Category :
Article
article id 5623,
category
Article
Harry T. Valentine.
(1997).
Height growth, site index, and carbon metabolism.
Silva Fennica
vol.
31
no.
3
article id 5623.
https://doi.org/10.14214/sf.a8524
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A metabolic model of height growth and site index is derived from a parametrization of the annual carbon balance of a tree. The parametrization is based on pipe-model theory. Four principal variants of the height-growth model correspond to four combinations of assumptions regarding carbon allocation: (a) the apical shoot is autonomous or (b) it is not; and (A) the specific rate of elongation of a shoot equals that of a woody root or (B) it does not. The bB model is the most general as it includes the aA, bA, and aB models as special cases. If the physiological parameters are constant, then the aA model reduces to the form of the Mitscherlich model and the bA model to the form of a Bertalanffy model. Responses of height growth to year-to-year variation in atmospheric conditions are rendered through adjustments of a subset of the model's parameters, namely, the specific rate of production of carbon substrate and three specific rates of maintenance respiration. As an example, the effect of the increasing atmospheric concentration of CO2 on the time-course of tree height of loblolly pine (Pinus taeda) is projected over 50-year span from 1986. Site index is predicted to increase and, more importantly, the shape of the site-index curve is predicted to change.
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Valentine,
E-mail:
hv@mm.unknown
article id 5518,
category
Article
Heikki Hänninen,
Seppo Kellomäki,
Kaisa Laitinen,
Brita Pajari,
Tapani Repo.
(1993).
Effect of increased winter temperature on the onset of height growth of Scots pine: a field test of a phenological model.
Silva Fennica
vol.
27
no.
4
article id 5518.
https://doi.org/10.14214/sf.a15679
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According to a recently presented hypothesis, the predicted climatic warming will cause height growth onset of trees during mild spells in winter and heavy frost damage during subsequent periods of frost in northern conditions. The hypothesis was based on computer simulations involving a model employing air temperature as the only environmental factor influencing height growth onset. In the present study, the model was tested in the case of eastern Finnish Scots pine (Pinus sylvestris L.) saplings. Four experimental saplings growing on their natural site were surrounded by transparent chambers in autumn 1990. The air temperature in the chambers was raised during the winter to present an extremely warm winter under the predicted conditions of a double level of atmospheric carbon dioxide. The temperature treatment hastened height growth onset by two months as compared to the control saplings, but not as much as expected on the basis of the previous simulation study. This finding suggests that 1) the model used in the simulation study needs to be developed further, either by modifying the modelled effect of air temperature or by introducing other environmental factors, and 2) the predicted climatic warming will not increase the risk of frost damage in trees as much as suggested by the previous simulation study.
The PDF includes an abstract in Finnish.
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Hänninen,
E-mail:
hh@mm.unknown
-
Kellomäki,
E-mail:
sk@mm.unknown
-
Laitinen,
E-mail:
kl@mm.unknown
-
Pajari,
E-mail:
bp@mm.unknown
-
Repo,
E-mail:
tr@mm.unknown
article id 5469,
category
Article
Risto Heikkilä,
Kari Löyttyniemi.
(1992).
Growth response of young Scots pines to artificial shoot breaking simulating moose damage.
Silva Fennica
vol.
26
no.
1
article id 5469.
https://doi.org/10.14214/sf.a15627
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The main stem of young Scots pine (Pinus sylvestris L.) trees was cut off halfway along the current leading shoot and the two previous years’ leading shoots to simulate moose (Alces alces) damage. Trees of the same size were chosen as controls before treatments. The experiment was inspected ten years after artificial stem breakage. Removing the current leading shoot and the second shoot did not essentially affect the height and diameter growth of the trees. Removal down to the third shoot reduced the height as well as diameter growth. The average loss in growth was equivalent to less than one year’s growth. When the stem was cut off at the second or third shoot, stem crookedness and the presence of knots resulted in stem defects that will subsequently reduce the sawtimber quality. A high proportion of the stem defects will obviously still be visible at the first thinning cutting. Removing injured trees as pulpwood and pruning the remaining parts of cut stems evidently improves the quality of pine stand with moose damage.
The PDF includes an abstract in Finnish.
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Heikkilä,
E-mail:
rh@mm.unknown
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Löyttyniemi,
E-mail:
kl@mm.unknown
article id 5445,
category
Article
Taneli Kolström.
(1991).
Kuusen kylvö- ja istutuskoe viljavilla kivennäismailla Pohjois-Karjalassa.
Silva Fennica
vol.
25
no.
2
article id 5445.
https://doi.org/10.14214/sf.a15599
English title:
Results from the sowing and planting experiment of Norway spruce (Picea abies) on fertile sites in North Karelia, Finland.
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Different methods of sowing and planting of Norway spruce (Picea abies (L.) H. Karst.) were compared on fertile sites in North Karelia (62°20’N, 29°35’E, 85–120 m a.s.l.). The planting material were 4-year-old bare-rooted transplants, 2-year-old bare-rooted seedlings, and 2-year-old containerized seedlings raised in plastic greenhouse. The sowing methods were band sowing and shelter sowing. Ground vegetation was controlled during the first growing season mechanically or chemically, or the control was omitted totally.
Planting of spruce gave better results than sowing. After eight growing seasons there were sowed seedlings left in 30% of the sowing pots. The average height of them was 35 cm. Seedling survival was best with large bare-rooted transplants (91%). Survival of containerized seedlings was 79% and of small bare-rooted transplants 71%. The average height of large bare-rooted transplants was 131 cm, of containerized seedlings 86 cm and small bare-rooted seedlings 68 cm.
Sowing is not an advisable method for regeneration of spruce due to the small survival rate and slow initial development when ground vegetation is controlled only once. Also 2-year-old seedlings gave a satisfactory result in regeneration. Seedlings raised in greenhouse were more sensitive to frost damage than seedlings grown on open ground.
The PDF includes an abstract in English.
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Kolström,
E-mail:
tk@mm.unknown
article id 5254,
category
Article
Kristina Palmgren,
Anna Saarsalmi,
Assi Weber.
(1985).
Nitrogen fixation and biomass production in some alder clones.
Silva Fennica
vol.
19
no.
4
article id 5254.
https://doi.org/10.14214/sf.a15433
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In a greenhouse experiment that lasted for two years, nitrogenase activity, height growth and biomass production was compared in six clones of alder of which four were clones of Alnus incana and two A. incana x A. glutinos hybrids. In addition, the effect of a fertilizer nitrogen gradient was tested on one of the clones.
Clonal differences in height growth and nitrogenase activity were recorded at the end of the first growing season. The growth rhythm of some of the clones changed markedly during the second growing season but differences in nitrogenase activity between clones levelled out. Nitrogen fertilization suppressed nodulation during the first growing season, and also the following year the nitrogenase activity was significantly higher in alders grown without nitrogen supplement. Height growth and total biomass production was also depressed at rather low nitrogen levels.
The PDF includes a summary in Finnish.
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Palmgren,
E-mail:
kp@mm.unknown
-
Saarsalmi,
E-mail:
as@mm.unknown
-
Weber,
E-mail:
aw@mm.unknown
article id 5249,
category
Article
Jari Parviainen.
(1985).
Istuttamalla perustetun nuoren männikön, kuusikon, siperianlehtikuusikon ja rauduskoivikon kasvu.
Silva Fennica
vol.
19
no.
4
article id 5249.
https://doi.org/10.14214/sf.a15428
English title:
Growth of young Scots pine, Norway spruce, siberian larch and silver birch plantations.
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Early growth of four different tree species (Pinus sylvestris L., Picea abies (L.) H. Karst., Larix sibirica Ledeb and Betula pendula Roth) 16–23 years after planting were compared in a field experiment of 16 square plots established on a stony, grove-like upland (Oxalis-Myrtillus forest type) in Southern Finland. This study gives additional results to the publication Folia Forestalia 386/1979.
At this early stage, the growth of the spruce stand was clearly slower than that of the other species for all parameters to be measured (height, diameter, and volume growth). Height growth was most rapid in the silver birch stand and diameter growth in the larch stand. No clear differences were found in the mean volume of the 100 thickest trees in the stand between the larch and silver birch.
The PDF includes a summary in English.
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Parviainen,
E-mail:
jp@mm.unknown
article id 5198,
category
Article
Leo Heikurainen,
Jukka Laine,
Jarmo Lepola.
(1983).
Lannoitus- ja sarkaleveyskokeita karujen rämeiden uudistamisessa ja taimikoiden kasvatuksessa.
Silva Fennica
vol.
17
no.
4
article id 5198.
https://doi.org/10.14214/sf.a15181
English title:
Fertilization and ditch spacing experiments concerned with regeneration and growth of young Scots pine stands on nutrient poor pine bogs.
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The effects of variations in the intensity of drainage and NPK fertilization on the natural regeneration and planting results and the subsequent development of seedling stands under various climatic conditions on drained nutrient poor pine bogs was investigated in a 16-year-old study.
Comparison of height development of Scots pine (Pinus sylvestris L.) stands on drained peatlands to that of pine stands growing in mineral soil sites show that in Southern Finland the most efficient forest improvement measures (10 m ditch spacing and 1,000 kg/ha NPK-fertilization) resulted in growth that corresponds a to a height index of a stand in a Vaccinium type site. Less efficient treatment (30 m ditch spacing and no fertilizer) resulted in growth corresponding the development of young stand in a Calluna type site. In Northern Finland the effect of fertilization on height growth was almost negligible. This is possibly due to a decrease in the nitrogen mobilization from south to north of Finland. Thus, it seems evident that fertilization of young Scots pine stands on nutrient poor drained peatlands can be recommended only in the southern part of the country.
The effect of ditch spacing is same in the whole country. The narrower the spacing the better the height growth. In the south planted stands thrive better than naturally regenerated stands, but the situation is reversed in the north.
The PDF includes a summary in English.
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Heikurainen,
E-mail:
lh@mm.unknown
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Laine,
E-mail:
jl@mm.unknown
-
Lepola,
E-mail:
jl@mm.unknown
article id 5197,
category
Article
Kari Heliövaara,
Erkki Annila,
Eero Terho.
(1983).
Effect of nitrogen fertilization and insecticides on the population density of pine bark bug, Aradus cinnamomeus (Heteroptera, Aradidae).
Silva Fennica
vol.
17
no.
4
article id 5197.
https://doi.org/10.14214/sf.a15180
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The effect of nitrogen fertilization and two insecticides on the occurrence of the plant pine bark bug, Aradus cinnamomeus Panzer, was investigated in a young Scots pine (Pinus sylvestris L.) stand in Southern Finland. Three years after the treatment the bug density was lowest in the trees treated with lindane or dimethoate. However, in spite of the increasing height growth of the trees, they did not grow significantly faster than the control trees. Nitrogen fertilization increased both bug density and the height growth of the trees. Thus, the value of nitrogen fertilization against Aradus cinnamomeus remains obscure.
The PDF includes a summary in Finnish.
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Heliövaara,
E-mail:
kh@mm.unknown
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Annila,
E-mail:
ea@mm.unknown
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Terho,
E-mail:
et@mm.unknown
article id 5168,
category
Article
Kari Heliövaara.
(1982).
The pine bark bug, Aradus cinnamomeus (Heteroptera, Aradidae) and the height growth rate of young Scots pines.
Silva Fennica
vol.
16
no.
4
article id 5168.
https://doi.org/10.14214/sf.a15083
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Relationships between densities of pine bark bug, Aradus cinnamomeus Panzer and the height growth of young Scots pine (Pinus sylvestris L.) were studied in several habitats, including a highly infested area in Southern Finland. The slower the growth of the pines was, the greater was the height up to which bugs were found. On the average, maximum bug density was noted at a height corresponding to a fifth of the height of the tree. In stands restocked by natural generation, the greatest bug densities were noted in pines about three metres high and over twenty years old. Bug densities in trees whose height growth had been decelerating for five years were twice those in trees whose growth was accelerating. A significant negative correlation was found between the bug density and the last-year height increment.
The PDF includes a summary in Finnish.
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Heliövaara,
E-mail:
kh@mm.unknown
article id 5125,
category
Article
Ilppo Greis,
Seppo Kellomäki.
(1981).
Crown structure and stem growth of Norway spruce undergrowth under varying shading.
Silva Fennica
vol.
15
no.
3
article id 5125.
https://doi.org/10.14214/sf.a15066
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The crown structure and stem growth of Norway spruce (Picea abies (L.) H. Karst.) undergrowth was studied in relation to the prevailing light conditions and potential photosynthesis. Shading decreased the stem height growth more than the length increment of laterals, producing a plate-shaped crown in deep shade. Needles responded to shading by adopting a horizontal inclination in deep shade. The needles were wide and thin respectively in shade. In the open the needle cross-section was almost square. Stem radial growth and height growth were both affected by shading exhibiting a linear response to the prevailing light conditions and the potential photosynthesis. Light conditions under dominating trees were closely correlated with the basal area of the dominating trees.
The PDF includes a summary in Finnish.
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Greis,
E-mail:
ig@mm.unknown
-
Kellomäki,
E-mail:
sk@mm.unknown
article id 5123,
category
Article
English title:
Effect of foliar application of dilute sulphuric acid on Scots pine seedlings.
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Needle damages, transpiration, photosynthesis and needle and stem height growth of Scots pine (Pinus sylvestris L.) seedlings treated with dilute sulphuric acid were studied. The acidity of the solution was pH 3. Application of a dilute solution of sulphuric acid equivalent to the normal amount of precipitation occurring during the growing season damaged the surface of two-year-old needles but not that of the current-year needles. A reduction in the photosynthetic rate of 10–30% was observed compared with the untreated seedlings. Transpiration of the seedlings was not affected by the treatment. Needle growth and stem height growth of the seedlings growing on a substrate representing poor sandy soil were reduced. Increased needle growth and stem height growth were characteristic for the seedlings growing on substrate representing fertile moraine.
The PDF includes a summary in English.
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Katainen,
E-mail:
hk@mm.unknown
-
Kellomäki,
E-mail:
sk@mm.unknown
article id 5092,
category
Article
Seppo Kellomäki,
Markku Kanninen.
(1980).
Eco-physiological studies on young Scots pine stands. IV.
Silva Fennica
vol.
14
no.
4
article id 5092.
https://doi.org/10.14214/sf.a15033
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Crown and stem growth of young Scots pines (Pinus sylvestris L.) were studied in relation to photosynthate supply and light condition in a stand. The magnitude of needle and bud formation, and radial and height growth were to a great extent dependent on the photosynthate supply. However, in shaded conditions the growth of each characteristics was greater than expected on the basis of photosynthate supply. In the stem system this was especially apparent for height growth. Consequently, height growth was favoured at the expense of radial growth in shaded conditions. It also appeared that the basic density of wood was negatively related to both tree position and photosynthate supply.
The PDF includes a summary in Finnish.
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Kellomäki,
E-mail:
sk@mm.unknown
-
Kanninen,
E-mail:
mk@mm.unknown
article id 5069,
category
Article
Jouni Mikola.
(1980).
The effect of seed size and duration of growth on the height of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.) provenances and progenies at the nursery stage.
Silva Fennica
vol.
14
no.
1
article id 5069.
https://doi.org/10.14214/sf.a15010
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In this paper the connection between seed weight and amount and duration of growth are studied at the progeny level within stands or climatically uniform areas, and at the provenance level within larger geographic areas. The material consists of materials of several experiments in the nursery of Maisala in Southern Finland in 1971–76. The origins of the plant material used in the different experiments ranged from progenies of individual Scots pine (Pinus sylvestris L.) or Norway spruce (Picea abies (L.) H. Karst.) trees to provenance selections covering almost the whole natural range of these species.
The effect of seed weight on plant height is strongest immediately after germination and subsequently decreases steadily, when the genetic growth properties of the plants themselves become effective. The effect is usually visible at least until the end of the 1st growing season. This relationship varies considerably depending on the material studied. The connections between the duration and the amount of height growth also proved to differ according to the nature of genetic variation. In wide selection of provenances, which show clear genetic differentiation in annual growth rhythm. The variation in the duration of growth accounts for most of the differences in total height growth. At the individual and family level or between provenances of a limited area, there seems to be no clear connection between the duration and the amount of growth. It seems that the duration of the annual growing period is a genetic property, which is not affected by seed weight.
The total height alone in 1-year old test material grown in a greenhouse had hardly any value in the forecasting of growth capacity. The growth differences were caused mainly by the variation in seed size and growth rate differences during the growing period.
The PDF includes a summary in Finnish.
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Mikola,
E-mail:
jm@mm.unknown
article id 5021,
category
Article
Antti Koskimäki,
Pertti Hari,
Markku Kanninen,
Seppo Kellomäki.
(1979).
Inherent growth rythm of some Larix-species grown in a plastic greenhouse.
Silva Fennica
vol.
13
no.
1
article id 5021.
https://doi.org/10.14214/sf.a14875
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The daily height growth rate of several larch species and progenies (Larix decidua, Larix sibirica, Larix laricina, Larix leptolepis) grown in a plastic greenhouse and in the open was measured. The growth pattern indoors was completely different compared with the normal outdoor growth pattern. The onset of growth took place in the greenhouse much earlier than outdoors and the phase of increasing growth was much shorter, as was expected. However, the phase of maximum growth was unexpectedly long. This fact suggests that there is great potential for using greenhouse cultivation to change the growth pattern of cultivated plants in order to obtain more complete utilization of the potential growing season.
The PDF includes a summary in Finnish.
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Koskimäki,
E-mail:
ak@mm.unknown
-
Hari,
E-mail:
ph@mm.unknown
-
Kanninen,
E-mail:
mk@mm.unknown
-
Kellomäki,
E-mail:
sk@mm.unknown
article id 4924,
category
Article
Teklé Kapustinskaité.
(1975).
Puuston kasvu ja turpeen tuhkapitoisuus ojitetuilla soilla.
Silva Fennica
vol.
9
no.
3
article id 4924.
https://doi.org/10.14214/sf.a14766
English title:
Ash content of peatland soils and stand growth in connection with drainage.
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The ash content has been found to correlate with the fertility of peatlands. Relationship between height of 80-year-old stands and ash content of peat in topmost 30 cm layer was examined in Lithuanian conditions. On drained peatlands with ash content of peat from 3% to 8% pine stands increase in height. Ash content of peat being about 7% Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) stands on drained sites are found to be of equal height. Ash content of peat more than 8–9% has no significant effect on growth of pine or spruce stands. Birch (Betula verrucosa (B. Pendula Roth.) and Betula pubescens Erhrh.), stands are less sensitive to ash content of peat compared with other species. Black alder (Alnus glutinosa L. Gaertn.) stands occurred in sites with ash content of peat more than 8–10%. The height of the stands become equal both in drained and undrained sites in the cases where ash content of peat is about 16–18%. Ash (Fraxinus exelsior L.) stands attain high productivity on drained sites with ash content of peat about 20%.
The PDF includes a summary in English.
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Kapustinskaité,
E-mail:
tk@mm.unknown
article id 4908,
category
Article
Juhani Päivänen.
(1974).
Sarkaleveyden ja naveroinnin vaikutus pohjavesipinnan syvyyteen ja männyntaimiston kehitykseen lyhytkortisella nevalla.
Silva Fennica
vol.
8
no.
4
article id 4908.
https://doi.org/10.14214/sf.a14752
English title:
The effect of ditch spacing and furrowing on depth of ground water table and on development of a Scots pine plantation on small-sedge bog.
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The paper describes the results obtained from an investigation into the effect of ditch spacing, ditch depth and furrowing on ground water table and on development of a Scots pine (Pinus sylvestris L.) plantation on open small-sedge bog in Central Finland (60° 50’ N; 24° 20’ E), drained in 1967. The area was planted in 1968 with 2+1 Scots pine transplants, and fertilized with Y fertilizer for peat soils. The seedlings were measured in 1972.
The depth of the ground water table was greater, the narrower the ditch spacing. The water furrows shortened the duration of the high ground water and lowered the ground water table particularly in the case of ineffective drainage. The narrower the ditch spacing within the blocks, the higher were the young trees. On the other hand, the differences in the height of the trees between the ditch spacings were eliminated by the effect of the furrows.
The PDF includes a summary in English.
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Päivänen,
E-mail:
jp@mm.unknown
article id 4901,
category
Article
Veli Pohjonen.
(1974).
Istutustiheyden vaikutus eräiden lyhytkiertoviljelyn puulajien ensimmäisen vuoden satoon ja pituuskasvuun.
Silva Fennica
vol.
8
no.
2
article id 4901.
https://doi.org/10.14214/sf.a14745
English title:
Effect of spacing on the first-year yield and height increment of some species undergoing short rotation culture.
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The effect of spacing on the first-year yield and height increment of Alnus incana (L.) Moench, Populus tremula L. x Populus tremuloides Michx. (Populus x wettsteinii), Salix ’Aquatica Gigantea’, and Salix phylicifolia L. was studied at the Arctic Circle Agricultural Experimental Station in Northern Finland. S. ’Aquatica Gigantea’ gave yields which were twice as high as those of the other species in the study. The highest yields were of the order of 60 tons per hectare (fresh yield including foliage). The annual height growth in S. ’Aquatica Gigantea’ was about 100 cm, in the others about 30–50 cm. S. ’Aquatica Gigantea’ had a maximal height growth when the distance between the seedlings was 25 cm.
The PDF includes a summary in English.
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Pohjonen,
E-mail:
vp@mm.unknown
article id 4859,
category
Article
Eljas Pohtila.
(1972).
Istutuskuoppaan annetun kuparihienofosfaatin vaikutus männyn ja kuusen taimien elossapysymiseen ja pituuskasvuun eräällä kulotetulla ja auratulla uudistusalalla Koillis-Suomessa.
Silva Fennica
vol.
6
no.
1
article id 4859.
https://doi.org/10.14214/sf.a14662
English title:
Effect of fine-grounded copper rock phosphate placed in the planting hole on the survival and height growth of Scots pine and Norway spruce in a burnt and furrowed reforestation are in northeast Finland.
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The paper describes the results of a fertilization experiment, in which transplants of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) were fertilized with various doses of fine-ground copper rock phosphate (33% P2O5, 4% Cu) placed direct in the planting hole. The experiment was made in northeast Finland on a clear-cut, burnt-over and furrowed moraine heath. The fertilization increased especially the survival and condition of the Scots pines and increased to some extent also the height growth of the plants. The spruce survived better than the pines.
The PDF includes a summary in English
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Pohtila,
E-mail:
ep@mm.unknown
article id 4856,
category
Article
Pekka Rautiainen.
(1971).
Ympäristö- ja perintötekijöiden vaikutus männyn ilmiasuun Pohjois-Karjalan piirimetsälautakunnan siemenviljelyksessä Tohmajärvellä.
Silva Fennica
vol.
5
no.
4
article id 4856.
https://doi.org/10.14214/sf.a14658
English title:
The effect of environmental and genetical factors on the phenotype of Scots pine in a seed orchard in North Karelia.
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The study material included 600 Scots pine (Pinus sylvestris L.) grafts from the Tohmajärvi seed orchard in Eastern Finland. Their broad sense heritability for the height growth was 0.92, for the number of branches 0.87 and for the angle of branching 0.84. Grafts from Central Finland had cones more often than the southern ones, the frequencies being 26.3% and 11.2%. It seems that dominance plays a significant role in the genetical variation of this seed orchard and that height growth is probably more rewarding breeding characteristic than quality, the difference being small, however.
The PDF includes a summary in English.
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Rautiainen,
E-mail:
pr@mm.unknown
article id 4841,
category
Article
Hannu Mannerkoski.
(1971).
Lannoituksen vaikutus kylvösten ensi kehitykseen turvealustalla.
Silva Fennica
vol.
5
no.
2
article id 4841.
https://doi.org/10.14214/sf.a14643
English title:
Effect of fertilization on the initial development of Scots pine and Norway spruce plantations established by sowing on peat.
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Experiments were carried out to find out the effect of fertilizer application on germination, seedling emmergnece and initial development in conifer plantations established on peat by sowing, with a special reference to Scots pine (Pinus sylvestris L.). The experiments were carried out in 1968–70 in laboratory, in greenhouse and in the field.
In the greenhouse experiments with Y fertilizer for peat soils (14% N, 18% P2O5, 10% K2O) it was shown that germination and seedling emergence decreased markedly with increased fertilizer application. Mortality among seedlings that had emerged was the higher the larger quantities of fertilizer had been applied. The effect of fertilization was the greater, the drier the substrate. Fine ground rock phosphate (33% P2O5) promoted seedling emergence on a dry substrate but not on a wet one.
The field experiments carried out in Central Finland included dry and wet sites. Y fertilizer, Oulu Saltpeter (25% N), fine-ground rock phosphate and potassium salt (50% K2O) were used. According to the results, easily soluble fertilizers decreased seedling emergence. On wet sites the effect of Y fertilizer was weaker than on drier sites. Fine-ground rock phosphate slightly increased the number of seedlings emerging. Height growth was increased during the first three growing seasons only by those fertilizers containing phosphorus.
The PDF includes a summary in English.
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Mannerkoski,
E-mail:
hm@mm.unknown
article id 4838,
category
Article
Kustaa Seppälä.
(1971).
Metsityslannoituksessa käytetyn lannoitemäärän ja levitystavan merkitys istutustaimiston alkukehitykselle ojitetuilla avosoilla.
Silva Fennica
vol.
5
no.
2
article id 4838.
https://doi.org/10.14214/sf.a14640
English title:
The quantity of fertilizer and application methods used in afforestation of open bogs.
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The paper describes the results obtained from an experiment of fertilization of drained treeless peatlands in connection of planting in three sites in Central Finland. Scots pine (Pinus sylvestris L.) seedlings 2+0 was used. The fertilizer (Y-fertilizer for peat soils, 14% N, 18% P2O5, 10% K2O) was applied in rates of 0, 20, 40 and 80 g/transplant. The fertilizer was strewn either around the plant within a circular patch of 20 cm in diameter, in a ring with a radius of 10 cm and in a ring with a radius of 20 cm. The seedlings were measured two and five years after planting.
The greater the quantity of fertilizer applied and the closer it was applied to the plant the higher was the mortality of transplants. Fertilization increased the mortality during the first two growing seasons after application. Later, however, the mortality decreased to a similar level irrespective the way the fertilizer was applied. In the beginning of the second growing season the fertilized plants showed considerably better height growth than the control plants. The smallest quantity of fertilizer applied produced almost full increase in growth. The pattern of application of the fertilizer had little effect on the growth.
It was concluded that a use of small amounts of fertilizer can be recommended in connection with planting and that it should not be applied very near the seedlings.
The PDF includes a summary in English.
-
Seppälä,
E-mail:
ks@mm.unknown
article id 4835,
category
Article
Kari Löyttyniemi.
(1971).
Havupunkin, Oligonychus ununguis (Jacobi), aiheuttaman neulasvioituksen vaikutuksesta kuusen taimien kasvuun.
Silva Fennica
vol.
5
no.
1
article id 4835.
https://doi.org/10.14214/sf.a14624
English title:
Influence of damage caused to needles of Norway spruce by spruce spider mite, Oligonychus ununguis, on seedling growth.
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The study was carried out in order to establish the possible influence of damage caused to the needles of Picea abies (L.) H. Karst. by the spruce spider mite, Oligonychus ununguis (Jacobi), and the growth of the damaged seedlings. The study was carried out in 1968–1970 by comparing growth of seedlings infected with spruce spider mite with that of seedlings where mites had been killed with acaricide (Eradex®). In the seedlings that had not been treated with acaricide, the number of wintering eggs were 60, 20 and 5 per shoot in the various years of the study. When the experiment was laid out, before planting and acaricide treatment, the seedlings were four years old, all seadlings were heavily infected, the number of wintering eggs being 100 per shoot. The growth of infected seedlings was 3, 20 and 15% smaller than that obtained for the seedlings which had been treated with acaricide.
The PDF includes a summary in English.
-
Löyttyniemi,
E-mail:
kl@mm.unknown
article id 4823,
category
Article
Christel Palmberg.
(1970).
Heritabiliteetin arvioiminen eräässä männyn (Pinus silvestris L.) jälkeläiskokeessa.
Silva Fennica
vol.
4
no.
3
article id 4823.
https://doi.org/10.14214/sf.a14612
English title:
Estimation of heritability in open-pollinated plus tree progenies of Pinus sylvestris L.
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Field experiments of Scots pine (Pinus sylvestris L.) was established by planting seedlings grown from seeds collected from open-pollinated plus trees throughout the country. The 36 progenies represented were planted in 4 blocks as 2+2 transplants in 1960. The main characteristics of the seedlings were measured in 1966 and 1968. Considerable damage had been caused to the stands by moose (Alces alces) and Melampsora pinitorqua Rostr., consequently, therefore, only normally developed seedlings were measured.
Highly significant differences between progenies were found in the number of branches in 1968 and in the ratio of height of tree to the length of the longest branch. In 1968, the differences in height between progenies were not significant, but there were significant differences between blocks both in tree height and length of terminal shoot. Obviously, the edaphic heterogeneity of the site has influenced mainly the juvenile growth of the plants, because in the length of the terminal shoot there could be seen also significant differences between the progenies. There were no significant differences between the progenies in the length of the longest branch, in the angles of the thickest branches, in stem taper and in the diameter of the thickest branch.
The PDF includes a summary in English.
-
Palmberg,
E-mail:
cp@mm.unknown
article id 4817,
category
Article
English title:
Height growth of seedling stands growing on peatland.
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This paper presents the results of a contest performed on behalf of the Finnish bank Kansallis-Osake-Pankki and the Central Forestry Board Tapio on growing trees on peatlands. Over 5,000 sample plots were established on drained peatlands in various parts of Finland. The aim was to achieve a best possible growth of seedling stands on peatland. The factors influencing the growth of 85 best Scots pine (Pinus sylvestris L.) and 60 best Norway spruce (Picea abies (L.) H.Karst.) sample plots were studied.
The height growth of the seedling stands decreased towards the north. Fertilization seemed not to decrease the regional differences; rather on the contrary. On the other hand, fertilization increased height growth, but evidently so that the increase obtained was greater in the southern than in the northern parts of the country. Light fertilization (50 kg/ha of K2O and 60 kg/ha of O2P5) caused a clear increase in height growth while heavy fertilization (100 g/ha of K2O and 120 kg/ha of O2P5), had same effect but to much greater extent than the former. Spruce seedling stands in particular benefitted of the heavy fertilization.
Fertilization did not eliminate the original differences in the quality of the sites in question, but these could still be seen in the height growth after fertilization. The effect of drain spacing on the height growth was not very clear. In dense seedling stands (800 seedlings/ha) the height growth of the dominant seedlings was greater than that obtained in stands of lower density. Hold-overs caused a decrease in the growth of the seedling stands.
The PDF includes a summary in English.
-
Heikurainen,
E-mail:
lh@mm.unknown
-
Ouni,
E-mail:
jo@mm.unknown
article id 4771,
category
Article
J. G. Gordon,
G. E. Gatherum.
(1968).
Photosynthesis and growth of selected Scots pine populations.
Silva Fennica
vol.
2
no.
3
article id 4771.
https://doi.org/10.14214/sf.a14556
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Eight Scots pine (Pinus sylvestris L.) seed sources, ranging from 42° to 66° north latitude, were grown under a constant, 16-hour photoperiod in a greenhouse for approximately 6 months. Rates of photosynthesis, as measured by an IRGA, and growth, as measured by increase in height and fresh and dry weight, differed among seed sources at the end of the six-month growing period. Photosynthetic capacity and growth were strongly related to latitude of seed source, and were greatest in the seed sources coming from a parent environment in which maximum photoperiods are about 16 hours.
Photosynthetic efficiency (rate of photosynthesis per gram needle weight) was also strongly related to latitude of seed source, but was lowest in the seedlings which exhibited the greatest growth and photosynthetic capacity. This may have been due to (1) more mutual shading of needles on the larger seedlings and (2) a lesser proportion of juvenile needles on the larger seedlings or (3) biochemical differences in the use of photosynthate in the needles. Seed source and light intensity had an interacting effect on rates of photosynthesis only in seedlings of the two northernmost seed sources.
The PDF includes a summary in Finnish.
-
Gordon,
E-mail:
jg@mm.unknown
-
Gatherum,
E-mail:
gg@mm.unknown
article id 4576,
category
Article
Bo-Eric Blumenthal.
(1942).
Distribution and properties of aspen in Finland.
Silva Fennica
no.
56
article id 4576.
https://doi.org/10.14214/sf.a9080
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Aspen (Populus tremula L.) is a common tree in Finland, and has been used, for instance, in matchstick industry. However, there has been little studies on its distribution and properties. In this study, 142 sample trees in different forest site types in Valtimo and Onkamo in Eastern Finland were measured in detail in 1935.
According to the results, during the first 10 years aspens height growth is fastest of the Finnish tree species surpassing, for instance, Scots pine (Pinus sylvestris L.) and birch (Betula sp.) . The diameter growth is similar to Scots pine up to the age of 50 years, after which the growth of aspen exceeds Scots pine. Branchless portion of the stem compared to the height of the tree increases until it reaches about 50% of the height of the tree. In poorer sites aspen is prone to decay.
Aspen regenerates easily both by root shoots and seeds. If root shoots are left to grow, the mother tree should be free of decay. In general, seedlings are of better quality. Good quality aspen stands require thinning and a rich forest type. If an old aspen stand has decay, the trees should be ring-barked and the site regenerated with a new tree species.
The article includes an abstract in German.
-
Blumenthal,
E-mail:
bb@mm.unknown
Category :
Article
article id 7231,
category
Article
Martti Hertz.
(1929).
Huomioita männyn ja kuusen pituuskehityksen "vuotuisesta" ja vuorokautisesta jaksosta.
Acta Forestalia Fennica
vol.
34
no.
18
article id 7231.
https://doi.org/10.14214/aff.7231
English title:
Observations on annual and daily cycles in the height growth of Scots pine and Norway spruce.
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The height growth of Scots pine (Pinus sylvestris L.) seedlings were observed in Korkeakoski and Evo in Southern Finland in 1925-1928. The growth was slow in the beginning of the growing season, increased after that to decrease again towards the end of the growing season. The height growth begun in May, reached the fastest growth rates in June, and ended in June-July. According to the earlier studies, the length of the height growth of Scots pine is dependent on the temperature of the previous summer. This study showed that warm temperatures of the same summer promote height growth, and low temperatures slow it down. Also the daily growth fluctuates, being highest during the afternoon and slowest during the early morning. The daily growth is dependent on temperature.
Norway spruce (Picea abies (L.) H. Karst.) begin the height growth in average 9 days later than Scots pine. Compared to pine, the speed of growth in spruce decreases slower towards the late summer.
The volume 34 of Acta Forestalia Fennica is a jubileum publication of professor Aimo Kaarlo Cajander. The PDF includes a summary in German.
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Hertz,
E-mail:
mh@mm.unknown
article id 7094,
category
Article
S. E. Multamäki.
(1923).
Tutkimuksia ojitettujen turvemaiden metsänkasvusta.
Acta Forestalia Fennica
vol.
27
no.
1
article id 7094.
https://doi.org/10.14214/aff.7094
English title:
Studies on the growth of drained peatlands in Finland.
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Only about 24,000 hectares of peatlands have been drained in the state lands by the 1921. The aim of this study was to define how much the growth of the trees in the drained peatland revives. Sample plots were measured in previously drained peatlands that had sufficient Scots pine (Pinus sylvestris L.) tree stand. A stem analysis was performed to one of the sample trees. The evenness of the stands was dependent on how evenly the peatlands had dried when the stand was regenerated. Thus, the sample stands were not always fully stocked. However, they had capacity to develop towards evenly structured forests as the peatlands continued to dry further. The diameter and height growth of the dried peatlands have corresponded the similar stands in mineral soil sites. In trees that have grown stunted in the peatlands, the diameter growth seems to increase faster than the height growth. The volume growth is slightly smaller than in the similar mineral soil sites due to less favorable stem form. After the draining, the roots of the trees continued to grow from the old branches of root, but start then to form new roots. When the ground water level drops, the root layer grows deeper.
The PDF includes a summary in German.
-
Multamäki,
E-mail:
sm@mm.unknown
article id 7049,
category
Article
Erkki Laitakari.
(1920).
Tutkimuksia sääsuhteiden vaikutuksesta männyn pituus- ja paksuuskasvuun.
Acta Forestalia Fennica
vol.
17
no.
1
article id 7049.
https://doi.org/10.14214/aff.7049
English title:
Studies on the effect of weather conditions on diameter and height growth of Scots pine.
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The height and diameter growth measured from different Scots pine (Pinus sylvestris L.) stands in Southern Finland was compared with meteorological information. The height growth benefits from warm weather in late summer in the previous year, and especially from high temperatures in June. Precipitation or the temperature in the same year did not affect the height growth. Diameter growth benefits from high temperatures in the spring of the same year, especially in April. High precipitation in the spring has in some cases negative impact on the diameter growth. The different combinations of precipitation and temperature can have variable effects on tree growth. In the diameter growth was seen a periodicity that coincides with sunspots.
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Laitakari,
E-mail:
el@mm.unknown
article id 7014,
category
Article
Raf. Björkenheim.
(1919).
Writings on knowledge on forest types in spruce forests of German “Mittelgebirge” (mountainous areas).
Acta Forestalia Fennica
vol.
6
no.
3
article id 7014.
https://doi.org/10.14214/aff.7014
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The data has been collected in spruce forests in mountainous areas of Germany: Fichtelgebirge and Böhmerwald in Bavaria, Erzgebirge in Saxonia. The studied characteristics of the stand were: growth of the trees in height and diameter, and the ground vegetation. The stands were classified according Cajander’s forest site classification. The article presents the most common plants and other characteristics of every forest site type and studied stands. The relation of the height of the trees and their age is represented in diagrams for every forest type.
The presence of indicator plants is somewhat dependent on the stand age and crown coverage. The amount of species is lowest when the crown coverage is at the greatest.
As conclusion of the study it can be seen that the growth of the stand differentiates clearly depending on the forest site type, being greater at the more nutritious sites. Since the differently growing stands need different management, it would be natural to direct the management of the stand according the forest site type.
-
Björkenheim,
E-mail:
rb@mm.unknown
article id 7012,
category
Article
Yrjö Ilvessalo.
(1916).
Mäntymetsikköjen valtapuitten kasvusta mustikka- ja kanervatyyppien kankailla Salmin kruununpuistossa.
Acta Forestalia Fennica
vol.
6
no.
1
article id 7012.
https://doi.org/10.14214/aff.7012
English title:
The growth of the dominant trees in Scots pine forests of Myrtillus and Calluna typesailla Salmin kruununpuistossa.
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Tree growth is one of the factors that have been used to determine the site quality. The aim of the study was to show that growth of single trees growing on a same forest site class are similar, but differ from trees growing on a different site type. To compare the tree growth, a stem analysis was performed to dominant trees in Scots pine (Pinus sylvestris L.) stands, measured in 15 Myrtillus type sample plots and in 15 Calluna type sample plots in state forests in Salmi, situated in north side of Lake Ladoga. The height growth when the tree was young was higher in the trees growing in the Myrtillus type than in the Calluna type. Also, the trees of same age are higher in Myrtillus type stand than in the Calluna type. In Calluna type, the height growth, however, evens out later in age than in the Myrtillus type. The volume growth of the trees begins to increase earlier in Myrtillus type, and is higher than in Calluna type. Similarly, the diameter growth in breast height is higher in the Myrtillus type.
The PDF includes a summary in German.
-
Ilvessalo,
E-mail:
yi@mm.unknown
article id 7611,
category
Article
Kustaa Seppälä.
(1969).
Kuusen ja männyn kasvun kehitys ojitetuilla turvemailla.
Acta Forestalia Fennica
no.
93
article id 7611.
https://doi.org/10.14214/aff.7611
English title:
Post-drainage growth rate of Norway spruce and Scots pine on peat.
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The present study is an attempt to establish the response to drainage of Norway spruce (Picea abies (L.) H. Karst.) and Scots pine (Pinus sylvestris L.) on some peatland sites, and to determine the revival of the trees and continuance of the growth after drainage. Growth of trees in four types of peatland types of drained peatlands drained between 1908-1918 were studied, and the results were compared with corresponding mineral soil sites
In pine the response to drainage was faster than in spruce in all age classes. Even the oldest groups of trees showed as good growth as trees of the same size growing on mineral soils. The rapidity of revival and the radial growth maximum are affected by the age of the tree at the time of ditching and the site fertility. The size of the trees, too, is of importance for the magnitude of post-drainage radial growth; the influence is similar in different sites. The basal area growth of trees growing on peat usually showed an unbroken increase during the entire post-drainage period. Neither the height growth indicates a decline in growth over time.
In the light of the results from sample tree analysis, it seems that tree growth gradually rises even after the revival period in peatlands originally covered by forest. The are some errors in the comparisons made, but it can be observed that aging of drainage areas as such does not mean that growth conditions become poorer.
The PDF includes a summary in English.
-
Seppälä,
E-mail:
ks@mm.unknown
Category :
Research article
article id 1680,
category
Research article
Highlights:
High water potential and carbon gain during bud forming favoured height growth; High water potential during the elongation period favoured height growth; A spring with high carbon gain favoured diameter growth; The obtained regression models had generally low generalization performance.
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Despite the numerous studies on year-to-year variation of tree growth, the physiological mechanisms controlling annual variation in growth are still not understood in detail. We studied the applicability of data-driven approach i.e. different regression models in analysing high-dimensional data set including continuous and comprehensive measurements over meteorology, ecosystem-scale water and carbon fluxes and the annual variation in the growth of app. 50-year-old Scots pine stand in southern Finland. Even though our dataset covered only 16 years, it is the most extensive collection of interactions between a Scots pine ecosystem and atmosphere. The analysis revealed that height growth was favoured by high water potential of the tree and carbon gain during the bud forming period and high water potential during the elongation period. Diameter growth seemed to be favoured by a winter with high precipitation and deep snow cover and a spring with high carbon gain. The obtained models had low generalization performance and they would require more evaluation and iterative validation to achieve credibility perhaps as a mixture of data-driven and first principle modeling approaches.
-
Kulmala,
University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland
E-mail:
liisa.kulmala@helsinki.fi
-
Žliobaitė,
Aalto University, Department of Computer Science and Helsinki Institute for Information Technology, P.O. Box 11000, FI-00076 Aalto, Finland; University of Helsinki, Department of Geosciences and Geography, P.O. Box 64, FI-00014 University of Helsinki, Finland
E-mail:
zliobaite@gmail.com
-
Nikinmaa,
University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland
E-mail:
eero.nikinmaa@helsinki.fi
-
Nöjd,
Natural Resources Institute Finland (Luke), Bio-based business and industry, Tietotie 2, FI-02150 Espoo, Finland
E-mail:
pekka.nojd@luke.fi
-
Kolari,
University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland; University of Helsinki, Department of Physics, P.O. Box 64, FI-00014 University of Helsinki, Finland
E-mail:
pasi.kolari@helsinki.fi
-
Kabiri Koupaei,
University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland
E-mail:
kourosh.kabiri@helsinki.fi
-
Hollmén,
Aalto University, Department of Computer Science and Helsinki Institute for Information Technology, P.O. Box 11000, FI-00076 Aalto, Finland; University of Helsinki, Department of Geosciences and Geography, P.O. Box 64, FI-00014 University of Helsinki, Finland
E-mail:
jaakko.hollmen@aalto.fi
-
Mäkinen,
Natural Resources Institute Finland (Luke), Bio-based business and industry, Tietotie 2, FI-02150 Espoo, Finland
E-mail:
harri.makinen@luke.fi
article id 1262,
category
Research article
Āris Jansons,
Roberts Matisons,
Māra Zadiņa,
Linards Sisenis,
Jānis Jansons.
(2015).
The effect of climatic factors on height increment of Scots pine in sites differing by continentality in Latvia.
Silva Fennica
vol.
49
no.
3
article id 1262.
https://doi.org/10.14214/sf.1262
Highlights:
Height increment-climate relationships of Scots pine were assessed using dendrochronological techniques; Annual height increment was significantly affected by climatic factors; In western Latvia, temperature in preceding summer mainly affected height increment; In eastern Latvia height increment was affected by previous autumn temperature; During the 20th century, the effect of climatic factors has altered likely dues to climate change.
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Height growth of trees is a crucial parameter that influences the composition and productivity of forest stands and quality of timber; however, the relationships between annual height increment (HI) and climatic factors have been poorly studied. In this study, the effect of monthly mean temperature and precipitation sums on the HI of Scots pine in two sites in Latvia have been determined using dendrochronological techniques. Correlation and response function analyses were conducted for entire chronologies of HI and for 50-year intervals within them. Climatic factors significantly affected the HI of Scots pine; however, not only did the suite of significant factors differ between the sites, but the influence of these factors changed during the 20th century. In the site in western Latvia where climate is milder, temperature in the preceding summer was the main climatic determinant of HI. The effect of temperature in the dormant period and spring was significant during the first part of the 20th century, while the effect of temperature in the previous September and November has become significant since the second half of the 20th century. In the site in eastern Latvia where summers are hotter, HI has been affected by both temperature and water deficit related factors in the summer. However, since the later part of the 20th century, the effect of temperature in the previous October has intensified and become the main climatic determinant of HI.
-
Jansons,
LSFRI “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169
E-mail:
aris.jansons@silava.lv
-
Matisons,
LSFRI “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169
E-mail:
robism@inbox.lv
-
Zadiņa,
LSFRI “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169
E-mail:
mara.zadina@silava.lv
-
Sisenis,
LUA Forestry Faculty, Akadēmijas str. 11, Jelgava, Latvia
E-mail:
linards.sisenis@llu.lv
-
Jansons,
Forest Competence Centre, Dzērbenes str. 27, Riga, Latvia, LV1006
E-mail:
janis.jansons@silava.lv
article id 1300,
category
Research article
Jaana Luoranen,
Risto Rikala.
(2015).
Post-planting effects of early-season short-day treatment and summer planting on Norway spruce seedlings.
Silva Fennica
vol.
49
no.
1
article id 1300.
https://doi.org/10.14214/sf.1300
Highlights:
Summer planting and short-day treatment advanced the bud burst and increased the height of Norway spruce seedlings after planting, compared to autumn and spring planted or untreated seedlings.
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Effects of short-day (SD) treatment on bud burst, growth and survival of Norway spruce (Picea abies [L.] Karst.) container seedlings after summer planting were studied in an experiment established in Suonenjoki, Central Finland. One-year-old seedlings were SD-treated for three weeks starting on 18 June, 24 June and 8 July 2004 and then planted on 22 July, 5 August, 6 September 2004 and, as a normal spring planting, on 10 May, 2005. Untreated control seedlings were also planted on these dates. Second flush on the planting year and bud burst the following spring was monitored in planted seedlings, whereas seedling height and survival were determined at the end of growing seasons 2004–2006. We observed a non-significant risk of a second flush if seedlings were SD-treated on 18 June. Also, SD-treated seedlings planted in July or August showed advanced bud burst and increased height the following growing season without significant effects on survival, compared to autumn and spring planted seedlings. Planting in July or early August was associated with a significant increase in the incidence of multiple leaders in later years. Based on our results, to begin a three-week SD treatment in late June or early July and then plant seedlings in late July or early August could be a good practice.
-
Luoranen,
Natural Resources Institute Finland (Luke), Natural resources and bioproduction, Juntintie 154, FI-77600 Suonenjoki, Finland
E-mail:
jaana.luoranen@luke.fi
-
Rikala,
The Finnish Forest Research Institute, Suonenjoki Unit, Suonenjoki, Finland
E-mail:
rikala@dnainternet.fi
article id 1192,
category
Research article
Hannu Hökkä,
Hanna Mäkelä.
(2015).
Post-harvest height growth of Norway spruce seedlings in northern Finland peatland forest canopy gaps and comparison to partial and complete canopy removals and plantations.
Silva Fennica
vol.
48
no.
5
article id 1192.
https://doi.org/10.14214/sf.1192
Highlights:
Norway spruce seedlings’ height growth recovered within four years after the cutting of canopy gaps; Growth was linearly related to tree height, being highest for tallest seedlings; Seedlings in the 20 m diameter gap and in the central and northern parts in the 15 m diameter gap showed the best growth; In gaps early height growth was 60% of that in peatland spruce plantations but 2–3 times higher than in uneven-aged cut forests.
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Recent studies have shown the establishment of Norway spruce (Picea abies (L.) Karst.) to be successful in small canopy gaps cut in drained spruce mire stands in northern Finland. The aim of this study was to quantify seedling height growth in gaps and compare it to that observed in other canopy cuttings and plantations. We sampled spruce crop seedlings (maximum density ca. 3000 ha–1) in the spring of 2013 in a field experiment in which canopy gaps of 10, 15 and 20 m in diameter had been cut in winter 2004. The total seedling height in 2013 and the length of annual shoots over the past five years (2012–2008) were recorded in the survey. Seedling height varied from 20 cm to 2.7 m, with an average of 65 cm. The average annual height growth was 7.1 cm. A mixed linear model analysis was carried out to investigate seedling height growth variation. Seedling height was linearly and positively related to growth. Height growth started to increase in the fifth growing season after cutting. Seedling height growth in the 20 m gap was slightly better than in the smaller ones. In the 15 m gap, both the centrally located seedlings and those located at the northern edge grew best. In the 20 m gap, southerly located seedlings grew more slowly than seedlings in all other locations. The average seedling height growth in this study was about 60% of that in peatland plantations, but comparable to that in mineral soil gaps, and 2–3 times higher than in uneven-age cut stands.
-
Hökkä,
Finnish Forest Research Institute, Rovaniemi Unit, Eteläranta 55, FI-96300 Rovaniemi, Finland
E-mail:
hannu.hokka@metla.fi
-
Mäkelä,
Lapland University of Applied Sciences, Jokiväylä 11 C, FI-96300 Rovaniemi, Finland
E-mail:
hanna.makela@edu.ramk.fi
article id 1124,
category
Research article
Āris Jansons,
Mārtiņš Zeps,
Juris Rieksts-Riekstiņš,
Roberts Matisons,
Oskars Krišāns.
(2014).
Height increment of hybrid aspen Populus tremuloides x P. tremula as a function of weather conditions in central part of Latvia.
Silva Fennica
vol.
48
no.
5
article id 1124.
https://doi.org/10.14214/sf.1124
Highlights:
Intra-annual height growth of hybrid aspen was monitored; Clones with early leaf flushing dates showed faster height growth; Height growth was generally controlled by temperature; Fast-growing hybrids were more robust to weather conditions than slow-growing ones; Potential evapotranspiration (moisture regime) negatively affected height growth of clones with delayed phenology.
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Height growth of young hybrid aspen (Populus tremula L. × P. tremuloides Michx.) was studied in relation to weather conditions. Height of clones with different leaf flushing phenology (early, intermediate and late) was monitored during the growing periods of 2010 and 2011 in a plantation established on former agricultural land. Mean daily height increment (HI) was calculated. Multiple linear regression was used to determine which weather factors (variables) had significant effect on HI. Mean seasonal height growth (mean seasonal HI) between clones (groups) was compared by ANOVA. In both years, HI was significantly higher for clones with early and intermediate leaf flushing compared to clones with late leaf flushing. The effect of weather factors also differed between clones according to their leaf flushing phenology; it was the weakest for HI of clones with early leaf flushing compared to clones with intermediate and late leaf flushing. Mean temperature was the main factor, which positively affected HI of all clones, suggesting that warmer climate might be beneficial for height growth of young hybrid aspen in Latvia. Nevertheless, significant negative relationship between HI and potential evapotranspiration (PET) was observed for clones with delayed leaf flushing, suggesting negative effect of increasing variability of precipitation on growth. Thus, the differences in height growth intensity might be related to growth sensitivity to weather conditions. On the other hand, such differences in height growth between clones might be caused by competition (i.e. with herbs), as trees with early leaf flushing might conquer more resources and become more robust against the environmental fluctuation.
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Jansons,
LSFRI „SILAVA”, Rigas Str. 111, Salaspils, Latvia, LV2169
E-mail:
aris.jansons@silava.lv
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Zeps,
LSFRI „SILAVA”, Rigas Str. 111, Salaspils, Latvia, LV2169
E-mail:
martins.zeps@silava.lv
-
Rieksts-Riekstiņš,
LSFRI „SILAVA”, Rigas Str. 111, Salaspils, Latvia, LV2169
E-mail:
Juris.Riekstins@silava.lv
-
Matisons,
LSFRI „SILAVA”, Rigas Str. 111, Salaspils, Latvia, LV2169
E-mail:
robism@inbox.lv
-
Krišāns,
LSFRI „SILAVA”, Rigas Str. 111, Salaspils, Latvia, LV2169
E-mail:
oskars.krisans@silava.lv
article id 107,
category
Research article
Jaana Luoranen,
Risto Rikala,
Heikki Smolander.
(2011).
Machine planting of Norway spruce by Bracke and Ecoplanter: an evaluation of soil preparation, planting method and seedling performance.
Silva Fennica
vol.
45
no.
3
article id 107.
https://doi.org/10.14214/sf.107
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We evaluated the effects of planting date and planting machine (Bracke: three machines, 69 regeneration areas in three years; Ecoplanter: six areas, two years) on the quality and field performance one and three years after planting of Norway spruce (Picea abies (L.) Karst.) seedlings in central Finland. Both machine types planted on average 1800 seedlings per hectare, and after three years approximately 1600 (Bracke) and 1200 (Ecoplanter) were still alive. This study suggests that planting with a Bracke machine can achieve better regeneration rates than those observed in privately-owned Finnish forests. We characterized the quality of mounding and planting with the Bracke machine as excellent and that of the Ecoplanter as good. The soil preparation method of the Ecoplanter produced humus-rich mounds where seedlings were susceptible to pine weevils and consequently suffered higher mortality. Different machines were used in different regional areas and each machine was operated by different driver/s which may have influenced the results. No negative effects of planting date were observed. Seedling growth decreased if they were tall in relation to their root plug volume, grown too densely in the nursery, and if stored in the field for several months prior to planting. We conclude that mechanized planting is successful when the soil preparation method produces mounds covered by purely mineral soil. Planting from May to the end of September is suitable for seedlings intended for use during this period.
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Luoranen,
Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland
E-mail:
jaana.luoranen@metla.fi
-
Rikala,
Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland
E-mail:
rr@nn.fi
-
Smolander,
Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland
E-mail:
hs@nn.fi
article id 120,
category
Research article
Antti J. Lukkarinen,
Seppo Ruotsalainen,
Teijo Nikkanen,
Heli Peltola.
(2010).
Survival, height growth and damages of Siberian (Larix sibirica Ledeb.) and Dahurian (Larix gmelinii Rupr.) larch provenances in field trials located in southern and northern Finland.
Silva Fennica
vol.
44
no.
5
article id 120.
https://doi.org/10.14214/sf.120
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The aim of this study was to analyse differences in the survival and height growth of, as well as damages to Siberian (Larix sibirica Ledeb.) and Dahurian (Larix gmelinii Rupr.) larch provenances over four growing seasons in field trials established in 2006 in southern (Punkaharju) and northern Finland (Kivalo). In this context, the study also investigated if the geographical and climatic conditions of the origin of the provenance could explain the differences between the provenances. The study material consisted of 20 Russian Siberian and Dahurian larch provenances and five seed sources from Finland (4) and Russia (1) as comparison lots. It was found that the Finnish seed sources of Siberian larch survived well in both the Kivalo and Punkaharju trials. Five northern latitude Russian provenances, of which one was Dahurian and the remainder were Siberian larches, had the highest survival in Kivalo. However, the differences observed in survival between provenances were only significant (p < 0.05) in Kivalo. Regardless of the trial, the differences, however, in height growth were significant and large between provenances. The southern Dahurian larches had a superior height growth in Punkaharju. The northern Dahurian larch provenance from Magadan (59°50′N, 150°40′E) had the largest height growth in Kivalo, among some northern Siberian larches. Damages were diverse, though Dahurian larches had less mammal damage than the Siberian larches. In general, the differences between provenances were not significant. Latitude and altitude best explained the differences between provenances, but also mean temperature, temperature sum and continentality index affected them (p < 0.05).
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Lukkarinen,
University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland
E-mail:
antti.lukkarinen@pp.inet.fi
-
Ruotsalainen,
Finnish Forest Research Institute, FI-58450 Punkaharju, Finland
E-mail:
sn@nn.fi
-
Nikkanen,
Finnish Forest Research Institute, FI-58450 Punkaharju, Finland
E-mail:
tn@nn.fi
-
Peltola,
University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland
E-mail:
hp@nn.fi
article id 209,
category
Research article
Jaana Luoranen,
Kyösti Konttinen,
Risto Rikala.
(2009).
Frost hardening and risk of a second flush in Norway spruce seedlings after an early-season short-day treatment.
Silva Fennica
vol.
43
no.
2
article id 209.
https://doi.org/10.14214/sf.209
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There have been years in Finland when container seedlings of Norway spruce (Picea abies (L.) Karsten) planted in the summer have been damaged by early-autumn frosts. For August and September plantings, the seedlings can be hardened by means of short-day (SD) treatment, but little information is available about its usability for earlier plantings. We studied the effects of early-season SD treatment on the frost hardiness and risk of a second flush of Norway spruce seedlings. In three successive years, second-year seedlings were grown in a greenhouse or outdoors in the spring and early summer and then subjected to two or three-week SD treatment beginning on the second, third, or fourth week of June. We monitored the height growth cessation, bud formation, and frost hardiness of the seedlings in the nursery. All SD treatments made the height growth cease, but the risk of a second flush increased if the temperature sum was less than 300 d.d. before the beginning of the SD treatment or more than 450 d.d. between the end of the treatment and mid-August. Clearly, then, SD treatment reduced the risk of a second flush in seedlings that had been grown in a greenhouse in the spring. Early-season SD treatment increased the frost hardiness of both needles and stems for late July to early September in comparison with untreated seedlings. Later in the autumn, however, the differences disappeared. Before recommending the use of early-season SD-treated seedlings for summer planting, the method has to be tested in practical field conditions.
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Luoranen,
The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland
E-mail:
jaana.luoranen@metla.fi
-
Konttinen,
The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland
E-mail:
kk@nn.fi
-
Rikala,
The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland
E-mail:
rr@nn.fi
article id 215,
category
Research article
Antti J. Lukkarinen,
Seppo Ruotsalainen,
Teijo Nikkanen,
Heli Peltola.
(2009).
The growth rhythm and height growth of seedlings of Siberian (Larix sibirica Ledeb.) and Dahurian (Larix gmelinii Rupr.) larch provenances in greenhouse conditions.
Silva Fennica
vol.
43
no.
1
article id 215.
https://doi.org/10.14214/sf.215
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The aim of this study was to determine whether the growth rhythm and height growth differ between various larch provenances grown in greenhouse conditions. We also investigated whether the geographic and climatic conditions at the origin of the provenance could explain the possible differences between the provenances. The study material consisted of 16 Russian Siberian (Larix sibirica Ledeb.) and Dahurian larch (Larix gmelinii Rupr.) provenances and four seed sources from Finland as comparison lots. The growth rhythm differences were clearest between the southern and northern provenances; the southern provenances grew for a longer period and the proportion of late summer height growth was larger. Autumn colouration also developed later in these provenances. In the Russian larch material the provenances with a longer growing period had greater height growth. In the whole material the relationship was not so linear due to the deviating behaviour of the comparison material. Several of the Russian larch provenances were taller than the Finnish comparison seed lots, although the difference was not statistically significant. However, the growth of these provenances must be followed for a much longer period in field trials before any conclusions can be drawn about their usability in practical forestry.
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Lukkarinen,
University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland
E-mail:
antti.lukkarinen@pp.inet.fi
-
Ruotsalainen,
Finnish Forest Research Institute, Punkaharju Research Unit, Finlandiantie 18, FI-58450 Punkaharju, Finland
E-mail:
sr@nn.fi
-
Nikkanen,
Finnish Forest Research Institute, Punkaharju Research Unit, Finlandiantie 18, FI-58450 Punkaharju, Finland
E-mail:
tn@nn.fi
-
Peltola,
University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland
E-mail:
hp@nn.fi
article id 344,
category
Research article
Juha Lappi.
(2006).
Smooth height/age curves from stem analysis with linear programming.
Silva Fennica
vol.
40
no.
2
article id 344.
https://doi.org/10.14214/sf.344
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Stem analysis data defines a range of possible heights for each age. A smooth stem/age curve is obtained with linear programming (LP) when the sum of the absolute second differences of heights is minimized subject to constraints obtained from the stem analysis. The method is analogous to cubic splines. A LP problem can include additional constraints that are based on the assumption that the crosscut is randomly located within the annual height increment. The method produces smoother height curves than Issa method which is utilizing second order differences of ring counts. It was found using simulated data that the method provides better results than earlier methods for short bolts if height growth is sufficiently regular.
-
Lappi,
Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland
E-mail:
juha.lappi@metla.fi
article id 362,
category
Research article
Hannu Salminen,
Risto Jalkanen.
(2005).
Modelling the effect of temperature on height increment of Scots pine at high latitudes.
Silva Fennica
vol.
39
no.
4
article id 362.
https://doi.org/10.14214/sf.362
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The effect of temperature and precipitation on the height increment of Pinus sylvestris (L.) was modelled using data gathered from a total of 49 felled sample trees from five stands of Scots pine located along a latitudinal transect from the Arctic Circle up to the northern timberline in Finland. A multilevel mixed effects model and cross-correlation analysis of prewhitened time series was used to analyse the dependence between height increment and monthly meteorological observations. The effect of the mean July temperature of the previous year on height increment proved to be very strong at high latitudes (r > 0.7). The mean November temperature of the year before the previous affected statistically significantly on height increment in the three northernmost stands. There was no correlation between height increment and precipitation in any of the sites. The final height increment model based on all stands included tree age, long-term mean temperature sum of site, and the mean July temperature of the previous year as independent variables. According to the model, one degree’s change in July temperature results on average in 1.8 cm change in the next year’s height increment. There was a modest but significant polynomial age-effect. The proportion of explained variance (at the year level) was 74%. The July temperature dependence on height increment was shown to be very strong, suggesting a high value of height increment in climate modelling at the tree line.
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Salminen,
Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland
E-mail:
hannu.salminen@metla.fi
-
Jalkanen,
Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland
E-mail:
rj@nn.fi
article id 361,
category
Research article
Jaana Luoranen,
Risto Rikala,
Kyösti Konttinen,
Heikki Smolander.
(2005).
Extending the planting period of dormant and growing Norway spruce container seedlings to early summer.
Silva Fennica
vol.
39
no.
4
article id 361.
https://doi.org/10.14214/sf.361
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In order to make mechanized planting economically viable, the present spring planting period for Norway spruce (Picea abies (L.) Karst.) seedlings in Scandinavia needs to be extended. To evaluate the possibilities to extend the planting period, six field experiments were established in four years during which frozen-stored, dormant seedlings and actively growing seedlings targeted for spring planting were planted regularly from mid-May to mid-July or the end of August. The survival of actively growing seedlings did not differ between planting dates from mid-May to mid-July. For dormant seedlings, however, the later in summer they were planted the lower was the survival. Oversized seedlings grown in the nursery in containers of too small volume, which were usually planted after mid-June, resulted in reduced growth of seedlings after planting. Root egress (growth of roots from root plugs into the surrounding soil) was most rapid in July and early August and slowest in May and September. Results showed that with dormant seedlings the planting period can be extended from May to mid-June without increasing mortality or reducing growth. The planting period for seedlings stored outdoors and those seedlings that were already growing in June for the purpose of spring plantings can be extended even longer, but it must be kept in mind that the risk of mechanical damage and reduced growth increase due to brittleness of the shoot and increased height. Further research is needed to evaluate the risks in practical scale plantings and with seedlings that are specially targeted for planting after mid-June.
-
Luoranen,
Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland
E-mail:
jaana.luoranen@metla.fi
-
Rikala,
Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland
E-mail:
rr@nn.fi
-
Konttinen,
Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland
E-mail:
kk@nn.fi
-
Smolander,
Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland
E-mail:
hs@nn.fi
article id 360,
category
Research article
Timo Kurkela,
Tarmo Aalto,
Martti Varama,
Risto Jalkanen.
(2005).
Defoliation by the common pine sawfly (Diprion pini) and subsequent growth reduction in Scots pine: a retrospective approach.
Silva Fennica
vol.
39
no.
4
article id 360.
https://doi.org/10.14214/sf.360
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The foliage status in the main stem of Scots pines (Pinus sylvestris) was studied retrospectively using the needle trace method (NTM) on a stand, seriously defoliated by the pine sawfly (Diprion pini) in the 1980s. Needle density increased abruptly in the seasons following the defoliation. The strongest reduction in annual needle production occurred one year later. As a consequence of lower needle production, the annual number of attached needles decreased three to five years after the defoliation. Needle retention and the average age of attached needles tended to increase after defoliation. In analyses of covariance with the NTM variables, needle density (logarithmic transformed values) and average age of attached needles, had the highest, significant, negative relationship with radial and height increments both in the period prior to the defoliation and in the time when the trees were suffering from defoliation. The relationships between height increment and the number of needles and needle loss were positive and significant. Also radial increment had a positive relationship with the number of needles but not with needle loss. Interestingly, an abrupt increase in the needle density gave a good indication of the effects of a sudden defoliation in pines.
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Kurkela,
Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland
E-mail:
timo.kurkela@metla.fi
-
Aalto,
Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland
E-mail:
ta@nn.fi
-
Varama,
Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland
E-mail:
mv@nn.fi
-
Jalkanen,
Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland
E-mail:
rj@nn.fi
article id 373,
category
Research article
Anna Saarsalmi,
Pekka Tamminen.
(2005).
Boron, phosphorus and nitrogen fertilization in Norway spruce stands suffering from growth disturbances.
Silva Fennica
vol.
39
no.
3
article id 373.
https://doi.org/10.14214/sf.373
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Growth disturbance symptoms typical of B deficiency have been reported on Norway spruce (Picea abies (L.) Karst.) trees in many parts of eastern Finland. In order to test the B deficiency hypothesis and explore the possibilities of curing the disturbed trees with B fertilization, three experiments were established in October 1999 in young Norway spruce stands growing on fertile sites in eastern Finland. All the stands contained healthy, slightly and severely damaged trees with growth disturbances typical of B deficiency (B < 5 mg kg–1). 40 healthy, 40 slightly damaged, and 40 severely damaged trees were selected as sample trees in each stand. In May 2000, the trees were fertilized with 2.0 kg B ha–1 as borax (B), 2.0 kg B ha–1 and 40 kg P ha–1 as superphosphate (B+P) or 200 kg N ha–1 as urea (N). The control trees were not fertilized (0). The needle response to B fertilization was rapid, relatively high B concentrations being achieved already after one growing season. Boron fertilization cured the growth disorders and increased height growth within four years, but had no effect on diameter growth. The trees also recovered without B fertilization, but to a lesser extent compared to the B fertilized trees. Compared to the control, boron fertilization increased the height growth in all the disorder classes, i.e. 5, 17 and 19 cm yr–1 for healthy, slightly and severely damaged trees, respectively. As the healthy trees also seemed to benefit from B fertilization, this indicates that B deficiency in fact retards height growth before any disorder symptoms become apparent in individual trees. Compared with B alone, the application of P together with B gave no additional benefit. Nitrogen fertilization alone appeared to have a detrimental effect on height growth in the severely disturbed trees.
-
Saarsalmi,
The Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FI-01301 Vantaa, Finland
E-mail:
anna.saarsalmi@metla.fi
-
Tamminen,
The Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FI-01301 Vantaa, Finland
E-mail:
pt@nn.fi
Category :
Research note
article id 403,
category
Research note
Timo Saksa,
Juha Heiskanen,
Jari Miina,
Jaakko Tuomola,
Taneli Kolström.
(2005).
Multilevel modelling of height growth in young Norway spruce plantations in southern Finland.
Silva Fennica
vol.
39
no.
1
article id 403.
https://doi.org/10.14214/sf.403
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Height development of Norway spruce (Picea abies (L.) Karst.) transplants was studied on 22 sites prepared by disc trenching or mounding. At the age of 4–9 years the plantations were surveyed using a multistage sampling design. For every planted spruce on a plot, the past annual height increments were measured as far into the past as possible. Multilevel mixed linear modelling was used to analyse the variation in growth at different levels (year, stand, cluster, plot, tree) and the effects of climatic and site characteristics on height growth. The within-plantation variation in height growth was higher on mounded sites than on disc-trenched sites. The mean temperature and the precipitation sum of the summer months affected height growth positively. Soil characteristics measured from undisturbed soil did not explain the height growth of seedlings on mounded sites, whereas on disc-trenched sites, the depth of the organic layer and the soil temperature had a positive effect and the depth of the eluvial horizon a negative effect. The modelling approach used proved to be a useful method for examining the sources of variation in development of young plantations.
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Saksa,
The Finnish Forest Institute, Suonenjoki Research Station, FI-77600 Suonenjoki, Finland
E-mail:
ts@nn.fi
-
Heiskanen,
The Finnish Forest Institute, Suonenjoki Research Station, FI-77600 Suonenjoki, Finland
E-mail:
jh@nn.fi
-
Miina,
The Finnish Forest Institute, Joensuu Research Centre, P. O. Box 68, FI-80101 Joensuu, Finland
E-mail:
jm@nn.fi
-
Tuomola,
The University of Joensuu, Mekrijärvi Research Station, FI-82900 Ilomantsi, Finland
E-mail:
jt@nn.fi
-
Kolström,
The University of Joensuu, Mekrijärvi Research Station, FI-82900 Ilomantsi, Finland
E-mail:
tk@nn.fi
article id 421,
category
Research note
Pekka Helenius,
Jaana Luoranen,
Risto Rikala.
(2004).
Effect of thawing duration and temperature on field performance of frozen-stored Norway spruce container seedlings.
Silva Fennica
vol.
38
no.
3
article id 421.
https://doi.org/10.14214/sf.421
Abstract |
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Increasing use of frozen storage in nurseries at northern latitudes calls for thawing methods that are safe, economical and easy to apply on a large scale. The easiest and most economical method would be to thaw seedlings in the same boxes they were stored in. However, doing this safely requires more knowledge about how long and at what temperatures seedlings should or can be kept in the boxes without reducing field performance. In this study, 1-yr-old frozen-stored Norway spruce (Picea abies (L.) Karst.) container seedlings were thawed for 4, 8 or 16 days at 4 or 12 °C in cardboard boxes before planting on a reforestation site and on experimental field in mid-June. Some seedlings were also planted on these locations after thawing for only 7 hours at 12 °C in order to separate frozen root plugs. We found some evidence that planting seedlings after short thawing periods (7 hours at 12 °C and 4 days at 4 °C), under which conditions the root plugs remain completely or partly frozen, has a negative effect on field performance of Norway spruce seedlings. Thawing over a 4-8 day period in cardboard boxes at ca. 12 °C appears to ensure complete thawing of the root plugs and unaffected field performance, but is short enough to prevent the growth of mould.
-
Helenius,
Finnish Forest Research Institute, Suonenjoki Research Station, Juntintie 154, FI-77600 Suonenjoki, Finland
E-mail:
pekka.helenius@metla.fi
-
Luoranen,
Finnish Forest Research Institute, Suonenjoki Research Station, Juntintie 154, FI-77600 Suonenjoki, Finland
E-mail:
jl@nn.fi
-
Rikala,
Finnish Forest Research Institute, Suonenjoki Research Station, Juntintie 154, FI-77600 Suonenjoki, Finland
E-mail:
rr@nn.fi