The purpose of this article was to collate the literature on fungal diseases that occur on seedlings in forest nurseries. It describes the symptoms of the diseases, the infection pattern of each fungus and the possibilities of controlling the diseases. As background a short introduction is given on forests and nursery practices in Finland.
Investigation on the relationship between the size of the Megastigmus suspectus Borr. (Hymenoptera, Torymidae) and the size of Abies alba Mill. seeds from which those insects emerged was conducted. The examined insects (n = 46) originated from southern Poland, and were obtained from cones collected in 1993. M. suspectus thorax length was used as an index of adult insect size, because that was correlated with 8 other body measurements. In addition, the diameter of the emergence hole of this insect was measured. All data were log-transformed and analysed using regression analysis. There was no linear correlation between adult size and seed size, using either length of thorax or any of the other 8 body traits tested.
The prescribed burning of a 7.3 ha clear-cut and a 1.7 ha partially cut forest (volume 150 m3/ha) was carried out in Evo (61 °12'N, 25°07'E) on 1 June 1992. The forest was a mesic Myrtillus site type forest dominated by Norway spruce (Picea abies (L.) H. Karst.). Practically all the trees and the above-ground parts of the understorey vegetation died in the fire, while the mor layer was thinned by an average of 1.5 cm.
A study was made on the change of germinated seedling population in time and their dependence on environmental factors. Seedlings of Norway spruce, Scots pine (Pinus sylvestris L.), silver birch (Betula pendula Roth), pubescent birch (B. pubescens Ehrh.) and rowan (Sorbus aucuparia L.) were inventoried in 1993 and in 1994 on permanent plots, four times per growing season. Autoregression models were used to compare regeneration of tree species in the burned forest with regeneration in the burnt clear-cut area, and to study the effect of distance from nearest seed source to regeneration.
The average number of seedlings germinating in 1993 was higher than in 1994, probably because of differences between these consecutive years in regard to the amount of seed rain and weather conditions. The number of Norway spruce and rowan seedling was higher inside the forest area than in the clear-cut area. The distance to the bordering forest and to the closest seed tree did not explain the result. It is suggested that the more stable microclimatic conditions under the shade of dead tree promote germination and seedling establishment in the forest area. As rowan is a bird-dispersed species, it is likely that dead trees help the dispersal of rowan seed by providing birds place to sit and defecate. The shade provided by dead trees may influence the further succession of the tree stand and vegetation composition and diversity.
The effects of wood ash and PK fertilization on natural regeneration and sowing of Scots pine (Pinus sylvestris L.) were studied in field experiments on nitrogen-poor (Ntot 0.87–1.26%) peat substrates. The study material was derived from three drained, nutrient-poor pine mires (64°52’ N, 25°08’ E) at Muhos, near Oulu, Finland. The experimental fields were laid out in 1985 as a split-split-plot design including the following treatments; mounding, natural regeneration and sowing and fertilization; PK (400 kg ha-1) and wood ash (5,000 kg ha-1). The seedlings were inventoried in circles in July–August 1991.
Changes in the vegetation were small and there were no statistical differences due to the fertilization treatments in the ground vegetation. PK or ash fertilization did not cause vegetation changes harmful to Scots pine regeneration on nitrogen-poor peatlands. Both sowing and fertilization significantly increased the number of pine seedlings, but not their height. Wood ash increased seedling number more than PK fertilizer. The number of seedlings varied from 7,963 (control) to 42,781 ha-1 (mounding + sowing + ash). The seedling number was adequate for successful regeneration even on non-mounded, non-fertilized naturally regenerated plots.
The number of birch seedlings varied more than that of pine (370–25,927 ha-1). Mounding especially increased the number of birches. The difference between PK fertiliser and ash was less pronounced than that for pine. In addition, to the field studies the effects of ash and PK fertilizer on the germination of Scots pine seeds was studied in a greenhouse experiment. Soaking in ash solutions strongly reduced seed germination, while the PK solution was less harmful.
Fill planting is a common procedure following reforestation in Finland. In 1990, 13% of the total of seedlings planted was used for fill planting. The objective of this study is (i) to survey the survival of fill-in seedlings and (ii) to estimate the spatial pattern of stands to evaluate the importance of fill-in seedlings in constituting the stocking of Scots pine (Pinus sylvestris L.) stands in Central and Northern Finland.
A survey of 63 artificially regenerated Scots pine stands was conducted in 1990. Stand densities varied from 950 to 3,925 seedlings/ha. The mean densities of originally planted, fill planted and naturally regenerated seedlings were 863, 639 and 791/ha, respectively. The survival of originally planted seedlings was 36% and that of fill-in seedlings 48%. Death rate of fill-in seedlings of Scots pine increased with longer times between original and fill planting. The survival rate of Norway spruce (Picea abies) seedlings was correlated with temperature sum. Height of the fill-in seedlings was less than that of the originally planted ones. Most stands had an even spatial distribution with the exception of sparsely populated stands, which were somewhat clustered. This indicates that dying of seedlings is not randomly spread. Because of poor survival, fill planting seems to be a risky business in most cases.
The following treatments were compared in three Scots pine (Pinus sylvestris L.) reforestation areas on a scarified moist mineral-soil site in southern Finland, planted with 1+1 bareroot stock in spring 1987: (a) no weed control treatment; (b) mulching with a fibre slurry produced by mixing wastepaper with water and applied 1 cm deep to an area of 60 cm in diameter around the seedling soon after planting; (c) glyphosate (at 2 kg ha-1) sprayed on a 1 m2 spot around the seedling in early August 1987; (d) terbuthylazine (at 10 kg ha-1) applied as (c). Monitoring of the trials over a 4-year period between 1987–90 showed that none of the treatments reduced surface vegetation to an extent that would have benefited pine. The percentage cover development of the vegetation, dominated by Agrostis capillaris, Calamagrostis arundinaceae, Deschampsia flexuosa, Festuca ovina, Epilobioum angustifolium and Pteridium aquillinum, followed much the same pattern in all treatments, with (c) slightly favouring forbs. Survival of pine at the end of the study period was about 90%, with non-significant differences between treatments. Mulching and terbuthylazine treatment slightly reduced seedling height growth in the second year. Growth was better in glyphosate treatment than in terbuthylazine treatment in the lowest (<30%) and the highest (>60%) pre-treatment weed cover classes, and in the latter also better than in untreated control. Mulching gave variable results; at its best it provided also good control of weeds for several years, without, however, improving the initial development of pine in these trials.
Growth, crown structure, flowering and seed production of silver birch (Betula pendula Roth) seedlings, grafts and micropropagated plants was compared during four years in a polythene greenhouse experiment. The growth of the seedlings was clearly the most vigorous and that of the grafts the weakest, the micropropagated plants being intermediate. The seedlings had the highest and the grafts the lowest number of branches before cutting the tops of the plants, but the differences between the material types were no more significant after cutting the tops. The grafts had significantly shorter and thinner branches than the seedlings and the micropropagated plants, whereas the differences in branch length and branch thickness between the latter two groups were not significant. The grafts started flowering at the age of two years, one year earlier than the other two types of material. At the age of four years the micropropagated plants had abundant seed production, about 75% of that of the seedlings and about two times higher than that of the grafts. Thus, the micropropagated plants can be used instead of grafts when establishing polythene greenhouse seed orchards of birch.
Micropropagated and seed-borne plants of sliver birch (Betula pendula Roth) were compared for survival and growth in a field trial at the age of six years. Three clones for micropropagation were selected from open-pollinated progenies of selected southern Finnish plus trees at the age of 17 and 20. The three seed-borne lots were of southern Finnish stand origin. The best two lots of the experiment as regards the height and diameter growth at the age of six were the clones. The best of these differed significantly from the best-growing seed-grown lot. The weakest lot of the experiment was also a clone which was clearly slow-growing with a dense and bushy crown. Survival of the material was high (mean = 94%), and there was no damage caused by voles and elks, for example. The results clearly show that the selection of material for clonal propagation should be done carefully. The clones should also be tested for performance in the field before propagation on a large scale.
After a presentation of basic biodiversity concepts, reviews are made of studies reporting genetic implications of tree improvement activities: seed treatments, seedling production, provenance transfers, plus tree selection, seed production in seed orchards and progeny testing.
Several of the activities may influence the genetic structure and diversity of the planted forests. The general conclusion is, however, that planted forests are at least as genetically diverse as the natural stands that they replace. The diversity in forest management and use is best assurance for the future adaptability of the forests.
Seed production of micropropagated plants, seedlings and grafts of Silver birch (Betula pendula Roth) in a polyethylene greenhouse experiment was followed for five years. The grafts started flowering and seed production at the age of two years, one year earlier than other two types of material. At the age of three the seed production of both micropropagated plants and seedlings was already more than two times higher than that of the grafts. Variation between the clones was high and plant type x clone interaction was significant. At the age of four, in 1993, seed production was high in all three types of material. Seed production of the micropropagated plants was two times higher than that of the grafts but about 75% of that of the seedlings. In 1994 seed production of all three plant types was very low, which shows large variation between different years. The early development of the plant material types suggests that micropropagated plants have higher seed production than grafts and could well be used instead of grafts in polythene greenhouse seed orchards.
Seedlings of Picea abies (L.) H. Karst. full-sib families of contrasting origins were cultivated in a phytotron under different photoperiodic, light-intensity and temperature treatments during their first growth period. The effects of the treatments on juvenile growth traits – whether enhanced or delayed maturation was induces – were observed during the two subsequent growth periods. The following hypotheses were tested: (A) Enhanced maturation can be induced in the first growth period from sowing with (i) a long period of continuous light during active growth (24 weeks vs. 8 weeks); (ii) a shorter night during bud maturation (12 h vs. 16 h); high temperature (25°C vs. 20°C) during (iii) active growth, growth cessation and bud maturation; and during (iv) the latter part of growth cessation and bud maturation only. (B) Delayed maturation can be induced after (i) low light intensity during growth cessation and bud maturation (114 μmol m-2 s-1 vs. 340 μmol m-2 s-1); low temperature (15°C vs. 20°C) during (ii) active growth, growth cessation and bud maturation; and during (iii) the latter part of growth cessation and bud maturation only.
The most dramatic effect was observed after 24 weeks of continuous light during active growth. All traits showed a significantly more mature performance in the second growth period compared with the control. The effect for all but one trait was carried over to the third growth period. This is in accordance with the hypothesis that the activity of apical shoot meristems controls the maturation process. For the other treatments there was only weak or no support for the hypothesis of induction of enhanced or delayed maturation. Strong family effects were observed for all traits. Differential responses of the various latitudinal families were observed, suggesting that family effects must be considered to predict and interpret correctly how plants will respond to environmental effects.
The aim of the study was to obtain information needed in preparing more precise fertilization recommendations for seed orchards. The fertilization requirement was estimated on basis of soil and needle analyses, and by investigating the effects of different fertilization treatments on male and female flowering, size of seed crop and seed quality.
The study was carried out in two Scots pine (Pinus sylvestris L.) seed orchards in Southern Finland, one established on forest soil in 1971 and 1972, and the other on a peat field in 1974. 10 clones and 39 grafts from each clone were selected randomly from both orchards in autumn 1985. The treatments consisted of N, P and K in various combinations, micronutrients, wood ash and grass control. The orchards were fertilized in spring 1986.
There were statistically significant differences between the clones in both orchards as regards amount of flowers, size of the seed crop and seed quality. The annual variation in flowering and the size of the seed crop was also large. In general, the seeds from cones collected in October matured well and their germination percentage was high. The effects of fertilization on flowering, the size of seed crop and seed quality were small. It would appear that the size of the crop can be affected to a much greater extent by favouring clones with a high seed-producing capacity than through fertilization. Fertilization is unnecessary if the nutrient status of the soil is satisfactory.
Seedlings from four Norway spruce (Picea abies (L.) H. Karst.) stands originating from areas with effective temperature sums ranging from 710 d.d. to 1,150 d.d. were raised under artificial light and temperature treatment. After a 10-week growing period the hardening process was started by subjecting the seedlings to +8°C night temperature and +15°C day temperature, and increasing the night length by 1.5 hour/week. Hardiness was measured by means of artificial freezing treatment (-10°C or -15°C), followed by visual estimation of the degree of needle injury. The stem height, lignification and bud development were measured before the freezing treatment. The amount of injury increased the more southern the origin of the tested material was. Furthermore, the proportion of non-lignified part of the seedling stem was negatively correlated with the latitude of the provenances. The proportion of seedlings with clearly visible buds was more than 90% in the northernmost entry and less than 1% in the southernmost one. The overall correlation coefficient between the needle injuries and the proportion of non-lignified part of the stem was rather high, but varied considerably from 0.3 in the northernmost material to over 0.6 in the southern provenances. According to the results, it seems to be possible to use growth characteristics as an indicator of frost hardiness at the provenance level.
The PDF includes an abstract in Finnish.
The ability of Lygus bugs to cause growth disorders in permethrin-treated Scots pine (Pinus sylvestris L.) seedlings after planting was studied in two regeneration areas. There were three treatments: exposure of the seedlings to Lygus, mechanical protection of the seedlings from insects, and control seedlings. There were no significant differences in the rate of growth disorders between the treatments. The permethrin application protected the seedlings against Lygus bugs in the early summer, as well as when the bug abundance was low. The development of these seedlings, as well as the multiple-leadered and bushy seedlings on a third regeneration area, was followed for two years. Multiple leaders reduced height growth and bud number of a seedling, but caused marked losses in growth only when the seedlings still had multiple leaders the following year, or when they formed several, equally developed stems.
The PDF includes an abstract in Finnish.
The interactive effects of potassium deficit and foliar application with acid water (pH 5.5, 4.5, 4.0 and 3.0 given consecutively) on CO2 exchange rate of Pinus sylvestris L. seedlings was investigated in field conditions. No reduction of the CO2 exchange rate was observed in the seedlings supplied with sufficient potassium. Only the seedlings having the lowest needle K concentration (2.4 mgg-1) had an apparently low CO2 exchange rate before the applications with acid water. The CO2 exchange rate of most of the seedlings with low needle K concentration (3.9–6.0 mgg-1) decreased after the acid water application. The threshold acidity for the reduction varied between pH 4,0 and 3.0 depending on the needle K concentration. The reduction was more apparent at high irradiance. It was concluded that acid precipitation disturbs the CO2 exchange only in conditions of mineral nutrient deficit.
The PDF includes an abstract in Finnish.
Model computations on the management of Scots pine (Pinus sylvestris L.) at the seedling stage showed that a rising temperature due to the suggested climate change could increase the competition capacity of birch species (Betula pendula) more than Scots pine, whose growth could even decline during the course of a rise in temperature. A temperature rise could, thus, bring the time of removal of birches forward when aiming at Scots pine timber stands composed of these tree species. The increasing proportion of birches makes the removal of birches even more urgent and emphasizes the need for careful management of Scots pine stands under rising temperatures. The first thinning of Scots pine is generally brought forward; this is particularly the case when wide spacing is applied in planting. A furthrer rise in temperature magnifies the above patterns by reducing further the competitive capacity of Scots pine in relation to birches.
The PDF includes an abstract in Finnish.
Two-year-old containerized Scots pine (Pinus sylvestris L.) seedlings, raised under different fertilization and watering regimes, were subjected to feeding preference tests with pine weevils (Hylobius abietis L.) in a bioassay. In the tests carried out with pairs of seedlings, the weevil preferred water-stressed seedlings to well-watered ones. In the case of well-watered seedlings, the weevil caused significantly more damage to NPK-fertilized seedlings than those given pure PK fertilization, or no fertilization at all. It is apparent that PK fertilization reduces, and water stress increases seedling susceptibility to weevil damage. The results support findings from field trials that water stress (planting shock) predisposes seedlings to weevil damage. Weevil resistance is discussed with respect to fertilization and water stress as determinants of seedling quality.
The PDF includes an abstract in Finnish.
The potential alleopathic inhibitive effects of aqueous extracts of 13 peatland plant species on germination, radicle and seedling growth of Scots pine (Pinus sylvestris L.), silver and downy birch (Betula pendula Roth., B. pubescens Ehrh.) were studied. Freshly cut plant parts were finely ground, mixed with distilled water and agitated. The proportions of fresh plant mass in the mass-based extracts varied within the range of 1, 5, 10 and 20% (w/w). The seeds were germinated in petri dishes moistened with the plant extracts. In a separate experiment growth of birch seedlings irrigated with the extracts was studied.
Ledum palustre, Vaccinium uliginosum and Empetrum nigrum extracts, and in certain experiments extracts of other species, inhibited the germination of Scots pine and birch seeds. Results from the different experiments were not, however, fully consistent. None of the low (1% w/w) extract concentrations had any effect on germination. Strong extract concentrations (20% w/w) inhibited germination of pine seedlings significantly. The extracts affected only slightly the growth of potted birch seedlings.
The PDF includes an abstract in Finnish.
During the past 30 years, genetics research has accumulated much information on black spruce (Picea mariana). The adoption of less intensive and faster plustree selection, establishment of seedling seed orchards and family selection significantly increased the rate of progress in improvement of the species. In New Brunswick this approach made it possible to obtain substantial quantities of seeds 10 years after the initiation of the program, and now all the seeds used in reforestation are derived from seed orchards.
Fourteen years after beginning the black spruce breeding program, second generation breeding is underway. The possibility of implementing alternative breeding strategies using ”breeding – cloning” procedures are explored for the advanced generation. Until somatic embryogenesis techniques become fully operational, ”backward selection” schemes and crossing in breeding halls followed by vegetative multiplication using serial rooted cuttings can be adopted. Larger genetic gains than those from conventional breeding are expected not only from the utilization of both additive and non-additive variances, but also from the elimination of inefficiencies of large conventional seed orchards.
The paper gives an introduction of the tree breeding program of Sweden that started in 1936 by the establishment of an association for the tree breeding. In 1967 the Institute of Forest Improvement was founded and it replaced the earlier association. The main species in the programme have been Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies (L.) H. Karst.), lately also birch (mainly Betula pendula Roth.) and lodgepole pine (Pinus contorta). In addition, limited breeding has been done also with hybrid aspen (Populus tremula x P. tremuloides), oak (Quercus), larch (Larix), black spruce (Picea mariana) and a few other native and exotic species. The dominating initial effort has been to select plustrees in natural stands and use them for production of reforestation material. In addition, a considerable body of tests was built. The paper lists the status of breeding material of the different tree species and introduces the medium and short-term breeding programmes.
Application of growth regulators, primarily gibberellins, has become an efficient method of flowering regulation in many coniferous forest trees. In this paper some results are discussed with respect to various genetic processes in seed orchards. It is concluded that there are still several unsolved problems in the regulation of flowering in forest seed orchards. These problems are related to different genetic processes occurring in the seed orchards and they should be taken into account in endeavouring to produce genetically improved seeds.
In two separate studies, seedlings from 20 loblolly pine (Pinus taeda L.) families and Virginian pine (Pinus virginiana Mill.) were used as root stocks for grafting loblolly pine seed orchard clones. The rootstocks were open-pollinated seedlings from orchard clones chosen to represent a wide range of flowering and survival capabilities, based on their performance in a first-generation seed orchard. Scions were derived from the same 20 loblolly clones. The effects of scion clone were significant and large for nearly all measured traits. Rootstock significantly affected survival, growth, flowering and foliar nutrients of the grafted ramets. Neither survival nor growth of the crafts was related to survival of growth of the orchard clones from which their rootstocks were derived, however. Survival of incompatible clones was enhanced by grafting on genetically related rootstocks.
The PDF includes an abstract in Finnish.
The mating system was analysed in the upper and lower crown of two groups of European larch (Larix decidua Mill.) clones divided according to the percentage of full seeds in the upper and lower crown parts. The overall multilocus estimate of outcrossing rate (t) was calculated to be 0.929. The differences of outcrossing rates between crown levels and clonal groups respectively were not statistically significant. The t estimates were greater for the upper crown level and for clones with higher percentage of full seeds in the upper crown level. However, among all observations there was no correlation between outcrossing rates and percentages of full seeds for particular crown levels and groups of clones. Observed similarity of outcrossing among grafts of the same clones may indicate genetic control of self-fertilization rate in individual European larch.
The PDF includes an abstract in Finnish.
This paper summarises the main topics and conclusions from the joint meeting of two IUFRO working parties, S2.04-02, Breeding theory and progeny testing, and S2.02-16, Seed orchards, held in Tuusula, Finland on September 10–15, 1991. It concludes the main topics that need more research in these disciplines and future trends in the research.
This issue of Silva Fennica includes the presentations given in a joint meeting of two IUFRO working parties, S2.04-02, Breeding theory and progeny testing, and S2.02-16, Seed orchards. The sessions were held in Tuusula, Finland on September 10–15, 1991.
The effect of photon flux density on bud dormancy release in two-year-old seedlings of Norway spruce (Picea abies (L.) H. Karst.) was examined. The seedlings were first chilled for 0–21 weeks under natural conditions and then forced in a warm greenhouse either in low (15 μEm-2s-1) or in high (300 μEm-2s-1) photon flux density. Occurrence of bud burst was observed in the forcing conditions, and the observations were used for estimating the cumulative frequency distribution of the chilling requirement for growth competence. The estimated distribution had greater variance in the low photon flux density than in the high photon flux density forcing. This finding suggests that unnaturally low photon flux densities during forcing may yield overestimates of the genetic within-population variation in the chilling requirement for growth competence.
The PDF includes an abstract in Finnish.
The effect of three pesticides containing either dimethoate (0.5% a.i.), permitrin (0.5%) or triadimephon (0.5%) on the cone pests and flowering biology of Norway spruce (Picea abies (L.) H. Karst.) was tested in a seed orchard in mid-May or in the beginning of June. The pesticide treatments significantly reduced infestation by Laspeyresia strobiella and Kaltenbachiella strobi only. Variation in the number of cones infested by both insects and cone rusts was high between the spruce clones. Generally, the pesticides did not affect flower viability, seed quality or seed germination, but reduced drastically the germination capacity of pollen in vitro. In practice, sufficient control cannot be achieved with concentrations or methods used in the present study.
The PDF includes an abstract in Finnish.
Containerized tree seedlings will be used on an increasing scale in the future in different parts of the world. There are number of techniques for the production of small one-year-old seedlings but it has not been possible to develop a completely satisfactory methods for large containerized seedlings production. In the long-term development of pine plantations established with containerized seedlings the greatest problem has been deformation of the root system. With a new method, based on a sheet of peat and root pruning, it has been possible to produce conifer seedlings with a good root regeneration potential and favourable morphological root system development. The use of small containerized seedlings allows an increase in planting density without any marked increase in regeneration costs.
The PDF includes an abstract in English.
Seed mass within any plant species is one of the least plastic components of plant structure. The aim of this study was to analyse the variation in the seed mass of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) in relation to three environmental factors: soil fertility, mean temperature and precipitation during seed filling period. Data published earlier on seed mass of these species on different sites and different years was used in the study.
The seed mass of both species was independent of soil fertility (forest type) but did vary between different years. It is hypothesized that if the seed-ripening summer is warmer than average, Scots pine seed mass tends to be smaller. In this study, seed mass varied independently of the amount of precipitation during the ripening summer. However, generalization of the results requires further study.
The PDF includes an abstract in English.
The author first introduced the cut-block seedling production method to Finland in 1969. The aim is to raise seedlings whose lateral roots do not become deformed as a result of a restricting container or other external pressure. The seedlings are raised in a large, fairly compact substrate block where the roots can freely develop in a normal fashion. The blocks are then cut up into individual cubes, each containing a seedling. The precise positioning of the sowing point permits mechanization of the work.
The PDF includes an abstract in English.
The study uses the methodology of ecological field theory to model the effect of Scots pine (Pinus sylvestris L.) seed trees on the density of tree seedlings and other plants in the field layer. The seed trees had a clear effect on the expected value of the amount and distribution of the ground vegetation. The vicinity of seed trees had an adverse effect on the growth of grasses, herbs and seedlings, while mosses were most abundant near the trees. Models based on the ecological field approach were derived to describe the effect of seed trees on the ground vegetation.
The PDF includes an abstract in Finnish.
In Northern Finland as well as in Northern Sweden there is a shortage of high-quality Scots pine (Pinus sylvestris L.) seeds, mostly due to unsuitable temperatures during the development and maturation of cones and seeds. Methods have been developed for elimination of non-productive seeds and for invigoration of seeds. In the present work, these methods were tested on poorly developed seed lot from Rovaniemi, Northern Finland (66°15’–66°30’; 180 m a.s.l.). The seeds were conditioned using the following treatments:
1. PREVAC method (5 min, 97 kPa below atmospheric pressures) for removal of mechanically damaged seeds (7%)
2. Invigoration using incubation at controlled moisture content (30% f.w) and continuous air supply, for 14 days at 5°C.
3. Additional water supply for 16 hours at 5°C.
4. Drying in dehumidified air until a near maximum difference in density between viable and dead seeds was obtained
5. Separation in a sedimentation flume to achieve a gradient of fractions of different germination rate and capacity.
The treatments resulted in an improvement of germination percentage from 33 to about 95% and a reduction in mean germination time from 8.8 days to 6 days if the control and the best fractions (32% seeds) were compared.
The effect of root exposure on the shoot and root development of Pinus sylvestris (L.) seedlings was studied at two soil temperatures. Roots of bare-rooted three-year-old seedlings were exposed to the temperature of 32°C at relative humidity of 50–40% for 85, 155 and 270 minutes which corresponds to accumulated water pressure deficit of 24, 47 and 91 mbar·h, respectively. Thereafter, seedlings were grown for 65 days at the soil temperatures of 12 and 23°C. Drought exposures inhibited new root initiation, delayed shoot elongation, and reduced shoot and needle growth. The stronger the exposure the larger the proportion of needles from the lower part of current shoot that remained undeveloped. Low soil temperature increased the effect of exposures so that needle elongation and initiation of new root tips of seedlings in cold soil with the longest exposure were inhibited totally. Root growth assessments made in warm soil may overestimate the acclimation potential of planted seedlings.
The PDF includes an abstract in English.
The effect of different environmental conditions (four outdoor localities and one greenhouse locality in Northern Sweden) on cold hardening of 29 one-year-old full-sib families from plus-trees of Scots pine (Pinus sylvestris L.) were studied by artificial freeze testing. Plants exposed to low night temperatures during August achieved faster cold hardening than plants raised in milder localities. The family ranking for rate of winter hardening was consistent among outdoor localities if freeze testing was performed at times when plants from different localities had attained similar levels of cold hardiness. However, significant family x locality interactions were obtained when plants from the outdoor localities were freeze tested on the same occasion. Freeze damage was positively correlated with plant height but not correlated with dry matter content in the autumn. Freezing damage of greenhouse raised plus-tree families was uncorrelated with damage of plants raised outdoors. Possible implications for hardiness breeding are suggested.
The PDF includes an abstract in Finnish.
Visible frost damage to forest trees in Sweden seldom occurs in winter but is frequent in late spring, summer and early autumn. Frosts are frequent in all seasons in various parts of Sweden, even in the southernmost part (lat. 56°, N) and temperatures may be as low as -10°C even around mid-summer. Ice crystal formation within the tissues, which in most seedlings takes place at around -2°C, causes injury, not the sub-zero temperatures themselves.
The apical meristem, the elongated zone, and the needles of seedlings of Picea abies (L.) H. Karst. in a growing phase were damaged at about -3°C and those of Pinus sylvestris L. at about -6°C. Other species of the genus Pinus were tested and most were found to be damaged at about -6°C, with some variations. Picea species tested were damaged at about -3°C to -4°C.
A method has been designed to compare the response of different species to winter desiccation, which occurs under conditions of (1) low night temperature, (2) very high irradiation and increase in needle temperature during the photoperiod, (3) frozen soil, and (4) low wind speed. There were differences in response to winter desiccation between pine and spruce species. Seedlings of Pinus contorta tolerated these winter desiccation conditions much better than those of P. sylvestris or Picea abies. Picea mariana was the least tolerant of the species tested.
The PDF includes an abstract in Finnish.
The occurrence of Caleopsis bifida on clear-cut and burned forest soil and its disappearance in 4–6 years after disturbance is attributed to its germination ecology. Initially the seeds are dormant 96–100% and remain dormant in nylon gaze bags in different types of forest humus layers at least 10 years. Dormancy is released in laboratory (1) by treatment of 100 ppm aqueous solution of GA3, (2) by heating the dormant seeds to 40–55°C for 1–5 h, and (3) by 1% KNO3 solution. It is concluded that conditions in clear-cut and burned areas favour germination of seeds in regard to temperature and content of nitrates in contrast to humus of closed vegetation where the seeds remain dormant.
The PDF includes an abstract in Finnish.
The study material consisted of 13 rather old Norway spruce (Picea abies (L.) H. Karst.) and 17 Scots pine (Pinus sylvestris L.) stands located in different parts of Finland. In each stand the seed crops, radial growth and amount of latewood were measured during a period of about ten years. Seed production reduces the radial growth of spruce and pine in the year of seed maturing. In Southern and Central Finland also the proportion of latewood is reduced. Seed production accounts for about 14% of the variation in radial growth of a spruce stand growing in Lapland, and 27% in other parts of Finland. In pine stands the seed crop explains 19% of the variation in radial growth in Lapland, and only 7% in the rest of Finland. In spruce stands an average seed crop reduces radial growth by 14% in Lapland and 5% in the rest of the country. An abundant seed production causes a reduction of about 20%. In southern parts of Finland, the proportion of latewood is reduced by 5% in an average seed year and by 24% in a good seed year. In pine stands an average seed crop decreases the width of annual ring by 5%, and a good seed crop by 15%. Outside Lapland, also the proportion of latewood is reduced: in an average seed year by 5%, and in a good seed year by 16%. The reduction in volume growth of spruce stands due to an average seed crop was estimated to be about 10% in Lapland, and 6% in other parts of Finland. A prolific seed production causes a reduction of 20%. In old pine stands the reduction is 5% in an average seed year, and 15% in a good seed year.
The PDF includes an abstract in English.
The seed crop of Norway spruce (Picea abies (L.) H. Karst.) and Scots pine (Pinus sylvestris L.) is predicted with the help of mean monthly temperatures during May–August one and two years before the flowering year. The prediction models were made separately for Lapland and for the rest of Finland. The models are based on 10-year periods of seed crop measurements and climatic data. The total number of time series was 59.
In Lapland, Norway spruce flowered abundantly and produced an abundant seed crop after warm July–August and two years after cool July–August. In other parts of Finland, warm June and July produced a good flowering year, especially if these months were cool two years before the flowering year.
In Lapland, Scots pine flowered abundantly if the whole previous growing season was warm. Elsewhere in Finland, a cool June preceded prolific flowering in the coming year if the rest of the growing season was considerably warmer than the average.
The prediction models explained 37–49 % of the variation in the size of the seed crop. The occurrence of good and poor seed years was usually predicted correctly. Using the presented models, the prediction of the seed crop is obtainable 1.5 year for Norway spruce and 2.5 year for Scots pine before the year of seed fall.
The PDF includes an abstract in English.
In the model the regeneration process is derived into three subprocesses: birth, growth and mortality of seedlings. The main emphasis is on the birth process where the following phases are simulated: seed crop, quality of seeds, maturity of seeds, predation of seeds and germination. The parameters are based on data published in Finland. Part of the parameters are obtained directly from the investigations and part is proposed by the author. The model can be calibrated by changing parameter values. The simulation is made with the help of random numbers which have the same means as the estimates and the same distributions as the residuals of the equations used in simulation. The time step of the model is one year. The number of emerged seedlings in one year is obtained by multiplying the seed crop with the probabilities that the seed passes different phases of the birth process. Because of stochasticity the regeneration period is simulated several times. From the results it is possible to evaluate the risk and succeeding probability of the regeneration. The main drawbacks of the simulation method are the lack of empirical parameters and the difficulty of testing. The model could be further developed by including spatiality into the model.
The PDF includes an abstract in Finnish.
The effect of the size of seed crop, dispersal of seeds and the early development of seedlings on the density and spatial distribution of young Scots pine (Pinus sylvestris L.) stands are evaluated on the basis of theoretical models. The models include (i) number and spatial distribution of parent trees on the regeneration area, (ii) size of annual seed crop, (iii) seed dispersal from a particular parent tree, (iv) germination of the seeds (germination percentage), (v) death of ageing seedlings after the establishment process, and (vi) height growth of the seedlings.
As expected, stand density and spatial distribution varied within a large range in relation to the density of the parent trees and the distance from them. The simulations also showed that natural seedling stands can be expected to be heterogenous due to the geometry of seed dispersal, emphasizing the frequency of young and small trees. The properties of the seedling stands were, however, greatly dependent on the density of the parent trees and the length of the regeneration period.
The PDF includes an abstract in Finnish.
Six seed collections were made in September–December 1984 in a natural Scots pine (Pinus sylvestris L.) stand in Southern Finland. The seeds were germinated immediately after the cone collection and three photoperiods (0.8 and 24 hours) were used in germination tests.
The seeds collected in September and October possessed relative dormancy, i.e. they did not germinate in darkness and at 10°C. Later in November and December the seeds were capable to germinate in darkness and at low temperature also. The gradual change in germination capacity is attributed to chilling temperatures in natural environments or in cone storage.
The PDF includes a summary in English.
The size, nutrient contents and terpene composition of needles of Scots pine (Pinus sylvestris L.) saplings untouched and repeatedly browsed by the moose (Alces alces L.) were compared. Material was collected from a 14-years old and 2.5 m high pine stand in Bromarv, Southern Finland. The average length and fresh and dry weight of the needles were measured, and nutrient content (N, P, K, Ca, Mg, B, Cu) was determined.
The needles of repeatedly browsed pines became long and robust. There was, however, no difference between the dry matter percentage between the needles. The average nitrogen content was higher in the rebrowsed trees. Nitrogen content is, however, not directly correlated with the palatability of pine needles. Even phosphorus and boron content were higher in the damaged trees. No difference was found in Ca, K, Mg and Cu contents of the browsed and control pine saplings.
The PDF includes a summary in Finnish.
Early growth of four different tree species (Pinus sylvestris L., Picea abies (L.) H. Karst., Larix sibirica Ledeb and Betula pendula Roth) 16–23 years after planting were compared in a field experiment of 16 square plots established on a stony, grove-like upland (Oxalis-Myrtillus forest type) in Southern Finland. This study gives additional results to the publication Folia Forestalia 386/1979.
At this early stage, the growth of the spruce stand was clearly slower than that of the other species for all parameters to be measured (height, diameter, and volume growth). Height growth was most rapid in the silver birch stand and diameter growth in the larch stand. No clear differences were found in the mean volume of the 100 thickest trees in the stand between the larch and silver birch.
The PDF includes a summary in English.
The aim of the present study was to survey the occurrence of Otiorrhynchus nodosus Müller weevils and their significance for the natural regeneration of Scots pine (Pinus sylvestris L.). The study was carried out during summer 1982 at Inari in northern Lapland.
There were two sample plots, one situated in a Scots pine seed-tree area and the other, the control sample plot, in an area with a coverage of mountain birch (Betula pubescens subsp. tortuosa, now subsp. czerepanovii). A total of 177 Otiorhynchus weevils were caught. Movement of the weevils reached its climax in July. There were 86% more individuals in the seed-tree area than in the mountain birch area. No damage to the pine germlings or seedlings was not observed, although the situation could be different during the peaks of the veewil populations.
The PDF includes a summary in English.
An inventory of Scots pine (Pinus sylvestris L.) graft collection in Kolari (67°15’ N, 23°45’ S) showed that severe damage by arctic hare (Lepus timidus L.), root and bank vole (Microtus oeconomus Pallas and M. agrestis L.) and moose (Alces alces L.) was done to grafts in size and in rather poor condition. Furthermore, the damage by arctic hare was dependent on the dry matter content of the needles. Another inventory in a fertilization experiment in a pine pole-stage forest showed that nitrogen fertilization increased the damage by arctic hare. On the basis of the present results, an assumption was made that the formation of repellent substances against herbivorous mammals is connected with wintering process of northern pines.
The PDF includes a summary in Finnish.
The choice of sampling method is of prime importance when seeking relevant information about the height distribution and spatial arrangement of seedlings in the regeneration surveys. It is suggested that the size of sampling plots should depend on the height of the seedlings. Tall seedlings should be sampled from a larger area than short ones since tall seedlings are more important for the future development of the stand. We suggest principles for a technical development task to construct a device which is easy to use in practical regeneration surveys and by which sampling can be made proportional to plant height or any desired function of height.
The PDF includes a summary in Finnish.
The aim of the paper was to describe the development of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) seedling stands on drained peatlands and to find out the principal factors influencing their growth. The material under survey consists of 180 sample plots distributed from southern coast of Finland to the Polar Circle.
The most important growth factors have been the accumulated temperature sum, site quality, drainage intensity and silvicultural condition, such as the density of the stand, the proportion of birch in the stand, and the amount of possible shelterwoods. The influence of these factors, and to some extent the influence of fertilizing, and the disturbing effects of some forest damages, such as frost, growth disturbances and elk damages were investigated. Comparisons of the development in the seedling stands on drained peatlands with the known development of seedling stands in mineral soils were made.
The PDF includes a summary in English.
The publication comprises proceedings of a conference held in Helsinki in 1981. Forest tree populations are investigated for population genetic structure, mating systems, mechanisms of genetic adaptation and ecological adaptation. Methods and techniques used in population genetic research of forest trees are presented. Much concern is given to applications by means of forest tree breeding, particularly the seed orchard breeding technique. Generally, the application of population genetics in cultivated forests is discussed.
The PDF includes a preface and the presentations of the conference (25 short papers) in English, and a comprehensive summary of the themes of the conference in Finnish.
Needle damages, transpiration, photosynthesis and needle and stem height growth of Scots pine (Pinus sylvestris L.) seedlings treated with dilute sulphuric acid were studied. The acidity of the solution was pH 3. Application of a dilute solution of sulphuric acid equivalent to the normal amount of precipitation occurring during the growing season damaged the surface of two-year-old needles but not that of the current-year needles. A reduction in the photosynthetic rate of 10–30% was observed compared with the untreated seedlings. Transpiration of the seedlings was not affected by the treatment. Needle growth and stem height growth of the seedlings growing on a substrate representing poor sandy soil were reduced. Increased needle growth and stem height growth were characteristic for the seedlings growing on substrate representing fertile moraine.
The PDF includes a summary in English.
The applicability of MCPA- and 2,4,5-T-herbicides for use in the management of sapling stands and the possibilities of carrying out foliar spraying at an earlier date than at precent with smaller doses of the active ingredient were examined in this study. The results were obtained from foliage spraying experiments carried out in Central Finland in summer 1976. MCPA and 2,4,5-T were as effective as each other against deciduous tree species. However, MCPA was slightly more effective against aspen (Populus tremula L.) than 2,4,5-T. The spraying date had no effect on the mortality rate of aspen or birch (Betula pendula Roth and B. pubescens Ehrh.) There were only very slight differences between the results for different dosage levels. The damage caused to Scots pine (Pinus sylvestris L.) was very slight. The temperature conditions prevailing during spraying affected spraying effectiveness in such way that the mortality rate decreased during cold period.
The PDF includes a summary in English.
In order to be recorded in the national list of untested seed orchards, set by law, a seed orchard must attain certain objectives. It is impossible to define universal requirements for all categories of seed orchards of all species. The present paper suggests minimum requirements for a clonal region seed orchards of Pinus sylvestris L in Finland. The clonal composition as well as factors influencing on the mating pattern have been given precise numerical threshold values. To be approved the seed orchard must fulfil all the requirements at the same time. The goal is to control the genetic superiority of the forest reproductive material obtained from registered untested seed orchards.
The PDF includes a summary in English.
Soft X-ray radiation (Grenz rays) has been used in the X-ray radiography of seeds. It is based on the principle that the different parts of the seed absorb X-rays to a slightly varying extent. Empty seeds and empty regions in the seeds appears as dark areas on the film. X-ray radiography without staining and X-ray contrast radiography were compared to study the stored seeds of Scots pine (Pinus sylvestris L.). The results were compared with the results of germination in a Jacobsen germinator according to ISTA norms.
Normal X-ray radiography gave too good estimates of germinative capacity of the seeds, and was shown to suit only when it is used to study fully ripe seeds which have recently been collected in an undamaged condition. The results of X-ray contrast radiography when barium chloride was used as the stain, however, followed closely the germination results.
The PDF includes a summary in Finnish.
The aim of the present study was to establish whether hormone treatments would promote flowering in Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) grafts under Finnish conditions. Also, an attempt was made to test the efficiency of hormones as well as the variation in response among different clones. Six Scots pine and six Norway spruce clones were selected in each seed orchard based on their flowering intensity, and treated with growth hormones (GA, NAA) of different dosages by spraying. Flowering was observed one year after the treatments.
None of the treated or untreated spruce grafts flowered. However, poor flowering in the natural stands indicated that the environmental conditions during the previous years did not favour flowering. On the other hand, a distinct increase in flowering in Scots pine was observed as a result of spraying with hormone solutions. Treatments with gibberellin had a distinct promoting effect both on male and female flowering in the Scots pine grafts, although the responses varied between the different hormones or clones. The relative effect was generally stronger in male flowering.
The PDF includes a summary in Finnish.
In this paper the connection between seed weight and amount and duration of growth are studied at the progeny level within stands or climatically uniform areas, and at the provenance level within larger geographic areas. The material consists of materials of several experiments in the nursery of Maisala in Southern Finland in 1971–76. The origins of the plant material used in the different experiments ranged from progenies of individual Scots pine (Pinus sylvestris L.) or Norway spruce (Picea abies (L.) H. Karst.) trees to provenance selections covering almost the whole natural range of these species.
The effect of seed weight on plant height is strongest immediately after germination and subsequently decreases steadily, when the genetic growth properties of the plants themselves become effective. The effect is usually visible at least until the end of the 1st growing season. This relationship varies considerably depending on the material studied. The connections between the duration and the amount of height growth also proved to differ according to the nature of genetic variation. In wide selection of provenances, which show clear genetic differentiation in annual growth rhythm. The variation in the duration of growth accounts for most of the differences in total height growth. At the individual and family level or between provenances of a limited area, there seems to be no clear connection between the duration and the amount of growth. It seems that the duration of the annual growing period is a genetic property, which is not affected by seed weight.
The total height alone in 1-year old test material grown in a greenhouse had hardly any value in the forecasting of growth capacity. The growth differences were caused mainly by the variation in seed size and growth rate differences during the growing period.
The PDF includes a summary in Finnish.
The paper discusses the variation of flowering and seed crop of Scots pine (Pinus sylvestris L.) stands based on literature published in Finland. According to the studies, the list of good seed years given by different authors do not completely match, and no essentially new about the periodicity of the seed crop or the causes of variation has been found. The difficulty in classification of the seed crops in different years in different stands and locations can be handled through the concept of capacity. It is suggested in the paper that to be able to compare the occurrence of good flowering and seed years of different stands and localities, a case exceeding 50% of the capacity is described as abundant, and a case exceeding 75% as very heavy.
The PDF includes a summary in Finnish.
Establishment of seed orchards to produce genetically improved seed started in the USSR in the 1960’s. The aim is that within 10–15 years the total seed production will reach 50% of the seed used. The paper describes seed production in seed orchards of Scots pine (Pinus sylvestris L.) in the forest-steppe of the European part of the USSR. The orchards have been established either by grafting or by planting of seedlings originating from plus trees. The grafts begin flowering relatively early, however, only at the age of 8–10 years seed crops become relatively regular and abundant. The cone yield in young seed orchards have great variability, and the yield of cones varies between clones. Cone yield does not always correlate with a good seed yield. One of the reasons for high amount of empty seeds is the difference in the periods of flowering between the clones.
The seed crops can be improved by establishing optimal conditions for the growth of the trees. Also, when a seed orchard is founded, the peculiarities of the generative activities of the clones and their reactions to changes in the environmental conditions must be taken account by taking an individual and selective measure approach to improvement measures. The orchards can be established with clones for their genetic combining ability and their requirements for the stimulation of seed-bearing.
The PDF includes a summary in Finnish.
Planting of forest trees takes place in USSR on millions of hectares but without permanent forest seed bases having yet been established in every region of the country. Consequently, local seed does not suffice the need, and it is necessary to secure it from other geographical regions. To avoid negative consequences of seed transfer it is necessary to study geographic variation of the genetic characteristics of forest trees and construct scientifically based division into seed zones. Geographic transfer of seed can also be regarded as a breeding measure. A large research program is being carried out in the USSR on the subject. Most of the existing trials are provenience tests of Scots pine (Pinus sylvestris L.). Over 2,000 hectares of new provenience experiments is to be established in near future.
The PDF includes a summary in Finnish.
The development of the scientific basis for the production of high-quality seed led to the introduction of a large part of the research findings in forest genetics and forest tree breeding into practical forest seed production in the USSR. Since 1971, work has begun in forest enterprises on the establishment of a permanent seed supply for the main tree species – Pinus sylvestris, Picea ssp., Larix ssp. Quercus robur, Haloxylon ssp. and nut trees.
The basis for forest tree improvement is a gene pool which is built up using mass and individual selection of valuable forms in natural populations. In accordance with a long-term programme up to 1990, an inventory of 13.2 million ha of the best high-productive stands has been carried out for breeding purposes in the state forests. About 7,000 ha of plus stands have been selected, and a total of 9,453 ha of seed orchards and 141,253 ha of seed collection stands have been established. The first stage of the programme is planned to be complete in 1980, and in the second stage clonal high-quality seed orchards will be established.
The PDF includes a summary in Finnish
As a part of the scientific and technical cooperation between Finland and the USSR a symposium and an excursion on forest genetics and seed production was organized in Finland in August 1978. The symposium paper presented at Punkaharju are published here in order to bring them available for a wider audience.
The 12 symposium reports deal with the following subjects: Practical application of forest genetic research (A.I. Novoselceva), geographical variability and provenance transfer (E.P. Prokazin, M. Hagman, I. Etverk), variation in wood density (P. Velling), variation of flowering and seed crops in seed orchards (Y.P. Efimov), and natural stands (V. Koski), vegetative propagation (J. Niiranen), seed size effects and early test problems (J. Mikola), quantitative analysis of genotypic and environmental effects (S.A. Petrov), hormonal induction of flowering (O. Luukkanen) and x-ray photography analysis of the ageing of seeds during storage (M. Ryynänen).
The PDF includes a summary in Finnish.
The development of Scots pine (Pinus sylvestris L.) seedlings and damage caused by Hylobius abietis L. (Coleoptera, Curculionidae) were studied during a three-year period. Olfactory responses of H. abietis was studied in laboratory with several volatile oils isolated from different kinds of P. sylvestris seedlings. Resistance of seedlings against H. Abietis was evaluated in terms of their monoterpene composition. Three aspects of resistance (preference, antibiosis and tolerance) were evaluated separately. Seedling chemotype was found to be associated with these aspects of host resistance on only minor scale. Discussion was attached to a further search for host resistance arising from other properties and constituents of oleoresin. Height growth of the seedlings recovering from weevil damage was 86–91% compared to healthy seedlings.
The PDF includes a summary in English.
The daily height growth rate of several larch species and progenies (Larix decidua, Larix sibirica, Larix laricina, Larix leptolepis) grown in a plastic greenhouse and in the open was measured. The growth pattern indoors was completely different compared with the normal outdoor growth pattern. The onset of growth took place in the greenhouse much earlier than outdoors and the phase of increasing growth was much shorter, as was expected. However, the phase of maximum growth was unexpectedly long. This fact suggests that there is great potential for using greenhouse cultivation to change the growth pattern of cultivated plants in order to obtain more complete utilization of the potential growing season.
The PDF includes a summary in Finnish.
A theoretical nomogram was made for estimating the costs of fully mechanized thinning and the driving speed of the machine. Based on this nomogram and the previous studies three harvesting methods were compared; systematic fully mechanized harvesting, selective fully mechanized harvesting, and manual felling combined with whole-tree chipping.
The third method was cheaper than the fully mechanized methods in a pole-stage stand. The choice of the most advantageous chipping station depended on conditions, but the smaller tree size and possibly the reduced damage on the remaining stand favour chipping on the strip road rather than chipping on the intermediate landing or at the mill.
Mechanized systematic thinning was the cheapest method for harvesting in the sapling stand. The required driving speed were so low that ergonomic factors should not hinder its use. Factors related to the future production of the stand do, however, limit its use. Mechanized selective thinning does not seem to be an economic method for harvesting in a sapling or pole-stage stand.
The PDF includes a summary in English.
The success of certain Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) provenances from Northern Finland was studied in a tree damage area occasioned by air pollution in the town of Oulu over the period of 1972-76. The pine strains from more northerly and easterly areas and the spruce strains from the immediate vicinity of the site itself were being observed to thrive best. The results point in a similar direction to those of other comparable experiments, except that the mortality rate amongst the saplings was exceptionally high and the proportion of healthy saplings in good condition was found to be unusually low. Structural properties suggestive of resistance to pollution were observable selectively in certain provenances, these including the xeromorphy of needles or thickness of the epidermis. The chief cause of mortality amongst the saplings was found to be the damage inflicted by pollution during the winter, while that arising in the summer months was relatively slight.
The PDF includes a summary in Finnish.
This study, comprising three experiments, aims to determine the effect of the geographical origin of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) seeds, and the duration of the time lag between the moisture treatment and subsequent irradiation on the gamma-irradiation sensitivity of seeds.
The studies showed that the greater the irradiation dose seeds were subjected to the slower the rate of germination. In general, small radiation doses (250–1,000 rad) had a stimulating effect and the final germination percentage (36–40 days) increased. However, when the level was further increased, the germination percentage decreased. Air-dry and moistened seeds withstood irradiation better than others. In a study with moistened seeds from different geographical sources, pine and spruce seeds from Northern Finland were less able to withstand irradiation than those originating from the south.
The PDF includes a summary in English.
The paper describes an attempt to determine whether ammonium, nitrate and urea nitrogen are bound in peat used as a filling material in containerized seedling production, what is the effect of the nutrients on certain chemical properties in the peat, and what is the effect of the nitrogen fertilizers on the primary growth of containerized (paper-pot VH 608) Scots pine (Pinus sylvestris L.) seedlings in connection with planting out. The seedlings were fertilized with ammonium sulphate, potassium nitrate and urea.
The results show that none of the fertilizers used were bound in the peat. The nitrogen content in the above ground part of the seedlings increased clearly. Fertilization with ammonium sulphate resulted in the greatest increment and this increase appears to be permanent. The wintering process was somewhat delayed by the fertilization. The seedling mortality rate for all the treatments has been quite appreciable. However, fertilization particularly with ammonium sulphate on the poorer of the two sites studied has had a positive effect on seedling survival. Furthermore, it appears that fertilizer treatments have decreased growth after planting, but in the case of ammonium sulphate this decrease has changed into a clear growth increment.
The PDF includes a summary in English.
The study was carried out in order to find out the changes taking place in germination of seeds in certain tree species as a function of gamma irradiation, the height growth of the seedlings produced and the types of phenotypic mutants possibly found in the generation that had received radiation. The tree species studied were Pinus sylvestris L., Picea abies (L.) H. Karst., Betula verrucosa (Betula pendula Roth), B. Pubescens Ehrh., Alnus glutinosa (L.) Gaertn. and Alnus incana (L.) Moench.
Soaked seeds that had received a rather small dose of radiation germinated usually better than storage-dry seeds, B. pubescens being an exception. The damages observed in germination, height growth and the relative number of mutants were greater the higher the irradiation doses. The LD50 dose (germination, 28 days) was as follows in the case of the different tree species (storage dry/soaked): P. Sylvestris 1,500-2,000/2500-3,000, P. abies 1,000-1,500/4,000-4,500, B. pendula 9,500-10,000/7,000-7,500, B. pubescens >10,000/7,500-8,000 and A. Glutinosa 10,000/8,500-9,000 rad. Mass production of different mutants of deciduous trees for ornamental purposes, for example, appears to be easy using gamma-irradiation. On the other hand, the possibility of increasing tree growth remains open for further study.
The PDF includes a summary in English.
The aim of the present study was to collect information presented in literature concerning the development and germination of the seeds of coniferous trees with special reference to those species which thrive in the conditions prevailing in Finland. Along with the increase in the importance of direct seeding as a silvicultural means in Northern Finland, there is a growing demand for methods by means of which the germination of seeds could be promoted. According to the results obtained from previous studies, such a method can be found, provided that a practically usable and reliable pre-germination method is developed.
The PDF includes a summary in English.
The paper describes the results obtained from an investigation into the effect of ditch spacing, ditch depth and furrowing on ground water table and on development of a Scots pine (Pinus sylvestris L.) plantation on open small-sedge bog in Central Finland (60° 50’ N; 24° 20’ E), drained in 1967. The area was planted in 1968 with 2+1 Scots pine transplants, and fertilized with Y fertilizer for peat soils. The seedlings were measured in 1972.
The depth of the ground water table was greater, the narrower the ditch spacing. The water furrows shortened the duration of the high ground water and lowered the ground water table particularly in the case of ineffective drainage. The narrower the ditch spacing within the blocks, the higher were the young trees. On the other hand, the differences in the height of the trees between the ditch spacings were eliminated by the effect of the furrows.
The PDF includes a summary in English.
The aim of the study was to assess, through field experiments, the possibilities of using peat briquettes in the seeding of Scots pine (Pinus sylvestris L.) in Southern Finland. The briquettes were dug into the soil in the middle of patches of mineral soil. The seeds were covered by a 2-5 mm layer of mineral soil. The seedings were inventoried in the three following autumns.
According to the results, the briquettes were clearly inferior to the control, which was ordinary drill seeding. This was mainly due to the fact that no rain was received after the seeding, and that the third summer from seeding was extremely dry. Abundant germination was observed during the second summer after seeding in both briquette seeding and the control. During more rainy summers the result might have been better.
The PDF includes a summary in English.
The purpose of this study was to compare the development of Scots pine (Pinus sylvestris L.) seedlings sown on substrates off milled peat and milled bark. Mille peat, ordinary milled bark, milled inner bark waste, and a mixture of milled peat and milled bark in the ratio of 1:1, were all compared in the plastic greenhouse. In addition, two fertilization applications were used with milled park: ordinary surface fertilization and double surface fertilization. The germination and development were measured twice during the summer.
It is concluded that milled bark seems to be a rather useful substrate for use in plastic greenhouses, as long as its special requirements are taken into consideration. In the first measurement, there were no differences between the treatments, in the second measurements seedlings growing on a mixture of peat and bark were slightly more developed than the others. Growth of the seedlings was slightly better in ordinary milled bark. Double surface fertilization increased disease and mortality compared to ordinary fertilization.
The PDF includes a summary in English.
The aim of the present study was to assess whether two-year old Betula verrucosa Ehrh. (now Betula pendula Roth.) transplants can be used in afforestation of drained peatlands and what factors affect the development of the young trees. The seedlings were planted in 1967. The site was repair planted next spring due to mortality caused by a undefined fungal disease, and the plantations were fertilized with NPK fertilizer (soil application. The seedlings were measured twice a year until the autumn 1970.
Only 28% of the original transplants, and 73.4% of the repair plantations were alive in 1970. In some cases, fertilization improved the results, while in others it was detrimental to the trees or had no effect on survival. According to peat analysis, the poor survival and development of the plants could be due to the too high ratios of N/Ca and N/P. Stunted or dead trees displayed often necrosis caused by Godronia multispora. According to the experiences, Betula verrucosa plantations are inferior to those obtained with Scots pine (Pinus sylvestris L.). In addition, the results indicate that in old draining areas calcium and phosphorus are often too low in comparison to nitrogen.
The PDF includes a summary in English.
The aim of this study was to estimate the genetic gain of volume growth in Scots pine (Pinus sylvestris L.) selected seed stands. To obtain highest possible accuracy, the estimations are based on a large statistical material comprising 197 separate seed stands. It is concluded that the genetic gain of volume growth ranges between 7.4–15.0%. Unwanted pollen contaminations may, however, in the worst case halve this genetic gain.
The PDF includes a summary in English.
Male and female flowering, cone crop, and some vegetative characteristics were studied in grafts 10 to 16 years of age in a clonal seed orchard of Scots pine (Pinus sylvestris L.). Genetic variation was found between clones in flowering as well as in cone production. Clone evaluation resulted in similar classifications of clones in different years. A regression analysis showed that crown size clearly increased but previous height growth slightly decreased flowering and cone production. The percentage of pollinated female strobili did not differ between clones.
The PDF includes a summary in English.
The paper describes the results of a fertilization experiment, in which transplants of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) were fertilized with various doses of fine-ground copper rock phosphate (33% P2O5, 4% Cu) placed direct in the planting hole. The experiment was made in northeast Finland on a clear-cut, burnt-over and furrowed moraine heath. The fertilization increased especially the survival and condition of the Scots pines and increased to some extent also the height growth of the plants. The spruce survived better than the pines.
The PDF includes a summary in English
The study material included 600 Scots pine (Pinus sylvestris L.) grafts from the Tohmajärvi seed orchard in Eastern Finland. Their broad sense heritability for the height growth was 0.92, for the number of branches 0.87 and for the angle of branching 0.84. Grafts from Central Finland had cones more often than the southern ones, the frequencies being 26.3% and 11.2%. It seems that dominance plays a significant role in the genetical variation of this seed orchard and that height growth is probably more rewarding breeding characteristic than quality, the difference being small, however.
The PDF includes a summary in English.
About 4,000 seedlings of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) were planted in 1965 both on a clear-cut and sheltered area in Central Finland. In the autumn of 1966 needle colour was determined by using Muncell Color Charts which allowed a quantitative measurement of three colour dimensions (hue, value, and chroma). Terminal shoot growth was recorded for two years after colour measurements. In both species, fertilization (NPK in the spring of the year of colour measurement) as well as other site factors caused differences in all three dimensions of needle colour. A regression of shoot growth on needle colour was found in both species. In most cases colour value (darkness) and, in spruce, also chroma, predicted the subsequent growth almost as well as did these two-colour variables together.
The PDF includes a summary in English.
Seedlings of three different Scots pine (Pinus sylvestris L.) nursery stock, 1+0 ,1+1, and 2+0, were kept over the winter, after they had been packed in polythene bags, in three different ways: 1) In a refrigerated storage room, 2) in a wooden crate in the ground, 3) submerged in a lake. The seedling to which they were to be compared with were left over the winter in a nursery bed. The 1,800 seedlings were planted out in the spring 1966 in 15 random blocks. Their development was scrutinized during the three subsequent falls.
The seedlings which had been stored in the lake all died. The seedlings which had been stored along the 1st and 2nd method, managed almost as well as the ones which had been kept over the winter in the nursery bed, except for those of 1+0 stock.
The PDF includes a summary in English.
Scots pine (Pinus sylvestris L.) seedlings were stored for five days in an ordinary wood shed. One half of the seedlings were planted out directly, and another half after soaking the roots of the seedlings for 3–6 hours in water to compensate the possible water deficit developed. According to the results of the experiment, the effect of watering was extremely small. The difference observed, which was in favour of the trees that had been watered during storage, was discernible only in the needle length and in the number of lateral buds; in mortality or in the growth of the seedlings no difference could be observed.
The PDF includes a summary in English.
The paper describes the results obtained from an experiment of fertilization of drained treeless peatlands in connection of planting in three sites in Central Finland. Scots pine (Pinus sylvestris L.) seedlings 2+0 was used. The fertilizer (Y-fertilizer for peat soils, 14% N, 18% P2O5, 10% K2O) was applied in rates of 0, 20, 40 and 80 g/transplant. The fertilizer was strewn either around the plant within a circular patch of 20 cm in diameter, in a ring with a radius of 10 cm and in a ring with a radius of 20 cm. The seedlings were measured two and five years after planting.
The greater the quantity of fertilizer applied and the closer it was applied to the plant the higher was the mortality of transplants. Fertilization increased the mortality during the first two growing seasons after application. Later, however, the mortality decreased to a similar level irrespective the way the fertilizer was applied. In the beginning of the second growing season the fertilized plants showed considerably better height growth than the control plants. The smallest quantity of fertilizer applied produced almost full increase in growth. The pattern of application of the fertilizer had little effect on the growth.
It was concluded that a use of small amounts of fertilizer can be recommended in connection with planting and that it should not be applied very near the seedlings.
The PDF includes a summary in English.
The aim of this study was to investigate how the weight loss and water content of cold stored plants depend on the storage conditions, and if there is a clear connection between these factors and the field survival of the planting stock. The experiments were carried out in a climate chamber at about +2°C and at three moisture levels (about 70, 85, and 95%) from November 1968 to May 1969. Three-year-old seedlings of Scots pine (Pinus sylvestris L.) average length 127 mm, diameter 3.5 mm and the top/root-ratio of fresh weight 1.93, were stored in open and sealed plastic bags. In addition, a transpiration retardant (Silvaplast) was used. The plastic bags (10 plants each) were weighted every 4. week. The remaining 270 seedlings were planted out and inspected after one growing season.
Although the experiment was made in a small scale, the results showed clearly that plant mortality, varying between 3 and 97%, was due to the storage conditions. The weight loss ranged between 2 and 50%, and the correlation between the weight loss and the mortality in the field was high. The water content of the seedlings was about 61%. The correlation between water content and survival was very high. Thus, the determination of weight loss or water content could be a useful method in observing the changes of water balance of the seedling stock during winter-storage. Further investigations are needed to show the tolerable rate of drying out for different sorts of plants. The Silvaplast-treatment had no visible effect either on the drying out or on the field survival.
The PDF includes a summary in English.
This paper reports spot sowing experiments of Scots pine (Pinus sylvestris L.). The seeds were either covered with coarse sand, tramped in the substrate or sowed without any covering, 30 seeds in each treatment in 70 replications. The site was of Vaccinium type with sandy soil. The germination percentage was 81 and 91 on the respective years. The development of seedlings was observed for 3–4 years.
The results indicate that both tramping and covering the seeds to some extent increased the number of seedlings and improved the early development. The highest numbers of seedlings were recorded in the first growing season, after which there was 23 seedlings/100 seeds in the uncovered spots, 27 seedlings in the covered spots and 31 seedlings in the tramped spots in the experiment sowed in 1965.
Mortality of the seedlings was highest between the first and second growing season, and empty spots increased with the time. There was no difference in mortality between the sowing methods, but the number of seedlings after first growing season affected the result. Under favourable conditions four seedlings per spot seemed enough to secure the survival of minimum one seedling per spot during the three first growing seasons. In poor conditions seven seedlings was needed.
The PDF includes a summary in English.
This paper presents the results of a contest performed on behalf of the Finnish bank Kansallis-Osake-Pankki and the Central Forestry Board Tapio on growing trees on peatlands. Over 5,000 sample plots were established on drained peatlands in various parts of Finland. The aim was to achieve a best possible growth of seedling stands on peatland. The factors influencing the growth of 85 best Scots pine (Pinus sylvestris L.) and 60 best Norway spruce (Picea abies (L.) H.Karst.) sample plots were studied.
The height growth of the seedling stands decreased towards the north. Fertilization seemed not to decrease the regional differences; rather on the contrary. On the other hand, fertilization increased height growth, but evidently so that the increase obtained was greater in the southern than in the northern parts of the country. Light fertilization (50 kg/ha of K2O and 60 kg/ha of O2P5) caused a clear increase in height growth while heavy fertilization (100 g/ha of K2O and 120 kg/ha of O2P5), had same effect but to much greater extent than the former. Spruce seedling stands in particular benefitted of the heavy fertilization.
Fertilization did not eliminate the original differences in the quality of the sites in question, but these could still be seen in the height growth after fertilization. The effect of drain spacing on the height growth was not very clear. In dense seedling stands (800 seedlings/ha) the height growth of the dominant seedlings was greater than that obtained in stands of lower density. Hold-overs caused a decrease in the growth of the seedling stands.
The PDF includes a summary in English.
An attempt was made to estimate critically the genetic gain in clonal seed orchards of Scots pine (Pinus sylvestris L.) in Finland. The selection differential of Scots pine and Norway spruce (Picea abies (L.) H. Karst.) is calculated on the basis of filed information on selected plus trees which has been kept by the genetic register at the Finnish Forest Research Institute. The differentials were computed as realized differences in height between plus-trees and normal stand characteristics on respectively site class and as a function of age.
The genetic gain in height growth of Pinus sylvestris was computed on the basis of information on selection differential and heritability. This genetic gain is between 2.6–4.4% provided there is no pollen contamination from unknown sources outside the seed orchard. The genetic gain of volume growth in Scots pine is about 7–15%, provided there is no pollen contamination in the seed orchard. However, according to investigations, there is invariably some pollen contamination in this kind of seed orchards. The contamination decreases to about 30–50% as the orchard matures and starts to produce endemic pollen. If the pollination would be entirely due to foreign pollen sources, the mathematically calculated genetic gain would be 3.5–7.5%.
The PDF includes a summary in English.
The aim of the study was to investigate the effect of four packing methods on the field survival and growth of seedlings and transplants of Scots pine (Pinus sylvestris L.) stored over the winter in a cold-storage cellar. The following sorts of plants were used: one-year-old seedlings (1+0) grown in a plastic greenhouse, two-year-old (2+0) open grown seedlings and three-year-old open grown transplants. These plants were stored in open wooden boxes, in sealed plastic bags, in boxes with wet peat on the bottom and in plastic-laminated paper bags.
The control plants were of the same types and were kept in a nursery over the winter. The storage was carried out in a mantle-chilled cold-storage from October 1966 to May 1967. The temperature in the cold-storage was kept around -2 °C and the relative humidity of the air over 90%. The water content of a randomly selected sample plants showed no increase in water deficit after the storing. Part of the seedlings were transplanted in the nursery and the rest were planted in a clear-cut area. A number of the latter plants were treated with an insecticide (1% Intaktol, which contains DDT, Lindane and dieldrin) before planting. All the experiments were examined after one growing season and the planting experiments the next fall.
The transplants (2+1) in the nursery, and in the forest had survived and grown better than the seedlings. In the nursery the 1+0 seedlings survived and grew better than the 2+0 seedlings. There was no difference in mortality between the seedlings. After the first growing season occasional significant differences between the packing methods were observed, but they disappeared during the second growing season. Thus, all packing methods proved to be as successful as the control method without winter storage.
Transplants were more often attacked by the large pine weevil (Hylobius abietis L.) than the smaller seedlings. The damage, however, was considerably greater on the seedlings because of their lower resistance. No significant differences in the Hylobius-attack between the packing methods could be observed. The Intaktol-treated plants were as often attacked as the untreated ones, but the damage was slighter on the treated ones.
The PDF includes a summary in English.
The aim of the present study was to establish, by means of planting experiments, the influence of different packing, heeling-in and watering as well as the length of the storage period on the development of Scots pine (Pinus sylvestris L.) seedlings, in all 2,090 seedlings, that had been lifted from the nursery bed in spring. The plants were packed in bundles and into plastic sacks in 1965 (6 storage methods) and in 1966 (3 storage methods). Control seedlings were planted without storing at the time when storage of the test material begun. The plantations were followed 3–4 years.
Storage for two weeks in the different ways and planting without storage gave similar results when seedling survival was compared. Storage in plastic sack proved to be as good as storage in bundles in a cellar, and healing-in in moist soil or in a drain were both usable methods. Watering the seedlings did not improve the results, which indicates that the storage caused no serious lack of water.
After four growing seasons an average of 19,6% of the seedlings of the 1965 experiment died, the bulk of them by the end of the first growing season. Despite control treatment, Hylobious abietis caused serious damages. In the plantations of the year 1966 mortality of the seedlings was under 5% by the end of third growing season. During the first two growing seasons after planting differences in growth of the seedlings stored in different ways could be observed in the plantations of the year 1965, but the differences levelled out later. In the plantations established in 1966 no differences in growth occurred.
The PDF includes a summary in English.
The aim of the study was to establish how the cold storage of cones of Norway spruce (Picea abies (L.) H. Karst.) affects the viability of the seeds and the percentage ratio in 7 days. A parallel study was made of the longevity of seed in barn-stored cones subject to weather fluctuations and the longevity of seed extracted immediately and stored in the conventional way in an air-tight container. The cones were collected near Kuopio in Central Finland and near Tampere.
The viability and germination rate of the control sample was constant throughout the storage period. This storage method proved the best. The viability of seeds kept in cones declined in cold storage after 3 ½ months. The cones collected in Tampere were damaged by Laspeyresia strobilella, which affected the viability of the seeds.
The viability of seeds stored in cones in a barn had not weakened by the end of May, however, they deteriorated during the summer, as did the seeds stored in cones in the cold storage. Viability of the seeds was still 94% in October. The germination rate was constant in each lot up to the end of May, after which it decreased to 81.7–86.1% in October.
The results show that healthy spruce cones can be stored in paper sacks in a single layer in cold storage and in an ordinary barn for several months without it affecting the viability of the seeds.
The PDF includes a summary in English.
This project studied the value of various shoot and root-system characteristics as indicators of plantability of transplants. Correlation and regression analysis was used to compare these characteristics. The study material consisted of two-year Scots pine (Pinus sylvestris L.) transplants that had grown in a plastic greenhouse for the first year and then been transplanted in the open. The seedlings had been transplanted in the field without treatment or with the roots cut to a length of 8 cm. A part was transplanted without treatment into plastic pails. A gravimetric and photometric method was used to obtain a description of the surface area of the root systems.
The results show that the photometric value gives a good picture of the surface area of the root system. The greatest advantage offered by the method is the simplicity and rapidity of measurement. The gravimetric, and especially the titrimetric, measurement takes much more time per plant. Photometric measurement affects plantability little, and measured and planted transplants can be followed up in the field. In gravimetric measurements, it was found that fresh and dry weight of the plants were closely correlated.
Mycorrhizal frequency in the root systems gave a good picture of the surface area of the root system. The number of living roots-tips was also rather closely correlated with the surface area of the root system. The other morphological characteristics failed to serve as a satisfactory index for the surface area of root systems. The one closest correlated was the annual leader growth. The second best was stem diameter; the height of the plant, on the contrary, was rather poorly correlated with the other characteristics.
The PDF includes a summary in English.
Certain biocides used in production of tree nursery stock exterminate undesirable organisms but cause an abnormal growth stimulation of plants. The reforestation material has decreased survival potential because of high degree of succulence, top:root and height:diameter ratios, and low specific gravity and root surface area. Some fumigants impede mycorrhizae development and arrest phosphorus uptake. Recovery of growth potential was achieved by aluminium sulphate and/or fermented compost inoculated with mycorrhiza-forming fungi.
The PDF includes a summary in Finnish.
The aim of this study was to establish the need of treatment of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) seeds to be sown in greenhouse. 3 x 100 seeds of each treatment (soaking in water, treatment with Pb3O4, treatment with tiram-containing coating substance) were sown in a glasshouse on a fertilized garden peat, and covered with peat layer of 6 mm thickness. The development of seedlings was followed for 100 days before the final measurement.
Soaking the seeds with water made germination somewhat faster. In spruce the germination percentage increased, but the opposite was observed in pine. No difference could be observed between the results from soaking with acid water from peat soil and lake water. Drying the soaked seeds for a week before sowing had no harmful influence on the germination or the early development of the seedlings. Treatment with Pb3O4 did not affect the germination speed or the seedling percentage of pine or spruce, but increased the germination percentage of spruce. Coating decreased germination and seedling percentages in pine. However, the differences between the treatments were so small that their practical significance is negligible.
Germination of both the species initiated on an average in 8 days, and 16 days after sowing 80% of the seeds had germinated. Seedling mortality was about 10% of the total number of seedlings, the most common reason being damping-off.
The PDF includes a summary in English.
In Finland the mite Nalepella is found in Norway spruce (Picea abies (L.) H. Karst.) in forests practically in every tree, and even in the nurseries. The paper reports on the occurrence of Nalepella Haarlovi var. picea-abietis Löyttyniemi in Finland in tree nurseries in Finland. The study is based on a large material, collected in connection with an investigation into spruce spider mites.
Nalepella lives vagrantly on the needles. Due to the sucking of the mites, the needles turn yellow, become dry an die. Single patches from sucking cannot be seen by the naked eye. They occur on all sides of the needles. The worst damage to spruce seedlings in nurseries is caused to the needles located in the top of the seedling. Sometimes the terminal bud dryes and the whole terminal shoot can die. However, the whole seedlings seldom die in consequence of the Nalepella mite alone. Subsequent damage to the injured needles is often caused by fungus Cladosporium herbarum.
The study shows that the mite causes economically significant damages only in the nurseries. In forests no such damages were observed in seedlings or in older trees. In 1965–68, significant damages occurred in 16 nurseries in Finland. About 600,000 four-year-old seedlings were destroyed in 1967. The damages were economically important only in the 4-year-old seedlings.
According to the study, seedlings damaged by Nalepella can be used for planting as they recover rather well after planting in the forest. Moreover, the damages end after planting, and density of the mite population decreases during the first summer.
The mite overwinters as egg on needles. The eggs hatch in Southern Finland in the end of April and in the beginning of May.
The PDF includes a summary in English.
In early spring 1968 it was noticed that the black grouses (Lyrurus tetris L.) was eating terminal shoots of Scots pine (Pinus sylvestris L.) seedlings in a tree nursery in Luumäki, Southern Finland. The terminal shoots were picked 1–4 cm from the top of the seedlings. In total some thousands of two-year-old seedlings were damaged. The depth of the snow was 10–15 cm deep and only the tops of the seedlings could be seen above the surface of the snow.
The PDF includes a summary in English.
The time interval between final felling and plantation means a waste of time and thus a production loss, and may lead to lush growth of ground cover and hardwood sprouts, which increases expenses in forest management. The objective of this study was to determine the length of time between final felling and artificial regeneration in private forests in the forest districts of Uusimaa-Häme in Southern Finland and Pohjois-Häme in Central Finland. The material consists of a sample of 150 plans of the 952 cutting and regeneration plans in the district of Uusimaa-Häme and a sample of 140 plans of the 1,102 plans in Pohjois-Häme.
The time lag between final cutting and seeding or planting was on average 1.4 years in Pohjois-Häme district and 0.7 years in Uusimaa-Häme. In the latter district, 56% of the logged area was regenerated in the spring immediately following the cutting, and 84% not later than in the second spring. In Pohjois-Häme, 29% of the harvested area was regenerated immediately in the first spring following cutting, and 79% not later than in the third spring following cutting.
In Pohjois-Häme, the interval was shortest in the smallest forest holdings, and longest in the largest holdings with the largest regeneration areas. The length seems to depend mainly on the size of the regeneration area. In Uusimaa-Häme district, the interval was shortest in the smallest holdings, rather short in the largest, and longest in the intermediate-size forest properties. Seeding with Scots pine (Pinus sylvestris L.) was used in almost all regeneration areas.
The forest owners had mainly carried out the regeneration work themselves. In the Pohjois-Häme area, the interval was shorter when the district forestry board regenerated the area. 35% of the regenerated areas had required supplementary planting in Pohjois-Häme, and 47% in the Uusimaa-Häme area. Supplementary planting was more common in areas regenerated later after the cutting. In Pohjois-Häme, according to the reports of the forest owners, 75% of the regenerated areas required tending during the first three years, in Uusimaa-Häme, 80%.
The PDF includes a summary in English.
The paper outlines the information about forest tree nurseries in the Roman Empire, found in ancient writings. According to the author, it cannot be stated that actual forest cultivation was practiced in the times of the Roman Empire, even if tree seedlings were used for a variety of purposes, such as embellishment of cities, parks and gardens, and raising supporting trees in forest vineyards. Nurseries were usually established on farms to fill the owner’s needs. For instance, Gato, Varro, Virgil, Pliny and Colulmella have given instructions about establishment and management of nurseries, and methods to sowing seeds of different tree species. Except for seeds, both root- and branch-cuttings were used in cultivation of trees. Also, grafting was known.
The PDF includes a summary in English.
Transfer of southern provenances of trees to the north leads to an increased growth until the limit of hardiness is exceeded, which may be utilized in practical forestry. Selection from certain local provenances are important both for the immediate supply of seed and on a somewhat longer view. A certain degree of improvement can be achieved by avoiding minus regions and concentrating seed collection on better areas. Also, seeds can be collected from the best stands only, and by accepting only seed from the best trees of such stands. The selection can be strengthened considerably by production of seed orchards. The seed production is based on a small number of trees of particular superiority that are reproduced vegetatively. The clones are planted in an orchard, which pollinate each other.
Hybridization of two different provenances might result in an increased capacity of production. Such hybridized seed can be produced in orchards established, for instance, as a combination of selection and hybridization orchards. As regards the major Scandinavian tree species there are only small prospects of advancement through species hybridization. The method available at present for efficient racial improvement of our forest trees is individual selection in connection with seed production in orchards. In special cases, however, other methods such as transfer of provenances, provenance hybridization, species hybridization, and polyploidization will result in considerable progress.
The PDF includes a summary in English.
The apparent toxicity of soil-incorporated monuron, picloram, CDEC, EPTC, CDAA, and sesone to young Pinus resinosa Ait. seedlings was studied over a temperature range of 10–30 °C in growth chambers. The herbicides were first applied to the surface of autoclaved soil at 1 1b/A and later mixed into the soil. Thereafter pine seeds were planted and subsequent seedling development was studied. The effect of CDEC, EPTC, CDAA, and sesone were also studied at dosages of 2 and 3 1b/A (soil surface basis).
Under the conditions of this study, picloram and monuron were persistent in the soil and toxic to pine seedlings, whereas CDEC, EPTC, CDAA, and sesone appeared to be non-toxic. However, the apparent lack of phytotoxicity of the latter group apparently was caused largely by lack of activation of sesone by autoclaving soil and large losses from the soil of CDEC, EPTC, and CDAA even before seeds were planted.
High toxicity of picloram and monuron was showed by reductions in seedling survival, total dry weight increment of plants, and dry weight increment of surviving seedlings. Various temperature regimes greatly affected growth of herbicide treated plants and controls. In control plants both high and low temperatures adversely affected seedling survival and dry weight more than shoot growth. Temperature extremes generally inhibited root growth more than shoot growth. The high temperatures, 25 and 30 °C, markedly enhanced phytotoxicity of picloram and muron.
The PDF includes a summary in Finnish.
The aim of this present study was to elucidate the quality of seed of foreign tree species grown in Finland, and the factors which have affected the quality of the seed yields. Due to the smallness of the material, however, no far-fetching conclusions can be drawn. The bulk of the seeds were collected in the fall of 1964. The samples of seeds were X-rayed and their classification to empty seeds and full seeds of four quality classes was done on the basis of the anatomical structures. The species studied (a total of 34 species) belonged to the following genera: Abies, Chamaecyparis, Larix, Picea, Pinus, Pseudotsuga, Thuja and Tsuga.
The percentage of empty seeds was throughout quite high. The reasons for the generation of empty seeds probably originate from the special nature of the stands from which the seeds were collected. As a rule, the stands were young and small in area, which may have caused weak pollination and self-pollination leading to embryo mortality. Also, insect damages were observed.
Seeds with albumen still discernible, although the embryo had died, occurred to some extent. In some Larix species, even the bulk of the seeds recorded as full belonged in this group.
The ripening of seeds with embryos was quite successful in spite of the fact that the temperature sum of the year of ripening was slightly below the average in Finland. For instance, all Abies species ripened almost completely.
According to the results, it can be expected that the tree species examined in this study are able to produce rich yields of good-quality seed in Finland, provided that the ovules are well pollinated and self-pollination does not take place to a large extent.
The PDF includes a summary in English.
This paper describes the preliminary results of Scots pine (Pinus sylvestris L.) seeding and planting trials on drained peat soils.
The results showed that a perpared peat surface was a better surface for seeding than the unprepared one. Planting of 2+1-year seedlings succeeded better than planting 1-year seedlings. Planting on the turf gave better survival than planting on the unprepared soil surface. The whole growing season was suitable time for planting Scots pine seedlings except May when the peat soil under the surface was still frozen.
Using fertilizers in connection with planting was surveyed in two ways. Mortality of seedlings increased when they were top-dressed with NPK fertilizer. Using a so-called spot fertilizing with several combinations of fertilizers resulted in K and N tending to increase the mortality of seedlings, but P decreasing mortality.
The PDF includes a summary in English.
When the seed harvest of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) are low, pine and spruce buds are among the secondary food items of squirrel (Sciurus vulgaris L.) in Finland. In this study, conducted in Nokia in Southern Finland in 1962-1963, eating of pine buds by squirrel is described. The eaten buds in 15-years old Scots pine seedlings were recorded in two seedling stands.
According to the results, the squirrels selected the largest buds of the best seedlings in the studied stands. In over 50% of the cases the squirrels chose only the buds of the leading shoot, especially the terminal bud. In half of the trees, a side bud of the leading shoot continued the growth, which causes form defects in the trees. In 35% of the damaged trees, a lateral branch continued the growth. Well-growing seedling stands may be especially susceptible for damages caused by, for instance, squirrels.
The PDF includes a summary in Finnish.
Since 1954 studies have been carried out by the Department of Plant Pathology of Agricultural Research Centre on occurrence of low-temperature parasitic fungi in nurseries in Finland. This paper reports analysis of the damage caused by the fungus to Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) seedlings.
In Southern and southwestern Finland, scarcely any damage caused by low-temperature parasitic fungi to coniferous seedlings was found. On the other hand, in Central, Eastern and Northern Finland, considerable injuries were present in the seedlings. The extent of damage varies between different localities and in a same location from year to year. The extent of damage is mostly dependent on snow cover which is heaviest in Central and Northern Finland. Damages are largest in wooded areas and in places where snow accumulates abundantly and remains until late in the spring.
The principal cause of winter damage to spruce seedlings is Hepotricia nigra (Hartig) which causes black snow mould. Depending on the amount of infestation, the damage can be limited to scattered groups or consist of large areas of dead seedlings. The fungus is unable to infect the plants during warm months of the growing season. The most damaging parasitic fungus in Scots pine is Phacidium infestans (Karst.) causing snow blight. The infestation varies from reddish-brown patches of infected seedlings to large areas of infected plants. Also, Botrytis cinerea has been determined from one- and two-year plants of pine and spruce.
In trials of chemical control by PCNB (pentachloronitrobenzene) gave nearly complete control of low-temperature parasitic fungi in one-year spruce seedlings. In addition, a compound of zineb (Dithane Z-78) gave similar results. Chemical control of the fungi is now common in the nurseries.
The PDF includes a summary in English.
The aim of the study was to find out more about pine weevil (Hylobious abietis L.) injuries in Scots pine (Pinus sylvestris L.) seedling stands and their control by means of DDT. For this purpose, inventories were made of seedling stands established earlier. Control experiments were made on burnt areas by planting seedlings dipped in a DDT emulsion.
The results of the inventories show that injuries caused by pine weevils can, in certain circumstances, especially in seedling stands established by planting, cause the complete failure in artificial regeneration. The extent and quality of the injuries vary greatly according to planting method, treatment of the cutting area, age of the seedling stand, environmental factors, and weather conditions. The most extensive injuries occur in regeneration areas of old Norway spruce stands burnt after clear cutting and planted with Scots pine seedlings. Injuries are greater in seedling stands established by planting, especially after broadcast burning, than in seedling stands originating either from artificial or natural seeding. The quality of the patch for sowing or planting has a considerable effect on the quantity and character of the injuries: in a patch from which organic matter has been removed, injuries do not appear or they are slighter. Seedlings can be protected effectively and economically by dipping their tops up to the root collar, in a DDT emulsion before planting.
The PDF includes a summary in English.
The article reviews the occurrence of damage causes by elk (Alces alces L.) in young Scots pine (Pinus sylvestris L.) stands established by direct seeding in the Ostrobothia region in Finland. The data was collected by random sampling, and consists of 110 sample plots in pine stands, established in 1930-1944.
Signs of elk damages could be observed in 20% of the stands. In more than half of the damaged stands pine seedlings were damaged by elk, on the rest of the stands the damage was targeted to hardwood saplings only. With the present density of elk population, the damage has an insignificant bearing upon the development of pine seedling stands in Ostrobothnia. The weaknesses of silvicultural state of the stands have been caused by other factors than elk.
Silviculturally weak stands were more liable to elk damage than strong ones. The occurrence of elk damage was more usual in stands with hardwood mixture than in pure pine stands. Especially goat willow, mountain ash and aspen, but also to some decree birch, seem to attract elk. Those factors that promote hardwood growth: fertility of the site, swampiness and the presence of seeding hardwoods in the area, increase the stand’s liability to elk damage.
The article includes an abstract in English and a summary in Swedish.
Increase in the elk (Alces alces L.) population and the problems of its grazing has called for detailed research. The present study concentrated on three observation areas representing northern, western and eastern parts of Finland. There were 28 field observers watching 68 elks.
Earlier investigations in Finland indicate that aspen (Populus tremula L.) is the staple diet of elk. This study reached different conclusions, probably largely because of aspen is gradually becoming an increasingly rare tree species in Finland. According to this study, the principal food of elk in the winter is willow (Salix sp.). In the whole country, willow accounts for about 70% of elk’s nutrition. In the Far North the percentage is approx. 90. Of the other tree species, the order of preference is: aspen, Scots pine, mountain ash, juniper and birch. In addition, in Western Finland where snow is less deep, lingonberry and blueberry shrubs are on the menu. Beard moss on the spruce was frequently eaten locally. Elk seems to have eaten mainly the last annual shoot of trees and bushes. In few cases it has gnawed the bark of Scots pine, aspen and willow. Elk consumes in average 340 twigs or terminal shoots per day in the winter. This corresponds to about 1.8 kg of food.
The problem of elks damaging Scots pine seedlings has been observed in Western Finland, were the elk population is higher. The article suggests that suitable feeding places would be left for elk in places that are unsuitable for agriculture or forestry. Leaving, for instance, birch seedlings in Scots pine stands has been noticed to attract elks and to increase the damage to pine.
The article includes an abstract in English and a summary in Swedish.
Prescribed burning has reported to avail forest regeneration, for instance, by releasing nutrients for the use of seedlings, changing the pH of the soil and decreasing competition of ground vegetation. The aim of the study was to find out if the effects could be verified. Sample plots were measured in the experimental area of Tuomarniemi, in Central Finland, both in previously burned and untreated seedling stands and young forests. The main species in the sample plots was Scots pine (Pinus sylvestris L.).
According to the results, prescribed burning prepares the soil for regeneration. Germination percentage of the seeds is higher on the burned soil. All the species, Scots pine, Norway spruce (Picea abies (L.) Karst.) and birch species (Betula sp.) grow faster. Prescribed burning increases the amount of birch seedlings by improving its regeneration compared to unburned sites. The seed trees survive burning better if they are tall and have short crown, and have thick bark. In general, prescribed burning improves regeneration in seed tree stands.
The article includes a summary in German.
Silva Fennica Issue 80 includes presentations held in 1952 in the 7th professional development courses, arranged for foresters working in the Forest Service. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.
This presentation describes principles of breeding of forest trees and the selection of fast growing, healthy and qualitatively good trees, named plus trees, for seed production in Finland.
Silva Fennica Issue 80 includes presentations held in 1952 in the 7th professional development courses, arranged for foresters working in the Forest Service. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.
This presentation describes a field experiment on the use of herbicides to prevent growth of ground vegetation in cutting areas before sowing. The study suggests that the chemicals used in the experiment were not effective enough to prevent growth of ground vegetation in the more fertile lands, but were effective in poorer lands. However, the treatment affected also growth of tree seedlings.
Silva Fennica issue 52 includes presentations held in professional development courses, arranged for foresters working in public administration in 1938. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.
This presentation describes a study of forest management practices suitable for in Hylocomnium-Myrtillus site type forests, typical for Northern Finland. A special problem for the forest site type is poor regeneration due weak seed production of Norway spruce (Picea abies L. Karst.) and thick moss layer.
Finland has a long tradition of grazing cattle in the forests and common land. There are also reports of degradation of forests by grazing already in 1600th century. The aim of the survey was to study which positive and negative effects grazing has in forests.
The study concludes that grazing has caused considerable economic losses through damages to forests. In addition, woodland pastures cannot give the yields required in modern animal husbandry. The quality of woodland pastures have decreased after the woodlands used in slash and burn culture have become wooded.
Grazing has also some positive effects to forests. It increases the diversity of vegetation in the woodland pastures and spreads species to new areas. This is supported by the lists of species found in different woodland pastures. Cattle destroy large grasses like Calamagrostis, which may avail growth of tree seedlings in the pastures. Grazing can also prepare the site for tree seedlings. On the other hand, prolonged grazing destroys tree seedlings and prevents regeneration.
The article includes a German summary.
Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.
This presentation describes production of forest tree seedlings.
Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.
This presentation describes inspection of forest regeneration of mineral soil forest types and drained peatlands, and inspection of ditches.
Silva Fennica Issue 39 includes presentations held in professional development courses in 1935 that were arranged for foresters working in public administration. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.
This presentation describes cultivation of seedlings in forest nurseries.
Regeneration of large open areas in dry mineral soil forest sites that usually grow Scots pine (Pinus sylvestris L.) have several problems. For instance, soil frost, snow, ground vegetation and dryness can prevent germination and growth of seedlings. The damages caused by insects and fungi in seedlings of a large burned area in Siikakangas in Southern Finland was studied. A forest fire burned the area nearly completely in 1909, and 310 hectares have been sowed or planted with mostly Scots pine during the following years. Minor areas have been regenerated with Pinus montana Noll, Pinus excelsa Lamb., Pinus murrayana Balf. and Larix sibirica Ledeb.
No completely healthy pine seedling stands could be found in the area. About 41% of the seedlings in the sample plots were damaged. The most common causes for damage were Evetria resinella (now Retinia resinella L.), Luperus pinicola (now Calomicrus pinicola (Duft.)), Pissodes notatus (now Pissodes castaneus Degeer), Evetria turionana Hb. and Hylobious abietis L. The most usual fungal disease was Lophodermium sp. Evetria resinella caused damages in all the area. Evetria turionana, Pissodes notatus and Hylobius abietina were found in the older seedling stands. Other damages were more localized. The slacks in the terrain seemed to have most damages, the original cause being probably soil frost. Some damages, as Lophodermium, were related to the density of the seedlings, especially in the sown areas. Cleaning of seedling stands could decrease these damages. Planting seems to have succeeded better than patch sowing.
The PDF includes a summary in German.
Norway spruce (Picea abies (L.) H. Karst.) is rarely the dominant species on dry mineral soil sites in Northern Finland. These sites are, in general, too poor and dry for spruce, and suit better for Scots pine (Pinus sylvestris L.). According to the study, the natural regeneration of spruce is in Northern Finland poor. In the sample plots, cones could be found in 35% of spruce trees in the stands in natural state and 46% in the harvested stands. Compared to the spruce areas in Northern Finland, or fresh mineral soil sites in Southern Finland, cone and seed production of Norway spruce was in dry mineral soil sites very low due to scarcity of seed trees and their low cone number. There were few spruce seedlings in the sample plots, but according to the observations, spruce is able to regenerate on lichen and heath covered sites. The seedling growth was, however, poor on dry sites. Spruce seedlings were often found near fallen trees and stumps. The growing trees prevent growth of seedlings of all species. Norway spruce seems, however, to be able to spread also to the poor sites. The success depends on the vegetation and dryness of the site. For instance, spruce can spread to dry mineral soil sites from seed trees of nearby peatlands.
The PDF includes a summary in German.
In this study an attempt was made to use manometric Warburg technique in studying the growing season variations in the respiration rates of the roots of 1–3-year-old seedlings of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.). The respiration rates in both short-roots and long-roots have also been investigated.
According to the results, respiration intensity was the greatest in Scots pine and Norway spruce short-roots but also considerable in the long-root tips at the points of elongation. When the oxygen uptake rate per weight unit in the pine short-roots is given value of 100, the rate in the long-root tips is 61 and in the basal area 36. The corresponding values for spruce are 100, 69 and 43. The relative carbon dioxide release rates are different for the basal parts of the long-roots: pine 53 and spruce 57, when the CO2 release from the short-roots is 100. The CO2 release rate in the basal parts of the long-roots is relatively greater than the oxygen uptake. The respiration rate of the root systems of pine was larger than that of spruce due to the larger size of the root system.
The respiration rate per unit weight of pine roots of the 1- to 3-year-old seedlings decreases significantly with the increasing age. In spruce, the decrease was smaller. The result could have been different if only the short-roots of the same growing season were studied from all seedlings.
During the first growing season the root respiration rate decreased from the middle of the summer towards autumn. An experiment with pine seedlings grown in the mineral soil showed a very rapid increase in respiration rate in the spring. The rate, especially oxygen uptake, is at its greatest in the roots at the time of fastest growth.
Growth-promoting effects of enhanced caron dioxide levels upon forest tree seedlings grown in plastic houses was studied in 1964 and 1965 in the Forest Breeding Foundation in Haapastensyrjä near Loppi in Southern Finland. In both years more vigorous height and weight growth, and development of root system was achieved when the CO2 concentration was increased to 0.2% than in the normal conditions (CO2 0.03%). The CO2 concentration was increased by burning propane in the plastic houses. Burning continued for four hours per day either at 8–10 and 14–16 a clock or 6–10 a clock. Growth was not affected by the time of the treatment, and it was equally high in 0.1% and 0.2% concentrations.
Treatment of the seedlings with 100–200 ppm gibberellic acid (GA) increased the height growth of healthy, well-rooted seedlings. Treatment with a concentrated (600 ppm) dosage, as well as treatment with a combination of GA and 1-naphtyl acetic acid (NAA) caused serious defects in grafts of Scots pine (Pinus sylvestris L.). GA treatments did not induce flower formation in pine. Red light during the night seemed to enhance growth of grafts of silver birch (Betula pendula Roth) and Norway spruce (Picea abies (L.) H. Karst.).
The PDF includes a summary in Finnish.
There has not been complete agreement as to what is meant by ectendotrophic mycorrhizae, and there is a wide variety of opinion among authors on mycorrhizal terminology. In this paper ectendotrophic mycorrhizae are defined to be short roots with Hartig net and intracellular hyphae in the cortex. A mantle and digestion of intracellular hyphae may be found but are not necessary. In the study of Mikola (1965) ectendotrophic mycorrhiza was found to be common in Scots pine (Pinus sylvestris L.) seedlings in Finnish nurseries. The mycorrhizae had always similar structure and the mycelium isolated from the seedlings (E-strains) was similar. The aim of this study was to find out what kind of ectendotrophic mycorrhizae exist in forests and nurseries outside Finland, what kind of mycorrhizae do the E-strains isolated from Scots pine form with other tree species, and are these associations symbiotic.
Only one type of ectendotrophic mycorrhiza was found on the 600 short roots collected from the continents of Europa and America. The type was similar to the one described by Mikola: the mycelium is coarse and forms a strong Hartig net, and intracellular infection is heavy. Evidence is convincing that this structure was formed by the same fungus species. The species is unidentified. Mycorrhizae synthesized by E-strain with six spruce species, fir, hemloch and Douglas fir were all ectotrophic.
The E-type ectendotrophic mycorrhizae proved to be a balanced symbiosis. The seedlings of 13 tree species inoculated with the E-strain grew in the experiment better than the controls. The observation that ectendotrophic mycorrhizae dominates in the nurseries but is seldom found in forests, and then only in seedlings growing in the forest, was confirmed in the study. In synthesis experiments E-strain formed either ecto- or ectendotrophic mycorrhiza depending on the tree species.
The differences between different types of mycorrhiza; endomycorrhiza, ectomycorrhiza and ectendomycorrhiza, and the use of the terms have been variable in the earlier research. Studied of mycorrhiza in Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) seedlings may suggest that the conditions affect which kind of mycorrhiza develops in the seedlings. This study is aimed mainly at finding out whether the difference of ectotrophic and ectendotrophic mycorrhizae depends on fungal symbionts or envirionmental conditions. Furthermore, the occurrence of ectendotrophic mycorrhiza in Finland under various conditions was studied, and experiments on the physiology and ecology of the mycorrhiza and the fungal partner were conducted.
The ectendotrophic mycorrhiza as described in this paper has proved to be very common on Scots pine in Finnish nurseries, but it was not found in Norway spruce seedlings. The results did not support the hypothesis presented in some earlier studies that ectendotrophic mycorrhiza is more parasitic than the other mycorrhizal fungi. The nursery survey showed that no correlation existed between the size and vigour of the seedlings and the presence of ectendotrophic mycorrhiza. Furthermore, greenhouse-grown seedlings with and without the fungus grew equally well. The type of mycorrhiza was, however, almost exclusively confined to young (1–3-years-old) seedlings and to nursery soils. The experiments indicates also that ectendomycorrhizal fungus has a very wide ecological amplitude in regard to light intensity, soil fertility, acidity, and humus content. It has, however, a weak competitive ability in natural forest soils against the indigenous fungal population. When the seedlings were transplanted from the nursery to forest soil, their mycorrhizal population was largely changed.
Seed storing experiments with cones of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) were conducted in Oitti seed extracting plant in Southern Finland from February to December 1955. The pine cones were stores for 267 and the spruce coned for 304 days. In four of the storage methods the cones were packed in sacks and another four in wooden boxes. Sample of cones were taken once a month, seeds were extracted and the germinative capacity was tested. The remaining extracted seeds were placed in storage, and in January 1956 moved to cold seed cellar until 1962, when the viability of the seeds was tested.
According to the results, cleaned pine cones can be stores for at least nine months using almost all methods of storage which are commonly used at our seed traction plants, without hazarding the usability of the seeds. The seeds in spruce cones, however, seemed to be more sensitive to conditions during the storage. The germinative capacity of the spruce seeds began to decrease after the beginning of May. Later the seeds were infected with mould, which increased towards the end of the experiment.
Thus, preservation of the germinative capacity of the seeds of pine and spruce requires storage in different conditions. The results suggest that extraction of spruce seeds should be finished during the cold winter months. It seems that seed in the cones of pine and spruce endure storage in piles of paper or cloth sacks at least as well as in wooden boxes. Occasional warming of the storage, snow and foreign material among the cones and an over meter thick cone layer decreased the germinative capacity of spruce seeds during spring and summer. Spruce seeds that had been extracted immediately after collecting of the cones preserved their germinative capacity well during an eight years storage period.
The PDF includes a summary in English.
Prescribed burning is a common silvicultural practice in northern Europe, intended to destroy the slash and ground vegetation and to reduce the thickness of the raw humus layer prior reforestation. The purpose of the experiments was to study whether there are any differences in the commencement and early development of mycorrhizal infection between burned and unburned areas. A clear-cutting area was burned on May 1961. The soil was rocky moraine, the forest type was Vaccinium type. Two weeks after burning Scots pine (Pinus sylvestris L.) was sown in patches.
According to the results, mycorrhizal infection took place on the unburned area earlier than on the burned. The difference was relatively small, perhaps 1–2 weeks. Although burning kills mycorrhizal fungi, it did not cause serious harm to the seedlings, on the contrary, the favourable influence of burning was more distinct. The high temperatures caused by the fire are restricted in the soil in a prescribed burning only a few centimetres deep. Although the mycorrhizal fungi are concentrated in a very thin surface layer of the soil, some mycorrhizae are situated deeper, and from there the fungi are able to infect roots and spread back to the surface layer. The fire also rises the pH of the soil, which can be harmful for mycorrhizal fungi. Even this effect, however, is limited to a thin surface layer.
The PDF includes a summary in Finnish.
Mycorrhizal association is a characteristic feature of the trees of the northern coniferous forests. The purpose of the present study was to determine what influence some fungicides and herbicides regularly used in Finnish nurseries have on formation and development mycorrhizal in Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) seedlings. The results are based mainly on field experiments in nurseries. First the initiation of mycorrhiza was described in untreated seedlings.
In the first growing season mycorrhizal infection commences fairly late even under normal conditions, i.e. 6–7 weeks after seeding and 3–4 weeks after the formation of the first short roots. Soil disinfectants are commonly used in nurseries before seeding, and they are supposed to evaporate or disintegrate in a few days or 1–2 weeks. In pure culture experiments mycorrhizal fungi proved several times more sensitive than parasitic and indifferent soil moulds to herbicides and fungicides, but in field experiments the delay of mycorrhizal infection caused by them does not seem to harm the seedlings. In the second summer differences of mycorrhizal relations between treated and control plots disappeared. Accordingly, the influence of biocides on mycorrhizae, when applied in the customary concentrations, does not extend beyond the first growing season.
Methyl bromide and SMDC retarded mycorrhiza formation distinctly, while formaldehyde and allyl alcohol had no effect. Apart from not retarding mycorrhizae, formaldehyde and allyl alcohol promoted seedling growth and favoured Trichoderma viride in the soil. Trichoderma is known to be antagonistic to many fungi.
The PDF includes a summary in Finnish.
This paper aims at studying regeneration of Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) by sowing and natural regeneration of birch (Betula sp.) in Western Finland.
Germination of spruce and pine seeds may be prevented by dryness and temperatures below the optimum for germination. In natural conditions, when temperature and moisture is insufficient for germination, the type of seedbed generally has en effect on germination result. Trenching of the seeding spots showed that root competition during the early stage of regeneration was not of decissive importance. It seemed to, however, improve the preservation of the seedlings later. It is common that it can take long before the seeds germinate, and during that time the number of viable seeds decrease strongly.
Also, the seedling stock quickly began to decrease in number after germination, especially during the first growing season and the following winter. The decrease was larger in intact vegetation than on mineral soil or in the humus layer. The emerging seedlings were destroyed by drought very easily, but their tolerance to drought improved later on.
The PDF includes a summary in English.
In this paper the development of sown Scots pine (Pinus sylvestris L.) seedling stands into forests is studied. The material was collected in stands sown in 1930–1940 in private forests in the Forestry Board districts of Central and Northern Ostrobothnia. The 119 areas, including both burned and other areas, were studied in 1955.
Most seedings had been carried out on relatively poor soils, mostly representing Vaccinium and Calluna type forests. 71% of the areas consisted of large forest fires, mostly from 1933. The most burned areas did not have seed producing trees nearby. The other sown areas were in general small, 1–2 ha, and near forests capable of producing seeds. The species of previous tree generation, in the older areas mostly pine and in the younger areas Norway spruce, affected tree species composition of the new tree generation.
Over 90% of the burned areas were in silviculturally good or satisfactory condition, while the main part of the other sown stands was in fair or poor condition. Weeding and thinning had been done only in the oldest stands. Most stands had been left untended. Natural new trees often competed with the sown pines, and cull-trees and border forest increased natural regeneration in the areas. In Calluna type the poor soil limited regeneration and growth of broadleaf trees. The worst competitors were naturally regenerated pine seedlings both on Calluna and Vaccinium type. On Vaccinium type also birch and sometimes also aspen (Populus tremula L.) competed with sown pine. On better sites and paludified areas competition by broadleaf trees increased. The rhythm of development of broadleaved trees is so different from pine that only those broadleaved trees that are formed in the stand when the pine seedlings are larger can develop harmoniously with pine. Due to the harmful competition, the seedling stands should be tended early on. In addition, it may be advisable to abandon the practise to leave trees on sowing areas.
The PDF includes a summary in Finnish.
Prescribed burning has been used in regeneration areas in Finland as a method to treat the humus layer and creating more favourable chemical, physical and biological conditions for the seedlings. At the same time, fire clears away seedlings and shoots of unwanted trees and other vegetation. Direct sowing or planting, mostly Scots pine (Pinus sylvestris L.), seldom natural regeneration, is used. In this paper, the initial stages of the formation of a new tree generation of Scots pine and Norway spruce (Picea abies (L.) Karst.) on prescribed burned areas is studied in Central Finland in 1956–1960.
The burned area remains almost without vegetation for about two growing seasons. Conditions on a burned area which has not been tiled are very unfavourable for germination of seeds of coniferous and deciduous trees. On the other hand, shoots of deciduous trees occur soon after burning. Conditions for regeneration were found to be better 3–5 years after burning. Removal of humus layer in spots improved regeneration. However, the patches facilitated also natural regeneration of Norway spruce and especially birch (Betula sp.), which compete with Scots pine seedlings.
Continuous rainy periods improved the germination of Scots pine and Norway spruce seeds sown on the humus layer. Pine and spruce developed more rapidly on the exposed soil, however, young seedlings were easily destroyed. Seed eaters destroyed the pine and spruce seeds sown on the humus layer of newly burned areas completely or almost completely. The viability of pine seeds sown on the burned humus layer did not decrease for three weeks, but the viability greatly weakened after six or more weeks. Spruce seeds lost their viability faster than pine seeds.
The PDF includes a summary in Finnish.
This paper deals with two machines designed for abrading seed wings, and their influence on the germinative capacity of seed of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.). Both machines are commonly used in Finland.
The results of the study indicate that the act of abrading may cause slight or even serious injuries to the seed. Slight injuries of about 3% are probably not easily avoided if mechanical abrading is resorted to. It must be noted, however, that even this reduction in germinative capacity causes significant yearly loss. If the reduction in germinative capacity is greater, which seems to be possible, it is advisable to test the mechanism of the machine and its method of abrading. As the clearance of the machines can affect the extent of injuries, all machines should be tested. If possible, a continual operation control should be arranged. It could, at the same time, to supply material for improving the abrading method and equipment.
The PDF includes a summary in English.
Litters of different plant species vary greatly in regard to their nutrient content and other properties. The aim of the study was to compare different litters from the standpoint of their value as soil fertilizer. In an experiment Scots pine (Pinus sylvestris L.) seedlings were grown in pot cultures in which known amounts of different litters had been mixed with the soil. The tested litters were Pinus sylvestris (L.), Larix sibirica (Ledeb.), Betula sp., Populus tremula (L.), Alnus incana (L.) Moench, A. glutinosa (L.) (Gaertn.), Sorbus aucuparia (L.), Tilia cordata (Mill.), Acer platanoides (L.), Corylus avellana (L.), Eupteris aquilina (L.), and Deschampsia flexuosa (L.) Trin.
A striking difference was found between alder (Alnus sp.) leaf litter and all the other litters tested. The difference can be seen from the second growth season on, becaus the young seedling uses mainly the nutrients included in the seed. The leaf litter has mainly unfavourable effect on the growth of the pine seedlings. Only both alder species improve the growth. This is mainly due to the nitrogen content of alder leaves. Tree leaves and other forest litter are often composted in the forest nurseries. It seems that adding nitrogen to the compost is necessary, otherwise compost added to the soil may have a harmful effect on the seedlings. Alder, on the other hand, has nitrogen binding Actinomyces growing in symbiosis in its root nodules, and is able to utilize atmospheric nitrogen.
The PDF includes a summary in Finnish.
Natural regeneration of Scots pine (Pinus sylvestris L.) by leaving a seed tree stand on a cutting area has long been the most popular regeneration method in Finland. Results of the method have, however, been unsatisfactory. The aim of the investigation was to study the basic problems of natural regeneration of Scots pine. Regeneration success was studied in 144 sample plots in pine stands at different stages of regeneration in Southern Finland. In addition, the data included information of 42 previously investigated areas.
According to the results, Scots pine can be successfully regenerated naturally on sandy and gravelly soils in Southern Finland. Preparing the ground surface by breaking or burning considerably facilitates the establishment of a seedling stand. The number of seedlings was considerably lower in the ground vegetation than in the mineral soil. Considering growth of the seedlings, root competition of the mother trees was heavy in dense stands, but insignificant in thin stands. The stand density did not affect germination of the seeds. In regeneration areas proper, where the density of mother trees usually is under 50 per hectare, there was in average 4,700 seedlings per hectare in Calluna type forests and 5,200 in Vaccinium type forests.
The PDF includes a summary in English.
Germlings and small tree seedlings are exposed to extreme conditions in the forest floor. In this study the influence of climatic factors to seeds and seedlings were studied experimentally, and an attempt was made to estimate the importance of various factors in several sowing experiments in Finland.
Seeds of Scots pine (Pinus sylvestris L.) were subjected to temperature variations which simulated those of exposed forest sites. The seeds lost some of their germinative capacity during the five-day treatments. Succulent seedlings died when subjected to immersion for 15 minutes at temperatures from 51.5 to 55 ºC. After a hardening pretreatments the seedlings tolerated 2-3 ºC higher temperatures. In artificial humus soil exposed to strong insolation for 15 minutes, temperatures in the range of 54-65 ºC proved to be critical for the seedlings. In natural conditions, also little lower temperatures may prove fatal. Exposure of succulent seedlings of Scots pine and Norway spruce (Picea abies (L.) Karst.) to insolation showed that most damage occurred on humus, quartz sand, and humus-sand mixture, due to rapid evaporation. Seeds of Scots pine, Norway spruce, Betula pendula and Betula pubescens tolerated poorly drought if germination had progressed to a 5–10 mm long radicle. Succulent seedlings tolerated 53-77 days long drought better in humus than in fine silty sand. Seedlings of Pinus sylvestris, Picea abies, Alnus incana and A. glutinosa tolerated cold variably. The developmental stage of the seedling affected cold resistance. Pine seeds sown in furrows germinated well after rain and the survival was high. Frost heaving, snail and insects caused some damages. Germination was lowest at the shallowest furrows. Sowing on natural surfaces gave poor results. Largest damages were caused by birds and ants.
The PDF includes a summary in Finnish.
Root systems of a Scots pine (Pinus sylvestris L.) stands of seed trees on a Vaccinium sites in Southern Finland were studied by taking soil samples around the seed trees. The results show that root system of an old Scots pine spreads relatively evenly around the tree up to at least 10 meters from the stem. The densest part of the root system is near the stem, which part is often acentric. This is probably due to root competition in the early stages of growth of the tree.
Root systems of the seed trees affect stocking of the site with seedlings and the growth of the seedlings. The root competition can cause, for instance, uneven grouping of the seedlings. It seems that the largest trees of a stand have the most even root system. It is therefore recommended to choose the strongest trees of the stand as seed trees, to ensure even distribution of seedlings.
The Acta Forestalia Fennica issue 61 was published in honour of professor Eino Saari’s 60th birthday.
The PDF includes a summary in German.
Pine swamps are easily regenerated by natural regeneration of Scots pine (Pinus sylvestris L.). Usually seeding felling is used, but also strip system or clear cutting and regeneration along stand edge has been suggested. This article discusses the regeneration by clear cutting and sparing the existing undergrowth. The article focuses on pine swamps to be drained and the ones in natural state.
Pine swamps in natural state usually have plenty of trees of smaller diameter classes, that can be trusted to form the future tree generation after the felling. This shortens the rotation by 20-30 years. The undergrowth has been shown to recover quickly. The method suits for regeneration of drained peatlands but could fit also for regeneration of pine swamps in natural state.
The seedlings in the pine swamps are mainly 1-5 years old, and the stock is changing. It seems that larger trees produce a wider selection of age groups, but the seedlings survive longer under smaller mother trees. Part of the younger generations of seedlings seem to be destroyed when the peatland is drained. Further studies are needed to investigate how the draining and felling are to be performed to spare the young seedlings.
The Acta Forestalia Fennica issue 61 was published in honour of professor Eino Saari’s 60th birthday.
The PDF includes a summary in German.
The cone crop of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.) has been assessed in Finland since 1930 annually by sending a questionnaire to forest professionals around the country. Based on the result it is decided if the crop is good enough for collection of the cones next winter. This article presents the results of cone surveys in 1950-1953, and suggest improvements in the method of the investigation.
According to the survey, Scots pine crop was best in 1952, when the crop was intermediate in the whole country, and relatively abundant in the county of Lapland. Norway spruce crop was best in 1951, when the crop was better than in average in the whole country. The evaluators had variable opinions whether the crop was good enough for cone collection or not. They assessed the pine cone collection more often as profitable than the spruce cone collection. Usual reasons to regard spruce cone collection as unprofitable were seed damages and the sites being too far away. To make the results more uniform and accurate, a suggestion to change the evaluation method is presented. The evaluation should be focused on the cone crop of mature stands.
The Acta Forestalia Fennica issue 61 was published in honour of professor Eino Saari’s 60th birthday.
The PDF includes a summary in German.
Planning of large central tree nurseries, which has become topical in Finland, means that the seedlings will be used in a wide geographical area. The nursery must decide which proveniences of seeds of the different tree species it will use. This concerns also the customer that buys the seedlings. The planting and lifting of the seedlings in the nursery have to be timed so that the seedlings are in a right state of growth at the time of planting.
The growth of the seedlings can, under certain conditions, be promoted by using a slightly southerly seed provenience, and large-sized seeds. There are, however, limitations to how much the seeds can be transferred northwards. If the nursery lies much south of the planting site, the seedlings have started height growth at the time of planting. This applies especially larch (Larix sp.), Scots pine (Pinus sylvestris L.) and birch (Betula sp.), but affects less Norway spruce (Picea abies (L.) Karst.). The problem can be handled by using a cool storage space for the seedlings waiting for a delivery in the nursery.
According to an international study, seedlings grown from seeds collected in countries south from Finland usually die already during the first two years in the nursery. Within Finland the seeds can be transferred at least by two latitudes. Spruce seems to tolerate longer transfer. Seed orchards should be planted south of the seed’s origin to ensure better yield and better quality seeds.
The Silva Fennica issue 61 was published in honour of professor Eino Saari‘s 60th birthday.
The PDF includes a summary in German.
There are contrary opinions on the ability of Scots pine (Pinus sylvestris L.) seedlings to withstand oppression by hold-overs and recover after their felling. The recovery potential of oppressed pine stands in Southern and Northern Finland was studied using two kinds of material, fully recovered Scots pine stands and stands recently released. The volume and volume increment of the stand were measured, and the health of the sample trees was determined.
The study showed that those released pine stands that had been in oppressed state very long (25-60 years) had recovered after clear-cutting. After the release the stands grew at first slowly, but after recovery at about the same rate as natural normal stands of a similar height. The smaller, younger, and less stunted the seedlings were when they were released, and the better the site, the faster was the recovery. At the base of released pine stands various defects was detected. When the trees were released, the defects decrease their technical value. A heavy partial cutting had generally a disadvantageous effect on the stand. Recovering seedlings were found clearly to hinder the development of younger seedlings nearby. This inhibition seemed to be a result of the rapid spread of the root system of released pine trees.
The PDF includes a summary in English.
The study is based on observations in a Scots pine (Pinus sylvestris L.) stand on a dry upland forest site in Karhumäki, where a 10-15-year old seedling stand grew under a hold-overs of larger trees that had been left in the site in a previous felling. The root systems of 80-120 cm tall seedlings growing around single mother trees were unearthed. Root maps were drawn of the root systems of 120 seedlings.
No seedlings grew around old, large hold-overs. It seems that seedlings could not compete with their root system. If the hold-overs were stunted in their growth, seedlings grew also under the canopy of the mother tree. 90% of the seedlings had a tap root. Rest of the roots grew horisontally in the topsoil. Around a vigorous mother tree, the seedlings grew their roots away from the mother tree. Hold-overs that had belonged originally to the lower canopy layer of the old forest did not have similar effect on the root orientation of the seedlings. Their roots had been previously affected by trees of higher canopy layer, later removed in the felling.
The PDF includes a summary in German.
Temperatures needed in extracting Scots pine (Pinus sylvestris L.) seeds is relatively high, however, there is little information on its effect on germination of the seeds. This survey aimed at studying how different temperatures affect both extraction result and germination of Scots pine seeds. Comparisons between different temperatures (20, 30, 40, 50, 60 and 70 ºC) were made from cones collected from same sample trees, three trees in total.
Temperatures 20 and 30 ºC resulted in incomplete opening of the cones, and gave thus smaller amount of seeds. Complete extraction requires at the least the temperature of 40 ºC. The result is slightly better in 50 ºC, but germination of the seeds is little lower. Temperatures 60 and 70 ºC improve the results, but in the cost of germination. The main reason for lower germination percentage was that the higher temperatures release more empty and defective seeds from the cones. Results of different sample trees were different due to, for instance, quality and size of cones. Higher temperatures accelerated the extraction. According to the study, perfect extraction in 40 ºC requires longer extraction time than when the temperature of 50 ºC is used. In practice, 50 ºC temperature or even little higher temperatures can be used when the extraction time is shorter. Decessive factors in choosing the temperature would be the humidity of cones and length of extraction time.
The PDF includes a summary in German.
The regeneration of forests in Hämeenkangas area in Southern Finland has been difficult due to various damages from the middle of the 1800s. Few seed trees were left in the area, and artificial regeneration has been used since 1880s. The area became an experimental area of the Forest Research Institute in 1924. The aim of the study was to survey the area before it was transferred to the Finnish Defense Forces.
The original Scots pine (Pinus sylvestris L.) forest of the esker area suffered from many forest fires. The total area is 13,000-14,000 ha, of which the experimental forests of Forest Research Institute cover 6,000 ha. The area is dry upland forest, and drought affects the survival of germlings. Soil frost is a major cause of loss of young seedlings. Sowing method affects the early development of the seedlings. Band sowing proved to be the best method regarding the soil frost. A total of 39 different harmful insect species, 8 pathogen species and 7 other causes of damages have been detected in the area.
The development of seedling stands follow a certain pattern, reported also in other studies. Many of the pine seedling stands develop well until they reach a certain height. After that seedlings begin to suffer from damages, but after reaching another stage develop normally. The damages affect the height growth of the seedlings. Some common damages are caused by Pissoides weevils, needle damages caused by certain beetles, shoot damages by Evetria resinella, and pine blister rust (Peridermium pini and Cronartium flaccidum).
The PDF includes a summary in GermanNatural regeneration has been common in Northern Finland, where forest fires have been usual, and the large areas make artificial regeneration expensive. The regeneration, and for instance tree species composition and density of the stand, cannot been controlled. In Northern Finland there is little demand for Betula sp. which is often abundant in the burnt areas. The unburned forests are generally Scots pine (Pinus sylvestris L.) or Norway spruce (Picea abies (L.) H. Karst.) dominated mixed forests with single Betula sp. trees.
The fire destroys birch for the most part in the Vaccinium site type, but the surviving trees produce enough seeds to regenerate the areas. The largest trees of Scots pine usually survive the fires. Pine has good seed years in the north only every 8th or 10th year. Spruce is totally destroyed in the forest fire and the seedlings grow poorly as primary species. The seedling stands are usually dominated by Scots pine and birch, but birch seedlings grow in batches, and do not hinder growth of pine. The drier Calluna site type stands are dominated by Scots pine. Birch seedlings may be abundant in the beginning, but most of them do not survive. Abundant emergent pine trees prevent the growth of seedlings especially in the dry site types, and they should be thinned to guarantee regeneration. Sowing results are better few years after the fire. The birch seedling should be removed from the seedling stands.
The PDF includes a summary in German.
The development roots of Norway spruce (Picea abies (L.) H. Karst.) seedlings was studied in sample seedlings grown in different kinds of sites. In the early stage, the seedling roots grow primarily length. The main root is usually long. If the growth of the root is hindered, the tip of the root dies, and the root system growing from the original root collar remains relatively small; in these cases, the secondary root system becomes more important. In unfavourable conditions the root branches can early on replace the main root. The main root of a germling seems to be less able to seek for free growing space than the main and side roots of older seedlings. When the growth of the root is blocked by some kind of obstacle, it does not often hinder the growth of the seedling. The type of soil influences strongly how the root system grows. In good soil and in humus the root system is regular and richly branched, while in clay and coarse sand the root system was small. Spahgnum moss was good substrate for seedlings, Dicranum undulatum moss little less good, while the seedlings grew poorly on Pleurozium Schreberi.
The PDF includes a summary in German.
The height growth of Scots pine (Pinus sylvestris L.) seedlings were observed in Korkeakoski and Evo in Southern Finland in 1925-1928. The growth was slow in the beginning of the growing season, increased after that to decrease again towards the end of the growing season. The height growth begun in May, reached the fastest growth rates in June, and ended in June-July. According to the earlier studies, the length of the height growth of Scots pine is dependent on the temperature of the previous summer. This study showed that warm temperatures of the same summer promote height growth, and low temperatures slow it down. Also the daily growth fluctuates, being highest during the afternoon and slowest during the early morning. The daily growth is dependent on temperature.
Norway spruce (Picea abies (L.) H. Karst.) begin the height growth in average 9 days later than Scots pine. Compared to pine, the speed of growth in spruce decreases slower towards the late summer.
The volume 34 of Acta Forestalia Fennica is a jubileum publication of professor Aimo Kaarlo Cajander. The PDF includes a summary in German.
Natural regeneration has been studied so far mostly on the perspective of regeneration, while the production capacity of the seed crop stand has been of little interest in the earlier studies. This paper studied volume growth of the seed trees and the economic impact of this regeneration method both in a literature review and measuring sample stands consisting of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) dominated stands and mixed forests in Norway.
It is concluded that in the most cases the seed tree stands give better pulpwood and timber yield than the stands in the average. The reason might be that the elite stems can better use their production capacity, and, thus, reach bigger and more valuable dimensions. Even though the growing stock diminishes in the seeding felling, the growth of the stand does not decrease or decreases only little. At the same time, the quality growth improves. In addition, the new tree regeneration is achieved usually without extra costs, and there is no unproductive time period in the stand.
The northern range of small leaved lime (Tilia cordata Mill.) in Finland has remained the same since the end of 1800s, but according to the studies of subfossiles of lime found in peatlands, the northern limit has once been higher than at the present. The northernmost natural specimens of the species in Reisjärvi do not produce seed, and are therefore probably a relict. The article includes a review on the distribution of the species in Finland and its capacity to regenerate. The natural regeneration of lime in Finland is at present very rare. The species has lost its ability to sexual reproduction, but reproduces readily vegetatively.
The PDF includes a summary in German.
The article begins on the page 8/122 of the PDF file.
Data for this paper have been collected during years between 1859 and 1862 in the communes of Asikkala, Hollola, Padasjoki, Lammi and Koski.
The first part of the text deals with a general presentation of the geographical, biological and cultural conditions of the studied areas. The second part presents the vegetation according their habitats; they are classified by the degree of moisture. The last is further classified as forested areas or open lands. The examination concentrates on phanerogams (seed plants).
The characteristics of vegetation of forests, open lands and paludified places (swampy or swamp forming places) are presented with the most influential characters and lists of species.
The frequency of years when Scots pine (Pinus sylvestris L.) produces cones and seeds affects its reproduction in the north. The study area covered most of the pine lands in Northern Finland. Scots pine seems to be able to produce cones relatively often in the north. The amount of seeds produced in one year was, however, not sufficient to produce a dense seedling stand. Thus, the natural pine stands contain usually trees in different ages-classes, which have germinated in different years. The cone production is highest in 150-170 years old trees. Pine also needs warm summers to produce viable seeds. Brush fires avail the growth of seedlings, because they clear of ground vegetation that hinders germination of seeds. The seedlings need also moisture to survive; good regeneration years have often had rainy summers.
The PDF includes a summary in German.
The success of natural regeneration of Scots pine (Pinus sylvestris L.) is dependent on the amount and quality of seeds produced in each year. While seed production of Scots pine in Northern Finland is well known, there is little information about frequency of good seed years in Southern and Central Finland. The success of regeneration of Scots pine was studied by defining from the growth rings of felled sample trees in which year they started to grow. The aim was to choose sample trees so that they would form a continual series from seedlings to 100 years old trees. In the material there was too little sample trees from certain years. However, within the period of 1827-1910 it could be identified 13 good regeneration years of pine in Southern Finland. According to the study, pine has regenerated very well every 6th year and at least moderately well every 4th year. When compared to the previous studies made in Northern Finland, part of the good regeneration years seems to be same both in Northern and Southern Finland. Regeneration of Scots pine is affected also by the type and condition of the site.
The PDF includes a summary in German.
The data has been collected from northern Finland between about 64th and 68th north latitudes, excluding the polar tree line areas. The sample plots have been selected to represents greatest variation of the habitats in altitude, rainfall and other site factors. The selected trees were cut, their age, height, and diameter at various heights examined and possible signs of forest fires noticed. The age of the trees was examined also with microscope. The age class distribution of the sample plots was studied. The determination of historical seed years was based on age class distribution of the sample plots.
Grouping of the trees into different age classes was too weak a method to make any conclusions about the periodicity or even less about the frequency of the seed years. There were also noticeable differences in determining the age of the trees between macroscopic and microscopic age examination. There are also differences in the amount of seedlings between different forest types.
The growth of pines in northern Finland is very slow particularly in the young ages but the growth increases after they reach 1,3 meters, which may take up to 30 years. The development of the forest has been similar through last centuriesThe study deals with the preconditions for natural regeneration of the pine at the polar tree line in northern Finland and Sweden. The data has been collected in three summers from 1909 to 1911 in areas of Inari and Utsjoki in Finland and Kaaresuvanto, Jukkasjärvi und Pajala in Sweden. The yearly variation of seed production of pine is studied and compared with the age of the stand, site factors and weather conditions.
Intensity of reproduction is dependent on different preconditions for pistils and stamens and hence the total reproduction (formation of cones) can vary very much depending on the weather.
According the study, the formation of cones varies yearly and is particularly strong every three to four years. Supposedly the same model applies to whole polar tree line of the Fennoscandia. There seem to be no difference between northern and southern parts of Scandinavia when it comes to frequency of good cone years though they not always occur in the same years.Microcatchment water harvesting (MCWH) improved the survival and growth of planted trees on heavy soils in eastern Kenya five to six years after planting. In the best method, the cross-tied furrow microcatchment, the mean annual increment (MAI; based on the average biomass of living trees multiplied by tree density and survival) of the total and usable biomass of Prosopis juliflora (Sw.) DC. were 2,787 and 1,610 kg ha-1 a-1 respectively, when the initial tree density was 500 to 1,667 trees per hectare. Based on survival, the indigenous Acacia horrida Span., A. mellifera (Vahl) Benth. and A. zanzibarica (S. Moore) Taub. were the most suitable species for planting using MCWH. When both survival and the yield were considered, a local seed source of P. Juliflora was superior to all other species. The MAI in MCWH was at best distinctly higher than that in the natural vegetation (163–307 and 66–111 kg ha-1 a-1 for total and usable biomass respectively); this cannot satisfy the fuelwood demand of concentrated populations, such as towns or irrigation schemes.
The density of seeds of woody species in the topsoil was 40.1 seeds/m2 in the Acacia-Commiphora bushland and 12.6 seeds/m2 in the zone between the bushland and the Tana riverine forest. Rehabilitation of woody vegetation using the soil seed bank alone proved difficult due to the lack of seeds of desirable species.
The regeneration and dynamics of woody vegetation were also studied both in cleared and undisturbed bushland. A sub-type of Acacia-Commiphora bushland was identified as Acacia reficiens bushland, in which the dominant Commiphora species is C. campestris. Most of the woody species did not have even-aged population but cohort structures that were skewed towards young individuals. The woody vegetation and the status of soil nutrients were estimated to recover in 15–20 years on Vertic Natrargid soils after total removal of above-ground vegetation.
The utilization of available food resources by the moose (Alces alces L.) was studied in a Scots pine (Pinus sylvestris L.) plantation containing an admixture of deciduous species. Rowan (Sorbus aucuparia L.) and aspen (Populus tremula L.) were highly utilized compared to pine and both silver birch (Betula pendula Roth) and downy birch (B. pubescens Ehrh.). However, they were not capable of withstanding continuous browsing by moose owing to their diminished biomass. In total, the browsing intensity (number of browsed twigs/tree) on pine and birch was about double of that on rowan and aspen.
The number of browsed twigs per tree increased as the amount of available main branches increased. The number of bites per available branch, as well as the maximum diameter of the bites, decreased as the density of the plantation increased. Silver birch was more used by moose than pubescent birch as well as planted silver birch compared with naturally regenerated trees.
Main stem breakage was especially common in winter 1988, the average height of the pine and birch trees being over two meters. The tops of broken stems were commonly utilized as food. The increase in moose density and the relatively deep snow cover evidently promoted the incidence of serious damage. The number of undamaged trees/ha was greater in dense than in sparse parts of the stand.
The PDF includes a summary in Finnish.
The establishment of moose (Alces alces L.) winter feeding sites, their utilization and their effect on damage to young Scots pine (Pinus sylvestris L.) plantations was studied in Ruokolahti-Imatra area in Eastern Finland in 1987–89. During the period, the density in the area was about 3–5 moose/ 1,000 ha.
Six feeding sites were established by fertilization, offering mineral lics and the tops of aspen and Scot pine and by salting the tops of pine. The moose preferred the feeding site to control areas during both summer and winter. In winter browsing was very heavy, especially in those areas located in or close to traditional wintering areas. In winter no moose were seen in the summer habitats.
The extent of, and fluctuations in moose damage were studied in 47 Scots pine plantations in the immediate surroundings of the feeding sites (29 plantations), control areas (18 plantations) and also 68 randomly selected pine plantations. Before the experiment began in 1987 four plantations had been seriously damaged. During the study period only one plantation was seriously damaged. However, it could not be conclusively proved that damage to the pine plantations had been reduced as a result of the feeding sites. The results of the study can be put into practice elsewhere to create better living conditions for moose in their winter habitats. However, the food offered at the feeding site should be in the right proportion to the number of animals wintering in the area, so that the risk of damage to nearby plantations would be kept as small as possible.
The PDF includes a summary in Finnish.
Germination tests in varying photoperiod- and temperature-regimes showed that for early autumn collections, germination of Scots pine (Pinus sylvestris L.) seeds is delayed, especially at low incubation temperature (+10°C) and in darkness. The presence of light during germination (8- or 24-hour photoperiod) or high incubation temperature (+20°C) enhanced germination. As autumn proceeded, a greater proportion of seed were able of germinate in darkness and also in low temperature regime. This result was consistent in both populations studied – in seeds from natural stand (Hyytiälä, Southern Finland) and in seeds from the Hyytiälä clone archive trees, growing in the same site.
An attempt was made to relate the development of germinability during autumn to previously accumulated chilling unit (optimum temperature +3.5°C) sum. Germination percent variation in subsequent cone-collection could not, however, be explained with accumulated chilling.
The PDF includes a summary in Finnish.
This paper is the final report of a study on the damage of tree transplants induced before the trees are planted in the field. The injury development is viewed as a transient rather than a ”step-like” process. The main objective of the study is to develop concepts and methods for recognizing and analysing the dynamic aspects of that process. The report consists of three parts: i) characterization of the environments to which the plants are exposed, ii) theoretical part of model development, and iii) experiments with bare-rooted Scots pine (Pinus sylvestris L.) transplants in which the model was applied.
The results of the first part indicate fast temporal variation of the environmental stress factors, and a slow response of the plant in terms of visible injury symptoms. A model is developed in the second part of the paper for analysing the relationship between the fast stress variables and the slow injury variables. The model is employed in the third part of the study for developing experiments in which the transplants were subjected to different stress conditions. The conclusions emphasize the hazardous role of root desiccation. Similar injury development was observed over a range of different planting sites. Controlling high temperatures during the transportation and storage of the plants was introduced as an indirect method for avoiding the risk of root desiccation.
The PDF includes a summary in Finnish.
An attempt was made in the study to determine the annual periods available for foliage spraying when cleaning Scots pine (Pinus sylvestris L.) dominated seedling stands. The study was made in nine experimental fields which were established in different parts of Finland. The spraying was applied throughout the growing season by DM, MCPA and Roundup. The results were inventoried one year after the treatments.
The results showed that there were big differences both in the destruction of hardwood sprouts and in the survival of pine seedlings due to the time period of the spraying. Threshold points were observed in the range of effect of DM and MCPA. By means of these it is possible to time the spraying treatments in such a way that there remains only slight damage to pine, but hardwood sprouts are destroyed totally. The results varied with Roundup so much, among other things due to rain, that such threshold points could not be determined. This preparation both had a milder effect on the hardwood seedlings and caused slighter damage to pine than the other preparations.
In Sodankylä in Northern Finland, the pines attained a good resistance to arboricides when the efficient temperature sum of the growing season was 550, but in Punkaharju in Central Finland only when it was 850. The seed provenance of the seedlings had an effect on the resistance. The threshold temperature sums of resistance in pine were on the average 70–74% from the long-term average number of degree days at the origin of the seed. The effect on the hardwood trees grew weaker as the long-term average was filled. Resistance of pine followed with a specific lag the lignification of the shoot and the ceasing of the growth of the needles.
The PDF includes a summary in English.
Flowering time and characteristics of cones and seed development of Scots pine (Pinus sylvestris L.) plus tree clones originating from various parts of Finland and planted (grafts) in Southern Finland (61° 48’ N, 29° 19’ E) were studied during 1976-1978.
The flowering time (in terms of period unit (p.u.) sums for flowering) of the Scots pine plus tree clones showed characteristics specific to each population and the characteristics appear mainly adapted to the local temperature factor within Finland. Generally, the development of floral organs, cones and seed in the spring and summer seasons also showed a temperature dependence in that the reproductive organs are developed rapidly and/or favourably under higher temperature conditions within its optimum range.
In this respect, establishment of northern Scots pine seed orchards in Central or Southern Finland or an optimum flowering, and a favourable seed development with an optimum physiological reproductive isolation from surrounding Scots pine populations can be justified. Problem arising from the north-south transfer of seed orchards and the significance of trees’ growth rhythm are discussed in connection with tree improvement.
The PDF includes a summary in Finnish.
An attempt was made in this study to determine which nutrients and in what amounts should be used in the fertilization of Scots pine (Pinus sylvestris L.) seedling stands on nutrient-poor open bogs in order to obtain optimum seedling growth and to minimize the risk of elk damage.
The most important nutrient to improve seedling growth in the experiments was phosphorus. Already rather small amounts produced a significant effect although the effect of higher dosages seemed to be longer lasting. After fertilization also nitrogen gave significant increase in growth. The number of seedlings damaged by elk increased the most on N-fertilized plots. Also, phosphorus increased the occurrence of elk damage, but effect seemed to be related to the better growth and more suitable size of P-fertilized seedlings. The effect of potassium on seedling growth and on occurrence of elk damage was negligible.
The PDF includes a summary in English.
In the experiment Scots pine (Pinus sylvestris L.) seedlings were transplanted out in the field. The effect of the treatments on gas metabolism and daily height increment were examined. The seedlings were 5-year old Scots pine plants growing in clay pots, covered with plastic bags. Transpiration and photosynthetic rates were monitored with open IRGA measuring system for a few days before being subjected to the treatments and for one month after. In addition, the daily amounts of transpired water and daily height increments were measured. A model for the potential rate of each metabolic process was constructed.
Planting and additional exposure had a strong and rather permanent effect on the self-regulation of the processes. This effect is very similar to that caused by water deficit. Exposure makes the disturbance more pronounced. Transpiration of the transplanted seedlings decreased in a few days after planting to less than half of the potential value and that of the exposed ones decreased to a quarter of the potential value. The daily amounts of photosynthesis decreased to half of the potential value. There was no recovery in photosynthesis during the whole monitoring period of four weeks. There was a slight recovery in transpiration about five weeks after transplanting.
Thus, the treatment probably generated stress conditions throughout the whole growing period, which is characterized by strong self-regulation of photosynthesis and transpiration, thus causing an essential decrease in the total amount of CO2 fixed. The photosynthesis was depressed especially at elevated temperatures after planting, as during water deficit. Planting and additional exposure did not produce any detectable changes in the dependence of the growth rate on temperature or in the effect of self-regulation on height growth. On the other hand, the level of growth was decreased as a result of planting out.
The PDF includes a summary in Finnish.
The paper presents some preliminary results of a 10-year-old study the purpose of which is to determine the effect of simultaneous variations in the intensity of drainage and fertilization on the development of planted and natural seedlings on peatlands under various climatic conditions. The development of the Scots pine (Pinus sylvestris L.) seedlings appeared to be better the more intensive the degree of drainage and fertilization used. The increase in the temperature sum had a positive effect on the development of pine seedlings and decreased the mortality rate.
The best growth result was obtained with a 10 m ditch spacing and strong fertilization. As it is difficult to decrease the 10 m ditch spacing for cost reasons, it can be concluded that on such oligotrophic peatlands as were used in this experiment, only an average growth level in the seedling stands can be reached even with the most efficient forest improvement measures. Broadcast fertilization used in the experiment, at least in large doses, increases seedling mortality, as well as the coverage of the ground vegetation, particularly that of cottongrass and fireweed, and also the shrub height, thus increasing competition. It cannot be recommended for afforestation, and today spot fertilization is used. According to this experiment natural seedlings seem once they have recovered after the first years, to grow better than the planted seedlings. This was true especially in the north and in areas, where drainage was not efficient. The height and height growth of the seedlings were to a large extent dependent on the temperature sum.
The PDF includes a summary in English.
In 1965 and 1966 a total of 25 experiments were laid out in various parts of Finland in order to find out the effect of simultaneous variation in the intensity of drainage and fertilization on the development of plantations and natural seedling stands of Scots pine (Pinus sylvestris L.) growing on pine swamps. The fertilizer used was Y fertilizer for peat soils, a fertilizer mixture containing 14 % N, 18 % P2O5 and 10 % K2O. It was applied in rates of 500, 1,000 and 1,500 kg/ha. The ditch spacings studied were 10, 20 and 30 m. The present paper is a preliminary report on a series of studies, the experiments will be observation for a total of 15–20 years.
Mortality of the planted seedlings was found to be the higher after the first growing season, the larger the quantity of fertilizer that had been applied. Fertilizing caused an increase in seedling mortality even after the first growing season following application. At the end of the fifth growing season the height of both natural and planted seedlings is the greater, the larger the quantity of fertilizer that has been applied. Analysis of the height growth of the seedlings showed that larger quantities of fertilizer did not increase growth in the same proportion. The occurrence of growth disturbances is the greater, the more fertilizer has been applied.
Fertilization also changed the composition of ground vegetation. The in the beginning of the experiment birch (Betula sp.) was absent in the area, but was found in the stands the greater abundance the higher application of the fertilizer.
From the viewpoint of growth of the seedlings the best results were obtained with the greatest intensity of fertilization and the narrowest ditch spacing used in the study. The results also show that strong fertilization and a high degree of drainage intensity are not capable of bringing about any particularly good growth on peatlands which originally are relatively poor in nutrients. The growth values now obtained equal only one third of those obtained on peat soils of greater fertility.
The PDF includes a summary in English.
This study was carried out in 1966 in the nursery at Hyytiälä, Korkeakoski Forest District, in Southern Finland. The influence of lifting date (two liftings), way of packing (two methods) and length of storage (one, three and six weeks) on the development of Scots pine (Pinus sylvestris L.), 2 + 1, during the four years following planting was assessed. On the seedlings stored for six weeks, the influence of compensating for the water deficit was also studied.
According to the results, the lifting carried out later, i.e. when the seedlings had already started growth, gave slightly better results than when seedlings were lifted earlier. No difference could be observed for seedlings stored for one week, but for the seedlings stored for three or six weeks, mortality in the lot lifted earlier was 6- to 7-fold that of the seedlings lifted later. The main reason for this was probably that the seedlings of the earlier lifting suffered from lack of water at the time of lifting.
The growth of the seedlings lifted earlier and stored for three weeks showed a decrease compared to those lifted later. For the seedlings stored for six weeks, on the other hand, faster growth was recorded for both the seedlings of the earlier and the later lifting in comparison with those stored for shorter times. Watering increased to some extent the growth of the seedlings stored for six weeks.
During the normal, one- and three-week storing periods, seedlings were well preserved when packed both in bundles and in polythene sacks. Three years after the planting the average mortality was about 10%. Effect of watering was large for those seedlings that had been longer in the storage.
The PDF includes a summary in English.
Heritability of first and second-year height growth of Pinus sylvestris (L.) plants was studied using three different mating designs. Plants grew in a plastic greenhouse during the first growth period. During the second growth period they were exposed to open air conditions.
Heritability values varied from zero to 42%. An average heritability based on six separate calculations (two from each mating design) was computed and its reliability was discussed. This heritability value was 18%.
Different ways of improving heritability estimates was discussed and the need of more investigations on trait quality in relation to fitness and on the effect of natural selection and population strategy was emphasized.
This paper is a report of the authors visits to over 80 forestry nurseries in 20 countries mostly in the tropics or subtropics. The article aim is to describe the methods used in the various countries and compares them to the conventional methods of cool and temperate countries. The article introduces nurseries of Africa south of the Sahara, Mediterranean area, Australian and New Zealand and Latin America.
A complete revolution has taken place in the Finnish nursery practice, which used to raise the seedlings in natural field soil in open-air nurseries. The seedlings were usually transplanted into transplant beds at the age of two years. Now the use of plastic greenhouses of light construction and an artificial soil substrate (fertilized peat) are essential. The new technique has some similarities to the practises of the tropical and subtropical nurseries. In Finland cultivation in greenhouses has hastened the development of the seedlings and shortened the nursery rotation from four to two years, and provided better control of watering and fertilization.
Peat beds in greenhouses are used also in Swaziland. The advantage of peat is that it is free of weed seeds, which eliminates weeding. Peat substrate gives also better yield of seedlings, which decreases the need of seeds, which is important in Finland. Another technique common with tropical silviculture is the production of potted seedlings, which are easy to handle and transport. In tropics, peat pots (jiffy pots) have made it possible to grow plantable seedlings in one season without transplanting. The present Finnish technique means a decreased degree of mechanization compared to the conventional technique of modern European and American nurseries.
The aim of this study was to find out the planting vigour of Scots pine (Pinus sylvestris L.) stored over the winter either in winter storage mainly in the temperature of 4 – -6 °C or in nursery beds. The experimental planting included about 4,500 of 2+1 transplants in Northern Finland. In spring 1965 the control plants were lifted in the spring before budbreak and stored in closed bags in a cold store, in the following year the control plants were lifted in June when the growth had started.
Winter storage of pine transplants in a cold store, tightly closed into bags for the major period, did not, according to the results, increase plant mortality as compared to lifting in the spring. Soaking the stored-plant roots did not affect plant mortality. Mortality was rather small in all treated lots and probably more dependent on planting site and other local factors.
No consistent difference on the leader growth, needle length, bud number and plant grade was found between the plants stored over winter and those lifted in the spring. Sealing the plants into tight bags for winter proved to be suitable method, efficiently preventing water shortage in the plants. No moulds or fungal diseases were found in the plants. In the exceptionally cold 1965–1966 winter, temperature in the cold store sank to -15 °C, but in spite of the temperatures below the recommended storing temperature, the plants survived well. The reason was that the plants froze slowly in the fall and thawed out slowly in the spring.
The value of vigour grade in predicting plant-characteristic development proved to be good, and predicted plant development also in the following year fairly well.
The PDF includes a summary in English.
After a decades-long increasing trend, the recent results of the National Forest Inventory (NFI) reported a decline of forest growth in North Finland. The aim of this study was to assess climatic and reproduction influences behind the growth decline. We used tree-ring data that had been collected by NFI using systematic sampling. The tree-ring width series were detrended using the regional curve standardisation (RCS) removing age-related trends. The resulting tree-ring indices of Scots pine (Pinus sylvestris L.) showed decadal variations with low increment in the 1990s, and high increment in the 1980s and the early years of the current century. Thereafter, a prolonged growth reduction for pine started both on the mineral soil sites and peatlands. The tree-ring indices of Norway spruce (Picea abies (L.) Karst.) had less pronounced decadal variations and no trend-like reduction over the last 15 years. High spring and summer temperatures were found to enhance radial growth, but high winter temperatures were related to low growth for pine and spruce in the following summer. Temperature variation, accompanied by variables indicating years of drought and intensive flowering, accounted for 34% annual growth variance of pine and 21–44% for spruce. Thus, the results imply that climatic factors may have to some extent contributed to the recent growth reduction of pine. Due to its ecological and economic consequences growth decline needs to be further monitored and investigated. Moreover, analyses of stand and age structure, potentially affecting the growth decline, were beyond the scope of this paper, but also warrant further investigation.
Because today’s tree planting machines do a good job silviculturally, the Nordic forest sector is interested in finding ways to increase the planting machines’ productivity. Faster seedling reloading increases machine productivity, but that solution might require investments in specially designed seedling packaging. The objective of our study was to compare the cost-efficiency of cardboard box concepts that increase the productivity of tree planting machines with that of today’s two most common seedling packaging systems in southern Sweden. We modelled the total cost of these five different seedling packaging systems using data from numerous sources including manufacturers, nurseries, contractors, and forest companies. Under these southern Swedish conditions, the total cost of cardboard box concepts that increase the productivity of intermittently advancing tree planting machines was higher than the cost of the cultivation tray system (5–49% in the basic scenario). However, the conceptual packaging system named ManBox_fast did show promise, especially with increasing primary transport distances and increased planting machine productivities and hourly costs. Thus, our results show that high seedling packing density is of fundamental importance for cost-efficiency of cardboard box systems designed for mechanized tree planting. Our results also illustrate how different factors in the seedling supply chain affect the cost-efficiency of tree planting machines. Consequently, our results underscore that the key development factor for mechanized tree planting in the Nordic countries is the development of cost-efficient seedling handling systems between nurseries and planting machines.
Our main objective was to determine whether various genetically improved reproductive materials of Scots pine (Pinus sylvestris L.) differ in growth rhythm, autumn cold acclimation and resilience from unimproved materials. The study consisted of two successive indoor experiments with Scots pine seedlings representing four levels of genetic gain (unimproved natural stands, first-generation seed orchards, 1.5-generation seed orchards and seed orchards established with freezing-tested parents) and a wide range of geographical origins within Finland. The seedlings were assessed for terminal shoot elongation, growth cessation, bud set, freezing injuries and bud flushing over the first growth period. All the adaptive traits showed a latitudinal trend regardless of the genetic level. Seed orchard progenies and natural stand progenies did not differ significantly in the timing of growth cessation, bud set, and the flushing rate of the frost-injured seedlings, after the trait variation was adjusted to the latitude of origin. The differences in autumn frost hardiness were insignificant, too, except for the somewhat higher injury rate displayed by the first-generation seed orchard materials. The finding was not conclusive due to ambiguous results from the two experiments. Overall, we did not find evidence of alarming compromises in the adaptive performance of genetically improved materials.
Handeliodendron bodinieri (H. Lév.) Rehder is a rare, endangered, and therefore, protected tree species native to China. However, there are serious limitations to the effective protection of the species, including a low seed germination-rate and difficult storage due to a high seed oil-content. Here, we evaluated the feasibility of ultra-dry seed storage and its effects on seedling growth. We used the silica gel method to prepare ultra-dry seeds with different moisture contents to find an optimal moisture content range (2.54%–4.77%). Ultra-dry treatment improved storability of H. bodinieri seeds. Furthermore, seeds with a moisture content of 4.77% stored at room temperature, and seeds with a moisture content of 3.97% stored at 4 °C yielded the best results. Priming with an appropriate concentration of polyethylene glycol had a certain repairing effect on ultra-dry stored seeds and improved seed vigor, with a two-day priming treatment with 20% polyethylene glycol having the best effect. Finally, compared with sand storage at 4 °C, ultra-dry storage promoted seedling growth and root development; furthermore, it alleviated storage damage to H. bodinieri seeds, promoted soluble sugar and soluble protein accumulation, and increased seedling nitrogen, phosphorus, and potassium uptake. Therefore, ultra-dry storage can be effectively used to preserve H. bodinieri seeds. Specifically, low-temperature storage of ultra-dry seeds with a moisture content of 3.97% enhanced H. bodinieri seed vigor, and seedling growth and development.
Mechanical site preparation methods that used tools mounted on lightweight excavators and that provided localised intensive preparation were tested in eight experimental sites across France where the vegetation was dominated either by Molinia caerulea (L.) Moench or Pteridium aquilinum (L.) Kuhn. Two lightweight tools (Deep Scarifier: DS; Deep Scarifier followed by Multifunction Subsoiler: DS+MS) were tested in pine (Pinus sylvestris L., Pinus nigra var. corsicana (Loudon) Hyl. or Pinus pinaster Aiton) and oak (Quercus petraea (Matt.) Liebl. or Quercus robur L.) plantations. Regional methods commonly used locally (herbicide, disk harrow, mouldboard plow) and experimental methods (repeated herbicide application; untreated control) were used as references in the experiments. Neighbouring vegetation cover, seedling survival, height and basal diameter were assessed over three to five years after plantation. For pines growing in M. caerulea, seedling diameter after four years was 37% and 98% greater in DS and DS+MS, respectively, than in the untreated control. For pines growing in P. aquilinum, it was 62% and 107% greater in the same treatments. For oak, diameter was only 4% and 15% greater in M. caerulea, and 13% and 25% greater in P. aquilinum, in the same treatments. For pines, the survival rate after four years was 26% and 32% higher in M. caerulea and 64% and 70% higher in P. aquilinum, in the same treatments. For oak, it was 3% and 29% higher in M. caerulea and 37% and 31% higher in P. aquilinum. Herbicide, when applied for three or four years after planting, provided the best growth performances for pines growing in M. caerulea and P. aquilinum and for oaks growing in P. aquilinum. For these species and site combinations, DS+MS and DS treatments reduced the neighbouring vegetation cover for one to four years following site preparation.
In Nordic countries, tree planting of seedlings is mainly performed during spring and early summer. Interest has increased in extending the planting window throughout the unfrozen growing season. This study compared the success of one-year-old spring, summer and autumn plantings in practical forestry in Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) in southern and central Finland. Planting success was based on the number of viable seedlings per hectare relative to a species-specific target density. The influence of different factors to poor planting results were determined, including quality of site preparation and planting, and sources of natural damage. Overall, in Norway spruce, 85, 69 and 84% and in Scots pine 53, 55 and 40% of spring, summer and autumn plantings succeeded. In Norway spruce, the planting results were consistent between the southern and central regions, whereas in Scots pine, the success was slightly lower in the south. The poor work quality and a low density of appropriate planting spots, contributed to poor planting results, regardless of planting season, region or tree species. Considering different damages, especially mammal damage contributed to the failure of Scots pine spring plantings, whereas in summer plantings, corresponding single failure reason could not be identified. Based on our findings, extending the planting season of Norway spruce could be recommended in both regions. For Scots pine, there is still significant uncertainty about the success of summer and autumn plantings, partially due to the limited number of plantings available for analyses.
We studied variations on different traits of Parkia timoriana (D.C.) Merr. in twelve provenances systematically from their source of origin to a controlled environment where representative seedlings were grown. Among the provenances, P1 gave the best result for seed traits including germination traits, P7 for pod traits and P10 for seedling vigour. Effects of seasonal distribution of rainfall and temperature on seed and pod traits were also determined by computing multiple regression analysis. The results displayed winter rainfall and summer temperature as the most important factor determining pod and seed traits. Latitude also significantly (P < 0.001) affected PWT (r = 0.52), SWP (r = 0.46) and SW (r = 0.50). A common garden study for germination and seedling growth indicated P1 and P10 provenance as the best among all. Seeds drawn from P10 gave the highest seedling vigour with an average growth rate of 0.61 cm/day from 90th to 180th day. Highest broad-sense heritability values (h2) were observed in germination traits, followed by seedling collar diameter. The lowest h2 was observed for seedling height.
In the boreal forest of eastern Canada, a large proportion of black spruce (Picea mariana [Mill.] Britton, Sterns & Poggenb.) stands are affected by paludification. Edaphic conditions that are created by paludification processes, including an abundance of microsites with high moisture and low nutrient contents, hinder forest regeneration. Disturbance of paludified sites by mechanical soil preparation (MSP) reduces organic layer thickness, while generating a range of substrates for regeneration establishment. Yet, little information is available regarding the effects of these substrates on tree growth. Our objective was to determine the effect of organic, mineral and organo-mineral substrates that are created following MSP of a paludified site on the growth and root development of black spruce seedlings in a semi-controlled environment. We demonstrated that substrate exerted a significant effect on seedling growth and foliar concentrations of N, P and K. Increase in height and diameter were respectively greatest on clay (mineral) and mesic substrates. Substrate effects did not affect total biomass increases or final root biomass. Foliar nutrients (N, P, K) were relatively high in seedlings that were established on mesic substrates and relatively low for those established on clay substrates. To ensure successful seedling establishment, we recommend the application of MSP techniques that expose organic-mesic substrates on sites that are susceptible to paludification.
The genus Betula L. is composed of several species, which are difficult to distinguish in the field on the basis of morphological traits. The aim of this study was to evaluate the taxonomic importance of using visible + near infrared (Vis + NIR) spectra of single seeds for differentiating Betula pendula Roth and Betula pubescens Ehrh. Seeds from several families (controlled crossings of known parent trees) of each species were used and Vis + NIR reflectance spectra were obtained from single seeds. Multivariate discriminant models were developed by Orthogonal Projections to Latent Structures – Discriminant Analysis (OPLS-DA). The OPLS-DA model fitted on Vis + NIR spectra recognized B. pubescens with 100% classification accuracy while the prediction accuracy of class membership for B. pendula was 99%. However, the discriminant models fitted on NIR spectra alone resulted in 100% classification accuracies for both species. Absorption bands accounted for distinguishing between birch species were attributed to differences in color and chemical composition, presumably polysaccharides, proteins and fatty acids, of the seeds. In conclusion, the results demonstrate the feasibility of NIR spectroscopy as taxonomic tool for classification of species that have morphological resemblance.
The mean temperature during the potential growing season (April–September) may increase by 1 °C by 2030, and by 4 °C, or even more, by 2100, accompanied by an increase in atmospheric CO2 concentrations of 536–807 ppm, compared to the current climate of 1981–2010, in which atmospheric CO2 is at about 350 ppm. This may affect both the growth and frost hardiness of boreal trees. In this work, we studied the responses of height and autumn frost hardiness development in 22 half-sib genotypes of one-year-old Norway spruce (Picea abies (L.) Karst.) seedlings to elevated temperatures and atmospheric CO2 concentration under greenhouse conditions. The three climate treatments used were: T+1 °C above ambient and ambient CO2; T+4 °C above ambient and ambient CO2; and T+4 °C above ambient and elevated CO2 (700 ppm). The height growth rate and final height were both higher under T+4 °C compared to T+1 °C. Temperature increase also delayed the onset, and shortened the duration, of autumn frost hardiness development. Elevated CO2 did not affect the development of height or frost hardiness, when compared to the results without CO2 elevation under the same temperature treatment. Higher temperatures resulted in greater variation in height and frost hardiness development among genotypes. Three genotypes with different genetic backgrounds showed superior height growth, regardless of climate treatment; however, none showed a superior development of autumn frost hardiness. In future studies, clonal or full-sib genetic material should be used to study the details of autumn frost hardiness development among different genotypes.
Light is an important environmental factor for all green plants. Its intensity, spectral composition and photoperiod can affect the regulatory pathways in plants that lead to floral initiation. In this report, results are presented from three experiments in which supplemental light with metal halide lamps (250 µmol m–2 s–1, 20 hours day–1, approx. 6 weeks) was tested as a complement to other flowering stimulation treatments (elevated temperature, treatment with gibberellin A4 and A 7 (GA4/7), restricted water supply) applied to potted Picea abies (L.) Karst. in the greenhouse. Flower stimulation in a greenhouse resulted in more floral initiation compared to flower stimulation outdoors. Supplemental light treatment increased floral initiation further, and to a larger extent in female than in male flowers. It also increased the proportion of trees and genotypes that induced reproductive buds. In a practical application of the supplemental light treatment to potted Picea abies breeding material, 90.6% of the clones produced either female or male flowers, or both. A subset of the same material kept outdoors, and thus subjected to natural light and temperatures, produced no flowers despite being treated with GA4/7 and receiving a restricted water supply. In conclusion, supplemental light treatment facilitates breeding programmes, and seed production of highly improved base material from new selections for vegetative production programmes, to be more efficient.
The pine weevil Hylobius abietis L. is an economically important pest insect that kills high proportions of conifer seedlings in reforestation areas. It is present in conifer forests all over Europe but weevil abundance and risk for damage varies considerably between areas. This study aimed to obtain a useful model for predicting damage risks by analyzing survey data from 292 regular forest plantations in northern Sweden. A model of pine weevil attack was constructed using various site characteristics, including both climatic factors and factors related to forest management activities. The optimal model was rather imprecise but showed that the risk of pine weevil attack can be predicted approximatively with three principal variables: 1) the proportion of seedlings expected to be planted in mineral soil rather than soil covered with duff and debris, 2) age of clear-cut at the time of planting, and 3) calculated temperature sum at the location. The model was constructed using long-run average temperature sums for epoch 2010, and so effects of climate change can be inferred from the model by adjustment to future epochs. Increased damage risks with a warmer climate are strongly indicated by the model. Effects of a warmer climate on the geographical distribution and abundance of the pine weevil are also discussed. The new tool to better estimate the risk of damage should provide a basis for foresters in their choice of countermeasures against pine weevil damage in northern Europe.
Silver birch (Betula pendula Roth) seed origins from the Baltic countries and from Finland were compared in terms of growth, wood density, bark thickness and the incidence of darkened core wood, frost cracks and decay, and the effect of seed origin latitude was examined in two Finnish provenance trials. The material consisted of 21 stand and single tree origins ranging from latitudes 54° to 63°N from the Baltic countries and Finland. The trials, measured at the age of 22 years, were located at Tuusula (60°21´N), southern Finland and at Viitasaari (63°11´N), central Finland. The Baltic origins were superior to the Finnish ones in the southern trial regarding height, whereas in central Finland the Finnish origins grew better. There was no consistent difference between the Baltic and the Finnish group of origins in wood density. Bark thickness decreased with increasing seed origin latitude. The Baltic origins had significantly thicker bark than the Finnish origins. A moderate positive correlation was detected between the seed origin latitude and the incidence of darkened core wood in the southern trial, where the darkened core wood was more common in the Finnish origins than in the Baltic ones. The highest proportion of trees with frost cracks was detected in the south-western Latvian origins growing in central Finland. Seed transfers from the Baltic would have an increasing effect on the bark thickness of birch logs, but no or only minor effects on wood density. Based on our results, there is no reason to recommend the use of non-native Baltic seed origins in Finland instead of the native locally adapted seed sources.
In the present study, the impact of geographical isolation and habitat conditions on genetic diversity and sexual reproduction was tested in four relict populations of dwarf birch Betula nana L. in Poland and Belarus. Amplified fragment length polymorphism (AFLP) method revealed that the endangered central European stands were not genetically extirpated compared with the widespread localities from Finland and Russia, which can result from infrequent outcrossing events in long-living clonal populations. However, genetic clustering methods indicated significant differentiation of the Polish populations because of their small sizes and long-term geographical isolation. Considerable numbers of empty seeds were observed in both relict and central locations, although fully developed seeds germinated better in widespread populations. Analysis of groundwater chemical parameters indicated that two relict populations were significantly different from the remaining samples with respect to pH, electrical conductivity and concentrations of phosphorus ions, which can also influence the efficiency of sexual reproduction. In the light of results obtained it seems that endangered B. nana localities are relatively stable.
Stored nutrient reserves are closely correlated with survival and growth of transplanted seedlings. Previous studies have proven that combining pre-hardening fertilization (PF) with fall fertilization (FF) built seedling nutrient reserves more effectively; however, their effect on transplanting performance is poorly documented. We investigated the independent and interacting effects of 2 levels of PF and 4 levels of FF on seedling growth, nutrient acquisition and accumulation during different growth phases 1 year after transplanting of Quercus variabilis Blume in a nursery. High PF benefited nutrient reserves and subsequent transplanted seedling growth and tissue nutrient storage at the end of the rapid growth and hardening phases. Fall fertilization with 36 mg N increased stem dry mass and tissue nutrient content at the end of the hardening phase. At the conclusion of establishment, PF and FF showed a significant interaction for N and K uptake from soil. At the end of the rapid growth and hardening phases, high PF consistently increased nutrient uptake. Enhanced N and K uptake occurred following application of 36 mg N of FF at the end of the hardening phase. Distinct roles for PF and FF on 3 phases of transplanted seedlings demonstrated the necessity to evaluate fertilization in terms of nutrient reserves and subsequent transplanting performance in consecutive phases. Combining 100 mg N seedling–1 during pre-hardening with 36 mg N seedling–1 during fall yielded ideal transplanting performance for Quercus variabilis seedlings.
The occurrence of moose damage was studied using data from three National Forest Inventories (NFIs) accomplished between 1986 and 2008 in Finland. The combined data included a total of 97 390 young stands. The proportion of moose damage increased from 3.6% to 8.6% between the 8th NFI (1986–1994) and the 10th NFI (2004–2008). The majority (75%) of the damage occurred in Scots pine-dominated stands. The proportion of damage was higher in aspen-dominated stands than in stands dominated by any other tree species. The tree species mixture also had a clear effect on the occurrence of damage. Pure Scots pine stands had less damage than mixed Scots pine stands, and moose damage decreased linearly with the increasing proportion of Scots pine. Stands on mineral soil had more frequent moose damage than stands on peatlands. The fertility class of the site had no straightforward effect on the damage frequency. Artificially regenerated stands had more damage than naturally regenerated stands. Accomplished soil preparation measures and the need for thinning or clearing operations increased moose damage. High proportions of moose damage in young stands were found around the country. In the 10th NFI, the largest concentration of damage was found in southwestern Finland. Our study shows the temporal and spatial changes in the occurrence of moose damage and pinpoints some important silvicultural factors affecting the relative risk of young stands over a large geographical area.
Larix sibirica Ledeb. is one of the promising timber species for planting in the boreal ecosystem; but poor seed lot quality is the major hurdle for production of sufficient quantity of planting stocks. Here, we evaluated the potential of Near Infrared (NIR) Spectroscopy for sorting viable and non-viable seeds, as the conventional sorting technique is inefficient. NIR reflectance spectra were collected from single seeds, and discriminant models were developed with Orthogonal Projections to Latent Structure – Discriminant Analysis (OPLS-DA). The computed model predicted the class membership of filled-viable, empty and petrified seeds in the test set with 98%, 82% and 87% accuracy, respectively. When two-class OPLS-DA model was fitted to discriminate viable and non-viable (empty and petrified seeds combined), the predicted class membership of test set samples was 100% for both classes. The origins of spectral differences between non-viable (petrified and empty) and viable seeds were attributed to differences in seed moisture content and storage reserves. In conclusion, the result provides evidence that NIR spectroscopy is a powerful non-destructive method for sorting non-viable seeds of Larix sibirica; thus efforts should be made to develop on-line sorting system for large-scale seed handling.
A large quantity of non-viable (empty, insect-attacked and shriveled) seeds of Juniperus polycarpos (K. Koch) is often encountered during seed collection, which should be removed from the seed lots to ensure precision sowing in the nursery or out in the field. The aims of this study were to evaluate different modelling approaches and to examine the sensitivity of the change in detection system (Silicon-detector in the shorter vis-a-vis InGsAs-detector in the longer NIR regions) for discriminating non-viable seeds from viable seeds by Near Infrared (NIR) spectroscopy. NIR reflectance spectra were collected from single seeds, and discriminant models were developed by Partial Least Squares – Discriminant Analysis (PLS-DA) and Orthogonal Projection to Latent Structures – Discriminant Analysis (OPLS-DA) using the entire or selected NIR regions. Both modelling approaches resulted in 98% and 100% classification accuracy for viable and non-viable seeds in the test set, respectively. However, OPLS-DA models were superb in terms of model parsimony and information quality. Modelling in the shorter and longer wavelength region also resulted in similar classification accuracy, suggesting that prediction of class membership is insensitive to change in the detection system. The origins of spectral differences between non-viable and viable seeds were attributed to differences in seed coat chemical composition, mainly terpenoids that were dominant in non-viable seeds and storage reserves in viable seeds. In conclusion, the results demonstrate that NIR spectroscopy has great potential as seed sorting technology to upgrade seed lot quality that ensures precision sowing.
Seed orchards are the link between tree breeding and reforestation. This paper presents data on cone, seed and pollen production and seed quality gathered over 21 years in a Pinus sylvestris (L.) experimental seed orchard containing plots with 14 different combinations of stand density and targeted pruning height. The treatments’ stand densities ranged from 267 to 4000 stems ha-1, and the target graft heights ranged from 2 to 6 meters. Pollen production began at the same orchard age for all studied combinations of stand density and target height but the level of pollen production per hectare increased more rapidly in treatments with higher stand densities. In treatments with dense spacing, cone and seed production initially increased more rapidly than in treatments with wider spacing, thereby providing an earlier return on investment and a shorter seed production lag time. However, the levels of cone and seed production in such treatments over the entire study period were not appreciably different to those achieved in treatments with wider spacing and higher target height. The treatments did not differ substantially with respect to seed quality. These results show that comparable levels of seed production can be obtained with different combinations of stand density and target height, giving seed orchard owners and managers a wide range of viable management options.
We studied the effect of soak-sorting Norway spruce (Picea abies (L.) H. Karst.) seeds on emergence, development and quality of container seedlings in two commercial seed lots. The seeds, separated by soaking into bottom and surface fractions, were sown in June, and the seedlings were grown during two growing seasons under typical Finnish nursery conditions. The first summer seedlings were grown in a greenhouse and outdoors for the second, full growing season. All sunken seeds were full and viable according to radiography, whereas the floating seeds contained 2% and 13% larvae-filled and 8% and 11% anatomically immature seeds, depending on the seed lot. Seedlings grown from the bottom fraction seed emerged 2.5–3.5 days earlier than seedlings of storage dry (i.e. control) seed. Height, diameter, and shoot and root dry mass of the seedlings were affected by soaking after both the first and second growing seasons. The largest seedlings originated from the bottom fraction. The proportion of saleable seedlings was four percentage points higher in the bottom fraction than in the other seedlings. The effects of soaking found in this study are more notable than as previously reported for Norway spruce seedlings. This suggests that soaking and soak-sorting may be most useful when the growing conditions are stressful, i.e. when seeds are sown in summer rather than 1-year-old seedling crops sown in spring under the climate conditions typical of Finland.