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Silva Fennica vol. 43 | 2009

Category : Research article

article id 460, category Research article

Hong Ling, Sandhya Samarasinghe, G. Don Kulasiri. (2009). Modelling variability in full-field displacement profiles and Poisson ratio of wood in compression using stochastic neural networks. Silva Fennica vol. 43 no. 5 article id 460. https://doi.org/10.14214/sf.460

Keywords: wood; digital image correlation; displacement profiles; variability; micro structure; stochastic neural networks; Poisson ratio

Abstract | View details | Full text in PDF | Author Info

Vertical and horizontal displacement profiles in compression parallel-to-grain in a 20 x 20 mm area (30 x 21 or 630 points) in the Tangential–Longitudinal (T–L) and Radial Longitudinal (R–L) sections of small wood columns were obtained from digital image correlation applied to simultaneously captured images of the two surfaces. These consisted of 21 displacement realisations of 30 points along the length of the specimen. They revealed considerable local variations. Stochastic neural networks were successfully developed to simulate trends and noise across and along a specimen in both displacements as well as Poisson ratios in T–L and R–L sections for two selected load levels of 20kN and 40kN. These networks specifically embed noise characteristics extracted from data to generate realistic displacement and Poisson ratio realisations with inherent variability. Models were successfully validated using independent data extracted based on bootstrapping method with high accuracy with R² ranging from 0.79 to 0.91. The models were further validated successfully using a second approach involving Confidence Intervals generated from the data extracted from the models. Models and experimental results revealed that for 20kN load, both vertical and horizontal displacements in T–L section were less heterogeneous across the specimen (smaller vertical shearing and horizontal strain, respectively) than those in the R–L section. For the 40kN load, both displacement profiles in the T–L section were less noisy and more compact than those for the 20kN load indicating less heterogeneity due to compaction of structure. In the R–L section, larger vertical shearing and horizontal strains persisted at 40 kN load. Poisson ratio decreased with load and it was nonlinear in both sections but T–L section showed much less noise across the specimen than the R–L section.

Ling, Centre for Advanced Computational Solutions (C-fACS), Lincoln University, Canterbury, New Zealand E-mail: hl@nn.nz
Samarasinghe, Centre for Advanced Computational Solutions (C-fACS), Lincoln University, Canterbury, New Zealand E-mail: sandhya.samarasinghe@lincoln.ac.nz
Kulasiri, Centre for Advanced Computational Solutions (C-fACS), Lincoln University, Canterbury, New Zealand E-mail: gdk@nn.nz

article id 178, category Research article

Raili Hokajärvi, Teppo Hujala, Leena A. Leskinen, Jukka Tikkanen. (2009). Effectiveness of sermon policy instruments: forest management planning practices applying the activity theory approach. Silva Fennica vol. 43 no. 5 article id 178. https://doi.org/10.14214/sf.178

Keywords: forest policy; forest planning; activity theory; advisory; extension; non-industrial private forests; qualitative study

Abstract | View details | Full text in PDF | Author Info

Recent and ongoing societal changes have brought about a need to foster multiple-use forestry and to strengthen customer orientation in family forestry outreach. The study assesses how forest management planning in family forest holdings could be developed to tackle these challenges. The approach introduces a new way of evaluating the effectiveness of information- and communication-based policy instruments. Here, the cultural-historical activity theory is applied in studying the interwoven practices of present-day planning and the associated advisory services targeted at landowners. The data, comprising semi-structured in-depth interviews with 19 professional planners, were qualitatively examined, and a forest management planning activity model was constructed with the emphasis placed on the inherent contradictions of planning work. As the main contradiction, the forest and the forest owner compete as objects. The aims of making the forest productive and advising the landowners towards an increased activeness motivate forest management planning but the planners feel that they lack the opportunity to respond to the needs of the landowner. A wood-production-emphasizing interpretation of the benefits to the national economy frustrates the policy goal of genuinely promoting the goals of multiple-use forestry. The conclusion drawn is that the actors engaged in forest management planning can reveal the needs for change by discussing their opinions and practical innovations. This can be done with the aid of facilitation by e.g. researchers oriented to developmental work study.

Hokajärvi, Oulu University of Applied Sciences, School of Renewable Natural Resources, Metsäkouluntie 4–6, FI-90650 Oulu, Finland E-mail: raili.hokajarvi@oamk.fi
Hujala, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: th@nn.fi
Leskinen, University of Eastern Finland, Department of Geographical and Historical Studies, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: lal@nn.fi
Tikkanen, Oulu University of Applied Sciences, School of Renewable Natural Resources, Metsäkouluntie 4–6, FI-90650 Oulu, Finland E-mail: jt@nn.fi

article id 177, category Research article

Tuomo Nurminen, Heikki Korpunen, Jori Uusitalo. (2009). Applying the activity-based costing to cut-to-length timber harvesting and trucking. Silva Fennica vol. 43 no. 5 article id 177. https://doi.org/10.14214/sf.177

Keywords: logistics; bucking; cutting; forest transport; long-distance transportation; trucking; time consumption; timber assortment; cost driver

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The supply chain of the forest industry has increasingly been adjusted to the customer’s needs for precision and quality. This has changed the operative environment both in the forest and on the roads. As the total removal of timber is increasingly divided into more log assortments, the lot size of each assortment decreases and the time consumed in sorting the logs increases. In this respect, the extra assortments have made harvesting work more difficult and affected the productivity of both cutting and forest transport; this has thus increased the harvesting costs. An activity-based cost (ABC) management system is introduced for timber harvesting and long-distance transport, based on the cut-to-length (CTL) method, in which the logistic costs are assigned to timber assortments and lots. Supplying timber is divided into three main processes: cutting, forest transport, and long-distance transportation. An ABC system was formulated separately for each of these main operations. Costs were traced to individual stands and to timber assortment lots from a stand. The cost object of the system is thus a lot of timber that makes up one assortment that has been cut, forwarded, and transported from the forest to the mill. Application of the ABC principle to timber harvesting and trucking was found to be relatively easy. The method developed gives estimates that are realistic to actual figures paid to contractors. The foremost use for this type of costing method should be as a tool to calculate the efficiency of an individual activity or of the whole logistic system.

Nurminen, Metsätoimisto Tuomo Nurminen, Joensuuntie 5 B 8, FI-41800 Korpilahti, Finland E-mail: tn@nn.fi
Korpunen, Finnish Forest Research Institute, Parkano Research Unit, Kaironiementie 54, FI-39700 Parkano, Finland E-mail: hk@nn.fi
Uusitalo, Finnish Forest Research Institute, Parkano Research Unit, Kaironiementie 54, FI-39700 Parkano, Finland E-mail: ju@nn.fi

article id 176, category Research article

María Pasalodos-Tato, Timo Pukkala, Antonio Rigueiro-Rodríguez, Esther Fernández-Nunez, María Rosa Mosquera-Losada. (2009). Optimal management of Pinus radiata silvopastoral systems established on abandoned agricultural land in Galicia (north-western Spain). Silva Fennica vol. 43 no. 5 article id 176. https://doi.org/10.14214/sf.176

Keywords: simulation; economic profitability; optimization; risk reduction; salvage

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Timber production has been the main objective in forest production in Galicia for a long time. Nevertheless, factors such as fire risk and the need to obtain non-timber benefits make other production alternatives like silvopastoral systems worth of consideration. Integration of grazing in the production system not only diversifies products and benefits, but also decreases fire risk by enhancing fuel control. Nonetheless, few studies have examined the economic profitability of these systems. This article analyses the economics of silvopastoral systems established on abandoned agricultural soils afforested with Pinus radiata D. Don. Different tree planting densities, discounting rates, grass values and fire risk scenarios were analysed. The technique employed is based on the combination of an optimization algorithm and a simulator of stand growth and grass yield. The most profitable schedules were obtained with initial stand densities of 1500 trees per hectare. However, with high unit values of pasture production (high value of grass), schedules with an initial stand density of 500 trees per hectare were the most profitable. When the risk of fire was included in the analyses, silvopastoral systems were always more profitable than timber production systems. With an assumption that grazing reduces fire risk thinnings should be done earlier and heavier to reduce the expected losses due to fire and to promote grass production. This lengthens the pasture period. In general, rotation lengt

Pasalodos-Tato, INIA, Instituto Nacional de Investigación y Tecnología Agraria y Alimentaria. Madrid, Spain E-mail: pasalodos.maria@inia.es
Pukkala, University of East Finland, Joensuu, Finland E-mail: tp@nn.fi
Rigueiro-Rodríguez, University of Santiago de Compostela, Lugo, Spain E-mail: arr@nn.es
Fernández-Nunez, University of Santiago de Compostela, Lugo, Spain E-mail: efn@nn.es
Mosquera-Losada, University of Santiago de Compostela, Lugo, Spain E-mail: mrml@nn.es

article id 175, category Research article

Heikki Hänninen, Jaana Luoranen, Risto Rikala, Heikki Smolander. (2009). Late termination of freezer storage increases the risk of autumn frost damage to Norway spruce seedlings. Silva Fennica vol. 43 no. 5 article id 175. https://doi.org/10.14214/sf.175

Keywords: Norway spruce; simulation; autumn frost damage; day degrees; freezer storage; growing season

Abstract | View details | Full text in PDF | Author Info

Over the last few years it has become increasingly common in artificial forest regeneration to extend the planting period by using freezer-stored seedlings for early summer plantings. Developmentally, however, planted freezer-stored seedlings lag behind seedlings planted earlier in the spring. As freezer-stored seedlings also start hardening later, they are more susceptible to early autumn frosts, especially in years when the thermal growing season ends and the first autumn frosts come earlier than usual. By means of computer simulations with a simple temperature sum model and long-term air-temperature data from three locations in Finland, we examined the effect of the freezer-storage termination date on the risk of autumn frost damage to the seedlings. The long-term simulations revealed a drastic effect of year-to-year variation in the thermal conditions during the growing season on the occurrence of autumn frost damage. Such results provide crucial information complementary to those obtained in field experiments, which are always restricted to a relatively short time period. Together with earlier field data, the present results suggest that at an average regeneration site in central Finland, the planting of seedlings whose storage has terminated on 15 June and 22 June involve autumn frost damage every tenth and every fifth year, respectively. The sensitivity analysis revealed that the temperature sum requirement of maturation has a great effect on the risk of autumn frost damage, thus pinpointing the need for experimental studies addressing this ecophysiological trait of the seedlings.

Hänninen, Plant Ecophysiology and Climate Change Group (PECC), Department of Biological and Environmental Sciences, Box 65, FI-00014 University of Helsinki, Finland E-mail: heikki.hanninen@helsinki.fi
Luoranen, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: jl@nn.fi
Rikala, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: rr@nn.fi
Smolander, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: hs@nn.fi

article id 174, category Research article

Akihiro Sumida, Taro Nakai, Masahito Yamada, Kiyomi Ono, Shigeru Uemura, Toshihiko Hara. (2009). Ground-based estimation of leaf area index and vertical distribution of leaf area density in a Betula ermanii forest. Silva Fennica vol. 43 no. 5 article id 174. https://doi.org/10.14214/sf.174

Keywords: allometry; leaf area density; LAI; leaf inclination angle; MacArthur–Horn method; pipe model; Betula ermanii

Abstract | View details | Full text in PDF | Author Info

We developed a ground-based method for estimating leaf area index (LAI) and vertical distribution of leaf area density (LAD) for two Betula ermanii plots, combining an allometric method for tree leaf area with the MacArthur–Horn (MH) method using a portable laser rangefinder, including a correction for changes in leaf inclination angle along the vertical gradient measured with a portable digital protractor from a canopy access tower in each plot. Vertical distribution of projected leaf area density obtained by the MH method (LADMH) was transformed to relative distribution for allotting fixed LAI to different heights. Hence, we first developed an allometric method for estimating tree leaf area for LAI determination. Trunk cross-sectional area at branching height (AB) was accurately estimated (r² = 0.97) from ground-based measurements of tree dimensions. We used this method to apply pipe model allometry between tree leaf area and AB, and estimated LAI (4.56 and 4.57 m² m^–2). We then examined how leaf inclination angle affected estimation of the vertical distribution of actual LAD. Leaf inclination angle measurements revealed that actual LAD in the upper canopy was 1.5–1.8-times higher than LADMH, because of steep leaf inclination, while the correction factor was 1.15–1.25 in the lower canopy. Due to the difference among heights, vertical distribution of LAD estimated with correction for vertical change in leaf inclination was more skewed to the upper canopy than that without correction. We also showed that error in LAD distribution can result if horizontal canopy heterogeneity is neglected when applying the MH method.

Sumida, Institute of Low Temperature Science, Hokkaido University, N19W8, Sapporo 060-0819, Japan E-mail: asumida@lowtem.hokudai.ac.jp
Nakai, International Arctic Research Center, University of Alaska Fairbanks, 930 Koyukuk Drive, P.O. Box 757340, Fairbanks, Alaska 99775-7340, USA E-mail: tn@nn.jp
Yamada, International Meteorological & Oceanographic Consultants Co., Ltd. Kawaguchi-cho 2-6528-87, Choshi, Chiba 288-0001, Japan E-mail: my@nn.jp
Ono, Institute of Low Temperature Science, Hokkaido University, N19W8, Sapporo 060-0819, Japan E-mail: ko@nn.jp
Uemura, Field Science Center for Northern Biosphere, Hokkaido University, Tokuda 250, Nayoro, Hokkaido 096-0071, Japan E-mail: su@nn.jp
Hara, Institute of Low Temperature Science, Hokkaido University, N19W8, Sapporo 060-0819, Japan E-mail: th@nn.jp

article id 173, category Research article

Erik Hellberg, Torbjörn Josefsson, Lars Östlund. (2009). The transformation of a Norway spruce dominated landscape since pre-industrial times in northern Sweden: the influence of modern forest management on forest structure. Silva Fennica vol. 43 no. 5 article id 173. https://doi.org/10.14214/sf.173

Keywords: birch; disturbance dynamics; forest history; deciduous forest; fire

Abstract | View details | Full text in PDF | Author Info

Logging history and the study of reference conditions in Scandinavian boreal forests has tended to focus on Scots pine dominated ecosystems. This paper presents a regional study of pre-industrial forest conditions and examines the effects of the industrial exploitation of ecosystems dominated by Norway spruce in northern Sweden. Historical records covering a period which preceded industrial logging in the study area (1917–1927) were used to obtain quantitative data on forest structure and influence of forest fires. These data were compared with a modern data set (2003) to analyse changes due to the industrial transformation of the forest. The early 20th century landscape was dominated by old, multi-cohorted spruce forests and mixed coniferous forests. It was found that fire affected both the structure and composition of the landscape. In post-burnt areas, even-aged forests dominated by deciduous species were the principal forest type. Between the early and modern data sets, profound changes in tree-species composition and age structure were documented. While the total volume of deciduous species increased substantially, the coverage of forests dominated by deciduous species decreased. There was also a significant increase in pine-dominated forests and in the total volume of pine. The industrial transformation of the studied landscapes has had profound effects on the structure of spruce forests, but much less so on deciduous forests. The study concludes that the present forest structure is a function of past management regimes, and that future transformations of the landscape will continue, thus affecting the natural variability and biodiversity of the forests.

Hellberg, Tunstigen 10, SE-831 43 Östersund, Sweden E-mail: eh@nn.se
Josefsson, Swedish University of Agricultural Sciences, Department of Forest Ecology and Management, SE-901 83 Umeå, Sweden E-mail: torbjorn.josefsson@svek.slu.se
Östlund, Swedish University of Agricultural Sciences, Department of Forest Ecology and Management, SE-901 83 Umeå, Sweden E-mail: lo@nn.se

article id 172, category Research article

Guntis Brumelis, Maris Strazds, Zanna Eglava. (2009). Stand structure and spatial pattern of regeneration of Pinus sylvestris in a natural treed mire in Latvia. Silva Fennica vol. 43 no. 5 article id 172. https://doi.org/10.14214/sf.172

Keywords: Pinus sylvestris; regeneration; natural forest; mire

Abstract | View details | Full text in PDF | Author Info

We examined the regeneration patterns of Pinus sylvestris L. in a natural treed mire in the hemiboreal zone in Latvia. Data on tree stem age and size was collected in 207 fine-scale plots (10 m²) and 4 medium-scale plots (400 m²). Size structure of living and dead trees was also estimated on transects 180-m and 250-m length and 10-m width. In addition, the vegetation was described in 1-m² plots to determine preferred microsites for P. sylvestris establishment. Pinus sylvestris showed continuous regeneration by an inverse J-shaped age and size structure. Pulses of mortality induced by fire were also evident. Regeneration of P. sylvestris was mostly on Sphagnum magellanicam Brid. hummocks free of competition from Ledum palustre L. and graminoids, which were found in small interspersed patches throughout the mire. The spatial pattern of tree cohorts differed, probably due to changing moisture conditions altering seedbed conditions and by fire-induced mortality in specific areas. Recent rapid invasion by Betula pubescens Ehrh. and Picea abies (L.) Karst. in a post-fire period since 1971 is probably associated with drier conditions.

Brumelis, Faculty of Biology, University of Latvia, Kronvalda bulv. 4, Riga LV1586, Latvia E-mail: guntis.brumelis@lu.lv
Strazds, Faculty of Biology, University of Latvia, Kronvalda bulv. 4, Riga LV1586, Latvia E-mail: ms@nn.lv
Eglava, Faculty of Biology, University of Latvia, Kronvalda bulv. 4, Riga LV1586, Latvia E-mail: ze@nn.lv

article id 171, category Research article

Takuo Nagaike. (2009). Snag abundance and species composition in a managed forest landscape in central Japan composed of Larix kaempferi plantations and secondary broadleaf forests. Silva Fennica vol. 43 no. 5 article id 171. https://doi.org/10.14214/sf.171

Keywords: snags; long-rotation plantation; plantation age; species composition

Abstract | View details | Full text in PDF | Author Info

Larix kaempferi is the main plantation species in the low-snow, cool-temperate zone of Japan. I studied L. kaempferi plantations of various stand ages in central Japan to examine and compare the effect of stand age on the abundance, size, and species composition of snags (standing dead trees) compared to those in secondary broadleaf forests. Plantations that were older than the standard rotation age had more and larger snags than young plantations, and the species diversity of snags was positively correlated with stand age. Because the density of living planted L. kaempferi showed little correlation with snag variables, whereas that of naturally regenerated tree species was positively correlated with snag variables, the density dependence of snag occurrence was stronger in naturally regenerated trees than in planted L. kaempferi. Snag species that were positively correlated with stand age were the main species in secondary broadleaf forests in this area. Basal area, density, and number of species of snags in standard-rotation plantations were significantly lower than in long-rotation plantations and secondary broadleaf forests. Long-rotation plantations are useful for retaining snags compared to standard-rotation plantations.

Nagaike, Yamanashi Forest Research Institute, Masuho, Yamanashi 400-0502, Japan E-mail: nagaike-zty@pref.yamanashi.lg.jp

article id 170, category Research article

Ryoichi Doi, Senaratne L. Ranamukhaarachchi. (2009). Community-level physiological profiling in monitoring rehabilitative effects of Acacia auriculiformis plantation on degraded land in Sakaerat, Thailand. Silva Fennica vol. 43 no. 5 article id 170. https://doi.org/10.14214/sf.170

Keywords: multivariate analysis; Acacia auriculiformis; community-level physiological profile; land degradation and rehabilitation; soil bacterial community

Abstract | View details | Full text in PDF | Author Info

This study was conducted to investigate the rehabilitative effects of planting Acacia auriculiformis trees on degraded land by observing variations in soil bacterial community profiles provided by community-level physiological profiling. Soil bacterial and physicochemical comparisons between an original evergreen forest and the Acacia plantation plot, established on an area severely degraded as a result of deforestation, showed that most soil characteristics were rehabilitated 18 to 19 years after the plantation of Acacia according to single variables, Shannon and Simpson diversity indices based on the community-level physiological profiles, principal component analysis and redundancy analysis. However, a more strict statistical comparison, discriminant analysis, completely discriminated between the Acacia plantation and the evergreen forest soils when the community-level physiological profiles were compared. Thus, the Acacia plantation soil was shown to still be in the process to full recovery. Here, we discuss the relevance of planting A. auriculiformis in land rehabilitation schemes in savanna regions.

Doi, AFE Building, School of Environment, Resources and Development, Asian Institute of Technology, Pathumthani 12120, Thailand E-mail: roird@aeiou.pt
Ranamukhaarachchi, AFE Building, School of Environment, Resources and Development, Asian Institute of Technology, Pathumthani 12120, Thailand E-mail: slr@nn.th

article id 169, category Research article

Susanne von Bargen, Elise Grubits, Risto Jalkanen, Carmen Büttner. (2009). Cherry leaf roll virus – an emerging virus in Finland? Silva Fennica vol. 43 no. 5 article id 169. https://doi.org/10.14214/sf.169

Keywords: Betula pendula; Betula pubescens; Betula nana; IC-RT-PCR; 3' non-coding region; CLRV; phylogenetic relationship

Abstract | View details | Full text in PDF | Author Info

Cherry leaf roll virus, CLRV, is a plant pathogen that infects a variety of deciduous trees and shrubs in temperate regions. Little is known about its occurrence at high latitudes and especially in Finnish birch species. Still, symptoms that seemed to be associated with CLRV such as vein banding, leaf roll and decline have been observed in birch trees throughout the country since the summer of 2002. Six different birch species, subspecies or varieties, i.e. Betula pubescens subsp. pubescens (downy birch), B. pendula (silver birch), B. nana (dwarf birch), B. pubescens var. appressa (Kiilopää birch), B. pubescens subsp. czerepanovii (mountain birch) and B. pendula var. carelica (curly birch) originating from all over Finland were assessed by immunocapture-reverse transcription-polymerase chain reaction (IC-RT-PCR) for CLRV infection. It was shown that CLRV is widely distributed in B. pendula and B. pubescens throughout the country. Furthermore, dwarf birch, mountain birch, Kiilopää birch and curly birch were confirmed to be previously unkown hosts of CLRV. Genetic analysis of virus sequence variants originating from Finnish birch trees revealed atypical phylogenetic relationships. In contrast to CLRV isolates from birches growing in the United Kingdom and Germany which clustered exclusively within group A, Finnish CLRV isolates belonged either to group B, D or E. Thus, virus population structure in Finnish birches seems to be more variable and host plant dependency seems not to apply for Finnish CLRV isolates.

Bargen, Humboldt-Universität zu Berlin, Department für Nutzpflanzen- und Tierwissenschaften, Fachgebiet Phytomedizin, Lentzeallee 55/57, 14195 Berlin, Germany E-mail: susanne.von.bargen@agrar.hu-berlin.de
Grubits, Humboldt-Universität zu Berlin, Department für Nutzpflanzen- und Tierwissenschaften, Fachgebiet Phytomedizin, Lentzeallee 55/57, 14195 Berlin, Germany E-mail: eg@nn.de
Jalkanen, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: risto.jalkanen@metla.fi
Büttner, Humboldt-Universität zu Berlin, Department für Nutzpflanzen- und Tierwissenschaften, Fachgebiet Phytomedizin, Lentzeallee 55/57, 14195 Berlin, Germany E-mail: cb@nn.de

article id 168, category Research article

Saila Varis, Anne Pakkanen, Aina Galofré, Pertti Pulkkinen. (2009). The extent of south-north pollen transfer in Finnish Scots pine. Silva Fennica vol. 43 no. 5 article id 168. https://doi.org/10.14214/sf.168

Keywords: Pinus sylvestris; gene flow; adaptation; coniferous phenology; plant population biology; boreal forest dynamics

Abstract | View details | Full text in PDF | Author Info

In order to evaluate the possibility of long distance gene flow in Scots pine (Pinus sylvestris L.), we measured the amount and germinability of airborne pollen and flowering phenology in central, northern, and northernmost Finland during 1997–2000. Totally 2.3% of the detected germinable pollen grains were in the air prior to local pollen shedding. The mean number of germinable pollen grains m^–3 air per day was lower prior to local pollen shedding, but in the year 2000 there were more germinable pollen grains in the air of central study site prior to local pollen shedding. Prior to the onset of pollen shedding, 7.5% of female strobili which we observed were receptive. On average female strobili became receptive three days earlier than local pollen shedding started. During the period of pollen shedding in the central study site, we detected germinable airborne pollen in the northern site in years 1997, 1999 and 2000. At the northermost site, we detected germinable airborne pollen during the pollen-shedding period of the northern site in 2000. Our detection of germinable airborne pollen and synchrony of strobili maturation from south to north suggest that populations of Scots pine in central and northern Finland may provide genetic material to populations in northern and northernmost Finland, respectively.

Varis, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: saila.varis@metla.fi
Pakkanen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: ap@nn.fi
Galofré, Passeig de l’estació 21, 5-1, 43800 Valls, Tarragona, Spain E-mail: ag@nn.fi
Pulkkinen, Finnish Forest Research Institute, Haapastensyrjä Breeding Station, Karkkilantie 247, FI-12600 Läyliäinen, Finland E-mail: pp@nn.fi

article id 190, category Research article

Katri Luostarinen, Veikko Möttönen. (2009). Effect of felling season, storage and drying on colour of silver birch (Betula pendula) wood from four different growing sites. Silva Fennica vol. 43 no. 4 article id 190. https://doi.org/10.14214/sf.190

Keywords: birch; age; colour darkening; conventional drying; environmental factors; plantation; wood

Abstract | View details | Full text in PDF | Author Info

Darkening of birch wood during artificial drying is a significant problem regarding the use of its timber as raw material by the mechanical wood industry. In the future, an increasing proportion of birch timber will be obtained from plantation forests, which differ from natural forests in many respects. In this investigation sample boards of Betula pendula, both from naturally regenerated stands and plantations, were sawn into the dimensions used as raw material for parquet billets. Growing site, felling season, and storage of logs were taken into account as possible factors affecting wood colour changes during drying. The wood of birches from fertile plantations remained lighter-coloured during conventional drying than the wood of naturally regenerated birches from low- and medium-fertile stands. The reason may be the difference in tree age and growth rate between natural and planted stands. Thus, it could be beneficial to grow birch in fertile stands so that the trees reach log size as young as possible. The results of this study emphasise the good quality of the birch wood from planted stands compared to natural stands with regard to its colour.

Luostarinen, Faculty of Forest Sciences, University of Joensuu, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: katri.luostarinen@joensuu.fi
Möttönen, Department of Mechanical Engineering, Lappeenranta University of Technology, P.O. Box 20, FI-53851 Lappeenranta, Finland E-mail: vm@nn.fi

article id 189, category Research article

Lars Rytter, Gunnar Jansson. (2009). Influence of pruning on wood characters in hybrid aspen. Silva Fennica vol. 43 no. 4 article id 189. https://doi.org/10.14214/sf.189

Keywords: Populus tremula P. tremuloides; cicatrisation; faultless wood; twig angle; twig diameter

Abstract | View details | Full text in PDF | Author Info

Fast-growing hybrid aspens (Populus tremula L. x P. tremuloides Michx.) are currently of great interest in Sweden since they can produce biomass at high rates and, at the same time, can be used to produce higher value wood products. This study focuses on the effects of pruning hybrid aspen to improve its wood quality. About 50% of the trees in the experimental stand were pruned by removing twigs, at heights up to 4 m, when they were 7–8 years old. Ten years later, 20 pruned and 20 unpruned trees, representing four clones, were randomly selected. Ten knots or twig/stem junctions, respectively, per tree were exposed for inspection using a chain saw and examined. The results revealed that pruned trees cicatrised the knots within about three years and thereafter produced substantial amounts of faultless wood. In contrast, unpruned trees (which had retained almost 80% of their twigs, often as dry twigs with bark pockets) had produced small uneven amounts of quality wood. Removal of twigs with acute angles and/or large diameters resulted in greater colour defects and rot in annual rings outside the pruning position, but the time of cicatrisation was not significantly affected. The results show that pruning can be used to enhance the wood quality of hybrid aspen over a short time period, and that pruning should be performed early during the rotation period when branches are small, in order to minimize discolouration and rot in the new annual rings.

Rytter, The Forestry Research Institute of Sweden (Skogforsk), Ekebo 2250, SE-26890 Svalöv E-mail: lars.rytter@skogforsk.se
Jansson, The Forestry Research Institute of Sweden (Skogforsk), Uppsala Science Park, SE-751 83 Uppsala E-mail: gj@nn.se

article id 188, category Research article

Mikko Havimo, Juha Rikala, Jari Sirviö, Marketta Sipi. (2009). Tracheid cross-sectional dimensions in Scots pine (Pinus sylvestris) – distributions and comparison with Norway spruce (Picea abies). Silva Fennica vol. 43 no. 4 article id 188. https://doi.org/10.14214/sf.188

Keywords: Norway spruce; Scots pine; cell wall thickness; radial diameter; tangential diameter; distribution

Abstract | View details | Full text in PDF | Author Info

Cell wall thickness and tracheid radial and tangential diameter are important characteristics in papermaking. These fibre cross-sectional dimensions affect paper properties such as light scattering, and tear and tensile indexes. In the authors’ previous article, the mean values and distributions of tracheid cross-sectional dimensions were obtained for Norway spruce (Picea abies). This article characterises the cross-sectional tracheid properties of Scots pine (Pinus sylvestris) using exactly the same methodology as in the previous study on Norway spruce, which enables the comparison between the tree species. The distributions for Scots pine cell wall thickness and tracheid radial diameter were similar: a narrow peak due to earlywood tracheids, and a wide peak due to latewood tracheids. The tangential diameter distributions for Scots pine were very similar in both earlywood and latewood, having one wide peak. Also, the distributions in whole stem, top pulpwood and sawmill chip assortments were quite similar. The differences between Scots pine and Norway spruce tracheid cross-sectional dimensions were fairly marginal. This is at least the case when comparing large tracheid populations, in which differences tend to even out. The situation may be different on a more detailed level of observation, for example, when individual annual rings in the different tree species are compared.

Havimo, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: mikko.havimo@helsinki.fi
Rikala, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: jr@nn.fi
Sirviö, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: js@nn.fi
Sipi, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ms@nn.fi

article id 187, category Research article

Tobias Biechele, Leif Nutto, Gero Becker. (2009). Growth strain in Eucalyptus nitens at different stages of development. Silva Fennica vol. 43 no. 4 article id 187. https://doi.org/10.14214/sf.187

Keywords: growth stress; growth strain; Eucalyptus nitens; stages of development; sawnwood

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Eucalypts are renowned for their high growth stress levels. These stresses cause splitting, warping and dimensional instability when cutting, processing and drying the wood. In Chile, large Eucalyptus nitens plantations can be found, which, due to these problems, are scarcely utilised for solid wood products (veneer, sawn wood). This study aims to determine the factors influencing growth stress at different stages of tree’s development, and to identify whether the factors influencing growth stress change over time. In five stands of different ages, growth strain, as an indicator of growth stress, was measured at different tree heights with the single hole drilling method. The tree variables DBH, tree height, slenderness (height/diameter ratio) and crown parameters also were measured. A correlation analysis of tree variables and growth strains was undertaken. The results obtained indicate a high variability in growth strain values. It was concluded that growth strain is not correlated with a single growth parameter, but with a combination of factors that variously influence it at different ages and tree heights.

Biechele, University of Freiburg, Institute of Forest Utilisation and Work Science, Werthmannstr. 6, DE-79085 Freiburg, Germany E-mail: tobias.biechele@fobawi.uni-freiburg.de
Nutto, University of Freiburg, Institute of Forest Utilisation and Work Science, Werthmannstr. 6, DE-79085 Freiburg, Germany E-mail: ln@nn.de
Becker, University of Freiburg, Institute of Forest Utilisation and Work Science, Werthmannstr. 6, DE-79085 Freiburg, Germany E-mail: gb@nn.de

article id 186, category Research article

Juho Rantala, Pertti Harstela, Veli-Matti Saarinen, Leo Tervo. (2009). A techno-economic evaluation of Bracke and M-planter tree planting devices. Silva Fennica vol. 43 no. 4 article id 186. https://doi.org/10.14214/sf.186

Keywords: productivity; cost-efficiency; mechanization; planting cost

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Techno-economically reasonable mechanization of tree planting has proved to be a difficult task in the Nordic working conditions. Although planting machines and combinations of base machine and planting device have been developed since the 1970s, mechanized planting has not been cost-competitive to manual planting. The aim of this study was to find out work time distributions, productivities, costs and effects of different work difficulty factors on productivities and costs of the state-of-the-art Nordic planting devices, Swedish Bracke and Finnish M-Planter, and to compare the devices with each other. The theory of comparative time studies was the base for the experimental design of this study. In the average working conditions, productivity (E15) of M-Planter (236 seedlings/hour) was 36.0% higher than that of Bracke (174 seedlings/hour). Here, M-Planter performed planting work 23.4% cheaper than Bracke. However, the difference depended greatly on the working conditions; the more stones or stumps the smaller the difference, and the more slash the bigger the difference.

Rantala, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: juho.rantala@metla.fi
Harstela, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: ph@nn.fi
Saarinen, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: vms@nn.fi
Tervo, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: lt@nn.fi

article id 185, category Research article

Bianca N. I. Eskelson, Tara M. Barrett, Hailemariam Temesgen. (2009). Imputing mean annual change to estimate current forest attributes. Silva Fennica vol. 43 no. 4 article id 185. https://doi.org/10.14214/sf.185

Keywords: forest inventory and analysis; forest monitoring; national forest inventories; nearest neighbor imputation; Pacific Northwest; paneled inventory data

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When a temporal trend in forest conditions is present, standard estimates from paneled forest inventories can be biased. Thus methods that use more recent remote sensing data to improve estimates are desired. Paneled inventory data from national forests in Oregon and Washington, U.S.A., were used to explore three nearest neighbor imputation methods to estimate mean annual change of four forest attributes (basal area/ha, stems/ha, volume/ha, biomass/ha). The randomForest imputation method outperformed the other imputation approaches in terms of root mean square error. The imputed mean annual change was used to project all panels to a common point in time by multiplying the mean annual change with the length of the growth period between measurements and adding the change estimate to the previously observed measurements of the four forest attributes. The resulting estimates of the mean of the forest attributes at the current point in time outperformed the estimates obtained from the national standard estimator.

Eskelson, Oregon State University, Department of Forest Engineering, Resources and Management, 204 Peavy Hall, Corvallis, Oregon 97331, USA E-mail: bianca.eskelson@oregonstate.edu
Barrett, Oregon State University, Department of Forest Engineering, Resources and Management, 204 Peavy Hall, Corvallis, Oregon 97331, USA E-mail: tmb@nn.us
Temesgen, Oregon State University, Department of Forest Engineering, Resources and Management, 204 Peavy Hall, Corvallis, Oregon 97331, USA E-mail: ht@nn.us

article id 184, category Research article

Jaakko Repola. (2009). Biomass equations for Scots pine and Norway spruce in Finland. Silva Fennica vol. 43 no. 4 article id 184. https://doi.org/10.14214/sf.184

Keywords: pine; tree biomass; biomass equations; spruce

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In this study, biomass equations for the above- and below-ground tree components of Scots pine (Pinus sylvestris) and Norway spruce (Picea abies [L.] Karst.) were developed. The models were based on 908 pine trees and 613 spruce trees collected in 77 stands located on mineral soil, and represented a wide range of stand and site conditions in Finland. The whole data set consisted of three sub data sets: 33 temporary sample plots, five thinning experiments, and the control plots of 39 fertilization experiments. The biomass equations were estimated for the individual tree components: stem wood, stem bark, living and dead branches, needles, stump, and roots. In the data analysis, a multivariate procedure was applied in order to take into account the statistical dependence among the equations. Three multivariate models for above-ground biomass and one for below-ground biomass were constructed. The multivariate model (1) was mainly based on tree diameter and height, and additional commonly measured tree variables were used in the multivariate models (2) and (3). Despite the unbalanced data in terms of the response variables, the statistical method generated equations that enable more flexible application of the equations, and ensure better biomass additivity compared to the independently estimated equations. The equations provided logical biomass predictions for a number of tree components, and were comparable with other functions used in Finland and Sweden even though the study material was not an objective, representative sample of the tree stands in Finland.

Repola, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: jaakko.repola@metla.fi

article id 183, category Research article

Stella Bogino, María José Fernández Nieto, Felipe Bravo. (2009). Climate effect on radial growth of Pinus sylvestris at its southern and western distribution limits. Silva Fennica vol. 43 no. 4 article id 183. https://doi.org/10.14214/sf.183

Keywords: Scots pine; dendroclimatology; tree ring; pulse and interpulse

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The associations between tree-ring width and climatic variables, the tree age effect on climate-growth relationship and the drought index impact on radial growth of Scots pine (Pinus sylvestris L.) were studied in Mediterranean environments in the Iberian Peninsula. Dendrochronological techniques were applied to construct six tree-ring width chronologies for P. sylvestris. The association between tree growth and climate was analysed with correlation coefficients and bootstrapped response functions. Drought index (DRI) was used to detect the effects of aridity. Tree-ring width was positively correlated with rainfall in the growing season but this association stated earlier at the lowest altitudinal sites. The effects of temperature varied according to the site. The response to climatic variables depended on the age of the trees: more of the variability was explained by climatic variables in young stands than in old stands. The significant association between radial growth and DRI suggests that drought is a determining factor in the radial growth of P. sylvestris. Climate forecast scenarios show an increase in rainfall irregularity in the Mediterranean region so the differential tree response to rainfall at different elevations can be used to predict tree growth for determining silvicultural treatments.

Bogino, Universidad de Valladolid, Departamento de Producción Vegetal y Recursos Forestales, Joint Unit UVa-INIA ‘Sustainable Forest Management’, Avda. de Madrid 44, 34004 Palencia, Spain E-mail: sbogino@fices.unsl.edu.ar
Fernández Nieto, Universidad de Valladolid, Departamento de Producción Vegetal y Recursos Forestales, Joint Unit UVa-INIA ‘Sustainable Forest Management’, Avda. de Madrid 44, 34004 Palencia, Spain E-mail: mjfn@nn.es
Bravo, Universidad de Valladolid, Departamento de Producción Vegetal y Recursos Forestales, Joint Unit UVa-INIA ‘Sustainable Forest Management’, Avda. de Madrid 44, 34004 Palencia, Spain E-mail: fb@nn.es

article id 182, category Research article

Janne Miettinen, Pekka Helle, Ari Nikula, Pekka Niemelä. (2009). Changes in landscape-scale habitat selection of capercaillie (Tetrao urogallus) in managed north-boreal forest. Silva Fennica vol. 43 no. 4 article id 182. https://doi.org/10.14214/sf.182

Keywords: forestry; landscape composition; capercaillie; wildlife triangle

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We studied changes in habitat selection of capercaillie in northern Finnish boreal forests at two spatial scales during two time periods, 1989–1992 and 2000–2003. We studied capercaillie densities and their changes between the study periods in relation to the landscape class proportions within 3-km buffer zones around the wildlife triangle center points. Furthermore, we compared the landscape class proportions in 800-meter buffer zones around capercaillie wildlife triangle count observations and around the counted wildlife triangle transects using t-tests and compositional analysis. At the local population scale (3 km) the change in adult density between the study periods was associated positively with the proportion of young thinning stands in 2003 and reversely with the mature stand (1992 and 2003) and clear-cut (1992) proportions. Capercaillie juvenile and pooled densities during 2000–2003 were positively associated with the advanced thinning stand proportion in 2003. At the capercaillie home range scale (800 m) habitats were rich in mature stands during 1989–1992 in relation to available habitats, but not during 2000–2003 when young thinning stands were more abundant in relation to available habitats. Relatively young managed forests can be suitable for capercaillie, but mature managed forests as capercaillie habitats may have deteriorated between the study periods. Spatial planning may help to form suitable areas that are large enough for the species, but the highest potential may lay in the forest stand scale, where increased cover on the ground could promote the habitat quality.

Miettinen, Kankurinhaka 14, FI-90450 Kempele, Finland E-mail: janne.miettinen@rktl.fi
Helle, Finnish Game and Fisheries Research Institute, Tutkijantie 2 E, FI-90570 Oulu, Finland E-mail: ph@nn.fi
Nikula, Finnish Forest Research Institute, Rovaniemi Research Unit, Eteläranta 55, FI-99600 Rovaniemi, Finland E-mail: an@nn.fi
Niemelä, University of Turku, Department of Biology, FI-20014 University of Turku, Finland E-mail: pn@nn.fi

article id 181, category Research article

Jari Miina, Juha-Pekka Hotanen, Kauko Salo. (2009). Modelling the abundance and temporal variation in the production of bilberry (Vaccinium myrtillus L.) in Finnish mineral soil forests. Silva Fennica vol. 43 no. 4 article id 181. https://doi.org/10.14214/sf.181

Keywords: vegetation; generalized linear mixed model; heath forest

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Miina, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jari.miina@metla.fi
Hotanen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jph@nn.fi
Salo, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: ks@nn.fi

article id 180, category Research article

Hongzhang Kang, Björn Berg, Chunjiang Liu, Carl J. Westman. (2009). Variation in mass-loss rate of foliar litter in relation to climate and litter quality in Eurasian forests: differences among functional groups of litter. Silva Fennica vol. 43 no. 4 article id 180. https://doi.org/10.14214/sf.180

Keywords: carbon sequestration; climate; Eurasian forests; leaf litter; mass-loss; litter quality

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With a data set of litter decomposition collected by means of literature survey, our objectives are 1) to determine the differences in the variation in the first-year mass loss (%) of leaf litter with regard to climate and litter quality among different functional groups of tree species in Eurasian forests, and 2) to determine the difference in effect of mean annual temperature (°C), annual precipitation (dm), as well as concentration of nitrogen (%), and lignin (%) on first-year mass loss over a wide range in climate and litter quality. The main results are as follows. 1) The significant differences between litter types in the relationships between first-year mass loss and climatic factors plus litter quality revealed clearly different decomposition patterns over the continent. Thus, differences were found between coniferous and broadleaf litter, between deciduous broadleaf and evergreen broadleaf as well as between genera and even within a genus, viz. between deciduous and evergreen Quercus. 2) With a change in a relative unit of climate and litter quality variables, there were clear differences in effects of mean annual temperature, annual precipitation, and nitrogen on first-year mass loss for different functional groups of trees. 3) We identified some broadleaf litter species that decomposed to 100% in one year and thus did not contribute to carbon sequestration in a humus layer. Thus, the variation in pattern of foliar litter decomposition with climate and litter quality across functional groups in Eurasian forests showed different decomposition strategies for litter of different groups and genera.

Kang, School of Agriculture and Biology, Shanghai Jiao Tong University, Dongchuan Rd. 800, Shanghai 200240, P. R. China E-mail: hk@nn.cn
Berg, Department of Forest Ecology, University of Helsinki, Latokartanonkaari 7, FIN-00014 Finland; Dipartimento Biologia Strutturale e Funzionale. Complesso Universitario, Monte S. Angelo, Via Cinthia, IT-80126 Napoli, Italy E-mail: bb@nn.fi
Liu, School of Agriculture and Biology, Shanghai Jiao Tong University, Dongchuan Rd. 800, Shanghai 200240, P. R. China; Key Laboratory of Urban Agriculture (South), Ministry of Agriculture, P. R. China, Shanghai Jiao Tong University, 800 Dongchuan Rd., Shanghai 200240, P. R. China E-mail: chjliu@sjtu.edu.cn
Westman, Dipartimento Biologia Strutturale e Funzionale. Complesso Universitario, Monte S. Angelo, Via Cinthia, IT-80126 Napoli, Italy E-mail: cjw@nn.it

article id 179, category Research article

Henna Vartiamäki, Jarkko Hantula, Antti Uotila. (2009). Susceptibility of silver birch pruning wounds to infection by white-rot fungus (Chondrostereum purpureum), a potential bioherbicide. Silva Fennica vol. 43 no. 4 article id 179. https://doi.org/10.14214/sf.179

Keywords: birch; vegetation management; biological control; fungal decay; pruning

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We artificially inoculated pruning wounds of silver birch (Betula pendula Roth) to study seasonal variation in their vulnerability to infection by the fungal decomposer Chondrostereum purpureum (Pers. ex Fr.) Pouzar. This information is critical to the assessment of incidental infection risks in areas where C. purpureum may be used as a bioherbicide. On seven monthly occasions between April and October 2005, 30 birch trees were pruned to yield a total of 210 experimental trees. On each occasion, 10 trees were inoculated immediately with C. purpureum mycelium, 10 were inoculated with blank inoculum and 10 were only pruned. In the summer of 2007, a survey of 129 experimental trees showed that pruning wounds were most susceptible to infection during May. Treatment with C. purpureum at other times during the growing season also increased the extent of discoloration or decay but the effect was considerably less.

Vartiamäki, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: henna.vartiamaki@metla.fi
Hantula, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: jh@nn.fi
Uotila, University of Helsinki, Hyytiälä Forestry Field Station, Hyytiäläntie 124, FI-35500 Korkeakoski, Finland E-mail: au@nn.fi

article id 204, category Research article

Gaby Deckmyn, Bostjan Mali, Hojka Kraigher, Niko Torelli, Maarten Op de Beeck, Reinhart Ceulemans. (2009). Using the process-based stand model ANAFORE including Bayesian optimisation to predict wood quality and quantity and their uncertainty in Slovenian beech. Silva Fennica vol. 43 no. 3 article id 204. https://doi.org/10.14214/sf.204

Keywords: forest management; wood quality; Bayesian calibration; beech; forest model; mechanistic; red heart

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The purpose of this study was to expand an existing semi-mechanistic forest model, ANAFORE (ANAlysing Forest Ecosystems), to allow for the prediction of log quality and the accompanying uncertainty as influenced by climate and management. The forest stand is described as consisting of trees of different cohorts, either of the same or of different species (deciduous or coniferous). In addition to photosynthesis, transpiration, total growth and yield, the model simulates the daily evolution in vessel biomass and radius, parenchyma and branch development. From these data early and latewood biomass, wood tissue composition, knot formation and density are calculated. The new version presented here, includes the description of log quality, including red heart formation of beeches. A Bayesian optimisation routine for the species parameters was added to the stand model. From a given range of input parameters (prior), the model calculates an optimised range for the parameters (posterior) based on given output data, as well as an uncertainty on the predicted values. A case study was performed for Slovenian beech forests to illustrate the main model functioning and more in particular the simulation of the wood quality. The results indicate that the ANAFORE model is a useful tool for analyzing wood quality development and forest ecosystem functioning in response to management, climate and stand characteristics. However, the Bayesian optimization showed that the remaining uncertainty on the input parameters for the chosen stand was very large, due to the large number of input parameters in comparison to the limited stand data.

Deckmyn, Research Group Plant and Vegetation Ecology, University of Antwerpen, Universiteitsplein 1, 2610 Antwerpen, Belgium E-mail: gaby.deckmyn@ua.ac.be
Mali, Slovenian Forestry Institute, Vecna pot 2, 1000 Ljubljana, Slovenia E-mail: bm@nn.si
Kraigher, Slovenian Forestry Institute, Vecna pot 2, 1000 Ljubljana, Slovenia E-mail: hk@nn.si
Torelli, Slovenian Forestry Institute, Vecna pot 2, 1000 Ljubljana, Slovenia E-mail: nt@nn.si
Op de Beeck, Research Group Plant and Vegetation Ecology, University of Antwerpen, Universiteitsplein 1, 2610 Antwerpen, Belgium E-mail: modb@nn.be
Ceulemans, Research Group Plant and Vegetation Ecology, University of Antwerpen, Universiteitsplein 1, 2610 Antwerpen, Belgium E-mail: rc@nn.be

article id 203, category Research article

Matti Maltamo, Jussi Peuhkurinen, Jukka Malinen, Jari Vauhkonen, Petteri Packalén, Timo Tokola. (2009). Predicting tree attributes and quality characteristics of Scots pine using airborne laser scanning data. Silva Fennica vol. 43 no. 3 article id 203. https://doi.org/10.14214/sf.203

Keywords: LIDAR; alpha shape; crown height; height metrics; k-MSN; timber quality

Abstract | View details | Full text in PDF | Author Info

The development of airborne laser scanning (ALS) during last ten years has provided new possibilities for accurate description of the living tree stock. The forest inventory applications of ALS data include both tree and area-based plot level approaches. The main goal of such applications has usually been to estimate accurate information on timber quantities. Prediction of timber quality has not been focused to the same extent. Thus, in this study we consider here the prediction of both basic tree attributes (tree diameter, height and volume) and characteristics describing tree quality more closely (crown height, height of the lowest dead branch and sawlog proportion of tree volume) by means of high resolution ALS data. The tree species considered is Scots pine (Pinus sylvestris), and the field data originate from 14 sample plots located in the Koli National Park in North Karelia, eastern Finland. The material comprises 133 trees, and size and quality variables of these trees were modeled using a large number of potential independent variables calculated from the ALS data. These variables included both individual tree recognition and area-based characteristics. Models for the dependent tree characteristics to be considered were then constructed using either the non-parametric k-MSN method or a parametric set of models constructed simultaneously by the Seemingly Unrelated Regression (SUR) approach. The results indicate that the k-MSN method can provide more accurate tree-level estimates than SUR models. The k-MSN estimates were in fact highly accurate in general, the RMSE being less than 10% except in the case of tree volume and height of the lowest dead branch.

Maltamo, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: matti.maltamo@joensuu.fi
Peuhkurinen, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: jp@nn.fi
Malinen, Finnish Forest Research Institute, Joensuu Research Unit, FI-80101 Joensuu, Finland E-mail: jm@nn.fi
Vauhkonen, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: jv@nn.fi
Packalén, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: pp@nn.fi
Tokola, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: tt@nn.fi

article id 202, category Research article

Tuula Nuutinen, Antti Kilpeläinen, Hannu Hirvelä, Kari Härkönen, Veli-Pekka Ikonen, Reetta Lempinen, Heli Peltola, Lars Wilhelmsson, Seppo Kellomäki. (2009). Future wood and fibre sources – case North Karelia in eastern Finland. Silva Fennica vol. 43 no. 3 article id 202. https://doi.org/10.14214/sf.202

Keywords: National Forest Inventory; fibre length; wood density; proportion of latewood; forest scenario analysis

Abstract | View details | Full text in PDF | Author Info

Information on the potential wood supply is important for the wood industry. In this study, the future development of growing stock, cutting potential and wood properties corresponding to the regional scenario of North Karelian Forest Programme 2006–2010 was analysed. The simulations were performed by employing the Finnish MELA system together with the sample plot and tree data of the 9th Finnish National Forest Inventory (NFI9) as initial data for the simulations. Disc-based models for basic wood density, proportion of latewood and fibre length of Norway spruce and Scots pine in Sweden were calibrated and integrated into the MELA system. The wood properties at breast height of both harvested and standing trees were analysed in different strata (age, site type and cutting method) during the scenario period of 50 years (2002–2052). The average wood properties within the same strata varied only slightly over time. However, the results for different strata differed considerably. In general, wood density, fibre length and proportion of latewood increased, on average, as a function of tree age and along with a decrease in site fertility (excl. wood density and proportion of latewood in harvested Norway spruce in the first case and fibre length in the latter case for both species). For trees less than 80 years, properties in harvested trees were equal to or slightly greater than those of standing trees. The values for clear-cuttings were greater or equal to those of thinnings (excl. wood density and proportion of latewood in Norway spruce). The study demonstrates the value of model-based analyses utilising NFI tree measurements in regions that are considered to be sources of raw material.

Nuutinen, European Forest Institute, Torikatu 34, FI-80100 Joensuu, Finland E-mail: tuula.nuutinen@efi.int
Kilpeläinen, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: ak@nn.fi
Hirvelä, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: hh@nn.fi
Härkönen, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: kh@nn.fi
Ikonen, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: vpi@nn.fi
Lempinen, Finnish Forest Research Institute, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: rl@nn.fi
Peltola, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: hp@nn.fi
Wilhelmsson, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: lw@nn.fi
Kellomäki, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: sk@nn.fi

article id 201, category Research article

Rui Qi, Véronique Letort, Mengzhen Kang, Paul-Henry Cournède, Philippe de Reffye, Thierry Fourcaud. (2009). Application of the GreenLab model to simulate and optimize wood production and tree stability: a theoretical study. Silva Fennica vol. 43 no. 3 article id 201. https://doi.org/10.14214/sf.201

Keywords: wood quality; optimization; biomechanics; FSPM; Particle Swarm Optimization; source-sink dynamics; biomass allocation

Abstract | View details | Full text in PDF | Author Info

The GreenLab model was used to study the interaction between source-sink dynamics at the whole tree level, wood production and distribution within the stem, and tree mechanical stability through simulation and optimization. In this first promising numerical attempt, two GreenLab parameters were considered in order to maximize wood production: the sink strength for cambial growth and a coefficient that determines the way the biomass assigned to cambial growth is allocated to each metamer, through optimization and simulation respectively. The optimization procedure that has been used is based on a heuristic optimization algorithm called Particle Swarm Optimization (PSO). In the first part of the paper, wood production was maximized without considering the effect of wood distribution on tree mechanical stability. Contrary to common idea that increasing sink strength for cambial growth leads to increasing wood production, an optimal value can be found. The optimization results implied that an optimal source and sink balance should be considered to optimize wood production. In a further step, the mechanical stability of trees submitted to their self weight was taken into account based on simplified mechanical assumptions. Simulation results revealed that the allocation of wood at the stem base strongly influenced its global deformation. Such basic mechanical criterion can be an indicator of wood quality if we consider further the active biomechanical processes involved in tree gravitropic responses, e.g. formation of reaction wood.

Qi, Ecole Centrale Paris, Laboratory of Applied Mathematics, Grande Voie des Vignes, 92295 Chatenay-Malabry, France; Institute of Automation, Chinese Academy of Sciences, LIAMA/NLPR, P.O.Box 2728, Beijing, China E-mail: qiruitree@gmail.com
Letort, Ecole Centrale Paris, Laboratory of Applied Mathematics, Grande Voie des Vignes, 92295 Chatenay-Malabry, France E-mail: vl@nn.fr
Kang, Institute of Automation, Chinese Academy of Sciences, LIAMA/NLPR, P.O.Box 2728, Beijing, China E-mail: mk@nn.cn
Cournède, Ecole Centrale Paris, Laboratory of Applied Mathematics, Grande Voie des Vignes, 92295 Chatenay-Malabry, France; INRIA saclay Ile-de-France, EPI Digiplant, Parc Orsay Université, 91893 Orsay cedex, France E-mail: phc@nn.fr
Reffye, INRIA saclay Ile-de-France, EPI Digiplant, Parc Orsay Université, 91893 Orsay cedex, France; CIRAD, UMR AMAP, Montpellier, F-34000 France E-mail: pdr@nn.fr
Fourcaud, CIRAD, UMR AMAP, Montpellier, F-34000 France E-mail: tf@nn.fr

article id 200, category Research article

Peder Gjerdrum, Mauro Bernabei. (2009). Three-dimensional spiral grain pattern in five large Norway spruce stems. Silva Fennica vol. 43 no. 3 article id 200. https://doi.org/10.14214/sf.200

Keywords: grain angle; 3D-model; tree helix; split disk

Abstract | View details | Full text in PDF | Author Info

There is a great deal of interest involved in investigating and understanding grain angle in trees. The objective of the study presented in this paper has been to identify a joint, three-dimensional model for grain angle in stems of Norway spruce (Picea abies (L.) Karst.) Five large spruce trees were sampled. Transverse disks were extracted at regular intervals along the stem, split through the pith with a blunt knife, and observed for grain angle and cambial age along the north radius, setting pith observation to zero angle. The overall finding confirmed grain pattern congruent to distance from pith along the stem, a pattern that varies from tree to tree. Models expressing distance from the pith in cambial age performed slightly better than models in spatial distance. Grain pattern changed slightly along the stem, and this change was found to be consistent for the five stems: the left-handed grain angle in the juvenile wood was more pronounced upwards in the stem, and the angle changed faster towards right-handed spiralling. The model did not account for grain variation related to knots, undulating pith or other irregularities. Model residuals were normally distributed and seemingly homogeneous for all trees and for all height levels. The model is an extension of existing two-dimensional models for stem cross-cuts. After verification in a wider sample, the results should be applicable in tree and wood modelling.

Gjerdrum, Norwegian Forest and Landscape Institute, P.O. Box 115, N-1431 Ås, Norway E-mail: peder.gjerdrum@skogoglandskap.no
Bernabei, Norwegian Forest and Landscape Institute, P.O. Box 115, N-1431 Ås, Norway E-mail: mb@nn.no

article id 199, category Research article

R. Edward Thomas. (2009). Modeling the relationships among internal defect features and external Appalachian hardwood log defect indicators. Silva Fennica vol. 43 no. 3 article id 199. https://doi.org/10.14214/sf.199

Keywords: modeling; hardwood; log defects; surface scan

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As a hardwood tree grows and develops, surface defects such as branch stubs and wounds are overgrown. Evidence of these defects remain on the log surface for decades and in many instances for the life of the tree. As the tree grows the defect is encapsulated or grown over by new wood. During this process the appearance of the defect in the tree’s bark changes. The defect becomes flatter and its dimension changes. This progressional change in appearance is predictable, permitting the size and location of the internal defect to be reliably estimated. This paper concerns the development and analysis of models for the prediction of internal features. With the advent of surface scanning and external detection systems, the prediction of internal features promises to significantly improve the quality, yield, and value of sawn wood products.

Thomas, USDA Forest Service, 241 Mercer Springs Road, Princeton, WV 24740, USA E-mail: edthomas@gmail.com

article id 198, category Research article

Henrik Heräjärvi. (2009). Effect of drying technology on aspen wood properties. Silva Fennica vol. 43 no. 3 article id 198. https://doi.org/10.14214/sf.198

Keywords: stability; heat treatment; modification; press drying; strength

Abstract | View details | Full text in PDF | Author Info

Heräjärvi, Metla, Joensuu Research Unit, Box 68, FI-80101 Joensuu, Finland E-mail: henrik.herajarvi@metla.fi

article id 197, category Research article

Veli-Pekka Ikonen, Seppo Kellomäki, Heli Peltola. (2009). Sawn timber properties of Scots pine as affected by initial stand density, thinning and pruning: a simulation based approach. Silva Fennica vol. 43 no. 3 article id 197. https://doi.org/10.14214/sf.197

Keywords: three-dimensional modeling; sawing simulator; knots; Pinus sylvestris L.; shading

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The aim of this work was to analyze how different management schedules with varying initial stand density, thinning and artificial pruning of branches affect the quality, quantity and value of sawing yield in Scots pine (Pinus sylvestris L.). For this purpose, an integrated model system was employed and further developed to simulate: i) the three dimensional structure of the crown and stem of an average tree grown in a stand related to the changes in the within-stand light conditions as caused by the stand management, and ii) the sawing of logs into pieces and their quality grading based on the size and number of living and dead knots on the surfaces of sawn pieces. To maximize the quality of sawn timber, relatively dense stand is desired in the early phase of the rotation to reduce, especially in the lower part of stem, the growth of branches, and to increase the rate of dying and pruning-off of branches. In the later phase, a relatively sparse stand is desired to increase the self-pruning of branches and the occlusion of knots. However, in any case, artificial pruning is needed to maximize the knot-free zone of the stem. Also the value optimization of individual sawn pieces affects the quality and value of sawn timber. Because, only average tree was simulated, the differences between scenarios for stem volume were small. In the future, further model development is needed to analyze the development of crown and stem properties of trees with different status in a stand.

Ikonen, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: veli-pekka.ikonen@joensuu.fi
Kellomäki, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: sk@nn.fi
Peltola, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: hp@nn.fi

article id 196, category Research article

Aaron R. Weiskittel, Laura S. Kenefic, Robert S. Seymour, Leah M. Phillips. (2009). Long-term effects of precommercial thinning on the stem dimensions, form and branch characteristics of red spruce and balsam fir crop trees in Maine, USA. Silva Fennica vol. 43 no. 3 article id 196. https://doi.org/10.14214/sf.196

Keywords: Acadian Forest; growth and yield; Abies balsamea; Picea rubens; precommercial thinning

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The effects of precommercial thinning (PCT) on stem dimensions, form, volume, and branch attributes of red spruce [Picea rubens Sarg.] and balsam fir [Abies balsamea (L.) Mill.] crop trees were assessed 25 years after treatment in an even-aged northern conifer stand. Treatments were a uniform 2.4 x 2.4-m spacing and a control (no PCT). The PCT treatment significantly increased individual tree diameter at breast height (DBH), height growth, crown ratio, and crown width, while it reduced the tree height to DBH ratio. PCT also significantly increased stem taper and consequently, regional volume equations overpredicted observed stem volume by 2 to 15%, particularly for the spaced trees. PCT also increased the number and maximum size of branches on the lower bole. The sizes of knots on half of the sampled spruce crop trees in the spaced plots precluded them from being used as select structural lumber; there were no other effects on log grade. Our findings indicate that PCT can have a long-term influence on the structural attributes of individual trees, and that improved stem-volume prediction equations are needed in the Acadian region of North America.

Weiskittel, School of Forest Resources, University of Maine, Orono, ME 04469, USA E-mail: aaron.weiskittel@umit.maine.edu
Kenefic, U.S. Forest Service, Northern Research Station, Bradley, ME 04411, USA E-mail: lsk@nn.us
Seymour, School of Forest Resources, University of Maine, Orono, ME 04469, USA E-mail: rss@nn.us
Phillips, School of Forest Resources, University of Maine, Orono, ME 04469, USA E-mail: lmp@nn.us

article id 195, category Research article

John R. Moore, Andrew J. Lyon, Gregory J. Searles, Leena E. Vihermaa. (2009). The effects of site and stand factors on the tree and wood quality of Sitka spruce growing in the United Kingdom. Silva Fennica vol. 43 no. 3 article id 195. https://doi.org/10.14214/sf.195

Keywords: wood quality; acoustics; timber properties; wood stiffness

Abstract | View details | Full text in PDF | Author Info

The extent and sources of variation in the wood quality of Sitka spruce (Picea sitchensis (Bong.) Carr.) were quantified using data collected from 64 stands in northern Britain. These stands were selected on the basis of elevation, latitude, longitude, yield class, initial spacing and the presence or absence of thinning. Dynamic modulus of elasticity (MOE) was calculated from measurements of stress wave velocity made on standing trees and qualitative descriptions were made of stem form. Dynamic MOE of individual trees ranged from 3.81 kN/mm² up to 12.29 kN/mm², with a mean of 7.71 kN/mm². Approximately 55 percent of the variation in dynamic MOE was due to differences between individual trees within a site, while 35 percent was due to differences between sites. The remaining 10 percent was due to differences between the measurements made on opposite sides of each tree. Variation in dynamic MOE at the site level was significantly influenced by yield class, elevation as well as by a number of the interactions between these factors and latitude, longitude and initial spacing. A multiple regression model incorporating these variables was able to explain 45 percent of the variation in dynamic MOE. Ramicorn branches were the most commonly recorded defect (37.2% of all live trees), followed by stem scarring and basal sweep (6.9% and 6.3%, respectively). Dynamic MOE was not influenced by stem straightness (p = 0.10) which indicates the utility of stress wave velocity measurements for segregating Sitka spruce stands based on potential grade recovery.

Moore, Centre for Timber Engineering, Napier University, 10 Colinton Road, Edinburgh EH10 5DT, United Kingdom E-mail: j.moore@napier.ac.uk
Lyon, Centre for Timber Engineering, Napier University, 10 Colinton Road, Edinburgh EH10 5DT, United Kingdom E-mail: ajl@nn.uk
Searles, Centre for Timber Engineering, Napier University, 10 Colinton Road, Edinburgh EH10 5DT, United Kingdom E-mail: gjs@nn.uk
Vihermaa, Department of Analytical and Environmental Chemistry, University of Glasgow, Glasgow, UK E-mail: lev@nn.uk

article id 194, category Research article

Ane Zubizarreta Gerendiain, Heli Peltola, Pertti Pulkkinen. (2009). Growth and wood property traits in narrow crowned Norway spruce (Picea abies f. pendula) clones grown in southern Finland. Silva Fennica vol. 43 no. 3 article id 194. https://doi.org/10.14214/sf.194

Keywords: fibre length; stem volume; height; diameter; wood density; earlywood; latewood

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We investigated the growth, yield, wood density traits and fibre properties in 13 narrow crowned Norway spruce (Picea abies f. pendula) clones grown at a spacing of 2 m x 1.5 m (about 3300 seedlings/ha) in a field trial established in 1988 in southern Finland on a forest soil. For comparison, we used 3 normal crowned Norway spruce (Picea abies (L.) Karst.) genetic entries grown as a mixture in the same trial representing southern Finnish breeding regions. We found that wood density traits and fibre properties showed, on average, lower phenotypic variation than growth and yield traits regardless of crown type. Narrow crowned clones also had, on average, lower stem volume and fibre length, but higher overall wood density. More over, the phenotypic correlations between studied growth and wood properties ranged, on average, from moderate (normal crown) to high (narrow crown). These results were opposite to previous findings for narrow and normal crowned genetic entries grown in narrower spacing (1 m x 1 m) in southern Finland. Thus, this indicates lower plasticity of narrow crowned clones to the increasing growing space compared to normal crowned ones, so, they should be grown at denser spacing in order to fully utilise its space efficiency capacity. However, this field trial was established as a mixture of normal and narrow crown trees, so that 90% of genetic entries were narrow crowned ones, and therefore the crown competition would be much higher for normal crowned trees when the whole trial would consist of that entry alone. In the latter case, we could expect significantly lower productivity of normal crowned genetic entries with this spacing.

Zubizarreta Gerendiain, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: ane.zubizarreta@joensuu.fi
Peltola, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: hp@nn.fi
Pulkkinen, Finnish Forest Research Institute, Haapastensyrjä Breeding Station, FI-12600 Läyliäinen, Finland E-mail: pp@nn.fi

article id 193, category Research article

Jaume Gort, Ane Zubizarreta Gerendiain, Heli Peltola, Pertti Pulkkinen, Johanna Routa, Raimo Jaatinen. (2009). Differences in fibre properties in Scots pine (Pinus sylvestris L.) genetic entries grown at different spacing and sites. Silva Fennica vol. 43 no. 3 article id 193. https://doi.org/10.14214/sf.193

Keywords: fibre length; fibre width; genetic entry; growth; wood density; coarseness; spacing

Abstract | View details | Full text in PDF | Author Info

In forest breeding, stem volume growth and sawn timber quality indicators have been used as the most important selection traits for Scots pine, whereas less attention has been given to characteristics such as fibre properties. In the above context, we investigated the differences in fibre properties (i.e. fibre length, fibre width and coarseness) in 20 year old Scots pine (Pinus sylvestris L.) genetic entries as affected by spacing and site, but also the phenotypic correlations between fibre properties, yield and wood density. The study was based on materials harvested from 10 genetic entries grown in a spacing trial (site 1) in central Finland, with a current stand density of 2000 (spacing 1), 2000–2500 (spacing 2) and 4000 trees/ha (spacing 3). In order to study the effects of site, we harvested additional material (4 of 7 genetic entries same as on site 1) from a trial located in southern Finland with a corresponding stand density of 2000 trees/ha (site 2). On site 1, spacing 1 and 3, all average values for analysed fibre properties were similar. In spacing 2 average values were slightly higher. On site 2, the average values for different fibre properties were similar compared to the corresponding spacing 1 on site 1. Spacing affected (p < 0.05) all average fibre properties on site 1; as did also site, when comparing same genetic entries grown on both sites. Regardless of spacing and site, the phenotypic correlations between average fibre length, fibre width and coarseness showed, on average, moderate to strong correlation (p < 0.05). Fibre width showed, in general, low and positive phenotypic correlation with diameter at breast height, stem volume and wood density on site 1. However, as a whole, the ranking of genetic entries changed depending on the trait and spacing considered. Thus, no overall ranking between genetic entries was possible.

Gort, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: jaume.gort@joensuu.fi
Zubizarreta Gerendiain, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: azg@nn.fi
Peltola, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: hp@nn.fi
Pulkkinen, Finnish Forest Research Institute, Haapastensyrjä Breeding Station, Karkkilantie E-mail: pp@nn.fi
Routa, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: jr@nn.fi
Jaatinen, Finnish Forest Research Institute, Haapastensyrjä Breeding Station, Karkkilantie E-mail: rj@nn.fi

article id 192, category Research article

Heli Peltola, Jaume Gort, Pertti Pulkkinen, Ane Zubizarreta Gerendiain, Jouni Karppinen, Veli-Pekka Ikonen. (2009). Differences in growth and wood density traits in Scots pine (Pinus sylvestris L.) genetic entries grown at different spacing and sites. Silva Fennica vol. 43 no. 3 article id 192. https://doi.org/10.14214/sf.192

Keywords: genetic entry; stem volume; height; diameter; wood property traits; phenotypic correlation

Abstract | View details | Full text in PDF | Author Info

In forest breeding, stem volume has typically taken as the most important selection trait, whereas less attention has been given to wood density traits. In this work, we investigated the effects of spacing and genetic entry on the growth, yield and wood density traits in 20 year old Scots pines (Pinus sylvestris L.) based on 10 genetic entries harvested from a spacing trial (stand density range 2000–4000 trees/ha) in central Finland. In order to study also the site effects, we harvested additional material from a trial located in southern Finland (stand density of 2000 trees/ha). Compared to growth and yield properties, wood density traits showed a lower phenotypic variation. Phenotypic correlations among different traits were negative, and mostly moderate to high, suggesting that selection for one trait would simultaneously affect the others. In addition, moderate to strong phenotypic correlations were found among different wood density traits. Stem volume (V) and breast height diameter (DBH) were the largest in widest spacing, whereas in the densest one tree height (H) and latewood percentage were the highest. Genetic entry affected H and wood density traits regardless of spacing. When comparing two sites (with same stand density), genetic entry affected H, whereas site affected DBH and wood density traits. Ranking between genetic entries changed depending on the trait, spacing or site considered. Therefore, no overall ranking was possible. However, we could identify genetic entries having a high V and a relatively high wood density, showing potential for future forest regeneration material.

Peltola, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: heli.peltola@joensuu.fi
Gort, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: jg@nn.fi
Pulkkinen, Finnish Forest Research Institute, Haapastensyrjä Breeding Station, Karkkilantie 247, FI-12600 Läyliäinen, Finland E-mail: pp@nn.fi
Zubizarreta Gerendiain, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: azg@nn.fi
Karppinen, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: jk@nn.fi
Ikonen, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: vpi@nn.fi

article id 191, category Research article

Katri Luostarinen, Noora Huotari, Eila Tillman-Sutela. (2009). Effect of regeneration method on growth, wood density and fibre properties of downy birch (Betula pubescens Ehrh.). Silva Fennica vol. 43 no. 3 article id 191. https://doi.org/10.14214/sf.191

Keywords: coppice; fibre length; fibre width; regeneration methods; wood properties

Abstract | View details | Full text in PDF | Author Info

Short rotation tree stands, established by coppicing, are nowadays used mainly for energy purposes in Fennoscandia, but their usage for pulp raw material may increase in the future. Downy birch (Betula pubescens), which is commonly used for pulp production in boreal zone, has good sprouting capacity. However, it is not known if the fibre properties of sprout-originated downy birches differ from those of seed-originated ones. Therefore, fibre length and width of sprout- and seed-originated downy birches grown on fertile soil were measured at a stand age of 25 years. Additionally annual ring width, stem height and diameter, and wood density were studied to compare the growth and wood properties of sprout- and seed-originated birches. Annual rings were slightly wider in sprout- than in seed-originated birches, whereas no differences were observed in wood density. Fibres, too, were slightly longer and wider in sprout- than in seed-originated trees. Still these minor differences observed here are hardly significant for the industries using birch wood. Consequently downy birch wood from coppiced stands is well suited for pulp. The advantages of coppice, i.e. rapidity and low costs of establishment, productivity, and the ability of downy birch to grow on untypical forest sites, may even increase the importance of the wood coming from coppiced birch stands in the future.

Luostarinen, Faculty of Forest Sciences, University of Joensuu, P.O. Box 111, FI-80101, Joensuu, Finland E-mail: katri.luostarinen@joensuu.fi
Huotari, Finnish Forest Research Institute, Muhos Research Unit, Kirkkosaarentie, FI-91500 Muhos, Finland E-mail: nh@nn.fi
Tillman-Sutela, Finnish Forest Research Institute, Muhos Research Unit, Kirkkosaarentie, FI-91500 Muhos, Finland E-mail: ets@nn.fi

article id 213, category Research article

Bum-Jin Park, Yuko Tsunetsugu, Tamami Kasetani, Takeshi Morikawa, Takahide Kagawa, Yoshifumi Miyazaki. (2009). Physiological effects of forest recreation in a young conifer forest in Hinokage Town, Japan. Silva Fennica vol. 43 no. 2 article id 213. https://doi.org/10.14214/sf.213

Keywords: blood pressure; heart rate variability; pulse rate; relaxation; Shinrin-yoku; therapeutic effects of forest; well-being

Abstract | View details | Full text in PDF | Author Info

It is widely believed that coming into contact with forest environments is somehow beneficial to human well-being and comfort. In Japan, “Shinrin-yoku” (taking in the atmosphere of a forest) has been proposed to be a relaxation activity associated with forest recreation. The purpose of this study was to examine the physiological effects of forest recreation on the autonomic nervous activity. The subjects were twelve male university students (21.8 ± 0.8 years old). On the first day of the experiment, six subjects were sent to a forest area, and the other six to a city area. On the second day, each subject was sent to the area he did not visit on the first day as a cross check. The subjects walked (15 minutes) around their assigned areas before noon, and sat on chairs viewing (15 minutes) the landscapes of their assigned areas in the afternoon. Heart rate variability (HRV), blood pressure, and pulse rate were measured as physiological indices. Measurements were taken at the place of accommodation in the morning, before and after walking, and before and after viewing at their assigned field areas. Pulse rate, diastolic blood pressure and LF/(LF+HF) (LF – low frequency, HF – high frequency) components of HRV were significantly lower in the forest area than in the city area. HF components of HRV tended to be higher in the forest than in the city. In conclusion, the results of the physiological measurements show that forest recreation enabled effective relaxation in people, both of the mind and body.

Park, Chiba University, Center for Environment, Health and Field Sciences, Kashiwa-no-ha 6-2-1, 277-0882 Kashiwa, Chiba, Japan E-mail: bjpark@faculty.chiba-u.jp
Tsunetsugu, Chiba University, Center for Environment, Health and Field Sciences, Kashiwa-no-ha 6-2-1, 277-0882 Kashiwa, Chiba, Japan E-mail: yt@nn.jp
Kasetani, Chiba University, Center for Environment, Health and Field Sciences, Kashiwa-no-ha 6-2-1, 277-0882 Kashiwa, Chiba, Japan E-mail: tk@nn.jp
Morikawa, Chiba University, Center for Environment, Health and Field Sciences, Kashiwa-no-ha 6-2-1, 277-0882 Kashiwa, Chiba, Japan E-mail: tm@nn.jp
Kagawa, Chiba University, Center for Environment, Health and Field Sciences, Kashiwa-no-ha 6-2-1, 277-0882 Kashiwa, Chiba, Japan E-mail: tk@nn.jp
Miyazaki, Chiba University, Center for Environment, Health and Field Sciences, Kashiwa-no-ha 6-2-1, 277-0882 Kashiwa, Chiba, Japan E-mail: ym@nn.jp

article id 212, category Research article

Sandhya Samarasinghe. (2009). Exploration of fracture dynamics properties and predicting fracture toughness of individual wood beams using neural networks. Silva Fennica vol. 43 no. 2 article id 212. https://doi.org/10.14214/sf.212

Keywords: Pinus radiata; wood properties; cracks; initiation; New Zealand; peak stress; speed; video imaging

Abstract | View details | Full text in PDF | Author Info

In this study, the time to crack initiation (T_init), duration of crack propagation (T_frac), crack initiation stress, peak stress as well as crack speed and fracture toughness were investigated for three Rates of Loading (ROL) and four sizes of notched wood beams using high-speed video imaging and neural networks. T_init was consistent for all volumes and the average T_init was nonlinearly related to volume and ROL. For the smallest ROL, there was a distinct volume effect on T_init and the effect was negligble at the largest ROL. However, the stress at crack initiation was not consistent. Contrasting these, T_frac for all volumes appeared to be highly variable but the peak stress carried prior to catastrophic failure was consistent. The crack propagation was a wave phenomenon with positive and negative (crack closure) speeds that varied with the ROL. As accurate estimation of crack initiation load (or stress) and its relationship to peak load (or stress) is important for determining fracture toughness, Artificial Neural Networks (ANN) models were developed for predicting them from volume, Young’s modulus, face and grain angles, density, moisture content and ROL. Models for crack initiation load and peak load showed much higher predictive power than those for the stresses with correlation coefficients of 0.85 and 0.97, respectively, between the actual and predicted loads. Neural networks were also developed for predicting fracture toughness of individual wood specimens and the best model produced a statistically significant correlation of 0.813 between the predicted and actual fracture toughness on a validation dataset. The inputs captured 62% of variability of fracture toughness. Volume and Young’s modulus were the top two contributing variables with others providing lesser contributions.

Samarasinghe, Centre for Advanced Computational Solutions (C-fACS), Lincoln University, Canterbury, New Zealand E-mail: sandhya.samarasinghe@lincoln.ac.nz

article id 211, category Research article

Timo Pukkala. (2009). Population-based methods in the optimization of stand management. Silva Fennica vol. 43 no. 2 article id 211. https://doi.org/10.14214/sf.211

Keywords: Particle Swarm Optimization; differential evolution; evolution strategy; Nelder-Mead method; amoeba search; polytope search; evolutionary computing

Abstract | View details | Full text in PDF | Author Info

In Finland, the growth and yield models for tree stands are simulation programs that consist of several sub-models. These models are often non-smooth and non-differentiable. Direct search methods such as the Hooke-Jeeves algorithm (HJ) are suitable tools for optimizing stand management with this kind of complicated models. This study tested a new class of direct search methods, namely population-based methods, in the optimization of stand management. The tested methods were differential evolution, particle swarm optimization, evolution strategy, and the Nelder-Mead method. All these methods operate with a population of solution vectors, which are recombined and mutated to obtain new candidate solutions. The management schedule of 719 stands was optimized with all population-based methods and with the HJ method. The population-based methods were competitive with the HJ method, producing 0.57% to 1.74% higher mean objective function values than HJ. On the average, differential evolution was the best method, followed by particle swarm optimization, evolution strategy, and Nelder-Mead method. However, differences between the methods were small, and each method was the best in several stands. HJ was alone the best method in 7% of stands, and a population based method in 3% (Nelder-Mead) to 29% (differential evolution) of stands. All five methods found the same solution in 18% of stands.

Pukkala, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: timo.pukkala@joensuu.fi

article id 210, category Research article

Juha Heiskanen, Markku Lahti, Jaana Luoranen, Risto Rikala. (2009). Nutrient loading has a transitory effect on the nitrogen status and growth of outplanted Norway spruce seedlings. Silva Fennica vol. 43 no. 2 article id 210. https://doi.org/10.14214/sf.210

Keywords: nursery fertilization; plantation performance; soil fertility

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In recent years increased fertilization provided to tree seedlings in the nursery in the previous autumn has been introduced in order to promote good outplanting performance. In this paper this nutrient loading has been studied in order to determine how the increased seedling nutrient status with unaffected seedling size affects both the growth and the nutrient concentration, content and uptake of two-year-old Norway spruce container seedlings (Picea abies (L.) Karsten) after outplanting. Seedling development was monitored for three years at two contrasting soil fertility levels on a sandy test field in two planting years and on one natural forest outplanting site in central Finland. Nutrient loading was shown to increase shoot and root growth in a poor fertility soil during the first growing season after planting, while, after the first growing season, nutrient loading was not found to affect seedling performance. However, although nutrient loading cannot compensate for the availability of nutrients to the seedlings from the soil, it may provide an additional input for fast plantation establishment on poorer sites during the first crucial growing season after outplanting.

Heiskanen, The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland E-mail: juha.heiskanen@metla.fi
Lahti, The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland E-mail: ml@nn.fi
Luoranen, The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland E-mail: jl@nn.fi
Rikala, The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland E-mail: rr@nn.fi

article id 209, category Research article

Jaana Luoranen, Kyösti Konttinen, Risto Rikala. (2009). Frost hardening and risk of a second flush in Norway spruce seedlings after an early-season short-day treatment. Silva Fennica vol. 43 no. 2 article id 209. https://doi.org/10.14214/sf.209

Keywords: frost hardiness; Picea abies; bud formation; container seedlings; height growth cessation; second flush

Abstract | View details | Full text in PDF | Author Info

There have been years in Finland when container seedlings of Norway spruce (Picea abies (L.) Karsten) planted in the summer have been damaged by early-autumn frosts. For August and September plantings, the seedlings can be hardened by means of short-day (SD) treatment, but little information is available about its usability for earlier plantings. We studied the effects of early-season SD treatment on the frost hardiness and risk of a second flush of Norway spruce seedlings. In three successive years, second-year seedlings were grown in a greenhouse or outdoors in the spring and early summer and then subjected to two or three-week SD treatment beginning on the second, third, or fourth week of June. We monitored the height growth cessation, bud formation, and frost hardiness of the seedlings in the nursery. All SD treatments made the height growth cease, but the risk of a second flush increased if the temperature sum was less than 300 d.d. before the beginning of the SD treatment or more than 450 d.d. between the end of the treatment and mid-August. Clearly, then, SD treatment reduced the risk of a second flush in seedlings that had been grown in a greenhouse in the spring. Early-season SD treatment increased the frost hardiness of both needles and stems for late July to early September in comparison with untreated seedlings. Later in the autumn, however, the differences disappeared. Before recommending the use of early-season SD-treated seedlings for summer planting, the method has to be tested in practical field conditions.

Luoranen, The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland E-mail: jaana.luoranen@metla.fi
Konttinen, The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland E-mail: kk@nn.fi
Rikala, The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland E-mail: rr@nn.fi

article id 208, category Research article

Mikko Räisänen, Tapani Repo, Tarja Lehto. (2009). Cold acclimation of Norway spruce roots and shoots after boron fertilization. Silva Fennica vol. 43 no. 2 article id 208. https://doi.org/10.14214/sf.208

Keywords: boron deficiency; dieback; freezing tolerance; mineral nutrition

Abstract | View details | Full text in PDF | Author Info

Boron deficiency, manifested as shoot dieback, is a problem in conifer stands growing on soils with high nitrogen availability in Fennoscandia. Earlier observations on Norway spruce (Picea abies L. Karst.) suggest that freezing tolerance is decreased by boron deficiency. Here, the effect of boron fertilization on cold acclimation of Norway spruce was studied in a young stand with initially low boron status two years after fertilization. Buds, stems, needles and roots were collected at five sampling times during cold acclimation and subsequently exposed to series of freezing temperatures. Lethal temperatures of organs were assessed by electrolyte leakage method (EL) and visual scoring of damage (VS). Freezing tolerance of buds was measured also by differential thermal analysis (DTA). The mean boron (B) concentration in needles was 4 mg kg–1 in unfertilized and 21 mg kg–1 in B-fertilized trees while critical level of B deficiency is considered to be 5 mg kg–1. The risk for increased freezing injuries in the low-B trees was not evident since all trees achieved cold hardiness that would be sufficient in central Finland. At two sampling times out of five, shoots or stem of B-fertilized trees were slightly more freezing tolerant than non-fertilized trees. However, the present study does not give strong evidence for the hypothesis that decreased freezing tolerance in B deficiency would be a triggering factor for leader dieback in Norway spruce at the B levels studied.

Räisänen, University of Joensuu, Faculty of Forest Sciences, P.O.Box 111, FI-80101 Joensuu, Finland; (present address), FAForest, FI-83480 Ahonkylä, Finland E-mail: mr@nn.fi
Repo, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: tr@nn.fi
Lehto, University of Joensuu, Faculty of Forest Sciences, P.O.Box 111, FI-80101 Joensuu, Finland E-mail: tarja.lehto@joensuu.fi

article id 207, category Research article

Mats T. Olsson, Maria Erlandsson, Lars Lundin, Torbjörn Nilsson, Åke Nilsson, Johan Stendahl. (2009). Organic carbon stocks in Swedish Podzol soils in relation to soil hydrology and other site characteristics. Silva Fennica vol. 43 no. 2 article id 207. https://doi.org/10.14214/sf.207

Keywords: temperature sum; soil organic carbon; forest land; soil hydrological class; N deposition; latitude; site capacity; mean annual precipitation; altitude

Abstract | View details | Full text in PDF | Author Info

Site characteristics influence soil organic carbon (SOC) stocks. In Podzols under Swedish forest land, SOC stocks were related to latitude, altitude, soil hydrological class categorized by mean groundwater level, mean annual precipitation, temperature sum during the growing season, total annual nitrogen (N) deposition and site capacity. SOC stocks were determined for the O-horizon and for total soil (O-horizon + mineral soil to a depth of 50 cm). Data from the Swedish National Forest Soil Inventory 1993–2001 were used (1477 field plots). The O-horizon was sampled with a core sampler and carbon (C) stocks were determined. For the mineral soil layers the SOC stock was calculated based on the SOC concentrations, bulk density and content of rock fragments. The results showed that the overall mean SOC stock was 2.8 and 8.2 kg C m^–2 for O-horizon and total soil, respectively. Soil hydrological class strongly affected SOC stocks, which increased from on average 6.7 kg C m^–2 at dry sites to 9.7 kg C m^–2 at slightly moist sites. Corresponding values for the O-horizon were 2.0 to 4.4 kg C m^–2. The correlation coefficients for the linear relationship between SOC stock and site characteristics were highest for N deposition, which explained up to 25% of variation, and latitude, which explained up to 20% of variation. Altitude had the lowest degree of explanation.

Olsson, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden E-mail: mto@nn.se
Erlandsson, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden E-mail: me@nn.se
Lundin, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden E-mail: ll@nn.se
Nilsson, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden E-mail: torbjorn.nilsson@mark.slu.se
Nilsson, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden E-mail: an@nn.se
Stendahl, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden E-mail: js@nn.se

article id 206, category Research article

Qiaoying Zhang, Yunchun Zhang, Shaolin Peng, Eshetu Yirdaw, Ning Wu. (2009). Spatial structure of alpine trees in Mountain Baima Xueshan on the southeast Tibetan Plateau. Silva Fennica vol. 43 no. 2 article id 206. https://doi.org/10.14214/sf.206

Keywords: treeline; timberline; Abies forestii var. georgei (Orr) Farjon; Juniperus saltuaria Rehder & E.H. Wilson; spatial point pattern

Abstract | View details | Full text in PDF | Author Info

Zhang, State Key Laboratory of Biocontrol, Sun Yat-Sen University, Guangzhou 510275, Guangdong Province, China; Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, Sichuan Province, China E-mail: lsspsl@mail.sysu.edu.cn
Zhang, Shandong Institute of Light Industry, Jinan 250353, Shandong Province, China E-mail: yz@nn.cn
Peng, State Key Laboratory of Biocontrol, Sun Yat-Sen University, Guangzhou 510275, Guangdong Province, China E-mail: sp@nn.cn
Yirdaw, Viikki Tropical Resources Institute (VITRI), P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ey@nn.fi
Wu, Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, Sichuan Province, China E-mail: nw@nn.cn

article id 205, category Research article

Pekka Vakkari, Mari Rusanen, Katri Kärkkäinen. (2009). High genetic differentiation in marginal populations of European white elm (Ulmus laevis). Silva Fennica vol. 43 no. 2 article id 205. https://doi.org/10.14214/sf.205

Keywords: marginal populations; genetic differentiation; genetic conservation; Ulmus

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Studies on the amount of genetic variation in marginal populations and differentiation between them are essential for assessment of best gene conservation strategies and sampling schemes. Thirteen marginal populations of Ulmus laevis in southern Finland and one in Estonia were investigated for genetic variation in 20 allozyme loci. Population differentiation among Finnish stands was high, F_st = 0.290, and mean genetic diversity low, H_e = 0.088. The differentiation follows the isolation-by-distance structure within the core of the distribution area (lake Vanajavesi). Fairly high frequency of recurrent genotypes was observed, but this did not have an influence on the genetic parameters. The observed genetic structure is consistent with the central-marginal hypothesis. In the light of the results, the Finnish gene conservation strategy for U. laevis seems to be on a sound basis.

Vakkari, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: pekka.vakkari@metla.fi
Rusanen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: mr@nn.fi
Kärkkäinen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: kk@nn.fi

article id 464, category Research article

Christian Kanzian, Franz Holzleitner, Karl Stampfer, Sarah Ashton. (2009). Regional energy wood logistics – optimizing local fuel supply. Silva Fennica vol. 43 no. 1 article id 464. https://doi.org/10.14214/sf.464

Keywords: logistics; energy wood; transport optimization; GIS

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The promotion of electric energy production from solid biomass by the Austrian government has lead to a boom in the construction of new combined heat and power plants. The current total demand for wood chips in the research area for energy purposes is 70 400 m³ of loose volume chips per year. The expected increase in demand due to these new plants is more than 4 times greater than current demand: up to 302 700 m³ of loose volume per year. Even if the energy wood feedstock potential is satisfactory, the design of the supply chain is still unresolved. The aim of this study is to give decision-makers a base for further development. To accomplish this, we designed and tested four different supply scenarios: one for 9 plants and one for 16 plants. The scenarios were developed using a combination of geographic information systems (GIS) and linear programming methods. The results indicate that direct transport of solid fuel wood as round wood and chipping at the plant is the cheapest supply system with a resulting cost of 5.6–6.6 EUR/m³ loose. Using harvesting residues can only be recommended for large plants because of poor fuel quality. In this case, residues would be chipped at or near the landing, piled and transported via self-loading trucks at a cost between 8.4 and 9.1 EUR/m³ loose. In order to meet increasing demand and to ensure a continuous supply, especially during the winter and spring seasons it is necessary to optimize the supply chain by including storage terminals. However, using terminals and increased demand both lead to higher logistical costs. For example, if the total volume is handled via terminals, the average supply costs including storage will increase by 26%. Higher demand increases the costs by 24%.

Kanzian, University of Applied Life Sciences Vienna, Institute of Forest Engineering, Peter Jordan Strasse 82, A-1190 Vienna, Austria E-mail: christian.kanzian@boku.ac.at
Holzleitner, University of Applied Life Sciences Vienna, Institute of Forest Engineering, Peter Jordan Strasse 82, A-1190 Vienna, Austria E-mail: fh@nn.at
Stampfer, University of Applied Life Sciences Vienna, Institute of Forest Engineering, Peter Jordan Strasse 82, A-1190 Vienna, Austria E-mail: ks@nn.at
Ashton, Southern Regional Extension Forestry, Forestry Bldg. 4-420, University of Georgia, Athens, GA 30602, USA E-mail: sa@nn.us

article id 222, category Research article

Yi Pan, Yaoqi Zhang, Indrajit Majumdar. (2009). Population, economic welfare and holding size distribution of private forestland in Alabama, USA. Silva Fennica vol. 43 no. 1 article id 222. https://doi.org/10.14214/sf.222

Keywords: consolidation; private forestland owners; parcellation; land tenure

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This paper analyzes structural differences in holding size distribution of private forestland at the county level in Alabama, USA. The following categories are used to describe the timberland holding distribution: mean holding size, Gini coefficient of timberland holdings, and share of county timberland in holding sizes of fewer than 80 hectares and more than 800 hectares. Regressions are conducted to explore the socio-economic correlates of forestland holding size distribution. Results indicate that population density, age structure, urban-rural population structure, income structure and land tenure type are strongly associated with the forest holding size distribution.

Pan, Auburn University, School of Forestry and Wildlife Sciences, Auburn, AL 36849, USA E-mail: yp@nn.us
Zhang, Auburn University, School of Forestry and Wildlife Sciences, Auburn, AL 36849, USA E-mail: yaoqi.zhang@auburn.edu
Majumdar, Auburn University, School of Forestry and Wildlife Sciences, Auburn, AL 36849, USA E-mail: im@nn.us

article id 221, category Research article

Glen Murphy, Rod Brownlie, Mark Kimberley, Peter Beets. (2009). Impacts of forest harvesting related soil disturbance on end-of-rotation wood quality and quantity in a New Zealand radiata pine forest. Silva Fennica vol. 43 no. 1 article id 221. https://doi.org/10.14214/sf.221

Keywords: Pinus radiata; harvesting; compaction; wood density; tree growth; stiffness; litter removal; nitrogen deficiency

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The long-term effect of soil disturbance (litter removal, topsoil removal and compaction) from forest harvesting on wood quality and quantity of second-rotation Pinus radiata growing on a clay loam soil, was assessed at the end the rotation, 26 years after planting. Relative to Control plots, average tree and stand total volume at rotation end was not significantly affected by litter removal and nil or light compaction, but was significantly reduced by 28% by litter and topsoil removal and moderate subsoil compaction, and further reduced by 38% by heavy compaction. Wood density at breast height in the inner rings of trees in the most disturbed treatments was elevated by up to 30 kg m^–3. This occurred because these treatments were more N deficient as reflected by foliar N levels during the first 11 years of growth relative to the Control. However, no treatment differences in wood density were evident in outer rings, and by rotation age overall mean density did not differ significantly between treatments. Neither acoustic velocity of standing trees, nor acoustic velocity of logs, was significantly affected by soil disturbance, indicating that stiffness of lumber cut from trees in the trial was likely to be similar for all treatments. Economic impacts of soil disturbance and compaction on this soil type will therefore result largely from the considerable negative impacts on final tree size, with little or no compensation from improved wood properties.

Murphy, Forest Engineering, Resources and Management Department, Oregon State University, Corvallis, Oregon, USA E-mail: glen.murphy@oregonstate.edu
Brownlie, Scion Research, Rotorua, New Zealand E-mail: rb@nn.nz
Kimberley, Scion Research, Rotorua, New Zealand E-mail: mk@nn.nz
Beets, Scion Research, Rotorua, New Zealand E-mail: pb@nn.nz

article id 220, category Research article

Jani Heikkilä, Matti Sirén, Anssi Ahtikoski, Jari Hynynen, Tiina Sauvula, Mika Lehtonen. (2009). Energy wood thinning as a part of stand management of Scots pine and Norway spruce. Silva Fennica vol. 43 no. 1 article id 220. https://doi.org/10.14214/sf.220

Keywords: energy wood thinning; stand management; MOTTI simulator

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The effects of combined production of industrial and energy wood on yield and harvesting incomes, as well as the feasibility of energy wood procurement, were studied. Data for 22 Scots pine (Pinus sylvestris L.) and 21 Norway spruce (Picea abies (L.) Karst.) juvenile stands in Central and Southern Finland were used to compare six combined production regimes to conventional industrial wood production. The study was based on simulations made by the MOTTI stand simulator, which produces growth predictions for alternative management regimes under various site and climatic conditions. The combined production regimes included precommercial thinning at 4–8 m dominant height to a density of 3000–4000 stems ha^–1 and energy wood harvesting at 8, 10 or 12 m dominant height. Combined production did not decrease the total yield of industrial wood during the rotation period. Differences in the mean annual increment (MAI) were small, and the rotation periods varied only slightly between the alternatives. Combined production regime can be feasible for a forest owner if the price of energy wood is 3–5 EUR m^–3 in pine stands, and 8–9 EUR m^–3 in spruce stands. Energy wood procurement was not economically viable at the current energy price (12 EUR MWh^–1) without state subsidies. Without subsidies a 15 EUR MWh^–1 energy price would be needed. Our results imply that the combined production of industrial and energy wood could be a feasible stand management alternative.

Heikkilä, L&T Biowatti Oy, P.O. Box 738, FI-60101 Seinäjoki, Finland E-mail: jani.heikkila@biowatti.fi
Sirén, Finnish Forest Research Institute, Vantaa Research Unit, P.O.Box 18, FI-01301 Vantaa, Finland E-mail: ms@nn.fi
Ahtikoski, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O.Box 16, FI-96301 Rovaniemi, Finland E-mail: aa@nn.fi
Hynynen, Finnish Forest Research Institute, Vantaa Research Unit, P.O.Box 18, FI-01301 Vantaa, Finland E-mail: jh@nn.fi
Sauvula, Seinäjoki University of Applied Sciences, School of Agriculture and Forestry, Tuomarniementie 55, FI-63700 Ähtäri, Finland E-mail: ts@nn.fi
Lehtonen, Finnish Forest Research Institute, Vantaa Research Unit, P.O.Box 18, FI-01301 Vantaa, Finland E-mail: ml@nn.fi

article id 219, category Research article

Arto Haara, Pekka Leskinen. (2009). The assessment of the uncertainty of updated stand-level inventory data. Silva Fennica vol. 43 no. 1 article id 219. https://doi.org/10.14214/sf.219

Keywords: uncertainty; measurement error; simulation; non-parametric methods; observed error; stand-level inventory

Abstract | View details | Full text in PDF | Author Info

Predictions of growth and yield are essential in forest management planning. Growth predictions are usually obtained by applying complex simulation systems, whose accuracy is difficult to assess. Moreover, the computerised updating of old inventory data is increasing in the management of forest planning systems. A common characteristic of prediction models is that the uncertainties involved are usually not considered in the decision-making process. In this paper, two methods for assessing the uncertainty of updated forest inventory data were studied. The considered methods were (i) the models of observed errors and (ii) the k-nearest neighbour method. The derived assessments of uncertainty were compared with the empirical estimates of uncertainty. The practical utilisation of both methods was considered as well. The uncertainty assessments of updated stand-level inventory data using both methods were found to be feasible. The main advantages of the two studied methods include that bias as well as accuracy can be assessed.

Haara, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: arto.haara@joensuu.fi
Leskinen, Finnish Environment Institute, Research Programme for Production and Consumption, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: pl@nn.fi

article id 218, category Research article

Md. Nurul Islam, Mikko Kurttila, Lauri Mehtätalo, Arto Haara. (2009). Analyzing the effects of inventory errors on holding-level forest plans: the case of measurement error in the basal area of the dominated tree species. Silva Fennica vol. 43 no. 1 article id 218. https://doi.org/10.14214/sf.218

Keywords: inoptimality loss; dominated tree species; erroneous inventory data; forest plan

Abstract | View details | Full text in PDF | Author Info

Accurate inventory data are required for ensuring optimal net return on investment from the forest. Erroneous data can lead to the formulation of a non-optimal plan that can cause inoptimality losses. Little is known of the effect of using erroneous stand inventory data in preparing holding-level forest plans. This study reports on an approach for analyzing such inoptimality losses. Furthermore, inoptimality losses caused by measurement errors in the basal area of the dominated tree species were investigated in a case study. Based on the inventory data including routine measurements by 67 measurers, four measurer groups were created with different measurement error profiles for the basal area of the dominated tree species. This was followed by measurement error simulations for each group and by adding these to the accurate control inventory data to create erroneous data of different error profiles. Three different forest plans were then constructed by using erroneous data of each group. The plans were then analyzed and compared with plans based on correct data. The effect of measurement errors on the net present value from the whole planning period, and on the amount of remaining growing stock at the end of planning period, were analyzed and utilized in calculating the inoptimality losses. It was concluded that even errors involving dominated tree species can cause significant changes in the holding-level forest plans.

Islam, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: nurul.islam@joensuu.fi
Kurttila, Finnish Forest Research Institute, Joensuu Research Unit, FI-80101 Joensuu, Finland E-mail: mk@nn.fi
Mehtätalo, University of Helsinki, Dept. of Forest Resource Management, FI-00014 University of Helsinki, Finland E-mail: lm@nn.fi
Haara, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: ah@nn.fi

article id 217, category Research article

Ville Kankaanhuhta, Timo Saksa, Heikki Smolander. (2009). Variation in the results of Norway spruce planting and Scots pine direct seeding in privately-owned forests in southern Finland. Silva Fennica vol. 43 no. 1 article id 217. https://doi.org/10.14214/sf.217

Keywords: Pinus sylvestris; Picea abies; quality control; forest regeneration survey; forest regeneration inventory; quality management

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This study describes the variation in the planting results for 3-year-old Norway spruce (Picea abies (L.) Karst.) and 4-year-old Scots pine (Pinus sylvestris L.) using direct seeding in privately-owned regeneration areas in southern Finland. The study material consists of operative forest regeneration quality management inventory areas from the years 2000–2006. The effect of both the regional and the administrative levels as well as ecological factors was modelled on the basis of the hierarchy structure forestry centre, Forest Owners’ Association (= FOA), forestry professional, regeneration area and sample plot. The major part of the variation occurred at the sample plot and regeneration area level. Particular attention was paid to observation of the clustered spatial distribution of Scots pine seedlings. The FOA and forestry professional levels explained 5% of the variation in Norway spruce planting and 11% of the variation in Scots pine direct seeding. Applied forest regeneration operations, site and soil characteristics were included in the fixed effects. In the planting of Norway spruce the most important factor explaining the regeneration result was soil preparation. Mounding produced better results than patching and disc trenching. The site and soil characteristics were other important factors in the operations. The selection of direct seeding of Scots pine on too fertile, fine textured or moist sites yielded poor results.

Kankaanhuhta, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: ville.kankaanhuhta@metla.fi
Saksa, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: ts@nn.fi
Smolander, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: hs@nn.fi

article id 216, category Research article

Juha Siitonen, Jenni Hottola, Auli Immonen. (2009). Differences in stand characteristics between brook-side key habitats and managed forests in southern Finland. Silva Fennica vol. 43 no. 1 article id 216. https://doi.org/10.14214/sf.216

Keywords: dead wood; woodland key habitats; stand structure; CWD; WKH; Forest Act

Abstract | View details | Full text in PDF | Author Info

Preservation of small habitat patches termed as “woodland key habitats” or “especially important habitats” in the Finnish Forest Act has become an integral part of biodiversity-oriented forest management. Forest Act habitats belong to particular habitat types defined in the act, and they are supposed to have natural-like stand characteristics. However, very little is known about the actual stand structure in the designated habitats. Our aim was to compare stand characteristics between brook-side key habitats and comparable managed forests as controls. Seven study areas were selected from four regions across southern Finland. Within each study area ten key habitats and ten controls (140 stands) were randomly selected. Living and dead trees and cut stumps were measured in each stand within a 0.2 ha plot. The average degree of previous cutting was significantly lower whereas the volume of dead wood, volume of deciduous trees, and stand diversity were each significantly higher in key habitats than controls. The average volume of dead wood was 11.7 m³ ha^–1 in key habitats and 6.5 m³ ha^–1 in controls. However, there was considerable variation among individual stands, and a large part of key habitats could not be distinguished from randomly selected control stands with respect to stand characteristics. The preservation of natural brook channels with their immediate surroundings is undoubtedly important for maintaining aquatic and semiaquatic biodiversity. Nevertheless, when complementing the forest conservation network in the future, main emphasis in selecting potentially valuable stands should be placed on important structural features such as dead wood and old trees.

Siitonen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: juha.siitonen@metla.fi
Hottola, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: jh@nn.fi
Immonen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: ai@nn.fi

article id 215, category Research article

Antti J. Lukkarinen, Seppo Ruotsalainen, Teijo Nikkanen, Heli Peltola. (2009). The growth rhythm and height growth of seedlings of Siberian (Larix sibirica Ledeb.) and Dahurian (Larix gmelinii Rupr.) larch provenances in greenhouse conditions. Silva Fennica vol. 43 no. 1 article id 215. https://doi.org/10.14214/sf.215

Keywords: height growth; larch; Larix; provenance; growth rhythm

Abstract | View details | Full text in PDF | Author Info

The aim of this study was to determine whether the growth rhythm and height growth differ between various larch provenances grown in greenhouse conditions. We also investigated whether the geographic and climatic conditions at the origin of the provenance could explain the possible differences between the provenances. The study material consisted of 16 Russian Siberian (Larix sibirica Ledeb.) and Dahurian larch (Larix gmelinii Rupr.) provenances and four seed sources from Finland as comparison lots. The growth rhythm differences were clearest between the southern and northern provenances; the southern provenances grew for a longer period and the proportion of late summer height growth was larger. Autumn colouration also developed later in these provenances. In the Russian larch material the provenances with a longer growing period had greater height growth. In the whole material the relationship was not so linear due to the deviating behaviour of the comparison material. Several of the Russian larch provenances were taller than the Finnish comparison seed lots, although the difference was not statistically significant. However, the growth of these provenances must be followed for a much longer period in field trials before any conclusions can be drawn about their usability in practical forestry.

Lukkarinen, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: antti.lukkarinen@pp.inet.fi
Ruotsalainen, Finnish Forest Research Institute, Punkaharju Research Unit, Finlandiantie 18, FI-58450 Punkaharju, Finland E-mail: sr@nn.fi
Nikkanen, Finnish Forest Research Institute, Punkaharju Research Unit, Finlandiantie 18, FI-58450 Punkaharju, Finland E-mail: tn@nn.fi
Peltola, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: hp@nn.fi

article id 214, category Research article

Michelle de Chantal, Hannu Rita, Urban Bergsten, Mikaell Ottosson Löfvenius, Harald Grip. (2009). Frost heaving of Picea abies seedlings as influenced by soil preparation, planting technique, and location along gap-shelterwood gradients. Silva Fennica vol. 43 no. 1 article id 214. https://doi.org/10.14214/sf.214

Keywords: deep planting; normal planting; mobile container; fixed container; HuMinMix; soil horizon

Abstract | View details | Full text in PDF | Author Info

The effects of soil preparation, planting technique and location along gap-shelterwood gradients (position and orientation) on frost heaving damage to seedlings were studied in Vindeln Experimental Forests, northern Sweden. The forest was harvested in a grid pattern in winter 2004–2005, forming gaps and shelterwood areas of 30 x 40 m each. Gap-shelterwood gradients were delimited in four orientations and subdivided into five positions: 7 m and 15 m into the gap and shelterwood, and at the gap edge. At each position, three replicates of three soil preparations were made: exposed E and B horizons and HuMinMix (milled vegetation and humus layers mixed with surface mineral soil). In early October 2005, one-year-old containerized Picea abies (L.) Karst. seedlings were planted using four techniques: normal and deep planting, and mobile and fixed experimental containers. After one winter, frost heaving damage was highest for seedlings on B horizon combined with the mobile container (51 ± 6%) and normal planting (43±6%). Normal- or deep-planted seedlings in HuMinMix had the least damage (5–6.6 ± 2.5%). Compared to normal planting, deep planting reduced frost heaving damage only on B horizon. When considering the orientation, seedlings in the experimental containers had more or similar frost heaving damage than normal- or deep-planted seedlings. Along the eastern gradient, seedlings incurred more frost heaving damage in the center of the gap than under the canopy.

Chantal, University of Helsinki, Dept of Forest Ecology & Dept of Forest Resource Management, Helsinki, Finland E-mail: michelle.dechantal@helsinki.fi
Rita, University of Helsinki, Dept of Forest Ecology & Dept of Forest Resource Management, Helsinki, Finland E-mail: hr@nn.fi
Bergsten, Swedish University of Agricultural Sciences, Dept of Forest Ecology and Management, Umeå, Sweden E-mail: ub@nn.se
Löfvenius, Swedish University of Agricultural Sciences, Dept of Forest Ecology and Management, Umeå, Sweden E-mail: mol@nn.se
Grip, Swedish University of Agricultural Sciences, Dept of Forest Ecology and Management, Umeå, Sweden E-mail: hg@nn.se

Category : Review article

article id 463, category Review article

Thomas Gschwantner, Klemens Schadauer, Claude Vidal, Adrian Lanz, Erkki Tomppo, Lucio di Cosmo, Nicolas Robert, Daisy Englert Duursma, Mark Lawrence. (2009). Common tree definitions for national forest inventories in Europe. Silva Fennica vol. 43 no. 2 article id 463. https://doi.org/10.14214/sf.463

Keywords: harmonisation; COST Action E43; tree definitions; shrub definitions; tree elements; tree variables; tree characteristics

Abstract | View details | Full text in PDF | Author Info

At the international level, various definitions have been established for the compilation and publication of forest resources assessment results over the last decade. These international definitions frequently rely on terms that are not precisely specified for inventory purposes and do not completely cover the requirements arising from the application of National Forest Inventory (NFI) data. Also, with respect to conventional topics such as forest area and growing stock estimation, several terms and expressions referring to individual trees are not, or are only vaguely, defined until now. Since the individual tree is the basic element of any forest resources assessment, the clarification of tree-related terms is an important part of COST Action E43 to harmonise common reporting of National Forest Inventories. Based on a review of existing definitions and on the requirements for harmonised reporting, common tree-related definitions are established. One objective of this study is to refine and enhance the applicability of available tree and shrub definitions, in particular with regard to the distinction between trees and shrubs. The study also focuses on the parts or “elements” of trees and on the distinction between these elements as they are of particular importance in growing stock and biomass definitions. Furthermore, several definitions for tree characteristics such as “living” and “standing”, as well as tree variables such as height, length, diameter at breast height, and crown projection area are adjusted with respect to NFI purposes. A concluding discussion reflects upon the reviewed, refined and newly established definitions. The definitions presented in this paper provide a firm basis for a common set of harmonised reference definitions developed by COST Action E43 and contribute to the precise and consistent use of terms.

Gschwantner, Federal Research and Training Centre for Forests, Natural Hazards and Landscape, Wien, Austria E-mail: thomas.gschwantner@bfw.gv.at
Schadauer, Federal Research and Training Centre for Forests, Natural Hazards and Landscape, Wien, Austria E-mail: ks@nn.at
Vidal, French National Forest Inventory, Nogent-sur-Vernisson, France E-mail: cv@nn.fr
Lanz, Swiss Federal Institute for Forest, Snow and Landscape Research, Birmensdorf, Switzerland E-mail: al@nn.ch
Tomppo, Finnish Forest Research Institute, Vantaa, Finland E-mail: et@nn.fi
di Cosmo, Agricultural Research Council – Forest Monitoring and Planning Research Unit, Trento, Italy E-mail: ldc@nn.it
Robert, French National Forest Inventory, Nogent-sur-Vernisson, France E-mail: nr@nn.fr
Englert Duursma, Finnish Forest Research Institute, Vantaa, Finland E-mail: ded@nn.fi
Lawrence, Northern Research Station, Roslin, Midlothian, Great Britain E-mail: ml@nn.uk

Category : Research note

article id 223, category Research note

Bum-Jin Park, Takeshi Morikawa, Tomohiro Ogata, Kenji Washida, Mario Iwamoto, Hirohiko Nakamura, Yoshifumi Miyazaki. (2009). Physiological effects of ingesting eucalyptus essential oil with milk casein peptide. Silva Fennica vol. 43 no. 1 article id 223. https://doi.org/10.14214/sf.223

Keywords: relaxation; eucalyptus essential oil; POMS; salivary cortisol; type A behavior pattern

Abstract | View details | Full text in PDF | Author Info

This study was conducted to clarify the effect of eucalyptus essential oil mixed with milk casein peptide food for human physiological relaxation. Fifteen male university students (21.2 ± 0.9 yr) participated in study as subjects. The subjects were given one of two types of experimental drink (peptide + eucalyptus flavor (Pep + EF), and peptide + grapefruit·orange flavor (Pep + G·O), each flavor contains natural essential oil). We measured the change in salivary cortisol concentration and POMS scores before and two hours after taking experimental drink. The results of a Type A behavior pattern test were used to classify subjects. The concentration of salivary cortisol decreased significantly two hours after taking Pep + EF. And Type B showed bigger change than Type A. In conclusion, the results show that eucalyptus essential oil has the effect of relaxation, and that the effects on Type A and Type B are different.

Park, Chiba University, Center for Environment, Health and Field Sciences, 277-0882 Kashiwa, Chiba, Japan E-mail: bjpark@faculty.chiba-u.jp
Morikawa, Chiba University, Center for Environment, Health and Field Sciences, 277-0882 Kashiwa, Chiba, Japan E-mail: tm@nn.jp
Ogata, Chiba University, Center for Environment, Health and Field Sciences, 277-0882 Kashiwa, Chiba, Japan E-mail: to@nn.jp
Washida, Chiba University, Center for Environment, Health and Field Sciences, 277-0882 Kashiwa, Chiba, Japan E-mail: kw@nn.jp
Iwamoto, Chiba University, Center for Environment, Health and Field Sciences, 277-0882 Kashiwa, Chiba, Japan E-mail: mi@nn.jp
Nakamura, Chiba University, Center for Environment, Health and Field Sciences, 277-0882 Kashiwa, Chiba, Japan E-mail: hn@nn.jp
Miyazaki, Chiba University, Center for Environment, Health and Field Sciences, 277-0882 Kashiwa, Chiba, Japan E-mail: ym@nn.jp

Category : Commentary

article id 461, category Commentary

Sari Palmroth. (2009). Boreal forest and climate change – from processes and transport to trees, ecosystems and atmosphere. Silva Fennica vol. 43 no. 4 article id 461. https://doi.org/10.14214/sf.461

Keywords: carbon; climate; growth; forests; ecology; book reviews

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Book review: Hari, P. & Kulmala, L. (eds.). 2008. Boreal Forest and Climate Change. Advances in Global Change Research 34. Springer. 582 p. ISBN 978-1-4020-8717-2.

Palmroth, Duke University, Nicholas School of the Environment, Box 90328 Duke University, Durham, USA E-mail: sari.palmroth@duke.edu

article id 462, category Commentary

Jean-Michel Leban, Heli Peltola, Sanna Leinonen, Christine Todoroki, Sophie D'Amours, Barry Gardiner, Jim Goudie, Eini Lowell, Thomas Seifert, Erkki Verkasalo, Lars Wilhelmsson, Tuula Nuutinen. (2009). Preface. Silva Fennica vol. 43 no. 3 article id 462. https://doi.org/10.14214/sf.462

Abstract | View details | Full text in PDF | Author Info

The 6th IUFRO Workshop on Connection between Forest Resources and Wood Quality: Modelling Approaches and Simulation Software.

Leban, E-mail: jml@nn.fr
Peltola, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: heli.peltola@uef.fi
Leinonen, E-mail: sl@nn.fi
Todoroki, E-mail: ct@nn.nz
D'Amours, E-mail: sda@nn.ca
Gardiner, E-mail: bg@nn.uk
Goudie, E-mail: jg@nn.ca
Lowell, E-mail: el@nn.us
Seifert, E-mail: ts@nn.de
Verkasalo, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101, Joensuu, Finland E-mail: erkki.verkasalo@metla.fi
Wilhelmsson, E-mail: lw@nn.se
Nuutinen, European Forest Institute, Torikatu 34, FI-80100 Joensuu, Finland E-mail: tuula.nuutinen@efi.int

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