article id 377,
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                            Stand structural characteristics were examined in old Pinus sylvestris  dominated sites in three regions along a broad geographic and human  impact gradient in mid-boreal Fennoscandia. The study regions were: 1)  Häme in south-western Finland, with a long history of forest  utilization, 2) Kuhmo in north-eastern Finland, with a more recent  history of intensive forest utilization, and 3) Vienansalo in Russian  Karelia, still characterized by a large near-natural forest landscape.  Within each region the sampled sites were divided into three human  impact classes: 1) near-natural stands, 2) stands selectively logged in  the past, and 3) managed stands treated with thinnings. The near-natural  and selectively logged stands in Häme and Kuhmo had a significantly  higher Picea proportion compared to stands in Vienansalo. In comparison,  the proportions of deciduous tree volumes were higher in near-natural  stands in Vienansalo compared to near-natural stands in Häme. The pooled  tree diameter distributions, both in near-natural and selectively  logged stands, were descending whereas managed stands had a bimodal  diameter distribution. Structural diversity characteristics such as  broken trunks were most common in near-natural stands and in stands  selectively logged in the past. The results demonstrate the higher  structural complexity of near-natural stands and stands selectively  logged in the past compared to managed stands, and highlight that old  near-natural stands and stands selectively logged in the past vary  widely in their structures. This obviously reflects both their natural  variability but also various combinations of pre-industrial land use and  human impact on fire disturbance. These factors need to be acknowledged  when using “natural” forest structures as a reference in developing  strategies for forest management, restoration and nature conservation.
                        
                
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                            Lilja,
                            University of Helsinki, Department of Forest Ecology, P.O. Box 27, FI-00014 University of Helsinki, Finland
                                                        E-mail:
                                                            saara.lilja@helsinki.fi
                                                                                        
                                                     
                                            - 
                            Kuuluvainen,
                            University of Helsinki, Department of Forest Ecology, P.O. Box 27, FI-00014 University of Helsinki, Finland
                                                        E-mail:
                                                            tk@nn.fi