Category :
                    
                    Article
                                    
                            
                    
        
            
            article id 5583,
                            category
                        Article
                    
        
        
                            Ilkka Leinonen,
                            Heikki Hänninen,
                            Tapani Repo.
                    
                    
                (1996).
            
                            
                                    Testing of frost hardiness models for Pinus sylvestris in natural conditions and in elevated temperature.
                            
                            
                Silva Fennica
                                                            vol.
                                        30
                                                                            no.
                                        2–3
                                article id 5583.
            
                            
                https://doi.org/10.14214/sf.a9228
            
             
        
                                    
                                    
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                            Two dynamic models predicting the development of frost hardiness of Finnish Scots pine (Pinus sylvestris L.) were tested with frost hardiness data obtained from trees growing in the natural conditions of Finland and from an experiment simulating the predicted climatic warming. The input variables were temperature in the first model, and temperature and night length in the second. The model parameters were fixed on the basis of previous independent studies. The results suggested that the model which included temperature and photoperiod as input variables was more accurate than the model using temperature as the only input variable to predict the development of frost hardiness in different environmental conditions. Further requirements for developing the frost hardiness models are discussed.
                
                                            - 
                            Leinonen,
                            
                                                        E-mail:
                                                            il@mm.unknown
                                                                                          
- 
                            Hänninen,
                            
                                                        E-mail:
                                                            hh@mm.unknown
                                                                                
- 
                            Repo,
                            
                                                        E-mail:
                                                            tr@mm.unknown
                                                                                
 
         
     
 
            
        
            
            article id 5433,
                            category
                        Article
                    
        
        
                                    
                                    
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                            The relationships between bud dormancy and frost hardiness were examined using two-year-old Pinus sylvestris L. seedlings. The chilling temperatures used were +4 and -2°C. To examine the dormancy release of the seedlings, a forcing technique was used. Frost hardiness was determined by artificial freezing treatments and measurements of electrical impedance. At the start of the experiment, the frost hardiness of the seedlings was about -25°C. After the rest break, the seedlings kept at +4°C dehardened until after eight weeks their frost hardiness reached -5°C. At the lower chilling temperature (-2°C) the frost hardiness remained at the original level. When moved from +4 to -2°C, seedlings were able to reharden only after the time required for bud burst in the forcing conditions had reached the minimum.
The PDF includes an abstract in Finnish
                
                                            - 
                            Valkonen,
                            
                                                        E-mail:
                                                            mv@mm.unknown
                                                                                          
- 
                            Hänninen,
                            
                                                        E-mail:
                                                            hh@mm.unknown
                                                                                
- 
                            Pelkonen,
                            
                                                        E-mail:
                                                            pp@mm.unknown
                                                                                
- 
                            Repo,
                            
                                                        E-mail:
                                                            tr@mm.unknown
                                                                                
 
         
     
 
            
        
            
            article id 5369,
                            category
                        Article
                    
        
        
                            Helen J. Jozefek.
                    
                    
                (1989).
            
                            
                                    The effect of varying levels of potassium on the frost resistance of birch seedlings.
                            
                            
                Silva Fennica
                                                            vol.
                                        23
                                                                            no.
                                        1
                                article id 5369.
            
                            
                https://doi.org/10.14214/sf.a15528
            
             
        
                                    
                                    
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                            Seven hundred one-year-old Betula pendula Roth seedlings were given different concentrations of potassium fertilizer. Over the study period seedlings were subjected to artificial growing and dormant phases. Frost resistance of the seedlings was assessed by artificial freezing tests and electrical impedance measurements on stem cuttings. In general, high concentrations of potassium fertilizer reflected a low tolerance to frost. Pre-freezing impedance readings decreased with increasing potassium fertilizer dosages. Results from pre-freezing impedance measurements were found to be in broad agreement with the hypothesis that high impedance readings indicate a frost hardy tissue whereas low readings imply the opposite.
The PDF includes an abstract in Finnish.
                
                                            - 
                            Jozefek,
                            
                                                        E-mail:
                                                            hj@mm.unknown
                                                                                          
 
         
     
 
            
        
            
            article id 5353,
                            category
                        Article
                    
        
        
                            Øystein Johnsen,
                            Inger Apeland.
                    
                    
                (1988).
            
                            
                                    Screening early autumn frost hardiness among progenies from Norway spruce seed orchards.
                            
                            
                Silva Fennica
                                                            vol.
                                        22
                                                                            no.
                                        3
                                article id 5353.
            
                            
                https://doi.org/10.14214/sf.a15510
            
             
        
                                    
                                    
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                            Nursery grown Norway spruce (Picea abies (L.) H. Karst.) seedlings from 12 different seed orchards were tested for early autumn frost hardiness using artificial freezing tests. Seed orchards containing grafted parent clones originating from high altitudes produced seedlings showing higher damage than commercial control seed lots of the commercial controls. A seed orchard containing both German and Norwegian clones produced seedlings showing high damage. The correlation between bud-set and frost damage was high at the provenance level, but lower at the half- and full-sib-levels. Families with good growth capacity in progeny field tests showed large between-family variation in frost damage in the artificial freezing tests. This indicates the possibility to combine high growth rate with acceptable autumn frost hardiness in the selection of parent trees.
The PDF includes an abstract in Finnish.
                
                                            - 
                            Johnsen,
                            
                                                        E-mail:
                                                            –
                                                                                
- 
                            Apeland,
                            
                                                        E-mail:
                                                            ia@mm.unknown
                                                                                
 
         
     
 
            
        
            
            article id 5352,
                            category
                        Article
                    
        
        
                            Lars Christersson,
                            Heinrich A. von Fircks.
                    
                    
                (1988).
            
                            
                                    Injuries to conifer seedlings caused by simulated summer frost and winter desiccation.
                            
                            
                Silva Fennica
                                                            vol.
                                        22
                                                                            no.
                                        3
                                article id 5352.
            
                            
                https://doi.org/10.14214/sf.a15509
            
             
        
                                    
                                    
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                            Visible frost damage to forest trees in Sweden seldom occurs in winter but is frequent in late spring, summer and early autumn. Frosts are frequent in all seasons in various parts of Sweden, even in the southernmost part (lat. 56°, N) and temperatures may be as low as -10°C even around mid-summer. Ice crystal formation within the tissues, which in most seedlings takes place at around -2°C, causes injury, not the sub-zero temperatures themselves.
The apical meristem, the elongated zone, and the needles of seedlings of Picea abies (L.) H. Karst. in a growing phase were damaged at about -3°C and those of Pinus sylvestris L. at about -6°C. Other species of the genus Pinus were tested and most were found to be damaged at about -6°C, with some variations. Picea species tested were damaged at about -3°C to -4°C.
A method has been designed to compare the response of different species to winter desiccation, which occurs under conditions of (1) low night temperature, (2) very high irradiation and increase in needle temperature during the photoperiod, (3) frozen soil, and (4) low wind speed. There were differences in response to winter desiccation between pine and spruce species. Seedlings of Pinus contorta tolerated these winter desiccation conditions much better than those of P. sylvestris or Picea abies. Picea mariana was the least tolerant of the species tested.
The PDF includes an abstract in Finnish.
                
                                            - 
                            Christersson,
                            
                                                        E-mail:
                                                            lc@mm.unknown
                                                                                          
- 
                            Fircks,
                            
                                                        E-mail:
                                                            hf@mm.unknown
                                                                                
 
         
     
 
            
        
            
            article id 5350,
                            category
                        Article
                    
        
        
                            Heikki Hänninen,
                            Paavo Pelkonen.
                    
                    
                (1988).
            
                            
                                    Frost hardiness and over-wintering of forest trees.
                            
                            
                Silva Fennica
                                                            vol.
                                        22
                                                                            no.
                                        3
                                article id 5350.
            
                            
                https://doi.org/10.14214/sf.a15507
            
             
        
                                    
                                    
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                            This issue of Silva Fennica consists of eight articles, which are based on a co-nordic conference ”Frost hardiness and over-wintering in forest tree seedlings”, held in Joensuu, Finland, during December 1–3, 1986. The whole annual cycle of the trees is considered. Emphasis is given on methods for the study of frost hardiness, genetic variation in frost hardiness, nitrogen metabolism, bud dormancy release, and joint effect of natural and anthropogenic stress factors in the winter damage of forest trees. Practical implications for tree breeding and nursery management are discussed.
The PDF includes an abstract in Finnish.
                
                                            - 
                            Hänninen,
                            
                                                        E-mail:
                                                            –
                                                                                
- 
                            Pelkonen,
                            
                                                        E-mail:
                                                            –
                                                                                
 
         
     
 
                        
                
                
                                            Category :
                    
                    Research article
                                    
                            
                    
        
            
            article id 45,
                            category
                        Research article
                    
        
        
                            Guolei Li,
                            Yong Liu,
                            Yan Zhu,
                            Qing Mei Li,
                            R. Karsten Dumroese.
                    
                    
                (2012).
            
                            
                                    Effect of fall-applied nitrogen on growth, nitrogen storage and frost hardiness of bareroot Larix olgensis seedlings.
                            
                            
                Silva Fennica
                                                            vol.
                                        46
                                                                            no.
                                        3
                                article id 45.
            
                            
                https://doi.org/10.14214/sf.45
            
             
        
                                    
                                    
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                            Nursery response of evergreen trees to fall fertilization has been  studied widely, but little attention has been given to deciduous trees.  Bareroot Olga Bay larch (Larix olgensis Henry) seedlings were fertilized  in the nursery with urea at four rates (0, 30, 60, 90 kg N ha–1),  with half of each rate applied on two dates (September 16 and October  1, 2009). The seedlings were excavated for evaluation on October 15. In  the unfertilized (control) treatment, root and shoot dry mass increased  by 100% and 57% respectively, while N concentration in the roots and  shoots increased by 43% and 40% during the 30 day period. This indicated  that substantial biomass growth during this period did not lead to  internal nutrient dilution. Root dry mass increased when fall  fertilization rates were ≥ 60 kg N ha–1. Fall fertilization  increased N concentrations in root tissue by 48–73%. Compared with the  control, shoot tissues of fall fertilized seedlings had slightly higher N  concentration and content and significantly higher frost hardiness.
                        
                
                                            - 
                            Li,
                            Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing 100083, China
                                                        E-mail:
                                                            gl@nn.cn
                                                                                
- 
                            Liu,
                            Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing 100083, China
                                                        E-mail:
                                                            lyong@bjfu.edu.cn
                                                                                          
- 
                            Zhu,
                            Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing 100083, China
                                                        E-mail:
                                                            yz@nn.cn
                                                                                
- 
                            Li,
                            Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Forest Silviculture of State Forestry Administration, Beijing 100091, China
                                                        E-mail:
                                                            qml@nn.cn
                                                                                
- 
                            Dumroese,
                            US Department of Agriculture, Forest Service, Rocky Mountain Research Station, Moscow, ID, USA
                                                        E-mail:
                                                            rkd@nn.us
                                                                                
 
         
     
 
            
        
            
            article id 209,
                            category
                        Research article
                    
        
        
                            Jaana Luoranen,
                            Kyösti Konttinen,
                            Risto Rikala.
                    
                    
                (2009).
            
                            
                                    Frost hardening and risk of a second flush in Norway spruce seedlings after an early-season short-day treatment.
                            
                            
                Silva Fennica
                                                            vol.
                                        43
                                                                            no.
                                        2
                                article id 209.
            
                            
                https://doi.org/10.14214/sf.209
            
             
        
                                    
                                    
                            Abstract |
                        
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                            There have been years in Finland when container seedlings of Norway  spruce (Picea abies (L.) Karsten) planted in the summer have been  damaged by early-autumn frosts. For August and September plantings, the  seedlings can be hardened by means of short-day (SD) treatment, but  little information is available about its usability for earlier  plantings. We studied the effects of early-season SD treatment on the  frost hardiness and risk of a second flush of Norway spruce seedlings.  In three successive years, second-year seedlings were grown in a  greenhouse or outdoors in the spring and early summer and then subjected  to two or three-week SD treatment beginning on the second, third, or  fourth week of June. We monitored the height growth cessation, bud  formation, and frost hardiness of the seedlings in the nursery. All SD  treatments made the height growth cease, but the risk of a second flush  increased if the temperature sum was less than 300 d.d. before the  beginning of the SD treatment or more than 450 d.d. between the end of  the treatment and mid-August. Clearly, then, SD treatment reduced the  risk of a second flush in seedlings that had been grown in a greenhouse  in the spring. Early-season SD treatment increased the frost hardiness  of both needles and stems for late July to early September in comparison  with untreated seedlings. Later in the autumn, however, the differences  disappeared. Before recommending the use of early-season SD-treated  seedlings for summer planting, the method has to be tested in practical  field conditions.
                        
                
                                            - 
                            Luoranen,
                            The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland
                                                        E-mail:
                                                            jaana.luoranen@metla.fi
                                                                                          
- 
                            Konttinen,
                            The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland
                                                        E-mail:
                                                            kk@nn.fi
                                                                                
- 
                            Rikala,
                            The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland
                                                        E-mail:
                                                            rr@nn.fi
                                                                                
 
         
     
 
                        
                
                
                                            Category :
                    
                    Review article
                                    
                            
                    
        
            
            article id 313,
                            category
                        Review article
                    
        
        
                            Heikki Hänninen,
                            Koen Kramer.
                    
                    
                (2007).
            
                            
                                    A framework for modelling the annual cycle of trees in boreal and temperate regions.
                            
                            
                Silva Fennica
                                                            vol.
                                        41
                                                                            no.
                                        1
                                article id 313.
            
                            
                https://doi.org/10.14214/sf.313
            
             
        
                                    
                                    
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                            Models of the annual development cycle of trees in boreal and temperate  regions were reviewed and classified on the basis of their  ecophysiological assumptions. In our classification we discern two main  categories of tree development: 1) fixed sequence development, which  refers to irreversible ontogenetic development leading to visible  phenological events such as bud burst or flowering, and 2) fluctuating  development, which refers to reversible physiological phenomena such as  the dynamics of frost hardiness during winter. As many of the  physiological phenomena are partially reversible, we also describe  integrated models, which include aspects of both fixed-sequence and  fluctuating development. In our classification we further discern simple  E-models, where the environmental response stays constant, and more  comprehensive ES-models, where the environmental response changes  according to the state of development. On the basis of this model  classification, we have developed an operational modelling framework, in  which we define an explicit state variable and a corresponding rate  variable for each attribute of the annual cycle considered. We introduce  a unifying notation, which we also use when presenting a selection of  previously published models. To illustrate the various developmental  phenomena and their modelling, we have carried out model simulations.  Finally, we discuss the ecophysiological interpretation of the model  variables, methodological aspects of the empirical development and  testing of the models, the introduction of new aspects to the modelling,  other closely related models, and applications of the models.
                        
                
                                            - 
                            Hänninen,
                            Plant Ecophysiology and Climate Change Group (PECC), Department of Biological and Environmental Sciences, Box 65, FI-00014 University of Helsinki, Finland
                                                        E-mail:
                                                            heikki.hanninen@helsinki.fi
                                                                                          
- 
                            Kramer,
                            Alterra, P.O. Box 47, 6700 AA Wageningen, The Netherlands
                                                        E-mail:
                                                            kk@nn.nl