Current issue: 58(5)
In soil profiles from a series of uplands of different site types bulk density, density of solids and porosity of soil were clearly related to soil organic matter content and its distribution in the soil profile. Soil organic matter contents were also strongly correlated to effective cation exchange capacity (CEC) and soil acidity. Site fertility was primarily related to the fine fraction (ø<0.06 mm) content in the C horizon and related properties (i.e. CEC). In the humus layer, the content of exchangeable bases and base saturation most strongly related to site fertility.
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A close relationship between photosynthetic capacity and nitrogen concentration of leaves is known to exist. In conifers, nitrogen also affects the pattern of mutual shading within a shoot, which is a basic unit used in studying photosynthesis of coniferous trees. These effects of needle nitrogen concentration on photosynthetic capacity and mutual shading of needles were analysed for Scots pine (Pinus sylvestris L.) shoots taken from five young stands growing on sites of different fertility. The effect of nitrogen concentration on needle photosynthesis was studied based on measurements of the photosynthetic radiation response of shoots from which two thirds of the needles were removed in order to eliminate the effect of within shading.
An increase of one percentage unit in nitrogen concentration of needles increased the photosynthetic capacity of needles by 25 mg CO2 dm-2h-1. The effect of nitrogen on within-shoot shading was quantified in terms of the silhouette area to total needle area ratio of a shoot (STAR), which determines the relative interception rate per unit of needle area on the shoot. Although nitrogen promoted needle growth, an increase in nitrogen concentration decreased the within-shoot shading. This effect resulted from a decrease in needle density on the shoot and an increased needle angle with increasing nitrogen content.
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A field data set representing boreal forest-floor vegetation in Southern Finland was analysed using a simultaneous equation model. Some physical and chemical characteristics of the soil and some structural characteristics of the tree stand were treated as predictors in such a way that the tree stand factor was specified to be dependent on the soil variables. Alpha diversity, measured as the total number of species per plot, was treated as a criterion variable.
The model explains 60% of variance in the alpha diversity indicating markedly strong relationships with the site characteristics. Alpha diversity appears to increase with increases in site fertility characteristics. On the other hand, measured characteristics of the tree stand indicate no significant independent effects on the alpha diversity.
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Vegetation data collected from a random sample of mature forest stands representing mesic upland forest sites in Southern Finland were analysed and classified using two-way indicator species analysis (TWINSPAN). The variation of some characteristics of the tree stand and soil fertility between the produced vegetation units were analysed statistically.
Both the species list and the sample list arranged by TWINSPAN reflects the overall site fertility considerably well. The results are in agreement with the main division of mesic forest sites in the Finnish forest site type classification: vegetation units which can be assigned to the Oxalis-Myrtillus site type are clearly separated from the remaining units, and the overall site fertility indicates a statistically significant difference. The within-type variation in the vegetation composition in the Oxalis-Myrtillus site type reflects the variation in site fertility, whereas the within-type variation in the Myrtillus site type is mainly caused by the tree stand factor.
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The objective of the study was to investigate the factors affecting the deterioration of urban forests based on literature. Regression analysis is applied to the material obtained from unpublished material of the latter author. The size of forest stand, and its fertility proved to exercise the greater effect on the deterioration of urban forests. A model based on these factors has been developed and its area of applying is discussed.
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The paper deals with the nutrient status of surface peat layer from virgin sedge-pine swamps and its relationship to peatland types. When the nutrients are expressed in mg/100 g peat, only easily extractable Ca and Mg correspond to the productivity status of the peatland type. N, P, and K levels in the herb rich sedge-pine swamp are generally lower than in the small sedge-pine swamps, which are the least productive ones. The differences between the site types in all the five nutrients become much clearer when the results are expressed in kg/ha. P, K, and Ca are significantly different between the site types, and correspond to the productivity of the site type. For N and Mg the same tendency can be seen. The organically bound nutrients N, and to a lesser extent, P appear to comply with the hypothesis of an increase in nutrient availability in Southern Finland.
The PDF includes a summary in Finnish.
The ash content has been found to correlate with the fertility of peatlands. Relationship between height of 80-year-old stands and ash content of peat in topmost 30 cm layer was examined in Lithuanian conditions. On drained peatlands with ash content of peat from 3% to 8% pine stands increase in height. Ash content of peat being about 7% Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) stands on drained sites are found to be of equal height. Ash content of peat more than 8–9% has no significant effect on growth of pine or spruce stands. Birch (Betula verrucosa (B. Pendula Roth.) and Betula pubescens Erhrh.), stands are less sensitive to ash content of peat compared with other species. Black alder (Alnus glutinosa L. Gaertn.) stands occurred in sites with ash content of peat more than 8–10%. The height of the stands become equal both in drained and undrained sites in the cases where ash content of peat is about 16–18%. Ash (Fraxinus exelsior L.) stands attain high productivity on drained sites with ash content of peat about 20%.
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Growth capacity of peatlands after draining depends largely on quality of the surface peat. The future growth capacity can be determined with considerable accuracy by the vegetation of the peatland. The aim of this study was to draw guidelines to identify the fertility of a peatland and its potential for draining based on its vegetation.
Certain species and plant associations were identified to describe the fertility of different peatland types. Brown mosses indicate an abundant nutrient content of the site, certain herbs moderately abundant nutrient content, sedges (Carex sp.) moderate nutrient content, shrubs poor nutrient content and vegetation indicating an oligotrophic peatland indicates excessively poor nutrient content. The article includes detailed descriptions of the vegetation of different peatland types.
Supplementary features, such as thin peat, flooded parts or abundance of Spangnum fuscum can be used as additional indications to determine the drainability of the site. The article describes an identification tool to determine the drainability of a peatland based on vegetation and the supplementary features of the peatland.
The article includes a summary in English.
Typical for the area of Längälmävesi, in Satakunta in Southern Finland, are densely populated fertile lowland areas near the waterways and poorer sparcely populated upland areas. The changes in fertility of the land influence the vegetation, and has directed where the population has settled. A vegetation survey was made using specific plant species as indication to fertility. A detailed description of the species composition and distribution of the plant species indicative for fertile land is presented in the article.
Analyzing the fertility only based on the plant species indicative for rich soils underestimates the proportion of fertile lands. Farmlands, on the other hand, have been cleared also on less fertile soils, which would give an overestimate of the fertile lands. A map of fertile lands was drawn based on both vegetation and location of the fields. The changes in fertility influenced also bird species observed in the area.
The PDF includes a summary in German.
The article is a review on methods used in Central Europe and in Finland to construct growth and yield tables, and on their defects and advantages. One of the main defects of growth and yield tables prepared previously in Central Europe is that the site quality classes for different tree species have been formed independently and using different principles. Thus, the yields of different tree species on a similar site can’t be compared. In addition, the quality classes of this kind of growth and yield tables and growth series are artificial.
The Society of Forestry in Finland (now Finnish Society of Forest Sciences) started to prepare new yield tables for the most important tree species in Finland in 1916. The new yield tables aimed to improve the yield tables made in Central Europe in two aspects. Firstly, the quality of the site of each sample plot was assessed on the spot and independently from the standing crop. Thus the sample plots of each site class in setting up the growth series can be treated as independent groups. Consequently, the quality classes are be the same for all tree species. Secondly, mathematical-statistical methods were used to determine which of the stands that belong to the same quality class belong also to the same growth series.
The PDF includes a summary in Finnish.
Result of a survey of soils supporting forest plantations in Wisconsin in the United States indicated a close correlation between the levels of fertility of non-phreatic, coarse-textured soils and the growth of red pine (Pinus resinosa Roezl) stands aged from 15 to 32. This relationship, however was not observed in plantations established on deep-gley soils, underlain at a depth of 3–9 fl by ground water.
The survey encountered 20 red pine plantations on soils underlain by a deep ground water table accessible to tree roots thorough their contact with gley horizon or with extended capillary fringe. The average growth of the stands was 80 cubic feet/acre (5.6 m3/ha) at the age of 22 years. Thus, mensuration analysis suggested that the soils are the choice grounds for forestry enterprise. However, the analysis of soil samples showed that in many instances the soils are extremely low in mineral colloids, organic matter and nutrients. Many of the sites would be regarded as critically deficient in nitrogen, phosphorus and potassium.
The following hypothesis are suggested to explain this discrepancy:
a) The moisture content of coarse-textured non-phreatic soils remain near the wilting point during a large apart of the growing season with subsequent reduction of transpiration and uptake of nutrients. If a capillary fringe provides a supply of water for the root system, trees may derive an adequate supply of salts and exchangeable ions from comparatively infertile substrata.
b) The suitably located ground water provides adequate aeration of the surface soil layers which is not impeded by capillary fringe, increasing activity of mycorrhiza, and a mycotrophic uptake of nutrients from unweathered minerals.
c) The above effects of natural subirrigation should change the concept of soil fertility based on mere chemical analysis. The time during which the roots are engaged in active absorption appears to be of equal importance as the concentration of nutrients in available form.
The PDF includes a summary in Finnish.
According to the theory of peatland types, particular peatland types, after sufficient drainage, change into certain forest types. It has been found, that the range of forest types in peatlands in different stages of draining is as large as on mineral soil; and comprise Cladina, Calluna, Vaccinium, Myrtillus Oxalis-myrtillus and grove types. Poor peatland types change into poor forest types, better for better types. However, a Swedish scientist Mellin suggested that after effective drainage especially oligotrophic bogs, when well drained, change usually into Myrtillus type.
The different conclusions are due to the fact that the same bog type may develop into different forest types according to the effectiveness and duration of the drainage. Greater the decay of the peat bog layer, the more exacting is the type of vegetation which appears. Bog types of classes V, IV and III (Finnish classification of site quality) change into a Myrtillus type, as do the poorer peatland types of class II. The types vary, however, in their economical drainage value. The fact that bogs which in their natural stage are clearly different in their site quality change after through drainage into the same forest type, is explained by the chemical quality of the peat. However, class I and the best types of class II bogs change into better forest types because they as eutropchic bogs are richer in nitrogen and lime. This difference persists despite of effective draining.
It has been shown that the development of eutrophic peatland types at the forest type stage also differs clearly from the development of oligotrophic peatland types. The Finnish classification of drainage value shows correctly the relative drainability when using normal spacing of ditches. The notes on forest types on mineral soil should, however, be replaced by corresponding notes on the transitive types between bog and forest types.
The PDF includes a summary in Swedish and English.
The relation between the occurrence of forest site types and swamp types was studied using data collected in the national line survey of Finnish forests carried out in 1921‒1924. The majority of peatlands in Finland has been formed by forest land becoming paludificated. When the peat layer is thin, the fertility of the peatland depends on the underlying mineral soil, consequently, good swamp types occur on fertile subsoil and poor types on poor subsoil. When the peat layer becomes thicker, the relationship weakens. The surrounding mineral soils influence the quality of the peatland by the runoff from the catchment area. The direct comparison of forest site and swamp types is not possible, because for one forest site type there are several swamp types that have different levels of humidity. According to the study, a very distinct mutual relation can be seen between occurrence of forest site types and swamp types.
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The article presents a survey on distribution of fertile lands, soil types and site classes in Savo and Karelia in the central and eastern parts of Finland. The survey was based both in existing publications and statistics, a line survey, and visual observations during field trips. The site quality classification is based on the vegetation and occurrence of indicator plant species. The article lists distribution of indicator species in different forest site types on maps of the area. In addition, a review of history of land use and agriculture give indications of the location of the fertile lands in the area. A map of the forest site types in different parts of the area illustrate the data collected from the different sources.
The PDF includes a summary in German.
The vegetation and number of physical and chemical soil properties were studied on a random sample of closed upland forest stands in Southern Finland. The material consists of a total of 410 sample plots. Two-way indicator species analysis (TWINSPAN) was carried out in order to produce a hierarchical clustering of samples on the basis of the vegetation data. Discriminant analysis and analysis of variance were applied in order to find environmental correlations of the vegetation clustering.
The vegetation was found to indicate the nutrient regime of the humus layer well, but to a less extent the properties of the sub-soil. The understorey vegetation was found to be jointly dependent on the site fertility and on the properties of the tree stand, especially on the tree species composition. Although the forest vegetation appears to be distributed rather continuously along an axis of increasing site fertility, relatively unambiguous classification can be based on the appearance of indicator species and species groups.
The results of the study were interpreted as indication that operational site classification done using the vegetation is rather good method for classification in closed forest stands. Different methods produce relatively consistent, natural and ecologically interpretable classifications. The results also imply that the vegetation cover and the humus layer develop concurrently during the development of the ecosystem, but the differentiation of the site type is regulated simultaneously by a number of interacting factors ranging from mineralogical properties of the parent material to the topographical exposition of the site. As the plant cover depicts all these primary factors simultaneously, only a relatively rough ecological site classification can be based on the vegetation.
The PDF includes a summary in Finnish.
The aims of the present study were to determine physical and physio-chemical properties of some Finnish forest tree nursery soils, and to examine relationships between these properties and the amount of organic matter in the soil.
The following soil tillage layer properties of 33 fields belonging to 8 forest tree nurseries were determined: soil particle size distribution, organic matter content, bulk density and density of solids, total pore space, soil water volume at potentials pF 2.0 and 4.2, available water content and air space at potential pF 2.0, active acidity, electrical conductivity index and cation exchange capacities at pH 4.5 and 8.0. The soil texture class of the tillage layer parent material was sand, only in a few cases did higher percentage of silt and clay indicate a morainic nature of parent material. The amount of organic material in the soils varied within wide limits, reflecting differences in amelioration policy between the single nurseries.
Relationships between the physical properties of the soil parent material and those related to fertility were in most cases strongly influenced by the amount of soil organic matter. Soil density values decreased as the organic matter content increased from 2 to 25%, giving rise to the increase in the total pore space. However, the amount of water held at potential pF 2.0 and the available water content did not increase with increasing organic matter content. This was due to the absence of the particle fraction in the sand. Nursery soil amelioration, involving in most cases a mixture of Sphagnum peat with sand, thus gives rise to an increase in the content of drainable water.
Cation exchange capacities were positively correlated with the organic matter content. However, the absolute number of exchange sites expressed as equivalents in the tillage layer did not increase in accordance with the increase in organic matter content due to the influence of the organic matter content upon the ratio of solids in the voids.
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Fertility of surface peat from sedge pine mires was studied by measuring several edaphic growth factors: bulk density, volume weight of organic matter, ash content, acidity, electric conductivity, effective cation exchange capacity, degree of base saturation, and total contents of N, P, K, Ca and Mg. The 168 temporal sample plots were situated on virgin sedge mires in different parts of Finland, and the 30 permanent sample plots on two uniform sedge mires.
The results showed that peat bulk density and volume weight of organic matter tend to increase with increasing site quality. Ash content increased gradually in the site series from small sedge mire to the herb-rich sedge mire. The relationship between the total content of macronutrients in peat and the site quality is clear. The importance of bulk density in evaluating the site quality is further emphasized when taking into account its significant correlation to contents of N and P. The soil variables follow the accepted quality gradient of the site series. Consequently, the plant sociologically based site classification seems to reflect satisfactorily the average soil properties. However, the within site variation was significant.