Articles containing the keyword 'model'

A process-oriented tree and stand growth model is extended to be applicable to the analysis of timber quality, and how it is influenced by silvicultural treatments. The tree-level model is based on the carbon balance and it incorporates the dynamics of five biomass variables as well as tree height, crown base, and breast height diameter. Allocation of carbon is based on the conservation of structural relationships, in particular, the pipe model. The pipe-model relationships are extended to the whorl level, but in order to avoid a 3-dimensional model of entire crown structure, the branch module is largely stochastic and aggregated. In model construction, a top-down hierarchy is used where at each step down, the upper level sets constraints for the lower level. Some advantages of this approach are model consistency and efficiency of calculations, but probably at the cost of reduced flexibility. The detailed structure related with the branching module is preliminary and will be improved when more data becomes available. Model parameters are identified for Scots pine (Pinus sylvestris L.) in Southern Finland, and example simulations are carried out to compare the development of quality characteristics in different stocking densities.

• Mäkelä, E-mail: am@mm.unknown
• Ikonen, E-mail: vi@mm.unknown
• Vanninen, E-mail: pv@mm.unknown

In the last decades, architectural analysis has been used to understand and to model plant development. These studies have led us to reconsider the problem of measuring plants while taking into account their topological structure at several scales of detail. A computational platform, called AMAPmod, was created to work on such plant representations. This paper outlines the general methodology used in AMAPmod to represent plant topological structures and to explore these special types of databases. Plant structures are first encoded in order to build corresponding formal representations. Then, a dedicated language, AML, enables the user to extract various types of information from the plant databases and provides appropriate analysing tools.

• Godin, E-mail: cg@mm.unknown
• Caraglio, E-mail: yc@mm.unknown
• Costes, E-mail: ec@mm.unknown

A shoot-root carbon:nitrogen allocation model, based on the two processes of transport and chemical conversion, is described and explored. The view is proposed that all allocation models, whether built for the purposes of theoretical investigation or practical application, should start with this irreducible framework. In the present implementation, the processes operate according to: for substrate sources, dependence on shoot and root sizes, with possible product inhibition; for transport, movement down a substrate concentration gradient; for substrate sinks or utilization, linear bisubstrate kinetics. The dynamic and equilibrium properties of the model are explored. Failure of this approach to allocation will indicate to the modeller that additional mechanisms to control the processes are needed, and the mode of failure will indicate the type of mechanisms required. Additional mechanisms are discussed which may involve hormones or teleonomic (goal-seeking) controls, and may be added to the irreducible framework. However, these additions should not replace the irreducible framework of transport and chemical conversion, because they do not in reality. Modifications to the basic model to reflect some possibilities such as ontogenesis with the transition from exponential growth towards a steady state or with the scaling of within-plant transport resistances, the influence of hormones, and active transport, are described.

• Thornley, E-mail: jt@mm.unknown

The effects of two alternative formulations of sapwood senescence on the behaviour of model LIGNUM (with parameter values adjusted for Scots pine (Pinus sylvestris L.) growing southern Finland) were studied. The two alternatives were autonomous sapwood senescence assuming a maximum age for the tree ring, and sapwood senescence that is controlled by the mortality of foliage. For the latter alternative two hypothetical further mechanisms were stipulated. All the formulations were implemented in LIGNUM. Simulations were made with all model variants for fertile and poor soil conditions using high, normal and low rates of foliage mortality. The simulation results were compared against of a data set consisting of 11 open grown Scots pine trees from southern Finland. Observations of heartwood proportion were used in this study. They show that heartwood starts to increase in trees from age of approximately 20 years onwards. The simulation results showed no differences between fertile and poor soil conditions as regards heartwood formation. Of the variants of foliage-controlled sapwood senescence the one where death of sapwood in a tree segment induces sapwood senescence in the tree parts below only slightly was the best. This and the autonomous sapwood senescence corresponded equally well to the observations. In order to make more refined conclusions additional data and simulations are necessary.

• Sievänen, E-mail: rs@mm.unknown
• Nikinmaa, E-mail: en@mm.unknown
• Perttunen, E-mail: jp@mm.unknown

A new approach for modelling plant growth using the software AMAPpara is presented. This software takes into consideration knowledge about plant architecture which has been accumulated at the Plant Modelling Unit of CIRAD for several years, and introduces physiological concepts in order to simulate the dynamic functioning of trees. The plant is considered as a serial connection of vegetative organs which conduct water from the roots to the leaves. Another simple description of the plant as a network of parallel pipes is also presented which allows an analytical formulation of growth to be written. This recurring formula is used for very simple architectures and is useful to understand the role of each organ in water transport and assimilate production. Growth simulations are presented which show the influence of modifications in architecture on plant development.

• de Reffye, E-mail: pd@mm.unknown
• Barthélémy, E-mail: db@mm.unknown
• Blaise, E-mail: fb@mm.unknown
• Fourcaud, E-mail: tf@mm.unknown
• Houllier, E-mail: fh@mm.unknown

The model HYDRA, which simulates water flow in the branched tree architecture, is characterized. Empirical studies of the last decades give strong evidence for a close structure-function linkage in the case of tree water flow. Like stomatal regulation, spatial patterns of leaf specific conductivity can be regarded as a strategy counteracting conductivity losses, which may arise under drought. Branching-oriented water flow simulation may help to understand how damaging and compensating mechanisms interact within the hydraulic network of trees. Furthermore, a coupling of hydraulic to morphological modelling is a prerequisite if water flow shall be linked to other processes. Basic assumptions of the tree water flow model HYDRA are mass conservation, Darcy's law and the spatial homogeneity of capacitance and axial conductivity. Soil water potential is given as a one-sided border condition. Water flow is driven by transpiration. For unbranched regions these principles are condensed to a nonlinear diffusion equation, which serves as a continuous reference for the discrete method tailored to the specific features of the hydraulic network. The mathematical derivation and model tests indicate that the realization of the basic assumptions is reproducible and sufficiently exact. Moreover, structure and function are coupled in a flexible and computationally efficient manner. Thus, HYDRA may serve as a tool for the comparative study of different tree architectures in terms of hydraulic function.

• Früh, E-mail: tf@mm.unknown

A metabolic model of height growth and site index is derived from a parametrization of the annual carbon balance of a tree. The parametrization is based on pipe-model theory. Four principal variants of the height-growth model correspond to four combinations of assumptions regarding carbon allocation: (a) the apical shoot is autonomous or (b) it is not; and (A) the specific rate of elongation of a shoot equals that of a woody root or (B) it does not. The bB model is the most general as it includes the aA, bA, and aB models as special cases. If the physiological parameters are constant, then the aA model reduces to the form of the Mitscherlich model and the bA model to the form of a Bertalanffy model. Responses of height growth to year-to-year variation in atmospheric conditions are rendered through adjustments of a subset of the model's parameters, namely, the specific rate of production of carbon substrate and three specific rates of maintenance respiration. As an example, the effect of the increasing atmospheric concentration of CO₂ on the time-course of tree height of loblolly pine (Pinus taeda) is projected over 50-year span from 1986. Site index is predicted to increase and, more importantly, the shape of the site-index curve is predicted to change.

• Valentine, E-mail: hv@mm.unknown

Process-based tree growth models are recognized to be flexible tools which are valuable for investigating tree growth in relation to changing environment or silvicultural treatments. In the context of forestry, we address two key modelling problems: allocation of growth which determines total wood production, and distribution of wood along the stem which determines stem form and wood quality. Growth allocation and distribution are the outcome of carbon translocation, which may be described by the Munch theory. We propose a simpler gradient process to describe the carbon distribution in the phloem of conifers. This model is a reformulation of a carbon diffusion-like process proposed by Thornley in 1972. By taking into account the continuity of the cambium along the stem, we obtain a one-dimensional reaction-diffusion model which describes both growth allocation between foliage, stem and roots, and growth distribution along the stem. Distribution of wood along the stem is then regarded as an allocation process at a smaller scale. A preliminary sensitivity analysis is presented. The model predicts a strong relationship between morphology and foliage-root allocation. It also suggests how empirical data, such as stem analysis, could be used to calibrate and validate allocation rules in process-based growth models.

• Deleuze, E-mail: cd@mm.unknown
• Houllier, E-mail: fh@mm.unknown

Within the European Community snow damage affects an estimated 4 million m³ of timber every year, causing significant economic losses to forest owners. In Northern Europe, for example, the occurrence of snow damage has increased over the last few decades mainly due to the increase in total growing stock. The most common form of damage is stem breakage, but trees can also be bent or uprooted. Trees suffering snow damage are also more prone to consequential damage through insect or fungal attacks.

Snow accumulation on trees is strongly dependent upon weather and climatological conditions. Temperature influences the moisture content of snow and therefore the degree to which it can accumulate on branches. Wind can cause snow to be shed, but can also lead to large accumulations of wet snow, rime or freezing rain. Wet snow is most likely in late autumn or early spring. Geographic location and topography influence the occurrence of damaging forms of snow, and coastal locations and moderate to high elevations experience large accumulations. Slope plays a less important role and the evidence on the role of aspect is contradictory. The occurrence of damaging events can vary from every winter to once every 10 years or so depending upon regional climatology. In the future, assuming global warming in northern latitudes, the risk of snow damage could increase, because the relative occurrence of snowfall near temperatures of zero could increase.

The severity of snow damage is related to tree characteristics. Stem taper and crown characteristics are the most important factors controlling the stability of trees. Slightly tapering stems, asymmetric crowns, and rigid horizontal branching are all associated with high risk. However, the evidence on species differences is less clear due to the interaction with location. Management of forests can alter risk through choice of regeneration, tending, thinning and rotation. However, quantification and comparison of the absolute effect of these measures is not yet possible. An integrated risk model is required to allow the various locational and silvicultural factors to be assessed. Plans are presented to construct such a model, and gaps in knowledge are highlighted.

• Nykänen, E-mail: mn@mm.unknown
• Broadgate, E-mail: mb@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown
• Peltola, E-mail: hp@mm.unknown
• Quine, E-mail: cq@mm.unknown

Models for individual-tree basal area growth were constructed for Scots pine (Pinus sylvestris L.), pubescent birch (Betula pubescens Ehrh.) and Norway spruce (Picea abies (L.) Karst.) growing in drained peatland stands. The data consisted of two separate sets of permanent sample plots forming a large sample of drained peatland stands in Finland. The dependent variable in all models was the 5-year basal area growth of a tree. The independent tree-level variables were tree dbh, tree basal area, and the sum of the basal area of trees larger than the target tree. Independent stand-level variables were stand basal area, the diameter of the tree of median basal area, and temperature sum. Categorical variables describing the site quality, as well as the condition and age of drainage, were used. Differences in tree growth were used as criteria in reclassifying the a priori site types into new yield classes by tree species. All models were constructed as mixed linear models with a random stand effect. The models were tested against the modelling data and against independent data sets.

• Hökkä, E-mail: hh@mm.unknown
• Alenius, E-mail: va@mm.unknown
• Penttilä, E-mail: tp@mm.unknown

This paper reports on the possibility and difficulties in building growth models from past Forest Administration records on cut and growth in the Italian Alps. As a case study, a matrix model was calibrated for uneven-aged forests in the Valsugana valley of the Trentino province. The model gave reliable predictions over 30 years, and plausible long-term forest dynamics, including steady-states that are similar to virgin forests. The results support the view that the current forests are deeply altered as to composition, relative to what would obtain from natural growth. They also support the concept of long cyclic changes in natural stands, gradually approaching a climax state. Shortcomings of the data are that they do not come from an experimental design, they are not always accurate, and they must be supplemented with other information, especially concerning mortality. Still, these cheap and available data can lead to workable models adapted to local conditions, with many management applications.

• Virgilietti, E-mail: pv@mm.unknown
• Buongiorno, E-mail: jb@mm.unknown

The present study is the first attempt to carry out an inventory of greenhouse gas (GHG) fluxes in the forests of Estonia. The emission and uptake of CO₂ as a result of forest management, forest conversion and abandonment of cultivated lands in Estonia was estimated. The removal of GHG by Estonian forests in 1990 exceeded the release about 3.3 times. Changes in the species composition and productivity of forest sites under various simulated climate change scenarios have been predicted by using the Forest Gap Model for the central and coastal areas of Estonia. The computational examples showed that the changes in forest community would be essential.

• Mandre, E-mail: mm@mm.unknown
• Klõseiko, E-mail: jk@mm.unknown
• Reisner, E-mail: vr@mm.unknown
• Tullus, E-mail: ht@mm.unknown

A gap-model was used with forest inventory data in taking ground-true site, soil and tree characteristics into account in predicting the effects of climate change on forests. A total of 910 permanent sample plots established in the course of national forest inventory (NFI) in Finland and located on mineral soil sites in southern Finland were selected as the input data. The climatological input used in the simulations consisted of interpolated means of and deviations from long-term local temperature and precipitation records. The policy-oriented climate scenarios of SILMU (Finnish Research Programme on Climate Change) were used to describe the climate change. The temperature changes in the climate scenarios were increases of ca. +1.1 °C (low), +4.4 °C (medium) and +6.6 °C (high) compared to the current climate in 110 years. The simulation period was 110 years covering the time years 1990–2100.

Southern Finland, divided into fifteen forestry board districts, was used as the study region. Regional development of stand volume, cutting yield, and total wood production of forests under different climate scenarios were examined. The annual average growth in simulations under current climate was close to that observed in NFL Forests benefited from a modest temperature increase (Scenario 2), but under Scenario 1 the growing stock remained at a lower level than under the current climate in all parts of the study region. In wood production and cutting yield there were regional differences. In the southern part of the study regional wood production under Scenario 1 was ca. 10% lower than under the current climate, but in the eastern and western parts wood production was 5–15% higher under Scenario 1 than under the current climate. The relative values of total wood production and cutting yield indicated that the response of forests to climate change varied by geographical location and the magnitude of climate change. This may be a consequence of not just varying climatic (e.g. temperature and precipitation) and site conditions, but of varying responses by different kind of forests (e.g. forests differing in tree species composition and age).

• Talkkari, E-mail: at@mm.unknown

To project the changes in wood production of Norway spruce (Picea abies (L.) H. Karst.) and Scots pine (Pinus sylvestris L.) in Finland as a result of climate change, two separate studies were made. The first study, at the Faculty of Forestry, University of Joensuu, based its projections on mathematical models; the second one, at the Finnish Forest Research Institute, based projections on measurements of wood production in two series of aged provenance experiments. The results of the two studies were similar for both species: after a 4°C increase of the annual mean temperature a drastic increase in wood production in northern Finland, but little effect, or even some decrease in the southern part of the country. However, the assumptions used in the two studies differed. One important difference was that in the models the temperature is assumed to be increasing gradually over the years, whereas in the provenance experiments, climate changed immediately when the seedlings were transferred to the planting sites. Another problem with the provenance experiments is that when material is moved in a north-south direction in Finland, not only temperature but also photoperiod changes markedly. To compare these two studies, site factors (e.g. soil type, temperature, precipitation) and silvicultural factors (e.g. plant spacing, survival, time of thinning, thinning intensity) from the provenance experiments were included a variable in the mathematical models.

• Beuker, E-mail: eb@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown
• Kolström, E-mail: mk@mm.unknown

The productivity of Scots pine (Pinus sylvestris L.) under changing climatic conditions in the southern part of Finland was studied by scenario analysis with a gap-type forest ecosystem model. Standard simulations with the model predicted an increased rate of growth and hence increased productivity as a result of climatic warming. The gap-type model was refined by introducing an overwintering sub-model describing the annual growth cycle, frost hardiness, and frost damage of the trees. Simulations with the refined gap-type model produced results conflicting with those of the standard simulation, i.e., drastically decreased productivity caused by mortality and growth-reducing damage due to premature dehardening in the changing climate. The overwintering sub-model was tested with frost hardiness data from Scots pine saplings growing at their natural site 1) under natural conditions and 2) under elevated temperature condition, both in open-top chambers. The model predicted the frost hardiness dynamics quite accurately for the natural conditions while underestimating the frost hardiness of the saplings for the elevated temperature conditions. These findings show that 1) the overwintering sub-model requires further development, and 2) the possible reduction of productivity caused by frost damage in a changing climate is less drastic than predicted in the scenario analysis. The results as a whole demonstrated the need to consider the overwintering of trees in scenario analysis carried out with ecosystem model for boreal conditions. More generally, the results revealed a problem that exists in scenario analysis with ecological models: the accuracy of a model in predicting the ecosystem functioning under present climatic condition does not guarantee the realism of the model, nor for this reason the accuracy for predicting the ecosystem functioning under changing climatic conditions. This finding calls for the continuous rigorous experimental testing of ecological models used for assessing the ecological implications of climatic change.

• Hänninen, E-mail: hh@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown
• Leinonen, E-mail: il@mm.unknown
• Repo, E-mail: tr@mm.unknown

A system of zonality in Siberia has been formed under the control of continentality, which provides the heat and humidity regimes of the forest provinces. Three sectors of continentality and four to six boreal sub-zone form a framework for the systematization of the different features of land cover in Siberia. Their climatic ordination provides the fundamental basis for the principal potential forest types (composition, productivity) forecasting the current climate. These are useful in predicting the future transformations and succession under global change.

• Nazimova, E-mail: dn@mm.unknown
• Polikarpov, E-mail: np@mm.unknown

An equilibrium model driven by climatic parameters, the Siberian Vegetation Model, was used to estimate changes in the phytomass of Siberian vegetation under climate change scenarios (CO₂ doubling) from four general circulation models (GCM's) of the atmosphere. Ecosystems were classified using a three-dimensional climatic ordination of growing degree days (above a 5 °C threshold), Budyko's dryness index (based on radiation balance and annual precipitation), and Conrad's continentality index. Phytomass density was estimated using published data of Bazilevich covering all vegetation zones in Siberia. Under current climate, total phytomass of Siberia is estimated to be 74.1 ± 2.0 Pg (petagram = 1,015 g). Note that this estimate is based on the current forested percentage in each vegetation class compiled from forest inventory data.

Moderate warming associated with the GISS (Goddard Institute for Space Studies) and OSU (Oregon State Univ.) projections resulted in a 23–26 % increase in phytomass (to 91.3 ± 2.1 Pg and 93.6 ± 2.4 Pg, respectively), primarily due to an increase in the productive Southern Taiga and Sub-taiga classes. Greater warming associated with the GFDL (General Fluid Dynamics Laboratory) and UKMO (United Kingdom Meteorological Office) projections resulted in a small 3–7 % increase in phytomass (to 76.6 ± 1.3 Pg and 79.6 ± 1.2 Pg, respectively). A major component of predicted change using GFDL and UKMO is the introduction of a vast Temperate Forest-Steppe class covering nearly 40% of the area of Siberia, at the expense of Taiga; with current climate, this vegetation class is nearly non-existent in Siberia. In addition, Sub-boreal Forest-Steppe phytomass double with all GCM predictions. In all four climate change scenarios, the predicted phytomass stock of all colder, northern classes is reduced considerably (viz., Tundra, Fore Tundra, northern Taiga, and Middle Taiga). Phytomass in Sub-taiga increases greatly with all scenarios, from a doubling with GFDL to quadrupling with OSU and GISS. Overall, phytomass of the Taiga biome (Northern, Middle, Southern and Sub-taiga) increased 15% in the moderate OSU and GISS scenarios and decreased by a third in the warmer UKMO and GFDL projections. In addition, a sensitivity analysis found that the percentage of a vegetation class that is forested is a major factor determining phytomass distribution. From 25 to 50% more phytomass is predicted under climate change if the forested proportion corresponding to potential rather than current vegetation is assumed.

• Monserud, E-mail: rm@mm.unknown
• Denissenko, E-mail: od@mm.unknown
• Kolchugina, E-mail: tk@mm.unknown
• Tchebakova, E-mail: nt@mm.unknown

Two dynamic models predicting the development of frost hardiness of Finnish Scots pine (Pinus sylvestris L.) were tested with frost hardiness data obtained from trees growing in the natural conditions of Finland and from an experiment simulating the predicted climatic warming. The input variables were temperature in the first model, and temperature and night length in the second. The model parameters were fixed on the basis of previous independent studies. The results suggested that the model which included temperature and photoperiod as input variables was more accurate than the model using temperature as the only input variable to predict the development of frost hardiness in different environmental conditions. Further requirements for developing the frost hardiness models are discussed.

• Leinonen, E-mail: il@mm.unknown
• Hänninen, E-mail: hh@mm.unknown
• Repo, E-mail: tr@mm.unknown

The effect of species mixture was studied in a mixed stand of Norway spruce (Picea abies (L.) H. Karst.) and Scots pine (Pinus sylvestris L.) by simulating around 100 different treatment schedules during the rotation in a naturally regenerated even-aged stand located on a site of medium fertility in North Karelia, Finland. Both thinning from below and thinning from above were applied. Optimum rotations were determined by maximising the net present value calculated to infinity and different treatment schedules were compared with the net present value over one rotation as per rotation applied. In the optimum treatment programme, the proportion of pines was decreased by half of the basal area in the first thinning stage and by the end of the rotation to about one third. In thinning from above, the proportion of pines can be maintained at a slightly higher level. It is economically profitable to maintain the growing stock capital at approximately the level recommended by Forest Centre Tapio, a semi-governmental forestry authority. With non-optimum species composition, the loss in net present value over one rotation can be about 10 % in thinning from below and about 20 % in thinning from above.

• Vettenranta, E-mail: jv@mm.unknown

The prescribed burning of a 7.3 ha clear-cut and a 1.7 ha partially cut forest (volume 150 m³/ha) was carried out in Evo (61 °12'N, 25°07'E) on 1 June 1992. The forest was a mesic Myrtillus site type forest dominated by Norway spruce (Picea abies (L.) H. Karst.). Practically all the trees and the above-ground parts of the understorey vegetation died in the fire, while the mor layer was thinned by an average of 1.5 cm.

A study was made on the change of germinated seedling population in time and their dependence on environmental factors. Seedlings of Norway spruce, Scots pine (Pinus sylvestris L.), silver birch (Betula pendula Roth), pubescent birch (B. pubescens Ehrh.) and rowan (Sorbus aucuparia L.) were inventoried in 1993 and in 1994 on permanent plots, four times per growing season. Autoregression models were used to compare regeneration of tree species in the burned forest with regeneration in the burnt clear-cut area, and to study the effect of distance from nearest seed source to regeneration.

The average number of seedlings germinating in 1993 was higher than in 1994, probably because of differences between these consecutive years in regard to the amount of seed rain and weather conditions. The number of Norway spruce and rowan seedling was higher inside the forest area than in the clear-cut area. The distance to the bordering forest and to the closest seed tree did not explain the result. It is suggested that the more stable microclimatic conditions under the shade of dead tree promote germination and seedling establishment in the forest area. As rowan is a bird-dispersed species, it is likely that dead trees help the dispersal of rowan seed by providing birds place to sit and defecate. The shade provided by dead trees may influence the further succession of the tree stand and vegetation composition and diversity.

• Vanha-Majamaa, E-mail: iv@mm.unknown
• Suominen, E-mail: rs@mm.unknown
• Tonteri, E-mail: tt@mm.unknown
• Tuittila, E-mail: et@mm.unknown

Individual tree-growth models for diameter and height, and a model for the cylindrical stem form factor are presented. The aims of the study were to examine modelling methods in predicting growth response to thinning, and to develop individual-tree, distance-independent growth models for predicting the development of thinned and unthinned stands of Scots pine (Pinus sylvestris L.). The models were constructed to be applicable in simulation systems used in practical forest management planning. The models were based on data obtained from eleven permanent thinning experiments located in even-aged Scots pine stands in Southern and Central Finland.

Two alternative models were developed to predict tree diameter growth in thinned and unthinned stands. In the first model, the effect of stand density was described using stand basal area. In the alternative model, an explicit variable was incorporated referring to the relative growth response due to thinning. The magnitude of the growth response was expressed as a function of thinning intensity. The Weibull function was employed to describe the temporal distribution of the thinning response. Both models resulted in unbiased predictions in unthinned and in moderately thinned stands. An explicit thinning variable was needed for unbiased growth prediction in heavily thinned stands, and in order to correctly predict the dynamics of the growth response.

In the height growth model, no explicit thinnning variable referring thinning was necessary for growth prediction in thinned stands. The stem form factor was predicted using the model that included tree diameter and tree height as regressor variables. According to the results obtained, the information on the changes in the diameter/height ratio following the thinning is sufficient to predict the change in stem form.

• Hynynen, E-mail: jh@mm.unknown

Semiparametric models, ordinary regression models and mixed models were compared for modelling stem volume in National Forest Inventory data. MSE was lowest for the mixed model. Examination of spatial distribution of residuals showed that spatial correlation of residuals is lower for semiparametric and mixed models than for parametric models with fixed regressors. Mixed models and semiparametric models can both be used for describing the effect of geographic location on stem form.

• Kangas, E-mail: ak@mm.unknown
• Korhonen, E-mail: kk@mm.unknown

Much of forestry data is characterized by a longitudinal or repeated measures structure where multiple observations taken on some units of interest are correlated. Such dependencies are often ignored in favour of an apparently simpler analysis at the cost of invalid inferences. The last decade has brought to light many new statistical techniques that enable one to successfully deal with dependent observations. Although apparently distinct at first, the theory of Estimating Functions provides a natural extension of classical estimation that encompasses many of these new approaches. This contribution introduces Estimating Function Theory as a principle with potential for unification and presents examples covering a variety of modelling issues to demonstrate its applicability.

• Schabenberger, E-mail: os@mm.unknown
• Gregoire, E-mail: tg@mm.unknown

A spatial growth model is presented for Scots pine (Pinus sylvestris L.) on a dwarf-shrub pine mire drained 14 years earlier. The growth model accounts for the variation in tree diameter growth owing to the competition between trees, the distance between tree and ditch, and the time passed since drainage. The model was used to study the effect of tree arrangement on the post-drainage growth of a pine stand. Clustering of trees decreased the volume growth by 9–20% as compared to a regular spatial distribution. Stand volume growth, for a given number of stems, was at its maximum and variation in diameter growth at its minimum when the stand density on the ditch border was 1.5–5 higher than midway between two adjacent ditches.

• Miina, E-mail: jm@mm.unknown

In this study, model-based and design-based inference methods are used for estimating mean volume and its standard error for systematic cluster sampling. Results obtained with models are compared to results obtained with classical methods. The data are from the Finnish National Forest Inventory. The variation of volume in ten forestry board districts in Southern Finland is studied. The variation is divided into two components: trend and correlated random errors. The effect of the trend and the covariance structure on the obtained mean volume and standard error estimates is discussed. The larger the coefficient of determination of the trend model, the smaller the model-based estimates of standard error, when compared to classical estimates. On the other hand, the wider the range and level of autocorrelation between the sample plots, the larger the model-based estimates of standard error.

• Kangas, E-mail: ak@mm.unknown

Regression models for estimating stem volume of Scots pine (Pinus sylvestris L.) were constructed using sample tree data measured in the 7th and 8th National Forest Inventory of Finland. Stem volume were regressed on diameter, basal area of growing stock, and geographic location. The results of the study show that using second order trend surface to describe the geographic variation of the residuals gives satisfactory results. Using mixed estimation for combining old and new sample tree data improves the efficiency of an inventory. The weight of the prior information must be low, because remarkable differences in stem form was found in the two inventories.

The PDF includes an abstract in Finnish.

• Korhonen, E-mail: kk@mm.unknown

In the method presented in this study, a group of experts evaluate, in a pairwise manner, a set of forest areas with respect to the game species considered. On the basis of these comparisons, relative priorities of forest areas are estimated using the eigenvalue technique. Using regression analysis, a habitat suitability function is estimated in which the priority is predicted by measures already familiar in forest planning. As a case study, a habitat suitability function was estimated for black grouse (Tetrao tetrix, Lururus tetrix L.). The function is applicable in forestry planning carried out using modern planning techniques.

The PDF includes an abstract in Finnish.

• Kangas, E-mail: jk@mm.unknown
• Karsikko, E-mail: jk@mm.unknown
• Laasonen, E-mail: ll@mm.unknown
• Pukkala, E-mail: tp@mm.unknown

Yield of Cupressus lusitanica Mill. was modelled by predicting the diameter distribution of trees at given stand ages. The beta distribution was used as a theoretical distribution. The models used for the calculation of diameter distribution were based on 66 temporal sample plots with varying age, site and stand density. The growing sites of Cupressus lusitanica were divided into four classes on the basis of age and dominant height. Using the stand models developed in the study, the yield and profitability of different thinning schedules was evaluated by a simulation technique. In the simulated treatment regimes, the mean annual increment varied from 6.6 m³/ha in the poorest site class to 16.6 m³/ha in the best class with rotation lengths ranging from 25 years (best sites) to 34 years (poorest sites). With typical planting densities (1,600 trees/ha), thinnings increase the total harvest by a few percentage points and improved the profitability of plantation forestry.
The PDF includes an abstract in Finnish.

• Pohjonen, E-mail: vp@mm.unknown
• Pukkala, E-mail: tp@mm.unknown

Mixed linear models were constructed to describe the height development of Norway spruce (Picea abies (L.) H. Karst.) and Scots pine (Pinus sylvestris L.) advance growth after release cutting. The models related density of the overstory, time elapsed since release cutting and tree size with annual height increment. Parameters of preliminary models were estimated from a limited data set to judge the feasibility of the approach for further studies.

The PDF includes an abstract in Finnish.

• Koistinen, E-mail: ek@mm.unknown
• Valkonen, E-mail: sv@mm.unknown

A model for the mechanism of windfall and stem breakage was constructed for single Scots pine (Pinus sylvestris L.) at the stand edge. The total turning moment arising from the wind drag and from the bending of stem and crown was calculated along with the breaking stress of the stem. Similarly, the support given by the root -soil plate anchorage was calculated. Windspeed variation within the crown and the vertical distribution of stem and crown weight were taken into account. Model computations showed that trees having a large height to diameter ratio were subjected to greater risk of falling down or breaking than trees with a small height to diameter ratio. The windspeed required to blow down a tree or break the stem of a tree decreased if the height to diameter ratio or the crown to stem ratio of trees increased.
The PDF includes an abstract in Finnish.

• Peltola, E-mail: hp@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown

Models for estimating the upper diameter of trees were constructed using sample tree data measured in the 7th National Forest Inventory in Finland. Calibration of the models was tested with data from the 8th National Forest Inventory. The results showed that using mixed estimation for combining the two data sets improves the reliability of the models. Models and methods used in this study can be recommended for use in forest inventories.

The PDF includes an abstract in Finnish.

• Korhonen, E-mail: kk@mm.unknown

An alternative approach to formulating a forestry goal programming problem is presented. First, single objective optima levels are solved. The Analytical Hierarchy Process is applied in the estimation of a priori weights of deviations from the goal target levels. The ratios of the weights can be interpreted as relative importance of the goals, respectively. The sum of the weighted deviations from all single optima levels associated with the management goals is minimized. Instead of absolute deviations, relative ones are used. A case study problem of forest management planning with several objectives, measured in different units, is analysed.

The PDF includes an abstract in Finnish.

• Kangas, E-mail: jk@mm.unknown
• Pukkala, E-mail: tp@mm.unknown

Model computations on the management of Scots pine (Pinus sylvestris L.) at the seedling stage showed that a rising temperature due to the suggested climate change could increase the competition capacity of birch species (Betula pendula) more than Scots pine, whose growth could even decline during the course of a rise in temperature. A temperature rise could, thus, bring the time of removal of birches forward when aiming at Scots pine timber stands composed of these tree species. The increasing proportion of birches makes the removal of birches even more urgent and emphasizes the need for careful management of Scots pine stands under rising temperatures. The first thinning of Scots pine is generally brought forward; this is particularly the case when wide spacing is applied in planting. A furthrer rise in temperature magnifies the above patterns by reducing further the competitive capacity of Scots pine in relation to birches.
The PDF includes an abstract in Finnish.

• Kellomäki, E-mail: sk@mm.unknown
• Kolström, E-mail: mk@mm.unknown

A model for the succession of the forest ecosystem is described. The growth and development of trees and ground cover are controlled by temperature and light conditions and the availability of nitrogen and water. In addition, the effects of the annual cycle of trees including the risk of frost damage, wild fire, and wind damages are contained in the model as factors which control the survival and productivity of trees. The model also makes it possible to evaluated the risk of insect attack assuming that this risk is inversely related to the growth efficiency of trees.

The PDF includes an abstract in Finnish.

• Kellomäki, E-mail: sk@mm.unknown
• Hänninen, E-mail: hh@mm.unknown
• Kolström, E-mail: tk@mm.unknown
• Lauhanen, E-mail: rl@mm.unknown
• Mattila, E-mail: um@mm.unknown
• Pajari, E-mail: bp@mm.unknown
• Väisänen, E-mail: hv@mm.unknown

Old inventory data has widely been used as prior information in forest inventory using the method of sampling with partial replacement (SPR). In this method knowledge about forest growth has not been utilized. However, the accuracy of the inventory results can be improved if this knowledge is utilized. The usability of the inventory results can be improved if the prior information is updated by treewise growth models. In this paper a statistical basis is presented for a method in which such information can be used. The applicability of the method is also discussed. An example is given to demonstrate the method.

The PDF includes an abstract in Finnish.

• Kangas, E-mail: ak@mm.unknown

The simulation model consists of a method to generate theoretical Norway spruce (Picea abies (L.) H. Karst.) stands, and a spatial growth model to predict the growth of these stands. The stand generation procedure first predicts the tree diameters from a few stand characteristics and from tree locations. Tree age and height are predicted using spatial models. Spatial growth models were made for both diameter growth and basal area growth. Past growth was used as a predictor in one pair of models and omitted in another pair. The stand generation method and the growth models were utilized in studying the effect of tree arrangement and thinning method on the growth of a Norway spruce stand.

The PDF includes an abstract in Finnish.

• Pukkala, E-mail: tp@mm.unknown
• Kolström, E-mail: tk@mm.unknown

Decision making in the forest protection involves diagnosing the pest, making predictions of the effects of the pest on forest, knowing the possible control tactics, and cost/benefit integration. To cope with all that, a generalist forest manager needs a tool like an expert system to support decisions.

This paper presents an expert system that approaches the goals of integrated pest management. With the systm, the user can make diagnosis and predictions of 12 North European bark beetles. Written in Common LISP and Flavors, the expert system has a combined frame- and rule-based knowledge representation. Frames are used to represent the hierarchy of insect taxonomy in diagnosis. Prediction is made with qualitative reasoning with rules. The interface engine applies both forward and backward chaining. The system has a graphical user interface that supports exploring the sensitivity of advice on input.

It is concluded that expert systems and artificial intelligence have high applicability everywhere in forestry where complicated decisions have to be made. Especially, an integrated pest management system in forestry is largely equivalent to a computerized decision-making aid.

The PDF includes an abstract in Finnish.

• Saarenmaa, E-mail: hs@mm.unknown

Linear programming was used to analyse the land use alternatives in the Debre Birhan Fuelwood Plantation area, in the central highlands of Ethiopia. The region represents a rural, high-altitude area, where the main land uses are grazing and cultivation of barley, wheat and pulses. To alleviate fuelwood shortage, large plantations of Eucalyptus globulus Labill. have been established. Livestock has traditionally used the major part of the production capacity of the sites. A decrease in the number of cattle would facilitate a considerable increase in the production of cereals, pulses, fuelwood and construction timber. The optimal share of the land for arable crops, grazing and tree plantations would be about 40, 45 and 15% respectively.

The PDF includes an abstract in Finnish.

• Pukkala, E-mail: tp@mm.unknown
• Pohjonen, E-mail: vp@mm.unknown

The paper presents a simple model of long-term forest management planning in tree plantations. The model is particularly suitable for developing countries where the research resources are limited. The management plan is prepared in two steps. First, one or several treatment schedules are simulated for each calculation unit (age class, compartment, etc.) over the selected planning period. Second, an optimal combination treatment schedules according to the selected objectives and constraints is searched by mathematical programming. The simulation of growth is based on the prediction of the diameter distribution at the desired time point. All stand characteristics are derived from this distribution. The models needed in the yield simulation can be estimated from temporary sample plots. A case study management plan for 13,000 ha of Pinus kesiya (Royle ex Gordon) plantations in Zambia is presented to demonstrate the system.

The PDF includes an abstract in Finnish.

• Pukkala, E-mail: tp@mm.unknown
• Mubita, E-mail: om@mm.unknown
• Saramäki, E-mail: js@mm.unknown

Ring width at breast height is presented as a function of stem radius at breast height, the ratio between the diameter of a tree and the basal area median diameter, site index, and density of stand. By means of a conversion model ring width at stump height can be estimated as a function of ring width at breast height.

According to previous studies substantially better wood quality can be expected if mean width near the pith at stump height decreases from 3 to 2 mm. According to the present study only on the poorest sites suitable for Scots pine (Pinus sylvestris L.) planting (poor Vaccinium type) the ring width is less than 3 mm at stump height even in the thickest trees. On more fertile sites a substantial increase in the recommended planting density is required, if the mean ring width is aimed to be less than 3 mm. On the best sites it is impossible to reach mean ring width of less than 2 mm, when the density is less than 4,000 stems/ha. Only the thinnest trees on the poorest sites can have a mean ring width less than 2mm.

The PDF includes an abstract in English.

• Varmola, E-mail: mv@mm.unknown

The study uses the methodology of ecological field theory to model the effect of Scots pine (Pinus sylvestris L.) seed trees on the density of tree seedlings and other plants in the field layer. The seed trees had a clear effect on the expected value of the amount and distribution of the ground vegetation. The vicinity of seed trees had an adverse effect on the growth of grasses, herbs and seedlings, while mosses were most abundant near the trees. Models based on the ecological field approach were derived to describe the effect of seed trees on the ground vegetation.

The PDF includes an abstract in Finnish.

• Kuuluvainen, E-mail: tk@mm.unknown
• Pukkala, E-mail: tp@mm.unknown

The single tree growth models presented in this study were based on about 4,000 trees measured in 50 even-aged Scots pine (Pinus sylvestris L.) sample plots with varying density, spatial pattern of trees and stand age. Predictors that used information about tree locations decreased the relative standard error of estimate by 10 percentage points (15%), if past growth was not used as a predictor, and about 15 percentage points (30%) when past growth was one of the predictors. When ranked according to the degree of determination, the best growth models were obtained for the basal area increment, the next best for relative growth, and the poorest for diameter increment. The past growth decreased the relative standard error of estimate by 15–20 percentage points, but did not make the spatial predictors unnecessary. The degree of determination of the spatial basal area growth model was almost 80% if the past growth was unknown and almost 90% if the past growth was known. Variables that described the amount of removed competition did not improve the growth models.

The PDF includes an abstract in Finnish.

• Pukkala, E-mail: tp@mm.unknown

The study presents two methods of predicting tree dimensions in a Scots pine (Pinus sylvestris L.) stand if only the location of trees is known. The first method predicts the tree diameter from the spatial location of neighbours. In the second method the diameter distribution of a subarea is estimated from the local stand density. This distribution is then sampled to obtain diameters. In both methods the tree height is predicted with a spatial model on the basis of diameters and locations of trees. The main purpose of the presented models is to generate realistic stands for simulation studies.

The PDF includes an abstract in Finnish.

• Pukkala, E-mail: tp@mm.unknown

Models concerning the effects of temperature on dormancy release in woody plants were tested using two-year old seedlings of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.). Chilling experiments suggest that the rest period has a distinct end point. Before the attainment of this end point high temperatures do not promote bud development towards dormancy release, and after it further chilling does not affect the subsequent bud development. A new hypothesis of dormancy release is suggested on the basis of a comparison between present and earlier findings. No difference in the proportion of growth commencing seedlings were detected between the forcing temperatures of 17°C and 22°C. The rest break of 50% of Norway spruce and Scots pine seedlings required six and eight weeks of chilling, respectively. Great variation in the chilling requirement was found, especially for Scots pine.

The PDF includes an abstract in Finnish.

• Hänninen, E-mail: hh@mm.unknown
• Pelkonen, E-mail: pp@mm.unknown

The effect of grouping on 5-year old volume increment was studied by a simulation technique using spatial growth models estimated in Scots pine (Pinus sylvestris L.) stands in the phase of the first commercial thinning. A total of 24 model stands were regenerated by applying 12 spatial processes for two different diameter distributions. In addition to model stands, 6 different thinnings were simulated in two real stands. The clustering of trees was described with Fisher’s grouping index and by estimating the relative interception of diffuse radiation. In model stands with constant diameter distribution the correlation between the grouping index and volume increment ranged from -0.81 to -0.91. The correlation between volume increment and interception was 0.81–0.83 with one diameter distribution and 0.70 if both distributions were combined. In one thinned stand the correlation between the growth estimate and grouping index varied between -0.33 and 0.76. The correlation between interception and growth was about 0.30 in one stand and 0.72 if both stands were combined. Small irregularities do not decrease the volume production of a young Scots pine stand, but if the clustering is considerable or there are reasonably wide harvest strips, growth will be reduced by 10–20%.

The PDF includes a summary in Finnish.

• Pukkala, E-mail: tp@mm.unknown

Logical structure of three simulation models and one conceptual model concerning effects of temperature on dormancy release in woody plants was examined. The three basic types of simulation models differed in their underlying assumptions. Contrasting implications of the models were inferred by deduction. With the aid of these implications, the model types can be tested using experiments with continuous and interrupted chilling. Similarly, implications of the conceptual model of rest phases were inferred, by which the model can be tested using experiments with continuous chilling and forcing in multiple temperatures. The possibilities to synthetize the conceptual model with any of the three simulation model types, as well as the biological interpretation of the model variables, were discussed.

The PDF includes an abstract in Finnish.

• Hänninen, E-mail: hh@mm.unknown

The seed crop of Norway spruce (Picea abies (L.) H. Karst.) and Scots pine (Pinus sylvestris L.) is predicted with the help of mean monthly temperatures during May–August one and two years before the flowering year. The prediction models were made separately for Lapland and for the rest of Finland. The models are based on 10-year periods of seed crop measurements and climatic data. The total number of time series was 59.

In Lapland, Norway spruce flowered abundantly and produced an abundant seed crop after warm July–August and two years after cool July–August. In other parts of Finland, warm June and July produced a good flowering year, especially if these months were cool two years before the flowering year.

In Lapland, Scots pine flowered abundantly if the whole previous growing season was warm. Elsewhere in Finland, a cool June preceded prolific flowering in the coming year if the rest of the growing season was considerably warmer than the average.

The prediction models explained 37–49 % of the variation in the size of the seed crop. The occurrence of good and poor seed years was usually predicted correctly. Using the presented models, the prediction of the seed crop is obtainable 1.5 year for Norway spruce and 2.5 year for Scots pine before the year of seed fall.

The PDF includes an abstract in English.

• Pukkala, E-mail: tp@mm.unknown

In the model the regeneration process is derived into three subprocesses: birth, growth and mortality of seedlings. The main emphasis is on the birth process where the following phases are simulated: seed crop, quality of seeds, maturity of seeds, predation of seeds and germination. The parameters are based on data published in Finland. Part of the parameters are obtained directly from the investigations and part is proposed by the author. The model can be calibrated by changing parameter values. The simulation is made with the help of random numbers which have the same means as the estimates and the same distributions as the residuals of the equations used in simulation. The time step of the model is one year. The number of emerged seedlings in one year is obtained by multiplying the seed crop with the probabilities that the seed passes different phases of the birth process. Because of stochasticity the regeneration period is simulated several times. From the results it is possible to evaluate the risk and succeeding probability of the regeneration. The main drawbacks of the simulation method are the lack of empirical parameters and the difficulty of testing. The model could be further developed by including spatiality into the model.

The PDF includes an abstract in Finnish.

• Pukkala, E-mail: tp@mm.unknown

This study deals with the evaluation of logging machines. The analyses were based on the results of a productivity study with special reference to a PIKA 35 processor, a delimber-bucker, working in Kyröskoski forest area in Finland. Factors affecting the productivity of the machine were surveyed. Mathematical models for determining the productivity were developed and their practical applications to the particular problem under study was demonstrated.

The PDF includes an abstract in Finnish.

• Tan, E-mail: jt@mm.unknown

The effect of the size of seed crop, dispersal of seeds and the early development of seedlings on the density and spatial distribution of young Scots pine (Pinus sylvestris L.) stands are evaluated on the basis of theoretical models. The models include (i) number and spatial distribution of parent trees on the regeneration area, (ii) size of annual seed crop, (iii) seed dispersal from a particular parent tree, (iv) germination of the seeds (germination percentage), (v) death of ageing seedlings after the establishment process, and (vi) height growth of the seedlings.

As expected, stand density and spatial distribution varied within a large range in relation to the density of the parent trees and the distance from them. The simulations also showed that natural seedling stands can be expected to be heterogenous due to the geometry of seed dispersal, emphasizing the frequency of young and small trees. The properties of the seedling stands were, however, greatly dependent on the density of the parent trees and the length of the regeneration period.

The PDF includes an abstract in Finnish.

• Kellomäki, E-mail: sk@mm.unknown
• Hänninen, E-mail: hh@mm.unknown
• Kolström, E-mail: tk@mm.unknown
• Kotisaari, E-mail: ak@mm.unknown
• Pukkala, E-mail: tp@mm.unknown

The usefulness of forest sector models in forest policy analysis is discussed, mainly based on experiences from Norway. Forest sector modelling is contrasted to two alternative approaches: (i) Intuitive, verbal analysis, and (ii) econometric models. It is concluded that forest sector models, properly developed in contact with the policy makers, should be of considerable value in forest policy analysis.

• Solberg, E-mail: bs@mm.unknown

This paper designs an Economy-Welfare-Environment Adjustment System model (EWEAS model or EWE model in short) which consists of the circular flow of the economic, the welfare, and the environment system of forestry. That is, this model builds the relationship between the systems for material wealth and that for mental wealth.

The EWE model is designed as a complete open system model which describes the economy-welfare-environment circular system in forestry by linking up the internal system of forestry with the surrounding external systems. The EWE model can be manipulated as a policy formation or a policy decision model, and it is available for policy evaluation in the economic, the welfare and the environmental phase of forestry. The model is a basic simulation system model which is reliable in its reproductive fitness, stability and universality. Thus, this model ought to be useful in any country in the world as well as in Japan.

• Kishine, E-mail: tk@mm.unknown

The issue of Silva Fennica comprises 22 papers on forest policy and program analysis, and evaluation presented at the XVIII IUFRO World Congress in 1986 arranged in Ljubljana. The papers discuss the future and role of policy and program analysis, the effectiveness of policy programs on timber supply and private forestry investments as well as the application of forest sector models to policy analysis.

The PDF includes the preface and list of authors in English and an abstract in Finnish.

• Tikkanen, E-mail: –

The paper discusses the evaluation of timber production policies with different income (timber drain) schedules. Special attention is given to the temporal smoothness of the income flow. A utility model is formulated in which the objective is to maximize a fixed consumption pattern, and money can be saved and borrowed at different interest rates. We thus have smoothness requirements only for consumption, the capital market then determines the smoothness of the optimal income flow. Present discounted value and maximization of even income flow criteria are special cases of the utility model. Consumption can be maximized by linear programming. A sample problem is presented.

The PDF includes a summary in Finnish.

• Lappi, E-mail: jl@mm.unknown
• Siitonen, E-mail: ms@mm.unknown

Thinning models are generally based on the density of the stand measured by the average basal area per hectare, for instance. These models are handicapped by the uneven structure of the stands. In uneven stands the averages are inadequate indicators for the need and amount of thinnings.

Small relascope plots were tested in the measurement of the spatial distribution of trees and in the determination of the need and amount of thinnings. The thinning quantity was determined as the difference between the actual distribution of the relascope plots into basal area classes and the ideal distribution after thinning. Sequential sampling was used in the derivation of the decision equations. A respective BASIC-program for a programmable pocket calculator is given.

The PDF includes a summary in English.

• Kilkki, E-mail: pk@mm.unknown
• Pohjola, E-mail: tp@mm.unknown
• Pohtila, E-mail: ep@mm.unknown

Two Japanese models regarding the within-stand competition have been reviewed on the basis of relevant literature. Competition-density and 3/2 th power models seem to be applicable also into tree stands. The latter model has been applied into the material obtained from literature. Computations showed consistancy with the results obtained elsewhere in the world. It is concluded that also in Finnish conditions the 3/2 th power law may have great potentials in describing the effects of stand density on tree size.

The PDF includes a summary in English.

• Kellomäki, E-mail: sk@mm.unknown
• Nevalainen, E-mail: tn@mm.unknown

A dynamic programming approach toward stem value estimation for standing Scots pine (Pinus sylvestris L.) trees was developed. The determination of the saw log value was based on the sawing pattern and on the final products composition. The combination of taper curve models and bark models providing taper curves both over bark and under bark, which constituted the basis of the optimum stem scaling. A computer program was developed to determine the optimum log sequence of the stem aiming at maximizing the value of the final products. To examine the reliability of the computation system, 445 Scots pine sample trees from 29 stands were used as a test material. The stem values of sample trees were calculated in two ways: 1) with 12 measured diameters, and 2) with 12 estimated diameters derived from measured tree characteristics. In both cases the values of the intermediate diameters were calculated via cubic spline interpolation.

The PDF includes a summary in Finnish.

• Luo, E-mail: fl@mm.unknown

Methods involving the use of moving averages, trend surfaces and their combination are compared in deriving local values of monthly mean temperatures and precipitation sums from the observations made by the Finnish Meteorological Office. Correlation between meteorological variables and sea index, lake index and height above sea level were used in the trend surface method and in the combined method. Combined method, with a trend surface calculated from means of a long time period, was the most reliable method to estimate long local time series.

A method to calculate unbiased estimates of effective temperature sums from monthly mean temperatures is presented.

The PDF includes a summary in English.

• Ojansuu, E-mail: ro@mm.unknown
• Henttonen, E-mail: hh@mm.unknown

A model was constructed, the aim of which was to predict growth under conditions where air pollutants are present. The model is based on photosynthesis and on the allocation of photosynthetic products for growth. It is assumed that air pollutants released during energy production mainly affect photosynthesis in two ways: 1) directly by injuring the photosynthetic mechanism, and 2) indirectly by leaching nutrients. The two ways were studied empirically in order to identify a sub-model for the photosynthesis of a plant exposed to air pollutants. 

The stand model will be applied to two purposes. The present stage of forests in Finland is compared with the simulated state based on the assumption that no pollutants are present. In addition, the decrease in forest yield under different conditions derived from predictions about long-range pollutant transport in Europe is analysed.

• Mäkelä, E-mail: am@mm.unknown
• Hari, E-mail: ph@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown

A semi-statistical model is suggested for monitoring injuries of plants for long-time field exposures (months). The model is based on the following assumptions:

1. The concentrations of air pollutants in the atmosphere follow the Johnson SB distribution.

2. The degree of plant injury is proportional to the logarithm of air pollutant dose.

3. No injuries occur below a certain dose level.

4. A dose is defined as the air pollutant concentration multiplied by the duration of exposure raised to an exponent.

Based on the air pollutant frequency distribution a total dose for the exposure period is calculated by integration, and the total dose is related to the observed plant injury by non-linear regression. The model is tested for long-time exposures of sulphur dioxide to transplant lichen in natural environment.

• Moseholm, E-mail: lm@mm.unknown

This paper presents the principles of a unified data processing system suitable for derivation of the most variables of interest in forest mensuration. The precedence (succedence) relations between the tree and forest stand variables are analysed and a block-wise simultaneous recursive multi-equation model is suggested to describe these relations. Regression analysis is used in the estimation of the model parameters and Taylor’s series and Monte Carlo simulation are available in the derivation of the unbiased results.

The PDF includes a summary in Finnish.

• Kilkki, E-mail: pk@mm.unknown

In the original set of equations derived by regression analysis, relative-height diameters (endogenous variables) are presented as nonlinear functions of the other relative-height diameters and of the height of the tree (an exogenous variable). Any of the original equations can be replaced by an interpolation formula which links a measured diameter to the four closest relative-height diameters. The solution of the simultaneous equation model yields 10 relative-height diameters. Intermediate values are obtained to avoid biases due to the nonlinearity of the simultaneous model equations.

The PDF includes a summary in Finnish.

• Kilkki, E-mail: pk@mm.unknown
• Varmola, E-mail: mv@mm.unknown

A simultaneous equation model to determine taper curve for Scots pine is presented. The diameters at relative heights are endogenous variables and height an exogenous variable. Any equations may be substituted by the measured value of diameter. Solution of the system of equations yields 11 diameters at relative heights. Intermediate values are obtained by interpolation. Interpolation allows the use of diameters measured at absolute heights, too.

The PDF includes a summary in Finnish.

• Kilkki, E-mail: pk@mm.unknown
• Saramäki, E-mail: ms@mm.unknown
• Varmola, E-mail: mv@mm.unknown

The purpose of this study was to analyse economic models used in certain studies and to evaluate whether the economic theory envisaged by a model is consistent with its aims in forest policy. Second, the paper will point out assumed improvements for models that can be used as a basis for forest policy.

There are two types of economic models according to their purpose. One type, the marketing models, can be used for explaining or forecasting the consumer’s behaviour with no intension to affect the economies to be gained from the alternative patterns of behaviour explained. Others, the policy models, are meant to serve as guideposts which by means of normative forecasts point out the programme to be followed in order to attain certain aims. The majority of the policy models are static. The paper assesses the static models, and evaluates how well they fit their purpose. Special attention is given to dynamic economic models. A dynamic model can, at least in principle, be used to explain the course of events during the adjustment period required to achieve a production goal.

The PDF includes a summary in Finnish.

• Riihinen, E-mail: pr@mm.unknown

The mean has a great importance in statistics in general and also in forest statistical calculations. The meaning of the average tree and its characteristics is important also for the practical forest mensuration work. However, the question is how are the statistical numbers of a mean tree related to the statistical numbers of the stand.   

Study is based on the strip-wise survey of forests in southern Finland. From that information the 30 sample plots were chosen, 10 of each of most typical forest site types, MT, VT and CT. The stands are of different ages and development classes, varying from 14 to 159 years.

For the determination of the average tree are the statistical numbers of five characteristics needed: volume, basal area, diameter, height and form factor. The stereometric mean tree of the stand can be calculated with only one statistical method and that solution is absolute.

Theoretically and statistically absolute solution for the problem is the continuous solution by the mean that is weighted with the number of stems. This solution however is not very useful in practical sense.

A simple, practical and adequately exact solution for determining the average tree by approximation procedure of a certain arithmetic mean. 

• Lönnroth, E-mail: el@mm.unknown

In this study we analyse how the ion concentrations in forest soil solution are determined by hydrological and biogeochemical processes. A dynamic mode ACIDIC was developed, including processes common to dynamic soil acidification models. The model treats one to eight interacting layers and simulates soil hydrology, transpiration, root water and nutrient uptake, cation exchange, dissolutions and reaction of Al hydroxides in solution, and the formation of carbonic acid and its dissociation products. It includes also a possibility to a simultaneous use of preferential and matrix flow paths, enabling the throughfall water to enter the deeper soil layers in macropores without first reacting with the upper layers. Three different combinations of routing the throughfall water via macro- and micropores through the soil profile is presented. The large vertical gradient in the observed total charge was simulated successfully. According to the simulations, gradient is mostly caused by differences in the intensity of water uptake, sulphate adsorption and organic anion retention at the various depths. The temporal variations in Ca and Mg concentrations were simulated fairly well in all soil layers. For H⁺, Al and K there were much more variation in the observed than in the simulated concentrations. Flow in macropores is a possible explanation for the apparent disequilibrium of the cation exchange for H⁺ ad K, as the solution H⁺ and K concentrations have great vertical gradients in soil. The amount of exchangeable H⁺ increased in O and E horizons and decreased in the Bs1 and Bs2 horizons, the net change in whole soil profile being a decrease. A large part of the decrease of the exchangeable H⁺ in the illuvial B horizon was caused by sulphate adsorption. The model produces soil water amounts and solution ion concentrations which are comparable to the measured values, and it can be used in both hydrological and chemical studies of soils.

• Kareinen, E-mail: tk@mm.unknown
• Nissinen, E-mail: an@mm.unknown
• Ilvesniemi, E-mail: hi@mm.unknown

Productive coexistence and gain of populations were studied using nine years’ data from field experiments of Taxodium ascendens-intercrop systems in Lixiahe, Jiangsu province, China. A theoretical framework for productive coexistence in agroforestry was developed. Interaction patterns between trees and intercrops were presented within the framework. A model framework was developed to describe the coexistence gain and interaction of populations in T. ascedens-intercrop systems. Facilitation and resource sharing were identified as main contribution to the advantage of species combination in agroforestry. The model of population interaction developed in the present study was accepted for describing the interaction of populations in T. ascendens -intercrop system, because it explained a high proportion of the variance of experimental data and fitted well the observations in most intercropping types. The model provides flexibility for describing different patterns of intra- and inter-specific interactions. Model coefficients were applied to the determination of the ecological compatibility of species.

Managed T. ascendens-intercrop systems were advantageous as compared to a monoculture of trees or arable crops. In T. ascendens stands up to the age of three, arable crops contributed about 50–80% of the total biomass yield of agroforestry. The diameter height growth of T. ascendens was not significantly influenced by intercrops. When the trees were young (during the first three years), T. ascendens did not depress the crop yields, and a land equivalent ratio greater than unity was obtained together with a high yield of both components. The diameter and height of the trees were similar in four spacing configurations with an equal number of trees per hectare up to the age of 8, but wider between-rows open range were beneficial for the intercrops. The relationship between open-ranges and species coexistence was also analysed and the distribution of soil nutrients studied.

• Huang, E-mail: wh@mm.unknown

To enhance the utilization of the wood, the sawmills are forced to place more emphasis on planning to master the whole production chain from the forest to the end product. One significant obstacle to integrating the forest-sawmill-market production chain is the lack of appropriate information about forest stands. Since the wood procurement point of view in forest planning systems has been almost totally disregarded there has been a great need to develop an easy and efficient pre-harvest measurement method, allowing separate measurement of stands prior to harvesting. The main purpose of this study was to develop a measurement method for Scots pine (Pinus sylvestris L.) stands which forest managers could use in describing the properties of the standing trees for sawing production planning.

Study materials were collected from ten Scots pine stands located in North Häme and South Pohjanmaa, in Southern Finland. The data comprise test sawing data on 314 pine stems, diameter at breast height (dbh) and height measures of all trees and measures of the quality parameters of pine sawlog stems in all ten study stands as well as the locations of all trees in six stands. The study was divided into four sub-studies which deal with pine quality prediction, construction of diameter and dead branch height distributions, sampling designs and applying height and crown height models. The final proposal for the pre-harvest measurement method is a synthesis of the individual sub-studies.

Quality analysis resulted in choosing dbh, distance from stump height to the first dead branch, crown height and tree height as the most appropriate quality characteristics of Scots pine. Dbh and dead branch height are measured from each pine sample tree while height and crown height are derived from dbh measures by aid of mixed height and crown height models. Pine and spruce diameter distribution as well as dead branch height distribution are most effectively predicted by the kernel function. Roughly 25 sample trees seem to be appropriate in pure pine stands. In mixed stands the number of sample trees needs to be increased in proportion to the intensity of pines in order to attain the same level of accuracy.

• Uusitalo, E-mail: ju@mm.unknown

A sensitive framework has been developed for modelling young radiata pine (Pinus radiata D. Don) survival, its growth and size class distribution, from time of planting to age 5 or 6 years. The data and analysis refer to the Central North Island region of New Zealand. The survival function is derived from a Weibull probability density function, to reflect diminishing mortality with the passage of time in young stands. An anamorphic family of trends was used, as very little between-tree competition can be expected in young stands. An exponential height function was found to fit best the lower portion of its sigmoid form. The most appropriate basal area/ha exponential function included an allometric adjustment which resulted in compatible mean height and basal area/ha models. Each of these equations successfully represented the effects of several establishment practices by making coefficients linear functions of site factors, management activities and their interactions. Height and diameter distribution modelling techniques that ensured compatibility with stand values were employed to represent the effects of management practices on crop variation. Model parameters for this research were estimated using data from site preparation experiments in the region and were tested with some independent data sets.

• Mason, E-mail: em@mm.unknown
• Whyte, E-mail: aw@mm.unknown

The accompanying collective research report is the result of the research project in 1986–90 between the Finnish Academy and the former Soviet Academy of Sciences. The project was organized around common field work in Finland and in the former Soviet Union, and theoretical analyses of tree growth determining processes. Based on theoretical analyses, dynamic stand growth models were made and their parameters were determined utilizing the field results.

Annual cycle effects the tree growth. Our theoretical approach was based on adaptation to local climate conditions from Lapland to South Russia. The initiation of growth was described as a simple low and high temperature accumulation driven model. The model was linked with long-term temperature data.

Analysis of field measurements of CO₂ exchange showed that irradiance is the dominating factor causing variation in photosynthetic rate in natural conditions during summer. The penetration of irradiance into Scots pine (Pinus sylvestris L.) canopy is a complicated phenomenon. A moderately simple but balanced forest radiation regime sub-model was constructed.

The common field excursions in different geographical regions resulted in a lot of experimental data of regularities of woody structure. The water transport seems to be a good common factor to analyse these properties of tree structure. The produced theoretical and experimental material was utilized in the development of stand growth model that describes the growth and development of Scots pine stands in Finland and the former Soviet Union. The core of the model is carbon and nutrient balances. This means that carbon obtained in photosynthesis is consumed or growth and maintenance and nutrients are taken according to the metabolic need. Since the fundamental metabolic processes are the same in all locations, the same growth model structure can be applied in the large range of Scots pine. The model enables the analysis of geographical differences in the growth of Scots pine. The present approach enables utilization of structural and functional knowledge gained in places of intensive research, in the analysis of growth and development of any stand.

• Hari, E-mail: ph@mm.unknown
• Ross, E-mail: jr@mm.unknown
• Mecke, E-mail: ph@mm.unknown

The presence/absence data of 27 forest insect taxa (Retinia resinella, Formica spp., Pissodes spp., several scolytids) and recorded environmental variation were used to investigate the applicability of modelling insect occurrence based on satellite imagery. The sampling was based on 1,800 sample plots (25 m by 25 m) placed along the sides of 30 equilateral triangles (side 1 km) in a fragmented forest area (approximately 100 km²) in Evo, Southern Finland. The triangles were overlaid on land use maps interpreted from satellite images (Landsat TM 30 m multispectral scanner imagery 1991) and digitized geological maps. Insect occurrence was explained using either environmental variables measured in the field or those interpreted from the land use and geological maps. The fit of logistic regression models carried between species, possibly because some species may be associated with characteristics of single trees while other species with stand characteristics. The occurrence of certain insect species at least, especially those associated with Scots pine, could be relatively accurately assessed indirectly on the basis of satellite imagery and geological maps. Models based on both remotely sensed and geological data better predicted the distribution of forest insects except in the case of Xylechinus pilosus, Dryocetes sp. and Trypodendron lineatum, where the differences were relatively small in favour of the models based on field measurements. The number of species was related to habitat compartment size and distance from the habitat edge calculated from the land use maps, but logistic regressions suggested that other environmental variables in general masked the effect of these variables in species occurrence at the present scale.

• Väisänen, E-mail: rv@mm.unknown
• Heliövaara, E-mail: kh@mm.unknown

Diameter and volume increment as well as change in stem form of Scots pine (Pinus sylvestris L.) were analysed to predict tree increment variables. A stem curve set model is presented, based on prediction of the diameters at fixed angles in a polar coordinate system. This model consists of three elementary stem curves: 1) with bark, 2) without bark, and 3) without bark five years earlier. The differences between the elementary stem curves are the bark curve and the increment curve. The error variances at fixed angles and covariances between the fixed angles are divided into between-stand and within-stand components. Using principal components, the between-stand and within-stand covariance matrices are condensed separately for stem curve with bark, bark curve and increment curve. The two first principal components of the bark curve describe the vertical change in Scots pine bark type and the first principal component of the increment curve describes the increment rate. The elementary stem curves, bark curve and increment curve as well as corresponding stem volumes, bark volume and volume increment can be predicted for all trees in the stand with free choice of sample tree measurements. When only a few sample trees are measured, the stem curve set model gives significantly more accurate predictions of bark volume and volume increment for tally trees than does the volume method, which is based on the differences between two independent predictions of volume. The volume increment of tally trees can be predicted as reliably with as without measurement of sample tree height increment.

The PDF includes a summary in English.

• Ojansuu, E-mail: ro@mm.unknown

A theoretical framework to analyse the growth of Scots pine (Pinus sylvestris L.) is presented. Material exchange processes and internal processes that transport, transform and consume materials are identified as the components of growth. Hierarchical system is lined out. Momentary uptake of material at a single exchange site depends on the environmental condition next to the exchange site, the internal state of the biochemical system of the plant and the structure of the plant. The internal state depends on the exchange flows over period of time and the structural growth depends on the internal state. The response of these processes to the fluxes is controlled by the genetic composition of the plant.

The theoretical framework is formulated into a mathematical model. A concept of balanced internal state was applied to describe the poorly known internal processes. Internal substrate concentrations were assumed to remain constant but tissue-specific. A linear relationship between the quantity of foliage and wood cross-sectional area was assumed to describe balanced formation of structure. The exchange processes were thus described as a function of external conditions. The stand level interactions were derived from shading and effects of root density on nutrient uptake.

The approach was tested at different levels of hierarchy. Field measurements indicated that the hypothesis of the linear relationship described well the regularities between foliage and sapwood of a tree within a stand when measured at functionally corresponding height. There was considerable variation in the observed regularities in the range of geographic occurrence of Scots pine. Model simulations gave a realistic description of stand development in Southern Finland. The same model was also able to describe growth differences in Lapland after considering the effect of growing season length in the parameter values. Simulations to South Russia indicate stronger deviation from the observed patterns.

The simulations suggest interesting features of stand development. They indicate strong variability in the distribution of carbohydrates between tree parts during stand development. Internal circulation of nutrients and the reuse of the same transport structure by various needle generations had a strong influence on the simulation results.

The PDF includes a summary in Finnish.

• Nikinmaa, E-mail: en@mm.unknown

The effects and relative efficiency of alternative forest taxes are analysed theoretically. The Fisherian two-period model of consumption, savings and timber harvesting is extended by incorporating the management intensity decision and deriving the concept of long-run timber supply. The effects of lump-sum (site productivity), realized income (yield) and ad valorem property taxes on short-run timber supply, management intensity, and long-run timber supply are established. As the core of the study, the alternative taxes are compared in order to determine the appropriate forest tax regime in terms of production efficiency. The efficiency criterion generally requires that the excess burden of taxation at any given tax revenue should be kept to a minimum. The study distinguishes between an initially undistorted economy and an economy with pre-existing distortions due to capital income taxation (interest charge deductions). When the effects on forest management decisions of forest and capital income taxes are considered as a whole, a neutral forest taxation is no longer efficient. The non-timber benefits of a forest are incorporated to examine the robustness of the tax results with respect to the objective function. Finally, forest tax issues specific to Finland are considered, and administrational and equity aspects are discussed.

The PDF includes a summary in Finnish.

• Ovaskainen, E-mail: vo@mm.unknown

The study presents a growth and yield prediction model for a Pinus kesiya (Royle ex Gordon) stand by diameter classes. The material consists of temporary sample plots taken from a plantation inventory, of permanent sample plots established in commercial compartments and of an espacement trial. The mean basal area of the stand, variance and skewness of the diameter distribution are predicted. From these variables the parameters of the Weibull function are derived. Site class is assumed to be known or is calculated from measured information. Mortality is also predicted by estimating the number and mean size of dead trees. Thinnings are defined by the number of trees removed and by their relative size. If measured tree level data at the initial situation is available it can be utilized in the predictions, however, simulations can also be performed with stand level information. The minimum information needed for the prediction is planting density, site class as well as the times and removals of thinnings.

The calculations show that by decreasing the planting density of P. Kesiya from the present 1,330 stems/ha or by conducting early precommercial thinning both the relative and absolute amount of large sawlogs in the total production increase. An increase in the present planting density only slightly increases total yield. It is obvious that the presently recommended rotation of 25 years is too short for producing large sawlogs, especially on poor sites. This rotation period is suitable for small sawlog production while for pulpwood regimes shorter rotation periods can be used. If thinnings are done before the maximum current annual growth is reached stands will react well, but later on the ability to respond to thinnings decreases rapidly. Thinnings from below accelerates the production of large sawlogs more than thinning from above or systematic thinning. If all sawlog sizes are considered no great differences between thinning type exist. The study recommends different thinning regimes according to site class. Separate programs are recommended for the production of sawlogs and pulpwood.

The used thinning reaction model needs refinement and further studies with annual measured thinning trial material.

The PDF includes a summary in Finnish.

• Saramäki, E-mail: js@mm.unknown

Biogeographical patterns of the Scolytidae in Fennoscandia and Denmark, based on species incidence data from the approximately 70 km x 70 km quadrats (n = 221) used by Lekander et al. (1977), were classified to environmental variables using multivariate methods (two-way indicator species analysis, detrended correspondence analysis, canonical correspondence analysis).

The distributional patterns of scolytid species composition showed similar features to earlier presented zonations based on vegetation composition. One major difference, however, was that the region was more clearly divided in an east-west direction. Temperature variables associated with the location of the quadrat had the highest canonical coefficient values on the first axis of the CCA. Although these variables were the most important determinants of the biogeographical variation in the beetle species assemblages, annual precipitation and the distribution of Picea abies also improved the fit of the species data.

Samples with the most deviant rarity and typicality indices for the scolytid species assempblages in each quadrat were concentrated in several southern Scandinavian quadrats, in some quadrats in northern Sweden, and especially on the Swedish islands (Öland, Gotland, Gotska Sandön) in the Baltic Sea. The use of rarity indices which do not take the number of species per quadrat, also resulted high values for areas near Stockholm and Helsinki with well-known faunas. Methodological tests in which the real changes in the distribution of Ips acuminatus and I. amitinus were used as indicators showed that the currently available multivariate methods are sensitive to small faunal shifts even, and thus permit analysis of the fauna in relation to environmental changes. However, this requires more detailed monitoring of the species’ distributions over longer time spans.

Distribution of seven species (Scolytus intricatus, S. laevis, Hylurgops glabratus, Crypturgus cinereus, Pityogenes salasi, Ips typographus, and Cyleborus dispar) were predicted by logistic regression models using climatic variables. In spite of the deficiencies in the data and the environmental variables selected, the models were relatively good for several but not for all species. The potential effects of climate change on bark beetles are discussed.

The PDF includes a summary in Finnish.

• Heliövaara, E-mail: kh@mm.unknown
• Väisänen, E-mail: rv@mm.unknown
• Immonen, E-mail: ai@mm.unknown

Factors determining newsprint consumption in Finland in 1960–1986 were analysed. An econometric recursive multi-equation model describing the structure of the newspaper industry was formulated and estimated to obtain information on direct factors influencing newsprint demand. Short-term and long-term demand elasticities for newspapers and newspaper advertising were estimated.

The results indicate that the main factors affecting newsprint consumption are total circulation of newspapers, volume of newspaper advertising and the change in newsprint substance weight. Total newspaper circulation was found to depend on the rate of household formation and real household income changes. Demand for newspapers was shown to be price-inelastic. Structural analysis indicates that income elasticity of newspaper demand has increased slightly over time.

The volume of newspaper advertising was shown to affect newsprint consumption via the effects on pagination. Newspaper and television advertising were found to be independent of each other. The impact of the reduction in the basis weight was found to be substantial. The estimation of long-term elasticities of demand for newspapers and newspaper advertising using dynamic models revealed that demand rigidities exist.

The case study of Finland proposes three reasons why newsprint demand has not shown clear signs of reaching a saturation level. First, although population growth has stagnated in major consuming countries, the number of households has been increasing continuously. Second, income elasticity of newspaper demand does not show a declining trend. Third, the main driving force behind the buoyant demand is the resurgence of demand for newspaper as an advertising medium. In forecasting newsprint consumption, in addition to projections of economic growth, attention must be paid to the rate of household formation, the development of the advertising sector, the factors affecting competition between alternative media and the resulting media-mix in advertising, and changes in the substantial weight.

The PDF includes a summary in Finnish

• Katila, E-mail: mk@mm.unknown
• Riihinen, E-mail: pr@mm.unknown

At the beginning of the investigation period the total biomass of the Scots pine (Pinus sylvestris L.) stands on the ordinary sedge pine mire was 48 t/ha. The biomass of the mixed stands of Scots pine and birch (Betula pubescens Erhr.) on the herbrich sedge pine mire was 91 t/ha, out of which 60% was from pine. The biomass of the Norway spruce (Picea abies (L.) H. Karst.) on the Vaccinium-Myrtillus spruce mire was 148 t/ha. The average annual net increment of the stand biomass was 5.8 t/ha in the unfertilized pine stand and 6.7 t/ha in the NPK and micronutrient fertilized one during the six-year investigation period. The corresponding figures in the mixed stand were 7.2 t/ha and 7.6 t/ha. The net increment of the biomass in the unfertilized spruce stand was 6.9 t/ha and in the fertilized 8.4 t/ha. A considerable proportion of the net increment was lost to the ground as litter in all stands.

The nitrogen, phosphorus, potassium, magnesium, iron, manganese, zinc, copper and boron cycles were investigated. The annual nitrogen uptake from the soil was 26–42 kg/ha, that of phosphorus 2.5–3.4 kg/ha, potassium 4.5–12 kg/ha, calcium 12–29 kg/ha, magnesium 2–4 kg/ha, iron 1.4–6.6 kg/ha, manganese less than 2 kg/ha and the other nutrients only some grams. Only part of the fertilized nutrients was fixed in the stand.

The PDF includes a summary in Finnish.

• Finér, E-mail: lf@mm.unknown

The relationship between canopy structure and photosynthetic radiation regime are studied in a theoretical basis. In modelling the canopy structure, a statistical approach is applied and the radiation field inside a stand is described in terms of random variables and their distribution. A comparison is made between horizontally homogenous stands and grouped forest stands in order to assess the influence of grouping of foliage on the irradiance distribution in a forest stand. Results show that grouping considerably reduces the interception of radiation and causes a large spatial variation. In coniferous stands the grouping of needles into shoots and the effect of penumbra are shown to have an important influence on the distribution of radiation on the needle area.

The PDF includes a summary in Finnish.

• Oker-Blom, E-mail: po@mm.unknown

Taper curve models based on simultaneous equations were derived. The data consisted of 492 Scots pines (Pinus sylvestris L.) from Southern Finland. Two systems of simultaneous equations were constructed, one without the crown ratio and the other with the crown ratio as an exogenous variable. The endogenous variables consisted of 24 relative-height diameters and the height of the tree. The parameters of the model were derived by the ordinary least squares method.

In most applications, the height of the tree was exogenised. The logarithmically linear relationships between the relative-height diameters were utilized in the solution algorithm. The algorithm included both standard matrix operations and an iterative part in which the taper curve was fitted to any measured diameters by the natural cubic spline interpolation formula.

The models were applied to the derivation of taper curves, stem volumes, timber assortment percentages, and stem values. An experiment was also made to derive diameter and height increments from the taper curve model.

The reliability of the models was tested on the original data.

The PDF includes a summary in Finnish.

• Kilkki, E-mail: pk@mm.unknown
• Varmola, E-mail: mv@mm.unknown

A long-term timber production model was developed. The model is based upon numerical simulation and it is viewed only as a means of providing the decision-maker with values of the predicting variables in his utility function. Special attention was paid to the development of automatic cutting decision rules. The model was applied to the area of 2,752,000 hectares of forest land in Central Finland. The measurement data were extracted from the Sixth National Forest Inventory, which was made in 1973. Utilities from a hypothetical utility function were attained to a number of feasible timber production policies. The Bayes and maximin criteria were employed to evaluate these policies.

The PDF includes a summary in Finnish.

• Kilkki, E-mail: pk@mm.unknown
• Pökälä, E-mail: rp@mm.unknown

With developing possibilities to analyse data automatically there is a need to develop the mathematical and statistical practices for calculations. The article presents the basis about the growth of trees and the existing models of growth, the basics on growth functions,   and the construction of a regression model to analysis the growth. The theoretical model development has been tested with three existing data sets.

The analysis of growth should be considered with dynamic models. The model need to take into account various aspects and growth factors. The model should have practical implications.

The PDF contains a summary in German. 

• Kangas, E-mail: yk@mm.unknown

• Peltoniemi, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: mikko.peltoniemi@metla.fi
• Thürig, Swiss Federal Institute for Forest, Snow and Landscape Research (WSL), Birmensdorf, Switzerland; European Forest Institute, Joensuu, Finland E-mail: et@nn.ch
• Ogle, Natural Resources Ecology Laboratory, Colorado State University, Fort Collins, USA E-mail: so@nn.us
• Palosuo, European Forest Institute, Joensuu, Finland E-mail: tp@nn.fi
• Schrumpf, Max-Planck-Institute for Biogeochemistry, Jena, Germany E-mail: ms@nn.de
• Wutzler, Max-Planck-Institute for Biogeochemistry, Jena, Germany E-mail: tw@nn.de
• Butterbach-Bahl, Institute for Meteorology and Climate Research, Forschungszentrum Karlsruhe GmbH, Garmisch-Partenkirchen, Germany E-mail: kbb@nn.de
• Chertov, St. Petersburg State University, St. Petersburg-Peterhof, Russia E-mail: oc@nn.ru
• Komarov, Institute of Physicochemical and Biological Problems in Soil Science of Russian Academy of Sciences, Pushchino, Russia E-mail: ak@nn.ru
• Mikhailov, Institute of Physicochemical and Biological Problems in Soil Science of Russian Academy of Sciences, Pushchino, Russia E-mail: am@nn.ru
• Gärdenäs, Dept. of Soil Sciences, Swedish University of Agricultural Sciences, Uppsala, Sweden E-mail: ag@nn.se
• Perry, USDA Forest Service, Northern Research Station, St. Paul, MN USA E-mail: cp@nn.us
• Liski, Finnish Environment Institute, Helsinki, Finland E-mail: jl@nn.fi
• Smith, School of Biological Sciences, University of Aberdeen, Aberdeen, UK E-mail: ps@nn.uk
• Mäkipää, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: raisa.makipaa@metla.fi

• López, Inland Norway University of Applied Sciences, Faculty of Applied Ecology, Agricultural Sciences and Biotechnology, P.O. Box 2400, Koppang, Norway https://orcid.org/0009-0006-6860-3408 E-mail: lucas.lopez@inn.no
• Sjølie, Inland Norway University of Applied Sciences, Faculty of Applied Ecology, Agricultural Sciences and Biotechnology, P.O. Box 2400, Koppang, Norway https://orcid.org/0000-0001-8099-3521 E-mail: hanne.sjolie@inn.no
• Nabhani, Inland Norway University of Applied Sciences, Faculty of Applied Ecology, Agricultural Sciences and Biotechnology, P.O. Box 2400, Koppang, Norway E-mail: abbas.nabhani@inn.no
• Aguilar, Swedish University of Agricultural Sciences, Department of Forest Economics, SE-901 83 Umeå, Sweden E-mail: francisco.aguilar@slu.se

Mixed forests are known for their ability to provide a wide range of ecosystem services. Such forests have higher biodiversity compared to monocultures, are resilient against disturbances and may mitigate the effects of climate change. Despite well-known benefits, there is still little information on how these forests should be established and managed. The aim of this study was to describe the early growth dynamics of current boreal young mixed stands of planted Norway spruces (Picea abies (L.) Karst.) and naturally regenerated birches (Betula spp.). We collected data from 9  stands planted for spruce 8–14 years ago in Southern and Central Finland. Stem analysis was conducted to 144 spruces and to 144 birches to determine previous growth. We modelled the height and diameter development of individual trees in relation to tree age at stump height using non-linear mixed Chapman-Richards model. There were no significant differences between spruce and seed-origin birch in diameter growth at stump height, but the initial height increments of natural birches were larger than those of planted spruces. However, planted spruces were able to keep up with the height development of birches, if spruces received a head start over naturally regenerated seed-origin birch for two growing seasons. Thus, naturally regenerated birch admixture can be utilized to establish single-storied spruce-birch mixtures, and the admixture should be retained during the early cleaning of planted spruce stands.

• Männistö, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, FI-00790 Helsinki, Finland https://orcid.org/0009-0009-0213-1972 E-mail: lauri.mannisto@luke.fi
• Miina, Natural Resources Institute Finland (Luke), Natural resources, Yliopistokatu 6 B, FI-80100 Joensuu, Finland https://orcid.org/0000-0002-8639-4383 E-mail: jari.miina@luke.fi
• Huuskonen, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, FI-00790 Helsinki, Finland https://orcid.org/0000-0001-8630-3982 E-mail: saija.huuskonen@luke.fi

• Jääskeläinen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0009-0004-4127-7863 E-mail: johanna.jaaskelainen@uef.fi
• Korhonen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0000-0002-9352-0114 E-mail: lauri.korhonen@uef.fi
• Kukkonen, Natural Resources Institute Finland, Yliopistokatu 6 B, FI-80100 Joensuu, Finland https://orcid.org/0000-0003-4206-1680 E-mail: mikko.kukkonen@luke.fi
• Packalen, Natural Resources Institute Finland, Latokartanonkaari 9, FI-00790 Helsinki, Finland https://orcid.org/0000-0003-1804-0011 E-mail: petteri.packalen@luke.fi
• Maltamo, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0000-0002-9904-3371 E-mail: matti.maltamo@uef.fi

We developed tree level biomass (dry weight) models for Norway spruce (Picea abies [L.] H. Karst.), silver birch (Betula pendula Roth), rowan (Sorbus aucuparia L.) and aspen (Populus tremula L.) growing in young spruce dominated seedling stands with high mixture of broadleaves. The study material was collected from three planted Norway spruce seedling stands located on mineral soil in southern Finland. Biomass models were estimated by individual tree component (stem, living branches, foliage, stump, and roots with diameter of 2 mm) by using a multi-response approach (seemingly unrelated regression), which estimated the parameters of the sub-models (tree component) simultaneously. Even though the application and generalization of the developed models can be restricted by the limited material, they provide new information of seedling biomass allocation and more reliable biomass predictions for spruce and birch growing in young seedling stand compared with those of the commonly applied biomass models (Repola 2008, 2009) in Finland. Repola’s models (2008, 2009) tended to produce biased predictions for crown and below-ground biomasses of seedlings by allocating too much biomass to roots and too little to needle and branches. In addition, this study provides biomass models for aspen and rowan, which were not previously available.

• Repola, Natural Resources Institute Finland (Luke), Ounasjoentie 6, FI-96200 Rovaniemi, Finland https://orcid.org/0000-0001-7086-0549 E-mail: jaakko.repola@luke.fi
• Luoranen, Natural Resources Institute Finland (Luke), Juntintie 154, FI-77600 Suonenjoki, Finland https://orcid.org/0000-0002-6970-2030 E-mail: jaana.luoranen@luke.fi
• Huuskonen, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland https://orcid.org/0000-0001-8630-3982 E-mail: saija.huuskonen@luke.fi
• Peltoniemi, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland https://orcid.org/0000-0003-2028-6969 E-mail: mikko.peltoniemi@luke.fi
• Väänänen, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: paivi.vaananen@luke.fi
• Uotila, Natural Resources Institute Finland (Luke), Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: karri.uotila@luke.fi

Forest stands, crucial for inventory, planning, and management, traditionally rely on time-consuming visual analysis by forest managers. To enhance efficiency, there is a growing need for automated methods that take into account essential forest attributes. In response, we propose a novel approach utilizing airborne Light Detection and Ranging (LiDAR) and hyperspectral data for automated forest stand delineation. Our approach initiates with over-segmentation of the Canopy Height Model (CHM), followed by attribute calculation for each segment using both CHM and hyperspectral data. Two rules are applied to merge homogeneous segments and eliminate others based on calculated attributes. The effectiveness of our method was validated using three types of reference forest stands with two indices: the explained variance (R²) and Intersection over Union (IoU). Results from our study demonstrated notable accuracy, with a R² of 97.35% and 97.86% for mean tree height and mean diameter at breast height (DBH), respectively. The R² for mean canopy height is 81.80%, outperforming manual delineation by 7.31% and multi-scale segmentation results by 2.13%. Furthermore, our approach achieved high IoU values, which indicates a strong spatial agreement with manually delineated forest stands and leading to fewer manual adjustments when applied directly to forest management. In conclusion, our forest stand delineation method enhances both internal consistency and spatial accuracy. This method contributes to improving practical performance and forest management efficiency.

• Xiong, Institute of Forest Resource Information Techniques, Chinese Academy of Forestry, Beijing 100091, China; Key Laboratory of Forestry Remote Sensing and Information System, National Forestry and Grassland Administration, Beijing 100091, China; School of Geospatial Engineering and Science, Sun Yat-sen University, Zhuhai 519082, China https://orcid.org/0000-0003-4432-2485 E-mail: xiongh29@mail2.sysu.edu.cn
• Pang, Institute of Forest Resource Information Techniques, Chinese Academy of Forestry, Beijing 100091, China; Key Laboratory of Forestry Remote Sensing and Information System, National Forestry and Grassland Administration, Beijing 100091, China https://orcid.org/0000-0002-9760-6580 E-mail: pangy@ifrit.ac.cn
• Jia, Institute of Forest Resource Information Techniques, Chinese Academy of Forestry, Beijing 100091, China; Key Laboratory of Forestry Remote Sensing and Information System, National Forestry and Grassland Administration, Beijing 100091, China E-mail: jiawen@ifrit.ac.cn
• Bai, Institute of Forest Resource Information Techniques, Chinese Academy of Forestry, Beijing 100091, China; Key Laboratory of Forestry Remote Sensing and Information System, National Forestry and Grassland Administration, Beijing 100091, China E-mail: baiyu9224@163.com

• Gartner, University of Natural Resources and Life Sciences, Department of Economics and Social Sciences, Institute of Production and Logistics, Feistmantelstrasse 4, 1180 Vienna, Austria https://orcid.org/0000-0001-8547-718X E-mail: maria.gartner@boku.ac.at
• Kaltenbrunner, improvem GmbH, Holzinnovationszentrum 1a, 8740 Zeltweg, Austria https://orcid.org/0000-0002-1178-0087 E-mail: matthias.kaltenbrunner@improvem.at
• Gronalt, University of Natural Resources and Life Sciences, Department of Economics and Social Sciences, Institute of Production and Logistics, Feistmantelstrasse 4, 1180 Vienna, Austria https://orcid.org/0000-0003-0944-4911 E-mail: manfred.gronalt@boku.ac.at

• Böhm, Department of Forest and Soil Sciences, Institute of Forest Engineering, University of Natural Resources and Life Sciences, Vienna (BOKU), Peter-Jordan-Strasse 82, 1190 Vienna, Austria https://orcid.org/0000-0001-7803-6618 E-mail: stephan.boehm@boku.ac.at
• Baier, Department of Forest and Soil Sciences, Institute of Forest Entomology, Forest Pathology and Forest Protection, University of Natural Resources and Life Sciences, Vienna (BOKU), Peter-Jordan-Strasse 82, 1190 Vienna, Austria https://orcid.org/0000-0002-1029-5637 E-mail: peter.baier@boku.ac.at
• Kirisits, Department of Forest and Soil Sciences, Institute of Forest Entomology, Forest Pathology and Forest Protection, University of Natural Resources and Life Sciences, Vienna (BOKU), Peter-Jordan-Strasse 82, 1190 Vienna, Austria https://orcid.org/0000-0002-9918-3593 E-mail: thomas.kirisits@boku.ac.at
• Kanzian, Department of Forest and Soil Sciences, Institute of Forest Engineering, University of Natural Resources and Life Sciences, Vienna (BOKU), Peter-Jordan-Strasse 82, 1190 Vienna, Austria https://orcid.org/0000-0002-1198-9788 E-mail: christian.kanzian@boku.ac.at

• Vauhkonen, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, FI-80101 Joensuu, Finland; University of Helsinki, Department of Forest Sciences, Latokartanonkaari 7, FI-00014 Helsingin yliopisto, Finland E-mail: jari.vauhkonen@uef.fi
• Matala, Natural Resources Institute Finland (Luke), Natural resources, Yliopistokatu 6 B, FI-80100 Joensuu, Finland E-mail: juho.matala@luke.fi
• Nikula, Natural Resources Institute Finland (Luke), Natural resources, Ounasjoentie 6, FI-96200 Rovaniemi, Finland https://orcid.org/0000-0001-8372-8440 E-mail: ari.nikula@luke.fi

• Kangas, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Yliopistokatu 6, 80101 Joensuu, Finland https://orcid.org/0000-0002-8637-5668 E-mail: annika.kangas@luke.fi
• Myllymäki, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, FI-00790 Helsinki, Finland https://orcid.org/0000-0002-2713-7088 E-mail: mari.myllymaki@luke.fi
• Mehtätalo, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Yliopistokatu 6, 80101 Joensuu, Finland https://orcid.org/0000-0002-8128-0598 E-mail: lauri.mehtatalo@luke.fi

• Jåstad, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway https://orcid.org/0000-0002-1089-0284 E-mail: eirik.jastad@nmbu.no
• Nagel, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway https://orcid.org/0000-0002-3171-0262 E-mail: niels.oliver.nagel@nmbu.no
• Hu, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway https://orcid.org/0000-0003-0001-5993 E-mail: junhui.hu@nmbu.no
• Rørstad, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: per.kristian.rorstad@nmbu.no

• Korpela, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland 0000-0002-1665-3984 E-mail: ilkka.korpela@helsinki.fi
• Polvivaara, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: –
• Papunen, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland 0000-0001-5383-4314 E-mail: saija.papunen@outlook.com
• Jaakkola, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: laura.jaakkola@helsinki.fi
• Tienaho, University of Eastern Finland, Faculty of Science and Forestry, P.O. Box 111, FI-80101 Joensuu, Finland 0000-0002-6574-5797 E-mail: noora.tienaho@uef.fi
• Uotila, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: johannes.uotila@helsinki.fi
• Puputti, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland 0000-0003-1972-1636 E-mail: tuomas.puputti@helsinki.fi
• Flyktman, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland 0000-0002-5235-317X E-mail: aleksi.flyktman@helsinki.fi

Norway’s most common tree species, Picea abies (L.) Karst. (Norway spruce), is often infected with Heterobasidion parviporum Niemelä & Korhonen and Heterobasidion annosum (Fr.) Bref.. Because Pinus sylvestris L. (Scots pine) is less susceptible to rot, it is worth considering if converting rot-infested spruce stands to pine improves economic performance. We examined the economically optimal choice between planting Norway spruce and Scots pine for previously spruce-dominated clear-cut sites of different site indexes with initial rot levels varying from 0% to 100% of stumps on the site. While it is optimal to continue to plant Norway spruce in regions with low rot levels, shifting to Scots pine pays off when rot levels get higher. The threshold rot level for changing from Norway spruce to Scots pine increases with the site index. We present a case study demonstrating a practical method (“Precision forestry”) for determining the tree species in a stand at the pixel level when the stand is heterogeneous both in site indexes and rot levels. This method is consistent with the concept of Precision forestry, which aims to plan and execute site-specific forest management activities to improve the quality of wood products while minimising waste, increasing profits, and maintaining environmental quality. The material for the study includes data on rot levels and site indexes in 71 clear-cut stands. Compared to planting the entire stand with a single species, pixel-level optimised species selection increases the net present value in almost every stand, with average increase of approximately 6%.

• Aza, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, PO Box 5003, NO-1432, Ås, Norway https://orcid.org/0000-0002-6416-6697 E-mail: anfe@nmbu.no
• Kallio, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, PO Box 5003, NO-1432, Ås, Norway E-mail: maarit.kallio@nmbu.no
• Pukkala, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: timo.pukkala@uef.fi
• Hietala, Norwegian Institute of Bioeconomy Research, PO Box 115, NO-1431 Ås, Norway E-mail: ari.hietala@nibio.no
• Gobakken, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, PO Box 5003, NO-1432, Ås, Norway E-mail: terje.gobakken@nmbu.no
• Astrup, Norwegian Institute of Bioeconomy Research, PO Box 115, NO-1431 Ås, Norway E-mail: rasmus.astrup@nibio.no

The widespread cork oak (Quercus suber L.) mortality and reduced afforestation /regeneration are causing an overall reduction in cork production. To enhance trees’ growth and vitality, afforestation techniques using fertirrigation were tested. The main objective was the promotion of trees’ growth on new dense plantations using minimum water requirements until reaching productive forests. The experimental plot – Irricork – was installed in 2017 in a ≈1 ha stand with 14 years’ age cork oaks summer-fertirrigated since plantation. Four fertirrigation treatments were applied during fertirrigation campaigns. Radial growth, meteorological parameters and fertirrigation volume were measured every 15–30 days over four years. It was observed that weather, tree size, debarking and trees’ intra-competition had a significant effect on radial increments. Fertirrigation significantly enhanced growth during summer drought and decoupled increments from air vapor pressure deficit constraints. There was a linear relationship between trees’ radial increments and fertirrigation volume up to 140 m³ week^–1. Above this value, increments were smoother. In conclusion, summer fertirrigation of 140 m³ week^–1 efficiently enhanced the radial growth of trees with 50–75 circumference at breast height, under the particular edaphoclimatic conditions of the stand. This study showed to be, therefore, promising in the use of efficient fertirrigation the enhance cork oaks’ radial growth.

• Camilo-Alves, MED – Mediterranean Institute for Agriculture, Environment and Development & CHANGE – Global Change and Sustainability Institute, Institute for Advanced Studies and Research, University of Evora, Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal https://orcid.org/0000-0001-5156-172X E-mail: calves@uevora.pt
• Nunes, Department of Plant Science, School of Science and Technology, University of Evora, Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal https://orcid.org/0000-0002-6144-3484 E-mail: jain@uevora.pt
• Poeiras, MED – Mediterranean Institute for Agriculture, Environment and Development & CHANGE – Global Change and Sustainability Institute, Institute for Advanced Studies and Research, University of Evora, Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal https://orcid.org/0000-0002-6049-807X E-mail: apcp@uevora.pt
• Ribeiro, Department of Plant Science, School of Science and Technology, University of Evora, Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal https://orcid.org/0000-0001-7242-5866 E-mail: jmrpr@uevora.pt
• Dinis, https://orcid.org/0000-0001-6984-1033 E-mail: dinis.cati@gmail.com
• Barroso, MED – Mediterranean Institute for Agriculture, Environment and Development & CHANGE – Global Change and Sustainability Institute, and Department of Plant Science, School of Science and Technology, University of Evora, Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal https://orcid.org/0000-0002-0160-3845 E-mail: jmmb@uevora.pt
• Vaz, MED – Mediterranean Institute for Agriculture, Environment and Development & CHANGE – Global Change and Sustainability Institute, and Department of Biology, School of Science and Technology, University of Evora. Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal https://orcid.org/0000-0002-3368-757X E-mail: mvaz@uevora.pt
• Almeida-Ribeiro, ICT – Institute of Earth Sciences and Department of Plant Science, School of Science and Technology, University of Evora, Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal https://orcid.org/0000-0002-0160-3845 E-mail: nmcar@uevora.pt

Terrestrial laser scanning (TLS) has been applied to estimate forest wood volume based on detailed 3D tree reconstructions from point cloud data. However, sources of uncertainties in the point cloud data (alignment and scattering errors, occlusion, foliage...) and the reconstruction algorithm type and parameterisation are known to affect the reconstruction, especially around finer branches. To better understand the impacts of these uncertainties on the accuracy of TLS-derived woody volume, high-quality TLS scans were collected in leaf-off conditions prior to destructive harvesting of two forest-grown common ash trees (Fraxinus excelsior L.; diameter at breast height ~28 cm, woody volume of 732 and 868 L). We manually measured branch diameters at 265 locations in these trees. Estimates of branch diameters and tree volume from Quantitative Structure Models (QSM) were compared with these manual measurements. The accuracy of QSM branch diameter estimates decreased with smaller branch diameters. Tree woody volume was overestimated (+336 L and +392 L) in both trees. Branches measuring < 5 cm in diameter accounted for 80% and 83% of this overestimation respectively. Filtering for scattering errors or improved coregistration approximately halved the overestimation. Range filtering and modified scanning layouts had mixed effects. The small branch overestimations originated primarily in limitations in scanner characteristics and coregistration errors rather than suboptimal QSM parameterisation. For TLS-derived estimates of tree volume, a higher quality point cloud allows smaller branches to be accurately reconstructed. Additional experiments need to elucidate if these results can be generalised beyond the setup of this study.

• Demol, CAVElab – Computational and Applied Vegetation Ecology, Department of Environment, Faculty of Bioscience Engineering, Ghent University, Coupure Links 653, B-9000 Ghent, Belgium; PLECO – Plants and Ecosystems, Faculty of Science, Antwerp University, Universiteitsplein 1, B-2610 Wilrijk, Belgium https://orcid.org/0000-0002-5492-2874 E-mail: miro.demol@ugent.be
• Wilkes, UCL Department of Geography, Gower Street, London WC1E 6BT, UK; NERC National Centre for Earth Observation (NCEO), UK https://orcid.org/0000-0001-6048-536X E-mail: p.wilkes@ucl.ac.uk
• Raumonen, Mathematics, Tampere University, FI-33101 Tampere, Finland https://orcid.org/0000-0001-5471-0970 E-mail: pasi.raumonen@tuni.fi
• Krishna Moorthy, CAVElab – Computational and Applied Vegetation Ecology, Department of Environment, Faculty of Bioscience Engineering, Ghent University, Coupure Links 653, B-9000 Ghent, Belgium https://orcid.org/0000-0002-6838-2880 E-mail: Sruthi.KrishnaMoorthyParvathi@ugent.be
• Calders, CAVElab – Computational and Applied Vegetation Ecology, Department of Environment, Faculty of Bioscience Engineering, Ghent University, Coupure Links 653, B-9000 Ghent, Belgium https://orcid.org/0000-0002-4562-2538 E-mail: kim.calders@ugent.be
• Gielen, PLECO – Plants and Ecosystems, Faculty of Science, Antwerp University, Universiteitsplein 1, B-2610 Wilrijk, Belgium https://orcid.org/0000-0002-4890-3060 E-mail: bert.gielen@uantwerpen.be
• Verbeeck, CAVElab – Computational and Applied Vegetation Ecology, Department of Environment, Faculty of Bioscience Engineering, Ghent University, Coupure Links 653, B-9000 Ghent, Belgium https://orcid.org/0000-0003-1490-0168 E-mail: hans.verbeeck@ugent.be

For constructing growth and yield models the concept of site index as measure of productivity is crucial. Here, we use nonlinear mixed-effects models (NLME) with random individual effects and nonlinear models with dummy variables as fixed individual effects (NLFE) to fit mechanistic growth functions to stem analysis data of the economically most important tree species in Zhongtiaoshan forest region, China. The Richards and Lundqvist function are formulated into five dynamic equations (R1, R2, L1, L2 and L3) applying the generalized algebraic difference approach (GADA), which inherit polymorphism, varying asymptotes and base-age invariance. According to Akaike information criterion the R1 model as NLFE fits height growth data of Pinus tabuliformis Carrière, Pinus armandii Franch., Quercus liaotungensis Koidz., Quercus aliena Blume and Betula platyphylla Sukaczev best, while for Quercus variabilis Blume R2 as NLFE fits height growth data best. For Larix principis-rupprechtii Mayr L1 as NLME has been selected as best model, as R1 and R2 both as NLFE and NLME are not extrapolating the comparably short length of height growth data well enough. However, according to the root mean square error and bias differences between model fits of both the selected equation and the chosen model fitting approach are not so clear. Presented families of height growth curves serve as planning tools to identify site index and therefore assess productivity of forest stands in the studied region. A direct comparison of the productivity of forest stands of the same tree species is possible due to base-age invariance of the selected models.

• Sprengel, Chair of Forest Growth and Dendroecology, Albert-Ludwigs-University Freiburg, Tennenbacher Str. 4, 79106 Freiburg, Germany https://orcid.org/0000-0002-6332-7911 E-mail: lars.sprengel@iww.uni-freiburg.de
• Spiecker, Chair of Forest Growth and Dendroecology, Albert-Ludwigs-University Freiburg, Tennenbacher Str. 4, 79106 Freiburg, Germany E-mail: instww@uni-freiburg.de
• Wu, Research Institute of Forest Policy and Information, Chinese Academy of Forestry, No. 1 Dongxiaofu, Haidian District, Beijing 100091, China E-mail: shuirongwu@126.com

Hybrid aspen (Populus tremula L. × P. tremuloides Michx.) is known with outstanding growth rate and some favourable wood characteristics, but models for stand management have not yet been prepared in northern Europe. This study introduces methods and models to predict tree dimensions, diameter at breast height (dbh) and tree height for a hybrid aspen plantation using data from repeatedly measured permanent sample plots established in clonal plantations in southern Finland. Dbh distributions using parameter recovery method for the Weibull function was used with Näslund’s height curve to model tree heights. According to the goodness-of-fit statistics of Kolmogorov-Smirnov and the Error Index, the arithmetic mean diameter (D) and basal area-weighted mean diameter (DG) provided more stable parameter recovery for the Weibull distribution than the median diameter (DM) and basal area-weighted median diameter (DGM), while DG showed the best overall fit. Thus, Näslund’s height curve was modelled using DG with Lorey’s height (HG), age, basal area (BA), and tree dbh (Model 1). Also, Model 2 was tested using all predictors of Model 1 with the number of trees per ha (TPH). All predictors were shown to be significant in both Models, showing slightly different behaviour. Model 1 was sensitive to the mean characteristics, DG and HG, while Model 2 was sensitive to stand density, including both BA and TPH as predictors. Model 1 was considered more reasonable to apply based on our results. Consequently, the parameter recovery method using DG and Näslund’s models were applicable for predicting tree diameter and height.

• Lee, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, FI-00790 Helsinki, Finland https://orcid.org/0000-0003-1586-9385 E-mail: daesung.lee@luke.fi
• Siipilehto, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: jouni.siipilehto@luke.fi
• Hynynen, Natural Resources Institute Finland (Luke), Natural resources, Vipusenkuja 5, FI-57200 Savonlinna, Finland https://orcid.org/0000-0002-9132-8612 E-mail: jari.hynynen@luke.fi

Bilberry (Vaccinium myrtillus L.) and lingonberry (V. vitis-idaea L.) can be a part of healthy diet and are important for many animals. Two approaches are described to assessing their vegetation cover and berry yield via national forest inventory (NFI) observations. The aim was to provide estimates and predictions of the abundance and yield of the species at regional and national levels in Finland and Sweden. In Finland, the model-based predictions are used in evaluating the impacts of cutting intensity on forest berries needed in forest-related decision making. In Sweden, seasonal inventory-based estimates are used to evaluate the annual national and regional berry yields, and in a forecasting system aimed at large public and berry enterprises. Based on the NFI sample plots measured between 2014 and 2018, the total annual yields are estimated to be 208 Mkg of bilberry and 246 Mkg of lingonberry on productive forest land (increment at least 1 m³ ha^–1 year^–1) in Finland, and 336 and 382 Mkg respectively in Sweden (average of NFI inventories in 2015–2019). The predicted development of berry yields is related to the intensity of cuttings in alternative forest management scenarios: lower removals favoured bilberry, and higher removals lingonberry. The model-based method describes the effects of stand development and management on berry yields, whereas the inventory-based method can calibrate seasonal estimates through field observations. In providing spatially and timely more accurate information concerning seasonal berry yields, an assessment of berry yields should involve the elements of both inventory-based and model-based approaches described in this study.

• Miina, Natural Resources Institute Finland (Luke), Yliopistokatu 6 B, FI-80100 Joensuu, Finland https://orcid.org/0000-0002-8639-4383 E-mail: jari.miina@luke.fi
• Bohlin, Swedish University of Agricultural sciences (SLU), Department of Forest Resource Management, Skogsmarksgränd, S-901 83 Umeå, Sweden E-mail: inka.bohlin@slu.se
• Lind, Swedish University of Agricultural sciences (SLU), Department of Forest Resource Management, Skogsmarksgränd, S-901 83 Umeå, Sweden E-mail: torgny.lind@slu.se
• Dahlgren, Swedish University of Agricultural sciences (SLU), Department of Forest Resource Management, Skogsmarksgränd, S-901 83 Umeå, Sweden E-mail: jonas.dahlgren@slu.se
• Härkönen, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: kari.harkonen@luke.fi
• Packalen, Natural Resources Institute Finland (Luke), Yliopistokatu 6 B, FI-80100 Joensuu, Finland; Ministry of Agriculture and Forestry of Finland, P.O. Box 30, FI-00023 Government, Finland E-mail: tuula.packalen@mmm.fi
• Tolvanen, Natural Resources Institute Finland (Luke), Paavo Havaksentie 3, FI-90014 University of Oulu, Finland E-mail: anne.tolvanen@luke.fi

Studies of intra-annual growth are particularly useful for understanding tree growth because of their high temporal resolution. This study was performed in Austria and included hourly band dendrometer data of 244 annual tree recordings from six tree species (Picea abies (L.) Karst., Pinus sylvestris L., Larix decidua Mill., Abies alba Mill., Fagus sylvatica L., Quercus spp. (Quercus petraea (Matt.) Liebl., Quercus robur L.) sampled on five sites with contrasting site conditions in pure and mixed stands and on trees of different social position. Measurements encompassed 1–7 years. Cumulative diameter increment was modelled by logistic mixed-effects models with random effects at the tree and year level. The results showed large differences in seasonal growth patterns between sites, with a clearly shorter growing season at the drier sites. Species specific response on dry sites could be linked to drought characteristics, whereas response on more humid sites was related to light requirements or successional status. The deciduous trees showed earlier growth culmination and shorter growing periods than the evergreen species. Individual tree growth of Quercus spp., P. abies, and F. sylvatica was positively affected by mixture whereas L. decidua, P. sylvestris and A. alba showed no or adverse mixture effects. Mixture effects differed between years and social position. Furthermore, increment culmination was earlier in mixed stands, but shifts were minor. Tree growth differed by social position with dominant trees showing the largest increment and the longest growth duration, with shifts in tree growth patterns due to social position being as large as those between different sites.

• Strieder, Boku, University of Natural Resources and Life Sciences Vienna, Department of Forest- and Soil Sciences, Institute of Forest Growth, Peter-Jordan-Str. 82, A-1190 Vienna, Austria https://orcid.org/0000-0001-6398-8536 E-mail: emanuel.strieder@students.boku.ac.at
• Vospernik, Boku, University of Natural Resources and Life Sciences Vienna, Department of Forest- and Soil Sciences, Institute of Forest Growth, Peter-Jordan-Str. 82, A-1190 Vienna, Austria http://orcid.org/0000-0002-4201-6444 E-mail: sonja.vospernik@boku.ac.at

Dalbergia latifolia Roxb., commonly known as rosewood, is one of the highly valuable tropical timber species of Nepal. The tree species was widely distributed in the past, however, over-exploitation of natural habitat, deforestation, forest conversion for agriculture, illegal logging and the invasion of alien species resulted in the classification of this species as vulnerable by the IUCN (International Union for Conservation of Nature) category. So, the prediction of habitat suitability and potential distribution of the species is required to develop restoration mechanisms and conservation interventions. In this study, we modelled the suitable habitat of D. latifolia over the entire possible range of Nepal using a Maxent model. We compiled 23 environmental variables (19 bioclimatic, 3 topographic and a vegetative layer), however, only 12 least correlated variables along with 43 spatially representative presence locations were retained for model prediction. We used a receiver operating characteristic (ROC) curve to assess the model’s performance and a Jackknife procedure to evaluate the relative importance of predictor variables. The model was statistically significant with an area under the curve (AUC) value of 0.969. The internal Jackknife test indicated that elevation was the most important variable for the model prediction with 71.3% contribution followed by mean temperature of driest quarter (9.8%). The most (>0.6) suitable habitat for the D. latifolia was 235 484 hectares with large sections of area in two provinces whereas, the western most provinces were not suitable for D. latifolia as per Maxent model. The information presented here can provide a framework for nature conservation planning, monitoring and habitat management of this rare and endangered species.

• Mahatara, Forest Research and Training Centre, Government of Nepal, P.O. Box 3339, Babarmahal, Kathmandu 44600, Nepal E-mail: honeystar73@gmail.com
• Acharya, Forest Research and Training Centre, Government of Nepal, P.O. Box 3339, Babarmahal, Kathmandu 44600, Nepal E-mail: acharya.amulkumar@gmail.com
• Dhakal, Forest Research and Training Centre, Government of Nepal, P.O. Box 3339, Babarmahal, Kathmandu 44600, Nepal E-mail: dhakalbp.shorea@gmail.com
• Sharma, Forest Research and Training Centre, Government of Nepal, P.O. Box 3339, Babarmahal, Kathmandu 44600, Nepal E-mail: dipeshsharmadiyu2015@gmail.com
• Ulak, Forest Research and Training Centre, Government of Nepal, P.O. Box 3339, Babarmahal, Kathmandu 44600, Nepal E-mail: sunitaulak@gmail.com
• Paudel, Agriculture Forestry University, P.O. Box 13712 Rampur, Chitwan, Nepal E-mail: prashant.paudel88@gmail.com

We used a process-based hydrological model SUSI to improve guidelines for ditch network maintenance (DNM) operations on drained peatland forests. SUSI takes daily weather data, ditch depth, strip width, peat properties, and forest stand characteristics as input and calculates daily water table depth (WTD) at different distances from ditch. The study focuses on Scots pine (Pinus sylvestris L.) dominated stands which are the most common subjects of DNM. Based on a literature survey, and consideration of the tradeoffs between forest growth and detrimental environmental impacts, long term median July–August WTD of 0.35 m was chosen as a target WTD. The results showed that ditch depths required to reach such WTD depends strongly on climatic locations, stand volume, ditch spacing, and peat thickness and type. In typical ditch cleaning areas in Finland with parallel ditches placed about 40 m apart and tree stand volumes exceeding 45 m³ ha^–1, 0.3–0.8 m deep ditches were generally sufficient to lower WTD to the targeted depth of 0.35 m. These are significantly shallower ditch depths than generally recommended in operational forestry. The main collector ditch should be naturally somewhat deeper to permit water outflow. Our study brings a firmer basis on environmentally sound forestry on drained peatlands.

• Hökkä, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: hannu.hokka@luke.fi
• Laurén, University of Eastern Finland, Faculty of Science and Forestry, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: ari.lauren@uef.fi
• Stenberg, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: leena.stenberg@luke.fi
• Launiainen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: samuli.launiainen@luke.fi
• Leppä, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: kersti.leppa@luke.fi
• Nieminen, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: mika.nieminen@luke.fi

Ageing and competition reduce trees’ ability to capture resources, which predisposes them to death. In this study, the effect of senescence on the survival probability of Norway spruce (Picea abies (L.) Karst.) was analysed by fitting alternative survival probability models. Different model formulations were compared in the dataset, which comprised managed and unmanaged plots in long-term forest experiments in Finland and Norway, as well as old-growth stands in Finland. Stand total age ranged from 19 to 290 years. Two models were formulated without an age variable, such that the negative coefficient for the squared stem diameter described a decreasing survival probability for the largest trees. One of the models included stand age as a separate independent variable, and three models included an interaction term between stem diameter and stand age. According to the model including stand age and its interaction with stem diameter, the survival probability curves could intersect each other in stands with a similar structure but a different mean age. Models that did not include stand age underestimated the survival rate of the largest trees in the managed stands and overestimated their survival rate in the old-growth stands. Models that included stand age produced more plausible predictions, especially for the largest trees. The results supported the hypothesis that the stand age and senescence of trees decreases the survival probability of trees, and that the ageing effect improves survival probability models for Norway spruce.

• Siipilehto, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, P.O. Box 2, FI-00790 Helsinki, Finland E-mail: jouni.siipilehto@luke.fi
• Mäkinen, Natural Resources Institute Finland (Luke), Production systems, Latokartanonkaari 9, P.O. Box 2, FI-00790 Helsinki, Finland https://orcid.org/0000-0002-1820-6264 E-mail: harri.makinen@luke.fi
• Andreassen, Norwegian Institute of Bioeconomy Research (NIBIO), NO-1431 Ås, Norway E-mail: kjellandreassen@gmail.com
• Peltoniemi, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, P.O. Box 2, FI-00790 Helsinki, Finland https://orcid.org/0000-0003-2028-6969 E-mail: mikko.peltoniemi@luke.fi

Tree height-diameter allometry reflects the response of specific species to above and belowground resource allocation patterns. However, traditional methods (e.g. stepwise regression (SR)) may ignore model uncertainty during the variable selection process. In this study, 450 trees of Chinese fir (Cunninghamia lanceolata (Lamb.) Hook.) grown at five spacings were used. We explored the height-diameter allometry in relation to stand and climate variables through Bayesian model averaging (BMA) and identifying the contributions of these variables to the allometry, as well as comparing with the SR method. Results showed the SR model was equal to the model with the third highest posterior probability of the BMA models. Although parameter estimates from the SR method were similar to BMA, BMA produced estimates with slightly narrower 95% intervals. Heights increased with increasing planting density, dominant height, and mean annual temperature, but decreased with increasing stand basal area and summer mean maximum temperature. The results indicated that temperature was the dominant climate variable shaping the height-diameter allometry for Chinese fir plantations. While the SR model included the mean coldest month temperature and winter mean minimum temperature, these variables were excluded in BMA, which indicated that redundant variables can be removed through BMA.

• Lu, Key Laboratory of Tree Breeding and Cultivation of the National Forestry and Grassland Administration, Research Institute of Forestry, Chinese Academy of Forestry, Beijing 100091, P. R. China; Collaborative Innovation Center of Sustainable Forestry in Southern China, Nanjing Forestry University, Nanjing, 210037, P. R. China E-mail: 18556439861@163.com
• Chhin, Division of Forestry and Natural Resources, West Virginia University, 322 Percival Hall, 1145 Evansdale Dr, Morgantown, West Virginia, 26506, USA E-mail: steve.chhin@mail.wvu.edu
• Zhang, Key Laboratory of Tree Breeding and Cultivation of the National Forestry and Grassland Administration, Research Institute of Forestry, Chinese Academy of Forestry, Beijing 100091, P. R. China E-mail: xqzhang85@caf.ac.cn
• Zhang, Key Laboratory of Tree Breeding and Cultivation of the National Forestry and Grassland Administration, Research Institute of Forestry, Chinese Academy of Forestry, Beijing 100091, P. R. China; Collaborative Innovation Center of Sustainable Forestry in Southern China, Nanjing Forestry University, Nanjing, 210037, P. R. China E-mail: xqzhang85@yahoo.com

The diameter at any point on a stem and tree volume are some of the most important types of information used in forest management planning. One of the methods to predict the diameter at any point on a stem is to develop taper models. Black locust (Robinia pseudoacacia L.) occurs in almost all forests in Poland, with the largest concentration in the western part of the country. Using empirical data obtained from 13 black locust stands (48 felled trees), seven taper models with different numbers of estimated parameters were analysed for section diameters both over and under bark using fixed and mixed-effects modelling approaches. Assuming a lack of additional measurements, the best fitted taper models were used for the prediction of over bark volume using both methods. The predicted volume was compared with the results from different volume equations available for black locust. The variable-form taper model with eight estimated parameters fitted the data the best. The lowest root mean square error for volume prediction was achieved for the elaborated fixed-effects taper model (0.0476), followed by the mixed-effects taper model (0.0489). At the same time, the difference between the volume relative errors achieved based on the taper models does not differ significantly from the results obtained using the volume equations already available for black locust (two of the three analysed).

• Bronisz, Institute of Forest Sciences, Warsaw University of Life Sciences-SGGW, Nowoursynowska 159, PL 02-776 Warsaw, Poland E-mail: karol.bronisz@wl.sggw.pl
• Zasada, Institute of Forest Sciences, Warsaw University of Life Sciences-SGGW, Nowoursynowska 159, PL 02-776 Warsaw, Poland https://orcid.org/0000-0002-4881-296X E-mail: Michal.Zasada@wl.sggw.pl