thinning stumps of Norway spruce and birch. Silva Fennica vol. 46 no. 5 article id 917. https://doi.org/10.14214/sf.917
An alternative precommercial thinning technique results in trees being cut higher up the stem compared to the normal method using a brush saw. The aims of this study were to investigate if primary infection of Heterobasidion spp. on precommercial thinning stumps of Norway spruce and birch is influenced by stump height and to test the control efficacy of stump treatment with Phlebiopsis gigantea on precommercial thinning stumps of Norway spruce. Small Norway spruce and birch trees were felled on five sites in southern Sweden and their stumps subjected to natural spore infection. For each species, two treatments of stump height were created: 15 and 100 cm. Half of the Norway spruce stumps were treated with P. gigantea. After two months, 896 stumps were sampled and infection by Heterobasidion spp. was quantified. The height of stumps did not significantly influence infection frequency or size of infection on either tree species. Untreated Norway spruce stumps had an infection frequency of 55% while 31% of the treated stumps were infected. The control efficacy of stump treatment in terms of reduced relative infected area was 61–65%. The area occupied by Heterobasidion spp. on birch stumps was generally small, on average 0.4 cm2 per infected stump, although 15% of the stumps were infected. The risk of primary infection in Norway spruce dominated stands should be considered when precommercial thinning is conducted but the control efficacy and economy of stump treatment warrants further investigation before practical recommendations can be made.
  • Gunulf, Swedish University of Agricultural Sciences, Department of Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53 Alnarp, Sweden ORCID ID:E-mail: anna.gunulf@slu.se (email)
  • Mc Carthy, Skogsforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden ORCID ID:E-mail: Rebecka.McCarthy@skogforsk.se
  • Rönnberg, Swedish University of Agricultural Sciences, Department of Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53 Alnarp, Sweden ORCID ID:E-mail: jonas.ronnberg@slu.se
article id 913, category Research article
Han Zhang, Joseph Buongiorno. (2012). Markets, government policy, and China's timber supply. Silva Fennica vol. 46 no. 4 article id 913. https://doi.org/10.14214/sf.913
China's domestic demand and exports of wood products are rising rapidly compared to domestic supply. The determinants of timber supply in China were investigated with panel data from 25 provinces from 1999 to 2009. The results indicated that China’s timber supply had responded to both market forces, reflected by timber prices largely determined by world demand and supply, and to government policies expressed by production quotas and the tenure reform on collective forestland. The price elasticity of China’s timber supply was estimated at 0.31±0.12. The inelastic response of production to the quota (0.20±0.02) suggested that government had a limited, though significant, control of timber supply. Other things being equal, the land tenure reform increased timber supply by 18±8 percent, where and when it had been implemented.
  • Zhang, Research Center for Resource Economics and Environment Management, Northwest A&F University. No. 3 Taicheng Road, Yangling, Shaanxi, China 712100 ORCID ID:E-mail:
  • Buongiorno, Department of Forest and Wildlife Ecology, University of Wisconsin-Madison, Madison, WI 53706, U.S.A. ORCID ID:E-mail: jbuongio@wisc.edu (email)
article id 912, category Research article
Heidi Hallongren, Juho Rantala. (2012). Commercialisation and international market potential of Finnish silvicultural machines. Silva Fennica vol. 46 no. 4 article id 912. https://doi.org/10.14214/sf.912
Recently, the need to mechanise silvicultural operations has increased in the Nordic countries. While several new machine concepts have been developed, the manufacture of silvicultural devices remains at the market introduction stage. Information is required in support of existing and forthcoming Finnish small-scale producers of silvicultural devices, who wish to commercialise and further market their innovations in domestic and export markets. The aim of this study was to identify the opportunities, challenges and market potential of business activities that develop in connection with device or machine production. Small-scale Finnish manufacturers of silvicultural devices, Finnish large-scale harvester manufacturers and international silvicultural experts participated in the study. The results show that participant groups have varying opinions of cooperation methods and export activities, as well as of the export markets with the best potential. According to international silvicultural experts, mechanised planting and pre-commercial thinning have the greatest potential worldwide. However, demand for mechanised pre-commercial thinning and planting has been mainly confined to the Nordic countries. For a foreign firm marketing a new silvicultural machine concept, the most important customers and cooperation partners are locally operating forest firms, machine contractors and research organisations. The results of the study provide a useful overview of the current state of silvicultural device manufacturing in Finland.
  • Hallongren, Finnish Forest Research Institute, Suonenjoki Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: heidi.hallongren@metla.fi (email)
  • Rantala, Metsä Group, Tampere, Finland ORCID ID:E-mail: juho.rantala@metsagroup.com
article id 912, category Research article
Heidi Hallongren, Juho Rantala. (2012). Commercialisation and international market potential of Finnish silvicultural machines. Silva Fennica vol. 46 no. 4 article id 912. https://doi.org/10.14214/sf.912
Recently, the need to mechanise silvicultural operations has increased in the Nordic countries. While several new machine concepts have been developed, the manufacture of silvicultural devices remains at the market introduction stage. Information is required in support of existing and forthcoming Finnish small-scale producers of silvicultural devices, who wish to commercialise and further market their innovations in domestic and export markets. The aim of this study was to identify the opportunities, challenges and market potential of business activities that develop in connection with device or machine production. Small-scale Finnish manufacturers of silvicultural devices, Finnish large-scale harvester manufacturers and international silvicultural experts participated in the study. The results show that participant groups have varying opinions of cooperation methods and export activities, as well as of the export markets with the best potential. According to international silvicultural experts, mechanised planting and pre-commercial thinning have the greatest potential worldwide. However, demand for mechanised pre-commercial thinning and planting has been mainly confined to the Nordic countries. For a foreign firm marketing a new silvicultural machine concept, the most important customers and cooperation partners are locally operating forest firms, machine contractors and research organisations. The results of the study provide a useful overview of the current state of silvicultural device manufacturing in Finland.
  • Hallongren, Finnish Forest Research Institute, Suonenjoki Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: heidi.hallongren@metla.fi (email)
  • Rantala, Metsä Group, Tampere, Finland ORCID ID:E-mail: juho.rantala@metsagroup.com
article id 907, category Research article
Chunyu Zhang, Yazhou Zhao, Xiuhai Zhao, Klaus von Gadow. (2012). Species-habitat associations in a northern temperate forest in China. Silva Fennica vol. 46 no. 4 article id 907. https://doi.org/10.14214/sf.907
This contribution identifies species-habitat associations in a temperate forest in north-eastern China, based on the assumption that habitats are spatially autocorrelated and species are spatially aggregated due to limited seed dispersal. The empirical observations were obtained in a large permanent experimental area covering 660 x 320 m. The experimental area was subdivided into four habitat types using multivariate regression tree (MRT) analysis. According to an indicator species analysis, 38 of the 47 studied species were found to be significant indicators of the MRT habitat types. The relationships between species richness and topographic variables were found to be scale-dependent, while the great majority of the species shows distinct habitat-dependence. There are 188 potential species-habitat associations, and 114 of these were significantly positive or negative based on habitat randomization. We identified 139 significant associations using a species randomization. A habitat is not a closed system it may be both, either a sink or a source. Therefore, additional to the randomization, the Poisson Cluster Model (PCM) was applied. PCM considers the spatial autocorrelation of species and habitats, and thus appears to be more realistic than the traditional randomization processes. It identified only 37 associations that were significant. In conclusion, the deviation from the random process, i.e. the high degree of species spatial mingling may be explained by persistent immigration across habitats.
  • Zhang, Key Laboratory for Forest Resources & Ecosystem Processes of Beijing, Beijing Forestry University, Beijing 100083, China ORCID ID:E-mail: zcy_0520@163.com (email)
  • Zhao, Department of Landscape Architecture, School of Architecture, Tsinghua University, Beijing 100084, China ORCID ID:E-mail:
  • Zhao, Key Laboratory for Forest Resources & Ecosystem Processes of Beijing, Beijing Forestry University, Beijing 100083, China ORCID ID:E-mail:
  • von Gadow, Faculty of Forestry and Forest Ecology, Georg-August-University Göttingen, Büsgenweg 5, D-37077 Göttingen, Germany ORCID ID:E-mail: KGadow@gwdg.de
article id 907, category Research article
Chunyu Zhang, Yazhou Zhao, Xiuhai Zhao, Klaus von Gadow. (2012). Species-habitat associations in a northern temperate forest in China. Silva Fennica vol. 46 no. 4 article id 907. https://doi.org/10.14214/sf.907
This contribution identifies species-habitat associations in a temperate forest in north-eastern China, based on the assumption that habitats are spatially autocorrelated and species are spatially aggregated due to limited seed dispersal. The empirical observations were obtained in a large permanent experimental area covering 660 x 320 m. The experimental area was subdivided into four habitat types using multivariate regression tree (MRT) analysis. According to an indicator species analysis, 38 of the 47 studied species were found to be significant indicators of the MRT habitat types. The relationships between species richness and topographic variables were found to be scale-dependent, while the great majority of the species shows distinct habitat-dependence. There are 188 potential species-habitat associations, and 114 of these were significantly positive or negative based on habitat randomization. We identified 139 significant associations using a species randomization. A habitat is not a closed system it may be both, either a sink or a source. Therefore, additional to the randomization, the Poisson Cluster Model (PCM) was applied. PCM considers the spatial autocorrelation of species and habitats, and thus appears to be more realistic than the traditional randomization processes. It identified only 37 associations that were significant. In conclusion, the deviation from the random process, i.e. the high degree of species spatial mingling may be explained by persistent immigration across habitats.
  • Zhang, Key Laboratory for Forest Resources & Ecosystem Processes of Beijing, Beijing Forestry University, Beijing 100083, China ORCID ID:E-mail: zcy_0520@163.com (email)
  • Zhao, Department of Landscape Architecture, School of Architecture, Tsinghua University, Beijing 100084, China ORCID ID:E-mail:
  • Zhao, Key Laboratory for Forest Resources & Ecosystem Processes of Beijing, Beijing Forestry University, Beijing 100083, China ORCID ID:E-mail:
  • von Gadow, Faculty of Forestry and Forest Ecology, Georg-August-University Göttingen, Büsgenweg 5, D-37077 Göttingen, Germany ORCID ID:E-mail: KGadow@gwdg.de
article id 440, category Research article
Chen-Jung Lin, Chih-Hsin Chung, Te-Hsin Yang, Far-Ching Lin. (2012). Detection of electric resistivity tomography and evaluation of the sapwood-heartwood demarcation in three Asia Gymnosperm species. Silva Fennica vol. 46 no. 3 article id 440. https://doi.org/10.14214/sf.440
The proportions of sapwood and heartwood of trees have significant impacts on various uses. Electric resistivity tomography (ERT) and corresponding electrical resistance (ER) value maps were examined in Japanese cedar (Cryptomeria japonica D. Don), Taiwania (Taiwania cryptomerioides Hayata), and Luanta fir (Cunninghamia konishii Hayata) trees. The position of the sapwood-heartwood demarcation was measured on incremental cores from living trees and the corresponding ER of the sapwood-heartwood boundary was acquired from the ER map. A positive significant relationship was found between the maximum ER plus minimum ER values (ERmax + ERmin) and ER of the sapwood-heartwood demarcation from the tomographic data. The position of the sapwood-heartwood demarcation was determined by corresponding ER, and the critical ER can be established by the ERmax + ERmin value of the tomographic data. The results from this study indicate that ERT technique can be used to determine the position of the sapwood-heartwood boundary and can serve as a methodology in undamaged living trees of Gymnosperm species.
  • Lin, Taiwan Forestry Research Institute, Taipei Taiwan ORCID ID:E-mail: d88625002@yahoo.com.tw (email)
  • Chung, Taiwan Forestry Research Institute, Taipei Taiwan ORCID ID:E-mail:
  • Yang, Taiwan Forestry Research Institute, Taipei Taiwan ORCID ID:E-mail:
  • Lin, Taiwan Forestry Research Institute, Taipei Taiwan ORCID ID:E-mail:
article id 440, category Research article
Chen-Jung Lin, Chih-Hsin Chung, Te-Hsin Yang, Far-Ching Lin. (2012). Detection of electric resistivity tomography and evaluation of the sapwood-heartwood demarcation in three Asia Gymnosperm species. Silva Fennica vol. 46 no. 3 article id 440. https://doi.org/10.14214/sf.440
The proportions of sapwood and heartwood of trees have significant impacts on various uses. Electric resistivity tomography (ERT) and corresponding electrical resistance (ER) value maps were examined in Japanese cedar (Cryptomeria japonica D. Don), Taiwania (Taiwania cryptomerioides Hayata), and Luanta fir (Cunninghamia konishii Hayata) trees. The position of the sapwood-heartwood demarcation was measured on incremental cores from living trees and the corresponding ER of the sapwood-heartwood boundary was acquired from the ER map. A positive significant relationship was found between the maximum ER plus minimum ER values (ERmax + ERmin) and ER of the sapwood-heartwood demarcation from the tomographic data. The position of the sapwood-heartwood demarcation was determined by corresponding ER, and the critical ER can be established by the ERmax + ERmin value of the tomographic data. The results from this study indicate that ERT technique can be used to determine the position of the sapwood-heartwood boundary and can serve as a methodology in undamaged living trees of Gymnosperm species.
  • Lin, Taiwan Forestry Research Institute, Taipei Taiwan ORCID ID:E-mail: d88625002@yahoo.com.tw (email)
  • Chung, Taiwan Forestry Research Institute, Taipei Taiwan ORCID ID:E-mail:
  • Yang, Taiwan Forestry Research Institute, Taipei Taiwan ORCID ID:E-mail:
  • Lin, Taiwan Forestry Research Institute, Taipei Taiwan ORCID ID:E-mail:
article id 45, category Research article
Guolei Li, Yong Liu, Yan Zhu, Qing Mei Li, R. Karsten Dumroese. (2012). Effect of fall-applied nitrogen on growth, nitrogen storage and frost hardiness of bareroot Larix olgensis seedlings. Silva Fennica vol. 46 no. 3 article id 45. https://doi.org/10.14214/sf.45
Nursery response of evergreen trees to fall fertilization has been studied widely, but little attention has been given to deciduous trees. Bareroot Olga Bay larch (Larix olgensis Henry) seedlings were fertilized in the nursery with urea at four rates (0, 30, 60, 90 kg N ha–1), with half of each rate applied on two dates (September 16 and October 1, 2009). The seedlings were excavated for evaluation on October 15. In the unfertilized (control) treatment, root and shoot dry mass increased by 100% and 57% respectively, while N concentration in the roots and shoots increased by 43% and 40% during the 30 day period. This indicated that substantial biomass growth during this period did not lead to internal nutrient dilution. Root dry mass increased when fall fertilization rates were ≥ 60 kg N ha–1. Fall fertilization increased N concentrations in root tissue by 48–73%. Compared with the control, shoot tissues of fall fertilized seedlings had slightly higher N concentration and content and significantly higher frost hardiness.
  • Li, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing 100083, China ORCID ID:E-mail:
  • Liu, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing 100083, China ORCID ID:E-mail: lyong@bjfu.edu.cn (email)
  • Zhu, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing 100083, China ORCID ID:E-mail:
  • Li, Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Forest Silviculture of State Forestry Administration, Beijing 100091, China ORCID ID:E-mail:
  • Dumroese, US Department of Agriculture, Forest Service, Rocky Mountain Research Station, Moscow, ID, USA ORCID ID:E-mail:
article id 43, category Research article
Anni Markkanen, Panu Halme. (2012). Polypore communities in broadleaved boreal forests. Silva Fennica vol. 46 no. 3 article id 43. https://doi.org/10.14214/sf.43
The cover and extent of boreal broadleaved forests have been decreasing due to modern forest management practices and fire suppression. As decomposers of woody material, polypores are ecologically important ecosystem engineers. The ecology and conservation biology of polypores have been studied intensively in boreal coniferous forests. However, only a few studies have focused on the species living on broadleaved trees. To increase knowledge on this species group we conducted polypore surveys in 27 broadleaved forests and 303 forest compartments (539 ha) on the southern boreal zone in Finland and measured dead wood and forest characteristics. We detected altogether 98 polypore species, of which 13 are red-listed in Finland. 60% of the recorded species are primarily associated with broadleaved trees. The number of species in a local community present in a broadleaved forest covered approximately 50 species, of which 30–40 were primarily associated with broadleaved trees. The size of the inventoried area explained 67% of the variation in the species richness, but unlike in previous studies conducted in coniferous forests, dead wood variables as well as forest structure had very limited power in explaining polypore species richness on forest stand level. The compartments occupied by red listed Protomerulius caryae had an especially high volume of living birch, but otherwise the occurrences of red-listed species could not be predicted based on the forest structure.
  • Markkanen, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland ORCID ID:E-mail: anni.e.markkanen@gmail.com (email)
  • Halme, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland ORCID ID:E-mail:
article id 43, category Research article
Anni Markkanen, Panu Halme. (2012). Polypore communities in broadleaved boreal forests. Silva Fennica vol. 46 no. 3 article id 43. https://doi.org/10.14214/sf.43
The cover and extent of boreal broadleaved forests have been decreasing due to modern forest management practices and fire suppression. As decomposers of woody material, polypores are ecologically important ecosystem engineers. The ecology and conservation biology of polypores have been studied intensively in boreal coniferous forests. However, only a few studies have focused on the species living on broadleaved trees. To increase knowledge on this species group we conducted polypore surveys in 27 broadleaved forests and 303 forest compartments (539 ha) on the southern boreal zone in Finland and measured dead wood and forest characteristics. We detected altogether 98 polypore species, of which 13 are red-listed in Finland. 60% of the recorded species are primarily associated with broadleaved trees. The number of species in a local community present in a broadleaved forest covered approximately 50 species, of which 30–40 were primarily associated with broadleaved trees. The size of the inventoried area explained 67% of the variation in the species richness, but unlike in previous studies conducted in coniferous forests, dead wood variables as well as forest structure had very limited power in explaining polypore species richness on forest stand level. The compartments occupied by red listed Protomerulius caryae had an especially high volume of living birch, but otherwise the occurrences of red-listed species could not be predicted based on the forest structure.
  • Markkanen, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland ORCID ID:E-mail: anni.e.markkanen@gmail.com (email)
  • Halme, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland ORCID ID:E-mail:
article id 58, category Research article
Ilona Suuriniemi, Jukka Matero, Harri Hänninen, Jussi Uusivuori. (2012). Factors affecting enlargement of family forest holdings. Silva Fennica vol. 46 no. 2 article id 58. https://doi.org/10.14214/sf.58
This study contributes to the research of enlargement – a counterforce of parcelization – of forest holdings. To help planning policy measures aiming at increased average size of forest holdings, we study the characteristics of family forest owners who acquired additional forestland area during the years 2004–2008. Increases of forestland area due to purchases on the open market, purchases from parents or other relatives, inheritance or gift are studied. Survey data, containing information of 6318 forest owners, are analyzed with logistic regression analysis in order to establish a relationship between the probability of increasing the forestland area and the characteristics of landowners. The results indicate that young male owners, who appreciate economic values of the ownership and are active users of their forest estates, most often expand their forest property. This can be considered as an encouraging result from the point of view of the political objective to boost forest management activity through enlarging family forest holdings.
  • Suuriniemi, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail:
  • Matero, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: jukka.matero@uef.fi (email)
  • Hänninen, Finnish Forest Research Institute, Vantaa Unit, P.O. Box 18, FI-01301 Vantaa, Finland ORCID ID:E-mail:
  • Uusivuori, Finnish Forest Research Institute, Vantaa Unit, P.O. Box 18, FI-01301 Vantaa, Finland ORCID ID:E-mail:
article id 56, category Research article
Johan Holmgren, Andreas Barth, Henrik Larsson, Håkan Olsson. (2012). Prediction of stem attributes by combining airborne laser scanning and measurements from harvesters. Silva Fennica vol. 46 no. 2 article id 56. https://doi.org/10.14214/sf.56
In this study, a new method was validated for the first time that predicts stem attributes for a forest area without any manual measurements of tree stems by combining harvester measurements and Airborne Laser Scanning (ALS) data. A new algorithm for automatic segmentation of tree crowns from ALS data based on tree crown models was developed. The test site was located in boreal forest (64°06’N, 19°10’E) dominated by Norway spruce (Picea abies) and Scots Pine (Pinus sylvestris).The trees were harvested on field plots, and each harvested tree was linked to the nearest tree crown segment derived from ALS data. In this way, a reference database was created with both stem data from the harvester and ALS derived features for linked tree crowns. To estimate stem attributes for a tree crown segment in parts of the forest where trees not yet have been harvested, tree stems are imputed from the most similar crown segment in the reference database according to features extracted from ALS data. The imputation of harvester data was validated on a sub-stand-level, i.e. 2–4 aggregated 10 m radius plots, and the obtained RMSE of stem volume, mean tree height, mean stem diameter, and stem density (stems per ha) estimates were 11%, 8%, 12%, and 19%, respectively. The imputation of stem data collected by harvesters could in the future be used for bucking simulations of not yet harvested forest stands in order to predict wood assortments.
  • Holmgren, Swedish University of Agricultural Sciences, Forest Resource Management, Umeå, Sweden ORCID ID:E-mail: johan.holmgren@slu.se (email)
  • Barth, The Forestry Research Institute of Sweden, Uppsala, Sweden ORCID ID:E-mail:
  • Larsson, Swedish University of Agricultural Sciences, Forest Resource Management, Umeå, Sweden ORCID ID:E-mail:
  • Olsson, Swedish University of Agricultural Sciences, Forest Resource Management, Umeå, Sweden ORCID ID:E-mail:
article id 444, category Research article
Sovu, Mulualem Tigabu, Patrice Savadogo, Per Christer Odén. (2012). Facilitation of forest landscape restoration on abandoned swidden fallows in Laos using mixed-species planting and biochar application. Silva Fennica vol. 46 no. 1 article id 444. https://doi.org/10.14214/sf.444
The cessation of swidden cultivation and the increasing trend of abandonment of swidden fallows have created an opportunity for forest landscape restoration. However, ways need to be found to improve the poor soil fertility at these sites with affordable materials and to generate short-term socio-economic benefits for small-scale swidden fallow holders. This study assessed the feasibility of using mixed-planting of eight native species and application of rice husk biochar as soil amendment measure at a site in Laos. The effect of biochar application was compared against addition of inorganic (NPK) fertilizer and the control. The establishment and growth of the planted seedlings was then monitored for four years. The addition of rice husk biochar and NPK fertilizer did not significantly (p = 0.578) improve the survival rate of planted seedlings, which ranged from 72% to 91% (depending on the species) compared to the control. No significant growth responses to the soil amendments were observed for most of the species during the first year after planting compared to the control. The biochar effect was, however, more evident at the fourth year for diameter (p < 0.01) and height (p < 0.01) of sapling for all species; particularly its effect was more vivid on the diameter of slow-growing species. The results indicate that the species tested in the mixed-planting showed marked growth variation while application of rice husk biochar boosted their growth. Thus, planting mixed-species in swidden fallows has potential to provide continuous supplies of wood from different species to diversify the livelihood of swidden field owners, while maintaining ecosystem services.
  • -, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, Alnarp, Sweden ORCID ID:E-mail:
  • Tigabu, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, Alnarp, Sweden ORCID ID:E-mail: mulualem.tigabu@slu.se (email)
  • Savadogo, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, Alnarp, Sweden ORCID ID:E-mail:
  • Odén, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, Alnarp, Sweden ORCID ID:E-mail:
article id 68, category Research article
Maria Villikka, Petteri Packalén, Matti Maltamo. (2012). The suitability of leaf-off airborne laser scanning data in an area-based forest inventory of coniferous and deciduous trees. Silva Fennica vol. 46 no. 1 article id 68. https://doi.org/10.14214/sf.68
This study examined the suitability of airborne laser scanner (ALS) data collected under leaf-off conditions in a forest inventory, in which deciduous and coniferous trees need to be separated. All analyses were carried out with leaf-on and leaf-off ALS data collected from the same study area. Additionally, aerial photographs were utilized in the Nearest Neighbor (NN) imputations. An area-based approach was used in this study. Regression estimates of plot volume were more accurate in the case of leaf-off than leaf-on data. In addition, regression models were more accurate in coniferous plots than in deciduous plots. The results of applying leaf-on models with leaf-off data, and vice versa, indicate that leaf-on and leaf-off data should not be combined since this causes serious bias. The total volume and volume by coniferous and deciduous trees was estimated by the NN imputation. In terms of total volume, leaf-off data provided more accurate estimates than leaf-on data. In addition, leaf-off data discriminated between coniferous and deciduous trees, even without the use of aerial photographs. Accurate results were also obtained when leaf-off ALS data were used to classify sample plots into deciduous and coniferous dominated plots. The results indicate that the area-based method and ALS data collected under leaf-off conditions are suitable for forest inventory in which deciduous and coniferous trees need to be distinguished.
  • Villikka, University of Easten Finland, Department of Forest Sciences, Joensuu, Finland ORCID ID:E-mail:
  • Packalén, University of Easten Finland, Department of Forest Sciences, Joensuu, Finland ORCID ID:E-mail: petteri.packalen@uef.fi (email)
  • Maltamo, University of Easten Finland, Department of Forest Sciences, Joensuu, Finland ORCID ID:E-mail:
article id 65, category Research article
Kristóf Kelemen, Barbara Mihók, László Gálhidy, Tibor Standovár. (2012). Dynamic response of herbaceous vegetation to gap opening in a Central European beech stand. Silva Fennica vol. 46 no. 1 article id 65. https://doi.org/10.14214/sf.65
Herbaceous ground vegetation in artificially-created gaps was studied in a managed beech (Fagus sylvatica L.) forest over a period of eight years in Northern Hungary, Central Europe. These gaps were being used as an alternative to the regular shelterwood system to create uneven-aged stands. The effects of gap size (15 and 40 m diameter) and canopy openness on herbaceous species colonization and persistence were assessed in a systematic grid of 5 5 m. Overall, herbaceous cover was low before gap creation, increased soon afterwards, and continued to rise over time. The number of herb species increased in the gaps and, to a lesser extent, in adjacent areas under the remaining tree canopy. Colonization of gaps was rapid and there was substantial turnover of species i.e. various species disappeared from the gaps over time whilst others colonized. Species with both long-term persistent seed banks and long distance dispersal abilities were the most successful types colonizing gaps. Six species occurred preferentially in large gaps, while only one species was found to prefer small gaps. Species present before gap creation survived in both gap sizes. Smaller gaps with a diameter of half the height of canopy trees also tended to remain free of common weed species, whereas large cover of Rubus fruticosus L. and Calamagrostis epigejos (L.) Roth could hamper natural regeneration in larger gaps. For the successful regeneration of beech we recommend the use of small gaps complemented by few large gaps.
  • Kelemen, Loránd Eötvös University, Institute of Biology, Department of Plant Systematics, Ecology and Theoretical Biology, Budapest, Hungary ORCID ID:E-mail:
  • Mihók, Loránd Eötvös University, Institute of Biology, Department of Plant Systematics, Ecology and Theoretical Biology, Budapest, Hungary ORCID ID:E-mail:
  • Gálhidy, Loránd Eötvös University, Institute of Biology, Department of Plant Systematics, Ecology and Theoretical Biology, Budapest, Hungary ORCID ID:E-mail:
  • Standovár, Institute of Biology, Department of Plant Systematics, Ecology and Theoretical Biology, Budapest, Hungary ORCID ID:E-mail: standy@ludens.elte.hu (email)
article id 90, category Research article
Per Angelstam, Kjell Andersson, Robert Axelsson, Marine Elbakidze, Bengt Gunnar Jonsson, Jean-Michel Roberge. (2011). Protecting forest areas for biodiversity in Sweden 1991–2010: the policy implementation process and outcomes on the ground. Silva Fennica vol. 45 no. 5 article id 90. https://doi.org/10.14214/sf.90
Swedish forest and environmental policies imply that forests should be managed so that all naturally occurring species are maintained in viable populations. This requires maintenance of functional networks of representative natural forest and cultural woodland habitats. We first review the policy implementation process regarding protected areas in Sweden 1991–2010, how ecological knowledge was used to formulate interim short-term and strategic long-term biodiversity conservation goals, and the development of a hierarchical spatial planning approach. Second, we present data about the amount of formally protected and voluntarily set aside forest stands, and evaluate how much remains in terms of additional forest protection, conservation management and habitat restoration to achieve forest and environmental policy objectives in the long-term. Third, a case study in central Sweden was made to estimate the functionality of old Scots pine, Norway spruce and deciduous forest habitats, as well as cultural woodland, in different forest regions. Finally, we assess operational biodiversity conservation planning processes. We conclude that Swedish policy pronouncements capture the contemporary knowledge about biodiversity and conservation planning well. However, the existing area of protected and set-aside forests is presently too small and with too poor connectivity. To bridge this gap, spatial planning, management and restoration of habitat, as well as collaboration among forest and conservation planners need to be improved.
  • Angelstam, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden ORCID ID:E-mail: per.angelstam@slu.se (email)
  • Andersson, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden ORCID ID:E-mail:
  • Axelsson, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden ORCID ID:E-mail:
  • Elbakidze, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden ORCID ID:E-mail:
  • Jonsson, Dept of Natural Science, Engineering and Mathematics, Mid Sweden University, Sundsvall, Sweden ORCID ID:E-mail:
  • Roberge, Dept of Wildlife, Fish and Environmental Studies, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Umeå, Sweden ORCID ID:E-mail:
article id 90, category Research article
Per Angelstam, Kjell Andersson, Robert Axelsson, Marine Elbakidze, Bengt Gunnar Jonsson, Jean-Michel Roberge. (2011). Protecting forest areas for biodiversity in Sweden 1991–2010: the policy implementation process and outcomes on the ground. Silva Fennica vol. 45 no. 5 article id 90. https://doi.org/10.14214/sf.90
Swedish forest and environmental policies imply that forests should be managed so that all naturally occurring species are maintained in viable populations. This requires maintenance of functional networks of representative natural forest and cultural woodland habitats. We first review the policy implementation process regarding protected areas in Sweden 1991–2010, how ecological knowledge was used to formulate interim short-term and strategic long-term biodiversity conservation goals, and the development of a hierarchical spatial planning approach. Second, we present data about the amount of formally protected and voluntarily set aside forest stands, and evaluate how much remains in terms of additional forest protection, conservation management and habitat restoration to achieve forest and environmental policy objectives in the long-term. Third, a case study in central Sweden was made to estimate the functionality of old Scots pine, Norway spruce and deciduous forest habitats, as well as cultural woodland, in different forest regions. Finally, we assess operational biodiversity conservation planning processes. We conclude that Swedish policy pronouncements capture the contemporary knowledge about biodiversity and conservation planning well. However, the existing area of protected and set-aside forests is presently too small and with too poor connectivity. To bridge this gap, spatial planning, management and restoration of habitat, as well as collaboration among forest and conservation planners need to be improved.
  • Angelstam, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden ORCID ID:E-mail: per.angelstam@slu.se (email)
  • Andersson, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden ORCID ID:E-mail:
  • Axelsson, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden ORCID ID:E-mail:
  • Elbakidze, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden ORCID ID:E-mail:
  • Jonsson, Dept of Natural Science, Engineering and Mathematics, Mid Sweden University, Sundsvall, Sweden ORCID ID:E-mail:
  • Roberge, Dept of Wildlife, Fish and Environmental Studies, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Umeå, Sweden ORCID ID:E-mail:
article id 90, category Research article
Per Angelstam, Kjell Andersson, Robert Axelsson, Marine Elbakidze, Bengt Gunnar Jonsson, Jean-Michel Roberge. (2011). Protecting forest areas for biodiversity in Sweden 1991–2010: the policy implementation process and outcomes on the ground. Silva Fennica vol. 45 no. 5 article id 90. https://doi.org/10.14214/sf.90
Swedish forest and environmental policies imply that forests should be managed so that all naturally occurring species are maintained in viable populations. This requires maintenance of functional networks of representative natural forest and cultural woodland habitats. We first review the policy implementation process regarding protected areas in Sweden 1991–2010, how ecological knowledge was used to formulate interim short-term and strategic long-term biodiversity conservation goals, and the development of a hierarchical spatial planning approach. Second, we present data about the amount of formally protected and voluntarily set aside forest stands, and evaluate how much remains in terms of additional forest protection, conservation management and habitat restoration to achieve forest and environmental policy objectives in the long-term. Third, a case study in central Sweden was made to estimate the functionality of old Scots pine, Norway spruce and deciduous forest habitats, as well as cultural woodland, in different forest regions. Finally, we assess operational biodiversity conservation planning processes. We conclude that Swedish policy pronouncements capture the contemporary knowledge about biodiversity and conservation planning well. However, the existing area of protected and set-aside forests is presently too small and with too poor connectivity. To bridge this gap, spatial planning, management and restoration of habitat, as well as collaboration among forest and conservation planners need to be improved.
  • Angelstam, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden ORCID ID:E-mail: per.angelstam@slu.se (email)
  • Andersson, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden ORCID ID:E-mail:
  • Axelsson, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden ORCID ID:E-mail:
  • Elbakidze, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden ORCID ID:E-mail:
  • Jonsson, Dept of Natural Science, Engineering and Mathematics, Mid Sweden University, Sundsvall, Sweden ORCID ID:E-mail:
  • Roberge, Dept of Wildlife, Fish and Environmental Studies, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Umeå, Sweden ORCID ID:E-mail:
article id 90, category Research article
Per Angelstam, Kjell Andersson, Robert Axelsson, Marine Elbakidze, Bengt Gunnar Jonsson, Jean-Michel Roberge. (2011). Protecting forest areas for biodiversity in Sweden 1991–2010: the policy implementation process and outcomes on the ground. Silva Fennica vol. 45 no. 5 article id 90. https://doi.org/10.14214/sf.90
Swedish forest and environmental policies imply that forests should be managed so that all naturally occurring species are maintained in viable populations. This requires maintenance of functional networks of representative natural forest and cultural woodland habitats. We first review the policy implementation process regarding protected areas in Sweden 1991–2010, how ecological knowledge was used to formulate interim short-term and strategic long-term biodiversity conservation goals, and the development of a hierarchical spatial planning approach. Second, we present data about the amount of formally protected and voluntarily set aside forest stands, and evaluate how much remains in terms of additional forest protection, conservation management and habitat restoration to achieve forest and environmental policy objectives in the long-term. Third, a case study in central Sweden was made to estimate the functionality of old Scots pine, Norway spruce and deciduous forest habitats, as well as cultural woodland, in different forest regions. Finally, we assess operational biodiversity conservation planning processes. We conclude that Swedish policy pronouncements capture the contemporary knowledge about biodiversity and conservation planning well. However, the existing area of protected and set-aside forests is presently too small and with too poor connectivity. To bridge this gap, spatial planning, management and restoration of habitat, as well as collaboration among forest and conservation planners need to be improved.
  • Angelstam, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden ORCID ID:E-mail: per.angelstam@slu.se (email)
  • Andersson, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden ORCID ID:E-mail:
  • Axelsson, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden ORCID ID:E-mail:
  • Elbakidze, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden ORCID ID:E-mail:
  • Jonsson, Dept of Natural Science, Engineering and Mathematics, Mid Sweden University, Sundsvall, Sweden ORCID ID:E-mail:
  • Roberge, Dept of Wildlife, Fish and Environmental Studies, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Umeå, Sweden ORCID ID:E-mail:
article id 88, category Research article
Seppo Rouvinen, Jari Kouki. (2011). Tree regeneration in artificial canopy gaps established for restoring natural structural variability in a Scots pine stand. Silva Fennica vol. 45 no. 5 article id 88. https://doi.org/10.14214/sf.88
In Finland and elsewhere in Europe, many protected forest areas include also stands that were previously managed and that lack several naturally occurring stand characteristics. In these areas, ecosystem restoration can be used to facilitate and accelerate the formation of structural and habitat features resembling those of natural forests. For example, by creating small gaps it could be possible to diversify forest structure and tree species composition and to produce dead wood while still maintaining mostly continous canopy coverage. We examined experimentally the effects of artificial gap formation on post-disturbance tree regeneration in the gaps in a young protected, but formerly commercially managed pine (Pinus sylvestris L.) dominated forest. In the experimental sites, gap size and the portion of girdled trees out of all treated trees (girdled and felled trees combined) in the gaps varied. Natural and artificial (direct seeding of silver birch Betula pendula Roth) tree regeneration and development was monitored both on disturbed (scarified soil patches) and undisturbed forest floor during three growing seasons. Results show that gaps can be valuable in diversifying stand structure but to be successful and rapid, tree regeneration needs disturbed forest floor. Pine regenerated numerously, but birch had clearly lower regeneration, especially on small-sized gaps. In conclusion, increasing tree diversity in young pine-dominated forests seems to be difficult when only small artificial gaps are used. But even small gaps can be used to create and maintain diverse cohort structure of the dominant species and thus they can contribute to restoration goals.
  • Rouvinen, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland ORCID ID:E-mail: seppo.rouvinen@uef.fi (email)
  • Kouki, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland ORCID ID:E-mail:
article id 86, category Research article
Mats Jonsell, Jesper Hansson. (2011). Logs and stumps in clearcuts support similar saproxylic beetle diversity: implications for bioenergy harvest. Silva Fennica vol. 45 no. 5 article id 86. https://doi.org/10.14214/sf.86
Stumps from clear cuts are increasingly used for bioenergy. Extracting this wood will reduce the habitat available for saproxylic (wood-living) organisms. As little is known about the species assemblages that will be affected, we investigated the diversity of saproxylic beetles in stumps on clear-felled sites and as a reference, we compared it with the diversity in downed logs. Stumps and logs of aspen (Populus tremula L.), birch (Betula pubescens Ehrh. and B. verrucosa Ehrh.[syn. B. pendula Roth]), spruce (Picea abies (L.) Karst.) and pine (Pinus sylvestris L.) were examined in clear cuts of two different ages: one summer old and 4–5 years old. The beetles were sampled by sieving bark (0.25 m2) peeled from the wood. The samples were taken in pairs of one log and one stump situated close together and of the same tree species, age since death and diameter. In total 3348 saproxylic beetles belonging to 124 species were found in 176 samples. The stumps had a similar number of species to the logs both as measured per sample and as an accumulated number. Exceptions were 4–5 years old wood of birch and pine where the number was significantly higher in the stumps. The number of red-listed species was also similar between stumps and logs. Species composition was more different between the stumps and logs of conifers than of deciduous trees. We conclude that clear-felled stumps have a diverse saproxylic insect fauna. This has to be taken into account if large scale extraction of logging stumps is implemented.
  • Jonsell, Swedish University of Agrarian Sciences, Dept of Ecology, Uppsala, Sweden ORCID ID:E-mail: mats.jonsell@ekol.slu.se (email)
  • Hansson, Swedish University of Agrarian Sciences, Dept of Ecology, Uppsala, Sweden ORCID ID:E-mail:
article id 85, category Research article
Ville A.O. Selonen, Maija Mussaari, Tero Toivanen, Janne S. Kotiaho. (2011). The conservation potential of brook-side key habitats in managed boreal forests. Silva Fennica vol. 45 no. 5 article id 85. https://doi.org/10.14214/sf.85
Today, maintaining biodiversity is included in the targets of boreal forest management. A widespread approach in northern Europe is to identify and preserve woodland key habitats within managed forests. Woodland key habitats are expected to be patches that host populations of threatened and declining species, and the preservation of these patches is assumed to enable the persistence of the focal species in the landscape. In Finland, the criteria for selecting woodland key habitats are defined in the Finnish Forest Act, and the selection has been done by forest practitioners. Our objective was to determine whether the surroundings of boreal brooks and rivulets qualified as key habitats are truly different from brook-side habitats not granted the key habitat status, and whether the brook-side habitats of the two types differ from the forest matrix managed for timber production. We found that the two brook-side habitats were in most aspects rather alike but there was a difference in the composition of ground vegetation assemblages. In contrast, the control forests were distinct from the brook-sides in terms of dead wood, species richness and assemblages of polypores, species richness of epiphytic mosses, and the composition of beetle assemblages. We conclude that brook-sides in general provide an important habitat clearly diverging from the surrounding matrix but that the conservation value of the brook-sides granted the key habitat status may not be substantially larger than that of the brook-sides without the status.
  • Selonen, University of Jyväskylä, Department of Biological and Environmental Sciences, Lahti, Finland ORCID ID:E-mail: ville.selonen@juy.fi (email)
  • Mussaari, University of Jyväskylä, Department of Biological and Environmental Sciences, Lahti, Finland ORCID ID:E-mail:
  • Toivanen, University of Jyväskylä, Department of Biological and Environmental Sciences, Lahti, Finland ORCID ID:E-mail:
  • Kotiaho, University of Jyväskylä, Department of Biological and Environmental Sciences, Lahti, Finland ORCID ID:E-mail:
article id 83, category Research article
Inari Ylläsjärvi, Håkan Berglund, Timo Kuuluvainen. (2011). Relationships between wood-inhabiting fungal species richness and habitat variables in old-growth forest stands in the Pallas-Yllästunturi National Park, northern boreal Finland. Silva Fennica vol. 45 no. 5 article id 83. https://doi.org/10.14214/sf.83
Indicators for biodiversity are needed for efficient prioritization of forests selected for conservation. We analyzed the relationships between 86 wood-inhabiting fungal (polypore) species richness and 35 habitat variables in 81 northern boreal old-growth forest stands in Finland. Species richness and the number of red-listed species were analyzed separately using generalized linear models. Most species were infrequent in the studied landscape and no species was encountered in all stands. The species richness increased with 1) the volume of coarse woody debris (CWD), 2) the mean DBH of CWD and 3) the basal area of living trees. The number of red-listed species increased along the same gradients, but the effect of basal area was not significant. Polypore species richness was significantly lower on western slopes than on flat topography. On average, species richness was higher on northern and eastern slopes than on western and southern slopes. The results suggest that a combination of habitat variables used as indicators may be useful in selecting forest stands to be set aside for polypore species conservation.
  • Ylläsjärvi, Rovaniemi University of Applied Sciences, School of Forestry and Rural Industries, Jokiväylä 11 c, FI-96300 Rovaniemi, Finland ORCID ID:E-mail: inari.yllasjarvi@ramk.fi (email)
  • Berglund, University of Helsinki, Department of Forest Sciences, Helsinki, Finland ORCID ID:E-mail:
  • Kuuluvainen, University of Helsinki, Department of Forest Sciences, Helsinki, Finland ORCID ID:E-mail:
article id 83, category Research article
Inari Ylläsjärvi, Håkan Berglund, Timo Kuuluvainen. (2011). Relationships between wood-inhabiting fungal species richness and habitat variables in old-growth forest stands in the Pallas-Yllästunturi National Park, northern boreal Finland. Silva Fennica vol. 45 no. 5 article id 83. https://doi.org/10.14214/sf.83
Indicators for biodiversity are needed for efficient prioritization of forests selected for conservation. We analyzed the relationships between 86 wood-inhabiting fungal (polypore) species richness and 35 habitat variables in 81 northern boreal old-growth forest stands in Finland. Species richness and the number of red-listed species were analyzed separately using generalized linear models. Most species were infrequent in the studied landscape and no species was encountered in all stands. The species richness increased with 1) the volume of coarse woody debris (CWD), 2) the mean DBH of CWD and 3) the basal area of living trees. The number of red-listed species increased along the same gradients, but the effect of basal area was not significant. Polypore species richness was significantly lower on western slopes than on flat topography. On average, species richness was higher on northern and eastern slopes than on western and southern slopes. The results suggest that a combination of habitat variables used as indicators may be useful in selecting forest stands to be set aside for polypore species conservation.
  • Ylläsjärvi, Rovaniemi University of Applied Sciences, School of Forestry and Rural Industries, Jokiväylä 11 c, FI-96300 Rovaniemi, Finland ORCID ID:E-mail: inari.yllasjarvi@ramk.fi (email)
  • Berglund, University of Helsinki, Department of Forest Sciences, Helsinki, Finland ORCID ID:E-mail:
  • Kuuluvainen, University of Helsinki, Department of Forest Sciences, Helsinki, Finland ORCID ID:E-mail:
article id 79, category Research article
Jean-Philippe Légaré, Christian Hébert, Jean-Claude Ruel. (2011). Alternative silvicultural practices in irregular boreal forests: response of beetle assemblages. Silva Fennica vol. 45 no. 5 article id 79. https://doi.org/10.14214/sf.79
In the process of implementing sustainable management in the eastern Canadian boreal forest, we tested two selection cutting methods and compared them with two widely used practices in the boreal forest: clearcutting with protection of the advanced growth and soils and irregular shelterwood cutting leaving small merchantable stems. We used old-growth irregular stands as references in comparing the impact of these silvicultural treatments on the diversity and abundance of beetles. Three groups were targeted: saproxylic flying beetles, epigaeic saproxylic beetles and epigaeic non-saproxylic beetles. A sampling design including 320 pitfall traps and 80 multidirectional flight-interception traps was deployed in 2007. A total of 26 906 beetles was captured including 407 taxa distributed among 52 families. We found that clearcutting with protection of the advanced growth and soils and irregular shelterwood cutting leaving small merchantable stems had a greater impact on beetle communities than both selection cuttings. Canopy opening as well as the presence of snags and downed woody debris appear as important attributes for several saproxylic and non-saproxylic species. Beetle communities in selection cuttings remained more similar to those found in controls; these silvicultural treatments are new tools to implement ecosystemic and sustainable management in irregular boreal forests.
  • Légaré, Université Laval, Faculté de foresterie, de géographie et de géomatique, Pavillon Abitibi-Price, Québec, Canada ORCID ID:E-mail:
  • Hébert, Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, 1055 du P.E.P.S., P.O. Box 10380, Stn. Sainte-Foy, Québec (Québec), G1V 4C7, Canada ORCID ID:E-mail: christian.hebert@rncan.gc.ca (email)
  • Ruel, Université Laval, Faculté de foresterie, de géographie et de géomatique, Pavillon Abitibi-Price, Québec, Canada ORCID ID:E-mail:
article id 78, category Research article
Kris Vandekerkhove, Luc De Keersmaeker, Ruben Walleyn, Frank Köhler, Luc Crevecoeur, Leen Govaere, Arno Thomaes, Kris Verheyen. (2011). Reappearance of old-growth elements in lowland woodlands in northern Belgium: Do the associated species follow? Silva Fennica vol. 45 no. 5 article id 78. https://doi.org/10.14214/sf.78
The forest cover of the western European lowland plain has been very low for centuries. Remaining forests were intensively managed, and old-growth elements like veteran trees and coarse woody debris became virtually absent. Only over the last decades have these old-growth elements progressively redeveloped in parks, lanes and forests, and have now reached their highest level over the last 500–1000 years. Biodiversity associated with these old-growth elements makes up an important part of overall forest biodiversity. The ability of species to recolonise the newly available habitat is strongly determined by limitations in their dispersal and establishment. We analyse the current status and development of old-growth elements in Flanders (northern Belgium) and the process of recolonisation by means of specific cases, focussing on saproxylic fungi and saproxylic beetles. Our results show that ‘hotspots’ of secondary old growth, even isolated small patches, may have more potential for specialised biodiversity than expected, and may provide important new strongholds for recovery and recolonisation of an important share of old-growth related species.
  • Vandekerkhove, INBO, Gaverstraat 4, B-9500 Geraardsbergen, Belgium ORCID ID:E-mail: kris.vandekerkhove@inbo.be (email)
  • De Keersmaeker, INBO, Gaverstraat 4, B-9500 Geraardsbergen, Belgium ORCID ID:E-mail:
  • Walleyn, INBO, Gaverstraat 4, B-9500 Geraardsbergen, Belgium ORCID ID:E-mail:
  • Köhler, Koleopterologisches Forschungsbüro, Bornheim, Germany ORCID ID:E-mail:
  • Crevecoeur, Genk, Belgium ORCID ID:E-mail:
  • Govaere, Agency of Nature and Forests, Brussels, Belgium ORCID ID:E-mail:
  • Thomaes, INBO, Gaverstraat 4, B-9500 Geraardsbergen, Belgium ORCID ID:E-mail:
  • Verheyen, Ghent University, Laboratory of Forestry, Gontrode, Belgium ORCID ID:E-mail:
article id 77, category Research article
Annie Claude Bélisle, Sylvie Gauthier, Dominic Cyr, Yves Bergeron, Hubert Morin. (2011). Fire regime and old-growth boreal forests in central Quebec, Canada: an ecosystem management perspective. Silva Fennica vol. 45 no. 5 article id 77. https://doi.org/10.14214/sf.77
Boreal forest management in Eastern Canada has caused depletion and fragmentation of old-growth ecosystems, with growing impacts on the associated biodiversity. To mitigate impacts of management while maintaining timber supplies, ecosystem management aims to narrow the gap between natural and managed landscapes. Our study describes the fire history and associated natural old-growth forest proportions and distribution of a 5000 km2 area located in the black spruce-feather moss forest of central Quebec. We reconstructed a stand-origin map using archival data, aerial photos and dendrochronology. According to survival analysis (Cox hazard model), the mean fire cycle length was 247 years for the 1734–2009 period. Age-class distribution modelling showed that old-growth forests were present on an average of 55% of the landscape over the last 275 years. The mean fire size was 10 113 ha, while most of the burned area was attributable to fires larger than 10 000 ha, leading to old-growth agglomerations of hundreds of square kilometres. In regards to our findings, we propose ecosystem management targets and strategies to preserve forest diversity and resilience.
  • Bélisle, Centre for Forest Research, Université du Québec à Montréal, Montréal, Québec, Canada ORCID ID:E-mail: annieclaude_b@hotmail.com (email)
  • Gauthier, Laurentian Forestry Centre, Canadian Forest Service, Natural Resources Canada, Sainte-Foy, Québec, Canada ORCID ID:E-mail:
  • Cyr, Institut Québécois d’Aménagement de la Fort Feuillue, Université du Québec en Outaouais, Ripon, Québec, Canada ORCID ID:E-mail:
  • Bergeron, Centre for Forest Research, Université du Québec à Montréal, Montréal, Québec, Canada & NSERC-UQAT-UQAM Industrial Chair in Sustainable Forest Management, Université du Québec en Abitibi-Témiscamingue, Rouyn-Noranda, Québec, Canada ORCID ID:E-mail:
  • Morin, Département des Sciences Fondamentales, Université du Québec à Chicoutimi, Chicoutimi, Québec, Canada ORCID ID:E-mail:
article id 76, category Research article
Alessandra Bottero, Matteo Garbarino, Vojislav Dukic, Zoran Govedar, Emanuele Lingua, Thomas A. Nagel, Renzo Motta. (2011). Gap-phase dynamics in the old-growth forest of Lom, Bosnia and Herzegovina. Silva Fennica vol. 45 no. 5 article id 76. https://doi.org/10.14214/sf.76
We investigated forest canopy gaps in the mixed beech (Fagus sylvatica L.), silver fir (Abies alba Miller), and Norway spruce (Picea abies (L.) Karst.) old-growth forest of Lom in the Dinaric Mountains of Bosnia and Herzegovina. Gap size, age, gap fraction, gapmaker characteristics and the structure and composition of gapfillers were documented to investigate gap dynamics. The percentages of forest area in canopy and expanded gaps were 19% and 41%, respectively. The median canopy gap size was 77 m2, and ranged from 11 to 708 m2. Although there were many single tree-fall gaps, the majority had multiple gapmakers that were often in different stages of decay, suggesting gap expansion is important at the study site. Of the gapmakers recorded, 14% were uprooted stems, 60% snapped stems, and 26% were standing dead trees. Dendroecological analysis suggests that gap formation varied in time. The density of gapfillers was not correlated to gap size, and the species composition of gapfillers varied between seedling, sapling, and tree life stages. The results suggest that gaps are mainly formed by endogenous senescence of single canopy trees. Exogenous disturbance agents, most likely related to wind and snow, act mainly as secondary agents in breaking weakened trees and in expanding previously established gaps. Although the findings are partially consistent with other studies of gap disturbance processes in similar old-growth forests in central Europe, the observed gap dynamic places the Lom core area at the end of a gradient that ranges from forests controlled by very small-scale processes to those where large, stand replacing disturbances predominate.
  • Bottero, University of Turin, Department Agroselviter, Via Leonardo da Vinci 44, 10095 Grugliasco (TO), Italy ORCID ID:E-mail: alessandra.bottero@unito.it (email)
  • Garbarino, University of Turin, Department Agroselviter, Via Leonardo da Vinci 44, 10095 Grugliasco (TO), Italy ORCID ID:E-mail:
  • Dukic, University of Banja Luka, Faculty of Forestry, Banja Luka, Bosnia and Herzegovina ORCID ID:E-mail:
  • Govedar, University of Banja Luka, Faculty of Forestry, Banja Luka, Bosnia and Herzegovina ORCID ID:E-mail:
  • Lingua, University of Padua, Department of TeSAF, Legnaro (PD), Italy ORCID ID:E-mail:
  • Nagel, University of Ljubljana, Department of Forestry and Renewable Forest Resources, Ljubljana, Slovenia ORCID ID:E-mail:
  • Motta, University of Turin, Department Agroselviter, Via Leonardo da Vinci 44, 10095 Grugliasco (TO), Italy ORCID ID:E-mail:
article id 100, category Research article
Annika Kangas, Lauri Mehtätalo, Antti Mäkinen, Kalle Vanhatalo. (2011). Sensitivity of harvest decisions to errors in stand characteristics. Silva Fennica vol. 45 no. 4 article id 100. https://doi.org/10.14214/sf.100
In forest planning, the decision maker chooses for each stand a treatment schedule for a predefined planning period. The choice is based either on optimization calculations or on silvicultural guidelines. Schedules for individual stands are obtained using a growth simulator, where measured stand characteristics such as the basal area, mean diameter, site class and mean height are used as input variables. These characteristics include errors, however, which may lead to incorrect decisions. In this study, the aim is to study the sensitivity of harvest decisions to errors in a dataset of 157 stands. Correct schedules according to silvicultural guidelines were first determined using error-free data. Different amounts of errors were then generated to the stand-specific characteristics, and the treatment schedule was selected again using the erroneous data. The decision was defined as correct, if the type of harvest in these two schedules were similar, and if the timings deviated at maximum ±2 for thinning and ±3 years for clear-cut. The dependency of probability of correct decisions on stand characteristics and the degree of errors was then modelled. The proposed model can be used to determine the required level of measurement accuracy for each characteristics in different kinds of stands, with a given accuracy requirement for the timing of treatments. This information can further be utilized in selecting the most appropriate inventory method.
  • Kangas, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID ID:E-mail: annika.kangas@helsinki.fi (email)
  • Mehtätalo, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland ORCID ID:E-mail:
  • Mäkinen, Simosol Oy, Riihimäki, Finland ORCID ID:E-mail:
  • Vanhatalo, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID ID:E-mail:
article id 98, category Research article
Meeri Pearson, Markku Saarinen, Kari Minkkinen, Niko Silvan, Jukka Laine. (2011). Mounding and scalping prior to reforestation of hydrologically sensitive deep-peated sites: factors behind Scots pine regeneration success. Silva Fennica vol. 45 no. 4 article id 98. https://doi.org/10.14214/sf.98
Watering up typically ensues after clearcutting forestry-drained peatland forests. Thus, the effectiveness of maintenance drainage and soil preparation procedures becomes paramount for establishing a new generation of commercial forest. Mounding is the primary method of soil preparation applied in regeneration sites lying on deep peat. As raised planting spots, mounds are resistant to waterlogging and assumed to be beneficial for organic matter (OM) decomposition via, e.g., increased soil aeration and temperature, which would also enhance seedling growth. In recent years, however, less intensive and cheaper alternatives like scalping have been sought with some reported cases of success. Our case study investigated the survival and growth of Scots pine outplants in mounds, scalps, and unprepared microsites along a moisture gradient. After three growing seasons, mounding accelerated neither seedling growth nor OM decomposition relative to the unprepared treatment. Survival in mounds was nonetheless superior overall. Scalps behaved as water collecting depressions leading to a catastrophic regeneration result. Based on our findings, water table level (WTL) overrides other growth-controlling factors in excess moisture conditions. To combat watering up coupled with greater than normal rainfall, we recommend reforestation strategies which provide elevated, prepared planting spots (i.e., mounds) or utilize unprepared, higher microforms.
  • Pearson, Finnish Forest Research Institute, Western Finland Regional Unit, Kaironiementie 15, FI-39700 Parkano, Finland ORCID ID:E-mail: meeri.pearson@metla.fi (email)
  • Saarinen, Finnish Forest Research Institute, Western Finland Regional Unit, Kaironiementie 15, FI-39700 Parkano, Finland ORCID ID:E-mail:
  • Minkkinen, University of Helsinki, Department of Forest Sciences, Helsinki, Finland ORCID ID:E-mail:
  • Silvan, Finnish Forest Research Institute, Western Finland Regional Unit, Kaironiementie 15, FI-39700 Parkano, Finland ORCID ID:E-mail:
  • Laine, Finnish Forest Research Institute, Western Finland Regional Unit, Kaironiementie 15, FI-39700 Parkano, Finland ORCID ID:E-mail:
article id 96, category Research article
Scott R. Abella. (2011). How well do U.S. Forest Service terrestrial ecosystem surveys correspond with measured vegetation properties? Silva Fennica vol. 45 no. 4 article id 96. https://doi.org/10.14214/sf.96
Reliable estimates of species composition that forest sites are capable of supporting – specific to ecosystem mapping units across landscapes – are useful for many purposes in forest science and management. Like forestry agencies in numerous countries, the U.S. Forest Service has invested in ecological land classification (termed terrestrial ecosystem survey [TES] in the study region of Arizona) that includes ecosystem-explicit species lists taken to be estimated potential natural vegetation (PNV). Using multivariate community analyses, PNV in the TES was compared to measured species composition on 66 sites representing among the least-disturbed vegetation (considered this study’s measured PNV) spanning 11 ecosystem types on a Pinus ponderosa P. & C. Lawson landscape in northern Arizona, USA. Agreement between the TES PNV and measured species composition was lowest for forbs and shrubs (compared to graminoids), and species composition differed significantly between the TES and this study for at least one plant lifeform in 73% of ecosystems. Reasons for differences between the TES and this study are difficult to resolve, but in some cases appear to result from identification of different species pools in the region. This study suggests that the TES is a useful starting point in understanding vegetation-environment relationships, but further work is needed to refine species lists and more thoroughly account for the influences of fire, grazing, and climate that can influence both PNV and current vegetation. Refining and updating ecosystem-specific species lists may benefit existing forest site classifications and could be planned for when new site classifications are developed, especially with changing climates.
  • Abella, Department of Environmental and Occupational Health, University of Nevada Las Vegas, Las Vegas, Nevada 89154-3064 USA ORCID ID:E-mail: scott.abella@unlv.edu (email)
article id 93, category Research article
Pertti Pulkkinen, Saila Varis, Raimo Jaatinen, Aulis Leppänen, Anne Pakkanen. (2011). Increasing survival and growth of Scots pine seedlings with selection based on autumn coloration. Silva Fennica vol. 45 no. 4 article id 93. https://doi.org/10.14214/sf.93
This study evaluates the possibility of using autumn coloration of young Scots pine (Pinus sylvestris L.) seedlings as an indicator of adaptation to harsh climate conditions. One-year old seedlings from natural stands with different origins and seed orchards were classified as “red/reddish” and “green” based on the needle color after artificially increased night length in nursery and then measured after 14 years in field trials. In almost all the studied groups seedlings classified as “red/reddish” had significantly higher survival rate than seedlings classified as “green”. The survival of “red/reddish” was 14.2% higher than “green” among natural stand seed material and 56.2% among seed orchard material. During the study period the survival difference between “red/reddish” and “green” seedlings tended to increase. The seedling color had limited connection with the height growth, even though the trees classified as “red/reddish” were slightly taller than those classified as “green”. However, the total productivity over all field trials, described here as a heightsum, of “red/reddish” trees was 15% higher than productivity of “green” trees from natural stand material, and 61% higher than those from seed orchard material. It seems that controlled selection based on autumn color can be utilized within seed crops of different types with the aim to increase the adaptability of seed material to different environmental conditions.
  • Pulkkinen, Metla, Haapastensyrjä, Läyliäinen, Finland ORCID ID:E-mail: pertti.pulkkinen@metla.fi (email)
  • Varis, Metla, Haapastensyrjä, Läyliäinen, Finland ORCID ID:E-mail:
  • Jaatinen, Metla, Haapastensyrjä, Läyliäinen, Finland ORCID ID:E-mail:
  • Leppänen, Metla, Haapastensyrjä, Läyliäinen, Finland ORCID ID:E-mail:
  • Pakkanen, Metla, Haapastensyrjä, Läyliäinen, Finland ORCID ID:E-mail:
article id 108, category Research article
Back Tomas Ersson, Urban Bergsten, Ola Lindroos. (2011). The cost-efficiency of seedling packaging specifically designed for tree planting machines. Silva Fennica vol. 45 no. 3 article id 108. https://doi.org/10.14214/sf.108
Today’s crane-mounted planting heads plant seedlings with biologically similar or better results than operational manual planting. However, the total cost of mechanized tree planting in southern Sweden must decrease at least 25% to compete economically with manual planting. Although seedlings packed in machine-specific packaging increase the productivity of planting machines by reducing seedling reloading time, they also increase logistics and investment costs. In this study, we analyzed the total cost of outplanting seedlings with an excavator-mounted Bracke Planter and seedlings packed according to four different concepts: cultivation trays, cardboard boxes, band-mounted seedlings in cardboard boxes and linked pots in container modules. The total cost per planted seedling was calculated for each packaging system as the sum of all costs from nursery to the recovery of empty packaging. The results showed that today’s system of transporting seedlings in cultivation trays is the most cost-efficient of the four alternatives. Machine-specific seedling packaging was 16–23% costlier per planted seedling than cultivation trays when trucking distances were 100 km. Sensitivity analyses indicated that machine-specific seedling packaging increased in cost-efficiency relative to cultivation trays primarily when more planting machines were contracted, but also as planting machine fixed costs and productivity increased. Moreover, the relative cost-efficiency of band-mounted seedlings, but not seedlings in container modules, increased with increasing trucking distance. Thus, we show that investments in machine-specific seedling packaging for today’s planting machines are justified only when the fixed costs, productivity and number of contracted planting machines increase substantially.
  • Ersson, Swedish University of Agricultural Sciences, Department of Forest Resource Management, SE-90183 Umeå, Sweden ORCID ID:E-mail: back.tomas.ersson@slu.se (email)
  • Bergsten, Swedish University of Agricultural Sciences, Department of Forest Resource Management, SE-90183 Umeå, Sweden ORCID ID:E-mail:
  • Lindroos, Swedish University of Agricultural Sciences, Department of Forest Resource Management, SE-90183 Umeå, Sweden ORCID ID:E-mail:
article id 108, category Research article
Back Tomas Ersson, Urban Bergsten, Ola Lindroos. (2011). The cost-efficiency of seedling packaging specifically designed for tree planting machines. Silva Fennica vol. 45 no. 3 article id 108. https://doi.org/10.14214/sf.108
Today’s crane-mounted planting heads plant seedlings with biologically similar or better results than operational manual planting. However, the total cost of mechanized tree planting in southern Sweden must decrease at least 25% to compete economically with manual planting. Although seedlings packed in machine-specific packaging increase the productivity of planting machines by reducing seedling reloading time, they also increase logistics and investment costs. In this study, we analyzed the total cost of outplanting seedlings with an excavator-mounted Bracke Planter and seedlings packed according to four different concepts: cultivation trays, cardboard boxes, band-mounted seedlings in cardboard boxes and linked pots in container modules. The total cost per planted seedling was calculated for each packaging system as the sum of all costs from nursery to the recovery of empty packaging. The results showed that today’s system of transporting seedlings in cultivation trays is the most cost-efficient of the four alternatives. Machine-specific seedling packaging was 16–23% costlier per planted seedling than cultivation trays when trucking distances were 100 km. Sensitivity analyses indicated that machine-specific seedling packaging increased in cost-efficiency relative to cultivation trays primarily when more planting machines were contracted, but also as planting machine fixed costs and productivity increased. Moreover, the relative cost-efficiency of band-mounted seedlings, but not seedlings in container modules, increased with increasing trucking distance. Thus, we show that investments in machine-specific seedling packaging for today’s planting machines are justified only when the fixed costs, productivity and number of contracted planting machines increase substantially.
  • Ersson, Swedish University of Agricultural Sciences, Department of Forest Resource Management, SE-90183 Umeå, Sweden ORCID ID:E-mail: back.tomas.ersson@slu.se (email)
  • Bergsten, Swedish University of Agricultural Sciences, Department of Forest Resource Management, SE-90183 Umeå, Sweden ORCID ID:E-mail:
  • Lindroos, Swedish University of Agricultural Sciences, Department of Forest Resource Management, SE-90183 Umeå, Sweden ORCID ID:E-mail:
article id 107, category Research article
Jaana Luoranen, Risto Rikala, Heikki Smolander. (2011). Machine planting of Norway spruce by Bracke and Ecoplanter: an evaluation of soil preparation, planting method and seedling performance. Silva Fennica vol. 45 no. 3 article id 107. https://doi.org/10.14214/sf.107
We evaluated the effects of planting date and planting machine (Bracke: three machines, 69 regeneration areas in three years; Ecoplanter: six areas, two years) on the quality and field performance one and three years after planting of Norway spruce (Picea abies (L.) Karst.) seedlings in central Finland. Both machine types planted on average 1800 seedlings per hectare, and after three years approximately 1600 (Bracke) and 1200 (Ecoplanter) were still alive. This study suggests that planting with a Bracke machine can achieve better regeneration rates than those observed in privately-owned Finnish forests. We characterized the quality of mounding and planting with the Bracke machine as excellent and that of the Ecoplanter as good. The soil preparation method of the Ecoplanter produced humus-rich mounds where seedlings were susceptible to pine weevils and consequently suffered higher mortality. Different machines were used in different regional areas and each machine was operated by different driver/s which may have influenced the results. No negative effects of planting date were observed. Seedling growth decreased if they were tall in relation to their root plug volume, grown too densely in the nursery, and if stored in the field for several months prior to planting. We conclude that mechanized planting is successful when the soil preparation method produces mounds covered by purely mineral soil. Planting from May to the end of September is suitable for seedlings intended for use during this period.
  • Luoranen, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: jaana.luoranen@metla.fi (email)
  • Rikala, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail:
  • Smolander, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail:
article id 104, category Research article
Eeva Terhonen, Teresa Marco, Hui Sun, Risto Jalkanen, Risto Kasanen, Martti Vuorinen, Fred Asiegbu. (2011). The effect of latitude, season and needle-age on the mycota of Scots pine (Pinus sylvestris) in Finland. Silva Fennica vol. 45 no. 3 article id 104. https://doi.org/10.14214/sf.104
The seasonal and latitudinal influences on the diversity and abundance of mycota of Pinus sylvestris needles were investigated. A sample of 1620 needles resulted in a total of 3868 fungal isolates, which were assigned to 68 operational taxonomic units (OTUs). The majority of these OTUs (65%) belong to Ascomycota and only 0.03% was grouped as Basidiomycota. The dominant and most frequently isolated OTU was Hormonema dematioides. Other well-known species with a saprotrophic nutritional mode such as Lophodermium spp. were also observed. The abundance of fungi increased from fall to spring. Frequencies varied significantly in Northern and Southern Finland suggesting that factors associated with latitudinal differences have an impact on the abundance of fungi.
  • Terhonen, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID ID:E-mail:
  • Marco, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID ID:E-mail:
  • Sun, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID ID:E-mail:
  • Jalkanen, Finnish Forest Research Institute, Rovaniemi Research Unit, Rovaniemi, Finland ORCID ID:E-mail:
  • Kasanen, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID ID:E-mail:
  • Vuorinen, Finnish Forest Research Institute, Suonenjoki Research Unit, Suonenjoki, Finland ORCID ID:E-mail:
  • Asiegbu, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID ID:E-mail:
article id 117, category Research article
Harri Kilpeläinen, Jari Lindblad, Henrik Heräjärvi, Erkki Verkasalo. (2011). Saw log recovery and stem quality of birch from thinnings in southern Finland. Silva Fennica vol. 45 no. 2 article id 117. https://doi.org/10.14214/sf.117
The objective of this study was to examine the timber quality of silver birch (Betula pendula Roth) and European white birch (Betula pubescens Ehrh.) trees in the first and second thinnings in southern parts of Finland, from the viewpoint of sawing of small-diameter, short logs, in particular. The average stem volume of birch was 0.140 m3 in the first thinning stands and 0.206 m3 in the second thinning stands. In planted stands, the trees were larger in the first thinnings but slightly smaller in the second thinnings, compared with naturally regenerated pure birch stands or mixed stands of Norway spruce and birch species. Almost 60% of the harvested and 35% of the remaining stems that could provide saw logs were graded as pulpwood for timber quality due to the occurrence of stem defects. The most common stem defects were multiple crooks and middle crooks. Only minor between-stratum differences were detected in the numbers of defects. Depending on the bucking option, the total percentage of saw and plywood logs from the total birch recovery in the thinning of the sample stands varied between 11.7 and 18.2. The recovery of saw logs was clearly higher in the second thinnings, 12–19%, than in the first thinnings, 8–14%. Of the stand types, saw log recovery was the highest in planted birch stands, 12–19%, but lower in naturally regenerated pure birch stands and mixed stands of Norway spruce and birch. The highest share of saw logs was in the second thinning of planted stands, 17–25%. This study shows that the harvesting recoveries of end-use based timber assortments can be estimated in different kinds of thinning birch stands. Based on tree and log dimensions and stem squality, silver birch firstly from plantations and secondly from mixed stands should be the most interesting source of raw material for the saw milling, furniture and interior product sectors.
  • Kilpeläinen, Finnish Forest Research Institute, Eastern Finland Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail:
  • Lindblad, Finnish Forest Research Institute, Eastern Finland Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail:
  • Heräjärvi, Finnish Forest Research Institute, Eastern Finland Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail: henrik.herajarvi@metla.fi (email)
  • Verkasalo, Finnish Forest Research Institute, Eastern Finland Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail:
article id 117, category Research article
Harri Kilpeläinen, Jari Lindblad, Henrik Heräjärvi, Erkki Verkasalo. (2011). Saw log recovery and stem quality of birch from thinnings in southern Finland. Silva Fennica vol. 45 no. 2 article id 117. https://doi.org/10.14214/sf.117
The objective of this study was to examine the timber quality of silver birch (Betula pendula Roth) and European white birch (Betula pubescens Ehrh.) trees in the first and second thinnings in southern parts of Finland, from the viewpoint of sawing of small-diameter, short logs, in particular. The average stem volume of birch was 0.140 m3 in the first thinning stands and 0.206 m3 in the second thinning stands. In planted stands, the trees were larger in the first thinnings but slightly smaller in the second thinnings, compared with naturally regenerated pure birch stands or mixed stands of Norway spruce and birch species. Almost 60% of the harvested and 35% of the remaining stems that could provide saw logs were graded as pulpwood for timber quality due to the occurrence of stem defects. The most common stem defects were multiple crooks and middle crooks. Only minor between-stratum differences were detected in the numbers of defects. Depending on the bucking option, the total percentage of saw and plywood logs from the total birch recovery in the thinning of the sample stands varied between 11.7 and 18.2. The recovery of saw logs was clearly higher in the second thinnings, 12–19%, than in the first thinnings, 8–14%. Of the stand types, saw log recovery was the highest in planted birch stands, 12–19%, but lower in naturally regenerated pure birch stands and mixed stands of Norway spruce and birch. The highest share of saw logs was in the second thinning of planted stands, 17–25%. This study shows that the harvesting recoveries of end-use based timber assortments can be estimated in different kinds of thinning birch stands. Based on tree and log dimensions and stem squality, silver birch firstly from plantations and secondly from mixed stands should be the most interesting source of raw material for the saw milling, furniture and interior product sectors.
  • Kilpeläinen, Finnish Forest Research Institute, Eastern Finland Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail:
  • Lindblad, Finnish Forest Research Institute, Eastern Finland Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail:
  • Heräjärvi, Finnish Forest Research Institute, Eastern Finland Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail: henrik.herajarvi@metla.fi (email)
  • Verkasalo, Finnish Forest Research Institute, Eastern Finland Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail:
article id 116, category Research article
Eva-Maria Nordström, Ljusk Ola Eriksson, Karin Öhman. (2011). Multiple criteria decision analysis with consideration to place-specific values in participatory forest planning. Silva Fennica vol. 45 no. 2 article id 116. https://doi.org/10.14214/sf.116
The combination of multiple criteria decision analysis (MCDA) and participatory planning is an approach that has been applied in complex planning situations where multiple criteria of very different natures are considered, and several stakeholders or social groups are involved. The spatial character of forest planning problems adds further to the complexity, because a large number of forest stands are to be assigned different treatments at different points in time. In addition, experience from participatory forest planning indicates that stakeholders may think about the forest in terms of place-specific values rather than in forest-wide terms. The objective of this study was to present an approach for including place-specific values in MCDA-based participatory forest planning and illustrate the approach by a case study where the objective was to choose a multipurpose forest plan for an area of urban forest in northern Sweden. Stakeholder values were identified in interviews, and maps were used to capture place-specific spatial values. The nonspatial and nonplace-specific spatial values were formulated as criteria and used to build an objective hierarchy describing the decision situation. The place-specific spatial values were included in the creation of a map showing zones of different silvicultural management classes, which was used as the basis for creation of forest plan alternatives in the subsequent process. The approach seemed to work well for capturing place-specific values, and the study indicates that formalized methods for including and evaluating place-specific values in participatory forest planning processes should be developed and tested further.
  • Nordström, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd 1, SE-901 83 Umeå, Sweden ORCID ID:E-mail: eva-maria.nordstrom@slu.se (email)
  • Eriksson, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd 1, SE-901 83 Umeå, Sweden ORCID ID:E-mail:
  • Öhman, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd 1, SE-901 83 Umeå, Sweden ORCID ID:E-mail:
article id 114, category Research article
Roy V. Rea. (2011). Impacts of moose (Alces alces) browsing on paper birch (Betula papyrifera) morphology and potential timber quality. Silva Fennica vol. 45 no. 2 article id 114. https://doi.org/10.14214/sf.114
Although moose browsing effects on the growth and morphology of birch are well studied, effects of moose browsing on potential timber quality of birch have received little attention. Here, an assessment was made of the impacts of moose (Alces alces L.) damage to Paper Birch (Betula papyrifera Marsh.) trees from a 20-year old clear cut area in a sub-boreal spruce forest within the Aleza Lake Research Forest, near Prince George, British Columbia, Canada. Specifically, differences in overall tree architecture and in the internal characteristics of trees that had been severely damaged and suppressed by moose winter browsing were compared to birch trees that had not been damaged by moose in this way and were considered free-to-grow. The average stem diameter, number of annular growth rings, and height of stem breaks made by moose on suppressed birches at the point of breakage was 17.9 ± 6.6 mm, 4.6 ± 1.2, and 141.8 ± 32.0 cm, respectively. Stem diameters and the heights above-the-ground of stem breaks made by moose during sequential breakage events were not significantly different (all p ≥ 0.05) from one another. Decay was significantly (all p ≤ 0.001) more extensive in trees where branches had been broken off by moose than in trees with no breaks or where breaks were from unknown agents. Suppressed birches were significantly (p = 0.048) more exposed (farther from their nearest tree neighbor) when compared to birches that were free-to-grow. The distance from birch trees to species-specific neighbors (of any species) did not differ (all p ≥ 0.05) between suppressed and free-to-grow birches. Suppressed birches damaged from intense browsing and stem breakage were significantly (≤ 0.001) farther away from other birches showing signs of slight to moderate browsing than free-to-grow birches were from similar conspecifics. Because moose appear to impact the potential wood quality of birch, forest managers should consider the impacts that browsing and stem breakage can have on birch timber where these trees co-occur with and are eaten by moose.
  • Rea, University of Northern British Columbia, 3333 University Way, Prince George, BC, Canada V2N 4Z9 ORCID ID:E-mail: reav@unbc.ca (email)
article id 112, category Research article
Mike R. Saunders, Shawn Fraver, Robert G. Wagner. (2011). Nutrient concentration of down woody debris in mixedwood forests in central Maine, USA. Silva Fennica vol. 45 no. 2 article id 112. https://doi.org/10.14214/sf.112
Both nutrient concentrations and pre- and post-harvest pool sizes were determined across down woody debris decay classes of several hardwood and softwood species in a long-term, natural disturbance based, silvicultural experiment in central Maine. Concentrations of N, P, Ca, Mg, Cu, Fe, and Zn generally increased 2- to 5-fold with increasing decay class. Concentrations of Mn, Al and B did not differ among decay classes, while K decreased by 20–44% from decay class 1 to class 4. C:N-ratios declined with increasing decay class, while N:P-ratios increased from decay class 1 to 2 and then plateaued with further decay. Within decay classes, softwoods generally had lower nutrient concentrations and higher C:N-ratios than hardwoods; N:P-ratios did not differ between hardwoods and softwoods. Although gap harvesting increased the size of the overall down woody debris nutrient pools, mostly through a large pulse of decay class 1 material, harvesting generally reduced the nutrients held in advanced decay classes. Pre-harvest down woody debris pools for N, P, K and Ca were 11.0, 0.6, 2.1 and 21.1 kg ha–1, respectively, while postharvest were 20.0, 1.3, 6.2 and 46.2 kg ha–1, respectively. While the gap-based silvicultural systems sampled in this study doubled the size of the pre-harvest, downed woody debris nutrient pools, the post-harvest pools were estimated to be only 3.2–9.1% of aboveground nutrients.
  • Saunders, Department of Forestry and Natural Resources, Purdue University, 715 State Street, West Lafayette, IN, USA ORCID ID:E-mail: msaunder@purdue.edu (email)
  • Fraver, USFS Northern Research Station, Grand Rapids, MN, USA ORCID ID:E-mail:
  • Wagner, School of Forest Resources, University of Maine, Orono, ME, USA ORCID ID:E-mail:
article id 41, category Research article
Ying Hou, Jintao Qu, Zukui Luo, Chao Zhang, Kaiyun Wang. (2011). Morphological mechanism of growth response in treeline species Minjiang fir to elevated CO2 and temperature. Silva Fennica vol. 45 no. 2 article id 41. https://doi.org/10.14214/sf.41
To test whether and how morphological traits are linked with growth responses of plants to temperature and CO2 is important for understanding the mechanism underlying how plant growth will respond to global warming. In this study, using closed-top chambers to mimic future elevated CO2 and temperature, the growth response, morphological traits of Minjiang fir (Abies faxoniana Rehd.et Wils.) and the relationship of the two were investigated after two years of exposure to the single and combined elevation of CO2 and temperature. The results showed that biomass of Minjiang fir was 21%, 31%, and 35% greater than the control in elevated CO2, elevated temperature and the combination of elevated CO2 and temperature treatments, respectively. Elevated CO2 and temperature significantly affected the morphology of Minjiang fir, and a few morphological traits were highly correlated with growth responses. Larger branch angles at the upper layer, crown volume, and relative crown length contributed to positive growth responses to elevated CO2, while decreased specific leaf area (SLA) constricted any further growth response. Leaf morphological traits were more closely correlated with the response ratio than crown did in the elevated temperature, while in the combination of elevated CO2 and temperature, crown was more correlated with the response ratio than the leaf morphological traits. Thus, our results indicate that morphological traits may contribute differently to growth responses under different experimental conditions.
  • Hou, Department of Life Sciences, Shangqiu Normal University, Shangqiu, China ORCID ID:E-mail:
  • Qu, Department of Life Sciences, Shangqiu Normal University, Shangqiu, China ORCID ID:E-mail:
  • Luo, School of Environment and Life Sciences, Kaili University, Kaili, China ORCID ID:E-mail:
  • Zhang, Shanghai Key Laboratory of Urbanization and Ecological Restoration, East China Normal University, Shanghai, China, and University of Eastern Finland, School of Forest Sciences, Joensuu, Finland ORCID ID:E-mail:
  • Wang, Shanghai Key Laboratory of Urbanization and Ecological Restoration, East China Normal University, Shanghai, China, and University of Eastern Finland, School of Forest Sciences, Joensuu, Finland ORCID ID:E-mail: kywang@re.ecnu.edu.cn (email)
article id 34, category Research article
Natalie Macias, Chris Knowles. (2011). Examining the effect of environmental certification, wood source, and price on architects’ preferences of hardwood flooring. Silva Fennica vol. 45 no. 1 article id 34. https://doi.org/10.14214/sf.34
This article examines the importance architects place on three factors, environmental certification, wood source, and price, when specifying hardwood flooring. Architects were presented with nine flooring scenarios, in which the three factors were present in varying levels. They were asked to rank the scenarios from the least preferred to the most preferred. Data were obtained from a mail survey of architects in Oregon and Washington, U.S.A. (n = 402). Conjoint analyses determined that architects consider price and wood source to be the most important factors when specifying hardwood flooring. Interestingly, environmental certification was considered the least important factor. The respondents were then separated into three groups for further analysis based on whether they identified themselves as more influenced by environmental factors (biocentric) or human needs (anthropocentric). This analysis showed that the biocentric group favored wood source over price and environmental certification, while the anthropocentric group favored price.
  • Macias, Oregon State University, Department of Wood Science and Engineering, 119 Richardson Hall, Corvallis, OR 97331, USA ORCID ID:E-mail: natalie.macias@gmail.com
  • Knowles, Oregon State University, Department of Wood Science and Engineering, 119 Richardson Hall, Corvallis, OR 97331, USA ORCID ID:E-mail: chris.knowles@oregonstate.edu (email)
article id 29, category Research article
Piotr Androsiuk, Roman Zielinski, Kornelia Polok. (2011). B-SAP markers derived from the bacterial KatG gene differentiate populations of Pinus sylvestris and provide new insights into their postglacial history. Silva Fennica vol. 45 no. 1 article id 29. https://doi.org/10.14214/sf.29
The aim of the studies was to evaluate the efficiency of the KatG gene based B-SAP markers as a tool to distinguish morphologically diversified and geographically distant Scots pine populations and to track the routes of migrations. The 19 populations growing in the IUFRO 1982 provenance experiment and representing the natural distribution of the species in Europe were scored using 103 B-SAP loci. Among them 26% loci were polymorphic. The level of polymorphism was associated with the location of primers on the KatG template. The diversity was low, He = 0.086, and deposited mostly among populations. Seven unique markers were found that identified populations and likely they were associated with morphology. The overall genetic identity was relatively low, I = 0.933 (D = 0.069). The block of six B-SAP markers discriminated populations into two groups in agreement with their geographic origin and thereby further described as the North and the South. The North group was uniform with genetic diversity, He = 0.026 and the overall genetic distance D = 0.022. Presumably, it migrated from refugia in the Alps via France, northern Germany and Denmark, to Scandinavia and Russia. The South group was heterogeneous with He = 0.063 and D = 0.047. This group migrated from the Carpathians via Slovakia to Germany and Poland. The Balkans and Asian refugia did not take part in recolonization of Europe. The block of six B-SAP/KatG markers can be recommended for tracking postglacial history of Scots pine.
  • Androsiuk, University of Warmia and Mazury in Olsztyn, Department of Genetics, Plac Lodzki 3, 10-967 Olsztyn, Poland ORCID ID:E-mail:
  • Zielinski, University of Warmia and Mazury in Olsztyn, Department of Genetics, Plac Lodzki 3, 10-967 Olsztyn, Poland ORCID ID:E-mail:
  • Polok, University of Warmia and Mazury in Olsztyn, Department of Genetics, Plac Lodzki 3, 10-967 Olsztyn, Poland ORCID ID:E-mail: kpolok@moskit.uwm.edu.pl (email)
article id 451, category Research article
Thomas P. Sullivan, Druscilla S. Sullivan, Pontus M. F. Lindgren, Douglas B. Ransome. (2010). Green-tree retention and life after the beetle: stand structure and small mammals 30 years after salvage harvesting. Silva Fennica vol. 44 no. 5 article id 451. https://doi.org/10.14214/sf.451
We report on a retrospective investigation of the impacts of salvage harvesting of lodgepole pine (Pinus contorta var. latifolia Engelm. ex S. Wats.), killed by an outbreak of mountain pine beetle (Dendroctonus ponderosae Hopk.) in the 1970s, with variable retention of Douglas-fir (Pseudotsuga menziesii (Mirbel) Franco). Our inference to biodiversity was coniferous stand structure and four mammal species: the southern red-backed vole (Myodes gapperi Vigors), common shrew (Sorex cinereus Kerr), red squirrel (Tamiasciurus hudsonicus Erxleben) and northern flying squirrel (Glaucomys sabrinus Shaw). We tested hypotheses that, at 30 years after salvage harvest of beetle-killed lodgepole pine trees, (1) abundance and diversity of stand structure, and (2) abundance of mammal species, will increase with higher levels of green-tree retention (GTR). Stand structure attributes and small mammals were sampled during 2005–2008 in young pine stands, with a range of GTR seed-trees (none, dispersed, and aggregated Douglas-fir), and uncut forest in south-central British Columbia, Canada. Diameters and heights of Douglas-fir and lodgepole pine and basal area of total conifers supported hypothesis (1). Mean abundance of the red-backed vole was consistently higher (2.3 to 6.4 times) in the uncut forest than other stands. Overall mean patterns of abundance for common shrews, red squirrels, and northern flying squirrels were similar among treatment stands. Mean abundance of the red-backed vole supported hypothesis (2), but numbers of the other three species did not. There is “life after the beetle” at 30 years after salvage harvesting, and this was enhanced by GTR.
  • Sullivan, Department of Forest Sciences, Faculty of Forestry, University of BC, 2424 Main Mall, Vancouver, BC, Canada V6T 1Z4 ORCID ID:E-mail: tom.sullivan@ubc.ca (email)
  • Sullivan, Department of Forest Sciences, Faculty of Forestry, University of BC, 2424 Main Mall, Vancouver, BC, Canada V6T 1Z4 ORCID ID:E-mail:
  • Lindgren, Applied Mammal Research Institute, 11010 Mitchell Avenue, Summerland, BC, Canada V0H 1Z8 ORCID ID:E-mail:
  • Ransome, Applied Mammal Research Institute, 11010 Mitchell Avenue, Summerland, BC, Canada V0H 1Z8 ORCID ID:E-mail:
article id 126, category Research article
Anders Roos, Lotta Woxblom, Denise McCluskey. (2010). The influence of architects and structural engineers on timber in construction – perceptions and roles. Silva Fennica vol. 44 no. 5 article id 126. https://doi.org/10.14214/sf.126
This study considers structural engineers’ and architects’ perceptions of structural timber in multi-story construction contexts. Qualitative approaches – interviews and focus groups – were used to investigate attitudes, perceived norms, and perceived factors that hamper or facilitate the prescription of wood use in construction. Wood was perceived as an appropriate building material. Architects’, and even more so engineers’, perceptions of negative aspects of wood focused on decay, instability and sound transmission. Although wood-based construction was seen as a required professional skill, it was not expected to improve one’s professional status. Positive aspects of wood in construction included its strength, environmental friendliness, simple handling and appropriateness for use in conjunction with industrial methods, whereas knowledge gaps and weak support from the wood industry have reduced the use of wood among structural engineers and architects. Both professions perceived their influence on material selection to be weak. They sensed that most of the influence over material selection rested with developers and contractors. The paper contains suggestions on how to make these two professions more influential advocates for wood in construction.
  • Roos, Swedish University of Agricultural Sciences, Department of Forest Products, P.O. Box 7008, SE-750 07 Uppsala, Sweden ORCID ID:E-mail: anders.roos@sprod.slu.se (email)
  • Woxblom, Swedish University of Agricultural Sciences, Department of Forest Products, P.O. Box 7008, SE-750 07 Uppsala, Sweden ORCID ID:E-mail:
  • McCluskey, Swedish University of Agricultural Sciences, Department of Forest Products, P.O. Box 7008, SE-750 07 Uppsala, Sweden ORCID ID:E-mail:
article id 125, category Research article
Juho Rantala, Tiina Laine. (2010). Productivity of the M-Planter tree-planting device in practice. Silva Fennica vol. 44 no. 5 article id 125. https://doi.org/10.14214/sf.125
Need to mechanise tree-planting work have recently increased for many reasons. The newest planting and soil scarification device performing work in Nordic forests is the Finnish M-Planter. This study aims to clarify M-Planter’s productivity in practice and show how various factors affect it. The follow-up data set covers 607 work shifts, of 13 operators with, in total, five M-Planters. The average productivity figures for the operators were 143 and 169 seedlings per effective working hour during the first and second planting season, respectively. Overall, the measured average productivity was 34.2% lower than that observed in an earlier work study of the M-Planter based on an experimental study design. On average, the operators learned to use the combination of the M-Planter and a base machine more efficiently while their experience in using it increased during the follow-up. Increasing number of stones and stumps as well as a thicker humus layer decreased productivity of the M-Planter. The study concludes that utilisation of the full productivity potential of the M-Planter requires not only good operators but also development of the whole planting service supply chain.
  • Rantala, Finnish Forest Research Institute, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: juho.rantala@metla.fi (email)
  • Laine, Finnish Forest Research Institute, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail:
article id 125, category Research article
Juho Rantala, Tiina Laine. (2010). Productivity of the M-Planter tree-planting device in practice. Silva Fennica vol. 44 no. 5 article id 125. https://doi.org/10.14214/sf.125
Need to mechanise tree-planting work have recently increased for many reasons. The newest planting and soil scarification device performing work in Nordic forests is the Finnish M-Planter. This study aims to clarify M-Planter’s productivity in practice and show how various factors affect it. The follow-up data set covers 607 work shifts, of 13 operators with, in total, five M-Planters. The average productivity figures for the operators were 143 and 169 seedlings per effective working hour during the first and second planting season, respectively. Overall, the measured average productivity was 34.2% lower than that observed in an earlier work study of the M-Planter based on an experimental study design. On average, the operators learned to use the combination of the M-Planter and a base machine more efficiently while their experience in using it increased during the follow-up. Increasing number of stones and stumps as well as a thicker humus layer decreased productivity of the M-Planter. The study concludes that utilisation of the full productivity potential of the M-Planter requires not only good operators but also development of the whole planting service supply chain.
  • Rantala, Finnish Forest Research Institute, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: juho.rantala@metla.fi (email)
  • Laine, Finnish Forest Research Institute, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail:
article id 132, category Research article
Mats Berlin, Lars Lönnstedt, Gunnar Jansson, Öje Danell, Tore Ericsson. (2010). Developing a Scots pine breeding objective: a case study involving a Swedish sawmill. Silva Fennica vol. 44 no. 4 article id 132. https://doi.org/10.14214/sf.132
The aim of this study was to develop a Scots pine (Pinus sylvestris L.) breeding objective for a vertically integrated sawmill in northern Sweden. The production system was defined as comprising the sawmill and the forests supplying it. Volume per hectare, wood density, survival and straightness were used as objective traits and the related selection criteria were measurements, collected at young tree age, of height, diameter, pilodyn penetration, vitality and straightness. A bio-economic model was used to calculate economic weights for the objective traits identified. We also investigated the efficiency of different selection indices based on these economic weights, in combination with available data on genetic parameters. Furthermore, we studied the effect of different discount rates on the calculated economic weights. The results showed that, compared to the full index (which included all selection criteria), omitting either vitality or straightness had a negligible effect, reducing predicted profit gain per hectare by less than one per cent. Height or diameter each had a greater effect, with a loss of predicted profit gain per hectare of up to 6%. Excluding pilodyn penetration from the selection index caused the largest reduction in predicted profit gain per hectare, amounting to over 10%. However, when both height and diameter were removed the predicted profit gain per hectare dropped to one-third of that based on the full index. Finally, ranking and genetic selection for the developed breeding objective was insensitive to changes in the discount rate.
  • Berlin, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail: mats.berlin@skogsforsk.se (email)
  • Lönnstedt, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail:
  • Jansson, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail:
  • Danell, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail:
  • Ericsson, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail:
article id 132, category Research article
Mats Berlin, Lars Lönnstedt, Gunnar Jansson, Öje Danell, Tore Ericsson. (2010). Developing a Scots pine breeding objective: a case study involving a Swedish sawmill. Silva Fennica vol. 44 no. 4 article id 132. https://doi.org/10.14214/sf.132
The aim of this study was to develop a Scots pine (Pinus sylvestris L.) breeding objective for a vertically integrated sawmill in northern Sweden. The production system was defined as comprising the sawmill and the forests supplying it. Volume per hectare, wood density, survival and straightness were used as objective traits and the related selection criteria were measurements, collected at young tree age, of height, diameter, pilodyn penetration, vitality and straightness. A bio-economic model was used to calculate economic weights for the objective traits identified. We also investigated the efficiency of different selection indices based on these economic weights, in combination with available data on genetic parameters. Furthermore, we studied the effect of different discount rates on the calculated economic weights. The results showed that, compared to the full index (which included all selection criteria), omitting either vitality or straightness had a negligible effect, reducing predicted profit gain per hectare by less than one per cent. Height or diameter each had a greater effect, with a loss of predicted profit gain per hectare of up to 6%. Excluding pilodyn penetration from the selection index caused the largest reduction in predicted profit gain per hectare, amounting to over 10%. However, when both height and diameter were removed the predicted profit gain per hectare dropped to one-third of that based on the full index. Finally, ranking and genetic selection for the developed breeding objective was insensitive to changes in the discount rate.
  • Berlin, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail: mats.berlin@skogsforsk.se (email)
  • Lönnstedt, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail:
  • Jansson, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail:
  • Danell, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail:
  • Ericsson, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail:
article id 132, category Research article
Mats Berlin, Lars Lönnstedt, Gunnar Jansson, Öje Danell, Tore Ericsson. (2010). Developing a Scots pine breeding objective: a case study involving a Swedish sawmill. Silva Fennica vol. 44 no. 4 article id 132. https://doi.org/10.14214/sf.132
The aim of this study was to develop a Scots pine (Pinus sylvestris L.) breeding objective for a vertically integrated sawmill in northern Sweden. The production system was defined as comprising the sawmill and the forests supplying it. Volume per hectare, wood density, survival and straightness were used as objective traits and the related selection criteria were measurements, collected at young tree age, of height, diameter, pilodyn penetration, vitality and straightness. A bio-economic model was used to calculate economic weights for the objective traits identified. We also investigated the efficiency of different selection indices based on these economic weights, in combination with available data on genetic parameters. Furthermore, we studied the effect of different discount rates on the calculated economic weights. The results showed that, compared to the full index (which included all selection criteria), omitting either vitality or straightness had a negligible effect, reducing predicted profit gain per hectare by less than one per cent. Height or diameter each had a greater effect, with a loss of predicted profit gain per hectare of up to 6%. Excluding pilodyn penetration from the selection index caused the largest reduction in predicted profit gain per hectare, amounting to over 10%. However, when both height and diameter were removed the predicted profit gain per hectare dropped to one-third of that based on the full index. Finally, ranking and genetic selection for the developed breeding objective was insensitive to changes in the discount rate.
  • Berlin, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail: mats.berlin@skogsforsk.se (email)
  • Lönnstedt, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail:
  • Jansson, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail:
  • Danell, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail:
  • Ericsson, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail:
article id 456, category Research article
Sam B. Coggins, Nicholas C. Coops, Michael A. Wulder. (2010). Improvement of low level bark beetle damage estimates with adaptive cluster sampling. Silva Fennica vol. 44 no. 2 article id 456. https://doi.org/10.14214/sf.456
Detection of low level infestation in forest stands is of principle importance to determine effective control strategies before the attack spread to large areas. Of particular concern is the ongoing mountain pine beetle, Dendroctonus ponderosae (Hopkins) epidemic, which has caused approximately 14 million hectares of damage to lodgepole pine (Pinus contorta Dougl. ex. Loud var. latifolia Engl.) forests in western Canada. At the stand level attacked trees are often difficult to locate and can remain undetected until the infestation has become established beyond a small number of trees. As such, methods are required to detect and characterise low levels of attack prior to infestation expansion, to inform management, and to aid mitigation activities. In this paper, an adaptive cluster sampling approach was applied to very fine-scale (20 cm) digital aerial imagery to locate mountain pine beetle damaged trees at the leading edge of the current infestation. Results indicated a mean number of 7.36 infested trees per hectare with a variance of 18.34. In contrast a non-adaptive approach estimated the mean number of infested trees in the same area to be 61.56 infested trees per hectare with a variance of 41.43. Using a relative efficiency estimator the adaptive cluster sampling approach was found to be over two times more efficient when compared to the non-adaptive approach.
  • Coggins, Department of Forest Resources Management, University of British Columbia, 2424 Main Mall, Vancouver, B.C., Canada V6T 1Z4 ORCID ID:E-mail: scoggins@interchange.ubc.ca (email)
  • Coops, Department of Forest Resources Management, University of British Columbia, 2424 Main Mall, Vancouver, B.C., Canada V6T 1Z4 ORCID ID:E-mail:
  • Wulder, Canadian Forest Service (Pacific Forestry Centre), Natural Resources Canada, 506 West Burnside Rd., Victoria, B.C., Canada V8Z 1M5 ORCID ID:E-mail:
article id 455, category Research article
Seppo Kellomäki, Matti Maajärvi, Harri Strandman, Antti Kilpeläinen, Heli Peltola. (2010). Model computations on the climate change effects on snow cover, soil moisture and soil frost in the boreal conditions over Finland. Silva Fennica vol. 44 no. 2 article id 455. https://doi.org/10.14214/sf.455
This study considered how climate change affects the accumulation of snow, the soil moisture and soil frost at sites without tree cover in boreal conditions in Finland (60°–70°N). An increase of 4.5 °C in annual mean temperature and 20 % in annual precipitation were assumed for Finland by the year 2100 according to A2 emission scenario. Along with climate, the soil type of the permanent inventory plots of the Finnish National Forest Inventory was used. Soil and climate data were combined by using a process-based ecosystem model. Calculations were done for four periods: current climate (1971–2000), near future (2001–2020), mid-term future (2021–2050) and long-term future (2071–2100). According to our simulations, the average monthly duration and depth of snow decreased over the simulation period. However, the increasing precipitation may locally increase the snow depths in the mid-term calculations. In the autumn and winter, the average volumetric soil moisture content slightly increased in southern Finland during the near future, but decreased towards the end of the century, but still remained on a higher level than presently. In northern Finland, the soil moisture in the autumn and winter increased by the end of this century. In the summertime soil moisture decreased slightly regardless of the region. Throughout Finland, the length and the depth of soil frost decreased by the end of the century. In the south, the reduction in the depth was largest in the autumn and spring, while in the mid-winter it remained relatively deep in the middle of the century. In the north, the depth tended to increase during the first two calculation periods, in some areas, even during the third calculation period (2071–2100) due to reduced insulation effects of snow during cold spells. The wintertime increase in soil moisture and reduced soil frost may be reflected to reduced carrying capacity of soil for timber harvesting.
  • Kellomäki, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland ORCID ID:E-mail: seppo.kellomaki@uef.fi (email)
  • Maajärvi, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland ORCID ID:E-mail:
  • Strandman, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland ORCID ID:E-mail:
  • Kilpeläinen, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland ORCID ID:E-mail:
  • Peltola, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland ORCID ID:E-mail:
article id 158, category Research article
Ville Hallikainen, Mikko Hyppönen, Leena Pernu, Jouni Puoskari. (2010). Family forest owners’ opinions about forest management in northern Finland. Silva Fennica vol. 44 no. 2 article id 158. https://doi.org/10.14214/sf.158
Forest management guidelines changed at the end of the 1990’s in Finland. Biodiversity, visual landscape, water systems, and different forms of forest use are now better taken into account. The objectives, outdoor recreation motives, and attitudes towards the present forest management activities of the non-industrial private forest owners called family forest owners in this article, whose forest holdings are located in northern Finland, were studied. In addition, a forest owner typology based on the above-mentioned motives, objectives, and attitudes was created, and the relationship between the typology and the forest owners’ background was tested. Principal component analysis, log-linear models, canonical correlations, and K-means cluster analysis were used in the data analysis. The results showed that especially commercial timber production, but also multiple-use forestry, is important for forest owners. Non-timber products such as game, berries, and forest mushrooms were considered more important than biodiversity, conservation of endangered species, tourism, and reindeer herding. The current, more ecological forest management activities were widely accepted by the owners. The changes had been perceived in the forest management activities. Close relationships were found between the objectives, attitudes and motives of the forest owners. Those owners who emphasized ecological tourism and multiple-use forestry, more frequently accepted detailed conservation and other “softer” management methods than those who emphasized commercial timber production. Typologies, called conservationists, timber producers, and multi-objective forest owners, were identified. Forest owner’s education and source of income were closely related to their typology. Highly educated forest owners and those who gained their money from tourism belonged to the groups named conservationists or multi-objective owners, whereas those who lived on forestry income represented timber producers.
  • Hallikainen, Finnish Forest Research Institute, Eteläranta 55, FI-96301 Rovaniemi, Finland ORCID ID:E-mail: ville.hallikainen@metla.fi (email)
  • Hyppönen, Finnish Forest Research Institute, Eteläranta 55, FI-96301 Rovaniemi, Finland ORCID ID:E-mail:
  • Pernu, Finnish Forest Research Institute, Eteläranta 55, FI-96301 Rovaniemi, Finland ORCID ID:E-mail:
  • Puoskari, Finnish Forest Research Institute, Eteläranta 55, FI-96301 Rovaniemi, Finland ORCID ID:E-mail:
article id 157, category Research article
Ville Kankaanhuhta, Timo Saksa, Heikki Smolander. (2010). The effect of quality management on forest regeneration activities in privately-owned forests in southern Finland. Silva Fennica vol. 44 no. 2 article id 157. https://doi.org/10.14214/sf.157
This study describes the effect of quality management on forest regeneration activities in privately-owned forests in southern Finland. The study material consists of two mail surveys conducted in 2006 and 2007. The questionnaires were sent to forestry professionals in Forest Owners’ Associations (FOAs), who had either participated (participants) or not (non-participants) in the forest regeneration quality management interventions in years 2000–2006. Quality management interventions of the FOAs included field inventories, feedback meetings, and education sessions about quality techniques. The activities of participant and non-participant FOAs were compared in terms of the available resources, the actions taken, and the aspirations for improvement. In the FOAs, which had participated in quality management, the number of excavator contractors had increased by 16% more than in the non-participant FOAs. The contractors had acquired 23% more soil preparation equipment under the supervision of the participant forestry professionals. The soil preparation method used in conjunction with Norway spruce that had most increased in use was patch mounding. Seedlings that were 1.5 years and older were used by participant forestry professionals in the planting of Norway spruce 11% more than by non-participants. The planting workers had attended 14% more educational sessions, while the use of self-control measurements in soil preparation and planting density had been adopted 10% more frequently under the supervision of the forestry professionals participating in the quality management. A greater interest in obtaining feedback by using quality control inventories was also found among forestry professionals who participated in quality management.
  • Kankaanhuhta, Finnish Forest Research Institute, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: ville.kankaanhuhta@metla.fi (email)
  • Saksa, Finnish Forest Research Institute, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail:
  • Smolander, Finnish Forest Research Institute, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail:
article id 156, category Research article
Ilkka Korpela, Hans Ole Ørka, Matti Maltamo, Timo Tokola, Juha Hyyppä. (2010). Tree species classification using airborne LiDAR – effects of stand and tree parameters, downsizing of training set, intensity normalization, and sensor type. Silva Fennica vol. 44 no. 2 article id 156. https://doi.org/10.14214/sf.156
Tree species identification constitutes a bottleneck in remote sensing-based forest inventory. In passive images the differentiating features overlap and bidirectional reflectance hampers analysis. Airborne LiDAR provides radiometric and geometric information. We examined the single-trees-level response of two LiDAR sensors in over 13 000 forest trees in southern Finland. We focused on the commercially important species. Our aims were to 1) explore the relevant LiDAR features and study their dependencies on stand and tree variables, 2) examine two sensors and their fusion, 3) quantify the gain from intensity normalizations, 4) examine the importance of the size of the training set, and 5) determine the effects of stand age and site fertility. A set of 570 semiurban broad-leaved trees and exotic conifers was analyzed to 6) examine the LiDAR signal in the economically less important species. An accuracy of 88 90% was achieved in the classification of Scots pine, Norway spruce, and birch, using intensity variables. Spruce and birch showed the highest levels of confusion. Downsizing the training set from 30% to 2.5% of all trees had only a marginal effect on the performance of classifiers. The intensity features were dependent on the absolute and relative sizes of trees, especially for birch. The results suggest that leaf size, orientation, and foliage density affect the intensity, which is thus not affected by reflectance only. Some of the ecologically important species in Finland may be separable, since they gave rise to high intensity values. Comparison of the sensors implies that performance of the intensity data for species classification varies between sensors for reasons that remained uncertain. Both range and gain receiver normalization improved species classification. Weighting of the intensity values improved the fusion of two LiDAR datasets.
  • Korpela, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID ID:E-mail: ilkka.korpela@helsinki.fi (email)
  • Ørka, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, P.O.Box 5003, NO-1432 Ås, Norway ORCID ID:E-mail:
  • Maltamo, University of Eastern Finland, School of Forest Science, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail:
  • Tokola, University of Eastern Finland, School of Forest Science, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail:
  • Hyyppä, Finnish Geodetic Institute, Department of Photogrammetry and Remote Sensing, P.O.Box 15, FI-02431 Masala, Finland ORCID ID:E-mail:
article id 155, category Research article
Minna Räty, Annika Kangas. (2010). Segmentation of model localization sub-areas by Getis statistics. Silva Fennica vol. 44 no. 2 article id 155. https://doi.org/10.14214/sf.155
Models for large areas (global models) are often biased in smaller sub-areas, even when the model is unbiased for the whole area. Localization of the global model removes the local bias, but the problem is to find homogenous sub-areas in which to localize the function. In this study, we used the eCognition Professional 4.0 (later versions called Definies Pro) segmentation process to segment the study area into homogeneous sub-areas with respect to residuals of the global model of the form height and/or local Getis statistics calculated for the residuals, i.e., Gi*-indices. The segmentation resulted in four different rasters: 1) residuals of the global model, 2) the local Gi*-index, and 3) residuals and the local Gi*-index weighted by the inverse of the variance, and 4) without weighting. The global model was then localized (re-fitted) for these sub-areas. The number of resulting sub-areas varied from 4 to 366. On average, the root mean squared errors (RMSEs) were 3.6% lower after localization than the global model RMSEs in sub-areas before localization. However, the localization actually increased the RMSE in some sub-areas, indicating the sub-area were not appropriate for local fitting. For 56% of the sub-areas, coordinates and distance from coastline were not statistically significant variables, in other words these areas were spatially homogenous. To compare the segmentations, we calculated an aggregate standard error of the RMSEs of the single sub-areas in the segmentation. The segmentations in which the local index was present had slightly lower standard errors than segmentations based on residuals.
  • Räty, University of Helsinki, Department of Forest Sciences, P.O. Box 27 (Latokartanonkaari 7), FI-00014 University of Helsinki, Finland ORCID ID:E-mail: minna.s.raty@helsinki.fi (email)
  • Kangas, University of Helsinki, Department of Forest Sciences, P.O. Box 27 (Latokartanonkaari 7), FI-00014 University of Helsinki, Finland ORCID ID:E-mail:
article id 459, category Research article
Tomi Tulokas, Jawdat Tannous. (2010). Research method and improvement of log rotation in sawmills. Silva Fennica vol. 44 no. 1 article id 459. https://doi.org/10.14214/sf.459
Log rotation studies were performed at 14 Finnish sawmills during the years 2003–2005. In the investigation of automatic log rotation, the materialized degree of log rotation for each log was calculated from photos captured from a digital video recording. Rotation errors (D) for individual logs and again the accuracy of the automatic log rotator was determined from optimized angle values of the log measuring system and materialized angle values calculated from the still photos. The accuracy of the log rotation varied considerably between sawmills. The rotation error average of the automatic log rotation varied from –23.6° to +11.4°. This means that in some cases the logs were under-rotated and in some they were over-rotated, on an average. Standard deviation of the rotation error (s) of the automatic rotator varied from 4.4° to 22.9°. The results of the simulation indicated that the performance of the log rotation system can be improved by adjusting the log rotator control. In addition to the zero degree error on an average rotation (or near to zero), the corrected values have a significantly smaller standard deviation of the rotation error, and the number of correct rotations was significantly higher compared to the situation before the adjusted rotation commands. At Sawmill 1, standard deviation of the rotation error was reduced by 40.9% from 14.9 degrees to 8.8 degrees. At the same time the number of correct rotations (–10°≤ D ≤ +10°) increased 4.0 fold from 20.1% to 79.4%. At Sawmill 2, standard deviation of the rotation error was reduced by 23.8% from 10.5 degrees to 8.0 degrees. At this sawmill, the rate of accepted rotations increased 1.9 fold from 42.6% to 81.0%. According to previous research, 2.5° decrease in standard deviation of the rotation error (from 10.5 to 8.0°) in square sawing means about 0.5% increase in value yield. For example with 10 million Û annual sales of sawn timber this means 50 000 Û extra profit.
  • Tulokas, Centre for Timber Engineering, Edinburgh Napier University, 10 Colinton Road, Edinburgh EH10 5DT, UK ORCID ID:E-mail: tomi.tulokas@lut.fi (email)
  • Tannous, School of Engineering and the Built Environment, Edinburgh Napier University, 10 Colinton Road, Edinburgh EH10 5DT, UK ORCID ID:E-mail:
article id 167, category Research article
Berit H. Lindstad, Birger Solberg. (2010). Assessing national compliance with international forest policy processes – the role of subjective judgments. Silva Fennica vol. 44 no. 1 article id 167. https://doi.org/10.14214/sf.167
Several international policy processes with sustainable forest management (SFM) as a common goal have emerged during the past two decades. Based on an empirical study from Norway, this paper analyses the role of subjective judgments in assessing national compliance with three international forest policy processes, and the implications for determination of the effects of these processes. The Expanded Programme of Work on Forest Biological Diversity, the Ministerial Conference on the Protection of Forests in Europe and the United Nations Forum on Forests, including its predecessors, collectively provide more than 600 recommendations for SFM. While it is nothing new that SFM encompasses value questions, this paper is a systematic review of where in a process of assessing national compliance the role of judgments is most profound. The paper shows that the multiple objectives of the forest recommendations, references to national circumstances and provisions for stakeholder involvement lead to differing opinions about the degree of conformity between international recommendations and national situation, i.e. compliance. These differing opinions mean different prospects for the international processes to have effects, because only implementation, or active responses to international recommendations, constitutes effects. The roles of judgments and values are recommended topics for further investigation. Factors influencing how compliance is assessed, and consequently the degree to which implementation is deemed necessary, require specific attention. Due consideration to substantive and methodological choices in determining national changes and in separation of other sources of influence will provide a better basis for informed discussion of compliance with and effects of international forest-related policy processes.
  • Lindstad, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, Ås, Norway ORCID ID:E-mail: berit.lindstad@umb.no (email)
  • Solberg, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, Ås, Norway ORCID ID:E-mail:
article id 159, category Research article
Johan Stendahl, Maj-Britt Johansson, Erik Eriksson, Åke Nilsson, Ola Langvall. (2010). Soil organic carbon in Swedish spruce and pine forests – differences in stock levels and regional patterns. Silva Fennica vol. 44 no. 1 article id 159. https://doi.org/10.14214/sf.159
The selection of tree species is one factor to consider if we want to mitigate carbon dioxide emissions to the atmosphere through forest management. The objectives of this study were to estimate the differences in soil organic carbon (SOC) stocks under Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) forests and to examine causes of differences in the accumulation of carbon in the forest soil. Large-scale inventory data was used to quantify variations in SOC stock in relation to stand type and the accumulation of carbon for spruce and pine stands was analysed by simulation. Based on field data, the national mean SOC stock was 9.2 kg m–2 in spruce dominated stands and 5.7 kg m–2 in pine dominated stands. For both species, the SOC stock, measured in the field inventory, increased significantly with increasing temperature, although at different rates. The SOC stock was larger for spruce under all temperature conditions, but the difference between species diminished with increasing temperature. The simulations indicated that the build-up of SOC over several rotations was 22% higher in spruce stands than in pine stands under similar environmental conditions. The main difference was found to be the greater input of harvest residues for spruce. Further, the simulations showed that ground vegetation contributed considerably more to the litter production under pine than under spruce. On sites where both Scots pine and Norway spruce are considered suitable, the latter should be selected if the aim of the forest management policy is to maximize the accumulation of SOC in the forest. Further, spruce is more favourable for SOC accumulation in areas with cold temperatures and on sites with low productivity.
  • Stendahl, Department of Soil and Environment, P.O. Box 7001, Swedish University of Agricultural Sciences, SE-75007 Uppsala, Sweden ORCID ID:E-mail: johan.stendahl@mark.slu.se (email)
  • Johansson, Department of Soil and Environment, P.O. Box 7001, Swedish University of Agricultural Sciences, SE-75007 Uppsala, Sweden ORCID ID:E-mail:
  • Eriksson, Department of Energy and Technology, P.O. Box 7061, Swedish University of Agricultural Sciences, SE-75007 Uppsala, Sweden ORCID ID:E-mail:
  • Nilsson, Department of Soil and Environment, P.O. Box 7001, Swedish University of Agricultural Sciences, SE-75007 Uppsala, Sweden ORCID ID:E-mail:
  • Langvall, Unit for Field-based Forest Research, Asa Experimental Forest and Research Station, Swedish University of Agricultural Sciences, SE-36030 Lammhult, Sweden ORCID ID:E-mail:
article id 176, category Research article
María Pasalodos-Tato, Timo Pukkala, Antonio Rigueiro-Rodríguez, Esther Fernández-Nunez, María Rosa Mosquera-Losada. (2009). Optimal management of Pinus radiata silvopastoral systems established on abandoned agricultural land in Galicia (north-western Spain). Silva Fennica vol. 43 no. 5 article id 176. https://doi.org/10.14214/sf.176
Timber production has been the main objective in forest production in Galicia for a long time. Nevertheless, factors such as fire risk and the need to obtain non-timber benefits make other production alternatives like silvopastoral systems worth of consideration. Integration of grazing in the production system not only diversifies products and benefits, but also decreases fire risk by enhancing fuel control. Nonetheless, few studies have examined the economic profitability of these systems. This article analyses the economics of silvopastoral systems established on abandoned agricultural soils afforested with Pinus radiata D. Don. Different tree planting densities, discounting rates, grass values and fire risk scenarios were analysed. The technique employed is based on the combination of an optimization algorithm and a simulator of stand growth and grass yield. The most profitable schedules were obtained with initial stand densities of 1500 trees per hectare. However, with high unit values of pasture production (high value of grass), schedules with an initial stand density of 500 trees per hectare were the most profitable. When the risk of fire was included in the analyses, silvopastoral systems were always more profitable than timber production systems. With an assumption that grazing reduces fire risk thinnings should be done earlier and heavier to reduce the expected losses due to fire and to promote grass production. This lengthens the pasture period. In general, rotation lengt
  • Pasalodos-Tato, INIA, Instituto Nacional de Investigación y Tecnología Agraria y Alimentaria. Madrid, Spain ORCID ID:E-mail: pasalodos.maria@inia.es (email)
  • Pukkala, University of East Finland, Joensuu, Finland ORCID ID:E-mail:
  • Rigueiro-Rodríguez, University of Santiago de Compostela, Lugo, Spain ORCID ID:E-mail:
  • Fernández-Nunez, University of Santiago de Compostela, Lugo, Spain ORCID ID:E-mail:
  • Mosquera-Losada, University of Santiago de Compostela, Lugo, Spain ORCID ID:E-mail:
article id 173, category Research article
Erik Hellberg, Torbjörn Josefsson, Lars Östlund. (2009). The transformation of a Norway spruce dominated landscape since pre-industrial times in northern Sweden: the influence of modern forest management on forest structure. Silva Fennica vol. 43 no. 5 article id 173. https://doi.org/10.14214/sf.173
Logging history and the study of reference conditions in Scandinavian boreal forests has tended to focus on Scots pine dominated ecosystems. This paper presents a regional study of pre-industrial forest conditions and examines the effects of the industrial exploitation of ecosystems dominated by Norway spruce in northern Sweden. Historical records covering a period which preceded industrial logging in the study area (1917–1927) were used to obtain quantitative data on forest structure and influence of forest fires. These data were compared with a modern data set (2003) to analyse changes due to the industrial transformation of the forest. The early 20th century landscape was dominated by old, multi-cohorted spruce forests and mixed coniferous forests. It was found that fire affected both the structure and composition of the landscape. In post-burnt areas, even-aged forests dominated by deciduous species were the principal forest type. Between the early and modern data sets, profound changes in tree-species composition and age structure were documented. While the total volume of deciduous species increased substantially, the coverage of forests dominated by deciduous species decreased. There was also a significant increase in pine-dominated forests and in the total volume of pine. The industrial transformation of the studied landscapes has had profound effects on the structure of spruce forests, but much less so on deciduous forests. The study concludes that the present forest structure is a function of past management regimes, and that future transformations of the landscape will continue, thus affecting the natural variability and biodiversity of the forests.
  • Hellberg, Tunstigen 10, SE-831 43 Östersund, Sweden ORCID ID:E-mail:
  • Josefsson, Swedish University of Agricultural Sciences, Department of Forest Ecology and Management, SE-901 83 Umeå, Sweden ORCID ID:E-mail: torbjorn.josefsson@svek.slu.se (email)
  • Östlund, Swedish University of Agricultural Sciences, Department of Forest Ecology and Management, SE-901 83 Umeå, Sweden ORCID ID:E-mail:
article id 171, category Research article
Takuo Nagaike. (2009). Snag abundance and species composition in a managed forest landscape in central Japan composed of Larix kaempferi plantations and secondary broadleaf forests. Silva Fennica vol. 43 no. 5 article id 171. https://doi.org/10.14214/sf.171
Larix kaempferi is the main plantation species in the low-snow, cool-temperate zone of Japan. I studied L. kaempferi plantations of various stand ages in central Japan to examine and compare the effect of stand age on the abundance, size, and species composition of snags (standing dead trees) compared to those in secondary broadleaf forests. Plantations that were older than the standard rotation age had more and larger snags than young plantations, and the species diversity of snags was positively correlated with stand age. Because the density of living planted L. kaempferi showed little correlation with snag variables, whereas that of naturally regenerated tree species was positively correlated with snag variables, the density dependence of snag occurrence was stronger in naturally regenerated trees than in planted L. kaempferi. Snag species that were positively correlated with stand age were the main species in secondary broadleaf forests in this area. Basal area, density, and number of species of snags in standard-rotation plantations were significantly lower than in long-rotation plantations and secondary broadleaf forests. Long-rotation plantations are useful for retaining snags compared to standard-rotation plantations.
  • Nagaike, Yamanashi Forest Research Institute, Masuho, Yamanashi 400-0502, Japan ORCID ID:E-mail: nagaike-zty@pref.yamanashi.lg.jp (email)
article id 186, category Research article
Juho Rantala, Pertti Harstela, Veli-Matti Saarinen, Leo Tervo. (2009). A techno-economic evaluation of Bracke and M-planter tree planting devices. Silva Fennica vol. 43 no. 4 article id 186. https://doi.org/10.14214/sf.186
Techno-economically reasonable mechanization of tree planting has proved to be a difficult task in the Nordic working conditions. Although planting machines and combinations of base machine and planting device have been developed since the 1970s, mechanized planting has not been cost-competitive to manual planting. The aim of this study was to find out work time distributions, productivities, costs and effects of different work difficulty factors on productivities and costs of the state-of-the-art Nordic planting devices, Swedish Bracke and Finnish M-Planter, and to compare the devices with each other. The theory of comparative time studies was the base for the experimental design of this study. In the average working conditions, productivity (E15) of M-Planter (236 seedlings/hour) was 36.0% higher than that of Bracke (174 seedlings/hour). Here, M-Planter performed planting work 23.4% cheaper than Bracke. However, the difference depended greatly on the working conditions; the more stones or stumps the smaller the difference, and the more slash the bigger the difference.
  • Rantala, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: juho.rantala@metla.fi (email)
  • Harstela, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail:
  • Saarinen, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail:
  • Tervo, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail:
article id 179, category Research article
Henna Vartiamäki, Jarkko Hantula, Antti Uotila. (2009). Susceptibility of silver birch pruning wounds to infection by white-rot fungus (Chondrostereum purpureum), a potential bioherbicide. Silva Fennica vol. 43 no. 4 article id 179. https://doi.org/10.14214/sf.179
We artificially inoculated pruning wounds of silver birch (Betula pendula Roth) to study seasonal variation in their vulnerability to infection by the fungal decomposer Chondrostereum purpureum (Pers. ex Fr.) Pouzar. This information is critical to the assessment of incidental infection risks in areas where C. purpureum may be used as a bioherbicide. On seven monthly occasions between April and October 2005, 30 birch trees were pruned to yield a total of 210 experimental trees. On each occasion, 10 trees were inoculated immediately with C. purpureum mycelium, 10 were inoculated with blank inoculum and 10 were only pruned. In the summer of 2007, a survey of 129 experimental trees showed that pruning wounds were most susceptible to infection during May. Treatment with C. purpureum at other times during the growing season also increased the extent of discoloration or decay but the effect was considerably less.
  • Vartiamäki, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland ORCID ID:E-mail: henna.vartiamaki@metla.fi (email)
  • Hantula, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland ORCID ID:E-mail:
  • Uotila, University of Helsinki, Hyytiälä Forestry Field Station, Hyytiäläntie 124, FI-35500 Korkeakoski, Finland ORCID ID:E-mail:
article id 204, category Research article
Gaby Deckmyn, Bostjan Mali, Hojka Kraigher, Niko Torelli, Maarten Op de Beeck, Reinhart Ceulemans. (2009). Using the process-based stand model ANAFORE including Bayesian optimisation to predict wood quality and quantity and their uncertainty in Slovenian beech. Silva Fennica vol. 43 no. 3 article id 204. https://doi.org/10.14214/sf.204
The purpose of this study was to expand an existing semi-mechanistic forest model, ANAFORE (ANAlysing Forest Ecosystems), to allow for the prediction of log quality and the accompanying uncertainty as influenced by climate and management. The forest stand is described as consisting of trees of different cohorts, either of the same or of different species (deciduous or coniferous). In addition to photosynthesis, transpiration, total growth and yield, the model simulates the daily evolution in vessel biomass and radius, parenchyma and branch development. From these data early and latewood biomass, wood tissue composition, knot formation and density are calculated. The new version presented here, includes the description of log quality, including red heart formation of beeches. A Bayesian optimisation routine for the species parameters was added to the stand model. From a given range of input parameters (prior), the model calculates an optimised range for the parameters (posterior) based on given output data, as well as an uncertainty on the predicted values. A case study was performed for Slovenian beech forests to illustrate the main model functioning and more in particular the simulation of the wood quality. The results indicate that the ANAFORE model is a useful tool for analyzing wood quality development and forest ecosystem functioning in response to management, climate and stand characteristics. However, the Bayesian optimization showed that the remaining uncertainty on the input parameters for the chosen stand was very large, due to the large number of input parameters in comparison to the limited stand data.
  • Deckmyn, Research Group Plant and Vegetation Ecology, University of Antwerpen, Universiteitsplein 1, 2610 Antwerpen, Belgium ORCID ID:E-mail: gaby.deckmyn@ua.ac.be (email)
  • Mali, Slovenian Forestry Institute, Vecna pot 2, 1000 Ljubljana, Slovenia ORCID ID:E-mail:
  • Kraigher, Slovenian Forestry Institute, Vecna pot 2, 1000 Ljubljana, Slovenia ORCID ID:E-mail:
  • Torelli, Slovenian Forestry Institute, Vecna pot 2, 1000 Ljubljana, Slovenia ORCID ID:E-mail:
  • Op de Beeck, Research Group Plant and Vegetation Ecology, University of Antwerpen, Universiteitsplein 1, 2610 Antwerpen, Belgium ORCID ID:E-mail:
  • Ceulemans, Research Group Plant and Vegetation Ecology, University of Antwerpen, Universiteitsplein 1, 2610 Antwerpen, Belgium ORCID ID:E-mail:
article id 204, category Research article
Gaby Deckmyn, Bostjan Mali, Hojka Kraigher, Niko Torelli, Maarten Op de Beeck, Reinhart Ceulemans. (2009). Using the process-based stand model ANAFORE including Bayesian optimisation to predict wood quality and quantity and their uncertainty in Slovenian beech. Silva Fennica vol. 43 no. 3 article id 204. https://doi.org/10.14214/sf.204
The purpose of this study was to expand an existing semi-mechanistic forest model, ANAFORE (ANAlysing Forest Ecosystems), to allow for the prediction of log quality and the accompanying uncertainty as influenced by climate and management. The forest stand is described as consisting of trees of different cohorts, either of the same or of different species (deciduous or coniferous). In addition to photosynthesis, transpiration, total growth and yield, the model simulates the daily evolution in vessel biomass and radius, parenchyma and branch development. From these data early and latewood biomass, wood tissue composition, knot formation and density are calculated. The new version presented here, includes the description of log quality, including red heart formation of beeches. A Bayesian optimisation routine for the species parameters was added to the stand model. From a given range of input parameters (prior), the model calculates an optimised range for the parameters (posterior) based on given output data, as well as an uncertainty on the predicted values. A case study was performed for Slovenian beech forests to illustrate the main model functioning and more in particular the simulation of the wood quality. The results indicate that the ANAFORE model is a useful tool for analyzing wood quality development and forest ecosystem functioning in response to management, climate and stand characteristics. However, the Bayesian optimization showed that the remaining uncertainty on the input parameters for the chosen stand was very large, due to the large number of input parameters in comparison to the limited stand data.
  • Deckmyn, Research Group Plant and Vegetation Ecology, University of Antwerpen, Universiteitsplein 1, 2610 Antwerpen, Belgium ORCID ID:E-mail: gaby.deckmyn@ua.ac.be (email)
  • Mali, Slovenian Forestry Institute, Vecna pot 2, 1000 Ljubljana, Slovenia ORCID ID:E-mail:
  • Kraigher, Slovenian Forestry Institute, Vecna pot 2, 1000 Ljubljana, Slovenia ORCID ID:E-mail:
  • Torelli, Slovenian Forestry Institute, Vecna pot 2, 1000 Ljubljana, Slovenia ORCID ID:E-mail:
  • Op de Beeck, Research Group Plant and Vegetation Ecology, University of Antwerpen, Universiteitsplein 1, 2610 Antwerpen, Belgium ORCID ID:E-mail:
  • Ceulemans, Research Group Plant and Vegetation Ecology, University of Antwerpen, Universiteitsplein 1, 2610 Antwerpen, Belgium ORCID ID:E-mail:
article id 201, category Research article
Rui Qi, Véronique Letort, Mengzhen Kang, Paul-Henry Cournède, Philippe de Reffye, Thierry Fourcaud. (2009). Application of the GreenLab model to simulate and optimize wood production and tree stability: a theoretical study. Silva Fennica vol. 43 no. 3 article id 201. https://doi.org/10.14214/sf.201
The GreenLab model was used to study the interaction between source-sink dynamics at the whole tree level, wood production and distribution within the stem, and tree mechanical stability through simulation and optimization. In this first promising numerical attempt, two GreenLab parameters were considered in order to maximize wood production: the sink strength for cambial growth and a coefficient that determines the way the biomass assigned to cambial growth is allocated to each metamer, through optimization and simulation respectively. The optimization procedure that has been used is based on a heuristic optimization algorithm called Particle Swarm Optimization (PSO). In the first part of the paper, wood production was maximized without considering the effect of wood distribution on tree mechanical stability. Contrary to common idea that increasing sink strength for cambial growth leads to increasing wood production, an optimal value can be found. The optimization results implied that an optimal source and sink balance should be considered to optimize wood production. In a further step, the mechanical stability of trees submitted to their self weight was taken into account based on simplified mechanical assumptions. Simulation results revealed that the allocation of wood at the stem base strongly influenced its global deformation. Such basic mechanical criterion can be an indicator of wood quality if we consider further the active biomechanical processes involved in tree gravitropic responses, e.g. formation of reaction wood.
  • Qi, Ecole Centrale Paris, Laboratory of Applied Mathematics, Grande Voie des Vignes, 92295 Chatenay-Malabry, France; Institute of Automation, Chinese Academy of Sciences, LIAMA/NLPR, P.O.Box 2728, Beijing, China ORCID ID:E-mail: qiruitree@gmail.com (email)
  • Letort, Ecole Centrale Paris, Laboratory of Applied Mathematics, Grande Voie des Vignes, 92295 Chatenay-Malabry, France ORCID ID:E-mail:
  • Kang, Institute of Automation, Chinese Academy of Sciences, LIAMA/NLPR, P.O.Box 2728, Beijing, China ORCID ID:E-mail:
  • Cournède, Ecole Centrale Paris, Laboratory of Applied Mathematics, Grande Voie des Vignes, 92295 Chatenay-Malabry, France; INRIA saclay Ile-de-France, EPI Digiplant, Parc Orsay Université, 91893 Orsay cedex, France ORCID ID:E-mail:
  • Reffye, INRIA saclay Ile-de-France, EPI Digiplant, Parc Orsay Université, 91893 Orsay cedex, France; CIRAD, UMR AMAP, Montpellier, F-34000 France ORCID ID:E-mail:
  • Fourcaud, CIRAD, UMR AMAP, Montpellier, F-34000 France ORCID ID:E-mail:
article id 199, category Research article
R. Edward Thomas. (2009). Modeling the relationships among internal defect features and external Appalachian hardwood log defect indicators. Silva Fennica vol. 43 no. 3 article id 199. https://doi.org/10.14214/sf.199
As a hardwood tree grows and develops, surface defects such as branch stubs and wounds are overgrown. Evidence of these defects remain on the log surface for decades and in many instances for the life of the tree. As the tree grows the defect is encapsulated or grown over by new wood. During this process the appearance of the defect in the tree’s bark changes. The defect becomes flatter and its dimension changes. This progressional change in appearance is predictable, permitting the size and location of the internal defect to be reliably estimated. This paper concerns the development and analysis of models for the prediction of internal features. With the advent of surface scanning and external detection systems, the prediction of internal features promises to significantly improve the quality, yield, and value of sawn wood products.
  • Thomas, USDA Forest Service, 241 Mercer Springs Road, Princeton, WV 24740, USA ORCID ID:E-mail: edthomas@gmail.com (email)
article id 196, category Research article
Aaron R. Weiskittel, Laura S. Kenefic, Robert S. Seymour, Leah M. Phillips. (2009). Long-term effects of precommercial thinning on the stem dimensions, form and branch characteristics of red spruce and balsam fir crop trees in Maine, USA. Silva Fennica vol. 43 no. 3 article id 196. https://doi.org/10.14214/sf.196
The effects of precommercial thinning (PCT) on stem dimensions, form, volume, and branch attributes of red spruce [Picea rubens Sarg.] and balsam fir [Abies balsamea (L.) Mill.] crop trees were assessed 25 years after treatment in an even-aged northern conifer stand. Treatments were a uniform 2.4 x 2.4-m spacing and a control (no PCT). The PCT treatment significantly increased individual tree diameter at breast height (DBH), height growth, crown ratio, and crown width, while it reduced the tree height to DBH ratio. PCT also significantly increased stem taper and consequently, regional volume equations overpredicted observed stem volume by 2 to 15%, particularly for the spaced trees. PCT also increased the number and maximum size of branches on the lower bole. The sizes of knots on half of the sampled spruce crop trees in the spaced plots precluded them from being used as select structural lumber; there were no other effects on log grade. Our findings indicate that PCT can have a long-term influence on the structural attributes of individual trees, and that improved stem-volume prediction equations are needed in the Acadian region of North America.
  • Weiskittel, School of Forest Resources, University of Maine, Orono, ME 04469, USA ORCID ID:E-mail: aaron.weiskittel@umit.maine.edu (email)
  • Kenefic, U.S. Forest Service, Northern Research Station, Bradley, ME 04411, USA ORCID ID:E-mail:
  • Seymour, School of Forest Resources, University of Maine, Orono, ME 04469, USA ORCID ID:E-mail:
  • Phillips, School of Forest Resources, University of Maine, Orono, ME 04469, USA ORCID ID:E-mail:
article id 213, category Research article
Bum-Jin Park, Yuko Tsunetsugu, Tamami Kasetani, Takeshi Morikawa, Takahide Kagawa, Yoshifumi Miyazaki. (2009). Physiological effects of forest recreation in a young conifer forest in Hinokage Town, Japan. Silva Fennica vol. 43 no. 2 article id 213. https://doi.org/10.14214/sf.213
It is widely believed that coming into contact with forest environments is somehow beneficial to human well-being and comfort. In Japan, “Shinrin-yoku” (taking in the atmosphere of a forest) has been proposed to be a relaxation activity associated with forest recreation. The purpose of this study was to examine the physiological effects of forest recreation on the autonomic nervous activity. The subjects were twelve male university students (21.8 ± 0.8 years old). On the first day of the experiment, six subjects were sent to a forest area, and the other six to a city area. On the second day, each subject was sent to the area he did not visit on the first day as a cross check. The subjects walked (15 minutes) around their assigned areas before noon, and sat on chairs viewing (15 minutes) the landscapes of their assigned areas in the afternoon. Heart rate variability (HRV), blood pressure, and pulse rate were measured as physiological indices. Measurements were taken at the place of accommodation in the morning, before and after walking, and before and after viewing at their assigned field areas. Pulse rate, diastolic blood pressure and LF/(LF+HF) (LF – low frequency, HF – high frequency) components of HRV were significantly lower in the forest area than in the city area. HF components of HRV tended to be higher in the forest than in the city. In conclusion, the results of the physiological measurements show that forest recreation enabled effective relaxation in people, both of the mind and body.
  • Park, Chiba University, Center for Environment, Health and Field Sciences, Kashiwa-no-ha 6-2-1, 277-0882 Kashiwa, Chiba, Japan ORCID ID:E-mail: bjpark@faculty.chiba-u.jp (email)
  • Tsunetsugu, Chiba University, Center for Environment, Health and Field Sciences, Kashiwa-no-ha 6-2-1, 277-0882 Kashiwa, Chiba, Japan ORCID ID:E-mail:
  • Kasetani, Chiba University, Center for Environment, Health and Field Sciences, Kashiwa-no-ha 6-2-1, 277-0882 Kashiwa, Chiba, Japan ORCID ID:E-mail:
  • Morikawa, Chiba University, Center for Environment, Health and Field Sciences, Kashiwa-no-ha 6-2-1, 277-0882 Kashiwa, Chiba, Japan ORCID ID:E-mail:
  • Kagawa, Chiba University, Center for Environment, Health and Field Sciences, Kashiwa-no-ha 6-2-1, 277-0882 Kashiwa, Chiba, Japan ORCID ID:E-mail:
  • Miyazaki, Chiba University, Center for Environment, Health and Field Sciences, Kashiwa-no-ha 6-2-1, 277-0882 Kashiwa, Chiba, Japan ORCID ID:E-mail:
article id 209, category Research article
Jaana Luoranen, Kyösti Konttinen, Risto Rikala. (2009). Frost hardening and risk of a second flush in Norway spruce seedlings after an early-season short-day treatment. Silva Fennica vol. 43 no. 2 article id 209. https://doi.org/10.14214/sf.209
There have been years in Finland when container seedlings of Norway spruce (Picea abies (L.) Karsten) planted in the summer have been damaged by early-autumn frosts. For August and September plantings, the seedlings can be hardened by means of short-day (SD) treatment, but little information is available about its usability for earlier plantings. We studied the effects of early-season SD treatment on the frost hardiness and risk of a second flush of Norway spruce seedlings. In three successive years, second-year seedlings were grown in a greenhouse or outdoors in the spring and early summer and then subjected to two or three-week SD treatment beginning on the second, third, or fourth week of June. We monitored the height growth cessation, bud formation, and frost hardiness of the seedlings in the nursery. All SD treatments made the height growth cease, but the risk of a second flush increased if the temperature sum was less than 300 d.d. before the beginning of the SD treatment or more than 450 d.d. between the end of the treatment and mid-August. Clearly, then, SD treatment reduced the risk of a second flush in seedlings that had been grown in a greenhouse in the spring. Early-season SD treatment increased the frost hardiness of both needles and stems for late July to early September in comparison with untreated seedlings. Later in the autumn, however, the differences disappeared. Before recommending the use of early-season SD-treated seedlings for summer planting, the method has to be tested in practical field conditions.
  • Luoranen, The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: jaana.luoranen@metla.fi (email)
  • Konttinen, The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland ORCID ID:E-mail:
  • Rikala, The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland ORCID ID:E-mail:
article id 207, category Research article
Mats T. Olsson, Maria Erlandsson, Lars Lundin, Torbjörn Nilsson, Åke Nilsson, Johan Stendahl. (2009). Organic carbon stocks in Swedish Podzol soils in relation to soil hydrology and other site characteristics. Silva Fennica vol. 43 no. 2 article id 207. https://doi.org/10.14214/sf.207
Site characteristics influence soil organic carbon (SOC) stocks. In Podzols under Swedish forest land, SOC stocks were related to latitude, altitude, soil hydrological class categorized by mean groundwater level, mean annual precipitation, temperature sum during the growing season, total annual nitrogen (N) deposition and site capacity. SOC stocks were determined for the O-horizon and for total soil (O-horizon + mineral soil to a depth of 50 cm). Data from the Swedish National Forest Soil Inventory 1993–2001 were used (1477 field plots). The O-horizon was sampled with a core sampler and carbon (C) stocks were determined. For the mineral soil layers the SOC stock was calculated based on the SOC concentrations, bulk density and content of rock fragments. The results showed that the overall mean SOC stock was 2.8 and 8.2 kg C m–2 for O-horizon and total soil, respectively. Soil hydrological class strongly affected SOC stocks, which increased from on average 6.7 kg C m–2 at dry sites to 9.7 kg C m–2 at slightly moist sites. Corresponding values for the O-horizon were 2.0 to 4.4 kg C m–2. The correlation coefficients for the linear relationship between SOC stock and site characteristics were highest for N deposition, which explained up to 25% of variation, and latitude, which explained up to 20% of variation. Altitude had the lowest degree of explanation.
  • Olsson, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden ORCID ID:E-mail:
  • Erlandsson, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden ORCID ID:E-mail:
  • Lundin, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden ORCID ID:E-mail:
  • Nilsson, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden ORCID ID:E-mail: torbjorn.nilsson@mark.slu.se (email)
  • Nilsson, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden ORCID ID:E-mail:
  • Stendahl, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden ORCID ID:E-mail:
article id 218, category Research article
Md. Nurul Islam, Mikko Kurttila, Lauri Mehtätalo, Arto Haara. (2009). Analyzing the effects of inventory errors on holding-level forest plans: the case of measurement error in the basal area of the dominated tree species. Silva Fennica vol. 43 no. 1 article id 218. https://doi.org/10.14214/sf.218
Accurate inventory data are required for ensuring optimal net return on investment from the forest. Erroneous data can lead to the formulation of a non-optimal plan that can cause inoptimality losses. Little is known of the effect of using erroneous stand inventory data in preparing holding-level forest plans. This study reports on an approach for analyzing such inoptimality losses. Furthermore, inoptimality losses caused by measurement errors in the basal area of the dominated tree species were investigated in a case study. Based on the inventory data including routine measurements by 67 measurers, four measurer groups were created with different measurement error profiles for the basal area of the dominated tree species. This was followed by measurement error simulations for each group and by adding these to the accurate control inventory data to create erroneous data of different error profiles. Three different forest plans were then constructed by using erroneous data of each group. The plans were then analyzed and compared with plans based on correct data. The effect of measurement errors on the net present value from the whole planning period, and on the amount of remaining growing stock at the end of planning period, were analyzed and utilized in calculating the inoptimality losses. It was concluded that even errors involving dominated tree species can cause significant changes in the holding-level forest plans.
  • Islam, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland ORCID ID:E-mail: nurul.islam@joensuu.fi (email)
  • Kurttila, Finnish Forest Research Institute, Joensuu Research Unit, FI-80101 Joensuu, Finland ORCID ID:E-mail:
  • Mehtätalo, University of Helsinki, Dept. of Forest Resource Management, FI-00014 University of Helsinki, Finland ORCID ID:E-mail:
  • Haara, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland ORCID ID:E-mail:
article id 466, category Research article
Ilkka Korpela, Tuukka Tuomola, Timo Tokola, Bo Dahlin. (2008). Appraisal of seedling stand vegetation with airborne imagery and discrete-return LiDAR – an exploratory analysis. Silva Fennica vol. 42 no. 5 article id 466. https://doi.org/10.14214/sf.466
The potential for combined use of airborne discrete-return LiDAR and digital imagery in the classification and measurement of common seedling stand vegetation was examined in southern Finland (61°50’N, 24°20’E). Classification was based on spectral and textural image features in addition to geometric and radiometric features of the LiDAR. The accuracy of leaf-on, LiDAR-based terrain elevation models was tested as well as the accuracy of LiDAR in the measurement of vegetation heights. LiDAR-based canopy height and the range-normalized intensity of the LiDAR were strong explanatory variables in vegetation classification. Interspecies variation was observed in the height measurement accuracy of LiDAR for different tree, shrub and low vegetation canopies. Elevation models derived with 1–15 pulses per m2 showed an inherent noise of app. 15–25 cm, which restricts the use of LiDAR in regeneration assessment of very young stands. The spatial pattern of the competing vegetation was reproduced in classification-based raster surfaces, which could be useful in deriving meaningful treatment proposals.
  • Korpela, University of Helsinki, Dept of Forest Management, P.O. Box 27, FI-00014 University of Finland ORCID ID:E-mail: ilkka.korpela@helsinki.fi (email)
  • Tuomola, University of Helsinki, Dept of Forest Management, P.O. Box 27, FI-00014 University of Finland ORCID ID:E-mail:
  • Tokola, University of Helsinki, Dept of Forest Management, P.O. Box 27, FI-00014 University of Finland ORCID ID:E-mail:
  • Dahlin, University of Helsinki, Dept of Forest Management, P.O. Box 27, FI-00014 University of Finland ORCID ID:E-mail:
article id 465, category Research article
Ken Olaf Storaunet, Jørund Rolstad, Målfrid Toeneiet, Erlend Rolstad. (2008). Effect of logging on the threatened epiphytic lichen Usnea longissima: a comparative and retrospective approach. Silva Fennica vol. 42 no. 5 article id 465. https://doi.org/10.14214/sf.465
Usnea longissima is a conspicuous circumboreal “beard lichen” draping tree canopies in moist coastal and mountainous forests. It is extirpated in many European and North-American localities, presumably due to industrial forestry and air pollution, but still has a stronghold in parts of Scandinavia and U.S. and Canadian Pacific Northwest. Using a retrospective approach, we reconstructed the stand histories in 24 plots (0.1 ha) of mature Norway spruce (Picea abies) forest stands in Lillehammer, Norway, of which 21 was selected due to the presence of U. longissima. Number of trees with U. longissima present within plots varied from 4 to 37 and number of visible thalli from 12 to 469. The detailed stand reconstructions were done by means of tree-ring analysis of 517 living trees and the size and decay stage of 1423 stumps from logging and 467 dead trees. Total harvested volume during the last 100 years ranged 100–370 m3ha–1 (representing 40–350% of the present-day standing volume), and present amount of dead wood ranged 2–87 m3ha–1 (1.0–37% of the standing volume). All stands had been selectively logged 2–4 times during the last 100 years, of which 5 stands almost to a clearcut appearance. We used a variety of present-day and historic forest structural variables, both at the scale of study plots and individual trees, to predict the occurrence and abundance of U. longissima. Although most forest stand variables failed in this respect, there were indications of a certain negative influence of the historic logging activity. Number of thalli present on trees showed a unimodal relationship to present-day tree density, indicating that medium dense forest stands are most favorable for U. longissima. We tentatively suggest that selective logging, securing lichen-rich trees, may be a viable management option to keep tree density at a moderate level in the long run, thereby enhancing growth and establishment of U. longissima.
  • Storaunet, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway ORCID ID:E-mail: ken.storaunet@skogoglandskap.no (email)
  • Rolstad, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway ORCID ID:E-mail:
  • Toeneiet, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway ORCID ID:E-mail:
  • Rolstad, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway ORCID ID:E-mail:
article id 465, category Research article
Ken Olaf Storaunet, Jørund Rolstad, Målfrid Toeneiet, Erlend Rolstad. (2008). Effect of logging on the threatened epiphytic lichen Usnea longissima: a comparative and retrospective approach. Silva Fennica vol. 42 no. 5 article id 465. https://doi.org/10.14214/sf.465
Usnea longissima is a conspicuous circumboreal “beard lichen” draping tree canopies in moist coastal and mountainous forests. It is extirpated in many European and North-American localities, presumably due to industrial forestry and air pollution, but still has a stronghold in parts of Scandinavia and U.S. and Canadian Pacific Northwest. Using a retrospective approach, we reconstructed the stand histories in 24 plots (0.1 ha) of mature Norway spruce (Picea abies) forest stands in Lillehammer, Norway, of which 21 was selected due to the presence of U. longissima. Number of trees with U. longissima present within plots varied from 4 to 37 and number of visible thalli from 12 to 469. The detailed stand reconstructions were done by means of tree-ring analysis of 517 living trees and the size and decay stage of 1423 stumps from logging and 467 dead trees. Total harvested volume during the last 100 years ranged 100–370 m3ha–1 (representing 40–350% of the present-day standing volume), and present amount of dead wood ranged 2–87 m3ha–1 (1.0–37% of the standing volume). All stands had been selectively logged 2–4 times during the last 100 years, of which 5 stands almost to a clearcut appearance. We used a variety of present-day and historic forest structural variables, both at the scale of study plots and individual trees, to predict the occurrence and abundance of U. longissima. Although most forest stand variables failed in this respect, there were indications of a certain negative influence of the historic logging activity. Number of thalli present on trees showed a unimodal relationship to present-day tree density, indicating that medium dense forest stands are most favorable for U. longissima. We tentatively suggest that selective logging, securing lichen-rich trees, may be a viable management option to keep tree density at a moderate level in the long run, thereby enhancing growth and establishment of U. longissima.
  • Storaunet, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway ORCID ID:E-mail: ken.storaunet@skogoglandskap.no (email)
  • Rolstad, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway ORCID ID:E-mail:
  • Toeneiet, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway ORCID ID:E-mail:
  • Rolstad, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway ORCID ID:E-mail:
article id 465, category Research article
Ken Olaf Storaunet, Jørund Rolstad, Målfrid Toeneiet, Erlend Rolstad. (2008). Effect of logging on the threatened epiphytic lichen Usnea longissima: a comparative and retrospective approach. Silva Fennica vol. 42 no. 5 article id 465. https://doi.org/10.14214/sf.465
Usnea longissima is a conspicuous circumboreal “beard lichen” draping tree canopies in moist coastal and mountainous forests. It is extirpated in many European and North-American localities, presumably due to industrial forestry and air pollution, but still has a stronghold in parts of Scandinavia and U.S. and Canadian Pacific Northwest. Using a retrospective approach, we reconstructed the stand histories in 24 plots (0.1 ha) of mature Norway spruce (Picea abies) forest stands in Lillehammer, Norway, of which 21 was selected due to the presence of U. longissima. Number of trees with U. longissima present within plots varied from 4 to 37 and number of visible thalli from 12 to 469. The detailed stand reconstructions were done by means of tree-ring analysis of 517 living trees and the size and decay stage of 1423 stumps from logging and 467 dead trees. Total harvested volume during the last 100 years ranged 100–370 m3ha–1 (representing 40–350% of the present-day standing volume), and present amount of dead wood ranged 2–87 m3ha–1 (1.0–37% of the standing volume). All stands had been selectively logged 2–4 times during the last 100 years, of which 5 stands almost to a clearcut appearance. We used a variety of present-day and historic forest structural variables, both at the scale of study plots and individual trees, to predict the occurrence and abundance of U. longissima. Although most forest stand variables failed in this respect, there were indications of a certain negative influence of the historic logging activity. Number of thalli present on trees showed a unimodal relationship to present-day tree density, indicating that medium dense forest stands are most favorable for U. longissima. We tentatively suggest that selective logging, securing lichen-rich trees, may be a viable management option to keep tree density at a moderate level in the long run, thereby enhancing growth and establishment of U. longissima.
  • Storaunet, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway ORCID ID:E-mail: ken.storaunet@skogoglandskap.no (email)
  • Rolstad, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway ORCID ID:E-mail:
  • Toeneiet, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway ORCID ID:E-mail:
  • Rolstad, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway ORCID ID:E-mail:
article id 227, category Research article
Ola Lindroos. (2008).