Distribution and occurrence of bark beetles and other forest insects in relation to environmental variation were analysed by multivariate methods. Eight different forest edges were studied using 10 x 10 m sample plots that formed 200 m linear transects perpendicular to the forest edge. Forest edge affected the distribution of insect species only in the edges between mature, non-managed spruce stands and clear cuts or young seedling stands, but not in the pine stands. The occurrence of the selected forest insects mainly depended on variables associated with the amount and quality of suitable woody material. The most significant environmental variables were forest site type, crown canopy coverage, tree species, number of stumps, number of dead spruce trunks and amount of logging waste at site. Quantitative classification of species and sample plots showed that some specialized species (Xylechinus pilosus, Cryphalus saltuarius, Polygraphus poligraphus and P. subopacus) adapted to mature spruce forests, tended to withdraw from the forest edge to interior stand sites. By contrast many generalized species (Pityogenes chalcographus, P. quadridens, Pissodes spp., Hylurgops palliatus, Tomicus piniperda, Dryocoetes spp. and Trypodendron lineatum) benefitted from cuttings and spread over stand borders into mature forest.
Four dispersal stages of the nematode Bursaphelenchus mucronatus (Nematoda: Aphelenchoididae) were surveyed in Finland during the summers 1989 and 1990 examining field collected conifer beetle (Coleopterea) adults of the following species: Monochamus sutor, M. galloprovincialis, Acanthocinus aedilis, Rhagium inquisitor, Asemum striatum, Spondylis buprestoides and Hylobius abietis. All but the last one (Curculionidae) belong to genus Cerambycidae. The two Monochamus species were the only ones carrying B. mucronatus fourth dispersal stages, total number of nematode larvae per beetle being higher in M. galloprovincialis. The frequency of infestation was 24% in M. galloprovincialis and 14% in M. sutor but the difference eas not statistically significant.
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In the 1980 and 1981, windthrown and felled Scots pine (Pinus sylvestris L.) were examined at 8 localities in Sweden. The number and length of egg galleries as well as the number of exit holes of Tomicus piniperda (L.) and T. minor (Hart.) were recorded on sample sections (30 m in length) distributed at 3 m intervals on the 37 fallen pine stems, which were successfully colonized by the beetles. In addition, 78 uprooted pines were surveyed in 6 localities. Most trees were attacked by T. piniperda, but only a few by T. minor. Successful colonization often resulted in the production of several thousand beetles per tree, the maximum being approximately 1,800. The attack density of T. piniperda seldom exceeded 200 egg galleries/m3 bark area, and the brood production usually remained below 1,000 beetles/m3. Much higher figures were obtained or T. minor. In T. piniperda, the rate of reproduction (i.e. the number of exit holes /egg gallery) decreased rapidly with increasing attack density, whereas T. minor seemed to be less sensitive to intraspecific competition.
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Ips acuminatus Gyll. (Coleoptera, Scolytidae) is a bark beetle that causes deep bluing in thin-barked Scots pine (Pinus sylvestris L.) pulpwood. It has been shown that this decreases pulp yield. The purpose of this study was to map the southern border of the distribution of Ips acuminatus in Finland. It was found that there have been changes in the distribution of this species during the last three decades. Ips acuminatus has now disappeared from southern Finland. On the basis of the sample plots (134 cutting areas) the southern border of this pest lies on the line running through the towns Vaasa–Seinäjoki–Äänekoski–Jyväskylä–Pieksämäki–Savonlinna–Punkaharju. A certain degree of localisation was observed in the occurrence of I. acuminatus in its distribution area, for instance, differences in its occurrence frequency in cutting areas and even in log and cutting residue piles in the sample cutting area.
It is considered that the most important reasons for these changes in distribution are the increase in logging and changes in the location of cutting sites, and resulting competition for breeding material for the increased population of bark beetles. Furthermore, the long-distance transport of unpeeled logs from the north across the present southern borders may, in the future, contribute to local changes in the southern distribution of Ips acuminatus.
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The dying-off of more trees in an over-aged Norway spruce (Picea abies (L.) H. Karst.) stand caused by Ips-bark beetles was reduced by a pheromone preparation, ipslure. 20 preparations placed in trapping bolts captured more than 13,700 specimens of Ips typographus L. and Ips duplicatus Sahlb., which alone corresponded to a saving of five old trees in this valuable exhibition and seed collection stand. Attractance of ipslure to the following predators of bark beetles was also examined; Thanasimus formicarius, T. rufipes, Epuracea bickhardti, Rhizophagus ferrugineus, Pityophagus ferrugineus.
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The aim of the study was to determine which kinds of insects had infected the Norway spruce (Picea abies (L.) H. Karst.) in different stands killed by flooding caused by beavers (Castor canadensis Kuhl), and if there was any danger that they would subsequently cause damage in the surrounding forests. The effect of tree diameter and certain stand characteristics on the fauna of dead trees are discussed. The occurrence of different insect combinations and qualifications for their coexistence were studied.
Pityogenes chalcographus L., Trypodendron lineatum O., Hylurgops palliatus Gyll. and Dryocetes autographus Ratz. occurred most abundantly. 20 phloem or wood boring species were observed in 5 regular succession types. Secondary species occurred in a virgin stand while Ips typographus L. was found at the edge of a felling area. Owing to the flooding, species preferring moist conditions were abundant. In this case damages had not spread to the surrounding forests which, however, might be possible under certain conditions.
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The paper studied the effect of felling time and conditions in the forest depot of timber to damages caused by spruce ambrosia beetle (Trypodendron lineatum Oliv.) to coniferous timber with bark, both experimentally and observing forest depots in Finland. Effects of fellings was studied by studying the abundance of the beetles in logging residue.
The results show that the spruce ambrosia beetles favour timber felled during the late autumn and winter, stored in a shaded place in the forest. In addition, new spruce stumps maintain and increase the beetle population. Fellings in the forest will increase population during the next year and cause damages in forest depot of timber nearby, because the insect breeds in the stumps. The experiments showed that it is possible to diminish the damages caused by the beetle to timber with bark by spraying with insecticides, and timing the fellings and transport of timber so that there is no timber in the forest in the spring during the time when the insect swarms.
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Degree of decomposition of logging residue, and decay in stumps have been used in forest inventories to estimate the time of the felling. In this paper, a method was developed to use insects breeding in the logging residue to determine how long ago the felling took place. The method is based on the arrival and rate of development of the different species of bark beetles that breed in the logging residue.
The most suitable insect species to be used in the purpose of timing the age of logging residue were defined, and their occurrence in different tree species and fellings performed at different times of the year were described. The species can be easily identified by gallery systems characteristic to the species.
It is concluded that the method does not suit for broadleaved species, because there is no common insects suitable for this purpose. Also, the time of swarming of the insects depends on the weather conditions in the spring, which makes it difficult to give definite dateshe progress of the spring has to be taken into account when the occurrence of the insects is used in the determination of the time of the felling. In addition, local conditions, such as shading, affect drying of the branches, and can influence the occurrence of the insects. For Scots pine and Norway spruce the age of the logging residue can be determined precisely only at most two years back.
Bark beetle populations live usually in a balance in natural forests, and outbreaks occur seldom. The populations have been found to increase in managed forests. Fellings affect the structure of the forests, which influence the living conditions of the insects, and produce material for reproduction. In this study the occurrence of bark beetles was studied in a forest area in Etelä-Häme in Southern Finland using line plot survey.
The forests in the area were Norway spruce (Picea abies L. Karst.) dominated. Over third of the 140 sample plots studied were in forests which had never been cut or it was over ten years to the last logging. Bark beetles of 26 different species were found in 66 of the sample plots. The most common species was six-toothed spruce bark beetle (Pityogenes chalcographus L.), which was due to the abundance of growth material suitable for the species in the area. New species in the area were common double-eyed spruce bark beetle (Polygraphus polygraphus L.), Pityophthorus micrographus L., and Dryocetes-beetle (either Dryocetes autographus or D. hectographus). The fellings increased the occurrence of beetles. The amount and quality of logging residue affected the abundance of the insects.
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