Current issue: 54(2)
Under compilation: 54(3)
Competitive interactions in clonal forestry are not well understood and this needs to be addressed to develop better deployment strategies. Eight juvenile Sitka spruce (Picea sitchensis (Bong.) Carriére) clones were grown in monoclonal and clonal mixtures in a field experiment for three years to assess the effects of genetic diversity on shoot growth, above- and below-ground biomass partitioning and crown characteristics. Shoot elongation was measured throughout the growing season, while diameter was measured twice annually in May and December. After the third year, crown silhouette area was estimated from digitised images for one ramet per plot and ramets were then destructively harvested. Deployment × clone interactions were observed for tree height and diameter with reductions observed in mixed plots. Mixed plots had significantly greater height and diameter heterogeneity and more asymmetrical competition than monoclonal plots. Results from this study demonstrate that stem growth can be significantly altered when clones are planted in multi-clonal mixtures but for most clones, deployment-type will not significantly reduce their productivity.
Farm forestry in northern Ostrobothnia has met different kinds of obstacles that decrease its profitability, some national, some local. One of the later is partitioning of land. The purpose of this investigation was to survey the division of farm land in the coastal municipalities of northern Ostrobothnia in Finland, where the conditions are among the most unfavourable in the country in this respect. The material used in the investigation was collected in a previous study about the structure of the farms in the area. First part of the paper summarises the history of partitioning of land in Finland.
The results show that division of the woodlots of a farm are in the coastal municipalities of northern Ostrobothnia very disadvantageous for forestry. The average distance of a woodlot to the farmhouse is 8.3 km, but there is a great variation between the municipalities, and the distance varies from 30 to 1.9 km. The form of the lots, as the long ribbon-like woodlots in the municipality of Liminka, complicates often practical forestry. In addition, the number of separate woodlots is high, in average 9.2 per farm.
The great distance of the woodlots from the main farms hinders the use of forests and diminishes the financial result of forestry. Unfavourable form of the woodlots posts similar hindrances to harvesting of timber and forest management as the long distances and high number of separate plots. The problem is heightened by the abundance of peatlands in relation to productive forest lands in the area.
The PDF includes a summary in German.
A shoot-root carbon:nitrogen allocation model, based on the two processes of transport and chemical conversion, is described and explored. The view is proposed that all allocation models, whether built for the purposes of theoretical investigation or practical application, should start with this irreducible framework. In the present implementation, the processes operate according to: for substrate sources, dependence on shoot and root sizes, with possible product inhibition; for transport, movement down a substrate concentration gradient; for substrate sinks or utilization, linear bisubstrate kinetics. The dynamic and equilibrium properties of the model are explored. Failure of this approach to allocation will indicate to the modeller that additional mechanisms to control the processes are needed, and the mode of failure will indicate the type of mechanisms required. Additional mechanisms are discussed which may involve hormones or teleonomic (goal-seeking) controls, and may be added to the irreducible framework. However, these additions should not replace the irreducible framework of transport and chemical conversion, because they do not in reality. Modifications to the basic model to reflect some possibilities such as ontogenesis with the transition from exponential growth towards a steady state or with the scaling of within-plant transport resistances, the influence of hormones, and active transport, are described.
Process-based tree growth models are recognized to be flexible tools which are valuable for investigating tree growth in relation to changing environment or silvicultural treatments. In the context of forestry, we address two key modelling problems: allocation of growth which determines total wood production, and distribution of wood along the stem which determines stem form and wood quality. Growth allocation and distribution are the outcome of carbon translocation, which may be described by the Munch theory. We propose a simpler gradient process to describe the carbon distribution in the phloem of conifers. This model is a reformulation of a carbon diffusion-like process proposed by Thornley in 1972. By taking into account the continuity of the cambium along the stem, we obtain a one-dimensional reaction-diffusion model which describes both growth allocation between foliage, stem and roots, and growth distribution along the stem. Distribution of wood along the stem is then regarded as an allocation process at a smaller scale. A preliminary sensitivity analysis is presented. The model predicts a strong relationship between morphology and foliage-root allocation. It also suggests how empirical data, such as stem analysis, could be used to calibrate and validate allocation rules in process-based growth models.
Change in dry matter partitioning, 14C-incorporation, and sink 14C-activity of 1.5-year-old Scots pine (Pinus sylvestris L.) seedlings grown in growth chamber conditions were studied during a 91-day experiment. On five sampling dates, seedlings were labelled with 14CO2 and whole-plant allocation patterns were determined. Intensively growing shoots modified the dry matter partitioning: during shoot growth the proportion of roots decreased but after that it increased. Based on their large proportion of dry matter, the needles (excluding current needles) were the strongest sink of carbon containing 40% of the incorporated 14C. Despite their small initial sink size, the elongating shoots (current main shoot + current branch) and their needles were the second strongest sink (30–40% of the total 14C) which reflects their high physiological activity. The proportion of 14C in the current year’s main shoot increased during shoot growth but decreased as the growth began to decline after 70 days. 10–20% of the total assimilated 14C was translocated to the roots. Laterals above 2nd order were the strongest sink in the root system, containing twice as much 14C as the other roots together. Alternation between shoot and root growth can be seen clearly: carbon allocation to roots was relatively high before and after the period of intensive shoot growth. Changes in root sink strength resulted primarily from changes in root sink activity rather than sink size.
Questions of the small size of non-industrial private forest (NIPF) holdings in Finland are considered and factors affecting their partitioning are analysed. This work arises out of Finnish forest policy statements in which the small average size of holdings has been seen to have negative influence on the economics of forestry. A literature survey indicates that the size of holdings is an important factor determining the costs of logging and silvicultural operations, while its influence on the timber supply is slight.
The empirical data are based on a sample of 314 holdings collected by interviewing forest owners in 1980–86. In 1990–91 the same holdings were resurveyed by means of a postal inquiry and partly by interviewing the forest owners. The principal objective was to collect data to assist in quantifying ownership factors that influence partitioning among different kinds of NIPF holdings. Thus, the mechanism of partitioning was described and a maximum likelihood logistic regression model was constructed using seven independent holding and ownership variables.
One out of four holdings had undergone partitioning in conjunction with a change in ownership, one fifth among family owned holdings and nearly a half among jointly owned holdings. The results of the logistic regression model indicate, for instance, that the odds on partitioning is about three times greater for jointly owned holdings than for family owned ones. Also, the probabilities of partitioning were estimated and the impact of independent dichotomous variables on the probability of partitioning ranged between 0.02 and 0.01. The low value of Hosmer-Lemeshow test statistics indicates a good fit of the model, and the rate of correct classification was estimated to be 88% with a cut-off point of 0.5.
The average size of holdings undergoing ownership changes decreased from 29.9 ha to 28.7 ha over the approximate interval 1983–90. In addition, the transition probability matrix showed that the trends towards smaller size categories mostly concerned the small size categories. The results can be used in considering the effects of the small size holdings for forestry and if the purpose is to influence partitioning through forest or rural policy.
A theoretical framework to analyse the growth of Scots pine (Pinus sylvestris L.) is presented. Material exchange processes and internal processes that transport, transform and consume materials are identified as the components of growth. Hierarchical system is lined out. Momentary uptake of material at a single exchange site depends on the environmental condition next to the exchange site, the internal state of the biochemical system of the plant and the structure of the plant. The internal state depends on the exchange flows over period of time and the structural growth depends on the internal state. The response of these processes to the fluxes is controlled by the genetic composition of the plant.
The theoretical framework is formulated into a mathematical model. A concept of balanced internal state was applied to describe the poorly known internal processes. Internal substrate concentrations were assumed to remain constant but tissue-specific. A linear relationship between the quantity of foliage and wood cross-sectional area was assumed to describe balanced formation of structure. The exchange processes were thus described as a function of external conditions. The stand level interactions were derived from shading and effects of root density on nutrient uptake.
The approach was tested at different levels of hierarchy. Field measurements indicated that the hypothesis of the linear relationship described well the regularities between foliage and sapwood of a tree within a stand when measured at functionally corresponding height. There was considerable variation in the observed regularities in the range of geographic occurrence of Scots pine. Model simulations gave a realistic description of stand development in Southern Finland. The same model was also able to describe growth differences in Lapland after considering the effect of growing season length in the parameter values. Simulations to South Russia indicate stronger deviation from the observed patterns.
The simulations suggest interesting features of stand development. They indicate strong variability in the distribution of carbohydrates between tree parts during stand development. Internal circulation of nutrients and the reuse of the same transport structure by various needle generations had a strong influence on the simulation results.
The PDF includes a summary in Finnish.