Articles containing the keyword 'variation'

The horizontal and vertical stand structure of living trees was examined in a managed and in a primeval Norway spruce-dominated forest in Southern Finland. Tree size distributions (DBHs, tree height) were compared using frequency histograms. The vertical distribution of tree heights was illustrated as tree height plots and quantified as the tree height diversity (THD) using the Shannon-Weaver formula. The horizontal spatial pattern of trees was described with stem maps and quantified with Ripley's K-function. The spatial autocorrelation of tree sizes was examined with semivariogram analysis. In the managed forest the DBH and height distributions of trees were bimodal, indicating a two-layered vertical structure with a single dominant tree layer and abundant regeneration in the understory. The primeval forest had a much higher total number of trees which were rather evenly distributed in different diameter and tree height classes. The K-function summaries for trees taller than 15 m indicated that the primeval stand was close to complete random pattern. The managed stand was regular at small distances (up to 4 m). The semivariograms of tree sizes (DBH tree height) showed that the managed forest had a clear spatial dependence in tree sizes up to inter-tree distances of about 12 meters. In contrast, the primeval spruce forest had a variance peak at very short inter-tree distances (< 1 m) and only weak spatial autocorrelation at short inter-tree distances (1–5 m). Excluding the understory trees (h < 15 m) from the analysis drastically changed the spatial structure of the forest as revealed by semivariograms. ln general, the structure of the primeval forest was both horizontally and vertically more variable and heterogeneous compared to the managed forest. The applicability of the used methods in describing fine-scale forest structure i discussed.

• Kuuluvainen, E-mail: tk@mm.unknown
• Leinonen, E-mail: kl@mm.unknown
• Nygren, E-mail: mn@mm.unknown
• Penttinen, E-mail: ap@mm.unknown

Spatial variation in the density of soil organic carbon (kg/m²) and the thickness of soil horizons (F/H, E) were investigated in a 6 m x 8 m area in Scots pine (Pinus sylvestris L.) stand in Southern Finland for designing an effective sampling for the C density and studying the effect of trees on the variation. The horizon thickness of the podzolized soil were measured on a total of 126 soil cores (50 cm deep) and the C density of the organic F/H and 0–10 cm, 10–20 cm and 20–40 cm mineral soil layers was analysed.

The C density varied 3–5 fold within the layers and the coefficients of variation ranged from 22 % to 40%. Considering the gain in confidence per sample, 8–10 samples were suggested for estimating the mean C density in the F/H and 0–40 cm layers, although about 30 samples are needed for 10% confidence in the mean. The C densities and horizon thicknesses were spatially dependent within the distances of 1–8 m, the spatial dependence accounting for 43–86% of the total variance. The F/H layer was thicker and contained more C within 1–3 m radius from trees. In the 10–20 cm and 20–40 cm layers (B horizon) the C density also increased towards the trees, but more pronouncedly in the immediate vicinity of the stems. Because the spatial patterning of the E horizon thickness was similar, the increase was attributed to stemflow and precipitation of organic compounds in the podzol B horizon.

• Liski, E-mail: jl@mm.unknown

Spatial and age-related variation in nutrient concentrations of Scots pine (Pinus sylvestris L.) needles was studied during 1984–86 in three stands of different stages of development. The dry weight of current needles was significantly (p < 0.05) higher in the tree top than in a composite sample representing the whole crown. However, there were no significant differences in the concentrations of nutrients in needles between upper and lower crown levels. The concentrations of mobile nutrients N, P, K and Mg decreased with increasing needle age whereas the concentrations of poorly mobile nutrients Ca, Mn and Fe increased during needle ageing. The coefficient of variation for nutrient concentrations varied irregularly when only a few trees were sampled but stabilized when tree number was ten or more.
The PDF includes an abstract in Finnish.

• Helmisaari, E-mail: hh@mm.unknown

Forest tree breeding involves manipulation of genetic composition of populations and individuals, and biotechnology focuses on selected individuals. The new techniques cannot replace the conventional breeding techniques but both need effective cooperation of each other. Thus, the distinction between conventional breeding and biotechnology is artificial. The biotechnology methods are new and fast developing and the future with field and progeny testing will show which techniques will be permanently adopted into tree breeding. For instance, the earlier hope of the use of somaclonal variation as a new source of variability and a powerful tool for the breeder seem today quite the opposite. Somaclonal variation constituting a major problem in present-day micropropagation is due to the unpredictable variation. Based on knowledge of today, especially micropropagation via somatic embryos, transgenic trees and the identification of major genes seem to be good candidates to be permanently adopted into tree breeding.

• Häggman, E-mail: hh@mm.unknown

Shoot elongation of Pinus kesiya Royle ex Gordon was studied using 2-year old grafts in a clonal seed orchard of the Pine Improvement Centre, located at the Huey Bong Experimental Station near Chiangmai, Thailand (19° 17’ N, 99° 15’ E, 900 m a.s.l.).

The seed orchard had a completely randomized block design with 30 blocks and 80 single-tree plots (clones) in each block. Eleven clones in four blocks were selected out of the total of 80 grafts (clones). From each graft, three lateral branches at the height of 1.6 m from the ground level were selected. Thus, total of 109 branches were measured. Shoot length of branches was measured between July 3, 1983 and March 11, 1984 at approximately bi-weekly intervals. Method of classical growth analysis were used in describing the shoot growth.

The annual shoot growth pattern of P. kesiya exhibited two consecutive sigmoid growth curves, i.e. it consisted of two flushes of shoot elongation, both formed by free growth. Thus, the pattern of shoot growth resembled the caribaea pattern. However, the annual shoot was composed of summer and winter shoots. These could be distinguished from each other by the reproductive organs, which always occur on winter shoot. The shoot contributed 61% of the total annual shoot length.

There were significant differences in the pattern of shoot elongation between the studied clones, which may reflect differences in the adaptation to different environmental conditions.

The PDF includes an abstract in Finnish.

• Sirikul, E-mail: ws@mm.unknown
• Kanninen, E-mail: mk@mm.unknown

The study uses the methodology of ecological field theory to model the effect of Scots pine (Pinus sylvestris L.) seed trees on the density of tree seedlings and other plants in the field layer. The seed trees had a clear effect on the expected value of the amount and distribution of the ground vegetation. The vicinity of seed trees had an adverse effect on the growth of grasses, herbs and seedlings, while mosses were most abundant near the trees. Models based on the ecological field approach were derived to describe the effect of seed trees on the ground vegetation.

The PDF includes an abstract in Finnish.

• Kuuluvainen, E-mail: tk@mm.unknown
• Pukkala, E-mail: tp@mm.unknown

This issue of Silva Fennica consists of eight articles, which are based on a co-nordic conference ”Frost hardiness and over-wintering in forest tree seedlings”, held in Joensuu, Finland, during December 1–3, 1986. The whole annual cycle of the trees is considered. Emphasis is given on methods for the study of frost hardiness, genetic variation in frost hardiness, nitrogen metabolism, bud dormancy release, and joint effect of natural and anthropogenic stress factors in the winter damage of forest trees. Practical implications for tree breeding and nursery management are discussed.

The PDF includes an abstract in Finnish.

• Hänninen, E-mail: –
• Pelkonen, E-mail: –

The purpose of the study was to determine the effects of the origin of seeds and the location of cultivation of Scots pine (Pinus sylvestris L.) on certain properties particularly important to the pulp industry. The research material consisted of six parallel trials of the same 12 provenances. Increment cores were taken of a total of 1,267 sample trees, 19 years old. The location of the trial site generally affected the properties to a larger extent than the origin of the seed. The effect of the variation of wood density and fibre yield on the cultivation values of the provenances was only a few percentages on average, however, at most the effect was nearly 10%. Eastern Finnish provenances adapted well to western Finnish conditions.

The PDF includes an abstract in Finnish and French.

• Velling, E-mail: pv@mm.unknown
• Nepveu, E-mail: gn@mm.unknown

Variation of monoterpene composition of Pinus sylvestris L. was studied in Southern, Central and Northern Finland using data from both natural stands and plus trees. The natural stands were analysed using different techniques and for fewer terpenes than the plus trees.

There were large differences between areas in the proportion of 3-carene in trees from natural stands, as has been discussed by previous authors. The proportion of 3-carene is bimodally distributed and believed to be controlled by a single gene with large effect. For this reason, we stratified our samples into high carene (>10%) and low carene (<10%) groups. Univariate analysis did not reveal any additional differences between natural populations in different zones for components other than 3-carene. In plus trees, several components showed significant differences, but the proportion of 3-carene did not differ between areas. Multivariate discrimination analysis did not distinguish between areas for natural stands. However, for the plus trees discriminant analysis allowed us to discriminate between the zones relatively efficiently. The proportion of correct classification was greater than 64% using the best methods. The central zone was most distinct, and 80% of its trees were correctly classified. Broad generalizations are not possible due to the limitations imposed by our data. Our analysis of phenotypic variation does not support the suggestion that plus trees selected from the north represent a southern type.

The PDF includes an abstract in Finnish.

• Muona, E-mail: om@mm.unknown
• Hiltunen, E-mail: rh@mm.unknown
• Morén, E-mail: em@mm.unknown
• Shaw, E-mail: ds@mm.unknown

The publication comprises proceedings of a conference held in Helsinki in 1981. Forest tree populations are investigated for population genetic structure, mating systems, mechanisms of genetic adaptation and ecological adaptation. Methods and techniques used in population genetic research of forest trees are presented. Much concern is given to applications by means of forest tree breeding, particularly the seed orchard breeding technique. Generally, the application of population genetics in cultivated forests is discussed.

The PDF includes a preface and the presentations of the conference (25 short papers) in English, and a comprehensive summary of the themes of the conference in Finnish.

• Tigerstedt, E-mail: –

The productivity and several morphological features of Estonian Norway spruce (Picea abies L. Karst.) show significant geographical variation. This is no doubt because of differences in the climatic and soil conditions, which in spite of the small area of the country is quite large. In order to check the possible geographical variability of the gene pool, preliminary experiments were carried out in 1969, when seeds from 93 spruce stands originating from 14 forest enterprises were sowed in a nursery. After two years, the seedlings originating from south-eastern Estonia were the tallest. The seedlings from northern origins were smallest. However, it cannot be maintained that spruces from Southern Estonia are of better genotype than genotype from Northern Estonia since the genotypes are evaluated on the basis of ecological conditions under which the experiments are carried out. Another study suggests that an average shift of 7° to the east of the territory for spruces are suitable for cultivation in Estonia.

The PDF includes a summary in Finnish.

• Etverk, E-mail: ie@mm.unknown

Male and female flowering, cone crop, and some vegetative characteristics were studied in grafts 10 to 16 years of age in a clonal seed orchard of Scots pine (Pinus sylvestris L.). Genetic variation was found between clones in flowering as well as in cone production. Clone evaluation resulted in similar classifications of clones in different years. A regression analysis showed that crown size clearly increased but previous height growth slightly decreased flowering and cone production. The percentage of pollinated female strobili did not differ between clones.

The PDF includes a summary in English.

• Tormilainen, E-mail: mt@mm.unknown

In this study an attempt was made to use manometric Warburg technique in studying the growing season variations in the respiration rates of the roots of 1–3-year-old seedlings of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.). The respiration rates in both short-roots and long-roots have also been investigated.

According to the results, respiration intensity was the greatest in Scots pine and Norway spruce short-roots but also considerable in the long-root tips at the points of elongation. When the oxygen uptake rate per weight unit in the pine short-roots is given value of 100, the rate in the long-root tips is 61 and in the basal area 36. The corresponding values for spruce are 100, 69 and 43. The relative carbon dioxide release rates are different for the basal parts of the long-roots: pine 53 and spruce 57, when the CO₂ release from the short-roots is 100. The CO₂ release rate in the basal parts of the long-roots is relatively greater than the oxygen uptake. The respiration rate of the root systems of pine was larger than that of spruce due to the larger size of the root system.

The respiration rate per unit weight of pine roots of the 1- to 3-year-old seedlings decreases significantly with the increasing age. In spruce, the decrease was smaller. The result could have been different if only the short-roots of the same growing season were studied from all seedlings.

During the first growing season the root respiration rate decreased from the middle of the summer towards autumn. An experiment with pine seedlings grown in the mineral soil showed a very rapid increase in respiration rate in the spring. The rate, especially oxygen uptake, is at its greatest in the roots at the time of fastest growth.

• Lähde, E-mail: el@mm.unknown

Seasonal variation in the sawmill industry of Finland was studied in an investigation based on questionnaires answered by a random sample of sawmills concerning the time period of 1958-1960. The method is described in detail in a separate article in Acta Forestalia Fennica issue 75 no. 1.

The seasonal variations in purchase of roundwood was largest in big sawmills, which purchase the main part of the timber as standing sales and buy most of the wood from the State Forest auctions at the end of September. Also, they can afford to reserve their material earlier than the smaller companies. The saw logs are mainly felled in the winter in Finland because the climatic conditions and availability of labour are best at that time. Small sawmills begin fellings a little earlier than the larger ones.

In long-transport of timber the proportion of floating decreased from 47% in 1958 to 38% in 1960. At the same time, proportion of truck transport increased from 48% to 55%. Small sawmills use almost exclusively land transport. They received almost three-fourths of their logs between January and May, because the sawing is concentrated in the first half of the year. Therefore, floating does not suit for their transport method. The larger the sawmill, the later is the seasonal peak of log deliveries. The output of the big sawmills is distributed more evenly thoughout the year. The smaller the sawmill, the quicker is the turnover of raw material and the smaller the sawlog inventories.

The seasonal variation in output is sharper at small sawmills where sawing is concentrated in the first half of the year. The seasonal peak of the early spring is due to the aim at getting the sawn wood to dry early enough for shipments in the summer. Air drying takes an average of 4 ½ months. Kiln drying is more common at the larger sawmills, and gives them more flexibility. Due to the large seasonal variation in operation, the capacity of the small mills is poorly utilized. Domestic sales of sawn wood levels up the seasonality of the deliveries. Export sales are concentrated at the end and turn of the year. Also, the seasonal peak of expenditure occurs in the winter, but that of income in the summer.

The PDF includes a summary in English.

• Ervasti, E-mail: se@mm.unknown

The purpose of this paper is 1) to establish the possible seasonal variation of the different phases of work in the sawmill industry in Finland, 2) to study the internal and external factors influencing the seasonal character of the sawmill industry, 3) to study ha time lags between the work phases of the sawmill industry, 4) to analyse the seasonal nature of the industry’s money transactions, and 5) to give information concerning the factors influencing employment. The investigation is based on questionnaires of a random sample of sawmills concerning the time period of 1958-1960. This paper concentrates on the methods of the study, the results are reported in a separate paper in the Acta Forestalia Fennica issue 75 vol 1.

It was concluded that if seasonal variation in the sawmill industry is to be analysed on the scale it has been in the present work, sampling is the cheapest and most practicable method of collecting the material. If seasonal fluctuation of the industry is to be calculated by size classes, the sample must be allocated into strata by measuring the heterogeneity of the classes with a parameter illustrating seasonal variation. It might be useful to apply these parameters already when the total size of the sample is determined. For the smallest sawmills, for which practically all data have to be collected from primary documents, the most practical method is perhaps to send collectors to the spots. To the larger sawmills the questionnaire can perhaps be sent by mail. A moving index should be obtained for calculation of a seasonal index, but this demands long time series. The collection of the data is described in detail.

The PDF includes a summary in English.

• Ervasti, E-mail: se@mm.unknown

The article presents the results of the studies about the horizontal root systems of pine. The results have been obtained with the method developed by the author.

The size of the root system varies very strongly during the growing period. The amount of roots is at the smallest in the spring, increasing then rapidly and peaking at the end of July. After that the amount of roots decreases again against the winter to the same size than in the spring. The differences are due the changes in the amount of the smallest roots. There seem to be no big differences in the amount of roots between stand of different ages.

After the thinning there is a drop in the amount of roots on the stand level, but after two growing periods then standing trees have taken over the unused land. However after selection felling the less vital trees are not capable of utilizing the vacant resource as effectively.     

The PDF contains a summary in Finnish. 

• Kalela, E-mail: ek@mm.unknown

The purpose of the investigation was to examine the seasonal pattern in Finnish export shipments and export sales of sawn softwood in 1927-1953. Statistics concerning shipments have been obtained from the Board of Customs, and material relating to sales has been provided by the Finnish Sawmill Owner’s Association (now Finnish Sawmills Association). On the basis of original monthly statistics, 13-month moving averages were computed. Finally, a seasonal index was calculated.

According to the results, the export shipments have a fairly apparent seasonal pattern with very low figures from January to April, a peak from June to August, and thereafter a gradual decline up to the end of the year. There are also considerable variations from year to year but in general the exports follow this rhythm. In contrast to export shipments the seasonal pattern of export sales is characterised by significant irregularity. Market developments and speculation play a far greater role than the seasonal factors. Indeed, a seasonal character in export sales can scarcely be discerned.

The Acta Forestalia Fennica issue 61 was published in honour of professor Eino Saari’s 60th birthday.

The PDF includes a summary in English.

• Holopainen, E-mail: vh@mm.unknown

The relationship between canopy structure and photosynthetic radiation regime are studied in a theoretical basis. In modelling the canopy structure, a statistical approach is applied and the radiation field inside a stand is described in terms of random variables and their distribution. A comparison is made between horizontally homogenous stands and grouped forest stands in order to assess the influence of grouping of foliage on the irradiance distribution in a forest stand. Results show that grouping considerably reduces the interception of radiation and causes a large spatial variation. In coniferous stands the grouping of needles into shoots and the effect of penumbra are shown to have an important influence on the distribution of radiation on the needle area.

The PDF includes a summary in Finnish.

• Oker-Blom, E-mail: po@mm.unknown

The problems caused by the temporal and spatial microvariation in irradiance during field measurements of photosynthesis are studied. It is concluded on the basis of variation analyses based on irradiance data measured in a Scots pine (Pinus sylvestris L.) stand that the microvariation should be measured by integrating it over the measurement time and space.

However, the curvlinearity of the light response of photosynthesis results in biased estimates when linear integration (mean irradiance) is used. The significance of the bias is examined using a simulation technique on irradiance material. Whether the actual integral of photosynthesis can be approximated with mathematical method is next studied. The method gave satisfactory results only for a low curvature response, but the latter method was applicable also to the high curvature response. However, both methods presuppose that the mean and variance are known. Measurement of the variance is based on integration of the second power.

A new method, where the nonlinearity problem is avoided, is presented to measure fluctuation of the irradiance. The method enables the shoot geometry to be taken into account and it is also applicable to transpiration studies.

The PDF includes a summary in Finnish.

• Smolander, E-mail: hs@mm.unknown

Studies on Finnish Scots pine (Pinus sylvestris L.) plus tree clones by monoterpene and isozyme analyses was undertaken to further investigate mating system, population structure and pollination. Six allozyme systems (3 GOT, 1 GDH and 2 LAP) were properly analysed on the basis of segregation. Monoterpenes were analysed from needle material and segregation in high and low 3-carene content was found to depend on two alleles C and c. Thus, six allozyme systems and one monoterpene system were used as markers in this study.

It was shown that the northern clonal group maintains a much genetic variation as the central or southern clonal groups. The conditional probability of self-fertilization in about 20-year old clones estimated by the multilocus model was 14.1%, of which 8% originate from mating between trees that carry the same alleles to one of the maternal parent at some loci and 6% through self-fertilization.

There was no prominent difference in allele frequency of male gametes that pollinated the very early or very late flowering clones. The northern clonal group has higher a lower frequency of alleles GOT B2 and B3 respectively than of the southern clonal groups. The artificial plus tree selection, particularly in northern Finland, appears to favour heterozygous genotypes for the alleles that control 3-carene content n Scots pine.

The PDF includes a summary in Finnish.

• Chung, E-mail: mc@mm.unknown

Variation in the radial growth of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) was studied in 10 localities in Southern Finland. The aim of the study was to utilize the information on variation in the growth indices for growth estimation, and to evaluate the relative accuracy of growth estimation based on the data from increment boring, in comparison to other methods.

The climatic variation in periodic growth (5-year-period) in Southern Finland is about 11% of the normal level. The results suggest that the data from 10 localities can be used in the computation of the index series for Southern Finland and that the data from 10 relascope plots are required in the study of the climatic variation in tree growth in a given locality. The standard error of the estimate in actual growth estimation for past and future periods by the stand function method is about 23%, the errors of the method involving increment boring being 6% and 16% respectively in past and future growth estimation. The respective methods yield about 20% and 13% error in average growth estimation. With aid of the average growth indices for Southern Finland in local growth estimation the accuracy is distinctly improved, the error being equal to 12% only, while the accuracy in other cases is slightly improved.

The PDF includes a summary in Finnish.

• Thammincha, E-mail: st@mm.unknown

The study deals with the development during the 1950s and 1960s of a stand growing on peatlands which had been drained in the 1930s. The following characters were determined by measurements: the volume of the growing stock, the volume increment, the relative increment, the increment percent and the increment curves. Moreover, the possible changes taking place in the difference between tree growth along the ditches and in the middle of the strip between ditches were studied. In addition, the regional variation in increment was studied; this question was studied as the regression between the relative growth and the temperature sum. The results were compared with other Finnish investigations into the regional variation of increment.

The volumes of the growing stock had increased during the course of twelve years by 70–10 m³ /ha depending on the site type and climatic zone concerned. The relative increment had dropped in each case studied. As a matter of fact, this is only to be expected because the volumes had increased and the absolute growth had remained more or less unchanged. The development of the increment percent was compared with mineral soil stands in the case of Southern Finland, both uncut stands and stands treated with cuttings. According to the results obtained, the development of the increment percent was better in the present material than in uncut forests, but in some cases it did not reach the level of tended stands. The revival of the tree crop after draining takes place at different rates in the vicinity of and, on the other hand, at greater distances from the ditches and that this relationship is dependent on the fertility of the site.

The PDF includes a summary in English.

• Heikurainen, E-mail: lh@mm.unknown
• Seppälä, E-mail: ks@mm.unknown

The present study deals with the ground water table in the soil of drained peatlands and with the technique used for its determination. The terms depth and height of the ground water table are defined in the paper. Because of the fact that the surface of peatlands moves under the influence of a great number of different factors, the depth of the ground water table and the height of the ground water table are not parallel concepts. The present paper concentrates on the depth of the ground water table.

Observations on the depth of the ground water table in the sample plots in 1966-67 and 1968-69 show that the maximum of the late summer usually exceeds that of the early spring, and that the minimum occurring in the period of snow melting is more clearly discernible than that of the fall. Great differences occur in the depths of the ground water table in different sample plots. These differences are due to the specific properties of the peat of different peat layers, which are expressed in terms of the ground water coefficient. The duration of the depth of the ground water table proved to be a useful way to express the long-term changes.

Four kinds of short-term fluctuations in the ground water table were observed: a) the ground water table falls during the night hours, although the rate of falling is slower than in daytime, b) the ground water table rests at the same depth during the night, whereas during the day it clearly falls, c) the descent of the ground water table is similar throughout the whole 24-hour period, d) the ground water table rises during the night hours and falls in daytime. Occurrence of these types are discussed. Typical short-term fluctuation is the fall due to evaporation in the daytime.

The third part of the paper discusses the techniques used to measure the changes in ground water table.

The PDF includes a summary in English.

• Heikurainen, E-mail: lh@mm.unknown

After a decades-long increasing trend, the recent results of the National Forest Inventory (NFI) reported a decline of forest growth in North Finland. The aim of this study was to assess climatic and reproduction influences behind the growth decline. We used tree-ring data that had been collected by NFI using systematic sampling. The tree-ring width series were detrended using the regional curve standardisation (RCS) removing age-related trends. The resulting tree-ring indices of Scots pine (Pinus sylvestris L.) showed decadal variations with low increment in the 1990s, and high increment in the 1980s and the early years of the current century. Thereafter, a prolonged growth reduction for pine started both on the mineral soil sites and peatlands. The tree-ring indices of Norway spruce (Picea abies (L.) Karst.) had less pronounced decadal variations and no trend-like reduction over the last 15 years. High spring and summer temperatures were found to enhance radial growth, but high winter temperatures were related to low growth for pine and spruce in the following summer. Temperature variation, accompanied by variables indicating years of drought and intensive flowering, accounted for 34% annual growth variance of pine and 21–44% for spruce. Thus, the results imply that climatic factors may have to some extent contributed to the recent growth reduction of pine. Due to its ecological and economic consequences growth decline needs to be further monitored and investigated. Moreover, analyses of stand and age structure, potentially affecting the growth decline, were beyond the scope of this paper, but also warrant further investigation.

• Mäkinen, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, 00790 Helsinki https://orcid.org/0000-0002-1820-6264 E-mail: harri.makinen@luke.fi
• Nöjd, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, 00790 Helsinki E-mail: pekka.nojd@luke.fi
• Helama, Natural Resources Institute Finland (Luke), Ounasjoentie 6, 96200 Rovaniemi, Finland https://orcid.org/0000-0002-9777-3354 E-mail: samuli.helama@luke.fi

Forests in the western United States generally have increased in tree density since Euro-American settlement, particularly through increases in fire-sensitive species, such as spruces, firs, and junipers. Like most areas, the Black Hills region in western South Dakota and eastern Wyoming was logged for forest products and underwent agricultural conversion before historical forests were documented. To supplement historical reconstructions and accounts, we compared tree composition and densities (diameters ≥12.7 cm at 1.37 m above ground height) from historical General Land Office (GLO) records (years 1878 to 1915) and current Forest Inventory and Analysis (FIA) tree surveys (years 2011 to 2016) in the Black Hills Highlands of South Dakota. For composition, ponderosa pine (Pinus ponderosa P. Lawson & C. Lawson) decreased from 95% to 86% of all trees, with a consequent increase specifically of white spruce (Picea glauca (Moench) Voss) from 1.5% to 6.7% of all trees. Ponderosa pine currently is smaller in mean diameter by 7.4 cm, while white spruce is larger in mean diameter by 2.4 cm than historically. When the 35% of historical survey points without recorded trees were excluded, historical tree densities indicated an overall forested structure of savannas and open woodlands with tree densities ranging from 66 trees ha^–1 to 162 trees ha^–1. However, historical forests of the Black Hills incorporated dense stands. Tree densities have increased two- to more than four-fold, to 311 trees ha^–1 currently. These comparisons provide another source of information, paralleling changes documented in surface fire-dependent pine and oak forests throughout the United States, of transitions in forest composition and structure since Euro-American settlement.

• Tatina, Department of Biological Sciences, Dakota Wesleyan University, Mitchell, SD 57301 USA E-mail: rotatina@dwu.edu
• Hanberry, USDA Forest Service, Rocky Mountain Research Station, Rapid City, SD 57702 USA E-mail: brice.hanberry@usda.gov

We studied variations on different traits of Parkia timoriana (D.C.) Merr. in twelve provenances systematically from their source of origin to a controlled environment where representative seedlings were grown. Among the provenances, P1 gave the best result for seed traits including germination traits, P7 for pod traits and P10 for seedling vigour. Effects of seasonal distribution of rainfall and temperature on seed and pod traits were also determined by computing multiple regression analysis. The results displayed winter rainfall and summer temperature as the most important factor determining pod and seed traits. Latitude also significantly (P < 0.001) affected PWT (r = 0.52), SWP (r = 0.46) and SW (r = 0.50). A common garden study for germination and seedling growth indicated P1 and P10 provenance as the best among all. Seeds drawn from P10 gave the highest seedling vigour with an average growth rate of 0.61 cm/day from 90th to 180th day. Highest broad-sense heritability values (h²) were observed in germination traits, followed by seedling collar diameter. The lowest h² was observed for seedling height.

• Thangjam, Department of Forestry, School of Earth Sciences & Natural Resource Management, Mizoram University, Aizawl-796004, Mizoram, India https://orcid.org/0000-0003-1263-5348 E-mail: thangjam1987@gmail.com
• Sahoo, Department of Forestry, School of Earth Sciences & Natural Resource Management, Mizoram University, Aizawl-796004, Mizoram, India https://orcid.org/0000-0002-6524-1775 E-mail: uksahoo_2003@rediffmail.com
• Thong, Department of Forestry, School of Earth Sciences & Natural Resource Management, Mizoram University, Aizawl-796004, Mizoram, India https://orcid.org/0000-0002-5358-181X E-mail: pentilethong@gmail.com

Progenies from open pollinated cones collected in natural populations of Norway spruce (Picea abies (L.) Karst.) distributed along two altitudinal transects in Mid-Norway were tested in the nursery, in short term tests and in long-term field trials. The populations showed clinal variation related to the mean annual temperatures of the populations, with the earliest bud flush and cessation of shoot elongation and lowest height at age nine years for the high altitude populations. Within population variation was considerable as the narrow sense heritability for these traits was 0.67, 0.31 and 0.09 in one transect and 0.55, 0.18 and 0.14 in the other transect, respectively. Lammas shoots occurred in the short term trials with large variation in frequency between years. There was significant family variation for this trait, but also interactions between populations and year. The variance within populations was considerably larger in the populations from low altitude compared to the high-altitude populations. Significant genetic correlations between height and phenology traits and damage scores indicate that families flushing early and ceasing growth late were taller. Taller families also had higher frequencies of damages. Selection of the top 20% families for height growth in short term tests at age nine years gave a simulated gain of 11% increased height growth at age 18 years in long term trials at altitudes similar to those of origin of the populations. The gain was negative when high altitude populations were selected based on testing in the lowland.

• Skrøppa, Norwegian Institute of Bioeconomy Research, P.O. Box 115, 1431 Ås, Norway E-mail: tore.skroppa@nibio.no
• Steffenrem, E-mail: as@nn.no

Riparian trees, especially relict trees, are attractive and important for research to understand both past and recent biogeographical and evolutionary processes. Our work is the first study to elucidate the genetic diversity and spatial genetic structure of the canopy-dominating riparian Pterocarya fraxinifolia (Juglandaceae) along two altitudinal gradients in different river systems of the Hyrcanian forest, which is one of the most important refugium of relict trees in Western Eurasia. Altitudinal gradients were chosen along two river systems at 100, 400 and 900 m a.s.l. Leaf samples were collected from 116 trees, and the genetic diversity was evaluated with eight SSR markers. Overall, 39 alleles were identified for all of the populations studied. The observed heterozygosity (Ho) varied from 0.79 to 0.87 (with a mean of 0.83). The results of the AMOVA analysis indicated that the variation within populations was 88%, whereas the variation among populations was 12% for all of the gradients. A structure analysis indicated that 93% of the trees were grouped in the same gradient. The genetic distance based on Fst confirmed the structure result and indicated a high rate of gene flow among the investigated populations. Based on high gene flow (low differentiation of the population along the same river) and the clearly distinct genetic structure of the investigated gradients, it can be concluded that rivers are the main seed dispersal vector among P. fraxinifolia populations. The genetic diversity of P. fraxinifolia did not show any trend from upstream to downstream. The high level of gene flow and uniform genetic diversity along each river suggest the “classical” metapopulation structure of the species.

• Yousefzadeh, Department of Environmental Science, Faculty of Natural Resources and Marine Sciences, Tarbiat Modares University, Noor, Mazandaran, Iran E-mail: h.yousefzadeh@modares.ac.ir
• Rajaei, Department of Forestry, Faculty of Natural Resources and Marine Sciences, Tarbiat Modares University, Noor, Mazandaran, Iran E-mail: r.rajaei@modares.ac.ir
• Jasińska, Laboratory of Systematics and Geography, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, PL-62-035 Kornik, Poland E-mail: jasiak9@wp.pl
• Walas, Laboratory of Systematics and Geography, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, PL-62-035 Kornik, Poland E-mail: lukaswalas@wp.pl
• Fragnière, Department of Biology and Botanic Garden, University of Fribourg, Chemin du Musée 10, CH-1700 Fribourg, Switzerland E-mail: yann.fragniere@unifr.ch
• Kozlowski, Department of Biology and Botanic Garden, University of Fribourg, Chemin du Musée 10, CH-1700 Fribourg, Switzerland; Natural History Museum Fribourg, Chemin du Musée 6, CH-1700 Fribourg, Switzerland E-mail: gregor.kozlowski@unifr.ch

Hybrid aspen (Populus tremula × P. tremuloides) is one of the fastest growing tree species in Finland. During the mid-1990s, a breeding programme was started with the aim of selecting clones that were superior in producing pulpwood. Hybrid aspen can also be grown as a short-rotation crop for bioenergy. To study clonal variation in wood and bark properties, seven clones were selected from a 12-year-old field trial located in southern Finland. From each clone, five trees were harvested and samples were taken from stem wood, stem bark and branches to determine basic density, effective heating value, moisture and ash content. Vertical within-tree variation in moisture content and basic density was also studied. The differences between clones were significant for almost all studied properties. For all studied properties there was a significant difference between wood and bark. Wood had lower ash content (0.5% vs. 3.9%), basic density (378 kg m^–3 vs. 450 kg m^–3) and effective heating value (18.26 MJ kg^–1 vs. 19.24 MJ kg^–1), but higher moisture content (55% vs. 49%) than bark. The values for branches were intermediate. These results suggest that the properties of hybrid aspen important for energy use could be improved by clonal selection. However, selecting clones based on fast growth only may be challenging since it may lead to a decrease in hybrid aspen wood density.

• Hytönen, Natural Resources Institute Finland (Luke), Natural resources, Teknologiakatu 7, FI-67100 Kokkola, Finland E-mail: jyrki.hytonen@luke.fi
• Beuker, Natural Resources Institute Finland (Luke), Production systems, Vipusenkuja 6, FI-57200 Savonlinna, Finland E-mail: egbert.beuker@luke.fi
• Viherä-Aarnio, Natural Resources Institute Finland (Luke), Production systems, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: anneli.vihera-aarnio@luke.fi

Salix viminalis L. is an important shrub that has potential for use as a bioenergy crop, for phytoremediation of heavy metal contaminated soil and sewage sludge treatment. It is mainly distributed in the northeast of China, but the species has not yet been used a resource here. We examined the genetic diversity and population structure of populations from the Ergun basin and West Liao basin using 20 microsatellite markers. A high level of genetic diversity (N_a = 16.45, H_e = 0.742) was detected for S. viminalis, and populations from the Ergun basin exhibited higher genetic diversity and private alleles numbers than the West Liao basin. The 12 populations could be divided into two clusters by both Bayesian analysis and UPGMA clustering which were consistent with the populations derived from the two basins. Moderate population differentiation (F_ST = 0.076) was shown in S. viminalis, and AMOVA analysis confirmed that most of the genetic variation (86.13%) was attributed to individual differences within populations, while 11.49% was attributed to differences between basins and 2.38% to differences within each basin. Significant correlations of F_ST/(1–F_ST) with log (geographic distance) among 12 populations (r = 0.634, p < 0.00) and 10 populations within the Ergun basin (r = 0.482, p = 0.0002) indicated that geographical distance was the principal factor influencing genetic structure. As most of genetic variation exist within populations, so protection measures should be focused on populations with higher genetic diversity and unique alleles, such as Tuli, Mordaga downstream, Zhadun1 and Genhe.

• Zhai, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China; School of Architectural and Artistic Design, Henan Polytechnic University, Century Avenue, Jiaozuo, Henan, 454000, PR China E-mail: lkyzff@163.com
• Liu, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China E-mail: liu-jx295@163.com
• Li, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China E-mail: zhenjianli@163.com
• Mao, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China; Research Institute of Economic Forest, Xinjiang Academy of Forestry, Anjunanlu, Urumqi, Xinjiang, PR China E-mail: 350512173@qq.com
• Qian, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China E-mail: qianyq@caf.ac.cn
• Han, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China E-mail: hdd@caf.ac.cn
• Sun, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China E-mail: lkyszy@126.com

Polymorphisms at a set of eighteen nuclear (nSSR) and chloroplast (cpSSR) microsatellite loci were investigated in sixteen populations of Scots pine (Pinus sylvestris L.) derived from the provenance trial experiment and representative of the species distribution range and climatic zones in Poland. The patterns of genetic variation were compared to the reference samples from the species distribution in Europe and Asia. A similar level of genetic variation and no evidence of population structure was found among the Polish stands. They showed genetic similarity and homogenous patterns of allelic frequency spectra compared to the Northern European populations. Those populations were genetically divergent compared to the marginal populations from Turkey, Spain and Scotland. The population structure patterns reflect the phylogeography of the species and the divergence of populations that most likely do not share recent history. As the analysed provenance trial populations from Poland are diverged in phenotypic traits but are genetically similar, they could be used to test for selection at genomic regions that influence variation in quantitative traits.

• Hebda, University of Agriculture in Krakow, Faculty of Forestry, Institute of Forest Ecology and Silviculture, Department of Genetics and Forest Tree Breeding, 29 Listopada 46, 31-425 Kraków, Poland http://orcid.org/0000-0002-3149-8644 E-mail: ana.hebda@gmail.com
• Wójkiewicz, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland E-mail: bwojkiew@man.poznan.pl
• Wachowiak, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland; Institute of Environmental Biology, Faculty of Biology, Adam Mickiewicz University, Umultowska 89, 61-614 Poznań, Poland E-mail: witoldw@man.poznan.pl

• Jansons,  LSFRI “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 E-mail: aris.jansons@silava.lv
• Matisons,  LSFRI “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 E-mail: robism@inbox.lv
• Zadiņa,  LSFRI “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 E-mail: mara.zadina@silava.lv
• Sisenis, LUA Forestry Faculty, Akadēmijas str. 11, Jelgava, Latvia E-mail: linards.sisenis@llu.lv
• Jansons, Forest Competence Centre, Dzērbenes str. 27, Riga, Latvia, LV1006 E-mail: janis.jansons@silava.lv

• Miranda, Centro de Estudos Florestais, Instituto Superior de Agronomia, Universidade Lisboa, 1349-017 Lisboa, Portugal E-mail: Imiranda@isa.ulisboa.pt
• Gominho, Centro de Estudos Florestais, Instituto Superior de Agronomia, Universidade Lisboa, 1349-017 Lisboa, Portugal E-mail: Jgominho@isa.utl.pt
• Pereira, Centro de Estudos Florestais, Instituto Superior de Agronomia, Universidade Lisboa, 1349-017 Lisboa, Portugal E-mail: Hpereira@isa.utl.pt

• Turtiainen, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: marjut.turtiainen@uef.fi
• Salo, Finnish Forest Research Institute, Joensuu Research Unit, Joensuu, Finland E-mail: ks@nn.fi
• Saastamoinen, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: os@nn.fi

• Persson, Forestry Research Institute of Sweden, Sävar, Sweden E-mail: torgny.persson@skogforsk.se
• Andersson, Forestry Research Institute of Sweden, Sävar, Sweden E-mail: ba@nn.se
• Ericsson, Forestry Research Institute of Sweden, Sävar, Sweden E-mail: te@nn.se

• Laiho, University of Helsinki, Department of Forest Ecology, FI-00014 University of Helsinki, Finland E-mail: raija.laiho@helsinki.fi
• Sarkkola, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: ss@nn.fi
• Kaunisto, Finnish Forest Research Institute, Parkano Research Unit, Kaironiementie 54, FI-39700 Parkano, Finland E-mail: sk@nn.fi
• Laine, Finnish Forest Research Institute, Parkano Research Unit, Kaironiementie 54, FI-39700 Parkano, Finland E-mail: jl@nn.fi
• Minkkinen, University of Helsinki, Department of Forest Ecology, FI-00014 University of Helsinki, Finland E-mail: km@nn.fi

• Jyske, The Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: tuula.jyske@metla.fi
• Mäkinen, The Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: hm@nn.fi
• Saranpää, The Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: ps@nn.fi

• Steinke, Plant Ecology and Systematics, Lund University, Ecology Department, 223 62 Lund, Sweden E-mail: lks@nn.se
• Premoli, Universidad Nacional del Comahue, Laboratorio Ecotono – CRUB, Quintral 1250, 8400 Bariloche, Argentina E-mail: apremoli@crub.uncoma.edu.ar
• Souto, Universidad Nacional del Comahue, Laboratorio Ecotono – CRUB, Quintral 1250, 8400 Bariloche, Argentina E-mail: cps@nn.ar
• Hedrén, Plant Ecology and Systematics, Lund University, Ecology Department, 223 62 Lund, Sweden E-mail: mh@nn.se

• Kosinska, Department of Human Medical Genetics, Medical University of Warsaw, Oczki 1, 02-007 Warsaw, Poland E-mail: jk@nn.pl
• Lewandowski, Polish Academy of Sciences, Institute of Dendrology, 62-035 Kórnik, Poland E-mail: al@nn.pl
• Chalupka, Polish Academy of Sciences, Institute of Dendrology, 62-035 Kórnik, Poland E-mail: wc@nn.pl

• Zhang, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O. Box 416, Chengdu 610041, P. R. China; Graduate School of the Chinese Academy of Sciences, Beijing 100039, P. R. China E-mail: xz@nn.cn
• Korpelainen, Department of Applied Biology, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: hk@nn.fi
• Li, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O. Box 416, Chengdu 610041, P. R. China E-mail: licy@cib.ac.cn

• Pukkala, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: timo.pukkala@joensuu.fi
• Miina, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jm@nn.fi

• Laiho, Univ. of Helsinki, Dept. of Forest Ecology, Peatland Ecology Group, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: raija.laiho@helsinki.fi
• Penttilä, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: tp@nn.fi
• Laine, Univ. of Helsinki, Dept. of Forest Ecology, Peatland Ecology Group, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: jl@nn.fi

• Rouvinen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: seppo.rouvinen@forest.joensuu.fi
• Kuuluvainen, University of Helsinki, Department of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: tk@nn.fi
• Siitonen, Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: juha.siitonen@metla.fi

• Viherä-Aarnio, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box. 18, FIN-01301 Vantaa, Finland E-mail: anneli.vihera-aarnio@metla.fi
• Velling, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box. 18, FIN-01301 Vantaa, Finland E-mail: pv@nn.fi

• Leskinen, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: pekka.leskinen@metla.fi
• Kangas, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: jk@nn.fi

• Alía, CIFOR-INIA, Unidad de Mejora Forestal, 28080 Madrid, Spain E-mail: ra@nn.es
• Moro-Serrano, CIFOR-INIA, Unidad de Mejora Forestal, 28080 Madrid, Spain E-mail: jmoro@inia.es
• Notivol, Unidad de Recursos Forestales, SIA-DGA, Ca de Montañana 179, 50080 Zaragoza, Spain E-mail: en@nn.es

• Eriksson, Swedish University of Agricultural Sciences, Department of Forest Genetics, Uppsala, Sweden E-mail: gosta.eriksson@sgen.slu.se

The objective of this study was to investigate basic density and its within-stem variation by studying 84 European aspen stems from 28 forest stands in Latvia. The studied forest stands covered all age classes from young stands to matured forests in representative growth conditions of European aspen. The densities of 2722 wood and 1022 bark specimens were measured from the sampled trees. Only the knot-free wood specimens without obvious wood defects were chosen for analyses. A map of basic density summarizing its radial and axial variations was constructed to show species-specific, within-stem variability and the relationships between density and tree and stand variables were examined. Stem wood and bark of the European aspen show different patterns of basic density variation along the tree stem. Wood density increases from pith to bark up to certain dimensions and shows a slight decrease afterwards. The weighted basic density of bark (446 ± 39.6 kg m^–3) was higher than stem wood density (393 ± 30.4 kg m^–3). Our results suggest that wood and bark density measurements obtained at breast height can be used for reliable estimation of the densities of whole-tree stem components, while tree parameters such as diameter at breast height (DBH), tree height and social status or stand parameters, including number of trees, basal area and age, are weak predictors in this context.

• Liepiņš, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: janis.liepins@silava.lv
• Ivanovs, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: janis.ivanovs@silava.lv
• Lazdiņš, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: andis.lazdins@silava.lv
• Jansons, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: jurgis.jansons@silava.lv
• Liepiņš, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: kaspars.liepins@silava.lv

• Boratynski, Polish Academy of Sciences, Institute of Dendrology, 5 Parkowa str., 62-035 Kórnik, Poland E-mail: borata@man.poznan.pl
• Marcysiak, Kazimierz Wielki University, Institute of Biology and Environment Protection, 12 Ossolinskich str., 85-064 Bydgoszcz, Poland E-mail: km@nn.pl
• Lewandowska, Kazimierz Wielki University, Institute of Biology and Environment Protection, 12 Ossolinskich str., 85-064 Bydgoszcz, Poland E-mail: al@nn.pl
• Jasinska, Polish Academy of Sciences, Institute of Dendrology, 5 Parkowa str., 62-035 Kórnik, Poland E-mail: aj@nn.pl
• Iszkulo, Polish Academy of Sciences, Institute of Dendrology, 5 Parkowa str., 62-035 Kórnik, Poland E-mail: gi@nn.pl
• Burczyk, Kazimierz Wielki University, Institute of Biology and Environment Protection, 12 Ossolinskich str., 85-064 Bydgoszcz, Poland E-mail: jb@nn.pl