Articles containing the keyword 'frost'

The productivity of Scots pine (Pinus sylvestris L.) under changing climatic conditions in the southern part of Finland was studied by scenario analysis with a gap-type forest ecosystem model. Standard simulations with the model predicted an increased rate of growth and hence increased productivity as a result of climatic warming. The gap-type model was refined by introducing an overwintering sub-model describing the annual growth cycle, frost hardiness, and frost damage of the trees. Simulations with the refined gap-type model produced results conflicting with those of the standard simulation, i.e., drastically decreased productivity caused by mortality and growth-reducing damage due to premature dehardening in the changing climate. The overwintering sub-model was tested with frost hardiness data from Scots pine saplings growing at their natural site 1) under natural conditions and 2) under elevated temperature condition, both in open-top chambers. The model predicted the frost hardiness dynamics quite accurately for the natural conditions while underestimating the frost hardiness of the saplings for the elevated temperature conditions. These findings show that 1) the overwintering sub-model requires further development, and 2) the possible reduction of productivity caused by frost damage in a changing climate is less drastic than predicted in the scenario analysis. The results as a whole demonstrated the need to consider the overwintering of trees in scenario analysis carried out with ecosystem model for boreal conditions. More generally, the results revealed a problem that exists in scenario analysis with ecological models: the accuracy of a model in predicting the ecosystem functioning under present climatic condition does not guarantee the realism of the model, nor for this reason the accuracy for predicting the ecosystem functioning under changing climatic conditions. This finding calls for the continuous rigorous experimental testing of ecological models used for assessing the ecological implications of climatic change.

• Hänninen, E-mail: hh@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown
• Leinonen, E-mail: il@mm.unknown
• Repo, E-mail: tr@mm.unknown

Two dynamic models predicting the development of frost hardiness of Finnish Scots pine (Pinus sylvestris L.) were tested with frost hardiness data obtained from trees growing in the natural conditions of Finland and from an experiment simulating the predicted climatic warming. The input variables were temperature in the first model, and temperature and night length in the second. The model parameters were fixed on the basis of previous independent studies. The results suggested that the model which included temperature and photoperiod as input variables was more accurate than the model using temperature as the only input variable to predict the development of frost hardiness in different environmental conditions. Further requirements for developing the frost hardiness models are discussed.

• Leinonen, E-mail: il@mm.unknown
• Hänninen, E-mail: hh@mm.unknown
• Repo, E-mail: tr@mm.unknown

Budburst timing and the relationship to storage temperature and duration were investigated in four varieties (entries) of 1–2 metres tall silver birch (Betula pendula Roth) trees. A total of 2,160 shoots were sampled, and the material stores in darkness at 0, 3 or 6 °C from November 29, 1993. When the shoots were placed in storage, they had been through a period of 29 days with temperatures below 0°C (since October 15). By that time the autumn dormancy was assumed already broken, and the trees were expected to respond to increased temperature by bud development. On January 4, 1994, and on four subsequent dates, January 19, February 1, March 4 and March 17, shoots were taken out of storage and set in growth chambers at 9, 12 or 15°C. The time to budburst was recorded.

Duration of storage, storage temperatures and varieties were all highly significant for budburst. The interaction terms were of less statistical importance. Based on the contrast between the three different growth chamber environments, three different methods were used to calculate the threshold temperatures for each entry. In spite of the pre-selection of variable budburst performers, the threshold values, varying between 0°C to -2°C, could not be shown to be statistically different. According to the results, the time of budburst changes in accordance with both winter and spring temperatures, being extremely early after a mild winter and warm spring, given sufficient autumn chilling. The similarities in the threshold temperatures indicate that the ranking in earliness between varieties will most likely be the same from year to year without regard to climate change.

• Skuterud, E-mail: rs@mm.unknown
• Dietrichson, E-mail: jd@mm.unknown

Temperature sums required for budburst in various Norway spruce (Picea abies (L.) H. Karst.) provenances were determined, and weather statistics were then used to predict the risk of potentially damaging frosts at 11 locations in Sweden. Frost risk was quantified as the probability of a frost occurring within 100 day-degrees (two weeks) after budburst. The examples provided show that a spruce seedling from central Sweden has to sustain almost twice as many frost occassions as a seedling from Belorussia, when planted in southern and central Sweden. The method presented here can be used for mapping early summer frost risk in Sweden and for supporting provenance transfer guidelines.

• Hannerz, E-mail: mh@mm.unknown

According to a recently presented hypothesis, the predicted climatic warming will cause height growth onset of trees during mild spells in winter and heavy frost damage during subsequent periods of frost in northern conditions. The hypothesis was based on computer simulations involving a model employing air temperature as the only environmental factor influencing height growth onset. In the present study, the model was tested in the case of eastern Finnish Scots pine (Pinus sylvestris L.) saplings. Four experimental saplings growing on their natural site were surrounded by transparent chambers in autumn 1990. The air temperature in the chambers was raised during the winter to present an extremely warm winter under the predicted conditions of a double level of atmospheric carbon dioxide. The temperature treatment hastened height growth onset by two months as compared to the control saplings, but not as much as expected on the basis of the previous simulation study. This finding suggests that 1) the model used in the simulation study needs to be developed further, either by modifying the modelled effect of air temperature or by introducing other environmental factors, and 2) the predicted climatic warming will not increase the risk of frost damage in trees as much as suggested by the previous simulation study.

The PDF includes an abstract in Finnish.

• Hänninen, E-mail: hh@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown
• Laitinen, E-mail: kl@mm.unknown
• Pajari, E-mail: bp@mm.unknown
• Repo, E-mail: tr@mm.unknown

Seedlings from four Norway spruce (Picea abies (L.) H. Karst.) stands originating from areas with effective temperature sums ranging from 710 d.d. to 1,150 d.d. were raised under artificial light and temperature treatment. After a 10-week growing period the hardening process was started by subjecting the seedlings to +8°C night temperature and +15°C day temperature, and increasing the night length by 1.5 hour/week. Hardiness was measured by means of artificial freezing treatment (-10°C or -15°C), followed by visual estimation of the degree of needle injury. The stem height, lignification and bud development were measured before the freezing treatment. The amount of injury increased the more southern the origin of the tested material was. Furthermore, the proportion of non-lignified part of the seedling stem was negatively correlated with the latitude of the provenances. The proportion of seedlings with clearly visible buds was more than 90% in the northernmost entry and less than 1% in the southernmost one. The overall correlation coefficient between the needle injuries and the proportion of non-lignified part of the stem was rather high, but varied considerably from 0.3 in the northernmost material to over 0.6 in the southern provenances. According to the results, it seems to be possible to use growth characteristics as an indicator of frost hardiness at the provenance level.
The PDF includes an abstract in Finnish.

• Pulkkinen, E-mail: pp@mm.unknown

The ability of one-year old Scots pine (Pinus sylvestris L.) seedlings to reharden during the dehardening period was studied. Naturally hardened quiescent seedlings were preconditioned at 0°C for ten days and then placed in chambers at different forcing temperatures with different light regimes. The forcing periods were followed by cool periods. Changes in frost hardiness were monitored at intervals using freeze tests of whole plants. Frost hardiness was assessed by three methods: impedance, survival and growth retardation. Dehardening seemed to be a partially reversible process, i.e. in some growing conditions slight rehardening was found.

The PDF includes an abstract in Finnish.

• Repo, E-mail: tr@mm.unknown

The relationships between bud dormancy and frost hardiness were examined using two-year-old Pinus sylvestris L. seedlings. The chilling temperatures used were +4 and -2°C. To examine the dormancy release of the seedlings, a forcing technique was used. Frost hardiness was determined by artificial freezing treatments and measurements of electrical impedance. At the start of the experiment, the frost hardiness of the seedlings was about -25°C. After the rest break, the seedlings kept at +4°C dehardened until after eight weeks their frost hardiness reached -5°C. At the lower chilling temperature (-2°C) the frost hardiness remained at the original level. When moved from +4 to -2°C, seedlings were able to reharden only after the time required for bud burst in the forcing conditions had reached the minimum.

The PDF includes an abstract in Finnish

• Valkonen, E-mail: mv@mm.unknown
• Hänninen, E-mail: hh@mm.unknown
• Pelkonen, E-mail: pp@mm.unknown
• Repo, E-mail: tr@mm.unknown

The winter 1986–87 was unusually cold; the snow cover remained thin and consequently the soil froze to a considerable depth. In spite of the severe frost, the lowest temperatures measured at the ground surface was -10.3°C and in the soil at the depth of 10 cm -5.8°C. The temperature sum of the following summer was unusually small and the soil frost melted more slowly than usual. The winter frosts did not have a decisive influence on the survival of planted seedlings.

The PDF includes an abstract in English.

• Kubin, E-mail: ek@mm.unknown

Seven hundred one-year-old Betula pendula Roth seedlings were given different concentrations of potassium fertilizer. Over the study period seedlings were subjected to artificial growing and dormant phases. Frost resistance of the seedlings was assessed by artificial freezing tests and electrical impedance measurements on stem cuttings. In general, high concentrations of potassium fertilizer reflected a low tolerance to frost. Pre-freezing impedance readings decreased with increasing potassium fertilizer dosages. Results from pre-freezing impedance measurements were found to be in broad agreement with the hypothesis that high impedance readings indicate a frost hardy tissue whereas low readings imply the opposite.

The PDF includes an abstract in Finnish.

• Jozefek, E-mail: hj@mm.unknown

Frost resistance during shoot elongation in seedlings of Norway spruce (Picea abies (L.) H. Karst.) was studied in two experiments. The aim of the first study was to evaluate the effect of varying mineral nutrition. Except for potassium, only minor differences in mineral elements concentrations were established, presumably due to low levels of irradiance and thus a low rate of dry matter production. No significant differences in frost injuries were found between the treatments in the experimental series, but the control seedlings were significantly less injured. It is assumed that poor hardiness development at the end of one growth period resulting from low levels of irradiance may decrease the frost resistance during the next shoot elongation phase. Observations from the second experiment with Norway spruce nursery stocks representing different seedling ages and production systems, support this assumption.

The PDF includes an abstract in Finnish.

• Rostad, E-mail: hr@mm.unknown

Nursery grown Norway spruce (Picea abies (L.) H. Karst.) seedlings from 12 different seed orchards were tested for early autumn frost hardiness using artificial freezing tests. Seed orchards containing grafted parent clones originating from high altitudes produced seedlings showing higher damage than commercial control seed lots of the commercial controls. A seed orchard containing both German and Norwegian clones produced seedlings showing high damage. The correlation between bud-set and frost damage was high at the provenance level, but lower at the half- and full-sib-levels. Families with good growth capacity in progeny field tests showed large between-family variation in frost damage in the artificial freezing tests. This indicates the possibility to combine high growth rate with acceptable autumn frost hardiness in the selection of parent trees.

The PDF includes an abstract in Finnish.

• Johnsen, E-mail: –
• Apeland, E-mail: ia@mm.unknown

Visible frost damage to forest trees in Sweden seldom occurs in winter but is frequent in late spring, summer and early autumn. Frosts are frequent in all seasons in various parts of Sweden, even in the southernmost part (lat. 56°, N) and temperatures may be as low as -10°C even around mid-summer. Ice crystal formation within the tissues, which in most seedlings takes place at around -2°C, causes injury, not the sub-zero temperatures themselves.

The apical meristem, the elongated zone, and the needles of seedlings of Picea abies (L.) H. Karst. in a growing phase were damaged at about -3°C and those of Pinus sylvestris L. at about -6°C. Other species of the genus Pinus were tested and most were found to be damaged at about -6°C, with some variations. Picea species tested were damaged at about -3°C to -4°C.

A method has been designed to compare the response of different species to winter desiccation, which occurs under conditions of (1) low night temperature, (2) very high irradiation and increase in needle temperature during the photoperiod, (3) frozen soil, and (4) low wind speed. There were differences in response to winter desiccation between pine and spruce species. Seedlings of Pinus contorta tolerated these winter desiccation conditions much better than those of P. sylvestris or Picea abies. Picea mariana was the least tolerant of the species tested.

The PDF includes an abstract in Finnish.

• Christersson, E-mail: lc@mm.unknown
• Fircks, E-mail: hf@mm.unknown

Electrical impedance characteristics of plant cells are dependent on such physiological factors as physiological condition, developmental stage, cell structure, nutrient status, water balance and temperature acclimation. In the measurements also such technical and physical factors as type of electrodes, frequency, geometry of the object, inter-electrode distance and temperature have an effect. These factors are discussed especially with respect to the impedance method in frost resistance studies of plants.

The PDF includes an abstract in Finnish.

• Repo, E-mail: tr@mm.unknown

This issue of Silva Fennica consists of eight articles, which are based on a co-nordic conference ”Frost hardiness and over-wintering in forest tree seedlings”, held in Joensuu, Finland, during December 1–3, 1986. The whole annual cycle of the trees is considered. Emphasis is given on methods for the study of frost hardiness, genetic variation in frost hardiness, nitrogen metabolism, bud dormancy release, and joint effect of natural and anthropogenic stress factors in the winter damage of forest trees. Practical implications for tree breeding and nursery management are discussed.

The PDF includes an abstract in Finnish.

• Hänninen, E-mail: –
• Pelkonen, E-mail: –

The distribution of the minimum temperatures after the beginning of the active period (one temperature for each spring) have been calculated for each principle using daily meteorological data collected during the years 1883–1980. The efficiency criterion is the variance of the minimum temperature distributions and the length of the active period. The most efficient regulation principle is found to be based on the temperature sum which includes a feedback component.

The PDF includes a summary in Finnish.

• Häkkinen, E-mail: rh@mm.unknown
• Hari, E-mail: ph@mm.unknown

Young Norway spruce (Picea abies (L.) H. Karst.) are susceptible to early summer frost damage. Birch (Betula pubescens Ehrh.) naturally colonize rich or fairly rich drained peatlands after clear cutting, and can provide protection for developing seedlings. The report describes the development of spruce stands after various types of handing of the birch nurse crops.

Different proportions of birch and spruces did not have any influence on the spruce stand production. In cases where the nurse crop stand is removed when the spruce stand age was 20 years and height 4 m the spruce suffered badly but recovered with time, reaching the spruce stand growing under a nurse stand within the next 20 years. The height growth of spruce depends on the density of the nurse stand, especially on fertile sites. The development of diameter growth also depends on the density of the nurse trees. Removal of the nurse stand in spruce stands on the sites concerned should be done when the spruce stand is 20 years old and at the height of 4 m.

The PDF includes a summary in English.

• Heikurainen, E-mail: lh@mm.unknown

During the next decade there will be a marked increase in the allowable cut in drained peatlands. At the same time, the mechanization in logging proceeds, and in short-distance haulage the use of forwarders will increase. This study, based on literature and some observations, deals with logging conditions in drained peatlands with special reference to the suitability of heavy logging machines for use in such terrain. In addition, soil frost and the bearing capacity of the frozen peat soil were studied.

Freezing of the soil in a drained peatland area depends prevailingly on the weather conditions during early winter. The factors influencing soil freezing of a drained peatland are completely different from those regulating the freezing of natural peat soils. The frost penetrates in general deeper in the drained than virgin peatland. The topmost peat layer does not, however, freeze uniformly. Generally speaking, the bearing capacity of a drained peat soil is lower than that of undrained peat due to lower water content.

It is concluded that heavy logging machines are probably not fitted for use in drained areas on peatland even if the average soil frost values recorded would suggest it. Moreover, because of their extremely superficial root systems, peatland forests are exposed to damages by heavy machines in thinning operations.

The PDF includes a summary in English.

• Hannelius, E-mail: sh@mm.unknown

The paper describes the results obtained from an investigation into the effect of thinning of different intensity and fertilization on the depth and water equivalent of the snow cover as well as on the depth of the soil frost in a young Scots pine (Pinus sylvestris L.) stand growing on drained peatland in Central Finland. Thinnings and fertilization was carried out in 1968, and the snow cover was followed in the winters 1970/71 and 1971/72.

Only extremely heavy thinnings (60% of the volume) seemed to increase the depth and water equivalent of the snow cover. The indirect effect of fertilization on the snow cover was insignificant. In the clear-cut sample plot of the study, soil frost was either not found at all or the depths of the frozen soil layer was smaller than in the other plots. When deciding the silvicultural measures to be taken in the case of tree stands growing on drained peatlands, there seems to be reason to avoid radical thinnings. Otherwise, the favourable influence of the trees on a site on its water relationships will be diminished.

The PDF includes a summary in English.

• Päivänen, E-mail: jp@mm.unknown

In the eastern parts of South-Finland the growing season of 1967 was highly favourable, which resulted in good height growth during the following year. During the summer 1968, temperature conditions were unfavourable, while the middle of summer was cold and the later part of the growing season unusually hot. The following winter had exceptionally cold spells from January to March, which caused Norway spruce (Picea abies (L.) H. Karst.) abundant winter frost damages such as dead shoots and buds, and destroyed needles.

These damages occurred particularly in stands with height of 0.5–3 m, and the occurrence of damages seemed to concentrate to the parts of saplings that had been immediately above the snow cover. Detailed observations on spruce plantations growing under a dense nurse stand of alder (Alnus sp.) indicated that explicitly the top shoots suffered from damages and not so much the laterals. When the needles of the leader suffered from minor damages, the shoot continued to grow normally. Still, sometimes a branch took over and became a new leader. If only the leader bud was killed, further stem development became dependent on one of the topmost lateral buds. When the upper part of the leaded died, one of the lateral shoots at its base usually became the new leader.

The PDF includes a summary in English.

• Yli-Vakkuri, E-mail: py@mm.unknown

The objective of this investigation was to study the influence of stand density of white birch (Betula pubescens Ehrl.) on the minimum temperatures in the stand during the growing season, and the actual minimum temperatures of the leading shoot of Norway spruce (Picea abies (L.) H. Karst.) seedlings growing in the open. The 40-year-old uniform white birch stand was situated in 142 m above the sea level in Southern Finland. The stand was treated with thinnings of three different densities in 1961.

Air temperature was recorded in four sample plots at heights of 0.1 m, 0.5 m, 1.0 m, 2 m and 4 m. In the stand of moderate density, temperatures were measured at heights of 6.0 m, and in the stand of full density at 6.0 m, 8.0 m and 10.0 m.

The temperature differences between stands of various densities proved to be rather small. Especially the thinnest stand differed very little from the open area. The soil surface has in all cases been warm compared with the higher air layers indicating meadow-fog-type by Geier (1965). On cloudy or windy weather all the temperature profiles in the various stands resembled each other. The difference between the air temperature and temperature of the spruce shoot was greatest at midnight and decreased steadily thereafter.

The problem in using shelter stands for spruce regeneration areas is that optimum shelter stand density is difficult to define. Already a thin shelter stand causes drawbacks to the young seedlings, but in order to be effective enough against early frosts, the shelter stand should be comparatively dense.

The PDF includes a summary in English.

• Leikola, E-mail: ml@mm.unknown
• Pylkkö, E-mail: pp@mm.unknown

The aim of the investigation was to obtain by snow and soil frost observations sufficient material for determination of regional springtime snow and soil frost values, because the water equivalent of snow and the frost depth affect runoff. The present paper elaborates a method by which the observations along a survey line can be corrected to be valid for a basin. Along the line 50 measurement points were arranged at specific intervals. Snow depth was recorded at each point, and snow density and frost depth at every fifth point. The terrain was studied along the line and the terrain of the survey points were classified in eight classes depending on the vegetation. The classes ranged from cultivated lands and open bogs to wooded areas according to volume of the growing stock and tree species composition.

The mean snow depth was 51.9 cm and mean snow density 0.235 g/m². Water equivalent of snow in class 4 terrain (forest with small growing stock) was 30% higher than in class 8 (forest with high growing stock). An ample stand increases evaporation in wintertime. The difference can be partly caused by the different accumulation of snow in the different types of stands.

Soil type was not found to have any distinct influence on the frost depth in the present material. On cultivated lands the soil frost clearly penetrates to greater depth than in the forest. The growing stock of wooded areas influences the snow depth

The PDF includes a summary in English.

• Mustonen, E-mail: sm@mm.unknown

Trees and shrubs of foreign origin have been grown in Finland at least from the 1700th century. At the State Horticultural Institute in the neighbourhood of the town Turku in southwestern coast of the country, a number of ornamental trees and shrubs have been planted since 1927. During the first decade, weather conditions were quite favourable, but the winters in 1939–1940 were so severe, that only the hardiest plants survived.

It would be important to study hardiness and suitability of the various woody plants cultivated in the different parts of the country. This paper includes notes of the survival of the tree species and shrubs so far planted at the Institute.

The PDF includes a summary in English.

• Meurman, E-mail: om@mm.unknown

Snow cover and ground frost was studied in 29 forest stands in Southern and Central Finland in 1957–1959. The tree species influenced greatly accumulation of snow on the forest floor. Norway spruce (Picea abies (L.) Karst.) retains snow in its crown. In addition, snow and water falling from the branches compress the snow cover under the trees, and the ground freezes deeper because of the shallow snow cover. In the spring, the dense crown prevents rain and radiation reaching the ground, which remains cold longer. However, ground frost may protect spruce, which has a weak root system, from wind damages.

Scots pine (Pinus sylvestris L.) has similar, but milder, effects on snow cover within the forest. The crowns of pine seedlings and young trees pass snow easily, but later the crowns intercept it considerably. The lower branches are, however, high up and the snow is evenly spread on the ground. The deciduous trees intercept little snow and in the spring the snow smelts and the frozen soil thaws early. The snow conditions of deciduous forests are, however, changed by a spruce undergrowth.

It can be assumed that the unfavourable conditions in spruce forests can be alleviated by thinning. Also, mixture of pine and deciduous trees can transform the conditions more favourable in the spruce stands.

The PDF includes a summary in English.

• Yli-Vakkuri, E-mail: py@mm.unknown

The aim of the investigation was to study natural regeneration of Norway spruce (Picea abies (L.) Karst.) in drained peatlands and frost injuries in seedlings, and to compare microclimates of the regeneration areas. The experiments included peatlands in Satakunta in Western Finland. Restocking of the areas with seedlings and their survival was followed in 1935-40 at sample plots that were mainly 1 are large.

Susceptibility to freezing was shown to be dependent on the stage of development of the shoots. Shoots that have just begun to grow contain little water, and withstand better freezing temperatures than shoots in later stages of growth. Damages to the seedlings were observed when the temperatures decreased to -2.8–-4.3 °C. The most severe damage to a seedling was caused by the death of the leading shoot by spring frost.

Norway spruce regenerates easily on moist peatlands, but peatlands with dry surface tend to have little or no seedlings. The species regenerated better in marshy sites than correspondingly fertile mineral soil sites. However, it needs shelter to avoid frost damage. On clear cut spruce swamp the undergrowth spruce seedlings that were left in the site got severe frost damage. If the site had birch (Betula sp.) coppice or undergrowth, spruce seedlings survived in their shelter depending on the height and density of the birch trees. To be effective, the protective forest should have relatively even crown cover. Young spruce seedlings could grow well even under relatively dense birch stand.

The PDF includes a summary in German.

• Multamäki, E-mail: sm@mm.unknown

Soil frost from the winter 1941-1942 was of a special form. To create such a strong soil frost there need to be capillary soil form and long period of permanent and intense cold, meaning minus temperatures.  The frost layer was thicker than normally.

Such a frost causes many problems on the fields and roads when the soil rises because of the frost and then falls unevenly back when the frost melts.

• Kokkonen, E-mail: pk@mm.unknown

The drained peatlands regenerate usually well, and artificial regeneration by sowing or planting has been rare. Field trials of Norway spruce (Picea abies (L.) H. Karst.) were established in northern Satakunta in Western Finland in three drained peatlands in 1934. Sowing trials of Norway spruce consisted of patch and broadcast sowed sample sites in treeless bogs and under protective forest. The seedlings of spruce were planted either under protective forest or in treeless peatland.

The results show that artificial regeneration of Norway spruce succeeds best under protective forest. The best tree species for upper storey is Betula sp. which grows fast and controls growth of ground vegetation. The peat is relatively decomposed on those peatlands that are suitable for spruce, and breaking of the surface of the peat is not recommended. In the sowing trials, breaking of the upper layer of the peat caused frost heaving, cracking of the dried surface and sticking of mud in the seedlings in the patch sown sample site. The shoot and root growth of seedlings of the broadcast sown site was better than seedlings of the patch sown site. The planted spruce seedlings seemed to be more susceptible for spring frost than the seedlings in the sown site. The plants of seed origin succeeded in general better than the planted seedlings.

The PDF includes a summary in German.

• Multamäki, E-mail: sm@mm.unknown

The article presents some basic elements of soil frost and its occurrence.  The data contains observations from different regions and soil types in Finland. Different forms of soil frost and factors affecting its formulation are discussed. 

• Kokkonen, E-mail: pk@mm.unknown

Silver birch (Betula pendula Roth) seed origins from the Baltic countries and from Finland were compared in terms of growth, wood density, bark thickness and the incidence of darkened core wood, frost cracks and decay, and the effect of seed origin latitude was examined in two Finnish provenance trials. The material consisted of 21 stand and single tree origins ranging from latitudes 54° to 63°N from the Baltic countries and Finland. The trials, measured at the age of 22 years, were located at Tuusula (60°21´N), southern Finland and at Viitasaari (63°11´N), central Finland. The Baltic origins were superior to the Finnish ones in the southern trial regarding height, whereas in central Finland the Finnish origins grew better. There was no consistent difference between the Baltic and the Finnish group of origins in wood density. Bark thickness decreased with increasing seed origin latitude. The Baltic origins had significantly thicker bark than the Finnish origins. A moderate positive correlation was detected between the seed origin latitude and the incidence of darkened core wood in the southern trial, where the darkened core wood was more common in the Finnish origins than in the Baltic ones. The highest proportion of trees with frost cracks was detected in the south-western Latvian origins growing in central Finland. Seed transfers from the Baltic would have an increasing effect on the bark thickness of birch logs, but no or only minor effects on wood density. Based on our results, there is no reason to recommend the use of non-native Baltic seed origins in Finland instead of the native locally adapted seed sources.

• Viherä-Aarnio, Natural Resources Institute Finland (Luke), Green technology, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: anneli.vihera-aarnio@luke.fi
• Velling, E-mail: pike.velling@phnet.fi

Topographic complexity in mountainous ecosystems strongly influences plant growth and as such also wood formation. This wood formation can possibly be used to understand topographic variation of the main climatic drivers, e.g. by modulating frost events. Here we test the influence of different slope exposures on the spatio-temporal distribution of frost rings in Siberian spruce (Picea obovata Ledeb.) in the Southern Urals, Russia. We sampled on two opposite slopes, northeast (NE) and southwest (SW), on three elevation levels from the highest single trees to closed canopy forest and analysed frost ring occurrence and their seasonal timing. Frost ring formation at all exposure-elevation combinations was common and mainly concentrated in the early part of the growing season. The age until trees record frost rings was equally similar (until about 35 years) on both slopes and different elevational levels with the exception of the climatically harshest site, the highest elevation on the NE slope. While we could not deduce a direct, easily identifiable climatic driver for the formation of frost rings, our analysis shows high potential to disentangle the complex interplay between climate, site condition and tree growth in mountainous ecosystems.

• Gurskaya, Institute of Plant and Animal Ecology (IPAE), Ural Branch of Russian Academy of Sciences , 8 Marta St. 202, 620144 Ekaterinburg, Russia E-mail: mgurskaya@yandex.ru
• Moiseev, Institute of Plant and Animal Ecology (IPAE), Ural Branch of Russian Academy of Sciences , 8 Marta St. 202, 620144 Ekaterinburg, Russia E-mail: moiseev@ipae.uran.ru
• Wilmking, Institute of Botany and Landscape Ecology, Soldmanstrasse 15, Ernst-Moritz-Arndt-University Greifswald, 17487 Greifswald, Germany E-mail: wilmking@uni-greifswald.de

• Li, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing 100083, China E-mail: gl@nn.cn
• Liu, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing 100083, China E-mail: lyong@bjfu.edu.cn
• Zhu, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing 100083, China E-mail: yz@nn.cn
• Li, Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Forest Silviculture of State Forestry Administration, Beijing 100091, China E-mail: qml@nn.cn
• Dumroese, US Department of Agriculture, Forest Service, Rocky Mountain Research Station, Moscow, ID, USA E-mail: rkd@nn.us

• Gregow, Finnish Meteorological Institute, Helsinki, Finland E-mail: hilppa.gregow@fmi.fi
• Peltola, University of Eastern Finland, Faculty of Forest Sciences, Joensuu, Finland E-mail: heli.peltola@uef.fi
• Laapas, Finnish Meteorological Institute, Helsinki, Finland E-mail: ml@nn.fi
• Saku, Finnish Meteorological Institute, Helsinki, Finland E-mail: ss@nn.fi
• Venäläinen, Finnish Meteorological Institute, Helsinki, Finland E-mail: av@nn.fi

• Kellomäki, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: seppo.kellomaki@uef.fi
• Maajärvi, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: mm@nn.fi
• Strandman, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: hs@nn.fi
• Kilpeläinen, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: ak@nn.fi
• Peltola, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: hp@nn.fi

• Hänninen, Plant Ecophysiology and Climate Change Group (PECC), Department of Biological and Environmental Sciences, Box 65, FI-00014 University of Helsinki, Finland E-mail: heikki.hanninen@helsinki.fi
• Luoranen, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: jl@nn.fi
• Rikala, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: rr@nn.fi
• Smolander, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: hs@nn.fi

• Luoranen, The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland E-mail: jaana.luoranen@metla.fi
• Konttinen, The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland E-mail: kk@nn.fi
• Rikala, The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland E-mail: rr@nn.fi

• Gregow, Finnish Meteorological Institute, P.O. Box 503, FI-00101 Helsinki, Finland E-mail: hilppa.gregow@fmi.fi
• Puranen, Finnish Meteorological Institute, P.O. Box 503, FI-00101 Helsinki, Finland E-mail: up@nn.fi
• Venäläinen, Finnish Meteorological Institute, P.O. Box 503, FI-00101 Helsinki, Finland E-mail: av@nn.fi
• Peltola, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: heli.peltola@uef.fi
• Kellomäki, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: seppo.kellomaki@uef.fi
• Schultz, Finnish Meteorological Institute, P.O. Box 503, FI-00101 Helsinki, Finland E-mail: ds@nn.fi

• Hytönen, Finnish Forest Research Institute, Kannus Research Unit, P.O. Box 44, FI-69101, Kannus, Finland E-mail: jyrki.hytonen@metla.fi
• Jylhä, Finnish Forest Research Institute, Kannus Research Unit, P.O. Box 44, FI-69101, Kannus, Finland E-mail: pj@nn.fi

• Hänninen, Plant Ecophysiology and Climate Change Group (PECC), Department of Biological and Environmental Sciences, Box 65, FI-00014 University of Helsinki, Finland E-mail: heikki.hanninen@helsinki.fi
• Kramer, Alterra, P.O. Box 47, 6700 AA Wageningen, The Netherlands E-mail: kk@nn.nl

• Leinonen, University of Oklahoma, Department of Botany and Microbiology, Norman, OK 73019, USA E-mail: leinonen@ou.edu
• Hänninen, University of Helsinki, Department of Ecology and Systematics, FIN-00014 Helsinki, Finland E-mail: hh@nn.fi