Articles by Olavi Luukkanen

Ethanolic sprays of GA₄ or GA₇ on 9-year old Scots pine (Pinus sylvestris L.) grafts, repeated four times during the shoot elongation period, resulted in a statistically significant increase in female flowering one year after the treatment. Of the two compounds, GA₄ seemed to be somewhat more efficient, yielding 47 female strobili/100 shoots vs. 36 and 6 strobili/100 shoots in GA₇ and control treatments respectively. The mixture of GA₄ and GA₇ compounds was also applied and seemed to have an effect intermediary to those of the pure compounds. However, due to the limited amounts of material, none of the differences between the gibberellins could be statistically confirmed. Male flowering frequencies were also too low to allow any firm conclusions, but the numerical results suggested that the purified gibberellins may promote male and female flowering in different ways.

The PDF includes a summary in Finnish.

• Luukkanen, E-mail: ol@mm.unknown

The paper consists of a report of a study tour made by Finnish forestry students, under the leadership of the author, to Harbin, Changchun, Peking, Nanking and Shanghai in December 1977. In addition, some earlier literature sources concerning forestry in China are briefly reviewed. The paper presents the general geographic characteristics of north-eastern China, as well as the vegetation zones and timber species of this region. Silvicultural methods and the main features of forest technology and forest industry are also discussed. The last chapters describe the forestry administration and current trends in forestry education and research in north-eastern China as observed during the tour.

The PDF includes a summary in Finnish.

• Luukkanen, E-mail: ol@mm.unknown

The aim of the present study was to establish whether hormone treatments would promote flowering in Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) grafts under Finnish conditions. Also, an attempt was made to test the efficiency of hormones as well as the variation in response among different clones. Six Scots pine and six Norway spruce clones were selected in each seed orchard based on their flowering intensity, and treated with growth hormones (GA, NAA) of different dosages by spraying. Flowering was observed one year after the treatments.

None of the treated or untreated spruce grafts flowered. However, poor flowering in the natural stands indicated that the environmental conditions during the previous years did not favour flowering. On the other hand, a distinct increase in flowering in Scots pine was observed as a result of spraying with hormone solutions. Treatments with gibberellin had a distinct promoting effect both on male and female flowering in the Scots pine grafts, although the responses varied between the different hormones or clones. The relative effect was generally stronger in male flowering.

The PDF includes a summary in Finnish.

• Luukkanen, E-mail: ol@mm.unknown
• Johansson, E-mail: sj@mm.unknown

A system for measuring the net photosynthesis, transpiration and environmental factors within the canopy and ground cover vegetation is described. The system operates continuously throughout the growing season in a young Scots pine (Pinus sylvestris L.) stand. A data-logging unit controls the system and carries out the measurements on the readings of the sensors of photosynthesis, transpiration, light intensity outside the canopy, light climate inside the assimilation chambers, and dry and wet temperatures from selected points. These measurements are shown digitally and automatically punched onto paper tape.

The PDF includes a summary in Finnish.

• Hari, E-mail: ph@mm.unknown
• Kanninen, E-mail: mk@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown
• Luukkanen, E-mail: ol@mm.unknown
• Pelkonen, E-mail: pp@mm.unknown
• Salminen, E-mail: rs@mm.unknown
• Smolander, E-mail: hs@mm.unknown

The main forest research institutes in USSR as well as schools for higher education in forestry are presented in this article, on the basis of observations made during study tours and the relevant literature. Research in forest genetics, tree physiology and related fields i stressed. The State Forestry Committee leads the regional units of forestry administration and also governs a number of research institutes. Among these, the All-Union Forest Research Institute near Moscow has a central position. Regional institutes as well as those specializing in a particular field within forest research may belong to this category, too. Basic research, on the other hand, is conducted under the auspices of the Academy of Sciences, the Academy of Agricultural Sciences or academies of sciences of the Soviet republics. This type of institute also includes those devoted purely to forestry. Forest education at the university level takes place in schools usually called institutes of forest technology.

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• Luukkanen, E-mail: ol@mm.unknown

Since the year 1966, fourteen specialists, most of them forestry graduates from Kasetsari University in Bangkok, have studied or will start studying in Finland. The visiting graduate students have completed their program successfully, and are interested, as well as the Thai authorities, in continuing the educational cooperation. However, the Finnish Government now seems to have changed the list of countries which are included in this kind of program. The Thai authorities will possibly grant a limited number of scholarships for studies carried out in Finland. The possible need to develop the study program further is discussed if the educational cooperation with Thailand will continue.

The PDF includes a summary in English.

• Luukkanen, E-mail: ol@mm.unknown

After 64 days of aseptic culture, germlings of Pinus Syvestris L. were cut at the middle of the hypocotyl and above the root. The upper and lower halves of the hypocotyls were transferred onto agar medium RM-196 of Linsmaier & Skoog (1964) including 2 mg/l IAA and 0.1, 1 or 10 mg/l kinetin, one or both halves being put in each vial. Callus growth and root formation was observed after 55 days.

The lower ends of basally cut seedlings generally formed callus tissue and 20% of them also formed roots from this callus. No roots and less callus growth were observed in the lower hypocotyle halves excised at both ends. In the latter hypocotyles callus growth was promoted by the presence in the same vial of a basally excised germling, including cotyledons and plumule. Increasing amounts of kinetin slightly enhanced callus formation of basally excised germlings but seemed to inhibit callus growth in hypocotyls excised at both ends and placed alone on the growth medium. The total amount of callus was greatest in hypocotyls which included intact cotyledons and plumule.

The PDF includes a summary in English.

• Luukkanen, E-mail: ol@mm.unknown

Photosynthesis and dark respiration in five families of autochtonous Norway spruce (Picea abies (L.) H. Karst.) and in seedlings from twenty Finnish stands of Scots pine (Pinus sylvestris L.) were investigated in constant environmental conditions. Values of CO₂ exchange were compared with the height growth and weight of seedlings in Norway spruce and with the weight alone in Scots pine. No statistically significant differences were found in CO₂ exchange among progenies or stands. Photosynthetic efficiency and photosynthetic capacity showed a positive correlation both in spruce and in pine. Growth and net photosynthetic capacity were linearly and positively correlated in pine. Spruce and a higher light compensation point than pine. The use of an open IRGA system with several simultaneous measurements and the trap-type cuvette construction in genetic work are discussed.

The PDF includes a summary in English.

• Luukkanen, E-mail: ol@mm.unknown

In the literature review the current status of information on the genetic variation of CO₂ exchange and some reviews and investigations on this subject are listed. Photorespiration is separately discussed and unpublished data of an electron microscope study of poplar leaf microbodies are presented.

Considerable genetic inter- and intraspecific variation is found in several characteristics that affect CO₂ exchange in trees. Photosynthesis in young trees does not correlate well with growth through the whole rotation cycle. A special interest has been shown to marginal environmental conditions (e.g. water deficit, low temperature, and low light intensity), as opposed to optimal conditions often employed in laboratory studies of CO₂ exchange in trees.

In an unpublished poplar studies by the author et.al. a preliminary experiment with poplar clones showed variation in the CO₂ competition point. This variation was negatively correlated with the photosynthesis efficiency of these clones.

The PDF includes a summary in English.

• Luukkanen, E-mail: ol@mm.unknown

About 4,000 seedlings of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) were planted in 1965 both on a clear-cut and sheltered area in Central Finland. In the autumn of 1966 needle colour was determined by using Muncell Color Charts which allowed a quantitative measurement of three colour dimensions (hue, value, and chroma). Terminal shoot growth was recorded for two years after colour measurements. In both species, fertilization (NPK in the spring of the year of colour measurement) as well as other site factors caused differences in all three dimensions of needle colour. A regression of shoot growth on needle colour was found in both species. In most cases colour value (darkness) and, in spruce, also chroma, predicted the subsequent growth almost as well as did these two-colour variables together.

The PDF includes a summary in English.

• Luukkanen, E-mail: ol@mm.unknown
• Räsänen, E-mail: pr@mm.unknown
• Yli-Vakkuri, E-mail: py@mm.unknown

The profitability of fast-growing trees (Eucalyptus camaldulensis Dehnh., Acacia mangium Willd. and Melia azedarach L.) was investigated in the north-eastern and eastern provinces of Thailand. The financial, economic, and tentative environmental-economic profitability was determined separately for three fast-growing plantation tree species and for three categories of plantation managers: the private industry, the state (the Royal Forest Department) and the farmers. Fast-growing tree crops were also compared with teak (Tectona grandis L. f.), a traditional medium or long rotation species, and Para rubber (Hevea brasiliensis (Willd. ex A. Juss.) Müll. Arg.) which presently is the most common cultivated tree in Thailand.

The optimal rotation for Eucalyptus camaldulensis pulpwood production was eight years. This was the most profitable species in pulpwood production. In sawlog production Acacia mangium and Melia azedarach showed a better financial profitability. Para rubber was more profitable and teak less profitable than the three fast-growing species. The economic profitability was higher than the financial one, and the tentative environmental-economic profitability was slightly higher than the economic profitability.

The profitability of tree growing is sensitive to plantation yields and labour cost changes and especially to wood prices. Management options which aim at pulpwood production are more sensitive to input or output changes than those options which include sawlog production. There is an urgent need to improve the growth and yield data and to study the environmental impacts of tree plantations for all species and plantation types.

The PDF includes a summary in Finnish.

• Niskanen, E-mail: an@mm.unknown
• Luukkanen, E-mail: ol@mm.unknown
• Saastamoinen, E-mail: os@mm.unknown
• Bhumibhamon, E-mail: sb@mm.unknown

The aim of the present work was to clarify the structure of selected natural Pinus kesiya Royale ex Gord. And P. merkusii Jungh. et de Vriese stands, with particular attention on the amount and quality of naturally-occurring seedlings and young trees. Furthermore, basic information required in the management of pine stands in watershed areas of northern Thailand was obtained.

Large number of pine seedlings were found in P. kesiyana stands only. Two of the examined four localities where this species occurred also included various intermediary height classes under the mature trees. In dense stands the number of seedlings was generally smaller as compared with lower seed tree densities. Neither one of the two localities where P. Merkusii was investigated indicated sufficient natural regeneration. Young pines (over 5 m height) seemed to survive the frequent ground fires quite well, whereas younger seedlings were destroyed and the ground layer vigour was lowered in these cases.

In situ sowing experiments at the beginning of the dry season indicated a faster development and better survival of emerging P. Kesiya seedlings as compared to P. Merkusii. Soil preparation and exposure to sun decreased the survival of seedlings in both species. Utilization of natural regeneration and direct sowing of pines as a silvicultural method as well as the general significance of the two pine species in the succession of plant communities under the influence of forest fires is also discussed.

The PDF includes a summary in Finnish.

• Turakka, E-mail: at@mm.unknown
• Luukkanen, E-mail: ol@mm.unknown
• Bhumibhamon, E-mail: sb@mm.unknown

The net photosynthetic rate per unit of foliage was studied in two-year old cuttings of Norway spruce (Picea abies (L.) H. Karst.), representing four clones, at varying temperature and soil moisture. The CO₂ compensation point (Γ), photorespiration, dark respiration, and water balance were also investigated. All these characteristics indicated differences among the clones. A correlation between CO₂ exchange and transpiration suggested that stomatal control determined at least a part of this variation during a favourable water balance. An inverse relationship existed between Γ and net photosynthetic rate, and the same curvilinear model explained this variation in unstressed as well as stressed plants at a given temperature. An increase in Γ seems to be a normal result of water stress, particularly at high temperature, indicating an increase in mesophyll resistance to CO₂ diffusion. This result was in agreement with calculated values of mesophyll resistance. It also supported our earlier conclusions about the significance of mesophyll resistance during water stress.

• Luukkanen, E-mail: ol@mm.unknown

• Li, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O. Box 416, Chengdu 610041, P. R. China E-mail: licy@cib.ac.cn
• Zhang, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O. Box 416, Chengdu 610041, P. R. China; Graduate School of the Chinese Academy of Sciences, Beijing 100039, P.R. China E-mail: xz@nn.cn
• Liu, Sichuan Academy of Forestry, Chengdu 610081, P. R. China E-mail: xl@nn.cn
• Luukkanen, Viikki Tropical Resources Institute, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ol@nn.fi
• Berninger, Département des sciences biologiques, Cp 8888 succ centre ville, Université du Québec à Montréal, Montréal (QC) H3C 3P8, Canada E-mail: fb@nn.ca

• Liu, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O. Box 416, Chengdu 610041, P. R. China; Sichuan Academy of Forestry, Chengdu 610081, P. R. China E-mail: xl@nn.cn
• Xu, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O.Box 416, Chengdu 610041, P. R. China E-mail: hx@nn.cn
• Berninger, Département des sciences biologiques, Cp 8888 succ centre ville, Université du Québec à Montréal, Montréal (QC) H3C 3P8, Canada E-mail: fb@nn.ca
• Luukkanen, Viikki Tropical Resources Institute, P.O. Box 28, FI-00014 University of Helsinki, Finland E-mail: ol@nn.fi
• Li, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O.Box 416, Chengdu 610041, P. R. China; Viikki Tropical Resources Institute, P.O. Box 28, FI-00014 University of Helsinki, Finland E-mail: licy@cib.ac.cn