Moose (Alces alces L.) browsing was studied in young Scots pine (Pinus sylvestris L.) stands mixed with deciduous trees in high-density winter ranges. The proportional use of twig biomass decreased as the availability increased. The total as well as proportional biomass consumption were higher on the moist than on the dry type of forest. The per tree consumption of pine was higher on the moist type, where the availability of pine was lower. Deciduous trees were more consumed on the moist type, where their availability was relatively high. The consumption of pine saplings increased as the availability of birch increased. Pine stem breakages were most numerous when birch occurred as overgrowth above pine and at high birch densities. The availability of other deciduous tree species did not correlate with browsing intensity of Scots pine. Moose browsing had seriously inhibited the development of Scots pines in 6% of the stands, over 60% of available biomass having been removed. Rowan and aspen were commonly over-browsed and their height growth was inhibited, which occurred rarely by birch. There was no difference in the proportion of young stands in forest areas with high and low moose density. A high proportion of peatland forests was found to indicate relatively good feeding habitats in the high-density areas.
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Two-year-old containerized Scots pine (Pinus sylvestris L.) seedlings, raised under different fertilization and watering regimes, were subjected to feeding preference tests with pine weevils (Hylobius abietis L.) in a bioassay. In the tests carried out with pairs of seedlings, the weevil preferred water-stressed seedlings to well-watered ones. In the case of well-watered seedlings, the weevil caused significantly more damage to NPK-fertilized seedlings than those given pure PK fertilization, or no fertilization at all. It is apparent that PK fertilization reduces, and water stress increases seedling susceptibility to weevil damage. The results support findings from field trials that water stress (planting shock) predisposes seedlings to weevil damage. Weevil resistance is discussed with respect to fertilization and water stress as determinants of seedling quality.
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The effects of plantation characteristics on moose (Alces alces) browsing intensity was studied in 82 Scots pine (Pinus sylvestris L.) plantations in Southern Finland. Moose browsing occurred most commonly in plantations established on relatively fertile soil, and the degree of damage was at highest in plantations with openings. A high amount of brush, especially aspen, increased the risk of damage. Furthermore, damage was intensified in plantations situated on hills, slopes or at a long distance from main roads or settlements.
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In early spring 1968 it was noticed that the black grouses (Lyrurus tetris L.) was eating terminal shoots of Scots pine (Pinus sylvestris L.) seedlings in a tree nursery in Luumäki, Southern Finland. The terminal shoots were picked 1–4 cm from the top of the seedlings. In total some thousands of two-year-old seedlings were damaged. The depth of the snow was 10–15 cm deep and only the tops of the seedlings could be seen above the surface of the snow.
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The winter food of moose (Alces alces L.) was examined in 1967-68 in the Saariselkä fell area in the communes of Inari and Sodankylä, in the northern parts of the communes of Salla and Savukoski, and in the central part of the commune of Salla in eastern Lapland in Finland.
In northern parts of Salla and Savukoski 25 moose were followed during 3.-13.4.1968. This area is typical wintering terrain of moose in north-east Lapland. According to the estimate, 45% food taken by the moose was Scots pine shoots and needles, 28% birch, 17% juniper sprigs and needles, 9% willow, and 1.5% bear moss. According to observations of the researchers in 26.1.-16.5.1968, moose seemed to avoid birch, even if it was available in the area, and eat Scots pine shoots and needles and juniper.
Moose seemed to prefer willow in as a winter feed in the southern part of the area studied, where it accounts according to the present and earlier studies 50-90% of the winter food. In the northern wintering areas of moose, where willow is not as common, willow seemed to account for less than 10% of the winter food. There Scots pine is the most important winter food for moose.
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The study was carried out at Padasjoki, Southern Finland, where moose (Alces alces L.) density on the winter range had been over 1.5 individuals/km2. Moose browsing intensity, expressed in terms of number of twigs eaten and biomass used, increased with stand density (biomass). Total biomass consumption (dry weight) per sample plot and per sapling. The number of bites increased, but the percentage biomass removed did not differ when stand density increased. A relatively large bite size was observed on the plots of low stand density. The quantity of food, which on average was of relatively low quality, was obviously important due to the benefit gained through reducing the search time.
The nutritive value of the browse, expressed in terms of chemical compounds indicating low food digestibility, was lower in the dense than in the sparse Scots pine stand. However, the amount of crude protein and arginine were relatively high in the dense stand. We concluded that shading affected the nutritional status of saplings on high density plots.
Although the biomass removed by moose per sapling was high for low density plots, the remaining biomass was larger than that on the high-density plots owing to the relatively large twig biomass of saplings. The number of saplings per hectare without main stem breakage increased significantly as stand density increased.
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The utilization of available food resources by the moose (Alces alces L.) was studied in a Scots pine (Pinus sylvestris L.) plantation containing an admixture of deciduous species. Rowan (Sorbus aucuparia L.) and aspen (Populus tremula L.) were highly utilized compared to pine and both silver birch (Betula pendula Roth) and downy birch (B. pubescens Ehrh.). However, they were not capable of withstanding continuous browsing by moose owing to their diminished biomass. In total, the browsing intensity (number of browsed twigs/tree) on pine and birch was about double of that on rowan and aspen.
The number of browsed twigs per tree increased as the amount of available main branches increased. The number of bites per available branch, as well as the maximum diameter of the bites, decreased as the density of the plantation increased. Silver birch was more used by moose than pubescent birch as well as planted silver birch compared with naturally regenerated trees.
Main stem breakage was especially common in winter 1988, the average height of the pine and birch trees being over two meters. The tops of broken stems were commonly utilized as food. The increase in moose density and the relatively deep snow cover evidently promoted the incidence of serious damage. The number of undamaged trees/ha was greater in dense than in sparse parts of the stand.
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The establishment of moose (Alces alces L.) winter feeding sites, their utilization and their effect on damage to young Scots pine (Pinus sylvestris L.) plantations was studied in Ruokolahti-Imatra area in Eastern Finland in 1987–89. During the period, the density in the area was about 3–5 moose/ 1,000 ha.
Six feeding sites were established by fertilization, offering mineral lics and the tops of aspen and Scot pine and by salting the tops of pine. The moose preferred the feeding site to control areas during both summer and winter. In winter browsing was very heavy, especially in those areas located in or close to traditional wintering areas. In winter no moose were seen in the summer habitats.
The extent of, and fluctuations in moose damage were studied in 47 Scots pine plantations in the immediate surroundings of the feeding sites (29 plantations), control areas (18 plantations) and also 68 randomly selected pine plantations. Before the experiment began in 1987 four plantations had been seriously damaged. During the study period only one plantation was seriously damaged. However, it could not be conclusively proved that damage to the pine plantations had been reduced as a result of the feeding sites. The results of the study can be put into practice elsewhere to create better living conditions for moose in their winter habitats. However, the food offered at the feeding site should be in the right proportion to the number of animals wintering in the area, so that the risk of damage to nearby plantations would be kept as small as possible.
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Adult pine weevils (Hylobius abietis (L.)) feed on the tender bark of branches and roots of mature conifer trees and on the stem bark of conifer seedlings. Their feeding on mature trees does not cause any economic damage, but their feeding on planted seedlings is so devastating that the pine weevil is considered one of the most important forest pest insects in Europe. We asked whether the pine weevil prefers seedlings over other regularly utilized food sources. This question is of particular interest because new approaches to seedling protection are based on decreasing any preference for seedlings by using less palatable plants or by enhancing their defence (by genetic selection or by methyl jasmonate treatment). In a laboratory choice experiment we tested pine weevil feeding preferences for seedlings compared with branches and roots from mature trees (separately for Norway spruce and Scots pine). Pine weevils preferred roots, but not branches, of Norway spruce over seedlings of the same species. With Scots pine there were no clear preferences, but the weevils showed a tendency to prefer roots over seedlings. These results provide support for seedling protection approaches that attempt to redirect pine feeding from planted seedlings to other food sources.