Current issue: 56(2)
The purpose of this paper is to review tests made on the basis of Finnish material with regard to the efficiency of the 10-point cluster in sampling a stand in forest inventory. Currently, this system is applied in field work in the national forest surveys in the United States of America. The paper reports on tests, made on the basis of Finnish material, for comparison of the 10-point cluster of variable plots with 13 other designs in sampling a stand in forest survey. The research material consists of 12 stands, with Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) as the main species.
The main results are concerned with the ability of different designs to provide gross volume estimates. As a measure of efficiency, three alternative series of variances were used, adjusted by three alternatives of time. The results are applicable, for instance, in double-sampling with photo and field classifications. In the comparisons, no attention was paid to the possibility of systematic errors in various designs.
For inventory volume, the 10-point cluster proved to be about 10 per cent less efficient than the best design of each alternative. The use of a single circular plot of 1,000 m2 can be recommended under the conditions of this test; furthermore, one or two 500 m2 plots were more efficient than any combination of variable plots.
The reason for the use of the 10-point cluster in forest surveying has been the ability of the design to provide simultaneous information on area condition classes. Among the designs tested, the 10-point cluster seems to be the only one capable of application in the estimation of condition classes.
Most of the information obtained by means of the 10-point cluster can be gained through ocular estimation, and from the sample trees to be measured in any design, but a cluster of several points appears to offer good means of estimation, for instance, of the presence of clumps and gasps in a stand.
There has not been complete agreement as to what is meant by ectendotrophic mycorrhizae, and there is a wide variety of opinion among authors on mycorrhizal terminology. In this paper ectendotrophic mycorrhizae are defined to be short roots with Hartig net and intracellular hyphae in the cortex. A mantle and digestion of intracellular hyphae may be found but are not necessary. In the study of Mikola (1965) ectendotrophic mycorrhiza was found to be common in Scots pine (Pinus sylvestris L.) seedlings in Finnish nurseries. The mycorrhizae had always similar structure and the mycelium isolated from the seedlings (E-strains) was similar. The aim of this study was to find out what kind of ectendotrophic mycorrhizae exist in forests and nurseries outside Finland, what kind of mycorrhizae do the E-strains isolated from Scots pine form with other tree species, and are these associations symbiotic.
Only one type of ectendotrophic mycorrhiza was found on the 600 short roots collected from the continents of Europa and America. The type was similar to the one described by Mikola: the mycelium is coarse and forms a strong Hartig net, and intracellular infection is heavy. Evidence is convincing that this structure was formed by the same fungus species. The species is unidentified. Mycorrhizae synthesized by E-strain with six spruce species, fir, hemloch and Douglas fir were all ectotrophic.
The E-type ectendotrophic mycorrhizae proved to be a balanced symbiosis. The seedlings of 13 tree species inoculated with the E-strain grew in the experiment better than the controls. The observation that ectendotrophic mycorrhizae dominates in the nurseries but is seldom found in forests, and then only in seedlings growing in the forest, was confirmed in the study. In synthesis experiments E-strain formed either ecto- or ectendotrophic mycorrhiza depending on the tree species.
The differences between different types of mycorrhiza; endomycorrhiza, ectomycorrhiza and ectendomycorrhiza, and the use of the terms have been variable in the earlier research. Studied of mycorrhiza in Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) seedlings may suggest that the conditions affect which kind of mycorrhiza develops in the seedlings. This study is aimed mainly at finding out whether the difference of ectotrophic and ectendotrophic mycorrhizae depends on fungal symbionts or envirionmental conditions. Furthermore, the occurrence of ectendotrophic mycorrhiza in Finland under various conditions was studied, and experiments on the physiology and ecology of the mycorrhiza and the fungal partner were conducted.
The ectendotrophic mycorrhiza as described in this paper has proved to be very common on Scots pine in Finnish nurseries, but it was not found in Norway spruce seedlings. The results did not support the hypothesis presented in some earlier studies that ectendotrophic mycorrhiza is more parasitic than the other mycorrhizal fungi. The nursery survey showed that no correlation existed between the size and vigour of the seedlings and the presence of ectendotrophic mycorrhiza. Furthermore, greenhouse-grown seedlings with and without the fungus grew equally well. The type of mycorrhiza was, however, almost exclusively confined to young (1–3-years-old) seedlings and to nursery soils. The experiments indicates also that ectendomycorrhizal fungus has a very wide ecological amplitude in regard to light intensity, soil fertility, acidity, and humus content. It has, however, a weak competitive ability in natural forest soils against the indigenous fungal population. When the seedlings were transplanted from the nursery to forest soil, their mycorrhizal population was largely changed.
The aim of the investigation was to obtain by snow and soil frost observations sufficient material for determination of regional springtime snow and soil frost values, because the water equivalent of snow and the frost depth affect runoff. The present paper elaborates a method by which the observations along a survey line can be corrected to be valid for a basin. Along the line 50 measurement points were arranged at specific intervals. Snow depth was recorded at each point, and snow density and frost depth at every fifth point. The terrain was studied along the line and the terrain of the survey points were classified in eight classes depending on the vegetation. The classes ranged from cultivated lands and open bogs to wooded areas according to volume of the growing stock and tree species composition.
The mean snow depth was 51.9 cm and mean snow density 0.235 g/m2. Water equivalent of snow in class 4 terrain (forest with small growing stock) was 30% higher than in class 8 (forest with high growing stock). An ample stand increases evaporation in wintertime. The difference can be partly caused by the different accumulation of snow in the different types of stands.
Soil type was not found to have any distinct influence on the frost depth in the present material. On cultivated lands the soil frost clearly penetrates to greater depth than in the forest. The growing stock of wooded areas influences the snow depth
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