Articles containing the keyword 'biomass'

Dry mass and nutrient (N, P, K, Ca, Mg, B) contents of field layer vegetation and a combination of bottom layer vegetation and litter (referred to as bottom/litter layer in the text) were studied one year before and three years after fertilization (NPK and PK) on a drained low-shrub pine bog in eastern Finland. The results of an earlier study on the tree layer were combined with those of this study in order to estimate the changes caused by fertilization in the total plant biomass and litter. Before fertilization the average dry mass of the field and bottom/litter layers was 8,400 kg ha^-1 and 7,650 kg ha^-1, respectively. The above-ground parts accounted for 25% of the total field layer biomass. The dry mass of the field and bottom/litter layers together was < 20% of the dry mass accumulated in the total plant biomass and litter. The corresponding figures for N, P, K, Ca, Mg and B were 44%, 38%, 30%, 38%, 31% and 17%, respectively. Fertilization did not significantly affect the dry mass of either the field layer vegetation or the bottom/litter layer. 33% of the applied P was accumulated in the total plant biomass and litter on the PK-fertilized plots, and 25% on the NPK-fertilized plots. For the other elements, the proportions on the PK-fertilized plots were K 31%, Ca 6%, Mg 11% and B 13%. On the NPK-fertilized plots, the corresponding figures were N 62%, K 32%, Ca 6%, Mg 9% and B 13%. Except for B and K, the accumulation of fertilizer nutrients in the understorey vegetation and litter was of the same magnitude or greater than the uptake by the tree layer.

• Finér, E-mail: lf@mm.unknown
• Nieminen, E-mail: mn@mm.unknown

A process-oriented tree and stand growth model is extended to be applicable to the analysis of timber quality, and how it is influenced by silvicultural treatments. The tree-level model is based on the carbon balance and it incorporates the dynamics of five biomass variables as well as tree height, crown base, and breast height diameter. Allocation of carbon is based on the conservation of structural relationships, in particular, the pipe model. The pipe-model relationships are extended to the whorl level, but in order to avoid a 3-dimensional model of entire crown structure, the branch module is largely stochastic and aggregated. In model construction, a top-down hierarchy is used where at each step down, the upper level sets constraints for the lower level. Some advantages of this approach are model consistency and efficiency of calculations, but probably at the cost of reduced flexibility. The detailed structure related with the branching module is preliminary and will be improved when more data becomes available. Model parameters are identified for Scots pine (Pinus sylvestris L.) in Southern Finland, and example simulations are carried out to compare the development of quality characteristics in different stocking densities.

• Mäkelä, E-mail: am@mm.unknown
• Ikonen, E-mail: vi@mm.unknown
• Vanninen, E-mail: pv@mm.unknown

The carbon reservoir of ecosystems was estimated based on field measurements for forests and peatlands on an area in Finland covering 263,000 km² and extending about 900 km across the boreal zone from south to north. More than two thirds of the reservoir was in peat, and less than ten per cent in trees. Forest ecosystems growing on mineral soils covering 144,000 km² contained 10–11 kg C m^-2 on an average, including both vegetation (3.4 kg C m^-2) and soil (uppermost 75 cm; 7.2 kg C m^-2). Mire ecosystems covering 65,000 km² contained an average of 72 kg C m^-2 as peat. For the landscape consisting of peatlands, closed and open forests, and inland water, excluding arable and built-up land, a reservoir of 24.6 kg C m^-2 was observed. This includes the peat, forest soil and tree biomass. This is an underestimate of the true total reservoir, because there are additional unknown reservoirs in deep soil, lake sediments, woody debris, and ground vegetation. Geographic distributions of the reservoirs were described, analysed and discussed. The highest reservoir, 35–40 kg C m^-2, was observed in sub-regions in central western and north western Finland. Many estimates given for the boreal carbon reservoirs have been higher than those of ours. Either the Finnish environment contains less carbon per unit area than the rest of the boreal zone, or the global boreal reservoir has earlier been overestimated. In order to reduce uncertainties of the global estimates, statistically representative measurements are needed especially on Russian and Canadian peatlands.

• Kauppi, E-mail: pk@mm.unknown
• Hänninen, E-mail: ph@mm.unknown
• Henttonen, E-mail: hh@mm.unknown
• Ihalainen, E-mail: ai@mm.unknown
• Lappalainen, E-mail: el@mm.unknown
• Posch, E-mail: mp@mm.unknown
• Starr, E-mail: ms@mm.unknown
• Tamminen, E-mail: pt@mm.unknown

A simple, manually-operated and easily portable device for sampling volumetric soil cores to a depth of 100 cm with a minimum soil disturbance is described. The device consists of a sample tube, a sampler and an extension tube. A dead blow nylon mallet is used to force the sampler into the soil and a small winch attached to an aluminium tube pulls the sampler from the soil. The total weight of the equipment (sampler, mallet and winch) is 18.5 kg and may be carried in the trunk of a small car. Sampling is easily done by one person in good physical condition but four-handed operation is recommended as more efficient. The sampling device has been in heavy use during the summers of 1993–95 when several hundred soil cores have been extracted on various sites all over Finland.

• Westman, E-mail: cw@mm.unknown

The biomass production and nutrient uptake of silver birch (Betula pendula Roth), downy birch (Betula pubescens Erhr.), grey alder (Alnus incana (L.) Moench), native willows Salix triandra L. and S. phylicifolia L. and exotic willows S. x dasyclados and S. ’Aquatica’ growing on a clay mineral soil field (Sukeva) and on two cut-away peatland areas (Piipsanneva, Valkeasuo) were investigated.

Biomass production of downy birch was greater than that of silver birch, and the biomass production of the native willows greater than that of the exotic ones. The performance of S. phylicifolia was the best of the studied willow species. Exotic willows were susceptible to frost damage and their winter hardiness was poor. The production of all species was lower on the clay mineral soil field than on the cut-away peatland areas. Fertilization of birches and alder – on the double dose given to the willows – increased biomass production. After 6 growing seasons the leafless biomass production of fertilized silver birch at Piipsanneca was 21 t ha^-1 (at Valkeasuo 34 t ha^-1) and of grey alder 24 t ha^-1, and that of S. triandra after five growing seasons 31 t ha^-1, S. phylicifolia 38 t ha^-1 and of S. x dasyclados 16 t ha^-1.

6-year-old stands of silver birch bound more nutrients per unit biomass than downy birch stands. Grey alder bound more N, Ca and Co but less Mn and Zn per unit biomass than silver and downy birch. On the field more P was bound in grey alder per unit biomass compared to downy birch. The willows had more K per unit biomass than the other tree species, and the exotic willow species more N than the native ones. Less N, K and Mg were bound per unit biomass of S. phylicifolia compared to the other tree species.

• Hytönen, E-mail: jh@mm.unknown
• Rossi, E-mail: pr@mm.unknown
• Saarsalmi, E-mail: as@mm.unknown

The effects of repeated fertilizer treatment on biomass production and nutrient status of willow (Salix ’Aquatica’) plantations established on two cut-away peatland areas in western Finland were studied over a rotation period of three years. Comparisons were made between single fertilizer applications and repeated annual fertilization.

The annually repeated fertilizer application increased the amounts of acid ammonium acetate extractable phosphorus and potassium in the soil as well as the concentrations of foliar nitrogen, phosphorus and potassium compared to single application. Depending on the fertilizer treatment and application rate, annual fertilizer application resulted in over two times higher biomass production when compared to single fertilizer application over a three-year rotation period. The effect of phosphorus fertilizer application lasted longer than that of nitrogen. The optimum fertilization regime for biomass production requires that nitrogen fertilizer should be applied annually, but the effect of phosphorus can last at least over a rotation of three years. Potassium fertilizer treatment did not increase the yield in any of the experiments during the first three years. The leafless, above-ground yield of three-year-old, annually NP-fertilized willow plantations was 9.5 t ha^-1 and the total biomass, including stems, leaves, roots and the stump, averaged 17 t ha^-1.

• Hytönen, E-mail: jh@mm.unknown

The effects of fertilized treatment on the soil nutrient concentrations, biomass production and nutrient consumption of Salix x dasyclados and Salix ’Aquatica’ were studied in five experiments on three cut-away peatland sites in western and eastern Finland during three years. Factorial experiments with all combinations of N (100 kg ha^-1 a-¹⁾, P (30 kg ha^-1 a^-1) and K (80 kg ha^-1 a^-1) were conducted.

The application of P and K fertilizers increased the concentrations of corresponding extractable nutrients in the soil as well as in willow foliage. N-fertilization increased foliar nitrogen concentration. An increase in age usually led to decreases in bark and wood N, P and K concentrations and increases in bark Ca concentrations. N-fertilization increased the three-year biomass yield 1.5–2.7 times when compared to control plots. P-fertilization increased the yield only in those experimental fields whose substrates had the lowest phosphorus concentration. K-fertilization did not increase the yield in any of the experimental fields. The highest total biomass yield of NPK-fertilized willow after three growing seasons, 23 t ha^-1, was distributed in the following way: wood 42%, bark 19%, foliage 17%, stumps 6% and roots 16%. As the yield and stand age increased, more biomass was allocated in above-ground wood. Three-year-old stands (above-ground biomass 18 t ha^-1) contained as much as 196 kg N ha^-1, 26 kg P ha^-1, 101 kg K ha^-1, 74 kg Ca ha^-1 and 37 kg Mg ha ^-1. By far the highest proportion of nutrients accumulated in the foliage. The bark and wood contained relatively high proportions of calcium and phosphorus. With an increase in age and size, the amount of nitrogen and potassium bound in one dry-mass ton of willow biomass decreased while that of phosphorus remained unchanged.

• Hytönen, E-mail: jh@mm.unknown

For biomass forestry in the inland parts of Southern and Central Finland, the obvious choice of willow species is Salix myrsinifolia. However, selection of clones of indigenous species has not yet been completed and more research and selection is needed. In the Piipsanneva old peatland trial, indigenous species of willow, mostly clones of S. myrsinifolia and S. phylicifolia, were compared in terms of biomass production, coppicing, height growth and diameter distributions. In this trial, the mean annual biomass production was not particularly high; more important results were attained in the ranking of clones. The trial strengthens the hypothesis that, over the long-term, the biomass production of S. myrsinifolia is higher than that of S. phylicifolia. It was supposed that behind the highest yield there was a clone with uniform quality, one whose diameter distribution would be narrow and positively skewed. Comparisons of parameters of Weibull functions showed that the distributions of the best clones were wide, indicating that those clones use the whole growth space better than those with narrow distribution.

• Honkanen, E-mail: ah@mm.unknown

Growth and nutrition of 20 clones representing different species and interspecific hybrids of willows (Salix spp.) growing on an abandoned field were studied. There were highly significant differences between the clones as regards the survival, number of sprouts per stool, sprout mean height and diameter and stem biomass production per stool. The differences between the clones in the concentration of all nutrients in both the leaves and stems were highly significant. 

• Viherä-Aarnio, E-mail: av@mm.unknown
• Saarsalmi, E-mail: as@mm.unknown

Crown characteristics and the distribution of three years’ (1986–88) biomass production of 20 pendulous Norway spruce (Picea abies f. pendula (Lawson) Sylvén) trees with heritable narrow crown and 15 normal-growned spruces (Picea abies (L.) H. Karst.) were studied in a 19-year-old mixed stand.

The form of the crown is conical in normal-crowned trees, columnar and narrow in pendulous trees. The partitioning of aboveground biomass to stems during the studied 3-year period was significantly higher in pendulous (0.281) than in normal-crowned trees (0.255) and also the ratio between growth of stemwood and growth of needle biomass during three years was higher in pendulous trees (0.67 g g^-1) than in normal-crowned trees (0.52 g g^-1). The needle biomass was distributed higher in the crown in pendula than in normal-crowned trees and they had a higher needle biomass/branchwood biomass ratio than normal trees. The difference in harvest increment between the two crown types are mostly due to the significantly lower branchwood biomass values in pendulous than in normal-crowned trees. The higher needle ’efficiency’ in pendulous trees is probably connected with high partitioning of needle biomass to the upper part of the crown in pendulous trees.

The PDF includes an abstract in Finnish.

• Pulkkinen, E-mail: pp@mm.unknown

Scots pine (Pinus sylvestris L.) living root biomass (ø≤ 10 mm) was 640 g/m² on the studied low-shrub pine bog before fertilization, and that of the ground vegetation almost the same. The total root necromass was 23% of the biomass of living roots. The length of the pine roots was 2,440 m/m². The biomass of living roots and root necromass were mostly located in the top 20 cm layer of the soil. The ø < 1 mm pine root fraction accounted for almost 90% of the pine root length; in contrast, over 50% of the biomass was in the 1–10 mm thick root biomass, pine root length and PK (MgB) fertilization did not affect total living root biomass, pine root length, nor the root necromass during the three-year observation period.

The PDF includes an abstract in Finnish.

• Finér, E-mail: lf@mm.unknown

The development of shoot number and shoot properties was examined in successive shoot cohorts of young widely-spaced Scots pine trees (Pinus sylvestris L.) growing in a progeny trial. This was accomplished by reconstructing the branching process of the trees over a period of five years, from tree age 4 to 8. During this time the number of shoots in successive shoot cohorts increased rapidly, while at the same time the mean shoot length decreased. The decrease in shoot lengths from older to younger shoots was accompanied by a decline in the bifuraction frequency of the shoots. In general, rapid changes occurred in the branching characteristics during the yearly development of the trees. The variation in the branching characteristics was reflected in the development of the architecture and biomass production of the trees.

The PDF includes an abstract in Finnish.

• Kuuluvainen, E-mail: tk@mm.unknown

The architecture of Scots pine (Pinus sylvestris L.) was studied in an eight-year-old progeny test. The measurements included characteristics of crown structure, spatial distribution of shoots and yield components. The spatial distribution of shoots showed striking between-tree differences, and two extreme distribution patterns were detected. One represented a non-layered structure with a vertically relative even shoot distribution, and the other a layered structure with a vertically highly uneven shoot distribution.

Close correlations existed between several components of tree architecture and it is suggested that changes in the phenotypic architecture in Scots pine follow an epigenetic pattern, which enables the prediction of adaptational changes in structural components. The structural characteristics related to high above-ground biomass were a long crown, high total shoot length, high number of branches per whorl and big shoots of low needle density occupying a big share of the crown volume.

The PDF includes a summary in Finnish.

• Kuuluvainen, E-mail: tk@mm.unknown
• Kanninen, E-mail: mk@mm.unknown
• Salmi, E-mail: js@mm.unknown

Possibilities of developing suitable willow (Salix spp.) clones for short-rotation forestry on mined peatlands in the north-western area of Finland were studied in a field experiment in which 300 willow clones were tested during 1985–87. Most of the tested clones started to grow from cuttings on limed and fertilized peat soil. Salix viminalis L. clones of southern origin had a higher leafless above ground biomass production than the well adapted control clone and the local Finnish willows, but their winter hardiness was not satisfactory. The growth habit of some southern willows was also better than that of the control clone. It was also possible to select clones with good sprouting capacity. There were few Salix myrsinifolia Calisb. clones of Finnish origin, which had better cold tolerance than all other willows tested and higher biomass production than that of the control clone. The most critical factor to be selected for this is the optimal combination of winter hardiness and biomass production. This is attempted by selecting clones on the basis of this experiment for a breeding program.

The PDF includes a summary in Finnish.

• Lumme, E-mail: il@mm.unknown
• Törmälä, E-mail: tt@mm.unknown

Salix 'Aquatica Gigantea', widely experimented and promising species for temperate zone short rotation forestry, has since 1950 recorded in Finland 23 times with different clone numbers. Salix x dasyclados Wimm., by morphological, cultivational and productivity characteristics similar willow has been recorded 16 times.

The nomenclature and origin of both willows have remained unclear in field research. Recent observations, based on morphological analyses and chromosome studies suggest that ’Aguatica gigantea’ and most S. x dasuclados clones can be collected under one Siberian species: Salix burjatica Nasarov. The true Salix x dasyclados Wimm. is a female hybrid S. x viminalis x cinerea, famous West-European basket willow that has been very little experimented in Finland.

The PDF includes an abstract in Finnish.

• Pohjonen, E-mail: vp@mm.unknown

In a greenhouse experiment that lasted for two years, nitrogenase activity, height growth and biomass production was compared in six clones of alder of which four were clones of Alnus incana and two A. incana x A. glutinos hybrids. In addition, the effect of a fertilizer nitrogen gradient was tested on one of the clones.

Clonal differences in height growth and nitrogenase activity were recorded at the end of the first growing season. The growth rhythm of some of the clones changed markedly during the second growing season but differences in nitrogenase activity between clones levelled out. Nitrogen fertilization suppressed nodulation during the first growing season, and also the following year the nitrogenase activity was significantly higher in alders grown without nitrogen supplement. Height growth and total biomass production was also depressed at rather low nitrogen levels.

The PDF includes a summary in Finnish.

• Palmgren, E-mail: kp@mm.unknown
• Saarsalmi, E-mail: as@mm.unknown
• Weber, E-mail: aw@mm.unknown

The study deals with the distribution of above-ground biomass of Vaccinium myrtillus L. along the vegetation continuum segregated by using phytosociological classification method composite clustering. The qualitative characteristics of forest sites corresponding to different vegetational clusters were defined on the basis of indirect gradient analysis of vegetation data and description of tree stand properties in stands in 160 sample areas in Southern Finland.

Six vegetation types differing from each other mainly in abundance of the most constant and dominant plant species were formed. Sample areas with rich grass-herb vegetation, as well as sample areas representing comparatively dry, barren site type were clearly separated from other groups of sample areas. Stand characteristics, particularly the proportional distribution of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) seems to be another important factor affecting the vegetation composition.

The lowest biomass values of blueberry were connected with herb-rich vegetation and shady spruce-dominated stands. Comparatively low values were connected to also other spruce-dominated sample areas. Pine-dominant or mixed pine-spruce stands the biomass values were significantly higher. Even if the site quality is essentially equal, the abundance relationships between Vaccinium myrtillus and other components of the ground vegetation may vary in wide ranges and cause difficulties in practical site classification.

The PDF includes a summary in English.

• Kuusipalo, E-mail: jk@mm.unknown

A study based on four young Scots pines (Pinus sylvestris L.) showed that the number of needle-covered shoots per crown volume unit was independent on tree position representing a constant value of 600–700 shoots/m³. This was true, even though the total shoot number decreased with deteriorating tree position. In tree crown there were fourth-order shoots in good light conditions but only first- and second-order-shoots, when light conditions were poor. The length of shoots decreased in accordance with increasing order of the shoot.

The share of the needle biomass and growth increased, when the shoot order increased. Similarly, the share of needles increased with deteriorating tree position. This was especially true in the upper crown. On the other hand, the share of the crown from the total biomass and growth increased with improving tree position. The percentage of crown system of a dominant tree in a sparse stand was 64% of that of biomass and 83% of that of growth. The corresponding values for a suppressed tree in a dense stand were 36% and 35%. The growth of wood, bark and needles in crown systems was linearly correlated with prevailing light conditions around the branch. It is evident that the tree position and light condition within the stand control the wood, bark and needle growth in the crown system and their interrelationships.

The PDF includes a summary in English.

• Viherä, E-mail: av@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown

The needle area distribution and crown structure of a young planted Scots pine (Pinus sylvestris L.) stand are described. The crown structure and crown shape showed apparent regularity in crown structure regardless of stand dynamics. Similarly, the shoot structure and individual needle area showed regularity in the number of needles per branch and shoot length unit, and consequent phytoarea density inside the needle cylinder. Also, the shoot area and needle area distributions were found to show a regular distribution of needle biomass throughout the crown, also inside the crown, in the dominant trees. In the suppressed trees the needle biomass was located in the upper crown and on the surface area of the crown. Estimates of the canopy needle area and distributions are given. The results were applied in calculations of the within-stand light regime. The results correlated well with the empirical results.

The PDF includes a summary in Finnish.

• Kellomäki, E-mail: sk@mm.unknown
• Oker-Blom, E-mail: po@mm.unknown

The proportions of needle and wood in current-year shoots in crown systems of young Scots pine (Pinus sylvestris L.) trees was studied. The proportion of needles out of the total shoot biomass increased according to the increasing number of the whorl counted from the apex. In the lower part of the crown system the needle biomass of newly-formed shoots was 50–60 fold compared to that of wood and bark biomass. In the upper part of the crown system the same ratio was 1–2. The variation in ratio between needle and wood biomass was whorl-specific and independent of tree class. The magnitude of the ratio was not related to the position of the tree in the stand nor to the prevailing light conditions within the state.

The PDF includes a summary in Finnish.

• Mäkelä, E-mail: am@mm.unknown
• Hari, E-mail: ph@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown

The technique of double normalizing, i.e. normalizing the relative needle biomass and the length of the living crown system, is applied to the modelling of the distribution of needle biomass in the canopy of young Scots pine (Pinus sylvestris L.) stands. The study based on the parameters of β-function shows that at the individual-tree level, the variance in needle distribution was not closely associated with any tree characteristics. A shift in the point of maximum needle biomass upwards unsuppressed trees was, however, evident. This was associated with an increase in the height of the trees. At the stand level, the stand mean height and stand density had an equal and a rather high potential for explaining the variance in the needle distribution. The normalized crowns are utilized in models for determining light extinction in the crown. A special technique for determining the amount of photosynthates available for growth in a particular tree is presented.

The PDF includes a summary in Finnish.

• Kellomäki, E-mail: sk@mm.unknown
• Hari, E-mail: ph@mm.unknown
• Ilonen, E-mail: pi@mm.unknown
• Kanninen, E-mail: mk@mm.unknown

The concept of tree class as indicated by dominant, codominant, intermediate and suppressed trees is analysed using empirical material representing 13 younger Scots pine (Pinus sylvestris L.) stands. The relative needle biomass, i.e. the needle biomass related to the maximum needle biomass in the stand, was closely related to the relative height of the tree and independent of the stand characteristics. Furthermore, the relative illumination of the crown system was related to the relative height of the trees as well as the relative photosynthetic capacity and tree-to-tree variation in growth. When calculated per needle biomass unit it appeared that height growth, radial growth, needle growth and other growth parameters were the highest in the suppressed trees. The suppressed trees thus appear to be more efficient in utilizing available resources than other tree classes. The ecologic significance of the results is discussed.

The PDF includes a summary in Finnish.

• Kellomäki, E-mail: sk@mm.unknown
• Hari, E-mail: ph@mm.unknown

The effect of nitrogen fertilizers on the photosynthetic capacity of conifers is assessed on the basis of literature. The review emphasizes the role of changes of needle mass as a factor affecting the result of nutrient application. In particular, the increase in needle mass results in a considerable increase in photosynthetic capacity. The effect of fertilization on the photosynthetic rate seems to be of minor importance. The effect on the photosynthetic rate is, however, poorly documented as is the case for the effect of fertilization on the respiration rate. There is evidence that proper application of nitrogen fertilizers may double the photosynthetic capacity of conifers, mainly as a result of increase in needle mass.

The PDF includes a summary in English.

• Kellomäki, E-mail: sk@mm.unknown

The distribution of the dry matter and nutrients in Scots pine (Pinus sylvestris L.) tree stock growing on a Vaccinium type site, ground vegetation, and humus were determined in the study. The greatest part of the dry matter in the tree was found in the stemwood. The living branches, roots, bark, needles and dead branches decreasing order of magnitude made up the rest of the biomass. The trees contained over 90%, the field layer vegetation 3% and the bottom layer vegetation 2% of the dry matter in the tree stand. The tree stock contained 86–95% of the total amount nutrients in the stand. The field layer vegetation contained less nutrients than the bottom layer vegetation. Nitrogen, however, was an exception, the amount being approximately the same in both vegetation layers. 

The PDF includes a summary in English.

• Lehtonen, E-mail: il@mm.unknown

The study deals with the relationship between biomass and coverage in ground vegetation of Vaccinium and Calluna forest types. The results show that the biomass of the field layer on both forest types can predict satisfactorily in both coverage values of some the most important species and groups of species. Their total height per plot was used as independent variables. The explaining power of the models constructed for Vaccinium type increased to approximately 57% and for the Calluna type to approximately 74% of the total variation in the amount of dry matter. When the biomass of the bottom layer was predicted using the same kind of variables, the explaining power for the Vaccinium type increased to approximately 35% of the total variation and for the Calluna type approximately 53% of the total variation in the amounts of dry matter. The models for field layer of both Vaccinium and Calluna types were quite well suited for describing the test material. In the case of the bottom layer, the constructed models were not suitable for describing the test material.

The PDF includes a summary in English.

• Kellomäki, E-mail: sk@mm.unknown

The possibilities of using results of coverage analyses for estimating the dry matter content of the ground vegetation has been examined in this study. The material covers 150 sample areas, 400 m² large, which represent ground vegetation of Myrtillus type in the final succession stage. When the material was subjected to linear regression analysis, the amount of dry matter in the field and ground layer has been used as dependant variables and the results of the coverage analysis as the independent variable.

The study shows that it is possible to predict satisfactorily the dry matter content of the ground vegetation by grouping the coverage, according to the coverage weight, into parts of equal size and the using them as independent explaining variables. In particular, the value of the prediction equation for the dry matter content of the field layer could be improved by using the height and density characteristics of the vegetation as explaining variables in addition to the coverage figures. Thus, slightly over 80% of the total variation of the dry matter content of the field layer could be predicted. In the case of the ground layer vegetation, the explaining power rose slightly above 70%.

The PDF includes a summary in English.

• Kellomäki, E-mail: sk@mm.unknown

This study estimates the supply of soil water required for the annual and total production of energy material by the biomass of 32-year-old plantation of red pine, Pinus resinosa Ait. in Wisconsin, the United States.

The supply of transpiration water was determined as the sum of summer precipitation, winter stored water, and condensed vapor, minus gravitational discharge and evaporation of intercepted rainfall. On the average, the 1,20 m root zone of coarse sandy soils of central Wisconsin receives 2,750 M.T. of water per hectare. During the 32nd year of plantation growth, the increment biomass, including 43% of merchantable timber, was 10,100 kg/ha, or 162 x 105 kcal/g. At this time of the culminating growth, the production of 1 kg of wood material consumed 272 kg of water. The corresponding transpiration coefficient 0,37% is near the maximum for the ecosystem of hard pines – sandy soils of glacial outwash with field capacity between 7 and 9%. On the weight basis, the annual leaf fall constituted 32% of the biomass and over 80% of merchantable timber.

The entire supply of water of 96,000 M.T./ha produces in 32 years 211,112 kg of total dry matter at a rate of 1 kg of wood per 455 kg of water, with corresponding transpiration coefficient of 0,22%. The evapogravitational losses during the early stages of the stand’s growth decreased the water utilization efficiency of trees about 40%.

The information obtained permitted to outline several hydrological relationships pertinent to forest culture, namely: maximum rate of forest growth as delineated by the supply of available transpiration water; content of available moisture in soils of high tension capacity; contribution to soil water rendered by natural subirrigation and condensation of athmospehric vapor; growth depressing effect of weeds.

The PDF includes a summary in Finnish.

• Wilde, E-mail: sw@mm.unknown

The effective heating values of the above and below ground biomass components of mature Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies (L.) H. Karst.), downy birch (Betula pubescens Ehrh), silver birch (B. pendula Roth), grey alder (Alnus incana (L.) Moench), black alder (A. glutinosa (L.) Gaertn.) and aspen (Populus tremula L.) were studied. Each sample tree was divided into wood, bark and foliage components. Bomb calorimetry was used to determine the calorimetric heating values.

The species is a significant factor in the heating value of individual tree components. The heating value of the wood proper is highest in conifers. Broadleaved species have a higher heating value of bark than conifers. The species factor diminishes when the weighted heating value of crown, whole stems or stump-root-system are considered. The crown material has a higher heating value per unit weight in comparison with fuelwood from small-sized stems or whole trees. The additional advantages of coniferous crown material are that it is non-industrial biomass resource and is readily available. The variability of both the chemical composition and the heating value is small in any given tree component of any species. However, lignin, carbohydrate and extractive content were found to vary from one part of the tree to another and to correlate with the heating value

• Nurmi, E-mail: jn@mm.unknown

The heating values of wood, inner and outer bark, and foliage components of seven small-size tree species (Pinus sylvestris L., Picea abies (L.) H. Karst., Betula pubescens Erhr., B. pendula Roth, Alnus incana (L.) Moench, A. glutinosa (L.) Gaertn., Populus tremula L.) were studied. Significant differences were found between species within each component. However, the differences between species for weighted stem, crown and whole-tree biomass are very small. The weighted heating value of the crown mass is slightly higher than that of the stem in all species. The heating value of stem, crown and whole-tree material was found to increase with increasing latitude.

The effective heating value of wood correlated best with the lignin content, inner bark with carbohydrate, and outer bark with carbohydrates and the extractives soluble in alkalic solvents. It is suggested that the determination of the heating value might be used as an indicator of the cellulose content of coniferous wood.

The PDF includes a summary in Finnish.

• Nurmi, E-mail: jn@mm.unknown

The first three-year effects of PK(MgB) and NPK(MgB) fertilization on the dry mass accumulation and nutrient cycling were studied in a Scots pine (Pinus sylvestris L.) stand growing on a drained low-shrub pine bog in Eastern Finland. The total dry mass of the tree stand before fertilization was 78 tn/ha, of which the above-ground compartments accounted for 69%. The annual above-ground dry mass production was 6.3 tn/ha, 51% of it accumulating in the tree stand.

The study period was too short for detecting any fertilization response in the stems. The total dry mass accumulation was not affected, because the increase in foliar and cone dry masses after both fertilization treatments, and that of the living branches after NPK fertilization, were compensated by the decrease in the dry mass of dead branches.

The nutrients studied accounted for 392 kg/ha (0.49%) of the total dry mass of the tree stand before fertilization. The amounts were as follows; N 173 kg/ha (44%), Ca 90 kg (23%), K 58 kg/ha (15%). The rest (18%) consisted of P, Mg, S and micronutrients combined. The unfertilized trees took up the following amounts of nutrients of the soil: N 15.6, Ca 12.8, K 4.1, P 1.3, MG 1.7, and S and Mn 1.5 kg/ha. The uptake of Fe and Zn was 510 and 130 g/ha and that of B and Cu less than 100 g/ha. More than 50% of the nutrient uptake, except for that of K and Fe, was released in litterfall. The results indicated very efficient cycling of K, Mn and B between the soil and trees.

The fertilized stands accumulated more N, P, K and B than the unfertilized ones during the tree-year study period. The increased accumulation corresponded to 35% (52 kg/ha) of the N applied on the NPK fertilized plots, 10% of the P, 25% of the K and 10% of the B on the PK and NPK fertilized plots. The increased amount of B released in litterfall after fertilization was equivalent to 4% of the applied B. Fertilization inhibited the uptake of Mn and Ca.

The PDF includes a summary in Finnish.

• Finér, E-mail: lf@mm.unknown

Willow (Salix sp.) species and clones have been selected in Finland, originally for basket willow husbandry, since 1910's. Screening for biomass willows started in 1973 by the Foundation for Forest Tree Breeding. Biomass willow research for energy started in 1978. The objective of the study was, based on theoretical background, on historical record of Finnish willow research between 1910–80 and on analysis of the Finnish biomass willow research of the 1980s, a further selection of exotic and indigenous willows for energy and chemicals.

Swedish selection of 63 exotics, mainly of Salix viminalis L. and S. burjatica Nazarov, was screened in Kompparnäs willow research site in the southern coast of Finland in 1983–89. S. viminalis showed both high yield potential and good crop certainty. The yield variation in S. burjatica were big due to rust (Melampsora sp.) infection followed by lowered winter hardiness. Three recommendable S. viminalis clones for Southern Finland were found.

Finnish indigenous species were screened based on collection (375 clones) in 1974–74 of the Foundation for Forest Tree Breeding, and the Finnish 4H-organization (566 clones) in 1978–79 in test sites in Suomusjärvi, Nurmijärvi, Kannus and Haapavesi. Salix myrsinifolia Salisb. was most productive of the indigenous willows. Five recommendable clones were selected. The second most productive indigenous was S. phylicifolia, with three recommendable clones. Based on willow hybridization studies in the Finnish Forest Research Institute, a considerable additional selection effect, boosted by heterosis, was found from the progenies. Further intraspecific crossings of geographically distant clones of S. myrsinifolia are recommended.

Based on the results, S. viminalis is recommended for practical biomass forestry application in the southernmost agroclimatic zone of Finland. S. myrsinifolia is recommended for further research and development in the other zones.

The PDF includes a summary in Finnish.

• Pohjonen, E-mail: vp@mm.unknown

The aim of this study was to investigate the ecophysiological and morphological characteristics of two salt-tolerant tree species, Eucalyptus camaldulensis Dehn. and Combretum quadrangulare Kurz. A greenhouse experiment with different levels of NaCl salinity (0, 0.5, 1.0, 1.5, and 2.0%) was set up and the results were compared with those of a field study on non-saline and saline soils. The determination of optimum gas exchange and the development and evaluation of photosynthetic models with and without water deficit were also included in this study.

Morphological characteristics under saline conditions showed that shoot height and diameter growth, shoot internode length, root length/biomass, leaf width and length, leaf area, number and biomass, and shoot/root and leaf/root ratios decreased with salinity, while leaf thickness increased with salinity. More growth was allocated to the roots than to the leaf canopy. Ecophysiological studies in laboratory showed that photosynthesis, stomatal conductance and water potential decreased with salinity, while the CO₂ compensation point increased with salinity. Transpiration, dark respiration and photorespiration increased at low salinity but decreased at high salinity levels. In the field study, however, there were no significant differences in stomatal conductance and opening between saline and non-saline soils. Model predictions supported the results of the field measurements. Adaptation to salinity was reflected in an acclimatization of tree structure in the field study. There were both functioning and structural changes of seedlings in the greenhouse experiment

In terms of ecophysiological and morphological characteristics, E. Camaldulensis showed better salt tolerance than C. Quadragulare both in the greenhouse experiment and field study

The PDF includes a summary in Finnish.

• Luangjame, E-mail: jl@mm.unknown

At the beginning of the investigation period the total biomass of the Scots pine (Pinus sylvestris L.) stands on the ordinary sedge pine mire was 48 t/ha. The biomass of the mixed stands of Scots pine and birch (Betula pubescens Erhr.) on the herbrich sedge pine mire was 91 t/ha, out of which 60% was from pine. The biomass of the Norway spruce (Picea abies (L.) H. Karst.) on the Vaccinium-Myrtillus spruce mire was 148 t/ha. The average annual net increment of the stand biomass was 5.8 t/ha in the unfertilized pine stand and 6.7 t/ha in the NPK and micronutrient fertilized one during the six-year investigation period. The corresponding figures in the mixed stand were 7.2 t/ha and 7.6 t/ha. The net increment of the biomass in the unfertilized spruce stand was 6.9 t/ha and in the fertilized 8.4 t/ha. A considerable proportion of the net increment was lost to the ground as litter in all stands.

The nitrogen, phosphorus, potassium, magnesium, iron, manganese, zinc, copper and boron cycles were investigated. The annual nitrogen uptake from the soil was 26–42 kg/ha, that of phosphorus 2.5–3.4 kg/ha, potassium 4.5–12 kg/ha, calcium 12–29 kg/ha, magnesium 2–4 kg/ha, iron 1.4–6.6 kg/ha, manganese less than 2 kg/ha and the other nutrients only some grams. Only part of the fertilized nutrients was fixed in the stand.

The PDF includes a summary in Finnish.

• Finér, E-mail: lf@mm.unknown

• Wutzler, Max Planck Institute for Biogeochemistry, Jena, Germany E-mail: tm@nn.de
• Mund, Max Planck Institute for Biogeochemistry, Jena, Germany E-mail: mm@nn.de

Forest fires pose a significant threat to forest carbon storage and sinks, yet they also play a crucial role in the natural dynamics of boreal forests. Accurate quantification of biomass changes resulting from forest fires is essential for damage assessment and controlled burning evaluation. This study utilized terrestrial laser scanning (TLS) to quantify changes in ground vegetation resulting from low-intensity surface fires. TLS data were collected before and after controlled burnings at eight one-hectare test sites in Scots pine (Pinus sylvestris L.) dominated boreal forests in Finland. A surface differencing-based method was developed to identify areas exposed to fire. Validation, based on visual interpretation of 1 × 1 m surface patches (n = 320), showed a recall, precision, and F1-score of 0.9 for the accuracy of identifying burned surfaces. The developed method allowed the assessment of the magnitude of fire-induced vegetation changes within the test sites. The proportions of burned 1 × 1 m areas within the test sites varied between 51–96%. Total volumetric change in ground vegetation was on average –1200 m³ ha^-1, with burning reducing the vegetation volume by 1700 m³ ha^-1 and vegetation growth increasing it by 500 m³ ha^-1. Substantial variations in the volumetric changes within and between the test sites were detected, highlighting the complex dynamics of surface fires, and emphasizing the importance of having observations from multiple sites. This study demonstrates that bitemporal TLS measurements provide a robust means for characterizing fire-induced changes, facilitating the assessment of the impact of surface fires on forest ecosystems.

• Tienaho, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland https://orcid.org/0000-0002-6574-5797 E-mail: noora.tienaho@uef.fi
• Saarinen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland https://orcid.org/0000-0003-2730-8892 E-mail: ninni.saarinen@uef.fi
• Yrttimaa, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland https://orcid.org/0000-0003-2648-523X E-mail: tuomas.yrttimaa@uef.fi
• Kankare, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland https://orcid.org/0000-0001-6038-1579 E-mail: ville.kankare@uef.fi
• Vastaranta, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland https://orcid.org/0000-0001-6552-9122 E-mail: mikko.vastaranta@uef.fi

We developed tree level biomass (dry weight) models for Norway spruce (Picea abies [L.] H. Karst.), silver birch (Betula pendula Roth), rowan (Sorbus aucuparia L.) and aspen (Populus tremula L.) growing in young spruce dominated seedling stands with high mixture of broadleaves. The study material was collected from three planted Norway spruce seedling stands located on mineral soil in southern Finland. Biomass models were estimated by individual tree component (stem, living branches, foliage, stump, and roots with diameter of 2 mm) by using a multi-response approach (seemingly unrelated regression), which estimated the parameters of the sub-models (tree component) simultaneously. Even though the application and generalization of the developed models can be restricted by the limited material, they provide new information of seedling biomass allocation and more reliable biomass predictions for spruce and birch growing in young seedling stand compared with those of the commonly applied biomass models (Repola 2008, 2009) in Finland. Repola’s models (2008, 2009) tended to produce biased predictions for crown and below-ground biomasses of seedlings by allocating too much biomass to roots and too little to needle and branches. In addition, this study provides biomass models for aspen and rowan, which were not previously available.

• Repola, Natural Resources Institute Finland (Luke), Ounasjoentie 6, FI-96200 Rovaniemi, Finland https://orcid.org/0000-0001-7086-0549 E-mail: jaakko.repola@luke.fi
• Luoranen, Natural Resources Institute Finland (Luke), Juntintie 154, FI-77600 Suonenjoki, Finland https://orcid.org/0000-0002-6970-2030 E-mail: jaana.luoranen@luke.fi
• Huuskonen, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland https://orcid.org/0000-0001-8630-3982 E-mail: saija.huuskonen@luke.fi
• Peltoniemi, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland https://orcid.org/0000-0003-2028-6969 E-mail: mikko.peltoniemi@luke.fi
• Väänänen, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: paivi.vaananen@luke.fi
• Uotila, Natural Resources Institute Finland (Luke), Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: karri.uotila@luke.fi

• Liepiņš, Latvian State Forest Research Institute “Silava,” Rigas Street 111, LV-2169 Salaspils, Latvia https://orcid.org/0000-0003-3030-1122 E-mail: janis.liepins@silava.lv
• Jaunslaviete, Latvian State Forest Research Institute “Silava,” Rigas Street 111, LV-2169 Salaspils, Latvia https://orcid.org/0009-0000-7322-2729 E-mail: ieva.jaunslaviete@silava.lv
• Liepiņš, Latvian State Forest Research Institute “Silava,” Rigas Street 111, LV-2169 Salaspils, Latvia https://orcid.org/0000-0002-1179-8586 E-mail: kaspars.liepins@silava.lv
• Jansone, Latvian State Forest Research Institute “Silava,” Rigas Street 111, LV-2169 Salaspils, Latvia https://orcid.org/0000-0003-2748-3797 E-mail: liga.jansone@silava.lv
• Matisons, Latvian State Forest Research Institute “Silava,” Rigas Street 111, LV-2169 Salaspils, Latvia E-mail: roberts.matisons@silava.lv
• Lazdiņš, Latvian State Forest Research Institute “Silava,” Rigas Street 111, LV-2169 Salaspils, Latvia https://orcid.org/0000-0002-7169-2011 E-mail: andis.lazdins@silava.lv
• Jansons, Latvian State Forest Research Institute “Silava,” Rigas Street 111, LV-2169 Salaspils, Latvia https://orcid.org/0000-0001-7981-4346 E-mail: aris.jansons@silava.lv

• Jåstad, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway https://orcid.org/0000-0002-1089-0284 E-mail: eirik.jastad@nmbu.no
• Nagel, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway https://orcid.org/0000-0002-3171-0262 E-mail: niels.oliver.nagel@nmbu.no
• Hu, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway https://orcid.org/0000-0003-0001-5993 E-mail: junhui.hu@nmbu.no
• Rørstad, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: per.kristian.rorstad@nmbu.no

Tree carbon (C) and nitrogen (N) concentrations and C:N ratio are critical for understanding the elemental compositions of forests, N use efficiency, productivity and the biogeochemical cycles. We evaluate differences in C and N allocation among biomass compartments of three N‑fixing tree species of Espinal Argentine eco-region; the scaling relationship between C and N and the C:N ratio variation among compartments and tree size. Neltuma affinis (Spreng.) C.E. Hughes & G.P. Lewis, Neltuma nigra (Griseb.) C.E. Hughes & G.P. Lewis and Vachellia caven (Molina) Seigler & Ebinger plants (n = 30 for each species) were felled, grouped by stem basal diameter-based size classes and partitioned into 3 biomass compartments: stem (st), large branches (lb) and small branches + leaves, flowers and fruits (sbl). C and N concentrations were markedly influenced by species and biomass compartments. In general, sbl compartment presented more N than the st and lb, while C concentrations in Neltuma stems were the highest. Overall, no isometric C–N scaling relationships were found in different compartments. C:N variations in compartments were positively correlated with N concentrations but did not exhibit any significant association with C concentrations. C:N ratios differed significantly among species and biomass compartments. The C:N ratio for compartments ranked in an order of st > lb > sbl. C:N ratio variability in sbl was the least. Only in N. affinis and V. caven stems C:N relationship differed among tree size. Our results provide evidence of the importance of using in situ C and N concentration per main tree species and biomass compartments, to more accurate estimates of C and N stocks.

• Sione, National University of Entre Ríos, Faculty of Agricultural Sciences, Ruta 11 Km 10,5 (3101) Oro Verde, Entre Ríos, Argentina; CICyTTP-CONICET (National Research and Technological Investigation Council), Materi y España (3105) Diamante, Entre Ríos, Argentina https://orcid.org/0000-0002-2119-0363 E-mail: silvana.sione@fca.uner.edu.ar
• Ledesma, National University of Entre Ríos, Faculty of Agricultural Sciences, Ruta 11 Km 10,5 (3101) Oro Verde, Entre Ríos, Argentina https://orcid.org/0000-0002-9264-1766 E-mail: silvia.ledesma@fca.uner.edu.ar
• Aceñolaza, National University of Entre Ríos, Faculty of Agricultural Sciences, Ruta 11 Km 10,5 (3101) Oro Verde, Entre Ríos, Argentina; CICyTTP-CONICET (National Research and Technological Investigation Council), Materi y España (3105) Diamante, Entre Ríos, Argentina https://orcid.org/0000-0002-5033-3466 E-mail: pablo.acenolaza@fca.uner.edu.ar
• Wilson, National University of Entre Ríos, Faculty of Agricultural Sciences, Ruta 11 Km 10,5 (3101) Oro Verde, Entre Ríos, Argentina https://orcid.org/0000-0002-1214-1041 E-mail: wilson.marcelo@inta.gob.ar

Currently, tools to predict the aboveground and belowground biomass (AGB and BGB) of woody species in Guinean savannas (and the data to calibrate them) are still lacking. Multispecies allometric equations calibrated from direct measurements can provide accurate estimates of plant biomass in local ecosystems and can be used to extrapolate local estimates of carbon stocks to the biome scale. We developed multispecies models to estimate AGB and BGB of trees and multi-stemmed shrubs in a Guinean savanna of Côte d’Ivoire. The five dominant species of the area were included in the study. We sampled a total of 100 trees and 90 shrubs destructively by harvesting their biometric data (basal stem diameter D_b, total stem height H, stump area S_S, as well as total number of stems n for shrubs), and then measured their dry AGB and BGB. We fitted log-log linear models to predict AGB and BGB from the biometric measurements. The most relevant model for predicting AGB in trees was fitted as follows: AGB = 0.0471 (ρD_b²H)^0.915 (with AGB in kg and ρD_b²H in g cm^–1 m). This model had a bias of 19%, while a reference model for comparison (fitted from tree measurements in a similar savanna ecosystem, Ifo et al. 2018) overestimated the AGB of trees of our test savannas by 132%. The BGB of trees was also better predicted from ρDb2H as follows: BGB = 0.0125 (ρD_b²H)^0.6899 (BGB in kg and ρD_b²H in g cm^–1 m), with 6% bias, while the reference model had about 3% bias. In shrubs, AGB and BGB were better predicted from ρD_b²H together with the total number of stems (n). The best fitted allometric equation for predicting AGB in shrubs was as follows: AGB = 0.0191 (ρD_b²H)^0.6227 n^0.9271. This model had about 1.5% bias, while the reference model overestimated the AGB of shrubs of Lamto savannas by about 79%. The equation for predicting BGB of shrubs is: BGB = 0.0228 (ρD_b²H)^0.7205 n^0.992 that overestimated the BGB of the shrubs of Lamto savannas with about 3% bias, while the reference model underestimated the BGB by about 14%. The reference model misses an important feature of fire-prone savannas, namely the strong imbalance of the BGB/AGB ratio between trees and multi-stemmed shrubs, which our models predict. The allometric equations we developed here are therefore relevant for C stocks inventories in trees and shrubs communities of Guinean savannas.

• Kouamé, UFR Sciences de la Nature, UFR-SN/ Station d’Ecologie de Lamto (CRE), Pôle de recherche Environnement et Développement Durable, Université NANGUI ABROGOUA, 02 BP 801 Abidjan 02, Côte d’Ivoire); Institute of Ecology and Environmental Sciences IEES-Paris (Sorbonne Université, CNRS, Université Paris Diderot, IRD, UPEC, INRA), 4 Place Jussieu, 75005, Paris, France https://orcid.org/0000-0002-0847-2569 E-mail: kouameyag@gmail.com
• Millan, Centre for African Ecology, School of Animal, Plant and Environmental Sciences, University of the Witwatersrand, Private Bag 3, Johannesburg, South Africa; Global Change Biology Group, Department of Botany and Zoology, Stellenbosch University, Private Bag X1, Matieland 7602, South Africa; Institute of Botany of the Czech Academy of Sciences, v.v.i, Dukelská 135, Třeboň, 379 01, Czech Republic https://orcid.org/0000-0002-0151-6055 E-mail: mathieu.millan@gmail.com
• N'Dri, UFR Sciences de la Nature, UFR-SN/ Station d’Ecologie de Lamto (CRE), Pôle de recherche Environnement et Développement Durable, Université NANGUI ABROGOUA, 02 BP 801 Abidjan 02, Côte d’Ivoire https://orcid.org/0000-0002-6333-6279 E-mail: brigitte.aya@gmail.com
• Charles-Dominique, Institute of Ecology and Environmental Sciences IEES-Paris (Sorbonne Université, CNRS, Université Paris Diderot, IRD, UPEC, INRA), 4 Place Jussieu, 75005, Paris, France https://orcid.org/0000-0002-5767-0406 E-mail: tristan.charles-dominique@sorbonne-universite.fr
• Konan, UFR Sciences de la Nature, UFR-SN/ Station d’Ecologie de Lamto (CRE), Pôle de recherche Environnement et Développement Durable, Université NANGUI ABROGOUA, 02 BP 801 Abidjan 02, Côte d’Ivoire E-mail: marcelkonan.lamto@gmail.com
• Bakayoko, UFR Sciences de la Nature, UFR-SN/ Station d’Ecologie de Lamto (CRE), Pôle de recherche Environnement et Développement Durable, Université NANGUI ABROGOUA, 02 BP 801 Abidjan 02, Côte d’Ivoire E-mail: bakadamaci@yahoo.fr
• Gignoux, Institute of Ecology and Environmental Sciences IEES-Paris (Sorbonne Université, CNRS, Université Paris Diderot, IRD, UPEC, INRA), 4 Place Jussieu, 75005, Paris, France https://orcid.org/0000-0003-3853-9282 E-mail: jacques.gignoux@upmc.fr

Terrestrial laser scanning (TLS) has been applied to estimate forest wood volume based on detailed 3D tree reconstructions from point cloud data. However, sources of uncertainties in the point cloud data (alignment and scattering errors, occlusion, foliage...) and the reconstruction algorithm type and parameterisation are known to affect the reconstruction, especially around finer branches. To better understand the impacts of these uncertainties on the accuracy of TLS-derived woody volume, high-quality TLS scans were collected in leaf-off conditions prior to destructive harvesting of two forest-grown common ash trees (Fraxinus excelsior L.; diameter at breast height ~28 cm, woody volume of 732 and 868 L). We manually measured branch diameters at 265 locations in these trees. Estimates of branch diameters and tree volume from Quantitative Structure Models (QSM) were compared with these manual measurements. The accuracy of QSM branch diameter estimates decreased with smaller branch diameters. Tree woody volume was overestimated (+336 L and +392 L) in both trees. Branches measuring < 5 cm in diameter accounted for 80% and 83% of this overestimation respectively. Filtering for scattering errors or improved coregistration approximately halved the overestimation. Range filtering and modified scanning layouts had mixed effects. The small branch overestimations originated primarily in limitations in scanner characteristics and coregistration errors rather than suboptimal QSM parameterisation. For TLS-derived estimates of tree volume, a higher quality point cloud allows smaller branches to be accurately reconstructed. Additional experiments need to elucidate if these results can be generalised beyond the setup of this study.

• Demol, CAVElab – Computational and Applied Vegetation Ecology, Department of Environment, Faculty of Bioscience Engineering, Ghent University, Coupure Links 653, B-9000 Ghent, Belgium; PLECO – Plants and Ecosystems, Faculty of Science, Antwerp University, Universiteitsplein 1, B-2610 Wilrijk, Belgium https://orcid.org/0000-0002-5492-2874 E-mail: miro.demol@ugent.be
• Wilkes, UCL Department of Geography, Gower Street, London WC1E 6BT, UK; NERC National Centre for Earth Observation (NCEO), UK https://orcid.org/0000-0001-6048-536X E-mail: p.wilkes@ucl.ac.uk
• Raumonen, Mathematics, Tampere University, FI-33101 Tampere, Finland https://orcid.org/0000-0001-5471-0970 E-mail: pasi.raumonen@tuni.fi
• Krishna Moorthy, CAVElab – Computational and Applied Vegetation Ecology, Department of Environment, Faculty of Bioscience Engineering, Ghent University, Coupure Links 653, B-9000 Ghent, Belgium https://orcid.org/0000-0002-6838-2880 E-mail: Sruthi.KrishnaMoorthyParvathi@ugent.be
• Calders, CAVElab – Computational and Applied Vegetation Ecology, Department of Environment, Faculty of Bioscience Engineering, Ghent University, Coupure Links 653, B-9000 Ghent, Belgium https://orcid.org/0000-0002-4562-2538 E-mail: kim.calders@ugent.be
• Gielen, PLECO – Plants and Ecosystems, Faculty of Science, Antwerp University, Universiteitsplein 1, B-2610 Wilrijk, Belgium https://orcid.org/0000-0002-4890-3060 E-mail: bert.gielen@uantwerpen.be
• Verbeeck, CAVElab – Computational and Applied Vegetation Ecology, Department of Environment, Faculty of Bioscience Engineering, Ghent University, Coupure Links 653, B-9000 Ghent, Belgium https://orcid.org/0000-0003-1490-0168 E-mail: hans.verbeeck@ugent.be

Young, dense forest in Finland and Sweden urgently need to receive first thinning. In such stands, conventional selective thinning methods make the harvester work time consuming and, thus, costly. To make small-sized trees economically competitive as raw material for bioenergy and biorefining, new harvesting technologies and/or thinning methods need to be developed. A potential solution is boom-corridor thinning (BCT), rendering effective cutting work. The aim of this study was to describe and compare the stand structure of two Scots pine stands (Pinus sylvestris L.) and one birch-dominated (Betula pendula Roth with natural downy birch, B. pubescens Ehrh.) stand after BCT and selective thinning at the first thinning phase. Furthermore, simulations were conducted to predict the future stand development after the first thinning treatments. The density of the growing stock was 16–46% higher after BCT treatment than after selective thinning because BCT stands included more small and supressed trees with a dbh < 100 mm. However, the numbers of future crop trees with a dbh > 140 mm per hectare were at the same level in both treatments. The stem volume removal per hectare did not differ between treatments. However, simulation of stand development and intermediate thinning and clearcutting revealed that the total removal volume was 10–18% higher in BCT stands compared to selectively thinned ones. The saw log volumes harvested did, however, not differ between treatments. This study shows that BCT generates stands with higher biodiversity compared to conventional thinning as higher levels of biomass removal can be reached throughout stand rotations.

• Nuutinen, Natural Resources Institute Finland (Luke), Production systems, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: yrjo.nuutinen@luke.fi
• Miina, Natural Resources Institute Finland (Luke), Natural resources, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jari.miina@luke.fi
• Saksa, Natural Resources Institute Finland (Luke), Natural resources, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@luke.fi
• Bergström, Swedish University of Agricultural Sciences (SLU), Dept of Forest Biomaterials and Technology, Section of Forest Operations, SE-90183 Umeå, Sweden E-mail: dan.bergstrom@slu.se
• Routa, Natural Resources Institute Finland (Luke), Production systems, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: johanna.routa@luke.fi

Accurate estimates of aboveground biomass (AGB) strongly depend on the suitability and precision of allometric models. Diospyros mespiliformis Hochst. ex A. DC. is a key component of most sub-Sahara agroforestry systems and, one of the most economically important trees in Africa. Despite its importance, very few scientific information exists regarding its biomass and carbon storage potential. In this study direct method was used to develop site-specific biomass models for D. mespiliformis tree components in Burkina Faso. Allometric models were developed for stem, branch and leaf biomass using data from 39 tree harvested in Sudanian savannas of Burkina Faso. Diameter at breast height (DBH), tree height, crown diameter (CD) and basal diameter (D₂₀) were regressed on biomass component using non-linear models with DBH alone, and DBH in combination with height and/or CD as predictor variables. Carbon content was estimated for each tree component using the ash method. Allometric models differed between the experimental sites, except for branch biomass models. Site-specific models developed in this study exhibited good model fit and performance, with explained variance of 81–98%. Using models developed from other areas would have underestimated or overestimated biomass by between –72% and +98%. Carbon content in aboveground components of D. mespiliformis in Tiogo, Boulon and Tapoa-Boopo was 55.40% ± 1.50, 55.52% ± 1.06 and 55.63% ± 1.00, respectively, and did not vary significantly (P-value = 0.909). Site-specific models developed in this study are useful tool for estimating carbon stocks and can be used to accurately estimate tree components biomass in vegetation growing under similar conditions.

• Ouédraogo, University Joseph Ki-Zerbo, UFR/SVT, Laboratory of Plant Biology and Ecology, 03 B.P. 7021 Ouagadougou 03, Burkina Faso E-mail: okorotimi@yahoo.fr
• Dimobe, University Joseph Ki-Zerbo, UFR/SVT, Laboratory of Plant Biology and Ecology, 03 B.P. 7021 Ouagadougou 03, Burkina Faso; University of Dédougou, Institut des Sciences de l’Environnement et du Développement Rural (ISEDR), BP 139 Dédougou, Burkina Faso; West African Science Service Center on Climate Change and Adapted Land Use, Competence Center, Avenue Muamar Ghadhafi, Ouagadougou, BP 9507, Burkina Faso https://orcid.org/0000-0001-5536-9700 E-mail: kangbenidimobe@gmail.com
• Thiombiano, University Joseph Ki-Zerbo, UFR/SVT, Laboratory of Plant Biology and Ecology, 03 B.P. 7021 Ouagadougou 03, Burkina Faso E-mail: adjima_thiombiano@yahoo.fr

The paper deals with production of above-ground biomass of silver birch (Betula pendula Roth) stands in the Czech Republic. One-year biomass dynamics was studied within chronosequence of birch stands at the age of 4–5, 8–9, 17–18 and 22–23 years. With the exception of the youngest stand, which was established by seeding, all experimental birch stands were regenerated naturally after the allochthonous spruce stands. Above-ground biomass (AB) was calculated from plot inventory data and biomass equations were parameterized from destructive sampling of biomass component of sampled trees. Results reveal that the peak of the mean annual increment (MAI_ABtotal) of birch stands can be expected at the age from 15 to 20 years. Additionally, the stand age, the value of basal area (BA) should be considered as a predictor of stand productivity. If the value of BA varied from 25 to 35 m² ha^–1, the MAI of the birch stands reached the range from 5.0 to 6.5 t of dry biomass per ha y^–1 at the age ranging between 15 and 25 years. The stem/branch proportion increased with stand age, the stem relative proportion ranging from 75 to 90% of total above-ground biomass. According to the results of this study, birch stand biomass production and utilization is one of the approaches in terms of forest recovery management in large disturbed areas. Although, no silvicultural treatments were occurred in all analysed stands, the pre-commercial thinning method could increase stand productivity and stability as well.

• Martiník, Department of Silviculture, Faculty of Forestry and Wood Technology, Mendel University in Brno, 613 00 Brno, Czech Republic https://orcid.org/0000-0002-5906-8830 E-mail: martinik@mendelu.cz
• Knott, Department of Silviculture, Faculty of Forestry and Wood Technology, Mendel University in Brno, 613 00 Brno, Czech Republic E-mail: robert.knott@mendelu.cz
• Krejza, Global Change Research Institute CAS, v.v.i., Bělidla 4a, 603 00 Brno, Czech Republic E-mail: krejza.j@czechglobe.cz
• Černý, The Forestry and Game Management Research Institute, Research Station at Opočno, Na Olivě 550, 517 73 Opočno, Czech Republic E-mail: cerny@vulhmop.cz

Industrial chipping is becoming increasingly popular, as the result of a growing demand for woody biomass. Industrial chippers are large, powerful machines that generate much noise and vibration. This study explored some factors that may affect exposure to noise and vibration, namely: feedstock type (branches vs. logs), work station characteristics (truck cab vs. separate cab) and knife wear (new knives vs. blunt knives). Exposure to noise was significantly affected by all three factors, and it was higher for branch feedstock, separate cabs and blunt knives. The higher exposure levels recorded for the separate cab were especially insidious, because they were below and above the hearing threshold and would elude immediate perception. Exposure to whole-body vibration (WBV) was significantly higher for branch feedstock and for the separate cab. Knife wear seemed to determine an increase in WBV, but this effect had no statistical significance and the result could not be taken as conclusive. Among the three factors studied, work station characteristics had the strongest effect. Further studies may extend the comparison to a wider range of options, as well as explore the use of exposure variation for machine diagnostics.

• Anton, University of Ljubljana, Dept. of Forestry and Renewable Resources, Večna pot 83, 1000 Ljubljana, Slovenia E-mail: anton.poje@bf.uni-lj.si
• Spinelli, CNR IVALSA, Via Madonna del Piano 10, I-50019 Sesto Fiorentino, Italy; AFORA, University of the Sunshine Coast, Locked Bag 4, Maroochydore DC, Queensland, 4558 Australia http://orcid.org/0000-0001-9545-1004 E-mail: spinelli@ivalsa.cnr.it
• Magagnotti, CNR IVALSA, Via Madonna del Piano 10, I-50019 Sesto Fiorentino, Italy; AFORA, University of the Sunshine Coast, Locked Bag 4, Maroochydore DC, Queensland, 4558 Australia E-mail: magagnotti@ivalsa.cnr.it
• Mihelic, University of Ljubljana, Dept. of Forestry and Renewable Resources, Večna pot 83, 1000 Ljubljana, Slovenia E-mail: matevz.mihelic@bf.uni-lj.si

Accurate estimates of both above-ground biomass (AGB) and below-ground biomass (BGB) are essential for estimating carbon (C) balances at various geographical scales and formulating effective climate change mitigation programs. However, estimating BGB is challenging, particularly for forest ecosystems, so robust allometric equations are needed. To obtain such equations for savanna-woodlands of the West African north sudanian zone, we selected four common native woody species (Anogeissus leiocarpa (DC.) Guill. & Perr., Detarium microcarpum Guill. & Perr., Piliostigma thonningii (Schumach.) Milne-Redh. and Vitellaria paradoxa C.F. Gaertn.). At two sites in Burkina Faso, we determined the BGB of 30 trees of each of these species by excavation, and measured various above-ground dimensional variables. The root:shoot ratio varied widely among the species, from 0.1 to 3.4. Depending on the species, allometric equations based on stem basal area at 20 cm height, basal area at breast height and tree height explained 50–95% of the variation in BGB. The best generic equation we obtained, based on basal area at 20 cm, explained 60% of the variation in BGB across the species. Three previously published generic allometric equations underestimated BGB by 8 to 63%. The presented equations should significantly improve the accuracy of BGB estimates in savanna-woodlands and help avoid costly needs to excavate root systems.

• Koala, Centre National de Recherche Scientifique et Technologique (CNRST), Institut de l’Environnement et de Recherches Agricoles (INERA), Département Productions Forestières, 03 BP 7047, Ouagadougou 03, Burkina Faso E-mail: ezeyamb@yahoo.fr
• Sawadogo, Centre National de Recherche Scientifique et Technologique (CNRST), Institut de l’Environnement et de Recherches Agricoles (INERA), Département Productions Forestières, 03 BP 7047, Ouagadougou 03, Burkina Faso E-mail: sawadogo_ls@hotmail.com
• Savadogo, World Agroforestry Centre & International Crop Research Institute for the Semi-Arid Tropics (ICRAF-ICRISAT), West and Central Africa Region-Sahel Node, BP 12404, Niamey, Niger E-mail: savadogo.patrice@gmail.com
• Aynekulu, World Agroforestry Centre (ICRAF), United Nations Avenue, P.O. Box 30677-00100, Nairobi, Kenya E-mail: e.betemariam@cgiar.org
• Heiskanen, University of Helsinki, Department of Geosciences and Geography, P.O. Box 68, FI-00014 University of Helsinki, Finland E-mail: janne.heiskanen@helsinki.fi
• Saïd, International Livestock Research Institute (ILRI). P.O. Box 30709, Nairobi, Kenya E-mail: m.said@cgiar.org

Competitive interactions in clonal forestry are not well understood and this needs to be addressed to develop better deployment strategies. Eight juvenile Sitka spruce (Picea sitchensis (Bong.) Carriére) clones were grown in monoclonal and clonal mixtures in a field experiment for three years to assess the effects of genetic diversity on shoot growth, above- and below-ground biomass partitioning and crown characteristics. Shoot elongation was measured throughout the growing season, while diameter was measured twice annually in May and December. After the third year, crown silhouette area was estimated from digitised images for one ramet per plot and ramets were then destructively harvested. Deployment × clone interactions were observed for tree height and diameter with reductions observed in mixed plots. Mixed plots had significantly greater height and diameter heterogeneity and more asymmetrical competition than monoclonal plots. Results from this study demonstrate that stem growth can be significantly altered when clones are planted in multi-clonal mixtures but for most clones, deployment-type will not significantly reduce their productivity.

• Donnelly, UCD Forestry, University College Dublin, Belfield, Dublin 4, Ireland E-mail: liam.donnelly@ucdconnect.ie
• Grant, UCD Forestry, University College Dublin, Belfield, Dublin 4, Ireland E-mail: olga.grant@ucd.ie
• O’Reilly, UCD Forestry, University College Dublin, Belfield, Dublin 4, Ireland E-mail: conor.oreilly@ucd.ie

In Finland the increasing use of biofuel in transport presupposes a search for new raw material sources for biorefining. The aim of this study was, at the regional level, to compare the procurement costs of logging residues, stumps, delimbed stems and cereal straw for biorefining. The accumulation and procurement costs of forest chips and cereal straw were estimated within a 100-kilometre transporting distance via existing road network from an end-use-facility located in Kouvola in South-East Finland. The analyses were performed as simulated treatments in thinnings of young stands, cereal fields and regeneration fellings using existing productivity and cost functions and yield calculations based on crop statistics, the forest industry stand data and the sample plots data of the National Forest Inventory of Finland. Accumulation of raw material assortments and costs of production stages were defined per dry tonnes. Subsidies and raw material prices were excluded from consideration in the study. The results indicate that recovering logging residues requires lower costs than utilization of stumps, delimbed stems or cereal straw. Cereal straw represents an important source of biomass in Kouvola but the cost competiveness is poor compared the procurement costs of forest chips. When the annual procurement volume of biomass was 50 000 dry tonnes the cost at the plant was 49 € dry tonne ^–1 and biomass was comprised totally of logging residues. Procurement cost grew to 59 € dry tonne ^–1 when the annual procurement volume of biomass was doubled to 100 000 dry tonnes. Of that amount, the proportion of logging residues was 58.4%, stumps 24.3% and delimbed stems 17.3%. First cereal straw dry tonnes were delivered to end-use-facility, when procurement cost grew to 60 € dry tonne ^–1 and annual procurement volume of biomass was 110 000 dry tonnes.

• Laitila, Natural Resources Institute Finland (Luke), Bio-based Business and Industry, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: juha.laitila@luke.fi
• Lehtonen, Natural Resources Institute Finland (Luke), Green Technology, Halolantie 31A, FI-71750 Maaninka, Finland E-mail: eeva.lehtonen@luke.fi
• Ranta, Lappeenranta University of Technology, LUT School of Energy Systems, Laboratory of Bioenergy, Lönnrotinkatu 7, FI-50100 Mikkeli, Finland E-mail: tapio.ranta@luke.fi
• Anttila, Natural Resources Institute Finland (Luke), Bio-based Business and Industry, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: perttu.anttila@luke.fi
• Rasi, Natural Resources Institute Finland (Luke), Bio-based Business and Industry, Survontie 9A, FI-40500 Jyväskylä E-mail: saija.rasi@luke.fi
• Asikainen, Natural Resources Institute Finland (Luke), Bio-based Business and Industry, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: antti.asikainen@luke.fi

Silver birch (Betula pendula Roth.) is one of the main pioneer tree species occupying large areas of abandoned agricultural lands under natural succession in Estonia. We estimated aboveground biomass (AGB) dynamics during 17 growing seasons, and analysed soil nitrogen (N) and carbon (C) dynamics for 10 year period in a silver birch stand growing on former arable land. Main N fluxes were estimated and nitrogen budget for 10-year-old stand was compiled. The leafless AGB and stem mass of the stand at the age of 17-years were 94 and 76 Mg ha^–1 respectively. The current annual increment (CAI) of stemwood fluctuated, peaking at 10 Mg ha^–1 yr^–1 at the age of 15 years; the mean annual increment (MAI) fluctuated at around 4–5 Mg ha^–1. The annual leaf mass of the stand stabilised at around 3 Mg ha^–1 yr^–1. The stand density decreased from 11600 to 2700 trees ha^–1 in the 8- and 17-year-old stand, respectively. The largest fluxes in N budget were net nitrogen mineralization and gaseous N₂-N emission. The estimated fluxes of N₂O and N₂ were 0.12 and 83 kg ha^–1 yr^–1, respectively; N leaching was negligible. Nitrogen retranslocation from senescing leaves was approximately 45 kg ha^–1, N was mainly retranslocated into stembark. The N content in the upper 0–10 cm soil layer increased significantly (145 kg ha^–1) from 2004 to 2014; soil C content remained stable. Both the woody biomass dynamics and the N cycling of the stand witness the potential for bioenergetics of such ecosystems.

• Aosaar, Estonian University of Life Sciences, Institute of Forestry and Rural Engineering, Kreutzwaldi 1, 51014 Tartu, Estonia E-mail: jyrgen.aosaar@emu.ee
• Mander, University of Tartu, Institute of Ecology and Earth Sciences, Ülikooli 18, 50090 Tartu, Estonia E-mail: ulo.mander@ut.ee
• Varik, Estonian University of Life Sciences, Institute of Forestry and Rural Engineering, Kreutzwaldi 1, 51014 Tartu, Estonia E-mail: mats.varik@emu.ee
• Becker, Estonian University of Life Sciences, Institute of Forestry and Rural Engineering, Kreutzwaldi 1, 51014 Tartu, Estonia E-mail: hardo.becker@emu.ee
• Morozov, Estonian University of Life Sciences, Institute of Forestry and Rural Engineering, Kreutzwaldi 1, 51014 Tartu, Estonia E-mail: gunnar.morozov@emu.ee
• Maddison, University of Tartu, Institute of Ecology and Earth Sciences, Ülikooli 18, 50090 Tartu, Estonia E-mail: martin.maddison@ut.ee
• Uri, Estonian University of Life Sciences, Institute of Forestry and Rural Engineering, Kreutzwaldi 1, 51014 Tartu, Estonia E-mail: veiko.uri@emu.ee

The biomass of a secondary evergreen and deciduous broad-leaved mixed forest was comprehensively inventoried in a permanent 2 ha plot in southwestern China. Biomass models, sub-sampling, soil pit method, and published data were utilized to determine the biomass of all components. Results showed that the total biomass of the forest was 158.1 Mg ha⁻¹; the total biomass included the major aboveground (137.7 Mg ha⁻¹) and belowground (20.3 Mg ha⁻¹) biomass components of vascular plants as well as the minor biomass components of bryophytes (0.078 Mg ha⁻¹) and lichens (0.043 Mg ha⁻¹). The necromass was 17.6 Mg ha⁻¹ and included woody debris (9.0 Mg ha⁻¹) and litter (8.6 Mg ha⁻¹). The spatial pattern of the aboveground biomass was determined by the spatial distribution of dominant trees with large diameter, tall height, and dense wood. The belowground biomass differed in terms of root diameter and decreased with increasing soil depth. The belowground biomass in each soil pit in local habitats was not related to the spatial distribution of woody plants and soil pit depth. The karst forest presented lower biomass compared than the nonkarst forests in the subtropical zone. Biomass carbon in the karst terrains would increase substantially if degraded karst vegetation could be successfully restored to the forest. Comprehensive site-based biomass inventory of karst vegetation will contribute not only to provide data for benchmarking global and regional vegetation and carbon models but also for regional carbon inventory and vegetation restoration.

• Liu, State Key Laboratory of Environmental Geochemistry, Institute of Geochemistry, Chinese Academy of Sciences, Lincheng West Road 99, 550081 Guiyang, China; Puding Karst Ecosystem Research Station, Chinese Academy of Sciences, 562100 Puding, China; University of Chinese Academy of Sciences, 100049 Beijing, China E-mail: liulibin@mail.gyig.ac.cn
• Wu, State Key Laboratory of Environmental Geochemistry, Institute of Geochemistry, Chinese Academy of Sciences, Lincheng West Road 99, 550081 Guiyang, China; Puding Karst Ecosystem Research Station, Chinese Academy of Sciences, 562100 Puding, China; University of Chinese Academy of Sciences, 100049 Beijing, China E-mail: wuyang2468@hotmail.com
• Hu, School of Chemistry and Life Science, Guangxi Teachers Education University, 530001 Nanning, China; Key Laboratory of Beibu Gulf Environment Change and Resources Utilization of Ministry of Education, Guangxi Teachers Education University, 530001 Nanning, China E-mail: ahhugang@gmail.com
• Zhang, School of Chemistry and Life Science, Guangxi Teachers Education University, 530001 Nanning, China; Key Laboratory of Beibu Gulf Environment Change and Resources Utilization of Ministry of Education, Guangxi Teachers Education University, 530001 Nanning, China E-mail: gxtczzh@gmail.com
• Cheng, State Key Laboratory of Environmental Geochemistry, Institute of Geochemistry, Chinese Academy of Sciences, Lincheng West Road 99, 550081 Guiyang, China; Puding Karst Ecosystem Research Station, Chinese Academy of Sciences, 562100 Puding, China E-mail: chenganyun@vip.skleg.cn
• Wang, State Key Laboratory of Environmental Geochemistry, Institute of Geochemistry, Chinese Academy of Sciences, Lincheng West Road 99, 550081 Guiyang, China; Puding Karst Ecosystem Research Station, Chinese Academy of Sciences, 562100 Puding, China E-mail: wangshijie@vip.skleg.cn
• Ni, State Key Laboratory of Environmental Geochemistry, Institute of Geochemistry, Chinese Academy of Sciences, Lincheng West Road 99, 550081 Guiyang, China; Puding Karst Ecosystem Research Station, Chinese Academy of Sciences, 562100 Puding, China E-mail: nijian@vip.skleg.cn

Understanding the characteristics of unutilized biomass resources, such as small-diameter trees from biomass-dense thinning forests (BDTF) (non-commercially-thinned forests), can provide important information for developing a bio-based economy. The aim of this study was to describe the areal distribution, characteristics (biomass of growing stock, tree height, etc.) and harvesting potential of BDTF in Sweden. A national forest inventory plot dataset was imported into a geographical information system and plots containing BDTF were selected by applying increasingly stringent constraints. Results show that, depending on the constraints applied, BDTF covers 9–44% (2.1–9.8 M ha) of the productive forest land area, and contains 7–34% of the total growing stock (119–564 M OD t), with an average biomass density of 57 OD t ha^–1. Of the total BDTF area, 65% is located in northern Sweden and 2% corresponds to set-aside farmlands. Comparisons with a study from 2008 indicate that BDTF area has increased by at least 4% (about 102 000 ha), in line with general trends for Sweden and Europe. Analyses revealed that the technical harvesting potential of delimbed stemwood (over bark, including tops) from BDTF ranges from 3.0 to 6.1 M OD t yr^–1 (7.5 to 15.1 M m³ yr^–1), while the potential of whole-tree harvesting ranges from 4.3 to 8.7 M OD t yr^–1 (10.2 to 20.6 M m³ yr^–1) depending on the scenario considered. However, further technological developments of the harvest and supply systems are needed to utilize the full potential of BDTF.

• Fernandez-Lacruz, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden http://orcid.org/0000-0001-9284-8911 E-mail: raul.fernandez@slu.se
• Di Fulvio, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden; International Institute for Applied Systems Analysis (IIASA), Ecosystems Services and Management Program (ESM), Schlossplatz 1, A-2361 Laxenburg, Austria E-mail: Fulvio.di.Fulvio@slu.se
• Athanassiadis, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: Dimitris.Athanassiadis@slu.se
• Bergström, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: Dan.Bergstrom@slu.se
• Nordfjell, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: Tomas.Nordfjell@slu.se

• Hytönen, Natural Resources Institute Finland, Silmäjärventie 2, FI-69100 Kannus, Finland E-mail: jyrki.hytonen@luke.fi
• Saarsalmi, Natural Resources Institute Finland, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: anna.saarsalmi@luke.fi

• Wolfsmayr, University of Natural Resources and Life Sciences, Vienna, Gregor Mendel Straße 33, A-1180 Wien, Austria E-mail: ulrich.wolfsmayr@boku.ac.at
• Rauch, University of Natural Resources and Life Sciences, Vienna, Gregor Mendel Straße 33, A-1180 Wien, Austria E-mail: peter.rauch@boku.ac.at

• Repola, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: jaakko.repola@metla.fi
• Ahnlund Ulvcrona, SLU, Forest Biomaterials and Technology, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: kristina.ulvcrona@slu.se

• Akujärvi, Finnish Environment Institute, Helsinki, Finland E-mail: anu.akujarvi@ymparisto.fi
• Hallikainen, Finnish Forest Research Institute, Northern Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: ville.hallikainen@metla.fi
• Hyppönen, Finnish Forest Research Institute, Northern Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: mikko.hypponen@metla.fi
• Mattila, Finnish Forest Research Institute, Northern Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: eero.mattila@metla.fi
• Mikkola, Finnish Forest Research Institute, Northern Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: kari.mikkola@metla.fi
• Rautio, Finnish Forest Research Institute, Northern Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: pasi.rautio@metla.fi

• Spinelli, CNR IVALSA,Via Madonna del Piano 10, Sesto Fiorentino (FI), Italy E-mail: spinelli@ivalsa.cnr.it
• Lombardini, CNR IVALSA,Via Madonna del Piano 10, Sesto Fiorentino (FI), Italy E-mail: lombardini@ivalsa.cnr.it
• Magagnotti, CNR IVALSA,Via Biasi 75, S. Michele all’Adige (TN), Italy E-mail: magagnotti@ivalsa.cnr.it

• Zamora-Cristales, Department of Forest Engineering, Resources, and Management, College of Forestry, Oregon State University, 280 Peavy Hall, Corvallis, OR 97331, USA E-mail: rene.zamora@oregonstate.edu
• Boston, Department of Forest Engineering, Resources, and Management, College of Forestry, Oregon State University, 280 Peavy Hall, Corvallis, OR 97331, USA E-mail: kevin.boston@oregonstate.edu
• Sessions, Department of Forest Engineering, Resources, and Management, College of Forestry, Oregon State University, 280 Peavy Hall, Corvallis, OR 97331, USA E-mail: john.sessions@oregonstate.edu
• Murphy, Waiariki Institute of Technology, Rotorua, New Zealand E-mail: glen.murphy@waiariki.ac.nz

• Muinonen, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: eero.muinonen@metla.fi
• Anttila, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: perttu.anttila@metla.fi
• Heinonen, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jaakko.heinonen@metla.fi
• Mustonen, Stora Enso, Talvikkitie 40 C, FI-01300 Vantaa, Finland E-mail: jukka.mustonen@storaenso.com

• Huttunen, Section of Ecology, Department of Biology, University of Turku, FI-20014 Turku, Finland E-mail: liisa.huttunen@utu.fi
• Ayres, Biological Sciences, Dartmouth College, Hanover, NH 03755, USA E-mail: matt.ayres@dartmouth.edu
• Niemelä, Section of Biodiversity and Environmental Science, Department of Biology, University of Turku, FI-20014 Turku, Finland E-mail: pekka.niemela@utu.fi
• Heiska, The Finnish Forest Research Institute, Punkaharju Unit, Finlandiantie 18, FI-58450 Punkaharju, Finland E-mail: susanne.heiska@metla.fi
• Tegelberg, Digitarium - Digitization Centre of the Finnish Museum of Natural History and the University of Eastern Finland, Joensuu Science Park, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: riitta.tegelberg@helsinki.fi
• Rousi, The Finnish Forest Research Institute, Vantaa Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: matti.rousi@metla.fi
• Kellomäki, Faculty of Science and Forestry, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: seppo.kellomaki@uef.fi

• Bijak, Warsaw University of Life Sciences – SGGW, Faculty of Forestry, Laboratory of Dendrometry and Forest Productivity, Nowoursynowska 159, 02-776 Warszawa, Poland E-mail: szymon.bijak@wl.sggw.pl
• Zasada, Warsaw University of Life Sciences – SGGW, Faculty of Forestry, Laboratory of Dendrometry and Forest Productivity, Nowoursynowska 159, 02-767 Warszawa, Poland E-mail: michal.zasada@wl.sggw.pl
• Bronisz, Warsaw University of Life Sciences – SGGW, Faculty of Forestry, Laboratory of Dendrometry and Forest Productivity, Warszawa, Poland E-mail: agnieszka.bronisz@wl.sggw.pl
• Bronisz, Warsaw University of Life Sciences – SGGW, Faculty of Forestry, Laboratory of Dendrometry and Forest Productivity, Warszawa, Poland E-mail: karol.bronisz@wl.sggw.pl
• Czajkowski, Warsaw University of Life Sciences – SGGW, Faculty of Forestry, Laboratory of Dendrometry and Forest Productivity, Warszawa, Poland E-mail: maciej.czajkowski@wl.sggw.pl
• Ludwisiak, Warsaw University of Life Sciences – SGGW, Faculty of Forestry, Laboratory of Dendrometry and Forest Productivity, Warszawa, Poland E-mail: lukasz.ludwisiak@wl.sggw.pl
• Tomusiak, Warsaw University of Life Sciences – SGGW, Faculty of Forestry, Laboratory of Dendrometry and Forest Productivity, Warszawa, Poland E-mail: robert.tomusiak@wl.sggw.pl
• Wojtan, Warsaw University of Life Sciences – SGGW, Faculty of Forestry, Laboratory of Dendrometry and Forest Productivity, Warszawa, Poland E-mail: rafal.wojtan@wl.sggw.pl

• Omoro, Viikki Tropical Resources Institute, Department of Forest Sciences, P.O. Box 27 (Latokartanonkaari 7), FI-00014 University of Helsinki, Finland E-mail: loice.omoro@helsinki.fi
• Starr, Department of Forest Sciences, P. O. Box 27 (Latokartanonkaari 7), FI-00014 University of Helsinki, Finland E-mail: mike.starr@helsinki.fi
• Pellikka, Department of Geosciences and Geography, P. O. Box 64 (Gustaf Hällströminkatu 2), FI-00014 University of Helsinki, Finland E-mail: petri.pellikka@helsinki.fi

• Spinelli, CNR IVALSA, Via Madonna del Piano 10, Sesto Fiorentino (FI), Italy E-mail: spinelli@ivalsa.cnr.it
• Cavallo, CNR IVALSA, Via Madonna del Piano 10, Sesto Fiorentino (FI), Italy E-mail: e.cavallo@imamoter.cnr.it
• Eliasson, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: lars.eliasson@skogforsk.se
• Facello, CNR IVALSA, Via Madonna del Piano 10, Sesto Fiorentino (FI), Italy E-mail: a.facello@ima.to.cnr.it

• Tuominen, Finnish Forest Research Institute, P.O.Box 18, FI-01301 Vantaa, Finland E-mail: sakari.tuominen@metla.fi
• Haapanen, Finnish Forest Research Institute, P.O.Box 18, FI-01301 Vantaa, Finland E-mail: reija.haapanen@gmail.com

• Hytönen, Finnish Forest Research Institute, Kannus, Finland E-mail: jyrki.hytonen@metla.fi
• Aro, Finnish Forest Research Institute, Kannus, Finland E-mail: lasse.aro@metla.fi

• Wang, Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing 100083, China E-mail: jw@nn.cn
• Zhang, Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing 100083, China E-mail: cz@nn.cn
• Xia, Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing 100083, China E-mail: fx@nn.cn
• Zhao, Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing 100083, China E-mail: zhaoxh@bjfu.edu.cn
• Wu, Rothamsted Research, Okehampton, Devon, UK E-mail: lw@nn.uk
• Gadow, Faculty of Forestry and Forest Ecology, Georg-August-University Göttingen, Göttingen, Germany E-mail: kvg@nn.de

• Spinelli, CNR IVALSA, Via Madonna del Piano 10, Sesto Fiorentino (FI), Italy E-mail: spinelli@ivalsa.cnr.it
• Magagnotti, CNR IVALSA, Via Biasi 75, S. Michele all’Adige (TN), Italy E-mail: nm@nn.it
• Paletto, CNR IMAMOTER, Strada delle Cacce 73, Torino, Italy E-mail: gp@nn.it
• Preti, CNR IMAMOTER, Strada delle Cacce 73, Torino, Italy E-mail: cp@nn.it

• Jacobson, Skogforsk, Uppsala Science Park, SE-75183 Uppsala, Sweden E-mail: staffan.jacobson@skogforsk.se
• Pettersson, Skogforsk, Uppsala Science Park, SE-75183 Uppsala, Sweden E-mail: fp@nn.se

• Laitila, The Finnish Forest Research Institute, Joensuu, Finland E-mail: juha.laitila@metla.fi
• Heikkilä, L&T Biowatti Oy, Seinäjoki, Finland E-mail: jh@nn.fi
• Anttila, The Finnish Forest Research Institute, Joensuu, Finland E-mail: pa@nn.fi

• Belbo, Norwegian Forest and Landscape Institute, Box 115, 1431 Ås, Norway E-mail: helmer.belbo@skogoglandskap.no

• Tuominen, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: sakari.tuominen@metla.fi
• Eerikäinen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: ke@nn.fi
• Schibalski, University of Potsdam, Karl-Liebknecht-Strasse 24–25, 14476 Potsdam, Germany E-mail: as@nn.de
• Haakana, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: mh@nn.fi
• Lehtonen, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: al@nn.fi

• Mola-Yudego, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: blas.mola@uef.fi

• Repola, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: jaakko.repola@metla.fi

• Qi, Ecole Centrale Paris, Laboratory of Applied Mathematics, Grande Voie des Vignes, 92295 Chatenay-Malabry, France; Institute of Automation, Chinese Academy of Sciences, LIAMA/NLPR, P.O.Box 2728, Beijing, China E-mail: qiruitree@gmail.com
• Letort, Ecole Centrale Paris, Laboratory of Applied Mathematics, Grande Voie des Vignes, 92295 Chatenay-Malabry, France E-mail: vl@nn.fr
• Kang, Institute of Automation, Chinese Academy of Sciences, LIAMA/NLPR, P.O.Box 2728, Beijing, China E-mail: mk@nn.cn
• Cournède, Ecole Centrale Paris, Laboratory of Applied Mathematics, Grande Voie des Vignes, 92295 Chatenay-Malabry, France; INRIA saclay Ile-de-France, EPI Digiplant, Parc Orsay Université, 91893 Orsay cedex, France E-mail: phc@nn.fr
• Reffye, INRIA saclay Ile-de-France, EPI Digiplant, Parc Orsay Université, 91893 Orsay cedex, France; CIRAD, UMR AMAP, Montpellier, F-34000 France E-mail: pdr@nn.fr
• Fourcaud, CIRAD, UMR AMAP, Montpellier, F-34000 France E-mail: tf@nn.fr

• Repola, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: jaakko.repola@metla.fi

• Kindermann, International Institute for Applied Systems Analysis, Laxenburg, Austria E-mail: kinder@iiasa.ac.at
• McCallum, International Institute for Applied Systems Analysis, Laxenburg, Austria E-mail: im@nn.at
• Fritz, International Institute for Applied Systems Analysis, Laxenburg, Austria E-mail: sf@nn.at
• Obersteiner, International Institute for Applied Systems Analysis, Laxenburg, Austria E-mail: mo@nn.at

• Mattson, Swedish University of Agricultural Sciences, Dept of Forest Ecology and Management, SE-901 83 Umeå, Sweden E-mail: sm@nn.se
• Bergsten, Swedish University of Agricultural Sciences, Dept of Forest Ecology and Management, SE-901 83 Umeå, Sweden E-mail: ub@nn.se
• Mörling, Swedish University of Agricultural Sciences, Dept of Forest Ecology and Management, SE-901 83 Umeå, Sweden E-mail: tommy.morling@ssko.slu.se

• Eriksson, SLU, Dept of Bioenergy, P.O. Box 7061, SE-750 07 Uppsala, Sweden E-mail: ee@nn.se
• Johansson, SLU, Dept of Bioenergy, P.O. Box 7061, SE-750 07 Uppsala, Sweden E-mail: tj@nn.se

• Aphalo, University of Helsinki, Department of Biological and Environmental Sciences E-mail: pja@nn.fi
• Lahti, The Finnish Forest Research Institute E-mail: ml@nn.fi
• Lehto, University of Joensuu, Faculty of Forestry, Box 111, FI-80101 Joensuu, Finland E-mail: tarja.lehto@joensuu.fi
• Repo, The Finnish Forest Research Institute E-mail: tr@nn.fi
• Rummukainen, University of Joensuu, Faculty of Forestry, Box 111, FI-80101 Joensuu, Finland E-mail: ar@nn.fi
• Mannerkoski, University of Joensuu, Faculty of Forestry, Box 111, FI-80101 Joensuu, Finland E-mail: hm@nn.fi
• Finér, The Finnish Forest Research Institute E-mail: lf@nn.fi

• Muukkonen, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: petteri.muukkonen@metla.fi
• Mäkipää, Finnish Forest Research Institute, Unioninkatu 40 A, FI-00170 Helsinki, Finland E-mail: rm@nn.fi
• Laiho, Department of Forest Ecology, P.O. Box 24, FI-00014 University of Helsinki, Finland E-mail: rl@nn.fi
• Minkkinen, Department of Forest Ecology, P.O. Box 24, FI-00014 University of Helsinki, Finland E-mail: km@nn.fi
• Vasander, Department of Forest Ecology, P.O. Box 24, FI-00014 University of Helsinki, Finland E-mail: hv@nn.fi
• Finér, Finnish Forest Research Institute, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: lf@nn.fi

• Spinelli, CNR/IVALSA, Via Madonna del Piano - Palazzo F, I-50019 Sesto Fiorentino (FI), Italy E-mail: spinelli@ivalsa.cnr.it
• Nati, CNR/IVALSA, Via Madonna del Piano - Palazzo F, I-50019 Sesto Fiorentino (FI), Italy E-mail: cn@nn.it
• Magagnotti, CNR/IVALSA, Via Madonna del Piano - Palazzo F, I-50019 Sesto Fiorentino (FI), Italy E-mail: nm@nn.it

• Långström, SLU, Dept. of Entomology, P.O.Box 7044, S-750 07 Uppsala, Sweden. Fax +46 18 672 890 E-mail: bo.langstrom@entom.slu.se
• Hellqvist, SLU, Dept. of Entomology, P.O.Box 7044, S-750 07 Uppsala, Sweden. Fax +46 18 672 890 E-mail: ch@nn.se
• Cedervind, SLU, Dept. of Entomology, P.O.Box 7044, S-750 07 Uppsala, Sweden. Fax +46 18 672 890 E-mail: jc@nn.se

• Kauppi, International Institute for Applied Systems Analysis (IIASA), A-2361 Laxenburg, Austria. Present address: Environmental Science and Policy, Department of Limnology and Environmental Protection, PO Box 27, FIN-00014 University of Helsinki, Finland E-mail: pekka.kauppi@helsinki.fi

• Dean, CRC for Greenhouse Accounting, The Australian National University, GPO Box 475, Canberra, ACT 2601, Australia E-mail: cdean@rsbs.anu.edu.au

• Uri, Institute of Silviculture, Estonian Agricultural University, Kreutzwaldi 5, 51014 Tartu, Estonia E-mail: vuri@eau.ee
• Tullus, Institute of Silviculture, Estonian Agricultural University, Kreutzwaldi 5, 51014 Tartu, Estonia E-mail: ht@nn.ee
• Lõhmus, Institute of Geography, University of Tartu, Vanemuise 46, 51014 Tartu, Estonia E-mail: kl@nn.ee

• Grote, TU München, Chair of Forest Yield Science, Am Hochanger 13, D-85354 Freising, Germany E-mail: ruediger.grote@lrz.tu-muenchen.de

• Légaré, Université du Québec en Abitibi-Témiscamingue, Groupe de recherche en écologie forestière interuniversitaire, 445, boulevard de l'Université, Rouyn-Noranda, QC, Canada J9X 5E4 E-mail: sonia.legare@uqat.uquebec.ca
• Bergeron, NSERC-UQAT-UQAM, Industrial Chair in sustainable forest management, CP 8888, succ. Centre-Ville, Montréal, QC, Canada H3C 3P8 E-mail: yb@nn.ca
• Paré, Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, P.O. Box 3800, Sainte-Foy, QC, Canada G1V 4C7 E-mail: dp@nn.ca

• Johansson, Swedish University of Agricultural Sciences, Department of Forest Management and Products, P.O. Box 7060, S-750 07 Uppsala, Sweden E-mail: tord.johansson@sh.slu.se

• Henry, IRD, UMR Eco&Sols, Montpellier SupAgro, 2 place Viala, 34060 Montpellier, France; Di.S.A.F.Ri, Università degli Studi della Tuscia, Viterbo, Italy; and AgroParisTech-ENGREF, GEEFT, Montpellier, France E-mail: henry@unitus.it
• Picard, CIRAD, Montpellier, France E-mail: np@nn.fr
• Trotta, Di.S.A.F.Ri, Università degli Studi della Tuscia, Viterbo, Italy E-mail: ct@nn.it
• Manlay, IRD, UMR Eco&Sols, Montpellier SupAgro, 2 place Viala, 34060 Montpellier, France; and AgroParisTech-ENGREF, GEEFT, Montpellier, France E-mail: rjm@nn.fr
• Valentini, Di.S.A.F.Ri, Università degli Studi della Tuscia, Viterbo, Italy E-mail: rv@nn.it
• Bernoux, IRD, UMR Eco&Sols, Montpellier SupAgro, 2 place Viala, 34060 Montpellier, France E-mail: mb@nn.fr
• Saint-André, CIRAD, Montpellier, France; and INRA, UR1138, Biogeochimie des Ecosystèmes Forestiers, Champenoux, France E-mail: lsa@nn.fr

Long-term (47 years) effect of experimental whole tree harvesting (WTH) with a heavy soil scarification on ground cover vegetation was assessed in a dry nutrient-poor Scots pine (Pinus sylvestris L.) stand in Latvia. Neighbouring conventionally managed young (10 years) and mature (119 years) stands of the same type were used for comparison. Higher species richness was observed in the WTH stand compared to conventionally managed young and mature stands (24, 18 and 16 species, respectively), likely due to the profound disturbance. The Shannon diversity index was higher in the young than in the WTH and mature stands (2.36, 1.77 and 1.63, respectively); still, the composition and structure of ground cover vegetation in WTH was more similar to the mature stand. Nevertheless, the occurrence of oligotrophic species in the WTH stand suggested decreased soil nutrient content and potential development of different plant community. Hence, such method might be considered for restoration of oligotrophic stands. Nevertheless, the period of 47 years appeared sufficient for the ground cover vegetation to recover after the WTH.

• Jansons, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: aris.jansons@silava.lv
• Robalte, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: robalte.l@gmail.com
• Čakšs, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: chakijs95@gmail.com
• Matisons, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: roberts.matisons@silava.lv

Species-specific allometric equations for shrubs and small trees are relatively scarce, thus limiting the precise quantification of aboveground biomass (AGB) in both shrubby vegetation and forests. Fourteen shrub and small tree species in Eastern China were selected to develop species-specific and multispecies allometric biomass equations. Biometric variables, including the diameter of the longest stem (D), height (H), wet basic density (BD), and crown area and shape were measured for each individual plant. We measured the AGB through a non-destructive method, and validated these measurements using the dry mass of the sampled plant components. The AGB was related to biometric variables using regression analysis. The species-specific allometric models, with D and H as predictors (D-H models) accounted for 70% to 99% of the variation in the AGB of shrubs and small trees. A multispecies allometric D-H model accounted for 71% of the variation in the AGB. Although BD, as an additional predictor, improved the fit of most models, the D-H models were adequate for predicting the AGB for shrubs and small trees in subtropical China without BD data.

• Ali, School of Ecological and Environmental Sciences, East China Normal University, Shanghai 200241, China; Tiantong National Forest Ecosystem Observations and Research Station, Ningbo 315114, Zhejiang, China; Department of Environmental Sciences, Abdul Wali Khan University Mardan, 23200, KPK, Pakistan E-mail: arshadforester@gmail.com
• Xu, School of Ecological and Environmental Sciences, East China Normal University, Shanghai 200241, China; Tiantong National Forest Ecosystem Observations and Research Station, Ningbo 315114, Zhejiang, China E-mail: yumsh09@lzu.edu.cn
• Zhao, School of Ecological and Environmental Sciences, East China Normal University, Shanghai 200241, China; Tiantong National Forest Ecosystem Observations and Research Station, Ningbo 315114, Zhejiang, China E-mail: zhaoyantao1991@163.com
• Zhang, School of Ecological and Environmental Sciences, East China Normal University, Shanghai 200241, China; Tiantong National Forest Ecosystem Observations and Research Station, Ningbo 315114, Zhejiang, China E-mail: qingzq@yeah.net
• Zhou, School of Ecological and Environmental Sciences, East China Normal University, Shanghai 200241, China; Tiantong National Forest Ecosystem Observations and Research Station, Ningbo 315114, Zhejiang, China E-mail: 792920738@qq.com
• Yang, School of Ecological and Environmental Sciences, East China Normal University, Shanghai 200241, China; Tiantong National Forest Ecosystem Observations and Research Station, Ningbo 315114, Zhejiang, China E-mail: xjyangxd@sina.com
• Yan, School of Ecological and Environmental Sciences, East China Normal University, Shanghai 200241, China; Tiantong National Forest Ecosystem Observations and Research Station, Ningbo 315114, Zhejiang, China E-mail: eryan@des.ecnu.edu.cn

• Ahnlund Ulvcrona, SLU, Forest Biomaterials and Technology, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: kristina.ulvcrona@slu.se
• Karlsson, SLU, Forest Biomaterials and Technology, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: lars.karlsson@slu.se
• Backlund, SLU, Forest Biomaterials and Technology, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: ingegerd.backlund@slu.se
• Bergsten, SLU, Forest Biomaterials and Technology, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: urban.bergsten@slu.se

• Backlund, Department of Forest Ecology and Management, Swedish University of Agricultural Sciences, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: ingegerd.backlund@slu.se
• Bergsten, Department of Forest Ecology and Management, Swedish University of Agricultural Sciences, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: urban.bergsten@slu.se

• Larjavaara, Smithsonian Tropical Research Institute, Unit 9100 Box 0948, DPO AA 34002-9998, USA, and University of Helsinki, Department of Forest Sciences, Finland E-mail: larjavaaram@si.edu
• Muller-Landau, Smithsonian Tropical Research Institute, Unit 9100 Box 0948, DPO AA 34002-9998, USA E-mail: hcml@nn.us

• Muukkonen, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: petteri.muukkonen@metla.fi
• Mäkipää, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: raisa.makipaa@metla.fi