The effect of nitrogen fertilizers on the photosynthetic capacity of conifers is assessed on the basis of literature. The review emphasizes the role of changes of needle mass as a factor affecting the result of nutrient application. In particular, the increase in needle mass results in a considerable increase in photosynthetic capacity. The effect of fertilization on the photosynthetic rate seems to be of minor importance. The effect on the photosynthetic rate is, however, poorly documented as is the case for the effect of fertilization on the respiration rate. There is evidence that proper application of nitrogen fertilizers may double the photosynthetic capacity of conifers, mainly as a result of increase in needle mass.
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The aim of the present study was to collect information presented in literature concerning the development and germination of the seeds of coniferous trees with special reference to those species which thrive in the conditions prevailing in Finland. Along with the increase in the importance of direct seeding as a silvicultural means in Northern Finland, there is a growing demand for methods by means of which the germination of seeds could be promoted. According to the results obtained from previous studies, such a method can be found, provided that a practically usable and reliable pre-germination method is developed.
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Since 1954 studies have been carried out by the Department of Plant Pathology of Agricultural Research Centre on occurrence of low-temperature parasitic fungi in nurseries in Finland. This paper reports analysis of the damage caused by the fungus to Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) seedlings.
In Southern and southwestern Finland, scarcely any damage caused by low-temperature parasitic fungi to coniferous seedlings was found. On the other hand, in Central, Eastern and Northern Finland, considerable injuries were present in the seedlings. The extent of damage varies between different localities and in a same location from year to year. The extent of damage is mostly dependent on snow cover which is heaviest in Central and Northern Finland. Damages are largest in wooded areas and in places where snow accumulates abundantly and remains until late in the spring.
The principal cause of winter damage to spruce seedlings is Hepotricia nigra (Hartig) which causes black snow mould. Depending on the amount of infestation, the damage can be limited to scattered groups or consist of large areas of dead seedlings. The fungus is unable to infect the plants during warm months of the growing season. The most damaging parasitic fungus in Scots pine is Phacidium infestans (Karst.) causing snow blight. The infestation varies from reddish-brown patches of infected seedlings to large areas of infected plants. Also, Botrytis cinerea has been determined from one- and two-year plants of pine and spruce.
In trials of chemical control by PCNB (pentachloronitrobenzene) gave nearly complete control of low-temperature parasitic fungi in one-year spruce seedlings. In addition, a compound of zineb (Dithane Z-78) gave similar results. Chemical control of the fungi is now common in the nurseries.
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Compression wood of the tree species studied in this investigation, Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies (L.) Karst.) and common juniper (Juniperus communis L.), was found to be characterized in its cross section by the thick walls and rounded shape of its tracheids and the profuse occurence of spaces. Tension wood of aspen (Populus tremula L.) and alder (Alnus incana (L.) Moench) was found in microscopic examination to be characterized by the gelatinous appearance of the wood fibres, by its small cell cavities and by the thickness and buckling of the inner layer of the cecondary wall. Tracheids of the compression wood were found to have shorter length than normal on an average, while the tension wood fibres were found to be longer.
The microchemical studies suggest a higher than normal lignin content in compression wood and lower than normal lignin content in tension wood, as compared to normal wood. The reverse would be true for the cellulose contents. Volume weight of absolute dry reaction wood was distinctly higher than that of normal wood. The longitudinal shrinkage of reaction wood, particularly of compression wood, is several times that of normal wood. Transversal shrinkage of compression wood is much less than normal wood. Swelling tests revealed pushing effect of compression wood on elongation and pulling effect on tension wood on constraction. Volume shrinkage of compression wood is less than that of normal wood, in contrast to tension wood. The strength of compression wood in absolutely dry condition was nearly same as that of normal wood.
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An attempt was made in the study to determine the annual periods available for foliage spraying when cleaning Scots pine (Pinus sylvestris L.) dominated seedling stands. The study was made in nine experimental fields which were established in different parts of Finland. The spraying was applied throughout the growing season by DM, MCPA and Roundup. The results were inventoried one year after the treatments.
The results showed that there were big differences both in the destruction of hardwood sprouts and in the survival of pine seedlings due to the time period of the spraying. Threshold points were observed in the range of effect of DM and MCPA. By means of these it is possible to time the spraying treatments in such a way that there remains only slight damage to pine, but hardwood sprouts are destroyed totally. The results varied with Roundup so much, among other things due to rain, that such threshold points could not be determined. This preparation both had a milder effect on the hardwood seedlings and caused slighter damage to pine than the other preparations.
In Sodankylä in Northern Finland, the pines attained a good resistance to arboricides when the efficient temperature sum of the growing season was 550, but in Punkaharju in Central Finland only when it was 850. The seed provenance of the seedlings had an effect on the resistance. The threshold temperature sums of resistance in pine were on the average 70–74% from the long-term average number of degree days at the origin of the seed. The effect on the hardwood trees grew weaker as the long-term average was filled. Resistance of pine followed with a specific lag the lignification of the shoot and the ceasing of the growth of the needles.
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