A close relationship between photosynthetic capacity and nitrogen concentration of leaves is known to exist. In conifers, nitrogen also affects the pattern of mutual shading within a shoot, which is a basic unit used in studying photosynthesis of coniferous trees. These effects of needle nitrogen concentration on photosynthetic capacity and mutual shading of needles were analysed for Scots pine (Pinus sylvestris L.) shoots taken from five young stands growing on sites of different fertility. The effect of nitrogen concentration on needle photosynthesis was studied based on measurements of the photosynthetic radiation response of shoots from which two thirds of the needles were removed in order to eliminate the effect of within shading.
An increase of one percentage unit in nitrogen concentration of needles increased the photosynthetic capacity of needles by 25 mg CO2 dm-2h-1. The effect of nitrogen on within-shoot shading was quantified in terms of the silhouette area to total needle area ratio of a shoot (STAR), which determines the relative interception rate per unit of needle area on the shoot. Although nitrogen promoted needle growth, an increase in nitrogen concentration decreased the within-shoot shading. This effect resulted from a decrease in needle density on the shoot and an increased needle angle with increasing nitrogen content.
The PDF includes an abstract in Finnish.
The effect of drainage on structure of tree stands is analysed by comparing the average structural characteristics (e.g. diameter distribution) of stands in the data for different drainage age classes and selected site types. The material consists of ca. 4,400 relascope sample plots, which are part of a large drainage area inventory project. The uneven-aged structure of the virgin peatland forest is preserved for several decades after drainage. This is enhanced by the post-drainage increase of small-diameter trees, especially birch. The number of trees per hectare increased during a period of ca. 30 years and levelled off thereafter. The increase in the number of saw log stems is clearly related to the fertility of the site and its geographical location.
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The earliest manor parks, which are a special form of cultivated forests, were created at the end of the 18th century. The surrounding of the main buildings was divided into two parts, an aesthetic park and yard serving household and economic purposes. Early in the 19th century, large parks were created which represented dominant aesthetic ideals but, on the other hand, formed a ”wild” counterpart to the structured inner world of the main building. A good example is Ratula Manor and its park, which represent the diversity of the cultivated forest of the 19th century manors.
The paper is based on a lecture given in the seminar ‘The forest as a Finnish cultural entity’, held in Helsinki in 1986. The PDF includes a summary in English.
Variation in tracheid morphology were examined for the bole wood of lodgepole pine (Pinus contorta Douglas ex Loudon) grown in Southern and Central Finland. Tracheid lengths were examined in a fast-grown and in slow-grown trees from three stands. Tracheid length increased with increasing height to 4–8 metres and decreased after that, and increased also with increasing age from the pith. The variation between stems was high. The shortest tracheids were about 1.11 mm near the piths and the longest about 4.10 mm near the bark.
Tracheid diameter and cell wall thickness were measured for the total number of 16 stems from Southern and Central Finland. Tracheid diameter increased with increasing distance from pith and the largest tracheids were at a height of 4–8 metres. Cell wall thickness varied independently of height in the stem. Summerwood cell wall thickness was twice that of springwood. There was a difference of 0.6 μm in springwood and 1.0 μm in summerwood double cell wall thickness between the two stands. Cell wall percentage was 29±4.7 in springwood and 69±7.3 in summerwood.
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A study based on four young Scots pines (Pinus sylvestris L.) showed that the number of needle-covered shoots per crown volume unit was independent on tree position representing a constant value of 600–700 shoots/m3. This was true, even though the total shoot number decreased with deteriorating tree position. In tree crown there were fourth-order shoots in good light conditions but only first- and second-order-shoots, when light conditions were poor. The length of shoots decreased in accordance with increasing order of the shoot.
The share of the needle biomass and growth increased, when the shoot order increased. Similarly, the share of needles increased with deteriorating tree position. This was especially true in the upper crown. On the other hand, the share of the crown from the total biomass and growth increased with improving tree position. The percentage of crown system of a dominant tree in a sparse stand was 64% of that of biomass and 83% of that of growth. The corresponding values for a suppressed tree in a dense stand were 36% and 35%. The growth of wood, bark and needles in crown systems was linearly correlated with prevailing light conditions around the branch. It is evident that the tree position and light condition within the stand control the wood, bark and needle growth in the crown system and their interrelationships.
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The publication comprises proceedings of a conference held in Helsinki in 1981. Forest tree populations are investigated for population genetic structure, mating systems, mechanisms of genetic adaptation and ecological adaptation. Methods and techniques used in population genetic research of forest trees are presented. Much concern is given to applications by means of forest tree breeding, particularly the seed orchard breeding technique. Generally, the application of population genetics in cultivated forests is discussed.
The PDF includes a preface and the presentations of the conference (25 short papers) in English, and a comprehensive summary of the themes of the conference in Finnish.
The effect of soil compaction on transpiration and height increment of Scots pine (Pinus sylvestris L.) seedlings following planting out were investigated in a small-scale pot experiment. Compacted and loosely-packed fine sand and fine-sand moraine were used as the planting substrates. The compacted soils used corresponded to the normal type of soil to be found in tilled forest soils in Finland. The effect of soil compaction on seedling transpiration during water stress was also studied in a separate experiment.
Seedlings planted in compact soil had a higher rate of transpiration than those in loosely-packed soil. The recovery in transpiration, which started halfway through the growing season, was faster, however, in the seedlings planted in loosely-packed soil. Under conditions of water stress, the seedlings planted in compact fine-sand moraine started to reduce the transpiration rate at higher soil moisture values than those planted in loosely-packed soil. No corresponding difference was observed for fine sand. Compaction was not found to affect the overall height growth, but it did at certain time during the growing season.
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In this study the width and height of 1,588 uniseriate and 454 fusiform rays were measured from tangential sections of four Scots pine (Pinus sylvestris L.) trunks. The samples represented various height levels and distances from the pith. The average width of the uniseriate rays was 19.7 μm and that of the fusiform rays, 51.9 μm. The average height of the uniseriate rays was 215.7 μm and that of the fusiform rays 406.2 μm. Due to this difference in height, it may be possible to develop an automatic system for distinguishing between uniseriate end fusiform rays on the basis of their height.
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The study is continuation of the earlier structure and growth studies of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) in Forest Research Institute. The material represents birch stands (Betula verrucosa, now B. pendula, and B. Pubescens L.) in Southern Finland. The stands were treated with different fellings, and in regard to their silvicultural condition classified as good, satisfactory and unsatisfactory. Height of the trees, height of living crown, volume, increment and volume increment and development of stem diameter series was measured.
The most characteristic difference between the silviculturally good and poor stands was that the the annual increment of the good stands concentrated into large size trees, and the increment of unsatisfactory stands into small and inferior trees.
It is concluded that if the aim of stand treatment is to produce large and high quality volume increment, the most favourable stand volume of birch stands, compared with naturally normal stand volume, seems to be 90-85% at the age of 41-55 years, and 80-70% at the age of 56-65 years. If growth of large size trees is aimed at, the maximum number of the dominant trees per hectares cannot be more than 400 at the age of 50-60 years.
The article includes a summary in English.
The investigation concerns with the strength of the eccentric growth accompanying formation of tension wood in silver birch (Betula pendula Roth.) and downy birch (Betula pubescens Ehrh.), behaviour of wood in wood-working machines and its macroscopic characteristics, its microscopic and sub-microscopic structure, chemical composition, resistance against certain chemicals, physical properties, and the strength characteristics of wood.
The most detrimental properties of tension wood used in wood working industry are high longitudinal shrinkage, warping, twisting and checking. The wooliness of the cut is unwanted, for instance, in plywood and furniture. In pulp industry tension wood is better raw material than normal wood because it yields more and purer cellulose than normal wood. However, it has poorer strength properties.
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Permanent sample plots are considered to be the most reliable basis for investigations into structure and development of stands. Such sample plots, established since 1924 in Finland, have been used to study thinnings of varying intensity. These studies are yet too short to give comprehensive conclusions. It is also possible to base the studies on sample plots measured in managed forests and gain in this way information suitable for practical purposes. In this investigation development of stands treated by two different methods, repeated thinnings and repeated selection cutting were studied in pure, even-aged Scots pine (Pinus sylvestris L.) stands in Southern Finland, on three forest types.
The results show that volume increment level of naturally normal stands seem to have been reached easily by stands treated with repeated thinnings. With advancing age, the growing stock of thinned stands fall short from the natural stands. As thinnings have removed primarily the poorest trees, the increment is distributed over trees of a larger size more in thinned than in naturally normal stands.
When intensive cuttings have resulted in a relatively small growing stock, the decrease in volume increment leads to considerable decrease in volume. The size of the tree has no essential effect – within certain limits - on the volume increment of the stand, if the volume removed is similar. However, every intermediate thinning removing largest-sized trees may result in the prolonged rotation. Since the volume increment of an older stand is much smaller than earlier, intermediate thinnings removing largest-sized trees should be avoided if the aim is the greatest volume yield. The growing stock of middle-aged or older stands untreated or treated with slight cuttings only can as a rule be considerably reduced without volume increment declining.
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The article includes a detailed review on the technical properties of wood. The weight, water content, strength and conductivity of the wood, and the factors affecting them are discussed. The mechanical and technical properties of wood are influenced, for instance, by tree species, age and part of the tree, geographical situation, site and growth conditions of the stand, anatomical structure of the wood, and temperature. The author summarizes the topics where further research should be addressed. For instance, the forest site types developed in Finland could be utilized in studies of the mechanic technical properties of wood.
The PDF includes a summary in English.
The paper is the final report of a study on the estimation of value increment and inherent variables of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) stands. The main aim was to obtain improved criteria for decision-making concerning the priority of stands for regeneration.
The construction of various estimation models and their reliability are discussed in detail. The study, together with some previous papers, has resulted in a system which on the basis of a number of easily assessed stand variables gives for the stands concerned the volume of stems, percentages of timber assortments, stumpage value, volume increment and value increment.
The following examples are given with regard to the practical application of the results, in addition to the determination of the relative maturity of stands: 1) The study of various trends in stand development; the comparison between the volume and value variables. 2) The estimation of timber assortments needed for a cutting budget, trees marked for felling etc. 3) The calculation of the value of forests.
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The report concludes a series of studies on the early development of young Scots pine (Pinus sylvestris L.) stands. The basis assumption made in the study series was that the within-stand light regime is the main driving force for total tree growth and its allocation of photosynthates for crown, stem and root growth. An individual tree growing in a stand under a varying light regime which is controlled by the stand structure, is the basic unit used in the study. The photosynthesis of an individual tree is determined by the light regime. The stand is formed from individual trees.
The model is applied in simulation of the growth and development of tree stands. Several computer runs representing various densities, height distributions and tree species mixtures were carried out. Potential application areas, properties of the model and future needs of investigations are discussed.
The PDF includes a summary in English.