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Articles by Mikko Hyppönen

Category : Research article

article id 24064, category Research article

Mikko Hyppönen, Ville Hallikainen, Urban Bergsten, Hans Winsa, Jari Miina, Sauli Valkonen, Pekka Helenius, Esko Jaskari, Pasi Rautio. (2025). Is it possible to extend the season for direct seeding of Scots pine and lodgepole pine in northern Fennoscandia? Silva Fennica vol. 59 no. 3 article id 24064. https://doi.org/10.14214/sf.24064

Keywords: Pinus sylvestris; Pinus contorta; artificial regeneration; autumn direct seeding; generalized linear mixed effects models

Highlights: Scots pine and lodgepole pine were direct seeded during June–November in a six-year experiment in Finnish Lapland; Both early summer and late autumn as well as early winter were appropriate periods for the successful seeding of both species; Seeding in late summer and early autumn (August and September) was much less successful for Scots pine; For lodgepole pine seeding time, site quality or cold-wet treatment of seeds were not critical for the seedling emergence or survival.

Abstract | Full text in HTML | Full text in PDF | Author Info

The aim of this study was to experimentally test the most favourable seeding times for Scots pine (Pinus sylvestris L.), and to compare Scots pine and lodgepole pine (Pinus contorta Douglas ex Loudon) in respect to suitable seeding times and site factors. The experimental area was located in Sodankylä, Central Finnish Lapland. Generalized linear mixed effects models with binomial distribution assumption were applied to model the presence or absence of a seedling at a seeding point. The study shows that in addition to spring and early summer, direct seeding of Scots pine can also be carried out during late autumn or even early winter (October and November) in northern Fennoscandia. On the contrary, seeding in late summer and early autumn (August and September) is much less successful and cannot be recommended as such, but can be used if the amount of seeding material is increased to compensate the loss. There is more flexibility for lodgepole pine for which the proper seeding period seems to be from spring through to late autumn. Whether lodgepole pine seeds were stratified or not had no statistically significant effect on regeneration success. Our results clearly indicate that lodgepole pine is less susceptible to unfavourable site and soil factors than Scots pine. Our results give support to extend the period of direct seeding from the present early summer to another period in late autumn and even early winter.

Hyppönen, Natural Resources Institute Finland (Luke), Ounasjoentie 6, FI-96200 Rovaniemi, Finland E-mail: mikkokthypponen@gmail.com
Hallikainen, Natural Resources Institute Finland (Luke), Ounasjoentie 6, FI-96200 Rovaniemi, Finland https://orcid.org/0000-0001-5384-8265 E-mail: ext.ville.hallikainen@luke.fi
Bergsten, Department of Forest Biomaterials and Technology, Swedish University of Agricultural Sciences (SLU), Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: burskog@gmail.com
Winsa, Natural Resources Institute Finland (Luke), Ounasjoentie 6, FI-96200 Rovaniemi, Finland E-mail: hans.winsa@gmail.com
Miina, Natural Resources Institute Finland (Luke), Yliopistokatu 6 B, FI-80100 Joensuu, Finland https://orcid.org/0000-0002-8639-4383 E-mail: jari.miina@luke.fi
Valkonen, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790, Helsinki, Finland https://orcid.org/0000-0002-2879-4821 E-mail: sauli.valkonen@luke.fi
Helenius, Natural Resources Institute Finland (Luke), Tietotie 4, FI-31600 Jokioinen, Finland E-mail: psjhelenius@elisanet.fi
Jaskari, Natural Resources Institute Finland (Luke), Ounasjoentie 6, FI-96200 Rovaniemi, Finland E-mail: ejaskari@gmail.com
Rautio, Natural Resources Institute Finland (Luke), Ounasjoentie 6, FI-96200 Rovaniemi, Finland https://orcid.org/0000-0003-0559-7531 E-mail: pasi.rautio@luke.fi

article id 1408, category Research article

Anna Katharina Franke, Pasi Aatsinki, Ville Hallikainen, Esa Huhta, Mikko Hyppönen, Vesa Juntunen, Kari Mikkola, Seppo Neuvonen, Pasi Rautio. (2015). Quantifying changes of the coniferous forest line in Finnish Lapland during 1983–2009. Silva Fennica vol. 49 no. 4 article id 1408. https://doi.org/10.14214/sf.1408

Keywords: Pinus sylvestris; Picea abies; tree line; forest regeneration; environmental change; stand volume

Highlights: Volume of the growing stock of spruce and pine has increased in forests and in timber lines during the past 26 years; Spruce stem numbers increased on average, while pine stem numbers remained stable and location-specific variation was observed; Presuming that the ongoing trend of increasing temperature will remain, the enhanced forest regeneration and growth may result in extension of forests in the future.

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The boreal timber- and tree-line forests grow in harsh environmental conditions in their outermost distribution limit. Here even small environmental changes may cause dramatic changes in the distribution of tree species. We examined changes of the forest lines of Norway spruce (Picea abies (L.) H. Karst.) and Scots pine (Pinus sylvestris L.) in Finnish Lapland five times during 1983–2009. We monitored the number of stems and the volume of the growing stock in thirteen different locations in forest-line areas. The linear temporal trends and the variations of these response variables were used as indicators of a possible change during the study period. Spruce showed a significant increase both in the volume of the growing stock (up to 40% increase) and in the total stem number (up to 100% increase). A significant increase in the volume of the growing stock was observed in the pine data as well (up to 70% increase), whereas the stem number stagnated or even decreased. The results suggest that spruce needs favourable conditions to have an abundant regeneration, but after the establishment the seedlings seem to be more resistant against biotic and abiotic disturbances than pine seedlings. The increasing stand volume might result in a climate-related northward and upward extension of forests in the future. However, our results show that responses in the boreal forest line are species and location specific and a more favourable climate does not necessarily lead to an advance of the coniferous forest line.

Franke, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: anna.franke@fau.de
Aatsinki, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: pasi.aatsinki@luke.fi
Hallikainen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: ville.hallikainen@luke.fi
Huhta, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: esa.huhta@luke.fi
Hyppönen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: mikko.hypponen@luke.fi
Juntunen, The Sámi Education Institute, Menesjärventie 4, P.O. Box 50, FI-99871 Inari, Finland E-mail: vesa.juntunen@sogsakk.fi
Mikkola, Natural Resources Institute Finland (Luke), Economics and society, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: kari.mikkola@luke.fi
Neuvonen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 68, FI-80101, Joensuu, Finland E-mail: seppo.neuvonen@luke.fi
Rautio, Natural Resources Institute Finland (Luke), Bio-based business and industry, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: pasi.rautio@luke.fi

article id 1153, category Research article

Anu Akujärvi, Ville Hallikainen, Mikko Hyppönen, Eero Mattila, Kari Mikkola, Pasi Rautio. (2014). Effects of reindeer grazing and forestry on ground lichens in Finnish Lapland. Silva Fennica vol. 48 no. 3 article id 1153. https://doi.org/10.14214/sf.1153

Keywords: forest management; timber harvesting; reindeer herding; pastures; lichen cover; lichen biomass

Highlights: Both reindeer grazing and forestry affect the cover and biomass of reindeer lichens; Reindeer grazing has bigger impact than forestry; The lichen cover was about five-fold and the biomass about fifteen-fold in the ungrazed (fenced) sites than in the grazed ones; The decrease of not only the biomass, but also the cover of lichens, is alarming.

Abstract | Full text in HTML | Full text in PDF | Author Info

Reindeer husbandry and forestry are practiced in the same areas in northern Fennoscandia. Reindeer pastures have largely deteriorated. We aimed to quantify the separate and combined effects of reindeer grazing and forestry on the amount of ground lichens. To do this, we mapped and inventoried all larger enclosures (49) in Finnish Lapland where forest management practices were similar in both sides of the fence. The average time since fencing was 43 years. We recorded the cover and estimated dry biomass of ground lichens, as well as parameters describing forest stand characteristics. The effect of reindeer grazing on both the cover and estimated dry biomass of lichens was clear: in the ungrazed (fenced) sites, the lichen cover (35.8%) was on average 5.3-fold and the dry biomass (1929 kg ha^–1) 14.8-fold compared with the corresponding estimates in the grazed sites (6.8% and 130 kg ha^–1). The effect of forestry on lichens was smaller. In the grazed stands the cover and biomass of lichens were higher in the mature stands compared to the younger stand development classes, whereas in the ungrazed stands there were no significant differences between the development classes. Both reindeer grazing and forestry affect the cover and biomass of ground lichens. The influence of reindeer grazing is, however, much heavier than that of forestry. The decrease of not only the biomass, but also the lichen cover, is alarming. The decrease of lichen cover may hinder the recovery of reindeer pastures, which in the long run endangers the sustainability of reindeer husbandry.

Akujärvi, Finnish Environment Institute, Helsinki, Finland E-mail: anu.akujarvi@ymparisto.fi
Hallikainen, Finnish Forest Research Institute, Northern Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: ville.hallikainen@metla.fi
Hyppönen, Finnish Forest Research Institute, Northern Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: mikko.hypponen@metla.fi
Mattila, Finnish Forest Research Institute, Northern Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: eero.mattila@metla.fi
Mikkola, Finnish Forest Research Institute, Northern Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: kari.mikkola@metla.fi
Rautio, Finnish Forest Research Institute, Northern Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: pasi.rautio@metla.fi

article id 903, category Research article

Mikko Hyppönen, Ville Hallikainen, Juhani Niemelä, Pasi Rautio. (2013). The contradictory role of understory vegetation on the success of Scots pine regeneration. Silva Fennica vol. 47 no. 1 article id 903. https://doi.org/10.14214/sf.903

Keywords: site preparation; seedling establishment; seedling mortality; field- and ground-layer vegetation; modelling approach; initial growth; prescribed burning

Abstract | Full text in HTML | Full text in PDF | Author Info

In North-East Finland, severe problems have been encountered in the natural regeneration of Scots pine (Pinus sylvestris L.) on sites where regeneration through site preparation usually is quite successful. We hypothesized that in that area understory vegetation, especially heather (Calluna vulgaris), crowberry (Empetrum hermaphroditum), mosses and lichens, could play a key role in this pattern. We found that in general, ground- and field-layer vegetation tends to be in a negative relationship with the establishment, growth and survival of pine seedlings. Some positive relationships were also observed. Lingonberry (Vaccinium vitis-idea) tended to improve seedling height growth. Heather, instead, seemed to have a contradictory role. It was positively related to seedling establishment but negatively to seedling growth. This dual role raises further questions about the primary reasons for the regeneration problems in North-East Finland. All in all, our results suggest that conventional methods of forest regeneration in these kinds of areas are not always effective enough and additional measures are needed. These might include severe prescribed burning along with site preparation in order to decrease the impact of the dominant ground- and field-layer vegetation on the success of Scots pine regeneration.

Hyppönen, Finnish Forest Research Institute, Rovaniemi Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: mikko.hypponen@metla.fi
Hallikainen, Finnish Forest Research Institute, Rovaniemi Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: ville.hallikainen@metla.fi
Niemelä, Finnish Forest Research Institute, Rovaniemi Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: jn@nn.fi
Rautio, Finnish Forest Research Institute, Rovaniemi Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: pasi.rautio@metla.fi

article id 158, category Research article

Ville Hallikainen, Mikko Hyppönen, Leena Pernu, Jouni Puoskari. (2010). Family forest owners’ opinions about forest management in northern Finland. Silva Fennica vol. 44 no. 2 article id 158. https://doi.org/10.14214/sf.158

Keywords: forest management; silviculture; family forest owner; northern Finland; forest owner typology; forest management objective; forest management activity; tourism

Abstract | View details | Full text in PDF | Author Info

Hallikainen, Finnish Forest Research Institute, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: ville.hallikainen@metla.fi
Hyppönen, Finnish Forest Research Institute, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: mh@nn.fi
Pernu, Finnish Forest Research Institute, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: lp@nn.fi
Puoskari, Finnish Forest Research Institute, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: jp@nn.fi

article id 235, category Research article

Ari Nikula, Ville Hallikainen, Risto Jalkanen, Mikko Hyppönen, Kari Mäkitalo. (2008). Modelling the factors predisposing Scots pine to moose damage in artificially regenerated sapling stands in Finnish Lapland. Silva Fennica vol. 42 no. 4 article id 235. https://doi.org/10.14214/sf.235

Keywords: Pinus sylvestris; boreal forest; forestry; Alces alces; damage; modelling

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Moose (Alces alces) damage in forest plantations have been at a high level in Finland in recent decades. Nowadays, moose is the most severe pest in Scots pine plantations also in Finnish Lapland. So far, despite the high level of damage and different bio-geographical conditions in Northern Finland, most of the moose-damage research has been carried out in Southern Finland. A number of research have also been performed to analyse factors affecting browsing but predictive models are rare. Data from 123 randomly selected and artificially regenerated pine plantations in Northern Finland were used in modelling the risk of moose browsing. The stands had been regenerated during 1984–1995. A total of 508 sample plots (range 2–8 plots per stand) were measured. Hierarchical logistic regression models with a random factor were constructed to predict the probability of leader-shoot browsing of pine on a plot. The number of planted pines and deciduous trees overtopping the pines were the most important predictors increasing the browsing probability. The results support earlier findings that deciduous trees overtopping or reaching the height of the pines should be cleaned from the immediate vicinity of the pines. Seedlings with a height ranging from 75 to 299 centimetres were more susceptible to browsing. Heavy soil scarification, such as ploughing or mounding, increased the browsing probability compared with lighter scarification methods. Soil type did not affect the browsing probability, but paludification decreased it. The within-stand variation in deciduous trees density and height should be taken into account in future moose browsing risk assessments. In Lapland, high moose damage risk areas are characterized by a low elevation and higher temperature sum.

Nikula, Finnish Forest Research Institute, Rovaniemi Research Unit, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: ari.nikula@metla.fi
Hallikainen, Finnish Forest Research Institute, Rovaniemi Research Unit, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: vh@nn.fi
Jalkanen, Finnish Forest Research Institute, Rovaniemi Research Unit, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: rj@nn.fi
Hyppönen, Finnish Forest Research Institute, Rovaniemi Research Unit, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: mh@nn.fi
Mäkitalo, Finnish Forest Research Institute, Rovaniemi Research Unit, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: km@nn.fi

article id 308, category Research article

Ville Hallikainen, Mikko Hyppönen, Juha Hyvönen, Juhani Niemelä. (2007). Establishment and height development of harvested and naturally regenerated Scots pine near the timberline in North-East Finnish Lapland. Silva Fennica vol. 41 no. 1 article id 308. https://doi.org/10.14214/sf.308

Keywords: natural regeneration; seedling establishment; Scots pine; timberline; seed-tree method; density model; height development model

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Researchers and professionals in practical forestry have faced problems concerning the regeneration success of Scots pine in natural regeneration near the timberline in North-East Lapland. The aim of the study was to analyze the seedling establishment and seedling height development of Scots pine in seed-tree stands in the area. The average number of living pine seedlings in the study stands was about 1000 ha^–1, but there was considerable variation between the stands. The seedling density was modelled using a multinomial logistic regression with a random factor. Forest site type and the time since seed-tree cutting were the most significant explanatory variables in the model. The probability of reaching the acceptable seedling density was higher on dry site types than on the more fertile ones. The probability increased with the time elapsed since the regeneration activities. Effective temperature sum and the number of intermediate pines also positively affected the probability, but the presence of residual trees negatively. On northern and eastern slopes the probability was lower than on southern and western ones. Seedling height was modelled using a linear mixed model. The age of a dominant seedling was the most positively effective explanatory variable in the height development model. Other positively affecting significant predictors were time since seed-tree cutting, number of intermediate birches, and distance between a seedling and the nearest seed tree. Degree of paludification had a negative effect. The study suggests that the regeneration of Scots pine in North-East Lapland is a relatively slow process.

Hallikainen, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: ville.hallikainen@metla.fi
Hyppönen, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: mikko.hypponen@metla.fi
Hyvönen, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: juha.hyvonen@metla.fi
Niemelä, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: jn@nn.fi

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