Articles by Hannu Hökkä

Models for individual-tree basal area growth were constructed for Scots pine (Pinus sylvestris L.), pubescent birch (Betula pubescens Ehrh.) and Norway spruce (Picea abies (L.) Karst.) growing in drained peatland stands. The data consisted of two separate sets of permanent sample plots forming a large sample of drained peatland stands in Finland. The dependent variable in all models was the 5-year basal area growth of a tree. The independent tree-level variables were tree dbh, tree basal area, and the sum of the basal area of trees larger than the target tree. Independent stand-level variables were stand basal area, the diameter of the tree of median basal area, and temperature sum. Categorical variables describing the site quality, as well as the condition and age of drainage, were used. Differences in tree growth were used as criteria in reclassifying the a priori site types into new yield classes by tree species. All models were constructed as mixed linear models with a random stand effect. The models were tested against the modelling data and against independent data sets.

• Hökkä, E-mail: hh@mm.unknown
• Alenius, E-mail: va@mm.unknown
• Penttilä, E-mail: tp@mm.unknown

The use of random parameter models in forestry has been proposed as one method of incorporating different levels of information into prediction equations. By explicitly considering the variance-covariance structure of observations and considering some model parameters as random rather than fixed, one can incorporate more complex error structures in analysing data.

Competition indices and variance component techniques were applied to 92 Scots pine (Pinus sylvestris L.) -dominated permanent sample plots on drained peatlands in Northern Finland. By quantifying stand, plot, and tree level variation, it was possible to identify the level (stand, plot or tree) at which the explanatory variables contributed to the model. The replication of plots within stands revealed little variation among plots within a single stand but significant variation occurred at stand and tree levels. Positive and negative effects of inter-tree competition are identified by examining simple correlation statistics and the random parameter model.

• Penner, E-mail: mp@mm.unknown
• Hökkä, E-mail: hh@mm.unknown
• Penttilä, E-mail: tp@mm.unknown

The effect of drainage on structure of tree stands is analysed by comparing the average structural characteristics (e.g. diameter distribution) of stands in the data for different drainage age classes and selected site types. The material consists of ca. 4,400 relascope sample plots, which are part of a large drainage area inventory project. The uneven-aged structure of the virgin peatland forest is preserved for several decades after drainage. This is enhanced by the post-drainage increase of small-diameter trees, especially birch. The number of trees per hectare increased during a period of ca. 30 years and levelled off thereafter. The increase in the number of saw log stems is clearly related to the fertility of the site and its geographical location.

The PDF includes a summary in English.

• Hökkä, E-mail: hh@mm.unknown
• Laine, E-mail: jl@mm.unknown

We used a process-based hydrological model SUSI to improve guidelines for ditch network maintenance (DNM) operations on drained peatland forests. SUSI takes daily weather data, ditch depth, strip width, peat properties, and forest stand characteristics as input and calculates daily water table depth (WTD) at different distances from ditch. The study focuses on Scots pine (Pinus sylvestris L.) dominated stands which are the most common subjects of DNM. Based on a literature survey, and consideration of the tradeoffs between forest growth and detrimental environmental impacts, long term median July–August WTD of 0.35 m was chosen as a target WTD. The results showed that ditch depths required to reach such WTD depends strongly on climatic locations, stand volume, ditch spacing, and peat thickness and type. In typical ditch cleaning areas in Finland with parallel ditches placed about 40 m apart and tree stand volumes exceeding 45 m³ ha^–1, 0.3–0.8 m deep ditches were generally sufficient to lower WTD to the targeted depth of 0.35 m. These are significantly shallower ditch depths than generally recommended in operational forestry. The main collector ditch should be naturally somewhat deeper to permit water outflow. Our study brings a firmer basis on environmentally sound forestry on drained peatlands.

• Hökkä, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: hannu.hokka@luke.fi
• Laurén, University of Eastern Finland, Faculty of Science and Forestry, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: ari.lauren@uef.fi
• Stenberg, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: leena.stenberg@luke.fi
• Launiainen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: samuli.launiainen@luke.fi
• Leppä, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: kersti.leppa@luke.fi
• Nieminen, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: mika.nieminen@luke.fi

The effects of wood ash fertilisation on tree nutrition and growth on forested peatlands has been studied using loose ash, but in practice, ash fertilisation is done almost exclusively with granulated ash. In this study, the effects of granulated ash and loose ash (both 5 Mg ha^–1) on the growth and nutrition of Scots pine (Pinus sylvestris L.) stands were compared between a nitrogen-poor and a nitrogen-rich site over 15 years. On the nitrogen-rich site, wood ash application was also compared with commercial PK fertilisation. On the nitrogen-rich site, mean stand volume growth increase over unfertilised control treatment during the 15 year study period using granulated ash and commercial PK fertiliser was of the same magnitude (on average, 2.2–2.3 m³ ha^–1 a^–1). However, when loose ash was used growth increase over control was higher (3.7 m³ ha^–1 a^–1). On the nitrogen-poor site, the mean growth increase gained by loose or granulated ash (1.4–1.5 m³ ha^–1 a^–1) over the unfertilised control treatment was not significant. Fertilisation with loose ash or PK increased foliar P, K and B concentrations already in the first or second growing season, following fertilisation on both sites. Granulated ash increased foliar P concentrations on the nitrogen-rich site less than loose ash. After an initial increase, foliar P, K and B concentrations decreased at the end of study period. On the nitrogen-poor site, foliar P concentrations were below the deficiency limit by the end of the study period.

• Hytönen, Natural Resources Institute Finland, Natural Resources, Teknologiakatu 7, FI-67100 Kokkola, Finland https://orcid.org/0000-0001-8475-3568 E-mail: jyrki.hytonen@luke.fi
• Hökkä, Natural Resources Institute Finland, Natural Resources, Paavo Havaksentie 3, FI-90570 Oulu, Finland E-mail: hannu.hokka@luke.fi

The aim of this study was to develop individual-tree diameter and height growth models for Scots pine, Norway spruce, and pubescent birch growing in drained peatlands in Finland. Trees growing in peatland sites have growth patterns that deviate from that of trees growing in mineral soil sites. Five-year growth was explained by tree diameter, different tree and stand level competition measures, management operations and site characteristics. The drainage status of the site was influencing growth directly or in interaction with other variables. Site quality had a direct impact but was also commonly related to current site drainage status (need for ditch maintenance). Recent thinning increased growth of all species and former PK fertilization increased growth of pine and birch. Temperature sum was a significant predictor in all models and altitude for spruce and birch. The data were a subsample of the 7th National Forest Inventory (NFI) sample plots representing northern and southern Finland and followed by repeated measurements for 15–20 yrs. Growth levels predicted by the models were calibrated using NFI11 data to remove bias originating from the sample of the modelling data. The mixed linear models technique was used in model estimation. The models will be incorporated into the MOTTI stand simulator to replace the current peatlands growth models.

• Repola, Natural Resources Institute Finland (Luke), Natural resources, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: jaakko.repola@luke.fi
• Hökkä, Natural Resources Institute Finland (Luke), Natural resources, Paavo Havaksen tie 3, FI-90014 University OF Oulu, Finland E-mail: hannu.hokka@luke.fi
• Salminen, Natural Resources Institute Finland (Luke), Natural resources, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: hannu.salminen@luke.fi

The amount of water in peat soil is one factor affecting its bearing capacity, which is a crucial aspect in planning peatland timber harvesting operations. We studied the influence of weather variables on the variation of drained peatland growing season water conditions, here the ground water table depth (WTD). WTD was manually monitored four times in 2014 and three times in 2015 in 10–30 sample plots located in four drained peatland forests in south-western Finland. For each peatland, precipitation and evapotranspiration were calculated from the records of the nearest Finnish Meteorological Institute field stations covering periods from one day to four weeks preceding the WTD monitoring date. A mixed linear model was constructed to investigate the impact of the weather parameters on WTD. Precipitation of the previous four–week period was the most important explanatory variable. The four-week evapotranspiration amount was interacting with the Julian day showing a greater effect in late summer. Other variables influencing WTD were stand volume within the three-metre radius sample plot and distance from nearest ditch. Our results show the potential of weather parameters, specifically that of the previous four-week precipitation and evapotranspiration, for predicting drained peatland water table depth variation and subsequently, the possibility to develop a more general empirical model to assist planning of harvesting operations on drained peatlands.

• Hökkä, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Paavo Havaksen tie 3, FI-90014 Oulun yliopisto, Finland E-mail: hannu.hokka@luke.fi
• Uusitalo, Natural Resources Institute Finland (Luke), Green Technology, Kaironiementie 15, FI-39700 Parkano, Finland E-mail: jori.uusitalo@luke.fi
• Lindeman, Natural Resources Institute Finland (Luke), Green Technology, Kaironiementie 15, FI-39700 Parkano, Finland E-mail: harri.lindeman@luke.fi
• Ala-Ilomäki, Natural Resources Institute Finland (Luke), Green Technology, Jokiniemenkuja 1, FI-01370 Vantaa, Finland E-mail: jari.ala-ilomaki@luke.fi

The long-term effects of fertilization on the needle nutrient concentrations, growth and financial performance of a Scots pine (Pinus sylvestris L.) stand was examined in a thick-peated drained peatland forest located in Central Finland. At the trial establishment in 1985, the trees were suffering from P and K deficiencies, but their N status was good. The fertilizer treatments were Control, PK (rock phosphate + potassium chloride), ApaBio (apatite phosphorus + biotite) and wood ash, applied both with and without N and replicated six times. All treatments containing phosphorus and potassium increased foliar P and K concentrations above the deficiency limits up to the end of the study period of 26 years. The effect of the fertilization on stand volume growth of Scots pine was strong and continued still at the end of the study period. The trees on ApaBio and PK plots grew nearly two-fold and those on Ash plots over two-fold compared with the control plots. In a thinning made at the end of the study period the total logging removal on fertilized plots was 1.5–2.2 times greater and included more saw logs than on the control plots. Ash fertilizer treatment outperformed other fertilizer treatments as well as the control. With a 5% discounted equivalent annual income (EAI) of Ash fertilizer treatment was statistically significantly (p=0.009) almost three times higher than that of control. As a conclusion, fertilization (either using PK fertilizers or Ash) in N-rich drained peatlands is a financially feasible method of management.

• Moilanen, Natural Resources Institute Finland, Natural resources and bioproduction, Paavo Havaksen tie 3, FI-90014 Oulu, Finland E-mail: mikko.moilanen@luke.fi
• Hytönen, Natural Resources Institute Finland, Natural resources and bioproduction, Silmäjärventie 2, FI-69100 Kannus, Finland E-mail: jyrki.hytonen@luke.fi
• Hökkä, Natural Resources Institute Finland, Natural resources and bioproduction, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: hannu.hokka@luke.fi
• Ahtikoski, Natural Resources Institute Finland, Natural resources and bioproduction, Paavo Havaksen tie 3, FI-90014 Oulu, Finland E-mail: anssi.ahtikoski@luke.fi

• Hökkä, Finnish Forest Research Institute, Rovaniemi Unit, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: hannu.hokka@metla.fi
• Mäkelä, Lapland University of Applied Sciences, Jokiväylä 11 C, FI-96300 Rovaniemi, Finland E-mail: hanna.makela@edu.ramk.fi

• Hökkä, Finnish Forest Research Institute, Rovaniemi Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: hannu.hokka@metla.fi
• Repola, Finnish Forest Research Institute, Rovaniemi Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: jaakko.repola@metla.fi
• Moilanen, Finnish Forest Research Institute, Oulu Unit, P.O. Box 413, FI-90014 University of Oulu, Finland E-mail: mikko.moilanen@metla.fi
• Saarinen, Finnish Forest Research Institute, Parkano Unit, Kaironiementie 15, FI-39700 Parkano, Finland E-mail: markku.saarinen@metla.fi

• Hökkä, Finnish Forest Research Institute, Rovaniemi Research Unit, FI-96301 Rovaniemi, Finland E-mail: hannu.hokka@metla.fi
• Repola, Finnish Forest Research Institute, Rovaniemi Research Unit, FI-96301 Rovaniemi, Finland E-mail: jr@nn.fi
• Moilanen, Finnish Forest Research Institute, Muhos Research Unit, Muhos, Finland E-mail: mm@nn.fi
• Saarinen, Finnish Forest Research Institute, Parkano Research Unit, Parkano, Finland E-mail: ms@nn.fi

• Hökkä, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: hannu.hokka@metla.fi
• Hyttinen, Metsänhoitoyhdistys Keski-Suomi, Viitasaari, Finland E-mail: hh@nn.fi
• Marttila, University of Oulu, Water Resources and Environmental Engineering Lab, Oulu, Finland E-mail: hm@nn.fi
• Jämsen, Forestry Centre Keski-Suomi, Pihtipudas, Finland E-mail: jj@nn.fi
• Kløve, University of Oulu, Water Resources and Environmental Engineering Lab, Oulu, Finland E-mail: bk@nn.fi

• Repola, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: jaakko.repola@metla.fi
• Hökkä, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: hh@nn.fi
• Penttilä, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: tp@nn.fi

• Sarkkola, University of Helsinki, Department of Forest Ecology, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: sakari.sarkkola@helsinki.fi
• Hökkä, Finnish Forest Research Institute, Rovaniemi Research Station, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: hh@nn.fi
• Penttilä, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: tp@nn.fi

• Jutras, Département des sciences du bois et de la forêt, Université Laval, Ste-Foy, Québec, G1K 7P4, Canada E-mail: sj@nn.ca
• Hökkä, Finnish Forest Research Institute, Rovaniemi Research Station, P.O. Box 16, FIN 96301 Rovaniemi, Finland E-mail: hannu.hokka@metla.fi
• Alenius, Finnish Forest Research Institute, Rovaniemi Research Station, P.O. Box 16, FIN 96301 Rovaniemi, Finland E-mail: va@nn.fi
• Salminen, Finnish Forest Research Institute, Rovaniemi Research Station, P.O. Box 16, FIN 96301 Rovaniemi, Finland E-mail: hs@nn.fi

• Minkkinen, University of Helsinki, Department of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: kari.minkkinen@helsinki.fi
• Laine, University of Helsinki, Department of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: jl@nn.fi
• Hökkä, Finnish Forest Research Institute, Rovaniemi Research Station, P.O. Box 16, FIN-96301 Rovaniemi, Finland E-mail: hh@nn.fi

• Nuutinen, Finnish Forest Research Institute, Joensuu Research Station, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: tuula.nuutinen@metla.fi
• Hirvelä, Finnish Forest Research Institute, Joensuu Research Station, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: hh@nn.fi
• Hynynen, Finnish Forest Research Institute, Joensuu Research Station, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: jh@nn.fi
• Härkönen, Finnish Forest Research Institute, Joensuu Research Station, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: kh@nn.fi
• Hökkä, Finnish Forest Research Institute, Joensuu Research Station, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: hh@nn.fi
• Korhonen, Finnish Forest Research Institute, Joensuu Research Station, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: ktk@nn.fi
• Salminen, Finnish Forest Research Institute, Joensuu Research Station, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: os@nn.fi

• Hökkä, Finnish Forest Research Institute, Rovaniemi Research Station, PO Box 16, 96301 Rovaniemi, Finland E-mail: hannu.hokka@metla.fi
• Penttilä, Finnish Forest Research Institute, Rovaniemi Research Station, PO Box 16, 96301 Rovaniemi, Finland E-mail: tp@nn.fi

At sites with either peat or mineral soils in large areas of boreal forests, high soil-water contents hamper tree growth and drainage can significantly increase growth. Hence, areas covering about 15 × 10⁶ ha of northern peatlands and wet mineral soils have been drained for forestry purposes. Usually ditches gradually deteriorate, thus reducing their functionality as drains, and ditch-network maintenance (DNM) might be needed to maintain stand growth rates enabled by the original ditching. This article reviews current knowledge on establishing the need for DNM in boreal forest stands, subsequent growth responses, and the financial outcome of the activity. The issues covered in the review are: (i) ditching, changes in ditches over time and the need for DNM; (ii) interactions between soil water and both stand properties and stand management; (iii) ground-water level (GWL) and tree growth responses to DNM; and (iv) financial viability of DNM. Conclusions about the current understanding of issues related to DNM are drawn and implications for DNM in practice are summarized. Finally, gaps in knowledge are identified and research needs are suggested.

• Sikström, The Forestry Research Institute of Sweden (Skogforsk), Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: ulf.sikstrom@skogforsk.se
• Hökkä, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: hannu.hokka@luke.fi