Current issue: 53(2)

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Silva Fennica 1926-1997
Acta Forestalia Fennica

Articles containing the keyword 'deciduous'.

Category: Research article

article id 45, category Research article
Guolei Li, Yong Liu, Yan Zhu, Qing Mei Li, R. Karsten Dumroese. (2012). Effect of fall-applied nitrogen on growth, nitrogen storage and frost hardiness of bareroot Larix olgensis seedlings. Silva Fennica vol. 46 no. 3 article id 45.
Nursery response of evergreen trees to fall fertilization has been studied widely, but little attention has been given to deciduous trees. Bareroot Olga Bay larch (Larix olgensis Henry) seedlings were fertilized in the nursery with urea at four rates (0, 30, 60, 90 kg N ha–1), with half of each rate applied on two dates (September 16 and October 1, 2009). The seedlings were excavated for evaluation on October 15. In the unfertilized (control) treatment, root and shoot dry mass increased by 100% and 57% respectively, while N concentration in the roots and shoots increased by 43% and 40% during the 30 day period. This indicated that substantial biomass growth during this period did not lead to internal nutrient dilution. Root dry mass increased when fall fertilization rates were ≥ 60 kg N ha–1. Fall fertilization increased N concentrations in root tissue by 48–73%. Compared with the control, shoot tissues of fall fertilized seedlings had slightly higher N concentration and content and significantly higher frost hardiness.
  • Li, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing 100083, China ORCID ID:E-mail:
  • Liu, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing 100083, China ORCID ID:E-mail: (email)
  • Zhu, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing 100083, China ORCID ID:E-mail:
  • Li, Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Forest Silviculture of State Forestry Administration, Beijing 100091, China ORCID ID:E-mail:
  • Dumroese, US Department of Agriculture, Forest Service, Rocky Mountain Research Station, Moscow, ID, USA ORCID ID:E-mail:
article id 43, category Research article
Anni Markkanen, Panu Halme. (2012). Polypore communities in broadleaved boreal forests. Silva Fennica vol. 46 no. 3 article id 43.
The cover and extent of boreal broadleaved forests have been decreasing due to modern forest management practices and fire suppression. As decomposers of woody material, polypores are ecologically important ecosystem engineers. The ecology and conservation biology of polypores have been studied intensively in boreal coniferous forests. However, only a few studies have focused on the species living on broadleaved trees. To increase knowledge on this species group we conducted polypore surveys in 27 broadleaved forests and 303 forest compartments (539 ha) on the southern boreal zone in Finland and measured dead wood and forest characteristics. We detected altogether 98 polypore species, of which 13 are red-listed in Finland. 60% of the recorded species are primarily associated with broadleaved trees. The number of species in a local community present in a broadleaved forest covered approximately 50 species, of which 30–40 were primarily associated with broadleaved trees. The size of the inventoried area explained 67% of the variation in the species richness, but unlike in previous studies conducted in coniferous forests, dead wood variables as well as forest structure had very limited power in explaining polypore species richness on forest stand level. The compartments occupied by red listed Protomerulius caryae had an especially high volume of living birch, but otherwise the occurrences of red-listed species could not be predicted based on the forest structure.
  • Markkanen, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland ORCID ID:E-mail: (email)
  • Halme, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland ORCID ID:E-mail:
article id 114, category Research article
Roy V. Rea. (2011). Impacts of moose (Alces alces) browsing on paper birch (Betula papyrifera) morphology and potential timber quality. Silva Fennica vol. 45 no. 2 article id 114.
Although moose browsing effects on the growth and morphology of birch are well studied, effects of moose browsing on potential timber quality of birch have received little attention. Here, an assessment was made of the impacts of moose (Alces alces L.) damage to Paper Birch (Betula papyrifera Marsh.) trees from a 20-year old clear cut area in a sub-boreal spruce forest within the Aleza Lake Research Forest, near Prince George, British Columbia, Canada. Specifically, differences in overall tree architecture and in the internal characteristics of trees that had been severely damaged and suppressed by moose winter browsing were compared to birch trees that had not been damaged by moose in this way and were considered free-to-grow. The average stem diameter, number of annular growth rings, and height of stem breaks made by moose on suppressed birches at the point of breakage was 17.9 ± 6.6 mm, 4.6 ± 1.2, and 141.8 ± 32.0 cm, respectively. Stem diameters and the heights above-the-ground of stem breaks made by moose during sequential breakage events were not significantly different (all p ≥ 0.05) from one another. Decay was significantly (all p ≤ 0.001) more extensive in trees where branches had been broken off by moose than in trees with no breaks or where breaks were from unknown agents. Suppressed birches were significantly (p = 0.048) more exposed (farther from their nearest tree neighbor) when compared to birches that were free-to-grow. The distance from birch trees to species-specific neighbors (of any species) did not differ (all p ≥ 0.05) between suppressed and free-to-grow birches. Suppressed birches damaged from intense browsing and stem breakage were significantly (≤ 0.001) farther away from other birches showing signs of slight to moderate browsing than free-to-grow birches were from similar conspecifics. Because moose appear to impact the potential wood quality of birch, forest managers should consider the impacts that browsing and stem breakage can have on birch timber where these trees co-occur with and are eaten by moose.
  • Rea, University of Northern British Columbia, 3333 University Way, Prince George, BC, Canada V2N 4Z9 ORCID ID:E-mail: (email)
article id 173, category Research article
Erik Hellberg, Torbjörn Josefsson, Lars Östlund. (2009). The transformation of a Norway spruce dominated landscape since pre-industrial times in northern Sweden: the influence of modern forest management on forest structure. Silva Fennica vol. 43 no. 5 article id 173.
Logging history and the study of reference conditions in Scandinavian boreal forests has tended to focus on Scots pine dominated ecosystems. This paper presents a regional study of pre-industrial forest conditions and examines the effects of the industrial exploitation of ecosystems dominated by Norway spruce in northern Sweden. Historical records covering a period which preceded industrial logging in the study area (1917–1927) were used to obtain quantitative data on forest structure and influence of forest fires. These data were compared with a modern data set (2003) to analyse changes due to the industrial transformation of the forest. The early 20th century landscape was dominated by old, multi-cohorted spruce forests and mixed coniferous forests. It was found that fire affected both the structure and composition of the landscape. In post-burnt areas, even-aged forests dominated by deciduous species were the principal forest type. Between the early and modern data sets, profound changes in tree-species composition and age structure were documented. While the total volume of deciduous species increased substantially, the coverage of forests dominated by deciduous species decreased. There was also a significant increase in pine-dominated forests and in the total volume of pine. The industrial transformation of the studied landscapes has had profound effects on the structure of spruce forests, but much less so on deciduous forests. The study concludes that the present forest structure is a function of past management regimes, and that future transformations of the landscape will continue, thus affecting the natural variability and biodiversity of the forests.
  • Hellberg, Tunstigen 10, SE-831 43 Östersund, Sweden ORCID ID:E-mail:
  • Josefsson, Swedish University of Agricultural Sciences, Department of Forest Ecology and Management, SE-901 83 Umeå, Sweden ORCID ID:E-mail: (email)
  • Östlund, Swedish University of Agricultural Sciences, Department of Forest Ecology and Management, SE-901 83 Umeå, Sweden ORCID ID:E-mail:
article id 575, category Research article
Sylvie Mussche, Roeland Samson, Lieven Nachtergale, An De Schrijver, Raoul Lemeur, Noël Lust. (2001). A comparison of optical and direct methods for monitoring the seasonal dynamics of leaf area index in deciduous forests. Silva Fennica vol. 35 no. 4 article id 575.
During the 1996 growing season the seasonal dynamics of the Leaf Area Index (LAI) were determined by 3 different methods in two forest types: a mixed oak (Quercus robur L.) – beech (Fagus sylvatica L.) stand and an ash dominated (Fraxinus excelsior L.) stand. The results obtained from the two indirect methods, i.e. hemispherical photography and LAI-2000 Plant Canopy Analyser (Li-COR), were compared with the results of the direct measurement of litter fall collected in litter trap systems. In this study the direct method is considered to be the reference, giving the most accurate LAI-values. Both the hemispherical photography and the LAI-2000 PCA introduced an underestimation of LAI when the actual canopy leaf distribution in the crown layer deviates from a random distribution of leaf area in space as is found in the mixed oak/beech stand. However, when the condition of random leaf distribution is nearly fulfilled (ash stand), the LAI-2000 PCA gave LAI-values which were close to the results obtained from the direct method. Regression curves with R2 > 0.93 could be calculated for both indirect methods.
  • Mussche, Laboratory of Forestry, Ghent University, Geraardsbergse Steenweg 267, B-9090 Melle, Belgium ORCID ID:E-mail:
  • Samson, Laboratory of Plant Ecology, Ghent University, Coupure Links 653, B-9000 Gent, Belgium ORCID ID:E-mail:
  • Nachtergale, Laboratory of Forestry, Ghent University, Geraardsbergse Steenweg 267, B-9090 Melle, Belgium ORCID ID:E-mail:
  • De Schrijver, Laboratory of Forestry, Ghent University, Geraardsbergse Steenweg 267, B-9090 Melle, Belgium ORCID ID:E-mail: (email)
  • Lemeur, Laboratory of Plant Ecology, Ghent University, Coupure Links 653, B-9000 Gent, Belgium ORCID ID:E-mail:
  • Lust, Laboratory of Forestry, Ghent University, Geraardsbergse Steenweg 267, B-9090 Melle, Belgium ORCID ID:E-mail:

Category: Review article

article id 535, category Review article
Thomas J. Givnish. (2002). Adaptive significance of evergreen vs. deciduous leaves: solving the triple paradox. Silva Fennica vol. 36 no. 3 article id 535.
Patterns in the dominance of evergreen vs. deciduous plants have long interested ecologists, biogeographers, and global modellers. But previous models to account for these patterns have significant weaknesses. Bottom-up, mechanistic models – based on physiology, competition, and natural selection – have often been non-quantitative or restricted to a small range of habitats, and almost all have ignored belowground costs and whole-plant integration. Top-down, ecosystem-based models have succeeded in quantitatively reproducing several patterns, but rely partly on empirically derived constants and thresholds that lack a mechanistic explanation. It is generally recognized that seasonal drought can favor deciduous leaves, and that infertile soils can favor long-lived evergreen leaves. But no model has yet explained three great paradoxes, involving dominance by 1) evergreens in highly seasonal, boreal forests, 2) deciduous larch in many nutrient-poor peatlands, and 3) evergreen leaf-exchangers in nutrient-poor subtropical forests, even though they shed their leaves just as frequently as deciduous species. This paper outlines a generalized optimality model to account for these and other patterns in leaf longevity and phenology, based on maximizing whole-plant carbon gain or height growth, and building on recent advances in our understanding of the quantitative relationships of leaf photosynthesis, nitrogen content, and mass per unit area to leaf life-span. Only a whole-plant approach can explain evergreen dominance under realistic ecological conditions, or account for the boreal paradox, the larch paradox, the leaf-exchanger paradox, and expected shifts in shade tolerance associated with leaf phenology. Poor soils favor evergreens not merely by increasing the costs of nutrient acquisition, but also by depressing the maximum rate of photosynthesis and thus the seasonal contrast in photosynthetic return between leaves adapted to favorable vs. unfavorable conditions. The dominance of evergreens in western North America beyond the coastal zone of mild winters and winter rainfall appears related to the unusually long photosynthetic season for evergreen vs. deciduous plants there. Future models for optimal leaf phenology must incorporate differences between evergreen and deciduous plants in allocation to photosynthetic vs. non-photosynthetic tissue, rooting depth, stem allometry, xylem anatomy, and exposure to herbivores and leaching, and analyze how these differences interact with the photosynthetic rate, transpiration, and nutrient demands of leaves with different life-spans to affect rates of height growth in specific microsites.
  • Givnish, Dept of Botany, University of Wisconsin, Madison, WI 53706, USA ORCID ID:E-mail: (email)

Category: Article

article id 7114, category Article
Paavo Yli-Vakkuri. (1960). Metsiköiden routa- ja lumisuhteista. Acta Forestalia Fennica vol. 71 no. 5 article id 7114.
English title: Snow cover and ground frost in Finnish forests.

Snow cover and ground frost was studied in 29 forest stands in Southern and Central Finland in 1957–1959. The tree species influenced greatly accumulation of snow on the forest floor. Norway spruce (Picea abies (L.) Karst.) retains snow in its crown. In addition, snow and water falling from the branches compress the snow cover under the trees, and the ground freezes deeper because of the shallow snow cover. In the spring, the dense crown prevents rain and radiation reaching the ground, which remains cold longer. However, ground frost may protect spruce, which has a weak root system, from wind damages.

Scots pine (Pinus sylvestris L.) has similar, but milder, effects on snow cover within the forest. The crowns of pine seedlings and young trees pass snow easily, but later the crowns intercept it considerably. The lower branches are, however, high up and the snow is evenly spread on the ground. The deciduous trees intercept little snow and in the spring the snow smelts and the frozen soil thaws early. The snow conditions of deciduous forests are, however, changed by a spruce undergrowth.

It can be assumed that the unfavourable conditions in spruce forests can be alleviated by thinning. Also, mixture of pine and deciduous trees can transform the conditions more favourable in the spruce stands.

The PDF includes a summary in English.

  • Yli-Vakkuri, ORCID ID:E-mail:
article id 7261, category Article
K. Linkola. (1930). Über die Halbhainwälder in Eesti. Acta Forestalia Fennica vol. 36 no. 3 article id 7261.
English title: On half-deciduous woodlands in Estonia.

Half-deciduous woodlands are the combination of deciduous forests and heath forests from their vegetation. The article describes three half-deciduous forest types from the commune of Hager, 40 kilometers south form Tallinn.

The occurrence of half-deciduous forests depends on the soil conditions, especially on lime content of the soil, when the occurrence of other forest types is more a question of climate. They are important as an original site for many continental plant species.   

The PDF contains a summary in Finnish. 

  • Linkola, ORCID ID:E-mail:
article id 5600, category Article
Anton K. Chtchoukine. (1996). North European platyphyllous forests: biodiversity dynamics and climate changes in northwest European Russia. Silva Fennica vol. 30 no. 2–3 article id 5600.

Distribution, biodiversity and reforestation dynamics of the platyphyllous forests in the Northwest European Russia were investigated. Data assembled from 21 landscape regions (250–350 km2 each) show special features of small-leaved lime (Tilia cordata Mill., Norway maple (Acer platanoides L.), mountain elm (Ulmus glabra Mill.) and English oak (Qurecus robur L.) reforestation during the last two decades. New tendencies were found for the taiga areas with natural Norway spruce (Picea abies (L.) H. Karst.) and Scots pine (Pinus sylvestris L.) vegetation. Natural platyphyllous reforestation in cut spruce areas poses as supposed a special question for forest management policy in the relationship to global climate changes. Feasible unsustainability of the common types of succession (Norway spruce - European birch (Betula pendula Roth); Norway spruce - European aspen (Populus tremula L.)) is discussed. Biodiversity of herbs, shrubs and tree species of platyphyllous forests is high and complex and is situated in 4–15 old-growth relics in each landscape region. Low-level genotype heterogeneity of nemoral flora species of such isolated populations is presumed. Special biodiversity conservation regulations are proposed.

  • Chtchoukine, ORCID ID:E-mail:
article id 5506, category Article
Risto Heikkilä, Sauli Härkönen. (1993). Moose (Alces alces L.) browsing in young Scots pine stands in relation to the characteristics of their winter habitats. Silva Fennica vol. 27 no. 2 article id 5506.

Moose (Alces alces L.) browsing was studied in young Scots pine (Pinus sylvestris L.) stands mixed with deciduous trees in high-density winter ranges. The proportional use of twig biomass decreased as the availability increased. The total as well as proportional biomass consumption were higher on the moist than on the dry type of forest. The per tree consumption of pine was higher on the moist type, where the availability of pine was lower. Deciduous trees were more consumed on the moist type, where their availability was relatively high. The consumption of pine saplings increased as the availability of birch increased. Pine stem breakages were most numerous when birch occurred as overgrowth above pine and at high birch densities. The availability of other deciduous tree species did not correlate with browsing intensity of Scots pine. Moose browsing had seriously inhibited the development of Scots pines in 6% of the stands, over 60% of available biomass having been removed. Rowan and aspen were commonly over-browsed and their height growth was inhibited, which occurred rarely by birch. There was no difference in the proportion of young stands in forest areas with high and low moose density. A high proportion of peatland forests was found to indicate relatively good feeding habitats in the high-density areas.

The PDF includes an abstract in Finnish.

  • Heikkilä, ORCID ID:E-mail:
  • Härkönen, ORCID ID:E-mail:
article id 4627, category Article
Paavo Jaakko Ollinmaa. (1952). Jalot lehtipuumme luontaisina ja viljeltyinä. Silva Fennica no. 77 article id 4627.
English title: Native and cultivated southern broadleaved tree species in Finland.

The aim of the study was to update knowledge of natural range of English oak (Quercus robur L.), European ash (Fraxinus exelsior L.), Norway maple (Acer platanoides L.), small-leaved lime (Tilia cordata Miller), wych elm (Ulmus glabra Mill.) and European white elm (Ulmus laevis Pall.) in Finland, and estimate how far north they could be grown as forest trees or as park trees. The study is based on literature and questionnaires sent to cities and towns, District Forestry Boards, districts of Forest Service, Forestry Management Associations and railway stations.

The northern borders in the natural range of the species succeed one another from south to north as follows: English oak, European ash, Norway maple, wych elm, and small-leaved lime. Occurrence of European white elm is sporadic. The English oak forms forests in the southernmost Finland, while the other species grow only as small stands, groups or solitary trees. According to experiences of planted stands or trees, the northern limits of the species succeed one another from south to north as follows: European ash, English oak, Norway maple, European white elm, wych elm and small-leaved lime. All the species are grown in parks fairly generally up to the district of Kuopio-Vaasa (63 °). The northern limits where the species can be grown as park trees reach considerably further north in the western part of the country than in the east.

The article includes a summary in English.

  • Ollinmaa, ORCID ID:E-mail:

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