Articles containing the keyword 'site'

A metabolic model of height growth and site index is derived from a parametrization of the annual carbon balance of a tree. The parametrization is based on pipe-model theory. Four principal variants of the height-growth model correspond to four combinations of assumptions regarding carbon allocation: (a) the apical shoot is autonomous or (b) it is not; and (A) the specific rate of elongation of a shoot equals that of a woody root or (B) it does not. The bB model is the most general as it includes the aA, bA, and aB models as special cases. If the physiological parameters are constant, then the aA model reduces to the form of the Mitscherlich model and the bA model to the form of a Bertalanffy model. Responses of height growth to year-to-year variation in atmospheric conditions are rendered through adjustments of a subset of the model's parameters, namely, the specific rate of production of carbon substrate and three specific rates of maintenance respiration. As an example, the effect of the increasing atmospheric concentration of CO₂ on the time-course of tree height of loblolly pine (Pinus taeda) is projected over 50-year span from 1986. Site index is predicted to increase and, more importantly, the shape of the site-index curve is predicted to change.

• Valentine, E-mail: hv@mm.unknown

Models for individual-tree basal area growth were constructed for Scots pine (Pinus sylvestris L.), pubescent birch (Betula pubescens Ehrh.) and Norway spruce (Picea abies (L.) Karst.) growing in drained peatland stands. The data consisted of two separate sets of permanent sample plots forming a large sample of drained peatland stands in Finland. The dependent variable in all models was the 5-year basal area growth of a tree. The independent tree-level variables were tree dbh, tree basal area, and the sum of the basal area of trees larger than the target tree. Independent stand-level variables were stand basal area, the diameter of the tree of median basal area, and temperature sum. Categorical variables describing the site quality, as well as the condition and age of drainage, were used. Differences in tree growth were used as criteria in reclassifying the a priori site types into new yield classes by tree species. All models were constructed as mixed linear models with a random stand effect. The models were tested against the modelling data and against independent data sets.

• Hökkä, E-mail: hh@mm.unknown
• Alenius, E-mail: va@mm.unknown
• Penttilä, E-mail: tp@mm.unknown

The purpose of this study was to test the benefits of a forest site quality map, when applying satellite image-based forest inventory. By combining field sample plot data from national forest inventories with satellite imagery and forest site quality data, it is possible to estimate forest stand characteristics with higher accuracy for smaller areas. The reliability of the estimates was evaluated using the data from a stand-wise survey for area sizes ranging from 0.06 ha to 300 ha. When the mean volume was estimated, a relative error of 14 per cent was obtained for areas of 50 ha; for areas of 30 ha the corresponding figure was below 20 per cent. The relative gain in interpretation accuracy, when including the forest site quality information, ranged between 1 and 6 per cent. The advantage increased according to the size of the target area. The forest site quality map had the effect of decreasing the relative error in Norway spruce (Picea abies) volume estimations, but it did not contribute to Scots pine (Pinus sylvestris) volume estimation procedure.

• Tokola, E-mail: tt@mm.unknown
• Heikkilä, E-mail: jh@mm.unknown

Growth data were collected from 40 European aspen (Populus tremula L.) stands growing on eight localities in Sweden. The stands ranged in latitude from 56 to 66°N. The mean age of the stands was 32 years (range, 12–63), the mean stand density 1978 stems ha^-1 (range, 300–6,000), and the mean diameter at breast height (on bark) 17 cm (range, 8–34).

Site index curves were constructed for total age. Curves for H₄₀ (dominant height at 40 years total age) were made for total Sweden. Curves fitted for H₄₀ total age have another shape than curves presented by other Nordic studies. The curves from the present study have slower growth for young aspens than curves from Norwegian and Finnish conditions. For 50–70-year-old aspen stands, curves from the present study indicate taller heights than from Nordic studies.

Classified soil types from the stands were grouped into three groups: sandy till (17), light clay (15) and medium clay till (4). As there was only one stand growing in the fine sand group and one stand in the heavy clay till group and two stands in the silty till group, these stands were not presented with growth curves. There were no statistically significant differences in site index between the three soil type groups. Some recommendations for management of aspen stand are given. Damages caused by moose, fungi and other injuries are discussed as a problem for height yield production and a good timber quality.

• Johansson, E-mail: tj@mm.unknown

Fresh and herb-rich upland forest sites in the north-western part of the central boreal vegetation zone in Finland were studied with respect to vegetation structure and vegetation-environment relationships (soil, stand characteristics). Two fresh heath vegetation data sets, one from the northern boreal zone and the other from the central boreal zone, were compared with the data of this study using multivariate methods.

The variation in heath forest vegetation within the climatically uniform area was mainly determined by the fertility of the soil (primarily Ca and Mg) and the stage of stand development. N, P and K content of the humus layer varied little between the vegetation classes. Fertile site types occurred, in general, on coarse-textured soils than infertile site types, may be due to the fact that the sample plots were located in various bedrock and glacial till areas, i.e. to sampling effects.

The place of the vegetational units of the study area in the Finnish forest site type system is discussed. The vegetation of the area has features in common with the northern boreal zone as well as the southern part of the central boreal vegetation zone. The results lend some support to the occurrence of a northern Myrtillus type or at least that intermediate form of fresh and herb-rich mineral soil sites commonly occur in the studied area. It is argued that the older name Dryopteris-Myrtillus type is more suitable than Geranium-Oxalis-Myrtillus type for herb-rich heath sites in the study area.

• Lehtelä, E-mail: ml@mm.unknown
• Hotanen, E-mail: jh@mm.unknown
• Sepponen, E-mail: ps@mm.unknown

It was examined whether the present site classification method, and especially its applicability to site productivity estimation, could be improved in upland Scots pine (Pinus sylvestris L.) forests in Southern Finland by developing a classification key based on Two-way Indicator Species Analysis (TWINSPAN), and/or by inclusion of soil texture, stoniness and the humus layer depth more closely in the classification method. TWINSPAN clusters (TW) explained 71%, and forest site types (FST) 64% of the variation in site index (SI) (H₁₀₀). When soil texture (TEXT) was added to the regression model, the explanatory power increased to 82% (SI = TW + TW * TEXT) and to 80% (SI = FST + FST * TEXT), respectively. Soil texture alone explained 69% of the variation in site index. The influence of stoniness on site index was significant (P <0.05) on sorted medium sand soils and on medium and fine sand moraine soils. The thickness of the humus layer (2–6 cm) was not significantly (P=0.1) related to site index.

It is suggested that the proposed TWINSPAN classification cannot replace the present forest site type system in Scots pine stands in Southern Finland. However, the TWINSPAN key may be used to aid the identification of forest types. The observation of dominant soil texture within each forest type is recommended.

The PDF includes an abstract in Finnish.

• Nieppola, E-mail: jn@mm.unknown

Five ploughed research areas from Finnish Norther Karelia were selected for comparison studies of plough ridges and untouched soil. Measurements were made at a depth of 10 cm in sample plots on both mineral and paludified mineral soil and peatland parts of these areas. In summer 1987 daily soil water matric potential was measured using tensiometers, and volumetric soil moisture content and density were determined from soil samples at two dates during the summer. Water characteristics of the core samples were also determined. On paludified mineral and peat soils the water table depth from the soil surface was measured.

The results indicated that in plough ridges matric potential was lowest. Plough ridges were also seen to dry and wet faster and to a greater degree than untouched soils. In untouched soils, soil water relations and aeration were not affected by the distance to the furrow. The effect of the plough ridge was smallest on peatland, where there was a good capillary connection from plough ridge to the ground water, if the ditches were not very effective. The soil in the ridges did not dry too much to restrict seedling growth. The untouched surface soil in poorly drained peat and paludified minear soil was, at least in a rainy growing season, often and also for long times so wet that 10% minimum air space required for good seedling root growth was not available.

The PDF includes an abstract in English.

• Mannerkoski, E-mail: hm@mm.unknown
• Möttönen, E-mail: vm@mm.unknown

In soil profiles from a series of uplands of different site types bulk density, density of solids and porosity of soil were clearly related to soil organic matter content and its distribution in the soil profile. Soil organic matter contents were also strongly correlated to effective cation exchange capacity (CEC) and soil acidity. Site fertility was primarily related to the fine fraction (ø<0.06 mm) content in the C horizon and related properties (i.e. CEC). In the humus layer, the content of exchangeable bases and base saturation most strongly related to site fertility.

The PDF includes an abstract in English.

• Westman, E-mail: cw@mm.unknown

A close relationship between photosynthetic capacity and nitrogen concentration of leaves is known to exist. In conifers, nitrogen also affects the pattern of mutual shading within a shoot, which is a basic unit used in studying photosynthesis of coniferous trees. These effects of needle nitrogen concentration on photosynthetic capacity and mutual shading of needles were analysed for Scots pine (Pinus sylvestris L.) shoots taken from five young stands growing on sites of different fertility. The effect of nitrogen concentration on needle photosynthesis was studied based on measurements of the photosynthetic radiation response of shoots from which two thirds of the needles were removed in order to eliminate the effect of within shading.

An increase of one percentage unit in nitrogen concentration of needles increased the photosynthetic capacity of needles by 25 mg CO₂ dm^-2h^-1. The effect of nitrogen on within-shoot shading was quantified in terms of the silhouette area to total needle area ratio of a shoot (STAR), which determines the relative interception rate per unit of needle area on the shoot. Although nitrogen promoted needle growth, an increase in nitrogen concentration decreased the within-shoot shading. This effect resulted from a decrease in needle density on the shoot and an increased needle angle with increasing nitrogen content.

The PDF includes an abstract in Finnish.

• Smolander, E-mail: hs@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown
• Oker-Blom, E-mail: po@mm.unknown

The winter 1986–87 was unusually cold; the snow cover remained thin and consequently the soil froze to a considerable depth. In spite of the severe frost, the lowest temperatures measured at the ground surface was -10.3°C and in the soil at the depth of 10 cm -5.8°C. The temperature sum of the following summer was unusually small and the soil frost melted more slowly than usual. The winter frosts did not have a decisive influence on the survival of planted seedlings.

The PDF includes an abstract in English.

• Kubin, E-mail: ek@mm.unknown

Paratization of Petrova resinella L. in the gall stage was studied in the surrounding of Harjavalta, south-western Finland, in relation to industrial air pollution. Of the studied 283 galls, 28% produced a moth, 52% of the larvae/pupae were paratisized, and 20% were empty or contained a dead larva. The proportion of paratisized galls did not depend on the distance from emission sources of industrial air pollutants.

The PDF includes an abstract in Finnish.

• Heliövaara, E-mail: kh@mm.unknown
• Väisänen, E-mail: rv@mm.unknown

A field data set representing boreal forest-floor vegetation in Southern Finland was analysed using a simultaneous equation model. Some physical and chemical characteristics of the soil and some structural characteristics of the tree stand were treated as predictors in such a way that the tree stand factor was specified to be dependent on the soil variables. Alpha diversity, measured as the total number of species per plot, was treated as a criterion variable.

The model explains 60% of variance in the alpha diversity indicating markedly strong relationships with the site characteristics. Alpha diversity appears to increase with increases in site fertility characteristics. On the other hand, measured characteristics of the tree stand indicate no significant independent effects on the alpha diversity.

The PDF includes a summary in Finnish.

• Kuusipalo, E-mail: jk@mm.unknown

Vegetation data collected from a random sample of mature forest stands representing mesic upland forest sites in Southern Finland were analysed and classified using two-way indicator species analysis (TWINSPAN). The variation of some characteristics of the tree stand and soil fertility between the produced vegetation units were analysed statistically.

Both the species list and the sample list arranged by TWINSPAN reflects the overall site fertility considerably well. The results are in agreement with the main division of mesic forest sites in the Finnish forest site type classification: vegetation units which can be assigned to the Oxalis-Myrtillus site type are clearly separated from the remaining units, and the overall site fertility indicates a statistically significant difference. The within-type variation in the vegetation composition in the Oxalis-Myrtillus site type reflects the variation in site fertility, whereas the within-type variation in the Myrtillus site type is mainly caused by the tree stand factor.

The PDF includes a summary in Finnish.

• Kuusipalo, E-mail: jk@mm.unknown

The particle size distribution affects several properties of the soil, thus, the ability to define the texture type of the soil as accurately as possible in field conditions is essential. The soil particle size classification devised by Atterberg (1912) is used in Finnish forestry. The study is based on a small laboratory material. The correlation between some characteristics of the soil particle size distribution, field capacity and cation exchange capacity were determined.

The particle size characteristics such as the relative proportion of different particle sizes, average particle size (Md) and parameters depicting the degree of sorting were determined. The relative proportion of soil particles below 0.06 mm correlated best with both field capacity and cation exchange capacity. Similarly, the average particle size and the degree of sorting correlated well with the field capacity and the cation exchange capacity.

The use of sorting characteristics is not well-suited to the type of soil sample material containing a high proportion of particles of varying size as was used in this material. Such characteristics are probably more easily applicable to the fine sand and sand sediments which are predominant in Finnish forest soils. The most useful particle size distribution characteristics in soils having a great variation in particle sizes were the average particle size and the relative proportion of silt and clay. Thus, the nutrient and water status of the soil can be predicted to some extent by examining the percentage of silt and clay, average particle size and the degree of sorting.

The PDF includes a summary in English.

• Sepponen, E-mail: ps@mm.unknown

The ground cover vegetation and tree growth at several camping sites in Southern Finland were investigated. The deterioration of ground vegetation appeared to be unavoidable in these areas, and only a few grass species were tolerant to heavy trampling. Only moderate deterioration was, however, detected when the trampling level was lower than 10,000–15,000 user days. On the other hand, the ground cover was completely destroyed when the trampling level exceeded 100,000 user days. A considerable decrease in radial growth appeared to be associated with destruction of the ground vegetation. The decrease was abrupt and was found to continue throughout the whole period of use. After ten years’ use the growth in the trampled areas was 35% lower than that that of the untrampled areas. A further decrease in radial growth is expected in the future. 

The PDF includes a summary in Finnish. 

• Nylund, E-mail: mn@mm.unknown
• Haapanen, E-mail: ah@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown
• Nylund, E-mail: ln@mm.unknown

The effect of nitrogen fertilizers on the photosynthetic capacity of conifers is assessed on the basis of literature. The review emphasizes the role of changes of needle mass as a factor affecting the result of nutrient application. In particular, the increase in needle mass results in a considerable increase in photosynthetic capacity. The effect of fertilization on the photosynthetic rate seems to be of minor importance. The effect on the photosynthetic rate is, however, poorly documented as is the case for the effect of fertilization on the respiration rate. There is evidence that proper application of nitrogen fertilizers may double the photosynthetic capacity of conifers, mainly as a result of increase in needle mass.

The PDF includes a summary in English.

• Kellomäki, E-mail: sk@mm.unknown

About one million hectares of forests are fertilized annually in Finland. The goal of the present study was to find out, by means of calculations, how the profitability of forest fertilization varies with variations in the stage of development of tree crops, the quality of the site and its geographical location. Calculations concerned bot fertilization of forests in mineral soil sites and in drained peatlands. The study is a part of a larger project concerning the order of profitability of different forest improvement measures in different conditions. The problems dealt with in this study were approached from the point of view of national economy.

On the basis of two empirical materials it is shown that there is a high correlation between the stand growth percentages before and after the fertilization. Applying the results to existing yield tables the authors calculate benefit/cost ratios showing the stage of development of the stand, the quality of the site and its geographical location. According to the results, fertilization is more profitable in sites of medium fertility than on poor sites. Profitability decreases rather fast from south to north and with decreasing timber prices. 

The PDF includes a summary in English.

• Keltikangas, E-mail: mk@mm.unknown
• Seppälä, E-mail: ks@mm.unknown

No other manifestation of life is allied more conspicuously to the theory of relativity as the growth of forest stands which is a function of the inherent growth potential of trees, the productive capacity of environment, and time.

The height over age quotient of a forest stand is usually the most reliable indicator of the productive forces of the habitat. Stem analysis have shown that increment of a tree at different ages is closely correlated with the extension of roots into individual geological horizons of different productive capacity. Growth curves of stands of a same tree species growing on different soils can be disparate due to different conditions. The temporal variety of tree growth on different sites is of prime importance in the construction of yield tables. Investigations of natural plant communities of Finland provided one rational approach towards the construction of yield tables. By confining mensuration analyses to define floristic types, the Finnish foresters harmonized their records with Einstein’s formula for space-time matrix of material events.

• Wilde, E-mail: sw@mm.unknown

The paper is a review on the topics of Symposium on forest types and forest ecosystems, held in connection to the IX internal botanical congress in Montreal in August 1959, the chairman of which was Ilmari Hustich. The article includes 18 preparatory papers that were distributed among the participants of the symposium. The common theme of the papers was the question of finding common platform for the different schools of forest types and forest ecosystems. In addition to the papers, the article includes a summary of the proceedings and discussions of the symposium.

The following papers were presented in the symposium:

Aichinger, E. Können wir eine gemeinsame Platform für die verscheidenen Schulen in der Waldtypenklassifikationen finden?

Arnborg, T. Can we find a common platform for the different schools of forest type classifications?

Dansereau, P. A combined structural and floristic approach to the definition of forest ecosystems.

Daubenmire, R. Some major problems in vegetation classification

Ellenberg, H. Können wir eine gemeinsame Platform für die verscheidenen Schulen in der Waldtypenklassifikationen finden?

Hills, G.A. Comparison of forest ecosystems (vegetation and soil) in different climatic zones

Kalela, A. Classification of the vegetation, especially of the forest, with particular reference to regional problems

Krajina, V.J. Can we find a common platform for the different schools of forest type classifications?

Kühler, A.W. Mapping tropical forest vegetation

Linteau, A. Y. a-t-il. Un terrain d’entente possible entre les différentes écoles au sujet de la classification de types forestiers?

Medvecka-Kornaś, A. Some problems of forest climaxes in Poland

Ovington, J.D. The ecosystem concept as aid to forest classification

Puri, G.S. The concept of climax in forest botany as applied in India

Rowe, J.S. Can we find a common platform for the different schools of forest type classifications?

Scamoni, A. Können wir eine gemeinsame Grundlage für die verscheidenen Schulen in der Waldtypenklassifikationen finden?

Sukachev, V.N. The correlation between the concept ’forest ecosystem’ and ’forest biogeocoenise’ and their importance for the classification of forests

Webb, L.J. A new attempt to classify Australian rain forest

• Heikurainen, E-mail: –
• Finnish Society of Forest Science, E-mail: –

Professor V. Sukachev, a Soviet developer of the doctrine of forest types, published a guide to the study of forest types in 1931. He states in the introduction of the guide that his aim is to facilitate the work of beginners in determining forest types and to instruct them in developing methods for their description and classification.
This Finnish edition of the guide includes new picture material and has an extensive supplementary preface, written by the author, that presents his view on forest types. The guide describes the concept of forest types, methods of describing forest types, guidelines to material collected in the forests and principles of classification of the different forest types.

• Sukatsev, E-mail: vs@mm.unknown

The article is a summary of a presentation by W. Wittich, held in the University of Helsinki in 26.4.1958.
The significance of fertility of the site in tree growh was known already for over 100 years ago in Germany, but after the First World War the view was abandoned in forestry. According to the Dauerwald system of managing forests that was introduced at the time, the type of site was considered relatively insignificant in forest management. Therefore, similar practices were used in all kinds of sites. The opposition against the use of this method resulted in new research on the site factors.
Knowledge of the relation of the site types and vegetation makes it possible to improve productivity: in regional planning the production that is considered to be necessary is assigned to the sites that have best conditions for it. For instance, in Niedersachsen county about 6% of the forest lands are reserved for cultivation of oak.
Another line of soil science studies the root causes behind the hands-on experiences of forest management. The aim is to abandon rigid approaches in forestry. Studying the effects of forest management practices on soil has been targeted, for instance, on effects of clear cutting on decomposition and vegetation, how the soil affects choice of tree species, and decomposition of litter from different tree species. Knowledge of soil and the trees’s demand of nutrients helps to mend disturbancies, such as nutrient deficiensies. Consequently, fertilization has become a new tool to improve productivity in forestry.
The article includes a German summary.

• Wittich, E-mail: ww@mm.unknown

A committee was appointed in 1940 to prepare a proposal for reform of forest taxation. The taxation based on net income of forestry was considered to have limitations, and the actual net income had been observed to be markedly higher than the income that was used in taxation.

The report describes in detail the principles and shortcomings of taxation used since 1922. These include inaccuracies in the forest areas of a woodland estate, and weaknesses in classification of forest land and demand zones. The committee suggests several improvements in calculating the taxable income, which in the new calculations is based on yield on terms of value.

The article includes an abstract in German.

• Metsäverotuksen uudistamiskomitea, E-mail: –

Aspen (Populus tremula L.) is a common tree in Finland, and has been used, for instance, in matchstick industry. However, there has been little studies on its distribution and properties. In this study, 142 sample trees in different forest site types in Valtimo and Onkamo in Eastern Finland were measured in detail in 1935.

According to the results, during the first 10 years aspens height growth is fastest of the Finnish tree species surpassing, for instance, Scots pine (Pinus sylvestris L.) and birch (Betula sp.) . The diameter growth is similar to Scots pine up to the age of 50 years, after which the growth of aspen exceeds Scots pine. Branchless portion of the stem compared to the height of the tree increases until it reaches about 50% of the height of the tree. In poorer sites aspen is prone to decay.

Aspen regenerates easily both by root shoots and seeds. If root shoots are left to grow, the mother tree should be free of decay. In general, seedlings are of better quality. Good quality aspen stands require thinning and a rich forest type. If an old aspen stand has decay, the trees should be ring-barked and the site regenerated with a new tree species.

The article includes an abstract in German.

• Blumenthal, E-mail: bb@mm.unknown

Silva Fennica issue 52 includes presentations held in professional development courses, arranged for foresters working in public administration in 1938. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.

This presentation describes growth and form of root systems of different tree species in different sites and how growth of roots affect forest management.

• Aaltonen, E-mail: va@mm.unknown

Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service. 

This presentation describes forest regeneration on poor forest sites.

• Laitakari, E-mail: el@mm.unknown

The aim of the study was to follow development of vegetation in dry upland forest sites after forest fire. The sample sites were situated in the counties of Muonio, Kolari, Sodankylä, Pelkosenniemi, Savukoski, Kemijärvi and Salla, in the northernmost Finland.
The growth of plant communities can arise either from the vegetation and seeds that survived the fire, or from seeds that spread from the surrounding areas. The development of vegetation in the burned areas was unexpectedly independent of the surrounding areas, which indicates that role of the seeds from the outside of the burned ares is small. The occurence of different species of lichens, moss, scale moss and vascular plants in the burned areas are described in detail. The development of vegetation was strongly dependent on the forest site type. The thin humus layer of Cladina site type burns usually evenly, and also the vegetation develops more evenly than in the more fresh site types. Vegetation typical for burned areas was fully developed within 10-15 years, and after 25 years it began to resemble the vegetation of Cladina site type forests. The ground vegetation of Calluna type burned area was more patchy. It developed quicker than in Cladina type. Absense of lichens made it seem more fertile than is usual for Calluna type. The humus layer of Empetrum-Myrtillus site type burned unevenly, and if the area was lightly burned, the vegetation recovered quickly. The vegetation was often patchy.
The PDF includes a summary in German.

• Sarvas, E-mail: rs@mm.unknown

• Finnish Society of Forest Science, E-mail: sms@mm.unknown

The article is a description of the 75th anniversary of Finnish Forest Service (now Metsähallitus, Forest Service), held on April 19^th 1934. The greetings of the quests, presented in the Commemoration festival and Banquet, are included in the article.

• Finnish Society of Forest Science, E-mail: fsfs@mm.unknown

Typical for the area of Längälmävesi, in Satakunta in Southern Finland, are densely populated fertile lowland areas near the waterways and poorer sparcely populated upland areas. The changes in fertility of the land influence the vegetation, and has directed where the population has settled. A vegetation survey was made using specific plant species as indication to fertility. A detailed description of the species composition and distribution of the plant species indicative for fertile land is presented in the article.

Analyzing the fertility only based on the plant species indicative for rich soils underestimates the proportion of fertile lands. Farmlands, on the other hand, have been cleared also on less fertile soils, which would give an overestimate of the fertile lands. A map of fertile lands was drawn based on both vegetation and location of the fields. The changes in fertility influenced also bird species observed in the area.

The PDF includes a summary in German.

• Cajander, E-mail: oc@mm.unknown

Metsähallitus (Forest Service) decided to protect two areas around Hiisijärvi lake in Eastern Finland already in 1916. Later, a natural park was suggested to be established in the area. A survey of the vegetation in the area was composed in 1931-1932. The total land area of the protected area was 3.5 km³. A vegetation map was drawn based on a nature inventory. A detailed description of the forest site types, peatland types, aquatic flora and the vegetation of the area are included in the article. The calcareous soil promotes rich vegetation. Typical for the area are also rich fens. The area can be divided to a eutrophic and a oligotrophic part.

The PDF includes a summary in German.

• Brandt, E-mail: ab@mm.unknown

The article is an obituary of Dr. Lauri Ilvessalo, who died in 1928. He worked as a professor of silviculture in 1918-24, and published numerous scientific articles on silviculture. The article includes his presentations on the state of forestry in Finland, and on the establishment of Forest Service in Finland and the public attitude towards the organization.

The PDF includes a summary in German.

• Finnish Society of Forest Science, E-mail: fs@mm.unknown

The article contains three presentations given about forest type classification at the University of Tarto in Estonia. The article has an introduction, a part about the meaning of the natural classification of forest sites and up to now conducted studies on site classification. The second part presents the characteristics of plant communities and the forest types, and practical and theoretical meaning of forest types.

Classifying the forest sites is important in practical forestry, because the forest growth and forest valuation are dependent on the productivity of the soil. The classification of the sites for forest management purposes needs to result in classes that are easily distinguished in the forest. This then leads to forest management that best fits to a certain forest site. 

• Cajander, E-mail: ac@mm.unknown

Norway spruce (Picea abies (L.) H. Karst.) is rarely the dominant species on dry mineral soil sites in Northern Finland. These sites are, in general, too poor and dry for spruce, and suit better for Scots pine (Pinus sylvestris L.). According to the study, the natural regeneration of spruce is in Northern Finland poor. In the sample plots, cones could be found in 35% of spruce trees in the stands in natural state and 46% in the harvested stands. Compared to the spruce areas in Northern Finland, or fresh mineral soil sites in Southern Finland, cone and seed production of Norway spruce was in dry mineral soil sites very low due to scarcity of seed trees and their low cone number. There were few spruce seedlings in the sample plots, but according to the observations, spruce is able to regenerate on lichen and heath covered sites. The seedling growth was, however, poor on dry sites. Spruce seedlings were often found near fallen trees and stumps. The growing trees prevent growth of seedlings of all species. Norway spruce seems, however, to be able to spread also to the poor sites. The success depends on the vegetation and dryness of the site. For instance, spruce can spread to dry mineral soil sites from seed trees of nearby peatlands.

The PDF includes a summary in German.

• Tikka, E-mail: pt@mm.unknown

The article is a review on methods used in Central Europe and in Finland to construct growth and yield tables, and on their defects and advantages. One of the main defects of growth and yield tables prepared previously in Central Europe is that the site quality classes for different tree species have been formed independently and using different principles. Thus, the yields of different tree species on a similar site can’t be compared. In addition, the quality classes of this kind of growth and yield tables and growth series are artificial.

The Society of Forestry in Finland (now Finnish Society of Forest Sciences) started to prepare new yield tables for the most important tree species in Finland in 1916. The new yield tables aimed to improve the yield tables made in Central Europe in two aspects. Firstly, the quality of the site of each sample plot was assessed on the spot and independently from the standing crop. Thus the sample plots of each site class in setting up the growth series can be treated as independent groups. Consequently, the quality classes are be the same for all tree species. Secondly, mathematical-statistical methods were used to determine which of the stands that belong to the same quality class belong also to the same growth series.

The PDF includes a summary in Finnish.

• Ilvessalo, E-mail: yi@mm.unknown

The article is a lecture given by A.K. Cajander in the International Congress of Plant Science. The lecture describes results of Finnish forest research that might be regarded significant also for North America. Because of similarities in nature and forest management, forest research may use similar methods in both areas.

For instance, line plot survey in the form used in Finland could well be applied in North America. In Finland, lines were drawn at 26 kilometer intervals. Visual estimates about, for instance, species, tree growth and productivity class, were made along the lines and sample plots were taken every other kilometer. To gain full advantage of the method, a productivity classification and yield tables are needed. When these are known, it is possible to find out how to increase the productivity of forests with suitable tree species and proper forest management. This kind of inventory of forest resources and the state of forests provides reliable information for forest policy. Another important issue for forest research is forest management, which requires understanding on their biology. At the same time, research must provide methods for practical forestry.

A summary in Finnish is included in the PDF.

• Cajander, E-mail: ac@mm.unknown

One of the difficulties in constructing growth and yield tables has been to determine which of the sample plots growing the same tree species and belonging to the same forest site type, with reference to the quality of stands, have to be included in the same growth series.

New growth and yield tables for the most important tree species were constructed in Finland in 1916–1919, using new principles that aim at avoiding some of the common weaknesses. There were two main differences to the earlier work. First, the site quality class (forest site type) was determined for each sample plot when the sample plot was measured, independently of the stand occupying the site. In this way it was possible to treat the sample plots of each site as an independent group from the beginning, and so that the quality classes were the same for all the tree species. Second, mathematic-statistical methods were used to deduct the so-called stem frequency distribution series, when studying which of the sample plots of the same quality class belong to the same growth series. They represent the average number of stems of the different diameter classes. A more detailed description of the method used to create the growth and yield tables is published in Acta Forestalia Fennica no. 15.

In the PDF is included a summary in Finnish.

• Ilvessalo, E-mail: yi@mm.unknown

The article highlights the need to classify the forest sites in objective and exact classes. This is important both from a practical and a scientific point of view as well as from a silvicultural point of view, for the forest management varíes for each tree species, and according to the site, even if the species remains the same. It is evident that the same classification of sites according to quality ought to be applicable to silviculture, forest mensuration and statistics. In Finland, a forest site type rating has been created for this purpose.

The PDF includes a summary in Finnish.

• Cajander, E-mail: ac@mm.unknown

Result of a survey of soils supporting forest plantations in Wisconsin in the United States indicated a close correlation between the levels of fertility of non-phreatic, coarse-textured soils and the growth of red pine (Pinus resinosa Roezl) stands aged from 15 to 32. This relationship, however was not observed in plantations established on deep-gley soils, underlain at a depth of 3–9 fl by ground water.

The survey encountered 20 red pine plantations on soils underlain by a deep ground water table accessible to tree roots thorough their contact with gley horizon or with extended capillary fringe. The average growth of the stands was 80 cubic feet/acre (5.6 m³/ha) at the age of 22 years. Thus, mensuration analysis suggested that the soils are the choice grounds for forestry enterprise. However, the analysis of soil samples showed that in many instances the soils are extremely low in mineral colloids, organic matter and nutrients. Many of the sites would be regarded as critically deficient in nitrogen, phosphorus and potassium.

The following hypothesis are suggested to explain this discrepancy:

a) The moisture content of coarse-textured non-phreatic soils remain near the wilting point during a large apart of the growing season with subsequent reduction of transpiration and uptake of nutrients. If a capillary fringe provides a supply of water for the root system, trees may derive an adequate supply of salts and exchangeable ions from comparatively infertile substrata.

b) The suitably located ground water provides adequate aeration of the surface soil layers which is not impeded by capillary fringe, increasing activity of mycorrhiza, and a mycotrophic uptake of nutrients from unweathered minerals.

c) The above effects of natural subirrigation should change the concept of soil fertility based on mere chemical analysis. The time during which the roots are engaged in active absorption appears to be of equal importance as the concentration of nutrients in available form.

The PDF includes a summary in Finnish.

• Wilde, E-mail: sw@mm.unknown
• Iyer, E-mail: ji@mm.unknown

This special volume of Acta Forestalia Fennica is published in memory of professor A.K. Cajander (1879-1943), who was one of the founders of the Finnish Society of Forest Science. It contains a short biography, a complete list of his literary works and one of the papers on forest types he has written.

This article contains professor A.K. Cajander’s paper on the theory of forest types in English. The article can be found in Finnish in separate PDF (article id 7695).

The idea of forest types was developed by professor Cajander first in 1904-1909, when he was working as a student in the Evo Forestry Institute. The first publication in 1909 was followed by intensive research and the findings of the investigations were published in 1929 in Acta Forestalia Fennica 29 (Metsätyyppiteoria, The theory of forest types).

When classifying forest growing sites, the purpose is to combine those having the same or approximately same yield capacity, and to separate into different classes those whose yield capacity is widely different. The article states that site quality classification is a necessity in organized forest management as it gives a basis for all forest calculations concerning with yield and profit. The principles of site classification and its meaning in forest management is discussed.

• Cajander, E-mail: ac@mm.unknown

This special volume of Acta Forestalia Fennica is published in memory of professor A.K. Cajander (1879-1943), who was one of the founders of the Finnish Society of Forest Science. It contains a short biography, a complete list of his literary works and one of the papers on forest types he has written.

This PDF includes his article on forest types in Finnish. The translation of the text in English can be found in a separate PDF (article id 7396).

The idea of forest types was developed by professor Cajander first in 1904-1909, when he was working as a student in the Evo Forestry Institute. The first publication in 1909 was followed by intensive research and the findings of the investigations were published in 1929 in Acta Forestalia Fennica 29 (Metsätyyppiteoria, The theory of forest types).

When classifying forest growing sites, the purpose is to combine those having the same or approximately same yield capacity, and to separate into different classes those whose yield capacity is widely different. The article states that site quality classification is a necessity in organized forest management as it gives a basis for all forest calculations concerning with yield and profit. The principles of site classification and its meaning in forest management is discussed.

• Cajander, E-mail: ac@mm.unknown

Department of plant pathology of University of Helsinki conducted studies in the surrounding forests of Viikki test farm and on the Hyytiälä forest training station to find about occurrence and spore production of pine needle rust.

The damages have been minor. Most often the infection did not cause yellowing of the needles, only individual needles might have dried up. There were no dead young trees.

The spore production was strongest at the more fertile sites, with abundant occurrence of cow-wheat (Melanpyrum spp.). The infection caused more harm on the cow-wheat than on the pines. In many cases the foliage died prematurely. Melanpyrum spp. were more strongly infected on sites with more light. However, there was no difference found with the fertility of the site.

The PDF contains a summary in Finnish. 

• Pohjakallio, E-mail: op@mm.unknown
• Vaartaja, E-mail: ov@mm.unknown

The article presents the characteristics of different vegetation areas (meadows and peatlands) by their distinctive vegetation. The study area is by the Barents Sea and is the northernmost part of continental European Russia. Different sites are classified by plant communities and/or vegetation units.

The article continues on the second PDF-file. 

• Kalela, E-mail: ak@mm.unknown

The valuation of forest land for financial accounting purposes is usually performed only when using methods that are based on wood resources. In the yield based methods, the book value of forest land and wood resources form one totality. In the first case, forest land in a separate land account usually has same value in the beginning and end of the accounting year. For instance, the costs of forest improvement are considered capital costs. Forest land can be valued either by multiplying the average hectare price of land with the hectares, or using separate unit prices for the different forest site types. Different ways to value forest land are presented, comparing the forest site type classification developed in Finland and the traditional method based on average height of the trees used in Central Europe. The study shows that values of forest land has relative nature.

The PDF includes a summary in German.

• Keltikangas, E-mail: vk@mm.unknown

The development roots of Norway spruce (Picea abies (L.) H. Karst.) seedlings was studied in sample seedlings grown in different kinds of sites. In the early stage, the seedling roots grow primarily length. The main root is usually long. If the growth of the root is hindered, the tip of the root dies, and the root system growing from the original root collar remains relatively small; in these cases, the secondary root system becomes more important. In unfavourable conditions the root branches can early on replace the main root. The main root of a germling seems to be less able to seek for free growing space than the main and side roots of older seedlings. When the growth of the root is blocked by some kind of obstacle, it does not often hinder the growth of the seedling. The type of soil influences strongly how the root system grows. In good soil and in humus the root system is regular and richly branched, while in clay and coarse sand the root system was small. Spahgnum moss was good substrate for seedlings, Dicranum undulatum moss little less good, while the seedlings grew poorly on Pleurozium Schreberi.

The PDF includes a summary in German.

• Hertz, E-mail: mh@mm.unknown

About 40% forest in Finland are mixed stands that have birch (Betula pubescens and B. verrucosa) as one of the species. The aim of this research was to study the structure of root system of birch and compare it to the other main tree species in Finland.

The root systems were dug out and measured in 28 sample plots in Southern and Central Finland, representing different forest site types. Birch roots correspond 30‒100% of the volume of the stem, the largest root systems being in the sandy soils or peatlands. Also the longest lateral roots can be found at these sites. The size variation of root system of birch is larger than in Scots pine (Pinus sylvestris L.), and the vertical root system is in general smaller in birch. Birch seems to be better than pine able to adapt its root system to the existing conditions. The smallest root systems were found in the good forest site types, but the roots grow in the good sites denser than in the poor sites. The lateral roots of the main tree species in Finland, birch, Scots pine and Norway spruce (Picea abies (L.) H. Karst.) grow in different depths, which decreases the competition between the species. This finding gives support to cultivation of mixed stands.

The PDF includes a summary in English.

• Laitakari, E-mail: el@mm.unknown

The abundance of Norway spruce (Picae abies (L.) H. Karst.) undergrowth is common for the state forests in Karelia near the Russian border, in Finland. In the survey, the occurrence of the undergrowth was studied. The article includes a review on the ownership of the forest, forest soils in the area, and the state of forests in the area. Scots pine (Pinus sylvestris L.) is the dominative species in 67%, Norway spruce in 27% and Betula sp. 6% of the state forests. Only 2% of the forests are 1‒20 years of age. Stands in the age group of 61‒80 years are the most common (25%). Norway spruce undergrowth is most abundant in the municipality of Salmi. The forests are typically moist forest site types or grass-herb site types. If the stands are allowed to develop naturally, even the Vaccinium sites become Norway spruce dominated. Spruce undergrowth is formed seldom under a spruce forest unless the stand is thin or has openings. Because Norway spruce is often rare in the mineral soil sites, the undergrowth is often regenerated from seeds that spread from spruce swamps. Earlier practiced shifting cultivation and its frequent fires prevented regeneration of spruce undergrowth. Similarly, the common felling method used, clear felling in strips, does not promote spruce undergrowth. Consequently, their occurrence is likely to decrease in the future.

The PDF includes a summary in German.

• Pöntynen, E-mail: vp@mm.unknown

After critically reviewing earlier studies on soil properties and their influence on forest growth and yield, it seems that defining the forest yield could be possible by means of soil properties. To be able to do so, the site needs to be defined and delineated in some other way. It is also necessary to decide the right soil properties to study for the purpose.

For the classification of forest sites the results of soil analyses need to be compared with growth and yield data from the site. To further the practice of classification of forest sites by means of soil studies, four aspects need to be taken into account:

1)  the site needs to be delineated beforehand according its vegetation, preferable with Cajander’s forest type classification

2) the experiments about soil needs to be done for as many properties as possible

3) the studied sites need to be as representative as possible in their class

4) there are as many samples for one site as possible studied

The volume 34 of Acta Forestalia Fennica is a jubileum publication of professor Aimo Kaarlo Cajander.

• Aaltonen, E-mail: va@mm.unknown

Forest site type classification based on the vegetation has not been developed in India. The classifications made by forest officers have been based on the upper storeys of trees. Shrubs have been used to class such sites where grasses are the dominant species. However, some observers in India have used grass and bamboo species to identify sites suitable on unsuitable for certain valuable tree species. In Burma, some bamboo species have been noticed to be good indicators for sites suitable or unsuitable for teak (Tectona grandis L. f.). Studies in the western sub-Himalayan area suggest that certain grasses could be used as indicators for sites suitable for sal (Shorea robusta Gaertn.). Grasses have also been identified as indicators for certain kinds of forests and soils in the area between Ganges and the Jumna.

The volume 34 of Acta Forestalia Fennica is a jubileum publication of professor Aimo Kaarlo Cajander.

• Osmaston, E-mail: ao@mm.unknown

The article is a congratulatory letter to professor Cajander. The author describes the work of Cajander about forest site classification and its importance to the development of forest sciences not only in Finland but worldwide.

The volume 34 of Acta Forestalia Fennica is a jubileum publication of professor Aimo Kaarlo Cajander.

• Wappes, E-mail: lw@mm.unknown

The vegetation in the Pacific coasts of British Columbia and Southeastern Alaska resemble the vegetation in the northern Fennoscandia. The national forests have been divided in two parts: Tongass and Chugach national forests. Both of the forests are fairly uniform in their vegetation. The forests have few coniferous tree species as the dominant species from south to north, Sitka spruce (Picea sitchensis (Bong.) Carrière) and western hemloch (Tsuga heterophylla (Raf.) Sarg.). It is difficult to distinguish forest site types, but it is probable that the forest lands in Alaska and British Columbia could be delimited to similar forest site types as professor Cajander established in Finland.

The volume 34 of Acta Forestalia Fennica is a jubileum publication of professor Aimo Kaarlo Cajander.

• Harshberger, E-mail: jh@mm.unknown

The primeval forests of Scotland were deforested in the Lowlands by the end of 1500th century, and in the end of the 1800th century also the best forests of the more inaccessible Highlands were exploited. The 1800th century witnessed an outburst of afforestation among the private land owners. With help of nursery work and use of exotic species, the work was successful. Silviculture of Scotland would benefit of a reliable method of site classification. The complexity of the geology and topography, and the lack of mature natural stands complicate the establish a forest type classification similar to the one Prof. Cajander has evolved in Finland. The aim is to establish forest site types which include similar types as in Finland, with possibly additional types in the grass-herb series.

Jubileum publication of professor Aimo Kaarlo Cajander.

• Anderson, E-mail: ma@mm.unknown

The root systems of 192 Scots pine (Pinus sylvestris L.) sample trees were dug out and measured in Säyneinen, Rautavaara and Pielijsäjvi in the Central Finland and Orivesi, Teisko and Hämeenkyrö in the Southern Finland. The volume of root system of Scots pine was always smaller than the stem, varying from 15% to 94% of the stem volume. The ratio is smaller in dense stands. The type of soil of the site affects how the central root system (tap root and the inner vertical roots) develop. This reflect the adaptability of the root system to different growth conditions. The root system may, for instance, substitute the tap root with stronger inner roots.

PDF includes a summary in English.

• Laitakari, E-mail: el@mm.unknown

The forest sites have typically been classified by two principles, either as stand quality classes or as locality (site) classes. This article describes the principles of Finnish forest site types (forest quality classes) which are based on classification of localities according to their forest plant associations. All the stands that belong to the same forest site type are characterized by a distinct, more or less identical plant species composition. The forest site types are independent of the tree species. The forest site types in a larger area are relatively numerous, but can be grouped according to their normal form. The Finnish forests are separated to dry moss forest class, the moist moss-forest forest class and grass-herb forest class. The different forest site types belonging to the classes are described in detail. Growth of the trees is different for the different forest site types, but varies little within a same site type. The forest site types suit therefore well for the purposes of forest mensuration and for yield tables. The forest site types reflect also the properties of the soil.

• Cajander, E-mail: ac@mm.unknown

The forest sites have typically been classified by two principles, either as stand quality classes or as locality (site) classes. This article describes the principles of Finnish forest site types (forest quality classes) which are based on classification of localities according to their forest plant associations. All the stands that belong to the same forest site type are characterized by a distinct, more or less identical plant species composition. The forest site types are independent of the tree species. The forest site types in a larger area are relatively numerous, but can be grouped according to their normal form. The Finnish forests are separated to dry moss forest class, the moist moss-forest forest class and grass-herb forest class. The different forest site types belonging to the classes are described in detail. Growth of the trees is different for the different forest site types, but varies little within a same site type. The forest site types suit therefore well for the purposes of forest mensuration and for yield tables. The forest site types reflect also the properties of the soil.

• Cajander, E-mail: ac@mm.unknown

The study is based on the results of the soil studies by Valmari (1921) and the growth inventories of respective areas. The aim is to show the connection of soil fertility (nutrient content) and forest growth with means of correlation calculations. The examined nutrients were nitrogen, calcium and phosphorus, also the electrolyte content was studied.

• Ilvessalo, E-mail: yi@mm.unknown

The vegetation of the forest and peatland site types in Northern Finland differ markedly from those in Southern Finland, also the vegetation of the subtypes in the north is distinctive. A line survey was conducted to study the distribution of forest and peatland site subtypes in Northern Finland.

The vegetation of rich grass-herb forest types differs little from the poorer grass-herb forest types in Northern Finland. They abundance decrease towards north. The main fresh mineral soil sites are Myrtillus site type, Hylocomnium-Myrtillus site type and their paludified forms. The abundance of the fresh mineral soil sites decreased towards north so that in Kemi the proportion was 20.5 and in Lapland 12.0%. A transition from the fresh to the drier site types is gradual. The Vaccinium site type that is dominant in the south, is rare in the north, where it is replaced by Empetrum-Vaccinium type. The proportion of dry forest sites increase towards north, in average their distribution is 25% of the lands. There are numerous subtypes, which can be merged in to four main site types: Calluna, Cladina-Calluna, Myrtillus-Cladina and Cladina site type. The peatlands are more abundant in the southern part of the study area. The most common peatland types are pine swamps.

The PDF includes a summary in German.

• Multamäki, E-mail: sm@mm.unknown

The article contains tree lectures given in the meeting of the Geographical Society in Finland on February 25^th 1921.  The titles of the lectures are I Forest types in general, II Forest types as a basis for new growth and yield tables in Finland, and III Other research on forest types.  

The first lecture is a follow-up of the Cajander’s 1909 published article on forest types. It deepens the theory on forest types. The classification into forest types represents primarily different plant communities of ground cover. The types are named after the characteristic plant species, indicator plants, however, many other species appear in different abundance.

The second lecture represents the research proceedings of mensuration of forest stands of different types to compile yield tables for pine.  The forest types differ from each other distinctly on their growing preconditions, but inside one class the variation of the growing conditions is so small, that the classification can be used for yield tables, determining the basis of taxation and for classification of forest based on height over age.

The third lecture is a summary of other studies about forest type classification.  They confirm the results presented in earlier lectures. 

• Cajander, E-mail: ac@mm.unknown
• Ilvessalo, E-mail: yi@mm.unknown

• Kujala, E-mail: vk@mm.unknown

The first proper growth and yield tables were prepared in Finland already in 1872, but they have been used little as the needs of forestry and forest sciences increased. One of the problems of the old yield tables was how the site quality classes are determined. The new growth and yield tables use the forest site type classification, which enables the use of same site types for all tree species. This makes it possible to compare the growth of different tree species in same kind of sites. The tables also use stem frequency distribution series. In the first stage, the tables were prepared for Southern and Central Finland.

The PDF includes a summary in German.

• Ilvessalo, E-mail: yi@mm.unknown

New growth and yield tables were prepared for Southern Finland. To finalize the tables, it had to be determined whether the forest site types developed by Cajanus could be used in mensurational research.

Comparative study was performed in 1916-1919 to study the growth of the trees in different forest site types. Total of 467 sample sites were measured in Southern and Central Finland. All the forest site types were found to have a distinctive vegetation typical to the site. It can be concluded that the ground vegetation can be used to determine the forest site type. The growth of trees was different in different forest site types, yet similar within each site type. The forest site types are uniform, natural and easy to determine, and can thus be used to classify the forest stands and used in mensurational research and a basis to growth and yield tables.

The PDF includes a summary in German.

• Ilvessalo, E-mail: yi@mm.unknown

The forest site classification system used in Finland is based on ground vegetation rather than the wood production capacity. A. K. Cajander has presented a detailed classification of different forest site classes in different parts of the country. This study focuses on the forest site types of Northern Finland, which are less well defined. The article presents detailed vegetation analysis and lists of plant species in different forest site types in Northern Finland. In contrast to southern parts of Finland, both the natural Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst) stands are uneven-aged in the north. The forests are also relatively thin. There is a marked difference in height of trees between the richest and poorest sites, but the dominant trees of the same site type were of similar height both in the north and south part of the study area. The differences in the height of dominant trees seem to be smaller than in Southern Finland. Also, in windy areas prone to snow damage, climate conditions can affect tree growth more than the forest site type. In more sheltered areas forest site type determines forest growth.

The PDF includes a summary in German.

• Lakari, E-mail: ol@mm.unknown

A strip survey was made to define the forest and peatland site class distribution and the condition of the forests in Savo and Karelia in central and eastern parts of Finland. According to the survey, 24% of the forested lands are peatlands. Fresh mineral soil sites (26%) were the most common mineral soil site type. Intermediately dry forest soil sites covered 22% of the area, forest sites with grass-herb vegetation 12,79%, rich grass-herb forest soil sites 3,16% and dry forest soil sites 9,59% of the forested area. The most common tree species were Scots pine (Pinus sylvestris L.), 39%, Betula sp., 26%, Norway spruce (Picea abies (L.) H. Karst.), 18%, and grey alder (Alnus incana (L.) Moench), 10% of the forest land. The article includes a review about the wood harvesting in the forests, and their present silvicultural state. According to the study, about 30% of the forested lands (not including peatlands) were unproductive; mostly mixed alder and birch stands of poor quality or open lands.

The PDF includes a summary in German.

• Multamäki, E-mail: sm@mm.unknown

The article presents a survey on distribution of fertile lands, soil types and site classes in Savo and Karelia in the central and eastern parts of Finland. The survey was based both in existing publications and statistics, a line survey, and visual observations during field trips. The site quality classification is based on the vegetation and occurrence of indicator plant species. The article lists distribution of indicator species in different forest site types on maps of the area. In addition, a review of history of land use and agriculture give indications of the location of the fertile lands in the area. A map of the forest site types in different parts of the area illustrate the data collected from the different sources.

The PDF includes a summary in German.

• Lukkala, E-mail: ol@mm.unknown

The data has been collected in spruce forests in mountainous areas of Germany: Fichtelgebirge and Böhmerwald in Bavaria, Erzgebirge in Saxonia. The studied characteristics of the stand were: growth of the trees in height and diameter, and the ground vegetation. The stands were classified according Cajander’s forest site classification. The article presents the most common plants and other characteristics of every forest site type and studied stands. The relation of the height of the trees and their age is represented in diagrams for every forest type.

The presence of indicator plants is somewhat dependent on the stand age and crown coverage. The amount of species is lowest when the crown coverage is at the greatest.

As conclusion of the study it can be seen that the growth of the stand differentiates clearly depending on the forest site type, being greater at the more nutritious sites. Since the differently growing stands need different management, it would be natural to direct the management of the stand according the forest site type. 

• Björkenheim, E-mail: rb@mm.unknown

Tree growth is one of the factors that have been used to determine the site quality. The aim of the study was to show that growth of single trees growing on a same forest site class are similar, but differ from trees growing on a different site type. To compare the tree growth, a stem analysis was performed to dominant trees in Scots pine (Pinus sylvestris L.) stands, measured in 15 Myrtillus type sample plots and in 15 Calluna type sample plots in state forests in Salmi, situated in north side of Lake Ladoga. The height growth when the tree was young was higher in the trees growing in the Myrtillus type than in the Calluna type. Also, the trees of same age are higher in Myrtillus type stand than in the Calluna type. In Calluna type, the height growth, however, evens out later in age than in the Myrtillus type. The volume growth of the trees begins to increase earlier in Myrtillus type, and is higher than in Calluna type. Similarly, the diameter growth in breast height is higher in the Myrtillus type.

The PDF includes a summary in German.

• Ilvessalo, E-mail: yi@mm.unknown

The study is based on research in Germany, Austria, Switzerland, north Russia and Siberia, and Finland in years 1906-1908. The objective of the study is to find means to create forest site classes or forest types to direct practical forest management.

The article presents the classification of forests into site classes (Oxalis-Majanthemum type, Myrtillus type, Vaccinium type and Calluna type). The second part of the article represents different methods to calculate growth and yield tables for different forest site types. The conclusion of the study is that forest areas with similar vegetation and forest type can be handled in one way for forest management. 

• Cajander, E-mail: ac@mm.unknown

The relationship between site characteristics and understorey vegetation composition was analysed with quantitative methods, especially from the viewpoint of site quality estimation. Theoretical models were applied to an empirical data set collected from the upland forests of Southern Finland comprising 104 sites dominated by Scots pine (Pinus sylvestris. L.) and 165 sites dominated by Norway spruce (Picea abies (L.) H. Karst.). Site index H₁₀₀ was used as an independent measure of site quality.

A new model for the estimation of site quality at sites with a known understorey vegetation composition was introduced. It is based on the application of Bayes’ theorem to the density function of site quality within the study area combined with the species-specific presence-absence response curves. The resulting probability density function may be used for calculating an estimate for the site variable

Using this method, a jackknife estimate of site index H₁₀₀ was calculated separately for pine- and spruce-dominated sites. The results indicated that the cross-validation root mean squared error (RMSE_cv) of the estimates improved from 2.98 m down to 2.34 m relative to the ”null” model (standard deviation of the sample distribution) in pine-dominated forests. In spruce-dominated forests RMSE_cv decreased from 3.94 m down to 3.19 m.

In order to assess these results, four other estimation methods based on understorey vegetation composition were applied to the same data set. The results showed that none of the methods was clearly superior to the others. In pine-dominated forests RMSE_cv varied between 2.34 and 2.47 m, and the corresponding range for spruce-dominated forest was from 3.13 to 3.57 m.

• Lahti, E-mail: tl@mm.unknown

Two operative forest site class estimation methods utilizing satellite images have been developed for forest income taxation purposes. For this, two pixelwise classification methods and two post-processing methods for estimating forest site fertility are compared using different input data. The pixelwise methods are discriminant analysis, based on generalized squared distances, and logistic regression analysis. The results of pixelwise classifications are improved either with mode filtering within forest stands or assuming a Markov random field type dependence between pixels. The stand delineation is obtained by using ordinary segmentation techniques. Optionally, known stand boundaries given by the interpreter can be applied. The spectral values of images are corrected using a digital elevation model of the terrain. Some textural features are preliminary tested in classification. All methods are justified by using independent test data.

A test of the practical methods was carried out and a cost-benefit analysis computed. The estimated cost saving in site quality classification varies from 14% to 35% depending on the distribution of the site classes of the area. This means a saving of about 2.0–4.5 million FMK per year in site fertility classification for income taxation purposes. The cost savings would rise even to 60% if that version of the method were chosen where field checking is totally omitted. The classification accuracy at the forest holding level would still be similar to that of traditional method.

The PDF includes a summary in Finnish.

• Tomppo, E-mail: et@mm.unknown

Mineral soil sites where Scots pine (Pinus sylvestris L.) were suffering from Gremmeniella abietina die-back (Lagerb.) M. Morelet. were characterized and classified in Central Finland. The tree stand, ground vegetation, soil type and site topography were described in 163 sample plots in 16 stands. The sites were classified according to system developed by Cajander and numerically using TWINSPAN analysis based on the ground vegetation. The site topography of severely damaged stands was checked from colour infrared aerial photographs. The disease was most severe in depressions and frost pockets. Apart from topography no significant correlations were found between disease severity and site factors. No typical vegetational pattern of forest type of the severely affected stands could be detected. Most of the stands were growing on medium-coarse, unfertile soil with a rather thick humus layer.

The PDF includes a summary in Finnish.

• Sairanen, E-mail: as@mm.unknown

An extensive field-based survey was conducted to establish the distribution of site types on drained peatlands, the condition of the drainage networks, the post-drainage development of the tree stands, their structure and silvicultural condition and the corresponding requirements for operational measures. The data is based on sampling of the forest drainage undertaking during 1930–78 and consists of 1,312 km inventory transect, 6,030 relascope sample plots and 21,700 studied ditches.

Of the studied peatlands more than 60% were Scots pine (Pinus sylvestris L.) mires, slightly under 20% Norway spruce (Picea abies) mires, and under 10% each treeless mires and paludified upland forest sites. The remaining peatland area that is to be considered suitable for forest drainage according to criteria used by Heikurainen (1960) now consists mainly of spruce mires and paludified upland forest types; about 1 million ha both groups still remain undrained.

The proportion of ditches in need of ditch cleaning was estimated to be under 10% in the youngest drained areas and under 30% in the oldest. The mean tree stand volumes of the drained peatlands of different site types show the same dependence on the trophic level as in earlier studies but the volumes seem to be some 5–10% lower. These results compare favourably with those of the 7th national forest inventory.

Trends in the post-drainage development of tree stand volumes and increment are also, generally, in accordance with earlier findings but have somewhat lower values. The development of the nutrient-poor site type stands, especially in Northern Finland, seems to be significantly poorer than was earlier assumed.

The PDF includes a summary in English.

• Keltikangas, E-mail: mk@mm.unknown
• Laine, E-mail: jl@mm.unknown
• Puttonen, E-mail: pp@mm.unknown
• Seppälä, E-mail: ks@mm.unknown

The vegetation and number of physical and chemical soil properties were studied on a random sample of closed upland forest stands in Southern Finland. The material consists of a total of 410 sample plots. Two-way indicator species analysis (TWINSPAN) was carried out in order to produce a hierarchical clustering of samples on the basis of the vegetation data. Discriminant analysis and analysis of variance were applied in order to find environmental correlations of the vegetation clustering.

The vegetation was found to indicate the nutrient regime of the humus layer well, but to a less extent the properties of the sub-soil. The understorey vegetation was found to be jointly dependent on the site fertility and on the properties of the tree stand, especially on the tree species composition. Although the forest vegetation appears to be distributed rather continuously along an axis of increasing site fertility, relatively unambiguous classification can be based on the appearance of indicator species and species groups.

The results of the study were interpreted as indication that operational site classification done using the vegetation is rather good method for classification in closed forest stands. Different methods produce relatively consistent, natural and ecologically interpretable classifications. The results also imply that the vegetation cover and the humus layer develop concurrently during the development of the ecosystem, but the differentiation of the site type is regulated simultaneously by a number of interacting factors ranging from mineralogical properties of the parent material to the topographical exposition of the site. As the plant cover depicts all these primary factors simultaneously, only a relatively rough ecological site classification can be based on the vegetation.

The PDF includes a summary in Finnish.

• Kuusipalo, E-mail: jk@mm.unknown

The aims of the present study were to determine physical and physio-chemical properties of some Finnish forest tree nursery soils, and to examine relationships between these properties and the amount of organic matter in the soil.

The following soil tillage layer properties of 33 fields belonging to 8 forest tree nurseries were determined: soil particle size distribution, organic matter content, bulk density and density of solids, total pore space, soil water volume at potentials pF 2.0 and 4.2, available water content and air space at potential pF 2.0, active acidity, electrical conductivity index and cation exchange capacities at pH 4.5 and 8.0. The soil texture class of the tillage layer parent material was sand, only in a few cases did higher percentage of silt and clay indicate a morainic nature of parent material. The amount of organic material in the soils varied within wide limits, reflecting differences in amelioration policy between the single nurseries.

Relationships between the physical properties of the soil parent material and those related to fertility were in most cases strongly influenced by the amount of soil organic matter. Soil density values decreased as the organic matter content increased from 2 to 25%, giving rise to the increase in the total pore space. However, the amount of water held at potential pF 2.0 and the available water content did not increase with increasing organic matter content. This was due to the absence of the particle fraction in the sand. Nursery soil amelioration, involving in most cases a mixture of Sphagnum peat with sand, thus gives rise to an increase in the content of drainable water.

Cation exchange capacities were positively correlated with the organic matter content. However, the absolute number of exchange sites expressed as equivalents in the tillage layer did not increase in accordance with the increase in organic matter content due to the influence of the organic matter content upon the ratio of solids in the voids.

The PDF includes a summary in English.

• Westman, E-mail: cw@mm.unknown

Fertility of surface peat from sedge pine mires was studied by measuring several edaphic growth factors: bulk density, volume weight of organic matter, ash content, acidity, electric conductivity, effective cation exchange capacity, degree of base saturation, and total contents of N, P, K, Ca and Mg. The 168 temporal sample plots were situated on virgin sedge mires in different parts of Finland, and the 30 permanent sample plots on two uniform sedge mires.

The results showed that peat bulk density and volume weight of organic matter tend to increase with increasing site quality. Ash content increased gradually in the site series from small sedge mire to the herb-rich sedge mire. The relationship between the total content of macronutrients in peat and the site quality is clear. The importance of bulk density in evaluating the site quality is further emphasized when taking into account its significant correlation to contents of N and P. The soil variables follow the accepted quality gradient of the site series. Consequently, the plant sociologically based site classification seems to reflect satisfactorily the average soil properties. However, the within site variation was significant.

• Westman, E-mail: cw@mm.unknown

The data used to control the developed methods are presented in chapter two. The third chapter presents the statistical analysis conducted. The factor analysis shows that tree stand characteristics must be divided into three factors to able the information needed for site indexing to distinguish different site classes. The phases of model development are presented and the results of calculations with control data are shown.   

The results indicate that for calculations of nature-normal forests the suitable amount of tree stand characters for creating the site indexing function, is three.  These are: number of trees (N), basal area of the stand (G) and mean height (H).

The PDF contains a summary in Finnish. 

• Kangas, E-mail: yk@mm.unknown

The article is the first part of a wider project aiming to study the measurement practices of stand’s height-over-age based classifications and to develop new methods for actually measuring the side index.  

The first part of the article presents the concepts of site indexing and discusses it with a mathematical model. The second part of the article examines the structure of site indexing system regarding the reliability of the method. Third part presents the factors affecting the method development and the fourth part discusses the possible site index classes and their characteristics.   

The PDF contains a summary in Finnish. 

• Kangas, E-mail: yk@mm.unknown

• López, Inland Norway University of Applied Sciences, Faculty of Applied Ecology, Agricultural Sciences and Biotechnology, P.O. Box 2400, Koppang, Norway https://orcid.org/0009-0006-6860-3408 E-mail: lucas.lopez@inn.no
• Sjølie, Inland Norway University of Applied Sciences, Faculty of Applied Ecology, Agricultural Sciences and Biotechnology, P.O. Box 2400, Koppang, Norway https://orcid.org/0000-0001-8099-3521 E-mail: hanne.sjolie@inn.no
• Nabhani, Inland Norway University of Applied Sciences, Faculty of Applied Ecology, Agricultural Sciences and Biotechnology, P.O. Box 2400, Koppang, Norway E-mail: abbas.nabhani@inn.no
• Aguilar, Swedish University of Agricultural Sciences, Department of Forest Economics, SE-901 83 Umeå, Sweden E-mail: francisco.aguilar@slu.se

The poor cost-effectiveness of mechanized planting (MECP) is the main reason for the low mechanization rate of planting. In this study, we investigated the productivity of the mechanized excavator-based planting of Norway spruce (Picea abies [L.] H. Karst.) seedlings based on data collected by the Risutec Asta documentation system. We also compared the costs of a MECP chain with two different manual planting (MAP) chains, where mounding was carried out by a crawler excavator (EXC) or a continuously advancing mounder (CONT). The MECP of seedlings was carried out using an EXC equipped with a Risutec PM-160 planting device. Generally, the nine study sites in western Finland contained few surface obstacles (e.g., the logging residues had mainly been harvested), which made the conditions very suitable for MECP. The average production time taken by the MECP was 9 h ha^-1. The operating hour (G₁₅-h) productivity averaged 215 seedlings G₁₅-h^-1 , with the mean planting time being 13.8 s seedling^-1. Loading 160 seedlings into the seedling cassette took approximately 10 min (3.8 s seedling^-1). Overall, the cost of the MECP was about 5% lower than for the EXC + MAP. However, when productivity was set at <200 seedlings G₁₅-h^-1 , the cost of the MECP was higher than that of the EXC + MAP. Based on our findings, the most cost-efficient planting chain was CONT + MAP. However, based on our results, the required level of productivity can be achieved if the sites are suitable for MECP and the machine operators are skilled.

• Kemppainen, School of Forest Sciences, University of Eastern Finland (UEF), P.O. Box 111, FI-80101 Joensuu, Finland; Natural Resources Institute Finland (Luke), Yliopistokatu 6, FI-80100 Joensuu, Finland https://orcid.org/0009-0000-6184-8812 E-mail: kalle.kemppainen@uef.fi
• Kärhä, School of Forest Sciences, University of Eastern Finland (UEF), P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0000-0002-8455-2974 E-mail: kalle.karha@uef.fi
• Laitila, Natural Resources Institute Finland (Luke), Yliopistokatu 6, FI-80100 Joensuu, Finland https://orcid.org/0000-0003-4431-3319 E-mail: juha.laitila@luke.fi
• Sairanen, School of Forest Sciences, University of Eastern Finland (UEF), P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0009-0008-8632-3797 E-mail: anttsair@student.uef.fi
• Kankaanhuhta, Natural Resources Institute Finland (Luke), Yliopistokatu 6, FI-80100 Joensuu, Finland https://orcid.org/0000-0001-5785-5972 E-mail: ville.kankaanhuhta@luke.fi
• Viiri, UPM-Kymmene Plc, UPM Forest, Peltokatu 26 C 4, FI-33100 Tampere, Finland https://orcid.org/0000-0003-3952-9481 E-mail: heli.viiri@upm.com
• Peltola, School of Forest Sciences, University of Eastern Finland (UEF), P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0000-0003-1384-9153 E-mail: heli.peltola@uef.fi

• Nordin, pcSKOG, Grisslevägen 15, 227 32 Lund, Sweden https://orcid.org/0000-0002-2156-6615 E-mail: per.nordin@skogforsk.se
• Olofsson, Linnaeus University, Department of Forestry and Wood Technology, 351 95 Växjö, Sweden https://orcid.org/0000-0001-5844-6775 E-mail: erika.olofsson@lnu.se
• Hjelm, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, Box 190, 234 22 Lomma, Sweden https://orcid.org/0000-0002-6144-8250 E-mail: karin.hjelm@slu.se

Mechanical site preparation (MSP) is a common practice that precedes the planting of Norway spruce (Picea abies (L.) H. Karst.) in Nordic forests. Mounding has become the most used method in spruce planting in recent years. This study examined the effects of different mounding treatments (spot and ditch mounding, inversion, unprepared control with or without herbicide application) and a mechanical vegetation control (MVC) treatment done 3–4 years after planting on the post-planting growth of spruce container seedlings and their development to saplings during the first 11–13 years on two forest till soils in central Finland, one on flat terrain and other on a southwest slope. On these fine-textured soils the spot and ditch mounding methods favoured spruce saplings development. Inversion and unprepared plots showed weakest growth. On the site with flat terrain, 11 years post planting, spruce saplings were 78–144 cm (38–80%) taller and their breast height diameters were 11–13 mm (60–74%) thicker for ditch or spot mounding than for inversion or herbicide treatment. On the site with sloped terrain the differences were minor between the MSP treatments. MVC improved spruce height growth on sites which did not have intensive MSP, especially on control saplings planted on unprepared soil in herbicide and inversion treatments. On the flat terrain, MVC reduced the density of resprouts to be removed later in pre-commercial thinning. As a conclusion, spot or ditch mounding favoured the growth of spruce over inversion especially on flat terrain with fine-textured soil.

• Uotila, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: karri.uotila@luke.fi
• Luoranen, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland https://orcid.org/0000-0002-6970-2030 E-mail: jaana.luoranen@luke.fi
• Saksa, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland https://orcid.org/0000-0002-1776-2357 E-mail: timo.saksa@luke.fi
• Laine, Metsä Forest, Revontulenpuisto 2, FI-02100 Espoo, Finland https://orcid.org/0000-0002-6448-8274 E-mail: tiina.laine@metsagroup.com
• Heiskanen, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland https://orcid.org/0000-0002-0549-835X E-mail: juha.heiskanen@luke.fi

Over 20% of regeneration operations will be on drained peatland in the next decade in Finland. There are only a few studies comparing the planting success and the risk of pine weevil (Hylobius abetis (L.) feeding damage on mineral soil and drained peatland. Thirty sites planted with Norway spruce (Picea abies (L.) H. Karst.) container seedlings in 2009 in Southern and Central Finland were inventoried three growing seasons after planting. Prediction models for the probability of survival, pine weevil damage and the presence of ground vegetation cover were done separately for peatland and mineral soil sites. The planting success was 17% lower on peatland sites (1379 surviving seedlings ha^–1) than on mineral soil (1654 seedlings ha^–1). The factors explaining the survival were the ground vegetation cover and type of the planting spot on the peatland, and the ground vegetation cover on mineral soil. On mineral soil, 76% of the planting spots were on cultivated mineral soil while on peatland only 28% of the seedlings were planted on similar spots. There were also fewer seedlings that were surrounded by dense ground vegetation on mineral soil (4%) than on peatland (14%). Pine weevil feeding damage did not differ significantly on peatland (23%) or mineral soil (18%). The more time there was from clear-cutting, the more the probability of pine weevil feeding damage was reduced on both soil classes. Additionally, cover vegetation in the vicinity of the seedlings increased on mineral soil. Cultivated planting spots, especially those covered by mineral soil, prevented pine weevil feeding and reduced the harmful effects of vegetation on the seedlings both on mineral soil and peatland.

• Luoranen, Natural Recourses Institute Finland (Luke), Production systems, Juntintie 154, FI-77600 Suonenjoki, Finland https://orcid.org/0000-0002-6970-2030 E-mail: jaana.luoranen@luke.fi
• Viiri, UPM Forest, Peltokatu 26 C, P.O. Box 85, FI-33100 Tampere, Finland https://orcid.org/0000-0003-3952-9481 E-mail: heli.viiri@upm.com

In Nordic forests, consistent evidence about better seedling survival rate and increased growth due to site preparation have been obtained in numerous studies. Proper site preparation method can reduce costs of the whole regeneration chain through its effects on survival of planted seedlings, abundance of natural regeneration and competition in early stand development. This study compared the natural regeneration of birches (silver birch (Betula pendula Roth) and downy birch (B. pubescens Ehrh.)), amount of exposed mineral soil, and growth of planted seedlings between spot mounding and inverting site preparation methods. Present study was conducted in eight forest stands established in 2012 or 2015. Even though difference was not statistically significant, inverting exposed less mineral soil than spot mounding and thus reduced the natural regeneration of birch seedlings by 6135 seedlings ha^–1 compared to spot mounding. However, the variation between regeneration areas was remarkable. There was no difference in seedling mortality or growth between the site preparation methods. In order to achieve high growth of conifers, moderate amount of exposed mineral soil and thus less naturally regenerated birch, inverting should be favored over spot mounding.

• Laine, Metsä Group, P.O. Box 208, FI-70101 Kuopio, Finland E-mail: tiina.laine@metsagroup.com
• Kankaanhuhta, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: ville.kankaanhuhta@luke.fi
• Rantala, Metsä Group, P.O. Box 10, FI-02020 METSÄ, Finland E-mail: juho.rantala@metsagroup.com
• Saksa, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@luke.fi

Hybrid aspen (Populus tremula L. × P. tremuloides Michx.) is a deciduous tree species suitable for producing large amounts of renewable biomass during short rotations. Its potential under North European conditions could be largely extended if not only agricultural land but also forest land was used for cultivation. Unfortunately, the knowledge of appropriate forest site conditions and effects of site preparation methods on hybrid aspen establishment is limited. In this paper, two studies that explore these questions are presented. In the first study, the sensitivity to acid soils was tested under greenhouse conditions in two type of soils: a) peat soil limed to certain pH levels (3.4–5.7) and b) collected forest soils where pH varied from 3.9 to 5.3. The lowest pH level resulted in reduced growth, elsewhere no significant differences were found. The second study was applied in the field and investigated the effect of four site preparation methods on survival and growth. The methods were: 1) control with no site preparation, 2) patch scarification, 3) mounding and 4) soil inversion. While no differences were found for survival, mounding was generally the method with the highest growth and patch scarification was least successful. The result was probably an effect of good soil aeration and less competition from vegetation after mounding. The field study also revealed clonal differences in growth performance, which stresses the importance of clone selection prior to planting. The results of these studies indicate that hybrid aspen is less sensitive to variation in pH and site preparation methods compared with other poplar species, as have been found in similar studies.

• Hjelm, Skogforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden E-mail: karin.hjelm@skogforsk.se
• Rytter, Skogforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden http://orcid.org/0000-0001-6183-4832 E-mail: lars.rytter@skogforsk.se

Physical soil properties have a marked influence on the quality of forest sites and on the preconditions for forest growth and management. In this study, water retention characteristics (WRC) and related physical soil properties in addition to vegetation coverage and tree stand data were studied at upland forest sites in Finland. Fixed and mixed models between soil and site characteristics were formed to estimate physical and hydrologic soil characteristics and the site quality with indirect co-varying variables. In the present data, the site quality index (H100) shows a high coefficient of determination in respect to the temperature sum. It is also related to soil fine fraction content, topsoil pH and water retention at field capacity. The thickness of the humus layer is predictable from the pH and cover of xeric and mesic plant species. The soil fine fraction content (clay + silt) is closely related to water retention at field capacity, the soil layer and site type, and without WRC to the temperature sum and site index and type, as well as the slope angle. The soil bulk density is related to organic matter, depth (layer) or alternatively to organic matter, slope and field estimated textural class (fine, medium, coarse). Water retention characteristics were found to be best determinable by the fine fraction content, depth and bulk density. Water content and air-filled porosity at field capacity are closely related to the fine fraction. This study provides novel models for further investigations that aim at improved prediction models for forest growth, hydrology and trafficability.

• Heiskanen, Natural Resources Institute Finland (Luke), Soil ecosystems, Neulaniementie 5, FI-70100 Kuopio, Finland E-mail: juha.heiskanen@luke.fi
• Hallikainen, Natural Resources Institute Finland (Luke), Applied statistical methods, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: ville.hallikainen@luke.fi
• Uusitalo, Natural Resources Institute Finland (Luke), Forest technology and logistics, Korkeakoulunkatu 7, FI-33720 Tampere, Finland E-mail: jori.uusitalo@luke.fi
• Ilvesniemi, Natural Resources Institute Finland (Luke), Biorefinery and bioproducts, Tietotie 2, FI-02150 Espoo, Finland E-mail: hannu.ilvesniemi@luke.fi

Different environmental factors were studied to determine which factors influence the species richness, composition and structure of vascular plants in Pinus sylvestris L. forests in a fixed dune landscape in south-western Estonia. In addition to site topographic factors, different environmental parameters were investigated. Thirty-four vascular plant species were recorded in 232 quadrats. The most abundant species was Vaccinium vitis-idaea L., which was in 82.8% of quadrats, followed by Vaccinium myrtillus L. (74.1%), Melampyrum pratense L. (71.1%) and Deschampsia flexuosa (L.) Trin. (69.8%). The multiple response permutation procedure (MRPP) showed considerable differences in species composition at the bottoms of dunes compared with that on the slopes and at the tops of dunes. Indicator species analysis (ISA) determined species exhibited characteristics specific to zone: V. myrtillus had the highest indicator value at the bottoms of dunes; Calluna vulgaris L., at the tops. Soils were Haplic Podzols, and the presence of humus horizon depended on zone. Soil conditions on the dunes were variable and site specific, in general, soils at the bottoms of the dunes were more acidic and moist compared with those of the slopes and tops of the dunes, and the nutrient content decreased toward the dune tops. According to non-metric multidimensional scaling (NMDS) and linear mixed model analyses, species coverage, composition and richness were controlled by site-specific factors such as absolute height, location and aspect of the quadrat on the dune; soil nitrogen, potassium and phosphorus contents; soil pH and moisture; light conditions; and the thickness of the litter horizon.

• Tilk, Department of Silviculture, Estonian University of Life Sciences, Kreutzwaldi 5, Tartu, Estonia, 51014; Tallinn Botanic Garden, Kloostrimetsa Road 52, Tallinn, Estonia, 11913 E-mail: Mari.Tilk@botaanikaaed.ee
• Tullus, Department of Silviculture, Estonian University of Life Sciences, Kreutzwaldi 5, Tartu, Estonia, 51014 E-mail: Tea.Tullus@emu.ee
• Ots, Department of Silviculture, Estonian University of Life Sciences, Kreutzwaldi 5, Tartu, Estonia, 51014 E-mail: Katri.Ots@emu.ee

To promote the growth and survival of regenerated forests, site preparation prior to tree planting on clearcuts is necessary. This is often performed with scarifiers, either through trenching or mounding. Mounding is generally considered better in a plant survival perspective but is inefficient on obstacle-rich clearcuts. By utilising machine vision through e.g. remote sensing methods, new strategies can enable efficient mound positioning. In this paper, three realistic strategies utilizing ideal clearcut object identification through machine vision have been developed that can be used for more efficient mounding. The results show that mounding efficiency can be significantly improved with a new mound positioning strategy that employs ideal object identification, especially on obstacle-rich clearcuts.

• Lideskog, Luleå University of Technology, Department of Engineering Sciences and Mathematics, Division of Product and Production Development, SE-971 87 Luleå, Sweden E-mail: hakan.lideskog@ltu.se
• Karlberg, Luleå University of Technology, Department of Engineering Sciences and Mathematics, Division of Product and Production Development, SE-971 87 Luleå, Sweden E-mail: magnus.karlberg@ltu.se

Stump wood is a possible alternative to fossil fuel. Its harvesting, however, disturbs the ground and this has not yet been quantified at stump level. Such disturbance is likely to be dependent on stump size, type of soil and timing of stump harvesting. Therefore, we measured ground disturbance and root breakage diameter at two Norway spruce sites with sandy glacial till soil. The sites were harvested with a fork type head, 6 and 18 months after clear cutting. Measurements were made within 2 weeks of harvest. No difference was found between the two sites. The mean area of disturbed ground was 6.06 (std 3.14) m² per stump and increased exponentially with stump size. A regression function modelling the relationship was constructed. Unexpectedly, many fine roots where extracted in the harvest. The arithmetic and basal area weighted mean root breakage diameter was 4.6 (std 2.2) and 29.5 (std 17.9) mm, respectively. There seems to be a limited increase in root breakage diameter with increased stump size. The small root breakage diameter is associated with reduced fuel quality and greater nutrient removal. It appears that much of the ground disturbance is associated with the creation of ruts rather than stump harvest per se. Stump harvesting disturbs a larger percentage of the area of a harvested site than mounding. Postponing stump harvest by one year did not decrease the ground disturbance or increase the root breakage diameter. To achieve less disturbance and larger root breakage diameter, probably new stump harvesting technology is required.

• Berg, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: phd.simon.berg@gmail.com
• Nordfjell, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: tomas.nordfjell@slu.se
• Bergström, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: dan.bergstrom@slu.se

• Hyppönen, Finnish Forest Research Institute, Rovaniemi Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: mikko.hypponen@metla.fi
• Hallikainen, Finnish Forest Research Institute, Rovaniemi Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: ville.hallikainen@metla.fi
• Niemelä, Finnish Forest Research Institute, Rovaniemi Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: jn@nn.fi
• Rautio, Finnish Forest Research Institute, Rovaniemi Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: pasi.rautio@metla.fi

• Hökkä, Finnish Forest Research Institute, Rovaniemi Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: hannu.hokka@metla.fi
• Repola, Finnish Forest Research Institute, Rovaniemi Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: jaakko.repola@metla.fi
• Moilanen, Finnish Forest Research Institute, Oulu Unit, P.O. Box 413, FI-90014 University of Oulu, Finland E-mail: mikko.moilanen@metla.fi
• Saarinen, Finnish Forest Research Institute, Parkano Unit, Kaironiementie 15, FI-39700 Parkano, Finland E-mail: markku.saarinen@metla.fi

• Meen, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, P.O. Box 5003, NO-1432 Ås, Norway E-mail: eivind.meen@umb.no
• Nielsen, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, P.O. Box 5003, NO-1432 Ås, Norway E-mail: an@nn.no
• Ohlson, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, P.O. Box 5003, NO-1432 Ås, Norway E-mail: mo@nn.no

• Nilsson, Swedish University of Agricultural Sciences (SLU), Southern Swedish Forest Research Centre, Alnarp, Sweden E-mail: urban.nilsson@slu.se
• Elfving, SLU, Forest Ecology and Management, Umeå, Sweden E-mail: be@nn.se
• Karlsson, SLU, Unit of Field Based Research, Asa, Sweden E-mail: kk@nn.se

• Strömgren, Swedish University of Agricultural Sciences, Department of Soil and Environment, Uppsala, Sweden E-mail: monika.stromgren@slu.se
• Mjöfors, Swedish University of Agricultural Sciences, Department of Soil and Environment, Uppsala, Sweden E-mail: km@nn.se
• Holmström, Swedish University of Agricultural Sciences, Department of Soil and Environment, Uppsala, Sweden E-mail: bh@nn.fi
• Grelle, Swedish University of Agricultural Sciences, Department of Ecology, Uppsala, Sweden E-mail: ag@nn.se

• Grenfell, University of Helsinki, Dept of Forest Sciences, Helsinki, Finland E-mail: russell.grenfell@helsinki.fi
• Aakala, University of Helsinki, Dept of Forest Sciences, Helsinki, Finland E-mail: ta@nn.fi
• Kuuluvainen, University of Helsinki, Dept of Forest Sciences, Helsinki, Finland E-mail: tk@nn.fi

• Bace, Czech University of Life Sciences, Faculty of Forestry and Wood Sciences, Prague, Czech Republic E-mail: bace@fld.czu.cz
• Svoboda, Czech University of Life Sciences, Faculty of Forestry and Wood Sciences, Prague, Czech Republic E-mail: ms@nn.cz
• Janda, Czech University of Life Sciences, Faculty of Forestry and Wood Sciences, Prague, Czech Republic E-mail: pv@nn.cz

• Martín-García, Sustainable Forest Management Research Institute, University of Valladolid – INIA, Avenida de Madrid 57, 34004 Palencia, Spain, and Forestry Engineering, University of Extremadura, Plasencia, Spain E-mail: jorgemg@pvs.uva.es
• Espiga, Sustainable Forest Management Research Institute, University of Valladolid – INIA, Palencia, Spain E-mail: ee@nn.es
• Pando, Statistics and Operations Research Department, University of Valladolid, Spain E-mail: vp@nn.es
• Diez, Sustainable Forest Management Research Institute, University of Valladolid – INIA, Palencia, Spain E-mail: jjd@nn.es

• Moilanen, Finnish Forest Research Institute, Muhos and Parkano Research Units, Finland E-mail: mikko.moilanen@metla.fi
• Saarinen, Finnish Forest Research Institute, Muhos and Parkano Research Units, Finland E-mail: ms@nn.fi
• Silfverberg, Finnish Forest Research Institute, Muhos and Parkano Research Units, Finland E-mail: ks@nn.fi

• Olsson, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden E-mail: mto@nn.se
• Erlandsson, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden E-mail: me@nn.se
• Lundin, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden E-mail: ll@nn.se
• Nilsson, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden E-mail: torbjorn.nilsson@mark.slu.se
• Nilsson, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden E-mail: an@nn.se
• Stendahl, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden E-mail: js@nn.se

• Kalliokoski, The Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: tuomo.kalliokoski@metla.fi
• Nygren, Department of Forest Ecology, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: pn@nn.fi
• Sievänen, The Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: rs@nn.fi

• Granhus, Norwegian University of Life Sciences, Dept. of Ecology and Natural Resource Management (INA), P.O.Box 5003, NO-1432 Ås, Norway E-mail: aksel.granhus@umb.no
• Fjeld, Swedish University of Agricultural Sciences, Dept. of Forest Resource Management, SE-901 83 Umeå, Sweden E-mail: df@nn.se

• Bolli, Swiss Federal Research Institute, Zürcherstrasse 111, CH-8903 Birmensdorf, Switzerland E-mail: jacqueline.bolli@wsl.ch
• Rigling, Swiss Federal Research Institute, Zürcherstrasse 111, CH-8903 Birmensdorf, Switzerland E-mail: ar@nn.ch
• Bugmann, Swiss Federal Institute of Technology (ETH) Zürich, Universitätstrasse 16, CH-8092 Zürich, Switzerland E-mail: hb@nn.ch

• Diéguez-Aranda, Departamento de Ingeniería Agroforestal, Universidad de Santiago de Compostela. Escuela Politécnica Superior, Campus universitario, 27002 Lugo, Spain E-mail: udieguez@lugo.usc.es
• Grandas-Arias, Departamento de Ingeniería Agroforestal, Universidad de Santiago de Compostela. Escuela Politécnica Superior, Campus universitario, 27002 Lugo, Spain E-mail: jaga@nn.es
• Álvarez-González, Departamento de Ingeniería Agroforestal, Universidad de Santiago de Compostela. Escuela Politécnica Superior, Campus universitario, 27002 Lugo, Spain E-mail: jgag@nn.es
• Gadow, Institut fur Waldinventur und Waldwachstum, George-Auguts-Universität Göttingen. Büsgenweg 5, D-37077 Göttingen, Germany E-mail: kg@nn.de

• Hotanen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: juha-pekka.hotanen@metla.fi
• Maltamo, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: mm@nn.fi
• Reinikainen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: ar@nn.fi

• Nieminen, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: mika.nieminen@metla.fi
• Penttilä, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: tp@nn.fi

• Spinelli, CNR - Timber and Tree Institute, Via Madonna del Piano - Pal. F, 50019 Sesto Fiorentino, Italy; University College Dublin, Ireland E-mail: spinelli@ivalsa.cnr.it
• Owende, Dept. of Agricultural and Food Engineering, University College Dublin, Earlsfort Terrace, Dublin 2, Ireland E-mail: pmoo@nn.ie
• Ward, Dept. of Agricultural and Food Engineering, University College Dublin, Earlsfort Terrace, Dublin 2, Ireland E-mail: smw@nn.ie
• Tornero, Servicios Forestales SA, Ctra. SE-184 Km 0.63, E-41970 Santiponce, Sevilla, Spain E-mail: mt@nn.es

• Hotanen, Finnish Forest Research Institute, Joensuu Research Centre, P.O.Box 68, FIN-80101 Joensuu, Finland E-mail: juha-pekka.hotanen@metla.fi

• Chantal, University of Helsinki, Dept. of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: michelle.dechantal@helsinki.fi
• Eskola, University of Helsinki, Dept. of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: le@nn.fi
• Ilvesniemi, University of Helsinki, Dept. of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: hi@nn.fi
• Leinonen, University of Helsinki, Dept. of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: kl@nn.fi
• Westman, University of Helsinki, Dept. of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: cjw@nn.fi

• Nigh, Ministry of Forests, Research Branch, P.O. Box 9519, Stn. Prov. Govt., Victoria, BC, Canada V8W 9C2 E-mail: gordon.nigh@gems2.gov.bc.ca

• Haeussler, C2 Site 81 RR#2 Monckton Rd., Smithers, B.C., Canada V0J 2N0 E-mail: skeena@bulkley.net
• Bedford, B.C. Ministry of Forests, P.O. Box 9513 Stn. Prov. Govt., Victoria, B.C., Canada, V8W 9C2 E-mail: lb@nn.ca
• Leduc, Groupe de recherche en écologie forestière interuniversitaire, Université du Québec à Montréal, CP 8888, Succursale A, Montréal, Québec, Canada, H3C 3P8 E-mail: al@nn.ca
• Bergeron, Groupe de recherche en écologie forestière interuniversitaire, Université du Québec à Montréal, CP 8888, Succursale A, Montréal, Québec, Canada, H3C 3P8 E-mail: yb@nn.ca
• Kranabetter, B.C. Ministry of Forests, Bag 5000, Smithers, B.C., Canada, V0J 2N0 E-mail: jmk@nn.ca

• Quine, Forestry Commission, Northern Research Station, Roslin, Midlothian EH25 9SY, United Kingdom E-mail: chris.quine@forestry.gsi.gov.uk
• Humphrey, Forestry Commission, Northern Research Station, Roslin, Midlothian EH25 9SY, United Kingdom E-mail: jwh@nn.uk
• Purdy, Forestry Commission, Northern Research Station, Roslin, Midlothian EH25 9SY, United Kingdom E-mail: kp@nn.uk
• Ray, Forestry Commission, Northern Research Station, Roslin, Midlothian EH25 9SY, United Kingdom E-mail: dr@nn.uk

• Moberg, Swedish University of Agricultural Sciences, Department of Forest Management and Products, P.O. Box 7060, S-750 07 Uppsala, Sweden E-mail: lennart.moberg@sh.slu.se

• Hökkä, Finnish Forest Research Institute, Rovaniemi Research Station, PO Box 16, 96301 Rovaniemi, Finland E-mail: hannu.hokka@metla.fi
• Penttilä, Finnish Forest Research Institute, Rovaniemi Research Station, PO Box 16, 96301 Rovaniemi, Finland E-mail: tp@nn.fi

• Karlsson, Swedish University of Agricultural Sciences, Department of Silviculture, S-901 83 Umeå, Sweden E-mail: anders.karlsson@ssko.slu.se
• Albrektson, Swedish University of Agricultural Sciences, Department of Silviculture, S-901 83 Umeå, Sweden E-mail: aa@nn.se
• Forsgren, Swedish University of Agricultural Sciences, Department of Silviculture, S-901 83 Umeå, Sweden E-mail: af@nn.se
• Svensson, Swedish University of Agricultural Sciences, Department of Silviculture, S-901 83 Umeå, Sweden E-mail: ls@nn.se

• Päätalo, Faculty of Forestry, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: mlp@nn.fi

The height growth of trees influences the productivity of stands and the competitiveness of species, shaping the range of their distribution. Dominant height growth was assessed for European beech (Fagus sylvatica L.), growing outside of its natural distribution range in the western part of Latvia. In 10 neighbouring experimental stands, 20 dominant trees were felled for stem analysis. Height growth was modelled using the generalised algebraic difference approach, applying several non-linear equations and mixed procedures. The Chapman-Richards and Sloboda models showed the best fit to the data. Height growth of the second generation (younger) trees exceeded that of the first generation, and followed curve for a higher site index, suggesting sufficient adaptation and improving conditions. Height growth of the studied beech exceeded predictions for beech in southern Sweden, which is considered to be the northern limit of the species range, yet the growth pattern differed. In Latvia, slower height growth was estimated for site indices < 32 m (in 100 years) during the first 60 years, yet larger maximal height was predicted, suggesting a longer establishment period. Nevertheless, the improving height growth indicated increasing potential for the application of the species in commercial forestry, and an expansion of the species within the region even during the 21th century.

• Matisons, Latvian State Forest Research Institute “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 E-mail: robism@inbox.lv
• Šņepsts, Latvian State Forest Research Institute “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 E-mail: guntars.snepsts@silava.lv
• Puriņa, Latvian State Forest Research Institute “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 E-mail: liga.purina@silava.lv
• Donis, Latvian State Forest Research Institute “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 E-mail: janis.donis@silava.lv
• Jansons, Latvian State Forest Research Institute “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 E-mail: aris.jansons@silava.lv

• Lideskog, Division of Product and Production Development, Luleå University of Technology, SE-97187 Luleå, Sweden E-mail: hakan.lideskog@ltu.se
• Ersson, Department of Forest Biomaterials and Technology, SLU, SE-90183 Umeå, Sweden E-mail: back.tomas.ersson@slu.se
• Bergsten, Department of Forest Biomaterials and Technology, SLU, SE-90183 Umeå, Sweden E-mail: urban.bergsten@slu.se
• Karlberg, Division of Product and Production Development, Luleå University of Technology, SE-97187 Luleå, Sweden E-mail: magnus.karlberg@ltu.se