Articles containing the keyword 'stem'

This study examined the relationships between forest management planning units and patches formed by forest habitat components. The test area used was a part of Koli National Park in North Karelia, eastern Finland. Forest management planning units (i.e. forest compartments) were defined by using a traditional method of Finnish forestry which applies aerial photographs and compartment-wise field inventory. Patches of forest habitat components were divided according to subjective rules by using a chosen set of variables depicting the edaphic features and vegetation of a forest habitat. The spatial distribution of the habitat components was estimated with the kriging-interpolation based on systematically located sample plots. The comparisons of the two patch mosaics were made by using the standard tools of GIS. The results of the study show that forest compartment division does not correlate very strongly with the forest habitat pattern. On average, the mean patch size of the forest habitat components is greater and the number of these patches lower compared to forest compartment division. However, if the forest habitat component distribution had been considered, the number of the forest compartments would have at least doubled after intersection.

• Uuttera, E-mail: ju@mm.unknown
• Hyppänen, E-mail: hh@mm.unknown

Dry mass and nutrient (N, P, K, Ca, Mg, B) contents of field layer vegetation and a combination of bottom layer vegetation and litter (referred to as bottom/litter layer in the text) were studied one year before and three years after fertilization (NPK and PK) on a drained low-shrub pine bog in eastern Finland. The results of an earlier study on the tree layer were combined with those of this study in order to estimate the changes caused by fertilization in the total plant biomass and litter. Before fertilization the average dry mass of the field and bottom/litter layers was 8,400 kg ha^-1 and 7,650 kg ha^-1, respectively. The above-ground parts accounted for 25% of the total field layer biomass. The dry mass of the field and bottom/litter layers together was < 20% of the dry mass accumulated in the total plant biomass and litter. The corresponding figures for N, P, K, Ca, Mg and B were 44%, 38%, 30%, 38%, 31% and 17%, respectively. Fertilization did not significantly affect the dry mass of either the field layer vegetation or the bottom/litter layer. 33% of the applied P was accumulated in the total plant biomass and litter on the PK-fertilized plots, and 25% on the NPK-fertilized plots. For the other elements, the proportions on the PK-fertilized plots were K 31%, Ca 6%, Mg 11% and B 13%. On the NPK-fertilized plots, the corresponding figures were N 62%, K 32%, Ca 6%, Mg 9% and B 13%. Except for B and K, the accumulation of fertilizer nutrients in the understorey vegetation and litter was of the same magnitude or greater than the uptake by the tree layer.

• Finér, E-mail: lf@mm.unknown
• Nieminen, E-mail: mn@mm.unknown

The rule-based formal language of "stochastic sensitive growth grammars" was designed to describe algorithmically the changing morphology of forest trees during their lifetime under the impact of endogenous and exogenous factors, and to generate 3-D simulations of tree structures in a systematic manner. The description in the form of grammars allows the precise specification of structural models with functional components. These grammars (extended L-systems) can be interpreted by the software GROGRA (Growth grammar interpreter) yielding time series of attributed 3-D structures representing plants. With some recent extensions of the growth-grammar language (sensitive functions, local variables) it is possible to model environmental control of shoot growth and some simple allocation strategies, and to obtain typical competition effects in tree stands qualitatively in the model.

• Kurth, E-mail: wk@mm.unknown
• Sloboda, E-mail: bs@mm.unknown

The model HYDRA, which simulates water flow in the branched tree architecture, is characterized. Empirical studies of the last decades give strong evidence for a close structure-function linkage in the case of tree water flow. Like stomatal regulation, spatial patterns of leaf specific conductivity can be regarded as a strategy counteracting conductivity losses, which may arise under drought. Branching-oriented water flow simulation may help to understand how damaging and compensating mechanisms interact within the hydraulic network of trees. Furthermore, a coupling of hydraulic to morphological modelling is a prerequisite if water flow shall be linked to other processes. Basic assumptions of the tree water flow model HYDRA are mass conservation, Darcy's law and the spatial homogeneity of capacitance and axial conductivity. Soil water potential is given as a one-sided border condition. Water flow is driven by transpiration. For unbranched regions these principles are condensed to a nonlinear diffusion equation, which serves as a continuous reference for the discrete method tailored to the specific features of the hydraulic network. The mathematical derivation and model tests indicate that the realization of the basic assumptions is reproducible and sufficiently exact. Moreover, structure and function are coupled in a flexible and computationally efficient manner. Thus, HYDRA may serve as a tool for the comparative study of different tree architectures in terms of hydraulic function.

• Früh, E-mail: tf@mm.unknown

Process-based tree growth models are recognized to be flexible tools which are valuable for investigating tree growth in relation to changing environment or silvicultural treatments. In the context of forestry, we address two key modelling problems: allocation of growth which determines total wood production, and distribution of wood along the stem which determines stem form and wood quality. Growth allocation and distribution are the outcome of carbon translocation, which may be described by the Munch theory. We propose a simpler gradient process to describe the carbon distribution in the phloem of conifers. This model is a reformulation of a carbon diffusion-like process proposed by Thornley in 1972. By taking into account the continuity of the cambium along the stem, we obtain a one-dimensional reaction-diffusion model which describes both growth allocation between foliage, stem and roots, and growth distribution along the stem. Distribution of wood along the stem is then regarded as an allocation process at a smaller scale. A preliminary sensitivity analysis is presented. The model predicts a strong relationship between morphology and foliage-root allocation. It also suggests how empirical data, such as stem analysis, could be used to calibrate and validate allocation rules in process-based growth models.

• Deleuze, E-mail: cd@mm.unknown
• Houllier, E-mail: fh@mm.unknown

Many kinds of planning systems have been labelled decision support systems (DSS), but few meet the most important features of real DSSs in planning and control of wood procurement. It has been concluded that many reasons exist to develop DSSs for wood procurement. The purchasing of timber seems to be one of the most promising areas for DSS, because there is no formal structure for these operations and decisions deal with human behaviour. Relations between DSSs and different features of the new approaches in wood procurement are also discussed, and hypotheses for future studies suggested.

• Harstela, E-mail: ph@mm.unknown

Within the European Community snow damage affects an estimated 4 million m³ of timber every year, causing significant economic losses to forest owners. In Northern Europe, for example, the occurrence of snow damage has increased over the last few decades mainly due to the increase in total growing stock. The most common form of damage is stem breakage, but trees can also be bent or uprooted. Trees suffering snow damage are also more prone to consequential damage through insect or fungal attacks.

Snow accumulation on trees is strongly dependent upon weather and climatological conditions. Temperature influences the moisture content of snow and therefore the degree to which it can accumulate on branches. Wind can cause snow to be shed, but can also lead to large accumulations of wet snow, rime or freezing rain. Wet snow is most likely in late autumn or early spring. Geographic location and topography influence the occurrence of damaging forms of snow, and coastal locations and moderate to high elevations experience large accumulations. Slope plays a less important role and the evidence on the role of aspect is contradictory. The occurrence of damaging events can vary from every winter to once every 10 years or so depending upon regional climatology. In the future, assuming global warming in northern latitudes, the risk of snow damage could increase, because the relative occurrence of snowfall near temperatures of zero could increase.

The severity of snow damage is related to tree characteristics. Stem taper and crown characteristics are the most important factors controlling the stability of trees. Slightly tapering stems, asymmetric crowns, and rigid horizontal branching are all associated with high risk. However, the evidence on species differences is less clear due to the interaction with location. Management of forests can alter risk through choice of regeneration, tending, thinning and rotation. However, quantification and comparison of the absolute effect of these measures is not yet possible. An integrated risk model is required to allow the various locational and silvicultural factors to be assessed. Plans are presented to construct such a model, and gaps in knowledge are highlighted.

• Nykänen, E-mail: mn@mm.unknown
• Broadgate, E-mail: mb@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown
• Peltola, E-mail: hp@mm.unknown
• Quine, E-mail: cq@mm.unknown

The purpose of this article was to collate the literature on fungal diseases that occur on seedlings in forest nurseries. It describes the symptoms of the diseases, the infection pattern of each fungus and the possibilities of controlling the diseases. As background a short introduction is given on forests and nursery practices in Finland.

• Lilja, E-mail: al@mm.unknown
• Kurkela, E-mail: tk@mm.unknown
• Lilja, E-mail: sl@mm.unknown
• Rikala., E-mail: rr@mm.unknown

The productivity of Scots pine (Pinus sylvestris L.) under changing climatic conditions in the southern part of Finland was studied by scenario analysis with a gap-type forest ecosystem model. Standard simulations with the model predicted an increased rate of growth and hence increased productivity as a result of climatic warming. The gap-type model was refined by introducing an overwintering sub-model describing the annual growth cycle, frost hardiness, and frost damage of the trees. Simulations with the refined gap-type model produced results conflicting with those of the standard simulation, i.e., drastically decreased productivity caused by mortality and growth-reducing damage due to premature dehardening in the changing climate. The overwintering sub-model was tested with frost hardiness data from Scots pine saplings growing at their natural site 1) under natural conditions and 2) under elevated temperature condition, both in open-top chambers. The model predicted the frost hardiness dynamics quite accurately for the natural conditions while underestimating the frost hardiness of the saplings for the elevated temperature conditions. These findings show that 1) the overwintering sub-model requires further development, and 2) the possible reduction of productivity caused by frost damage in a changing climate is less drastic than predicted in the scenario analysis. The results as a whole demonstrated the need to consider the overwintering of trees in scenario analysis carried out with ecosystem model for boreal conditions. More generally, the results revealed a problem that exists in scenario analysis with ecological models: the accuracy of a model in predicting the ecosystem functioning under present climatic condition does not guarantee the realism of the model, nor for this reason the accuracy for predicting the ecosystem functioning under changing climatic conditions. This finding calls for the continuous rigorous experimental testing of ecological models used for assessing the ecological implications of climatic change.

• Hänninen, E-mail: hh@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown
• Leinonen, E-mail: il@mm.unknown
• Repo, E-mail: tr@mm.unknown

BOREAS is a four-year, regional-scale experiment to study the forested continental interior of Canada. It aims at improving our understanding of the interaction between the earths' climate system and the boreal forests at short and intermediate time scales, in order to clarify their role in global change.

During the winter, spring and summer of 1994, five field campaigns were conducted. About 85 investigation teams including nearly 300 scientists participated, including forest ecologists and ecophysiologists, atmospheric physicists, boundary-layer meteorologists, hydrologists, biochemists, atmospheric chemists and remote sensing specialists.

The findings so far have been significant in terms of their implication for global change. The boreal ecosystem, occupying roughly 17 percent of the vegetated land surface and thus an important driver of global weather and climate, absorbs much more solar energy than is assumed by operational numerical weather prediction models. Albedo measurement show that this forest absorbs nearly 91% of the sun's incident energy. Additionally, while it is known that much of the boreal ecosystems consists of forested wetlands, lakes, bogs and fens, the measurements show that the atmosphere above was extremely dry; humidity and deep boundary layer convection (3,000 m) mimicked conditions found only over deserts. Physiological measurements of the trees show that this atmospheric desiccation was a result of the forests' strong biological control limiting surface evaporation. This tight control was linked to the low soil temperature and subsequently reduced rates of photosynthesis. BOREAS measurement also focused on net ecosystem carbon exchange. Data acquired during the late spring and summer, showed the boreal forests to be a net carbon sink. However, no measurements were taken in the early spring following thaw, and in the late fall, where the balance between photosynthesis and respiration is poorly understood. During 1996 additional data will be acquired to resolve the annual carbon budget and how it might depend on interannual climate differences.

• Hall, E-mail: fh@mm.unknown
• Sellers, E-mail: ps@mm.unknown
• Williams, E-mail: dw@mm.unknown

The eddy covariance technique is a novel micrometeorological method that enables the determination of the atmosphere-biosphere exchange rate of gases such as ozone and carbon dioxide on an ecosystem scale. This paper describes the technique and presents results from the first direct measurements of turbulent fluxes of O₃, CO₂ and H₂O above a forest in Finland. The measurements were performed during 15 July-5 August 1994 above a Scots pine (Pinus sylvestris L.) stand near the Mekrijärvi research station in Eastern Finland.

The expected diurnal cycles were observed in the atmospheric fluxes of O₃, CO₂ and H₂O. The data analysis includes interpretation of the O₃ flux in terms of the dry deposition velocity and evaluation the dependency of the net CO₂ flux on radiation. The eddy covariance method and the established measurement system has proved suitable for providing high-resolution data for studying ozone deposition to a forest as well as the net carbon balance and related physiological processes of an ecosystem.

• Aurela, E-mail: ma@mm.unknown
• Laurila, E-mail: tl@mm.unknown
• Tuovinen, E-mail: jt@mm.unknown

This paper deals with the testing of dynamic stratification for estimating stand level forest characteristics (basal areas, mean diameter, mean height and mean age) for a 117 ha study areas in Finland. The results do not show possibilities to achieve more accurate estimates using only Landsat TM principal components as auxiliary data opposed to static stratification. It was found that in dynamic stratification non-measured observations should be assigned the mean characteristics of the measured observations that belong to the same cube (class) instead of stratification variable classes until a certain limit. If only one principal component is used the number of classes has, however, little influence. Low field values are overestimated and high values underestimated.

The only successful results were obtained using two variables of different origin – the qualitative development stage class and the quantitative 1st principal component. The lowest root mean square error in estimating basal area was 6.40 m²/ha, mean diameter 3.34 cm, mean height 2.65 m and mean age 14.06 years. This increase of stratification accuracy is mainly resulted by the use of development stage class as an auxiliary variable.

• Mozgeris, E-mail: gm@mm.unknown

Individual tree-growth models for diameter and height, and a model for the cylindrical stem form factor are presented. The aims of the study were to examine modelling methods in predicting growth response to thinning, and to develop individual-tree, distance-independent growth models for predicting the development of thinned and unthinned stands of Scots pine (Pinus sylvestris L.). The models were constructed to be applicable in simulation systems used in practical forest management planning. The models were based on data obtained from eleven permanent thinning experiments located in even-aged Scots pine stands in Southern and Central Finland.

Two alternative models were developed to predict tree diameter growth in thinned and unthinned stands. In the first model, the effect of stand density was described using stand basal area. In the alternative model, an explicit variable was incorporated referring to the relative growth response due to thinning. The magnitude of the growth response was expressed as a function of thinning intensity. The Weibull function was employed to describe the temporal distribution of the thinning response. Both models resulted in unbiased predictions in unthinned and in moderately thinned stands. An explicit thinning variable was needed for unbiased growth prediction in heavily thinned stands, and in order to correctly predict the dynamics of the growth response.

In the height growth model, no explicit thinnning variable referring thinning was necessary for growth prediction in thinned stands. The stem form factor was predicted using the model that included tree diameter and tree height as regressor variables. According to the results obtained, the information on the changes in the diameter/height ratio following the thinning is sufficient to predict the change in stem form.

• Hynynen, E-mail: jh@mm.unknown

The Värriö environmental measurement station has been designed and constructed during 1991 and 1992. The measurement system consists of measurement units for gases (sulphur dioxide, ozone, carbon dioxide), particles, photosynthesis and irradiation. A meteorological station is also included. The preliminary measurement period was started on August, 1991. During the first year (1991–1992) some parts of the system were redeveloped and rebuilt. Full, continuous measurement started in August 1992. The system has been working quite reliably, with good accuracy. The preliminary results show that pollution episodes are observed when the wind direction is from Monchegorsk or Nikel, the main emission sources in Kola Peninsula.

• Hari, E-mail: ph@mm.unknown
• Aalto, E-mail: pa@mm.unknown
• Hämeri, E-mail: kh@mm.unknown
• Kulmala, E-mail: mk@mm.unknown
• Lahti, E-mail: tl@mm.unknown
• Luoma, E-mail: sl@mm.unknown
• Palva, E-mail: lp@mm.unknown
• Pohja, E-mail: tp@mm.unknown
• Pulliainen, E-mail: ep@mm.unknown
• Siivola, E-mail: es@mm.unknown
• Vesala, E-mail: tv@mm.unknown

In this study, model-based and design-based inference methods are used for estimating mean volume and its standard error for systematic cluster sampling. Results obtained with models are compared to results obtained with classical methods. The data are from the Finnish National Forest Inventory. The variation of volume in ten forestry board districts in Southern Finland is studied. The variation is divided into two components: trend and correlated random errors. The effect of the trend and the covariance structure on the obtained mean volume and standard error estimates is discussed. The larger the coefficient of determination of the trend model, the smaller the model-based estimates of standard error, when compared to classical estimates. On the other hand, the wider the range and level of autocorrelation between the sample plots, the larger the model-based estimates of standard error.

• Kangas, E-mail: ak@mm.unknown

Coniferous and non-coniferous fine root and rhizome production was measured after one growing season using the ingrowth bag method in Scots pine (Pinus sylvestris L.) stands differing in ages from 7 to 105 years in Southern Finland. Total fine-root production decreased from the 7-year to 20-year-old stands, and then increased slightly in the 85- to 105-year-old stands. Most of the total fine-root biomass in the youngest age groups came from non-conifer species, whereas most of the total fine-root biomass in the three older age groups came from conifer species. The maximum coniferous fine-root production was found to occur at crown closure in the 11- to 13-year-old stands. Rhizome production was the lowest and highest in the 20- and 85- to 105-year-old stands, respectively. The increase in rhizome production in the 85- to 105-year-old stands was associated with an abundant understory cover of Vaccinium myrtillus and V. vitis-idaea and an increase in light penetration. The ingrowth bag method was found to be useful in assessing the relative fine-root production among species-group and successional stages of Scots pine stands.

The PDF includes an abstract in Finnish.

• Messier, E-mail: cm@mm.unknown
• Puttonen, E-mail: pp@mm.unknown

A model for the mechanism of windfall and stem breakage was constructed for single Scots pine (Pinus sylvestris L.) at the stand edge. The total turning moment arising from the wind drag and from the bending of stem and crown was calculated along with the breaking stress of the stem. Similarly, the support given by the root -soil plate anchorage was calculated. Windspeed variation within the crown and the vertical distribution of stem and crown weight were taken into account. Model computations showed that trees having a large height to diameter ratio were subjected to greater risk of falling down or breaking than trees with a small height to diameter ratio. The windspeed required to blow down a tree or break the stem of a tree decreased if the height to diameter ratio or the crown to stem ratio of trees increased.
The PDF includes an abstract in Finnish.

• Peltola, E-mail: hp@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown

Seedlings from four Norway spruce (Picea abies (L.) H. Karst.) stands originating from areas with effective temperature sums ranging from 710 d.d. to 1,150 d.d. were raised under artificial light and temperature treatment. After a 10-week growing period the hardening process was started by subjecting the seedlings to +8°C night temperature and +15°C day temperature, and increasing the night length by 1.5 hour/week. Hardiness was measured by means of artificial freezing treatment (-10°C or -15°C), followed by visual estimation of the degree of needle injury. The stem height, lignification and bud development were measured before the freezing treatment. The amount of injury increased the more southern the origin of the tested material was. Furthermore, the proportion of non-lignified part of the seedling stem was negatively correlated with the latitude of the provenances. The proportion of seedlings with clearly visible buds was more than 90% in the northernmost entry and less than 1% in the southernmost one. The overall correlation coefficient between the needle injuries and the proportion of non-lignified part of the stem was rather high, but varied considerably from 0.3 in the northernmost material to over 0.6 in the southern provenances. According to the results, it seems to be possible to use growth characteristics as an indicator of frost hardiness at the provenance level.
The PDF includes an abstract in Finnish.

• Pulkkinen, E-mail: pp@mm.unknown

Managing forests and other natural resources requires merging of data and knowledge from many fields. Research efforts in many countries have simultaneously aimed at computer applications to help managing the large amounts of data involved and the complexities of decision making. This has invariably led to large integrated systems. An integrated system is software that consists of modules for various tasks in natural resource management, spatial analysis, simulation and optimization, diagnostic reasoning, levels, and communicating with the user.

The paper presents an overview of the need, levels and historical development of integrated systems. Newly emerged technologies, especially object-oriented programming and the X Window System with its associated environment have given new flexibility and transparency to the designs. The client-server architecture is found out as an ideal model for integrated systems. The paper describes an implementation of these ideas, the INFORMS system that supports the information needs of district level forest management planning.

The PDF includes an abstract in Finnish.

• Loh, E-mail: dl@mm.unknown
• Saarenmaa, E-mail: hs@mm.unknown

The main stem of young Scots pine (Pinus sylvestris L.) trees was cut off halfway along the current leading shoot and the two previous years’ leading shoots to simulate moose (Alces alces) damage. Trees of the same size were chosen as controls before treatments. The experiment was inspected ten years after artificial stem breakage. Removing the current leading shoot and the second shoot did not essentially affect the height and diameter growth of the trees. Removal down to the third shoot reduced the height as well as diameter growth. The average loss in growth was equivalent to less than one year’s growth. When the stem was cut off at the second or third shoot, stem crookedness and the presence of knots resulted in stem defects that will subsequently reduce the sawtimber quality. A high proportion of the stem defects will obviously still be visible at the first thinning cutting. Removing injured trees as pulpwood and pruning the remaining parts of cut stems evidently improves the quality of pine stand with moose damage.

The PDF includes an abstract in Finnish.

• Heikkilä, E-mail: rh@mm.unknown
• Löyttyniemi, E-mail: kl@mm.unknown

A model for the succession of the forest ecosystem is described. The growth and development of trees and ground cover are controlled by temperature and light conditions and the availability of nitrogen and water. In addition, the effects of the annual cycle of trees including the risk of frost damage, wild fire, and wind damages are contained in the model as factors which control the survival and productivity of trees. The model also makes it possible to evaluated the risk of insect attack assuming that this risk is inversely related to the growth efficiency of trees.

The PDF includes an abstract in Finnish.

• Kellomäki, E-mail: sk@mm.unknown
• Hänninen, E-mail: hh@mm.unknown
• Kolström, E-mail: tk@mm.unknown
• Lauhanen, E-mail: rl@mm.unknown
• Mattila, E-mail: um@mm.unknown
• Pajari, E-mail: bp@mm.unknown
• Väisänen, E-mail: hv@mm.unknown

Crown characteristics and the distribution of three years’ (1986–88) biomass production of 20 pendulous Norway spruce (Picea abies f. pendula (Lawson) Sylvén) trees with heritable narrow crown and 15 normal-growned spruces (Picea abies (L.) H. Karst.) were studied in a 19-year-old mixed stand.

The form of the crown is conical in normal-crowned trees, columnar and narrow in pendulous trees. The partitioning of aboveground biomass to stems during the studied 3-year period was significantly higher in pendulous (0.281) than in normal-crowned trees (0.255) and also the ratio between growth of stemwood and growth of needle biomass during three years was higher in pendulous trees (0.67 g g^-1) than in normal-crowned trees (0.52 g g^-1). The needle biomass was distributed higher in the crown in pendula than in normal-crowned trees and they had a higher needle biomass/branchwood biomass ratio than normal trees. The difference in harvest increment between the two crown types are mostly due to the significantly lower branchwood biomass values in pendulous than in normal-crowned trees. The higher needle ’efficiency’ in pendulous trees is probably connected with high partitioning of needle biomass to the upper part of the crown in pendulous trees.

The PDF includes an abstract in Finnish.

• Pulkkinen, E-mail: pp@mm.unknown

The mating system was analysed in the upper and lower crown of two groups of European larch (Larix decidua Mill.) clones divided according to the percentage of full seeds in the upper and lower crown parts. The overall multilocus estimate of outcrossing rate (t) was calculated to be 0.929. The differences of outcrossing rates between crown levels and clonal groups respectively were not statistically significant. The t estimates were greater for the upper crown level and for clones with higher percentage of full seeds in the upper crown level. However, among all observations there was no correlation between outcrossing rates and percentages of full seeds for particular crown levels and groups of clones. Observed similarity of outcrossing among grafts of the same clones may indicate genetic control of self-fertilization rate in individual European larch. 

The PDF includes an abstract in Finnish.

• Burczyk, E-mail: jb@mm.unknown
• Kosiński, E-mail: gk@mm.unknown
• Lewandowski, E-mail: al@mm.unknown

Decision making in the forest protection involves diagnosing the pest, making predictions of the effects of the pest on forest, knowing the possible control tactics, and cost/benefit integration. To cope with all that, a generalist forest manager needs a tool like an expert system to support decisions.

This paper presents an expert system that approaches the goals of integrated pest management. With the systm, the user can make diagnosis and predictions of 12 North European bark beetles. Written in Common LISP and Flavors, the expert system has a combined frame- and rule-based knowledge representation. Frames are used to represent the hierarchy of insect taxonomy in diagnosis. Prediction is made with qualitative reasoning with rules. The interface engine applies both forward and backward chaining. The system has a graphical user interface that supports exploring the sensitivity of advice on input.

It is concluded that expert systems and artificial intelligence have high applicability everywhere in forestry where complicated decisions have to be made. Especially, an integrated pest management system in forestry is largely equivalent to a computerized decision-making aid.

The PDF includes an abstract in Finnish.

• Saarenmaa, E-mail: hs@mm.unknown

Containerized tree seedlings will be used on an increasing scale in the future in different parts of the world. There are number of techniques for the production of small one-year-old seedlings but it has not been possible to develop a completely satisfactory methods for large containerized seedlings production. In the long-term development of pine plantations established with containerized seedlings the greatest problem has been deformation of the root system. With a new method, based on a sheet of peat and root pruning, it has been possible to produce conifer seedlings with a good root regeneration potential and favourable morphological root system development. The use of small containerized seedlings allows an increase in planting density without any marked increase in regeneration costs.

The PDF includes an abstract in English.

• Parviainen, E-mail: jp@mm.unknown

Properties of fibres in pulpwood, especially length, width and the thickness of walls in tracheids, are essential for strength properties of pulp and paper. Length and width of tracheids increase from pith to surface in radial direction. Young and small-sized stems have also smaller fibres. Small-sized Pinus sylvestris L. test trees had tracheids that were shorter both in stems and knot wood than those in normal sized trees. However, cell walls in test trees were as thick as in normal sized trees. It seems that especially the L/T -ratio (length/thickness) in small stems is worse than in normal sized pulp wood.

The PDF includes an abstract in English.

• Laurila, E-mail: rl@mm.unknown

The seed crop of Norway spruce (Picea abies (L.) H. Karst.) and Scots pine (Pinus sylvestris L.) is predicted with the help of mean monthly temperatures during May–August one and two years before the flowering year. The prediction models were made separately for Lapland and for the rest of Finland. The models are based on 10-year periods of seed crop measurements and climatic data. The total number of time series was 59.

In Lapland, Norway spruce flowered abundantly and produced an abundant seed crop after warm July–August and two years after cool July–August. In other parts of Finland, warm June and July produced a good flowering year, especially if these months were cool two years before the flowering year.

In Lapland, Scots pine flowered abundantly if the whole previous growing season was warm. Elsewhere in Finland, a cool June preceded prolific flowering in the coming year if the rest of the growing season was considerably warmer than the average.

The prediction models explained 37–49 % of the variation in the size of the seed crop. The occurrence of good and poor seed years was usually predicted correctly. Using the presented models, the prediction of the seed crop is obtainable 1.5 year for Norway spruce and 2.5 year for Scots pine before the year of seed fall.

The PDF includes an abstract in English.

• Pukkala, E-mail: tp@mm.unknown

Salix 'Aquatica Gigantea', widely experimented and promising species for temperate zone short rotation forestry, has since 1950 recorded in Finland 23 times with different clone numbers. Salix x dasyclados Wimm., by morphological, cultivational and productivity characteristics similar willow has been recorded 16 times.

The nomenclature and origin of both willows have remained unclear in field research. Recent observations, based on morphological analyses and chromosome studies suggest that ’Aguatica gigantea’ and most S. x dasuclados clones can be collected under one Siberian species: Salix burjatica Nasarov. The true Salix x dasyclados Wimm. is a female hybrid S. x viminalis x cinerea, famous West-European basket willow that has been very little experimented in Finland.

The PDF includes an abstract in Finnish.

• Pohjonen, E-mail: vp@mm.unknown

A team of 2 experienced workers was time-studied and their heart rate recorded under 4 days in clearcutting of a highly self-pruned Pinus patula Schltdl. & Cham. plantation. Task work and bonus payment systems were compared, but there was no difference in production rate, only the workplace time was extended from 2.3 h/d in task work to 3.9 h/d in bonus payment. The heart rate was 115–116 P/min in felling, 105–109 P/min in debranching and 109–114 P/min in bucking. The average heart rate in timber cutting was 108–109 P/min. Work load index was 34–37%, and the workers did not show any symptoms of accumulated stress. The production rate was 3.2 m³/h, (WPT, crew), which corresponds average piecework rate, the comparable walking speed being about 6.0 km/h. There are possibilities to increase the daily task by ergonomic grounds.

The PDF includes an abstract in Finnish.

• Saarilahti, E-mail: ms@mm.unknown
• Ole-Meiludie, E-mail: ro@mm.unknown

This paper designs an Economy-Welfare-Environment Adjustment System model (EWEAS model or EWE model in short) which consists of the circular flow of the economic, the welfare, and the environment system of forestry. That is, this model builds the relationship between the systems for material wealth and that for mental wealth.

The EWE model is designed as a complete open system model which describes the economy-welfare-environment circular system in forestry by linking up the internal system of forestry with the surrounding external systems. The EWE model can be manipulated as a policy formation or a policy decision model, and it is available for policy evaluation in the economic, the welfare and the environmental phase of forestry. The model is a basic simulation system model which is reliable in its reproductive fitness, stability and universality. Thus, this model ought to be useful in any country in the world as well as in Japan.

• Kishine, E-mail: tk@mm.unknown

In this paper, the author explains the characteristics of the Japanese forestry planning system and points out some of the problems found therein from the viewpoint of the management of privately-owned forests relating to the economic background and governmental policy.

The forestry planning system is a centralizes type of planning, the planning begins at the top and flows downward and outward the periphery. In order to make this planning system an effective instrument, the district forestry planning founded under the system must approach the problem of how to combine the resources of the forest with the district’s inhabitants and the forest owners; and further, the extent of the effective union of the district and the local timber manufacturing must be examined.

• Shimotori, E-mail: ss@mm.unknown

Development of timber sale accounting system for the U.S. Department of Agriculture, Forest Service ordered by the U.S. Congress, has entailed numerous evaluations and research projects. Critics of the current process claims administrative costs are not recovered by the prices paid for Federal timber. However, management of multiple resources for multiple uses makes traditional accounting difficult; i.e., keeping track of cash flows. A further complication involves allocating costs to the various resources (joint cost allocation), for which no nonarbitrary method currently exists. A concurrent issue involves the building of roads for timber harvest into areas released from wilderness consideration. Environmentalists see the road building program of the Federal land management agencies as an additional reason Federal management costs are not recovered from timber-generated revenues. The heart of the issue is which lands are economically suited for timber management, and what nonmarket benefits and costs accrue from the timber management.

• Risbrudt, E-mail: cr@mm.unknown

The structure of 20 Scots pine (Pinus sylvestris L.) trees was analysed as a water transport system. There is a tight linear regression between the cross-sectional area of the stem at the height of its lowest living branch and the cross-sectional area of its coarse roots, between the cross-sectional area of the stem at the height of its lowest living branch and the total cross-sectional area of its branches, and between the cross-sectional area of the base of a branch and the total cross-sectional area of subsidiary branches of that branch. The capacity of successive organs, measured as cross-sectional areas, to transport water was thus found to be regular within a tree.

The PDF includes an abstract in Finnish.

• Hari, E-mail: ph@mm.unknown
• Heikinheimo, E-mail: ph@mm.unknown
• Kaipiainen, E-mail: lk@mm.unknown
• Korpilahti, E-mail: eeva.korpilahti@luke.fi
• Mäkelä, E-mail: am@mm.unknown
• Samela, E-mail: js@mm.unknown

A mathematical model was developed for determining the value of Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies (L.) H. Karst.) stems on the basis of sawing and pulping. The model was based on selling prices of sawn goods, pulp and other products as well as processing costs. Sawing was applied to large-dimension parts of stems and pulping to other parts and small stems. Bark and other residues were burned. The quality of pine stems was described by the distance of the lowest dead branch. In spruce only stem size affected the quality-

According to the results, the size of stem affects considerably the value of pine stems and clearly that of spruce stems. The main reason is an increase in the productivity of frame sawing as the stem size increases. In pine another factor is the higher price of sawn goods. The effect of pulp price increases as the stem size decreases. Even in large sized stems the effect of pulp was notable as the value of chips and saw dust was determined on the basis of product values in export. The competition ability of mechanical pulp was greatly affected by the price of electricity.

The PDF includes an abstract in English.

• Kärkkäinen, E-mail: mk@mm.unknown

Especially in forest vegetation studies, the light climate below the canopy is of great interest. In extensive forest inventories, direct measurement of the light conditions is too time-consuming. Often only the standard tree stand parameters are available. The present study was undertaken with the aim to develop methods for estimation of the light climate on the basis of readily measurable tree stand characteristics. The study material includes 40 sample plots representing different kinds of more or less mature forest stands of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.).

In each forest stand, a set of hemipherical photographs was taken and standard tree stand measurements were performed. A regression approach was applied in order to elaborate linear models for predicting the canopy coverage. The total basal area of the stand explained 63% of variance in the canopy coverage computed from hemipherical photographs. A coefficient representing the relative proportion of Norway spruce in the stand increased the explanatory power into 75%. When either the stand density (stems/unit area) or dominant age of the stand was included into the model, increment of the explanatory power into 80% was achieved. By incorporating both of the preceding predictors, an explanatory power of 85% was reached.

The PDF includes a summary in Finnish.

• Kuusipalo, E-mail: jk@mm.unknown

The goal of this research was to study the position of forest supervisors, their education, number, pay system, professional organizations and work situations in Nordic countries. The study belonged to a joint Nordic project of the Nordic Research Council on Forest Operations. Participating in the actual work the number of forest supervisors were Denmark (year 1978) 715, Finland (1980) 8,000, Norway (1967) 1,055, and Sweden (1975) 6,400. In Denmark, 87% of the supervisors worked in forestry, in Finland 91%, in Norway 98% and in Sweden 86%.

Forest supervisor education started at the end of the last century. In the 1950s and 1960s the forest supervisors’ education has been renewed in all four countries. Supervisors have a special 1 to 4.5 years’ training, but many have a forest technician’s education, too. In Finland and Sweden forest supervisor education was reformed at the end of the 1970s. Supervisors work in functional as well as in regional organizations. In Nordic countries, supervisors are paid monthly salaries. Salaries are a little higher in private than in public sector companies. In 1981 the start salary in Finland was 3,107 FIM and in Sweden in 1980 about 4,425 FIM. Most supervisors in the Nordic countries belong to some union.

The PDF includes a summary in English.

• Juntunen, E-mail: mj@mm.unknown
• Engsås, E-mail: je@mm.unknown
• Gustafsson, E-mail: lg@mm.unknown
• Mäkinen, E-mail: pm@mm.unknown
• Theilby, E-mail: ft@mm.unknown
• Vik, E-mail: tv@mm.unknown

This article reviews experiments and practical experience of forest management by the selection system in Finland. In an experiment of 25-year duration the annual growth of uneven-aged Norway spruce (Picea abies (L.) H. Karst.) stands was only about 50% of the average annual yield of even-aged stands in normal rotation on the same site.

In Finland the selection system is applicable under exceptional conditions only, viz. In intensively managed park stands and, on the other hand, on very marginal sites, e.g. on peat bogs and mountains near the tree-line. Even normal silviculture, however, may include cuttings which somewhat resemble selection system, e.g. removal of standards or restoration of mismanaged forests.

The PDF includes a summary in English.

• Mikola, E-mail: pm@mm.unknown

A dynamic programming approach toward stem value estimation for standing Scots pine (Pinus sylvestris L.) trees was developed. The determination of the saw log value was based on the sawing pattern and on the final products composition. The combination of taper curve models and bark models providing taper curves both over bark and under bark, which constituted the basis of the optimum stem scaling. A computer program was developed to determine the optimum log sequence of the stem aiming at maximizing the value of the final products. To examine the reliability of the computation system, 445 Scots pine sample trees from 29 stands were used as a test material. The stem values of sample trees were calculated in two ways: 1) with 12 measured diameters, and 2) with 12 estimated diameters derived from measured tree characteristics. In both cases the values of the intermediate diameters were calculated via cubic spline interpolation.

The PDF includes a summary in Finnish.

• Luo, E-mail: fl@mm.unknown

Application of nitrogen at levels of 200, 400 and 600 kg ha^-1 resulted in increases of 35, 18 and 12% in the photosynthetic rate in young Scots pine (Pinus sylvestris L.). The number of buds, degree of branching, and needle size were positively related to the amount of nitrogen applied. A 10–40% increase in the average needle area was found. A positive correlation was found between total photosynthesis and stem growth.

The PDF includes a summary in Finnish.

• Kellomäki, E-mail: sk@mm.unknown
• Puttonen, E-mail: pp@mm.unknown
• Tamminen, E-mail: ht@mm.unknown
• Westman, E-mail: cw@mm.unknown

The share of stem, branch and needle growth was dependent on the within-stand light regime in a young Scots pine (Pinus sylvestris L.) stand. The share of needle growth increased at the expense of stem and branch growth in poor light conditions. In good light condition the share of branch wood increased substantially. The share of stem wood growth was greatest in moderate shading, emphasizing the role of an adequate stand density for growing high-quality timber. The basic density of the stem wood was considerably greater in suppressed trees than in dominating trees. The differences were related to the illumination of the crown system.

The PDF includes a summary in Finnish.

• Kellomäki, E-mail: sk@mm.unknown

Length variation of fibres and vessels was studied in the branches, stems and roots of Betula pendula Roth and B. pubescense Ehrh. The cells were significantly shorter in the branches and roots than in the stems. There was no significant difference in the cell length between the upper and lower radii of the branches and roots. The length increased from the pith to the surface and decreased in the branches and stems from the base onwards. In the roots the length increased in that direction. The differences between the tree species were small although the cells of B. pubescens were a little longer.

The PDF includes a summary in Finnish.

• Bhat, E-mail: kb@mm.unknown
• Kärkkäinen, E-mail: mk@mm.unknown

Variation of cellular proportion within the same growth rings counted from the pith of the stems and branches in four trees of Betula pendula Roth was studied. The fibre percentage decreased from breast height to the crown and then increased in the branches. The reverse trend was found in the percentage of vessels and parenchyma, although the latter varied relatively little. No statistically significant differences were found in the percentages of fibres, vessels and rays within the same growth rings counted from the pith between the stems and branches. In both the stem and the branches, the proportion of fibres increased and that of vessels and rays decreased from the pith to the surface. Even crown formed wood differed from that of stem formed. 

The PDF includes a summary in Finnish. 

• Bhat, E-mail: kb@mm.unknown
• Kärkkäinen, E-mail: mk@mm.unknown

The article examines the problems of interdisciplinary research and the Finnish participation in MAB Project 2, which concentrates on the influences of man’s activities on forests. From the Finnish point of view, the main research areas are the effects of forestry activities which affect large areas, multiple use of forests, forests and environmental pollution, and the effects of energy economy.

This paper was presented in the ‘Man and the Biosphere’ programme project 2 seminar held on August 24–25 1978 in Hyytiälä research station of University of Helsinki.

The PDF includes a summary in English.

• Paavilainen, E-mail: ep@mm.unknown

Man and the Biosphere (MAB) programme of UNESCO was launched in 1970. This interdisciplinary programme represents a new integrated approach to research, training and action aimed at improving man’s partnership with the environment. It consists of 14 project areas.

The Academy of Finland and the Finnish Committee for the MAB, in cooperation with the University of Helsinki and the city of Tampere organized a seminar with an aim of reviewing the execution of the Finnish participation in the MAB project No. 2. The seminar took place at Hyytiälä, a forest research station of the University of Helsinki, on August 24–25 1978.

During the seminar, an excursion was made to Pyynikki esker, a unique natural park close to the centre of the city of Tamper. Eight papers were presented and discussed in the seminar. The papers are published in this issue of Silva Fennica.

The PDF includes a summary in English.

• Pärnänen, E-mail: ap@mm.unknown

The distribution of the dry matter and nutrients in Scots pine (Pinus sylvestris L.) tree stock growing on a Vaccinium type site, ground vegetation, and humus were determined in the study. The greatest part of the dry matter in the tree was found in the stemwood. The living branches, roots, bark, needles and dead branches decreasing order of magnitude made up the rest of the biomass. The trees contained over 90%, the field layer vegetation 3% and the bottom layer vegetation 2% of the dry matter in the tree stand. The tree stock contained 86–95% of the total amount nutrients in the stand. The field layer vegetation contained less nutrients than the bottom layer vegetation. Nitrogen, however, was an exception, the amount being approximately the same in both vegetation layers. 

The PDF includes a summary in English.

• Lehtonen, E-mail: il@mm.unknown

The study is part of a project designed to elucidate the nutrient budget of a Scots pine (Pinus sylvestris L.) stand. Results of variation in the nutrient contents were compared with those obtained for the previous growing season.

The potassium and calcium contents varied the greatest in the humus layer. However, in the bottom and field layer vegetation and needles the variation in the nitrogen content was the greatest. The nutrient contents of the needles were affected by the physiological stage of development, needle age and the position in the crown of the tree. The nutrient content of the rainfall increased in the order: free rainfall, throughfall, and stem flow.

The PDF includes a summary in Finnish.

• Lehtonen, E-mail: il@mm.unknown

The amounts of harvestable logging residues and stump and root wood were examined in the area where 100,000 solid m³ of stemwood was cut in 1975. The cutting amounts of stemwood from work sites suitable for harvesting of logging residues was 35,000 m³, and suitable for harvesting of stump and root wood 38,000 m³. The increase in the yield of wood (without bark) from logging residues compared with the unbarked stemwood was 2.4%. The same percentage of wood from stump and root wood was 5.0–5.8% depending on the harvesting loss.

The PDF includes a summary in Finnish.

• Mäkelä, E-mail: mm@mm.unknown

This study forms part of a project designed to elucidate the total nutrient budget of a Scots pine (Pinus sylvestris L.) stand in Central Finland during the 1974 growing period. Precipitation has been divided up into precipitation, throughfall and stemflow.

The acidity of the rainwater was found to increase in the order – Precipitation, throughfall and stemflow. The electrolyte content of the rain water from throughfall and stemflow was higher than that of precipitation. The nutrient contents of precipitation were rather low. Throughfall and stemflow were found to leach some nutrient from the trees. This was most clear in the case of potassium. The total amounts of nutrients reaching the ground in throughfall were found to be smaller than those in precipitation. On the other hand, the amounts of potassium, calcium and magnesium in stemflow were clearly greater than those in precipitation.

The PDF includes a summary in English.

• Lehtonen, E-mail: il@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown
• Westman, E-mail: cw@mm.unknown

In the article some aspects concerning the measurement of environmental factors are discussed. Special attention is given to the following questions: The correct way of determining the active surface in a forest ecosystem, the time factor in measurement processes, and the mutual correlative relationships between the environmental factors. Analysis of the data is also taken into consideration.

The PDF includes a summary in English.

• Leikola, E-mail: ml@mm.unknown

The problem considered in this paper is how to analyse the usefulness of some average volume values of stems in predicting the average time consumption of some processors. In this connection, the usual arithmetic mean, the arithmetic mean weighted with volume and the geometric mean were used. The functions describing the dependence of time consumption per tree on the volume of the tree were obtained from previous studies, as were the stand properties.

The results were obtained with the aid of computer simulation techniques and the deterministic model described above. According to the results, the usual arithmetic mean is very applicable for predicting the time consumption. The geometric mean is nearly as good, or in some cases even better. However, as a rule its computation is much more tedious than that of the arithmetic average volume for the purpose.

The PDF includes a summary in English.

• Kärkkäinen, E-mail: mk@mm.unknown

The material of 78 damaged Norway spruce (Picea abies (L.) H. Karst.) trees was gathered in Southern Finland in order to clarify the advance of decay. The harvesting which had caused the scars had been carried out 12 years earlier and at the moment of the investigation the growing stand was 110 years old. It was noticed that the variables used could explain only a few per cent of the variation of the advance of decay. It was concluded that the only important thing in practice is whether the injuries are in roots or in stems.

The PDF includes a summary in English.

• Kärkkäinen, E-mail: mk@mm.unknown

The paper describes a recommendation of a coordinate system to be used in all biological disciplines to report observations, agreed on by the natural history museums and the scientific societies in Finland. In the recommended coordinate system, the p-axis of the Cartesian coordinate system joins the E 27° longitude. The recommendation describes in detail how the observations are reported using this system.

• Ruutujärjestelmätoimikunta, E-mail: –

The purpose of this study was to explain whether it is possible to affect, in practical working site conditions, by means of logging waste on the strip road, the depth of the track which is formed in terrain transportation and the injuries of the growing stand. Five 20 m long investigation areas with logging waste and five similar areas without logging waste were arranged on one strip road at Teisko logging site in Southern Finland. The logging waste layer was mainly Norway spruce and 10–15 cm thick. A KL–836 B forwarder was used. The type of soil was loam.

The logging waste affected the depth of the track only by decreasing the wear of humus layer. Even decreasing effect of logging waste on the injuries in the growing stand was minor. At Kitee working site in Eastern Finland strip roads were studied. The type of soil was thick, rather mouldered peat. The thickness of logging waste was 3–4 times greater than in Teisko, mainly spruce. A Volvo Nalle SM 460 forwarder was used. The effect of the logging waste on the depth of the tracks was clearly to be noticed. On basis of the appearance of the tracks one could assume that the difference was due to different wear of the humus, and not so much due to the quantity of logging waste that improves the carrying capacity of terrain.

In some extent logging waste was also found to affect the amount and quality of tree injuries. In practical working conditions, the importance might be small, since in the experiments an unrealistically great amount of logging waste was used.

The PDF includes a summary in English.

• Kärkkäinen, E-mail: mk@mm.unknown

This project studied the value of various shoot and root-system characteristics as indicators of plantability of transplants. Correlation and regression analysis was used to compare these characteristics. The study material consisted of two-year Scots pine (Pinus sylvestris L.) transplants that had grown in a plastic greenhouse for the first year and then been transplanted in the open. The seedlings had been transplanted in the field without treatment or with the roots cut to a length of 8 cm. A part was transplanted without treatment into plastic pails. A gravimetric and photometric method was used to obtain a description of the surface area of the root systems.

The results show that the photometric value gives a good picture of the surface area of the root system. The greatest advantage offered by the method is the simplicity and rapidity of measurement. The gravimetric, and especially the titrimetric, measurement takes much more time per plant. Photometric measurement affects plantability little, and measured and planted transplants can be followed up in the field. In gravimetric measurements, it was found that fresh and dry weight of the plants were closely correlated.

Mycorrhizal frequency in the root systems gave a good picture of the surface area of the root system. The number of living roots-tips was also rather closely correlated with the surface area of the root system. The other morphological characteristics failed to serve as a satisfactory index for the surface area of root systems. The one closest correlated was the annual leader growth. The second best was stem diameter; the height of the plant, on the contrary, was rather poorly correlated with the other characteristics.

The PDF includes a summary in English.

• Lähde, E-mail: el@mm.unknown
• Oksanen, E-mail: ao@mm.unknown

Stand precipitation and stemflow studies became necessary in connection with hydrologic studies, for instance, to explain the deviations resulting from rains in the ratios between the water content of peat and the groundwater level, throughfall during rains of variable heaviness, and effect of stand treatment on soil moisture level. In this project the distribution of rainfall in stands differing in species composition and density was studied in Central Finland in 1963–1965 in fifteen stand precipitation sample plots. In addition, rain gauges were situated under individual Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies (L.) H. Karst.) and birch (Betula sp.) trees.

The average precipitation in the open was 4.8 mm, the corresponding precipitation in the stand was 77% for birch, 71% for pine and 62% for spruce. Measurements of stemflow from individual sample trees showed that less than ¼ mm (about 1.5%) during a 15 mm rain in a pine stand. In the spruce stands stemflow is negligible. A part of the sample plots was in drained peatlands with a dense vegetation of small shrubs. The shrub layer retention was about 10% even during heavy rain. In a small forest clearing, the bordering effect of the forest was seen up to the distance of 5 metres from the edge of the forest. During the period of study, on an average 3% more precipitation was recorded in the clearing than in the open, the difference being probably due to the stronger wind effect in the open.

The PDF includes a summary in English.

• Päivänen, E-mail: jp@mm.unknown

Exceptionally widespread snow damages occurred in January 1959 in the southern coastal region of Finland. An inquiry showed that significant devastation had occurred over an area of 42,620 ha. The purpose of the present investigation was to study the susceptibility to snow damages of different stands in different locations. Only the stem breakage was recorded. 924 stands along 92 one-kilometre lines were studied in the western continuation of Salpausselkä ridge in the summer 1960. A supplementary study was carried out in 1961 in separate stands.

Most heavily damaged stands were found in a damage zone closest (31–40 km) to the coast of Gulf of Finland. The damages were 39% fewer in the zone 61–70 km from the coast. No stands over 140 m above sea level escaped damage. Stands on the edge of an open area such as a field, lake etc. fared better than areas within the forest. Eastern slopes were more susceptible for snow damages in these weather conditions. Also, conifers were more frequently damaged than deciduous trees. Dense stands, and stands aged 61–100 years had most damages.

The PDF includes a summary in English.

• Suominen, E-mail: os@mm.unknown

The paper is a review on the topics of Symposium on forest types and forest ecosystems, held in connection to the IX internal botanical congress in Montreal in August 1959, the chairman of which was Ilmari Hustich. The article includes 18 preparatory papers that were distributed among the participants of the symposium. The common theme of the papers was the question of finding common platform for the different schools of forest types and forest ecosystems. In addition to the papers, the article includes a summary of the proceedings and discussions of the symposium.

The following papers were presented in the symposium:

Aichinger, E. Können wir eine gemeinsame Platform für die verscheidenen Schulen in der Waldtypenklassifikationen finden?

Arnborg, T. Can we find a common platform for the different schools of forest type classifications?

Dansereau, P. A combined structural and floristic approach to the definition of forest ecosystems.

Daubenmire, R. Some major problems in vegetation classification

Ellenberg, H. Können wir eine gemeinsame Platform für die verscheidenen Schulen in der Waldtypenklassifikationen finden?

Hills, G.A. Comparison of forest ecosystems (vegetation and soil) in different climatic zones

Kalela, A. Classification of the vegetation, especially of the forest, with particular reference to regional problems

Krajina, V.J. Can we find a common platform for the different schools of forest type classifications?

Kühler, A.W. Mapping tropical forest vegetation

Linteau, A. Y. a-t-il. Un terrain d’entente possible entre les différentes écoles au sujet de la classification de types forestiers?

Medvecka-Kornaś, A. Some problems of forest climaxes in Poland

Ovington, J.D. The ecosystem concept as aid to forest classification

Puri, G.S. The concept of climax in forest botany as applied in India

Rowe, J.S. Can we find a common platform for the different schools of forest type classifications?

Scamoni, A. Können wir eine gemeinsame Grundlage für die verscheidenen Schulen in der Waldtypenklassifikationen finden?

Sukachev, V.N. The correlation between the concept ’forest ecosystem’ and ’forest biogeocoenise’ and their importance for the classification of forests

Webb, L.J. A new attempt to classify Australian rain forest

• Heikurainen, E-mail: –
• Finnish Society of Forest Science, E-mail: –

There are three theories regarding the stem form of trees. The stem form plays a role in ability of the tree to resist wind and wind breaks. This article presents the theory and experiments about mechanic stem form theory. The wind velocity in a forest stand and the coefficient of resistance at the tree crone and at the tree stem are calculated. The hypothesis about the point when the tree stem breaks is discussed. The approximate values of different calculations are presented. 

• Ylinen, E-mail: ay@mm.unknown

Silva Fennica issue 52 includes presentations held in professional development courses, arranged for foresters working in public administration in 1938. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.

This presentation describes growth and form of root systems of different tree species in different sites and how growth of roots affect forest management.

• Aaltonen, E-mail: va@mm.unknown

Finnish tree species have adapted differently to heavy snow loads that occur especially in fell areas in Kuusamo and Salla as well as Maanselkä area in Sotkamo and Rautavaara in Northern Finland. Norway spruce (Picea abies Karst. L) is adapted better than Scots pine (Pinus sylvestris L.). The aim of this study was to investigate how crown and stem form of Norway spruce in the snow damage area of Maanselkä area differ from other areas in the same region.  

Relatively broad crown at the base of the stem, quickly tapering crown and narrow and even upper crown were typical for trees growing in the snow damaged areas. The higher the altitude is, the stronger tapering the crown is. The tapering begins usually in a height of 4-5 meters. Even the stem diameter begins to taper strongly at this height. In the areas where heavy snow does not cause snow damage, top of crown is broader. Also, in the snow damage areas the damaged trees seem to have broader crown shape than the trees with little damages.  

Height of the trees decreases in the snow damage areas compared to forests in lower altitudes, which can be caused both by wind and snow load. 

The article includes a German summary. 

• Mikola, E-mail: pm@mm.unknown

Silva Fennica Issue 39 includes presentations held in professional development courses in 1935 that were arranged for foresters working in public administration. The presentations focus on practical issues in forest management and administration, especially in regional level.

This presentation gives guidelines for draining of peatlands.

• Cautón, E-mail: ac@mm.unknown

One of the difficulties in constructing growth and yield tables has been to determine which of the sample plots growing the same tree species and belonging to the same forest site type, with reference to the quality of stands, have to be included in the same growth series.

New growth and yield tables for the most important tree species were constructed in Finland in 1916–1919, using new principles that aim at avoiding some of the common weaknesses. There were two main differences to the earlier work. First, the site quality class (forest site type) was determined for each sample plot when the sample plot was measured, independently of the stand occupying the site. In this way it was possible to treat the sample plots of each site as an independent group from the beginning, and so that the quality classes were the same for all the tree species. Second, mathematic-statistical methods were used to deduct the so-called stem frequency distribution series, when studying which of the sample plots of the same quality class belong to the same growth series. They represent the average number of stems of the different diameter classes. A more detailed description of the method used to create the growth and yield tables is published in Acta Forestalia Fennica no. 15.

In the PDF is included a summary in Finnish.

• Ilvessalo, E-mail: yi@mm.unknown

This paper reports on tests made for the study of alternative methods in forest survey. Data were acquired by measurements in five areas in Finland and in Mexico, varying in size from 20 to 900 ha. The principal characteristics used in the analysis was the entire volume. By the combination of neighbouring plots, the variation could be studied for different plot sizes and survey strips. Variable (relascope) plots could be compared.

A starting point for the comparison of different sampling methods, calculations were made of the coefficients of variation for each plot type; total and within the strata. The amount of decrease of variation with an increasing plot size could be established. Comparisons have been made of the following sampling methods: simple random, stratified random, simple systematic, and stratified systematic sampling.

On comparisons of the standard error of sample mean it was found that in both stratified sampling and different types of systematic sampling there is, with increasing size and diminishing interval of sample plots, an increase in the relative improvement of the result against simple random sampling. Only in exceptional cases did systematic surveys give results which were less precise than those derived by other methods.

In discussion of some methods for determination of the precision of systematic sampling, possibilities of theoretical determination of the degree of precision was considered. An empirical study was made of the behaviour of some equations based on the sample itself. The larger the plot size and the shorter the plot interval, the more the equations overestimated in general the variance of sample mean.

As none of the equations studied gave reliable results, regression equations were calculated for the relative standard error on the basis of the data measured. The independent variables were plot size, plot or strip interval, area of survey unit and mean volume. The results arrived at are based mainly on the complete measurement of one area only. To enable extension of the scope of application, more material is needed with a complete enumeration of trees.

The PDF includes a summary in Finnish.

• Nyyssönen, E-mail: an@mm.unknown
• Kilkki, E-mail: pk@mm.unknown
• Mikkola, E-mail: em@mm.unknown

Distribution of rainfall in in a Scots pine (Pinus sylvestris L.) stand and in an open place in Alajärvi in Central Finland was studies in 1959–1960. Density of the about 80 years old stand was 0.36 and the height of the trees 8–14 m. The dependence of throughfall and dependence of stemflow on 24-hour precipitation, and dependence of the distribution of 24-hour precipitation on the amount and nature of precipitation was calculated.

The precipitation of the crown of the forest depended on the rainfall. When the rainfall in the open place was over 7 mm, the rainfall within the forest was in average 89% of the rainfall in the open place, but if the rainfall in the open place was less than 1 mm, the rainfall within the forest was only 64% of that in the open place. Total stemflow in the pine stand was only 0.4%, and interception loss was 13.6%.

The PDF includes a summary in English.

• Seppänen, E-mail: ms@mm.unknown

Draining transforms root systems of trees growing in peatlands towards the ones growing on mineral soil. However, even after efficient draining the root systems differ from the root systems of trees growing on mineral soil. This investigation concentrates on root systems of forests of similar mire types growing in similar draining conditions but having different tree species compositions. The peatland, situated in Pieksämäki in Southern Finland, was drained in 1937. Sample plots, measured in 1956, consisted of mixed forest of Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies L. Karst.) and birch (Betula sp.) in different compositions, and were in natural condition.

The sedge pine bog studied in this investigation was shown to have larger total amount of roots and mycorrhiza than in previously studied dwarf shrub pine bogs. This reflects better growth conditions of the better site. The depth of root system was, however, similar. Root systems of birch were deeper than those of the coniferous tree species. Differences between Scots pine and Norway spruce were small. Corresponding differences between the species were found in the density and total number of mycorrhizas. The abundance of mycorrhizas in the roots of birch increased in deeper layers of peat, but decreased especially in spruce roots. In earlier studies the abundance of mycorrhizas decreased in the roots growing in deeper layers in pure Scots pine stands, but no such variation was seen in this study. The result suggest that the deep root system of birch may affect also the root systems of the coniferous trees. On the other hand, birch roots can have advantage over the coniferous trees.

The PDF includes a summary in German.

• Heikurainen, E-mail: lh@mm.unknown

The root system of a Scots pine (Pinus sylvestris L.) growing on a peatland is restricted, according to earlier studies, on the top layers of the peat above the groundwater level. Drainage of the peatland affects growth of the root system. This investigation aims at studying the root systems on the point of view of draining of peatlands. The structure and distribution, and the growth of mycorrhiza in Scots pine roots in pine swamps varying from natural state to well drained state is studied.

The study shows that Scots pine on pine swamps has more extensive root system than has earlier assumed, it is common to find 1,000 m of roots in one cubic meter in a healthy stand. The trees reach this density of roots early on. In a drained peatland, the total root length is markedly higher than in a similar stand in natural state. The root systems proved to be very shallow. Even in a well-drained site the roots did not grow deeper than 20 cm. 70% of all roots were found in the upper 5 cm layer of peat, and 90% in the upper 10 cm layer. Root systems were deeper in drained peatlands, but the difference was small. In a site in natural state the average depth of the roots was 4 cm, and in a drained site 5 cm. About 85% of the roots were under 1 mm of diameter. Short roots were found only in the fine roots. Draining increases strongly the number of short roots. Mycorrhizas of the types A, B, C and D as well as pseudomychorrizas were found in the pine roots.

The PDF includes a summary in German.

• Heikurainen, E-mail: lh@mm.unknown

The seasonal changes of the roots systems of a pine stand on the peatlands have been studied with samples collected during summers 1952-1954 and winter 1955. There are altogether seven sample areas that are located in the district of Korkeakoski.  

The amount of roots is at the smallest in the spring, increasing then rapidly and peaking at the end of July. After that the amount of roots decreases again against the winter to the same size than in the spring. Variations seem to be similar in every year and also similar to other studies on mineral soils.

The PDF contains a summary in Finnish. 

• Heikurainen, E-mail: lh@mm.unknown

The article presents the results of the studies about the horizontal root systems of pine. The results have been obtained with the method developed by the author.

The size of the root system varies very strongly during the growing period. The amount of roots is at the smallest in the spring, increasing then rapidly and peaking at the end of July. After that the amount of roots decreases again against the winter to the same size than in the spring. The differences are due the changes in the amount of the smallest roots. There seem to be no big differences in the amount of roots between stand of different ages.

After the thinning there is a drop in the amount of roots on the stand level, but after two growing periods then standing trees have taken over the unused land. However after selection felling the less vital trees are not capable of utilizing the vacant resource as effectively.     

The PDF contains a summary in Finnish. 

• Kalela, E-mail: ek@mm.unknown

Root systems of a Scots pine (Pinus sylvestris L.) stands of seed trees on a Vaccinium sites in Southern Finland were studied by taking soil samples around the seed trees. The results show that root system of an old Scots pine spreads relatively evenly around the tree up to at least 10 meters from the stem. The densest part of the root system is near the stem, which part is often acentric. This is probably due to root competition in the early stages of growth of the tree.

Root systems of the seed trees affect stocking of the site with seedlings and the growth of the seedlings. The root competition can cause, for instance, uneven grouping of the seedlings. It seems that the largest trees of a stand have the most even root system. It is therefore recommended to choose the strongest trees of the stand as seed trees, to ensure even distribution of seedlings.

The Acta Forestalia Fennica issue 61 was published in honour of professor Eino Saari’s 60th birthday.

The PDF includes a summary in German.

• Kalela, E-mail: ek@mm.unknown

About 17.5% of forest lands of Finland is situated around the river and lake system of Lake Saimaa. Furthermore, the growth of the forests  of the area correspond about 25% of the total growth of forests in Finland. The watersystem is one of the most important portages of roundwood in the country. It consists 11,000 km of floating channels, 2,000 km of which suit for floating in bundles. Annually 30-35 million cu ft of saw logs, 7 million cu ft of veneer timber and 2,5-3 million cu ft of pulp wood is floated in the area.

Even if the water system at present still suits well for floating, there are many opportunities for development, which would improve its competitiveness against other modes of transport. Several different sites where building of floating channels or improving the floatways are needed are described in the article.

The PDF includes a summary in German.

• Roitto, E-mail: wr@mm.unknown

Observations of connections between the roots of living trees and root systems of stumps have been reported already in 1900s. In Finland root connections have been found in Birch (Betula sp.), Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.), but there are no studies on abundance of the connections. This investigation studied root connections in a series of naturally regenerated Scots pine stands from seedling stands to mature trees in Southern Finland, and some sown seedling stands.

Root connections were found to be common in naturally regenerated, older stands that had passed the thicket stage. Approximately 21-28% of the trees had at least one root connection to another living tree, dead tree or living stump. Connections were few or absent in seedling stands. Sown seedling groups had many root connections in contrary to naturally regenerated seedling stands. Trees belonging to the dominating canopy class had most root connections. The trees could form a network of up to twenty trees and living stumps. Root connections were more common the larger the tree was or the nearer the trees grew each other. The coalescent roots were often situated near the stem. Experiments showed that water and nutrients transferred in the roots could move from one tree to another. Living stumps from previous fellings were relatively common. In the sites studied, there was in average 178 stumps connected to a living tree per hectare.

The PDF includes a summary in German.

• Yli-Vakkuri, E-mail: py@mm.unknown

Norway spruce (Picea abies (L.) Karst.) invading sites is common in Finland. The species tends to establish itself as undergrowth, and takes over when it gets space to grow. To determine whether the undergrowth is suitable as the new generation requires knowledge on the biology of spruce undergrowth. One of the issues is determining the age of the stunted trees. In this investigation, 100 undergrown spruce trees, their crown and their root systems were studied. A method was developed to determine the age of the trees.

The root system of all trees in Vaccinium sites and of stunted trees in Myrtillius sites were superficial. The root systems of older spruces were purely of adventitious origin. The longer the period of stunting growth, the younger is the root system. In addition to acropetal and general adventitious ramification there is often adventitious branching of the roots of pathological causes. Mortality among the long roots is frequent.

A stunted tree has not the same ability as a viable tree to make use of already existing branches for building assimilating surface. When comparing trees with equally large assimilating surface, a stunted tree had greater sum of roots compared to a viable tree. The root system of a stunted undergrown spruce was very superficial compared with the other trees.

The PDF includes a summary in English.

• Sirén, E-mail: gs@mm.unknown

The purpose of the investigation was to study the amount, quality and distribution by layers of depth of horizontal roots in Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) stands in Southern Finland. The sample plots included stands on soil varying from sandy to stony, and stands of  varying ages from seedling stands to an old stand, in Myrtillus and Vaccinium type forests.

In a Norway spruce stand, the amount of roots increases rapidly and reaches its maximum, about 450 meters/m³, at an age of 100-110 years. In a Scots pine stand the maximum, about 370 m/m³, is reached earlier, at an age of 60-70 years. The root system of pine expands more rapidly than that of spruce. The total length of the horizontal root system of pine amounts to 1,000 m soon after 40 years of growth, of spruce at the age of 60. Later the situation changes, and at the age of 110 the root systems of both species are about the same size, but older trees of spruce have more extensive root system.

Majority of horizontal roots are under 1 mm in diameter. Of the horizontal roots of spruce stands the majority lie in the humus layer and in the topmost mineral soil stratum. Over half of horizontal spruce roots are, thus, at a maximum depth of 5 cm, while majority of the roots of Scots pine lie at maximum in depth of 10 cm. At the same layer grow also the roots of the ground vegetation, which may affect the competition between the species.

The PDF includes a summary in English.

• Kalela, E-mail: ek@mm.unknown

The study is based on observations in a Scots pine (Pinus sylvestris L.) stand on a dry upland forest site in Karhumäki, where a 10-15-year old seedling stand grew under a hold-overs of larger trees that had been left in the site in a previous felling. The root systems of 80-120 cm tall seedlings growing around single mother trees were unearthed. Root maps were drawn of the root systems of 120 seedlings.

No seedlings grew around old, large hold-overs. It seems that seedlings could not compete with their root system. If the hold-overs were stunted in their growth, seedlings grew also under the canopy of the mother tree. 90% of the seedlings had a tap root. Rest of the roots grew horisontally in the topsoil. Around a vigorous mother tree, the seedlings grew their roots away from the mother tree. Hold-overs that had belonged originally to the lower canopy layer of the old forest did not have similar effect on the root orientation of the seedlings. Their roots had been previously affected by trees of higher canopy layer, later removed in the felling.

The PDF includes a summary in German.

• Kalela, E-mail: ek@mm.unknown

The effect of different kinds of injuries in the amount of merchantable timber was studied in 57 sample plots in the northernmost Finland. Without any injuries the yield of timber would have been 72.3% in Scots pine (Pinus sylverstris L.) and 89.9% in Norway spruce (Picea abies (L.) H. Karst.). Butting, and removal of parts of the stems due to Injuries decreased the volume by 10.4% in pine and 13.5% in spruce. The main cause for butting of pine was fire wounds, and butt rot in spruce. Also pine blister rust (Peridermium pini and Cronartium flaccidum) causes injuries in Scots pine. The better the forest site type, the smaller is the timber discarded due to injuries. In pine 54% and in spruce 53% of the trees and were healthy. The forests in the northernmost Finland are over-mature which increase the occurrence of fire wounds and decay. Thus, forest fire control and the felling or thinning of over-mature stands will improve the quality of the timber in the long run.

The PDF includes a summary in German.

• Tikka, E-mail: pt@mm.unknown

The stem form influences the value and volume of the stem. Sample trees in homogenous mixed stand of Scots pine (Pinus sylvestris L.) and Betula sp. were measured to define the stem form of the trees, and to develop research methods. The height of butt swelling and the turning point of taper curve varies greatly. In Scots pine and Betula sp. it was typically between the 2/10 and 3/10 height of the tree. Consequently, the theoretical normal curves describing stem form, where the turning point of taper curve is situated under the breast height diameter, are not entirely generally applicable. There was a correlation between the base curve and the form of actual taper curve of the stem. The form of the top of the stem depends on the structure and dimensions of the crown. The most reliable measuring point to define taper curve would be a diameter that is above butt swelling, near the turning point of the taper curve. Length of the crown can be used to deduce the form of the top of the stem. According to the study, the volume tables could be based on diameter on breast height, slenderness of the stem (D_0,25h:h) and length of the crown. Age of the tree and position in the stand influence stem form, but the forest site type seemed not to have clear effect on the stem form.

The PDF includes a summary in German.

• Lappi-Seppälä, E-mail: ml@mm.unknown

The aim of the study was to find out what are the causes of damage in different parts of the trees and the frequency of different kinds of injuries. Sample plots were studied in over 80-year old forests in mineral soil sites and peatlands. All the trees over 1.5 m high were felled in the sample plots and the stem injuries were studied. The structure of the stand and the crown classes were recorded. The proportion of undamaged trees was largest in in dominant and codominant trees and increased towards the better forest site types. The typical injuries are listed for Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies (L). H. Karst.) and Betula sp. stands. The injuries were divided in inner and outer form defects and injuries, and defined in more detail by the part of the stem and tree species. Defects caused by decay were analyzed separately.

Healing over of injuries was faster in the better sites. Form defects and other injuries were more common in birch stands than in Scots pine and Norway spruce stands. Decay was most common in birch stands. The pine stands were the healthiest, followed by spruce stands. Fire wound were most usual in pine, butt rot for spruce, and crooks and general decay for birch.

The PDF includes a summary in German.

• Tikka, E-mail: pt@mm.unknown

Forest work in Finland has generally been paid by the job. The wages in 1930s were based on variable measuring units and principles in different parts of the country. The rates did not often take into account how difficult the individual stands are to harvest. The studies have shown that the productivity of logging depends on size of the stems, density of the stand, branchiness of the trees, terrain and depth of snow cover. The article suggests a wage system which takes into account the differences of the cutting areas.

Wages of felling and hauling logs that are payed per the trunk or cubic meter should be adjusted so that the earnings of an employee depends not on the quality of the stand. The quality of the stand influences the stumpage price of the wood, because of the versatile harvesting costs. If the demands of the job in a challenging stand are not compensated for the workers, the higher harvesting costs are inequitably balanced by lowering the wages of the forest workers.

The PDF includes a summary in German.

• Vuoristo, E-mail: iv@mm.unknown

Knowledge on the roots systems and their properties is needed when for example assessing the wintering properties of a plant. The article presents the studies on the roots and their functions made with rye, wheat, oat and barley.

The data has been collected during the whole growing season. The experiments took place in the green houses of the University of Helsinki and on the experiment field in Tikkurila, some kilometres north from Helsinki.

The roots of cultivable crop can be divided according their function, state of development, structure and position in the root system into four classes. The classes are sprouting roots, nutriment roots, nutriment-support roots and support roots.

The PDF contains a summary in Finnish. 

• Kokkonen, E-mail: pk@mm.unknown

In practical forestry being able to divide the stem into timber assortments in the best possible way is very important.  Defining the decrease in diameter of the stem plays an important role in that. The article aims to define the slenderness as the relation of the tree height to its breast height diameter. The decrease in diameter is taken into account by measurements of diameter above the breast height. The study is based on the measurements conducted in 1924 for the inventory of forest resources of Finland.

The influence of the stand density to the relationship between height of the pine and the breast height diameter is smaller in the more fertile sites than in the less fertile sites. In the more dense stands are pines more slender than in the sparser stand on all forest types.  

• Lappi-Seppälä, E-mail: ml@mm.unknown

There are several competing theories about forestry: school on continuous forest management system, school on profitability of forestry and a dynamic view on economics. The article discusses the pros and cons of the different theories or viewpoints and wider the essence of forest as an economic asset.

To serve the needs of the practical forestry and the economics related to forests the forestry needs to seek for standardization in relation to growth potential of the forests and market orientation of the timber. A modern theory of forestry needs to base itself on both.  

The volume 34 of Acta Forestalia Fennica is a jubileum publication of professor Aimo Kaarlo Cajander.

• Hagfors, E-mail: eh@mm.unknown

In attempts to improve the autumn cereals, wheat and rye, hibernation plays an essential role. Those varieties that hibernate well should be marketed and others rejected. Concerning roots, it seems that varieties that hibernate well have more extent root system than those hibernating poorly.

Four varieties of rye were chosen for experiments, two that knowingly hibernate well and two that don’t. The experiments were grown in the Botanical gardens of the university and at the same time in experiment field in Tikkurila.

The results proof that plant hibernating well have more extensible roots than others and hence they survive better in frosting soil that extents.

The PDF contains a summary in English and in Finnish.

• Kokkonen, E-mail: pk@mm.unknown

The growth of a tree or a forest stand can be only fully understood when the form, encroachment and development of the root s in different environments are known. Research on roots has already yielded in practical silvicultural improvements and the purpose of this study is to discuss different factors about the roots.  

The literature review deals with the depth of root system, extent of it, the relation between root system and the soil structure, the form and volume of roots and the phenomenon where roots from several tree individuals grow together.

The PDF contains the article also in Finnish.   

• Laitakari, E-mail: el@mm.unknown

The root systems of 192 Scots pine (Pinus sylvestris L.) sample trees were dug out and measured in Säyneinen, Rautavaara and Pielijsäjvi in the Central Finland and Orivesi, Teisko and Hämeenkyrö in the Southern Finland. The volume of root system of Scots pine was always smaller than the stem, varying from 15% to 94% of the stem volume. The ratio is smaller in dense stands. The type of soil of the site affects how the central root system (tap root and the inner vertical roots) develop. This reflect the adaptability of the root system to different growth conditions. The root system may, for instance, substitute the tap root with stronger inner roots.

PDF includes a summary in English.

• Laitakari, E-mail: el@mm.unknown

Because the cross-sectional area of a tree stem is ellipsis, it cannot be determined exactly with only the diameter measurement.  The article presents a formula and the calculation of the exact cross-sectional area. If the cross-sectional area is calculated as arithmetic mean of two diameter measurement, varies the error between two limit values. The error becomes smallest by calculating the area based on the biggest and the smallest diameter measurements. 

• Heikkilä, E-mail: th@mm.unknown

Sample trees of Betula sp. were felled in North Karelia in different forest site types. The stands, both mixed and pure stands, had been regenerated in areas where shifting cultivation had been practiced. Sample trees represented breast height diameters up to 43 cm. Diameter was measured in distances of 1/10 of the height of the tree to calculate the stem form. The form factor was higher for the good forest site types than the poor sites. The volume tables were calculated based on the assumption that diameter does not affect the form factor. Comparing the volume table to the original data, it was found that the table seems to form a successful fitting of the data. Control data proved that the method seems to give a good fitting to the used data. Thus, the volume table can be used to measure volume of birch stands in North Karelia.

The PDF includes a summary in German.

• Hildén, E-mail: nh@mm.unknown

Volume formulas have been developed already for long time but there are still some questions to be solved. The volume of a stem can be calculated based on measurements of diameter on various heights or based on mathematic formula. The article presents the formula development for that. The study is based on measurements of 24 pine stands.   

The results of the new formula can be seen satisfactory with the relatively small data of this study. Coincidentally the formally best mean solution of the here compared volume calculation formulas was the one based on Denzin’s formula. There the deviation from total mean is almost zero.   

• Lönnroth, E-mail: el@mm.unknown

Approximately 320,000 hectares of peatlands and paludified lands had been drained for agriculture and forestry purposes in Finland by 1920. The ditch network was not optimal in the early drained areas, and the condition of the ditches declined over time. In this study, the condition of drainage system, and the natural processes that affect them, was inspected in 18 drained peatlands.

Several processes can reduce of the size of the ditches. The process of crumbling of soil in the sides of the diches can last up to three years after the draining. Erosion of the main drains, caused by water flow can, however, continue longer. The ditches also sink when the peat dries. Soil frost can affect the ditches, but as the vegetation grows its effect gradually decreases. The vegetation that grows in the ditches may finally block the ditch completely.

The PDF includes a summary in English.

• Kokkonen, E-mail: pk@mm.unknown

Shrinking of timber when drying is a phenomenon that causes variation in measuring of timber in timber trade and on using the timber for construction or other purposes.   

The data for the article consists of 332 increment core samples from pine trees different ages, sizes and growth rate. There were collected in years 1910-1912 in Finnish Lapland, regions Utsjoki and Inari. The increment cores were collected on the height of 1.3 meters in south-north direction straight crosswise through the whole tree. The samples are 6mm thick. The diameter of the samples was measured immediately after making the sample and after several years’ storage in room temperatures. Also the age of the trees was determined.

The results are presented in tables. The degree of shrinkage varies heavily between the samples but stays anyhow between 1.5 and 3.9%. The mean degree of shrinkage for 314 samples was 2.9%. The results seem to indicate that the bigger the shrinkage the denser the annual growth ring system of the tree, meaning the slower the growth has been.  The older and of diameter bigger trees shrink less than younger and smaller trees. 

• Renvall, E-mail: ar@mm.unknown

The data has been collected during 1919 and 1920 in different region of Finland. The studied peatlands varied from fuscum pine swamps to pine swamps and partly to better sedge pine swamps.

The study presents five different forms of root systems. The root growth of pine on peatlands seems to vary strongly from the root form on mineral soils. On the peatlands, where the ground water near to soil cover is, can the roots grow only near the soil surface where the conditions are suitable. For the pine typical tap root is in most cases absent or grows along the soil surface. Also the frost heaving, snow and characteristics of peat affect the root system.   

• Kokkonen, E-mail: pk@mm.unknown

Crown class is useful tool both in forest management and forest mensuration. The study presents a detailed crown classification for Scots pine (Pinus sylvestris L.). It was used to classify the sample trees prior detailed measurements of the crown and stem form. The stem form of a tree was dependent on which canopy layer it belonged. This relation was detected on both Vaccinium and Calluna site type forests. In addition, the stem tapers faster in poorer forest site types compared to better sites. The shorter the self-pruned part of the stem is, the faster the stem tapers. According to the study, the stems of stunted trees taper faster than trees of other crown classes. Also the age group affects stem form.

The PDF includes a summary in German.

• Lakari, E-mail: ol@mm.unknown

The first proper growth and yield tables were prepared in Finland already in 1872, but they have been used little as the needs of forestry and forest sciences increased. One of the problems of the old yield tables was how the site quality classes are determined. The new growth and yield tables use the forest site type classification, which enables the use of same site types for all tree species. This makes it possible to compare the growth of different tree species in same kind of sites. The tables also use stem frequency distribution series. In the first stage, the tables were prepared for Southern and Central Finland.

The PDF includes a summary in German.

• Ilvessalo, E-mail: yi@mm.unknown

The study presents and describes the abundance and distribution of tree roots in specific stands of heathy forest types in Lapland. The data was collected in the Sodankylä commune.  

Due to the shortcomings in the data, conclusions can be drawn only regarding pine forests. The result of study states that the root competition plays an important role in the development of the forests, and most of the other observed phenomena are linked with root competition. The more infertile the soil the vertically and horizontally wider and more abundant the root system. It seems that the abundance of the root system is similar in forest of same fertility class and same density and age.      

• Aaltonen, E-mail: va@mm.unknown

Diameter and volume increment as well as change in stem form of Scots pine (Pinus sylvestris L.) were analysed to predict tree increment variables. A stem curve set model is presented, based on prediction of the diameters at fixed angles in a polar coordinate system. This model consists of three elementary stem curves: 1) with bark, 2) without bark, and 3) without bark five years earlier. The differences between the elementary stem curves are the bark curve and the increment curve. The error variances at fixed angles and covariances between the fixed angles are divided into between-stand and within-stand components. Using principal components, the between-stand and within-stand covariance matrices are condensed separately for stem curve with bark, bark curve and increment curve. The two first principal components of the bark curve describe the vertical change in Scots pine bark type and the first principal component of the increment curve describes the increment rate. The elementary stem curves, bark curve and increment curve as well as corresponding stem volumes, bark volume and volume increment can be predicted for all trees in the stand with free choice of sample tree measurements. When only a few sample trees are measured, the stem curve set model gives significantly more accurate predictions of bark volume and volume increment for tally trees than does the volume method, which is based on the differences between two independent predictions of volume. The volume increment of tally trees can be predicted as reliably with as without measurement of sample tree height increment.

The PDF includes a summary in English.

• Ojansuu, E-mail: ro@mm.unknown

The vegetation and number of physical and chemical soil properties were studied on a random sample of closed upland forest stands in Southern Finland. The material consists of a total of 410 sample plots. Two-way indicator species analysis (TWINSPAN) was carried out in order to produce a hierarchical clustering of samples on the basis of the vegetation data. Discriminant analysis and analysis of variance were applied in order to find environmental correlations of the vegetation clustering.

The vegetation was found to indicate the nutrient regime of the humus layer well, but to a less extent the properties of the sub-soil. The understorey vegetation was found to be jointly dependent on the site fertility and on the properties of the tree stand, especially on the tree species composition. Although the forest vegetation appears to be distributed rather continuously along an axis of increasing site fertility, relatively unambiguous classification can be based on the appearance of indicator species and species groups.

The results of the study were interpreted as indication that operational site classification done using the vegetation is rather good method for classification in closed forest stands. Different methods produce relatively consistent, natural and ecologically interpretable classifications. The results also imply that the vegetation cover and the humus layer develop concurrently during the development of the ecosystem, but the differentiation of the site type is regulated simultaneously by a number of interacting factors ranging from mineralogical properties of the parent material to the topographical exposition of the site. As the plant cover depicts all these primary factors simultaneously, only a relatively rough ecological site classification can be based on the vegetation.

The PDF includes a summary in Finnish.

• Kuusipalo, E-mail: jk@mm.unknown

The aim of the study was to assess the contents and quantities of macronutrients reaching the ground with precipitation, stemflow and throughfall in Scots pine (Pinus sylvestris L.) stands growing on drained peatland, one of which was unfertilized and two of which had been fertilized three growing seasons before the measurements were carried out.

According to the results, the quantities of nutrients reaching the ground with precipitation were relatively large as compared, for example, with those removed with the stem wood carried away from the forest in logging. The nutrient most exposed to leaching from the canopy is potassium. Both the content of potassium in rainwater penetrating the canopy and the quantities reaching the ground are highest in stemflow, decreasing when moving from under the tree crowns toward the edge of the crown projection and into openings in the canopy. The results for phosphorus were similar, although not as clear as for potassium.

The contents of NO₃-N were smaller in stemflow than in precipitation. The results did not support assumptions according to which nitrate nitrogen is leached from the canopy or is taken up by the canopy from precipitation. In the case both of precipitation and of throughfall and stemflow, the quantities of nitrite nitrogen recorded were smaller than the degree of precision applied in the determinations carried out (0.01 mg/1). The contents of NH₄-N were on average higher in stemflow and throughfall than in precipitation.

Fertilizer application (600 kg/ha of N-P₂O₅-K₂O, 14-18-10) increased the contents of potassium in stemflow and throughfall. A slight increase in phosphorus was also observed. Leaching of inorganic nitrogen was not affected by fertilization.

The PDF includes a summary in Finnish.

• Päivänen, E-mail: jp@mm.unknown

The study material was collected from 10 localities in South Finland in 1971–72. The material comprised 816 damaged Norway spruce (Picea abies (L.) H. Karst.) trees with a total of 978 injuries.

Decay (discoloration) spread upward from the damaged point was about three times as fast as downward. The mean rate of advance upward was 21 cm/year. The decay spreading at the quickest rate started from above-ground root collar injuries. The size of the damaged area (surface area, width and depth) correlated positively with the rate of increase in decay initiated by the injury. For the first 10 years the decay advanced at the same rate after which the advance became slower though not ceasing. Damage produced in the early summer caused a faster spread of decay than that produced in the late summer or winter. The rate of advance was the greater the larger the stem involved. When decay started from trunk damage its rate of advance was greater the faster the growth of the trees. With a better soil type, the rate of advance in decay increased. Fertilization increased the rate of advance.

The widest stem injuries reduced tree growth by about one-third, and severed roots by nearly half of the growth of trees where the width of the injuries was 0–4 cm. Fomes annosus (Heterobasidion annosum) infected spruce injuries especially in the southern coastal district. The farthest tips of discoloration proved in most cases to be sterile. The most common fungus isolated from these sites was Stereum sanguinolentum.

The PDF includes a summary in Finnish.

• Isomäki, E-mail: ai@mm.unknown
• Kallio, E-mail: tk@mm.unknown

Emphasis was laid on the finding of regression equations to indicate the dependence of standard error upon various variables in systematic sampling. As a result, the size of sample for a given precision could be computed, under varying alternatives of sample plot size and type. Another task was that of examining inventory costs by means of time studies. On combination of the results in regard to the sample size and survey time, the relative efficiency of different alternatives could be discussed, with a view to the precision of the total volume of growing stock.

The PDF includes a summary in Finnish.

• Nyyssönen, E-mail: an@mm.unknown
• Roiko-Jokela, E-mail: pr@mm.unknown
• Kilkki, E-mail: pk@mm.unknown

• Viljanen, University of Helsinki, Faculty of Agriculture and Forestry, Latokartanonkaari 7, 00014 Helsingin yliopisto, Finland https://orcid.org/0000-0001-7166-8022 E-mail: anne.viljanen@helsinki.fi
• Kurttila, Natural Resources Institute Finland (Luke), Research and Customer Relationships, Yliopistokatu 6, FI-80100 Joensuu, Finland https://orcid.org/0000-0001-5290-4771 E-mail: mikko.kurttila@luke.fi
• Toppinen, University of Helsinki, Faculty of Agriculture and Forestry, Latokartanonkaari 7, 00014 Helsingin yliopisto, Finland https://orcid.org/0000-0003-0910-1505 E-mail: anne.toppinen@helsinki.fi

The EU’s influence on national forest policies is growing, and the implementation of forest-related policies proposed by the Commission will affect the practice of forestry in Europe. For instance, the Nature Restoration Law sets concrete areal goals for restoring forest ecosystems and for conservation, the Deforestation Regulation requires meticulous tracking of wood’s origin, and the renewed Renewable Energy Directive (RED III) sets new criteria to sustainable forest biomass procurement. So far there have been no studies that have looked into the impacts from the economic and operational point of view. In this study, structural systems analysis was first performed to discover the relevant variables (and their functioning) associated with the roundwood harvesting operations and the operating environment. A scenario approach was then applied to capture the potential levels of implementation of the EU’s forest-related policies. Finally, using different scenarios (low-, moderate- and high-impact) and a systems analysis framework, the impact of alternative levels of implementation was quantified in terms of harvesting costs, measured in € m^–3. The results indicate that with the low- and moderate-impact scenarios the harvesting costs would increase by less than 10% from the current levels in three different regions in Finland. Such an increase (less than 10%) could be tolerated over a period of a few years, but a sudden increase is likely to lead to challenges to the running of businesses. With the high-impact scenario the harvesting costs would increase by between 15% and 18%, depending on the region. This magnitude of increase (of approximately a sixth) corresponds to a severe change in the roundwood harvesting operations and operating environment.

• Ahtikoski, Natural Resources Institute Finland (Luke), Tekniikankatu 1, FI-33720 Tampere, Finland https://orcid.org/0000-0003-1658-3813 E-mail: anssi.ahtikoski@luke.fi
• Väätäinen, Natural Resources Institute Finland (Luke), Yliopistonkatu 6, FI-80100 Joensuu, Finland https://orcid.org/0000-0002-6886-0432 E-mail: kari.vaatainen@luke.fi
• Anttila, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland https://orcid.org/0000-0002-6131-392X E-mail: perttu.anttila@luke.fi
• Laitila, Natural Resources Institute Finland (Luke), Yliopistonkatu 6, FI-80100 Joensuu, Finland E-mail: juha.laitila@luke.fi
• Mutanen, Natural Resources Institute Finland (Luke), Yliopistonkatu 6, FI-80100 Joensuu, Finland https://orcid.org/0000-0003-0533-9356 E-mail: antti.mutanen@luke.fi
• Lindblad, Natural Resources Institute Finland (Luke), Yliopistonkatu 6, FI-80100 Joensuu, Finland https://orcid.org/0009-0003-6766-6587 E-mail: jari.lindblad@luke.fi
• Sikanen, Natural Resources Institute Finland (Luke), Yliopistonkatu 6, FI-80100 Joensuu, Finland E-mail: lauri.sikanen@luke.fi
• Routa, Natural Resources Institute Finland (Luke), Yliopistonkatu 6, FI-80100 Joensuu, Finland https://orcid.org/0000-0001-7225-1798 E-mail: johanna.routa@luke.fi

We used forest ecosystem model simulations to study how forest conservation and management intensity affected timber yield, ecosystem carbon stocks, amount of dead wood, and habitat suitability area in a middle boreal forest region of Finland under changing climate over a 90-year simulation period. We used the following forest conservation and management scenarios: baseline forest management (BM), BM with 10 or 20% increase of conservation area with or without intensified forest management (i.e. improved forest regeneration material and forest fertilization). The simulations were done under current climate (reference period of 1981–2010), and Representative Concentration Pathway (RCP) climate change projections under the RCP2.6 and RCP4.5 forcing scenarios. Overall, increasing the forest conservation area decreased timber yield and increased the ecosystem carbon stock, the amount of dead wood and consequently the area of suitable habitat for saproxylic species. The use of intensified forest management reduced the loss of timber yield, increased ecosystem carbon stock, the amount of dead wood and area of suitable habitat for saproxylic species. At the end of simulation period, the use of intensified forest management even overcompensated (4–6% higher) the timber loss from 10% increase of conservation area. Under changing climate, timber yield, the amount of dead wood and the area of suitable habitats for saproxylic species increased. To conclude, with intensified forest management it is possible, in the short term, to decrease the loss of timber yield through increased forest conservation area and in the long term maintain or even increase it compared to baseline forest management.

• Pikkarainen, School of Forest Sciences, University of Eastern Finland, Yliopistokatu 7, FI-80100 Joensuu, Finland https://orcid.org/0000-0001-5301-3639 E-mail: laura.pikkarainen@uef.fi
• Strandman, School of Forest Sciences, University of Eastern Finland, Yliopistokatu 7, FI-80100 Joensuu, Finland https://orcid.org/0000-0002-9400-6424 E-mail: harri.strandman@uef.fi
• Vento, School of Forest Sciences, University of Eastern Finland, Yliopistokatu 7, FI-80100 Joensuu, Finland E-mail: eerik.vento@gmail.com
• Petty, School of Forest Sciences, University of Eastern Finland, Yliopistokatu 7, FI-80100 Joensuu, Finland https://orcid.org/0009-0006-6595-1386 E-mail: aaron.petty@uef.fi
• Tikkanen, School of Forest Sciences, University of Eastern Finland, Yliopistokatu 7, FI-80100 Joensuu, Finland https://orcid.org/0000-0002-3875-2772 E-mail: olli-pekka.tikkanen@uef.fi
• Kilpeläinen, School of Forest Sciences, University of Eastern Finland, Yliopistokatu 7, FI-80100 Joensuu, Finland https://orcid.org/0000-0003-4299-0578 E-mail: antti.kilpelainen@uef.fi
• Peltola, School of Forest Sciences, University of Eastern Finland, Yliopistokatu 7, FI-80100 Joensuu, Finland E-mail: heli.peltola@uef.fi

A new method for the co-registration of single tree data in forest stands and forest plots applicable to static as well as dynamic data capture is presented. This method consists of a stem diameter weighted linking algorithm that improves the linking accuracy when operating on diverse diameter stands with stem position errors in the single tree detectors. A co-registration quality metric threshold, Q_T, is also introduced which makes it possible to discriminate between correct and incorrect stem map co-registrations with high probability (>99%). These two features are combined to a simultaneous location and mapping-based co-registration method that operates with high linking accuracy and that can handle sensors with drifting errors and signal bias. A test with simulated data shows that the method has an 89.35% detection rate. The statistics of different settings in a simulation study are presented, where the effect of stem density and position errors were investigated. A test case with real sensor data from a forest stand shows that the average nearest neighbor distances decreased from 1.90 m to 0.51 m, which indicates the feasibility of this method.

• Olofsson, Section of Forest Remote Sensing, Department of Forest Resource Management, Swedish University of Agricultural Sciences, Umeå, Sweden https://orcid.org/0000-0002-2836-2316 E-mail: kenneth.olofsson@slu.se
• Holmgren, Section of Forest Remote Sensing, Department of Forest Resource Management, Swedish University of Agricultural Sciences, Umeå, Sweden https://orcid.org/0000-0002-7112-8015 E-mail: johan.holmgren@slu.se

The widespread cork oak (Quercus suber L.) mortality and reduced afforestation /regeneration are causing an overall reduction in cork production. To enhance trees’ growth and vitality, afforestation techniques using fertirrigation were tested. The main objective was the promotion of trees’ growth on new dense plantations using minimum water requirements until reaching productive forests. The experimental plot – Irricork – was installed in 2017 in a ≈1 ha stand with 14 years’ age cork oaks summer-fertirrigated since plantation. Four fertirrigation treatments were applied during fertirrigation campaigns. Radial growth, meteorological parameters and fertirrigation volume were measured every 15–30 days over four years. It was observed that weather, tree size, debarking and trees’ intra-competition had a significant effect on radial increments. Fertirrigation significantly enhanced growth during summer drought and decoupled increments from air vapor pressure deficit constraints. There was a linear relationship between trees’ radial increments and fertirrigation volume up to 140 m³ week^–1. Above this value, increments were smoother. In conclusion, summer fertirrigation of 140 m³ week^–1 efficiently enhanced the radial growth of trees with 50–75 circumference at breast height, under the particular edaphoclimatic conditions of the stand. This study showed to be, therefore, promising in the use of efficient fertirrigation the enhance cork oaks’ radial growth.

• Camilo-Alves, MED – Mediterranean Institute for Agriculture, Environment and Development & CHANGE – Global Change and Sustainability Institute, Institute for Advanced Studies and Research, University of Evora, Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal https://orcid.org/0000-0001-5156-172X E-mail: calves@uevora.pt
• Nunes, Department of Plant Science, School of Science and Technology, University of Evora, Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal https://orcid.org/0000-0002-6144-3484 E-mail: jain@uevora.pt
• Poeiras, MED – Mediterranean Institute for Agriculture, Environment and Development & CHANGE – Global Change and Sustainability Institute, Institute for Advanced Studies and Research, University of Evora, Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal https://orcid.org/0000-0002-6049-807X E-mail: apcp@uevora.pt
• Ribeiro, Department of Plant Science, School of Science and Technology, University of Evora, Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal https://orcid.org/0000-0001-7242-5866 E-mail: jmrpr@uevora.pt
• Dinis, https://orcid.org/0000-0001-6984-1033 E-mail: dinis.cati@gmail.com
• Barroso, MED – Mediterranean Institute for Agriculture, Environment and Development & CHANGE – Global Change and Sustainability Institute, and Department of Plant Science, School of Science and Technology, University of Evora, Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal https://orcid.org/0000-0002-0160-3845 E-mail: jmmb@uevora.pt
• Vaz, MED – Mediterranean Institute for Agriculture, Environment and Development & CHANGE – Global Change and Sustainability Institute, and Department of Biology, School of Science and Technology, University of Evora. Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal https://orcid.org/0000-0002-3368-757X E-mail: mvaz@uevora.pt
• Almeida-Ribeiro, ICT – Institute of Earth Sciences and Department of Plant Science, School of Science and Technology, University of Evora, Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal https://orcid.org/0000-0002-0160-3845 E-mail: nmcar@uevora.pt

Because today’s tree planting machines do a good job silviculturally, the Nordic forest sector is interested in finding ways to increase the planting machines’ productivity. Faster seedling reloading increases machine productivity, but that solution might require investments in specially designed seedling packaging. The objective of our study was to compare the cost-efficiency of cardboard box concepts that increase the productivity of tree planting machines with that of today’s two most common seedling packaging systems in southern Sweden. We modelled the total cost of these five different seedling packaging systems using data from numerous sources including manufacturers, nurseries, contractors, and forest companies. Under these southern Swedish conditions, the total cost of cardboard box concepts that increase the productivity of intermittently advancing tree planting machines was higher than the cost of the cultivation tray system (5–49% in the basic scenario). However, the conceptual packaging system named ManBox_fast did show promise, especially with increasing primary transport distances and increased planting machine productivities and hourly costs. Thus, our results show that high seedling packing density is of fundamental importance for cost-efficiency of cardboard box systems designed for mechanized tree planting. Our results also illustrate how different factors in the seedling supply chain affect the cost-efficiency of tree planting machines. Consequently, our results underscore that the key development factor for mechanized tree planting in the Nordic countries is the development of cost-efficient seedling handling systems between nurseries and planting machines.

• Ersson, SLU, School of Forest Management, SE-739 21 Skinnskatteberg, Sweden https://orcid.org/0000-0003-2442-7482 E-mail: back.tomas.ersson@slu.se
• Sundblad, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: lars-goran.sundblad@skogforsk.se
• Manner, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden https://orcid.org/0000-0002-4982-3855 E-mail: jussi.manner@skogforsk.se

Management of Scots pine (Pinus sylvestris L.) in Norway requires a forest growth and yield model suitable for describing stand dynamics of even-aged forests under contemporary climatic conditions with and without the effects of silvicultural thinning. A system of equations forming such a stand-level growth and yield model fitted to long-term experimental data is presented here. The growth and yield model consists of component equations for (i) dominant height, (ii) stem density (number of stems per hectare), (iii) total basal area, (iv) and total stem volume fitted simultaneously using seemingly unrelated regression. The component equations for stem density, basal area, and volume include a thinning modifier to forecast stand dynamics in thinned stands. It was shown that thinning significantly increased basal area and volume growth while reducing competition related mortality. No significant effect of thinning was found on dominant height. Model examination by means of various fit statistics indicated no obvious bias and improvement in prediction accuracy in comparison to existing models in general. An application of the developed stand-level model comparing different management scenarios exhibited plausible long-term behavior and we propose this is therefore suitable for national deployment.

• Kuehne, Norwegian Institute of Bioeconomy Research, Division of Forestry and Forest Resources, P.O. Box 115, NO-1431 Ås, Norway E-mail: christian.kuehne@nibio.no
• McLean, Norwegian Institute of Bioeconomy Research, Division of Forestry and Forest Resources, P.O. Box 115, NO-1431 Ås, Norway E-mail: paul.mclean@nibio.no
• Maleki, Norwegian Institute of Bioeconomy Research, Division of Forestry and Forest Resources, P.O. Box 115, NO-1431 Ås, Norway E-mail: kobra.maleki@nibio.no
• Antón-Fernández, Norwegian Institute of Bioeconomy Research, Division of Forestry and Forest Resources, P.O. Box 115, NO-1431 Ås, Norway E-mail: clara.anton.fernandez@nibio.no
• Astrup, Norwegian Institute of Bioeconomy Research, Division of Forestry and Forest Resources, P.O. Box 115, NO-1431 Ås, Norway E-mail: rasmus.astrup@nibio.no

For constructing growth and yield models the concept of site index as measure of productivity is crucial. Here, we use nonlinear mixed-effects models (NLME) with random individual effects and nonlinear models with dummy variables as fixed individual effects (NLFE) to fit mechanistic growth functions to stem analysis data of the economically most important tree species in Zhongtiaoshan forest region, China. The Richards and Lundqvist function are formulated into five dynamic equations (R1, R2, L1, L2 and L3) applying the generalized algebraic difference approach (GADA), which inherit polymorphism, varying asymptotes and base-age invariance. According to Akaike information criterion the R1 model as NLFE fits height growth data of Pinus tabuliformis Carrière, Pinus armandii Franch., Quercus liaotungensis Koidz., Quercus aliena Blume and Betula platyphylla Sukaczev best, while for Quercus variabilis Blume R2 as NLFE fits height growth data best. For Larix principis-rupprechtii Mayr L1 as NLME has been selected as best model, as R1 and R2 both as NLFE and NLME are not extrapolating the comparably short length of height growth data well enough. However, according to the root mean square error and bias differences between model fits of both the selected equation and the chosen model fitting approach are not so clear. Presented families of height growth curves serve as planning tools to identify site index and therefore assess productivity of forest stands in the studied region. A direct comparison of the productivity of forest stands of the same tree species is possible due to base-age invariance of the selected models.

• Sprengel, Chair of Forest Growth and Dendroecology, Albert-Ludwigs-University Freiburg, Tennenbacher Str. 4, 79106 Freiburg, Germany https://orcid.org/0000-0002-6332-7911 E-mail: lars.sprengel@iww.uni-freiburg.de
• Spiecker, Chair of Forest Growth and Dendroecology, Albert-Ludwigs-University Freiburg, Tennenbacher Str. 4, 79106 Freiburg, Germany E-mail: instww@uni-freiburg.de
• Wu, Research Institute of Forest Policy and Information, Chinese Academy of Forestry, No. 1 Dongxiaofu, Haidian District, Beijing 100091, China E-mail: shuirongwu@126.com

Young, dense forest in Finland and Sweden urgently need to receive first thinning. In such stands, conventional selective thinning methods make the harvester work time consuming and, thus, costly. To make small-sized trees economically competitive as raw material for bioenergy and biorefining, new harvesting technologies and/or thinning methods need to be developed. A potential solution is boom-corridor thinning (BCT), rendering effective cutting work. The aim of this study was to describe and compare the stand structure of two Scots pine stands (Pinus sylvestris L.) and one birch-dominated (Betula pendula Roth with natural downy birch, B. pubescens Ehrh.) stand after BCT and selective thinning at the first thinning phase. Furthermore, simulations were conducted to predict the future stand development after the first thinning treatments. The density of the growing stock was 16–46% higher after BCT treatment than after selective thinning because BCT stands included more small and supressed trees with a dbh < 100 mm. However, the numbers of future crop trees with a dbh > 140 mm per hectare were at the same level in both treatments. The stem volume removal per hectare did not differ between treatments. However, simulation of stand development and intermediate thinning and clearcutting revealed that the total removal volume was 10–18% higher in BCT stands compared to selectively thinned ones. The saw log volumes harvested did, however, not differ between treatments. This study shows that BCT generates stands with higher biodiversity compared to conventional thinning as higher levels of biomass removal can be reached throughout stand rotations.

• Nuutinen, Natural Resources Institute Finland (Luke), Production systems, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: yrjo.nuutinen@luke.fi
• Miina, Natural Resources Institute Finland (Luke), Natural resources, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jari.miina@luke.fi
• Saksa, Natural Resources Institute Finland (Luke), Natural resources, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@luke.fi
• Bergström, Swedish University of Agricultural Sciences (SLU), Dept of Forest Biomaterials and Technology, Section of Forest Operations, SE-90183 Umeå, Sweden E-mail: dan.bergstrom@slu.se
• Routa, Natural Resources Institute Finland (Luke), Production systems, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: johanna.routa@luke.fi