Articles by Juho Matala

The moose (Alces alces L.), a common large herbivore in the boreal region, impairs forest regeneration by browsing on tree seedlings and saplings. Moose prefer deciduous species, but during winter more coniferous seedlings are used. We used meta-analyses, separately for deciduous and coniferous seedlings, for evaluating whether excluding moose browsing affected seedling density and height. In addition, we compared (1) deciduous seedling proportion, (2) stand density, (3) elapsed time from fencing and (4) estimated moose density with moose exclusion effect sizes. Fencing had a positive effect on coniferous seedling height. With more deciduous trees in a seedling stand, the fencing effect for both seedling height and density of coniferous seedlings decreased. On the other hand, the fencing effects increased with denser stands. At some point effect sizes turned to negative, and conifer species varied in their response to browsing. This implies that deciduous seedlings can protect conifers from browsing by moose up to some mixing ratio, but when deciduous seedling densities are too high, their negative effect increases, presumably through increased competition. Our results suggest that a moderate deciduous admixture in conifer-dominated mixed seedling stands can decrease moose damage but also underline the significance of timely silvicultural measures to minimize the negative effects of excessive deciduous seedlings and too dense stands. Due to differences in coniferous and deciduous species, as well as their compositions and amounts in studied experiments, more studies adjusted to local conditions are still needed to give exact measures for silvicultural recommendations.

• Domisch, Natural Resources Institute Finland (Luke), Natural Resources, Yliopistokatu 6B, FI-80100 Joensuu, Finland https://orcid.org/0000-0001-7026-1087 E-mail: timo.domisch@luke.fi
• Huuskonen, Natural Resources Institute Finland (Luke), Natural Resources, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: saija.huuskonen@luke.fi
• Matala, Natural Resources Institute Finland (Luke), Natural Resources, Yliopistokatu 6B, FI-80100 Joensuu, Finland https://orcid.org/0000-0002-5867-5057 E-mail: juho.matala@luke.fi
• Nikula, Natural Resources Institute Finland (Luke), Natural Resources, Ounasjoentie 6, FI-96200 Rovaniemi, Finland https://orcid.org/0000-0001-8372-8440 E-mail: ari.nikula@luke.fi

An important modifier of forests and forestry practices is browsing by cervids. As high populations of moose (Alces alces L.) cause extensive forest damage in the Fennoscandian boreal forests, models should be able to predict the susceptibility of projected forest structures to browse damage. We augmented the European Forestry Dynamics Model (EFDM) for the area of seedling stands damaged by moose. The augmented model was tested in projecting both forest resources and moose damage for 18 million hectares of forest land in Finland, based on input data from the National Forest Inventory (NFI). Modeling the area of seedling stands damaged as a function of moose population density, forest characteristics, and region-specific interactions of these variables was found to work realistically for 30 years, predicting that the area of seedling stands damaged by moose would increase by up to a third from the last NFI observation. Our work lays the groundwork for modeling consequential, large-scale ecological and socio-economic effects of moose browsing.

• Vauhkonen, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, FI-80101 Joensuu, Finland; University of Helsinki, Department of Forest Sciences, Latokartanonkaari 7, FI-00014 Helsingin yliopisto, Finland E-mail: jari.vauhkonen@uef.fi
• Matala, Natural Resources Institute Finland (Luke), Natural resources, Yliopistokatu 6 B, FI-80100 Joensuu, Finland E-mail: juho.matala@luke.fi
• Nikula, Natural Resources Institute Finland (Luke), Natural resources, Ounasjoentie 6, FI-96200 Rovaniemi, Finland https://orcid.org/0000-0001-8372-8440 E-mail: ari.nikula@luke.fi

Traditional timber production may have negative effects on other ecosystem services. Therefore, new forest management guidelines have been developed in order to enhance a habitat suitable for wildlife. In Finland, a recent example of this is grouse-friendly forest management (GFFM) which emphasises the preservation of grouse species (Tetronidae) habitats. This study aimed to analyse the economic effects of these guidelines. An analysis was made on how the application of GFFM affected the Net Present Value (NPV) in a 30-year simulation of forest management of four large forest holdings located from south to north in Finland. In the simulations, traditional forest management practices were compared to two levels of GFFM. Five levels of interest rate were used, namely 1, 2, 3, 4, and 5%. In most of the simulations, the NPV was reduced by about 1% or less due to the application of GFFM in comparison to the traditional reference forest management. Only in one case with more intensive GFFM, was the reduction of NPV more than 5%. The interest rates had an impact on the differences between the management approaches. For example, a low interest rate resulted in a higher thinning intensity in GFFM in comparison to traditional forest management, which lead to a higher NPV in GFFM. To sum up, it seems that it would be possible to manage forest holdings in a grouse-friendly manner with minor effects on the economics.

• Haara, LUKE https://orcid.org/0000-0002-6895-5300 E-mail: arto.haara@luke.fi
• Matala, Natural Resources Institute Finland, (Luke), Natural resources, Yliopistokatu 6B, FI-80100 Joensuu, Finland E-mail: juho.matala@luke.fi
• Melin, Natural Resources Institute Finland, (Luke), Bioeconomy and environment, Yli opistokatu 6B, FI-80100 Joensuu, Finland E-mail: markus.melin@luke.fi
• Miettinen, Finnish Wildlife Agency, Ratatie 41, FI-91501 Muhos, Finland E-mail: janne.miettinen@riista.fi
• Korhonen, Natural Resources Institute Finland, (Luke), Bioeconomy and environment, Yli opistokatu 6B, FI-80100 Joensuu, Finland E-mail: kari.t.korhonen@luke.fi
• Packalen, Ministry of Agriculture and Forestry, P.O. Box 30, FI-00023 GOVERNMENT, Finland E-mail: tuula.packalen@mmm.fi
• Varjo, Finnish Wildlife Agency, Sompiontie 1, FI-00730 Helsinki, Finland E-mail: jari.varjo@riista.fi

Moose (Alces alces L.) browsing causes severe damage in Scots pine (Pinus sylvestris L.) seedling stands. The effects of this damage on the quality of sawlogs were studied in a long-term controlled experiment. This article reports the stem size and external quality characteristics of Scots pine stems 34 years after artificial moose browsing damage. Damaging the trees by clipping the main stem at the seedling stage reduced the diameter, height, and tree volume of the trees at the end of the experiment. The tree growth reduction was dependent on the severity of clipping. The differences between the damaged and the control trees were more obvious in diameter than in height at the time of final felling. Stem form defects and vertical branches were the most typical externally detectable defects caused by clipping. Defects in the butt logs were detected in 71–89% of the damaged trees, depending on the clipping treatment severity. The stronger the clipping treatment, the more likely the stem form was defected and the more commonly were vertical branches and crooks detected in the stems. The results indicate that both tree dimensions and stem quality suffer from moose browsing. The findings of this controlled experiment more likely underestimate than overestimate the damage in comparison to real moose browsing. Further analyses are required to assess the effects of browsing damage on the internal quality of sawlogs and subsequent economic outcomes.

• Matala, Natural Resources Institute Finland (Luke), Natural resources, Yliopistokatu 6B, FI-80101 Joensuu, Finland E-mail: juho.matala@luke.fi
• Kilpeläinen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Yliopistokatu 6B, FI-80101 Joensuu, Finland E-mail: harri.kilpelainen@luke.fi
• Heräjärvi, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6B, FI-80101 Joensuu, Finland E-mail: henrik.herajarvi@luke.fi
• Wall, Natural Resources Institute Finland (Luke), Research infrastructure services, Yliopistokatu 6B, FI-80101 Joensuu, Finland E-mail: tapio.wall@luke.fi
• Verkasalo, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6B, FI-80101 Joensuu, Finland E-mail: erkki.verkasalo@luke.fi

We modelled the effect of habitat composition and roads on the number and occurrence of moose (Alces alces L.) damage in Ostrobothnia and Lapland using a zero-inflated count model. Models were developed for 1 km², 25 km² and 100 km² landscapes consisting of equilateral rectangular grid cells. Count models predict the number of damage, i.e. the number of plantations and zero models the probability of a landscape being without damage for a given habitat composition. The number of moose damage in neighboring grid cells was a significant predictor in all models. The proportion of mature forest was the most frequent significant variable, and an increasing admixture of mature forests among plantations increased the number and occurrence of damage. The amount of all types of plantations was the second most common significant variable predicting increasing damage along with increasing amount of plantations. An increase in thinning forests as an admixture also increased damage in 1 km² landscapes in both areas, whereas an increase in pine-dominated thinning forests in Lapland reduced the number of damage in 25 km² landscapes. An increasing amount of inhabited areas in Ostrobothnia and the length of connecting roads in Lapland reduced the number of damage in 1 and 25 km² landscapes. Differences in model variables between areas suggest that models of moose damage risk should be adjusted according to characteristics that are specific to the study area.

• Nikula, Natural Resources Institute Finland (Luke), Bioeconomy and Environment, Ounasjoentie 6, FI-96200 Rovaniemi, Finland E-mail: ari.nikula@luke.fi
• Nivala, Natural Resources Institute Finland (Luke), Bioeconomy and Environment, Ounasjoentie 6, FI-96200 Rovaniemi, Finland E-mail: vesa.nivala@luke.fi
• Matala, Natural Resources Institute Finland (Luke), Natural resources, Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: juho.matala@luke.fi
• Heliövaara, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: kari.heliovaara@helsinki.fi

There is evidence that moose are attracted to fertile growth habitats apparently due to better quality and larger quantities of food. The nutrients in mineral soils originate from the weathering of bedrock and the composition of parental bedrock affects the fertility of produced mineral soil, thus affecting also the import of nutrients into the whole food web. We surveyed the connection between moose damage in forest plantations and the composition of bedrock and surficial deposits in Finnish Lapland. We used a database of compensated moose damage in private forests in years 1997−2010. Undamaged stands in National Forest Inventories (NFI) from years 1986–2008 served as a control data and moose-damaged NFI-stands as a reference data. Bedrock and surficial depositions and the location of studied stands in relation to ancient shorelines were explored by using the digital databases of the Geological Survey of Finland. Moose-damaged stands were concentrated in southwestern and east Lapland in the areas of the Peräpohja Schist Belt and Lapland’s Greenstone Belt that are both composed of nutrient-rich rocks. The bedrock of damaged stands contained a higher proportion of mafic and alkaline rocks than did the control stands. Moose-damaged stands were pine-dominated and grew in more fertile forest sites than did control stands. Part of pine stands probably located in soils formerly occupied by spruce, which may increase the stands’ vulnerability to biotic threats. Especially, there were relatively more moose damage in pine plantations regenerated on fine-grained mineral soils derived from nutrient rich rocks than in less fertile soils.

• Ruuhola, Natural Resources Institute Finland (Luke), Natural Resources and Bioproduction, Yliopistokatu 6, FI-80101 Joensuu, Finland; University of Eastern Finland, Faculty of Science and Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: teija.ruuhola@uef.fi
• Nikula, Natural Resources Institute Finland (Luke), Economics and Society, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: ari.nikula@luke.fi
• Vesa, Natural Resources Institute Finland (Luke), Economics and Society, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: vesa.nivala@luke.fi
• Nevalainen, Natural Resources Institute Finland (Luke), Natural Resources and Bioproduction, Yliopistokatu 6, FI-80101 Joensuu, Finland E-mail: seppo.nevalainen@luke.fi
• Matala, Natural Resources Institute Finland (Luke), Natural Resources and Bioproduction, Yliopistokatu 6, FI-80101 Joensuu, Finland E-mail: juho.matala@luke.fi

The occurrence of moose damage was studied using data from three National Forest Inventories (NFIs) accomplished between 1986 and 2008 in Finland. The combined data included a total of 97 390 young stands. The proportion of moose damage increased from 3.6% to 8.6% between the 8th NFI (1986–1994) and the 10th NFI (2004–2008). The majority (75%) of the damage occurred in Scots pine-dominated stands. The proportion of damage was higher in aspen-dominated stands than in stands dominated by any other tree species. The tree species mixture also had a clear effect on the occurrence of damage. Pure Scots pine stands had less damage than mixed Scots pine stands, and moose damage decreased linearly with the increasing proportion of Scots pine. Stands on mineral soil had more frequent moose damage than stands on peatlands. The fertility class of the site had no straightforward effect on the damage frequency. Artificially regenerated stands had more damage than naturally regenerated stands. Accomplished soil preparation measures and the need for thinning or clearing operations increased moose damage. High proportions of moose damage in young stands were found around the country. In the 10th NFI, the largest concentration of damage was found in southwestern Finland. Our study shows the temporal and spatial changes in the occurrence of moose damage and pinpoints some important silvicultural factors affecting the relative risk of young stands over a large geographical area.

• Nevalainen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: seppo.nevalainen@luke.fi
• Matala, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: juho.matala@luke.fi
• Korhonen, Natural Resources Institute Finland (Luke), Economics and society, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: kari.t.korhonen@luke.fi
• Ihalainen, Natural Resources Institute Finland (Luke), Economics and society, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: antti.ihalainen@luke.fi
• Nikula, Natural Resources Institute Finland (Luke), Economics and society, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: ari.nikula@luke.fi