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Under compilation: 55(5)

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Silva Fennica 1926-1997
Acta Forestalia Fennica

Articles containing the keyword 'structural diversity'.

Category: Research article

article id 84, category Research article
Asko Lõhmus, Piret Lõhmus. (2011). Old-forest species: the importance of specific substrata vs. stand continuity in the case of calicioid fungi. Silva Fennica vol. 45 no. 5 article id 84.
Appropriate conservation management of old-forest species depends on the causes of their old-forest affinity, which, however, are insufficiently known. Calicioid fungi are often considered old-forest dependent because of their special requirements for microhabitat, microclimate, and stand continuity for at least two tree generations. We demonstrate that, for several methodological or interpretational problems, published studies do not provide unequivocal evidence for such mechanisms and even for old-forest dependency of calicioids in general. We then analyse a large Estonian dataset (ca. 2300 records of 32 species) representing various management types and site types to answer whether old forests have more calicioid species, and any specific species, than could be expected for the substratum availability observed. Although old growth had more species and records than mature managed stands or cutover sites, those substratum types that occurred at roughly similar abundances also hosted comparable numbers of species in different management types. The characteristic substrata adding extra species to old growth were snags and root-plates of treefall mounds; wood surfaces in general comprised more than half of all calicioid records. Although substratum abundance did not fully explain the species-richness contrast between old growth and mature stands, additional evidence suggested that the unexplained variance is rather due to small-scale habitat characteristics than stand-scale continuity or microclimate. Finally, we review the evidence for old-forest affinity of calicioid species and distinguish a set of threatened species. We conclude that the availability of specific substrata is the main limiting factor for calicioid fungi in forests, and its quantitative and stochastic nature explains the large random and region-specific variation in the published lists of ‘old-forest species’.
  • Lõhmus, Department of Zoology, Institute of Ecology and Earth Sciences, University of Tartu, Vanemuise st. 46, EE-51014, Tartu, Estonia ORCID ID:E-mail: (email)
  • Lõhmus, Department of Botany, Institute of Ecology and Earth Sciences, University of Tartu, Tartu, Estonia ORCID ID:E-mail:
article id 352, category Research article
Juha-Pekka Hotanen, Matti Maltamo, Antti Reinikainen. (2006). Canopy stratification in peatland forests in Finland. Silva Fennica vol. 40 no. 1 article id 352.
Abundance and species number of the tree and shrub vegetation in different canopy layers were analysed according to site quality class and drainage succession phase on permanent sample plots on spruce mires (n = 268) and pine mires (n = 628) in the Finnish National Forest Inventory in 1995. The abundances based on the crown coverage were compared with the abundances based on the parallel basal area of the tree stand. The canopy coverages and species number for peatland forests were also compared with those for mineral soil forests on the permanent sample plots (n = 1725) in 1995. In general, effective temperature sum correlated positively, although not very strongly, with the coverages and species number in most of the canopy layers, as well as with the mean range of the diameter distribution. The effects of both site quality class and drainage phase were stronger on pine mires than on spruce mires, most probably due to the longer fertility gradient and large potential free growing space in the former group. On pine mires, drainage increased the abundances and species number in the different canopy layers, as well as the structural inequality of the tree stands. On spruce mires, the increase was principally allocated to the abundances of the dominant and intermediate tree layers. The correlations between the total crown coverage of the tree layers and stand basal area were r = 0.45 for spruce mires and r = 0.70 for pine mires. Compared to mineral soil forests, in addition to having a higher abundance of Betula pubescens, the dominant layer was not as pronounced in peatland forests. On spruce mires, the coverage of the shrub layer on mesotrophic and meso-oligotrophic sites was higher than that in mineral soil forests. The average species number in different canopy layers did not differ significantly between spruce mires and mineral soil forests in corresponding site quality classes. On pine mires, the species number was generally lower (except for the mesotrophic sites) than that in corresponding mineral soil forests.
  • Hotanen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland ORCID ID:E-mail: (email)
  • Maltamo, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland ORCID ID:E-mail:
  • Reinikainen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland ORCID ID:E-mail:

Category: Research note

article id 9899, category Research note
Linda Robalte, Diāna Jansone, Didzis Elferts, Roberts Matisons, Āris Jansons. (2018). Bilberry ramet dimensions in relation to stand age in oligotrophic conditions in Latvia. Silva Fennica vol. 52 no. 1 article id 9899.
Highlights: Bilberry ramet dimensions (age, diameter, height) and their structural diversity, as well as cover, increased with stand age; Active rejuvenation of ramets was observed in younger stands; The oldest bilberry ramets (>10 years of age) occurred in stands older than 70 years.

Dwarf shrub layer is an important component of boreal and hemiboreal forest ecosystems that has received little attention, particularly regarding its structural diversity, which, however, could serve as an additional proxy for habitat quality. Dimensions of bilberry (Vaccinium myrtillus L.) ramets were assessed in two sites in Latvia covered by dry oligotrophic Scots pine (Pinus sylvestris L.) stands 10–230 years of age. In total, 20 sampling plots (10×10 m) with 156 subplots (1×1 m) were sampled and 630 bilberry ramets analysed. The dimensions of ramets (age, diameter, and height) and cover of bilberry increased with stand age. The age of the studied ramets ranged 2–13 years; 5–6 years-old ramets were most frequent in all stands. The skewness of the distribution of the ramet dimensions shifted with stand age, leaning towards the higher values. Lower structural diversity of ramets was observed in stands 50–100 years of age. The highest diversity of ramet age structure occurred in stands younger than 150 years, whereas the oldest and largest ramets mostly occurred in the older stands (>150 years). Considering structural diversity of ramets, recovery of bilberry after stand-replacing disturbance (e.g. clearcut) was a continuous process, similarly to that observed in tree layer.

  • Robalte, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia ORCID ID:E-mail: (email)
  • Jansone, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia; University of Latvia, Faculty of Biology, Jelgavas Str. 1, LV 1004, Riga, Latvia ORCID ID:E-mail:
  • Elferts, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia; University of Latvia, Faculty of Biology, Jelgavas Str. 1, LV 1004, Riga, Latvia ORCID ID:E-mail:
  • Matisons, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia ORCID ID:E-mail:
  • Jansons, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia ORCID ID:E-mail:

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