Articles by Pentti K. Räsänen

A third generation of forest tree gas exchange measuring system design for the use in the field is described. The system is designed to produce data for determining the dependence of the rate of tree photosynthesis, respiration and transpiration on environmental factors. The system consists of eight cuvettes, a tubing system, two infrared gas analysers, an air flow controller, a data logger, and a computer. The measuring cuvette is a clap type, i.e. it is mostly open, only closing during measurement. CO₂ exchange is measured as the change in the cuvette concentration of CO₂, and, transpiration is measured as the increase in water vapour concentration while the cuvette is closed. The environmental factors measured are temperature, irradiance and air pressure. The system was planned in 1987 and constructed in 1988. It worked reliably in late summer 1988 and the quality of data seems to be satisfactory. 

The PDF includes an abstract in Finnish.

• Hari, E-mail: ph@mm.unknown
• Korpilahti, E-mail: eeva.korpilahti@luke.fi
• Pohja, E-mail: tp@mm.unknown
• Räsänen, E-mail: pr@mm.unknown

About 4,000 seedlings of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) were planted in 1965 both on a clear-cut and sheltered area in Central Finland. In the autumn of 1966 needle colour was determined by using Muncell Color Charts which allowed a quantitative measurement of three colour dimensions (hue, value, and chroma). Terminal shoot growth was recorded for two years after colour measurements. In both species, fertilization (NPK in the spring of the year of colour measurement) as well as other site factors caused differences in all three dimensions of needle colour. A regression of shoot growth on needle colour was found in both species. In most cases colour value (darkness) and, in spruce, also chroma, predicted the subsequent growth almost as well as did these two-colour variables together.

The PDF includes a summary in English.

• Luukkanen, E-mail: ol@mm.unknown
• Räsänen, E-mail: pr@mm.unknown
• Yli-Vakkuri, E-mail: py@mm.unknown

Seedlings of three different Scots pine (Pinus sylvestris L.) nursery stock, 1+0 ,1+1, and 2+0, were kept over the winter, after they had been packed in polythene bags, in three different ways: 1) In a refrigerated storage room, 2) in a wooden crate in the ground, 3) submerged in a lake. The seedling to which they were to be compared with were left over the winter in a nursery bed. The 1,800 seedlings were planted out in the spring 1966 in 15 random blocks. Their development was scrutinized during the three subsequent falls.

The seedlings which had been stored in the lake all died. The seedlings which had been stored along the 1st and 2nd method, managed almost as well as the ones which had been kept over the winter in the nursery bed, except for those of 1+0 stock.

The PDF includes a summary in English.

• Räsänen, E-mail: pr@mm.unknown
• Hänninen, E-mail: th@mm.unknown

Scots pine (Pinus sylvestris L.) seedlings were stored for five days in an ordinary wood shed. One half of the seedlings were planted out directly, and another half after soaking the roots of the seedlings for 3–6 hours in water to compensate the possible water deficit developed. According to the results of the experiment, the effect of watering was extremely small. The difference observed, which was in favour of the trees that had been watered during storage, was discernible only in the needle length and in the number of lateral buds; in mortality or in the growth of the seedlings no difference could be observed.

The PDF includes a summary in English.

• Räsänen, E-mail: pr@mm.unknown
• Hiltunen, E-mail: mh@mm.unknown

This paper reports spot sowing experiments of Scots pine (Pinus sylvestris L.). The seeds were either covered with coarse sand, tramped in the substrate or sowed without any covering, 30 seeds in each treatment in 70 replications. The site was of Vaccinium type with sandy soil. The germination percentage was 81 and 91 on the respective years. The development of seedlings was observed for 3–4 years.

The results indicate that both tramping and covering the seeds to some extent increased the number of seedlings and improved the early development. The highest numbers of seedlings were recorded in the first growing season, after which there was 23 seedlings/100 seeds in the uncovered spots, 27 seedlings in the covered spots and 31 seedlings in the tramped spots in the experiment sowed in 1965.

Mortality of the seedlings was highest between the first and second growing season, and empty spots increased with the time. There was no difference in mortality between the sowing methods, but the number of seedlings after first growing season affected the result. Under favourable conditions four seedlings per spot seemed enough to secure the survival of minimum one seedling per spot during the three first growing seasons. In poor conditions seven seedlings was needed.

The PDF includes a summary in English.

• Yli-Vakkuri, E-mail: py@mm.unknown
• Räsänen, E-mail: pr@mm.unknown

The aim of the present study was to establish, by means of planting experiments, the influence of different packing, heeling-in and watering as well as the length of the storage period on the development of Scots pine (Pinus sylvestris L.) seedlings, in all 2,090 seedlings, that had been lifted from the nursery bed in spring. The plants were packed in bundles and into plastic sacks in 1965 (6 storage methods) and in 1966 (3 storage methods). Control seedlings were planted without storing at the time when storage of the test material begun. The plantations were followed 3–4 years.

Storage for two weeks in the different ways and planting without storage gave similar results when seedling survival was compared. Storage in plastic sack proved to be as good as storage in bundles in a cellar, and healing-in in moist soil or in a drain were both usable methods. Watering the seedlings did not improve the results, which indicates that the storage caused no serious lack of water.

After four growing seasons an average of 19,6% of the seedlings of the 1965 experiment died, the bulk of them by the end of the first growing season. Despite control treatment, Hylobious abietis caused serious damages. In the plantations of the year 1966 mortality of the seedlings was under 5% by the end of third growing season. During the first two growing seasons after planting differences in growth of the seedlings stored in different ways could be observed in the plantations of the year 1965, but the differences levelled out later. In the plantations established in 1966 no differences in growth occurred.

The PDF includes a summary in English.

• Räsänen, E-mail: pr@mm.unknown
• Koukkula, E-mail: ak@mm.unknown
• Yli-Vakkuri, E-mail: py@mm.unknown

In the experiment Scots pine (Pinus sylvestris L.) seedlings were transplanted out in the field. The effect of the treatments on gas metabolism and daily height increment were examined. The seedlings were 5-year old Scots pine plants growing in clay pots, covered with plastic bags. Transpiration and photosynthetic rates were monitored with open IRGA measuring system for a few days before being subjected to the treatments and for one month after. In addition, the daily amounts of transpired water and daily height increments were measured. A model for the potential rate of each metabolic process was constructed.

Planting and additional exposure had a strong and rather permanent effect on the self-regulation of the processes. This effect is very similar to that caused by water deficit. Exposure makes the disturbance more pronounced. Transpiration of the transplanted seedlings decreased in a few days after planting to less than half of the potential value and that of the exposed ones decreased to a quarter of the potential value. The daily amounts of photosynthesis decreased to half of the potential value. There was no recovery in photosynthesis during the whole monitoring period of four weeks. There was a slight recovery in transpiration about five weeks after transplanting.

Thus, the treatment probably generated stress conditions throughout the whole growing period, which is characterized by strong self-regulation of photosynthesis and transpiration, thus causing an essential decrease in the total amount of CO₂ fixed. The photosynthesis was depressed especially at elevated temperatures after planting, as during water deficit. Planting and additional exposure did not produce any detectable changes in the dependence of the growth rate on temperature or in the effect of self-regulation on height growth. On the other hand, the level of growth was decreased as a result of planting out.

The PDF includes a summary in Finnish.

• Hallman, E-mail: eh@mm.unknown
• Hari, E-mail: ph@mm.unknown
• Räsänen, E-mail: pr@mm.unknown
• Smolander, E-mail: hs@mm.unknown

This study was carried out in 1966 in the nursery at Hyytiälä, Korkeakoski Forest District, in Southern Finland. The influence of lifting date (two liftings), way of packing (two methods) and length of storage (one, three and six weeks) on the development of Scots pine (Pinus sylvestris L.), 2 + 1, during the four years following planting was assessed. On the seedlings stored for six weeks, the influence of compensating for the water deficit was also studied.

According to the results, the lifting carried out later, i.e. when the seedlings had already started growth, gave slightly better results than when seedlings were lifted earlier. No difference could be observed for seedlings stored for one week, but for the seedlings stored for three or six weeks, mortality in the lot lifted earlier was 6- to 7-fold that of the seedlings lifted later. The main reason for this was probably that the seedlings of the earlier lifting suffered from lack of water at the time of lifting.

The growth of the seedlings lifted earlier and stored for three weeks showed a decrease compared to those lifted later. For the seedlings stored for six weeks, on the other hand, faster growth was recorded for both the seedlings of the earlier and the later lifting in comparison with those stored for shorter times. Watering increased to some extent the growth of the seedlings stored for six weeks.

During the normal, one- and three-week storing periods, seedlings were well preserved when packed both in bundles and in polythene sacks. Three years after the planting the average mortality was about 10%. Effect of watering was large for those seedlings that had been longer in the storage.

The PDF includes a summary in English.

• Räsänen, E-mail: pr@mm.unknown

The aim of this study was to find out the planting vigour of Scots pine (Pinus sylvestris L.) stored over the winter either in winter storage mainly in the temperature of 4 – -6 °C or in nursery beds. The experimental planting included about 4,500 of 2+1 transplants in Northern Finland. In spring 1965 the control plants were lifted in the spring before budbreak and stored in closed bags in a cold store, in the following year the control plants were lifted in June when the growth had started.

Winter storage of pine transplants in a cold store, tightly closed into bags for the major period, did not, according to the results, increase plant mortality as compared to lifting in the spring. Soaking the stored-plant roots did not affect plant mortality. Mortality was rather small in all treated lots and probably more dependent on planting site and other local factors.

No consistent difference on the leader growth, needle length, bud number and plant grade was found between the plants stored over winter and those lifted in the spring. Sealing the plants into tight bags for winter proved to be suitable method, efficiently preventing water shortage in the plants. No moulds or fungal diseases were found in the plants. In the exceptionally cold 1965–1966 winter, temperature in the cold store sank to -15 °C, but in spite of the temperatures below the recommended storing temperature, the plants survived well. The reason was that the plants froze slowly in the fall and thawed out slowly in the spring.

The value of vigour grade in predicting plant-characteristic development proved to be good, and predicted plant development also in the following year fairly well.

The PDF includes a summary in English.

• Yli-Vakkuri, E-mail: py@mm.unknown
• Räsänen, E-mail: pr@mm.unknown
• Hilli, E-mail: ah@mm.unknown