Current issue: 58(5)
The effects of wood ash and PK fertilization on natural regeneration and sowing of Scots pine (Pinus sylvestris L.) were studied in field experiments on nitrogen-poor (Ntot 0.87–1.26%) peat substrates. The study material was derived from three drained, nutrient-poor pine mires (64°52’ N, 25°08’ E) at Muhos, near Oulu, Finland. The experimental fields were laid out in 1985 as a split-split-plot design including the following treatments; mounding, natural regeneration and sowing and fertilization; PK (400 kg ha-1) and wood ash (5,000 kg ha-1). The seedlings were inventoried in circles in July–August 1991.
Changes in the vegetation were small and there were no statistical differences due to the fertilization treatments in the ground vegetation. PK or ash fertilization did not cause vegetation changes harmful to Scots pine regeneration on nitrogen-poor peatlands. Both sowing and fertilization significantly increased the number of pine seedlings, but not their height. Wood ash increased seedling number more than PK fertilizer. The number of seedlings varied from 7,963 (control) to 42,781 ha-1 (mounding + sowing + ash). The seedling number was adequate for successful regeneration even on non-mounded, non-fertilized naturally regenerated plots.
The number of birch seedlings varied more than that of pine (370–25,927 ha-1). Mounding especially increased the number of birches. The difference between PK fertiliser and ash was less pronounced than that for pine. In addition, to the field studies the effects of ash and PK fertilizer on the germination of Scots pine seeds was studied in a greenhouse experiment. Soaking in ash solutions strongly reduced seed germination, while the PK solution was less harmful.
Different methods of sowing and planting of Norway spruce (Picea abies (L.) H. Karst.) were compared on fertile sites in North Karelia (62°20’N, 29°35’E, 85–120 m a.s.l.). The planting material were 4-year-old bare-rooted transplants, 2-year-old bare-rooted seedlings, and 2-year-old containerized seedlings raised in plastic greenhouse. The sowing methods were band sowing and shelter sowing. Ground vegetation was controlled during the first growing season mechanically or chemically, or the control was omitted totally.
Planting of spruce gave better results than sowing. After eight growing seasons there were sowed seedlings left in 30% of the sowing pots. The average height of them was 35 cm. Seedling survival was best with large bare-rooted transplants (91%). Survival of containerized seedlings was 79% and of small bare-rooted transplants 71%. The average height of large bare-rooted transplants was 131 cm, of containerized seedlings 86 cm and small bare-rooted seedlings 68 cm.
Sowing is not an advisable method for regeneration of spruce due to the small survival rate and slow initial development when ground vegetation is controlled only once. Also 2-year-old seedlings gave a satisfactory result in regeneration. Seedlings raised in greenhouse were more sensitive to frost damage than seedlings grown on open ground.
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The study deals with the interaction of various soil preparation and reforestation methods. The most favourable time of the year for broadcast sowing and the effect of stabilization after soil preparation on restocking were studied as special problems.
Prescribed burning, scalping and disc ploughing made a better combination with sowing than planting, and ploughing better combination with planting than sowing. The longer the period was between sowing and germination the fewer seedlings emerged. The best stocking was clearly resulted with sowing in June. Stabilization of soil after preparation had a negative effect on reforestation results.
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The objective of the study was to compare different reforestation methods on ploughed areas in Finnish Lapland. Four species were compared: Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies (L.) H. Karst.), silver birch (Betula pendula Roth) and Siberian larch (Larix sibirica Ledeb.). The experiments were established in different parts of Lapland on different types of sites in 1970–72.
In Scots pine there was a difference of 15 percentage points in survival of seedlings between the best and worst methods of regeneration. Containerized seedlings and paper pot seedlings had the best survival rates. In Norway spruce the respective difference between sowing and planting was about 20 percentage points. In favour of planting. The survival rate can be increased by about 20 percentage points by selecting the right tree species. The average height varied from 25 cm (the sowed Norway spruce) to 179 cm (the planted silver birch) after 10 growing seasons. The birch was planted at the most fertile sites only. The longer time passed from the afforestation the clearer was the effect of the local growing conditions on the development of the seedlings. The elevation of the site was one factor seemed to influence the success of the seedlings.
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The aim of the present study was to collect information presented in literature concerning the development and germination of the seeds of coniferous trees with special reference to those species which thrive in the conditions prevailing in Finland. Along with the increase in the importance of direct seeding as a silvicultural means in Northern Finland, there is a growing demand for methods by means of which the germination of seeds could be promoted. According to the results obtained from previous studies, such a method can be found, provided that a practically usable and reliable pre-germination method is developed.
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The aim of the study was to assess, through field experiments, the possibilities of using peat briquettes in the seeding of Scots pine (Pinus sylvestris L.) in Southern Finland. The briquettes were dug into the soil in the middle of patches of mineral soil. The seeds were covered by a 2-5 mm layer of mineral soil. The seedings were inventoried in the three following autumns.
According to the results, the briquettes were clearly inferior to the control, which was ordinary drill seeding. This was mainly due to the fact that no rain was received after the seeding, and that the third summer from seeding was extremely dry. Abundant germination was observed during the second summer after seeding in both briquette seeding and the control. During more rainy summers the result might have been better.
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Experiments were carried out to find out the effect of fertilizer application on germination, seedling emmergnece and initial development in conifer plantations established on peat by sowing, with a special reference to Scots pine (Pinus sylvestris L.). The experiments were carried out in 1968–70 in laboratory, in greenhouse and in the field.
In the greenhouse experiments with Y fertilizer for peat soils (14% N, 18% P2O5, 10% K2O) it was shown that germination and seedling emergence decreased markedly with increased fertilizer application. Mortality among seedlings that had emerged was the higher the larger quantities of fertilizer had been applied. The effect of fertilization was the greater, the drier the substrate. Fine ground rock phosphate (33% P2O5) promoted seedling emergence on a dry substrate but not on a wet one.
The field experiments carried out in Central Finland included dry and wet sites. Y fertilizer, Oulu Saltpeter (25% N), fine-ground rock phosphate and potassium salt (50% K2O) were used. According to the results, easily soluble fertilizers decreased seedling emergence. On wet sites the effect of Y fertilizer was weaker than on drier sites. Fine-ground rock phosphate slightly increased the number of seedlings emerging. Height growth was increased during the first three growing seasons only by those fertilizers containing phosphorus.
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This paper reports spot sowing experiments of Scots pine (Pinus sylvestris L.). The seeds were either covered with coarse sand, tramped in the substrate or sowed without any covering, 30 seeds in each treatment in 70 replications. The site was of Vaccinium type with sandy soil. The germination percentage was 81 and 91 on the respective years. The development of seedlings was observed for 3–4 years.
The results indicate that both tramping and covering the seeds to some extent increased the number of seedlings and improved the early development. The highest numbers of seedlings were recorded in the first growing season, after which there was 23 seedlings/100 seeds in the uncovered spots, 27 seedlings in the covered spots and 31 seedlings in the tramped spots in the experiment sowed in 1965.
Mortality of the seedlings was highest between the first and second growing season, and empty spots increased with the time. There was no difference in mortality between the sowing methods, but the number of seedlings after first growing season affected the result. Under favourable conditions four seedlings per spot seemed enough to secure the survival of minimum one seedling per spot during the three first growing seasons. In poor conditions seven seedlings was needed.
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The time interval between final felling and plantation means a waste of time and thus a production loss, and may lead to lush growth of ground cover and hardwood sprouts, which increases expenses in forest management. The objective of this study was to determine the length of time between final felling and artificial regeneration in private forests in the forest districts of Uusimaa-Häme in Southern Finland and Pohjois-Häme in Central Finland. The material consists of a sample of 150 plans of the 952 cutting and regeneration plans in the district of Uusimaa-Häme and a sample of 140 plans of the 1,102 plans in Pohjois-Häme.
The time lag between final cutting and seeding or planting was on average 1.4 years in Pohjois-Häme district and 0.7 years in Uusimaa-Häme. In the latter district, 56% of the logged area was regenerated in the spring immediately following the cutting, and 84% not later than in the second spring. In Pohjois-Häme, 29% of the harvested area was regenerated immediately in the first spring following cutting, and 79% not later than in the third spring following cutting.
In Pohjois-Häme, the interval was shortest in the smallest forest holdings, and longest in the largest holdings with the largest regeneration areas. The length seems to depend mainly on the size of the regeneration area. In Uusimaa-Häme district, the interval was shortest in the smallest holdings, rather short in the largest, and longest in the intermediate-size forest properties. Seeding with Scots pine (Pinus sylvestris L.) was used in almost all regeneration areas.
The forest owners had mainly carried out the regeneration work themselves. In the Pohjois-Häme area, the interval was shorter when the district forestry board regenerated the area. 35% of the regenerated areas had required supplementary planting in Pohjois-Häme, and 47% in the Uusimaa-Häme area. Supplementary planting was more common in areas regenerated later after the cutting. In Pohjois-Häme, according to the reports of the forest owners, 75% of the regenerated areas required tending during the first three years, in Uusimaa-Häme, 80%.
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This paper describes the preliminary results of Scots pine (Pinus sylvestris L.) seeding and planting trials on drained peat soils.
The results showed that a perpared peat surface was a better surface for seeding than the unprepared one. Planting of 2+1-year seedlings succeeded better than planting 1-year seedlings. Planting on the turf gave better survival than planting on the unprepared soil surface. The whole growing season was suitable time for planting Scots pine seedlings except May when the peat soil under the surface was still frozen.
Using fertilizers in connection with planting was surveyed in two ways. Mortality of seedlings increased when they were top-dressed with NPK fertilizer. Using a so-called spot fertilizing with several combinations of fertilizers resulted in K and N tending to increase the mortality of seedlings, but P decreasing mortality.
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In 1933, forest fire caused by locomotive sparkle burned about 600 hectares of forest in a forest district named Vehkatallinmaa, in Central Finland. In 1934–36, the burned area was reforested, using different sowing and planting methods. At the same time, areas with poor runoff were drained. The results from reforestation throughout the area have been good. Also, natural regeneration of coniferous trees, especially Scots pine (Pinus sylvestris L.) has occurred. Even deciduous trees, especially birch (Betula sp.), have regenerated naturally in the area. The forests are an evidence of the adaptability of broadcast sowing on snow crust as a method of reproduction.
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The most common way to artificially regenerate cutting areas in Northern Finland has been sowing, which has, however, often given poor results. The aim of the study was to assess the success rate of sowing and study the causes of poor regeneration.
An inventory was made of 28 areas on Empetrum-Myrtillus and Hylocomnium-Myrtillus type sites sown with Scots pine (Pinus sylvestris L.) seeds in 1948-1950. In addition, to study the effect of substrata, a sowing experiment was established. According to the inventory, regeneration failed completely in 8 and partially in 6 of the 28 sown areas. Factors that explained the poor regeneration included unfavourable weather conditions in 1948-1952, insufficient clearing of sowing spot especially when the humus layer was thick, and insect damage by Otiorynchus dubius weevil.
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Silva Fennica issue 52 includes presentations held in professional development courses, arranged for foresters working in public administration in 1938. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.
This presentation describes preparation and forest regeneration methods of a burned area.
The issue 39 of Silva Fennica includes presentations held in professional development courses in 1935 that were arranged for foresters working in public administration. The presentations focus on practical issues in forest management and administration, especially in regional level.
This presentation describes the forest management work in the state forests.
Silva Fennica Issue 39 includes presentations held in professional development courses in 1935 that were arranged for foresters working in public administration. The presentations focus on practical issues in forest management and administration, especially in regional level.
This presentation describes the methods of artificial regeneration of forests.
Silva Fennica Issue 39 includes presentations held in professional development courses in 1935 that were arranged for foresters working in public administration. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.
This presentation describes natural regeneration in forests and factors affecting the success of regeneration.
Broadcast sowing on snow was relatively new method in the beginning of the 20th century in Finland, and the experiences of regeneration were diverse. The aim of the survey was to study the success rate of regeneration in the oldest and largest areas regenerated with this sowing method in Tuomarniemi district. Scots pine (Pinus sylvestris L.) was the most common tree species, but also Norway spruce (Picea abies (L.) H. Karst.) and European larch (Larix decidua Mill.) were used in broadcast sowing on snow.
According to the study, the success of broadcast sowing on snow was as good as patch sowing and sowing in furrows in the sites typical for Tuomarniemi. The regeneration areas were often drained peatlands or paludified lands. When sowing is done using Norway spruce seeds, site preparation either by broadcast burning or scalping with hoe is recommended. Mixed sowing with pine and spruce seldom succeeded due to the differences in site requirements of the species and growth of seedlings. Sowing of Scots pine succeeded well on the drained peatlands. Sowing should be done some years after draining to let the peat dry and sink. Site preparation is needed in sites growing Polytrichum-moss. Broadcast burned areas larger than 10 hectares seemed to regenerate poorer than sites in average, possibly due to dryness of the sites. Trials with European larch were successful, and the growth of the seedlings acceptable despite the sites being relatively poor for the species.
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The article is a review on early forest regeneration and management in Finland. Beginning of the 1900s marks change in attitudes and resources for forest management. The state increased the funding of forest regeneration and improvement in the state forests in 1928. State funding is directed also to forest improvement in the private forests, and organizations established to promote forest management in the private forests are reorganized. For instance, District Forestry Boards were appointed the forest improvement work in private lands, in addition to promotion of private forestry. Sowing increased in the state forests from 772 ha to 1,566 ha, in forests of the forest companies from 3,006 ha to 4,954, and in private forests from 1,417 ha to 1,566 ha in 1923-1926. The figures of private forests are, however, incomplete. The most usual methods are patch sowing and broadcast sowing on snow. Seeds used in sowing increased from 3,357 kg to 14,387 kg, and planting from 413 ha to 1,020 ha in 1923-1930. Almost half of the sown areas were in the state forests, and most of the planted area in the forests of the companies. Scots pine (Pinus sylvestris L.) was the main tree species in artificial regeneration, and Norway spruce (Picea abies (L.) H. Karst.) was more popular in planting.
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Regeneration of large open areas in dry mineral soil forest sites that usually grow Scots pine (Pinus sylvestris L.) have several problems. For instance, soil frost, snow, ground vegetation and dryness can prevent germination and growth of seedlings. The damages caused by insects and fungi in seedlings of a large burned area in Siikakangas in Southern Finland was studied. A forest fire burned the area nearly completely in 1909, and 310 hectares have been sowed or planted with mostly Scots pine during the following years. Minor areas have been regenerated with Pinus montana Noll, Pinus excelsa Lamb., Pinus murrayana Balf. and Larix sibirica Ledeb.
No completely healthy pine seedling stands could be found in the area. About 41% of the seedlings in the sample plots were damaged. The most common causes for damage were Evetria resinella (now Retinia resinella L.), Luperus pinicola (now Calomicrus pinicola (Duft.)), Pissodes notatus (now Pissodes castaneus Degeer), Evetria turionana Hb. and Hylobious abietis L. The most usual fungal disease was Lophodermium sp. Evetria resinella caused damages in all the area. Evetria turionana, Pissodes notatus and Hylobius abietina were found in the older seedling stands. Other damages were more localized. The slacks in the terrain seemed to have most damages, the original cause being probably soil frost. Some damages, as Lophodermium, were related to the density of the seedlings, especially in the sown areas. Cleaning of seedling stands could decrease these damages. Planting seems to have succeeded better than patch sowing.
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Prescribed burning is a common silvicultural practice in northern Europe, intended to destroy the slash and ground vegetation and to reduce the thickness of the raw humus layer prior reforestation. The purpose of the experiments was to study whether there are any differences in the commencement and early development of mycorrhizal infection between burned and unburned areas. A clear-cutting area was burned on May 1961. The soil was rocky moraine, the forest type was Vaccinium type. Two weeks after burning Scots pine (Pinus sylvestris L.) was sown in patches.
According to the results, mycorrhizal infection took place on the unburned area earlier than on the burned. The difference was relatively small, perhaps 1–2 weeks. Although burning kills mycorrhizal fungi, it did not cause serious harm to the seedlings, on the contrary, the favourable influence of burning was more distinct. The high temperatures caused by the fire are restricted in the soil in a prescribed burning only a few centimetres deep. Although the mycorrhizal fungi are concentrated in a very thin surface layer of the soil, some mycorrhizae are situated deeper, and from there the fungi are able to infect roots and spread back to the surface layer. The fire also rises the pH of the soil, which can be harmful for mycorrhizal fungi. Even this effect, however, is limited to a thin surface layer.
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This paper aims at studying regeneration of Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) by sowing and natural regeneration of birch (Betula sp.) in Western Finland.
Germination of spruce and pine seeds may be prevented by dryness and temperatures below the optimum for germination. In natural conditions, when temperature and moisture is insufficient for germination, the type of seedbed generally has en effect on germination result. Trenching of the seeding spots showed that root competition during the early stage of regeneration was not of decissive importance. It seemed to, however, improve the preservation of the seedlings later. It is common that it can take long before the seeds germinate, and during that time the number of viable seeds decrease strongly.
Also, the seedling stock quickly began to decrease in number after germination, especially during the first growing season and the following winter. The decrease was larger in intact vegetation than on mineral soil or in the humus layer. The emerging seedlings were destroyed by drought very easily, but their tolerance to drought improved later on.
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In this paper the development of sown Scots pine (Pinus sylvestris L.) seedling stands into forests is studied. The material was collected in stands sown in 1930–1940 in private forests in the Forestry Board districts of Central and Northern Ostrobothnia. The 119 areas, including both burned and other areas, were studied in 1955.
Most seedings had been carried out on relatively poor soils, mostly representing Vaccinium and Calluna type forests. 71% of the areas consisted of large forest fires, mostly from 1933. The most burned areas did not have seed producing trees nearby. The other sown areas were in general small, 1–2 ha, and near forests capable of producing seeds. The species of previous tree generation, in the older areas mostly pine and in the younger areas Norway spruce, affected tree species composition of the new tree generation.
Over 90% of the burned areas were in silviculturally good or satisfactory condition, while the main part of the other sown stands was in fair or poor condition. Weeding and thinning had been done only in the oldest stands. Most stands had been left untended. Natural new trees often competed with the sown pines, and cull-trees and border forest increased natural regeneration in the areas. In Calluna type the poor soil limited regeneration and growth of broadleaf trees. The worst competitors were naturally regenerated pine seedlings both on Calluna and Vaccinium type. On Vaccinium type also birch and sometimes also aspen (Populus tremula L.) competed with sown pine. On better sites and paludified areas competition by broadleaf trees increased. The rhythm of development of broadleaved trees is so different from pine that only those broadleaved trees that are formed in the stand when the pine seedlings are larger can develop harmoniously with pine. Due to the harmful competition, the seedling stands should be tended early on. In addition, it may be advisable to abandon the practise to leave trees on sowing areas.
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Prescribed burning has been used in regeneration areas in Finland as a method to treat the humus layer and creating more favourable chemical, physical and biological conditions for the seedlings. At the same time, fire clears away seedlings and shoots of unwanted trees and other vegetation. Direct sowing or planting, mostly Scots pine (Pinus sylvestris L.), seldom natural regeneration, is used. In this paper, the initial stages of the formation of a new tree generation of Scots pine and Norway spruce (Picea abies (L.) Karst.) on prescribed burned areas is studied in Central Finland in 1956–1960.
The burned area remains almost without vegetation for about two growing seasons. Conditions on a burned area which has not been tiled are very unfavourable for germination of seeds of coniferous and deciduous trees. On the other hand, shoots of deciduous trees occur soon after burning. Conditions for regeneration were found to be better 3–5 years after burning. Removal of humus layer in spots improved regeneration. However, the patches facilitated also natural regeneration of Norway spruce and especially birch (Betula sp.), which compete with Scots pine seedlings.
Continuous rainy periods improved the germination of Scots pine and Norway spruce seeds sown on the humus layer. Pine and spruce developed more rapidly on the exposed soil, however, young seedlings were easily destroyed. Seed eaters destroyed the pine and spruce seeds sown on the humus layer of newly burned areas completely or almost completely. The viability of pine seeds sown on the burned humus layer did not decrease for three weeks, but the viability greatly weakened after six or more weeks. Spruce seeds lost their viability faster than pine seeds.
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In Southern Finland Scots pine (Pinus sylvestris L.) is mainly sown on Vaccinium and Myrtillus-type sites. The material for the study was collected by measuring sample plots in pure, even-aged pine stand that had been sown. The sample stands had been thinned from below.
The volume of the stands was roughly the same as that of repeatedly thinned pine stands. The cubic volume of sown pine stands is 65–90%, varying according to age, of that of natural-normal pine stands. The current annual volume increment of stands on Myrtillus-type was 8–9 m3/ha at age of 20–30 years. The peak was reached at age of 35 years with 9 m3/ha, in the following years the increment is about 8 m3/ha until the age of 60 years. On Vaccinium type sites increment reaches 6–7 m3 level at age of 30 years, and attains the peak of 7 m3/ha at the age of 45 years. Annual increment was in young and middle-aged Myrtillus-type stands about 10% greater, and on Vaccinium-type stands 15–20% greater than in natural-normal pine stands.
The total volume increment in 70 years old Myrtillus-type stands was 580 m3/ha over bark, and in 80 years old Vaccinium-type stands 520 m3/ha. The total removal on Myrtillus-type sites totalled nearly 350 m3/ha in sown pine stands up to 70 years of age, and 280 m3/ha on Vaccinium-type stands. The total yield in sawn timber per hectare rises up to 6,300 cubic ft in a 70 years old stand on Myrtillus-type stands, and 5,300 cubic ft in Vaccinium-type stands. In conclusion, the volume and increment development of managed pine stands established by sowing up to 70–80 years of age is largely the same as in repeatedly thinned pine stands, but the structure and yield offer greater advantages. The investigation demonstrates that, in the case of Scots pine, sowing is an advantageous method of regeneration. Sowing is an advantage especially in the cases where natural regeneration is uncertain and slow.
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Following the example of Norway, which was the first nation to employ regular troops in silvicultural work, sowing and planting courses were organized for the Finnish troops in 1931. The initiative was taken by the Forestry Instruction Bureau, and the work was organized by the forestry committees. During three years, total of 8,189 men was involved in the work.
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Larix sibirica Ledeb. is one of the promising timber species for planting in the boreal ecosystem; but poor seed lot quality is the major hurdle for production of sufficient quantity of planting stocks. Here, we evaluated the potential of Near Infrared (NIR) Spectroscopy for sorting viable and non-viable seeds, as the conventional sorting technique is inefficient. NIR reflectance spectra were collected from single seeds, and discriminant models were developed with Orthogonal Projections to Latent Structure – Discriminant Analysis (OPLS-DA). The computed model predicted the class membership of filled-viable, empty and petrified seeds in the test set with 98%, 82% and 87% accuracy, respectively. When two-class OPLS-DA model was fitted to discriminate viable and non-viable (empty and petrified seeds combined), the predicted class membership of test set samples was 100% for both classes. The origins of spectral differences between non-viable (petrified and empty) and viable seeds were attributed to differences in seed moisture content and storage reserves. In conclusion, the result provides evidence that NIR spectroscopy is a powerful non-destructive method for sorting non-viable seeds of Larix sibirica; thus efforts should be made to develop on-line sorting system for large-scale seed handling.