Articles containing the keyword 'Salix'

Willows (Salix phylicifolia) were grown for four months in organic rich soil at four nutrient levels (fertilization with a micronutrient-macronutrient mixture of 0, 100, 500 and 1,000 kg ha^-1 per month) and four CO₂ concentrations (300, 500, 700 and 1,000 ppm). Nitrogen and phosphorus concentration of the willows were reduced at CO₂ enhancement, the decrease being larger in the leave and roots than in the stems. Nitrogen content of the willows plus extractable nitrate-N in the soil coincided well with the doses of nitrogen supplied, but the corresponding sum of phosphorus in the plants and soil were smaller. The total nitrogen content of willows grown in unfertilized soil was nearly two times higher than the sum of the extractable nitrate-N in soil and N content of the cutting at the beginning of the experiment. The contents of nitrogen and phosphorus of the unfertilized willows were independent of CO₂ concentration, suggesting that CO₂ concentration did not affect through increased mineralization the availability of those nutrients to the willows.

• Silvola, E-mail: js@mm.unknown
• Ahlholm, E-mail: ua@mm.unknown

The biomass production and nutrient uptake of silver birch (Betula pendula Roth), downy birch (Betula pubescens Erhr.), grey alder (Alnus incana (L.) Moench), native willows Salix triandra L. and S. phylicifolia L. and exotic willows S. x dasyclados and S. ’Aquatica’ growing on a clay mineral soil field (Sukeva) and on two cut-away peatland areas (Piipsanneva, Valkeasuo) were investigated.

Biomass production of downy birch was greater than that of silver birch, and the biomass production of the native willows greater than that of the exotic ones. The performance of S. phylicifolia was the best of the studied willow species. Exotic willows were susceptible to frost damage and their winter hardiness was poor. The production of all species was lower on the clay mineral soil field than on the cut-away peatland areas. Fertilization of birches and alder – on the double dose given to the willows – increased biomass production. After 6 growing seasons the leafless biomass production of fertilized silver birch at Piipsanneca was 21 t ha^-1 (at Valkeasuo 34 t ha^-1) and of grey alder 24 t ha^-1, and that of S. triandra after five growing seasons 31 t ha^-1, S. phylicifolia 38 t ha^-1 and of S. x dasyclados 16 t ha^-1.

6-year-old stands of silver birch bound more nutrients per unit biomass than downy birch stands. Grey alder bound more N, Ca and Co but less Mn and Zn per unit biomass than silver and downy birch. On the field more P was bound in grey alder per unit biomass compared to downy birch. The willows had more K per unit biomass than the other tree species, and the exotic willow species more N than the native ones. Less N, K and Mg were bound per unit biomass of S. phylicifolia compared to the other tree species.

• Hytönen, E-mail: jh@mm.unknown
• Rossi, E-mail: pr@mm.unknown
• Saarsalmi, E-mail: as@mm.unknown

The effects of repeated fertilizer treatment on biomass production and nutrient status of willow (Salix ’Aquatica’) plantations established on two cut-away peatland areas in western Finland were studied over a rotation period of three years. Comparisons were made between single fertilizer applications and repeated annual fertilization.

The annually repeated fertilizer application increased the amounts of acid ammonium acetate extractable phosphorus and potassium in the soil as well as the concentrations of foliar nitrogen, phosphorus and potassium compared to single application. Depending on the fertilizer treatment and application rate, annual fertilizer application resulted in over two times higher biomass production when compared to single fertilizer application over a three-year rotation period. The effect of phosphorus fertilizer application lasted longer than that of nitrogen. The optimum fertilization regime for biomass production requires that nitrogen fertilizer should be applied annually, but the effect of phosphorus can last at least over a rotation of three years. Potassium fertilizer treatment did not increase the yield in any of the experiments during the first three years. The leafless, above-ground yield of three-year-old, annually NP-fertilized willow plantations was 9.5 t ha^-1 and the total biomass, including stems, leaves, roots and the stump, averaged 17 t ha^-1.

• Hytönen, E-mail: jh@mm.unknown

The effects of fertilized treatment on the soil nutrient concentrations, biomass production and nutrient consumption of Salix x dasyclados and Salix ’Aquatica’ were studied in five experiments on three cut-away peatland sites in western and eastern Finland during three years. Factorial experiments with all combinations of N (100 kg ha^-1 a-¹⁾, P (30 kg ha^-1 a^-1) and K (80 kg ha^-1 a^-1) were conducted.

The application of P and K fertilizers increased the concentrations of corresponding extractable nutrients in the soil as well as in willow foliage. N-fertilization increased foliar nitrogen concentration. An increase in age usually led to decreases in bark and wood N, P and K concentrations and increases in bark Ca concentrations. N-fertilization increased the three-year biomass yield 1.5–2.7 times when compared to control plots. P-fertilization increased the yield only in those experimental fields whose substrates had the lowest phosphorus concentration. K-fertilization did not increase the yield in any of the experimental fields. The highest total biomass yield of NPK-fertilized willow after three growing seasons, 23 t ha^-1, was distributed in the following way: wood 42%, bark 19%, foliage 17%, stumps 6% and roots 16%. As the yield and stand age increased, more biomass was allocated in above-ground wood. Three-year-old stands (above-ground biomass 18 t ha^-1) contained as much as 196 kg N ha^-1, 26 kg P ha^-1, 101 kg K ha^-1, 74 kg Ca ha^-1 and 37 kg Mg ha ^-1. By far the highest proportion of nutrients accumulated in the foliage. The bark and wood contained relatively high proportions of calcium and phosphorus. With an increase in age and size, the amount of nitrogen and potassium bound in one dry-mass ton of willow biomass decreased while that of phosphorus remained unchanged.

• Hytönen, E-mail: jh@mm.unknown

For biomass forestry in the inland parts of Southern and Central Finland, the obvious choice of willow species is Salix myrsinifolia. However, selection of clones of indigenous species has not yet been completed and more research and selection is needed. In the Piipsanneva old peatland trial, indigenous species of willow, mostly clones of S. myrsinifolia and S. phylicifolia, were compared in terms of biomass production, coppicing, height growth and diameter distributions. In this trial, the mean annual biomass production was not particularly high; more important results were attained in the ranking of clones. The trial strengthens the hypothesis that, over the long-term, the biomass production of S. myrsinifolia is higher than that of S. phylicifolia. It was supposed that behind the highest yield there was a clone with uniform quality, one whose diameter distribution would be narrow and positively skewed. Comparisons of parameters of Weibull functions showed that the distributions of the best clones were wide, indicating that those clones use the whole growth space better than those with narrow distribution.

• Honkanen, E-mail: ah@mm.unknown

Growth and nutrition of 20 clones representing different species and interspecific hybrids of willows (Salix spp.) growing on an abandoned field were studied. There were highly significant differences between the clones as regards the survival, number of sprouts per stool, sprout mean height and diameter and stem biomass production per stool. The differences between the clones in the concentration of all nutrients in both the leaves and stems were highly significant. 

• Viherä-Aarnio, E-mail: av@mm.unknown
• Saarsalmi, E-mail: as@mm.unknown

Possibilities of developing suitable willow (Salix spp.) clones for short-rotation forestry on mined peatlands in the north-western area of Finland were studied in a field experiment in which 300 willow clones were tested during 1985–87. Most of the tested clones started to grow from cuttings on limed and fertilized peat soil. Salix viminalis L. clones of southern origin had a higher leafless above ground biomass production than the well adapted control clone and the local Finnish willows, but their winter hardiness was not satisfactory. The growth habit of some southern willows was also better than that of the control clone. It was also possible to select clones with good sprouting capacity. There were few Salix myrsinifolia Calisb. clones of Finnish origin, which had better cold tolerance than all other willows tested and higher biomass production than that of the control clone. The most critical factor to be selected for this is the optimal combination of winter hardiness and biomass production. This is attempted by selecting clones on the basis of this experiment for a breeding program.

The PDF includes a summary in Finnish.

• Lumme, E-mail: il@mm.unknown
• Törmälä, E-mail: tt@mm.unknown

The solubilityof various phosphorus and potassium compounds in a sedge peat soil was studied in an incubation experiment aimed at screening potential fertilizers for the cultivation of fast-growing willows. (KPO₃)n proved not to be a source of the slow-released K regarded as most desirable for this kind of cultivation as it was hydrolysed completely in the soil during incubation. Phosphorus from easily soluble or hydrolysable compounds (superphosphate, KH₂PO₄, (KPO₃)n) was bound in the soil largely by Al and Fe and elevated the level of readily soluble P considerably, whereas rock phosphates were found to be practically unchanged after incubation and did not contribute to the readily soluble P in the soil. Apatites proved to be quite insoluble and are therefore assumed to be unsuitable as P fertilizers for fast-growing willows, which have a high nutrient demand.

The PDF includes an abstract in Finnish.

• Lumme, E-mail: il@mm.unknown
• Yli-Halla, E-mail: my@mm.unknown

Salix 'Aquatica Gigantea', widely experimented and promising species for temperate zone short rotation forestry, has since 1950 recorded in Finland 23 times with different clone numbers. Salix x dasyclados Wimm., by morphological, cultivational and productivity characteristics similar willow has been recorded 16 times.

The nomenclature and origin of both willows have remained unclear in field research. Recent observations, based on morphological analyses and chromosome studies suggest that ’Aguatica gigantea’ and most S. x dasuclados clones can be collected under one Siberian species: Salix burjatica Nasarov. The true Salix x dasyclados Wimm. is a female hybrid S. x viminalis x cinerea, famous West-European basket willow that has been very little experimented in Finland.

The PDF includes an abstract in Finnish.

• Pohjonen, E-mail: vp@mm.unknown

The interactive effects of water stress and temperature on the CO₂ response of photosynthesis was studied in Salix sp. cv. Aquatica using the closed IRGA system. A semi-empirical model was used to describe the CO₂ response of photosynthesis. The interactive effect of water stress and temperature was divided into two components: the change in CO₂ conductance and the change in the photosynthetic capacity. The CO₂ conductance was not dependent on the temperature when the willow plant was well watered, but during water stress it decreased as the temperature increased. The photosynthetic capacity of the willow plant increased along with an increase in temperature when well-watered, but during water stress temperature had quite opposite effect.

The PDF includes a summary in Finnish.

• Smolander, E-mail: hs@mm.unknown
• Lappi, E-mail: jl@mm.unknown

Downy birch (Betula pubescens Ehrh.) trees growing on a drained peatland were cut during dormancy. The properties of the one-year old shoots produced by the stumps were measured in the autumn after one growing season. The one-year old willow shoots (a mixture of Salix phylicifolia L., S. pentandra L. and S. caprea L.) were collected from an abandoned field.

The basic density of unbarked shoots was 443 kg/m³ for birch and 346 kg/m³ for willow. The basic density of the bark was much higher than that of the wood. The effect of shoot length on the properties was small with the exception of cellular proportions. The fibre percentage increased and vessel percentage decreased with increasing shoot length.

The PDF includes a summary in Finnish.

• Bhat, E-mail: kb@mm.unknown
• Ferm, E-mail: af@mm.unknown
• Kärkkäinen, E-mail: mk@mm.unknown

The effect of spacing on the first-year yield and height increment of Alnus incana (L.) Moench, Populus tremula L. x Populus tremuloides Michx. (Populus x wettsteinii), Salix ’Aquatica Gigantea’, and Salix phylicifolia L. was studied at the Arctic Circle Agricultural Experimental Station in Northern Finland. S. ’Aquatica Gigantea’ gave yields which were twice as high as those of the other species in the study. The highest yields were of the order of 60 tons per hectare (fresh yield including foliage). The annual height growth in S. ’Aquatica Gigantea’ was about 100 cm, in the others about 30–50 cm. S. ’Aquatica Gigantea’ had a maximal height growth when the distance between the seedlings was 25 cm.

The PDF includes a summary in English.

• Pohjonen, E-mail: vp@mm.unknown

A review is made into the experiences and possibilities of willow breeding, with a particular consideration of their application under Finnish conditions.

In comparison with other tree genera, Salix has many advantages in breeding. The great number of species, large genetic variation between and within species and frequent crossability offer good possibilities for improvement by hybridization. The dioecious flowering, plentiful seed production and short generation cycle are valuable properties for breeding by controlled pollination. Individuals with desirable characteristics can be easily propagated vegetatively from cuttings. The propertied stated above are also characteristic of poplars, the breeding of which has resulted in great success. Corresponding results can be therefore expected from willow breeding, which has recently been initiated in several countries.

Today willows have no practical significance for forestry in Finland. There are, however, some suitable Salix species, both indigenous and exotic, for starting material for breeding, and plenty of favourable sites for willows. Therefore, the beginning of breeding research with willows should be well justified also in Finland.

The PDF includes a summary in English.

• Malmivaara, E-mail: em@mm.unknown
• Mikola, E-mail: jm@mm.unknown
• Palmberg, E-mail: cp@mm.unknown

There is demand for cultivated willow in toy and barrel making, and handcrafts in Finland. In Europe, Salix sp. have been cultivated on several kinds of sites. In Finland, suitable sites for willow plantations could be found, for instance, by the rivers and brooks, on other alluvial lands, wet meadows and other humid lands unsuitable for agriculture. The article suggests that the cultivation of willow should begin on the agricultural lands, because the choice of origin of the willow cuttings is less important when it is grown on fields, and the areas are more accessible. The desired qualities for cultivated willow are good growth rate and robust stems, frost resistance and resistance to annual cuttings. Of the different species, especially suitable is Salix sect. Amygdalinae. In field tests, all the tested Salix species have been frost resistant, which makes it possible to cut the stems annually.

The PDF includes a summary in German.

• Nordberg, E-mail: sn@mm.unknown

Willow (Salix sp.) species and clones have been selected in Finland, originally for basket willow husbandry, since 1910's. Screening for biomass willows started in 1973 by the Foundation for Forest Tree Breeding. Biomass willow research for energy started in 1978. The objective of the study was, based on theoretical background, on historical record of Finnish willow research between 1910–80 and on analysis of the Finnish biomass willow research of the 1980s, a further selection of exotic and indigenous willows for energy and chemicals.

Swedish selection of 63 exotics, mainly of Salix viminalis L. and S. burjatica Nazarov, was screened in Kompparnäs willow research site in the southern coast of Finland in 1983–89. S. viminalis showed both high yield potential and good crop certainty. The yield variation in S. burjatica were big due to rust (Melampsora sp.) infection followed by lowered winter hardiness. Three recommendable S. viminalis clones for Southern Finland were found.

Finnish indigenous species were screened based on collection (375 clones) in 1974–74 of the Foundation for Forest Tree Breeding, and the Finnish 4H-organization (566 clones) in 1978–79 in test sites in Suomusjärvi, Nurmijärvi, Kannus and Haapavesi. Salix myrsinifolia Salisb. was most productive of the indigenous willows. Five recommendable clones were selected. The second most productive indigenous was S. phylicifolia, with three recommendable clones. Based on willow hybridization studies in the Finnish Forest Research Institute, a considerable additional selection effect, boosted by heterosis, was found from the progenies. Further intraspecific crossings of geographically distant clones of S. myrsinifolia are recommended.

Based on the results, S. viminalis is recommended for practical biomass forestry application in the southernmost agroclimatic zone of Finland. S. myrsinifolia is recommended for further research and development in the other zones.

The PDF includes a summary in Finnish.

• Pohjonen, E-mail: vp@mm.unknown

• Püttsepp, Department of Ecology, Swedish University of Agricultural Sciences, P.O. Box 7072, SE-75007 Uppsala, Sweden; Estonian University of Life Sciences, Kreuzwaldi 64, Tartu 51014, Estonia E-mail: ulle.puttsepp@ekol.slu.se
• Lõhmus, Institute of Geography, University of Tartu, Vanemuise 46, Tartu 51014, Estonia E-mail: kl@nn.ee
• Koppel, Estonian University of Life Sciences, Kreuzwaldi 64, Tartu 51014, Estonia E-mail: ak@nn.ee