Articles containing the keyword 'stand'

The objectives of this study were to record residual stand damage during harvesting operations and evaluate the influence of factors such as distance of the tree from the strip road, machine parts, operational phase, on the occurrence of tree wounds. The machine was a farm tractor equipped with a crane mounted on the front axle and a single grip harvester head. The study was carried out in two stands located in Southeast Sweden. Stand 1 was a 30-year-old Norway spruce (Picea abies (L.) H. Karst.) plantation on an afforested pasture while stand 2 was a 90-year-old mixed stand of Norway spruce, Scots pine (Pinus sylvestris L.), birch (Betula pendula Roth) and aspen (Populus tremula L.).

The mean damage percentage was 6.3% for the first stand and 6.5% for the second stand. Sixty-five percent of the wounds were less than 50 cm², with 91% of the damage occurring on the stem and 91% of the damage on or below the root collar. Sixty-six percent of the wounds produced by the stem under processing or by the harvesting head while only 10% of the wounds were produced by the tractor wheel. Damaged trees were distributed evenly in the crane reach zone. Significant differences were found between rut depths after one, two, four and six passes of the tractor in stand 1.

• Athanassiadis, E-mail: da@mm.unknown

Within the European Community snow damage affects an estimated 4 million m³ of timber every year, causing significant economic losses to forest owners. In Northern Europe, for example, the occurrence of snow damage has increased over the last few decades mainly due to the increase in total growing stock. The most common form of damage is stem breakage, but trees can also be bent or uprooted. Trees suffering snow damage are also more prone to consequential damage through insect or fungal attacks.

Snow accumulation on trees is strongly dependent upon weather and climatological conditions. Temperature influences the moisture content of snow and therefore the degree to which it can accumulate on branches. Wind can cause snow to be shed, but can also lead to large accumulations of wet snow, rime or freezing rain. Wet snow is most likely in late autumn or early spring. Geographic location and topography influence the occurrence of damaging forms of snow, and coastal locations and moderate to high elevations experience large accumulations. Slope plays a less important role and the evidence on the role of aspect is contradictory. The occurrence of damaging events can vary from every winter to once every 10 years or so depending upon regional climatology. In the future, assuming global warming in northern latitudes, the risk of snow damage could increase, because the relative occurrence of snowfall near temperatures of zero could increase.

The severity of snow damage is related to tree characteristics. Stem taper and crown characteristics are the most important factors controlling the stability of trees. Slightly tapering stems, asymmetric crowns, and rigid horizontal branching are all associated with high risk. However, the evidence on species differences is less clear due to the interaction with location. Management of forests can alter risk through choice of regeneration, tending, thinning and rotation. However, quantification and comparison of the absolute effect of these measures is not yet possible. An integrated risk model is required to allow the various locational and silvicultural factors to be assessed. Plans are presented to construct such a model, and gaps in knowledge are highlighted.

• Nykänen, E-mail: mn@mm.unknown
• Broadgate, E-mail: mb@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown
• Peltola, E-mail: hp@mm.unknown
• Quine, E-mail: cq@mm.unknown

The purpose of this study was to test the benefits of a forest site quality map, when applying satellite image-based forest inventory. By combining field sample plot data from national forest inventories with satellite imagery and forest site quality data, it is possible to estimate forest stand characteristics with higher accuracy for smaller areas. The reliability of the estimates was evaluated using the data from a stand-wise survey for area sizes ranging from 0.06 ha to 300 ha. When the mean volume was estimated, a relative error of 14 per cent was obtained for areas of 50 ha; for areas of 30 ha the corresponding figure was below 20 per cent. The relative gain in interpretation accuracy, when including the forest site quality information, ranged between 1 and 6 per cent. The advantage increased according to the size of the target area. The forest site quality map had the effect of decreasing the relative error in Norway spruce (Picea abies) volume estimations, but it did not contribute to Scots pine (Pinus sylvestris) volume estimation procedure.

• Tokola, E-mail: tt@mm.unknown
• Heikkilä, E-mail: jh@mm.unknown

Possibilities of distance-independent and -dependent competition indices to describe the competition stress of an individual tree was studied in Southern Finland. Five half-sib open-pollinated families and one check lot of Scots pine (Pinus sylvestris L.) was used as study material in order to analyse competitive interactions of crown form and stand density variation. Almost all competition indices correlated strongly with radial increment. Thus distance-independent indices were adequate to describe competition in young row plantations, where distance effects between trees were implicitly eliminated. Correlations between indices and height increment were not significant. Along with the increase in competition, the width and length of the crown and the diameter increment of the stem of some narrow-crowned families decreased slowly compared to wide-crowned families.

• Mäkinen, E-mail: hm@mm.unknown

The horizontal and vertical stand structure of living trees was examined in a managed and in a primeval Norway spruce-dominated forest in Southern Finland. Tree size distributions (DBHs, tree height) were compared using frequency histograms. The vertical distribution of tree heights was illustrated as tree height plots and quantified as the tree height diversity (THD) using the Shannon-Weaver formula. The horizontal spatial pattern of trees was described with stem maps and quantified with Ripley's K-function. The spatial autocorrelation of tree sizes was examined with semivariogram analysis. In the managed forest the DBH and height distributions of trees were bimodal, indicating a two-layered vertical structure with a single dominant tree layer and abundant regeneration in the understory. The primeval forest had a much higher total number of trees which were rather evenly distributed in different diameter and tree height classes. The K-function summaries for trees taller than 15 m indicated that the primeval stand was close to complete random pattern. The managed stand was regular at small distances (up to 4 m). The semivariograms of tree sizes (DBH tree height) showed that the managed forest had a clear spatial dependence in tree sizes up to inter-tree distances of about 12 meters. In contrast, the primeval spruce forest had a variance peak at very short inter-tree distances (< 1 m) and only weak spatial autocorrelation at short inter-tree distances (1–5 m). Excluding the understory trees (h < 15 m) from the analysis drastically changed the spatial structure of the forest as revealed by semivariograms. ln general, the structure of the primeval forest was both horizontally and vertically more variable and heterogeneous compared to the managed forest. The applicability of the used methods in describing fine-scale forest structure i discussed.

• Kuuluvainen, E-mail: tk@mm.unknown
• Leinonen, E-mail: kl@mm.unknown
• Nygren, E-mail: mn@mm.unknown
• Penttinen, E-mail: ap@mm.unknown

This paper deals with the testing of dynamic stratification for estimating stand level forest characteristics (basal areas, mean diameter, mean height and mean age) for a 117 ha study areas in Finland. The results do not show possibilities to achieve more accurate estimates using only Landsat TM principal components as auxiliary data opposed to static stratification. It was found that in dynamic stratification non-measured observations should be assigned the mean characteristics of the measured observations that belong to the same cube (class) instead of stratification variable classes until a certain limit. If only one principal component is used the number of classes has, however, little influence. Low field values are overestimated and high values underestimated.

The only successful results were obtained using two variables of different origin – the qualitative development stage class and the quantitative 1st principal component. The lowest root mean square error in estimating basal area was 6.40 m²/ha, mean diameter 3.34 cm, mean height 2.65 m and mean age 14.06 years. This increase of stratification accuracy is mainly resulted by the use of development stage class as an auxiliary variable.

• Mozgeris, E-mail: gm@mm.unknown

Comparisons were made between artificially and naturally regenerated stands in the south-eastern part of North Karelia, Finland, and naturally regenerated stands in the western parts of the Republic of Karelia, Russian Federation. The effect of soil fertility and silvicultural operations on the stand structure was also investigated.

The results of the study show clearly that when forests are artificially regenerated the stand structure includes less variation when compared with the stands naturally regenerated. Differences between the regeneration methods are clearer the more fertile the forest site is. Within the regeneration method there is also a clear trend in stand structure, with the variation decreasing the poorer the site. The effect of silvicultural operations, i.e. the cleaning of the sapling stand, has disappeared by the time of first thinning, although it appears to have a permanent effect on the dynamics of the tree species within a stand.

The variation of the stand structure can be regarded as an essential factor for the potential biodiversity of the stand also at its young vegetation succession stage. This capacity for maintaining the forest biodiversity, developed at the young vegetation succession stage, becomes increasingly important in subsequent vegetation succession stages. Natural regeneration provides improved possibilities for the operations preserving forest biodiversity, as it generates more dense stands with a wider variation in stand structure, compared to artificial regeneration.

• Uuttera, E-mail: ju@mm.unknown
• Maltamo, E-mail: mm@mm.unknown

The following treatments were compared in three Scots pine (Pinus sylvestris L.) reforestation areas on a scarified moist mineral-soil site in southern Finland, planted with 1+1 bareroot stock in spring 1987: (a) no weed control treatment; (b) mulching with a fibre slurry produced by mixing wastepaper with water and applied 1 cm deep to an area of 60 cm in diameter around the seedling soon after planting; (c) glyphosate (at 2 kg ha^-1) sprayed on a 1 m² spot around the seedling in early August 1987; (d) terbuthylazine (at 10 kg ha^-1) applied as (c). Monitoring of the trials over a 4-year period between 1987–90 showed that none of the treatments reduced surface vegetation to an extent that would have benefited pine. The percentage cover development of the vegetation, dominated by Agrostis capillaris, Calamagrostis arundinaceae, Deschampsia flexuosa, Festuca ovina, Epilobioum angustifolium and Pteridium aquillinum, followed much the same pattern in all treatments, with (c) slightly favouring forbs. Survival of pine at the end of the study period was about 90%, with non-significant differences between treatments. Mulching and terbuthylazine treatment slightly reduced seedling height growth in the second year. Growth was better in glyphosate treatment than in terbuthylazine treatment in the lowest (<30%) and the highest (>60%) pre-treatment weed cover classes, and in the latter also better than in untreated control. Mulching gave variable results; at its best it provided also good control of weeds for several years, without, however, improving the initial development of pine in these trials.

• Siipilehto, E-mail: js@mm.unknown
• Lyly, E-mail: ol@mm.unknown

Model-based information systems have proved valuable planning tools for analysing the production possibilities of forests as well as for understanding forest resources dynamics, stand management practices and forest economics. Computerized forest models implemented in the users’ information systems facilitate the transfer and application of research results in practical forestry.

Conclusions and visions concerning modelling are drawn from experiences in developing the MELA system and its application in solving timber production problems on both the national and forest holding level in Finland. The precondition for predicting forest resource dynamics and for planning the utilization of forests is to accept conditions, uncertainties and a restricted period of time.

The interactive process of forest resource, growth and drain monitoring, and forest management planning supported by forest research and modelling, are the means to enable an operational information base for a dynamic regulation and adaptation strategy for forest resource management under changing conditions and uncertainty.

The PDF includes an abstract in Finnish.

• Siitonen, E-mail: ms@mm.unknown

The model predicts the base diameter of the thickest living branch of a tree growing in a planted or naturally regenerated even-aged stand. A mixed model type was used in which the residual variation was divided into within-stand and between-stand components. The study material consisted of 779 trees measured in 12 plots located in 20 to 35 years old Scots pine (Pinus sylvestris L.) stands (breast height age 10 to 20 years). Branch diameter was closely connected to the breast height diameter of the stem. In a stand of a certain age, competition by close neighbours slightly decreased branch diameter in a given diameter class. According to the model, the greatest difference is between trees subjected to very little competition and those subjected to normal competition. The model was used in simulated stands with varying age, density, and tree arrangement. The simulations showed that trees with rapid diameter growth at young age had thicker branches at a given breast height diameter than trees with slower diameter growth. However, a very slow growth rate did not produce trees with branches thinner than those possessing a medium growth rate.
The PDF includes an abstract in Finnish.

• Pukkala, E-mail: tp@mm.unknown
• Karsikko, E-mail: jk@mm.unknown
• Kolström, E-mail: tk@mm.unknown

Model computations on the management of Scots pine (Pinus sylvestris L.) at the seedling stage showed that a rising temperature due to the suggested climate change could increase the competition capacity of birch species (Betula pendula) more than Scots pine, whose growth could even decline during the course of a rise in temperature. A temperature rise could, thus, bring the time of removal of birches forward when aiming at Scots pine timber stands composed of these tree species. The increasing proportion of birches makes the removal of birches even more urgent and emphasizes the need for careful management of Scots pine stands under rising temperatures. The first thinning of Scots pine is generally brought forward; this is particularly the case when wide spacing is applied in planting. A furthrer rise in temperature magnifies the above patterns by reducing further the competitive capacity of Scots pine in relation to birches.
The PDF includes an abstract in Finnish.

• Kellomäki, E-mail: sk@mm.unknown
• Kolström, E-mail: mk@mm.unknown

The results of regenerating 54 and 36–38-year old strip cuttings were surveyed in the Kivalo Research Forest (N 66°20’, E 26°40’). 15 measurement plots were placed on each strip. The most common forest type was Hylocomnium-Myrtillus type. Regeneration of the strips proved to be slow. Most of the spruces growing on the strips probably originated from the time before the cutting. The average number of stems was 1,155 per ha, of which one third consisted of broadleaved trees. The average volume increment of stem wood after cutting had been about 1 m³/ha/yr, but it was increasing at the time of the inventory. Both the reforestation of the strips and the development of the emergent stands were dependent on elevation and site fertility. Site fertility was indicated by the abundance of Vaccinium myrtillus.

The PDF includes an abstract in English.

• Pohtila, E-mail: ep@mm.unknown

The effect of drainage on structure of tree stands is analysed by comparing the average structural characteristics (e.g. diameter distribution) of stands in the data for different drainage age classes and selected site types. The material consists of ca. 4,400 relascope sample plots, which are part of a large drainage area inventory project. The uneven-aged structure of the virgin peatland forest is preserved for several decades after drainage. This is enhanced by the post-drainage increase of small-diameter trees, especially birch. The number of trees per hectare increased during a period of ca. 30 years and levelled off thereafter. The increase in the number of saw log stems is clearly related to the fertility of the site and its geographical location.

The PDF includes a summary in English.

• Hökkä, E-mail: hh@mm.unknown
• Laine, E-mail: jl@mm.unknown

A method for using satellite data in forest inventories and updating is described and tested. The stand characteristics estimated by the method showed high correlation with the same characteristics measured in the field. The correlation coefficients for volume, age and mean height were about 0.85. It seems that the method is applicable to practical forestry. Extensive work in programming, however, is required.

The PDF includes an abstract in Finnish.

• Poso, E-mail: sp@mm.unknown
• Paananen, E-mail: rp@mm.unknown
• Similä, E-mail: ms@mm.unknown

The effect of competition on the radial growth of Scots pine (Pinus sylvestris L.) was studied in three naturally regenerated stands located in North Karelia, Finland. The competition situation of an individual tree was described with various competition indices which depended on the sizes and distances from the neighbouring trees. One competition index explained about 50% of the variation in 5-year radial growth in one stand. If all stands were combined, one index explained 43.5%, two indices 48.9% and three indices 51.2% of the variation. In one stand, the best competition indices accounted for about 20% of that variation which could not be explained by tree diameter. If all three stands were combined, the best index explained 11% of the residual variation. About 40% of the variation in 5-year radial growth could not be explained by the diameter and competition indices.
The PDF includes an abstract in Finnish.

• Pukkala, E-mail: tp@mm.unknown
• Kolström, E-mail: tk@mm.unknown

The effect of the size of seed crop, dispersal of seeds and the early development of seedlings on the density and spatial distribution of young Scots pine (Pinus sylvestris L.) stands are evaluated on the basis of theoretical models. The models include (i) number and spatial distribution of parent trees on the regeneration area, (ii) size of annual seed crop, (iii) seed dispersal from a particular parent tree, (iv) germination of the seeds (germination percentage), (v) death of ageing seedlings after the establishment process, and (vi) height growth of the seedlings.

As expected, stand density and spatial distribution varied within a large range in relation to the density of the parent trees and the distance from them. The simulations also showed that natural seedling stands can be expected to be heterogenous due to the geometry of seed dispersal, emphasizing the frequency of young and small trees. The properties of the seedling stands were, however, greatly dependent on the density of the parent trees and the length of the regeneration period.

The PDF includes an abstract in Finnish.

• Kellomäki, E-mail: sk@mm.unknown
• Hänninen, E-mail: hh@mm.unknown
• Kolström, E-mail: tk@mm.unknown
• Kotisaari, E-mail: ak@mm.unknown
• Pukkala, E-mail: tp@mm.unknown

Early growth of four different tree species (Pinus sylvestris L., Picea abies (L.) H. Karst., Larix sibirica Ledeb and Betula pendula Roth) 16–23 years after planting were compared in a field experiment of 16 square plots established on a stony, grove-like upland (Oxalis-Myrtillus forest type) in Southern Finland. This study gives additional results to the publication Folia Forestalia 386/1979.

At this early stage, the growth of the spruce stand was clearly slower than that of the other species for all parameters to be measured (height, diameter, and volume growth). Height growth was most rapid in the silver birch stand and diameter growth in the larch stand. No clear differences were found in the mean volume of the 100 thickest trees in the stand between the larch and silver birch.

The PDF includes a summary in English.

• Parviainen, E-mail: jp@mm.unknown

Young Norway spruce (Picea abies (L.) H. Karst.) are susceptible to early summer frost damage. Birch (Betula pubescens Ehrh.) naturally colonize rich or fairly rich drained peatlands after clear cutting, and can provide protection for developing seedlings. The report describes the development of spruce stands after various types of handing of the birch nurse crops.

Different proportions of birch and spruces did not have any influence on the spruce stand production. In cases where the nurse crop stand is removed when the spruce stand age was 20 years and height 4 m the spruce suffered badly but recovered with time, reaching the spruce stand growing under a nurse stand within the next 20 years. The height growth of spruce depends on the density of the nurse stand, especially on fertile sites. The development of diameter growth also depends on the density of the nurse trees. Removal of the nurse stand in spruce stands on the sites concerned should be done when the spruce stand is 20 years old and at the height of 4 m.

The PDF includes a summary in English.

• Heikurainen, E-mail: lh@mm.unknown

This article reviews experiments and practical experience of forest management by the selection system in Finland. In an experiment of 25-year duration the annual growth of uneven-aged Norway spruce (Picea abies (L.) H. Karst.) stands was only about 50% of the average annual yield of even-aged stands in normal rotation on the same site.

In Finland the selection system is applicable under exceptional conditions only, viz. In intensively managed park stands and, on the other hand, on very marginal sites, e.g. on peat bogs and mountains near the tree-line. Even normal silviculture, however, may include cuttings which somewhat resemble selection system, e.g. removal of standards or restoration of mismanaged forests.

The PDF includes a summary in English.

• Mikola, E-mail: pm@mm.unknown

The paper presents a method based on two phase sampling and applicable to forest inventories. The first phase estimates are obtained from satellite imagery and, if required, from extra material such as maps. Second phase estimates are measured in the field. The method is flexible and also applicable to compartmentwise forest inventories. The experiments were based on six study areas with 439 relascope plots. The correlation coefficients between first and second stage estimates varied largely according to the study area.

The PDF includes a summary in Finnish.

• Poso, E-mail: sp@mm.unknown
• Häme, E-mail: th@mm.unknown
• Paananen, E-mail: rp@mm.unknown

The study based on young Scots pine (Pinus sylvestris L.) of varying density showed that number of living branches per whorl and total number of living branches per tree were negatively correlated with stand density. On the contrary, the number of dead branches increased with increasing stand density. The diameter of living and dead branches decreased with increasing stand density. Consequently, the branchiness, i.e. the share of the branch cross-sectional area from the surface area of the stem, decreased in dense stands compared with the thin stands. At the densest stands the branchiness, however, levelled of indicating a greater decrease of the radial growth at stems than at branches. The 2/3 power law described relatively well the relationship between stand density and mean squared branch diameter of living branches.

The PDF includes a summary in English.

• Kellomäki, E-mail: sk@mm.unknown

Vegetation data collected from a random sample of mature forest stands representing mesic upland forest sites in Southern Finland were analysed and classified using two-way indicator species analysis (TWINSPAN). The variation of some characteristics of the tree stand and soil fertility between the produced vegetation units were analysed statistically.

Both the species list and the sample list arranged by TWINSPAN reflects the overall site fertility considerably well. The results are in agreement with the main division of mesic forest sites in the Finnish forest site type classification: vegetation units which can be assigned to the Oxalis-Myrtillus site type are clearly separated from the remaining units, and the overall site fertility indicates a statistically significant difference. The within-type variation in the vegetation composition in the Oxalis-Myrtillus site type reflects the variation in site fertility, whereas the within-type variation in the Myrtillus site type is mainly caused by the tree stand factor.

The PDF includes a summary in Finnish.

• Kuusipalo, E-mail: jk@mm.unknown

Two Japanese models regarding the within-stand competition have been reviewed on the basis of relevant literature. Competition-density and 3/2 th power models seem to be applicable also into tree stands. The latter model has been applied into the material obtained from literature. Computations showed consistancy with the results obtained elsewhere in the world. It is concluded that also in Finnish conditions the 3/2 th power law may have great potentials in describing the effects of stand density on tree size.

The PDF includes a summary in English.

• Kellomäki, E-mail: sk@mm.unknown
• Nevalainen, E-mail: tn@mm.unknown

The concepts of the terms compartment and compartment-wise forest inventory have been studied empirically by repeated delineation and intensive systematic plot samples. The material consisted of 16 study areas of some 8–90 hectares in size in Southern Finland and of more than 1,000 relascope plots. Stands and compartments were found to be rather heterogenous. Alternative photographs, working techniques and test persons were studied. An endeavour for better accuracy in compartment inventories is recommended.

The PDF includes a summary in English.

• Poso, E-mail: sp@mm.unknown

The choice of sampling method is of prime importance when seeking relevant information about the height distribution and spatial arrangement of seedlings in the regeneration surveys. It is suggested that the size of sampling plots should depend on the height of the seedlings. Tall seedlings should be sampled from a larger area than short ones since tall seedlings are more important for the future development of the stand. We suggest principles for a technical development task to construct a device which is easy to use in practical regeneration surveys and by which sampling can be made proportional to plant height or any desired function of height.

The PDF includes a summary in Finnish.

• Lappi, E-mail: jl@mm.unknown
• Kotisaari, E-mail: ak@mm.unknown
• Smolander, E-mail: hs@mm.unknown

The needle area distribution and crown structure of a young planted Scots pine (Pinus sylvestris L.) stand are described. The crown structure and crown shape showed apparent regularity in crown structure regardless of stand dynamics. Similarly, the shoot structure and individual needle area showed regularity in the number of needles per branch and shoot length unit, and consequent phytoarea density inside the needle cylinder. Also, the shoot area and needle area distributions were found to show a regular distribution of needle biomass throughout the crown, also inside the crown, in the dominant trees. In the suppressed trees the needle biomass was located in the upper crown and on the surface area of the crown. Estimates of the canopy needle area and distributions are given. The results were applied in calculations of the within-stand light regime. The results correlated well with the empirical results.

The PDF includes a summary in Finnish.

• Kellomäki, E-mail: sk@mm.unknown
• Oker-Blom, E-mail: po@mm.unknown

The aim of the paper was to describe the development of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) seedling stands on drained peatlands and to find out the principal factors influencing their growth. The material under survey consists of 180 sample plots distributed from southern coast of Finland to the Polar Circle.

The most important growth factors have been the accumulated temperature sum, site quality, drainage intensity and silvicultural condition, such as the density of the stand, the proportion of birch in the stand, and the amount of possible shelterwoods. The influence of these factors, and to some extent the influence of fertilizing, and the disturbing effects of some forest damages, such as frost, growth disturbances and elk damages were investigated. Comparisons of the development in the seedling stands on drained peatlands with the known development of seedling stands in mineral soils were made.

The PDF includes a summary in English.

• Heikurainen, E-mail: lh@mm.unknown

The share of stem, branch and needle growth was dependent on the within-stand light regime in a young Scots pine (Pinus sylvestris L.) stand. The share of needle growth increased at the expense of stem and branch growth in poor light conditions. In good light condition the share of branch wood increased substantially. The share of stem wood growth was greatest in moderate shading, emphasizing the role of an adequate stand density for growing high-quality timber. The basic density of the stem wood was considerably greater in suppressed trees than in dominating trees. The differences were related to the illumination of the crown system.

The PDF includes a summary in Finnish.

• Kellomäki, E-mail: sk@mm.unknown

The applicability of MCPA- and 2,4,5-T-herbicides for use in the management of sapling stands and the possibilities of carrying out foliar spraying at an earlier date than at precent with smaller doses of the active ingredient were examined in this study. The results were obtained from foliage spraying experiments carried out in Central Finland in summer 1976. MCPA and 2,4,5-T were as effective as each other against deciduous tree species. However, MCPA was slightly more effective against aspen (Populus tremula L.) than 2,4,5-T. The spraying date had no effect on the mortality rate of aspen or birch (Betula pendula Roth and B. pubescens Ehrh.) There were only very slight differences between the results for different dosage levels. The damage caused to Scots pine (Pinus sylvestris L.) was very slight. The temperature conditions prevailing during spraying affected spraying effectiveness in such way that the mortality rate decreased during cold period.

The PDF includes a summary in English.

• Meriluoto, E-mail: jm@mm.unknown

The technique of double normalizing, i.e. normalizing the relative needle biomass and the length of the living crown system, is applied to the modelling of the distribution of needle biomass in the canopy of young Scots pine (Pinus sylvestris L.) stands. The study based on the parameters of β-function shows that at the individual-tree level, the variance in needle distribution was not closely associated with any tree characteristics. A shift in the point of maximum needle biomass upwards unsuppressed trees was, however, evident. This was associated with an increase in the height of the trees. At the stand level, the stand mean height and stand density had an equal and a rather high potential for explaining the variance in the needle distribution. The normalized crowns are utilized in models for determining light extinction in the crown. A special technique for determining the amount of photosynthates available for growth in a particular tree is presented.

The PDF includes a summary in Finnish.

• Kellomäki, E-mail: sk@mm.unknown
• Hari, E-mail: ph@mm.unknown
• Ilonen, E-mail: pi@mm.unknown
• Kanninen, E-mail: mk@mm.unknown

The social circumstances of forest workers may be considered from the viewpoint of incomes and standard of living. The committee appointed by the U.N. did, however, define in 1961 the standard of living using the following partial factors: health, housing, working conditions, security, human rights, education, recreation, clothing and nutrition.

Research has given a rather gloomy picture of forest workers’ health conditions. The living of a forest worker is cramped and long distances to the work sites often compel him to live in working site accommodations, separated from the family. Transport to the working site and building of family quarters for forest workers in densely populated rural areas and villages can improve the worker’s living. The fall of worker’s income in contract work due to deterioration of his physical capacity of performance starts already at the age of 35. In forest work a man’s energy consumption may be higher than 20,000 kJ (4,800 kcal) per day; it is one of the most strenuous occupations. In addition, there is a high accident risk in the work. The heavy work and the working conditions should be taken into account in determining the correct nutrition. Being those who make objections and are primarily responsible for the forest workers’ conditions, forest technicians and forestry officers should – both in their capacity as forest authorities and as the employer’s representatives – see to the improvement of forest workers’ working conditions and social position.

The PDF includes a summary in Finnish.

• Hellman, E-mail: lh@mm.unknown

A system for measuring the net photosynthesis, transpiration and environmental factors within the canopy and ground cover vegetation is described. The system operates continuously throughout the growing season in a young Scots pine (Pinus sylvestris L.) stand. A data-logging unit controls the system and carries out the measurements on the readings of the sensors of photosynthesis, transpiration, light intensity outside the canopy, light climate inside the assimilation chambers, and dry and wet temperatures from selected points. These measurements are shown digitally and automatically punched onto paper tape.

The PDF includes a summary in Finnish.

• Hari, E-mail: ph@mm.unknown
• Kanninen, E-mail: mk@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown
• Luukkanen, E-mail: ol@mm.unknown
• Pelkonen, E-mail: pp@mm.unknown
• Salminen, E-mail: rs@mm.unknown
• Smolander, E-mail: hs@mm.unknown

The aim of the paper was to investigate the recreational potential of successional stand based on the suitability of the stands for different recreational activities and the trampling tolerance of ground cover. The relationship between selected recreational activities and the volume of the stands under study has been determined. These functions have been utilized in determining the potential of the successional stand for different recreational activities. Combining this information with trampling tolerance gave criteria for determining the recreational potential of a stand. The results emphasize the importance of varying the distribution of development stages in a recreational area and in its management.

The PDF includes a summary in Finnish.

• Kellomäki, E-mail: sk@mm.unknown

A theoretical nomogram was made for estimating the costs of fully mechanized thinning and the driving speed of the machine. Based on this nomogram and the previous studies three harvesting methods were compared; systematic fully mechanized harvesting, selective fully mechanized harvesting, and manual felling combined with whole-tree chipping.

The third method was cheaper than the fully mechanized methods in a pole-stage stand. The choice of the most advantageous chipping station depended on conditions, but the smaller tree size and possibly the reduced damage on the remaining stand favour chipping on the strip road rather than chipping on the intermediate landing or at the mill.

Mechanized systematic thinning was the cheapest method for harvesting in the sapling stand. The required driving speed were so low that ergonomic factors should not hinder its use. Factors related to the future production of the stand do, however, limit its use. Mechanized selective thinning does not seem to be an economic method for harvesting in a sapling or pole-stage stand.

The PDF includes a summary in English.

• Harstela, E-mail: ph@mm.unknown
• Tervo, E-mail: lt@mm.unknown

The aim of the present paper was to study the annual production of Pleurozium schreberi (Brid.) Mitt., Hylocomnium splendens (Hedw.) B.S.G and Dicranum polysetum Sw. as a function of light available for photosynthesis. The productivity of the above moss species is studied using the harvested quadrats method in Norway spruce (Picea abies) stands of the Myrtillus site type representing different stand density classes (basal area from 0 to 34 m2/ha) in Southern Finland.

The annual production of each species in different stands was correlated with the amount of light available for photosynthesis i.e. with the photosynthetic production. Functions for the dependence of productivity on light conditions were produced for each species. The individual functions and their ecological significance is discussed. The adaptation of each species to low light intensity is evident since no meaningful addition to production takes place when the photosynthetic light ratio reaches values greater than 0.3–0.4. In other words, the level of photosynthesis which is 30–40% of that possible in the open, provides sufficient supply of carbohydrates or the basic functions of the moss species studied. Pleurozium schreberi and Dicranum polysetum seem to have greater light requirements than Hylocomnium splendens.

The PDF includes a summary in Finnish.

• Kellomäki, E-mail: sk@mm.unknown
• Hari, E-mail: ph@mm.unknown
• Väisänen, E-mail: ev@mm.unknown

The aim of the study was to determine which characteristics of tree stands could be used in urban noise control and to develop guidelines for practical urban forestry. The attenuation of artificially produced noise was measured in various types of stands. The effect of tree stands on noise attenuation has been analysed using a model based on the physics of sound propagation.

The results show that the excess noise attenuation caused by trees can be in good situations 60% measured in energy units, compared with the attenuation caused by geometric spreading. This is 5–8 dB. The total amount of needles, leaves and branches of a stand proved to be the most important factors in noise attenuation. However, the density and height of a stand had rather high value in predicting the behaviour of noise in tree stands. Based on multiple regression between noise attenuation and tree species composition, density and height of a stand it was developed the nomograms with which the noise level at a certain point can be predicted when the tree stand characteristics, the distance and the noise level of the noise source are known and on the opposite way.

The PDF includes a summary in Finnish.

• Kellomäki, E-mail: sk@mm.unknown
• Haapanen, E-mail: ah@mm.unknown
• Salonen, E-mail: hs@mm.unknown

About one million hectares of forests are fertilized annually in Finland. The goal of the present study was to find out, by means of calculations, how the profitability of forest fertilization varies with variations in the stage of development of tree crops, the quality of the site and its geographical location. Calculations concerned bot fertilization of forests in mineral soil sites and in drained peatlands. The study is a part of a larger project concerning the order of profitability of different forest improvement measures in different conditions. The problems dealt with in this study were approached from the point of view of national economy.

On the basis of two empirical materials it is shown that there is a high correlation between the stand growth percentages before and after the fertilization. Applying the results to existing yield tables the authors calculate benefit/cost ratios showing the stage of development of the stand, the quality of the site and its geographical location. According to the results, fertilization is more profitable in sites of medium fertility than on poor sites. Profitability decreases rather fast from south to north and with decreasing timber prices. 

The PDF includes a summary in English.

• Keltikangas, E-mail: mk@mm.unknown
• Seppälä, E-mail: ks@mm.unknown

The aim of this study was to estimate the genetic gain of volume growth in Scots pine (Pinus sylvestris L.) selected seed stands. To obtain highest possible accuracy, the estimations are based on a large statistical material comprising 197 separate seed stands. It is concluded that the genetic gain of volume growth ranges between 7.4–15.0%. Unwanted pollen contaminations may, however, in the worst case halve this genetic gain.

The PDF includes a summary in English.

• Oskarsson, E-mail: oo@mm.unknown
• Tigerstedt, E-mail: pt@mm.unknown

This paper presents the results of a contest performed on behalf of the Finnish bank Kansallis-Osake-Pankki and the Central Forestry Board Tapio on growing trees on peatlands. Over 5,000 sample plots were established on drained peatlands in various parts of Finland. The aim was to achieve a best possible growth of seedling stands on peatland. The factors influencing the growth of 85 best Scots pine (Pinus sylvestris L.) and 60 best Norway spruce (Picea abies (L.) H.Karst.) sample plots were studied.

The height growth of the seedling stands decreased towards the north. Fertilization seemed not to decrease the regional differences; rather on the contrary. On the other hand, fertilization increased height growth, but evidently so that the increase obtained was greater in the southern than in the northern parts of the country. Light fertilization (50 kg/ha of K₂O and 60 kg/ha of O₂P₅) caused a clear increase in height growth while heavy fertilization (100 g/ha of K₂O and 120 kg/ha of O₂P₅), had same effect but to much greater extent than the former. Spruce seedling stands in particular benefitted of the heavy fertilization.

Fertilization did not eliminate the original differences in the quality of the sites in question, but these could still be seen in the height growth after fertilization. The effect of drain spacing on the height growth was not very clear. In dense seedling stands (800 seedlings/ha) the height growth of the dominant seedlings was greater than that obtained in stands of lower density. Hold-overs caused a decrease in the growth of the seedling stands.

The PDF includes a summary in English.

• Heikurainen, E-mail: lh@mm.unknown
• Ouni, E-mail: jo@mm.unknown

The objective of this investigation was to study the influence of stand density of white birch (Betula pubescens Ehrl.) on the minimum temperatures in the stand during the growing season, and the actual minimum temperatures of the leading shoot of Norway spruce (Picea abies (L.) H. Karst.) seedlings growing in the open. The 40-year-old uniform white birch stand was situated in 142 m above the sea level in Southern Finland. The stand was treated with thinnings of three different densities in 1961.

Air temperature was recorded in four sample plots at heights of 0.1 m, 0.5 m, 1.0 m, 2 m and 4 m. In the stand of moderate density, temperatures were measured at heights of 6.0 m, and in the stand of full density at 6.0 m, 8.0 m and 10.0 m.

The temperature differences between stands of various densities proved to be rather small. Especially the thinnest stand differed very little from the open area. The soil surface has in all cases been warm compared with the higher air layers indicating meadow-fog-type by Geier (1965). On cloudy or windy weather all the temperature profiles in the various stands resembled each other. The difference between the air temperature and temperature of the spruce shoot was greatest at midnight and decreased steadily thereafter.

The problem in using shelter stands for spruce regeneration areas is that optimum shelter stand density is difficult to define. Already a thin shelter stand causes drawbacks to the young seedlings, but in order to be effective enough against early frosts, the shelter stand should be comparatively dense.

The PDF includes a summary in English.

• Leikola, E-mail: ml@mm.unknown
• Pylkkö, E-mail: pp@mm.unknown

The technical term reduced yield stands – sometimes reduced yield soils – is widely used in the Finnish forest literature. However, there is no clear definition of the notion reduced yield and no explanation of how this notion is measured in the classifications. Some committee reports and bill proposals and some laws and statutes use the reduced yield forest soil. No definition can be found. In my opinion the term reduced yield forest soils ought to be completely abolished until forest soil experts can perhaps define what it means, if they consider such a concept useful.

Explanations for the term reduced yield stands can be found in the descriptions of classification systems of stands. According to them, the criterion is partly silvicultural (site, species of tree), partly mensurational (volume, sometimes growth), partly economic. No explanation is found as to how the economic aspect is measured, nor about the limit of a full yield and reduced yield.

In my opinion such a term is confusing. I therefore suggest that the term reduced yield stand ought to be abolished. If something is needed instead, I suggest the term understocked stand, defined as a growing stock under certain percentage of a fully stocked stand. The notion of economic reduced yield cannot be generally tied to certain silvicultural and mensurational characteristics of stands. The economic aspect of certain kinds of stands may differ, depending on the owner of the forest and his economic situation, the location of the forest, the composition of the whole forest ownership unit, etc.

The PDF includes a summary in English.

• Saari, E-mail: es@mm.unknown

The present paper is a preliminary report of a project designed to determine the order of profitability of various forest improvement measures – seeding and planting, drainage, and fertilization – in various types of stands and in different parts of the country on drained peatlands. Sample plot data on the effect of draining on increment was derived from areas drained 28– 36 years ago. The study was carried out in the southern half of Finland.

The observations on increment changes are based on two measurements of the sample stands 12 years apart. Supplementary calculations indicate that the stands on drained peatland, depending on site quality and tree species, have either continued to grow like mineral-soil sites of similar fertility or have somewhat increased their growth rate.

The effect of draining intensity was studied using strip measurements. It was found that both the total amount of wood produced (current stand + cutting removal + natural removal) and the current annual volume increment for the 5-year period systematically decrease as the ditch interval increases. The decrease is, however, relatively slight. In Eriophorum vaginatum pine swamps, the total amount of wood produced and the increment show a decrease of ca. 20% with an increase in ditch interval from 20 to 60 metres. In other sites, the decrease is ca. 5-10%

It can be concluded that if the increase in ditch interval do not result in considerably poorer timber assortment distributions than indicates by stand production and increment, it is profitable to pan for a relatively large ditch interval and a slightly smaller than maximum wood production. Supplementary data and check calculations may cause some changes in these preliminary results.

The PDF includes a summary in English.

• Seppälä, E-mail: ks@mm.unknown

The present study gives an account of the dependence prevailing between the actual volume of timber crops, and, on one hand, the sum of cylindrical volume of individual trees, and on the other hand, the utilization per cent. This dependence is rectlinear if the volumes in question are calculated on a running-foot basis. The cylindrical volume of individual trees was calculated on the basis of cross-section area at breast height and the height of the trees. On the basis of the results, it seems that it is possible to simplify timber measurement.

The PDF includes a summary in English.

• Nisula, E-mail: pn@mm.unknown

An explorative study on wind conditions in a well-managed Scots pine (Pinus sylvestris L.) stand was made in Southern Finland. The wind velocity was recorded continually with two cup anomometers from April to August, 1964. The two levels used were 2 m and 9 m. The wind velocity was lower at 2 m than within the canopy at 9 m. The dependence on the absolute wind velocity at 9 metres was logarithmic. The wind velocity did affect the difference between daily minimum temperatures at the two levels; the difference in the maximum temperatures was affected only in May and August.

The PDF includes a summary in English.

• Leikola, E-mail: ml@mm.unknown

The aim of the study was to compare the newly introduced brush cutter to ordinary hand tools in clearing of cutting areas and thinning young birch (Betula sp.) and Scots pine (Pinus Sylvestris L.) stands. Working with a prototype of the brush cutter, Brushmaster, reduced the total working time by 15-20% compared to bush knife and axe, in spite of the cutter’s weight. At rainy weather the advantage of Brushmaster dissapeared because of clogging of the cutter’s air filter. The prototype proved to be more effective when clearing a cutting area, and hand tools seemed to be faster if damage to the remaining trees have to be avoided.

In addition, eight either lighter or more effective motor saws or brush cutters were compared mutually, and the effect of motor effect, weight, handiness, arrangement of suspenders and handles is discussed. With these improved types of the cutter it was possible to shorten the total working time in thinning of stands (mostly clearing of young stands) to 30-40% of the total working time compared to hand tools. The most modern saws appeared to be best adapted in clearing cutting areas.

The article includes an abstract in English.

• Sirén, E-mail: gs@mm.unknown

The article reviews the occurrence of damage causes by elk (Alces alces L.) in young Scots pine (Pinus sylvestris L.) stands established by direct seeding in the Ostrobothia region in Finland. The data was collected by random sampling, and consists of 110 sample plots in pine stands, established in 1930-1944.

Signs of elk damages could be observed in 20% of the stands. In more than half of the damaged stands pine seedlings were damaged by elk, on the rest of the stands the damage was targeted to hardwood saplings only. With the present density of elk population, the damage has an insignificant bearing upon the development of pine seedling stands in Ostrobothnia. The weaknesses of silvicultural state of the stands have been caused by other factors than elk.

Silviculturally weak stands were more liable to elk damage than strong ones. The occurrence of elk damage was more usual in stands with hardwood mixture than in pure pine stands. Especially goat willow, mountain ash and aspen, but also to some decree birch, seem to attract elk. Those factors that promote hardwood growth: fertility of the site, swampiness and the presence of seeding hardwoods in the area, increase the stand’s liability to elk damage.

The article includes an abstract in English and a summary in Swedish.

• Yli-Vakkuri, E-mail: py@mm.unknown

Silva Fennica Issue 69 includes presentations held in 1948-1950 in the fourth professional development courses, arranged for foresters working in the Forest Service. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.

This presentation describes the development of standards for timber grades in Finland, and the present quality grades of raw timber for timber sales from state forests.

• Pöyhönen, E-mail: vp@mm.unknown

Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service. 

This presentation describes inspection of forest regeneration of mineral soil forest types and drained peatlands, and inspection of ditches. 

• Ahola, E-mail: va@mm.unknown

The issue 39 of Silva Fennica includes presentations held in professional development courses in 1935 that were arranged for foresters working in public administration. The presentations focus on practical issues in forest management and administration, especially in regional level.

This presentation describes the forest management work in the state forests.

• Ahola, E-mail: va@mm.unknown

Silva Fennica Issue 39 includes presentations held in professional development courses in 1935 that were arranged for foresters working in public administration. The presentations focus on practical issues in forest management and administration, especially in regional level.

This presentation describes how to make a document required in Metsähallitus (Forest Service) when a stand is marked for harvesting.

• Sopanen, E-mail: es@mm.unknown

Regeneration of large open areas in dry mineral soil forest sites that usually grow Scots pine (Pinus sylvestris L.) have several problems. For instance, soil frost, snow, ground vegetation and dryness can prevent germination and growth of seedlings. The damages caused by insects and fungi in seedlings of a large burned area in Siikakangas in Southern Finland was studied. A forest fire burned the area nearly completely in 1909, and 310 hectares have been sowed or planted with mostly Scots pine during the following years. Minor areas have been regenerated with Pinus montana Noll, Pinus excelsa Lamb., Pinus murrayana Balf. and Larix sibirica Ledeb.

No completely healthy pine seedling stands could be found in the area. About 41% of the seedlings in the sample plots were damaged. The most common causes for damage were Evetria resinella (now Retinia resinella L.), Luperus pinicola (now Calomicrus pinicola (Duft.)), Pissodes notatus (now Pissodes castaneus Degeer), Evetria turionana Hb. and Hylobious abietis L. The most usual fungal disease was Lophodermium sp. Evetria resinella caused damages in all the area. Evetria turionana, Pissodes notatus and Hylobius abietina were found in the older seedling stands. Other damages were more localized. The slacks in the terrain seemed to have most damages, the original cause being probably soil frost. Some damages, as Lophodermium, were related to the density of the seedlings, especially in the sown areas. Cleaning of seedling stands could decrease these damages. Planting seems to have succeeded better than patch sowing.

The PDF includes a summary in German.

• Kangas, E-mail: ek@mm.unknown

One of the difficulties in constructing growth and yield tables has been to determine which of the sample plots growing the same tree species and belonging to the same forest site type, with reference to the quality of stands, have to be included in the same growth series.

New growth and yield tables for the most important tree species were constructed in Finland in 1916–1919, using new principles that aim at avoiding some of the common weaknesses. There were two main differences to the earlier work. First, the site quality class (forest site type) was determined for each sample plot when the sample plot was measured, independently of the stand occupying the site. In this way it was possible to treat the sample plots of each site as an independent group from the beginning, and so that the quality classes were the same for all the tree species. Second, mathematic-statistical methods were used to deduct the so-called stem frequency distribution series, when studying which of the sample plots of the same quality class belong to the same growth series. They represent the average number of stems of the different diameter classes. A more detailed description of the method used to create the growth and yield tables is published in Acta Forestalia Fennica no. 15.

In the PDF is included a summary in Finnish.

• Ilvessalo, E-mail: yi@mm.unknown

Examination of stands developed under natural conditions can be used to provide basis for comparison for study of the development and yield of stands treated with intermediate fellings. In Finland, the first investigation and the yield and the structure of natural normal stands were published in 1920. This investigation on development and yield of the natural forests of Kainuu in southeastern Northern Finland is based on 92 sample plots on three forest types; Empetrum-Vaccinium type (EVT), Empetrum-Calluna type (ECT) and Vaccinium-Myrtillus type (VMT).

The Scots pine (Pinus sylvestris L.) sample plots represented variation of age classes for construction of mean development series. The Norway spruce (Picea abies (L.) H. Karst.) of the region are so old that development series could be obtained only for dominant trees based on stem analysis.

The average development of Scots pine stand on EMT type within the region is on average more rapid and the yield in cubic volume quantitatively larger and structurally better than that on ECT type. Self-thinning during the early decades of EVT is slower. The pine stands are denser in the age of 70 in Kainuu compared to Central Northern Finland, but the development and yield are similar.

The development, yield and structure of natural Norway spruce stand on VMT proved considerably inferior to the average level of pine stands on EVT, and to a major part on ECT. The mixed pines on spruce sample plots have developed better than spruces of equal age. Spruce stands on VMT in the area developed markedly better than Geranium-Dryopteris-Myrtillus (GDMT) in Central Northern Finland.

It seems that a spruce stand seems to require more fertile site type in north than in the southern part of Finland. These fertile types are relative rare in the north. In the region, the best results are received with pine. As a rule, also the yield of birch (Betula sp.) is poor in the region.

The PDF includes a summary in English.

• Ilvessalo, E-mail: yi@mm.unknown

The objective of the study was to establish the influence of the founding density of a stand and the intensity of intermediate cutting on the quality of pine saw logs stems, primarily on their branchiness. Measurements were carried out in 68 Myrtillus-type and 32 Vaccinium-type Scots pine (Pinus sylvestris L.) stands. The quality of 1,982 sample trees was assessed.

According to the results, the branchless part of the stem is longest in the older age classes of trees. In all age classes the percentage of the branchless part is highest in medium sized stems. The relative height of the crown limit is greatest in small diameter classes and continues as the thickness of the tree increases. The crown is longer in the thicker tree. The grade of the butt log is on average highest in medium sized stems. Knottiness of a log made it unsuitable for a saw log only among the thickest stems. The relative share of the u/s grade decreased as the thickness of the trees increased.

From the point of view of early development of the trees it was concluded that in all age classes the branchless part is the shorter the faster the tree has grown in diameter when it was young. Also, branches of the butt log are the bigger the faster the tree has developed when it was young. The grade of the butt log improves as the thickness of the annual rings diminishes.

To produce good quality sawn timber, the pine stands should be established dense, and the first thinnings should be delayed as much as possible. The best time for the thinning would be when the diameter of the dominant trees at stump height is 12–15 cm and when all the branches have died on the length of the butt log. After the first thinning, comparatively intense intermediate thinning may be applied.

The PDF includes a summary in English.

• Heiskanen, E-mail: vh@mm.unknown

The aim of this work was to study, on the basis of material published earlier (Heikurainen 1959), the effect of temperature on stand increment, to find out if there is any differences between Norway spruce (Picea abies (L.) H. Karst.) and Scots pine (Pinus sylvestris L.), and to study the effect of site quality on the relationship between stand increment and temperature. The calculations were based on data collected from 396 sample plots on drained peatlands in different parts of Finland.

There seemed to be no differences due to tree species or site quality in the relative amounts of growth under different climatic conditions. Thus, differences in the absolute growth between poor and fertile sites are noticeably smaller in Northern Finland than in Southern Finland. The author suggests that this implies that the lasting maximal increase of growth which can be produced, for instance, by using soil-improving agents must be less in unfavourable conditions than in favourable.

• Heikurainen, E-mail: lh@mm.unknown
• Seppälä, E-mail: ks@mm.unknown

In this paper the development of sown Scots pine (Pinus sylvestris L.) seedling stands into forests is studied. The material was collected in stands sown in 1930–1940 in private forests in the Forestry Board districts of Central and Northern Ostrobothnia. The 119 areas, including both burned and other areas, were studied in 1955.

Most seedings had been carried out on relatively poor soils, mostly representing Vaccinium and Calluna type forests. 71% of the areas consisted of large forest fires, mostly from 1933. The most burned areas did not have seed producing trees nearby. The other sown areas were in general small, 1–2 ha, and near forests capable of producing seeds. The species of previous tree generation, in the older areas mostly pine and in the younger areas Norway spruce, affected tree species composition of the new tree generation.

Over 90% of the burned areas were in silviculturally good or satisfactory condition, while the main part of the other sown stands was in fair or poor condition. Weeding and thinning had been done only in the oldest stands. Most stands had been left untended. Natural new trees often competed with the sown pines, and cull-trees and border forest increased natural regeneration in the areas. In Calluna type the poor soil limited regeneration and growth of broadleaf trees. The worst competitors were naturally regenerated pine seedlings both on Calluna and Vaccinium type. On Vaccinium type also birch and sometimes also aspen (Populus tremula L.) competed with sown pine. On better sites and paludified areas competition by broadleaf trees increased. The rhythm of development of broadleaved trees is so different from pine that only those broadleaved trees that are formed in the stand when the pine seedlings are larger can develop harmoniously with pine. Due to the harmful competition, the seedling stands should be tended early on. In addition, it may be advisable to abandon the practise to leave trees on sowing areas.

The PDF includes a summary in Finnish.

• Yli-Vakkuri, E-mail: py@mm.unknown

The article presents the background of increment calculations and periodic measurements of forests, as well the historical development of increment calculations in North-America, Middle-Europe, Scandinavia and Finland. The measurements and calculations are presented for individual trees, for a forest stand and for the total resource of a normal forest stand.   

The practice of increment calculations has still some problems regarding the measurements of standing and harvested trees. The article discusses some ways to overcome the problems. 

• Kuusela, E-mail: kk@mm.unknown

There are contrary opinions on the ability of Scots pine (Pinus sylvestris L.) seedlings to withstand oppression by hold-overs and recover after their felling. The recovery potential of oppressed pine stands in Southern and Northern Finland was studied using two kinds of material, fully recovered Scots pine stands and stands recently released. The volume and volume increment of the stand were measured, and the health of the sample trees was determined.

The study showed that those released pine stands that had been in oppressed state very long (25-60 years) had recovered after clear-cutting. After the release the stands grew at first slowly, but after recovery at about the same rate as natural normal stands of a similar height. The smaller, younger, and less stunted the seedlings were when they were released, and the better the site, the faster was the recovery. At the base of released pine stands various defects was detected. When the trees were released, the defects decrease their technical value. A heavy partial cutting had generally a disadvantageous effect on the stand. Recovering seedlings were found clearly to hinder the development of younger seedlings nearby. This inhibition seemed to be a result of the rapid spread of the root system of released pine trees.

The PDF includes a summary in English.

• Vaartaja, E-mail: ov@mm.unknown

The quality of birch (Betula sp.) stands in Perä-Pohjola in Northern Finland is low due to the harsh environment, unsuitable sites for the species and unsatisfactory silvicultural state. A total of 236 sample trees were felled and measured in 8 sample plots. The trees were over 80 years old.

Only third of the stand volume of birch in the stands had adequate quality for merchantable timber. This is due to birch growing often in sites unsuitable for the species, the low density of the stands, the small average size of stems, and the low amount of large sized trees. These problems may contribute to the fact that birch seem to be susceptible to decay. The trees have often grown from sprouts, which leads often to poor stem form and decay. The volume and quality of both pure and mixed birch stands was sufficient only in the most fertile sites. Also, decay was more common in poor sites.

The PDF includes a summary in German.

• Tikka, E-mail: pt@mm.unknown

The tendency of successional development from young vegetation, rich in species and exposed to chance factors, towards regular plant communities, is found all around the world. Thee ecological groups of trees seem to be present in all forest regions in the world, namely the pioneer and the climax species, and a group of pre-climax species that can be ecologically either near the pioneer or the climax species. The succession of tree species in forest always leads to a climax stand, determined by climate, quality of soil and the mutual biotical strengths of the tree species in the region.

This division into ecological groups greatly facilitates choosing among different methods of treating stands and understanding the silvicultural methods of foreign regions. Stands formed by species of the same group must follow the same lines in their silvicultural treatment. For instance, mixed stand consisting of both pioneer and climax species represents a transition stage, in which the climax species strive for dominating position, and preservation of pioneer species is difficult. This indicates the broad lines for management of the stand. Also, regeneration methods of pioneer and climax species must be different. Studying the succession of natural forests can be used as a means to reach the highest possible silvicultural level. This is one reason why the preservation and study of virgin forests still in existence is indispensable.

The PDF includes a summary in Finnish.

• Kalela, E-mail: ek@mm.unknown

Department of plant pathology of University of Helsinki conducted studies in the surrounding forests of Viikki test farm and on the Hyytiälä forest training station to find about occurrence and spore production of pine needle rust.

The damages have been minor. Most often the infection did not cause yellowing of the needles, only individual needles might have dried up. There were no dead young trees.

The spore production was strongest at the more fertile sites, with abundant occurrence of cow-wheat (Melanpyrum spp.). The infection caused more harm on the cow-wheat than on the pines. In many cases the foliage died prematurely. Melanpyrum spp. were more strongly infected on sites with more light. However, there was no difference found with the fertility of the site.

The PDF contains a summary in Finnish. 

• Pohjakallio, E-mail: op@mm.unknown
• Vaartaja, E-mail: ov@mm.unknown

The regeneration of forests in Hämeenkangas area in Southern Finland has been difficult due to various damages from the middle of the 1800s. Few seed trees were left in the area, and artificial regeneration has been used since 1880s. The area became an experimental area of the Forest Research Institute in 1924. The aim of the study was to survey the area before it was transferred to the Finnish Defense Forces.

The original Scots pine (Pinus sylvestris L.) forest of the esker area suffered from many forest fires. The total area is 13,000-14,000 ha, of which the experimental forests of Forest Research Institute cover 6,000 ha. The area is dry upland forest, and drought affects the survival of germlings. Soil frost is a major cause of loss of young seedlings. Sowing method affects the early development of the seedlings. Band sowing proved to be the best method regarding the soil frost. A total of 39 different harmful insect species, 8 pathogen species and 7 other causes of damages have been detected in the area.

The development of seedling stands follow a certain pattern, reported also in other studies. Many of the pine seedling stands develop well until they reach a certain height. After that seedlings begin to suffer from damages, but after reaching another stage develop normally. The damages affect the height growth of the seedlings. Some common damages are caused by Pissoides weevils, needle damages caused by certain beetles, shoot damages by Evetria resinella, and pine blister rust (Peridermium pini and Cronartium flaccidum).

• Kangas, E-mail: ek@mm.unknown

The plant populations of Finnish open bogs are typically formed of two layers. The layers normally consist of one or rarely two species. The structure of plant populations in open bogs is a consequence of the development where determining factors are different site requirements of the species, and the differences in the biotic vitality and capacity for reproduction.

Phytogenesis should be taken as a basic unit for describing the plant societies or vegetation of treeless bogs. However, acknowledging the sub-populations may be of advantage for describing the ecological, genetic and regional characters of open bogs.

The basic classification of open bogs must be done based on the ground layer. The more detailed classification follows mostly based on field layer, partly also based on the ground layer.

The PDF contains a summary in Finnish. 

• Paasio, E-mail: ip@mm.unknown

The article presents the characteristics of different vegetation areas (meadows and peatlands) by their distinctive vegetation. The study area is by the Barents Sea and is the northernmost part of continental European Russia. Different sites are classified by plant communities and/or vegetation units.

The article continues on the second PDF-file. 

• Kalela, E-mail: ak@mm.unknown

Natural regeneration has been common in Northern Finland, where forest fires have been usual, and the large areas make artificial regeneration expensive. The regeneration, and for instance tree species composition and density of the stand, cannot been controlled. In Northern Finland there is little demand for Betula sp. which is often abundant in the burnt areas. The unburned forests are generally Scots pine (Pinus sylvestris L.) or Norway spruce (Picea abies (L.) H. Karst.) dominated mixed forests with single Betula sp. trees.

The fire destroys birch for the most part in the Vaccinium site type, but the surviving trees produce enough seeds to regenerate the areas. The largest trees of Scots pine usually survive the fires. Pine has good seed years in the north only every 8^th or 10^th year. Spruce is totally destroyed in the forest fire and the seedlings grow poorly as primary species. The seedling stands are usually dominated by Scots pine and birch, but birch seedlings grow in batches, and do not hinder growth of pine. The drier Calluna site type stands are dominated by Scots pine. Birch seedlings may be abundant in the beginning, but most of them do not survive. Abundant emergent pine trees prevent the growth of seedlings especially in the dry site types, and they should be thinned to guarantee regeneration. Sowing results are better few years after the fire. The birch seedling should be removed from the seedling stands.

The PDF includes a summary in German.

• Sarvas, E-mail: rs@mm.unknown

Shifting cultivation, practiced earlier in Finland, was beneficial for grey alder (Alnus incana (L.) Moench). It can produce seeds early and the early growth of the seedlings is fast. Areas where shifting cultivation was intensive, the areas next to the fields were pure alder stands, next circle was Betula sp. dominated, beyond that could be found Scots pine (Pinus sylvestris L.), and finally Norway spruce (Picea abies (L.) H. Karst.). When shifting cultivation ended, Norway spruce became more common. Many young mixed stands had Norway spruce undergrowth and alder overgrowth. The aim of the study was to find out how the stands develop to spruce dominated stands, and how they should be managed.

The density of spruce undergrowth affects the further development of both spruce and alder. The number of alder stems decreases the faster the denser the spruce undergrowth is. Alder overgrowth slow down the early diameter and height growth of spruce compared to pure stands. Also the diameter and height growth of alder remains smaller in mixed stands. The basal area of spruce develops slowly in the beginning, increases significantly by the age of 30, and surpasses the growth of pure spruce stands in Oxalis-Myrtillus site type. Thus, Norway spruce do not suffer from growing in the undergrowth. In the first years, fast growing alder seedlings limits growth of ground vegetation and protects spruce seedlings from frost.  Later thinning or removal of alder benefits spruce growth. The density of spruce undergrowth decides how much alder can be leaved in the stand. If the spruce undergrowth is thin, more alder can be left in the stand.

The PDF includes a summary in German.

• Kalela, E-mail: ek@mm.unknown

About 40% forest in Finland are mixed stands that have birch (Betula pubescens and B. verrucosa) as one of the species. The aim of this research was to study the structure of root system of birch and compare it to the other main tree species in Finland.

The root systems were dug out and measured in 28 sample plots in Southern and Central Finland, representing different forest site types. Birch roots correspond 30‒100% of the volume of the stem, the largest root systems being in the sandy soils or peatlands. Also the longest lateral roots can be found at these sites. The size variation of root system of birch is larger than in Scots pine (Pinus sylvestris L.), and the vertical root system is in general smaller in birch. Birch seems to be better than pine able to adapt its root system to the existing conditions. The smallest root systems were found in the good forest site types, but the roots grow in the good sites denser than in the poor sites. The lateral roots of the main tree species in Finland, birch, Scots pine and Norway spruce (Picea abies (L.) H. Karst.) grow in different depths, which decreases the competition between the species. This finding gives support to cultivation of mixed stands.

The PDF includes a summary in English.

• Laitakari, E-mail: el@mm.unknown

Water and amount of light are the most important growth factors and the article discusses their relationship. It is knows that the more space is needed by a tree the worse the site is. The number of stems varies between tree species. Common understanding is that amount of light is decisive to self-thinning and regeneration of a stand. On a good site the adequacy of water may substitute the lack of light. However, the fertility of soil and moisture content is more important than light.

The question of the importance of light, soil fertility and water content, as well as their relationship is important when optimizing the forest management.       

The PDF contains a summary in Finnish.  

• Aaltonen, E-mail: va@mm.unknown

In a natural state growing closed forest the amount of timber grows yearly. However, in the same time the timber volume also decreases when part of the trees die in the competition for light and other growth factors.

There are many interactive functions and characteristic that influence the growth and drain in a stand. These can be illustrated as mathematical models. The article discusses a set of models.

The volume 34 of Acta Forestalia Fennica is a jubileum publication of professor Aimo Kaarlo Cajander.

• Lönnroth, E-mail: el@mm.unknown

To be able to exactly describe the similarities and differences of vegetation in certain areas, classifying the vegetation only in communities or formations is not enough. Therefore more classes are needed. The classification according the horizontal layers is based on the heights of plants and their relations to each other. Every population in one community has own area of height which extends to horizontal direction.

In comparison with populations the vegetation horizons create a biologically validated comparison of different vegetation groups and their parts. Defining the populations and vegetation horizons creates a division and systematization of plant communities on an ecological basis.       

The volume 34 of Acta Forestalia Fennica is a jubileum publication of professor Aimo Kaarlo Cajander.

• Kujala, E-mail: vk@mm.unknown

Natural regeneration has been studied so far mostly on the perspective of regeneration, while the production capacity of the seed crop stand has been of little interest in the earlier studies. This paper studied volume growth of the seed trees and the economic impact of this regeneration method both in a literature review and measuring sample stands consisting of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) dominated stands and mixed forests in Norway.

It is concluded that in the most cases the seed tree stands give better pulpwood and timber yield than the stands in the average. The reason might be that the elite stems can better use their production capacity, and, thus, reach bigger and more valuable dimensions. Even though the growing stock diminishes in the seeding felling, the growth of the stand does not decrease or decreases only little. At the same time, the quality growth improves. In addition, the new tree regeneration is achieved usually without extra costs, and there is no unproductive time period in the stand.

• Barth, E-mail: ab@mm.unknown

There has been earlier publications on calculating the standard error of the strips. However, they have been calculated with very small amount of data. In the article the confidence of standard error will be examined throughout. The article is a response to the article written by A. Langsaeter on the theme.  

• Lindeberg, E-mail: jl@mm.unknown

Study is based on the strip-wise survey of forests in southern Finland. From that information the 30 sample plots were chosen, 10 of each of most typical forest site types, MT, VT and CT. The stands are of different ages and development classes, varying from 14 to 159 years.

The article discusses the earlier literature on the factors effecting forest stands and presents the data in detail. The results section is divided into paragraphs on number of stems of the stand, height, crown, breast height diameter, basal area and volume. The statistical numbers are calculated and presented.    

• Lönnroth, E-mail: el@mm.unknown

• Valle, E-mail: kv@mm.unknown

The study is based on the results of the soil studies by Valmari (1921) and the growth inventories of respective areas. The aim is to show the connection of soil fertility (nutrient content) and forest growth with means of correlation calculations. The examined nutrients were nitrogen, calcium and phosphorus, also the electrolyte content was studied.

• Ilvessalo, E-mail: yi@mm.unknown

The article features a critical observation on used methods for calculating the errors and a trial to improve it. The article describes the calculation method used in Sweden, county of Värmland and another method used by Ilvessalo and developed by Cajanus. The shortcomings of these models are discussed. An improved calculation is presented.  

• Lindeberg, E-mail: jl@mm.unknown

The article is a presentation given by the author on occasion of visit from Austrian timber industry and foresters (August 17^th 1923), and again for the German visitors (August 20^th 1923) in Punkaharju, Finland. The speech gives an overview of Finland’s forest resources region by region starting from far north. Also the most important uses and changes of forest as well as forest industry locations are mentioned. 

• Cajander, E-mail: ac@mm.unknown

The article presents definitions for central concepts that are commonly used in the research of plant topography. The discipline has close connections to defining and classifying forest types.

The concepts defined involve concrete and abstract concepts linked to vegetation conditions and to habitat. 

• Cajander, E-mail: ac@mm.unknown

A method was developed for relative radiometric calibration of single multitemporal Landsat TM image, several multitemporal images covering each other, and several multitemporal images covering different geographical locations. The radiometrically calibrated different images were used for detecting rapid changes on forest stands. The nonparametric Kernel method was applied for change detection. The accuracy of the change detection was estimated by inspecting the image analysis results in field.

The change classification was applied for controlling the quality of the continuously updated forest stand information. The aim was to ensure that all the manmade changes and any forest damages were correctly updated including the attribute and stand delineation information. The image analysis results were compared with the registered treatments and the stand information base. The stands with discrepancies between these two information sources were recommended to be field inspected.

• Varjo, E-mail: jv@mm.unknown

The accompanying collective research report is the result of the research project in 1986–90 between the Finnish Academy and the former Soviet Academy of Sciences. The project was organized around common field work in Finland and in the former Soviet Union, and theoretical analyses of tree growth determining processes. Based on theoretical analyses, dynamic stand growth models were made and their parameters were determined utilizing the field results.

Annual cycle effects the tree growth. Our theoretical approach was based on adaptation to local climate conditions from Lapland to South Russia. The initiation of growth was described as a simple low and high temperature accumulation driven model. The model was linked with long-term temperature data.

Analysis of field measurements of CO₂ exchange showed that irradiance is the dominating factor causing variation in photosynthetic rate in natural conditions during summer. The penetration of irradiance into Scots pine (Pinus sylvestris L.) canopy is a complicated phenomenon. A moderately simple but balanced forest radiation regime sub-model was constructed.

The common field excursions in different geographical regions resulted in a lot of experimental data of regularities of woody structure. The water transport seems to be a good common factor to analyse these properties of tree structure. The produced theoretical and experimental material was utilized in the development of stand growth model that describes the growth and development of Scots pine stands in Finland and the former Soviet Union. The core of the model is carbon and nutrient balances. This means that carbon obtained in photosynthesis is consumed or growth and maintenance and nutrients are taken according to the metabolic need. Since the fundamental metabolic processes are the same in all locations, the same growth model structure can be applied in the large range of Scots pine. The model enables the analysis of geographical differences in the growth of Scots pine. The present approach enables utilization of structural and functional knowledge gained in places of intensive research, in the analysis of growth and development of any stand.

• Hari, E-mail: ph@mm.unknown
• Ross, E-mail: jr@mm.unknown
• Mecke, E-mail: ph@mm.unknown

A nonlinear programming algorithm was combined with two individual-tree growth simulators consisting of distance-independent diameter and height growth models and mortality models. Management questions that can be addressed by the optimization model include the timing, intensity and type of thinning, rotation age, and initial density. The results were calculated for Norway spruce (Picea abies (L.) H. Karst.) stands on Oxalis-Myrtillus site in Southern Finland, where the stand density after clearing of a seedling stand is about 2,000 trees/ha.

The optimum thinning programs were characterized by late first thinnings (at dominant height of 15–17 m) and relatively high growing stock levels. It was optimal to thin from above, unless mean annual increment was maximized instead of an economic objective. In most cases, the optimum number of thinnings was two or three. Compared to a no-thinning alternative, thinnings increased revenues by 15 –45% depending on the objective of stand management. Optimum rotation was strongly dependent on the interest rate.

Hooke and Jeeves’ direct search method was used for determining optimum solutions. The performance of the optimization algorithm was examined in terms of the number of functional evaluations and the equivalence of the objective function values of repeated optimizations.

The PDF includes a summary in Finnish.

• Valsta, E-mail: lv@mm.unknown

The study was carried out at Padasjoki, Southern Finland, where moose (Alces alces L.) density on the winter range had been over 1.5 individuals/km². Moose browsing intensity, expressed in terms of number of twigs eaten and biomass used, increased with stand density (biomass). Total biomass consumption (dry weight) per sample plot and per sapling. The number of bites increased, but the percentage biomass removed did not differ when stand density increased. A relatively large bite size was observed on the plots of low stand density. The quantity of food, which on average was of relatively low quality, was obviously important due to the benefit gained through reducing the search time.

The nutritive value of the browse, expressed in terms of chemical compounds indicating low food digestibility, was lower in the dense than in the sparse Scots pine stand. However, the amount of crude protein and arginine were relatively high in the dense stand. We concluded that shading affected the nutritional status of saplings on high density plots.

Although the biomass removed by moose per sapling was high for low density plots, the remaining biomass was larger than that on the high-density plots owing to the relatively large twig biomass of saplings. The number of saplings per hectare without main stem breakage increased significantly as stand density increased.

The PDF includes a summary in Finnish.

• Heikkilä, E-mail: rh@mm.unknown
• Mikkonen, E-mail: tm@mm.unknown

The establishment of moose (Alces alces L.) winter feeding sites, their utilization and their effect on damage to young Scots pine (Pinus sylvestris L.) plantations was studied in Ruokolahti-Imatra area in Eastern Finland in 1987–89. During the period, the density in the area was about 3–5 moose/ 1,000 ha.

Six feeding sites were established by fertilization, offering mineral lics and the tops of aspen and Scot pine and by salting the tops of pine. The moose preferred the feeding site to control areas during both summer and winter. In winter browsing was very heavy, especially in those areas located in or close to traditional wintering areas. In winter no moose were seen in the summer habitats.

The extent of, and fluctuations in moose damage were studied in 47 Scots pine plantations in the immediate surroundings of the feeding sites (29 plantations), control areas (18 plantations) and also 68 randomly selected pine plantations. Before the experiment began in 1987 four plantations had been seriously damaged. During the study period only one plantation was seriously damaged. However, it could not be conclusively proved that damage to the pine plantations had been reduced as a result of the feeding sites. The results of the study can be put into practice elsewhere to create better living conditions for moose in their winter habitats. However, the food offered at the feeding site should be in the right proportion to the number of animals wintering in the area, so that the risk of damage to nearby plantations would be kept as small as possible.

The PDF includes a summary in Finnish.

• Lääperi, E-mail: al@mm.unknown

At the beginning of the investigation period the total biomass of the Scots pine (Pinus sylvestris L.) stands on the ordinary sedge pine mire was 48 t/ha. The biomass of the mixed stands of Scots pine and birch (Betula pubescens Erhr.) on the herbrich sedge pine mire was 91 t/ha, out of which 60% was from pine. The biomass of the Norway spruce (Picea abies (L.) H. Karst.) on the Vaccinium-Myrtillus spruce mire was 148 t/ha. The average annual net increment of the stand biomass was 5.8 t/ha in the unfertilized pine stand and 6.7 t/ha in the NPK and micronutrient fertilized one during the six-year investigation period. The corresponding figures in the mixed stand were 7.2 t/ha and 7.6 t/ha. The net increment of the biomass in the unfertilized spruce stand was 6.9 t/ha and in the fertilized 8.4 t/ha. A considerable proportion of the net increment was lost to the ground as litter in all stands.

The nitrogen, phosphorus, potassium, magnesium, iron, manganese, zinc, copper and boron cycles were investigated. The annual nitrogen uptake from the soil was 26–42 kg/ha, that of phosphorus 2.5–3.4 kg/ha, potassium 4.5–12 kg/ha, calcium 12–29 kg/ha, magnesium 2–4 kg/ha, iron 1.4–6.6 kg/ha, manganese less than 2 kg/ha and the other nutrients only some grams. Only part of the fertilized nutrients was fixed in the stand.

The PDF includes a summary in Finnish.

• Finér, E-mail: lf@mm.unknown

The applicability of five mathematical programming methods, namely standard linear programming, parametric programming, goal programming, mixed integer programming and integer programming is discussed as a planning tool for the choice of wood procurement method.

Theoretically, the goal programming approach seems to be the best routine for mathematical handling of problems related to wood procurement. The parametric approach must include enough large post-optimality analysis routine. If the effect of the variables expressed with different measures is to be studied, interpretation of the economic information given by the approach becomes a problem. The other drawback is that the approach does not allow determination of the hierarchy of the goals objectively as they depend on the subjective preferences of the decision maker.

From the practical point of view, standard linear programming is the best method if the objective function can be formulated in economic terms, for instance. If there are several goals to be attained or satisfied the best method is goal programming.

According to the sub-studies, every method under consideration can be used as a solution routine for the minimization of wood procurement costs. In cost minimization the best methods are goal programming and standard linear programming. The best method for harvesting system evaluation purposes is parametric because it allows varied cost calculations within a certain cost range. The best method for harvesting equipment investment planning is mixed integer programming with binary decision variables.

The more complicated and restricted the problem environment is, the better the mathematical programming approach will be, also in harvesting related problems.

The PDF includes a summary in English.

• Mikkonen, E-mail: em@mm.unknown

Analysis of sample tree data showed that the major part of the residual variation of the stem volume estimates occurs within the forest stands. The division of the residual variation into the variation within and between populations is the basis for the mean-square error formulae of the volume estimators. Efficiency of different sample tree measurement combinations has been studied by comparing the errors of the volume estimates to the sampling costs. The measurement of the upper diameter (d6) is of less value than is generally suggested.

The PDF includes a summary in Finnish.

• Kilkki, E-mail: pk@mm.unknown

An attempt was made in the study to determine the annual periods available for foliage spraying when cleaning Scots pine (Pinus sylvestris L.) dominated seedling stands. The study was made in nine experimental fields which were established in different parts of Finland. The spraying was applied throughout the growing season by DM, MCPA and Roundup. The results were inventoried one year after the treatments.

The results showed that there were big differences both in the destruction of hardwood sprouts and in the survival of pine seedlings due to the time period of the spraying. Threshold points were observed in the range of effect of DM and MCPA. By means of these it is possible to time the spraying treatments in such a way that there remains only slight damage to pine, but hardwood sprouts are destroyed totally. The results varied with Roundup so much, among other things due to rain, that such threshold points could not be determined. This preparation both had a milder effect on the hardwood seedlings and caused slighter damage to pine than the other preparations.

In Sodankylä in Northern Finland, the pines attained a good resistance to arboricides when the efficient temperature sum of the growing season was 550, but in Punkaharju in Central Finland only when it was 850. The seed provenance of the seedlings had an effect on the resistance. The threshold temperature sums of resistance in pine were on the average 70–74% from the long-term average number of degree days at the origin of the seed. The effect on the hardwood trees grew weaker as the long-term average was filled. Resistance of pine followed with a specific lag the lignification of the shoot and the ceasing of the growth of the needles.

The PDF includes a summary in English.

• Pohtila, E-mail: ep@mm.unknown
• Pohjola, E-mail: tp@mm.unknown

The paper is the final report of a study on the estimation of value increment and inherent variables of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) stands. The main aim was to obtain improved criteria for decision-making concerning the priority of stands for regeneration.

The construction of various estimation models and their reliability are discussed in detail. The study, together with some previous papers, has resulted in a system which on the basis of a number of easily assessed stand variables gives for the stands concerned the volume of stems, percentages of timber assortments, stumpage value, volume increment and value increment.

The following examples are given with regard to the practical application of the results, in addition to the determination of the relative maturity of stands: 1) The study of various trends in stand development; the comparison between the volume and value variables. 2) The estimation of timber assortments needed for a cutting budget, trees marked for felling etc. 3) The calculation of the value of forests.

The PDF includes a summary in English.

• Nyyssönen, E-mail: an@mm.unknown
• Ojansuu, E-mail: ro@mm.unknown

The report concludes a series of studies on the early development of young Scots pine (Pinus sylvestris L.) stands. The basis assumption made in the study series was that the within-stand light regime is the main driving force for total tree growth and its allocation of photosynthates for crown, stem and root growth. An individual tree growing in a stand under a varying light regime which is controlled by the stand structure, is the basic unit used in the study. The photosynthesis of an individual tree is determined by the light regime. The stand is formed from individual trees.

The model is applied in simulation of the growth and development of tree stands. Several computer runs representing various densities, height distributions and tree species mixtures were carried out. Potential application areas, properties of the model and future needs of investigations are discussed.

The PDF includes a summary in English.

• Hari, E-mail: ph@mm.unknown
• Kellomäki, E-mail: sk@mm.unknown
• Mäkelä, E-mail: am@mm.unknown
• Ilonen, E-mail: pi@mm.unknown
• Kanninen, E-mail: mk@mm.unknown
• Korpilahti, E-mail: ek@mm.unknown
• Nygren, E-mail: mn@mm.unknown

The aim of this investigation was to examine the dependence of stand volume and increment on different growth factors on drained peatlands drained 20 years ago. Measurements were made in 1977-79 on 35 sample plots in Central Finland on relatively poor pine bogs with a thick peat layer.

It became evident that the stand volume, increment and radial growth and growth development are primarily functions of groundwater depth. Groundwater depth is dependent, in the first place, on ditch depth and ditch condition. With regard to the variation in ditch spacing (ca. 35-70 m) under examination, the effect of ditch spacing on the stand was insignificant. As a practical recommendation it was concluded that ditches should be kept deep enough (> 70 cm) in order to maintain undisturbed stand development.

The PDF includes a summary in English.

• Heikurainen, E-mail: lh@mm.unknown

The paper concerns the estimation of the increment of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) stands in the southern half of Finland. For the methods based on stand tables, tree functions forecasting the annual increment of diameter and height during the next 5-year period are presented. The main results of the study, however, are the functions for the volume increment percentage of pine and spruce stands. The independent variables are: forest site type, tree species, stand age and volume, and mean diameter. The standard error of estimate is about 17% in the best functions. Calculations were made also with regard to the application of the results in growth estimation of large forest areas.

The PDF includes a summary in English.

• Nyyssönen, E-mail: an@mm.unknown
• Mielikäinen, E-mail: km@mm.unknown

The data used to control the developed methods are presented in chapter two. The third chapter presents the statistical analysis conducted. The factor analysis shows that tree stand characteristics must be divided into three factors to able the information needed for site indexing to distinguish different site classes. The phases of model development are presented and the results of calculations with control data are shown.   

The results indicate that for calculations of nature-normal forests the suitable amount of tree stand characters for creating the site indexing function, is three.  These are: number of trees (N), basal area of the stand (G) and mean height (H).

The PDF contains a summary in Finnish. 

• Kangas, E-mail: yk@mm.unknown

The article is the first part of a wider project aiming to study the measurement practices of stand’s height-over-age based classifications and to develop new methods for actually measuring the side index.  

The first part of the article presents the concepts of site indexing and discusses it with a mathematical model. The second part of the article examines the structure of site indexing system regarding the reliability of the method. Third part presents the factors affecting the method development and the fourth part discusses the possible site index classes and their characteristics.   

The PDF contains a summary in Finnish. 

• Kangas, E-mail: yk@mm.unknown

The purpose of this study was that of providing a long-term timber production model (Kilkki and Pökkälä 1975) with growing stock models. The paper is divided into two parts; the first is concerned with generation of the stand data through Monte-Carlo simulation. The growing stock of each stand was described by a DBH-height distribution. The necessary information on the relationships between the stand characteristics was derived from sample plots measured in the national forest inventory of Finland. A total of 1,500 Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies (L.) H. Karst), and birch (Betula sp.) stands, each comprising 100 trees were provided by simulation.

In the second part, models predicting the form factor, timber assortment distribution, and value of the growing stock were derived through regression analysis for each species of tree. The predicting variables included the form factor of the basal area median tree, basal area median diameter, and height in the form factor models. In the timber assortment and value models, the only predicting variable was the volume of the basal area median tree. The Matchcurve-technique (Jensen 1973) was employed in derivation of the regression models.

The PDF includes a summary in English.

• Kilkki, E-mail: pk@mm.unknown
• Siitonen, E-mail: ms@mm.unknown

Production of timber in forest stands is described by a production function. The variable inputs of the function are land and growing stock and the output is the annual value growth. The partial derivatives of this production function express the marginal productivity of the land and of auction function express the marginal productivity of the land and of the growing stock. These marginal productivities can be utilized for determination of the need of regeneration and thinning. The stand should be regenerated when the marginal productivity of the land falls below the annual rent of a unit area of open land and thinned when the marginal productivity of the growing stock falls below the annual rent of one unit of growing stock.

The PDF includes a summary in Finnish.

• Kilkki, E-mail: pk@mm.unknown

Disturbances caused by the European spruce bark beetle (SBB; Ips typographus L.) on Norway spruce (Picea abies (L.) H. Karst.), have increased immensely across Central and Northern Europe, and are expected to increase further as a result of climate change. While this trend has been noted in Finland, so far limited research has been published. To support proper SBB risk management in Finland, we compared stand properties between salvage loggings due to SBB damage during 2012–2020 (4691 cases) and spruce stands free of SBB damage. Also, we explored the role of landscape attributes as drivers of SBB damage. We considered the forest stand attributes of site fertility class, stand development class, soil type, stand mean diameter at breast height and mean stand age. Considered forest landscape attributes were the distance from SBB-damaged stands to the closest clear-cut, to previous-year SBB-damaged stands and to the previous-year wind-damaged stand. We used nationwide forest logging and forest stock data, and analysed forest stand attributes using chi-squared and Mann-Whitney U tests and landscape attributes using generalised linear mixed models. Based on our findings, the SBB didn’t damage stands randomly, but prevailed in mature stands (high age and high mean diameter at breast height), in herb-rich heath forest site types and in semi-coarse or coarse heath forest soil soils. We found correlation between the landscape variables and the number of salvage loggings, with a higher number of loggings due to SBB damage close to clear-cuts. Our results help to find risk areas of SBB damage.

• Pulgarin Diaz, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0000-0003-0554-8254 E-mail: alexander.pulgarin.diaz@uef.fi
• Melin, Natural Resources Institute Finland (Luke), P.O. Box 68, FI-80101 Joensuu, Finland https://orcid.org/0000-0001-7290-9203 E-mail: markus.melin@luke.fi
• Ylioja, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-0079 Helsinki, Finland https://orcid.org/0000-0002-8840-7504 E-mail: tiina.ylioja@luke.fi
• Lyytikäinen-Saarenmaa, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0000-0003-1884-3084 E-mail: paivi.lyytikainen-saarenmaa@ef.fi
• Peltola, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: heli.peltola@uef.fi
• Tikkanen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland https://orcid.org/0000-0002-3875-2772 E-mail: olli-pekka.tikkanen@uef.fi

Uneven-aged forests set certain challenges for cut-to-length harvesting work. It is a challenge to cost-effectively remove larger trees while leaving a healthy understory for regrowth. The study’s aim was to evaluate productivity and costs of harvesting two-storied Silver birch (Betula pendula Roth) and Norway spruce (Picea abies (L.) H. Karst.) stands by creating time consumption models for cutting, and using existing models for forwarding. Damage to the remaining understory spruce was also examined. Four different harvesting methods were used: 1) all dominant birches were cut; 2) half of them thinned and understory was preserved; compared to 3) normal thinning of birch stand without understory; and 4) clear cutting of two-storied stand. Results showed the time needed for birch cutting was 26–30% lower when the understory was not preserved. Pulpwood harvesting of small sized spruces that prevent birch cutting was expensive, especially because of forwarding of small amounts with low timber density on the strip roads. Generally, when taking the cutting and forwarding into account, the unit cost at clear cuttings was lowest, due to lesser limitations on work. It was noted that with increasing removal from 100 to 300 m³ ha^–1, the relative share of initial undamaged spruces after the harvest decreased from 65 to 50% when the aim was to preserve them.  During summertime harvesting, the amount of stem damage was bigger than during winter. In conclusion, two-storied stands are possible to transit to spruce stands by accepting some losses in harvesting productivity and damages on remaining trees.

• Niemistö, Natural Resources Institute Finland (Luke), Natural resources, Kampusranta 9 C, FI-60320 Seinäjoki, Finland https://orcid.org/0000-0002-9152-2108 E-mail: ext.pentti.niemisto@luke.fi
• Korpunen, Norsk institutt for bioøkonomi (NIBIO), Division of Forest and Forest Resources, Department of Forest operations and digitalization, Divisjon for skog og utmark, Avdeling for Driftsteknikk og digitalisering, Høgskoleveien 8, 1433 Ås, Norway https://orcid.org/0000-0001-9749-5684 E-mail: heikki.korpunen@nibio.no
• Nuutinen, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6 B, FI-80100 Joensuu, Finland https://orcid.org/0000-0003-3360-4444 E-mail: yrjo.nuutinen@luke.fi

• Strîmbu, Norwegian University of Life Sciences, Faculty of Environmental Sciences and Natural Resource Management, NO-1432 Ås, Norway https://orcid.org/0000-0002-0588-2036 E-mail: victor.strimbu@nmbu.no
• Eid, Norwegian University of Life Sciences, Faculty of Environmental Sciences and Natural Resource Management, NO-1432 Ås, Norway E-mail: tron.eid@nmbu.no
• Gobakken, Norwegian University of Life Sciences, Faculty of Environmental Sciences and Natural Resource Management, NO-1432 Ås, Norway https://orcid.org/0000-0001-5534-049X E-mail: terje.gobakken@nmbu.no

• Saksa, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, 77600 Suonenjoki, Finland https://orcid.org/0000-0002-1776-2357 E-mail: ext.timo.saksa@luke.fi
• Miina, Natural Resources Institute Finland (Luke), Natural resources, Yliopistokatu 6 B, 80100 Joensuu, Finland https://orcid.org/0000-0002-8639-4383 E-mail: jari.miina@luke.fi
• Luukkonen, Metsäsydän Oy, Junninmäentie 53, 52200 Puumala, Finland E-mail: olli.p.j.luukkonen@gmail.com

Investing in planting genetically improved silver birch (Betula pendula Roth) in Swedish plantations requires understanding how birch stands will develop over their entire rotation. Previous studies have indicated relatively low production of birch compared to Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.). This could result from using unrepresentative basic data, collected from unimproved, naturally-regenerated birch (Betula spp.) growing on inventory plots often located in coniferous stands. The objective of this study was to develop a basal area development function of improved silver birch and evaluate production over a full rotation period. We used data from 52 experiments including planted silver birch of different genetic breeding levels in southern and central Sweden. The experimental plots were established on fertile forest sites and on former agricultural lands, and were managed with different numbers of thinnings and basal area removal regimes. The model best describing total stand basal area development was a dynamic equation derived from the Korf base model. The analysis of the realized gain trial for birch showed a good stability of the early calculated relative differences in basal area between tested genotypes over time. Thus, the relative difference in basal area might be with cautious used as representation of the realized genetic gain. On average forest sites in southern Sweden, improved and planted silver birch could produce between 6–10.5 m³ ha^–1 year^–1, while on fertile agriculture land the average productivity might be higher, especially with material coming from the improvement program. The performed analysis provided a first step toward predicting the effects of genetic improvement on total volume production and profitability of silver birch. However, more experiments are needed to set up the relative differences between different improved material.

• Liziniewicz, The Forestry Research Institute of Sweden, Ekebo, SE-268 90 Svalöv, Sweden E-mail: mateusz.liziniewicz@skogforsk.se
• Barbeito, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, Alnarp, Sweden; Université de Lorraine, AgroParisTech, INRAE, UMR Silva, Nancy, France E-mail: ignacio.barbeito@slu.se
• Zvirgzdins, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, Box 49, 23053 Alnarp, Sweden E-mail: andis.zvirgzdins@slu.se
• Stener, The Forestry Research Institute of Sweden, Ekebo, SE-268 90 Svalöv, Sweden E-mail: lg.stener@telia.com
• Niemistö, Natural Resources In-stitute Finland (Luke), Natural resources, Seinäjoki, Finland E-mail: pentti.niemisto@luke.fi
• Fahlvik, The Forestry Research Institute of Sweden, Ekebo, SE-268 90 Svalöv, Sweden E-mail: nils.fahlvik@skogforsk.se
• Johansson, Tönnersjöheden Experimental Forest, SLU, Simlångsdalen, Sweden E-mail: ulf.johansson@slu.se
• Karlsson, The Forestry Research Institute of Sweden, Ekebo, SE-268 90 Svalöv, Sweden E-mail: curly.birch@gmail.com
• Nilsson, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, Box 49, 23053 Alnarp, Sweden E-mail: urban.nilsson@slu.se

This study’s aim was to identify how the application season and the method of early cleaning (EC), the first stage of multistage pre-commercial thinning (PCT), affected the time consumption in EC and in the subsequent second PCT operation. The worktime in EC was recorded in the spring, summer, and autumn in 22 sites, which were either totally cleaned or point cleaned. Later, these sites were measured at the time of the second PCT. Time consumption was estimated in PCT, based on the removal of the sites. The time consumption in EC was 5.3 productive work hours (pwh) ha^–1, 7.3 pwh ha^–1, and 6.2 pwh ha^–1 respectively in the spring, summer, and autumn. EC in the spring instead of the summer saved 27–30% of working time, depending on the cleaning method. Point cleaning was 0.8 pwh ha^–1 quicker than total cleaning, but the difference was statistically insignificant. The second stage, PCT, was 1 pwh ha^–1 slower to conduct in sites which had been early cleaned in the spring instead of the summer. However, at the entire management program level, EC applied in the spring or autumn instead of the summer saved 11% or 5% respectively of the total discounted costs (3% interest rate) of multistage pre-commercial thinning. Today, the commonest time to conduct EC is in the summer, which was the most expensive of the analyzed management alternatives here. We can expect savings in juvenile stand management in forestry throughout boreal conifer forests by rethinking the seasonal workforce allocation.

• Uotila, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: karri.uotila@luke.fi
• Saksa, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@luke.fi

In Germany, management restrictions for Norway spruce (Picea abies (L.) H. Karst.) due to climate change lead to increasing interest in Douglas fir (Pseudotsuga menziesii (Mirb.) Franco) as a potential substituting species. However, Douglas fir requires cost-intensive silvicultural treatments, such as periodic thinnings and, in particular, pruning. In order to improve the efficiency of such treatments, a new tending system with an adapted two-step work system was analyzed. The new system, using electric pruning shears and the backpack clearing saw Husqvarna 535FBX ‘Spacer’, was compared to the conventional three-step work system, using handsaw and chainsaw and characterized by tree selection previously conducted as an independent work step. Time and motion studies to determine productivity and costs, as well as ergonomic analysis through heart rate measurements and posture analysis were conducted. Overall, the new system was found to be more productive and to have lower costs, with 8.9 trees per scheduled system hour (4.17 € tree^–1), compared to the conventional system with 8.1 trees per scheduled system hour (4.44 € tree^–1). Ergonomic improvements with the new system could be mainly observed during the felling of competing trees, when the level of heart rate reserve was reduced by 9.3 percent points, compared to the conventional system. However, significant advantages in reducing unfavorable body postures expected for the ‘Spacer’ could not be confirmed. Since time savings within the new system were mainly attributed to the adaptation of workflow and the use of the electric shears during pruning, it should be considered to replace the ‘Spacer’ within the new system by light chainsaws for best results under the conditions investigated.

• Schönauer, Department of Forest Work Science and Engineering, University of Göttingen, Göttingen/Germany E-mail: marian.schoenauer@uni-goettingen.de
• Hoffmann, Department of Forest Work Science and Engineering, University of Göttingen, Göttingen/Germany; School of Forestry, University of Canterbury, Christchurch/New Zealand https://orcid.org/0000-0002-8077-967X E-mail: stephan.hoffmann@uni-goettingen.de
• Nolte, Forest Education Center FBZ / State Enterprise Forestry and Timber NRW, Arnsberg/Germany E-mail: Martin.Nolte@wald-und-holz.nrw.de
• Jaeger, Department of Forest Work Science and Engineering, University of Göttingen, Göttingen/Germany E-mail: dirk.jaeger@uni-goettingen.de

Tree height-diameter allometry reflects the response of specific species to above and belowground resource allocation patterns. However, traditional methods (e.g. stepwise regression (SR)) may ignore model uncertainty during the variable selection process. In this study, 450 trees of Chinese fir (Cunninghamia lanceolata (Lamb.) Hook.) grown at five spacings were used. We explored the height-diameter allometry in relation to stand and climate variables through Bayesian model averaging (BMA) and identifying the contributions of these variables to the allometry, as well as comparing with the SR method. Results showed the SR model was equal to the model with the third highest posterior probability of the BMA models. Although parameter estimates from the SR method were similar to BMA, BMA produced estimates with slightly narrower 95% intervals. Heights increased with increasing planting density, dominant height, and mean annual temperature, but decreased with increasing stand basal area and summer mean maximum temperature. The results indicated that temperature was the dominant climate variable shaping the height-diameter allometry for Chinese fir plantations. While the SR model included the mean coldest month temperature and winter mean minimum temperature, these variables were excluded in BMA, which indicated that redundant variables can be removed through BMA.

• Lu, Key Laboratory of Tree Breeding and Cultivation of the National Forestry and Grassland Administration, Research Institute of Forestry, Chinese Academy of Forestry, Beijing 100091, P. R. China; Collaborative Innovation Center of Sustainable Forestry in Southern China, Nanjing Forestry University, Nanjing, 210037, P. R. China E-mail: 18556439861@163.com
• Chhin, Division of Forestry and Natural Resources, West Virginia University, 322 Percival Hall, 1145 Evansdale Dr, Morgantown, West Virginia, 26506, USA E-mail: steve.chhin@mail.wvu.edu
• Zhang, Key Laboratory of Tree Breeding and Cultivation of the National Forestry and Grassland Administration, Research Institute of Forestry, Chinese Academy of Forestry, Beijing 100091, P. R. China E-mail: xqzhang85@caf.ac.cn
• Zhang, Key Laboratory of Tree Breeding and Cultivation of the National Forestry and Grassland Administration, Research Institute of Forestry, Chinese Academy of Forestry, Beijing 100091, P. R. China; Collaborative Innovation Center of Sustainable Forestry in Southern China, Nanjing Forestry University, Nanjing, 210037, P. R. China E-mail: xqzhang85@yahoo.com

In Nordic forests, consistent evidence about better seedling survival rate and increased growth due to site preparation have been obtained in numerous studies. Proper site preparation method can reduce costs of the whole regeneration chain through its effects on survival of planted seedlings, abundance of natural regeneration and competition in early stand development. This study compared the natural regeneration of birches (silver birch (Betula pendula Roth) and downy birch (B. pubescens Ehrh.)), amount of exposed mineral soil, and growth of planted seedlings between spot mounding and inverting site preparation methods. Present study was conducted in eight forest stands established in 2012 or 2015. Even though difference was not statistically significant, inverting exposed less mineral soil than spot mounding and thus reduced the natural regeneration of birch seedlings by 6135 seedlings ha^–1 compared to spot mounding. However, the variation between regeneration areas was remarkable. There was no difference in seedling mortality or growth between the site preparation methods. In order to achieve high growth of conifers, moderate amount of exposed mineral soil and thus less naturally regenerated birch, inverting should be favored over spot mounding.

• Laine, Metsä Group, P.O. Box 208, FI-70101 Kuopio, Finland E-mail: tiina.laine@metsagroup.com
• Kankaanhuhta, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: ville.kankaanhuhta@luke.fi
• Rantala, Metsä Group, P.O. Box 10, FI-02020 METSÄ, Finland E-mail: juho.rantala@metsagroup.com
• Saksa, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@luke.fi

A major after-use option for former peat harvesting areas has been afforestation. The profitability of afforestation with Scots pine trees (Pinus sylvestris L.) was studied in two 31–32-year old experiments in southern and northern Finland. The stands were established by seeding and planting, and various fertilization treatments and drainage intensities were tested. The financial performance for each plot was assessed in three steps. First, the costs occurred during the measurement time were summed up according to their present value. Then, for the rest of the rotation (i.e., from the age of 31/32 onwards) the stand management was optimized in order to maximize the net present value (MaxNPV). Finally, bare land values (BLVs) were calculated by summing up the present value of costs and the MaxNPV and converting the sum of the series into infinity. The afforestation method did not affect the mean annual increment (MAI; 9.2–9.5 m³ ha^–1 a^–1) in the southern experiment. In the northern experiment the afforestation method, ditch spacing and fertilization had significant effects on the MAI of the stands. The average MAI of the planted pines was 8.9 m³ ha^–1 a^–1, and for seeded pines it was 7.5 m³ ha^–1 a^–1. The BLV at an interest rate of 3% was positive for all stands in both regions. In the northern region afforestation method, ditch spacing and fertilization also had a significant effect on the BLV. When the interest rate was 5%, almost two thirds of the stands had a negative BLV in both regions.

• Aro, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Itäinen Pitkäkatu 4A, FI-20520 Turku, Finland E-mail: lasse.aro@luke.fi
• Ahtikoski, Natural Resources Institute Finland (Luke), Natural resources, Paavo Havaksentie 3, FI-90570 Oulu, Finland https://orcid.org/0000-0003-1658-3813 E-mail: anssi.ahtikoski@luke.fi
• Hytönen, Natural Resources Institute Finland (Luke), Natural resources, Teknologiakatu 7, FI-67100 Kokkola, Finland http://orcid.org/0000-0001-8475-3568 E-mail: jyrki.hytonen@luke.fi

The resilience of closed-crown coniferous stands within the boreal forest of North America is highly dependent on successful re-establishment of tree species following fire. A shift from closed-crown forest to open lichen woodland is possible following poor natural regeneration during the initial establishment phase, followed by the development of extensive lichen cover, which may hinder ongoing recruitment. We examined the development of the crustose lichen Trapeliopsis granulosa (Hoffm.) 18 to 21 years following fire within six sites in the boreal forest of northwestern Quebec, and explored its potential to affect ongoing recruitment during early successional stages of stand development. Germination and survivorship trials were conducted within the laboratory to determine the establishment rate of Pinus banksiana Lamb. (jack pine) on T. granulosa, mineral soil, and burnt duff under two separate watering frequencies (observed and drought). Survival and establishment rates of jack pine were highest on burnt duff, and poor on both T. granulosa and mineral soil. Under the drought treatment, no seedlings survived on any substrates. In the field, T. granulosa cover had a positive relationship with mineral soil cover, and negative relationships with duff cover, ericaceous shrub cover, organic layer depth, other lichen cover, and Sphagnum moss cover. No discernable relationship was found between T. granulosa and tree density, rock cover, dead wood cover or other moss cover. The development of extensive T. granulosa cover in fire-initiated stands can impede ongoing recruitment of conifer species due to its poor seedbed quality, thereby maintaining open forests.

• Splawinski, Institut de recherche sur les forêts, Université du Québec en Abitibi-Témiscamingue, 445, boul. de l’Université, Rouyn-Noranda, QC, J9X 5E4, Canada E-mail: tsplawinski@gmail.com
• Gauthier, Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, 1055 rue du PEPS, P.O. Box 10380, Stn Sainte Foy, QC, G1V 4C7, Canada E-mail: sylvie.gauthier@rncan-nrcan.gc.ca
• Fenton, Institut de recherche sur les forêts (IRF), Université du Québec en Abitibi-Témiscamingue, 445 boul. de l’Université, Rouyn-Noranda, QC, J9X 5E4, Canada E-mail: nicole.fenton@uqat.ca
• Houle, Ministère des Forêts, de la Faune et des Parcs, Direction de la recherché forestière, Québec, QC, G1P 3W8, Canada; Ouranos Climate Change Consortium, Montréal, QC, H3A 1B9, Canada E-mail: daniel.houle@mffp.gouv.qc.ca
• Bergeron, Centre d’étude sur la forêt and Chaire industrielle en aménagement forestier durable, Université du Québec à Montréal, CP 8888 Succursale A, Montréal, QC, H3C 3P8, Canada E-mail: bergeron.yves@uqam.ca

• Haataja, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: lauritapiohaataja@gmail.com
• Kankaanhuhta, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: ville.kankaanhuhta@luke.fi
• Saksa, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@luke.fi