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Silva Fennica 1926-1997
Acta Forestalia Fennica

Articles containing the keyword 'biomass production'

Category: Article

article id 5551, category Article
Jyrki Hytönen, Pekka Rossi, Anna Saarsalmi. (1995). Biomass production and nutrient uptake of short-rotation plantations. Silva Fennica vol. 29 no. 2 article id 5551.
Keywords: Betula pendula; biomass production; Betula pubescens; Alnus incana; short-rotation forestry; Salix phylicifolia; Salix x dasyclados; Salix 'Aquatica'; nutrient uptake; Salix triandra
Abstract | View details | Full text in PDF | Author Info

The biomass production and nutrient uptake of silver birch (Betula pendula Roth), downy birch (Betula pubescens Erhr.), grey alder (Alnus incana (L.) Moench), native willows Salix triandra L. and S. phylicifolia L. and exotic willows S. x dasyclados and S. ’Aquatica’ growing on a clay mineral soil field (Sukeva) and on two cut-away peatland areas (Piipsanneva, Valkeasuo) were investigated.

Biomass production of downy birch was greater than that of silver birch, and the biomass production of the native willows greater than that of the exotic ones. The performance of S. phylicifolia was the best of the studied willow species. Exotic willows were susceptible to frost damage and their winter hardiness was poor. The production of all species was lower on the clay mineral soil field than on the cut-away peatland areas. Fertilization of birches and alder – on the double dose given to the willows – increased biomass production. After 6 growing seasons the leafless biomass production of fertilized silver birch at Piipsanneca was 21 t ha-1 (at Valkeasuo 34 t ha-1) and of grey alder 24 t ha-1, and that of S. triandra after five growing seasons 31 t ha-1, S. phylicifolia 38 t ha-1 and of S. x dasyclados 16 t ha-1.

6-year-old stands of silver birch bound more nutrients per unit biomass than downy birch stands. Grey alder bound more N, Ca and Co but less Mn and Zn per unit biomass than silver and downy birch. On the field more P was bound in grey alder per unit biomass compared to downy birch. The willows had more K per unit biomass than the other tree species, and the exotic willow species more N than the native ones. Less N, K and Mg were bound per unit biomass of S. phylicifolia compared to the other tree species.

  • Hytönen, E-mail: jh@mm.unknown (email)
  • Rossi, E-mail: pr@mm.unknown
  • Saarsalmi, E-mail: as@mm.unknown
article id 5550, category Article
Jyrki Hytönen. (1995). Effect of repeated fertilizer application on the nutrient status and biomass production of Salix ’Aquatica’ plantations on cut-away peatland areas. Silva Fennica vol. 29 no. 2 article id 5550.
Keywords: biomass production; fertilization; fuelwood; peatlands; short-rotation forestry; Salix 'Aquatica'
Abstract | View details | Full text in PDF | Author Info

The effects of repeated fertilizer treatment on biomass production and nutrient status of willow (Salix ’Aquatica’) plantations established on two cut-away peatland areas in western Finland were studied over a rotation period of three years. Comparisons were made between single fertilizer applications and repeated annual fertilization.

The annually repeated fertilizer application increased the amounts of acid ammonium acetate extractable phosphorus and potassium in the soil as well as the concentrations of foliar nitrogen, phosphorus and potassium compared to single application. Depending on the fertilizer treatment and application rate, annual fertilizer application resulted in over two times higher biomass production when compared to single fertilizer application over a three-year rotation period. The effect of phosphorus fertilizer application lasted longer than that of nitrogen. The optimum fertilization regime for biomass production requires that nitrogen fertilizer should be applied annually, but the effect of phosphorus can last at least over a rotation of three years. Potassium fertilizer treatment did not increase the yield in any of the experiments during the first three years. The leafless, above-ground yield of three-year-old, annually NP-fertilized willow plantations was 9.5 t ha-1 and the total biomass, including stems, leaves, roots and the stump, averaged 17 t ha-1.

  • Hytönen, E-mail: jh@mm.unknown (email)
article id 5544, category Article
Jyrki Hytönen. (1995). Effect of fertilizer treatment on the biomass production and nutrient uptake of short-rotation willow on cut-away peatlands. Silva Fennica vol. 29 no. 1 article id 5544.
Keywords: biomass production; fertilization; fuelwood; nutrients; peatlands; short-rotation forestry; Salix; consumption
Abstract | View details | Full text in PDF | Author Info

The effects of fertilized treatment on the soil nutrient concentrations, biomass production and nutrient consumption of Salix x dasyclados and Salix ’Aquatica’ were studied in five experiments on three cut-away peatland sites in western and eastern Finland during three years. Factorial experiments with all combinations of N (100 kg ha-1 a-1), P (30 kg ha-1 a-1) and K (80 kg ha-1 a-1) were conducted.

The application of P and K fertilizers increased the concentrations of corresponding extractable nutrients in the soil as well as in willow foliage. N-fertilization increased foliar nitrogen concentration. An increase in age usually led to decreases in bark and wood N, P and K concentrations and increases in bark Ca concentrations. N-fertilization increased the three-year biomass yield 1.5–2.7 times when compared to control plots. P-fertilization increased the yield only in those experimental fields whose substrates had the lowest phosphorus concentration. K-fertilization did not increase the yield in any of the experimental fields. The highest total biomass yield of NPK-fertilized willow after three growing seasons, 23 t ha-1, was distributed in the following way: wood 42%, bark 19%, foliage 17%, stumps 6% and roots 16%. As the yield and stand age increased, more biomass was allocated in above-ground wood. Three-year-old stands (above-ground biomass 18 t ha-1) contained as much as 196 kg N ha-1, 26 kg P ha-1, 101 kg K ha-1, 74 kg Ca ha-1 and 37 kg Mg ha -1. By far the highest proportion of nutrients accumulated in the foliage. The bark and wood contained relatively high proportions of calcium and phosphorus. With an increase in age and size, the amount of nitrogen and potassium bound in one dry-mass ton of willow biomass decreased while that of phosphorus remained unchanged.

  • Hytönen, E-mail: jh@mm.unknown (email)
article id 5459, category Article
Pertti Pulkkinen. (1991). Crown form and harvest increment in pendulous Norway spruce. Silva Fennica vol. 25 no. 4 article id 5459.
Keywords: biomass production; allocation; needle biomass; crown; stem; Picea abies f. pendula; branchwood biomass
Abstract | View details | Full text in PDF | Author Info

Crown characteristics and the distribution of three years’ (1986–88) biomass production of 20 pendulous Norway spruce (Picea abies f. pendula (Lawson) Sylvén) trees with heritable narrow crown and 15 normal-growned spruces (Picea abies (L.) H. Karst.) were studied in a 19-year-old mixed stand.

The form of the crown is conical in normal-crowned trees, columnar and narrow in pendulous trees. The partitioning of aboveground biomass to stems during the studied 3-year period was significantly higher in pendulous (0.281) than in normal-crowned trees (0.255) and also the ratio between growth of stemwood and growth of needle biomass during three years was higher in pendulous trees (0.67 g g-1) than in normal-crowned trees (0.52 g g-1). The needle biomass was distributed higher in the crown in pendula than in normal-crowned trees and they had a higher needle biomass/branchwood biomass ratio than normal trees. The difference in harvest increment between the two crown types are mostly due to the significantly lower branchwood biomass values in pendulous than in normal-crowned trees. The higher needle ’efficiency’ in pendulous trees is probably connected with high partitioning of needle biomass to the upper part of the crown in pendulous trees.

The PDF includes an abstract in Finnish.

  • Pulkkinen, E-mail: pp@mm.unknown (email)
article id 5385, category Article
Timo Kuuluvainen. (1989). Branching dynamics in young Scots pine. Silva Fennica vol. 23 no. 3 article id 5385.
Keywords: Pinus sylvestris; biomass production; branching; tree architecture; bifuraction; modular construction; shoot demography
Abstract | View details | Full text in PDF | Author Info

The development of shoot number and shoot properties was examined in successive shoot cohorts of young widely-spaced Scots pine trees (Pinus sylvestris L.) growing in a progeny trial. This was accomplished by reconstructing the branching process of the trees over a period of five years, from tree age 4 to 8. During this time the number of shoots in successive shoot cohorts increased rapidly, while at the same time the mean shoot length decreased. The decrease in shoot lengths from older to younger shoots was accompanied by a decline in the bifuraction frequency of the shoots. In general, rapid changes occurred in the branching characteristics during the yearly development of the trees. The variation in the branching characteristics was reflected in the development of the architecture and biomass production of the trees.

The PDF includes an abstract in Finnish.

  • Kuuluvainen, E-mail: tk@mm.unknown (email)
article id 5347, category Article
Timo Kuuluvainen, Markku Kanninen, Juha-Pekka Salmi. (1988). Tree architecture in young Scots pine: properties, spatial distribution and relationships of components of tree architecture. Silva Fennica vol. 22 no. 2 article id 5347.
Keywords: Pinus sylvestris; crown architecture; biomass production; branching pattern; tree ideotype
Abstract | View details | Full text in PDF | Author Info

The architecture of Scots pine (Pinus sylvestris L.) was studied in an eight-year-old progeny test. The measurements included characteristics of crown structure, spatial distribution of shoots and yield components. The spatial distribution of shoots showed striking between-tree differences, and two extreme distribution patterns were detected. One represented a non-layered structure with a vertically relative even shoot distribution, and the other a layered structure with a vertically highly uneven shoot distribution.

Close correlations existed between several components of tree architecture and it is suggested that changes in the phenotypic architecture in Scots pine follow an epigenetic pattern, which enables the prediction of adaptational changes in structural components. The structural characteristics related to high above-ground biomass were a long crown, high total shoot length, high number of branches per whorl and big shoots of low needle density occupying a big share of the crown volume.

The PDF includes a summary in Finnish.

  • Kuuluvainen, E-mail: tk@mm.unknown (email)
  • Kanninen, E-mail: mk@mm.unknown
  • Salmi, E-mail: js@mm.unknown
article id 5342, category Article
Ilari Lumme, Timo Törmälä. (1988). Selection of fast-growing willow (Salix spp.) clones for short-rotation forestry on mined peatlands in northern Finland. Silva Fennica vol. 22 no. 1 article id 5342.
Keywords: biomass production; breeding; Salix spp.; short-rotation forestry; willows; Salix viminalis; Salix myrsinifolia; winter hardiness; clone selection; mined peat areas
Abstract | View details | Full text in PDF | Author Info

Possibilities of developing suitable willow (Salix spp.) clones for short-rotation forestry on mined peatlands in the north-western area of Finland were studied in a field experiment in which 300 willow clones were tested during 1985–87. Most of the tested clones started to grow from cuttings on limed and fertilized peat soil. Salix viminalis L. clones of southern origin had a higher leafless above ground biomass production than the well adapted control clone and the local Finnish willows, but their winter hardiness was not satisfactory. The growth habit of some southern willows was also better than that of the control clone. It was also possible to select clones with good sprouting capacity. There were few Salix myrsinifolia Calisb. clones of Finnish origin, which had better cold tolerance than all other willows tested and higher biomass production than that of the control clone. The most critical factor to be selected for this is the optimal combination of winter hardiness and biomass production. This is attempted by selecting clones on the basis of this experiment for a breeding program.

The PDF includes a summary in Finnish.

  • Lumme, E-mail: il@mm.unknown (email)
  • Törmälä, E-mail: tt@mm.unknown

Category: Article

article id 7667, category Article
Veli Pohjonen. (1991). Selection of species and clones for biomass willow forestry in Finland. Acta Forestalia Fennica vol. 0 no. 221 article id 7667.
Keywords: biomass production; exotics; Salix viminalis; Salix myrsinifolia; indigenous species; Salicacea; heterosis
Abstract | View details | Full text in PDF | Author Info

Willow (Salix sp.) species and clones have been selected in Finland, originally for basket willow husbandry, since 1910's. Screening for biomass willows started in 1973 by the Foundation for Forest Tree Breeding. Biomass willow research for energy started in 1978. The objective of the study was, based on theoretical background, on historical record of Finnish willow research between 1910–80 and on analysis of the Finnish biomass willow research of the 1980s, a further selection of exotic and indigenous willows for energy and chemicals.

Swedish selection of 63 exotics, mainly of Salix viminalis L. and S. burjatica Nazarov, was screened in Kompparnäs willow research site in the southern coast of Finland in 1983–89. S. viminalis showed both high yield potential and good crop certainty. The yield variation in S. burjatica were big due to rust (Melampsora sp.) infection followed by lowered winter hardiness. Three recommendable S. viminalis clones for Southern Finland were found.

Finnish indigenous species were screened based on collection (375 clones) in 1974–74 of the Foundation for Forest Tree Breeding, and the Finnish 4H-organization (566 clones) in 1978–79 in test sites in Suomusjärvi, Nurmijärvi, Kannus and Haapavesi. Salix myrsinifolia Salisb. was most productive of the indigenous willows. Five recommendable clones were selected. The second most productive indigenous was S. phylicifolia, with three recommendable clones. Based on willow hybridization studies in the Finnish Forest Research Institute, a considerable additional selection effect, boosted by heterosis, was found from the progenies. Further intraspecific crossings of geographically distant clones of S. myrsinifolia are recommended.

Based on the results, S. viminalis is recommended for practical biomass forestry application in the southernmost agroclimatic zone of Finland. S. myrsinifolia is recommended for further research and development in the other zones.

The PDF includes a summary in Finnish.

  • Pohjonen, E-mail: vp@mm.unknown (email)

Category: Research article

article id 1260, category Research article
Jyrki Hytönen, Anna Saarsalmi. (2015). Biomass production of coppiced grey alder and the effect of fertilization. Silva Fennica vol. 49 no. 1 article id 1260.
Keywords: biomass production; coppicing; grey alder; fertilisation
Highlights: Fertilisation (wood ash, N, PK) did not affect grey alder biomass production; Leafless above-ground biomass of 17–20 year old stands was 52–57 Mg ha–1; MAI increased with increase of rotation length to the end of the follow-up period of 17–20 years; Coppicing increased stand density manifold.
Abstract | Full text in HTML | Full text in PDF | Author Info
We studied biomass production of two naturally originated grey alder (Alnus incana (L.) Moench) stands having a mixture of birch and willow located in central Finland. One of the stands was growing on a peatland site (Muhos) and the other on a mineral soil site (Juuka). The stands were clear-cut and fertilization experiments were laid out with several treatments. At Muhos, the treatments included nitrogen fertilisation with different amounts of wood ash and an unfertilized control. At Juuka, the treatments included nitrogen fertilisation either with ash or with PK, and ash and PK treatments alone and an unfertilized control. The sprouts at Muhos were grown for 17 years and at Juuka for 20 years. At Juuka the stand was clear-cut second time at the age of 20 years and grown for 8 years. The stands were measured several times and foliar samples were taken twice during the study period. Clear-cutting increased stem number manifold. The stand density of new coppiced forests after the clear-cutting decreased from 67 000–89 000 stems ha–1 at the age of 3–6 years to 10 000–12 000 stems ha–1 at the age of 17–20 years. On neither site fertilization affected biomass production of alders during the study period. Leafless above-ground biomass was 52–57 Mg ha–1 after 17–20 years. Mean annual leafless above-ground biomass production (MAI) increased with increase of rotation time. At the age of 17–20 years the MAI was 2.8–3.0 Mg ha a–1. At Muhos, ash increased foliar P and Ca concentrations, but decreased those of Mn.
  • Hytönen, Natural Resources Institute Finland, Silmäjärventie 2, FI-69100 Kannus, Finland E-mail: (email)
  • Saarsalmi, Natural Resources Institute Finland, P.O. Box 18, FI-01301 Vantaa, Finland E-mail:
article id 48, category Research article
Jyrki Hytönen, Lasse Aro. (2012). Biomass and nutrition of naturally regenerated and coppiced birch on cutaway peatland during 37 years. Silva Fennica vol. 46 no. 3 article id 48.
Keywords: Betula; biomass production; coppicing; fertilization; PK fertilizers; wood ash
Abstract | View details | Full text in PDF | Author Info
Biomass production and nutrient use of birch thickets with a mixture of willow on a cut away peatland in southern Finland over a period of 37 years was studied. Dense, naturally regenerated 16-year-old birch stands were cut down, fertilized with either wood ash (P 108 and K 339 kg ha–1) or PK fertilizer (P 50 and K 95 kg ha–1) or left unfertilized. The biomass production of the coppiced stands and one uncut stand was monitored for a period of 21 years. Soil nutrient and foliar nutrient concentrations were analyzed several times during the study period. Ash fertilization supplied more nutrients than PK fertilization and increased the soil nutrient amounts more. The foliar phosphorus concentration of birch on control plots indicated a severe phosphorus deficiency which was removed by PK and ash fertilization. Fertilization did not increase nutrient concentrations of the stem (wood + bark) nor the amount of nutrients bound in the biomass. Two energy wood rotations (16+21 years) produced 124–158 Mg ha–1 of leafless, above-ground biomass altogether corresponding to 61–78 Mg ha–1 of carbon. The highest biomass yield was achieved with a rotation of 37 years in the uncut stand (211 Mg ha–1). Corresponding values for mean annual increment (MAI) were 3.4–4.3 Mg ha–1 and 5.7 Mg ha–1. This study shows that the length of the rotation for birch in energy wood production should be longer than 21 years. PK and ash fertilization increased the biomass of coppiced 21-year-old birch by 23 Mg ha–1 and 33 Mg ha–1, respectively.
  • Hytönen, Finnish Forest Research Institute, Kannus, Finland E-mail: (email)
  • Aro, Finnish Forest Research Institute, Kannus, Finland E-mail:
article id 649, category Research article
Tord Johansson. (1999). Biomass production of Norway spruce (Picea abies (L.) Karst.) growing on abandoned farmland. Silva Fennica vol. 33 no. 4 article id 649.
Keywords: Norway spruce; biomass production; Picea abies; thinning; LAI; abandoned farmland; basic density
Abstract | View details | Full text in PDF | Author Info
Biomass production of forests has been studied for at least a century. Tree biomass is used in Sweden both as industrial raw material and an energy source. Few studies dealing with biomass yield from Norway spruce (Picea abies (L.) Karst.) growing on farmland are published. Practical recommendations are sparsely. The aim of this study was to construct dry weight equations for Norway spruce growing on farmland. Dry weight equations for fractions of Norway spruce trees were made. Biomass production was estimated in 32 stands of Norway spruce growing on abandoned farmland. The stands were located in Sweden at latitudes ranging from 58° to 64° N, and their total age varied from 17 to 54 years. A modified ‘mean tree technique’ was used to estimate biomass production; i.e. the tallest tree was chosen for sampling. The actual mean total dry weight above stump level for the 32 stands was 116 ton ha–1, with a range of 6.0 to 237.4 ton d.w. ha–1. When previous thinning removals were included, the mean biomass value was 127 ton ha–1 (6.0–262.8). In addition to estimating conventional dry weights of trees and tree components, basic density, specific leaf area, total surface area and leaf area index, among other measures, were estimated. Norway spruce biomass yields on plots subjected to different thinning were compared. The total harvested biomass was 75–120 ton d.w. ha–1 in heavy thinnings from below. Stands were thinned four to five times, with the first thinning at 23–27 years and the last at 51–64 years. The harvested biomass obtained in the first thinning was 18–38 ton d.w. ha–1. Total biomass production was 178–305 ton d.w. ha–1. Stands thinned from above supplied 71–130 ton d.w. ha–1 in total and 17–42 ton d.w. ha–1 in the first thinning. The total biomass supply was 221–304 ton d.w. ha–1. Unthinned stands produced a total of 155–245 ton d.w. ha–1.
  • Johansson, Swedish University of Agricultural Sciences, Department of Forest Management and Products, P.O. Box 7060, S-750 07 Uppsala, Sweden E-mail: (email)

Category: Research note

article id 914, category Research note
Ingegerd Backlund, Urban Bergsten. (2012). Biomass production of dense direct-seeded lodgepole pine (Pinus contorta) at short rotation periods. Silva Fennica vol. 46 no. 4 article id 914.
Keywords: biomass production; lodgepole pine; direct seeding; Pinus contorta; biorefinery; short rotation
Abstract | View details | Full text in PDF | Author Info
Lodgepole pine (Pinus contorta) is a fast-growing species that is suitable for producing woody biomass in Nordic countries. Direct seeding of this species is cheaper than planting and creates dense, stable stands. The objective of this study was to quantify the stem volume and biomass production of direct seeded lodgepole pine stands grown under different site conditions with different stem densities, at an age that would permit extensive harvesting of biomass. A circle-plot inventory was performed in 16 of the oldest direct seeded lodgepole pine stands in mid-northern Sweden. Stemwood production of almost 200 m3/ha was achieved on average on the best sites, rising to about 300 m3/ha for the best circle-plots within 30 years of direct seeding despite the fact that pre-commercial thinning was made once or twice. This corresponds to 100 and 140 tons of dry weight biomass/ha, respectively. Higher stand stem densities (≥3000 st/ha) yielded more biomass with only slight reductions in diameter at breast height. The development of stem volume with respect to dominant height in direct seeded stands was becoming comparable to that in planted stands with similar spacing. It therefore seems that there is an unutilized potential for cost-effectively growing lodgepole pine in dense stands for biomass production after direct seeding. It may be possible to devise regimes for short(er) rotation forestry that would yield substantial amount of inexpensive biomass for biorefineries within a few decades.
  • Backlund, Department of Forest Ecology and Management, Swedish University of Agricultural Sciences, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: (email)
  • Bergsten, Department of Forest Ecology and Management, Swedish University of Agricultural Sciences, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail:

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