Alpo Luomajoki 
Kuusen sopeutuminen Suomen ilmastoon hedekukkimisaikojen perusteella arvioituna
Luomajoki A. (1993). Kuusen sopeutuminen Suomen ilmastoon hedekukkimisaikojen perusteella arvioituna. Acta Forestalia Fennica no. 242 article id 7504. https://doi.org/10.14214/aff.7504
English title: Climatic adaptation of Norway spruce (Picea abies (L.) Karsten) in Finland based on male flowering phenologyAbstract
Anthesis was studied at the canopy level in 10 Norway spruce (Picea abies (L.) H. Karst.) stands from 9 localities in Finland was studied in 1963-74. Distribution of pollen catches were compared with the normal Gaussian distribution. The basis for the timing studies was the 50% point of the anthesis-fitted normal distribution. Development was characterized in calendar days, in degree days (>5°C) and in period units. The count of each unit began on March 19 (included). Male flowering in Norway spruce stands was found to have more annual variation in quantity than in Scots pine (Pinus sylvestris L.) stands studied earlier.
Anthesis in spruce in Northern Finland occurred at a later date than in the south. The heat sums needed for anthesis varied latitudinally less in spruce than in pine. The variation of pollen catches in spruce increased towards north-west as in the case of Scots pine. In the unprocessed data, calendar days were found to be the most accurate forecast of anthesis in Norway spruce. Locally, the period unit could be a more accurate parameter for the stand average. However, on a calendar day basis, when annual deviations between expected and measured heat sums were converted to days, period units were narrowly superior to days.
The geographical correlations respected to timing of flowering, calculated against distances measured along simulated post-glacial micgation routes, were stronger than purely latitudinal correlations. Effects of the reinvasion of Norway spruce into Finland are thus still visible in spruce populations just as they were in Scots pine populations.
The proportion of the average annual heat sum needed for spruce anthesis grew rapidly north of a latitude of ca. 63° and the heat sum needed for anthesis decreased only slightly towards the timberline. In light of flowering phenology, it seems probable that north-western third of Finnish Norway spruce populations are incompletely adapted to the prevailing cold climate. A moderate warming of the climate would therefore be beneficial for Norway spruce. This accords roughly with the adaptive situation in Scots pine
The PDF includes a summary in Finnish.
Keywords
Norway spruce;
Picea abies;
adaptation;
anthesis;
flowering phenology;
pollen recording;
heat sum;
reinvasion
Published in 1993
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