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Silva Fennica 1926-1997
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Acta Forestalia Fennica
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Articles containing the keyword 'mixed forest'.

Category: Research article

article id 970, category Research article
Zhen-Yu Du, Qing-Hua Wang, Shang-Jun Xing, Fang-Chun Liu, Bing-Yao Ma, Hai-Lin Ma, De-Xi Liu. (2013). Fine root distribution, characteristics and rhizosphere soil properties in a mixed stand of Robinia pseudoacacia and Fraxinus velutina in a saline soil. Silva Fennica vol. 47 no. 3 article id 970. https://doi.org/10.14214/sf.970
The spatial distribution and characteristics of fine roots (< 2 mm in diameter), and rhizosphere soil properties were studied in a mixed planted forest of black locust (Robinia pseudoacacia L.) and velvet ash (Fraxinus velutina Torr.) 27 years after planting in a coastal saline soil of the Yellow River delta, China. The results of fine root analysis showed that the fine roots of both black locust and velvet ash were mainly distributed in the soil layer at 0–20 cm depth and 50–150 cm from trees. The fine root distribution of both species suggests a strategy of avoiding salinity rather than salt –tolerance. The horizontal spread distance of fine roots of velvet ash was evidently longer than that of black locust. The fine root biomass, specific root length, specific root area, specific root volume and root activity were significantly higher for velvet ash in comparison with black locust. The results of soil analysis showed that rhizosphere soil pH of black locust and velvet ash were significantly lower compared with non-rhizosphere soil. The available N content in rhizosphere soil of black locust was higher than that of velvet ash. However, the contents of soluble salt, organic matter, available P and available K in rhizosphere soil of velvet ash were higher than those of black locust. The above results indicated that the differences between black locust and velvet ash in fine root distribution, characteristics and rhizosphere soil properties were the major reasons for that velvet ash showed stronger acclimation responses than black locust to the coastal saline soil.
  • Du, Shandong Academy of Forestry, 42 Wenhua East Road, Jinan 250014, P. R. China ORCID ID:E-mail: zydu@qq.com (email)
  • Wang, Shandong Academy of Forestry, Jinan, P. R. China ORCID ID:E-mail: wqh0228@163.com
  • Xing, Shandong Academy of Forestry, Jinan, P. R. China ORCID ID:E-mail: xingsj-126@126.com
  • Liu, Shandong Academy of Forestry, Jinan, P. R. China ORCID ID:E-mail: fchliu@126.com
  • Ma, Shandong Academy of Forestry, Jinan, P. R. China ORCID ID:E-mail: mby777@163.com
  • Ma, Shandong Academy of Forestry, Jinan, P. R. China ORCID ID:E-mail: mahlin@163.com
  • Liu, Shandong Academy of Forestry, Jinan, P. R. China ORCID ID:E-mail: llyldx@163.com
article id 351, category Research article
Jiaojun Zhu, Xiufen Li, Zugen Liu, Wei Cao, Yutaka Gonda, Takeshi Matsuzaki. (2006). Factors affecting the snow and wind induced damage of a montane secondary forest in northeastern China. Silva Fennica vol. 40 no. 1 article id 351. https://doi.org/10.14214/sf.351
In order to understand the processes of snow and wind induced damage in a natural montane, secondary forest in northeastern China, we examined the impacts of site conditions on the snow and wind damage; analyzed if the dominant tree species differed in their susceptibilities to the damage; and established the relationships between the characteristics of tree and stand and the damage. The results indicated that in regard to the topography factors, slope steepness and soil depth played a relatively important role for the damage. Damage ratios of all types combined were positively related with the composition of dominant tree species. The stand density was also important in determining resistance to the damage, i.e., the densely populated stand exhibited less overall damage ratios; however, the dominant tree species were commonly damaged easily by the snow and wind. Four damage modes found (uprooting, stem breakage, canopy damage and bending) were closely related to the stem taper (p < 0.05), and they could be ranked in following order: bending (92.0 ) > uprooting (85.3) > stem breakage (80.1) > canopy damage (65.0). In regard to differences in tree species’ susceptibilities to the damage, Betula costata exhibited the most uprooting, bending and overall damage ratios; while Quercus mongolica showed the highest breakage (both stem breakage and canopy damage) ratio, and Fraxinus mandshurica exhibited the least damage ratio (overall). The major six tree species could also be divided into two groups according to the overall damage ratios, i.e., more susceptible ones (B. costata, Ulmus laciniata and Q. mongolica), and less susceptible ones (F. mandshurica, Acer mono and Juglans mandshurica) to the snow and wind damage.
  • Zhu, Institute of Applied Ecology, Chinese Academy of Sciences, Wenhua Road 72, Shenyang 110016, China ORCID ID:E-mail: zrms29@yahoo.com (email)
  • Li, Institute of Applied Ecology, Chinese Academy of Sciences, Wenhua Road 72, Shenyang 110016, China; Graduate School of Chinese Academy of Sciences, Yuquan Road 19-A, Beijing, 100039, China ORCID ID:E-mail:
  • Liu, Institute of Applied Ecology, Chinese Academy of Sciences, Wenhua Road 72, Shenyang 110016, China; Graduate School of Chinese Academy of Sciences, Yuquan Road 19-A, Beijing, 100039, China ORCID ID:E-mail:
  • Cao, Institute of Applied Ecology, Chinese Academy of Sciences, Wenhua Road 72, Shenyang 110016, China ORCID ID:E-mail:
  • Gonda, Faculty of Agriculture, Niigata University, Ikarashi 2-8050, Niigata, 950-2181, Japan ORCID ID:E-mail:
  • Matsuzaki, Faculty of Agriculture, Niigata University, Ikarashi 2-8050, Niigata, 950-2181, Japan ORCID ID:E-mail:
article id 519, category Research article
Magnus Lindén, Gudmund Vollbrecht. (2002). Sensitivity of Picea abies to butt rot in pure stands and in mixed stands with Pinus sylvestris in southern Sweden. Silva Fennica vol. 36 no. 4 article id 519. https://doi.org/10.14214/sf.519
Repeatedly sampled data from permanent experimental plots in southern Sweden were used to model butt rot development in Norway spruce growing in pure stands and in mixed stands with Scots pine. The data come from 29 sites with pure spruce, altogether 100 plots, and from 15 sites of mixed spruce and pine, altogether 22 plots. A logistic model provided the best fit to the data. The study material revealed that in mixed stands the proportion of spruce trees with butt rot is lower than in pure Norway spruce stands. The difference in the incidence of butt rot cannot be explained by silviculture or windthrow since both factors are accounted for in the study. The most significant effect on butt rot development in Norway spruce by an admixture of Scots pine, was found when the Scots pine admixture was 50%. In order to reduce the incidence of butt rot in Norway spruce, the study material indicate that there is little to be gained by increasing the Scots pine admixture to much more than 50%.
  • Lindén, Southern Swedish Forest Research Centre c/o Asa Experimental Forest, SLU, S-360 30 Lammhult, Sweden ORCID ID:E-mail: magnus.linden@ess.slu.se (email)
  • Vollbrecht, Southern Swedish Forest Research Centre c/o Asa Experimental Forest, SLU, S-360 30 Lammhult, Sweden ORCID ID:E-mail:
article id 671, category Research article
Jouni Vettenranta, Jari Miina. (1999). Optimizing thinnings and rotation of Scots pine and Norway spruce mixtures. Silva Fennica vol. 33 no. 1 article id 671. https://doi.org/10.14214/sf.671
The study describes a simulation-optimization system which uses spatial models for diameter and height growth, crown ratio and tree mortality for Scots pine and Norway spruce mixtures. The optimal one- and two-thinning regimes of six initial stands with varing species composition were solved by using nonlinear optimization. The soil expectation value (SEV) at 3% interest rate was used as a management objective. The regimes are determined by taking into account the stand basal areas before the thinnings, the removal percentages for small, medium-sized and large pines and spruces, and the stand basal area before the final felling. The greatest SEV (8900 FIM ha–1) was attained with the initial stand where the proportion of pines was 65% of the number of the stems. In the two-thinning regime, the first thinning was conducted at the age of 39 years when the stand basal area was 37 m2 ha–1 and the dominant height was about 15 m. After the thinning, the basal area was 27 m2 ha–1. Spruces were thinned from below, but both small and large pines were removed. The second thinning was 8 years later and much heavier: the stand basal area was decreased from 35 m2 ha–1 to 18 m2 ha–1 by removing both small and large pines and spruces. When the optimal two-thinning regime was compared to the regime presented by Forest Centre Tapio, the loss of SEV was about 30% (6070 FIM ha–1) in the case of thinnings from below, and about 20% (7250 FIM ha–1) in the case of thinnings from above.
  • Vettenranta, Kivirinnanpolku 4, FIN-40950 Muurame, Finland ORCID ID:E-mail: vettenr@cc.joensuu.fi (email)
  • Miina, Faculty of Forestry, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland ORCID ID:E-mail:

Category: Article

article id 7476, category Article
Leo Heikurainen. (1958). Sekametsiköiden juuristoista ojitetulla suolla. Acta Forestalia Fennica vol. 67 no. 2 article id 7476. https://doi.org/10.14214/aff.7476
English title: Root systems of mixed forest in drained peatlands.

Draining transforms root systems of trees growing in peatlands towards the ones growing on mineral soil. However, even after efficient draining the root systems differ from the root systems of trees growing on mineral soil. This investigation concentrates on root systems of forests of similar mire types growing in similar draining conditions but having different tree species compositions. The peatland, situated in Pieksämäki in Southern Finland, was drained in 1937. Sample plots, measured in 1956, consisted of mixed forest of Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies L. Karst.) and birch (Betula sp.) in different compositions, and were in natural condition.

The sedge pine bog studied in this investigation was shown to have larger total amount of roots and mycorrhiza than in previously studied dwarf shrub pine bogs. This reflects better growth conditions of the better site. The depth of root system was, however, similar. Root systems of birch were deeper than those of the coniferous tree species. Differences between Scots pine and Norway spruce were small. Corresponding differences between the species were found in the density and total number of mycorrhizas. The abundance of mycorrhizas in the roots of birch increased in deeper layers of peat, but decreased especially in spruce roots. In earlier studies the abundance of mycorrhizas decreased in the roots growing in deeper layers in pure Scots pine stands, but no such variation was seen in this study. The result suggest that the deep root system of birch may affect also the root systems of the coniferous trees. On the other hand, birch roots can have advantage over the coniferous trees.

The PDF includes a summary in German.

  • Heikurainen, ORCID ID:E-mail:
article id 7361, category Article
M. Lappi-Seppälä. (1942). Siperian lehtikuusen kasvusta sekametsiköissä Evon valtionpuistossa. Acta Forestalia Fennica vol. 50 no. 8 article id 7361. https://doi.org/10.14214/aff.7361
English title: Growth of Siberian larch in mixed stands in state forests of Evo in Finland.

Of the foreign tree species Siberian larch (Larix sibirica Ledeb.) has the biggest economical potential in Finland. In its natural distribution the species grows mostly in mixed stands in other areas than the core of its range in Siberia, where it grows also in pure stands. However, growth studies have given contradictory results about how Siberian larch can manage competition of different tree species in mixed stands. In this study two-year old Siberian larch seedlings were planted in areas previously sown with Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.). The growth of the trees was measured when the stands were 50 years old.

It appears that the stands, about 3700 larch seedlings per hectare, have originally been too been too dense. In the two thinnings done in the area, larch has probably been favoured, which has resulted in varying mix of pine and spruce. In the 50-year old stands, Siberian larch has developed faster than Scots pine and Norway spruce. Contrary to some previous studies, the results show that Siberian larch can be grown also in mixed stands, but the growth will probably be slower than in pure stands. Best growth is achieved in pure stands that have been planted thinly enough.

The PDF includes a summary in German.

  • Lappi-Seppälä, ORCID ID:E-mail:
article id 7221, category Article
Konrad Rubner. (1929). Die Ziele der mitteleuropäischen Forstwirtschaft. Acta Forestalia Fennica vol. 34 no. 8 article id 7221. https://doi.org/10.14214/aff.7221
English title: Objectives of middle-European forestry.

The article presents middle-European forests and their characteristics, development and use. The main question is whether the current practice of planted coniferous tree forests should be kept or whether more natural mixed forests should be introduced instead. The article discusses the pros and cons of these and the possible felling and management practices.      

 The volume 34 of Acta Forestalia Fennica is a jubileum publication of professor Aimo Kaarlo Cajander.

  • Rubner, ORCID ID:E-mail:
article id 5574, category Article
Jouni Vettenranta. (1996). Effect of species composition on economic return in a mixed stand of Norway spruce and Scots pine. Silva Fennica vol. 30 no. 1 article id 5574. https://doi.org/10.14214/sf.a9219

The effect of species mixture was studied in a mixed stand of Norway spruce (Picea abies (L.) H. Karst.) and Scots pine (Pinus sylvestris L.) by simulating around 100 different treatment schedules during the rotation in a naturally regenerated even-aged stand located on a site of medium fertility in North Karelia, Finland. Both thinning from below and thinning from above were applied. Optimum rotations were determined by maximising the net present value calculated to infinity and different treatment schedules were compared with the net present value over one rotation as per rotation applied. In the optimum treatment programme, the proportion of pines was decreased by half of the basal area in the first thinning stage and by the end of the rotation to about one third. In thinning from above, the proportion of pines can be maintained at a slightly higher level. It is economically profitable to maintain the growing stock capital at approximately the level recommended by Forest Centre Tapio, a semi-governmental forestry authority. With non-optimum species composition, the loss in net present value over one rotation can be about 10 % in thinning from below and about 20 % in thinning from above.

  • Vettenranta, ORCID ID:E-mail:
article id 5506, category Article
Risto Heikkilä, Sauli Härkönen. (1993). Moose (Alces alces L.) browsing in young Scots pine stands in relation to the characteristics of their winter habitats. Silva Fennica vol. 27 no. 2 article id 5506. https://doi.org/10.14214/sf.a15667

Moose (Alces alces L.) browsing was studied in young Scots pine (Pinus sylvestris L.) stands mixed with deciduous trees in high-density winter ranges. The proportional use of twig biomass decreased as the availability increased. The total as well as proportional biomass consumption were higher on the moist than on the dry type of forest. The per tree consumption of pine was higher on the moist type, where the availability of pine was lower. Deciduous trees were more consumed on the moist type, where their availability was relatively high. The consumption of pine saplings increased as the availability of birch increased. Pine stem breakages were most numerous when birch occurred as overgrowth above pine and at high birch densities. The availability of other deciduous tree species did not correlate with browsing intensity of Scots pine. Moose browsing had seriously inhibited the development of Scots pines in 6% of the stands, over 60% of available biomass having been removed. Rowan and aspen were commonly over-browsed and their height growth was inhibited, which occurred rarely by birch. There was no difference in the proportion of young stands in forest areas with high and low moose density. A high proportion of peatland forests was found to indicate relatively good feeding habitats in the high-density areas.

The PDF includes an abstract in Finnish.

  • Heikkilä, ORCID ID:E-mail:
  • Härkönen, ORCID ID:E-mail:
article id 7580, category Article
Seppo Kellomäki. (1975). Forest stand preferences of recreationists. Acta Forestalia Fennica no. 146 article id 7580. https://doi.org/10.14214/aff.7580

The environmental preferences of recreationists were studied at the forest stand level in this study. The hypothesis which has been drawn up on the basis of the literature has been studied by attempting to elucidate the environmental preferences of groups using two recreation areas owned by the City of Helsinki using interviews and questionnaires. The material consisted of 1,323 interviews supplemented by questionnaires.

The replies to the questionnaires showed that recreationists consider birch and Scots pine to be more beautiful than Norway spruce, and stands made up of several tree species to be more beautiful than stands of single tree species. They also consider mature stands to be more beautiful than young stands.

During the interviews, the attention of the recreationists was directed at the view formed by the interview stand. The scenic preferences for the stands were measured using adjectives which the interviewee was asked to use in describing his or her impression of the view which was pointed out. First of all, the results clearly indicated that from the point of view of the scenic value of the stand, the way in which the stand is organised to form a scenic aspect or a stand view is more important than its ecological structure. However, it is obvious that stands containing large sized trees in particular are in many ways more preferred than stands which are younger in their development stage. This should therefore be the case when changes in the stand view resulting from management measures are insignificant or difficult to see. The main tree species in the stand does not seem to have from the point of view of scenic preference as much significance as would have been expected judging by the questionnaire material.

The PDF includes a summary in Finnish.

  • Kellomäki, ORCID ID:E-mail:
article id 4569, category Article
Martti Tertti. (1939). Näkökohtia kuusimetsän hoidosta. Silva Fennica no. 52 article id 4569. https://doi.org/10.14214/sf.a13976
English title: Forest management of Norway spruce forests.

Silva Fennica issue 52 includes presentations held in professional development courses, arranged for foresters working in public administration in 1938. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.

This presentation describes different types of fellings in Norway spruce (Picea abies (L.) H. Karst.) forests in different forest site types. The use of thinning from below and above, clear cutting of Norway spruce stands, and thinning of mixed forests with birch (Betula sp.) are discussed.

  • Tertti, ORCID ID:E-mail:
article id 4537, category Article
M. Lappi-Seppälä. (1938). Sekametsien kasvatuksesta. Silva Fennica no. 46 article id 4537. https://doi.org/10.14214/sf.a13946
English title: Management of mixed forests.

Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service. 

This presentation describes management of mixed forests.

  • Lappi-Seppälä, ORCID ID:E-mail:

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