Evaluation of the double normal distribution for tree diameter distribution modeling

Zasada M. (2013). Evaluation of the double normal distribution for tree diameter distribution modeling. Silva Fennica vol. 47 no. 2 article id 956. https://doi.org/10.14214/sf.956

Abstract

The double normal distribution consists of two normal distributions truncated at their means and then combined in such a way, that points of truncation now become the overall distribution mode. So far, parameters of the double normal distribution have been estimated exclusively using the method of moments. This study evaluates the maximum likelihood method for the estimation of the double normal distribution parameters in Scots pine stands in Poland, and compares it to the results of the method of moments and the two-parameter Weibull distribution fitted using the maximum likelihood method and the method of moments. Presented results show that it is not recommended to use the maximum likelihood method of parameter fitting with Nelder-Mead and quasi-Newton optimization algorithms for the double normal distribution for small samples. However, it can be used for large samples, giving the fit comparable to the two-parameter Weibull distribution and providing parameters having sound practical and biological meaning. In the case of smaller samples for the double normal distribution it is recommended to apply the maximum likelihood method with the alternative simulated annealing optimization algorithm, use the method of moments or substitute the described distribution with more the flexible and robust Weibull distribution. For the smaller samples, the method of moments was superior to the maximum likelihood method.

Keywords

diameter distribution; Weibull distribution; maximum likelihood method; method of moments; goodness-of-fit; simulated annealing

Author Info

Received 12 November 2012 Accepted 18 March 2013 Published 1 July 2013

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Available at https://doi.org/10.14214/sf.956 | Download PDF

For more than a hundred years foresters have written about how to describe stand diameter distributions using theoretical models, especially as a part of building volume and yield tables and growth models. Much research has been done on the utilization of various theoretical functions for diameter distribution modeling, such as Gram-Charlier series (Cajanus 1914), geometrical series, proposed by de Liocourt, an exponential form of the de Liocourt model (Meyer and Stevenson 1943; Meyer 1952; Leak 1964), Pearson distributions (Schnur 1934), Pearl-Reed growth curves (Osborne and Schumacher 1935; Nelson 1964), gamma distribution (Nelson 1935), lognormal distribution (Bliss and Reinker 1964), beta distribution (Clutter and Bennett 1965; McGee and Della-Bianca 1967; Lenhard and Clutter 1971), Weibull distribution (Bailey and Dell 1973) , logit-logistic distribution (Wang and Rennols 2005), SB Johnson distribution (Hafley and Shreuder 1977; Siekierski 1992) and Birnbaum–Saunders distribution (Podlaski 2008). Recently, there were also a few results published on the use of the mixture distribution for the modeling of tree diameter distribution in stands with more complex structure (Zhang et al. 2001; Liu et al. 2002; Zasada and Cieszewski 2005; Podlaski 2010; Podlaski 2011). Nowadays, because of its properties and advantages, one of the most frequently used has been the Weibull distribution, especially in its 3-parameter and, if necessary, truncated version (see Merganič and Sterba 2006). However, Maltamo et al. (1995) found 2-parameter Weibull to perform better than the 3-parameter one. Despite many successful applications of various functions, there is still a lot of research being done to find alternative functions and parameter estimation methods that could provide a better fit of models to the data or give more practical interpretations of their parameters.

One example of such an effort is the double normal distribution. Bruchwald (1988) used this newly developed distribution for tree diameter description in a growth model for even-aged stands of Scots pine (*Pinus sylvestris* L.). Siekierski (1991) provided the description of various methods of parameter estimation for the double normal distribution. The double normal distribution was later used for tree diameter modeling of other species such as true fir (*Abies alba* Mill.) (Zasada 1995), beech (*Fagus sylvatica*) (Rymer-Dudzińska and Dudzińska 2001), and birch (*Betula*) (Zasada 2000). The described distribution was also used as an element of growth models for Scots pine (Bruchwald 1986; Bruchwald 1991), Norway spruce (*Picea abies *L.) (Bruchwald et al. 1999), true fir (Zasada 1999), and oak (*Quercus*) (Bruchwald et al. 1996), developed in Poland.

So far, parameters of the double normal distribution have been estimated exclusively using the method of moments (Siekierski 1991), which in some cases reveals limitations. The aim of this study was to:

- evaluate the maximum likelihood method (MLM) for the estimation of the double normal distribution parameters in Scots pine stands,
- compare the maximum likelihood estimates to different variants of the method of moments (MOM) for the double normal distribution Siekierski (1991) and for the Weibull distribution,
- compare the double normal distribution to the two-parameter Weibull distribution, which has been the most widely used for tree diameter distribution modeling in forestry, and
- evaluate and compare the performance of the double normal and two-parameter Weibull distributions in estimation of parameters in cases where one has to deal with small samples.

The double normal distribution consists of two normal distributions N(*m*, *s*_{1}) and N(*m*, *s*_{2}) truncated at their means “*m*” and then combined in such a way, that points of truncation now become the new distribution mode “*m*_{0}“ (Fig. 1). The probability density function of this distribution, published by Siekierski (1991), has the following form:

where *x* is a random variable, *m*_{0} is the mode of the distribution, *s*_{1} is the standard deviation of the left half of the distribution, and *s*_{2} is the standard deviation of the right half of the distribution.

The cumulative distribution function (cdf) of the double normal distribution is not available in a closed form because Eq. 1 is not analytically integrable. However, values of the cdf can be determined numerically, as in the normal distribution, using the value of the “error function” (erf), and the following equations:

where = *F*(*x* = *m*_{0}).

In the study the diameter distribution data of 20 well-stocked, pure, even-aged Scots pine (*Pinus sylvestris *L.) stands was provided by the Department of Dendrometry and Forest Productivity at the Faculty of Forestry in Warsaw, Poland. The sample plots were located on typical pine sites (coniferous forest habitat) in Bory Tucholskie in north-central Poland. Areas of plots were set such that they comprised a few hundred trees. The diameters at breast height (dbh) of all trees were measured on the sample plots with an accuracy of 1 mm. Each plot was characterized by the following attributes: i) average total age, determined by counting rings on stumps of felled trees and sample cores; ii) average height, defined using the Lorey formula (basal area weighted average height); iii) site index, defined as the average height of the 100 thickest trees per hectare at base age 100 years, and calculated using the site index model for Scots pine in Poland (Bruchwald et al. 2000; Cieszewski and Zasada 2002); iv) basal area (BA), mean diameter (MD) and quadratic mean diameter (QMD). Table 1 summarizes the univariate statistics for the data used in the analysis.

Table 1. Summary statistics of the 20 sample plots used in the study. | ||||||||

Area | Age | N | QMD | MD | H_{L} | BA | SI | |

[ha] | [years] | [trees] | [cm] | [cm] | [m] | [m^{2 }ha^{–1}] | [m] | |

Minimum | 0.049 | 26 | 258 | 6.2 | 5.9 | 7.70 | 22.40 | 16.90 |

Maximum | 0.810 | 123 | 500 | 28.7 | 28.3 | 23.20 | 31.20 | 27.20 |

Average | 0.297 | 58 | 378 | 15.5 | 15.0 | 14.34 | 27.18 | 22.03 |

Std dev. | 0.235 | 25 | 67 | 6.0 | 5.9 | 3.70 | 2.55 | 3.53 |

Notation: Area – size of the sample plot; Age – average stand age; N – number of trees per plot; QMD – quadratic mean diameter of the stand (calculated from BA and N); MD – mean diameter (arithmetic mean of breast height diameter); H_{L} – average stand height calculated using the Lorey formula (basal area weighted average height); BA – basal area per hectare; SI – base-age 100 site index (Bruchwald et al. 2000; Cieszewski and Zasada 2002). |

As it was already mentioned, so far, parameters of the double normal distribution have been estimated exclusively using the method of moments (Siekierski 1991). Moments are numerical characteristics of a probability distributions or population, and can be calculated as raw (simple, ordinary, crude) moments, calculated about the origin (zero), and central ones, calculated about the arithmetic mean (or other value). Since we usually don’t have information on an entire population, we use a sample drawn from the population to estimate population moments.

The k^{th} ordinary moment of the probability distribution is defined as:

The k^{th} simple sample moment *m _{k}* is defined as:

The k^{th} central moment of the probability distribution is defined as:

The k^{th} central sample moment *m _{k}* is defined as:

where *k* is an integer number ≥1, *E*(●) is an expected value, *X* is a random variable, *F*(*x*) is a probability density function, *x*_{i} are elements sampled from the population, *m* is the arithmetic mean, and *n* is the sample size (Wooldridge 2001; Vrbik 2005; Encyclopedia 2012).

Note, that the first raw moment is the mean, the second central moment equals variance, and third moment corresponds to the distribution skewness. Sample moments are function of means of various powers of *x*_{i} derived from a sample, thus they are random variables. Theoretical moments (moments of the probability distribution) are related to the population, so they are non-random functions of the variable distribution and they are calculated from its parameters. Each k^{th} central moment can be expressed as a function of raw moments of the order ≤ *k*.

The method of moments is a technique for obtaining estimators of the distribution parameters that is based on matching the sample moments with the corresponding distribution moments and solving for a system of *k* equations with k unknowns. The method of moments in some cases reveals limitations. For example, the estimates given by the method of moments can be outside of the parameter space, especially for small samples. Moreover, estimates by the method of moments don’t take into account all information included in the sample. The MOM produces in most cases unbiased estimators, however they may not be the most efficient ones.

An alternative to the method of moments is the maximum likelihood method, first described by Fisher in 1922. It is a general method of parameter estimation for the population attribute distribution that maximizes the likelihood function of the sample (Fisher 1922). The likelihood function L of the n-element sample *x*_{1}, *x*_{2}, ..., *x*_{n}, is the joint probability function p(*x*_{1}, *x*_{2}, ..., *x*_{n}), where *x*_{1}, *x*_{2}, ..., *x*_{n} are the independent instances of a discrete random sample or a joint probability density function f(*x*_{1}, *x*_{2}, ..., *x*_{n}), when *x*_{1}, *x*_{2}, ..., *x*_{n} are detailed values of the continuous random variable. Likelihood function L can be written in the following parametrical form:

where ψ is a complete set of the density function parameters to be found. This function determines the chance of an occurrence that ψ is a „true” set of unknown parameters in such a situation, when the exact series of results *x*_{1}, *x*_{2}, …, *x*_{n} was obtained in the sample. The best estimation of the parameters occurs for such a value of ψ that the likelihood function reaches its maximum. Such parameters are called maximum likelihood estimates or MLE. If the likelihood function can be differentiated, such conditions are fulfilled if:

, assuming that

In practice, the maximum likelihood method maximizes the log of the likelihood function or, equivalently, minimizes the negative log of the likelihood function, for which the extremes are the same as for the original likelihood function L:

assuming that

The set of parameters ψ is usually estimated by minimizing the negative log-likelihood using some iterative method available in most statistical packages and languages.

In the presented study the distribution parameters for the double normal and for the Weibull distributions were estimated using the maximum likelihood method implemented using an optimization based on Nelder-Mead and quasi-Newton algorithms. The alternative simulated annealing optimization algorithm has also been applied for small samples. The initial values for the double normal distribution were calculated based on the analysis of the complete data compared to the mean *m*_{0}, *s*_{1} and *s*_{2} parameter values estimated for each sample plot from repeated samples. The starting value of m_{0} was set to 90% of the sample mean, *s*_{1} to 70% of the standard deviation of the sample and *s*_{2} to the standard deviation of the sample increased by 30%. The initial parameters of the Weibull distribution were calculated based on the mean and standard deviation of the sample using the *weibullpar* function from the *mixdist* package of the R software (Macdonald 2010).

Parameters of the double normal distribution were also fit by means of the method of moments (Siekierski 1991). The following variants of the MOM were used: MOM3, using first three moments of the sample (Eqs. 9, 10 and 11), and MOM2, using two first moments of the sample (Eqs. 9 and 10) and the distribution mode estimated using the approximate formula: *m*_{0} *= m* – 4.8 · (*m* – *m*_{e}), where *m*_{e} equals to the median. First three moments were calculated according to the following equations (Siekierski 1991):

where *m*_{k} is a k^{th} moment, *m* corresponds to the arithmetic mean of the truncated normal distributions (*m*_{0} of the joined double normal distribution), and *s*_{j} equals to the standard deviation of the left and right tail of the double normal distribution.

Also, for comparison, parameters of the two-parameter Weibull distribution were fit by means of the method of moments using two first moments of the sample (MOM2W), as described e.g. by Al-Fawzan (2000), Gove 2003 or Lei (2008). The k^{th} moment of the Weibull distribution is expressed by the following equation:

where *b* – scale parameter, *c* – shape parameter of the distribution, Γ – gamma function. Thus, the first and the second moments of the distribution (*m*_{1} and *m*_{2}) were calculated as follows:

By dividing *m*_{2} (Eq. 14) by the square of *m*_{1} (Eq. 13), and taking the square root of this ratio, we obtain the coefficient of variation (CV), which relies exclusively on the *c* parameter:

Eq. 15 was solved using the root-finding bisection procedure (Forsythe et al. 1977) to obtain the estimate of* *. The scale parameter *b* was finally obtained using the following equation (Shifley and Lentz 1985; Gorgoso et al. 2007):

Parameters were also calculated using the simplified method of moments described by García (1981).

In all cases the goodness-of-fit was tested using the Kolmogorov-Smirnov test (D_{n} statistic).

Sample plots used in the study consisted of a few hundred trees. This fact dramatically limits practical conclusions obtained from analyses, since in forest inventory foresters usually deal with much smaller numbers of measurements per plot. To address this problem, analyses were also performed using random subsets of 10 and 50 measurements taken 100 times from each sample plot. Samples drawn for each plot were characterized using the number of failed estimations at 0.05 significance level and the average Kolmogorov-Smirnov D_{n} statistic.

All analyses were performed using programs written in the R language (Ihaka and Gentleman 1996, see an example code for the maximum likelihood method for fitting parameters of the double normal distribution and the method of moments for the parameters of the Weibull distribution in the Appendix).

When analyzing all trees from sample plots, the hypothesis that empirical distributions match distributions with fitted parameters could not be rejected in any analyzed case. The maximum likelihood method for the double normal distribution was superior to the both variants of the method of moments in 9 out of 20 analyzed plots, however the differences were very small, sometimes almost negligible. The method of moments for the Weibull distribution was superior to the maximum likelihood method in 14 out of 20 analyzed plots, however, as in the case of the double normal distribution, the differences were very small, sometimes almost negligible. Results obtained from the simplified method of moments by García (1981) were practically identical to the classical method (Eqs. 13–16), so they were not reported nor analyzed separately. The double normal distribution slightly overtook the Weibull distribution in 9 cases out of 20 plots (regardless the fitting method). The average values of the D_{n} statistic of the Kolmogorov-Smirnov test were slightly higher for the double normal distribution than for the Weibull distribution, which indicates a slightly poorer overall performance (overall fit) of this distribution. The best overall performance revealed the Weibull distribution with parameters fit using the method of moments. Detailed values of the D_{n} statistic for various methods of parameter fitting for the double normal distribution and the Weibull distribution are presented in Table 2. The typical graphs of the fitted distributions are presented in Figs. 2 and 3. In both cases the double normal distribution is more positively skewed and has higher maximum frequency for the modal value than the Weibull distribution. The double normal distribution is inferior to the Weibull distribution in cases when the distribution becomes symmetrical or negatively skewed.

Table 2. Detailed values of the D_{n} statistics for various distributions and methods of fitting. MLE denotes the maximum likelihood method, MOM3 – method of moments based on first three moments of the sample, MOM2 – method of moments based on first two moments of the sample and the distribution mode estimated using the approximate formula: mode = mean – 4.8 · (mean – median), MOM2W – method of moments based on first two moments of the sample. Lower value of the D_{n} statistics denote better fit. The best fit is marked with bold numbers. | |||||

Double normal | Weibull | ||||

Plot | MLE | MOM3 ^{a)} | MOM2 ^{a)} | MLE | MOM2W |

BT112 | 0.0356 | 0.0327 | 0.0327 | 0.0324 | 0.0317 |

BT113 | 0.0347 | 0.0285 | 0.0285 | 0.0294 | 0.0282 |

BT114 | 0.0452 | 0.0542 | 0.0542 | 0.0568 | 0.0613 |

BT115 | 0.0391 | 0.0490 | 0.0490 | 0.0373 | 0.0412 |

BT116 | 0.0385 | 0.0413 | 0.0413 | 0.0422 | 0.0428 |

BT117 | 0.0344 | 0.0355 | 0.0355 | 0.0380 | 0.0378 |

BT118 | 0.0453 | 0.0294 | 0.0294 | 0.0255 | 0.0249 |

BT119 | 0.0244 | 0.0399 | 0.0399 | 0.0384 | 0.0343 |

BT120 | 0.0572 | 0.0396 | 0.0396 | 0.0345 | 0.0321 |

BT121 | 0.0277 | 0.0441 | 0.0441 | 0.0379 | 0.0375 |

BT122 | 0.0317 | 0.0355 | 0.0355 | 0.0310 | 0.0310 |

BT123 | 0.0315 | 0.0310 | 0.0310 | 0.0212 | 0.0211 |

BT124 | 0.0253 | 0.0197 | 0.0197 | 0.0275 | 0.0271 |

BT125 | 0.0221 | 0.0261 | 0.0261 | 0.0474 | 0.0406 |

BT126 | 0.0394 | 0.0476 | 0.0476 | 0.0552 | 0.0462 |

BT127 | 0.0322 | 0.0283 | 0.0283 | 0.0264 | 0.0302 |

BT128 | 0.0340 | 0.0323 | 0.0323 | 0.0289 | 0.0298 |

BT129 | 0.0391 | 0.0349 | 0.0349 | 0.0397 | 0.0386 |

BT130 | 0.0361 | 0.0354 | 0.0354 | 0.0273 | 0.0277 |

BT131 | 0.0499 | 0.0390 | 0.0390 | 0.0315 | 0.0270 |

Avg. | 0.0362 | 0.0362 | 0.0362 | 0.0354 | 0.0346 |

a) Values of the D_{n} statistics for MOM2 and MOM3 are not identical, but the differences exist below the resolution of the presentation of results. |

As the sample size decreases, the average goodness-of-fit decreases (Tables 3 and 4). For medium-size samples of 50 trees the best results were obtained for the Weibull distribution with parameters fitted using the maximum likelihood method. The Weibull distribution was superior to the double normal distribution on average in 16 cases out of 20. The maximum likelihood method gave on average a better fit than the method of moments, however it occasionally failed to converge. For the double normal distribution the method of moments gave on average a better fit than the maximum likelihood method, and the MOM3 variant (parameters fitted using first three moments of the sample) was better than the MOM2 one (method based on two first moments of the sample and the distribution mode estimated using the approximate formula), but comparison of detailed values for particular plots is not clear. The detailed values of the D_{n} statistic and the number of cases when the hypothesis of matching empirical and theoretical distributions was rejected are presented in Table 3.

Table 3. Detailed values of the D_{n} statistics and number of cases when the hypothesis about matching the empirical and theoretical distributions was rejected for various distributions and methods of fitting for 100 samples of 50 trees. For explanations of the abbreviations see Table 2. MLE(S) – the maximum likelihood method with simulated annealing optimization algorithm. The best fit without MLE(S) method is marked with bold numbers, and with the MLE(S) – with underlined numbers. | ||||||||||||

Plot | Double normal | Weibull | ||||||||||

MLE | 0.05 | MLE(S) | 0.05 | MOM3 | 0.05 | MOM2 | 0.05 | MLE | 0.05 | MOM2W | 0.05 | |

BT112 | 0.0973 | 7 | 0.0753 | 0 | 0.0811 | 0 | 0.0890 | 0 | 0.0674 | 0 | 0.0660 | 0 |

BT113 | 0.0967 | 8 | 0.0715 | 0 | 0.0698 | 0 | 0.0756 | 0 | 0.0670 | 0 | 0.0652 | 0 |

BT114 | 0.1054 | 7 | 0.0845 | 4 | 0.1533 | 0 | 0.0859 | 0 | 0.0855 | 1 | 0.0894 | 0 |

BT115 | 0.0906 | 2 | 0.0798 | 2 | 0.0985 | 0 | 0.0793 | 0 | 0.0802 | 0 | 0.0771 | 0 |

BT116 | 0.1329 | 12 | 0.0752 | 0 | 0.0733 | 0 | 0.0789 | 0 | 0.0670 | 0 | 0.0729 | 0 |

BT117 | 0.1352 | 14 | 0.0773 | 0 | 0.0867 | 0 | 0.0847 | 0 | 0.0757 | 0 | 0.0752 | 0 |

BT118 | 0.1168 | 9 | 0.0773 | 1 | 0.0691 | 0 | 0.0741 | 0 | 0.0697 | 0 | 0.0711 | 0 |

BT119 | 0.1078 | 6 | 0.0776 | 0 | 0.1251 | 0 | 0.0910 | 0 | 0.0789 | 0 | 0.0781 | 0 |

BT120 | 0.1563 | 18 | 0.0799 | 0 | 0.0877 | 0 | 0.0900 | 0 | 0.0758 | 0 | 0.0794 | 0 |

BT121 | 0.1193 | 12 | 0.0720 | 0 | 0.0753 | 0 | 0.0791 | 0 | 0.0687 | 1 | 0.0706 | 0 |

BT122 | 0.1738 | 20 | 0.0735 | 0 | 0.0785 | 0 | 0.0776 | 0 | 0.0660 | 0 | 0.0690 | 0 |

BT123 | 0.1794 | 22 | 0.0726 | 0 | 0.0890 | 0 | 0.0753 | 0 | 0.0627 | 0 | 0.0608 | 0 |

BT124 | 0.1350 | 15 | 0.0718 | 1 | 0.0978 | 0 | 0.0693 | 0 | 0.0703 | 2 | 0.0683 | 0 |

BT125 | 0.0756 | 2 | 0.0857 | 1 | 0.0743 | 0 | 0.0726 | 0 | 0.0768 | 1 | 0.0747 | 0 |

BT126 | 0.0825 | 1 | 0.0750 | 1 | 0.0896 | 0 | 0.0819 | 0 | 0.0889 | 2 | 0.0914 | 0 |

BT127 | 0.0838 | 2 | 0.0709 | 0 | 0.0783 | 0 | 0.0829 | 0 | 0.0723 | 0 | 0.0700 | 0 |

BT128 | 0.1013 | 5 | 0.0740 | 0 | 0.0778 | 0 | 0.0830 | 0 | 0.0684 | 0 | 0.0748 | 0 |

BT129 | 0.0971 | 5 | 0.0735 | 0 | 0.0855 | 0 | 0.0886 | 0 | 0.0769 | 1 | 0.0683 | 0 |

BT130 | 0.0946 | 5 | 0.0767 | 0 | 0.0697 | 0 | 0.0774 | 0 | 0.0716 | 0 | 0.0743 | 0 |

BT131 | 0.1133 | 8 | 0.0780 | 0 | 0.0912 | 0 | 0.0829 | 0 | 0.0727 | 0 | 0.0769 | 0 |

Avg. | 0.1147 | 0.0761 | 0.0876 | 0.0809 | 0.0731 | 0.0737 |

A further decrease in performance was observed for small samples of 10 trees (Table 4). In this case the worst appeared to be the double normal distribution, especially its variant with parameters fitted using the maximum likelihood method, which not only had the average value of the D_{n} statistic indicating lack of fit for 43% of samples at 0.05 significance level, but also in many cases did not reach the convergence of fitting. The Weibull distribution was able to provide a good fit to the data almost every time for each sample, and was followed by the method of moments for the double normal distribution, which did not provide good fitting only in a few cases. The average value of the D_{n} statistic for the double normal distribution and maximum likelihood method was more than two times higher than for the method of moments and for the Weibull function.

Table 4. Detailed values of the D_{n} statistics and number of cases when the hypothesis about matching the empirical and theoretical distributions was rejected for various distributions and methods of fitting for 100 samples of 10 trees. For explanations of the abbreviations see Table 2. The best fit is marked with bold numbers. | ||||||||||||

Plot | Double normal | Weibull | ||||||||||

MLE | 0.05 | MLE(S) | 0.05 | MOM3 | 0.05 | MOM2 | 0.05 | MLE | 0.05 | MOM2W | 0.05 | |

BT112 | 0.3519 | 45 | 0.1857 | 6 | 0.1678 | 1 | 0.1755 | 1 | 0.1472 | 0 | 0.1391 | 0 |

BT113 | 0.3210 | 37 | 0.2050 | 12 | 0.1555 | 2 | 0.1562 | 1 | 0.1387 | 0 | 0.1379 | 1 |

BT114 | 0.4111 | 53 | 0.2461 | 15 | 0.2076 | 4 | 0.1735 | 0 | 0.1577 | 0 | 0.1590 | 1 |

BT115 | 0.3605 | 39 | 0.1953 | 4 | 0.1823 | 2 | 0.1675 | 0 | 0.1482 | 0 | 0.1529 | 0 |

BT116 | 0.3478 | 40 | 0.1891 | 3 | 0.1745 | 2 | 0.1698 | 1 | 0.1394 | 0 | 0.1408 | 0 |

BT117 | 0.3947 | 50 | 0.2154 | 9 | 0.1894 | 4 | 0.1786 | 3 | 0.1528 | 2 | 0.1466 | 0 |

BT118 | 0.3348 | 38 | 0.1868 | 3 | 0.1530 | 2 | 0.1556 | 1 | 0.1478 | 0 | 0.1407 | 2 |

BT119 | 0.3604 | 44 | 0.2275 | 7 | 0.1867 | 3 | 0.1722 | 3 | 0.1536 | 0 | 0.1499 | 1 |

BT120 | 0.4047 | 50 | 0.2154 | 7 | 0.1698 | 3 | 0.1708 | 2 | 0.1510 | 1 | 0.1466 | 0 |

BT121 | 0.3522 | 42 | 0.2059 | 3 | 0.1690 | 1 | 0.1724 | 1 | 0.1290 | 0 | 0.1358 | 0 |

BT122 | 0.4264 | 56 | 0.1979 | 9 | 0.1782 | 2 | 0.1688 | 0 | 0.1473 | 0 | 0.1362 | 1 |

BT123 | 0.3514 | 39 | 0.2031 | 9 | 0.1742 | 1 | 0.1567 | 1 | 0.1419 | 0 | 0.1382 | 1 |

BT124 | 0.3946 | 50 | 0.1671 | 2 | 0.1657 | 1 | 0.1576 | 0 | 0.1325 | 0 | 0.1350 | 0 |

BT125 | 0.3530 | 40 | 0.1739 | 4 | 0.1642 | 2 | 0.1677 | 2 | 0.1611 | 0 | 0.1485 | 1 |

BT126 | 0.3227 | 36 | 0.1791 | 3 | 0.1806 | 3 | 0.1619 | 0 | 0.1636 | 0 | 0.1536 | 0 |

BT127 | 0.3840 | 49 | 0.1700 | 0 | 0.1451 | 0 | 0.1478 | 0 | 0.1565 | 0 | 0.1488 | 0 |

BT128 | 0.3413 | 38 | 0.1839 | 1 | 0.1616 | 5 | 0.1607 | 3 | 0.1626 | 1 | 0.1402 | 0 |

BT129 | 0.3259 | 35 | 0.1923 | 6 | 0.1644 | 2 | 0.1725 | 1 | 0.1648 | 0 | 0.1388 | 0 |

BT130 | 0.3322 | 38 | 0.1894 | 7 | 0.1716 | 2 | 0.1812 | 1 | 0.1668 | 1 | 0.1447 | 2 |

BT131 | 0.3324 | 36 | 0.1849 | 4 | 0.1669 | 1 | 0.1636 | 1 | 0.1586 | 2 | 0.1524 | 2 |

Avg. | 0.3602 | 0.1958 | 0.1714 | 0.1665 | 0.1511 | 0.1443 |

Maximum likelihood estimates can be heavily biased for small samples. Also, the parameter optimality property may not apply for small samples. In the presented study this was proven mostly for the double normal distribution, especially fitted to the data from 10-tree samples. Also, the maximum likelihood method appeared to be sensitive to the choice of starting values, again – mostly in the case of the double normal distribution. Even using starting values initially approximated by the method of moments did not assure a good fit to the data.

For big samples, the performance (measured by the D_{n} Kolmogorov statistic) of the analyzed distributions was comparable. The Weibull distribution was superior to the double normal one. The maximum likelihood method for the Weibull distribution appeared to be superior to the method of moments, which confirms results obtained by Shiver (1988). At the same time the MLM failed to converge in a few cases. After replacing the default Nelder-Mead (NM) optimization method with the simulated annealing (SANN) approach (chosen after checking also other possible fitting methods such as quasi-Newton (qN), gradient and box constraints) the overall D_{n} statistics did not improve, but the algorithm achieved the convergence in most cases. The maximum likelihood method of parameter estimation applied to the double normal distribution always gave the poorest performance. This was a surprising outcome taking into account results obtained from the Weibull distribution. After detailed check of the parameters it was noticed that worse overall fit statistics are caused mainly by the unrealistic parameter values which denote the failure of the fitting algorithm. Changing initial parameters did not improve fitting statistics, but optimization algorithm replaced with the SANN method gave desirable results: average fit statistics as well as the number of cases with unrealistic values decreased. The maximum likelihood method for the double normal distribution gave in this case better results than the method of moments, but worse than both analyzed methods of parameter calculations for the Weibull distribution. It outperformed the Weibull distribution only in the case of two plots (see values in the MLE(S) column in Table 3).

The trends of decreasing goodness-of-fit and increasing number of failed fits continued for smaller samples and finally practically disqualified the maximum likelihood method of parameter estimation for the double normal distribution. It was not possible to obtain a fit of the distribution to the data for most of the analyzed plots. Much more robust appeared to be the method of moments, which was able to provide satisfactory fits for medium-size, and also in many cases for the small-size samples.

In the case of the samples consisting of 10 element, poor performance of the maximum likelihood method applied to the double normal distribution can be caused by such factors as: small sample size, giving not enough information to fit the correct parameters, wrong choice of the starting parameters, failure in finding reasonable values of parameters and the optimization method applied in the MLM. In order to find the source of the problems, the extensive analysis was done, following the scheme described for 50-element samples.

First, the detailed values of parameters were checked. In many cases their values were clearly wrong (e.g. negative means and standard deviations), which denotes problems with the fitting process. In order to check whether the problem exists because of the inappropriate starting values of parameters, they were replaced with the values i) calculated using the method of moments from the sample and ii) calculated using the method of moments from the complete plot data. None of the approaches significantly changed the results: the average D_{n} statistics, as well as the number of failed fits, were the same.

Then, the influence of the optimization algorithm on the results was tested. The default NM and qN methods were replaced with the simulated annealing (SANN) method. The application of this algorithm lowered the average D_{n} statistics by about a half, but still placed the double normal distribution with parameters fitted using MLM behind the method of moments and the Weibull distribution. The number of failed fits (unrealistic parameters, mostly negative standard deviations of the left part of the distribution) significantly decreased (see values in the MLE(S) column in Table 4).

Clearly, one of the major factors influencing the results obtained for the double normal distribution analysis using small samples is the choice of the optimization method used in the MLM. Simulated annealing algorithm was much more resistant to the bad choice of starting values caused by a small sample size than the NM and qN. However, at the same time, the SANN approach appeared to be more time consuming than other methods. Another factor influencing the fitting process is the choice of the initial parameters. This is especially complicated process when small samples, providing very variable values, are taken into account. One of the solutions of this issue is the use of the method described by Mehtätalo et al. (2011).

Presented results discourage from using the maximum likelihood method of parameter fitting for the double normal distribution for small samples using NM and qN optimization methods. It can be used for larger samples, giving the fit comparable to the Weibull distribution, and providing, similarly to the Weibull distribution, parameters having sound practical and biological meaning. In case of smaller samples for the double normal distribution it is recommended to apply SANN optimization method, use the method of moments or substitute the described distribution with the more flexible and robust Weibull distribution.

The results presented in this study confirm that the most efficient procedure for fitting the parameters of the diameter distribution is to use the maximum likelihood method with initial values of parameters calculated using the method of moments from the sample data. This approach has proved to be useful for both analyzed distributions. It was also concluded that it is very important to choose a robust optimization algorithm of the applied MLM. This is especially important for small and medium samples, where SANN was much slower, but at the same time much more reliable than the NM and qN methods.

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#

# Sample R code for the double normal and Weibull distributions

#

# Probability density function of the 2-normal distribution

#

ddoublenormal <- function(x,m,s1,s2){

tty1 <- sqrt(2/pi)*(1/(s1+s2))*exp((-(x-m)^2)/(2*s1^2))

tty2 <- sqrt(2/pi)*(1/(s1+s2))*exp((-(x-m)^2)/(2*s2^2))

tty <- ifelse(x<=m,tty1,tty2)

return(tty)

}

#

# ML fitting using “optim” function

# for the double normal distribution

#

# x is a vector of diameter measurements

#

x <- (read.table(“bt112.dat”, header=F))$V1

loglike<-function(p) -2*sum(log(ddoublenormal(x,p[1],p[2],p[3])))

startm <- mean(x)*0.9

starts1 <- sd(x)*0.7

starts2 <- sd(x)*1.3

optim(c(startm,starts1,starts2),loglike)

#

# ML fitting can be alternatively done

# using “mle” function from the stats4 package

# “mle” calls “optim”, so results are identical

#

x <- (read.table(“bt112.dat”, header=F))$V1

loglike<-function(m,s1,s2) -2*sum(log(ddoublenormal(x,m,s1,s2)))

initial=list(m=mean(x)*0.9,s1=sd(x)*0.7,s2=sd(x)*1.3)

mle(loglike,start=initial)

#

# The method of moments for 2-parameter Weibull distribution

#

momweibull <- function(x) {

cv <- sd(x)/mean(x)

fun<-function(c) cv-sqrt(gamma(1+2/c)-(gamma(1+1/c))^2)/(gamma(1+1/c))

shape <- uniroot(fun, c(0.4, 5))$root

scale <- mean(x)/(gamma(1/shape+1))

list(shape=shape, scale=scale)

}

x <- (read.table(“bt112.dat”, header=F))$V1

momweibull(x)

#

# Modified simplified method of moments by Garcia (1981)

# http:// http://web.unbc.ca/~garcia/softw/weibullR.txt

#

mweibull <- function(x){

cv <- sd(x)/mean(x)

if (cv > 1.2 || cv <= 0)

stop(“Out of range”)

a <- 1/(cv*(((((0.007454537*cv*cv-0.08354348)*cv+0.153109251)*

cv-0.001946641)*cv-0.22000991)*(1-cv)*(1-cv)+1))

list(shape=a, scale=mean(x)/gamma(1+1/a))

}

x <- (read.table(“bt112.dat”, header=F))$V1

mweibull(x)

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Vrbik J. (2005). Population moments of sampling distributions. Computational Statistics 20: 611–621.