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Silva Fennica 1926-1997
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Acta Forestalia Fennica
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Articles containing the keyword 'LISA'.

Category: Research article

article id 1260, category Research article
Jyrki Hytönen, Anna Saarsalmi. (2015). Biomass production of coppiced grey alder and the effect of fertilization. Silva Fennica vol. 49 no. 1 article id 1260. https://doi.org/10.14214/sf.1260
Highlights: Fertilisation (wood ash, N, PK) did not affect grey alder biomass production; Leafless above-ground biomass of 17–20 year old stands was 52–57 Mg ha–1; MAI increased with increase of rotation length to the end of the follow-up period of 17–20 years; Coppicing increased stand density manifold.
We studied biomass production of two naturally originated grey alder (Alnus incana (L.) Moench) stands having a mixture of birch and willow located in central Finland. One of the stands was growing on a peatland site (Muhos) and the other on a mineral soil site (Juuka). The stands were clear-cut and fertilization experiments were laid out with several treatments. At Muhos, the treatments included nitrogen fertilisation with different amounts of wood ash and an unfertilized control. At Juuka, the treatments included nitrogen fertilisation either with ash or with PK, and ash and PK treatments alone and an unfertilized control. The sprouts at Muhos were grown for 17 years and at Juuka for 20 years. At Juuka the stand was clear-cut second time at the age of 20 years and grown for 8 years. The stands were measured several times and foliar samples were taken twice during the study period. Clear-cutting increased stem number manifold. The stand density of new coppiced forests after the clear-cutting decreased from 67 000–89 000 stems ha–1 at the age of 3–6 years to 10 000–12 000 stems ha–1 at the age of 17–20 years. On neither site fertilization affected biomass production of alders during the study period. Leafless above-ground biomass was 52–57 Mg ha–1 after 17–20 years. Mean annual leafless above-ground biomass production (MAI) increased with increase of rotation time. At the age of 17–20 years the MAI was 2.8–3.0 Mg ha a–1. At Muhos, ash increased foliar P and Ca concentrations, but decreased those of Mn.
  • Hytönen, Natural Resources Institute Finland, Silmäjärventie 2, FI-69100 Kannus, Finland ORCID ID:E-mail: jyrki.hytonen@luke.fi (email)
  • Saarsalmi, Natural Resources Institute Finland, P.O. Box 18, FI-01301 Vantaa, Finland ORCID ID:E-mail: anna.saarsalmi@luke.fi
article id 933, category Research article
Per-Ola Hedwall, Harald Grip, Sune Linder, Lars Lövdahl, Urban Nilsson, Johan Bergh. (2013). Effects of clear-cutting and slash removal on soil water chemistry and forest-floor vegetation in a nutrient optimised Norway spruce stand. Silva Fennica vol. 47 no. 2 article id 933. https://doi.org/10.14214/sf.933
Fertilisation with nutrient optimisation has in Sweden resulted in large increases in volume growth in young stands of Norway spruce. There are, however, environmental concerns about repeated fertilisation and one is the risk of nutrient leakage to ground water resources and aquatic ecosystems after clear-cutting of such forests. The present study followed soil-water chemistry in optimised fertilised stands after clear-cutting, as well as effects of harvest of slash on nutrient leakage. Parts of a 30-year-old stand of Norway spruce, which had been subject to a nutrient optimisation experiment for 17 years, were clear-cut. A split-plot design with whole-tree harvesting as the sub-plot treatment was applied. Lysimeters were installed and soil-water sampled at nine occasions during the following four years. No significant effects of fertilisation on nitrate leaching were found, while harvest of slash reduced the concentration of Ca, DOC, DON, K, Mg, ammonium and nitrate, as well as pH in the soil solution. While no effects of fertilisation could be seen on the soil water concentration of N, the results indicate an interaction between fertilisation and harvest of slash on the concentration of nitrate in the soil solution. The results indicate that forest-floor vegetation plays an important role in the retention of N after clear-cutting of fertilised forests.
  • Hedwall, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences (SLU), P.O. Box 49, SE-230 53 Alnarp, Sweden ORCID ID:E-mail: per-ola.hedwall@slu.se (email)
  • Grip, Department of Forest Ecology and Management, SLU, SE-901 83 Umeå, Sweden ORCID ID:E-mail: harald@grip2.se
  • Linder, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences (SLU), P.O. Box 49, SE-230 53 Alnarp, Sweden ORCID ID:E-mail: sune.linder@slu.se
  • Lövdahl, Department of Forest Ecology and Management, SLU, SE-901 83 Umeå, Sweden ORCID ID:E-mail:
  • Nilsson, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences (SLU), P.O. Box 49, SE-230 53 Alnarp, Sweden ORCID ID:E-mail: urban.nilsson@slu.se
  • Bergh, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences (SLU), P.O. Box 49, SE-230 53 Alnarp, Sweden ORCID ID:E-mail: johan.bergh@slu.se
article id 912, category Research article
Heidi Hallongren, Juho Rantala. (2012). Commercialisation and international market potential of Finnish silvicultural machines. Silva Fennica vol. 46 no. 4 article id 912. https://doi.org/10.14214/sf.912
Recently, the need to mechanise silvicultural operations has increased in the Nordic countries. While several new machine concepts have been developed, the manufacture of silvicultural devices remains at the market introduction stage. Information is required in support of existing and forthcoming Finnish small-scale producers of silvicultural devices, who wish to commercialise and further market their innovations in domestic and export markets. The aim of this study was to identify the opportunities, challenges and market potential of business activities that develop in connection with device or machine production. Small-scale Finnish manufacturers of silvicultural devices, Finnish large-scale harvester manufacturers and international silvicultural experts participated in the study. The results show that participant groups have varying opinions of cooperation methods and export activities, as well as of the export markets with the best potential. According to international silvicultural experts, mechanised planting and pre-commercial thinning have the greatest potential worldwide. However, demand for mechanised pre-commercial thinning and planting has been mainly confined to the Nordic countries. For a foreign firm marketing a new silvicultural machine concept, the most important customers and cooperation partners are locally operating forest firms, machine contractors and research organisations. The results of the study provide a useful overview of the current state of silvicultural device manufacturing in Finland.
  • Hallongren, Finnish Forest Research Institute, Suonenjoki Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: heidi.hallongren@metla.fi (email)
  • Rantala, Metsä Group, Tampere, Finland ORCID ID:E-mail: juho.rantala@metsagroup.com
article id 75, category Research article
Silvia Lamedica, Emanuele Lingua, Ionel Popa, Renzo Motta, Marco Carrer. (2011). Spatial structure in four Norway spruce stands with different management history in the Alps and Carpathians. Silva Fennica vol. 45 no. 5 article id 75. https://doi.org/10.14214/sf.75
In Europe most Norway spruce (Picea abies (L.) Karst.) mountain forests have been altered by human activities, leading to a lack of reference condition concerning their original structure. Nonetheless, remnants of Norway spruce primeval forests still exist in the Carpathians. Our objective was to assess the differences in structure between managed and unmanaged stands, concerning diameter distributions, amount of standing deadwood, spatial distributions and spatial structure of trees. We established four permanent plots: one in a virgin forest in the Eastern Carpathians and three in a previously managed forest in the Alps. In each plot, species, DBH, and position of the live and dead standing trees were collected. Spatial distribution and structure of all the trees was analysed through several indices. In the Carpathians forest there are clear signs of natural density-dependent mortality processes whereas in the Alpine plots such dynamics are less evident. In these latter plots, the lower snags volume and the random trees spatial distribution can be considered the legacies of past management. Nonetheless, despite the different history of the four stands, they all seem to converge towards a similar spatial structure with the presence of groups (30–40 m) of trees of similar size.
  • Lamedica, Forest Ecology Research Unit, Dept TeSAF, University of Padova, Legnaro (PD), Italy ORCID ID:E-mail:
  • Lingua, Forest Ecology Research Unit, Dept TeSAF, University of Padova, Legnaro (PD), Italy ORCID ID:E-mail:
  • Popa, Forest Research and Management Institute, Research Station for Norway Spruce Silviculture, Câmpulung Moldovenesc, Romania ORCID ID:E-mail:
  • Motta, Department AGROSELVITER, University of Torino, Grugliasco (TO), Italy ORCID ID:E-mail:
  • Carrer, Forest Ecology Research Unit, Dept TeSAF, University of Padova, Legnaro (PD), Italy ORCID ID:E-mail: marco.carrer@unipd.it (email)
article id 115, category Research article
Marjut Turtiainen, Kauko Salo, Olli Saastamoinen. (2011). Variations of yield and utilisation of bilberries (Vaccinium myrtillus L.) and cowberries (V. vitis-idaea L.) in Finland. Silva Fennica vol. 45 no. 2 article id 115. https://doi.org/10.14214/sf.115
So far, only rough estimates for the utilisation rates of wild berries in Finland have been available. One reason for this is that there has been a lack of empirical-knowledge-based studies concerning total yields of wild berries and their yield variations. This study had three aims: 1) total bilberry and cowberry yields of an average crop year were calibrated for different (abundant and poor) crop years using the inventory data on wild berries collected by the Finnish Forest Research Institute (1997–2008); 2) national utilisation rates of bilberries and cowberries were calculated for three different berry years 1997–1999; and 3) regional utilisation rates of these berry species were calculated for the year 1997. According to calculations, annual bilberry yields in Finland vary from 92 to 312 million kg. For cowberry, the range of variation in total berry yields is from 129 to 386 million kg. It was also found that approximately the same proportion of the total yield of bilberries (i.e. 5–6%) was collected between 1997 and 1999. Utilisation rates of cowberries were also quite constant varying from approximately 8% to nearly 10%. In 1997, bilberries and cowberries were utilised most intensively in the eastern parts of the country and in the Oulu-Kainuu region. The results of this present study describe the situation before the phenomenon of foreign pickers. It can be presumed that commercial wild berry picking by migrant collectors has so far affected both national and regional utilisation rates of wild berries.
  • Turtiainen, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: marjut.turtiainen@uef.fi (email)
  • Salo, Finnish Forest Research Institute, Joensuu Research Unit, Joensuu, Finland ORCID ID:E-mail:
  • Saastamoinen, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail:
article id 506, category Research article
Teijo Palander, Katja Turunen, Sanna Laukkanen. (2003). Attitude of Finnish timber buyers towards implementation of a forest computer visualisation. Silva Fennica vol. 37 no. 2 article id 506. https://doi.org/10.14214/sf.506
Timber buyers’ opinions as regards forest computer visualisation are studied. The results indicated that timber buyers are still rather conventional in their computer use since they mainly use only the information system of their own company. The majority of the buyers perceived computer visualisation to be slightly useful for their work, or they had no opinion concerning the usefulness of it. One third of the buyers considered computer visualisation to be a quality factor for timber trade, and the majority were willing to use it if a program is going to adopt by their company. In analysis, different personal characteristics were found for three timber buyer groups: qualified, neutral and reluctant. Qualified buyers were the largest one, about half of the buyers belonging to this group. The greatest barriers to adopting a positive attitude to computer visualisation turned out to be weak computer skills and a general lack of interest in computer use. On the other side, it was found that organisational factors did not influence the buyers’ computer skills or attitudes towards visualisation. The results of this study can be utilised by timber buying organisation in ensuring the successful adoption of a new computer system.
  • Palander, Faculty of Forestry, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland ORCID ID:E-mail: teijo.palander@joensuu.fi (email)
  • Turunen, Faculty of Forestry, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland ORCID ID:E-mail:
  • Laukkanen, Faculty of Forestry, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland ORCID ID:E-mail:
article id 540, category Research article
Finn H. Brække, Nagwa Salih. (2002). Reliability of foliar analyses of Norway spruce stands in a Nordic gradient. Silva Fennica vol. 36 no. 2 article id 540. https://doi.org/10.14214/sf.540
Norway spruce stands at eleven sites in Finland, Norway and Sweden have been studied under various climates, atmospheric deposition of N and S and fertilisation regimes. Nitrogen was growth restricting at eight inland sites, while P was growth restricting at three coastal sites. Liming and N fertilisation caused serious B deficiency on some of the inland sites. It is likely that liming affects uptake of B, whereas N fertilisation causes a dilution due to increased growth. Application of S combined with N probably caused K deficiency at one of the sites. The reliability of foliar analyses as a method to diagnose nutrient status and the likely changes after nutrient input to spruce forests in the Nordic countries, are discussed. The CR- and the DOP-method are evaluated for diagnostic purposes. Both methods seem to give reliable conclusions even if the CR-method often produces more specific results. Interpretation based on both current and one year old foliage improved the diagnostic prognoses. The accuracy of diagnosis also relies on knowledge and ability of the interpreter. Based on the results it is reason to be cautious about recommendations of single element fertilisations, e.g. with N alone, because the demand of other elements beyond available pools frequently occurs. Forest trees in the boreal region are probably well adapted to N deficiency, which means that they can handle the physiological consequences rather well, while deficiencies of other elements usually are more detrimental to growth vigour and stress related diseases.
  • Brække, Department of Forest Sciences, Agricultural University of Norway (AUN), P.O. Box 5044, N-1432 Ås, Norway ORCID ID:E-mail: finn.braekke@isf.nlh.no (email)
  • Salih, Department of Ecology and Environmental Research, Swedish University of Agricultural Sciences (SLU), P.O. Box 7072, S-750 07 Uppsala, Sweden ORCID ID:E-mail:
article id 625, category Research article
Hans-Örjan Nohrstedt. (2000). Effects of soil scarification and previous N fertilisation on pools of inorganic N in soil after clear-felling of a Pinus sylvestris (L.) stand. Silva Fennica vol. 34 no. 3 article id 625. https://doi.org/10.14214/sf.625
Previous analyses of soil water beneath mounds resulting from scarification have implied that this forestry measure increases leaching of inorganic N. However, more recent soil-water studies have not confirmed this assumption. The soil study presented here examined the pools of inorganic N in different microsites emanating from a simulated disc trenching, i. e. the mound with underlying soil, the furrow bottom and the undisturbed soil. The study was made five years after scarification. The mound itself with underlying soil had a larger pool of inorganic N than the undisturbed soil. This was mainly because of an increase in the embedded humus layer, thus implying a larger net N mineralisation and/or lower losses. However, when pools of inorganic N per hectare were calculated, taking into consideration that a scarified area comprises 25% mounds, 25% furrows and 50% undisturbed soil, there was no increase in pools of inorganic N when compared with an area not subjected to scarification. This observation supports the finding of the more recent soil-water studies mentioned, i. e., that leaching seems not to be influenced by soil scarification. The scarification was made as a split-plot treatment on main-plots in an old experiment with different N doses. Thus, the effect of the previous N fertilisation could also be evaluated. Two N doses were tested beside the unfertilised control: 720N (3 x 240 kg N ha–1 yr–1) and 1800N (3 x 600 kg N ha–1 yr–1). The last fertiliser application was made six years before the clearcutting and 13 years before the soil sampling. The previously fertilised main-plots had larger pools of inorganic N than the control plots.
  • Nohrstedt, The Forestry Research Institute of Sweden, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail: hans-orjan.nohrstedt@skogforsk.se (email)
article id 656, category Research article
Klaus Silfverberg, Markus Hartman. (1999). Effects of different phosphorus fertilisers on the nutrient status and growth of Scots pine stands on drained peatlands. Silva Fennica vol. 33 no. 3 article id 656. https://doi.org/10.14214/sf.656
The aim of the study was to compare the effects of phosphorus fertilisers of different solubility and different phosphorus doses. The material was collected from 8 field experiments situated on drained peatlands in southern and central Finland (60°–65°N). The sites were drained, oligotrophic pine fens and pine bogs, which had been fertilised between 1961 and 1977 with different combinations of N, K and P. In 1991–94 stand measurements and foliar and peat sampling were carried out on 162 sample plots. Apatite, rock phosphate and superphosphate affected basal area growth to a rather similar extent. However, apatite slightly surpassed superphosphate and rock phosphate at the end of the study period in two hollow-rich S. fuscum bogs. Higher doses of phosphorus did not significantly increase the basal area growth. The foliar phosphorus concentrations clearly reflected the effect of the P fertilisation. Especially on the pine bogs basic fertilisation with 66 kg P/ha maintained the needle phosphorus concentrations at a satisfactory level for more than 25 years after fertilisation. The amount of phosphorus in the 0–20 cm peat layer was not significantly increased either by basic fertilisation or refertilisation. The phosphorus reserves in the peat in the individual experiments were between 88 and 327 kg/ha. There was a strong correlation between the amounts of phosphorus and iron in the peat. Large amounts of iron in peat may reduce the solubility and availability of phosphorus. According to the foliar phosphorus concentrations in the basic-fertilised plots, the need for refertilisation seems to be unnecessary during the 25-year postfertilisation period at least. None of the basic fertilisation treatments seriously retarded the basal area growth compared to the refertilised treatments. There seems to be a greater shortage of potassium than of phosphorus, because the foliar potassium concentrations and the amounts of potassium in the 0–20 cm peat layer were very low in several of the experiments.
  • Silfverberg, The Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland ORCID ID:E-mail: klaus.silfverberg@metla.fi (email)
  • Hartman, The Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland ORCID ID:E-mail:
article id 662, category Research article
Lars Björklund. (1999). Identifying heartwood-rich stands or stems of Pinus sylvestris by using inventory data. Silva Fennica vol. 33 no. 2 article id 662. https://doi.org/10.14214/sf.662
Variations in heartwood percentage, heartwood radius and sapwood width, within and between stands of Scots pine (Pinus sylvestris L.), were analysed using a database of 198 CT-scanned (computer tomography) stems from 33 research plots (stands) throughout Sweden. Heartwood percentage varied greatly both between individual trees and between stands, and correlated poorly to site, stand and tree variables. This implies that it seems unfeasible to identify heartwood-rich stands or stems, e.g., for production of heartwood products, by using inventory data. Heartwood formation expressed as the number of new heartwood rings formed each year was found to increase with increasing cambial age, from about 0.5 rings per year at a cambial age of 45 years, to about 0.8 rings per year at a cambial age of 115 years.
  • Björklund, Swedish University of Agricultural Sciences, Department of Forest Management and Products, S-750 07 Uppsala, Sweden ORCID ID:E-mail: lars.bjorklund@sh.slu.se (email)

Category: Article

article id 7360, category Article
Viljo Kujala. (1942). Über die morphologische Deutung des Fichtenzapfens auf Grund eines Fertilisationsfalles. Acta Forestalia Fennica vol. 50 no. 7 article id 7360. https://doi.org/10.14214/aff.7360
English title: Morphologic interpretation of a spruce cone based on fertilization.
Original keywords: Fichte; Zapf; Fertilisation; Morfologie
English keywords: spruce; cone; fertilization; morphology

Article presents some rare cone formations found from a small spruce in region of Punkaharju, Finland. In this case all the scales have developed into perfect, green needle leaves. Instead of a normal scale of a cone there are formations of buds that on the basal part of a cone resemble a normal bud, and hence develop into a branch. On the upper part of the cone formation the bud develops into scale of a cone.

The morphological form of the cones has been interpreted many ways. These anomalies resemble earlier observations and hence support the former theory of inflorescence. According the theory, the scales and scales of a cone are separate leaves, the scale corresponding to one leaf, the scale of a cone corresponding to two leaves. A cone is hence rather to be compared with a branching inflorescence than only one flower.

The PDF contains a summary in Finnish. 

  • Kujala, ORCID ID:E-mail:
article id 5081, category Article
Reijo Jokinen. (1980). Metsälannoituksella saatavan kasvunlisäyksen arviointi. Silva Fennica vol. 14 no. 3 article id 5081. https://doi.org/10.14214/sf.a15022
English title: Estimation of growth response achieved through forest fertilization.

Dependence of the growth increase given by fertilization on different stand characteristics is examined in this article. The aim was to determine whether the volume growth increase can be accurately determined beforehand when fertilization is carried out on mineral soil sites at a dosage of 120 kg N/ha. The material consisted mostly on of mature stands ready for cutting, a total of 22 Scots pine (Pinus sylvestris L.) and 20 Norway spruce (Picea abies (L.) H. Karst.) stands. Increase in basal area, height quality class and basal area of the stand were found to best explain the increment and its increase in the regression equations calculated for different types of fertilizer and the control level.

The PDF includes a summary in English.

  • Jokinen, ORCID ID:E-mail:
article id 4886, category Article
Matti Keltikangas, Kustaa Seppälä. (1973). Metsälannoituksen edullisuuden vaihtelu. Silva Fennica vol. 7 no. 3 article id 4886. https://doi.org/10.14214/sf.a14723
English title: Variations in the profitability of forest fertilization.

About one million hectares of forests are fertilized annually in Finland. The goal of the present study was to find out, by means of calculations, how the profitability of forest fertilization varies with variations in the stage of development of tree crops, the quality of the site and its geographical location. Calculations concerned bot fertilization of forests in mineral soil sites and in drained peatlands. The study is a part of a larger project concerning the order of profitability of different forest improvement measures in different conditions. The problems dealt with in this study were approached from the point of view of national economy.

On the basis of two empirical materials it is shown that there is a high correlation between the stand growth percentages before and after the fertilization. Applying the results to existing yield tables the authors calculate benefit/cost ratios showing the stage of development of the stand, the quality of the site and its geographical location. According to the results, fertilization is more profitable in sites of medium fertility than on poor sites. Profitability decreases rather fast from south to north and with decreasing timber prices. 

The PDF includes a summary in English.

  • Keltikangas, ORCID ID:E-mail:
  • Seppälä, ORCID ID:E-mail:
article id 4886, category Article
Matti Keltikangas, Kustaa Seppälä. (1973). Metsälannoituksen edullisuuden vaihtelu. Silva Fennica vol. 7 no. 3 article id 4886. https://doi.org/10.14214/sf.a14723
English title: Variations in the profitability of forest fertilization.

About one million hectares of forests are fertilized annually in Finland. The goal of the present study was to find out, by means of calculations, how the profitability of forest fertilization varies with variations in the stage of development of tree crops, the quality of the site and its geographical location. Calculations concerned bot fertilization of forests in mineral soil sites and in drained peatlands. The study is a part of a larger project concerning the order of profitability of different forest improvement measures in different conditions. The problems dealt with in this study were approached from the point of view of national economy.

On the basis of two empirical materials it is shown that there is a high correlation between the stand growth percentages before and after the fertilization. Applying the results to existing yield tables the authors calculate benefit/cost ratios showing the stage of development of the stand, the quality of the site and its geographical location. According to the results, fertilization is more profitable in sites of medium fertility than on poor sites. Profitability decreases rather fast from south to north and with decreasing timber prices. 

The PDF includes a summary in English.

  • Keltikangas, ORCID ID:E-mail:
  • Seppälä, ORCID ID:E-mail:
article id 4822, category Article
Eino Oinonen. (1970). Metsäkasvien kloonikasvustot maanteiden iän arvioimisen apuneuvoina. Silva Fennica vol. 4 no. 3 article id 4822. https://doi.org/10.14214/sf.a14611
English title: Clone stands of forest plants as aids for estimating the age of roads.

Clone stands of bracken fern (Pteridium aquilinum (L.) Kuhn.) wood small reed (Calamagrostis epigeios L.) and lily-of-the-valley (Convallaria majalis L.) are often partly split into two by the road, but often encountered also unilaterally on the roadside in the shape of a semicircle. The unilateral stands can be at times 20–30 m wide and they are sometimes solitary stands of the species. A method to define the age of the solitary stands of six plant species including bracken, wood small reed and lily-of-the-valley was developed in a series of earlier studies.

These stands can be used to define the time the road was built. Clones that are bound by the road unilaterally are younger than the road. If there are several unilateral clones and they are of different sizes, the road is older than the largest clone. When the road is skirted bilaterally only by clones divided by the road, it is younger than the smallest clone. When there are by the road side both unilaterally delimited clones and clones split by the road, the age of the road comes in the range of time determined by the age difference between the largest unilateral and smallest bilateral clone.

The PDF includes a summary in English.

  • Oinonen, ORCID ID:E-mail:
article id 7187, category Article
Eino Oinonen. (1968). Lycopodium clavatum L.- ja L. annotium L.-kasvustojen laajuus rinnastettuna samanpaikkaisiin L. complanatum L.- ja Pteridium aquilinum (L.) Kuhn -esiintymiin sekä puuston ikään ja paloaikoihin. Acta Forestalia Fennica no. 87 article id 7187. https://doi.org/10.14214/aff.7187
English title: The size of Lycopodium clavatum and L. Annotinum stands as compared to that of L. Complanatum and Pteridium aquilinum stands, the age of the stand and the dates of fire, on the site.

The present study is a part of larger project into the size and age of certain forest plants. This study seeks to confirm the size-age relationship of Lycopodium clavatum L. and L. annotium L. stands, and the time of sporal regeneration. The stand dimensions were plotted against the size of bracken (Pteridium aquilinum (L.) Kuhn) and ground pine (Lycopodium complanatum L.) stands growing on the same site, and against the age of the timber and the time of fire on the site.

The method based on parallel measurements carried out in burned forests has proved to be suitable for establishing the rate of spreading of stands and their age. The three club-moss species proved to be much alike in their sporal regeneration. They, as well as bracken, regenerate under the condition created by fire. The largest detached patches are often clones. Both Lycopodium clavatum and L. annotium survive fire poorly. Thus, the maximum size of the clones is usually connected with the last forest fire. Pieces of the shoots may survive the fires.

Sporal regeneration of these species also occur without the aid of fire. As also variation in the annual growth of the shoots can be considerable, the stand size of these species is not as good indicator of the date of fire as with Pteridium aquilinum and L. complanatum. The winding appearance of the shoots increases the error in determination of stand age by the stand size further.

The results emphasize the importance of taking into account the time and spread of the plants in the traditional vegetation analysis. A central question is: what is an individual.

The PDF includes a summary in English.

  • Oinonen, ORCID ID:E-mail:

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