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Articles containing the keyword 'swamp'.

Category: Research article

article id 10013, category Research article
Hardo Becker, Jürgen Aosaar, Mats Varik, Gunnar Morozov, Kristiina Aun, Ülo Mander, Kaido Soosaar, Veiko Uri. (2018). Annual net nitrogen mineralization and litter flux in well-drained downy birch, Norway spruce and Scots pine forest ecosystems. Silva Fennica vol. 52 no. 4 article id 10013. https://doi.org/10.14214/sf.10013
Highlights: The net nitrogen mineralization (NNM) flux in drained peat soils depends largely on the C/N ratio and tree species; The soil NNM process is affected by trees through organic litter input into soil; Pine stand in low-fertility drained transitional bog is dominated by net ammonification; Birch and spruce stands on the fertile drained peat soil with higher pH and N content are dominated by net nitrification.

The main aim of the current study was to estimate the annual net nitrogen mineralization (NNM) flux in stands of different tree species growing on drained peatlands, as well as to clarify the effect of tree species, soil properties and litter on annual NNM dynamics. Three study sites were set up in May 2014: a downy birch (Betula pubescens Ehrh.) stand and a Norway spruce (Picea abies (L.) Karst.) stand in Oxalis full-drained swamp (ODS) and a Scots pine (Pinus sylvestris L.) stand in Myrtillus full-drained swamp (MDS). The NNM flux was estimated using the in situ method with incubated polyethylene bags. The highest value of NNM was found in stands that were growing on fertile ODS: 127.5 kg N ha–1 yr–1 and 87.7 kg N ha–1 yr–1, in the downy birch stand and in the Norway spruce stand, respectively. A significantly lower annual NNM flux (11.8 kg N ha–1 yr–1) occurred in the Scots pine stand growing in MDS. Nitrification was highest at fertile ODS sites and ammonification was the highest at the low fertility MDS site. For all study sites, positive correlation was found between soil temperature and NNM intensity. The difference in annual NNM between the downy birch stand and the Norway spruce stand growing on similar drained fertile peatlands was due to litter quality. The annual N input into the soil through leaf litter was the highest at the downy birch site where also the C/N ratio of litter was the lowest. The second highest N input into the soil was found in the spruce stand and the lowest in the pine stand.

  • Becker, Chair of Silviculture and Forest Ecology, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Friedrich Reinhold Kreutzwaldi 1, 51014 Tartu, Estonia ORCID ID:E-mail: hardo.becker@emu.ee (email)
  • Aosaar, Chair of Silviculture and Forest Ecology, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Friedrich Reinhold Kreutzwaldi 1, 51014 Tartu, Estonia ORCID ID:E-mail: jyrgen.aosaar@emu.ee
  • Varik, Chair of Silviculture and Forest Ecology, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Friedrich Reinhold Kreutzwaldi 1, 51014 Tartu, Estonia ORCID ID:E-mail: mats.varik@emu.ee
  • Morozov, Chair of Silviculture and Forest Ecology, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Friedrich Reinhold Kreutzwaldi 1, 51014 Tartu, Estonia ORCID ID:E-mail: gunnar.morozov@emu.ee
  • Aun, Chair of Silviculture and Forest Ecology, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Friedrich Reinhold Kreutzwaldi 1, 51014 Tartu, Estonia ORCID ID:E-mail: kristiina.aun@emu.ee
  • Mander, Department of Geography, Institute of Ecology & Earth Sciences, University of Tartu, Ülikooli 18, 50090 Tartu, Estonia ORCID ID:E-mail: ulo.mander@ut.ee
  • Soosaar, Department of Geography, Institute of Ecology & Earth Sciences, University of Tartu, Ülikooli 18, 50090 Tartu, Estonia ORCID ID:E-mail: kaido.soosaar@ut.ee
  • Uri, Chair of Silviculture and Forest Ecology, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Friedrich Reinhold Kreutzwaldi 1, 51014 Tartu, Estonia ORCID ID:E-mail: veiko.uri@emu.ee
article id 353, category Research article
Jaakko Repola, Hannu Hökkä, Timo Penttilä. (2006). Thinning intensity and growth of mixed spruce-birch stands on drained peatlands in Finland. Silva Fennica vol. 40 no. 1 article id 353. https://doi.org/10.14214/sf.353
The impacts of thinning at various intensities on the growth and mortality of mixed spruce-birch stands were investigated in thinning experiments on spruce swamps in northern and south-eastern Finland. At the time of establishment, three of the stands had recently reached the first commercial thinning stage and four were more advanced. The monitoring period was mainly 15 years, and the thinning intensity varied from heavy thinning (ca. 46 per cent of the basal area removed) to no thinning. Basal area removals of light and moderate thinning were ca. 22% and 39%, respectively. Unthinned plots had the highest volume increment. Light and moderate thinning slightly decreased the 15-year volume increment by, on an average, 1% and 8%, respectively. Heavy thinning led to a greater reduction (22%) in volume increment. The growth response to thinning intensity was evident as a higher relative volume and mean diameter increment of the living trees with decreasing stand density. Part of the volume increment on the unthinned plots was lost through natural mortality. Even light thinning significantly decreased natural mortality.
  • Repola, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland ORCID ID:E-mail: jaakko.repola@metla.fi (email)
  • Hökkä, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland ORCID ID:E-mail:
  • Penttilä, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland ORCID ID:E-mail:

Category: Article

article id 7466, category Article
Leo Heikurainen. (1955). Rämemännikön juuriston rakenne ja kuivatuksen vaikutus siihen. Acta Forestalia Fennica vol. 65 no. 3 article id 7466. https://doi.org/10.14214/aff.7466
English title: Structure of Scots pine root systems in a pine swamp and effect of draining on the structure.
Original keywords: räme; ojitus; juuristo; mänty; mykorritsa; suo

The root system of a Scots pine (Pinus sylvestris L.) growing on a peatland is restricted, according to earlier studies, on the top layers of the peat above the groundwater level. Drainage of the peatland affects growth of the root system. This investigation aims at studying the root systems on the point of view of draining of peatlands. The structure and distribution, and the growth of mycorrhiza in Scots pine roots in pine swamps varying from natural state to well drained state is studied.

The study shows that Scots pine on pine swamps has more extensive root system than has earlier assumed, it is common to find 1,000 m of roots in one cubic meter in a healthy stand. The trees reach this density of roots early on. In a drained peatland, the total root length is markedly higher than in a similar stand in natural state. The root systems proved to be very shallow. Even in a well-drained site the roots did not grow deeper than 20 cm. 70% of all roots were found in the upper 5 cm layer of peat, and 90% in the upper 10 cm layer. Root systems were deeper in drained peatlands, but the difference was small. In a site in natural state the average depth of the roots was 4 cm, and in a drained site 5 cm. About 85% of the roots were under 1 mm of diameter. Short roots were found only in the fine roots. Draining increases strongly the number of short roots. Mycorrhizas of the types A, B, C and D as well as pseudomychorrizas were found in the pine roots.

The PDF includes a summary in German.

  • Heikurainen, ORCID ID:E-mail:
article id 7439, category Article
Leo Heikurainen. (1954). Rämemänniköiden uudistamisesta paljaaksihakkausta käyttäen. Acta Forestalia Fennica vol. 61 no. 27 article id 7439. https://doi.org/10.14214/aff.7439
English title: Regeneration of Scots pine stands of pine swamps through clear cutting.

Pine swamps are easily regenerated by natural regeneration of Scots pine (Pinus sylvestris L.). Usually seeding felling is used, but also strip system or clear cutting and regeneration along stand edge has been suggested. This article discusses the regeneration by clear cutting and sparing the existing undergrowth. The article focuses on pine swamps to be drained and the ones in natural state.

Pine swamps in natural state usually have plenty of trees of smaller diameter classes, that can be trusted to form the future tree generation after the felling. This shortens the rotation by 20-30 years. The undergrowth has been shown to recover quickly. The method suits for regeneration of drained peatlands but could fit also for regeneration of pine swamps in natural state.

The seedlings in the pine swamps are mainly 1-5 years old, and the stock is changing. It seems that larger trees produce a wider selection of age groups, but the seedlings survive longer under smaller mother trees. Part of the younger generations of seedlings seem to be destroyed when the peatland is drained. Further studies are needed to investigate how the draining and felling are to be performed to spare the young seedlings.

The Acta Forestalia Fennica issue 61 was published in honour of professor Eino Saari’s 60th birthday.

The PDF includes a summary in German.

  • Heikurainen, ORCID ID:E-mail:
article id 5212, category Article
Heikki Kurimo. (1984). Simultaneous groundwater table fluctuation in different parts of virgin pine mires. Silva Fennica vol. 18 no. 2 article id 5212. https://doi.org/10.14214/sf.a15391

The study discusses the amplitude of the simultaneous groundwater table fluctuations in different parts of pine mires, and factors influencing it. The assumption generally used in hydrological computations that the simultaneous vertical fluctuation in the groundwater table in different parts of mires are equal does not hold good in detail. Numerous cases were detected where the fluctuation at one place did not correspond to that at another site to a statistically significant degree. The main reason for the unequal fluctuation at the different sites seems to be the difference in the microtopography and in the hydraulic conductivity between the sites.

The PDF includes a summary in Finnish.

  • Kurimo, ORCID ID:E-mail:
article id 4990, category Article
Michael Starr, Carl Johan Westman. (1978). Easily extractable nutrients in the surface peat layer of virgin sedge-pine swamps. Silva Fennica vol. 12 no. 2 article id 4990. https://doi.org/10.14214/sf.a14844

The paper deals with the nutrient status of surface peat layer from virgin sedge-pine swamps and its relationship to peatland types. When the nutrients are expressed in mg/100 g peat, only easily extractable Ca and Mg correspond to the productivity status of the peatland type. N, P, and K levels in the herb rich sedge-pine swamp are generally lower than in the small sedge-pine swamps, which are the least productive ones. The differences between the site types in all the five nutrients become much clearer when the results are expressed in kg/ha. P, K, and Ca are significantly different between the site types, and correspond to the productivity of the site type. For N and Mg the same tendency can be seen. The organically bound nutrients N, and to a lesser extent, P appear to comply with the hypothesis of an increase in nutrient availability in Southern Finland.

The PDF includes a summary in Finnish.

  • Starr, ORCID ID:E-mail:
  • Westman, ORCID ID:E-mail:
article id 7011, category Article
Antti Tanttu. (1915). Tutkimuksia ojitettujen soiden metsittymisestä. Acta Forestalia Fennica vol. 5 no. 2 article id 7011. https://doi.org/10.14214/aff.7011
English title: Studies on forest growth of drained peatlands.
Original keywords: ojitus; räme; neva; korpi; turvekangas

Draining of peatlands to improve forest growth started to increase in Finland in the beginning of 1900s. The aim of the study was to find out which kind of peatlands are suitable for draining. The peatlands examined in this study had been drained earlier in 1800s for other purposes, and the original peatland type was deduced afterwards. When the peatland is drained, its vegetation changes gradually towards that of mineral soil sites, depending on the original peatland type. The article includes detailed description of the vegetation on different drained peatland sites. Best represented in the study were different types of pine swamps, which change towards Calluna or Vaccinium forest site type depending on the original peatland type. The Sphangnum species and brushwood disappear gradually and Cladina sp. become common in some drained pine swamp types. Scots pine (Pinus sylvestris L.) regenerates well on most drained pine swamps, and also Betula sp may grow as dominant species. The richer pine swamp types develop to Vaccinium-myrtillus forest site type, which may grow also Norway spruce (Picea abies (L.) H. Karst). Drained treeless bogs change first towards pine swamps. However, trees regenerate poorly on these sites and the growth is low. Flark-bogs develop typically to treeless lichen heaths. Drained spruce swamps develop to forest with grass-herb vegetation or Myrtillus site type.

The PDF includes a summary in German.

  • Tanttu, ORCID ID:E-mail:
article id 7011, category Article
Antti Tanttu. (1915). Tutkimuksia ojitettujen soiden metsittymisestä. Acta Forestalia Fennica vol. 5 no. 2 article id 7011. https://doi.org/10.14214/aff.7011
English title: Studies on forest growth of drained peatlands.
Original keywords: ojitus; räme; neva; korpi; turvekangas

Draining of peatlands to improve forest growth started to increase in Finland in the beginning of 1900s. The aim of the study was to find out which kind of peatlands are suitable for draining. The peatlands examined in this study had been drained earlier in 1800s for other purposes, and the original peatland type was deduced afterwards. When the peatland is drained, its vegetation changes gradually towards that of mineral soil sites, depending on the original peatland type. The article includes detailed description of the vegetation on different drained peatland sites. Best represented in the study were different types of pine swamps, which change towards Calluna or Vaccinium forest site type depending on the original peatland type. The Sphangnum species and brushwood disappear gradually and Cladina sp. become common in some drained pine swamp types. Scots pine (Pinus sylvestris L.) regenerates well on most drained pine swamps, and also Betula sp may grow as dominant species. The richer pine swamp types develop to Vaccinium-myrtillus forest site type, which may grow also Norway spruce (Picea abies (L.) H. Karst). Drained treeless bogs change first towards pine swamps. However, trees regenerate poorly on these sites and the growth is low. Flark-bogs develop typically to treeless lichen heaths. Drained spruce swamps develop to forest with grass-herb vegetation or Myrtillus site type.

The PDF includes a summary in German.

  • Tanttu, ORCID ID:E-mail:
article id 7530, category Article
Aimo Kaarlo Cajander. (1913). Studien über die Moore Finnlands. Acta Forestalia Fennica vol. 2 no. 3 article id 7530. https://doi.org/10.14214/aff.7530
English title: Studies about Finnish peatlands.

The first part of the article presents an overview of peatlands and their classification on the basis of e.g. their nutrient content in Finland. The second part (a separate pdf file) represents the different types of peatland with their indicator plants in Finland. 

Peatlands can be classified in several ways, depending on the purpose of the classification. There are also differences whether an area is considered biologically or geographically as a peatland. For agricultural or forestry purposes the peatlands can be classified on the basis of their nutrient content.

Article also discusses the three ways of emergence of peatlands and the variations of water relations of peatlands.    

  • Cajander, ORCID ID:E-mail:
article id 4672, category Article
Leo Heikurainen. (1957). Lettoräme ja sen metsäojituskelpoisuus. Silva Fennica no. 93 article id 4672. https://doi.org/10.14214/sf.a9119
English title: Eutrophic pine bogs and their suitability for draining.

There are about 155,000 ha of fen-like pine swamps (eutrophic pine bogs) in Finland, major part of which are situated in Northern Finland. In the classification system for drainability of peatlands, this type of peatlands had been placed in the first class. The study presents a new evaluation for the peatland type, based on vegetation and tree growth.

According to a previous study, the vegetation of fen-like pine swamps can be characterised by distinctive plant communities that seem to reflect the fertility and high pH of the underlying soil. In this study, the fen-like pine swamps were divided in two subtypes based on the vegetation: proper fen-like pine swamps and fen-like pine swamps with ericaceous shrubs. Both have distinctive vegetation, which is described in the article. The distribution of the subtypes seems to be different: fen-like pine swamps with ericaceous shrubs are more common in eastern parts of Northern Finland. The two subtypes could be divided in different drainability classes according to tree growth, proper fen-like pine swamps belonging to class 1 and fen-like pine swamps with ericaceous shrubs to class 4.

The article includes a summary in English.

  • Heikurainen, ORCID ID:E-mail:
article id 4672, category Article
Leo Heikurainen. (1957). Lettoräme ja sen metsäojituskelpoisuus. Silva Fennica no. 93 article id 4672. https://doi.org/10.14214/sf.a9119
English title: Eutrophic pine bogs and their suitability for draining.

There are about 155,000 ha of fen-like pine swamps (eutrophic pine bogs) in Finland, major part of which are situated in Northern Finland. In the classification system for drainability of peatlands, this type of peatlands had been placed in the first class. The study presents a new evaluation for the peatland type, based on vegetation and tree growth.

According to a previous study, the vegetation of fen-like pine swamps can be characterised by distinctive plant communities that seem to reflect the fertility and high pH of the underlying soil. In this study, the fen-like pine swamps were divided in two subtypes based on the vegetation: proper fen-like pine swamps and fen-like pine swamps with ericaceous shrubs. Both have distinctive vegetation, which is described in the article. The distribution of the subtypes seems to be different: fen-like pine swamps with ericaceous shrubs are more common in eastern parts of Northern Finland. The two subtypes could be divided in different drainability classes according to tree growth, proper fen-like pine swamps belonging to class 1 and fen-like pine swamps with ericaceous shrubs to class 4.

The article includes a summary in English.

  • Heikurainen, ORCID ID:E-mail:
article id 4620, category Article
Leo Heikurainen. (1951). Eräs suokasvillisuuden analysoimismenetelmä. Silva Fennica no. 70 article id 4620. https://doi.org/10.14214/sf.a9092
English title: Determination method for peatland vegetation.

The aim of the study was to develop an determination method to define vegetation type of fen-like pine swamps, which are combinations of two peatland types, quagmire and pine swamp. Typical for this peatland type is that the vegetation is very heterogenous. Patches of different types of plant communities are found within a small area, but in a large scale, there are only few main types of plant communities. The commonly used way to use sample plots to study tree stands suit poorly to determine the type of this kind of heterogenous ground vegetation. The article compares strip survey and circular plot survey, of which circular plot survey is determined to be less time consuming. The article describes a way to choose the locations of sample points to achieve most correct areas for different plant communities.

The article includes an abstract in German.

  • Heikurainen, ORCID ID:E-mail:

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