Articles containing the keyword 'disturbance'

There is no doubt that tree survival, growth, and reproduction in North America's boreal forests would be directly influenced by the projected changes in climate if they occur. The indirect effects of climate change may be of even greater importance, however, because of their potential for altering the intensity, frequency, and perhaps even the very nature of the disturbance regimes which drive boreal forest dynamics. Insect defoliator populations are one of the dominating disturbance factors in North America's boreal forests and during outbreaks trees are often killed over vast forest areas. If the predicted shifts in climate occur, the damage patterns caused by insects may be considerably changed, particularly those of insects whose temporal and spatial distributions are singularly dependent on climatic factors. The ensuing uncertainties directly affect depletion forecasts, pest hazard rating procedures, and long-term planning for pest control requirements. Because the potential for wildfire often increases in stands after insect attack, uncertainties in future insect damage patterns also lead to uncertainties in fire regimes. In addition, because the rates of processes key to biogeochemical and nutrient recycling are influenced by insect damage, potential changes in damage patterns can indirectly affect ecosystem resilience and the sustainability of the multiple uses of the forest resource.

In this paper, a mechanistic perspective is developed based on available information describing how defoliating forest insects might respond to climate warming. Because of its prevalence and long history of study, the spruce budworm, Choristoneura fumiferana Clem. (Lepidoptera: Tortricidae), is used for illustrative purposes in developing this perspective. The scenarios that follow outline the potential importance of threshold behaviour, historical conditions, phenological relationships, infrequent but extreme weather, complex feedbacks, and natural selection. The urgency of such considerations is emphasized by reference to research suggesting that climate warming may already be influencing some insect lifecycles.

• Fleming, E-mail: rf@mm.unknown

Certain biocides used in production of tree nursery stock exterminate undesirable organisms but cause an abnormal growth stimulation of plants. The reforestation material has decreased survival potential because of high degree of succulence, top:root and height:diameter ratios, and low specific gravity and root surface area. Some fumigants impede mycorrhizae development and arrest phosphorus uptake. Recovery of growth potential was achieved by aluminium sulphate and/or fermented compost inoculated with mycorrhiza-forming fungi.

The PDF includes a summary in Finnish.

• Iyer, E-mail: ji@mm.unknown
• Chesters, E-mail: gc@mm.unknown
• Wilde, E-mail: sw@mm.unknown

Wildfires as natural disturbances have had important impacts on terrestrial ecosystems, including forests. We studied patterns of short-term vegetation recovery after surface fire in protected hemiboreal Pinus sylvestris L.-dominated forest. Our study was carried out near Stikli village in Western Latvia. Seven forest stands – middle-age and over-mature were sampled on nutrient-poor and mesic soils. Forest fire occurred in the summer of 2018 and covered 1440 ha of forested area. In each stand we established 16 sample plots (1 m × 1 m) in a radial pattern from the center. Every summer from 2019 till 2022 we surveyed these sample plots – recorded projective cover (%) and identified Ellenberg indicator values and species traits – plant strategy groups (C-S-R after Grime), Raunkiær life history forms and habitat types. Additionally, the occurrence of specialized fire-adapted plants was recorded. In total we identified 15 species in the ground layer, 47 species in the herbaceous layer, and 9 regenerating tree species. The colonization at the ground layer was the most rapid (projective cover increased overall by 67% in middle-aged stands and by 82% in over-mature stands). Species diversity was the highest at the herb layer during the third (middle-aged stands) and fourth (over-mature stands) after fire disturbance but showed overall declining trends. Betula spp. and Populus tremula L.-dominated regenerating tree species. The dominance of fire-adapted species declined rapidly after the fire except for moss Polytrichum spp. Overall, hemiboreal over-mature stands demonstrated higher vegetation cover and more rapid rate of initial colonization compared to middle-aged stands.

• Liepa, Latvian State Forest Research Institute “Silava”, Rīgas street 111, LV-2169, Salaspils, Latvia; Latvian University of Life Sciences and Technologies, Faculty of Forest and Environmental Sciences, Akadēmijas street 11, LV-3001, Jelgava, Latvia https://orcid.org/0000-0002-8270-6722 E-mail: liga.liepa@outlook.com
• Rendenieks, Latvian State Forest Research Institute “Silava”, Rīgas street 111, LV-2169, Salaspils, Latvia https://orcid.org/0000-0002-3511-1486 E-mail: zigmars.rendenieks@silava.lv
• Dubrovskis, Latvian University of Life Sciences and Technologies, Faculty of Forest and Environmental Sciences, Akadēmijas street 11, LV-3001, Jelgava, Latvia https://orcid.org/0000-0003-0810-5651 E-mail: edgars.dubrovskis@lbtu.lv
• Freimane, Latvian University of Life Sciences and Technologies, Faculty of Forest and Environmental Sciences, Akadēmijas street 11, LV-3001, Jelgava, Latvia E-mail: lasma.freimane@lbtu.lv
• Straupe, Latvian University of Life Sciences and Technologies, Faculty of Forest and Environmental Sciences, Akadēmijas street 11, LV-3001, Jelgava, Latvia https://orcid.org/0000-0002-2098-7194 E-mail: inga.straupe@lbtu.lv
• Jansons, Latvian State Forest Research Institute “Silava”, Rīgas street 111, LV-2169, Salaspils, Latvia https://orcid.org/0000-0001-7981-4346 E-mail: aris.jansons@silava.lv

Since fire frequency is expected to increase globally due to climate change, it is important to understand its effects on forest ecosystems. We studied the long-term patterns in species diversity, cover and composition of vascular plants and bryophytes after forest fire and the site-related factors behind them. Research was carried out in northwestern Estonia, using a chronosequence of Scots pine (Pinus sylvestris L.) stands, located on nutrient poor sandy soils, where fires had occurred 12, 23, 38, 69, 80 and 183 years ago. In every stand three 100 m² vegetation plots were established to collect floristic and environmental information. The effects on floristic characteristics of time since fire, light, and soil variables were evaluated with linear mixed models, followed by backward variable selection. Compositional variation was analysed with non-metric multidimensional scaling, Multi-response Permutation Procedures, and Indicator Species Analysis. Altogether, 31 vascular plant and 39 bryophyte species were found in vegetation plots. The cover of the vascular plant and bryophyte layers increased with a longer time since fire. Soil and light variables impacted the richness of several vascular plant and bryophyte groups, whereas only the richness of liverworts and dwarf-shrubs correlated with time since fire. Considerable compositional differences were observed in vascular plant and bryophyte assemblages between recently vs. long-time ago burned stands. To conclude, time since fire significantly impacted compositional patterns of vascular plants and bryophytes in pine forests on nutrient poor soils, although time-related trends in species richness were less evident.

• Orumaa, Institute of Forestry and Engineering, Estonian University of Life Sciences, Kreutzwaldi 5, 51006, Tartu, Estonia E-mail: argo.orumaa@emu.ee
• Köster, Department of Environmental and Biological Sciences, University of Eastern Finland, PO Box 111 (Yliopistokatu 7), 80130, Joensuu, Finland E-mail: kajar.koster@helsinki.fi
• Tullus, Department of Botany, Institute of Ecology and Earth Sciences, University of Tartu, Lai 40, Tartu, 51003, Estonia E-mail: arvo.tullus@ut.ee
• Tullus, Institute of Forestry and Engineering, Estonian University of Life Sciences, Kreutzwaldi 5, 51006, Tartu, Estonia E-mail: tea.tullus@emu.ee
• Metslaid, Institute of Forestry and Engineering, Estonian University of Life Sciences, Kreutzwaldi 5, 51006, Tartu, Estonia E-mail: marek.metslaid@emu.ee

Fire is a common disturbance in boreal forests causing changes in biological diversity at various spatial scales. In the past 100 years, forest management has limited fire outbreaks, but in the future, the fire-affected forest area is expected to increase in many regions due to climate change. Burned forests are typically salvage-logged, but the effect of this type of management versus natural regeneration on biological diversity is not well understood, particularly the mid-term effect to tree establishment and understory vegetation composition and diversity. Various management methods were used after a large fire in 1992 in a peatland-forest complex and neighbouring managed forests, which created an experimental setup for study of the effect of management after fire in the Sliteres National park, northwestern Latvia. Understory vegetation was described in plots using a design of four forest and three management types: natural regeneration (unmanaged) and managed sites with salvage logging followed by no further human intervention and salvage logging with planting. Post-fire management had different effect in each forest type. Species richness was higher in forest types with salvage logging than in natural regenerated sites on rich wet and rich dry forest types, but not for the poor forest types. Tree regeneration was generally greater in salvage-logged stands, but differed between forest types. Species composition was related to tree regeneration and canopy openness. In contrast to other studies, salvage logging had a positive mid-term effect to ground vegetation diversity and tree establishment in the studied stands, implying potential for concomitant management and conservation of ground cover vegetation in semi-natural stands.

• Kārkliņa, Latvian State Forest Research Institute “Silava”, 111 Rigas Street, LV-2169, Salaspils, Latvia E-mail: annija.karklina@silava.lv
• Brūmelis, University of Latvia, Faculty of Biology, Jelgavas street 1, LV-1004, Riga, Latvia E-mail: guntis.brumelis@lu.lv
• Dauškane, University of Latvia, Faculty of Biology, Jelgavas street 1, LV-1004, Riga, Latvia E-mail: iluta.dauskane@lu.lv
• Elferts, Latvian State Forest Research Institute “Silava”, 111 Rigas Street, LV-2169, Salaspils, Latvia; University of Latvia, Faculty of Biology, Jelgavas street 1, LV-1004, Riga, Latvia E-mail: didzis.elferts@lu.lv
• Freimane, Latvian State Forest Research Institute “Silava”, 111 Rigas Street, LV-2169, Salaspils, Latvia E-mail: lasma.freimane@silava.lv
• Kitenberga, Latvian State Forest Research Institute “Silava”, 111 Rigas Street, LV-2169, Salaspils, Latvia E-mail: mara.kitenberga@silava.lv
• Lībiete, Latvian State Forest Research Institute “Silava”, 111 Rigas Street, LV-2169, Salaspils, Latvia E-mail: zane.libiete@silava.lv
• Matisons, Latvian State Forest Research Institute “Silava”, 111 Rigas Street, LV-2169, Salaspils, Latvia E-mail: robism@inbox.lv
• Jansons, Latvian State Forest Research Institute “Silava”, 111 Rigas Street, LV-2169, Salaspils, Latvia E-mail: aris.jansons@silava.lv

Dead wood profile of a forest is a useful tool for describing forest characteristics and assessing forest disturbance history. Nevertheless, there are few studies on dead wood profiles, including both coarse and fine dead wood, and on the effect of sampling intensity on the dead wood estimates. In a semi-natural boreal forest, we measured every dead wood item over 2 cm in diameter from 80 study plots. From eight plots, we further recorded dead wood items below 2 cm in diameter. Based on these data we constructed the full dead wood profile, i.e. the overall number of dead wood items and their distribution among different tree species, volumes of different size and decay stage categories. We discovered that while the number of small dead wood items was immense, their number dropped drastically from the diameter below 1 cm to diameters 2–3 cm. Different tree species had notably different abundance-diameter distribution patterns: spruce dead wood comprised most strikingly the smallest diameter fractions, whereas aspen dead wood comprised a larger share of large-diameter items. Most of the dead wood volume constituted of large pieces (>10 cm in diameter), and 62% of volume was birch. The variation in the dead wood estimates was small for the numerically dominant tree species and smallest diameter categories, but high for the sub-dominant tree species and larger size categories. In conclusion, the more the focus is on rare tree species and large dead wood items, the more comprehensive should the sampling be.

• Halme, Department of Biological and Environmental Science, University of Jyväskylä, P.O. Box 35, FI-40014 University of Jyväskylä, Finland; School of Resource Wisdom, University of Jyväskylä, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: panu.halme@jyu.fi
• Purhonen, Department of Biological and Environmental Science, University of Jyväskylä, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: jenna.e.i.purhonen@jyu.fi
• Marjakangas, Centre for Biodiversity Dynamics, Department of Biology, Norwegian University of Science and Technology, N-7491 Trondheim, Norway E-mail: emma-liina.marjakangas@ntnu.no
• Komonen, Department of Biological and Environmental Science, University of Jyväskylä, P.O. Box 35, FI-40014 University of Jyväskylä, Finland; School of Resource Wisdom, University of Jyväskylä, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: atte.komonen@jyu.fi
• Juutilainen, Department of Biological and Environmental Science, University of Jyväskylä, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: kjuutilainen@yahoo.com
• Abrego, Department of Agricultural Sciences, University of Helsinki, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: nerea.abrego@helsinki.fi

Fire as disturbance of forests has an important ecological and economical role in boreal and hemiboreal forests. The occurrence of forest fires is both climatically and anthropogenically determined and shifts in fire regimes are expected due to climate change. Although fire histories have been well documented in boreal regions, there is still insufficient information about fire occurrence in the Baltic States. In this study, spatio-temporal patterns and climatic drivers of forest fires were assessed by means of spatial and time-series analysis. The efficiency of Canadian Fire Weather (FWI) indices as indicators for fire activity was tested. The study was based on data from the literature, archives, and the Latvian State Forest service database. During the period 1922–2014, the occurrence and area affected by forest fires has decreased although the total area of forest land has nearly doubled, suggesting improvement of the fire suppression system as well as changes in socioeconomic situation. The geographical distribution of forest fires revealed two pronounced clusters near the largest cities of Riga and Daugavpils, suggesting dominance of human causes of ignitions. The occurrence of fires was mainly influenced by drought. FWI appeared to be efficient in predicting the fire occurrence: 23–34% of fires occurred on days with a high or extremely high fire danger class, which overall had a relative occurrence of only 4.3–4.6%. During the 20th century, the peak of fire activity shifted from May to April, probably due to global warming and socioeconomic reasons. The results of this study are relevant for forest hazard mitigation and development of fire activity prediction system in Latvia.

• Donis, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: janis.donis@silava.lv
• Kitenberga, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: mara.kitenberga@gmail.com
• Šņepsts, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: guntars.snepsts@silava.lv
• Matisons, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: robism@inbox.lv
• Zariņš, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: juris.zarins@silava.lv
• Jansons, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: aris.jansons@silava.lv

Passive rewildening of forest ecosystems is commonly used for rehabilitating degraded habitats closer to their natural origin in addition to costly active restoration measures. However, it is not clear if passive processes are effective and how long the recovery of main ecosystem properties takes. We investigate the recovery of forest soil and tree stand characteristics a century after cessation of slash-and-burn cultivation, a major historical intensive disturbance regime that was applied widely in boreal zone of Finland until late 1800s. We systematically sampled soil and tree stand parameters within former slash-and-burn and nearby control areas. Humus layer thickness and soil organic matter (SOM) stocks were still lower in the historical slash-and-burn than in control areas. Slash-and-burn areas also had a larger volume of live birch trees and a higher standing dead wood volume than control areas. Accordingly, organic matter (humus layer thickness and SOM stocks) correlated negatively with birch standing live tree volume. Combined OM stock in humus and uppermost 10 cm mineral soil layer was positively correlated with lying dead wood volume. Overall, we observed clear recovery of several natural properties but we also found that a century after cessation of frequent anthropogenic burnings, clear legacies of disturbance in the above- and below-ground parts of boreal ecosystem were evident. Our results indicate that if only passive rewildening is applied as a restoration measure, the full recovery of boreal forest is slow and the effects of historical land-use may persist for over hundred years in soil and tree properties.

• Čugunovs, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: mihails.cugunovs@uef.fi
• Tuittila, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland http://orcid.org/0000-0001-8861-3167 E-mail: eeva-stiina.tuittila@uef.fi
• Sara-Aho, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: ida.sara-aho@mhy.fi
• Pekkola, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: laura.pekkola@gmail.com
• Kouki, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland http://orcid.org/0000-0003-2624-8592 E-mail: jari.kouki@uef.fi

Rut formation caused by logging operations has been recognised as a challenge for Swedish forestry. Frequent traffic with heavy machines on extraction roads, together with a warmer climate, is one of the factors that increases the risk of rut formation in forests. One possible way to control this impact of logging operations is to design and apply decision support tools that enable operators to take sensitive areas into account when planning extraction roads. In this study, 16 different logging sites in south-eastern Sweden were surveyed after clear-cut. Information was collected about extraction roads (i.e. traffic intensity, whether the roads had been reinforced with slash) and ruts. Digital maps such as cartographic depth-to-water (DTW) index and soil type were also examined for any connection to rut positions. Soil type and traffic intensity were found to be significant factors in rut formation, while DTW and slash reinforcement were not. However, the DTW map combined with other information, such as soil type, could contribute to decision support tools that improve planning of extraction roads.

• Mohtashami, The forestry research institute of Sweden, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: sima.mohtashami@skogforsk.se
• Eliasson, The forestry research institute of Sweden, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden http://orcid.org/0000-0002-2038-9864 E-mail: lars.eliasson@skogforsk.se
• Jansson, The forestry research institute of Sweden, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden http://orcid.org/0000-0002-3018-9161 E-mail: gunnar.jansson@skogforsk.se
• Sonesson, The forestry research institute of Sweden, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden http://orcid.org/0000-0002-2018-7496 E-mail: johan.sonesson@skogforsk.se

Forest ecological restoration by burning is widely applied to promote natural, early-successional sites and increase landscape biodiversity. Burning is also used as a forest management practice to facilitate forest regeneration after clearcutting. Besides the desired goals, restoration burnings also affect soil biogeochemistry, particularly soil organic matter (SOM) and related soil carbon stocks but the long-term effects are poorly understood. However, in order to study these effects, a reliable estimate of spatial variability is first needed for effective sampling. Here we investigate spatial variability of SOM and vegetation features 13 years after burnings and in combination with variable harvest levels. We sampled four experimental sites representing distinct management and restoration treatments with an undisturbed control. While variability of vegetation cover and biomass was generally higher in disturbed sites, soil parameter variability was not different between the four sites. The joint ecological patterns of soil and vegetation parameters across the whole sample continuum support well the prior assumptions on the characteristic disturbance conditions within each of the study sites. We designed and employed statistical simulations as a means to plan prospective sampling. Sampling six forest sites for each treatment type with 30 independent soil cores per site would provide enough statistical power to adequately capture the impacts of burning on SOM based on the data we obtained here and statistical simulations. In conclusion, we argue that an informed design-based approach to documenting the ecosystem effects of forest burnings is worth applying both through obtaining new data and meta-analysing the existing.

• Čugunovs, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: mihails.cugunovs@uef.fi
• Tuittila, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland http://orcid.org/0000-0001-8861-3167 E-mail: eeva-stiina.tuittila@uef.fi
• Mehtätalo, University of Eastern Finland, School of Computing, Science Park, Länsikatu 15, P.O. Box 111, 80101 Joensuu, Finland http://orcid.org/0000-0002-8128-0598 E-mail: lauri.mehtatalo@uef.fi
• Pekkola, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: laura.pekkola@gmail.com
• Sara-Aho, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: ida.sara-aho@mhy.fi
• Kouki, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland http://orcid.org/0000-0003-2624-8592 E-mail: jari.kouki@uef.fi

Boreal forest soil contains significant amounts of organic carbon. Soil disturbance, caused for example by site preparation or stump extraction, may increase decomposition and thus lead to higher CO₂ emissions, contributing to global warming. The aim of this study was to quantify responses of soil-surface CO₂ fluxes (R_s) and litter (needle and root) decomposition rates following various kinds of soil disturbance commonly caused by mechanical site preparation and stump harvest. For this purpose four treatments were applied in a clear-cut site in central Sweden: i) removal of the humus layer and top 2 cm of mineral soil, ii) placement of a humus layer and 2 cm of mineral soil upside down on top of undisturbed soil, forming a double humus layer buried under mineral soil, iii) heavy mixing of the humus layer and mineral soil, and iv) no disturbance (control). R_s measurements were acquired with a portable respiration system during two growing seasons. To assess the treatments’ effects on litter decomposition rates, needles or coarse roots (Ø = 6 mm) were incubated in litterbags at positions they would be located after the treatments (buried, or on top of the soil). The results indicate that site preparation-simulating treatments have no effect or may significantly reduce, rather than increase, CO₂ emissions during the following two years. They also show that buried litter decomposes more rapidly than litter on the surface, but in other respects the treatments have little effect on litter decomposition rates.

• Mjöfors, Swedish University of Agricultural Sciences (SLU), Department of Soil and Environment, P.O. Box 7014, 150 07 Uppsala, Sweden E-mail: kristina.mjofors@slu.se
• Strömgren, Swedish University of Agricultural Sciences (SLU), Department of Soil and Environment, P.O. Box 7014, 150 07 Uppsala, Sweden E-mail: Monika.stromgren@slu.se
• Nohrstedt, Swedish University of Agricultural Sciences (SLU), Department of Soil and Environment, P.O. Box 7014, 150 07 Uppsala, Sweden E-mail: Hans-orjan.nohrstedt@slu.se
• Gärdenäs, Swedish University of Agricultural Sciences (SLU), Department of Soil and Environment, P.O. Box 7014, 150 07 Uppsala, Sweden E-mail: Annemieke.gardenas@slu.se

• Franklin, Department of Renewable Resources, University of Alberta, 751 General Services Building, Edmonton, Alberta, T6G 2H1, Canada E-mail: cfrankli@ualberta.ca
• Harper, School for Resource and Environmental Studies, Dalhousie University, Suite 5010, 6100 University Ave., Halifax, Nova Scotia, B3H 3J5, Canada E-mail: Karen.Harper@dal.ca
• Murphy, Department of Environmental Science, Saint Mary’s University, 923 Robie St., Halifax, Nova Scotia, B3H 3C3, Canada E-mail: liamkmurphy@gmail.com

• Ezzati, Department of Forestry and Forest Engineering, Tarbiat Modares University, P.O. Box 64414-356, Iran E-mail: se@nn.ir
• Najafi, Department of Forestry and Forest Engineering, Tarbiat Modares University, P. Box 64414-356, Iran E-mail: a.najafi@modares.ac.ir
• Rab, Soil Physics Future Farming Systems Research Division, Department of Primary Industries, Victoria, Australia E-mail: mr@nn.ir
• Zenner, Department of Ecosystem Science and Management, Pennsylvania State University, University Park, PA, USA E-mail: eric.zenner@psu.edu

• Dragotescu, Université du Québec à Montréal, Centre d’Étude de la Forêt (CEF), Montreal, Quebec, Canada E-mail: idragot@hotmail.com
• Kneeshaw, Université du Québec à Montréal, Centre d’Étude de la Forêt (CEF), Montreal, Quebec, Canada E-mail: ddk@nn.ca

• Strömgren, Swedish University of Agricultural Sciences, Department of Soil and Environment, Uppsala, Sweden E-mail: monika.stromgren@slu.se
• Mjöfors, Swedish University of Agricultural Sciences, Department of Soil and Environment, Uppsala, Sweden E-mail: km@nn.se
• Holmström, Swedish University of Agricultural Sciences, Department of Soil and Environment, Uppsala, Sweden E-mail: bh@nn.fi
• Grelle, Swedish University of Agricultural Sciences, Department of Ecology, Uppsala, Sweden E-mail: ag@nn.se

• Angelstam, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden E-mail: per.angelstam@slu.se
• Andersson, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden E-mail: ka@nn.se
• Axelsson, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden E-mail: ra@nn.se
• Elbakidze, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden E-mail: me@nn.se
• Jonsson, Dept of Natural Science, Engineering and Mathematics, Mid Sweden University, Sundsvall, Sweden E-mail: bgj@nn.se
• Roberge, Dept of Wildlife, Fish and Environmental Studies, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Umeå, Sweden E-mail: jmr@nn.se

• Grenfell, University of Helsinki, Dept of Forest Sciences, Helsinki, Finland E-mail: russell.grenfell@helsinki.fi
• Aakala, University of Helsinki, Dept of Forest Sciences, Helsinki, Finland E-mail: ta@nn.fi
• Kuuluvainen, University of Helsinki, Dept of Forest Sciences, Helsinki, Finland E-mail: tk@nn.fi

• Fenton, Université du Québec en Abitibi-Témiscamingue, 445 Boulevard de l’Université, Rouyn-Noranda, Québec, Canada J9X 4E5 E-mail: nicole.fenton@uqat.ca
• Bergeron, Université du Québec en Abitibi-Témiscamingue, 445 Boulevard de l’Université, Rouyn-Noranda, Québec, Canada J9X 4E5 E-mail: yb@nn.ca

• Jönsson, Department of Ecology, SLU, P.O. Box 7044, SE-750 07 Uppsala, Sweden (current); Department of Natural Sciences, Engineering and Mathematics, Mid Sweden University, Sundsvall, Sweden E-mail: mari.jonsson@slu.se
• Fraver, U.S. Forest Service, Northern Research Station, Grand Rapids, Minnesota, USA (current); Department of Natural Sciences, Engineering and Mathematics, Mid Sweden University, Sundsvall, Sweden E-mail: sf@nn.us
• Jonsson, Department of Natural Sciences, Engineering and Mathematics, Mid Sweden University, Sundsvall, Sweden E-mail: bgj@nn.se

• Bottero, University of Turin, Department Agroselviter, Via Leonardo da Vinci 44, 10095 Grugliasco (TO), Italy E-mail: alessandra.bottero@unito.it
• Garbarino, University of Turin, Department Agroselviter, Via Leonardo da Vinci 44, 10095 Grugliasco (TO), Italy E-mail: mg@nn.it
• Dukic, University of Banja Luka, Faculty of Forestry, Banja Luka, Bosnia and Herzegovina E-mail: vd@nn.ba
• Govedar, University of Banja Luka, Faculty of Forestry, Banja Luka, Bosnia and Herzegovina E-mail: zg@nn.ba
• Lingua, University of Padua, Department of TeSAF, Legnaro (PD), Italy E-mail: el@nn.it
• Nagel, University of Ljubljana, Department of Forestry and Renewable Forest Resources, Ljubljana, Slovenia E-mail: tan@nn.si
• Motta, University of Turin, Department Agroselviter, Via Leonardo da Vinci 44, 10095 Grugliasco (TO), Italy E-mail: rm@nn.it

• Saunders, Department of Forestry and Natural Resources, Purdue University, 715 State Street, West Lafayette, IN, USA E-mail: msaunder@purdue.edu
• Fraver, USFS Northern Research Station, Grand Rapids, MN, USA E-mail: sf@nn.us
• Wagner, School of Forest Resources, University of Maine, Orono, ME, USA E-mail: rgw@nn.us

• Lafleur, NSERC-UQAT-UQAM Industrial Chair in Sustainable Forest Management, Université du Québec en Abitibi-Témiscamingue, 445 boul. de l’Université, Rouyn-Noranda, QC J9X 5E4, Canada E-mail: benoit.lafleur@uqat.ca
• Fenton, NSERC-UQAT-UQAM Industrial Chair in Sustainable Forest Management, Université du Québec en Abitibi-Témiscamingue, 445 boul. de l’Université, Rouyn-Noranda, QC J9X 5E4, Canada E-mail: njf@nn.ca
• Paré, Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, 1055 du P.E.P.S., P.O. Box 10380, Stn. Sainte-Foy, QC G1V 4C7, Canada E-mail: dp@nn.ca
• Simard, Département de Géographie, Université Laval, Pavillon Abitibi-Price, 2405 rue de la Terrasse, Québec, QC G1V 0A6, Canada E-mail: ms@nn.ca
• Bergeron, NSERC-UQAT-UQAM Industrial Chair in Sustainable Forest Management, Université du Québec en Abitibi-Témiscamingue, 445 boul. de l’Université, Rouyn-Noranda, QC J9X 5E4, Canada E-mail: yb@nn.ca

• Hellberg, Tunstigen 10, SE-831 43 Östersund, Sweden E-mail: eh@nn.se
• Josefsson, Swedish University of Agricultural Sciences, Department of Forest Ecology and Management, SE-901 83 Umeå, Sweden E-mail: torbjorn.josefsson@svek.slu.se
• Östlund, Swedish University of Agricultural Sciences, Department of Forest Ecology and Management, SE-901 83 Umeå, Sweden E-mail: lo@nn.se

• Lilja-Rothsten, University of Helsinki, Dept. of Forest Ecology, Finland E-mail: saara.lilja@helsinki.fi
• Chantal, University of Helsinki, Dept. of Forest Ecology, Finland E-mail: mdc@nn.fi
• Peterson, Dept. of Plant Biology, University of Georgia, Athens, GA, USA E-mail: cp@nn.us
• Kuuluvainen, University of Helsinki, Dept. of Forest Ecology, Finland E-mail: tk@nn.fi
• Vanha-Majamaa, The Finnish Forest Research Institute, Vantaa Unit, Finland E-mail: ivm@nn.fi
• Puttonen, The Finnish Forest Research Institute, Vantaa Unit, Finland E-mail: pp@nn.fi

• Lilja, University of Helsinki, Department of Forest Ecology, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: saara.lilja@helsinki.fi
• Kuuluvainen, University of Helsinki, Department of Forest Ecology, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: tk@nn.fi

• Weyenberg, University of Minnesota, Department of Forest Resources, 1530 Cleveland Ave. N., St. Paul, MN 55108, USA E-mail: saw@nn.us
• Frelich, University of Minnesota, Department of Forest Resources, 1530 Cleveland Ave. N., St. Paul, MN 55108, USA E-mail: freli001@umn.edu
• Reich, University of Minnesota, Department of Forest Resources, 1530 Cleveland Ave. N., St. Paul, MN 55108, USA E-mail: pbr@nn.us

• Spinelli, CNR - Timber and Tree Institute, Via Madonna del Piano - Pal. F, 50019 Sesto Fiorentino, Italy; University College Dublin, Ireland E-mail: spinelli@ivalsa.cnr.it
• Owende, Dept. of Agricultural and Food Engineering, University College Dublin, Earlsfort Terrace, Dublin 2, Ireland E-mail: pmoo@nn.ie
• Ward, Dept. of Agricultural and Food Engineering, University College Dublin, Earlsfort Terrace, Dublin 2, Ireland E-mail: smw@nn.ie
• Tornero, Servicios Forestales SA, Ctra. SE-184 Km 0.63, E-41970 Santiponce, Sevilla, Spain E-mail: mt@nn.es

• Drobyshev, SUFOR Project, Department of Plant Ecology and Systematics, Ecology Building, Sölvegatan 37, Lund University, SE-223 62 Lund, Sweden E-mail: igor.drobyshev@ekol.lu.se
• Niklasson, SUFOR Project, Southern Swedish Forest Research Centre, SLU, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: mn@nn.se
• Angelstam, Grimsö Wildlife Research Station, Department of Conservation Biology, Forest Faculty, SLU, SE-730 91 Riddarhyttan, Sweden E-mail: pa@nn.se

• Purdy, Dept. of Renewable Resources, University of Alberta, Edmonton, AB, Canada T6G 2E3 E-mail: bgp@nn.ca
• Macdonald, Dept. of Renewable Resources, University of Alberta, Edmonton, AB, Canada T6G 2E3 E-mail: ellen.macdonald@ualberta.ca
• Dale, Dale, Dept. of Biological Sciences, University of Alberta, Edmonton, AB, Canada T6G 2E9 E-mail: mrtd@nn.ca

• Quine, Forestry Commission, Northern Research Station, Roslin, Midlothian EH25 9SY, United Kingdom E-mail: chris.quine@forestry.gsi.gov.uk
• Humphrey, Forestry Commission, Northern Research Station, Roslin, Midlothian EH25 9SY, United Kingdom E-mail: jwh@nn.uk
• Purdy, Forestry Commission, Northern Research Station, Roslin, Midlothian EH25 9SY, United Kingdom E-mail: kp@nn.uk
• Ray, Forestry Commission, Northern Research Station, Roslin, Midlothian EH25 9SY, United Kingdom E-mail: dr@nn.uk

• Pennanen, Department of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: juho.pennanen@helsinki.fi

• Wallenius, Department of Ecology and Systematics, University of Helsinki, P.O. Box 56, FIN-00014 University of Helsinki, Finland E-mail: tuomo.wallenius@helsinki.fi

• Wallenius, Department of Ecology and Systematics, University of Helsinki, P.O. Box 65, FIN-00014, Helsinki, Finland E-mail: tuomo.wallenius@helsinki.fi
• Kuuluvainen, Department of Forest Ecology, University of Helsinki, P.O. Box 27, FIN-00014, Helsinki, Finland E-mail: tk@nn.fi
• Heikkilä, Research Centre of Friendship Park, Tönölä, FIN-88900 Kuhmo, Finland E-mail: rh@nn.fi
• Lindholm, Finnish Environment Institute, Nature and Land Use Division, P.O. Box 140, FIN-00251 Helsinki, Finland E-mail: tl@nn.fi

• Kuuluvainen, Department of Forest Ecology, University of Helsinki, P.O. Box 24, FIN-00014 University of Helsinki, Finland E-mail: timo.kuuluvainen@helsinki.fi
• Mäki, Department of Forest Ecology, University of Helsinki, P.O. Box 24, FIN-00014 University of Helsinki, Finland E-mail: jm@nn.fi
• Karjalainen, Department of Forest Ecology, University of Helsinki, P.O. Box 24, FIN-00014 University of Helsinki, Finland E-mail: lk@nn.fi
• Lehtonen, Faculty of Forestry, University of Joensuu, P.O. Box 24, FIN-80101 Joensuu, Finland E-mail: hl@nn.fi

• Karjalainen, Department of Forest Ecology, P.O. Box 24, FIN-00014 University of Helsinki, Finland E-mail: lk@nn.fi
• Kuuluvainen, Department of Forest Ecology, P.O. Box 24, FIN-00014 University of Helsinki, Finland E-mail: timo.kuuluvainen@helsinki.fi

• Rouvinen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: seppo.rouvinen@forest.joensuu.fi
• Kuuluvainen, University of Helsinki, Department of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: tk@nn.fi
• Siitonen, Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: juha.siitonen@metla.fi

Mechanical site preparation (MSP) is deliberate soil disturbance which is undertaken to improve the conditions for forest regeneration. Disc trenching and mounding are the dominant MSP practices currently used in Sweden and Finland. In this paper, the impacts of MSP on the soil, water quality, greenhouse gas (GHG) emissions and ground vegetation of mineral soil sites in Sweden and Finland are reviewed. The practices considered are patch scarification, mounding, inverting, disc trenching, and ploughing, which together represent a wide range of soil disturbance intensity. The environmental effects of MSP in this region have not been studied extensively. The environmental impact of MSP derives from the process of creating microsites which involves horizontal and/or vertical redistribution of soil and soil mixing. This typically affects decomposition, element circulation and leaching, vegetation coverage and uptake of nutrients and water, and possibly erosion and sediment exports. Following disc trenching or mounding the effects on GHG emissions appear to be minor over the first two years. For a few years after disc trenching concentrations in soil water collected below ridges are higher than that below furrows for some elements (e.g., NO₃^-, NH₄⁺, Mg²⁺, and total or dissolved organic C). The physical and chemical effects of ploughing remain detectable for several decades. There is little evidence about how the effects of forestry activities in upland areas on soil-water chemistry are transferred to adjacent surface water bodies, including what role streamside discharge areas play. MSP increases the tree biomass C store and may increase the total ecosystem C store. The impact of MSP on the cover and abundance of ground vegetation species depends on the composition of the original plant community, MSP intensity, and the establishment rate of different species. Species cover generally seems to decline for late succession understory species, while pioneer and ruderal species can benefit from the microsites created. Areas containing lichens which are used for reindeer forage require special consideration. More research is needed on the environmental effects of MSP, particularly regarding its long-term effects. Further efforts should be made to develop efficient site-preparation practices which better balance the disturbance intensity with what is needed for successful regeneration.

• Ring, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, 751 83, Uppsala, Sweden https://orcid.org/0000-0002-8962-9811 E-mail: eva.ring@skogforsk.se
• Sikström, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, 751 83, Uppsala, Sweden E-mail: ulf.sikstrom@skogforsk.se

• Brumelis, Faculty of Biology, University of Latvia, Kronvalda bulv. 4, Riga, LV-1586, Latvia; E-mail: guntis.brumelis@lu.lv
• Jonsson, Department of Natural Sciences, Engineering and Mathematics, Mid Sweden University, Sundsvall, Sweden E-mail: bgj@nn.se
• Kouki, School of Forest Sciences, University of Eastern Finland, Joensuu, Joensuu E-mail: jk@nn.fi
• Kuuluvainen, Department of Forest Sciences, University of Helsinki, Finland E-mail: tk@nn.fi
• Shorohova, Saint-Petersburg State Forest Academy, Saint-Petersburg, Russia & Finnish Forest Research Institute, Vantaa Research Unit, Vantaa, Finland E-mail: es@nn.ru

• Burton, Canadian Forest Service, Natural Resources Canada, 3333 University Way, Prince George, British Columbia, Canada V2N 4Z9 E-mail: Phil.Burton@NRCan-RNCan.gc.ca
• Macdonald, Department of Renewable Resources, University of Alberta, Edmonton, Alberta, Canada E-mail: sem@nn.ca

• Kuuluvainen, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: timo.kuuluvainen@helsinki.fi
• Aakala, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ta@nn.fi

• Shorohova, Saint-Petersburg State Forest University, Saint-Petersburg, Russia & Finnish Forest Research Institute, Vantaa Research Unit, Vantaa, Finland (ekaterina.shorohova@metla.fi) E-mail: shorohova@ES13334.spb.edu
• Kneeshaw, Université du Québec à Montréal, Centre d’étude de la forêt, Montreal, Canada E-mail: dk@nn.ca
• Kuuluvainen, University of Helsinki, Department of Forest Sciences, Helsinki, Finland E-mail: tk@nn.fi
• Gauthier, Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, Québec, Canada E-mail: sg@nn.fi

• Vasaitis, Department of Forest Mycology & Pathology, Swedish University of Agricultural Sciences, P.O. Box 7026, SE-75007 Uppsala, Sweden E-mail: rimvys.vasaitis@mykopat.slu.se
• Stenlid, Department of Forest Mycology & Pathology, Swedish University of Agricultural Sciences, P.O. Box 7026, SE-75007 Uppsala, Sweden E-mail: js@nn.se
• Thomsen, Forest & Landscape, University of Copenhagen, Hoersholm Kongevej 11, DK-2970 Hoersholm, Denmark E-mail: imt@nn.dk
• Barklund, Department of Forest Mycology & Pathology, Swedish University of Agricultural Sciences, P.O. Box 7026, SE-75007 Uppsala, Sweden E-mail: pb@nn.se
• Dahlberg, Swedish Species Information Centre, Swedish University of Agricultural Sciences, P.O. Box 7007, SE-75007 Uppsala, Sweden E-mail: ad@nn.se

• Bergeron, NSERC-UQAT-UQAM Industrial Chair in Sustainable Forest Management, C.P. 8888, Succursale Centre-ville, Montréal, Québec, Canada H3C 3P8 E-mail: bergeron.yves@uqam.ca
• Leduc, NSERC-UQAT-UQAM Industrial Chair in Sustainable Forest Management, C.P. 8888, Succursale Centre-ville, Montréal, Québec, Canada H3C 3P8 E-mail: al@nn.ca
• Harvey, NSERC-UQAT-UQAM Industrial Chair in Sustainable Forest Management, C.P. 8888, Succursale Centre-ville, Montréal, Québec, Canada H3C 3P8 E-mail: bdh@nn.ca
• Gauthier, NSERC-UQAT-UQAM Industrial Chair in Sustainable Forest Management, C.P. 8888, Succursale Centre-ville, Montréal, Québec, Canada H3C 3P8 E-mail: sg@nn.ca

• Kuuluvainen, Department of Forest Ecology, P.O. Box 24, FIN-00014 University of Helsinki, Finland E-mail: timo.kuuluvainen@helsinki.fi

• Edenius, SLU, Department of Animal Ecology, SE-901 83 Umeå, Sweden E-mail: lars.edenius@szooek.slu.se
• Bergman, SLU, Department of Animal Ecology, SE-901 83 Umeå, Sweden E-mail: mb@nn.se
• Ericsson, SLU, Department of Animal Ecology, SE-901 83 Umeå, Sweden E-mail: ge@nn.se
• Danell, SLU, Department of Animal Ecology, SE-901 83 Umeå, Sweden E-mail: kd@nn.se

• Gromtsev, Forest Research Institute, Karelian Research Centre of the Russian Academy of Sciences, P.O. Box 185610, Petrozavodsk, Pushkin st. 11, Russia E-mail: gromtsev@karelia.ru

• Ryan, USDA Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory, P.O. Box 8089, Missoula, Montana 59807, USA E-mail: kryan@fs.fed.us

In managed European hemiboreal forests, windstorms have a notable ecological and socio-economic impact. In this study, stand properties affecting windstorm damage occurrence at the stand-level were assessed using a Generalized Linear Mixed model. After 2005 windstorm, 5959 stands dominated by birch (Betula spp.), Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.), with mean height > 10 m were inventoried. Windstorm damage was positively associated with spruce and pine-dominated stands, increasing mean height, fresh forest edges, decreasing time since the last thinning and stronger wind gusts. Tree species composition – mixed or monodominant – was not statistically significant in the model; while, the admixture of spruce in the canopy layer was positively associated with higher windstorm damage. Stands on peat soils were more damaged than stands on mineral soils. Birch stands were more damaged than pine stands. This information could be used in forest management planning, selection of silvicultural treatments to increase forest resilience to natural disturbances.

• Donis, Latvian State Forest Research Institute “Silava”, Rigas 111, Salaspils, Latvia, LV-2169 E-mail: janis.donis@silava.lv
• Kitenberga, Latvian State Forest Research Institute “Silava”, Rigas 111, Salaspils, Latvia, LV-2169 E-mail: mara.kitenberga@gmail.com
• Šņepsts, Latvian State Forest Research Institute “Silava”, Rigas 111, Salaspils, Latvia, LV-2169 E-mail: guntars.snepsts@silava.lv
• Dubrovskis, Latvia University of Life Sciences and Technologies, Liela iela 2, Jelgava, Latvia, LV-3001 E-mail: edgars.dubrovskis@llu.lv
• Jansons, Latvian State Forest Research Institute “Silava”, Rigas 111, Salaspils, Latvia, LV-2169 E-mail: aris.jansons@silava.lv

Forest fire is one of the natural disturbances, which have important ecological and socioeconomical effect. Although fire activity is driven by weather conditions, during past two centuries forest fires have been strongly anthropogenically controlled. In this study, teleconnection between sea surface temperature (SST) in the Atlantic, which influences climate in Europe, and forest fire activity in Latvia and Estonia was assessed using “Climate explorer” web-tool. Factors affecting number and area of forest fires in Latvia and Estonia differed, suggesting regional specifics. In Estonia, the number of fires correlated with the SST in the North Atlantic in spring and summer, which affects the inflow of cool and dry air masses from the Arctic, hence the aridity and burnability. The area of fires in Estonia and in Latvia was associated with increased SST in Baltic Sea and near the European coast in summer, which likely were consequences of occurrence of warm high-pressure systems in summer, causing hot and dry conditions. Nevertheless, the observed teleconnections could be used to predict activity of forest fires in Latvia and Estonia.

• Kitenberga, Latvian State Forest Research Institute ‘Silava’, Rigas st. 111, Salaspils, Latvia, LV2169 E-mail: mara.kitenberga@gmail.com
• Matisons, Latvian State Forest Research Institute ‘Silava’, Rigas st. 111, Salaspils, Latvia, LV2169 E-mail: robism@inbox.lv
• Jansons, Latvian State Forest Research Institute ‘Silava’, Rigas st. 111, Salaspils, Latvia, LV2169 E-mail: aris.jansons@silava.lv
• Donis, Latvian State Forest Research Institute ‘Silava’, Rigas st. 111, Salaspils, Latvia, LV2169 E-mail: janis.donis@silava.lv

Long-term (47 years) effect of experimental whole tree harvesting (WTH) with a heavy soil scarification on ground cover vegetation was assessed in a dry nutrient-poor Scots pine (Pinus sylvestris L.) stand in Latvia. Neighbouring conventionally managed young (10 years) and mature (119 years) stands of the same type were used for comparison. Higher species richness was observed in the WTH stand compared to conventionally managed young and mature stands (24, 18 and 16 species, respectively), likely due to the profound disturbance. The Shannon diversity index was higher in the young than in the WTH and mature stands (2.36, 1.77 and 1.63, respectively); still, the composition and structure of ground cover vegetation in WTH was more similar to the mature stand. Nevertheless, the occurrence of oligotrophic species in the WTH stand suggested decreased soil nutrient content and potential development of different plant community. Hence, such method might be considered for restoration of oligotrophic stands. Nevertheless, the period of 47 years appeared sufficient for the ground cover vegetation to recover after the WTH.

• Jansons, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: aris.jansons@silava.lv
• Robalte, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: robalte.l@gmail.com
• Čakšs, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: chakijs95@gmail.com
• Matisons, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: roberts.matisons@silava.lv

• Kuuluvainen, Department of Forest Ecology, University of Helsinki, P.O.Box 27 FIN-00014, Finland E-mail: timo.kuuluvainen@helsinki.fi
• Aapala, Finnish Environment Institute, Expert Services Department, Nature Division, P.O. Box 140, FIN-00251 Helsinki E-mail: ka@nn.fi
• Ahlroth, University Museum, Section of Natural History, P.O. Box 35, FIN-40351, Jyväskylä, Finland E-mail: pa@nn.fi
• Kuusinen, Ministry of the Environment, Land Use Department, P.O.Box 380, FIN-00131 Helsinki, Finland E-mail: mk@nn.fi
• Lindholm, Finnish Environment Institute, Expert Services Department, Nature Division, P.O. Box 140, FIN-00251 Helsinki E-mail: tl@nn.fi
• Sallantaus, Pirkanmaa Regional Environment Centre, P.O. Box 297, FIN-33101 Tampere, Finland E-mail: ts@nn.fi
• Siitonen, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: juha.siitonen@metla.fi
• Tukia, Finnish Environment Institute, Expert Services Department, Nature Division, P.O. Box 140, FIN-00251 Helsinki E-mail: ht@nn.fi

• DeLong, Ministry of Forests, 1011 4th Ave, Prince George, BC, Canada V2L 3H9 E-mail: craig.delong@gems1.gov.bc.ca

• Jentsch, Conservation Biology and Ecological Modelling, UFZ – Centre for Environmental Research, Permoserstr. 15, D-04301 Leipzig, Germany E-mail: jentsch@pro.ufz.de
• Beierkuhnlein, Department of Landscape Ecology, University of Rostock, Justus-Liebig-Weg 6, D-18051 Rostock, Germany E-mail: cb@nn.de
• White, Department of Biology, Campus Box 3280, University of North Carolina at Chapel Hill, Chapel Hill, NC 27599-3280, USA E-mail: psw@nn.us

• Kuuluvainen, Department of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: timo.kuuluvainen@helsinki.fi