Silva Fennica vol. 48 | 2014

We tested the hypotheses (H) that on newly clearcut-harvested sites, (H1) abundance and species diversity of the forest-floor small mammal community, and (H2) abundance, reproduction, and recruitment of red-backed voles (Myodes gapperi Vigors), would increase with higher levels of structural retention via green-tree retention (GTR) and woody debris (dispersed and constructed into windrows). Study areas were located in three forest ecological zones in southern British Columbia, Canada. For H1, mean total abundance did generally increase with the gradient of retained habitat structure. Mean species richness and diversity were similar among treatment sites but did show an increasing gradient with structural compexity. For H2, mean abundance, reproduction, and recruitment of M. gapperi were higher in GTR and windrow sites than those without retained structures. There was a positive relationship between mean abundance of M. gapperi and total volume of woody debris across treatments. This study is the first investigation of the responses of forest-floor small mammals to an increasing gradient of retained habitat structure via GTR and woody debris on clearcuts. Our assessment of a combination of these two interventions suggested a potentially strong additive effect that could be cautiously extrapolated across three forest ecological zones. With the advent of low levels of GTR on clearcuts, woody debris structures should help provide some habitat to conserve forest mammals on harvest openings.

• Sullivan, Department of Forest and Conservation Sciences, Faculty of Forestry, University of BC, 2424 Main Mall, Vancouver, BC, Canada V6T 1Z4 E-mail: tom.sullivan@ubc.ca
• Sullivan, Applied Mammal Research Institute, 11010 Mitchell Avenue, Summerland, BC, Canada V0H 1Z8 E-mail: dru.sullivan@appliedmammal.com

Multimodal primary forest fuel (PFF) transport using the railroad for main haulage has been quite uncommon to present, although it could provide considerable advantages in terms of economical, ecological and social parameters. Accordingly, the objective of this paper is to assess barriers and drivers for the modal shift from truck to train. As methodological tool, we are using the concept of Quality Function Deployment (QFD) with the House of Quality (HoQ) – an approach that has not been used in forest management so far. As the most important barriers for the modal shift from truck to train in PFF transport in Austria, the following were identified: (i) bioenergy plant managers have a negative opinion and negative experience regarding the railroad in terms of high prices, a lot of bureaucracy, etc.; (ii) absence of rail sidings or relatively short rail sidings not suitable for block trains; and (iii) unwillingness to invest in new supply or unloading systems. On the contrary, the most important drivers for a modal shift are: (i) multimodal PFF supply chains using trains can provide high volumes; (ii) increasing catchment areas for larger CHP plants result in increasing transport distances; and (iii) rail transport has less negative environmental and social impact than road transport.

• Wolfsmayr, University of Natural Resources and Life Sciences, Vienna, Gregor Mendel Straße 33, A-1180 Wien, Austria E-mail: ulrich.wolfsmayr@boku.ac.at
• Rauch, University of Natural Resources and Life Sciences, Vienna, Gregor Mendel Straße 33, A-1180 Wien, Austria E-mail: peter.rauch@boku.ac.at

Recent studies have shown the establishment of Norway spruce (Picea abies (L.) Karst.) to be successful in small canopy gaps cut in drained spruce mire stands in northern Finland. The aim of this study was to quantify seedling height growth in gaps and compare it to that observed in other canopy cuttings and plantations. We sampled spruce crop seedlings (maximum density ca. 3000 ha^–1) in the spring of 2013 in a field experiment in which canopy gaps of 10, 15 and 20 m in diameter had been cut in winter 2004. The total seedling height in 2013 and the length of annual shoots over the past five years (2012–2008) were recorded in the survey. Seedling height varied from 20 cm to 2.7 m, with an average of 65 cm. The average annual height growth was 7.1 cm. A mixed linear model analysis was carried out to investigate seedling height growth variation. Seedling height was linearly and positively related to growth. Height growth started to increase in the fifth growing season after cutting. Seedling height growth in the 20 m gap was slightly better than in the smaller ones. In the 15 m gap, both the centrally located seedlings and those located at the northern edge grew best. In the 20 m gap, southerly located seedlings grew more slowly than seedlings in all other locations. The average seedling height growth in this study was about 60% of that in peatland plantations, but comparable to that in mineral soil gaps, and 2–3 times higher than in uneven-age cut stands.

• Hökkä, Finnish Forest Research Institute, Rovaniemi Unit, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: hannu.hokka@metla.fi
• Mäkelä, Lapland University of Applied Sciences, Jokiväylä 11 C, FI-96300 Rovaniemi, Finland E-mail: hanna.makela@edu.ramk.fi

Biomass models for the biomass of above-ground tree components of Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies [L.] Karst.) and birch (Betula pendula Roth and Betula pubescens Ehrh.) in young dense Scots pine dominated forest stands in northern Sweden were constructed. Destructive above-ground biomass sampling was conducted in naturally generated young, dense, Scots pine dominated mixed stands. Three sampling campaigns were undertaken, the first in 1997 and 1998. The second was six years later (2003), and the last 13 years after the first (2010). In total, 280 trees (126 Scots pine, 68 Norway spruce and 86 birches) were sampled from six different stands in northern Sweden. The sampled trees’ diameter at breast height (dbh) was in the range 1–22 cm (Scots pine), 1–21 cm (Norway spruce) and 1–11 cm (birch). Biomass predictions were tested using our models and the widely used biomass models originally constructed for managed stands. The results showed that the biomass allocation to tree components is different in unmanaged and managed young stands; higher foliage biomass and lower stem and branch biomass were detected in the unmanaged stands. The overall conclusion is that the biomass models for managed stands did not produce satisfactory biomass estimates in unthinned, dense, young stands.

• Repola, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: jaakko.repola@metla.fi
• Ahnlund Ulvcrona, SLU, Forest Biomaterials and Technology, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: kristina.ulvcrona@slu.se

The perception of the Finns about forests and forestry has been tracked over a period of more than 15 years. The results of this survey constitute the longest sequence of data of this type at the national level anywhere in the world. The people’s perception of reality represents a factor that influences decisions about policy. For this reason, it deserves monitoring and analysis. Forests in Finland are highly meaningful to the people, who are generally well informed and link their opinions to the facts that they are able to observe. The variability of the responses over the years of the survey is not significant. Silviculture and forest management are perceived as good by most Finns. Finns are aware that more forest grows than is harvested, and they also know that some raw material is still imported. However, they demand that more forest be protected. Finns are aware that their forest industry is not performing well at the international level. They also demand an increased wood supply for building construction. Forest harvesting is viewed as a source of employment and welfare.

• Fabra Crespo, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: mfabracrespo@yahoo.com
• Saastamoinen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: olli.saastamoinen@uef.fi
• Matero, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: materojukka@gmail.com
• Mäntyranta, Finnish Forest Association, Salomonkatu 17 A, FI-00100 Helsinki, Finland E-mail: hannes.mantyranta@smy.fi

Global efforts to decrease dependence on fossil fuels have increased interest in bioenergy production. One source of bioenergy is fast growing deciduous tree species, such as hybrid aspen (Populus × wettsteinii Hämet-Ahti). The majority of research on hybrid aspen which assesses biodiversity implications, has however primarily focused on agricultural lands as the reference condition. This has resulted in a substantial gap in our knowledge regarding the biodiversity implications of replacing production forest types with hybrid aspen, a form of reforestation taking place in northern Europe. In this study we address this knowledge gap by comparing the avian biodiversity of young hybrid aspen and spruce (Picea abies L.) plantations of similar age, the latter being the most prevalent forestry alternative in in southern Sweden. We found that hybrid aspen stands had higher bird species richness and abundance as well as a distinct community composition compared to the spruce stands. We suggest that the most likely driver was the greater structural and tree species complexity in the aspen stands, provided for by the fenced exclusion of ungulates from the regenerating hybrid aspen stands. Our results indicate that at least during early stages of regeneration, and in comparison to the dominating production forest type in the region, hybrid aspen stands can support relatively high levels of bird diversity, and a bird species composition more closely associated with broadleaf habitat types requiring restoration in this region.

• Lindbladh, Southern Swedish Forest Research Centre, SLU – Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: matts.lindbladh@slu.se
• Hedwall, Southern Swedish Forest Research Centre, SLU – Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: per-ola.hedwall@slu.se
• Wallin, Southern Swedish Forest Research Centre, SLU – Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: ida.wallin@slu.se
• Felton, Southern Swedish Forest Research Centre, SLU – Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: annika.felton@slu.se
• Böhlenius, Southern Swedish Forest Research Centre, SLU – Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: henrik.bohlenius@slu.se
• Felton, Southern Swedish Forest Research Centre, SLU – Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: adam.felton@slu.se

Height growth of young hybrid aspen (Populus tremula L. × P. tremuloides Michx.) was studied in relation to weather conditions. Height of clones with different leaf flushing phenology (early, intermediate and late) was monitored during the growing periods of 2010 and 2011 in a plantation established on former agricultural land. Mean daily height increment (HI) was calculated. Multiple linear regression was used to determine which weather factors (variables) had significant effect on HI. Mean seasonal height growth (mean seasonal HI) between clones (groups) was compared by ANOVA. In both years, HI was significantly higher for clones with early and intermediate leaf flushing compared to clones with late leaf flushing. The effect of weather factors also differed between clones according to their leaf flushing phenology; it was the weakest for HI of clones with early leaf flushing compared to clones with intermediate and late leaf flushing. Mean temperature was the main factor, which positively affected HI of all clones, suggesting that warmer climate might be beneficial for height growth of young hybrid aspen in Latvia. Nevertheless, significant negative relationship between HI and potential evapotranspiration (PET) was observed for clones with delayed leaf flushing, suggesting negative effect of increasing variability of precipitation on growth. Thus, the differences in height growth intensity might be related to growth sensitivity to weather conditions. On the other hand, such differences in height growth between clones might be caused by competition (i.e. with herbs), as trees with early leaf flushing might conquer more resources and become more robust against the environmental fluctuation.

• Jansons, LSFRI „SILAVA”, Rigas Str. 111, Salaspils, Latvia, LV2169 E-mail: aris.jansons@silava.lv
• Zeps, LSFRI „SILAVA”, Rigas Str. 111, Salaspils, Latvia, LV2169 E-mail: martins.zeps@silava.lv
• Rieksts-Riekstiņš, LSFRI „SILAVA”, Rigas Str. 111, Salaspils, Latvia, LV2169 E-mail: Juris.Riekstins@silava.lv
• Matisons, LSFRI „SILAVA”, Rigas Str. 111, Salaspils, Latvia, LV2169 E-mail: robism@inbox.lv
• Krišāns, LSFRI „SILAVA”, Rigas Str. 111, Salaspils, Latvia, LV2169 E-mail: oskars.krisans@silava.lv

Thinning alters litter quality and microclimate under forests. Both of these two changes after thinning induce alterations of litter decomposition rates and nutrient cycling. However, a possible interaction between these two changes remains unclear. We placed two types of litter (LN, low N concentration litter; HN, high N concentration litter) in a Chinese pine (Pinus tabulaeformis Carrière) plantation under four thinning treatments to test the impacts of litter quality, thinning or their combination on decomposition rate and N cycling. In our study, N was accumulated to approach an underlying critical acid-unhydrolyzable residue to nitrogen ratio (approximately 57–69) in litter. Moreover, an interaction between litter quality and thinning on decomposition rates, N accumulation and net release did exist. On one hand, one year decomposition rate of LN was elevated after thinning while that of HN remained the same or even lower (under light thinning); N accumulation of LN declined with light thinning and was restored with the increase of thinning intensity whereas that of HN did not decline with thinning and increased under heavy thinning; Net N release from LN was only found in light and heavy thinning while that from HN was found in all treatments, moreover net N release from LN and HN were both elevated under heavy thinning. On the other hand, HN decomposed faster, accumulated less and released more N than LN did under all treatments. Generally, high N concentration in litter and high-intensity thinning can lead to rapid litter decomposition and N cycling in coniferous plantations.

• Chen, Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing, 100083, China E-mail: chenxiao_0123@126.com
• Page-Dumroese, U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, 1221 South Main Street, Moscow, ID 83843, USA E-mail: ddumroese@fs.fed.us
• Lv, College of Plant Science and Technology, Tarim University, Alar Xinjiang, 843300, China E-mail: lvrh514723@126.com
• Wang, Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing, 100083, China E-mail: fuyuerdejia@126.com
• Li, Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing, 100083, China E-mail: glli226@163.com
• Liu, Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing, 100083, China E-mail: lyong@bjfu.edu.cn

The information about location and size of past human settlements can give new insights into the analysis of landscape structures. Vodlozero National Park (NP) is one the largest strictly protected areas in Northwestern Europe. We mapped the location of historic villages, which were abandoned in 1958, and studied the effect of past human activity in the forest landscape in two different scales using forest survey data. We assessed the possible change in age, volume and tree species composition from the edge of open fields up to the distance of 5 km from villages. We made a larger landscape analysis using a grid of forest stands covering the whole northern part of the NP. The past human activity was clearly visible in the present forest landscape. Distance from villages affected age, volume and tree species composition of the forest stands. This effect was the strongest within the first two kilometers from the villages. At the level of whole northern NP, the proportion of spruce markedly increased after approximately 15 km from the nearest old village. The changes in the forests surrounding the villages were most likely the result of the intensive use of wood for different commodities needed in households and farming, in addition to short rotation slash and burn agriculture. If the occurrence of forest fires was more frequent closer to villages than in more remote areas, it can well explain the observed pattern in the abundance of spruce in the larger landscape that is less tolerant to fire than pine.

• Tikkanen, Department of Biology, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland (Current: School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland) & Interdisciplinary Research and Educational Center of Cross-border Communication CARELICA, Institute of History, Political and Social Sciences, Petrozavodsk State University, 33 Lenin Prospectus, 185910 Petrozavodsk, Republic of Karelia, Russia E-mail: Olli-Pekka.Tikkanen@uef.fi
• Chernyakova, Interdisciplinary Research and Educational Center of Cross-border Communication CARELICA, Institute of History, Political and Social Sciences, Petrozavodsk State University, 33 Lenin Prospectus, 185910 Petrozavodsk, Republic of Karelia, Russia E-mail: irina.chernyakova@onego.ru

There is increasing interest in using introduced species in Swedish forestry in response to climate change, but it is important to assess their resistance to native pests. Thus, we compared the extent of pine weevil feeding on two dominant native conifers, Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.), the non-host deciduous broadleaf hybrid aspen (Populus × wettsteinii Hämet-Ahti) and four introduced conifers: Douglas fir (Pseudotsuga menziesii (Mirb.) Franco), hybrid larch (Larix × marschlinsii Coaz), Sitka spruce (Picea sitchensis (Bong.) Carriére) and lodgepole pine (Pinus contorta Douglas ex Loudon). The extent of feeding damage on seedlings and its effect on their vitality were examined in a field study in south-central Sweden and a laboratory experiment, which gave largely consistent results. Generally, the species most heavily attacked by the pine weevil, in both experiments, were Douglas fir and Sitka spruce. In the field experiment pine weevils killed or severely damaged significantly higher proportions of Douglas fir and Sitka spruce seedlings (60%) than any other species except Norway spruce (49%). Among conifer seedlings the proportions of killed or severely damaged seedlings were lowest for Scots pine and hybrid larch (27%) and Lodgepole pine (36%). The results indicate that most conifer species planted on young clear-cuttings in Sweden need some kind of pine weevil protection, and the possibility that introducing new tree species might increase damage caused by pests must be considered. For instance, widespread use of hybrid aspen could reduce damage by pine weevils, but increase damage by other, untested pests or pathogens.

• Wallertz, Unit for Field-based Forest Research, Swedish University of Agricultural Sciences (SLU), Asa Research Station, SE-36030 Lammhult, Sweden E-mail: kristina.wallertz@slu.se
• Nordenhem, Department of Ecology, Swedish University of Agricultural Sciences (SLU), P.O. Box 7044, SE-75007 Uppsala, Sweden E-mail: henrik.nordenhem@slu.se
• Nordlander, Department of Ecology, Swedish University of Agricultural Sciences (SLU), P.O. Box 7044, SE-75007 Uppsala, Sweden E-mail: goran.nordlander@slu.se

In afforestation, silvicultural processes of raising and planting seedlings under certain conditions can yield contrasting outcomes in tree stock performance. Moderate nursery conditions may select against stress tolerance whereas planting seedlings in stressful environments at afforestation sites may select for higher stress tolerance compared to natural populations. We compared germination performance between natural and afforested populations of Pinus brutia Ten. subsp. brutia and Cupressus sempervirens L. var. horizontalis (Mill.) under differing stress treatments. Seeds were collected from both natural stands and from afforested populations outside the natural distribution range, in Syria. Cold, intermediate and hot temperature regimes were simulated (8/4 °C, 20/10 °C and 32/20 °C) along with cold stratification, drought stress (–0.2 and –0.4 MPa), salt stress (50 and 100 mMol l^–1), and deionized water (control) conditions. In addition, we tested the effects of seed weight and climatic conditions on seed germination. In general, intermediate temperatures were optimal for both population types. Afforested P. brutia populations outperformed natural ones under drought stress levels at hot and/or intermediate temperatures. Conversely, in C. sempervirens, cold stratification at all temperatures and higher salt stress at intermediate temperatures significantly decreased germination in afforested populations. Seed weight did not significantly affect germination percentages, which were however significantly negatively related to annual precipitation in P. brutia, and to annual temperature in C. sempervirens. We infer that silvicultural processes led to divergent outcomes in our species: local adaptation to drought stress and hot temperatures in afforested P. brutia populations and lower salt-stress tolerance in C. sempervirens.

• Al-Hawija, Martin-Luther-University Halle-Wittenberg, Institute of Biology/Geobotany and Botanical Garden, Am Kirchtor 1, D-06108 Halle/Saale, Germany E-mail: batoulh@gmail.com
• Wagner, Department of Botany and Zoology, Masaryk University, Kotlářská 2, CZ-611 37 Brno, Czech Republic E-mail: wagner@sci.muni.cz
• Partzsch, Martin-Luther-University Halle-Wittenberg, Institute of Biology/Geobotany and Botanical Garden, Am Kirchtor 1, D-06108 Halle/Saale, Germany E-mail: monika.partzsch@botanik.uni-halle.de
• Hensen, Martin-Luther-University Halle-Wittenberg, Institute of Biology/Geobotany and Botanical Garden, Am Kirchtor 1, D-06108 Halle/Saale, Germany & German Centre for Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig, Deutscher Platz 5e, 04103 Leipzig, Germany E-mail: isabell.hensen@botanik.uni-halle.de

The aim of the study was to characterize repetitive cycles of harvester operation. The study was conducted in thinning, mature and post-disaster pine stands. The sequences of the activities characteristic of harvester operation were described as time series. In order to detect the cyclic variable structure of the analysed time series, the methodology of the single spectrum Fourier analysis was applied. In standard stands, post-disaster late-thinning stands and mature stands, the existence of stable operational cycles with the length of three activities was discovered while in post-disaster mature stands additional five-activity operational phases were noted. Described in this way, the lengths of the operational cycles of harvesters working in post-disaster areas were higher by about 55% and 30% respectively, as compared to standard thinning and mature stands.

• Szewczyk, University of Agriculture in Krakow, Faculty of Forestry, Institute of Forest Utilization and Forest Technology, Department of Forest and Wood Utilization, Al. 29-Listopada 46, 31-425 Krakow, Poland E-mail: rlszewcz@cyf-kr.edu.pl
• Sowa, University of Agriculture in Krakow, Faculty of Forestry, Institute of Forest Utilization and Forest Technology, Department of Forest and Wood Utilization, Al. 29-Listopada 46, 31-425 Krakow, Poland E-mail: rlsowa@cyf-kr.edu.pl
• Grzebieniowski, University of Agriculture in Krakow, Faculty of Forestry, Institute of Forest Utilization and Forest Technology, Department of Forest and Wood Utilization, Al. 29-Listopada 46, 31-425 Krakow, Poland E-mail: wrzoswj@interia.pl
• Kormanek, University of Agriculture in Krakow, Faculty of Forestry, Institute of Forest Utilization and Forest Technology, Department of Forest Work Mechanisation, Al. 29-Listopada 46, 31-425 Krakow, Poland E-mail: rlkorma@cyf-kr.edu.pl
• Kulak, University of Agriculture in Krakow, Faculty of Forestry, Institute of Forest Utilization and Forest Technology, Department of Forest and Wood Utilization, Al. 29-Listopada 46, 31-425 Krakow, Poland E-mail: rlkulak@cyf-kr.edu.pl
• Stańczykiewicz, University of Agriculture in Krakow, Faculty of Forestry, Institute of Forest Utilization and Forest Technology, Department of Forest and Wood Utilization, Al. 29-Listopada 46, 31-425 Krakow, Poland E-mail: rlstancz@cyf-kr.edu.pl

The purpose of this study was to examine how the age of a stand of post-agricultural Scots pine forests affects the species composition, biomass and the carbon stock of the forest understorey. The community structure and species composition were studied in 75 plots (100 m² in size), the amount of biomass, organic carbon and total nitrogen were analysed in 75 subplots (1/16 m² in size). The plots were located in 21 plantations with the stand age of 41–60, 61–80 and over 80-years. Results show that the understorey species diversity increased with the increasing age of Scots pine stands, and the structure and species composition of secondary forests (although managed for timber production) became similar to the fresh pine forest of the European temperate region (Leucobryo-Pinetum community). Despite the increasing species diversity, however, only six understorey vascular and moss species played an important role in the biomass accumulation and C sequestration. Due to the differences in the dominant species composition, the total amount of understorey biomass significantly differed among the forest stands. The mean moss biomass ranged from 3046 kg ha^–1 in 41–60-year-old stands, trough 2686 kg ha^–1 in 61–80-year-old stands to 2273 kg ha^–1 in over 80-year-old stands, and the mean understorey vascular plant biomass amounted to 2 kg ha^–1, 1924 kg ha^–1 and 3508 kg ha^–1, respectively. The concentration of organic C varied considerably between species; it was the highest in Vaccinium myrtillus (50.6%) and in Dicranum polysetum (49.5%). The total mass of C was nearly 800 kg ha^–1 in the youngest forests, in the subsequent age series it was two times higher and 3.5 times higher in the oldest ones. Differences in the species composition and in the C/N ratio in different species (generally higher for vascular plants and lower for mosses) were expressed in an increase in the understorey C/N ratio, which was 39.5, 46.6 and 48.6, respectively.

• Woziwoda, Department of Geobotany and Plant Ecology, Faculty of Biology and Environmental Protection, University of Lodz, Banacha 12/16, 90-237 Lodz, Poland E-mail: woziwoda@biol.uni.lodz.pl
• Parzych, Environmental Chemistry Research Unit, Institute of Biology and Environmental Protection, Pomeranian University in Słupsk, Arciszewskiego 22b, 76-200 Słupsk, Poland E-mail: parzycha1@op.pl
• Kopeć, Department of Geobotany and Plant Ecology, Faculty of Biology and Environmental Protection, University of Lodz, Banacha 12/16, 90-237 Lodz, Poland E-mail: domin@biol.uni.lodz.pl

We studied the physicochemical properties of stemwood in saplings of silver birch (Betula pendula Roth) and hybrid aspen (Populus tremula L. × P. tremuloides Michx.), grown for four years under artificially elevated relative air humidity (on average by 7%) in field conditions, using the Free Air Humidity Manipulation (FAHM) research facility in Estonia. Altogether 91 sample trees from three experimental plots with manipulated air humidity and from three control plots were cut in the dormant season and sampled for the analysis of cellulose, hemicellulose, acid detergent lignin, macronutrients (N, P, K), ash content, density, and calorific value of wood. The analysed trees grew significantly more slowly under elevated humidity conditions, with a more pronounced effect on aspens. Significantly higher concentrations of N and P were observed in the stemwood of both aspens and birches grown under elevated humidity. This could be the result of a change in the content of living parenchyma cells and/or enhanced retranslocation of nutrients into wood parenchyma. Additionally, humidification resulted in a significantly higher concentration of cellulose and a lower concentration of hemicellulose in aspen stemwood, and in significantly lower concentrations of cellulose and K in birch stemwood. Elevated humidity did not affect lignin concentration, ash content, basic density and calorific value of stemwood. Results from the FAHM experiment suggest that the increasing air humidity accompanying global warming at northern latitudes will affect the growth and functioning of deciduous trees and forests, with obvious consequences also for forest management and industry.

• Tullus, Department of Botany, Institute of Ecology and Earth Sciences, Faculty of Science and Technology, University of Tartu, Lai 40, Tartu 51005, Estonia E-mail: arvo.tullus@ut.ee
• Sellin, Department of Botany, Institute of Ecology and Earth Sciences, Faculty of Science and Technology, University of Tartu, Lai 40, Tartu 51005, Estonia E-mail: arne.sellin@ut.ee
• Kupper, Department of Botany, Institute of Ecology and Earth Sciences, Faculty of Science and Technology, University of Tartu, Lai 40, Tartu 51005, Estonia E-mail: priit.kupper@ut.ee
• Lutter, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Kreutzwaldi 5, Tartu 51014, Estonia E-mail: reimo.lutter@emu.ee
• Pärn, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Kreutzwaldi 5, Tartu 51014, Estonia E-mail: linnar.parn@emu.ee
• Jasinska, Department of Botany, Institute of Ecology and Earth Sciences, Faculty of Science and Technology, University of Tartu, Lai 40, Tartu 51005, Estonia & Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland E-mail: jasiak9@wp.pl
• Alber, Department of Botany, Institute of Ecology and Earth Sciences, Faculty of Science and Technology, University of Tartu, Lai 40, Tartu 51005, Estonia E-mail: meeli.alber@ut.ee
• Kukk, Department of Botany, Institute of Ecology and Earth Sciences, Faculty of Science and Technology, University of Tartu, Lai 40, Tartu 51005, Estonia E-mail: maarja.kukk@ut.ee
• Tullus, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Kreutzwaldi 5, Tartu 51014, Estonia E-mail: tea.tullus@emu.ee
• Tullus, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Kreutzwaldi 5, Tartu 51014, Estonia E-mail: hardi.tullus@emu.ee
• Lõhmus, Department of Botany, Institute of Ecology and Earth Sciences, Faculty of Science and Technology, University of Tartu, Lai 40, Tartu 51005, Estonia E-mail: krista.lohmus@ut.ee
• Sõber, Department of Botany, Institute of Ecology and Earth Sciences, Faculty of Science and Technology, University of Tartu, Lai 40, Tartu 51005, Estonia E-mail: anu.sober@ut.ee

The productivity of mechanized planting could be increased by minimizing the time spent manually reloading seedling cassettes. This study compared the work-time distribution, productivity and quality of the prototype Risutec APC fitted with an automatic feeding system and the commonly-used and manually-loaded Bracke P11.a. An approach of comparative time study was employed that compared performance of two operators using both machines in four sites where slash and stumps had been removed. Operating costs were estimated and compared for these two machines and an idealized machine with automatic feeding system (referred as AUT). AUT was assumed to be similar to the Bracke planting machine with the only difference being in automatic feeding. Productivity of the Risutec APC (196 seedlings per productive work hour [pl PWh₀^–1]) was lower than that of Bracke (244 pl PWh₀^–1), making the unit cost 35.7% higher. A large portion (17.6%) of the productive work time of Risutec APC was interrupted by malfunctions, so it cannot be considered robust and reliable yet. Quality of the planting work was reasonable for both machines. The results suggest that an idealized AUT could increase planting capacity (hectares per year [ha yr^–1]) by 15.4% and lower the unit cost (Euro per seedling [€ pl^–1]) by 4.7% compared to today’s machines. The importance of an automated feeding system increases with planting efficiency because relatively more time is spent reloading seedlings. Proper automatic feeding system could offer a cost-effective solution and could enhance productivity, but the Risutec APC has yet to meet the technical and economic standards required to be competitive.

• Laine, Finnish Forest Research Institute (METLA), Suonenjoki Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: tiina.laine@metla.fi
• Saarinen, Finnish Forest Research Institute (METLA), Suonenjoki Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: veli-matti.saarinen@metla.fi

Reindeer husbandry and forestry are practiced in the same areas in northern Fennoscandia. Reindeer pastures have largely deteriorated. We aimed to quantify the separate and combined effects of reindeer grazing and forestry on the amount of ground lichens. To do this, we mapped and inventoried all larger enclosures (49) in Finnish Lapland where forest management practices were similar in both sides of the fence. The average time since fencing was 43 years. We recorded the cover and estimated dry biomass of ground lichens, as well as parameters describing forest stand characteristics. The effect of reindeer grazing on both the cover and estimated dry biomass of lichens was clear: in the ungrazed (fenced) sites, the lichen cover (35.8%) was on average 5.3-fold and the dry biomass (1929 kg ha^–1) 14.8-fold compared with the corresponding estimates in the grazed sites (6.8% and 130 kg ha^–1). The effect of forestry on lichens was smaller. In the grazed stands the cover and biomass of lichens were higher in the mature stands compared to the younger stand development classes, whereas in the ungrazed stands there were no significant differences between the development classes. Both reindeer grazing and forestry affect the cover and biomass of ground lichens. The influence of reindeer grazing is, however, much heavier than that of forestry. The decrease of not only the biomass, but also the lichen cover, is alarming. The decrease of lichen cover may hinder the recovery of reindeer pastures, which in the long run endangers the sustainability of reindeer husbandry.

• Akujärvi, Finnish Environment Institute, Helsinki, Finland E-mail: anu.akujarvi@ymparisto.fi
• Hallikainen, Finnish Forest Research Institute, Northern Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: ville.hallikainen@metla.fi
• Hyppönen, Finnish Forest Research Institute, Northern Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: mikko.hypponen@metla.fi
• Mattila, Finnish Forest Research Institute, Northern Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: eero.mattila@metla.fi
• Mikkola, Finnish Forest Research Institute, Northern Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: kari.mikkola@metla.fi
• Rautio, Finnish Forest Research Institute, Northern Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: pasi.rautio@metla.fi

The spatial occurrence of heavy crown snow loads in Finland between 1961 and 2010 is studied by using for the first time a model that classifies the snow load into four different types: rime, dry snow, wet snow and frozen snow. In producing this climatology, we used meteorological observations made at 29 locations across Finland. The model performance is evaluated against classified daily images of canopy snow cover and with the help of two short case studies. The results are further compared to those achieved with a simpler method used in previous studies. The heaviest crown snow loads are found to occur typically in eastern Finland. The new method reveals that this holds not only for the total snow loads but also for the different snow load types, although there are certain differences in their geographical occurrence. The greatest benefit achieved with the new method is the inclusion of rime accretion. The forests most prone to heavy riming are those located on tree-covered hills in northern Finland, but as the terrain elevation affects riming efficiency greatly, these small-scale variations in the snow load amounts could not be described in this study in great detail. Moreover, the results are more inaccurate in northern Finland where variations in the terrain elevation are greater than elsewhere. Otherwise, the largest uncertainties in this study are related to wind speed measurements and possibly partly because of that, we were not able to detect any significant trends in the crown snow-load amounts over the study period.

• Lehtonen, Finnish Meteorological Institute, P.O. Box 503, FI-00101 Helsinki, Finland E-mail: ilari.lehtonen@fmi.fi
• Hoppula, Finnish Meteorological Institute, P.O. Box 503, FI-00101 Helsinki, Finland E-mail: petri.hoppula@fmi.fi
• Pirinen, Finnish Meteorological Institute, P.O. Box 503, FI-00101 Helsinki, Finland E-mail: pentti.pirinen@fmi.fi
• Gregow, Finnish Meteorological Institute, P.O. Box 503, FI-00101 Helsinki, Finland E-mail: hilppa.gregow@fmi.fi

Tree species identification using optical remote sensing is challenging. Modern digital photogrammetric cameras enable radiometrically quantitative remote sensing and the estimation of reflectance images, in which the observations depend largely on the reflectance properties of targets. Previous research has shown that there are species-specific differences in how the brightness observed changes when the viewing direction in an aerial image is altered. We investigated if accounting for such directional signatures enhances species classification, using atmospherically corrected, real and simulated multispectral Leica ADS40 line-camera data. Canopy in direct and diffuse illumination were differentiated and species-specific variance-covariance structures were analyzed in real reflectance data, using mixed-effects modeling. Species classification simulations aimed at elucidating the level of accuracy that can be achieved by using images of different quality, number and view-illumination geometry. In real data, a substantial variance component was explained by tree effect, which demonstrates that observations from a tree correlate between observation geometries as well as spectrally. Near-infrared band showed the strongest tree effect, while the directionality was weak in that band. The gain from directional signatures was insignificant in real data, while simulations showed a potential gain of 1–3 percentage points in species classification accuracy. The quality of reflectance calibration was found to be important as well as the image acquisition geometry. We conclude that increasing the number of image observations cancels out random observation noise and reflectance calibration errors, but fails to eliminate the tree effect and systematic calibration inaccuracy. Directional reflectance constitutes a marginal improvement in tree species classification.

• Korpela, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014, Finland E-mail: ilkka.korpela@helsinki.fi
• Mehtätalo, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: lauri.mehtatalo@uef.fi
• Markelin, Department of Remote Sensing and Photogrammetry, Finnish Geodetic Institute, P.O. Box 15, FI-02431 Masala, Finland E-mail: lauri.markelin@fgi.fi
• Seppänen, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: anne.seppanen@arbonaut.com
• Kangas, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014, Finland E-mail: annika.kangas@helsinki.fi

Buyers of standing timber take not only the market situation but also the harvest costs into consideration when making purchase offers. In Finland, 85% of all timber is sold as standing timber, but there is little information for forest owners and third parties regarding how differences in harvest costs are reflected in the stumpage prices. This article analyses the relationship between sale-specific factors and stumpage prices in Finland. Data on 4824 standing timber sales between 2008 and 2012 were gathered from five local Forest Management Associations. Regression analyses were run on the stumpage prices (euros m^–3) paid for sawlogs and pulpwood. Seasonal harvest restrictions, the volume of the sale and the timber assortment influenced stumpage prices, as did the presence of forest damages. Prices also differed over time and between locations. Furthermore, the forest haulage distance was statistically significant for pulpwood. The results suggest that the size of the individual sales and the composition of assortments affect the income owners earn from their forest. The results can be used to estimate stumpage prices and the monetary impacts on forest owners of procedures such as forest road network planning and land consolidation, as well as for valuation of forests.

• Kolis, Aalto University, Department of Real Estate, Planning and Geoinformatics, P.O. Box 12200, FI-00076 Aalto, Finland E-mail: karin.kolis@aalto.fi
• Hiironen, Aalto University, Department of Real Estate, Planning and Geoinformatics, P.O. Box 12200, FI-00076 Aalto, Finland E-mail: juhana.hiironen@aalto.fi
• Ärölä, National Land Survey of Finland, Production Support Services, P.O. Box 84, FI-00521 Helsinki, Finland E-mail: esa.arola@nls.fi
• Vitikainen, Aalto University, Department of Real Estate, Planning and Geoinformatics, P.O. Box 12200, FI-00076 Aalto, Finland E-mail: arvo.vitikainen@aalto.fi

Distribution ranges of spruces, European Picea abies and Siberian P. obovata intersect in the Eastern Europe and Urals, forming wide zone of introgressive hybridization where species discrimination is difficult. We applied both molecular (mtDNA and cpDNA sequences) and morphological approaches with goals of elucidating the origin of spruces in undisturbed forests of Russian Karelia (considered as part of the abovementioned introgression zone). Karelian spruces have morphology that is typical for P. obovata and characterized with genetic variation, described previously for P. abies. This contradiction between morphology and organellar DNA could be itself an evidence of hybrid origin because morphological data should have a genetic basis. If the genes responsible for the observed morphological differences are nuclear, that explains why we did not see any deviation of Karelian spruces towards P. obovata in organellar markers. In this situation nuclear gene flow from P. obovata should be performed via pollen. Thus, we should expect Karelian spruces to have cpDNA haplotypes (inherited paternally in conifers) typical for P. obovata. However, it is not the case for the majority of plants sampled and requires additional explanation like chloroplast capture in the introgresson zone. In all, we think that Karelian spruces evolved due to introgression between P. abies and P. obovata. However, it is unclear whether Karelian spruces could be treated as P. fennica, because unequivocal morphological and genetic characters of this hybridogenous species are still absent.

• Volkova, Moscow South-West High School (No. 1543), 26 Bakinskikh komissarov str. 3–5, RU-119571 Moscow, Russia E-mail: avolkov@orc.ru
• Shipunov, Department of Biology, Minot State University, Minot, North Dakota, USA 58707 E-mail: dactylorhiza@gmail.com
• Borisova, Biological Department, Moscow State University, Vorobjevy Gory, RU-119899, Moscow, Russia E-mail: salixhastata@ya.ru
• Moseng, Minot High School, Minot, North Dakota, USA 58701 E-mail: dactylorhiza@gmail.com
• Ivens, Department of Biology, Minot State University, Minot, North Dakota, USA 58707 E-mail: dactylorhiza@gmail.com

On Nordic clearcuts, today’s tree planting machines produce high-quality but costly regenerations. Much of this high cost is attributable to the planting machines’ low productivity. One promising way of raising productivity is to lessen the time spent manually reloading seedlings onto the carousels of crane-mounted planting devices. Using MagMat, a carousel test-rig designed by engineering students, we studied how much faster tray-wise seedling reloading is on the Bracke Planter compared to reloading with today’s seedling-wise-loaded carousel. The MagMat test-rig held eight Hiko cultivation trays from which seedlings were deplugged individually and dropped into the planting tube. The time study confirmed that seedling reloading was on average twice as fast with MagMat compared to today’s seedling carousel, thereby increasing assumed planting machine productivity by 8–9% depending on the planting device used. MagMat’s cost-efficiency was analysed to be particularly reliant on its added investment cost, mechanical availability and how quickly trays can be switched automatically. Nevertheless, MagMat’s field performance illustrated the overall potential of tray-wise loading compared to piecewise seedling loading for increasing the productivity of crane-mounted planting devices. Also, deplugging proved to be a reliable method of extracting seedlings from the rigid, copper-painted Hiko cultivation trays even when performed at the excavator’s boom-tip during mounding work. We conclude that, rather than piecewise seedling loading, tray-wise loading combined with deplugging seedlings from suitable cultivation trays is a reliable and much more time-efficient method to feed seedlings on probably any type of tree planting machine.

• Ersson, Department of Forest Biomaterials and Technology, SLU, SE-90183 Umeå, Sweden E-mail: back.tomas.ersson@slu.se
• Bergsten, Department of Forest Biomaterials and Technology, SLU, SE-90183 Umeå, Sweden E-mail: urban.bergsten@slu.se
• Lindroos, Department of Forest Biomaterials and Technology, SLU, SE-90183 Umeå, Sweden E-mail: ola.lindroos@slu.se

A simple and efficient photometric methodology is presented, covering all steps from field data acquisition to binarization and allowing for leaf contour modelling. This method comprises the modelling of area and size (correlated and modelled with a Chapman-Richards growth function, using final length as one parameter), and four shape descriptors, from which the entire contour can be reconstructed rather well using a specific spline methodology. As an improvement of this contour modelling method, a set of parameterized polynomials was used. To model the temporal kinetics of the shape, geodesics in shape spaces were employed. Finally it is shown how this methodology is integrated into the 3D modelling platform GroIMP.

• Henke, Department Ecoinformatics, Biometrics & Forest Growth, University of Göttingen, 37077 Göttingen, Germany E-mail: mhenke@uni-goettingen.de
• Huckemann, Institute of Mathematical Stochastics, University of Göttingen, 37077 Göttingen, Germany E-mail: huckeman@math.uni-goettingen.de
• Kurth, Department Ecoinformatics, Biometrics & Forest Growth, University of Göttingen, 37077 Göttingen, Germany E-mail: wk@informatik.uni-goettingen.de
• Sloboda, Department Ecoinformatics, Biometrics & Forest Growth, University of Göttingen, 37077 Göttingen, Germany E-mail: bslobod@web.de

The performance of growth models implemented in the Swedish Forest Planning System Heureka was evaluated. Four basal area growth models were evaluated by comparing their predictions to data from five-year growth records for 1711 permanent sample plots of the National Forest Inventory (NFI-data). Also, two alternative implementations of Heureka, including a combined stand- and tree-level basal area growth model and a single tree-level model, respectively, were evaluated using data from 57 blocks in a thinning experiment (GG-data) involving Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst) in which the trees were monitored for 30 years after the first thinning. The predicted volume growth was also compared to observed values. Growth models based on data from 1970’s and 1980’s overestimated growth in the NFI test plots from the early 2000’s by about 3%. Stand-level models had larger precision than tree-level models. Basal area growth was underestimated in dense NFI-plots and overestimated in non-thinned GG-plots, illustrating an un-solved modelling problem. Basal area growth was overestimated by 2–5% also in the GG-plots over the whole observation period. Volume growth was however accurately predicted for pine and underestimated by 2% for spruce. The relative prediction error did not increase with increasing length of prediction period. Thinning response models calibrated with GG-data worked well in the total application and produced growth levels for different thinning alternatives in line with observations.

• Fahlvik, Department of Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: nils.fahlvik@slu.se
• Elfving, Department of Forest Ecology and Management, Swedish University of Agricultural Sciences, SE-901 83 Umeå, Sweden E-mail: bjorn.elfving@slu.se
• Wikström, Peder Wikström Skogsanalys AB, c/o Peder Wikström, Huldrans väg 1, SE-907 52 Umeå, Sweden E-mail: peder.wikstrom@slu.se

In Finland, there are currently two, parallel sample-plot-based forest inventory systems, which differ in their methodologies, sampling designs, and objectives. One is the National Forest Inventory (NFI), aimed at unbiased inventory results at national and regional level. The other is the Forest Centre’s management-oriented forest inventory based on interpretation of airborne laser scanning and aerial images, with the aim of locally accurate stand-level forest estimates. The National Forest Inventory utilises relascope sample plots with systematic cluster sampling. This inventory method is optimised for accuracy of regional volume estimates. In contrast, the management-oriented forest inventory utilises circular sample plots with an allocation system covering certain pre-defined forest classes in the inventory area. This method is optimised to produce reference data for interpretation of the remote-sensing materials in use. In this study, we tested the feasibility of the National Forest Inventory sample plots in provision of additional reference data for the management-oriented inventory. Various combinations of NFI plots and management inventory plots were tested in the interpretation of the laser and aerial-image data. Adding NFI plots in the reference data generally improved the accuracy of the volume estimates by tree species but not the estimates of total volume or stand mean height and diameter. The difference between the plot types in the NFI and management inventories causes difficulties in combination of the two datasets.

• Tuominen, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: sakari.tuominen@metla.fi
• Pitkänen, Finnish Forest Research Institute, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: juho.pitkanen@metla.fi
• Balazs, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: andras.balazs@metla.fi
• Korhonen, Finnish Forest Research Institute, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: kari.t.korhonen@metla.fi
• Hyvönen, Finnish Forest Research Institute, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: pekka.hyvonen@metla.fi
• Muinonen, Finnish Forest Research Institute, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: eero.muinonen@metla.fi

Labour models were developed to estimate the time required to Pre-Commercially Thin (PCT) with a clearing saw 4- to 20-year-old stands of the main commercial tree species in Finland. Labour (i.e., work-time consumption) was estimated from the density and stem diameter of the removal of 448 stands via an existing work productivity function. The removal based estimator attained was used as the basis for a priori mixed linear regression models. The main finding was that when a young stand grows and gets older, the work time needed to make a PCT increases. The stands of Norway spruce (Picea abies (L.) Karst.), Scots pine (Pinus sylvestris L.) and hardwoods (Betula spp.) required an additional 8.2%, 5.2%, and 3.3% work-time per year, respectively. Site fertility also played a role in that the most fertile site (mesic OMT) had an estimated labour requirement 114% higher than that for dryish VT. We also note that, per unit area, small stands require less labour than large ones and soil preparation method had a minor effect on the labour estimate. The stands which had previously gone through PCT were separately analysed. In those stands, the only significant variable concerning the labour estimate was age. The a priori models described here can help foresters to develop economic management programmes and issue quotes for forestry services.

• Uotila, Finnish Forest Research Institute, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: karri.uotila@metla.fi
• Saksa, Finnish Forest Research Institute, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@metla.fi
• Rantala, Metsä Group, Lielahdenkatu 10, FI-33400 Tampere, Finland E-mail: juho.rantala@metsagroup.com
• Kiljunen, Metsähallitus, Asemakatu 7, FI-70107 Kuopio, Finland E-mail: nuutti.kiljunen@metsa.fi

Spatially explicit data on forest attributes is demanded for various research with landscape perspective. Existing datasets with estimates of continuous forest variables are often used as the basis for producing categorical forest type maps. Normally, this type of maps are used without knowing their accuracy. This paper presents a Bayesian network model for estimating pixel level class membership probabilities of thus derived categorical maps. Class membership probabilities can be used as a post-classification measure of map accuracy and in the process of map classification affecting the assignments of class labels. The method is applied in mapping deciduous dominated forests on the basis of the k-NN Sweden 2005 dataset in a study area in southern Sweden. The results indicate rather low accuracy for deciduous class regardless of the map classification method: 0.48 versus 0.50 in the maps classified without and with the use of the class membership probabilities given equal deciduous area. When probability-based classification is applied, the level of accuracy varies with the assumed map class proportions. Thus, when deciduous class area corresponding to the National Forest Inventory estimate was used, the accuracy of only 0.35 was obtained for the deciduous map class.

• Trubins, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, Box 49, 230 53 Alnarp, Sweden E-mail: renats.trubins@slu.se
• Sallnäs, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, Box 49, 230 53 Alnarp, Sweden E-mail: ola.sallnas@slu.se

Moose browsing damage from the winter preceding the study (recent damage) on Scots pine, birch and aspen was examined in relation to forage availability, an index of moose population density and site productivity in young forests in the hemiboreal zone. Recent damage was observed for 4.1 ± 0.54% (mean ± SE; Scots pine), 16.8 ± 1.89% (birch) and 67.6 ± 13.76% (aspen) of the trees. A multiple regression with five independent variables explained 19% (Scots pine) 14% (birch) and 33% (aspen) of the variation in recent damage. Cover of Scots pine browse was the most important variable for predicting damage to Scots pine and accounted for 44% of the explained variation. When birch was overtopping pine there was a significant increase in damage to pine. Moose index was the only significant variable to explain recent damage to birch, and accounted for 64% of the explained variation. For aspen, damage was negatively correlated to coverage of Scots pine and birch browse, each variable accounting for 38% of the explained variation. For Scots pine, increasing the number of pines ha^–1 and performing pre-commercial thinning in such a way that pines are not overtopped may be efficient ways of reducing damage proportions, whereas birch needs to be protected from moose (by a reduction of the moose population or otherwise) in order to escape damage. Increased amounts of Scots pine browse and birch browse may also reduce damage levels to aspen, according to this study.

• Bergqvist, Swedish Association for Hunting and Wildlife Management, Öster Malma, SE-611 91 Nyköping, Sweden E-mail: goran.bergqvist@jagareforbundet.se
• Bergström, Forestry Research Institute of Sweden, Uppsala Science Park, SE-751 83 Uppsala, Sweden; (present) Gropgränd 2 A, SE-753 10 Uppsala, Sweden E-mail: larsrogerbergstrom@yahoo.com
• Wallgren, Forestry Research Institute of Sweden, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: martha.wallgren@skogforsk.se

The study compared motor-manual cut-to-length (CTL) harvesting, motor-manual whole-tree (WT) harvesting, mechanized CTL harvesting and mechanized WT harvesting as applied to the production of energy chips from the second thinning of Mediterranean pine plantations in flat terrain. Mechanization increased productivity between 6 and 20 times, depending on process step. It also allowed reducing thinning cost by a factor 4. Shifting from CTL to WT harvesting resulted in a reduction of harvesting cost between 40 and 50%. Fuel consumption was between 40 and 100% higher for CTL harvesting than for WT harvesting. Mechanization entailed a reduction of fuel consumption between 10 and 40%. Stand damage was generally low, between 1.5 and 6%. Mechanized CTL harvesting resulted in the lowest incidence of wounding, and the difference between mechanized CTL and manual WT harvesting was statistically significant. Soil compaction was absent or very small, depending on treatment. Mechanized thinning may produce larger increases of soil bulk density, compared to motor-manual thinning, but the difference is small, although significant. CTL harvesting leaves a larger amount of biomass on the soil, which relieves possible concerns about soil nutrient depletion. On the other hand, heavy residue loads may increase fire risk especially in sensitive Mediterranean environments.

• Spinelli, CNR IVALSA,Via Madonna del Piano 10, Sesto Fiorentino (FI), Italy E-mail: spinelli@ivalsa.cnr.it
• Lombardini, CNR IVALSA,Via Madonna del Piano 10, Sesto Fiorentino (FI), Italy E-mail: lombardini@ivalsa.cnr.it
• Magagnotti, CNR IVALSA,Via Biasi 75, S. Michele all’Adige (TN), Italy E-mail: magagnotti@ivalsa.cnr.it

The response of surface soil after 45- to 52-years to repeated nitrogen and phosphorus fertilisation was studied. This study included 30 factorial experiments established in young (5- to 30-year-old) stands using plots of 900 m², on average, and by randomising treatments within each experiment. Total amount of N added varied from 534 to 1908 kg ha^–1 and that of P from 69 to 193 kg ha^–1, repeated at every second N fertilisation. Liming was performed twice; in total, 6000 kg ha^–1 of dolomite was applied. Nitrogen fertilisation increased the mass of the organic layer and the amount of carbon and consequently the amounts of most of the elements in the organic layer. In both the organic layer and the 0–10 cm layer of mineral soil, nitrogen fertilisation decreased the C/N ratio and tended to lower pH, although only slightly. Phosphorus fertilisation increased the amounts of P and Ca. Liming increased the total amounts of most elements in the organic layer, except for C and N. We were able to derive models to describe how changes in the chemical properties of the surface soil depended on doses of elements and on site and stand properties.

• Saarsalmi, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: anna.saarsalmi@metla.fi
• Tamminen, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: pekka.tamminen@metla.fi
• Kukkola, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: mikko.kukkola@metla.fi

Many protected areas have been under intensive forest management prior to protection and thus lack natural ecosystem structures and dynamics. Dead wood is a key structure in forests harboring hundreds of threatened species. We investigated the ecological success of dead wood creation as a boreal forest restoration measure. We analysed whether the polypore communities of chain-saw felled and girdled (subsequently fallen) Norway spruce (Picea abies (L.) H. Karst.) logs differ from naturally formed spruce logs of similar decay stage and size. The study was conducted in Leivonmäki National Park in central Finland 8 years after the restoration measures. The average number of polypore species was highest on the chain-saw felled logs and most of the common polypore species were most frequent on this substrate. However, among the natural logs, number of species increased more steeply with increasing number of logs, suggesting greater variation in community composition on this substrate. The old-growth forest indicator Phellinus ferrugineofuscus occurred frequently on natural logs, occupied a few girdled logs but was absent from chain-saw felled logs. Our results show that from the polypore perspective created logs do not fully mimic natural logs, suggesting that creating substrates for species may pose a challenge for restoration.

• Komonen, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland E-mail: atte.komonen@jyu.fi
• Halme, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland E-mail: panu.halme@jyu.fi
• Jäntti, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland E-mail: mari.j.jantti@student.jyu.fi
• Koskela, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland E-mail: tuuli.e.koskela@student.jyu.fi
• Kotiaho, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland E-mail: janne.kotiaho@jyu.fi
• Toivanen, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland; Current: Birdlife Finland, Annankatu 29 A 16, FI-00100 Helsinki, Finland E-mail: tero.toivanen@birdlife.fi

Contemporary forest harvesting practices, specifically harvesting woody biomass as a source of bioenergy feedstock, may remove more woody debris from a site than conventional harvesting. Woody debris, particularly smaller diameter woody debris, plays a key role in maintaining ecosystem nutrient stores following disturbance. Understanding nutrient concentrations within woody debris is necessary for assessing the long-term nutrient balance consequences of altered woody debris retention, particularly in forests slated for use as bioenergy feedstocks. Nutrient concentrations in downed woody debris of various sizes, decay classes, and species were characterized within one such forest type, Populus tremuloides Michx.-dominated forests of northern Minnesota, USA. Nutrient concentrations differed significantly between size and decay classes and generally increased as decay progressed. Fine woody debris (≤ 7.5 cm diameter) had higher nutrient concentrations than coarse woody debris (> 7.5 cm diameter) for all nutrients examined except Na and Mn, and nutrient concentrations varied among species. Concentrations of N, Mn, Al, Fe, and Zn in coarse woody debris increased between one and three orders of magnitude, while K decreased by an order of magnitude with progressing decay. The variations in nutrient concentrations observed here underscore the complexity of woody debris nutrient stores in forested ecosystems and suggest that retaining fine woody debris at harvest may provide a potentially important source of nutrients following intensive removals of bioenergy feedstocks.

• Klockow, Department of Forest Resources, University of Minnesota, St. Paul, MN 55108, USA E-mail: klock039@umn.edu
• D'Amato, Department of Forest Resources, University of Minnesota, St. Paul, MN 55108, USA E-mail: damato@umn.edu
• Bradford, US Geological Survey, Southwest Biological Science Center, Flagstaff, AZ 86001, USA E-mail: jbradford@usgs.gov
• Fraver, School of Forest Resources, University of Maine, Orono, ME 04469, USA E-mail: shawn.fraver@maine.edu

Usnea longissima Ach. is a circumboreal epiphytic lichen draping tree canopies in moist coastal and mountainous forests. It is extinct from many European and North-American localities, presumably due to industrial forestry and air pollution, but still has a stronghold in parts of Scandinavia and U.S. and Canadian Pacific Northwest. In 2005/06 we used a comparative and retrospective approach to evaluate how present and historic tree and stand characteristics influenced the occurrence and abundance of the lichen (Storaunet et al. 2008). In 2012, we re-inventoried ten Norway spruce forest stands with 401 U. longissima-bearing trees and recorded changes in the number of U. longissima thalli. Seven of the stands had been experimentally, selectively logged 5–8 years before, where the lichen-bearing trees had been marked in the field and were avoided during the logging operation. Total number of lichen-bearing trees decreased slightly (2.9%), whereas the number of thalli had increased with 34%. Number of thalli increased more where the forest was open (low basal area, m²ha^-1) whether or not the low tree density was caused by the logging events. At high tree densities the change in number of thalli was negligible. We suggest that selective logging, securing lichen-bearing trees, may be a viable management option to keep tree density from becoming too dense, thereby enhancing growth and establishment of U. longissima.

• Storaunet, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway E-mail: stk@skogoglandskap.no
• Rolstad, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway E-mail: roj@skogoglandskap.no
• Rolstad, Skogfaglig Rådgivning, Holmsida 126, NO-1488 Hakadal, Norway E-mail: roe@skogoglandskap.no

National Forest Inventories (NFIs) are becoming increasingly important worldwide in order to provide information about the multiple functions of forests, e.g. their provision of raw materials to industry, biodiversity and their capacity to store carbon for mitigating climate change. In several countries the history of NFIs is very long. For these countries a specific challenge is to keep the inventories up-to-date without sacrificing the advantages associated with long time series. At the turn of the 20th century European NFIs faced some major challenges. In this article we describe the history and the recent developments of the Swedish NFI as an example from which general observations are made and discussed. The Swedish NFI started in 1923 and has evolved from an inventory with a narrow focus on wood resources to an inventory today which aims to provide information about all major forest ecosystem services. It can be concluded that the traditional approaches of most European NFIs, e.g. to collect data through sample plot field inventories, has proved to be applicable even for a wide range of new information requirements. Specifically, detailed data about land use, trees, vegetation, and soils has found new important uses in connection with biodiversity assessments and the estimation of greenhouse gas emissions. Though time-consuming and difficult, making NFI information comparable across countries through harmonization appears to be a useful approach. The European National Forest Inventory Network (ENFIN) was formed in 2003 and has been successful in pan-European NFI harmonization.

• Fridman, Swedish University of Agricultural Sciences (SLU), SE-901 83 Umeå, Sweden E-mail: jonas.fridman@slu.se
• Holm, Swedish University of Agricultural Sciences (SLU), SE-901 83 Umeå, Sweden E-mail: soren.holm@slu.se
• Nilsson, Swedish University of Agricultural Sciences (SLU), SE-901 83 Umeå, Sweden E-mail: mats.nilsson@slu.se
• Nilsson, Swedish University of Agricultural Sciences (SLU), SE-901 83 Umeå, Sweden E-mail: per.nilsson@slu.se
• Ringvall, Swedish University of Agricultural Sciences (SLU), SE-901 83 Umeå, Sweden E-mail: Anna.Ringvall@slu.se
• Ståhl, Swedish University of Agricultural Sciences (SLU), SE-901 83 Umeå, Sweden E-mail: goran.stahl@slu.se

This paper presents a review of the most common methods in ecological studies aimed at spatial analysis of population structures (horizontal and vertical), based on point process statistics. Methods based on simple spatially explicit indices as well as more sophisticated methods relying on functions are described in a comprehensible manner. Simple indices revealing the information on spatial structure at the scale of the nearest neighbor can be easily implemented in practical forestry. On the other hand, spatial functions, based on much more detailed data, describe the spatial structure in terms of the spatial relationships between the natural processes and population structures and because of this complexity they are rarely used in forest practice. Including both methods in a single paper is also valuable from the potential reader’s point of view saving their time for searching and choosing the appropriate method to make their spatial analysis. This paper can also serve as an initial guide for young researchers or those who are going to start their studies on spatial aspects of bio-systems. Avoiding the statistical and mathematical details makes this paper understandable for readers who are not statisticians or mathematicians. Readers will find many references related to each method described here, allowing them to find solutions to different problems observed in practice. This paper ends with a list of the most common specific software packages available to support spatial analysis.

• Szmyt, Department of Silviculture, Faculty of Forestry, Poznań University of Life Sciences, ul. Wojska Polskiego 69, 60-625 Poznań, Poland E-mail: jszmyt@up.poznan.pl

Various applications currently available for Android allow the estimation of tree heights by using the 3D accelerometer on smartphones. Some make the estimation using the image on the screen, while in others, by pointing with the edges of the terminal. The present study establishes the measurement errors obtained with HTC Desire and Samsung Galaxy Note compared to those from Blume Leiss and Vertex IV. Six series of 12 measurements each were made with each hypsometer (for heights of 6 m, 8 m, 10 m and 12 m). A Kruskall Wallis test is applied to the relative errors to determine whether there are significant differences between the devices. The results indicate that the errors of the uncalibrated smartphones significantly exceed those of traditional forestry apparatus. However, calibration is a very easy procedure that can be done by means of a linear regression line between real angles (obtained with a Digital Angle Finder or with a series of measurements taken independently of the experiment), and the angles of the accelerometer. With this adjustment, the smartphones achieve adequate quality levels although the bias was not totally eliminated. The relative errors when pointing with the edges of the terminal show no significant differences compared to Blume Leiss. Applications that use the screen image give better results (no significant differences were detected with Vertex). There is currently no application that offers calibration of the linear regression slope, which is an essential requirement for ensuring the accuracy of height measurements obtained with smartphones.

• Villasante, Agroforestry Departament, Universitat de Lleida, Av. Rovira Roure 177, 25198 Lleida, Spain http://orcid.org/0000-0002-7549-7424 E-mail: avillasante@eagrof.udl.cat
• Fernandez, Agroforestry Departament, Universitat de Lleida, Av. Rovira Roure 177, 25198 Lleida, Spain E-mail: cfernandez@eagrof.udl.cat

Recently dead Scots pines (Pinus sylvestris L.) apparently killed by Ips acuminatus (Gyllenhal) were observed in Sipoo, southern Finland, in summer 2013. This record was unexpected and in contradiction with what is currently known about the distribution and aggressiveness of the species in Finland. The aim of this study was to survey a larger area in Uusimaa region, to find out whether I. acuminatus occurs frequently in recently dead pines, and whether inhabited trees share some common tree- or site-level characteristics. Galleries of I. acuminatus were found in most of the studied trees. A total of 96 inhabited trees were found in 21 separate sites. Colonized pines were typically large (average DBH 30 ± 9 cm) trees located in relatively open pine-dominated heathland stands at half-open, sun-exposed spots. The whole upper part of the trunk with thin bark was usually occupied. Galleries of Tomicus piniperda L. or T. minor Hartig occurred only in few cases in the same trees, indicating that the trees had died later in the summer. Galleries of the jewel beetle Phaenops cyanea F. were found in 13 trees. Trees colonized by I. acuminatus often occurred as small groups, with generally 1­–12 trees (average 3 trees), but in one exceptional group there were no less than 35 trees. It is possible that the hot and dry summers during the 2000s have increased the susceptibility of pines to insect damage, and have contributed to a population growth of I. acuminatus.

• Siitonen, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: juha.siitonen@metla.fi

After clearcutting, machines traffic the clearcut conducting different silvicultural activities. Many objects on a forest clearcut (slash residues, stones, stumps and roots) may disturb e.g. site preparation and planting. This paper describes properties and characteristics of these objects. A flowchart was developed that describes a possible computer-aided system that identifies the objects, and ultimately, makes a machine avoid or target them. A system for obstacle identification creates conditions for further technical development and (semi)automation of e.g., site preparation, mechanized planting, and stump removal.

• Lideskog, Division of Product and Production Development, Luleå University of Technology, SE-97187 Luleå, Sweden E-mail: hakan.lideskog@ltu.se
• Ersson, Department of Forest Biomaterials and Technology, SLU, SE-90183 Umeå, Sweden E-mail: back.tomas.ersson@slu.se
• Bergsten, Department of Forest Biomaterials and Technology, SLU, SE-90183 Umeå, Sweden E-mail: urban.bergsten@slu.se
• Karlberg, Division of Product and Production Development, Luleå University of Technology, SE-97187 Luleå, Sweden E-mail: magnus.karlberg@ltu.se

The interest in poplars (Populus spp.) has increased globally as a result of the current incentive to expand the use of biomass for energy. So far, knowledge about poplar stump sprouting is generally poor. The objective of this study was to investigate poplar stump sprouting as regeneration method in harvested plantations. A 19-year-old poplar clonal test, including 23 different clones, on former arable land was harvested in the winter. After one year, stumps were assessed for sprout survival, sprout straightness, number of sprouts, sprout height and basal diameter. Differences between clones were found for survival and sprout straightness. Clonal differences were also found in mean size of sprouts (basal diameter and height). However, no differences were found between clones when comparing size of the tallest sprout or number of sprouts produced on stumps. This study indicates that stump sprouts from various poplar clones differ in survival, sprout straightness and mean growth, which should be taken into account when planning for the next tree generation. The study needs to be complemented with other studies to better assess sprouting characteristics for different clones.

• Mc Carthy, Forestry Research Institute of Sweden, Skogforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden & Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: rebecka.mccarthy@skogforsk.se
• Ekö, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: Per.Magnus.Eko@slu.se
• Rytter, Forestry Research Institute of Sweden, Skogforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden E-mail: lars.rytter@skogforsk.se

We apply quantitative structure modelling to produce detailed information on branch-level metrics in trees. Particularly we are interested in the branch size distribution, by which we mean the total volume of branch parts distributed over the diameter classes of the parts. We investigate the possibility of predicting tree branch size distributions for trees in similar growing conditions. The quantitative structure model enables for the first time the comparisons of structure between a large number of trees. We found that the branch size distribution is similar for trees of different height in similar growing conditions. The results suggest that tree height could be used to estimate branch size distribution in areas with similar growing conditions and topography.

• Krooks, Finnish Geodetic Institute, Geodeetinrinne 2, FI–02431 Masala, Finland E-mail: Anssi.Krooks@fgi.fi
• Kaasalainen, Finnish Geodetic Institute, Geodeetinrinne 2, FI–02431 Masala, Finland E-mail: Sanna.Kaasalainen@fgi.fi
• Kankare, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014 Helsinki, Finland E-mail: ville.kankare@helsinki.fi
• Joensuu, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014 Helsinki, Finland E-mail: marianna.joensuu@alumni.helsinki.fi
• Raumonen, Tampere University of Technology, Department of Mathematics, P.O. Box 553, Tampere, FI-33101, Finland E-mail: Pasi.Raumonen@tut.fi
• Kaasalainen, Tampere University of Technology, Department of Mathematics, P.O. Box 553, Tampere, FI-33101, Finland E-mail: Mikko.Kaasalainen@tut.fi

Vitality and height of Larix species and provenances originating from Russia were estimated in a 5-yr field trial performed in the Komi Republic (north-west Russia) using a fully randomized single-tree plot design with 7–8 blocks. Tree provenance had no significant (p > 0.05) effect on tree vitality, though for Larix sukazcewii originating from the European part of Russia, trees from the northern regions were more vital than those from southern regions. Provenance was a significant (p < 0.05) factor for height, where the average height of 136 cm varied considerably (168 cm for trees from Nizhnij Novgorod and 111 cm for trees from Ufa). There were no significant correlations when vitality and height were compared to geographic and climatic variables for the locations.

• Fedorkov, Institute of Biology, Komi Science Center, Russian Academy of Sciences, Syktyvkar, 167982, Kommunisticheskaya st., 28, Russia E-mail: fedorkov@ib.komisc.ru