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Silva Fennica 1926-1997
Acta Forestalia Fennica

Articles containing the keyword 'branching'

Category: Article

article id 5632, category Article
Annikki Mäkelä, Veli-Pekka Ikonen, Petteri Vanninen. (1997). An application of process-based modelling to the development of branchiness in Scots pine. Silva Fennica vol. 31 no. 3 article id 5632.
Original keywords: puutavara; mänty; kasvu; hiili; läpimitta; biomassa; oksat; laatu; latvus; simulointi; kasvumallit; mallit; hiilitase; oksaisus
English keywords: Pinus sylvestris; carbon balance; simulation; pipe model; timber quality; growth model; branching; crown structure; whorl
Abstract | View details | Full text in PDF | Author Info

A process-oriented tree and stand growth model is extended to be applicable to the analysis of timber quality, and how it is influenced by silvicultural treatments. The tree-level model is based on the carbon balance and it incorporates the dynamics of five biomass variables as well as tree height, crown base, and breast height diameter. Allocation of carbon is based on the conservation of structural relationships, in particular, the pipe model. The pipe-model relationships are extended to the whorl level, but in order to avoid a 3-dimensional model of entire crown structure, the branch module is largely stochastic and aggregated. In model construction, a top-down hierarchy is used where at each step down, the upper level sets constraints for the lower level. Some advantages of this approach are model consistency and efficiency of calculations, but probably at the cost of reduced flexibility. The detailed structure related with the branching module is preliminary and will be improved when more data becomes available. Model parameters are identified for Scots pine (Pinus sylvestris L.) in Southern Finland, and example simulations are carried out to compare the development of quality characteristics in different stocking densities.

  • Mäkelä, E-mail: am@mm.unknown (email)
  • Ikonen, E-mail: vi@mm.unknown
  • Vanninen, E-mail: pv@mm.unknown
article id 5385, category Article
Timo Kuuluvainen. (1989). Branching dynamics in young Scots pine. Silva Fennica vol. 23 no. 3 article id 5385.
Keywords: Pinus sylvestris; biomass production; branching; tree architecture; bifuraction; modular construction; shoot demography
Abstract | View details | Full text in PDF | Author Info

The development of shoot number and shoot properties was examined in successive shoot cohorts of young widely-spaced Scots pine trees (Pinus sylvestris L.) growing in a progeny trial. This was accomplished by reconstructing the branching process of the trees over a period of five years, from tree age 4 to 8. During this time the number of shoots in successive shoot cohorts increased rapidly, while at the same time the mean shoot length decreased. The decrease in shoot lengths from older to younger shoots was accompanied by a decline in the bifuraction frequency of the shoots. In general, rapid changes occurred in the branching characteristics during the yearly development of the trees. The variation in the branching characteristics was reflected in the development of the architecture and biomass production of the trees.

The PDF includes an abstract in Finnish.

  • Kuuluvainen, E-mail: tk@mm.unknown (email)
article id 5347, category Article
Timo Kuuluvainen, Markku Kanninen, Juha-Pekka Salmi. (1988). Tree architecture in young Scots pine: properties, spatial distribution and relationships of components of tree architecture. Silva Fennica vol. 22 no. 2 article id 5347.
Keywords: Pinus sylvestris; crown architecture; biomass production; branching pattern; tree ideotype
Abstract | View details | Full text in PDF | Author Info

The architecture of Scots pine (Pinus sylvestris L.) was studied in an eight-year-old progeny test. The measurements included characteristics of crown structure, spatial distribution of shoots and yield components. The spatial distribution of shoots showed striking between-tree differences, and two extreme distribution patterns were detected. One represented a non-layered structure with a vertically relative even shoot distribution, and the other a layered structure with a vertically highly uneven shoot distribution.

Close correlations existed between several components of tree architecture and it is suggested that changes in the phenotypic architecture in Scots pine follow an epigenetic pattern, which enables the prediction of adaptational changes in structural components. The structural characteristics related to high above-ground biomass were a long crown, high total shoot length, high number of branches per whorl and big shoots of low needle density occupying a big share of the crown volume.

The PDF includes a summary in Finnish.

  • Kuuluvainen, E-mail: tk@mm.unknown (email)
  • Kanninen, E-mail: mk@mm.unknown
  • Salmi, E-mail: js@mm.unknown
article id 5049, category Article
Pirkko Ilonen, Pertti Hari, Markku Kanninen, Seppo Kellomäki. (1979). On distribution of growth in crown system of young Scots pine stands. Silva Fennica vol. 13 no. 4 article id 5049.
Keywords: Pinus sylvestris; light; Scots pine; needle; branching; terminal growth
Abstract | View details | Full text in PDF | Author Info

Branching and terminal growth of lateral shoots and needle growth of Scots pine (Pinus sylvestris L.) is investigated as a function of the whorl’s position and age and prevailing light climate. Number of buds per whorl was linearly and positively related to the whorl’s position and prevailing light climate. The growing whorl’s number counting from the apex was associated with declining bud number. The terminal growth of lateral shoots increased exponentially within the values 0.6–1.0 of the whorls position. Under these values the terminal growth was negligible. The growing whorl’s number indicated curvlinear decrease in shoot growth respectively, and only negligible growth occurred when the whorl’s age exceeded 10 years. The shoot growth was linearly related to the prevailing light climate but differences between dominating and dominated trees were apparent. The distribution of needle growth in the crown system was similar to that of shoot growth.

The PDF includes a summary in Finnish.

  • Ilonen, E-mail: pi@mm.unknown (email)
  • Hari, E-mail: ph@mm.unknown
  • Kanninen, E-mail: mk@mm.unknown
  • Kellomäki, E-mail: sk@mm.unknown
article id 4856, category Article
Pekka Rautiainen. (1971). Ympäristö- ja perintötekijöiden vaikutus männyn ilmiasuun Pohjois-Karjalan piirimetsälautakunnan siemenviljelyksessä Tohmajärvellä. Silva Fennica vol. 5 no. 4 article id 4856.
English title: The effect of environmental and genetical factors on the phenotype of Scots pine in a seed orchard in North Karelia.
Original keywords: mänty; oksikkuus; Suomi; periytyvyys; pituuskasvu; fenotyppi; heritabiliteetti; siemenviljelmä; Tohmajärvi; oksakulma
English keywords: Pinus sylvestris; Scots pine; height growth; heritability; branchiness; fenotyyppi; seed orchard; angle of branching
Abstract | View details | Full text in PDF | Author Info

The study material included 600 Scots pine (Pinus sylvestris L.) grafts from the Tohmajärvi seed orchard in Eastern Finland. Their broad sense heritability for the height growth was 0.92, for the number of branches 0.87 and for the angle of branching 0.84. Grafts from Central Finland had cones more often than the southern ones, the frequencies being 26.3% and 11.2%. It seems that dominance plays a significant role in the genetical variation of this seed orchard and that height growth is probably more rewarding breeding characteristic than quality, the difference being small, however.

The PDF includes a summary in English.

  • Rautiainen, E-mail: pr@mm.unknown (email)

Category: Article

article id 7372, category Article
Vilho Seppänen. (1942). Sahatukkien teosta aikatutkimuksen valossa. Acta Forestalia Fennica vol. 50 no. 19 article id 7372.
English title: Time studies on making of saw logs.
Original keywords: aikatutkimus; karsinta; kaato; hakkuutyö; kuoriminen; pölkytys; katkonta
English keywords: saw logs; time study; felling; branching; felling work; barking; cross-cutting
Abstract | View details | Full text in PDF | Author Info

A time study was conducted in saw log harvesting site in state forests of Evo in Southern Finland in 1934. Felling was performed in teams of two loggers. Two teams were observed. The work was divided into several stages of work: felling, branching, cross-cutting, barking and making of top log. On the site grew Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.).

The daily working hours not including breaks was in average 5 hours and 33 minutes. The most time-consuming stage of the work was barking of the stem (55% of working time for Scots pine and 47% for Norway spruce), followed by felling (22.5% for pine and 19.4% for spruce), branching (11.7% and 21.6%) and cross-cutting (11.3% and 11.8%). Temperature affects barking strongly. Scots pine is slower to bark than Norway spruce. Similarly, butt and middle logs are slower to bark than top logs. It took in average 79.02 min to process one solid m3 of timber with bark and 91.45 min without bark.

The PDF includes a summary in German.

  • Seppänen, E-mail: vs@mm.unknown (email)
article id 7532, category Article
August Renvall. (1914). Paper on knowledge about so-called partial variability of pine. Acta Forestalia Fennica vol. 3 no. 2 article id 7532.
Keywords: needles; branching; shoots; inflorescence; Wilhelm Johannsen; Pinus silvestris L. var. lapponica (Fr.) Hn.
Abstract | View details | Full text in PDF | Author Info

The article deals with outer characters of a pine from Patsjoki-river, in Finnish Lapland (Pinus silvestris L. var. lapponica (Fr.) Hn.). The tables describe the length of the needles, length of the shoots, branching and inflorescence. The statistical calculations of the data are based on W. Johannsen’s  (Elemente der exakten Erblichkeitslehre, Jena 1909)  (Elements of exact genetics) methods. The results cannot be generalized because of the insufficient amount of data. 

  • Renvall, E-mail: ar@mm.unknown (email)

Category: Research article

article id 252, category Research article
Tuomo Kalliokoski, Pekka Nygren, Risto Sievänen. (2008). Coarse root architecture of three boreal tree species growing in mixed stands. Silva Fennica vol. 42 no. 2 article id 252.
Keywords: Pinus sylvestris; Betula pendula; Picea abies; site type; branching pattern; influence area; developmental stage
Abstract | View details | Full text in PDF | Author Info
Root system architecture determines many of the vital functions of a tree, e.g. stability of anchorage and resource uptake. The shoot:root ratio is determined through the allocation of resources. Studies on below-ground architectural elements in boreal mixed forests are relatively scarce despite the fact that knowledge on below-ground interactions and allocation changes in relation to stand developmental stage and soil fertility is needed both in ecological and silvicultural research. In this study, sixty tree root systems of three different tree species, Betula pendula, Picea abies and Pinus sylvestris, were excavated in five mixed forest stands in order to quantify differences between the species and sites in terms of rooting behaviour. Root architecture differed greatly between the species, implying different solutions for the functions of root systems. Half of the P. sylvestris had developed a taproot as a response to anchorage needs, while P. abies correspondingly had pronounced secondary growth of proximal roots. Betula pendula had the most extensive root system, illustrating the greater demand of deciduous trees for water. Betula pendula was also the most sensitive to soil fertility: it favoured exploration on the poorest site, as illustrated by the high total root length, whereas on the most fertile site its strategy was to efficiently exploit soil resources through increased branching intensity. The results obtained in this study provide basic knowledge on the architectural characteristics of boreal tree root systems for use by forestry professionals and modellers.
  • Kalliokoski, The Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: (email)
  • Nygren, Department of Forest Ecology, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail:
  • Sievänen, The Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail:

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