Articles containing the keyword 'crown'

A process-oriented tree and stand growth model is extended to be applicable to the analysis of timber quality, and how it is influenced by silvicultural treatments. The tree-level model is based on the carbon balance and it incorporates the dynamics of five biomass variables as well as tree height, crown base, and breast height diameter. Allocation of carbon is based on the conservation of structural relationships, in particular, the pipe model. The pipe-model relationships are extended to the whorl level, but in order to avoid a 3-dimensional model of entire crown structure, the branch module is largely stochastic and aggregated. In model construction, a top-down hierarchy is used where at each step down, the upper level sets constraints for the lower level. Some advantages of this approach are model consistency and efficiency of calculations, but probably at the cost of reduced flexibility. The detailed structure related with the branching module is preliminary and will be improved when more data becomes available. Model parameters are identified for Scots pine (Pinus sylvestris L.) in Southern Finland, and example simulations are carried out to compare the development of quality characteristics in different stocking densities.

• Mäkelä, E-mail: am@mm.unknown
• Ikonen, E-mail: vi@mm.unknown
• Vanninen, E-mail: pv@mm.unknown

A method for the measurement of the three-dimensional (3D) architecture of trees was applied to describe two 20-year-old walnut trees, one of them is a timber tree while the other is a fruit tree. The method works at the shoot level and simultaneously describes the plant topology, the plant geometry and the shoot morphology. The method uses a 3D digitiser (3SPACE® FASTRAK®, Polhemus Inc.) associated with software DiplAmi designed for digitiser control and data acquisition management. Plant images may be reconstructed from the data set by using the ray tracing software POV-Ray. Visual comparison between photographs of the walnut trees and images synthesised from digitising was satisfactory. Distribution of basal shoot diameter, as well as leaf area and fruit distributions for both the timber and the fruit tree were non-uniformly distributed in the crown volume. Gradients were likely to be related to the light distribution within the tree. This is in agreement with previous experimental results on several tree species, and also with the predictions of tree architecture models based on light-vegetation interactions.

• Sinoquet, E-mail: hs@mm.unknown
• Godin, E-mail: cg@mm.unknown
• Rivet, E-mail: pr@mm.unknown

Variation in needle nutrient concentrations with age and vertical location in the crown was studied in three Scots pine (Pinus sylvestris L.) stands growing on peat soils in Eastern Finland. The concentrations of N, P, Fe and Zn decreased down the crown and those of Ca and Mn increased. Potassium and magnesium concentration patterns differed between sites.

Potassium and Mg concentrations were highest in the current needles at all heights in the crown, iron and manganese concentrations were highest in the oldest needles. The concentrations of N, P and Zn did not vary with needle age.

• Finér, E-mail: lf@mm.unknown

There is some evidence to support the hypothesis that crown form of Scots pine (Pinus sylvestris L.) can be inherited either mono- or polygenically. In Finland, a narrow, horizontal whorl-layer structure is striking in about one half of the offspring of a genotype E 1101, ”Kanerva pine”. The offspring are characterized by a narrow crown, short and thin branches at an angle of 90° to the stem, minimal tapering and by numerous long, lateral shoots, long needles and the common occurrence of three-needled fascicles among the dwarf shoots. These features are connected to a high growth rate, a high harvest index and to tortuosity of the stem. It is suggested that this complex of characters is determined by a single dominant gene.

In this study, several offspring of the E 1101 were classified into three form types in seven sets of progeny test data, including progenies of various ages having E 1101 as either maternal or paternal parent as well as open pollinated progenies of second-generation offspring. A segregation close to 1:1 was found both in the first and in the second-generation progenies when wilds and intermediates were combined and compared with Kanervas. The result indicates that the three types of Kanerva form can be due to a single dominant allele (K). Kanervas are heterozygous (Kk) for the allele and wilds are recessive homozygotes (kk) resulting 1:1 segregation in their progenies. However, there were also remarkable deviations from the expected distribution. The differences as well as the inheritance pattern are discussed.

The PDF includes an abstract in Finnish.

• Pöykkö, E-mail: tp@mm.unknown
• Velling, E-mail: pv@mm.unknown

A range of different indices are available for assessing the health of trees in forests. An even larger range can be used for the assessment of the health of forest ecosystems. Most studies made in connection with ”forest decline” and the impact of air pollution and other environmental stresses on forests have concentrated on the assessment of crown transparency and crown discoloration in individual trees. These are non-specific indicators which are now known to be sometimes of relatively little value when determining the health of a forest ecosystem. Numerous problems exist with both, and the standardisation of assessments between and even within countries has not been achieved. Consequently, studies claiming to compare ”defoliation” between different countries cannot be substantiated. The emphasis on crown transparency and crown discoloration has resulted in the neglect of a number of other indices that could be of considerable value. These include a variety of visual measures of crown condition and also several non-visual bioindicators. Some of these techniques are objective, reducing the present reliance on observed standardization. A large number of potential techniques are currently at the research stage and have yet to be adequately tested in field trials. This represent an area where a substantial amount of further research is required.

• Innes, E-mail: ji@mm.unknown

Crown characteristics and the distribution of three years’ (1986–88) biomass production of 20 pendulous Norway spruce (Picea abies f. pendula (Lawson) Sylvén) trees with heritable narrow crown and 15 normal-growned spruces (Picea abies (L.) H. Karst.) were studied in a 19-year-old mixed stand.

The form of the crown is conical in normal-crowned trees, columnar and narrow in pendulous trees. The partitioning of aboveground biomass to stems during the studied 3-year period was significantly higher in pendulous (0.281) than in normal-crowned trees (0.255) and also the ratio between growth of stemwood and growth of needle biomass during three years was higher in pendulous trees (0.67 g g^-1) than in normal-crowned trees (0.52 g g^-1). The needle biomass was distributed higher in the crown in pendula than in normal-crowned trees and they had a higher needle biomass/branchwood biomass ratio than normal trees. The difference in harvest increment between the two crown types are mostly due to the significantly lower branchwood biomass values in pendulous than in normal-crowned trees. The higher needle ’efficiency’ in pendulous trees is probably connected with high partitioning of needle biomass to the upper part of the crown in pendulous trees.

The PDF includes an abstract in Finnish.

• Pulkkinen, E-mail: pp@mm.unknown

The mating system was analysed in the upper and lower crown of two groups of European larch (Larix decidua Mill.) clones divided according to the percentage of full seeds in the upper and lower crown parts. The overall multilocus estimate of outcrossing rate (t) was calculated to be 0.929. The differences of outcrossing rates between crown levels and clonal groups respectively were not statistically significant. The t estimates were greater for the upper crown level and for clones with higher percentage of full seeds in the upper crown level. However, among all observations there was no correlation between outcrossing rates and percentages of full seeds for particular crown levels and groups of clones. Observed similarity of outcrossing among grafts of the same clones may indicate genetic control of self-fertilization rate in individual European larch. 

The PDF includes an abstract in Finnish.

• Burczyk, E-mail: jb@mm.unknown
• Kosiński, E-mail: gk@mm.unknown
• Lewandowski, E-mail: al@mm.unknown

The architecture of Scots pine (Pinus sylvestris L.) was studied in an eight-year-old progeny test. The measurements included characteristics of crown structure, spatial distribution of shoots and yield components. The spatial distribution of shoots showed striking between-tree differences, and two extreme distribution patterns were detected. One represented a non-layered structure with a vertically relative even shoot distribution, and the other a layered structure with a vertically highly uneven shoot distribution.

Close correlations existed between several components of tree architecture and it is suggested that changes in the phenotypic architecture in Scots pine follow an epigenetic pattern, which enables the prediction of adaptational changes in structural components. The structural characteristics related to high above-ground biomass were a long crown, high total shoot length, high number of branches per whorl and big shoots of low needle density occupying a big share of the crown volume.

The PDF includes a summary in Finnish.

• Kuuluvainen, E-mail: tk@mm.unknown
• Kanninen, E-mail: mk@mm.unknown
• Salmi, E-mail: js@mm.unknown

The utilization of direct radiation was studied in five model stands of Poisson-type tree distribution and cone-shaped crowns. The radiation extinction depended on the self-shading of the crown and the shading caused by other trees. The results indicate that at low sun elevation a stand populated by very narrow-crowned trees is most effective in light interception and photosynthesis. At high sun elevation a broad-crowned canopy is best illuminated and most favourable for photosynthesis. A stand with a two-storey canopy is effective in all latitudes when the crowns are moderately narrow. In two-storey canopies the foliage of the lower storey can be better illuminated than in the lower parts of the upper storey, because of the smaller self-shading in the small crowns of the lower storey. A canopy where the crown volume is concentrated on few big crowns is less effective than a canopy consisting of many small crowns.

The PDF includes an abstract in Finnish.

• Pukkala, E-mail: tp@mm.unknown
• Kuuluvainen, E-mail: tk@mm.unknown

A study based on four young Scots pines (Pinus sylvestris L.) showed that the number of needle-covered shoots per crown volume unit was independent on tree position representing a constant value of 600–700 shoots/m³. This was true, even though the total shoot number decreased with deteriorating tree position. In tree crown there were fourth-order shoots in good light conditions but only first- and second-order-shoots, when light conditions were poor. The length of shoots decreased in accordance with increasing order of the shoot.

The share of the needle biomass and growth increased, when the shoot order increased. Similarly, the share of needles increased with deteriorating tree position. This was especially true in the upper crown. On the other hand, the share of the crown from the total biomass and growth increased with improving tree position. The percentage of crown system of a dominant tree in a sparse stand was 64% of that of biomass and 83% of that of growth. The corresponding values for a suppressed tree in a dense stand were 36% and 35%. The growth of wood, bark and needles in crown systems was linearly correlated with prevailing light conditions around the branch. It is evident that the tree position and light condition within the stand control the wood, bark and needle growth in the crown system and their interrelationships.

The PDF includes a summary in English.

• Viherä, E-mail: av@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown

The needle area distribution and crown structure of a young planted Scots pine (Pinus sylvestris L.) stand are described. The crown structure and crown shape showed apparent regularity in crown structure regardless of stand dynamics. Similarly, the shoot structure and individual needle area showed regularity in the number of needles per branch and shoot length unit, and consequent phytoarea density inside the needle cylinder. Also, the shoot area and needle area distributions were found to show a regular distribution of needle biomass throughout the crown, also inside the crown, in the dominant trees. In the suppressed trees the needle biomass was located in the upper crown and on the surface area of the crown. Estimates of the canopy needle area and distributions are given. The results were applied in calculations of the within-stand light regime. The results correlated well with the empirical results.

The PDF includes a summary in Finnish.

• Kellomäki, E-mail: sk@mm.unknown
• Oker-Blom, E-mail: po@mm.unknown

The crown structure and stem growth of Norway spruce (Picea abies (L.) H. Karst.) undergrowth was studied in relation to the prevailing light conditions and potential photosynthesis. Shading decreased the stem height growth more than the length increment of laterals, producing a plate-shaped crown in deep shade. Needles responded to shading by adopting a horizontal inclination in deep shade. The needles were wide and thin respectively in shade. In the open the needle cross-section was almost square. Stem radial growth and height growth were both affected by shading exhibiting a linear response to the prevailing light conditions and the potential photosynthesis. Light conditions under dominating trees were closely correlated with the basal area of the dominating trees.

The PDF includes a summary in Finnish.

• Greis, E-mail: ig@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown

The specific needle area of young Scots pine (Pinus sylvestris L.) showed a substantial within-tree and between-tree variation which was associated with the position of the tree and the position of the whorl as indicated by the prevailing crown and branch illumination. In suppressed trees the values of the specific needle area were three to four times those in dominating trees. A similar morphogenesis was discernible in comparison of the lower and the upper part of the crown. The mean specific needle area value for the whole stand was 184 cm²/g.

The PDF includes a summary in Finnish.

• Kellomäki, E-mail: sk@mm.unknown
• Oker-Blom, E-mail: po@mm.unknown

Crown and stem growth of young Scots pines (Pinus sylvestris L.) were studied in relation to photosynthate supply and light condition in a stand. The magnitude of needle and bud formation, and radial and height growth were to a great extent dependent on the photosynthate supply. However, in shaded conditions the growth of each characteristics was greater than expected on the basis of photosynthate supply. In the stem system this was especially apparent for height growth. Consequently, height growth was favoured at the expense of radial growth in shaded conditions. It also appeared that the basic density of wood was negatively related to both tree position and photosynthate supply.

The PDF includes a summary in Finnish.

• Kellomäki, E-mail: sk@mm.unknown
• Kanninen, E-mail: mk@mm.unknown

The technique of double normalizing, i.e. normalizing the relative needle biomass and the length of the living crown system, is applied to the modelling of the distribution of needle biomass in the canopy of young Scots pine (Pinus sylvestris L.) stands. The study based on the parameters of β-function shows that at the individual-tree level, the variance in needle distribution was not closely associated with any tree characteristics. A shift in the point of maximum needle biomass upwards unsuppressed trees was, however, evident. This was associated with an increase in the height of the trees. At the stand level, the stand mean height and stand density had an equal and a rather high potential for explaining the variance in the needle distribution. The normalized crowns are utilized in models for determining light extinction in the crown. A special technique for determining the amount of photosynthates available for growth in a particular tree is presented.

The PDF includes a summary in Finnish.

• Kellomäki, E-mail: sk@mm.unknown
• Hari, E-mail: ph@mm.unknown
• Ilonen, E-mail: pi@mm.unknown
• Kanninen, E-mail: mk@mm.unknown

The Forest Research Institute of Finland has established permanent sample plots to survey the effect of thinnings on the stands. This study compares the development of tended and natural Scots pine (Pinus sylvestris L.) stands growing on three different forest types: Oxalis-Myrtillus, Vaccinium and Calluna site type. The effect of heavy thinning from below (Oxalis-Myrtillus and Vaccinium site types) and increment felling (Calluna site type) was assessed by dividing the trees of the stands in tree classification classes according to their crown storey and defects.

The results show that thinning from below and increment thinning increase the proportion of trees in the 1st crown storey, which is already large in the natural stands. Also the diameter distribution is more even and the mean diameter higher after the thinnings.

In Scots pine stands in natural state, volume increment per stem is highest in the 1st crown storey and diminishes strongly towards the lower crown storeys. Thinnings increased the increment. The study indicates that many of the objectives of the intermediate cuttings, including promoting the growth of the best trees and improving the quality of the stand, have in general been achieved. Consequently, the thinnings give means to achieve the most valuable yield in the stand.

The article includes a summary in English.

• Nyyssönen, E-mail: an@mm.unknown

Finnish tree species have adapted differently to heavy snow loads that occur especially in fell areas in Kuusamo and Salla as well as Maanselkä area in Sotkamo and Rautavaara in Northern Finland. Norway spruce (Picea abies Karst. L) is adapted better than Scots pine (Pinus sylvestris L.). The aim of this study was to investigate how crown and stem form of Norway spruce in the snow damage area of Maanselkä area differ from other areas in the same region.  

Relatively broad crown at the base of the stem, quickly tapering crown and narrow and even upper crown were typical for trees growing in the snow damaged areas. The higher the altitude is, the stronger tapering the crown is. The tapering begins usually in a height of 4-5 meters. Even the stem diameter begins to taper strongly at this height. In the areas where heavy snow does not cause snow damage, top of crown is broader. Also, in the snow damage areas the damaged trees seem to have broader crown shape than the trees with little damages.  

Height of the trees decreases in the snow damage areas compared to forests in lower altitudes, which can be caused both by wind and snow load. 

The article includes a German summary. 

• Mikola, E-mail: pm@mm.unknown

Norway spruce (Picea abies (L.) Karst.) invading sites is common in Finland. The species tends to establish itself as undergrowth, and takes over when it gets space to grow. To determine whether the undergrowth is suitable as the new generation requires knowledge on the biology of spruce undergrowth. One of the issues is determining the age of the stunted trees. In this investigation, 100 undergrown spruce trees, their crown and their root systems were studied. A method was developed to determine the age of the trees.

The root system of all trees in Vaccinium sites and of stunted trees in Myrtillius sites were superficial. The root systems of older spruces were purely of adventitious origin. The longer the period of stunting growth, the younger is the root system. In addition to acropetal and general adventitious ramification there is often adventitious branching of the roots of pathological causes. Mortality among the long roots is frequent.

A stunted tree has not the same ability as a viable tree to make use of already existing branches for building assimilating surface. When comparing trees with equally large assimilating surface, a stunted tree had greater sum of roots compared to a viable tree. The root system of a stunted undergrown spruce was very superficial compared with the other trees.

The PDF includes a summary in English.

• Sirén, E-mail: gs@mm.unknown

The aim of the study was to find out what are the causes of damage in Scots pine (Pinus sylvestris L.) stands and the frequency of different kinds of injuries, which are then discussed in relation to the silvicultural state and management of the stands in comparison to ideal forests. Sample plots were studied in over 80-year old Scots pine dominated stands in mineral soil sites of different forest types in Northern Finland in the area of Perä-Pohjola. 10–40 trees were chosen as sample trees in each sample plot. The sample trees were felled, and the diameter, height of crown and injuries outside and inside of the stem were recorded.

Length of knot-free part of the stem was higher in the dominant trees and in older age classes of the trees. The form of the stem becomes broader and rounder with the age. The crowns are, however, longer in Northern Finland compared to Southern Finland. In management of Scots pine stands, all trees diseased by Scots pine blister rust (Cronartium flaccidum) should be removed. The disease is common in Northern Finland, and the number of diseased trees increases as the stands get older. Decay was more common in trees that had fire wounds. In general, injuries decreased the length and diameter growth of the trees. From the dominant trees should only injured and diseased trees removed in the thinnigs. Codominant trees can be left to grow when spare trees are needed to replace missing dominant trees. Detailed instruction of selection of the removed trees are given for each age class.

The PDF includes a summary in German.

• Tikka, E-mail: pt@mm.unknown

The growth of a tree is influenced by inherited properties and external circumstances, including climate, soil, the position of the tree in the stand, and the position of the wood in the stem. The tree species have optimum climate and optimum conditions. The aim of this study was to determine if the summerwood content of the wood of Scots pine (Pinus sylvestris L.) is dependent on the rate of growth of the tree. Comparing the position of the sample trees in the stand, it seems that the position of the tree and the size of its crown influences strongly the quality of the wood. In a dense stand the summerwood content was higher in the trees that had small crowns. Thinning of the stand decreased the difference in summerwood content of the trees.

The PDF includes a summary in English.

• Jalava, E-mail: mj@mm.unknown

Doctor Lauri Ilvessalo modified tree classification developed by professor Gunnar Schotte to develop a tree classification and thinning system that suited Finnish conditions. His system was first applied when the Finnish Forest Research Institute began thinning experiments in a large scale in 1924. The system distinguishes four crown storeys: the predominating crown storey, dominated crown storey, emergent trees and undergrowth. Into the predominating storeys belong dominant trees and co-dominant trees, and into the dominated storeys the intermediate and ground trees. The trees in all the storeys can be divided in to normal trees with well-formed crown and stem, trees with defectively developed crown, trees with defective stem, and injured and diseased trees. The article describes different thinning methods (cleaning thinning, selective thinning from below, selective thinning from above, increment felling, freeing felling) using the tree-classification.

The volume 34 of Acta Forestalia Fennica is a jubileum publication of professor Aimo Kaarlo Cajander. The PDF includes a summary in English.

• Ilvessalo, E-mail: li@mm.unknown

Crown class is useful tool both in forest management and forest mensuration. The study presents a detailed crown classification for Scots pine (Pinus sylvestris L.). It was used to classify the sample trees prior detailed measurements of the crown and stem form. The stem form of a tree was dependent on which canopy layer it belonged. This relation was detected on both Vaccinium and Calluna site type forests. In addition, the stem tapers faster in poorer forest site types compared to better sites. The shorter the self-pruned part of the stem is, the faster the stem tapers. According to the study, the stems of stunted trees taper faster than trees of other crown classes. Also the age group affects stem form.

The PDF includes a summary in German.

• Lakari, E-mail: ol@mm.unknown

To enhance the utilization of the wood, the sawmills are forced to place more emphasis on planning to master the whole production chain from the forest to the end product. One significant obstacle to integrating the forest-sawmill-market production chain is the lack of appropriate information about forest stands. Since the wood procurement point of view in forest planning systems has been almost totally disregarded there has been a great need to develop an easy and efficient pre-harvest measurement method, allowing separate measurement of stands prior to harvesting. The main purpose of this study was to develop a measurement method for Scots pine (Pinus sylvestris L.) stands which forest managers could use in describing the properties of the standing trees for sawing production planning.

Study materials were collected from ten Scots pine stands located in North Häme and South Pohjanmaa, in Southern Finland. The data comprise test sawing data on 314 pine stems, diameter at breast height (dbh) and height measures of all trees and measures of the quality parameters of pine sawlog stems in all ten study stands as well as the locations of all trees in six stands. The study was divided into four sub-studies which deal with pine quality prediction, construction of diameter and dead branch height distributions, sampling designs and applying height and crown height models. The final proposal for the pre-harvest measurement method is a synthesis of the individual sub-studies.

Quality analysis resulted in choosing dbh, distance from stump height to the first dead branch, crown height and tree height as the most appropriate quality characteristics of Scots pine. Dbh and dead branch height are measured from each pine sample tree while height and crown height are derived from dbh measures by aid of mixed height and crown height models. Pine and spruce diameter distribution as well as dead branch height distribution are most effectively predicted by the kernel function. Roughly 25 sample trees seem to be appropriate in pure pine stands. In mixed stands the number of sample trees needs to be increased in proportion to the intensity of pines in order to attain the same level of accuracy.

• Uusitalo, E-mail: ju@mm.unknown

Diameter and volume increment as well as change in stem form of Scots pine (Pinus sylvestris L.) were analysed to predict tree increment variables. A stem curve set model is presented, based on prediction of the diameters at fixed angles in a polar coordinate system. This model consists of three elementary stem curves: 1) with bark, 2) without bark, and 3) without bark five years earlier. The differences between the elementary stem curves are the bark curve and the increment curve. The error variances at fixed angles and covariances between the fixed angles are divided into between-stand and within-stand components. Using principal components, the between-stand and within-stand covariance matrices are condensed separately for stem curve with bark, bark curve and increment curve. The two first principal components of the bark curve describe the vertical change in Scots pine bark type and the first principal component of the increment curve describes the increment rate. The elementary stem curves, bark curve and increment curve as well as corresponding stem volumes, bark volume and volume increment can be predicted for all trees in the stand with free choice of sample tree measurements. When only a few sample trees are measured, the stem curve set model gives significantly more accurate predictions of bark volume and volume increment for tally trees than does the volume method, which is based on the differences between two independent predictions of volume. The volume increment of tally trees can be predicted as reliably with as without measurement of sample tree height increment.

The PDF includes a summary in English.

• Ojansuu, E-mail: ro@mm.unknown

• Korpela, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland 0000-0002-1665-3984 E-mail: ilkka.korpela@helsinki.fi
• Polvivaara, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: –
• Papunen, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland 0000-0001-5383-4314 E-mail: saija.papunen@outlook.com
• Jaakkola, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: laura.jaakkola@helsinki.fi
• Tienaho, University of Eastern Finland, Faculty of Science and Forestry, P.O. Box 111, FI-80101 Joensuu, Finland 0000-0002-6574-5797 E-mail: noora.tienaho@uef.fi
• Uotila, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: johannes.uotila@helsinki.fi
• Puputti, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland 0000-0003-1972-1636 E-mail: tuomas.puputti@helsinki.fi
• Flyktman, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland 0000-0002-5235-317X E-mail: aleksi.flyktman@helsinki.fi

Terrestrial laser scanning (TLS) has been applied to estimate forest wood volume based on detailed 3D tree reconstructions from point cloud data. However, sources of uncertainties in the point cloud data (alignment and scattering errors, occlusion, foliage...) and the reconstruction algorithm type and parameterisation are known to affect the reconstruction, especially around finer branches. To better understand the impacts of these uncertainties on the accuracy of TLS-derived woody volume, high-quality TLS scans were collected in leaf-off conditions prior to destructive harvesting of two forest-grown common ash trees (Fraxinus excelsior L.; diameter at breast height ~28 cm, woody volume of 732 and 868 L). We manually measured branch diameters at 265 locations in these trees. Estimates of branch diameters and tree volume from Quantitative Structure Models (QSM) were compared with these manual measurements. The accuracy of QSM branch diameter estimates decreased with smaller branch diameters. Tree woody volume was overestimated (+336 L and +392 L) in both trees. Branches measuring < 5 cm in diameter accounted for 80% and 83% of this overestimation respectively. Filtering for scattering errors or improved coregistration approximately halved the overestimation. Range filtering and modified scanning layouts had mixed effects. The small branch overestimations originated primarily in limitations in scanner characteristics and coregistration errors rather than suboptimal QSM parameterisation. For TLS-derived estimates of tree volume, a higher quality point cloud allows smaller branches to be accurately reconstructed. Additional experiments need to elucidate if these results can be generalised beyond the setup of this study.

• Demol, CAVElab – Computational and Applied Vegetation Ecology, Department of Environment, Faculty of Bioscience Engineering, Ghent University, Coupure Links 653, B-9000 Ghent, Belgium; PLECO – Plants and Ecosystems, Faculty of Science, Antwerp University, Universiteitsplein 1, B-2610 Wilrijk, Belgium https://orcid.org/0000-0002-5492-2874 E-mail: miro.demol@ugent.be
• Wilkes, UCL Department of Geography, Gower Street, London WC1E 6BT, UK; NERC National Centre for Earth Observation (NCEO), UK https://orcid.org/0000-0001-6048-536X E-mail: p.wilkes@ucl.ac.uk
• Raumonen, Mathematics, Tampere University, FI-33101 Tampere, Finland https://orcid.org/0000-0001-5471-0970 E-mail: pasi.raumonen@tuni.fi
• Krishna Moorthy, CAVElab – Computational and Applied Vegetation Ecology, Department of Environment, Faculty of Bioscience Engineering, Ghent University, Coupure Links 653, B-9000 Ghent, Belgium https://orcid.org/0000-0002-6838-2880 E-mail: Sruthi.KrishnaMoorthyParvathi@ugent.be
• Calders, CAVElab – Computational and Applied Vegetation Ecology, Department of Environment, Faculty of Bioscience Engineering, Ghent University, Coupure Links 653, B-9000 Ghent, Belgium https://orcid.org/0000-0002-4562-2538 E-mail: kim.calders@ugent.be
• Gielen, PLECO – Plants and Ecosystems, Faculty of Science, Antwerp University, Universiteitsplein 1, B-2610 Wilrijk, Belgium https://orcid.org/0000-0002-4890-3060 E-mail: bert.gielen@uantwerpen.be
• Verbeeck, CAVElab – Computational and Applied Vegetation Ecology, Department of Environment, Faculty of Bioscience Engineering, Ghent University, Coupure Links 653, B-9000 Ghent, Belgium https://orcid.org/0000-0003-1490-0168 E-mail: hans.verbeeck@ugent.be

Airborne laser scanning (ALS) data is nowadays often available for forest inventory purposes, but adequate field data for constructing new forest attribute models for each area may be lacking. Thus there is a need to study the transferability of existing ALS-based models among different inventory areas. The objective of our study was to apply ALS-based mixed models to estimate the diameter, height and crown base height of individual sawlog sized Scots pines (Pinus sylvestris L.) at three different inventory sites in eastern Finland. Different ALS sensors and acquisition parameters were used at each site. Multivariate mixed-effects models were fitted at one site and the models were validated at two independent test sites. Validation was carried out by applying the fixed parts of the mixed models as such, and by calibrating them using 1–3 sample trees per plot. The results showed that the relative RMSEs of the predictions were 1.2–6.5 percent points larger at the test sites compared to the training site. Systematic errors of 2.4–6.2 percent points also emerged at the test sites. However, both the RMSEs and the systematic errors decreased with calibration. The results showed that mixed-effects models of individual tree attributes can be successfully transferred and calibrated to other ALS inventory areas in a level of accuracy that appears suitable for practical applications.

• Korhonen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland http://orcid.org/0000-0002-9352-0114 E-mail: lauri.korhonen@uef.fi
• Repola, Natural Resources Institute of Finland (Luke), Natural resources, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: jaakko.repola@luke.fi
• Karjalainen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: tomikar@uef.fi
• Packalen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: petteri.packalen@uef.fi
• Maltamo, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: matti.maltamo@uef.fi

This study examines the alternatives to include crown base height (CBH) predictions in operational forest inventories based on airborne laser scanning (ALS) data. We studied 265 field sample plots in a strongly pine-dominated area in northeastern Finland. The CBH prediction alternatives used area-based metrics of sparse ALS data to produce this attribute by means of: 1) Tree-level imputation based on the k-nearest neighbor (k-nn) method and full field-measured tree lists including CBH observations as reference data; 2) Tree-level mixed-effects model (LME) prediction based on tree diameter (DBH) and height and ALS metrics as predictors of the models; 3) Plot-level prediction based on analyzing the computational geometry and topology of the ALS point clouds; and 4) Plot-level regression analysis using average CBH observations of the plots for model fitting. The results showed that all of the methods predicted CBH with an accuracy of 1–1.5 m. The plot-level regression model was the most accurate alternative, although alternatives producing tree-level information may be more interesting for inventories aiming at forest management planning. For this purpose, k-nn approach is promising and it only requires that field measurements of CBH is added to the tree lists used as reference data. Alternatively, the LME-approach produced good results especially in the case of dominant trees.

• Maltamo, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: matti.maltamo@uef.fi
• Karjalainen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: tomimkarjalainen@gmail.com
• Repola, Natural Resources Institute of Finland (Luke), Natural resources, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: jaakko.repola@luke.fi
• Vauhkonen, Natural Resources Institute of Finland (Luke), Bioeconomy and environment, Yliopistokatu 6, 80100 Joensuu, Finland E-mail: jari.vauhkonen@luke.fi

The reach of different tree species’ crowns and the velocity of gap closure during the occupation of canopy gaps resulting from mortality and thinning during stand development determine species-specific competition and productivity within forest stands. However, classical dendrometric methods are rather inaccurate or even incapable of time- and cost-effectively measuring 3D tree structure, crown dynamics and space occupation non-destructively. Therefore, we applied terrestrial laser scanning (TLS) in order to measure the structural dynamics at tree and stand level from gap cutting in 2006 until 2012 in pure and mixed stands of Norway spruce (Picea abies [L.] Karst.) and European beech (Fagus sylvatica L.). In conclusion, our results suggest that Norway spruce invests newly available above-ground resources primarily into DBH as well as biomass growth and indicate a stronger resilience against loss of crown mass induced by mechanical damage. European beech showed a vastly different reaction, investing gains from additional above-ground resources primarily into faster occupation of canopy space. Whether our sample trees were located in pure or mixed groups around the gaps had no significant impact on their behavior during the years after gap cutting.

• Bayer, Address Technical University of Munich (TUM), Chair for Forest Growth and Yield Science, Hans-Carl-von-Carlowitz-Platz 2, 85354, Freising, Germany http://orcid.org/0000-0002-2084-3019 E-mail: dominik.bayer@lrz.tu-muenchen.de
• Pretzsch, Address Technical University of Munich (TUM), Chair for Forest Growth and Yield Science, Hans-Carl-von-Carlowitz-Platz 2, 85354, Freising, Germany E-mail: hans.pretzsch@lrz.tu-muenchen.de

The ongoing climate change may have a distinct effect on Norway spruce growth, one of the most important tree species in European forest management. Therefore, the understanding and assessment of climate-growth relationship can help to reveal relevant patterns in temporal variability that may result in lower tree vitality and decline. The main objective of our study was to evaluate the long-term climate-growth variability of Norway spruce in south-eastern Norway, at the northern edge of the temperate zone. We sampled in total 270 dominant and co-dominant trees from 18 plots in south-eastern Norway. We analysed stem cores and evaluated crown condition parameters to assess the retrospective tree growth and vitality. Despite considerable differences in the crown parameters, high similarity among tree-ring width (TRW) series allowed compiling the regional tree-ring width chronology. Correlations between TRW and climate parameters showed temporal instability in their relationship during the period 1915–2012. While we did not detect any significant relationships between TRW and climate parameters in the first half of the study period (1915–1963), a significant correlation between TRW and spring precipitation was observed for the period 1964–2012. This shift appeared concurrent with temperatures reaching above-average values compared to the average of the climate normal period 1961–1990.

• Čermák, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědělská 3, 613 00 Brno, Czech Republic E-mail: cermacek@mendelu.cz
• Rybníček, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědělská 3, 613 00 Brno, Czech Republic; Global Change Research Institute, The Czech Academy of Sciences, Bělidla 986/4a, 603 00 Brno, Czech Republic E-mail: michalryb@post.cz
• Žid, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědělská 3, 613 00 Brno, Czech Republic E-mail: tom.z@centrum.cz
• Andreassen, Norwegian Institute of Bioeconomy Research (NIBIO), P.O. Box 115, NO-1431 Ås, Norway E-mail: Kjell.Andressen@nibio.no
• Børja, Norwegian Institute of Bioeconomy Research (NIBIO), P.O. Box 115, NO-1431 Ås, Norway E-mail: Isabella.Borja@nibio.no
• Kolář, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědělská 3, 613 00 Brno, Czech Republic; Global Change Research Institute, The Czech Academy of Sciences, Bělidla 986/4a, 603 00 Brno, Czech Republic E-mail: koldatom@gmail.com

Gap formation and its effects on regeneration have been reported as being important in forest development, but seldom studies concentrated on the gap closure process by lateral extension growth of canopy trees surrounding gaps. We monitored the closure process of 12 artificial gaps for 7 years with three size classes: small (from 68 m² to 125 m²), middle (from 174 m² to 321 m²), and large (from 514 m² to 621 m²); and investigated the regeneration twice in a temperate secondary forest, Northeast China. The closure process can be described through quadratic functions, which showed the closure rates slowed down with gap ages. Large gaps had a higher closure rate (39 m² a^–1) than middle gaps (25 m² a^–1) and small gaps (11 m² a^–1). According to the quadratic equations, the lateral growth could last 11, 13 and 16 years for small, middle and large gaps with a remaining size of 12, 69 and 223 m², respectively. As expected, regeneration exhibited the highest seedling density and volume in large gaps. There was no significant difference in regeneration density between middle gaps, small gaps and forest understory in the final investigation; but the volume of regenerated woody species increased significantly from small gaps to large gaps compared with forest understory. These results may provide references on the choice of appropriate gap sizes to promote the regeneration in temperate secondary forests.

• Lu, State Key Laboratory of Forest and Soil Ecology, Qingyuan Forest CERN, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110016, China;  University of Chinese Academy of Sciences, Beijing 100049, China E-mail: delianglu14@hotmail.com
• Zhu, State Key Laboratory of Forest and Soil Ecology, Qingyuan Forest CERN, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110016, China E-mail: jiaojunzhu@iae.ac.cn
• Sun, State Key Laboratory of Forest and Soil Ecology, Qingyuan Forest CERN, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110016, China E-mail: yirongsun@iae.ac.cn
• Hu,  Chinese Research Academy of Environmental Sciences, Beijing 100012, China E-mail: lilehu@gmail.com
• Zhang, State Key Laboratory of Forest and Soil Ecology, Qingyuan Forest CERN, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110016, China;  University of Chinese Academy of Sciences, Beijing 100049, China E-mail: zgq04713@163.com

Seed orchards are the link between tree breeding and reforestation. This paper presents data on cone, seed and pollen production and seed quality gathered over 21 years in a Pinus sylvestris (L.) experimental seed orchard containing plots with 14 different combinations of stand density and targeted pruning height. The treatments’ stand densities ranged from 267 to 4000 stems ha^-1, and the target graft heights ranged from 2 to 6 meters. Pollen production began at the same orchard age for all studied combinations of stand density and target height but the level of pollen production per hectare increased more rapidly in treatments with higher stand densities. In treatments with dense spacing, cone and seed production initially increased more rapidly than in treatments with wider spacing, thereby providing an earlier return on investment and a shorter seed production lag time. However, the levels of cone and seed production in such treatments over the entire study period were not appreciably different to those achieved in treatments with wider spacing and higher target height. The treatments did not differ substantially with respect to seed quality. These results show that comparable levels of seed production can be obtained with different combinations of stand density and target height, giving seed orchard owners and managers a wide range of viable management options.

• Almqvist, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: curt.almqvist@skogforsk.se
• Jansson, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: gunnar.jansson@skogforsk.se

• Niemistö, Finnish Forest Research Institute, Parkano Unit, FI-39700 Parkano, Finland E-mail: pentti.niemisto@metla.fi

• Hou, Department of Life Sciences, Shangqiu Normal University, Shangqiu, China E-mail: yh@nn.cn
• Qu, Department of Life Sciences, Shangqiu Normal University, Shangqiu, China E-mail: jq@nn.cn
• Luo, School of Environment and Life Sciences, Kaili University, Kaili, China E-mail: zl@nn.cn
• Zhang, Shanghai Key Laboratory of Urbanization and Ecological Restoration, East China Normal University, Shanghai, China, and University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: cz@nn.cn
• Wang, Shanghai Key Laboratory of Urbanization and Ecological Restoration, East China Normal University, Shanghai, China, and University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: kywang@re.ecnu.edu.cn

• Schiessl, Department of Genetics, Research and Training Centre for Forests, Natural Hazards and Landscape, Hauptstrasse 7, A-1140 Vienna, Austria E-mail: es@nn.at
• Grabner, Department of Material Sciences and Process Engineering, Institute of Wood Science and Technology, University of Natural Resources and Applied Life Sciences, Peter-Jordan-Strasse 82, A-1190 Vienna, Austria E-mail: mg@nn.at
• Golesch, Department of Genetics, Research and Training Centre for Forests, Natural Hazards and Landscape, Hauptstrasse 7, A-1140 Vienna, Austria E-mail: gg@nn.at
• Geburek, Department of Genetics, Research and Training Centre for Forests, Natural Hazards and Landscape, Hauptstrasse 7, A-1140 Vienna, Austria E-mail: tg@nn.at
• Schueler, Department of Genetics, Research and Training Centre for Forests, Natural Hazards and Landscape, Hauptstrasse 7, A-1140 Vienna, Austria E-mail: silvio.schueler@bfw.gv.at

• Korpela, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ilkka.korpela@helsinki.fi
• Ørka, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, P.O.Box 5003, NO-1432 Ås, Norway E-mail: hoo@nn.no
• Maltamo, University of Eastern Finland, School of Forest Science, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: mm@nn.fi
• Tokola, University of Eastern Finland, School of Forest Science, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: tt@nn.fi
• Hyyppä, Finnish Geodetic Institute, Department of Photogrammetry and Remote Sensing, P.O.Box 15, FI-02431 Masala, Finland E-mail: jh@nn.fi

• Maltamo, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: matti.maltamo@joensuu.fi
• Peuhkurinen, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: jp@nn.fi
• Malinen, Finnish Forest Research Institute, Joensuu Research Unit, FI-80101 Joensuu, Finland E-mail: jm@nn.fi
• Vauhkonen, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: jv@nn.fi
• Packalén, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: pp@nn.fi
• Tokola, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: tt@nn.fi

• Rautiainen, Tartu Observatory, 61602 Tõravere, Estonia, and Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: miina.rautiainen@helsinki.fi
• Mõttus, Tartu Observatory, 61602 Tõravere, Estonia E-mail: mm@nn.ee
• Stenberg, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ps@nn.fi
• Ervasti, City of Vantaa, Land Use and Environment / Green Area Unit, Kielotie 13, FI-01300 Vantaa, Finland E-mail: se@nn.fi

• Hotanen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: juha-pekka.hotanen@metla.fi
• Maltamo, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: mm@nn.fi
• Reinikainen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: ar@nn.fi

• Kalliovirta, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: jk@nn.fi
• Tokola, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: timo.tokola@helsinki.fi

• Grote, TU München, Chair of Forest Yield Science, Am Hochanger 13, D-85354 Freising, Germany E-mail: ruediger.grote@lrz.tu-muenchen.de

• Pensa, Institute of Ecology, Department of Northeast Estonia, Pargi 15, EE-41537 Jõhvi, Estonia E-mail: margus@ecoviro.johvi.ee
• Jalkanen, Finnish Forest Research Institute, Rovaniemi Research Station, P.O. Box 16, FIN-96301 Rovaniemi, Finland E-mail: rj@nn.fi

• Vettenranta, Kivirinnanpolku 4, FIN-40950 Muurame, Finland E-mail: vettenr@cc.joensuu.fi

Fast and accurate estimates of canopy cover are central for a wide range of forestry studies. As direct measurements are impractical, indirect optical methods have often been used in forestry to estimate canopy cover. In this paper the accuracy of canopy cover estimates from two widely used canopy photographic methods, hemispherical photography (DHP) and cover photography (DCP) was evaluated. Canopy cover was approximated in DHP as the complement of gap fraction data at narrow viewing zenith angle range (0°–15°), which was comparable with that of DCP. The methodology was tested using artificial images with known canopy cover; this allowed exploring the influence of actual canopy cover and mean gap size on canopy cover estimation from photography. DCP provided robust estimates of canopy cover, whose accuracy was not influenced by variation in actual canopy cover and mean gap size, based on comparison with artificial images; by contrast, the accuracy of cover estimates from DHP was influenced by both actual canopy cover and mean gap size, because of the lower ability of DHP to detect small gaps within crown. The results were replicated in both DHP and DCP images collected in real forest canopies. Finally, the influence of canopy cover on foliage clumping index and leaf area index was evaluated using a theoretical gap fraction model. The main findings indicate that DCP can overcome the limits of indirect techniques for obtaining unbiased and precise estimates of canopy cover, which are comparable to those obtainable from direct, more labour-intensive techniques, being therefore highly suitable for routine monitoring and inventory purposes.

• Chianucci, Consiglio per la Ricerca in Agricoltura e l’Analisi dell’Economia Agraria – Forestry Research Centre, viale Santa Margherita 80, 52100 Arezzo, Italy http://orcid.org/0000-0002-5688-2060 E-mail: fchianucci@gmail.com