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Articles containing the keyword 'regeneration'.

Category: Research article

article id 10036, category Research article
Karin Hjelm, Lars Rytter. (2018). The demand of hybrid aspen (Populus tremula × P. tremuloides) on site conditions for a successful establishment on forest land. Silva Fennica vol. 52 no. 5 article id 10036. https://doi.org/10.14214/sf.10036
Highlights: Low pH (below 3.5) reduced growth, but not survival, in a greenhouse study; Site preparation methods did not affect survival in field, but differences were found for growth; Mounding had generally the best effect on growth; Clonal differences were found that could be useful for improving establishment and growth.

Hybrid aspen (Populus tremula L. × P. tremuloides Michx.) is a deciduous tree species suitable for producing large amounts of renewable biomass during short rotations. Its potential under North European conditions could be largely extended if not only agricultural land but also forest land was used for cultivation. Unfortunately, the knowledge of appropriate forest site conditions and effects of site preparation methods on hybrid aspen establishment is limited. In this paper, two studies that explore these questions are presented. In the first study, the sensitivity to acid soils was tested under greenhouse conditions in two type of soils: a) peat soil limed to certain pH levels (3.4–5.7) and b) collected forest soils where pH varied from 3.9 to 5.3. The lowest pH level resulted in reduced growth, elsewhere no significant differences were found. The second study was applied in the field and investigated the effect of four site preparation methods on survival and growth. The methods were: 1) control with no site preparation, 2) patch scarification, 3) mounding and 4) soil inversion. While no differences were found for survival, mounding was generally the method with the highest growth and patch scarification was least successful. The result was probably an effect of good soil aeration and less competition from vegetation after mounding. The field study also revealed clonal differences in growth performance, which stresses the importance of clone selection prior to planting. The results of these studies indicate that hybrid aspen is less sensitive to variation in pH and site preparation methods compared with other poplar species, as have been found in similar studies.

  • Hjelm, Skogforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden ORCID ID:E-mail: karin.hjelm@skogforsk.se (email)
  • Rytter, Skogforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden ORCID ID: http://orcid.org/0000-0001-6183-4832 E-mail: lars.rytter@skogforsk.se
article id 9946, category Research article
Frauke Fedderwitz, Niklas Björklund, Velemir Ninkovic, Göran Nordlander. (2018). Does the pine weevil (Hylobius abietis) prefer conifer seedlings over other main food sources? Silva Fennica vol. 52 no. 3 article id 9946. https://doi.org/10.14214/sf.9946
Highlights: Adult pine weevils feed on seedlings and mature conifers, but cause economic damage only on seedlings; Their feeding preferences for branches and roots over seedlings were tested in a laboratory experiment; The only clear preference was for Norway spruce roots; Results support new approaches of seedling protection attempting to redirect pine weevils from planted seedlings to other food sources.

Adult pine weevils (Hylobius abietis (L.)) feed on the tender bark of branches and roots of mature conifer trees and on the stem bark of conifer seedlings. Their feeding on mature trees does not cause any economic damage, but their feeding on planted seedlings is so devastating that the pine weevil is considered one of the most important forest pest insects in Europe. We asked whether the pine weevil prefers seedlings over other regularly utilized food sources. This question is of particular interest because new approaches to seedling protection are based on decreasing any preference for seedlings by using less palatable plants or by enhancing their defence (by genetic selection or by methyl jasmonate treatment). In a laboratory choice experiment we tested pine weevil feeding preferences for seedlings compared with branches and roots from mature trees (separately for Norway spruce and Scots pine). Pine weevils preferred roots, but not branches, of Norway spruce over seedlings of the same species. With Scots pine there were no clear preferences, but the weevils showed a tendency to prefer roots over seedlings. These results provide support for seedling protection approaches that attempt to redirect pine feeding from planted seedlings to other food sources.

  • Fedderwitz, Department of Ecology, Swedish University of Agricultural Sciences, Box 7044, SE-750 07 Uppsala, Sweden ORCID ID:E-mail: frauke.fedderwitz@slu.se (email)
  • Björklund, Department of Ecology, Swedish University of Agricultural Sciences, Box 7044, SE-750 07 Uppsala, Sweden ORCID ID:E-mail: niklas.bjorklund@slu.se
  • Ninkovic, Department of Ecology, Swedish University of Agricultural Sciences, Box 7044, SE-750 07 Uppsala, Sweden ORCID ID:E-mail: velemir.ninkovic@slu.se
  • Nordlander, Department of Ecology, Swedish University of Agricultural Sciences, Box 7044, SE-750 07 Uppsala, Sweden ORCID ID:E-mail: goran.nordlander@slu.se
article id 9933, category Research article
Timo Saksa, Jari Miina, Hilkka Haatainen, Kauko Kärkkäinen. (2018). Quality of spot mounding performed by continuously advancing mounders. Silva Fennica vol. 52 no. 2 article id 9933. https://doi.org/10.14214/sf.9933
Highlights: The number and quality of mounds varied considerably according to the operating conditions;The main factors reducing the quality of spot mounding were steep terrain, a thick humus layer, fresh logging residues, stoniness and soil texture;With careful selection of timing and conditions for mounding, the quality obtained by continuously advancing mounders can be improved.

Operating conditions affecting the quality of spot mounding by Bracke continuously advancing mounders were investigated on 66 regeneration areas (124 ha) in eastern Finland. The quality of mounds was classified as suitable (good or acceptable after additional compression) or unsuitable for planting. Models were constructed for the number of suitable planting spots obtained per hectare (good and acceptable mounds), the probability of successful mounding (≥1600 planting spots ha–1) and the probability of creating a suitable mound as a function of terrain, site and soil characteristics, as well as slash conditions (removed, fresh or dry logging residues). The average number of mounds created was 1892 ± 290 mounds ha–1, of which 1398 ± 325 mounds ha–1 (74%) were classified as suitable for planting. The quality of spot mounding was reduced by steep terrain, a thick humus layer and fresh logging residues. Stoniness and soil texture also affected the number of planting spots created. Mounding after logging residues had dried increased the number of planting spots by 191 spots ha–1 compared with mounding in the presence of fresh residues. Removing residues did not significantly increase the number of planting spots compared with mounding amongst dry residues. A thick humus layer, very stony soil, steep slopes and valley terrain decreased the number of planting spots by 150–450 spots ha–1. The number and quality of mounds varied considerably according to the operating conditions, but with careful selection of timing and sites the quality obtained by a continuously advancing mounder can be improved.

  • Saksa, Natural Resources Institute Finland (Luke), Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: timo.saksa@luke.fi (email)
  • Miina, Natural Resources Institute Finland (Luke), Yliopistokatu 6, FI-80100 Joensuu, Finland ORCID ID:E-mail: jari.miina@luke.fi
  • Haatainen, Faculty of Science and Forestry, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: hilkka.haatainen@storaenso.com
  • Kärkkäinen, Tornator Oyj, Muuntamontie 2, FI-80100 Joensuu, Finland ORCID ID:E-mail: kauko.karkkainen@tornator.fi
article id 1665, category Research article
Lauri Haataja, Ville Kankaanhuhta, Timo Saksa. (2018). Reliability of self-control method in the management of non-industrial private forests. Silva Fennica vol. 52 no. 1 article id 1665. https://doi.org/10.14214/sf.1665
Highlights: Self-control method was found reliable at the main stages of the forest regeneration process; Only slight overestimation was found in self-control results of soil preparation and planting and small underestimation in self-control of young stand management; Diverse utilizing of self-control data is possible in support of service providers operations.
This study seeks to determine the extent to which self-control data can be relied upon in the management of private forests. Self-control (SC) requires the forest workers to evaluate their own work quality to ensure the clients’ needs are met in terms of soil preparation, planting and young stand management. Self-control data were compared to an independent evaluation of the same worksites. Each dataset had a hierarchical structure (e.g., sample plot, regeneration area and contractor), and key quality indicators (i.e., number of prepared mounds, planted seedlings or crop trees) were measured for each plot.  Self-control and independent-assessments (IA) were analyzed by fitting a multi-level multivariate model containing explanatory variables. No significant differences were observed in terms of soil preparation (number of mounds) or young stand management (number of crop trees) between self-control and independent-assessments. However, the self-control planting data included a slight but significant overestimation of the number of planted seedlings. Discrepancies are discussed in terms of sampling error and other explanatory factors. According to overall results, self-control methods are reliable at every stage of the forest regeneration process. As such, the diverse utilizing of self-control data is possible in support of service providers operations.
  • Haataja, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: lauritapiohaataja@gmail.com (email)
  • Kankaanhuhta, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: ville.kankaanhuhta@luke.fi
  • Saksa, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: timo.saksa@luke.fi
article id 1564, category Research article
Stanislav Vacek, Zdeněk Vacek, Lukáš Bílek, Jaroslav Simon, Jiří Remeš, Iva Hůnová, Jan Král, Tereza Putalová, Miroslav Mikeska. (2016). Structure, regeneration and growth of Scots pine (Pinus sylvestris L.) stands with respect to changing climate and environmental pollution. Silva Fennica vol. 50 no. 4 article id 1564. https://doi.org/10.14214/sf.1564
Highlights: Pine forest stands showed positive development of stand structural characteristics related to their diversity, number of regeneration individuals and growth characteristics; Tree-ring width was positively correlated with precipitation, while it was negatively correlated with temperature in growing seasons; Mean NOx concentrations showed positive effect on radial growth of pine; Serious defoliation was caused by SO2 concentrations and N deposition in combination with extreme climate events.

Changes in the structure and development of managed Scots pine (Pinus sylvestris L.) stands with respect to changing environmental conditions were set for the period 1979–2015. The study was conducted in conditions of natural pinewoods and pine-oak sites on five permanent research plots (0.25 ha) in Eastern Bohemia, Czech Republic (CR). Studied forest stands showed positive development of stand structural characteristics related to their diversity, number of regeneration individuals and growth characteristics. The standing volume of regularly distributed tree layer in 2015 was in the range of 320–434 m3 ha–1, which indicates an increase by 5.9–20.0% over 10 years. Correlation between pine radial increment and the amount of precipitation was generally the strongest one. Positive statistically significant correlation between diameter increment and temperature was demonstrated only for the average March temperature of the current year. Within the CR, study site can be characterised as a medium polluted area both for sulphur and nitrogen, despite this SO2 concentrations and N deposition in combination with extreme climate events caused severe defoliation in pine stands. Conversely, radial growth was positively significantly correlated with mean NOx concentrations. Drought mainly in combination with even medium environmental pollution can further worsen the health status of pine stands in lowland areas of Central Europe. Thus, formulation of silvicultural techniques able to mitigate the impact of these stress factors is needed.

  • Vacek, Czech University of Life Sciences Prague, Faculty of Forestry and Wood Sciences, Kamýcká 129, 165 21 Prague, Czech Republic ORCID ID:E-mail: vacekstanislav@fld.czu.cz
  • Vacek, Czech University of Life Sciences Prague, Faculty of Forestry and Wood Sciences, Kamýcká 129, 165 21 Prague, Czech Republic ORCID ID:E-mail: vacekz@fld.czu.cz
  • Bílek, Czech University of Life Sciences Prague, Faculty of Forestry and Wood Sciences, Kamýcká 129, 165 21 Prague, Czech Republic ORCID ID:E-mail: bilek@fld.czu.cz
  • Simon, Mendel University in Brno, Faculty of Forestry and Wood Technology, Zemědělská 3, 613 00 Brno, Czech Republic ORCID ID:E-mail: jaroslav.simon@mendelu.cz
  • Remeš, Czech University of Life Sciences Prague, Faculty of Forestry and Wood Sciences, Kamýcká 129, 165 21 Prague, Czech Republic ORCID ID:E-mail: remesj@email.cz (email)
  • Hůnová, Czech Hydrometeorological Institute, Na Šabatce 17 143 06 Prague, Czech Republic ORCID ID:E-mail: hunova@chmi.cz
  • Král, Czech University of Life Sciences Prague, Faculty of Forestry and Wood Sciences, Kamýcká 129, 165 21 Prague, Czech Republic ORCID ID:E-mail: kraljan@fld.czu.cz
  • Putalová, Czech University of Life Sciences Prague, Faculty of Forestry and Wood Sciences, Kamýcká 129, 165 21 Prague, Czech Republic ORCID ID:E-mail: putalova@fld.czu.cz
  • Mikeska, University of Hradec Králové, Faculty of Science, Rokitanského 62, 500 03 Hradec Králové, Czech Republic ORCID ID:E-mail: Mikeska.Miroslav@uhul.cz
article id 1562, category Research article
Mats Erik Berlin, Torgny Persson, Gunnar Jansson, Matti Haapanen, Seppo Ruotsalainen, Lars Bärring, Bengt Andersson Gull. (2016). Scots pine transfer effect models for growth and survival in Sweden and Finland. Silva Fennica vol. 50 no. 3 article id 1562. https://doi.org/10.14214/sf.1562
Highlights: Scots pine transfer effect models for growth and survival, valid in both Sweden and Finland have been developed; The models use high-resolution gridded climate data and can predict performance in future climatic conditions; The models perform well both for unimproved and genetically improved material and can be used to develop deployment recommendations of contemporary forest regeneration material in Sweden and Finland.

In this study, we developed models of transfer effects for growth and survival of Scots pine (Pinus sylvestris L.) in Sweden and Finland using a general linear mixed-model approach. For model development, we used 378 provenance and progeny trials with a total of 276 unimproved genetic entries (provenances and stand seed check-lots) distributed over a wide variety of climatic conditions in both countries. In addition, we used 119 progeny trials with 3921 selected genetic entries (open- and control pollinated plus-tree families) for testing model performance. As explanatory variables, both climatic indices derived from high-resolution gridded climate datasets and geographical variables were used. For transfer, latitude (photoperiod) and, for describing the site, temperature sum were found to be main drivers for both survival and growth. In addition, interaction terms (between transfer in latitude and site altitude for survival, and transfer in latitude and temperature sum for growth) entail changed reaction patterns of the models depending on climatic conditions of the growing site. The new models behave in a way that corresponds well to previous studies and recommendations for both countries. The model performance was tested using selected plus-trees from open and control pollinated progeny tests. Results imply that the models are valid for both countries and perform well also for genetically improved material. These models are the first step in developing common deployment recommendations for genetically improved forest regeneration material in both Sweden and Finland.

  • Berlin, Uppsala Science Park, SE-75183 Uppsala, Sweden ORCID ID:E-mail: mats.berlin@skogforsk.se (email)
  • Persson, Skogforsk, Box 3, SE-91821 Sävar, Sweden ORCID ID:E-mail: torgny.persson@skogforsk.se
  • Jansson, Uppsala Science Park, SE-75183 Uppsala, Sweden ORCID ID:E-mail: gunnar.jansson@skogforsk.se
  • Haapanen, Natural Resources Institute Finland (Luke), Green Technology, Box 18, FI-01301 Vantaa, Finland ORCID ID:E-mail: matti.haapanen@luke.fi
  • Ruotsalainen, Natural Resources Institute Finland (Luke), Green Technology, Finlandiantie 18, FI-58450 Punkaharju, Finland ORCID ID:E-mail: seppo.ruotsalainen@luke.fi
  • Bärring, Rossby Centre, Swedish Meteorological and Hydrological Institute, Folkborgsvägen 17, SE-60176 Norrköping, Sweden ORCID ID:E-mail: lars.barring@smhi.se
  • Andersson Gull, Skogforsk, Box 3, SE-91821 Sävar, Sweden ORCID ID:E-mail: bengt.andersson@skogforsk.se
article id 1410, category Research article
Seppo Nevalainen, Juho Matala, Kari T. Korhonen, Antti Ihalainen, Ari Nikula. (2016). Moose damage in National Forest Inventories (1986–2008) in Finland. Silva Fennica vol. 50 no. 2 article id 1410. https://doi.org/10.14214/sf.1410
Highlights: Almost 100 000 stands were studied; The proportion of damage doubled during the study period; Tree species mixture had a clear effect on the damage frequency; The damage was more common in mineral soils than in peatlands, in artificially than in naturally regenerated stands and in stands that needed thinning or clearing or in which soil preparation was used.

The occurrence of moose damage was studied using data from three National Forest Inventories (NFIs) accomplished between 1986 and 2008 in Finland. The combined data included a total of 97 390 young stands. The proportion of moose damage increased from 3.6% to 8.6% between the 8th NFI (1986–1994) and the 10th NFI (2004–2008). The majority (75%) of the damage occurred in Scots pine-dominated stands. The proportion of damage was higher in aspen-dominated stands than in stands dominated by any other tree species. The tree species mixture also had a clear effect on the occurrence of damage. Pure Scots pine stands had less damage than mixed Scots pine stands, and moose damage decreased linearly with the increasing proportion of Scots pine. Stands on mineral soil had more frequent moose damage than stands on peatlands. The fertility class of the site had no straightforward effect on the damage frequency. Artificially regenerated stands had more damage than naturally regenerated stands. Accomplished soil preparation measures and the need for thinning or clearing operations increased moose damage. High proportions of moose damage in young stands were found around the country. In the 10th NFI, the largest concentration of damage was found in southwestern Finland. Our study shows the temporal and spatial changes in the occurrence of moose damage and pinpoints some important silvicultural factors affecting the relative risk of young stands over a large geographical area.

  • Nevalainen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail: seppo.nevalainen@luke.fi (email)
  • Matala, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail: juho.matala@luke.fi
  • Korhonen, Natural Resources Institute Finland (Luke), Economics and society, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail: kari.t.korhonen@luke.fi
  • Ihalainen, Natural Resources Institute Finland (Luke), Economics and society, P.O. Box 18, FI-01301 Vantaa, Finland ORCID ID:E-mail: antti.ihalainen@luke.fi
  • Nikula, Natural Resources Institute Finland (Luke), Economics and society, P.O. Box 16, FI-96301 Rovaniemi, Finland ORCID ID:E-mail: ari.nikula@luke.fi
article id 1323, category Research article
Tiina Laine, Kalle Kärhä, Antti Hynönen. (2016). A survey of the Finnish mechanized tree-planting industry in 2013 and its success factors. Silva Fennica vol. 50 no. 2 article id 1323. https://doi.org/10.14214/sf.1323
Highlights: In 2013, 31 planting machines were operated by 22 businesses and planted 4.7 million seedlings on 2663 hectares in Finland; Critical success factors included expertise of planting machine operators, high quality planting, adequate amount of work, stoniness, and removal of slash; Growth of the industry will depend on improved cost-efficiency, appropriate worksites, marketing, development of planting machines.

The aim of the study was to update the information pertaining to mechanized tree-planting activity in Finland in 2013 and its success factors. All businesses providing a mechanized tree-planting service were interviewed and asked to describe their equipment and activities, identify critical success factors (CSFs), and suggest areas for improvement. In 2013, 31 planting machines (18 Bracke P11.a, 11 M-Planter and 2 Risutec) operated by 22 businesses planted approximately 4.7 million seedlings on 2663 hectares. CSFs included expertise of planting machine operators, high quality planting, adequate amount of work, stoniness, and removal of slash. Based on the survey, some recommendations for improving mechanized planting work can be made. Firstly, improving the cost-efficiency of mechanized planting is necessary to enhance businesses’ profitability. Secondly, worksite selection is crucial as stoniness, stumps and slash debris diminish productivity. Lastly, the popularity of mechanized planting in the future will benefit from more marketing. Many businesses were unwilling to increase the area of service, invest in new equipment, or increase the volume of planting work but they believed that mechanized planting will become more popular in the near future.

  • Laine, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: tiina.laine@luke.fi (email)
  • Kärhä, Stora Enso Wood Supply Finland, P.O. Box 309, FI-00101 Helsinki, Finland ORCID ID:E-mail: kalle.karha@storaenso.com
  • Hynönen, University of Eastern Finland, Faculty of Science and Forestry, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail: hynonena@gmail.com
article id 1408, category Research article
Anna Katharina Franke, Pasi Aatsinki, Ville Hallikainen, Esa Huhta, Mikko Hyppönen, Vesa Juntunen, Kari Mikkola, Seppo Neuvonen, Pasi Rautio. (2015). Quantifying changes of the coniferous forest line in Finnish Lapland during 1983–2009. Silva Fennica vol. 49 no. 4 article id 1408. https://doi.org/10.14214/sf.1408
Highlights: Volume of the growing stock of spruce and pine has increased in forests and in timber lines during the past 26 years; Spruce stem numbers increased on average, while pine stem numbers remained stable and location-specific variation was observed; Presuming that the ongoing trend of increasing temperature will remain, the enhanced forest regeneration and growth may result in extension of forests in the future.

The boreal timber- and tree-line forests grow in harsh environmental conditions in their outermost distribution limit. Here even small environmental changes may cause dramatic changes in the distribution of tree species. We examined changes of the forest lines of Norway spruce (Picea abies (L.) H. Karst.) and Scots pine (Pinus sylvestris L.) in Finnish Lapland five times during 1983–2009. We monitored the number of stems and the volume of the growing stock in thirteen different locations in forest-line areas. The linear temporal trends and the variations of these response variables were used as indicators of a possible change during the study period. Spruce showed a significant increase both in the volume of the growing stock (up to 40% increase) and in the total stem number (up to 100% increase). A significant increase in the volume of the growing stock was observed in the pine data as well (up to 70% increase), whereas the stem number stagnated or even decreased. The results suggest that spruce needs favourable conditions to have an abundant regeneration, but after the establishment the seedlings seem to be more resistant against biotic and abiotic disturbances than pine seedlings. The increasing stand volume might result in a climate-related northward and upward extension of forests in the future. However, our results show that responses in the boreal forest line are species and location specific and a more favourable climate does not necessarily lead to an advance of the coniferous forest line.

  • Franke, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Eteläranta 55, FI-96301 Rovaniemi, Finland ORCID ID:E-mail: anna.franke@fau.de (email)
  • Aatsinki, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Eteläranta 55, FI-96301 Rovaniemi, Finland ORCID ID:E-mail: pasi.aatsinki@luke.fi
  • Hallikainen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Eteläranta 55, FI-96301 Rovaniemi, Finland ORCID ID:E-mail: ville.hallikainen@luke.fi
  • Huhta, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Eteläranta 55, FI-96301 Rovaniemi, Finland ORCID ID:E-mail: esa.huhta@luke.fi
  • Hyppönen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Eteläranta 55, FI-96301 Rovaniemi, Finland ORCID ID:E-mail: mikko.hypponen@luke.fi
  • Juntunen, The Sámi Education Institute, Menesjärventie 4, P.O. Box 50, FI-99871 Inari, Finland ORCID ID:E-mail: vesa.juntunen@sogsakk.fi
  • Mikkola, Natural Resources Institute Finland (Luke), Economics and society, Eteläranta 55, FI-96301 Rovaniemi, Finland ORCID ID:E-mail: kari.mikkola@luke.fi
  • Neuvonen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 68, FI-80101, Joensuu, Finland ORCID ID:E-mail: seppo.neuvonen@luke.fi
  • Rautio, Natural Resources Institute Finland (Luke), Bio-based business and industry, Eteläranta 55, FI-96301 Rovaniemi, Finland ORCID ID:E-mail: pasi.rautio@luke.fi
article id 1188, category Research article
Kristina Wallertz, Henrik Nordenhem, Göran Nordlander. (2014). Damage by the pine weevil Hylobius abietis to seedlings of two native and five introduced tree species in Sweden. Silva Fennica vol. 48 no. 4 article id 1188. https://doi.org/10.14214/sf.1188
Highlights: Both native and introduced confer species in Sweden can be highly susceptible to damage by the pine weevil; Douglas fir and Sitka spruce were generally the most damaged among six studied conifer species; The results highlight some of the risks in establishing exotic tree species for forest production.
There is increasing interest in using introduced species in Swedish forestry in response to climate change, but it is important to assess their resistance to native pests. Thus, we compared the extent of pine weevil feeding on two dominant native conifers, Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.), the non-host deciduous broadleaf hybrid aspen (Populus × wettsteinii Hämet-Ahti) and four introduced conifers: Douglas fir (Pseudotsuga menziesii (Mirb.) Franco), hybrid larch (Larix × marschlinsii Coaz), Sitka spruce (Picea sitchensis (Bong.) Carriére) and lodgepole pine (Pinus contorta Douglas ex Loudon). The extent of feeding damage on seedlings and its effect on their vitality were examined in a field study in south-central Sweden and a laboratory experiment, which gave largely consistent results. Generally, the species most heavily attacked by the pine weevil, in both experiments, were Douglas fir and Sitka spruce. In the field experiment pine weevils killed or severely damaged significantly higher proportions of Douglas fir and Sitka spruce seedlings (60%) than any other species except Norway spruce (49%). Among conifer seedlings the proportions of killed or severely damaged seedlings were lowest for Scots pine and hybrid larch (27%) and Lodgepole pine (36%). The results indicate that most conifer species planted on young clear-cuttings in Sweden need some kind of pine weevil protection, and the possibility that introducing new tree species might increase damage caused by pests must be considered. For instance, widespread use of hybrid aspen could reduce damage by pine weevils, but increase damage by other, untested pests or pathogens.
  • Wallertz, Unit for Field-based Forest Research, Swedish University of Agricultural Sciences (SLU), Asa Research Station, SE-36030 Lammhult, Sweden ORCID ID:E-mail: kristina.wallertz@slu.se (email)
  • Nordenhem, Department of Ecology, Swedish University of Agricultural Sciences (SLU), P.O. Box 7044, SE-75007 Uppsala, Sweden ORCID ID:E-mail: henrik.nordenhem@slu.se
  • Nordlander, Department of Ecology, Swedish University of Agricultural Sciences (SLU), P.O. Box 7044, SE-75007 Uppsala, Sweden ORCID ID:E-mail: goran.nordlander@slu.se
article id 1161, category Research article
Tiina Laine, Veli-Matti Saarinen. (2014). Comparative study of the Risutec Automatic Plant Container (APC) and Bracke planting devices. Silva Fennica vol. 48 no. 3 article id 1161. https://doi.org/10.14214/sf.1161
Highlights: As currently designed, the prototype Risutec APC fitted with an automatic feeding system offers no significant advantage over the Bracke planting device in terms of planting productivity or quality; Cost estimates suggest that an idealized automated feeding system could increase productivity and decrease unit costs.
The productivity of mechanized planting could be increased by minimizing the time spent manually reloading seedling cassettes. This study compared the work-time distribution, productivity and quality of the prototype Risutec APC fitted with an automatic feeding system and the commonly-used and manually-loaded Bracke P11.a. An approach of comparative time study was employed that compared performance of two operators using both machines in four sites where slash and stumps had been removed. Operating costs were estimated and compared for these two machines and an idealized machine with automatic feeding system (referred as AUT). AUT was assumed to be similar to the Bracke planting machine with the only difference being in automatic feeding. Productivity of the Risutec APC (196 seedlings per productive work hour [pl PWh0–1]) was lower than that of Bracke (244 pl PWh0–1), making the unit cost 35.7% higher. A large portion (17.6%) of the productive work time of Risutec APC was interrupted by malfunctions, so it cannot be considered robust and reliable yet. Quality of the planting work was reasonable for both machines. The results suggest that an idealized AUT could increase planting capacity (hectares per year [ha yr–1]) by 15.4% and lower the unit cost (Euro per seedling [€ pl–1]) by 4.7% compared to today’s machines. The importance of an automated feeding system increases with planting efficiency because relatively more time is spent reloading seedlings. Proper automatic feeding system could offer a cost-effective solution and could enhance productivity, but the Risutec APC has yet to meet the technical and economic standards required to be competitive.
  • Laine, Finnish Forest Research Institute (METLA), Suonenjoki Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: tiina.laine@metla.fi (email)
  • Saarinen, Finnish Forest Research Institute (METLA), Suonenjoki Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: veli-matti.saarinen@metla.fi
article id 1016, category Research article
Karin Johansson, Eva Ring, Lars Högbom. (2013). Effects of pre-harvest fertilization and subsequent soil scarification on the growth of planted Pinus sylvestris seedlings and ground vegetation after clear-felling. Silva Fennica vol. 47 no. 4 article id 1016. https://doi.org/10.14214/sf.1016
Highlights: Pre-harvest N fertilization had no significant effect on seedling growth and ground vegetation biomass; Scarification improved seedling survival and growth and reduced the amount of ground vegetation; Without scarification, pre-harvest fertilization increased the amount of damaged seedlings.
Fertilization and scarification are both performed to increase tree production at different stages of forest rotation periods. In this study, the effects of previous nitrogen fertilizations and scarification after clear felling on planted Pinus sylvestris L. seedlings and ground vegetation were investigated. Two fertilization experiments established around 1980 were harvested in 2006, after which the plots were scarified by disc trenching and re-planted. The plots had been repeatedly fertilized over a 20-year period before harvesting, with total N doses of 0, 450, 900 or 1800 kg N ha-1. After five growing seasons, the growth, survival and nutrient contents of the seedlings were measured, and ground vegetation was collected to estimate its biomass and nutrient content. Pre-harvest fertilization alone had only minor effects on the results, but scarification increased both the survival and growth of the planted seedlings. However, without scarification, seedling mortality increased with increasing fertilization intensity. The ground vegetation biomass was higher in plots without scarification, but the total biomass of seedlings and ground vegetation was similar in all treatments. Scarification thus favored seedling growth at the expense of ground vegetation. Only a few effects on nutrient content were found, but there were no signs of nutrient imbalance in any of the treatments. At higher levels of fertilization, the K:N ratio in the seedlings decreased while the K content in the ground vegetation increased. Overall, scarification had a greater impact than pre-harvest fertilization on the planted seedlings and the ground vegetation.
  • Johansson, Skogforsk, The Forestry Research Institute of Sweden, Ekebo 2250, SE-268 90 Svalöv, Sweden & Southern Swedish Forest Research Centre, SLU, Box 49, SE-230 53 Alnarp, Sweden ORCID ID:E-mail: karin.johansson@skogforsk.se (email)
  • Ring,  Skogforsk, The Forestry Research Institute of Sweden, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail: eva.ring@skogforsk.se
  • Högbom,  Skogforsk, The Forestry Research Institute of Sweden, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail: lars.hogbom@skogforsk.se
article id 920, category Research article
Hannu Hökkä, Jaakko Repola, Mikko Moilanen, Markku Saarinen. (2012). Seedling establishment on small cutting areas with or without site preparation in a drained spruce mire – a case study in northern Finland. Silva Fennica vol. 46 no. 5 article id 920. https://doi.org/10.14214/sf.920
A large proportion of drained spruce mire stands is currently approaching regeneration maturity in Finland. We studied the effect of cutting – small canopy openings (78, 177, and 314 m2) and small clear-cuts (0.25–0.37 ha) – with or without site preparation (scalping) on the establishment of natural Norway spruce seedlings in one experimental drained spruce mire stand in northern Finland. The cuttings were made in winter 2004–2005 and site preparation with scalping in early June 2005. The experimental design was composed of four blocks with altogether four clear-cuts and 33 canopy openings. The seedling establishment was surveyed annually (2006, 2008–2010) from five circular sample plots (one 10 m2 and four 5 m2 plots in size) located within the canopy openings and from 18 circular 5 m2 sample plots systematically located in the scalped and untreated halves of the clear-cuts. Site preparation was found unnecessary, because it resulted in a clearly lower number of seedlings in the openings. A slight negative effect was also found in the clear-cuts. In the two years following the cuttings, the number of seedlings increased quickly in the canopy openings, but more gradually in the clear-cut areas. In 2010, on average 15 500 new seedlings were observed in the canopy openings and 6700 in the clear-cut areas, of which 5050 and 1200, respectively, were >0.1 m tall spruces. The proportion of birch increased in the last two years, being ca. 22% in the openings and 45% in the clear-cuts in 2010. The spatial distribution of seedlings was more uneven in the clear-cuts than in the openings, with 41% and 20% of survey plots empty, respectively.
  • Hökkä, Finnish Forest Research Institute, Rovaniemi Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland ORCID ID:E-mail: hannu.hokka@metla.fi (email)
  • Repola, Finnish Forest Research Institute, Rovaniemi Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland ORCID ID:E-mail: jaakko.repola@metla.fi
  • Moilanen, Finnish Forest Research Institute, Oulu Unit, P.O. Box 413, FI-90014 University of Oulu, Finland ORCID ID:E-mail: mikko.moilanen@metla.fi
  • Saarinen, Finnish Forest Research Institute, Parkano Unit, Kaironiementie 15, FI-39700 Parkano, Finland ORCID ID:E-mail: markku.saarinen@metla.fi
article id 64, category Research article
Karin Johansson, Ola Langvall, Johan Bergh. (2012). Optimization of environmental factors affecting initial growth of Norway spruce seedlings. Silva Fennica vol. 46 no. 1 article id 64. https://doi.org/10.14214/sf.64
The purpose of the study was to create a near optimal environment for seedling establishment and growth, without the restrain of water and nutrients but under climate conditions typical for the region. This to give us valuable knowledge about the growth potential of different seedling types in the field. The experimental site was situated in southern Sweden. Six treatment combinations were applied including two site treatments; 1) soil inversion, i.e. the control treatment, and 2) soil inversion, drip irrigation and fertilization combined with plastic cover mulch, i.e. the optimization treatment, and three seedling types of Norway spruce (Picea abies L. Karst.), (a) a 2-year-old Plug+1 seedling, (b) a 1.5-year-old containerized seedling and (c) a 10-week-old mini seedling. Effects on seedling nutrient status and growth were studied during the first three years after planting. Height, diameter and biomass of the seedlings grown in the optimized environment were significantly greater than for seedlings grown in the control. The Plug+1 seedlings grown in the optimization treatment had, after three years, reached a height of 124 cm, while the containerized seedlings were 104 cm and the mini seedlings 45 cm. In practical plantations, this height is usually gained after 5–10 years depending on planting conditions. Biomass partitioning did not differ between optimization treatments, but between seedling types. The mini seedlings allocated less biomass to the roots and more biomass to needles and stem in comparison with the two other seedling types. Mini seedlings also broke bud earlier. Throughout the experimental period, seedling nutrient status for all treatment combinations was followed and a balanced nutrient supply of macro- and micronutrients was given in the optimization treatment. Nutrient concentrations were constantly higher in seedlings grown in the optimization treatment, but the difference decreased over time. Results from this study shows that, by improving site conditions associated with fast establishment, growth check can be avoided.
  • Johansson, Skogforsk, Svalöv, Sweden ORCID ID:E-mail: karin.johansson@skogforsk.se (email)
  • Langvall, SLU, Asa Forest Research Station, Sweden ORCID ID:E-mail:
  • Bergh, SLU, Southern Swedish Forest Research Centre, Sweden ORCID ID:E-mail:
article id 87, category Research article
Radek Bace, Miroslav Svoboda, Pavel Janda. (2011). Density and height structure of seedlings in subalpine spruce forests of Central Europe: logs vs. stumps as a favourable substrate. Silva Fennica vol. 45 no. 5 article id 87. https://doi.org/10.14214/sf.87
Decaying logs and stumps provide an important seedling substrate in natural subalpine forests. However, only stumps present such a role in managed forests. The aim of this study was to assess the differences in the process of seedling colonization between logs and stumps. The study was carried out in the Czech Republic, in two old-growth subalpine spruce forests located in the Bohemian Forest and Ash Mts., dominated by Athyrium distentifolium Opiz and Vaccinium myrtillus L. undergrowth, respectively. Norway spruce (Picea abies (L.) Karst.) regeneration growing on logs, stumps and non-coarse woody debris (CWD) microsites was surveyed. Regeneration (height 0–2.0 m) densities exceeded 5000 individuals per ha on both sites. The average density of P. abies regeneration per square meter of substrate was 0.3-5.7-19.6 and 0.5-3.8-11.0 on non-CWD microsites, logs and stumps, located in A. distentifolium and V. myrtillus undergrowth, respectively. Stumps and non-CWD microsites dominated by V. myrtillus, supported a higher proportion of taller seedlings per plot compared to the small seedlings growing on logs and non-CWD dominated by A. distentifolium ground-cover. The disproportion in regeneration densities between the stumps and the original logs decreased with increasing stages of decay. The tallest regeneration growing on stumps (root-soil plates) was significantly older than that growing on the logs (stems). Based on these two latter findings, the stumps appeared to provide suitable seedling substrates several years earlier than the logs did. Therefore, we conclude that the stumps play a more important role (relative to their covered area, 21–28 m2 ha–1) in terms of suitable microsites for regeneration, than the logs do.
  • Bace, Czech University of Life Sciences, Faculty of Forestry and Wood Sciences, Prague, Czech Republic ORCID ID:E-mail: bace@fld.czu.cz (email)
  • Svoboda, Czech University of Life Sciences, Faculty of Forestry and Wood Sciences, Prague, Czech Republic ORCID ID:E-mail:
  • Janda, Czech University of Life Sciences, Faculty of Forestry and Wood Sciences, Prague, Czech Republic ORCID ID:E-mail:
article id 98, category Research article
Meeri Pearson, Markku Saarinen, Kari Minkkinen, Niko Silvan, Jukka Laine. (2011). Mounding and scalping prior to reforestation of hydrologically sensitive deep-peated sites: factors behind Scots pine regeneration success. Silva Fennica vol. 45 no. 4 article id 98. https://doi.org/10.14214/sf.98
Watering up typically ensues after clearcutting forestry-drained peatland forests. Thus, the effectiveness of maintenance drainage and soil preparation procedures becomes paramount for establishing a new generation of commercial forest. Mounding is the primary method of soil preparation applied in regeneration sites lying on deep peat. As raised planting spots, mounds are resistant to waterlogging and assumed to be beneficial for organic matter (OM) decomposition via, e.g., increased soil aeration and temperature, which would also enhance seedling growth. In recent years, however, less intensive and cheaper alternatives like scalping have been sought with some reported cases of success. Our case study investigated the survival and growth of Scots pine outplants in mounds, scalps, and unprepared microsites along a moisture gradient. After three growing seasons, mounding accelerated neither seedling growth nor OM decomposition relative to the unprepared treatment. Survival in mounds was nonetheless superior overall. Scalps behaved as water collecting depressions leading to a catastrophic regeneration result. Based on our findings, water table level (WTL) overrides other growth-controlling factors in excess moisture conditions. To combat watering up coupled with greater than normal rainfall, we recommend reforestation strategies which provide elevated, prepared planting spots (i.e., mounds) or utilize unprepared, higher microforms.
  • Pearson, Finnish Forest Research Institute, Western Finland Regional Unit, Kaironiementie 15, FI-39700 Parkano, Finland ORCID ID:E-mail: meeri.pearson@metla.fi (email)
  • Saarinen, Finnish Forest Research Institute, Western Finland Regional Unit, Kaironiementie 15, FI-39700 Parkano, Finland ORCID ID:E-mail:
  • Minkkinen, University of Helsinki, Department of Forest Sciences, Helsinki, Finland ORCID ID:E-mail:
  • Silvan, Finnish Forest Research Institute, Western Finland Regional Unit, Kaironiementie 15, FI-39700 Parkano, Finland ORCID ID:E-mail:
  • Laine, Finnish Forest Research Institute, Western Finland Regional Unit, Kaironiementie 15, FI-39700 Parkano, Finland ORCID ID:E-mail:
article id 97, category Research article
Hannu Hökkä, Jaakko Repola, Mikko Moilanen, Markku Saarinen. (2011). Seedling survival and establishment in small canopy openings in drained spruce mires in Northern Finland. Silva Fennica vol. 45 no. 4 article id 97. https://doi.org/10.14214/sf.97
A large proportion of drained spruce mire stands is currently approaching regeneration maturity in Finland. Traditional regeneration methods with effective site preparation and planting generally result in satisfactory seedling stands also in spruce mires. However, natural regeneration methods may be more appropriate in protecting watercourses and minimizing regeneration costs. We studied the survival of advance growth and establishment of new seedlings in small canopy openings that were cut at three different diameters in two experimental drained spruce mire stands in Northern Finland (Tervola and Oulu) in 2004. The number of seedlings was repeatedly surveyed from five small circular plots (one 10 m2 and four 5 m2 plots in size) located within the opening. Advance growth which survived the cutting and new seedlings were separated in the surveys. The density of advance growth was on average 9000 ha–1 after cutting, and it decreased by 30% during the five-year monitoring period (2006–2010) due to natural mortality. The number of new seedlings increased rapidly within the three years after cutting the openings. In 2010, 11 000–26 000 new seedlings ha–1 in Tervola and 12 000–16 000 ha–1 in Oulu on average were observed. The size of the opening had no clear effect on the regeneration result. The proportion of birch of the new seedlings increased with time and opening size in Tervola. The results show that Norway spruce regenerates naturally in small canopy openings cut in mature drained spruce mire stands.
  • Hökkä, Finnish Forest Research Institute, Rovaniemi Research Unit, FI-96301 Rovaniemi, Finland ORCID ID:E-mail: hannu.hokka@metla.fi (email)
  • Repola, Finnish Forest Research Institute, Rovaniemi Research Unit, FI-96301 Rovaniemi, Finland ORCID ID:E-mail:
  • Moilanen, Finnish Forest Research Institute, Muhos Research Unit, Muhos, Finland ORCID ID:E-mail:
  • Saarinen, Finnish Forest Research Institute, Parkano Research Unit, Parkano, Finland ORCID ID:E-mail:
article id 146, category Research article
Karri Uotila, Juho Rantala, Timo Saksa, Pertti Harstela. (2010). Effect of soil preparation method on economic result of Norway spruce regeneration chain. Silva Fennica vol. 44 no. 3 article id 146. https://doi.org/10.14214/sf.146
Economic result of forest regeneration chains, based either on spot mounding or on disc trenching and planting of Norway spruce (Picea abies [L.] Karst.) seedlings, were clarified and compared to each other. First, effects of soil preparation method on early development of Norway spruce stands were measured from field experiments. Second, the effects of soil preparation method on stand level management programs were modelled. The modelling was based on growth simulation and investment calculations. The soil preparation methods substantially affected early development of a stand. The density of the removed trees in early cleaning was 56% higher on the disc-trenched area compared to the spot-mounded area. The difference was especially high (120%), close by (< 25 cm) the remained spruce seedlings. There was also a difference between the methods in the growth of crop spruces; at biological age of 8 years, the mean height of spruce was 110 cm on the spot-mounded area and 68 cm on the disc-trenched area. The differences led to divergent management programs between the areas. The disc-trenched area needed three young stand management operations whereas two was enough at the spot-mounded area. Although disc trenching is a less expensive method than spot mounding, the total management costs were higher in disc trenching than in spot mounding. Furthermore, incomes from the first commercial thinning were higher when regeneration based on spot mounding. At the interest rate of 3%, the investment in spot mounding had 329 EUR ha–1 higher net present value than the investment in disc trenching.
  • Uotila, Finnish Forest Research Institute, Suonenjoki, Finland ORCID ID:E-mail: karri.uotila@metla.fi (email)
  • Rantala, Finnish Forest Research Institute, Suonenjoki, Finland ORCID ID:E-mail:
  • Saksa, Finnish Forest Research Institute, Suonenjoki, Finland ORCID ID:E-mail:
  • Harstela, Finnish Forest Research Institute, Suonenjoki, Finland ORCID ID:E-mail:
article id 172, category Research article
Guntis Brumelis, Maris Strazds, Zanna Eglava. (2009). Stand structure and spatial pattern of regeneration of Pinus sylvestris in a natural treed mire in Latvia. Silva Fennica vol. 43 no. 5 article id 172. https://doi.org/10.14214/sf.172
We examined the regeneration patterns of Pinus sylvestris L. in a natural treed mire in the hemiboreal zone in Latvia. Data on tree stem age and size was collected in 207 fine-scale plots (10 m2) and 4 medium-scale plots (400 m2). Size structure of living and dead trees was also estimated on transects 180-m and 250-m length and 10-m width. In addition, the vegetation was described in 1-m2 plots to determine preferred microsites for P. sylvestris establishment. Pinus sylvestris showed continuous regeneration by an inverse J-shaped age and size structure. Pulses of mortality induced by fire were also evident. Regeneration of P. sylvestris was mostly on Sphagnum magellanicam Brid. hummocks free of competition from Ledum palustre L. and graminoids, which were found in small interspersed patches throughout the mire. The spatial pattern of tree cohorts differed, probably due to changing moisture conditions altering seedbed conditions and by fire-induced mortality in specific areas. Recent rapid invasion by Betula pubescens Ehrh. and Picea abies (L.) Karst. in a post-fire period since 1971 is probably associated with drier conditions.
  • Brumelis, Faculty of Biology, University of Latvia, Kronvalda bulv. 4, Riga LV1586, Latvia ORCID ID:E-mail: guntis.brumelis@lu.lv (email)
  • Strazds, Faculty of Biology, University of Latvia, Kronvalda bulv. 4, Riga LV1586, Latvia ORCID ID:E-mail:
  • Eglava, Faculty of Biology, University of Latvia, Kronvalda bulv. 4, Riga LV1586, Latvia ORCID ID:E-mail:
article id 191, category Research article
Katri Luostarinen, Noora Huotari, Eila Tillman-Sutela. (2009). Effect of regeneration method on growth, wood density and fibre properties of downy birch (Betula pubescens Ehrh.). Silva Fennica vol. 43 no. 3 article id 191. https://doi.org/10.14214/sf.191
Short rotation tree stands, established by coppicing, are nowadays used mainly for energy purposes in Fennoscandia, but their usage for pulp raw material may increase in the future. Downy birch (Betula pubescens), which is commonly used for pulp production in boreal zone, has good sprouting capacity. However, it is not known if the fibre properties of sprout-originated downy birches differ from those of seed-originated ones. Therefore, fibre length and width of sprout- and seed-originated downy birches grown on fertile soil were measured at a stand age of 25 years. Additionally annual ring width, stem height and diameter, and wood density were studied to compare the growth and wood properties of sprout- and seed-originated birches. Annual rings were slightly wider in sprout- than in seed-originated birches, whereas no differences were observed in wood density. Fibres, too, were slightly longer and wider in sprout- than in seed-originated trees. Still these minor differences observed here are hardly significant for the industries using birch wood. Consequently downy birch wood from coppiced stands is well suited for pulp. The advantages of coppice, i.e. rapidity and low costs of establishment, productivity, and the ability of downy birch to grow on untypical forest sites, may even increase the importance of the wood coming from coppiced birch stands in the future.
  • Luostarinen, Faculty of Forest Sciences, University of Joensuu, P.O. Box 111, FI-80101, Joensuu, Finland ORCID ID:E-mail: katri.luostarinen@joensuu.fi (email)
  • Huotari, Finnish Forest Research Institute, Muhos Research Unit, Kirkkosaarentie, FI-91500 Muhos, Finland ORCID ID:E-mail:
  • Tillman-Sutela, Finnish Forest Research Institute, Muhos Research Unit, Kirkkosaarentie, FI-91500 Muhos, Finland ORCID ID:E-mail:
article id 217, category Research article
Ville Kankaanhuhta, Timo Saksa, Heikki Smolander. (2009). Variation in the results of Norway spruce planting and Scots pine direct seeding in privately-owned forests in southern Finland. Silva Fennica vol. 43 no. 1 article id 217. https://doi.org/10.14214/sf.217
This study describes the variation in the planting results for 3-year-old Norway spruce (Picea abies (L.) Karst.) and 4-year-old Scots pine (Pinus sylvestris L.) using direct seeding in privately-owned regeneration areas in southern Finland. The study material consists of operative forest regeneration quality management inventory areas from the years 2000–2006. The effect of both the regional and the administrative levels as well as ecological factors was modelled on the basis of the hierarchy structure forestry centre, Forest Owners’ Association (= FOA), forestry professional, regeneration area and sample plot. The major part of the variation occurred at the sample plot and regeneration area level. Particular attention was paid to observation of the clustered spatial distribution of Scots pine seedlings. The FOA and forestry professional levels explained 5% of the variation in Norway spruce planting and 11% of the variation in Scots pine direct seeding. Applied forest regeneration operations, site and soil characteristics were included in the fixed effects. In the planting of Norway spruce the most important factor explaining the regeneration result was soil preparation. Mounding produced better results than patching and disc trenching. The site and soil characteristics were other important factors in the operations. The selection of direct seeding of Scots pine on too fertile, fine textured or moist sites yielded poor results.
  • Kankaanhuhta, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: ville.kankaanhuhta@metla.fi (email)
  • Saksa, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail:
  • Smolander, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail:
article id 217, category Research article
Ville Kankaanhuhta, Timo Saksa, Heikki Smolander. (2009). Variation in the results of Norway spruce planting and Scots pine direct seeding in privately-owned forests in southern Finland. Silva Fennica vol. 43 no. 1 article id 217. https://doi.org/10.14214/sf.217
This study describes the variation in the planting results for 3-year-old Norway spruce (Picea abies (L.) Karst.) and 4-year-old Scots pine (Pinus sylvestris L.) using direct seeding in privately-owned regeneration areas in southern Finland. The study material consists of operative forest regeneration quality management inventory areas from the years 2000–2006. The effect of both the regional and the administrative levels as well as ecological factors was modelled on the basis of the hierarchy structure forestry centre, Forest Owners’ Association (= FOA), forestry professional, regeneration area and sample plot. The major part of the variation occurred at the sample plot and regeneration area level. Particular attention was paid to observation of the clustered spatial distribution of Scots pine seedlings. The FOA and forestry professional levels explained 5% of the variation in Norway spruce planting and 11% of the variation in Scots pine direct seeding. Applied forest regeneration operations, site and soil characteristics were included in the fixed effects. In the planting of Norway spruce the most important factor explaining the regeneration result was soil preparation. Mounding produced better results than patching and disc trenching. The site and soil characteristics were other important factors in the operations. The selection of direct seeding of Scots pine on too fertile, fine textured or moist sites yielded poor results.
  • Kankaanhuhta, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: ville.kankaanhuhta@metla.fi (email)
  • Saksa, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail:
  • Smolander, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail:
article id 466, category Research article
Ilkka Korpela, Tuukka Tuomola, Timo Tokola, Bo Dahlin. (2008). Appraisal of seedling stand vegetation with airborne imagery and discrete-return LiDAR – an exploratory analysis. Silva Fennica vol. 42 no. 5 article id 466. https://doi.org/10.14214/sf.466
The potential for combined use of airborne discrete-return LiDAR and digital imagery in the classification and measurement of common seedling stand vegetation was examined in southern Finland (61°50’N, 24°20’E). Classification was based on spectral and textural image features in addition to geometric and radiometric features of the LiDAR. The accuracy of leaf-on, LiDAR-based terrain elevation models was tested as well as the accuracy of LiDAR in the measurement of vegetation heights. LiDAR-based canopy height and the range-normalized intensity of the LiDAR were strong explanatory variables in vegetation classification. Interspecies variation was observed in the height measurement accuracy of LiDAR for different tree, shrub and low vegetation canopies. Elevation models derived with 1–15 pulses per m2 showed an inherent noise of app. 15–25 cm, which restricts the use of LiDAR in regeneration assessment of very young stands. The spatial pattern of the competing vegetation was reproduced in classification-based raster surfaces, which could be useful in deriving meaningful treatment proposals.
  • Korpela, University of Helsinki, Dept of Forest Management, P.O. Box 27, FI-00014 University of Finland ORCID ID:E-mail: ilkka.korpela@helsinki.fi (email)
  • Tuomola, University of Helsinki, Dept of Forest Management, P.O. Box 27, FI-00014 University of Finland ORCID ID:E-mail:
  • Tokola, University of Helsinki, Dept of Forest Management, P.O. Box 27, FI-00014 University of Finland ORCID ID:E-mail:
  • Dahlin, University of Helsinki, Dept of Forest Management, P.O. Box 27, FI-00014 University of Finland ORCID ID:E-mail:
article id 250, category Research article
Saara Lilja-Rothsten, Michelle de Chantal, Chris Peterson, Timo Kuuluvainen, Ilkka Vanha-Majamaa, Pasi Puttonen. (2008). Microsites before and after restoration in managed Picea abies stands in southern Finland: effects of fire and partial cutting with dead wood creation. Silva Fennica vol. 42 no. 2 article id 250. https://doi.org/10.14214/sf.250
Different types of microsites, e.g. CWD (coarse woody debris), mounds, and uprooting pits, are important for tree regeneration and biodiversity. However, microsite diversity is greatly reduced in managed stands. We studied how restoration treatments changed microsite distribution in mature managed Picea abies stands. Four cutting treatments were used: uncut, low-CWD (5 m3 ha–1 of down retention trees, DRT, and 50 m3 ha–1 of standing retention trees), intermediate-CWD (as previous but leaving 30 m3 ha–1 of DRT), and high-CWD (as previous but with 60 m3 ha–1 of DRT). Timber harvested from stands ranged from 108–168 m3 ha–1. Half of the stands were burned, and half remained unburned. Sampling was stratified into upland and paludified biotopes within each stand. The pre-treatment microsite distributions were dominated by level ground in both biotopes; mounds and microsites on or next to CWD or a stump were slightly more abundant in the paludified than in the upland biotopes. Microsites were more diverse after cutting, with and without fire. The cutting treatment increased the relative abundances of microsites on or next to CWD. Fire consumed small diameter dead wood and flattened mounds. Microsites were more diverse in paludified than in upland biotopes. The results demonstrate that microsite diversity can rapidly be restored to structurally impoverished managed Picea stands despite a large portion of wood volume being harvested.
  • Lilja-Rothsten, University of Helsinki, Dept. of Forest Ecology, Finland ORCID ID:E-mail: saara.lilja@helsinki.fi (email)
  • Chantal, University of Helsinki, Dept. of Forest Ecology, Finland ORCID ID:E-mail:
  • Peterson, Dept. of Plant Biology, University of Georgia, Athens, GA, USA ORCID ID:E-mail:
  • Kuuluvainen, University of Helsinki, Dept. of Forest Ecology, Finland ORCID ID:E-mail:
  • Vanha-Majamaa, The Finnish Forest Research Institute, Vantaa Unit, Finland ORCID ID:E-mail:
  • Puttonen, The Finnish Forest Research Institute, Vantaa Unit, Finland ORCID ID:E-mail:
article id 308, category Research article
Ville Hallikainen, Mikko Hyppönen, Juha Hyvönen, Juhani Niemelä. (2007). Establishment and height development of harvested and naturally regenerated Scots pine near the timberline in North-East Finnish Lapland. Silva Fennica vol. 41 no. 1 article id 308. https://doi.org/10.14214/sf.308
Researchers and professionals in practical forestry have faced problems concerning the regeneration success of Scots pine in natural regeneration near the timberline in North-East Lapland. The aim of the study was to analyze the seedling establishment and seedling height development of Scots pine in seed-tree stands in the area. The average number of living pine seedlings in the study stands was about 1000 ha–1, but there was considerable variation between the stands. The seedling density was modelled using a multinomial logistic regression with a random factor. Forest site type and the time since seed-tree cutting were the most significant explanatory variables in the model. The probability of reaching the acceptable seedling density was higher on dry site types than on the more fertile ones. The probability increased with the time elapsed since the regeneration activities. Effective temperature sum and the number of intermediate pines also positively affected the probability, but the presence of residual trees negatively. On northern and eastern slopes the probability was lower than on southern and western ones. Seedling height was modelled using a linear mixed model. The age of a dominant seedling was the most positively effective explanatory variable in the height development model. Other positively affecting significant predictors were time since seed-tree cutting, number of intermediate birches, and distance between a seedling and the nearest seed tree. Degree of paludification had a negative effect. The study suggests that the regeneration of Scots pine in North-East Lapland is a relatively slow process.
  • Hallikainen, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland ORCID ID:E-mail: ville.hallikainen@metla.fi (email)
  • Hyppönen, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland ORCID ID:E-mail: mikko.hypponen@metla.fi
  • Hyvönen, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland ORCID ID:E-mail: juha.hyvonen@metla.fi
  • Niemelä, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland ORCID ID:E-mail:
article id 329, category Research article
Vesa Juntunen, Seppo Neuvonen. (2006). Natural regeneration of Scots pine and Norway spruce close to the timberline in northern Finland. Silva Fennica vol. 40 no. 3 article id 329. https://doi.org/10.14214/sf.329
Two different datasets were analyzed in order to clarify the factors that affect regeneration success of Scots pine (Pinus sylvestris) and Norway spruce (Picea abies) in the climatically extreme areas in northern Finland. First, pine seed maturity and the number of cones in the trees were investigated at five pairs of study sites during the period 1997–2003. Secondly, the rate of seedling establishment and seedling survival of Scots pine and Norway spruce were monitored and compared among three different timberline zones (forest zone, timberline, tree line) in 13 localities during the period 1983–1999. The first study showed that both cone production (bud formation) and seed maturity may be limiting factors for successful reproduction in the climatically marginal habitats. Seed maturity correlated well with the temperature sum of the summer, but variation in the number of cones had a periodic component rather than strictly following the temperature sum of the summer of bud formation. Monitoring surveys since 1983 showed that pine and spruce regenerated more or less regularly in all the zones during 1983–1999. However, seedling mortality of pines was much higher compared to spruce. In general, initially small sized seedlings showed higher mortality compared with larger ones. The results suggest that besides restrictions in reproduction, stand dynamics in the timberline habitats are strongly controlled by seedling mortality due to a variety of causes.
  • Juntunen, Finnish Forest Research Institute, Kolari Research Unit, Muoniontie 21, FI-95900 Kolari, Finland ORCID ID:E-mail: vesa.juntunen@metla.fi (email)
  • Neuvonen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail:
article id 376, category Research article
Nils Lexerød, Trond Eid. (2005). Recruitment models for Norway spruce, Scots pine, birch and other broadleaves in young growth forests in Norway. Silva Fennica vol. 39 no. 3 article id 376. https://doi.org/10.14214/sf.376
The objective of the present study was to develop recruitment models for Norway spruce, Scots pine, birch and other broadleaves in young growth forests in Norway. The models were developed from permanent sample plots established by the National Forest Inventory, and they will be included in a growth simulator that is part of a large-scale forestry scenario model. The modelling was therefore restricted to independent variables directly or indirectly available from inventories for practical forest management planning. A two-stage modelling approach that suited the stochastic nature of recruitment in boreal forests was used. Models predicting the probability of recruitment were estimated in a first stage, and conditional models for the number of recruits were developed in a second. The probability models as well as the conditional models were biologically realistic and logical. The goodness of fit tests revealed that the probability models fitted the data well, while the coefficients of determination for the conditional models were relatively low. No independent test data were available, but comparisons of predicted and observed number of recruits in different sub-groups of the data revealed few large deviations. The high level of large random errors was probably due to the great variability observed in number of recruits rather than inappropriate specifications of the models. Provided the generally high level of uncertainty connected to analysis performed with large-scale forestry scenario models and the stochastic nature of recruitment, the presented models seem to give satisfactory levels of accuracy.
  • Lexerød, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, P.O. Box 5003, NO-1432 Ås, Norway ORCID ID:E-mail: nils.lexerod@umb.no (email)
  • Eid, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, P.O. Box 5003, NO-1432 Ås, Norway. E-mail nils.lexerod@umb.no ORCID ID:E-mail:
article id 372, category Research article
Ulf Sikström. (2005). Pre-harvest soil acidification, liming or N fertilization did not significantly affect the survival and growth of young Norway spruce. Silva Fennica vol. 39 no. 3 article id 372. https://doi.org/10.14214/sf.372
Acidification, liming, and N fertilization affect a number of soil properties. Such changes may have an impact on forest regeneration and yield. The aim of this study was to investigate the survival and growth of Picea abies (L.) Karst. planted on plots that had been acidified (in 12 annual treatments totalling 600 or 1200 kg S ha–1 in the form of elemental sulphur), limed (12 x 500 = 6000 kg lime ha–1 in the form of CaCO3) or N-fertilized (3 x 200 = 600 kg N ha–1 in the form of urea) prior to harvest. Trees growing on plots treated with a combination of the N plus the lower S application were also tested. None of the treatments, applied in three replicate stands, significantly influenced either survival or growth of Picea abies trees during the first 11 growing seasons after planting.
  • Sikström, SkogForsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail: ulf.sikstrom@skogforsk.se (email)
article id 397, category Research article
Nuutti Kiljunen. (2005). Pricing the risk of the quality-guarantee in a stand establishment service. Silva Fennica vol. 39 no. 1 article id 397. https://doi.org/10.14214/sf.397
A stand-establishment service concept with quality guarantee was analysed. Here, the quality of stand establishment was assessed as the density of good quality seedlings evenly distributed on the plantation three years after planting. The amount of adequate premiums for the guarantee service and the risks accumulating to the service provider were studied. Monte Carlo simulation was used as a tool for analyzing the risks accumulating to the service provider of the stand-establishment in operational environments of different sizes. The premiums calculated to cover the expected amount of claims caused by the plantations not meeting the pre-set criteria were about 4–8% in addition to the approximated costs of stand establishment. The criteria used for determining the success or failure in a stand have a marked effect on the amount of premiums with a reasonable risk of ruin.
  • Kiljunen, Finnish Forest Research Institute, Suonenjoki Research Station, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: nuutti.kiljunen@metla.fi (email)
article id 405, category Research article
Timo Saksa. (2004). Regeneration process from seed crop to saplings – a case study in uneven-aged Norway spruce-dominated stands in southern Finland. Silva Fennica vol. 38 no. 4 article id 405. https://doi.org/10.14214/sf.405
The dynamics of spruce regeneration, from seed crop to saplings, was studied based on five permanent plots in uneven-aged, spruce-dominated, boreal forest stands, cut with single-tree selection in the beginning of the 1990’s. The annual fluctuation of the spruce seed crop was very similar in uneven-aged and even-aged stands. The correlation between seed crop and number of germinants was significant; but stem number, basal area or volume of the stand did not influence on seedling emergence. The effects of good seed crops were seen as peaks or an increase in the number of germinants and smallest seedlings. The mean number of ‘stabilised’ spruce seedlings (height 11 cm to 130 cm) varied from 6000 ha–1 to over 25 000 spruce seedlings ha–1 from one monitoring plot to another. On a monitoring plot the number of ‘stabilised’ spruce seedlings was stable over time. Neither stand basal area nor stand volume influenced the number of ‘stabilised’ spruce seedlings, but the height of these seedlings was higher on subplots with lower stand volume and smaller basal area. In this study the monitoring period, 5–10 years, was too short to obtain reliable figures for ingrowth, i.e. the transition of seedlings to the sapling stage (h > 130 cm). The adjusted mean ingrowth was 26 stems ha–1 year–1.
  • Saksa, Finnish Forest Research Institute, Suonenjoki Research Station, FI-77600 Suonenjoki, Finland ORCID ID:E-mail: timo.saksa@metla.fi (email)
article id 422, category Research article
Mika Nieminen. (2004). Export of dissolved organic carbon, nitrogen and phosphorus following clear-cutting of three Norway spruce forests growing on drained peatlands in southern Finland. Silva Fennica vol. 38 no. 2 article id 422. https://doi.org/10.14214/sf.422
The effect of clear-cutting on the concentrations of dissolved organic carbon (DOC), organic nitrogen (DON), NH4+, NO3–, and P in outflow water from three productive, Norway spruce dominated drained peatlands (RCC, VCC-1, VCC-2) were studied. Changes in runoff and transport loads (concentration x runoff) at two of the catchments during the frost-free period are also presented. Approximately 40% of the area was cut at RCC and VCC-2, and 72% at VCC-1. The volume removed was 250 m3 ha–1 at RCC, 259 m3 ha–1 at VCC-1, and for VCC-2, 317 m3 ha–1. The mean annual increase in outflow concentrations of DOC during the first four years after clear-cutting was 9.0 mg l–1 at RCC, 22.8 mg l–1 at VCC-1 and 8.4 mg l–1 at VCC-2. Corresponding increases in the forms of nitrogen were: 0.23, 0.51 and 0.16 mg DON l–1; 0.06, 0.31 and 0.04 mg NH4+-N l–1; and 0.05, 0.12 and 0.22 mg NO3–-N l–1. Clear-cutting did not significantly (p > 0.05) increase P concentrations. The increase in non-frost season runoff over the first three years after clear-cutting was 107 mm at RCC and 207 mm at VCC-1. The export loads of DOC during the non-frost season increased by 80 kg ha–1 at RCC and by 184 kg ha–1 at VCC-1 over the first three years. Corresponding increases for the other studied solutes were: 1.78 and 3.98 kg DON ha–1; 0.39 and 1.49 kg NH4+-N ha–1; 0.45 and 0.48 kg NO3–-N ha–1, and 0.09 and 0.06 kg P ha–1. The study demonstrated that clear-cutting may significantly increase the export of DOC and different forms of nitrogen from drained productive peatlands while only small increases in phosphorus export may occur.
  • Nieminen, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa, Finland ORCID ID:E-mail: mika.nieminen@metla.fi (email)
article id 518, category Research article
Mats Hannerz, Curt Almqvist, Roland Hörnfeldt. (2002). Timing of seed dispersal in Pinus sylvestris stands in central Sweden. Silva Fennica vol. 36 no. 4 article id 518. https://doi.org/10.14214/sf.518
The objective of the study presented here was to describe the variation in timing of seed dispersal from Scots pine (Pinus sylvestris L.) seed trees in central Sweden. Seeds were collected in traps at two sites, for three years at one, and four years at the other . The traps were emptied from March to August each year at 1–2 week intervals during the main period of seed dispersal. The annual seed fall varied between 200 000 and 1.6 million seeds per hectare. The seed fall started in mid to late April, shortly after the heat sum had started to accumulate. The most intensive seed fall took place in early to mid May. The peak period, when 50% of the total seed dispersal occurred, lasted for 18–28 days at the different sites and years. The variation in timing among years seemed to be mainly due to climatic factors – high temperatures promoted seed dispersal, for instance. The results may be useful for planning the time of scarification to optimise the natural regeneration of Scots pine. The data suggest that scarification in the spring, no later than mid-May, would generally create a good seed-bed for most of the current year’s seeds, whereas scarification in late May or June would bury a large proportion of this cohort.
  • Hannerz, The Forestry Research Institute of Sweden (SkogForsk) Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail: mats.hannerz@skogforsk.se (email)
  • Almqvist, The Forestry Research Institute of Sweden (SkogForsk) Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID ID:E-mail:
  • Hörnfeldt, Swedish University of Agricultural Sciences, Dept. of Forest Products and Markets, P.O. Box 7060, SE-750 07 Uppsala, Sweden ORCID ID:E-mail:
article id 564, category Research article
Brett G. Purdy, S. Ellen Macdonald, Mark R. T. Dale. (2002). The regeneration niche of white spruce following fire in the mixedwood boreal forest. Silva Fennica vol. 36 no. 1 article id 564. https://doi.org/10.14214/sf.564
Early establishment of white spruce (Picea glauca) in mixedwood boreal forest stands following fire was examined at several times-since-fire (1-, 2-, 4-, 6-, 14-years). Abiotic and biotic conditions in the stands were assessed at two scales, tree plot (5 m x 5 m) and microsite (1 m x 1 m), along with presence, density and height of white spruce seedlings. Germination and survival of seed sown 1- and 4-years post fire were quantified. Survival and growth of nursery-grown seedlings, and mycorrhizal colonization, survival and growth of sterile seedlings, planted 1-year post-fire were assessed. At the tree plot scale, presence of white spruce seedlings 1-year post-fire could be reliably predicted by organic layer depth and distance to and strength of seed source. In contrast, none of the biotic or abiotic factors measured was strongly correlated with occurrence or density of white spruce seedlings 6- and 14- years post-fire. At the microsite scale, seedling recruitment immediately post-fire was limited to a distinct subset of available microsites (greater % cover of downed wood and moss, lower % cover of litter and herbaceous vegetation). Likewise, seedling occurrence in older burns was associated with distinct microsite conditions; although this was more likely an effect of white spruce presence, rather than the cause. Less than 3% of seed sown 1 yr post-fire survived to become 1yr old germinants, survival was 41% over the next three years. Availability of suitable regeneration microsites declines rapidly with time-since-fire; less than 0.3% of seed sown 4 yrs post-fire survived one year. High rates of mycorrhizal colonization were found on white spruce seedlings planted 1-year post-fire, including early and late stage fungal species. Microsite characteristics were only weakly correlated with mycorrhizal fungal communities. We propose that immediate post-fire recruitment of white spruce is a key process in mixedwood boreal succession.
  • Purdy, Dept. of Renewable Resources, University of Alberta, Edmonton, AB, Canada T6G 2E3 ORCID ID:E-mail:
  • Macdonald, Dept. of Renewable Resources, University of Alberta, Edmonton, AB, Canada T6G 2E3 ORCID ID:E-mail: ellen.macdonald@ualberta.ca (email)
  • Dale, Dale, Dept. of Biological Sciences, University of Alberta, Edmonton, AB, Canada T6G 2E9 ORCID ID:E-mail:
article id 556, category Research article
Timo Kuuluvainen, Juha Mäki, Leena Karjalainen, Hannu Lehtonen. (2002). Tree age distributions in old-growth forest sites in Vienansalo wilderness, eastern Fennoscandia. Silva Fennica vol. 36 no. 1 article id 556. https://doi.org/10.14214/sf.556
The age and size of trees was sampled and measured on eight sample plots (0.2 ha each) within a Pinus sylvestris -dominated boreal forest landscape in Vienansalo wilderness, Russian Karelia. The fire history of these plots was obtained from a previous dendrochronological study. All the studied sample plots showed a wide and uneven distribution of tree ages, but the shape of the age distributions of trees as well as tree species composition varied substantially. Trees over 250 years of age occurred in every studied plot, despite its small size. This suggests that old Pinus were common and rather evenly distributed in the landscape matrix. The oldest Pinus tree was 525 years of age. The correlations between tree age and size were often weak or even nil. In Pinus the correlation between age and diameter was stronger than that between age and height. In the dominant tree species Pinus and Picea, the largest trees were not the oldest trees. The tree age distributions together with the fire history data indicated that the past fires have not been stand replacing, as many of the older Pinus had survived even several fires. Tree age classes that had regenerated after the last fire were most abundant and dominated by Picea and/or deciduous trees, while the trees established before the last fire were almost exclusively Pinus. The results suggest that periodic occurrence of fire is important for the maintenance of the Pinus-dominated landscape. This is because fire kills most Picea and deciduous trees and at the same time enhances conditions for Pinus regeneration, facilitated by available seed from the continuous presence of old fire-tolerant Pinus trees.
  • Kuuluvainen, Department of Forest Ecology, University of Helsinki, P.O. Box 24, FIN-00014 University of Helsinki, Finland ORCID ID:E-mail: timo.kuuluvainen@helsinki.fi (email)
  • Mäki, Department of Forest Ecology, University of Helsinki, P.O. Box 24, FIN-00014 University of Helsinki, Finland ORCID ID:E-mail:
  • Karjalainen, Department of Forest Ecology, University of Helsinki, P.O. Box 24, FIN-00014 University of Helsinki, Finland ORCID ID:E-mail:
  • Lehtonen, Faculty of Forestry, University of Joensuu, P.O. Box 24, FIN-80101 Joensuu, Finland ORCID ID:E-mail:

Category: Research note

article id 1136, category Research note
Håkan Lideskog, Back Tomas Ersson, Urban Bergsten, Magnus Karlberg. (2014). Determining boreal clearcut object properties and characteristics for identification purposes. Silva Fennica vol. 48 no. 3 article id 1136. https://doi.org/10.14214/sf.1136
Highlights: We define the quantitative properties and qualitative characteristics of stumps, stones, slash, and roots, the most important objects interacting with machine activities after clearcutting; We develop a flowchart showing how a computer-aided system using clearcut object identification should be executed.
After clearcutting, machines traffic the clearcut conducting different silvicultural activities. Many objects on a forest clearcut (slash residues, stones, stumps and roots) may disturb e.g. site preparation and planting. This paper describes properties and characteristics of these objects. A flowchart was developed that describes a possible computer-aided system that identifies the objects, and ultimately, makes a machine avoid or target them. A system for obstacle identification creates conditions for further technical development and (semi)automation of e.g., site preparation, mechanized planting, and stump removal.
  • Lideskog, Division of Product and Production Development, Luleå University of Technology, SE-97187 Luleå, Sweden ORCID ID:E-mail: hakan.lideskog@ltu.se (email)
  • Ersson, Department of Forest Biomaterials and Technology, SLU, SE-90183 Umeå, Sweden ORCID ID:E-mail: back.tomas.ersson@slu.se
  • Bergsten, Department of Forest Biomaterials and Technology, SLU, SE-90183 Umeå, Sweden ORCID ID:E-mail: urban.bergsten@slu.se
  • Karlberg, Division of Product and Production Development, Luleå University of Technology, SE-97187 Luleå, Sweden ORCID ID:E-mail: magnus.karlberg@ltu.se
article id 1126, category Research note
Rebecka Mc Carthy, Per Magnus Ekö, Lars Rytter. (2014). Reliability of stump sprouting as a regeneration method for poplars: clonal behavior in survival, sprout straightness and growth. Silva Fennica vol. 48 no. 3 article id 1126. https://doi.org/10.14214/sf.1126
Highlights: Clonal differences were found in stump sprout survival; Clonal differences occurred in sprout straightness; It is uncertain if the ability to produce a high number of stump sprouts is beneficial.
The interest in poplars (Populus spp.) has increased globally as a result of the current incentive to expand the use of biomass for energy. So far, knowledge about poplar stump sprouting is generally poor. The objective of this study was to investigate poplar stump sprouting as regeneration method in harvested plantations. A 19-year-old poplar clonal test, including 23 different clones, on former arable land was harvested in the winter. After one year, stumps were assessed for sprout survival, sprout straightness, number of sprouts, sprout height and basal diameter. Differences between clones were found for survival and sprout straightness. Clonal differences were also found in mean size of sprouts (basal diameter and height). However, no differences were found between clones when comparing size of the tallest sprout or number of sprouts produced on stumps. This study indicates that stump sprouts from various poplar clones differ in survival, sprout straightness and mean growth, which should be taken into account when planning for the next tree generation. The study needs to be complemented with other studies to better assess sprouting characteristics for different clones.
  • Mc Carthy, Forestry Research Institute of Sweden, Skogforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden & Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden ORCID ID:E-mail: rebecka.mccarthy@skogforsk.se (email)
  • Ekö, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden ORCID ID:E-mail: Per.Magnus.Eko@slu.se
  • Rytter, Forestry Research Institute of Sweden, Skogforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden ORCID ID:E-mail: lars.rytter@skogforsk.se
article id 1126, category Research note
Rebecka Mc Carthy, Per Magnus Ekö, Lars Rytter. (2014). Reliability of stump sprouting as a regeneration method for poplars: clonal behavior in survival, sprout straightness and growth. Silva Fennica vol. 48 no. 3 article id 1126. https://doi.org/10.14214/sf.1126
Highlights: Clonal differences were found in stump sprout survival; Clonal differences occurred in sprout straightness; It is uncertain if the ability to produce a high number of stump sprouts is beneficial.
The interest in poplars (Populus spp.) has increased globally as a result of the current incentive to expand the use of biomass for energy. So far, knowledge about poplar stump sprouting is generally poor. The objective of this study was to investigate poplar stump sprouting as regeneration method in harvested plantations. A 19-year-old poplar clonal test, including 23 different clones, on former arable land was harvested in the winter. After one year, stumps were assessed for sprout survival, sprout straightness, number of sprouts, sprout height and basal diameter. Differences between clones were found for survival and sprout straightness. Clonal differences were also found in mean size of sprouts (basal diameter and height). However, no differences were found between clones when comparing size of the tallest sprout or number of sprouts produced on stumps. This study indicates that stump sprouts from various poplar clones differ in survival, sprout straightness and mean growth, which should be taken into account when planning for the next tree generation. The study needs to be complemented with other studies to better assess sprouting characteristics for different clones.
  • Mc Carthy, Forestry Research Institute of Sweden, Skogforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden & Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden ORCID ID:E-mail: rebecka.mccarthy@skogforsk.se (email)
  • Ekö, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden ORCID ID:E-mail: Per.Magnus.Eko@slu.se
  • Rytter, Forestry Research Institute of Sweden, Skogforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden ORCID ID:E-mail: lars.rytter@skogforsk.se

Category: Article

article id 7181, category Article
Eino Oinonen. (1967). Keltalieon (Lycopodium complanatum L.) itiöllinen uudistuminen Etelä-Suomessa kloonien laajuutta ja ikää koskevan tutkimuksen valossa. Acta Forestalia Fennica vol. 83 no. 3 article id 7181. https://doi.org/10.14214/aff.7181
English title: Sporal regeneration of ground pine (Lycopodium complanatum L.) in Southern Finland in the light of the dimensions and the age of its clones.

In an earlier paper of the author it was established that sporal regeneration of bracken fern (Pteridium aquilinum (L.) Kuhn.) is almost entirely connected with fire, and that the size of the bracken fern clones, distinguished by their characteristics, was related to the time of the fire. In previous studies has been found that also sporal regeneration of ground pine (Lycopodium complanatum L., now Diphasiastrum complanatum) is rare. The occurrence of several plant species were studied in relation to bracken fern in the earlier investigation. This paper reports findings concerning ground pine.

When the size-age problem of bracken fern was solved, parallel measurements of ground pine stand on the same site led to the solution of the size-age problem and sporal regeneration of ground pine. The linkage is also valid when the size of ground pine stands was compared to the dates of fires. The ground pine stands are very long-lasting. The stands are fragmented more easily than bracken fern by environmental factors, such as fires, and tend to form large patch clusters with time. Large individual stands reveal the rarity of sporal regeneration of ground pine. The resemblance with bracken fern clones indicate a common factor of regeneration, fire, and a very even spreading rate. Though considerable variation of the colour and structure of ground pine was observed, the circular stands were identical patch by patch.

The PDF includes a summary in English.

  • Oinonen, ORCID ID:E-mail:
article id 7181, category Article
Eino Oinonen. (1967). Keltalieon (Lycopodium complanatum L.) itiöllinen uudistuminen Etelä-Suomessa kloonien laajuutta ja ikää koskevan tutkimuksen valossa. Acta Forestalia Fennica vol. 83 no. 3 article id 7181. https://doi.org/10.14214/aff.7181
English title: Sporal regeneration of ground pine (Lycopodium complanatum L.) in Southern Finland in the light of the dimensions and the age of its clones.

In an earlier paper of the author it was established that sporal regeneration of bracken fern (Pteridium aquilinum (L.) Kuhn.) is almost entirely connected with fire, and that the size of the bracken fern clones, distinguished by their characteristics, was related to the time of the fire. In previous studies has been found that also sporal regeneration of ground pine (Lycopodium complanatum L., now Diphasiastrum complanatum) is rare. The occurrence of several plant species were studied in relation to bracken fern in the earlier investigation. This paper reports findings concerning ground pine.

When the size-age problem of bracken fern was solved, parallel measurements of ground pine stand on the same site led to the solution of the size-age problem and sporal regeneration of ground pine. The linkage is also valid when the size of ground pine stands was compared to the dates of fires. The ground pine stands are very long-lasting. The stands are fragmented more easily than bracken fern by environmental factors, such as fires, and tend to form large patch clusters with time. Large individual stands reveal the rarity of sporal regeneration of ground pine. The resemblance with bracken fern clones indicate a common factor of regeneration, fire, and a very even spreading rate. Though considerable variation of the colour and structure of ground pine was observed, the circular stands were identical patch by patch.

The PDF includes a summary in English.

  • Oinonen, ORCID ID:E-mail:
article id 7180, category Article
Eino Oinonen. (1967). The correlation between the size of Finnish bracken (Pteridium aquilinum (L.) Kuhn.) clones and certain periods of site history. Acta Forestalia Fennica vol. 83 no. 2 article id 7180. https://doi.org/10.14214/aff.7180

In an earlier paper of the author it was established that bracken fern (Pteridium aquilinum (L.) Kuhn.) that the sporal regeneration of the species is almost entirely connected with fire, and that the size of the bracken fern clones, distinguished by their characteristics, was related to the time of the fire. The time of the fire was determined by the samples taken with auger from the trees on the sample plots. A large data on the occurrence of bracken fern was collected around Finland, but only part of it could be used in the previous study, as the trees of the site provided no means to find out it the site had been burned earlier.

In earlier studies, a mass occurrence of sporelings of bracken fern in England has been connected to cities bombed and burned during the World War II. In this study the data of the clones is connected to historical sources and the time of wars in Finland. The comparisons in this study have confirmed the results achieved by comparing the dimensions of the bracken clones to the fire dates. It was found that the wars have caused an increase in the regeneration frequency of bracken fern.

  • Oinonen, ORCID ID:E-mail:
article id 7179, category Article
Eino Oinonen. (1967). Sporal regeneration of bracken (Pteridium aquilinum (L.) Kuhn.) in Finland in the light of the dimensions and the age of its clones. Acta Forestalia Fennica vol. 83 no. 1 article id 7179. https://doi.org/10.14214/aff.7179

The objective of the present study was to obtain information on the regeneration by spores of bracken fern (Pteridium aquilinum (L.) Kuhn.) as well as the permanence and vegetative spreading of the clones. Prevention of bracken fern, which can hinder forest regeneration, is possible only when the basic facts of the life cycle is known. It would be useful to know if it is probable that bracken fern can become prevalent by spore regeneration, and to what extent it invades forest land by vegetative spreading.

No bracken fern sporelings were found, and the experiments of regeneration by spores were unsuccessful, therefore, the studies were concentrated on the clones. Due to the fire tolerance of the plant, it was expected that the clone material would show gradation according to the periods of fire. The first part of the work concerns individual variation in bracken, the second part concentrates on the data collected from sample plots, which latest fire could be dated from the trees.

The results of the studied clones confirm the correlation between the stands interpreted as clones and the length of the post-fire period. The stands were delimited to clones by their individual characteristics. The spreading on average sites (Vaccinium and Myrtillus type forests) was in average 35.8±2 cm annually. It is concluded that bracken fern regeneration by spores is very rare in Finland and that it has been rarer during the last 50 years compared to the preceding period. This is probably caused by the end of slash-and-burn cultivation and decrease of forest fires in Finland. Bracken fern clones often survive the fires through their rhizomes, and fragments of old clones originating before the fire could be found in the study areas. These clones may be even 300 years old. Especially in its northern areas of distribution, bracken fern is connected with settlement and old slash and burn sites. Regeneration by spores in the northern distribution areas seem to be less than in the southern areas.

  • Oinonen, ORCID ID:E-mail:
article id 7178, category Article
Matti Keltikangas, Pekka Tiililä. (1968). Koivun ja kuusen istutuksen keskinäinen edullisuusjärjestys käenkaali-mustikkatyypin metsämailla. Acta Forestalia Fennica vol. 82 no. 5 article id 7178. https://doi.org/10.14214/aff.7178
English title: The economic sequence of silver birch (Betula pendula) and Norway spruce (Picea abies) when planting Oxalis-Myrtillus type forest land.

The present study proposes to calculate the economic sequence of two of Finland’s three main tree species, Norway spruce (Picea abies (L.) H. Karst.) and silver birch (Betula pendula Roth) when planted on Oxalias-Myrtillus type sites where both species are equally suitable, on biological grounds. In addition, the accuracy and applicability of the present Finnish yield tables to an economic comparison is tested. Benefit/cost ratio was selected as criterion of profitableness. All future net incomes and costs were discounted into the planting time and added together. The ratio between the discounted net revenues and the discounted investment costs (later called profit ratio) was the criterion. There is no reliable method to forecast the future wood prices, therefore two price ratios, birch veneer timber to spruce pulpwood and birch cordwood to spruce pulpwood, were chosen as free variables. The economic sequence of the tree species was determined as the function of these variables.

The main conclusions are, first, that under the present price ratios spruce appears to be the better choice for the forest owner, and the most promising policy for changing the situation seems to decrease the production costs of plants in birch nurseries. Second, the present Finnish yield tables are not consistent or accurate enough to enable any sufficiently reliable economic comparisons of tree species in artificial regeneration. The possible error of difference between two rather uncertain estimates is big. More work is needed to construct a uniform system of yield tables covering all main tree species, all site types, all macro climate conditions and all types of regeneration.

The PDF includes a summary in English.

  • Keltikangas, ORCID ID:E-mail:
  • Tiililä, ORCID ID:E-mail:
article id 7135, category Article
Paavo Yli-Vakkuri. (1961). Kokeellisia tutkimuksia taimien syntymisestä ja ensi kehityksestä kuusikoissa ja männiköissä. Acta Forestalia Fennica vol. 75 no. 1 article id 7135. https://doi.org/10.14214/aff.7135
English title: Experimental studies on the emergence and development of tree seedlings in spruce and pine stands.

This paper aims at studying regeneration of Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) by sowing and natural regeneration of birch (Betula sp.) in Western Finland.

Germination of spruce and pine seeds may be prevented by dryness and temperatures below the optimum for germination. In natural conditions, when temperature and moisture is insufficient for germination, the type of seedbed generally has en effect on germination result. Trenching of the seeding spots showed that root competition during the early stage of regeneration was not of decissive importance. It seemed to, however, improve the preservation of the seedlings later. It is common that it can take long before the seeds germinate, and during that time the number of viable seeds decrease strongly.

Also, the seedling stock quickly began to decrease in number after germination, especially during the first growing season and the following winter. The decrease was larger in intact vegetation than on mineral soil or in the humus layer. The emerging seedlings were destroyed by drought very easily, but their tolerance to drought improved later on.

The PDF includes a summary in English.

  • Yli-Vakkuri, ORCID ID:E-mail:
article id 7130, category Article
Paavo Yli-Vakkuri. (1961). Tutkimuksia männyn kylvöalojen metsittymisvaiheesta. Acta Forestalia Fennica vol. 74 no. 3 article id 7130. https://doi.org/10.14214/aff.7130
English title: Studies on the development of young sown pine stands in Central Finland.
Original keywords: uudistaminen; kylvö; mänty; taimikko

In this paper the development of sown Scots pine (Pinus sylvestris L.) seedling stands into forests is studied. The material was collected in stands sown in 1930–1940 in private forests in the Forestry Board districts of Central and Northern Ostrobothnia. The 119 areas, including both burned and other areas, were studied in 1955.

Most seedings had been carried out on relatively poor soils, mostly representing Vaccinium and Calluna type forests. 71% of the areas consisted of large forest fires, mostly from 1933. The most burned areas did not have seed producing trees nearby. The other sown areas were in general small, 1–2 ha, and near forests capable of producing seeds. The species of previous tree generation, in the older areas mostly pine and in the younger areas Norway spruce, affected tree species composition of the new tree generation.

Over 90% of the burned areas were in silviculturally good or satisfactory condition, while the main part of the other sown stands was in fair or poor condition. Weeding and thinning had been done only in the oldest stands. Most stands had been left untended. Natural new trees often competed with the sown pines, and cull-trees and border forest increased natural regeneration in the areas. In Calluna type the poor soil limited regeneration and growth of broadleaf trees. The worst competitors were naturally regenerated pine seedlings both on Calluna and Vaccinium type. On Vaccinium type also birch and sometimes also aspen (Populus tremula L.) competed with sown pine. On better sites and paludified areas competition by broadleaf trees increased. The rhythm of development of broadleaved trees is so different from pine that only those broadleaved trees that are formed in the stand when the pine seedlings are larger can develop harmoniously with pine. Due to the harmful competition, the seedling stands should be tended early on. In addition, it may be advisable to abandon the practise to leave trees on sowing areas.

The PDF includes a summary in Finnish.

  • Yli-Vakkuri, ORCID ID:E-mail:
article id 7129, category Article
Erkki K. Kalela. (1961). Über die natürliche Bewaldung der Kulturböden im sog. Porkkala-Pachtgebiet. Acta Forestalia Fennica vol. 74 no. 2 article id 7129. https://doi.org/10.14214/aff.7129
English title: Natural regeneration of agricultural lands in so-called Porkkala-area of leased land.

The size of the field and the ditching on it as well as the condition of the field at the time of the surrender of Porkkala area (from Finland to Russia) play an important role on how far the natural regeneration of the fields has progressed.

Larger open fields have naturally regenerated only on sides where the surrounding forest can spread the seeds or the thicket of saplings has reached, whereas small parcels of fields have normally been fully forested. Most important species have been e.g. silver birch and pubescent birch, grey and common alders and European aspen as well as pine and spruce. The broad leaved species are dominant.     

The PDF contains a summary in Finnish.

 

  • Kalela, ORCID ID:E-mail:
article id 7128, category Article
Paavo Yli-Vakkuri. (1961). Emergence and initial development of tree seedlings on burnt-over forest land. Acta Forestalia Fennica vol. 74 no. 1 article id 7128. https://doi.org/10.14214/aff.7128

Prescribed burning has been used in regeneration areas in Finland as a method to treat the humus layer and creating more favourable chemical, physical and biological conditions for the seedlings. At the same time, fire clears away seedlings and shoots of unwanted trees and other vegetation. Direct sowing or planting, mostly Scots pine (Pinus sylvestris L.), seldom natural regeneration, is used. In this paper, the initial stages of the formation of a new tree generation of Scots pine and Norway spruce (Picea abies (L.) Karst.) on prescribed burned areas is studied in Central Finland in 1956–1960.

The burned area remains almost without vegetation for about two growing seasons. Conditions on a burned area which has not been tiled are very unfavourable for germination of seeds of coniferous and deciduous trees. On the other hand, shoots of deciduous trees occur soon after burning. Conditions for regeneration were found to be better 3–5 years after burning. Removal of humus layer in spots improved regeneration. However, the patches facilitated also natural regeneration of Norway spruce and especially birch (Betula sp.), which compete with Scots pine seedlings.

Continuous rainy periods improved the germination of Scots pine and Norway spruce seeds sown on the humus layer. Pine and spruce developed more rapidly on the exposed soil, however, young seedlings were easily destroyed. Seed eaters destroyed the pine and spruce seeds sown on the humus layer of newly burned areas completely or almost completely. The viability of pine seeds sown on the burned humus layer did not decrease for three weeks, but the viability greatly weakened after six or more weeks. Spruce seeds lost their viability faster than pine seeds.

The PDF includes a summary in Finnish.

  • Yli-Vakkuri, ORCID ID:E-mail:
article id 7478, category Article
Paavo Yli-Vakkuri. (1958). Tutkimuksia ojitettujen turvemaiden kulotuksesta. Acta Forestalia Fennica vol. 67 no. 4 article id 7478. https://doi.org/10.14214/aff.7478
English title: Studies on prescribed burning of drained peatlands.

Prescribed burning has been used to treat the mineral soil sites, but the method has been little used in drained peatlands. The course and methods of prescribed burning in drained peatlands, and the effect of burning on sprouting of broadleaved trees, growth of ground vegetation and regeneration of Scots pine (Pinus sylvestris L.) by sowing was studied in drained pine bogs in Southern Finland. The top layer of the peat was mostly Sphagnum peat. The material included a prescribed burned 12 ha drained peatland area in Tuomarniemi district, in addition to which ten previously burned areas were investigated.

The burning had succeeded mostly well, but also unsuccessfully burned sites were observed. Estinguishing of the fire was easy, and no peat fires occurred. The fire burned only the logging residue, ground vegetation and the dry top layer of the peat. The roots of brushwood and grasses survived in the peat that insulated the top layer from the heat. For instance, the abundance of cloudberry (Rubus chamaemorus L.) increased after the fire. Similarly, burning did not affect sprouting of the stumps of downy birch (Betula pubescens Ehrh.). It cannot thus be used as a method to restrict the growth of coppice in regenerated areas. The seeds of Scots pine germinated well on the burned surface. 46% of the seeds developed to seedlings on sphagnum-shrub vegetation and 16% in feathermoss-shrub vegetation.

The PDF includes a summary in German.

  • Yli-Vakkuri, ORCID ID:E-mail:
article id 7478, category Article
Paavo Yli-Vakkuri. (1958). Tutkimuksia ojitettujen turvemaiden kulotuksesta. Acta Forestalia Fennica vol. 67 no. 4 article id 7478. https://doi.org/10.14214/aff.7478
English title: Studies on prescribed burning of drained peatlands.

Prescribed burning has been used to treat the mineral soil sites, but the method has been little used in drained peatlands. The course and methods of prescribed burning in drained peatlands, and the effect of burning on sprouting of broadleaved trees, growth of ground vegetation and regeneration of Scots pine (Pinus sylvestris L.) by sowing was studied in drained pine bogs in Southern Finland. The top layer of the peat was mostly Sphagnum peat. The material included a prescribed burned 12 ha drained peatland area in Tuomarniemi district, in addition to which ten previously burned areas were investigated.

The burning had succeeded mostly well, but also unsuccessfully burned sites were observed. Estinguishing of the fire was easy, and no peat fires occurred. The fire burned only the logging residue, ground vegetation and the dry top layer of the peat. The roots of brushwood and grasses survived in the peat that insulated the top layer from the heat. For instance, the abundance of cloudberry (Rubus chamaemorus L.) increased after the fire. Similarly, burning did not affect sprouting of the stumps of downy birch (Betula pubescens Ehrh.). It cannot thus be used as a method to restrict the growth of coppice in regenerated areas. The seeds of Scots pine germinated well on the burned surface. 46% of the seeds developed to seedlings on sphagnum-shrub vegetation and 16% in feathermoss-shrub vegetation.

The PDF includes a summary in German.

  • Yli-Vakkuri, ORCID ID:E-mail:
article id 7472, category Article
Jaakko Lehto. (1956). Tutkimuksia männyn luontaisesta uudistumisesta Etelä-Suomen kangasmailla. Acta Forestalia Fennica vol. 66 no. 2 article id 7472. https://doi.org/10.14214/aff.7472
English title: Studies on the natural regeneration of Scots pine on the upland soils of Southern Finland.

Natural regeneration of Scots pine (Pinus sylvestris L.) by leaving a seed tree stand on a cutting area has long been the most popular regeneration method in Finland. Results of the method have, however, been unsatisfactory. The aim of the investigation was to study the basic problems of natural regeneration of Scots pine. Regeneration success was studied in 144 sample plots in pine stands at different stages of regeneration in Southern Finland. In addition, the data included information of 42 previously investigated areas.

According to the results, Scots pine can be successfully regenerated naturally on sandy and gravelly soils in Southern Finland. Preparing the ground surface by breaking or burning considerably facilitates the establishment of a seedling stand. The number of seedlings was considerably lower in the ground vegetation than in the mineral soil. Considering growth of the seedlings, root competition of the mother trees was heavy in dense stands, but insignificant in thin stands. The stand density did not affect germination of the seeds. In regeneration areas proper, where the density of mother trees usually is under 50 per hectare, there was in average 4,700 seedlings per hectare in Calluna type forests and 5,200 in Vaccinium type forests.

The PDF includes a summary in English.

  • Lehto, ORCID ID:E-mail:
article id 7472, category Article
Jaakko Lehto. (1956). Tutkimuksia männyn luontaisesta uudistumisesta Etelä-Suomen kangasmailla. Acta Forestalia Fennica vol. 66 no. 2 article id 7472. https://doi.org/10.14214/aff.7472
English title: Studies on the natural regeneration of Scots pine on the upland soils of Southern Finland.

Natural regeneration of Scots pine (Pinus sylvestris L.) by leaving a seed tree stand on a cutting area has long been the most popular regeneration method in Finland. Results of the method have, however, been unsatisfactory. The aim of the investigation was to study the basic problems of natural regeneration of Scots pine. Regeneration success was studied in 144 sample plots in pine stands at different stages of regeneration in Southern Finland. In addition, the data included information of 42 previously investigated areas.

According to the results, Scots pine can be successfully regenerated naturally on sandy and gravelly soils in Southern Finland. Preparing the ground surface by breaking or burning considerably facilitates the establishment of a seedling stand. The number of seedlings was considerably lower in the ground vegetation than in the mineral soil. Considering growth of the seedlings, root competition of the mother trees was heavy in dense stands, but insignificant in thin stands. The stand density did not affect germination of the seeds. In regeneration areas proper, where the density of mother trees usually is under 50 per hectare, there was in average 4,700 seedlings per hectare in Calluna type forests and 5,200 in Vaccinium type forests.

The PDF includes a summary in English.

  • Lehto, ORCID ID:E-mail:
article id 7453, category Article
Gustaf Sirén. (1955). The development of spruce forest on raw humus sites in Northern Finland and its ecology. Acta Forestalia Fennica vol. 62 no. 4 article id 7453. https://doi.org/10.14214/aff.7453

According to the National Forest Inventory of Finland, the age structure of forests in Northern Finland require large-scale fellings. One of the problems is regeneration of low-productive Norway spruce (Picea abies (L.) Karst.) forests in the area. The objectives of this investigation were to study the natural development of Hylocomnium-Myrtillus (HMT) type Norway spruce forests in Northern Finland and the ecological causes of the development. A total of 83 sample plots were studied in 1950-52. The stands were chosen to have a total effective temperature during growing season between 600-800 ºC.

According to the results, the HMT type forests represent secondary developmental stages of the northern Myrtillus type, characterized by tree stands of poor quality. By returning the habitats that have reached their secondary stages to their primary stage, e.g. by means of fires, the potential site quality is restored. The climax theory is inapplicable to the Northern Finland’s spruce stands on fresh site types. The natural development of HMT follows not the climax theory, i.e. a return in each succession to the same type stage expressing the same site quality. One reason is accumulation of the thick humus layer, caused by incomplete decomposition of litter. The humus binds nutrients inaccessible to the trees, and the substratum becomes cold, more acid, and moist.

These forests should be managed by clear-cutting, burning-over and artificial regeneration. When thinning the stands, at least a slight mixture of deciduous trees must be left in the stand.

The article is divided into two separate PDFs. The second PDF is in a supplementary file and includes a summary in Finnish.

  • Sirén, ORCID ID:E-mail:
article id 7440, category Article
Erkki K. Kalela. (1954). Mäntysiemenpuiden ja -puustojen juurisuhteista. Acta Forestalia Fennica vol. 61 no. 28 article id 7440. https://doi.org/10.14214/aff.7440
English title: Root systems of Scots pine seed trees and stands.

Root systems of a Scots pine (Pinus sylvestris L.) stands of seed trees on a Vaccinium sites in Southern Finland were studied by taking soil samples around the seed trees. The results show that root system of an old Scots pine spreads relatively evenly around the tree up to at least 10 meters from the stem. The densest part of the root system is near the stem, which part is often acentric. This is probably due to root competition in the early stages of growth of the tree.

Root systems of the seed trees affect stocking of the site with seedlings and the growth of the seedlings. The root competition can cause, for instance, uneven grouping of the seedlings. It seems that the largest trees of a stand have the most even root system. It is therefore recommended to choose the strongest trees of the stand as seed trees, to ensure even distribution of seedlings.

The Acta Forestalia Fennica issue 61 was published in honour of professor Eino Saari’s 60th birthday.

The PDF includes a summary in German.

  • Kalela, ORCID ID:E-mail:
article id 7440, category Article
Erkki K. Kalela. (1954). Mäntysiemenpuiden ja -puustojen juurisuhteista. Acta Forestalia Fennica vol. 61 no. 28 article id 7440. https://doi.org/10.14214/aff.7440
English title: Root systems of Scots pine seed trees and stands.

Root systems of a Scots pine (Pinus sylvestris L.) stands of seed trees on a Vaccinium sites in Southern Finland were studied by taking soil samples around the seed trees. The results show that root system of an old Scots pine spreads relatively evenly around the tree up to at least 10 meters from the stem. The densest part of the root system is near the stem, which part is often acentric. This is probably due to root competition in the early stages of growth of the tree.

Root systems of the seed trees affect stocking of the site with seedlings and the growth of the seedlings. The root competition can cause, for instance, uneven grouping of the seedlings. It seems that the largest trees of a stand have the most even root system. It is therefore recommended to choose the strongest trees of the stand as seed trees, to ensure even distribution of seedlings.

The Acta Forestalia Fennica issue 61 was published in honour of professor Eino Saari’s 60th birthday.

The PDF includes a summary in German.

  • Kalela, ORCID ID:E-mail:
article id 7439, category Article
Leo Heikurainen. (1954). Rämemänniköiden uudistamisesta paljaaksihakkausta käyttäen. Acta Forestalia Fennica vol. 61 no. 27 article id 7439. https://doi.org/10.14214/aff.7439
English title: Regeneration of Scots pine stands of pine swamps through clear cutting.

Pine swamps are easily regenerated by natural regeneration of Scots pine (Pinus sylvestris L.). Usually seeding felling is used, but also strip system or clear cutting and regeneration along stand edge has been suggested. This article discusses the regeneration by clear cutting and sparing the existing undergrowth. The article focuses on pine swamps to be drained and the ones in natural state.

Pine swamps in natural state usually have plenty of trees of smaller diameter classes, that can be trusted to form the future tree generation after the felling. This shortens the rotation by 20-30 years. The undergrowth has been shown to recover quickly. The method suits for regeneration of drained peatlands but could fit also for regeneration of pine swamps in natural state.

The seedlings in the pine swamps are mainly 1-5 years old, and the stock is changing. It seems that larger trees produce a wider selection of age groups, but the seedlings survive longer under smaller mother trees. Part of the younger generations of seedlings seem to be destroyed when the peatland is drained. Further studies are needed to investigate how the draining and felling are to be performed to spare the young seedlings.

The Acta Forestalia Fennica issue 61 was published in honour of professor Eino Saari’s 60th birthday.

The PDF includes a summary in German.

  • Heikurainen, ORCID ID:E-mail:
article id 7439, category Article
Leo Heikurainen. (1954). Rämemänniköiden uudistamisesta paljaaksihakkausta käyttäen. Acta Forestalia Fennica vol. 61 no. 27 article id 7439. https://doi.org/10.14214/aff.7439
English title: Regeneration of Scots pine stands of pine swamps through clear cutting.

Pine swamps are easily regenerated by natural regeneration of Scots pine (Pinus sylvestris L.). Usually seeding felling is used, but also strip system or clear cutting and regeneration along stand edge has been suggested. This article discusses the regeneration by clear cutting and sparing the existing undergrowth. The article focuses on pine swamps to be drained and the ones in natural state.

Pine swamps in natural state usually have plenty of trees of smaller diameter classes, that can be trusted to form the future tree generation after the felling. This shortens the rotation by 20-30 years. The undergrowth has been shown to recover quickly. The method suits for regeneration of drained peatlands but could fit also for regeneration of pine swamps in natural state.

The seedlings in the pine swamps are mainly 1-5 years old, and the stock is changing. It seems that larger trees produce a wider selection of age groups, but the seedlings survive longer under smaller mother trees. Part of the younger generations of seedlings seem to be destroyed when the peatland is drained. Further studies are needed to investigate how the draining and felling are to be performed to spare the young seedlings.

The Acta Forestalia Fennica issue 61 was published in honour of professor Eino Saari’s 60th birthday.

The PDF includes a summary in German.

  • Heikurainen, ORCID ID:E-mail:
article id 7402, category Article
Gustaf Sirén. (1950). Alikasvoskuusten biologiaa. Acta Forestalia Fennica vol. 58 no. 2 article id 7402. https://doi.org/10.14214/aff.7402
English title: On the biology of undergrown Norway spruce.

Norway spruce (Picea abies (L.) Karst.) invading sites is common in Finland. The species tends to establish itself as undergrowth, and takes over when it gets space to grow. To determine whether the undergrowth is suitable as the new generation requires knowledge on the biology of spruce undergrowth. One of the issues is determining the age of the stunted trees. In this investigation, 100 undergrown spruce trees, their crown and their root systems were studied. A method was developed to determine the age of the trees.

The root system of all trees in Vaccinium sites and of stunted trees in Myrtillius sites were superficial. The root systems of older spruces were purely of adventitious origin. The longer the period of stunting growth, the younger is the root system. In addition to acropetal and general adventitious ramification there is often adventitious branching of the roots of pathological causes. Mortality among the long roots is frequent.

A stunted tree has not the same ability as a viable tree to make use of already existing branches for building assimilating surface. When comparing trees with equally large assimilating surface, a stunted tree had greater sum of roots compared to a viable tree. The root system of a stunted undergrown spruce was very superficial compared with the other trees.

The PDF includes a summary in English.

  • Sirén, ORCID ID:E-mail:
article id 7402, category Article
Gustaf Sirén. (1950). Alikasvoskuusten biologiaa. Acta Forestalia Fennica vol. 58 no. 2 article id 7402. https://doi.org/10.14214/aff.7402
English title: On the biology of undergrown Norway spruce.

Norway spruce (Picea abies (L.) Karst.) invading sites is common in Finland. The species tends to establish itself as undergrowth, and takes over when it gets space to grow. To determine whether the undergrowth is suitable as the new generation requires knowledge on the biology of spruce undergrowth. One of the issues is determining the age of the stunted trees. In this investigation, 100 undergrown spruce trees, their crown and their root systems were studied. A method was developed to determine the age of the trees.

The root system of all trees in Vaccinium sites and of stunted trees in Myrtillius sites were superficial. The root systems of older spruces were purely of adventitious origin. The longer the period of stunting growth, the younger is the root system. In addition to acropetal and general adventitious ramification there is often adventitious branching of the roots of pathological causes. Mortality among the long roots is frequent.

A stunted tree has not the same ability as a viable tree to make use of already existing branches for building assimilating surface. When comparing trees with equally large assimilating surface, a stunted tree had greater sum of roots compared to a viable tree. The root system of a stunted undergrown spruce was very superficial compared with the other trees.

The PDF includes a summary in English.

  • Sirén, ORCID ID:E-mail:
article id 7377, category Article
S. E. Multamäki. (1942). Kuusen taimien paleltuminen ja sen vaikutus ojitettujen soiden metsittymiseen. Acta Forestalia Fennica vol. 51 no. 1 article id 7377. https://doi.org/10.14214/aff.7377
English title: Frost injuries of Norway spruce seedlings and their effect on afforestation of drained peatlands .

The aim of the investigation was to study natural regeneration of Norway spruce (Picea abies (L.) Karst.) in drained peatlands and frost injuries in seedlings, and to compare microclimates of the regeneration areas. The experiments included peatlands in Satakunta in Western Finland. Restocking of the areas with seedlings and their survival was followed in 1935-40 at sample plots that were mainly 1 are large.

Susceptibility to freezing was shown to be dependent on the stage of development of the shoots. Shoots that have just begun to grow contain little water, and withstand better freezing temperatures than shoots in later stages of growth. Damages to the seedlings were observed when the temperatures decreased to -2.8–-4.3 °C. The most severe damage to a seedling was caused by the death of the leading shoot by spring frost.

Norway spruce regenerates easily on moist peatlands, but peatlands with dry surface tend to have little or no seedlings. The species regenerated better in marshy sites than correspondingly fertile mineral soil sites. However, it needs shelter to avoid frost damage. On clear cut spruce swamp the undergrowth spruce seedlings that were left in the site got severe frost damage. If the site had birch (Betula sp.) coppice or undergrowth, spruce seedlings survived in their shelter depending on the height and density of the birch trees. To be effective, the protective forest should have relatively even crown cover. Young spruce seedlings could grow well even under relatively dense birch stand.

The PDF includes a summary in German.

  • Multamäki, ORCID ID:E-mail:
article id 7356, category Article
Peitsa Mikola. (1942). Koivun vesomisesta ja sen metsänhoidollisesta merkityksestä. Acta Forestalia Fennica vol. 50 no. 3 article id 7356. https://doi.org/10.14214/aff.7356
English title: Silvicultural usefulness of sprouting of birch.

Birches’ (Betula sp.) ability to grow sprouts is low. The stump grows root collar and stump shoots, but the stump shoots are not proper stump shoots that will grow from the space between wood and bark. The buds are situated very low in the base, even under the ground. In this study, no actual root shoots could be found. Also the bushy alpine birches seem to be formed from stump and root collar shoots.

In Southern Finland silver birch (Betula pendula Roth) is more common than downy birch (Betula pubescens Ehrh.) in dry upland forest sites, while downy birch is common in fresh mineral soil forests and peatlands. In Northern Finland downy birch is the dominant birch species. Of the two species downy birch has markedly better capacity to form stump and root collar shoots both in Northern and Southern Finland. In general, birches grow sprouts much more strongly in Northern Finland.

Growth of the shoots is fastest during the first year after the felling of the parent tree and slows down gradually. The stump shoots may get separated from the stump when the stump decays, and the decay may also spread to the shoots. It is common that the shoots have no own roots, and die along with the stump. The shoots may have own root system or use roots of the parent tree that have stayed alive, in the latter case decay spreads almost always from the stump to the shoot. Whether the tree was felled with axe or saw had no effect on sprouting, probably because the sprouting buds are situated in the base of the tree. The larger stumps had usually fewer sprouts than smaller stumps. The fertility of the site seemed to have little effect on sprouting, but more moist sites formed more sprouts.

Forest regeneration using sprouts may be possible in peatlands for firewood production. on mineral soil sites birch does not suit for coppicing. The proportion of trees originating from sprouts decreases strongly by the time. Consequently, in Southern Finland sprouts have little effect on regeneration of birch. In Northern Finland sprouting is the most important way of regeneration.

The PDF includes a summary in German.

  • Mikola, ORCID ID:E-mail:
article id 7351, category Article
Esko Kangas. (1940). Tuloksia Pohjankankaan ja Hämeenkankaan metsänviljelyksistä. Acta Forestalia Fennica vol. 49 no. 4 article id 7351. https://doi.org/10.14214/aff.7351
English title: Studies on artificial regeneration in Pohjankangas and Hämeenkangas in Southern Finland.

The regeneration of forests in Hämeenkangas area in Southern Finland has been difficult due to various damages from the middle of the 1800s. Few seed trees were left in the area, and artificial regeneration has been used since 1880s. The area became an experimental area of the Forest Research Institute in 1924. The aim of the study was to survey the area before it was transferred to the Finnish Defense Forces.

The original Scots pine (Pinus sylvestris L.) forest of the esker area suffered from many forest fires. The total area is 13,000-14,000 ha, of which the experimental forests of Forest Research Institute cover 6,000 ha. The area is dry upland forest, and drought affects the survival of germlings. Soil frost is a major cause of loss of young seedlings. Sowing method affects the early development of the seedlings. Band sowing proved to be the best method regarding the soil frost. A total of 39 different harmful insect species, 8 pathogen species and 7 other causes of damages have been detected in the area.

The development of seedling stands follow a certain pattern, reported also in other studies. Many of the pine seedling stands develop well until they reach a certain height. After that seedlings begin to suffer from damages, but after reaching another stage develop normally. The damages affect the height growth of the seedlings. Some common damages are caused by Pissoides weevils, needle damages caused by certain beetles, shoot damages by Evetria resinella, and pine blister rust (Peridermium pini and Cronartium flaccidum).

The PDF includes a summary in German
  • Kangas, ORCID ID:E-mail:
article id 7300, category Article
Olli Heikinheimo. (1934). Metsänviljelysmenetelmiä koskevista tutkimuksista. Acta Forestalia Fennica vol. 40 no. 23 article id 7300. https://doi.org/10.14214/aff.7300
English title: Research on forest regeneration methods.

The article is a review on the research about forest regeneration in Finland, executed in the Forest Research Institute. The climate affects the results of different regeneration methods. Thus, sample plots have been established in different parts of the country. In 1933 there was a total of 386 sample plots around the country. To compare the effect of weather conditions in the regeneration, sample plots have been established in successive years. Other conditions that affect forest regeneration are soil, forest site type, tree species, time from the felling of the stand, and tending of the seedling stand. Of the 386 sample plots 245 were planting experiments and 141 sowing experiments.

The PDF includes a summary in German.

  • Heikinheimo, ORCID ID:E-mail:
article id 7294, category Article
Martti Hertz. (1934). Tutkimuksia karjan vaikutuksesta hakkausalojen kasvillisuuteen. Acta Forestalia Fennica vol. 40 no. 17 article id 7294. https://doi.org/10.14214/aff.7294
English title: Studies on influence of grazing on the vegetation of cutting areas.

The aim of the study was to determine the effect of grazing of the cutting areas to the ground vegetation and regeneration of spruce. The cutting areas could be divided into two kinds of areas based on the vegetation. Hillocks were drier and poorer than other parts of the cutting areas. Their vegetation did suffer less from grazing than the other parts of the cutting areas. The shade-loving plant species decreased, but as the poorer sites have less edible plants, cattle caused less damage than in the better sites. The even spaces between the hillocks had both positive and negative changes. Cattle transport seeds, tile and fertilize the soil, promote paludification, and decrease competition by the primary species like large grasses. This is beneficial to new species. Grazing is directed to large grass species like Calamagrostis. Those species that cattle reject, become more abundant. Stamping damages especially shallow rooted species and perennial species, like Norway spruce (Picea abies (L.) H. Karst.). Larger tree seedlings may get injuries in the stem.

The PDF includes a summary in German.

  • Hertz, ORCID ID:E-mail:
article id 7256, category Article
V. Pöntynen. (1929). Tutkimuksia kuusen esiintymisestä alikasvoksina Raja-Karjalan valtionmailla. Acta Forestalia Fennica vol. 35 no. 1 article id 7256. https://doi.org/10.14214/aff.7256
English title: Studies on Norway spruce undergrowth in state forests in Karelia.

The abundance of Norway spruce (Picae abies (L.) H. Karst.) undergrowth is common for the state forests in Karelia near the Russian border, in Finland. In the survey, the occurrence of the undergrowth was studied. The article includes a review on the ownership of the forest, forest soils in the area, and the state of forests in the area. Scots pine (Pinus sylvestris L.) is the dominative species in 67%, Norway spruce in 27% and Betula sp. 6% of the state forests. Only 2% of the forests are 1‒20 years of age. Stands in the age group of 61‒80 years are the most common (25%). Norway spruce undergrowth is most abundant in the municipality of Salmi. The forests are typically moist forest site types or grass-herb site types. If the stands are allowed to develop naturally, even the Vaccinium sites become Norway spruce dominated. Spruce undergrowth is formed seldom under a spruce forest unless the stand is thin or has openings. Because Norway spruce is often rare in the mineral soil sites, the undergrowth is often regenerated from seeds that spread from spruce swamps. Earlier practiced shifting cultivation and its frequent fires prevented regeneration of spruce undergrowth. Similarly, the common felling method used, clear felling in strips, does not promote spruce undergrowth. Consequently, their occurrence is likely to decrease in the future.

The PDF includes a summary in German.

  • Pöntynen, ORCID ID:E-mail:
article id 7232, category Article
Edvard Wibeck. (1929). Erwägungen bei Wahl von Kulturmethoden in der schwedischen Nadelwaldwirtschaft. Acta Forestalia Fennica vol. 34 no. 19 article id 7232. https://doi.org/10.14214/aff.7232
English title: Discussions on the choice of cultivation method in coniferous forests in Sweden.

In Sweden lot of state owned forests or earlier mining districts or districts of ironworks have been afforested during the last decade. The amount of afforested areas sinks from south to north. Afforestation has taken place also in privately owned forests.

The article discusses the common economic questions related to afforestation work and the biological viewpoints related to it. The best cultivation methods are presented for several common forest types, such as herb-rich forest types, moss-grown forests types, swampy forests with heavy raw humus and barren pine forest sites (lichen type).

The volume 34 of Acta Forestalia Fennica is a jubileum publication of professor Aimo Kaarlo Cajander.

  • Wibeck, ORCID ID:E-mail:
article id 7228, category Article
Agnar Barth. (1929). Skjermforyngelsen i produksjonsökonomisk belysning. Acta Forestalia Fennica vol. 34 no. 15 article id 7228. https://doi.org/10.14214/aff.7228
English title: Natural regeneration from the production economic point of view.

Natural regeneration has been studied so far mostly on the perspective of regeneration, while the production capacity of the seed crop stand has been of little interest in the earlier studies. This paper studied volume growth of the seed trees and the economic impact of this regeneration method both in a literature review and measuring sample stands consisting of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) dominated stands and mixed forests in Norway.

It is concluded that in the most cases the seed tree stands give better pulpwood and timber yield than the stands in the average. The reason might be that the elite stems can better use their production capacity, and, thus, reach bigger and more valuable dimensions. Even though the growing stock diminishes in the seeding felling, the growth of the stand does not decrease or decreases only little. At the same time, the quality growth improves. In addition, the new tree regeneration is achieved usually without extra costs, and there is no unproductive time period in the stand.

  • Barth, ORCID ID:E-mail:
article id 7227, category Article
D. Fehér. (1929). Die Biologie des Waldbodens und ihre physiologische Bedeutung im Leben des Waldes. Acta Forestalia Fennica vol. 34 no. 14 article id 7227. https://doi.org/10.14214/aff.7227
English title: Biology of forest soil and its physiological meaning in the life of forest.

The article presents three studies about different aspects of the bio-chemical functions of forest soil. The three studies are i) research on microflora and microfauna of forest soils; ii) study on carbon nutrition of forests in relation to microbial functions of soil and effecting site factors and iii) study on nitrogen metabolism of forest soil. The results of the studies are summarized by every study.

The article discusses the meaning of the results for forest management in practice. The good biological and physiological condition of forest soil is important for the forest growth and it needs to be taken care in regeneration and other forest management. The natural regeneration seems to be better for soil functions. Favoring broadleaved trees as undergrowth enhances the biological processes of forest soil.   

The volume 34 of Acta Forestalia Fennica is a jubileum publication of professor Aimo Kaarlo Cajander.

  • Fehér, ORCID ID:E-mail:
article id 5573, category Article
Ilkka Vanha-Majamaa, Raili Suominen, Tiina Tonteri, Eeva-Stiina Tuittila. (1996). Seedling establishment after prescribed burning of a clear-cut and a partially cut mesic boreal forest in southern Finland. Silva Fennica vol. 30 no. 1 article id 5573. https://doi.org/10.14214/sf.a9218

The prescribed burning of a 7.3 ha clear-cut and a 1.7 ha partially cut forest (volume 150 m3/ha) was carried out in Evo (61 °12'N, 25°07'E) on 1 June 1992. The forest was a mesic Myrtillus site type forest dominated by Norway spruce (Picea abies (L.) H. Karst.). Practically all the trees and the above-ground parts of the understorey vegetation died in the fire, while the mor layer was thinned by an average of 1.5 cm.

A study was made on the change of germinated seedling population in time and their dependence on environmental factors. Seedlings of Norway spruce, Scots pine (Pinus sylvestris L.), silver birch (Betula pendula Roth), pubescent birch (B. pubescens Ehrh.) and rowan (Sorbus aucuparia L.) were inventoried in 1993 and in 1994 on permanent plots, four times per growing season. Autoregression models were used to compare regeneration of tree species in the burned forest with regeneration in the burnt clear-cut area, and to study the effect of distance from nearest seed source to regeneration.

The average number of seedlings germinating in 1993 was higher than in 1994, probably because of differences between these consecutive years in regard to the amount of seed rain and weather conditions. The number of Norway spruce and rowan seedling was higher inside the forest area than in the clear-cut area. The distance to the bordering forest and to the closest seed tree did not explain the result. It is suggested that the more stable microclimatic conditions under the shade of dead tree promote germination and seedling establishment in the forest area. As rowan is a bird-dispersed species, it is likely that dead trees help the dispersal of rowan seed by providing birds place to sit and defecate. The shade provided by dead trees may influence the further succession of the tree stand and vegetation composition and diversity.

  • Vanha-Majamaa, ORCID ID:E-mail:
  • Suominen, ORCID ID:E-mail:
  • Tonteri, ORCID ID:E-mail:
  • Tuittila, ORCID ID:E-mail:
article id 7195, category Article
Martti Hertz. (1925). Niinipuun uudistumisesta Suomessa. Acta Forestalia Fennica vol. 29 no. 5 article id 7195. https://doi.org/10.14214/aff.7195
English title: Regeneration of small leaved lime in Finland.

The northern range of small leaved lime (Tilia cordata Mill.) in Finland has remained the same since the end of 1800s, but according to the studies of subfossiles of lime found in peatlands, the northern limit has once been higher than at the present. The northernmost natural specimens of the species in Reisjärvi do not produce seed, and are therefore probably a relict. The article includes a review on the distribution of the species in Finland and its capacity to regenerate. The natural regeneration of lime in Finland is at present very rare. The species has lost its ability to sexual reproduction, but reproduces readily vegetatively.

The PDF includes a summary in German.

  • Hertz, ORCID ID:E-mail:
article id 7195, category Article
Martti Hertz. (1925). Niinipuun uudistumisesta Suomessa. Acta Forestalia Fennica vol. 29 no. 5 article id 7195. https://doi.org/10.14214/aff.7195
English title: Regeneration of small leaved lime in Finland.

The northern range of small leaved lime (Tilia cordata Mill.) in Finland has remained the same since the end of 1800s, but according to the studies of subfossiles of lime found in peatlands, the northern limit has once been higher than at the present. The northernmost natural specimens of the species in Reisjärvi do not produce seed, and are therefore probably a relict. The article includes a review on the distribution of the species in Finland and its capacity to regenerate. The natural regeneration of lime in Finland is at present very rare. The species has lost its ability to sexual reproduction, but reproduces readily vegetatively.

The PDF includes a summary in German.

  • Hertz, ORCID ID:E-mail:
article id 5562, category Article
Janne Uuttera, Matti Maltamo. (1995). Impact of regeneration method on stand structure prior to first thinning. Comparative study North Karelia, Finland vs. Republic of Karelia, Russian Federation. Silva Fennica vol. 29 no. 4 article id 5562. https://doi.org/10.14214/sf.a9213

Comparisons were made between artificially and naturally regenerated stands in the south-eastern part of North Karelia, Finland, and naturally regenerated stands in the western parts of the Republic of Karelia, Russian Federation. The effect of soil fertility and silvicultural operations on the stand structure was also investigated.

The results of the study show clearly that when forests are artificially regenerated the stand structure includes less variation when compared with the stands naturally regenerated. Differences between the regeneration methods are clearer the more fertile the forest site is. Within the regeneration method there is also a clear trend in stand structure, with the variation decreasing the poorer the site. The effect of silvicultural operations, i.e. the cleaning of the sapling stand, has disappeared by the time of first thinning, although it appears to have a permanent effect on the dynamics of the tree species within a stand.

The variation of the stand structure can be regarded as an essential factor for the potential biodiversity of the stand also at its young vegetation succession stage. This capacity for maintaining the forest biodiversity, developed at the young vegetation succession stage, becomes increasingly important in subsequent vegetation succession stages. Natural regeneration provides improved possibilities for the operations preserving forest biodiversity, as it generates more dense stands with a wider variation in stand structure, compared to artificial regeneration.

  • Uuttera, ORCID ID:E-mail:
  • Maltamo, ORCID ID:E-mail:
article id 5562, category Article
Janne Uuttera, Matti Maltamo. (1995). Impact of regeneration method on stand structure prior to first thinning. Comparative study North Karelia, Finland vs. Republic of Karelia, Russian Federation. Silva Fennica vol. 29 no. 4 article id 5562. https://doi.org/10.14214/sf.a9213

Comparisons were made between artificially and naturally regenerated stands in the south-eastern part of North Karelia, Finland, and naturally regenerated stands in the western parts of the Republic of Karelia, Russian Federation. The effect of soil fertility and silvicultural operations on the stand structure was also investigated.

The results of the study show clearly that when forests are artificially regenerated the stand structure includes less variation when compared with the stands naturally regenerated. Differences between the regeneration methods are clearer the more fertile the forest site is. Within the regeneration method there is also a clear trend in stand structure, with the variation decreasing the poorer the site. The effect of silvicultural operations, i.e. the cleaning of the sapling stand, has disappeared by the time of first thinning, although it appears to have a permanent effect on the dynamics of the tree species within a stand.

The variation of the stand structure can be regarded as an essential factor for the potential biodiversity of the stand also at its young vegetation succession stage. This capacity for maintaining the forest biodiversity, developed at the young vegetation succession stage, becomes increasingly important in subsequent vegetation succession stages. Natural regeneration provides improved possibilities for the operations preserving forest biodiversity, as it generates more dense stands with a wider variation in stand structure, compared to artificial regeneration.

  • Uuttera, ORCID ID:E-mail:
  • Maltamo, ORCID ID:E-mail:
article id 5562, category Article
Janne Uuttera, Matti Maltamo. (1995). Impact of regeneration method on stand structure prior to first thinning. Comparative study North Karelia, Finland vs. Republic of Karelia, Russian Federation. Silva Fennica vol. 29 no. 4 article id 5562. https://doi.org/10.14214/sf.a9213

Comparisons were made between artificially and naturally regenerated stands in the south-eastern part of North Karelia, Finland, and naturally regenerated stands in the western parts of the Republic of Karelia, Russian Federation. The effect of soil fertility and silvicultural operations on the stand structure was also investigated.

The results of the study show clearly that when forests are artificially regenerated the stand structure includes less variation when compared with the stands naturally regenerated. Differences between the regeneration methods are clearer the more fertile the forest site is. Within the regeneration method there is also a clear trend in stand structure, with the variation decreasing the poorer the site. The effect of silvicultural operations, i.e. the cleaning of the sapling stand, has disappeared by the time of first thinning, although it appears to have a permanent effect on the dynamics of the tree species within a stand.

The variation of the stand structure can be regarded as an essential factor for the potential biodiversity of the stand also at its young vegetation succession stage. This capacity for maintaining the forest biodiversity, developed at the young vegetation succession stage, becomes increasingly important in subsequent vegetation succession stages. Natural regeneration provides improved possibilities for the operations preserving forest biodiversity, as it generates more dense stands with a wider variation in stand structure, compared to artificial regeneration.

  • Uuttera, ORCID ID:E-mail:
  • Maltamo, ORCID ID:E-mail:
article id 5557, category Article
Klaus Silfverberg. (1995). Forest regeneration on nutrient-poor peatlands: Effects of fertilization, mounding and sowing. Silva Fennica vol. 29 no. 3 article id 5557. https://doi.org/10.14214/sf.a9208

The effects of wood ash and PK fertilization on natural regeneration and sowing of Scots pine (Pinus sylvestris L.) were studied in field experiments on nitrogen-poor (Ntot 0.87–1.26%) peat substrates. The study material was derived from three drained, nutrient-poor pine mires (64°52’ N, 25°08’ E) at Muhos, near Oulu, Finland. The experimental fields were laid out in 1985 as a split-split-plot design including the following treatments; mounding, natural regeneration and sowing and fertilization; PK (400 kg ha-1) and wood ash (5,000 kg ha-1). The seedlings were inventoried in circles in July–August 1991.

Changes in the vegetation were small and there were no statistical differences due to the fertilization treatments in the ground vegetation. PK or ash fertilization did not cause vegetation changes harmful to Scots pine regeneration on nitrogen-poor peatlands. Both sowing and fertilization significantly increased the number of pine seedlings, but not their height. Wood ash increased seedling number more than PK fertilizer. The number of seedlings varied from 7,963 (control) to 42,781 ha-1 (mounding + sowing + ash). The seedling number was adequate for successful regeneration even on non-mounded, non-fertilized naturally regenerated plots.

The number of birch seedlings varied more than that of pine (370–25,927 ha-1). Mounding especially increased the number of birches. The difference between PK fertiliser and ash was less pronounced than that for pine. In addition, to the field studies the effects of ash and PK fertilizer on the germination of Scots pine seeds was studied in a greenhouse experiment. Soaking in ash solutions strongly reduced seed germination, while the PK solution was less harmful.

  • Silfverberg, ORCID ID:E-mail:
article id 5552, category Article
Liisa Saarenmaa, Timo Leppälä. (1995). Fill-in seedlings in constituting the stocking of Scots pine stands in northern Finland. Silva Fennica vol. 29 no. 2 article id 5552. https://doi.org/10.14214/sf.a9203

Fill planting is a common procedure following reforestation in Finland. In 1990, 13% of the total of seedlings planted was used for fill planting. The objective of this study is (i) to survey the survival of fill-in seedlings and (ii) to estimate the spatial pattern of stands to evaluate the importance of fill-in seedlings in constituting the stocking of Scots pine (Pinus sylvestris L.) stands in Central and Northern Finland.

A survey of 63 artificially regenerated Scots pine stands was conducted in 1990. Stand densities varied from 950 to 3,925 seedlings/ha. The mean densities of originally planted, fill planted and naturally regenerated seedlings were 863, 639 and 791/ha, respectively. The survival of originally planted seedlings was 36% and that of fill-in seedlings 48%. Death rate of fill-in seedlings of Scots pine increased with longer times between original and fill planting. The survival rate of Norway spruce (Picea abies) seedlings was correlated with temperature sum. Height of the fill-in seedlings was less than that of the originally planted ones. Most stands had an even spatial distribution with the exception of sparsely populated stands, which were somewhat clustered. This indicates that dying of seedlings is not randomly spread. Because of poor survival, fill planting seems to be a risky business in most cases.

  • Saarenmaa, ORCID ID:E-mail:
  • Leppälä, ORCID ID:E-mail:
article id 5445, category Article
Taneli Kolström. (1991). Kuusen kylvö- ja istutuskoe viljavilla kivennäismailla Pohjois-Karjalassa. Silva Fennica vol. 25 no. 2 article id 5445. https://doi.org/10.14214/sf.a15599
English title: Results from the sowing and planting experiment of Norway spruce (Picea abies) on fertile sites in North Karelia, Finland.

Different methods of sowing and planting of Norway spruce (Picea abies (L.) H. Karst.) were compared on fertile sites in North Karelia (62°20’N, 29°35’E, 85–120 m a.s.l.). The planting material were 4-year-old bare-rooted transplants, 2-year-old bare-rooted seedlings, and 2-year-old containerized seedlings raised in plastic greenhouse. The sowing methods were band sowing and shelter sowing. Ground vegetation was controlled during the first growing season mechanically or chemically, or the control was omitted totally.

Planting of spruce gave better results than sowing. After eight growing seasons there were sowed seedlings left in 30% of the sowing pots. The average height of them was 35 cm. Seedling survival was best with large bare-rooted transplants (91%). Survival of containerized seedlings was 79% and of small bare-rooted transplants 71%. The average height of large bare-rooted transplants was 131 cm, of containerized seedlings 86 cm and small bare-rooted seedlings 68 cm.

Sowing is not an advisable method for regeneration of spruce due to the small survival rate and slow initial development when ground vegetation is controlled only once. Also 2-year-old seedlings gave a satisfactory result in regeneration. Seedlings raised in greenhouse were more sensitive to frost damage than seedlings grown on open ground.

The PDF includes an abstract in English.

  • Kolström, ORCID ID:E-mail:
article id 5420, category Article
Toyohiro Miyazava, Jukka Laine. (1990). Effect of macroclimate on the development of Scots pine seedling stands on drained oligotrophic pine mires. Silva Fennica vol. 24 no. 2 article id 5420. https://doi.org/10.14214/sf.a15574

The influence of different fertilization treatments and ditch spacings on the height growth of young Scots pine (Pinus sylvestris L.) seedling stands growing under various climatic regimes were determined. Comparisons were made between naturally regenerated and planted seedling stands. The effective temperature sum had a stronger effect on the height growth of planted seedlings, and in Northern Finland the planted seedlings seemed to be influenced to a greater degree by the adverse climatic conditions. The heavier the dose of fertilizer that had been applied, the greater the difference in growth caused by macroclimate. A considerably larger proportion of natural seedlings were located on hummocks compared with that of planted seedlings, irrespective of the region. On plots with wider ditch spacings, seedlings growing on hummocks were superior in height growth to those on flat surfaces.

The PDF includes an abstract in Finnish.

  • Miyazava, ORCID ID:E-mail:
  • Laine, ORCID ID:E-mail:
article id 5413, category Article
Eljas Pohtila. (1990). Metsän uudistuminen Kivalon vanhoilla kaistalehakkuualoilla. Silva Fennica vol. 24 no. 1 article id 5413. https://doi.org/10.14214/sf.a15565
English title: Forest regeneration of old strip cutting areas in Kivalo.

The results of regenerating 54 and 36–38-year old strip cuttings were surveyed in the Kivalo Research Forest (N 66°20’, E 26°40’). 15 measurement plots were placed on each strip. The most common forest type was Hylocomnium-Myrtillus type. Regeneration of the strips proved to be slow. Most of the spruces growing on the strips probably originated from the time before the cutting. The average number of stems was 1,155 per ha, of which one third consisted of broadleaved trees. The average volume increment of stem wood after cutting had been about 1 m3/ha/yr, but it was increasing at the time of the inventory. Both the reforestation of the strips and the development of the emergent stands were dependent on elevation and site fertility. Site fertility was indicated by the abundance of Vaccinium myrtillus.

The PDF includes an abstract in English.

  • Pohtila, ORCID ID:E-mail:
article id 5412, category Article
Jari Parviainen. (1990). Metsäpuiden paakkutaimituotannon nykynäkymät. Silva Fennica vol. 24 no. 1 article id 5412. https://doi.org/10.14214/sf.a15564
English title: Future trends for containerized tree seedling production: A literature review.

Containerized tree seedlings will be used on an increasing scale in the future in different parts of the world. There are number of techniques for the production of small one-year-old seedlings but it has not been possible to develop a completely satisfactory methods for large containerized seedlings production. In the long-term development of pine plantations established with containerized seedlings the greatest problem has been deformation of the root system. With a new method, based on a sheet of peat and root pruning, it has been possible to produce conifer seedlings with a good root regeneration potential and favourable morphological root system development. The use of small containerized seedlings allows an increase in planting density without any marked increase in regeneration costs.

The PDF includes an abstract in English.

  • Parviainen, ORCID ID:E-mail:
article id 5379, category Article
Timo Kuuluvainen, Timo Pukkala. (1989). Effect of Scots pine seed trees on the density of ground vegetation and tree seedlings. Silva Fennica vol. 23 no. 2 article id 5379. https://doi.org/10.14214/sf.a15536

The study uses the methodology of ecological field theory to model the effect of Scots pine (Pinus sylvestris L.) seed trees on the density of tree seedlings and other plants in the field layer. The seed trees had a clear effect on the expected value of the amount and distribution of the ground vegetation. The vicinity of seed trees had an adverse effect on the growth of grasses, herbs and seedlings, while mosses were most abundant near the trees. Models based on the ecological field approach were derived to describe the effect of seed trees on the ground vegetation.

The PDF includes an abstract in Finnish.

  • Kuuluvainen, ORCID ID:E-mail:
  • Pukkala, ORCID ID:E-mail:
article id 5379, category Article
Timo Kuuluvainen, Timo Pukkala. (1989). Effect of Scots pine seed trees on the density of ground vegetation and tree seedlings. Silva Fennica vol. 23 no. 2 article id 5379. https://doi.org/10.14214/sf.a15536

The study uses the methodology of ecological field theory to model the effect of Scots pine (Pinus sylvestris L.) seed trees on the density of tree seedlings and other plants in the field layer. The seed trees had a clear effect on the expected value of the amount and distribution of the ground vegetation. The vicinity of seed trees had an adverse effect on the growth of grasses, herbs and seedlings, while mosses were most abundant near the trees. Models based on the ecological field approach were derived to describe the effect of seed trees on the ground vegetation.

The PDF includes an abstract in Finnish.

  • Kuuluvainen, ORCID ID:E-mail:
  • Pukkala, ORCID ID:E-mail:
article id 5365, category Article
Pasi Puttonen. (1988). Metsänuudistamistutkimuksen arvioinnin seminaari 28.4.1988. Silva Fennica vol. 22 no. 4 article id 5365. https://doi.org/10.14214/sf.a15522
English title: Presentation report of evaluation group on forest regeneration research in Finland.

The Academy of Finland commissioned a review of forest regeneration research. This paper summarises the evaluation. The report deals with the individual research projects and, in addition, the international evaluation group gives several recommendations to develop the forest regeneration research further. The recommendations concern education and training, publications, peer review, experimental design, routine work and extension service, and technology transfer. The paper also includes comments of the Finnish Forest Institute, the Finnish Society of Forest Science and a research scientist on the results of the evaluation.

The PDF includes a summary in English.

  • Puttonen, ORCID ID:E-mail:
article id 5311, category Article
Timo Pukkala. (1987). Kuusen ja männyn siemensadon ennustemalli. Silva Fennica vol. 21 no. 2 article id 5311. https://doi.org/10.14214/sf.a15468
English title: Model for predicting the seed crop of Picea abies and Pinus sylvestris.

The seed crop of Norway spruce (Picea abies (L.) H. Karst.) and Scots pine (Pinus sylvestris L.) is predicted with the help of mean monthly temperatures during May–August one and two years before the flowering year. The prediction models were made separately for Lapland and for the rest of Finland. The models are based on 10-year periods of seed crop measurements and climatic data. The total number of time series was 59.

In Lapland, Norway spruce flowered abundantly and produced an abundant seed crop after warm July–August and two years after cool July–August. In other parts of Finland, warm June and July produced a good flowering year, especially if these months were cool two years before the flowering year.

In Lapland, Scots pine flowered abundantly if the whole previous growing season was warm. Elsewhere in Finland, a cool June preceded prolific flowering in the coming year if the rest of the growing season was considerably warmer than the average.

The prediction models explained 37–49 % of the variation in the size of the seed crop. The occurrence of good and poor seed years was usually predicted correctly. Using the presented models, the prediction of the seed crop is obtainable 1.5 year for Norway spruce and 2.5 year for Scots pine before the year of seed fall.

The PDF includes an abstract in English.

  • Pukkala, ORCID ID:E-mail:
article id 5302, category Article
Seppo Kellomäki, Heikki Hänninen, Taneli Kolström, Ahti Kotisaari, Timo Pukkala. (1987). A tentative model for describing the effects of some regenerative process on the properties of natural seedling stands. Silva Fennica vol. 21 no. 1 article id 5302. https://doi.org/10.14214/sf.a15459

The effect of the size of seed crop, dispersal of seeds and the early development of seedlings on the density and spatial distribution of young Scots pine (Pinus sylvestris L.) stands are evaluated on the basis of theoretical models. The models include (i) number and spatial distribution of parent trees on the regeneration area, (ii) size of annual seed crop, (iii) seed dispersal from a particular parent tree, (iv) germination of the seeds (germination percentage), (v) death of ageing seedlings after the establishment process, and (vi) height growth of the seedlings.

As expected, stand density and spatial distribution varied within a large range in relation to the density of the parent trees and the distance from them. The simulations also showed that natural seedling stands can be expected to be heterogenous due to the geometry of seed dispersal, emphasizing the frequency of young and small trees. The properties of the seedling stands were, however, greatly dependent on the density of the parent trees and the length of the regeneration period.

The PDF includes an abstract in Finnish.

  • Kellomäki, ORCID ID:E-mail:
  • Hänninen, ORCID ID:E-mail:
  • Kolström, ORCID ID:E-mail:
  • Kotisaari, ORCID ID:E-mail:
  • Pukkala, ORCID ID:E-mail:
article id 5243, category Article
Eljas Pohtila, Tapani Pohjola. (1985). Maan kunnostus männyn viljelyssä Lapissa. Silva Fennica vol. 19 no. 3 article id 5243. https://doi.org/10.14214/sf.a15422
English title: Soil preparation in reforestation of Scots pine in Lapland.

The study deals with the interaction of various soil preparation and reforestation methods. The most favourable time of the year for broadcast sowing and the effect of stabilization after soil preparation on restocking were studied as special problems.

Prescribed burning, scalping and disc ploughing made a better combination with sowing than planting, and ploughing better combination with planting than sowing. The longer the period was between sowing and germination the fewer seedlings emerged. The best stocking was clearly resulted with sowing in June. Stabilization of soil after preparation had a negative effect on reforestation results.

The PDF includes a summary in English.

  • Pohtila, ORCID ID:E-mail:
  • Pohjola, ORCID ID:E-mail:
article id 5227, category Article
Harri Vasander, Tapio Lindholm. (1985). Tulen voimakkuus ja maanpinnan lämpötila kulotuksen aikana. Silva Fennica vol. 19 no. 1 article id 5227. https://doi.org/10.14214/sf.a15406
English title: Fire intensities and surface temperatures during prescribed burning.

Surface temperature during two prescribed burnings were measured in 1983 in Evo, Southern Finland. Surface temperatures in relation to the amount of slash burned, energy released during the fires, and the fire intensities were studied. The fire intensity was also measured during a third burn. The Lake Nimetön site was burned int the end of May. Due to the uneven distribution of slash, colonization by Calamagrostis arundinacea and the spring moisture, the burning was very uneven. Surface temperatures varied between 410–809°C and the intensity of fire was low (range 0–900 kW/m).

The fire intensity on the other sites burned in May was also low (880 kW/m). During the burn in August the surface temperatures varied between 701–869°C and the intensity of fire was moderate (1,170 kW/m). Slash was burned more evenly and more thoroughly due to the dryness of the site and slash and the fact that grasses and other herbs were not abundant.

The PDF includes a summary in Finnish.

  • Vasander, ORCID ID:E-mail:
  • Lindholm, ORCID ID:E-mail:
article id 5218, category Article
Jouni Suoheimo. (1984). Isokorvakärsäkkään aikuisten esiintyminen ja merkitys männyn luontaiselle uudistamiselle Pohjois-Lapissa. Silva Fennica vol. 18 no. 3 article id 5218. https://doi.org/10.14214/sf.a15397
English title: The occurrence of Otiorrhynchus nodosus and its significance for the natural regeneration of Scots pine in Lapland.

The aim of the present study was to survey the occurrence of Otiorrhynchus nodosus Müller weevils and their significance for the natural regeneration of Scots pine (Pinus sylvestris L.). The study was carried out during summer 1982 at Inari in northern Lapland.

There were two sample plots, one situated in a Scots pine seed-tree area and the other, the control sample plot, in an area with a coverage of mountain birch (Betula pubescens subsp. tortuosa, now subsp. czerepanovii). A total of 177 Otiorhynchus weevils were caught. Movement of the weevils reached its climax in July. There were 86% more individuals in the seed-tree area than in the mountain birch area. No damage to the pine germlings or seedlings was not observed, although the situation could be different during the peaks of the veewil populations.

The PDF includes a summary in English.

  • Suoheimo, ORCID ID:E-mail:
article id 5198, category Article
Leo Heikurainen, Jukka Laine, Jarmo Lepola. (1983). Lannoitus- ja sarkaleveyskokeita karujen rämeiden uudistamisessa ja taimikoiden kasvatuksessa. Silva Fennica vol. 17 no. 4 article id 5198. https://doi.org/10.14214/sf.a15181
English title: Fertilization and ditch spacing experiments concerned with regeneration and growth of young Scots pine stands on nutrient poor pine bogs.

The effects of variations in the intensity of drainage and NPK fertilization on the natural regeneration and planting results and the subsequent development of seedling stands under various climatic conditions on drained nutrient poor pine bogs was investigated in a 16-year-old study.

Comparison of height development of Scots pine (Pinus sylvestris L.) stands on drained peatlands to that of pine stands growing in mineral soil sites show that in Southern Finland the most efficient forest improvement measures (10 m ditch spacing and 1,000 kg/ha NPK-fertilization) resulted in growth that corresponds a to a height index of a stand in a Vaccinium type site. Less efficient treatment (30 m ditch spacing and no fertilizer) resulted in growth corresponding the development of young stand in a Calluna type site. In Northern Finland the effect of fertilization on height growth was almost negligible. This is possibly due to a decrease in the nitrogen mobilization from south to north of Finland. Thus, it seems evident that fertilization of young Scots pine stands on nutrient poor drained peatlands can be recommended only in the southern part of the country.

The effect of ditch spacing is same in the whole country. The narrower the spacing the better the height growth. In the south planted stands thrive better than naturally regenerated stands, but the situation is reversed in the north.

The PDF includes a summary in English.

  • Heikurainen, ORCID ID:E-mail:
  • Laine, ORCID ID:E-mail:
  • Lepola, ORCID ID:E-mail:
article id 5198, category Article
Leo Heikurainen, Jukka Laine, Jarmo Lepola. (1983). Lannoitus- ja sarkaleveyskokeita karujen rämeiden uudistamisessa ja taimikoiden kasvatuksessa. Silva Fennica vol. 17 no. 4 article id 5198. https://doi.org/10.14214/sf.a15181
English title: Fertilization and ditch spacing experiments concerned with regeneration and growth of young Scots pine stands on nutrient poor pine bogs.

The effects of variations in the intensity of drainage and NPK fertilization on the natural regeneration and planting results and the subsequent development of seedling stands under various climatic conditions on drained nutrient poor pine bogs was investigated in a 16-year-old study.

Comparison of height development of Scots pine (Pinus sylvestris L.) stands on drained peatlands to that of pine stands growing in mineral soil sites show that in Southern Finland the most efficient forest improvement measures (10 m ditch spacing and 1,000 kg/ha NPK-fertilization) resulted in growth that corresponds a to a height index of a stand in a Vaccinium type site. Less efficient treatment (30 m ditch spacing and no fertilizer) resulted in growth corresponding the development of young stand in a Calluna type site. In Northern Finland the effect of fertilization on height growth was almost negligible. This is possibly due to a decrease in the nitrogen mobilization from south to north of Finland. Thus, it seems evident that fertilization of young Scots pine stands on nutrient poor drained peatlands can be recommended only in the southern part of the country.

The effect of ditch spacing is same in the whole country. The narrower the spacing the better the height growth. In the south planted stands thrive better than naturally regenerated stands, but the situation is reversed in the north.

The PDF includes a summary in English.

  • Heikurainen, ORCID ID:E-mail:
  • Laine, ORCID ID:E-mail:
  • Lepola, ORCID ID:E-mail:
article id 5191, category Article
Juha Lappi. (1983). Metsänuudistamisen vaatiman ajan merkitys uudistamispäätöksissä. Silva Fennica vol. 17 no. 3 article id 5191. https://doi.org/10.14214/sf.a15174
English title: Elevation of the time factor in reforestation decisions.

Length of the regeneration period is a criterion commonly used for comparing different reforestation methods. The time factor should be evaluated using a realistic system for long-term planning. In this paper the preliminary evaluation is made by simplified calculations based on the development series. The slow regeneration method is assumed to be otherwise equal to the rapid one but it has a 5- or 10-years delay at the beginning, and the rotation is thus the final cutting age plus 5- or 10-years delay. Cost of the time delay is taken to be the difference in reforestation costs that makes the rapid and the slow methods equivalent. Calculations are made using zero costs for the slow method; but if the cost of the slow method increases, the critical cost difference decreases very slowly. The final cutting age and the regeneration method must be decided simultaneously. Therefore, the cost of the time delay is presented as a function of final cutting age. By maximizing the average annual revenue, rotation can be even increased if more rapid but more expensive regeneration method is used.

The PDF includes a summary in English.

  • Lappi, ORCID ID:E-mail:
article id 5191, category Article
Juha Lappi. (1983). Metsänuudistamisen vaatiman ajan merkitys uudistamispäätöksissä. Silva Fennica vol. 17 no. 3 article id 5191. https://doi.org/10.14214/sf.a15174
English title: Elevation of the time factor in reforestation decisions.

Length of the regeneration period is a criterion commonly used for comparing different reforestation methods. The time factor should be evaluated using a realistic system for long-term planning. In this paper the preliminary evaluation is made by simplified calculations based on the development series. The slow regeneration method is assumed to be otherwise equal to the rapid one but it has a 5- or 10-years delay at the beginning, and the rotation is thus the final cutting age plus 5- or 10-years delay. Cost of the time delay is taken to be the difference in reforestation costs that makes the rapid and the slow methods equivalent. Calculations are made using zero costs for the slow method; but if the cost of the slow method increases, the critical cost difference decreases very slowly. The final cutting age and the regeneration method must be decided simultaneously. Therefore, the cost of the time delay is presented as a function of final cutting age. By maximizing the average annual revenue, rotation can be even increased if more rapid but more expensive regeneration method is used.

The PDF includes a summary in English.

  • Lappi, ORCID ID:E-mail:
article id 5191, category Article
Juha Lappi. (1983). Metsänuudistamisen vaatiman ajan merkitys uudistamispäätöksissä. Silva Fennica vol. 17 no. 3 article id 5191. https://doi.org/10.14214/sf.a15174
English title: Elevation of the time factor in reforestation decisions.

Length of the regeneration period is a criterion commonly used for comparing different reforestation methods. The time factor should be evaluated using a realistic system for long-term planning. In this paper the preliminary evaluation is made by simplified calculations based on the development series. The slow regeneration method is assumed to be otherwise equal to the rapid one but it has a 5- or 10-years delay at the beginning, and the rotation is thus the final cutting age plus 5- or 10-years delay. Cost of the time delay is taken to be the difference in reforestation costs that makes the rapid and the slow methods equivalent. Calculations are made using zero costs for the slow method; but if the cost of the slow method increases, the critical cost difference decreases very slowly. The final cutting age and the regeneration method must be decided simultaneously. Therefore, the cost of the time delay is presented as a function of final cutting age. By maximizing the average annual revenue, rotation can be even increased if more rapid but more expensive regeneration method is used.

The PDF includes a summary in English.

  • Lappi, ORCID ID:E-mail:
article id 5188, category Article
Eljas Pohtila, Tapani Pohjola. (1983). Vuosina 1970-1972 Lappiin perustetun aurattujen alueiden viljelykokeen tulokset. Silva Fennica vol. 17 no. 3 article id 5188. https://doi.org/10.14214/sf.a15171
English title: Results from the reforestation experiment on ploughed sites established in Finnish Lapland during 1970–1972.

The objective of the study was to compare different reforestation methods on ploughed areas in Finnish Lapland. Four species were compared: Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies (L.) H. Karst.), silver birch (Betula pendula Roth) and Siberian larch (Larix sibirica Ledeb.). The experiments were established in different parts of Lapland on different types of sites in 1970–72.

In Scots pine there was a difference of 15 percentage points in survival of seedlings between the best and worst methods of regeneration. Containerized seedlings and paper pot seedlings had the best survival rates. In Norway spruce the respective difference between sowing and planting was about 20 percentage points. In favour of planting. The survival rate can be increased by about 20 percentage points by selecting the right tree species. The average height varied from 25 cm (the sowed Norway spruce) to 179 cm (the planted silver birch) after 10 growing seasons. The birch was planted at the most fertile sites only. The longer time passed from the afforestation the clearer was the effect of the local growing conditions on the development of the seedlings. The elevation of the site was one factor seemed to influence the success of the seedlings.

The PDF includes a summary in English.

  • Pohtila, ORCID ID:E-mail:
  • Pohjola, ORCID ID:E-mail:
article id 5127, category Article
Simo Kaila, Juhani Päivänen. (1981). Metsämaanmuokkauksen suoritemäärät ja konekalusto vuosina 1976-1979. Silva Fennica vol. 15 no. 3 article id 5127. https://doi.org/10.14214/sf.a15068
English title: Forest soil cultivation areas and machinery in 1976–1979 in Finland.

The areas of soil cultivation in regeneration sites using different methods in 1976–1979 and the cultivation units in use in Finland in 1979 were studied by means of a questionnaire. Application of soil cultivation methods appears to have become strongly polarized: in Southern Finland disc plowing with wheel tractors are used, whilst in Northern Finland wing plowing with crawler tractors is the most popular method, each being practically the only alternative in the respective area.

The PDF includes a summary in English.

  • Kaila, ORCID ID:E-mail:
  • Päivänen, ORCID ID:E-mail:
article id 7036, category Article
I. Lassila. (1920). Tutkimuksia mäntymetsien synnystä ja kehityksestä pohjoisen napapiirin pohjoispuolella. Acta Forestalia Fennica vol. 14 no. 3 article id 7036. https://doi.org/10.14214/aff.7036
English title: Studies on the regeneration and development of Scots pine forests north of the arctic circle.

The frequency of years when Scots pine (Pinus sylvestris L.) produces cones and seeds affects its reproduction in the north. The study area covered most of the pine lands in Northern Finland. Scots pine seems to be able to produce cones relatively often in the north. The amount of seeds produced in one year was, however, not sufficient to produce a dense seedling stand. Thus, the natural pine stands contain usually trees in different ages-classes, which have germinated in different years. The cone production is highest in 150-170 years old trees. Pine also needs warm summers to produce viable seeds. Brush fires avail the growth of seedlings, because they clear of ground vegetation that hinders germination of seeds. The seedlings need also moisture to survive; good regeneration years have often had rainy summers.

The PDF includes a summary in German.

  • Lassila, ORCID ID:E-mail:
article id 7035, category Article
V. T. Aaltonen. (1920). Wasserverbrauch der Bäume und Feuchtigkeitsverhältnisse des Bodens. Acta Forestalia Fennica vol. 14 no. 2 article id 7035. https://doi.org/10.14214/aff.7035
English title: Use of water of the trees and the moisture conditions in the soil.

The data has been collected in the dry heathy forests in Harjavalta commune in south-west Finland and in Sodankylä commune in Lapland in summer time 1918 and 1919. The aim to the study was to find out how does the water consumption of the trees affect the moisture conditions of the soil and how they are linked to the regeneration of the forest.

Trees in one age class have a certain spatial distribution: the greater the distance between trees the older the trees and the smaller the distance, the younger the trees. This seem to rather be due to the development of the root system and the nutrition intake of a tree than the competition for light. The moisture content of the upper soil layers is higher in the open areas than in the closed canopy stands. Hence there are more seedlings growing in open areas.   However, it is not clear whether the results apply for other forest site types as well, and more research is needed.     

  • Aaltonen, ORCID ID:E-mail:
article id 5026, category Article
Alpo Enivaara. (1979). Pyynikin harjumännikön uudistaminen ja elvyttäminen. Silva Fennica vol. 13 no. 2 article id 5026. https://doi.org/10.14214/sf.a14881
English title: Regeneration and improvement of the Scots pine forest of Pyynikki esker in Southern Finland.

Pyynikki is a unique natural park in the centre of the city of Tampere in Southern Finland. Currently, the forest of Pyynikki is losing its vigour because of aging and environmental factors. In 1977 a policy was designed for the regrowth of Scots pine (Pinus sylvestris L.) forest and improvement of the condition of the trees to avoid the situation that the whole pine forest (34 ha) would be regrown at one time. The period of regrowth of pine forest is 60 years. It is accomplished by cutting small openings, on which 4-year old potted pine saplings are planted. Birch will be planted only on two spots. The aim is that the forest scenery will be maintained as undamaged as possible. The condition of the growing forest will be improved by fertilization and thinning.

This paper was presented in the ‘Man and the Biosphere’ programme project 2 seminar held on August 24–25 1978 in Hyytiälä research station of University of Helsinki.

The PDF includes a summary in English.

  • Enivaara, ORCID ID:E-mail:
article id 4984, category Article
Jyrki Raulo. (1978). Forestation chain for birch (Betula pendula Roth) in Finland. Silva Fennica vol. 12 no. 1 article id 4984. https://doi.org/10.14214/sf.a14838

During the 1970’s an average of 4 million Betula pendula (Roth) seedlings have been planted annually in Finland. The activities connected with the planting of this tree species, the selection of forestation sites, site preparation, planting out the seedlings and follow-up work on the forestation sites are briefly reviewed in the article. The manuscript is based on the studies into the breeding, seedling production and planting techniques of B. pendula started by the Finnish Forest Research Institute already in 1960’s, as well as on practical observations made at the planting sites. A list of some of the Finnish studies concerning B. pendula which have been published in English and studies with a summary in English is included.

The PDF includes a summary in Finnish.

  • Raulo, ORCID ID:E-mail:
article id 7024, category Article
August Renvall. (1919). Suojametsäkysymyksestä IV. Poronlaidunnan järjestely suojametsäalueella. Acta Forestalia Fennica vol. 11 no. 4 article id 7024. https://doi.org/10.14214/aff.7024
English title: Protection forests IV.

This fourth part of the six-article series about protection forest in the Northern Finland is a proposal for organizing reindeer grazing to enable scots pine (Pinus sylvestris L.) regeneration near the timber line. Protection forests in the Northern Finland cover third of the natural pastures of reindeer in the area. In these areas reindeer grazing can harm the young pine seedlings. The proposal suggests temporary restrictions in grazing in the coniferous forests. Also, in the northernmost parts of Lapland the pine timberline area would be used only as winter pastures. Regional limits should be set for the number of reindeer. Also the ownership of reindeer herds and herding coopearatives included problematic issues that should be solved.

The article is divided in six parts. A German summary is in a separate PDF.

  • Renvall, ORCID ID:E-mail:
article id 7022, category Article
August Renvall. (1919). Suojametsäkysymyksestä II. Suojametsäjärjestelmän tarkoitusperät ja edellytykset. Acta Forestalia Fennica vol. 11 no. 2-3 article id 7022. https://doi.org/10.14214/aff.7022
English title: Protection forests II.

The second part of the six-article series about protection forest in Northern Finland outlines the principles of protecting Scots pine (Pinus sylvestris L.) timber line. The protected areas should include at least all of the viable pine stands and those birch (Betula sp.) lands that are necessary to safeguard fuel wood supply in the pine forest zone. Outside the actual pine forest zone also those birch stands that are near populated regions should be protected to secure sustainable fuel wood supply. Third, south and down of the pine forest zone should be protected pine forest stands and adjacent open fell areas where forests are especially vulnerable. A new protection forest commission should be established to execute the protection forest legislation.

The article is divided in six parts. The parts II and III of the article series are included in the same PDF. A German summary is in a separate PDF.

  • Renvall, ORCID ID:E-mail:
article id 4962, category Article
Matti Keltikangas, Kustaa Seppälä. (1977). Ojitusalueiden hieskoivikoiden kasvatus taloudellisena vaihtoehtona. Silva Fennica vol. 11 no. 1 article id 4962. https://doi.org/10.14214/sf.a14813
English title: The economics of growing downy birch stands on drained peatlands in Finland.

The aim of this study was to determine under what conditions and with what premises the growing of Betula pubescens Ehrh. is an economically competitive alternative to the growing of Scots pine (Pinus sylvestris L.) in drained peatlands. The basic material consisted of all drainage projects in Ostrobothnia in Western Finland in 1937–38 and 1957–59, according to the archives of the Central Board of Forestry Tapio, including such areas that were at least moderately fertile and had birch dominated young stands or no tree cover. A total of 202 sample plots were measured.

According to the results, the discounted timber yield of the thinned B. pubescens stands is about 10% greater than that of untreated stands. The removing of birch seedlings and the subsequent growing of fully stocked Scots pine is more profitable than growing B. pubescens stands only if the establishment and subsequent development of the pine stand involve no costs. If the site in question is a fertile open drained peatland, establishment of a pine stand is obviously a better financial proposition than a naturally regenerated birch stand. However, if there is already a fully stocked young birch stand on the site, it is more economical to let it grow using a shortish rotation time.

The PDF includes a summary in English.

  • Keltikangas, ORCID ID:E-mail:
  • Seppälä, ORCID ID:E-mail:
article id 7018, category Article
Olli Heikinheimo. (1917). Metsänhävityksen ja polton vaikutuksesta metsämaahan. Acta Forestalia Fennica vol. 8 no. 3 article id 7018. https://doi.org/10.14214/aff.7018
English title: Effects of forest devastation and shifting cultivation on forest soil.
Original keywords: kaskeaminen; metsittyminen; maaperä

Soil of open and forest covered lands have different properties. Forest devastation has negative effect on forest soil, for instance, through leaching of nutrients. There are different views on how forest fire influences chemical properties of the soil. Burning of forests can have both positive and negative consequences.

The article publishes partial results from a study that aimed to find out how shifting cultivation affects the soil, and how it may change natural regeneration and forest growth of the stands. Soil samples were collected from four burnt-over sites. Of the samples, size classes of the soil particles, volatile solids and nutrient contents were determined. According to the results, the burning increased soluble nutrients, but the effect was not long-lasting. However, more sample sites would be needed to give definitive results of the differences.

  • Heikinheimo, ORCID ID:E-mail:
article id 4876, category Article
Tapio Lehtiniemi, Juhani Sarasto. (1973). Kokemuksia rauduksen istutuksesta ojitetuille soille. Silva Fennica vol. 7 no. 1 article id 4876. https://doi.org/10.14214/sf.a14706
English title: Betula verrucosa plantations on peat.

The aim of the present study was to assess whether two-year old Betula verrucosa Ehrh. (now Betula pendula Roth.) transplants can be used in afforestation of drained peatlands and what factors affect the development of the young trees. The seedlings were planted in 1967. The site was repair planted next spring due to mortality caused by a undefined fungal disease, and the plantations were fertilized with NPK fertilizer (soil application. The seedlings were measured twice a year until the autumn 1970.

Only 28% of the original transplants, and 73.4% of the repair plantations were alive in 1970. In some cases, fertilization improved the results, while in others it was detrimental to the trees or had no effect on survival. According to peat analysis, the poor survival and development of the plants could be due to the too high ratios of N/Ca and N/P. Stunted or dead trees displayed often necrosis caused by Godronia multispora. According to the experiences, Betula verrucosa plantations are inferior to those obtained with Scots pine (Pinus sylvestris L.). In addition, the results indicate that in old draining areas calcium and phosphorus are often too low in comparison to nitrogen.

The PDF includes a summary in English.

  • Lehtiniemi, ORCID ID:E-mail:
  • Sarasto, ORCID ID:E-mail:
article id 7013, category Article
Lauri Ilvessalo. (1917). Tutkimuksia mäntymetsien uudistumisvuosista Etelä- ja Keski-Suomessa. Acta Forestalia Fennica vol. 6 no. 2 article id 7013. https://doi.org/10.14214/aff.7013
English title: Studies on the regeneration years of Scots pine forests in Southern and Central Finland.

The success of natural regeneration of Scots pine (Pinus sylvestris L.) is dependent on the amount and quality of seeds produced in each year. While seed production of Scots pine in Northern Finland is well known, there is little information about frequency of good seed years in Southern and Central Finland. The success of regeneration of Scots pine was studied by defining from the growth rings of felled sample trees in which year they started to grow. The aim was to choose sample trees so that they would form a continual series from seedlings to 100 years old trees. In the material there was too little sample trees from certain years. However, within the period of 1827-1910 it could be identified 13 good regeneration years of pine in Southern Finland. According to the study, pine has regenerated very well every 6th year and at least moderately well every 4th year. When compared to the previous studies made in Northern Finland, part of the good regeneration years seems to be same both in Northern and Southern Finland. Regeneration of Scots pine is affected also by the type and condition of the site.

The PDF includes a summary in German.

  • Ilvessalo, ORCID ID:E-mail:
article id 4832, category Article
Paavo Yli-Vakkuri, Pentti K. Räsänen. (1971). Siementen peittämisen ja kylvökohdan polkaisun vaikutus männyn ruutukylvön tulokseen. Silva Fennica vol. 5 no. 1 article id 4832. https://doi.org/10.14214/sf.a14621
English title: The influence of covering and tramping the seeds into the soil on the success of spot sowing of Scots pine.

This paper reports spot sowing experiments of Scots pine (Pinus sylvestris L.). The seeds were either covered with coarse sand, tramped in the substrate or sowed without any covering, 30 seeds in each treatment in 70 replications. The site was of Vaccinium type with sandy soil. The germination percentage was 81 and 91 on the respective years. The development of seedlings was observed for 3–4 years.

The results indicate that both tramping and covering the seeds to some extent increased the number of seedlings and improved the early development. The highest numbers of seedlings were recorded in the first growing season, after which there was 23 seedlings/100 seeds in the uncovered spots, 27 seedlings in the covered spots and 31 seedlings in the tramped spots in the experiment sowed in 1965.

Mortality of the seedlings was highest between the first and second growing season, and empty spots increased with the time. There was no difference in mortality between the sowing methods, but the number of seedlings after first growing season affected the result. Under favourable conditions four seedlings per spot seemed enough to secure the survival of minimum one seedling per spot during the three first growing seasons. In poor conditions seven seedlings was needed.

The PDF includes a summary in English.

  • Yli-Vakkuri, ORCID ID:E-mail:
  • Räsänen, ORCID ID:E-mail:
article id 4784, category Article
Matti Leikola, Pentti Pylkkö. (1969). Verhopuuston tiheyden vaikutus metsikön minimilämpötiloihin hallaöinä. Silva Fennica vol. 3 no. 1 article id 4784. https://doi.org/10.14214/sf.a14570
English title: Influence of stand density on the minimum temperatures during frost nights.

The objective of this investigation was to study the influence of stand density of white birch (Betula pubescens Ehrl.) on the minimum temperatures in the stand during the growing season, and the actual minimum temperatures of the leading shoot of Norway spruce (Picea abies (L.) H. Karst.) seedlings growing in the open. The 40-year-old uniform white birch stand was situated in 142 m above the sea level in Southern Finland. The stand was treated with thinnings of three different densities in 1961.

Air temperature was recorded in four sample plots at heights of 0.1 m, 0.5 m, 1.0 m, 2 m and 4 m. In the stand of moderate density, temperatures were measured at heights of 6.0 m, and in the stand of full density at 6.0 m, 8.0 m and 10.0 m.

The temperature differences between stands of various densities proved to be rather small. Especially the thinnest stand differed very little from the open area. The soil surface has in all cases been warm compared with the higher air layers indicating meadow-fog-type by Geier (1965). On cloudy or windy weather all the temperature profiles in the various stands resembled each other. The difference between the air temperature and temperature of the spruce shoot was greatest at midnight and decreased steadily thereafter.

The problem in using shelter stands for spruce regeneration areas is that optimum shelter stand density is difficult to define. Already a thin shelter stand causes drawbacks to the young seedlings, but in order to be effective enough against early frosts, the shelter stand should be comparatively dense.

The PDF includes a summary in English.

  • Leikola, ORCID ID:E-mail:
  • Pylkkö, ORCID ID:E-mail:
article id 4769, category Article
Paavo Yli-Vakkuri, Aimo Autio, Juhani Vehkaoja. (1968). Päätehakkuun ja metsänviljelyn välinen aika. Silva Fennica vol. 2 no. 3 article id 4769. https://doi.org/10.14214/sf.a14554
English title: Time lag between final cutting and regeneration.

The time interval between final felling and plantation means a waste of time and thus a production loss, and may lead to lush growth of ground cover and hardwood sprouts, which increases expenses in forest management. The objective of this study was to determine the length of time between final felling and artificial regeneration in private forests in the forest districts of Uusimaa-Häme in Southern Finland and Pohjois-Häme in Central Finland. The material consists of a sample of 150 plans of the 952 cutting and regeneration plans in the district of Uusimaa-Häme and a sample of 140 plans of the 1,102 plans in Pohjois-Häme.

The time lag between final cutting and seeding or planting was on average 1.4 years in Pohjois-Häme district and 0.7 years in Uusimaa-Häme. In the latter district, 56% of the logged area was regenerated in the spring immediately following the cutting, and 84% not later than in the second spring. In Pohjois-Häme, 29% of the harvested area was regenerated immediately in the first spring following cutting, and 79% not later than in the third spring following cutting.

In Pohjois-Häme, the interval was shortest in the smallest forest holdings, and longest in the largest holdings with the largest regeneration areas. The length seems to depend mainly on the size of the regeneration area. In Uusimaa-Häme district, the interval was shortest in the smallest holdings, rather short in the largest, and longest in the intermediate-size forest properties. Seeding with Scots pine (Pinus sylvestris L.) was used in almost all regeneration areas.

The forest owners had mainly carried out the regeneration work themselves. In the Pohjois-Häme area, the interval was shorter when the district forestry board regenerated the area. 35% of the regenerated areas had required supplementary planting in Pohjois-Häme, and 47% in the Uusimaa-Häme area. Supplementary planting was more common in areas regenerated later after the cutting. In Pohjois-Häme, according to the reports of the forest owners, 75% of the regenerated areas required tending during the first three years, in Uusimaa-Häme, 80%.

The PDF includes a summary in English.

  • Yli-Vakkuri, ORCID ID:E-mail:
  • Autio, ORCID ID:E-mail:
  • Vehkaoja, ORCID ID:E-mail:
article id 7506, category Article
Eero Kubin, Lauri Kemppainen. (1994). Effect of soil preparation of boreal spruce forest on air and soil temperature conditions in forest regeneration areas. Acta Forestalia Fennica no. 244 article id 7506. https://doi.org/10.14214/aff.7506

The effect of scarification, ploughing and cross-directional ploughing on temperature conditions in the soil and adjacent air layer have been studied during 11 growth periods by using an unprepared clear-cut area as a control site. The development of seedling stand was followed to determine its shading effect on the soil surface.

Soil preparation decreased the daily temperature amplitude of the air at the height of 10 cm. The maximum temperatures on sunny days were lower in the tilts of the ploughed and in the humps of the cross-directional ploughed sites compared with the unprepared area. Correspondingly, the night temperatures were higher and so the soil preparation reduced the risk of night frost. In the soil at the depth of 5 cm, soil preparation increased daytime temperatures and reduced night temperatures compared with unprepared area. The maximum increase in monthly mean temperatures was almost 5°C, and the daily variation in the surface parts of the tilts and humps increased so that excessively high temperatures for the optimal growth of the root systems were measured from time to time. The temperature also rose at the depths of 50 and 100 cm.

Soil preparation also increased the cumulative temperature sum. The highest sums accumulated during the summer months were recorded at the depth of 5 cm in the humps of cross-directional ploughed area (1,127 dd.) and in the tilts of the ploughed area (1,106 dd.), while the corresponding figure in the unprepared soil was 718 dd. At the height of 10 cm the highest temperature sum was 1,020 dd. in the hump, and 925 dd. in the unprepared area.

The incidence of high temperature amplitudes and frequency of high temperatures at the depth of 5 cm decreased most rapidly in the humps of cross-sectional ploughed area and the ploughing tilts towards the end of the study period. The decrease was attributed principally to the compressing of tilts, the ground vegetation succession and the growth of seedlings. The difference between the prepared and unprepared area did not diminish. The increase in temperature due to soil preparation, thus, lasted at least over 10 years.

  • Kubin, ORCID ID:E-mail:
  • Kemppainen, ORCID ID:E-mail:
article id 7656, category Article
Kari Leinonen, Matti Leikola, Antti Peltonen, Pentti Räsä­nen. (1989). Kuusen luontainen uudistaminen Pirkka-­Hämeen metsälautakunnassa. Acta Forestalia Fennica no. 209 article id 7656. https://doi.org/10.14214/aff.7656
English title: Natural regeneration of Norway spruce in Pirkka-Häme Forestry Board District, Southern Finland.

The aim of the present study was to evaluate and develop the use of natural regeneration of Norway spruce (Picea abies (L.) H. Karst.) in private forestry. The study was carried out using a line-plot survey with permanent circular sample plots. In total 40 regeneration sites were measured. The study includes results from three successive inventories: prior to the shelterwood cutting, in the summer after the cutting, and one year after the cutting. Regression and logistic regression analyses were used to construct models describing the effect of various factors on the restocking of the stands.

The standing volume prior to the shelterwood cutting was on average 236 m3/ha (ranging from 80 to 428 m3/ha) and after the cutting 120 m3/ha (39–220 m3/ha). The average number of stems per hectare decreased from 435 to 186. Prior to the shelterwood cutting 22% of the stands were satisfactorily restocked. After the cutting and one year later these percentages were 6 and 29%, respectively. Prior to the shelterwood cutting the number of acceptable seedlings was 1,440/ha, in the summer and year later 1,308/ha and 1,546/ha, respectively. Prior to the shelterwood cutting the characteristics of the mother stands did not correlate well with the number of seedlings. The change in the number of seedlings during the initial stage of shelterwood method depended on height of the seedling stand, amount of logging waste and number of germlings prior to the cutting. The risk to fail in regeneration was highest in the poorly restocked, sparse shelterwood stands, where a fast expansion of grass vegetation took place.

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  • Leinonen, ORCID ID:E-mail:
  • Leikola, ORCID ID:E-mail:
  • Peltonen, ORCID ID:E-mail:
  • Räsä­nen, ORCID ID:E-mail:
article id 7625, category Article
Antti Turakka, Olavi Luukkanen, Suree Bhumibhamon. (1982). Notes on Pinus kesiya and P. merkusii and their natural regeneration in watershed areas of northern Thailand. Acta Forestalia Fennica no. 178 article id 7625. https://doi.org/10.14214/aff.7625

The aim of the present work was to clarify the structure of selected natural Pinus kesiya Royale ex Gord. And P. merkusii Jungh. et de Vriese stands, with particular attention on the amount and quality of naturally-occurring seedlings and young trees. Furthermore, basic information required in the management of pine stands in watershed areas of northern Thailand was obtained.

Large number of pine seedlings were found in P. kesiyana stands only. Two of the examined four localities where this species occurred also included various intermediary height classes under the mature trees. In dense stands the number of seedlings was generally smaller as compared with lower seed tree densities. Neither one of the two localities where P. Merkusii was investigated indicated sufficient natural regeneration. Young pines (over 5 m height) seemed to survive the frequent ground fires quite well, whereas younger seedlings were destroyed and the ground layer vigour was lowered in these cases.

In situ sowing experiments at the beginning of the dry season indicated a faster development and better survival of emerging P. Kesiya seedlings as compared to P. Merkusii. Soil preparation and exposure to sun decreased the survival of seedlings in both species. Utilization of natural regeneration and direct sowing of pines as a silvicultural method as well as the general significance of the two pine species in the succession of plant communities under the influence of forest fires is also discussed.

The PDF includes a summary in Finnish.

  • Turakka, ORCID ID:E-mail:
  • Luukkanen, ORCID ID:E-mail:
  • Bhumibhamon, ORCID ID:E-mail:
article id 7575, category Article
Robert T Brown, Peitsa Mikola. (1974). The influence of fruticose soil lichens upon the mycorrhizae and seedling growth of forest trees. Acta Forestalia Fennica no. 141 article id 7575. https://doi.org/10.14214/aff.7575

Water extracts of six common soil lichens, Cladonia alpestris, C. rangiferina, C. arbuscula (sylvatica), C. pleurota, Cetraria islandica, Stereocaulon paschale, inhibited growth of ectomycorrhizae of Pinus sylvestris (L.). Of 17 fungi (12 mycorrhizal) tested, many were inhibited while others were scarcely influenced or even occasionally stimulated by the extracts. Cladonia alpestris extract inhibited most fungi while C. rangiferina showed much less influence.

In pure culture synthesis experiments, 32P uptake of Pinus sylvestris was significantly reduced by C. alpestris extract. Different species of fungi showed widely variant abilities to pick up 32 P. in nursery experiments, much more vigorous growth of P. sylvestris and Picea abies (L.) H. Karst. was obtained on plots without C. alpestris than on paired plots covered with it. Betula verrucosa (B. pendula Roth) showed no difference. Under natural forest conditions, P. sylvestris seedlings grow much more rapidly where C. alpestris had been eliminated by road building or reindeer grazing than do seedlings only one meter distant under undisturbed C. alpestris cover. It is suggested that by properly controlled reindeer grazing, establishment and early growth of P. sylvestris on Cladonia sites can be much enhanced. By the time that C. alpestris could become re-established the pine seedlings would have grown large enough to suffer little from reindeer grazing. This study shows the continuity of the major components of the forest tundra biome – the dependence of pines, mycorrhizae, lichens, and reindeer and their predators (human or otherwise) upon each other for a healthy existence.

The PDF includes a summary in Finnish.

  • Brown, ORCID ID:E-mail:
  • Mikola, ORCID ID:E-mail:
article id 7561, category Article
Kalervo Salonen. (1973). On the life cycle, especially on the reproduction biology of Blastophagus piniperda L. ( Col., Scolytidae). Acta Forestalia Fennica no. 127 article id 7561. https://doi.org/10.14214/aff.7561

Blastophagus piniperda L. (now Tomicus piniperda) life cycle, especially temperature effects at various stages, has been studied in laboratory, outdoor development experiments and in natural conditions in the field in Finland, 1961–1970.

It was established that when a tree with overwintering beetles in its base is felled during the winter, the overwintered insects can establish an egg gallery into the stump without leaving for swarming flight. The egg gallery can be established in this case already about a month before swarming starts. Swarming starts after temperature in the shade exceeds 10° C. It is at its maximum at a temperature over 12° C and in conditions of no wind. The swarming starts in Southern Finland generally in the latter half of April, and in Lapland two to four weeks later than in Southern Finland, and it lasts from a few days to a few weeks. As the temperature rises, the rate of egg gallery construction and the rate of egg laying increase. The characteristics of the galleries and pupal stage are described in detail.

In Finnish conditions, B. Piniperda does not have a diapause. Movement into overwintering sites starts after the minimum daily temperature goes below 0° C. About 10% of the insects found in the overwintering sites in spring were estimated to have died during the winter. With increasing density, the number of new beetles which developed from each egg gallery decreased.

The PDF includes a summary in Finnish.

  • Salonen, ORCID ID:E-mail:
article id 7607, category Article
Eino Oinonen. (1969). The time table of vegetative spreading of the lily-of-the-valley (Convallaria majalis L.) and the wood small-reed (Calamagrostis epigeios (L.) Roth) in southern Finland. Acta Forestalia Fennica no. 97 article id 7607. https://doi.org/10.14214/aff.7607

The purpose of this study has been to compile a time-table for the vegetative spreading o the lily-of-the-valley and the wood small-reed. The diameters of the mainly solitary stands of these species have been compared to stand diameters of bracken (Lycopodium clavatum, L. annotium, and L. complanatum), to tree age determined by basal borings, and to times of fires in the site.

Regeneration of the lily-of-the-valley (Convallaria Majalis L.) and the wood small-reed (Calamagrostis epigeios (L.) Roth) from seed is relatively infrequent, and distance between stands may be considerable. Regeneration is more common after fire. The stands of the species are relatively fire resistant, especially on heathlands, where the subsurface parts are safe in the mineral soil. Large stands are generally due to vegetative spreading.

The rate of vegetative spreading of both of the species is practically constant, and about equal, 12.5 cm/year in diameter and 6.2 cm/year in the radial direction. When the actual regeneration time after the fire is not known, the rate of the spreading can be lower than this number.

  • Oinonen, ORCID ID:E-mail:
article id 7187, category Article
Eino Oinonen. (1968). Lycopodium clavatum L.- ja L. annotium L.-kasvustojen laajuus rinnastettuna samanpaikkaisiin L. complanatum L.- ja Pteridium aquilinum (L.) Kuhn -esiintymiin sekä puuston ikään ja paloaikoihin. Acta Forestalia Fennica no. 87 article id 7187. https://doi.org/10.14214/aff.7187
English title: The size of Lycopodium clavatum and L. Annotinum stands as compared to that of L. Complanatum and Pteridium aquilinum stands, the age of the stand and the dates of fire, on the site.

The present study is a part of larger project into the size and age of certain forest plants. This study seeks to confirm the size-age relationship of Lycopodium clavatum L. and L. annotium L. stands, and the time of sporal regeneration. The stand dimensions were plotted against the size of bracken (Pteridium aquilinum (L.) Kuhn) and ground pine (Lycopodium complanatum L.) stands growing on the same site, and against the age of the timber and the time of fire on the site.

The method based on parallel measurements carried out in burned forests has proved to be suitable for establishing the rate of spreading of stands and their age. The three club-moss species proved to be much alike in their sporal regeneration. They, as well as bracken, regenerate under the condition created by fire. The largest detached patches are often clones. Both Lycopodium clavatum and L. annotium survive fire poorly. Thus, the maximum size of the clones is usually connected with the last forest fire. Pieces of the shoots may survive the fires.

Sporal regeneration of these species also occur without the aid of fire. As also variation in the annual growth of the shoots can be considerable, the stand size of these species is not as good indicator of the date of fire as with Pteridium aquilinum and L. complanatum. The winding appearance of the shoots increases the error in determination of stand age by the stand size further.

The results emphasize the importance of taking into account the time and spread of the plants in the traditional vegetation analysis. A central question is: what is an individual.

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  • Oinonen, ORCID ID:E-mail:
article id 4451, category Article
Pekka Urho Sakari Tikka. (1928). Havaintoja kuusen esiintymisestä ja kehityksestä Pohjois-Suomen kuivissa kangasmetsissä. Silva Fennica no. 10 article id 4451. https://doi.org/10.14214/sf.a8399
English title: Observations on Norway spruce growth and development in dry mineral soil sites in Northern Finland.

Norway spruce (Picea abies (L.) H. Karst.) is rarely the dominant species on dry mineral soil sites in Northern Finland. These sites are, in general, too poor and dry for spruce, and suit better for Scots pine (Pinus sylvestris L.). According to the study, the natural regeneration of spruce is in Northern Finland poor. In the sample plots, cones could be found in 35% of spruce trees in the stands in natural state and 46% in the harvested stands. Compared to the spruce areas in Northern Finland, or fresh mineral soil sites in Southern Finland, cone and seed production of Norway spruce was in dry mineral soil sites very low due to scarcity of seed trees and their low cone number. There were few spruce seedlings in the sample plots, but according to the observations, spruce is able to regenerate on lichen and heath covered sites. The seedling growth was, however, poor on dry sites. Spruce seedlings were often found near fallen trees and stumps. The growing trees prevent growth of seedlings of all species. Norway spruce seems, however, to be able to spread also to the poor sites. The success depends on the vegetation and dryness of the site. For instance, spruce can spread to dry mineral soil sites from seed trees of nearby peatlands.

The PDF includes a summary in German.

  • Tikka, ORCID ID:E-mail:
article id 4731, category Article
Leo Heikurainen, Juhani Päivänen, Kustaa Seppälä. (1966). Koetuloksia männyn kylvöstä ja istutuksesta ojitetuilla soilla. Silva Fennica no. 119 article id 4731. https://doi.org/10.14214/sf.a14289
English title: Scots pine seeding and planting on drained peat soils.

This paper describes the preliminary results of Scots pine (Pinus sylvestris L.) seeding and planting trials on drained peat soils.

The results showed that a perpared peat surface was a better surface for seeding than the unprepared one. Planting of 2+1-year seedlings succeeded better than planting 1-year seedlings. Planting on the turf gave better survival than planting on the unprepared soil surface. The whole growing season was suitable time for planting Scots pine seedlings except May when the peat soil under the surface was still frozen.

Using fertilizers in connection with planting was surveyed in two ways. Mortality of seedlings increased when they were top-dressed with NPK fertilizer. Using a so-called spot fertilizing with several combinations of fertilizers resulted in K and N tending to increase the mortality of seedlings, but P decreasing mortality.

The PDF includes a summary in English.

  • Heikurainen, ORCID ID:E-mail:
  • Päivänen, ORCID ID:E-mail:
  • Seppälä, ORCID ID:E-mail:
article id 4731, category Article
Leo Heikurainen, Juhani Päivänen, Kustaa Seppälä. (1966). Koetuloksia männyn kylvöstä ja istutuksesta ojitetuilla soilla. Silva Fennica no. 119 article id 4731. https://doi.org/10.14214/sf.a14289
English title: Scots pine seeding and planting on drained peat soils.

This paper describes the preliminary results of Scots pine (Pinus sylvestris L.) seeding and planting trials on drained peat soils.

The results showed that a perpared peat surface was a better surface for seeding than the unprepared one. Planting of 2+1-year seedlings succeeded better than planting 1-year seedlings. Planting on the turf gave better survival than planting on the unprepared soil surface. The whole growing season was suitable time for planting Scots pine seedlings except May when the peat soil under the surface was still frozen.

Using fertilizers in connection with planting was surveyed in two ways. Mortality of seedlings increased when they were top-dressed with NPK fertilizer. Using a so-called spot fertilizing with several combinations of fertilizers resulted in K and N tending to increase the mortality of seedlings, but P decreasing mortality.

The PDF includes a summary in English.

  • Heikurainen, ORCID ID:E-mail:
  • Päivänen, ORCID ID:E-mail:
  • Seppälä, ORCID ID:E-mail:
article id 4723, category Article
Toivo Matilainen. (1965). Kasvutappiosta ja sen määrittämisestä Etelä-Suomen talousmetsissä. Silva Fennica no. 117 article id 4723. https://doi.org/10.14214/sf.a14281
English title: The increment loss and its determination in economically exploited forests in Southern Finland.

The present study is an attempt to clarify the decrease in growth, or the increment loss, caused by sudden reduction of growth of the growing stock below a certain level, and to find a method for its determination. Increment loss is defined as a decrease in growth during the rotation due to a deficient stock volume. The material consists of Koivisto’s yield tables for repeatedly thinned stands in Southern Finland, and the results of the Third National Forest Inventory concerning the mean volume and increment in the productive sites.

For the calculation of increment loss three formulae were constructed where the increment loss is calculated 1) as the difference between the removal by thinnings in normally developed stands during a time equal with the period of deficient stock and the suddenly removed stock, 2) according to the compound interest calculation principle as the sum of the differences which are obtained by subtracting from the removal in each thinning during the period of deficient stock its initial value, and 3) as the straight interest of the stock deficiency during the period of deficient stock.

According to the calculations, the increment loss is greatest in stands to be grown, viz. 50 m3 solid measure excluding bark per hectare tended Norway spruce stands on Oxalis-Myrtillus type sites at 40% deficiency below minimum stock. In stands to be regenerated the losses are, too, greatest in the similar stands. It exceeds 200 m3/ha when stands younger than 50 years have to been regenerated and the removal amounts to 50% of the stock. In stands to be regenerated the increment loss for spruce, due to the slow initial development by the species, is greater than for Scots pine and birch. The loss is the same at different period of age if the relative deficiency of the stock is of equal size.

According to the study, each stand has a characteristic variation in the increment loss which depends mainly on the relative degree of deficiency from the minimum stock. The formulae and methods can be used to determine the increment loss in average and better stands in Southern Finland when the stock suddenly decreases.

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  • Matilainen, ORCID ID:E-mail:
article id 4669, category Article
V. A. Kolehmainen. (1956). Vehkatallinmaa. Silva Fennica no. 90 article id 4669. https://doi.org/10.14214/sf.a9116
English title: Vehkatallinmaa, a successful reforestation area in Central Finland.

In 1933, forest fire caused by locomotive sparkle burned about 600 hectares of forest in a forest district named Vehkatallinmaa, in Central Finland. In 1934–36, the burned area was reforested, using different sowing and planting methods. At the same time, areas with poor runoff were drained. The results from reforestation throughout the area have been good. Also, natural regeneration of coniferous trees, especially Scots pine (Pinus sylvestris L.) has occurred. Even deciduous trees, especially birch (Betula sp.), have regenerated naturally in the area. The forests are an evidence of the adaptability of broadcast sowing on snow crust as a method of reproduction.

The PDF includes a summary in English.

  • Kolehmainen, ORCID ID:E-mail:
article id 4669, category Article
V. A. Kolehmainen. (1956). Vehkatallinmaa. Silva Fennica no. 90 article id 4669. https://doi.org/10.14214/sf.a9116
English title: Vehkatallinmaa, a successful reforestation area in Central Finland.

In 1933, forest fire caused by locomotive sparkle burned about 600 hectares of forest in a forest district named Vehkatallinmaa, in Central Finland. In 1934–36, the burned area was reforested, using different sowing and planting methods. At the same time, areas with poor runoff were drained. The results from reforestation throughout the area have been good. Also, natural regeneration of coniferous trees, especially Scots pine (Pinus sylvestris L.) has occurred. Even deciduous trees, especially birch (Betula sp.), have regenerated naturally in the area. The forests are an evidence of the adaptability of broadcast sowing on snow crust as a method of reproduction.

The PDF includes a summary in English.

  • Kolehmainen, ORCID ID:E-mail:
article id 4644, category Article
Vilho Antero Kolehmainen. (1955). Havaintoja kulotuksen merkityksestä metsiemme uudistamisessa. Silva Fennica no. 85 article id 4644. https://doi.org/10.14214/sf.a9106
English title: Effect of prescribed burning in the forest regeneration.

Prescribed burning has reported to avail forest regeneration, for instance, by releasing nutrients for the use of seedlings, changing the pH of the soil and decreasing competition of ground vegetation. The aim of the study was to find out if the effects could be verified. Sample plots were measured in the experimental area of Tuomarniemi, in Central Finland, both in previously burned and untreated seedling stands and young forests. The main species in the sample plots was Scots pine (Pinus sylvestris L.).

According to the results, prescribed burning prepares the soil for regeneration. Germination percentage of the seeds is higher on the burned soil. All the species, Scots pine, Norway spruce (Picea abies (L.) Karst.) and birch species (Betula sp.) grow faster. Prescribed burning increases the amount of birch seedlings by improving its regeneration compared to unburned sites. The seed trees survive burning better if they are tall and have short crown, and have thick bark. In general, prescribed burning improves regeneration in seed tree stands.

The article includes a summary in German.

  • Kolehmainen, ORCID ID:E-mail:
article id 4638, category Article
Risto Sarvas. (1953). Ohjeita pluspuiden valitsemista ja ilmoittamista varten. Silva Fennica no. 80 article id 4638. https://doi.org/10.14214/sf.a14054
English title: Instruction for the selection and registration of plus trees.

Silva Fennica Issue 80 includes presentations held in 1952 in the 7th professional development courses, arranged for foresters working in the Forest Service. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.

This presentation describes principles of breeding of forest trees and the selection of fast growing, healthy and qualitatively good trees, named plus trees, for seed production in Finland.

  • Sarvas, ORCID ID:E-mail:
article id 4637, category Article
Eero Paavola. (1953). Selostus kenttäkokeista, jotka koskivat eräiden kemiallisten aineiden käyttömahdollisuuksia metsänkylvössä. Silva Fennica no. 80 article id 4637. https://doi.org/10.14214/sf.a14053
English title: Review of field experiments on the fitness of certain chemicals for use at seeding.

Silva Fennica Issue 80 includes presentations held in 1952 in the 7th professional development courses, arranged for foresters working in the Forest Service. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.

This presentation describes a field experiment on the use of herbicides to prevent growth of ground vegetation in cutting areas before sowing. The study suggests that the chemicals used in the experiment were not effective enough to prevent growth of ground vegetation in the more fertile lands, but were effective in poorer lands. However, the treatment affected also growth of tree seedlings.

  • Paavola, ORCID ID:E-mail:
article id 4636, category Article
Tauno Hautamäki. (1953). Metsämaan laikutuksen koneellistamiskokeiluista. Silva Fennica no. 80 article id 4636. https://doi.org/10.14214/sf.a14052
English title: Mechanization of ground preparation.

Silva Fennica Issue 80 includes presentations held in 1952 in the 7th professional development courses, arranged for foresters working in the Forest Service. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.

Soil preparation improves formation of seedlings in natural regeneration sites. This presentation describes mechanization of the work using four different machines. According to the study, the effectiveness and quality of the work has improved compared to the earlier machines, and the method is becoming a competitive alternative to slash burning.

  • Hautamäki, ORCID ID:E-mail:
article id 4628, category Article
Gustaf Sirén. (1952). Havaintoja Peräpohjolan valtion mailla vuosina 1948-50 suoritetuista männyn kylvöistä. Silva Fennica no. 78 article id 4628. https://doi.org/10.14214/sf.a9100
English title: Observations on stands of Scots pine sown in state forests in Peräpohjola in northern Finland in 1948-50.

The most common way to artificially regenerate cutting areas in Northern Finland has been sowing, which has, however, often given poor results. The aim of the study was to assess the success rate of sowing and study the causes of poor regeneration.

An inventory was made of 28 areas on Empetrum-Myrtillus and Hylocomnium-Myrtillus type sites sown with Scots pine (Pinus sylvestris L.) seeds in 1948-1950. In addition, to study the effect of substrata, a sowing experiment was established. According to the inventory, regeneration failed completely in 8 and partially in 6 of the 28 sown areas. Factors that explained the poor regeneration included unfavourable weather conditions in 1948-1952, insufficient clearing of sowing spot especially when the humus layer was thick, and insect damage by Otiorynchus dubius weevil.

The article includes a summary in English.

  • Sirén, ORCID ID:E-mail:
article id 4578, category Article
Sakari Saarnijoki. (1942). Jalavan esiintymisestä Pyhäjärven, Kokemäenjoen vesistön keskusjärven tulvarannoilla. Silva Fennica no. 58 article id 4578. https://doi.org/10.14214/sf.a9082
English title: Distribution of European white elm in flooded shores of the central lake of Pyhäjärvi and Kokemäenjoki water system.

Distribution of European white elm (Ulmus laevis Pallas) is on its northernmost border in Pyhäjärvi and Kokemäenjoki area. This survey describes distribution of European white elm in the flooded shores of the central lake of Pyhäjärvi and Kokemäenjoki river water system.

Both Ulmus laevis and U. montana (now U. Glabra Hudson) can be found in the area, but most of the elms qrowing in thea area are U. laevis. U. laevis occurs around the lake in two separate areas, almost entirely in flooded shores of the lake. Regeneration of elm from seeds was limited on a narrow belt on the higher part of the flooded shore. Consequently, U. laevis can be found as zones around the lake, created by the changes in water level of the lake. The trees are judged to be native for the area.

The article includes an abstract in German.

  • Saarnijoki, ORCID ID:E-mail:
article id 4568, category Article
Atri S. Arimo. (1939). Paloalojen metsittämisestä. Silva Fennica no. 52 article id 4568. https://doi.org/10.14214/sf.a13975
English title: Forest regenration of burned areas.

Silva Fennica issue 52 includes presentations held in professional development courses, arranged for foresters working in public administration in 1938. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.

This presentation describes preparation and forest regeneration methods of a burned area.

  • Arimo, ORCID ID:E-mail:
article id 4559, category Article
A. A. Räsänen. (1939). Metsien uudistamisesta Perä-Pohjolassa. Silva Fennica no. 52 article id 4559. https://doi.org/10.14214/sf.a13966
English title: Forest regeneration in Northern Finland.

Silva Fennica issue 52 includes presentations held in professional development courses, arranged for foresters working in public administration in 1938. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.

This presentation describes the state of forests in Northern Finland, and suggests that fellings are targeted at the oldest forests until the over-presented over-aged forest are harvested. The preferred regeneration method is natural regeneration in most forest types.

  • Räsänen, ORCID ID:E-mail:
article id 4558, category Article
Olli Heikinheimo. (1939). Kokemuksia paksusammaltyypin metsien käsittelystä. Silva Fennica no. 52 article id 4558. https://doi.org/10.14214/sf.a13965
English title: Experiences in forest management of Hylocomnium-Myrtillus type forests in Northern Finland.

Silva Fennica issue 52 includes presentations held in professional development courses, arranged for foresters working in public administration in 1938. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.

This presentation describes a study of forest management practices suitable for in Hylocomnium-Myrtillus site type forests, typical for Northern Finland. A special problem for the forest site type is poor regeneration due weak seed production of Norway spruce (Picea abies L. Karst.) and thick moss layer.

  • Heikinheimo, ORCID ID:E-mail:
article id 4571, category Article
Tauno Lampimäki. (1939). Nautakarjan laiduntamisesta metsämailla. Silva Fennica no. 50 article id 4571. https://doi.org/10.14214/sf.a9074
English title: Grazing of cattle in forest land.

Finland has a long tradition of grazing cattle in the forests and common land. There are also reports of degradation of forests by grazing already in 1600th century. The aim of the survey was to study which positive and negative effects grazing has in forests.

The study concludes that grazing has caused considerable economic losses through damages to forests. In addition, woodland pastures cannot give the yields required in modern animal husbandry. The quality of woodland pastures have decreased after the woodlands used in slash and burn culture have become wooded.

Grazing has also some positive effects to forests. It increases the diversity of vegetation in the woodland pastures and spreads species to new areas. This is supported by the lists of species found in different woodland pastures. Cattle destroy large grasses like Calamagrostis, which may avail growth of tree seedlings in the pastures. Grazing can also prepare the site for tree seedlings. On the other hand, prolonged grazing destroys tree seedlings and prevents regeneration.

The article includes a German summary.

  • Lampimäki, ORCID ID:E-mail:
article id 4543, category Article
Erkki Laitakari. (1938). Metsän uudistamisesta laihoilla kangasmailla. Silva Fennica no. 46 article id 4543. https://doi.org/10.14214/sf.a13952
English title: Forest regeneration on poor forest sites.

Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service. 

This presentation describes forest regeneration on poor forest sites.

  • Laitakari, ORCID ID:E-mail:
article id 4543, category Article
Erkki Laitakari. (1938). Metsän uudistamisesta laihoilla kangasmailla. Silva Fennica no. 46 article id 4543. https://doi.org/10.14214/sf.a13952
English title: Forest regeneration on poor forest sites.

Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service. 

This presentation describes forest regeneration on poor forest sites.

  • Laitakari, ORCID ID:E-mail:
article id 4543, category Article
Erkki Laitakari. (1938). Metsän uudistamisesta laihoilla kangasmailla. Silva Fennica no. 46 article id 4543. https://doi.org/10.14214/sf.a13952
English title: Forest regeneration on poor forest sites.

Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service. 

This presentation describes forest regeneration on poor forest sites.

  • Laitakari, ORCID ID:E-mail:
article id 4542, category Article
Erkki Laitakari. (1938). Eräitä metsän uudistamisessa huomioon otettavia näkökohtia. Silva Fennica no. 46 article id 4542. https://doi.org/10.14214/sf.a13951
English title: Forest regeneration.

Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service. 

This presentation describes factors that should be taken into account in forest regeneration. 

  • Laitakari, ORCID ID:E-mail:
article id 4542, category Article
Erkki Laitakari. (1938). Eräitä metsän uudistamisessa huomioon otettavia näkökohtia. Silva Fennica no. 46 article id 4542. https://doi.org/10.14214/sf.a13951
English title: Forest regeneration.

Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service. 

This presentation describes factors that should be taken into account in forest regeneration. 

  • Laitakari, ORCID ID:E-mail:
article id 4536, category Article
V. K. Ahola. (1938). Metsän uudistumisen tarkkailusta kovilla metsämailla ja ojitetuilla soilla sekä ojien tarkkailusta. Silva Fennica no. 46 article id 4536. https://doi.org/10.14214/sf.a13945
English title: Inspection of regeneration of forests in mineral soil forests and drained peatlands and checking of ditches.

Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service. 

This presentation describes inspection of forest regeneration of mineral soil forest types and drained peatlands, and inspection of ditches. 

  • Ahola, ORCID ID:E-mail:
article id 4536, category Article
V. K. Ahola. (1938). Metsän uudistumisen tarkkailusta kovilla metsämailla ja ojitetuilla soilla sekä ojien tarkkailusta. Silva Fennica no. 46 article id 4536. https://doi.org/10.14214/sf.a13945
English title: Inspection of regeneration of forests in mineral soil forests and drained peatlands and checking of ditches.

Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service. 

This presentation describes inspection of forest regeneration of mineral soil forest types and drained peatlands, and inspection of ditches. 

  • Ahola, ORCID ID:E-mail:
article id 4534, category Article
Martti Tertti. (1938). Hakkausalan raivaamisesta. Silva Fennica no. 46 article id 4534. https://doi.org/10.14214/sf.a13943
English title: Clearing of a felling area.

Silva Fennica issue 46 includes presentations held in professional development coursesarranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administrationespecially in regional levelThe education was arranged by Forest Service. 
This presentation describes clearing of felling areas. 

  • Tertti, ORCID ID:E-mail:
article id 4530, category Article
O. J. Lukkala. (1938). Ojitettujen soiden metsittämisestä. Silva Fennica no. 46 article id 4530. https://doi.org/10.14214/sf.a13939
English title: Afforestation of drained peatlands.

Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service. 

This presentation describes afforestation of drained peatlands. 

  • Lukkala, ORCID ID:E-mail:
article id 4530, category Article
O. J. Lukkala. (1938). Ojitettujen soiden metsittämisestä. Silva Fennica no. 46 article id 4530. https://doi.org/10.14214/sf.a13939
English title: Afforestation of drained peatlands.

Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service. 

This presentation describes afforestation of drained peatlands. 

  • Lukkala, ORCID ID:E-mail:
article id 4529, category Article
V. T. Aaltonen. (1938). Maa ja metsän uudistuminen. Silva Fennica no. 46 article id 4529. https://doi.org/10.14214/sf.a13938
English title: Forest regeneration and soil.

Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service. 

This presentation describes the effect of properties of soil on forest regeneration. 

  • Aaltonen, ORCID ID:E-mail:
article id 4529, category Article
V. T. Aaltonen. (1938). Maa ja metsän uudistuminen. Silva Fennica no. 46 article id 4529. https://doi.org/10.14214/sf.a13938
English title: Forest regeneration and soil.

Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service. 

This presentation describes the effect of properties of soil on forest regeneration. 

  • Aaltonen, ORCID ID:E-mail:
article id 4492, category Article
V. K. Ahola. (1937). Metsän keinollisesta uudistamisesta. Silva Fennica no. 39 article id 4492. https://doi.org/10.14214/sf.a13902
English title: Artificial regeneration of forests.

Silva Fennica Issue 39 includes presentations held in professional development courses in 1935 that were arranged for foresters working in public administration. The presentations focus on practical issues in forest management and administration, especially in regional level.

 This presentation describes the methods of artificial regeneration of forests.

  • Ahola, ORCID ID:E-mail:
article id 4486, category Article
Martti Tertti. (1937). Metsien luontaisen uudistumisen edistämisestä. Silva Fennica no. 39 article id 4486. https://doi.org/10.14214/sf.a13896
English title: Facilitating natural regeneration.

Silva Fennica Issue 39 includes presentations held in professional development courses in 1935 that were arranged for foresters working in public administration. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.

This presentation describes natural regeneration in forests and factors affecting the success of regeneration.

  • Tertti, ORCID ID:E-mail:
article id 4479, category Article
L. E. T. Borg. (1936). Hankikylvöt Tuomarniemen hoitoalueessa vv. 1913-1930. Silva Fennica no. 38 article id 4479. https://doi.org/10.14214/sf.a9065
English title: Areas broadcast sown on snow in Tuomarniemi district in Finland in 1913-1930.

Broadcast sowing on snow was relatively new method in the beginning of the 20th century in Finland, and the experiences of regeneration were diverse. The aim of the survey was to study the success rate of regeneration in the oldest and largest areas regenerated with this sowing method in Tuomarniemi district. Scots pine (Pinus sylvestris L.) was the most common tree species, but also Norway spruce (Picea abies (L.) H. Karst.) and European larch (Larix decidua Mill.) were used in broadcast sowing on snow.

According to the study, the success of broadcast sowing on snow was as good as patch sowing and sowing in furrows in the sites typical for Tuomarniemi. The regeneration areas were often drained peatlands or paludified lands. When sowing is done using Norway spruce seeds, site preparation either by broadcast burning or scalping with hoe is recommended. Mixed sowing with pine and spruce seldom succeeded due to the differences in site requirements of the species and growth of seedlings. Sowing of Scots pine succeeded well on the drained peatlands. Sowing should be done some years after draining to let the peat dry and sink. Site preparation is needed in sites growing Polytrichum-moss. Broadcast burned areas larger than 10 hectares seemed to regenerate poorer than sites in average, possibly due to dryness of the sites. Trials with European larch were successful, and the growth of the seedlings acceptable despite the sites being relatively poor for the species.

The PDF includes a summary in German.

  • Borg, ORCID ID:E-mail:

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