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Silva Fennica 1926-1997
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Articles containing the keyword 'gene'

Category: Article

article id 5605, category Article
Matti Haapanen, Marja-Leena Annala, Pirkko Velling. (1997). Progeny trial estimates of genetic parameters for growth and quality traits in Scots pine. Silva Fennica vol. 31 no. 1 article id 5605. https://doi.org/10.14214/sf.a8506
Keywords: Pinus sylvestris; Scots pine; wood quality; genetic correlation; heritability; progeny testing
Abstract | View details | Full text in PDF | Author Info

Estimates of individual heritability and genetic correlation are presented for a set of 10 growth and quality traits based on data from 16 Scots pine (Pinus sylvestris L.) progeny trials in Finland. Seven of the traits (tree height, stem diameter, crown width, Pilodyn value, branch diameter, branch angle and branch number) were objectively measured, whereas three traits (stem straightness, branching score and overall score) were assessed visually. The genetic correlations were mostly moderate or low, and favourable from the tree breeder's point of view. All variables related to tree size correlated relatively strongly and positively. Tree height exhibited a more favourable genetic relationship with the crown form traits than diameter, the latter showing positive correlation with branch diameter. Except for the slight negative correlation between branch angle and branch diameter, the branching traits were not notably correlated. The pilodyn value was positively correlated with stem diameter, reflecting negative correlation between diameter growth and wood density. The highest genetic correlations occurred among the two visually evaluated quality scores and branch diameter. All of the heritabilities were less than 0.4. Overall score, Pilodyn, branch angle, branching score and tree height showed the highest heritability.

  • Haapanen, E-mail: mh@mm.unknown (email)
  • Annala, E-mail: ma@mm.unknown
  • Velling, E-mail: pv@mm.unknown
article id 5573, category Article
Ilkka Vanha-Majamaa, Raili Suominen, Tiina Tonteri, Eeva-Stiina Tuittila. (1996). Seedling establishment after prescribed burning of a clear-cut and a partially cut mesic boreal forest in southern Finland. Silva Fennica vol. 30 no. 1 article id 5573. https://doi.org/10.14214/sf.a9218
Keywords: Pinus sylvestris; Betula pendula; Picea abies; Betula pubescens; natural regeneration; seedling establishment; prescribed burning; controlled burning; Sorbus aucuparia; seed dispersal; mesic forest; seed rain; autoregression model; GLM
Abstract | View details | Full text in PDF | Author Info

The prescribed burning of a 7.3 ha clear-cut and a 1.7 ha partially cut forest (volume 150 m3/ha) was carried out in Evo (61 °12'N, 25°07'E) on 1 June 1992. The forest was a mesic Myrtillus site type forest dominated by Norway spruce (Picea abies (L.) H. Karst.). Practically all the trees and the above-ground parts of the understorey vegetation died in the fire, while the mor layer was thinned by an average of 1.5 cm.

A study was made on the change of germinated seedling population in time and their dependence on environmental factors. Seedlings of Norway spruce, Scots pine (Pinus sylvestris L.), silver birch (Betula pendula Roth), pubescent birch (B. pubescens Ehrh.) and rowan (Sorbus aucuparia L.) were inventoried in 1993 and in 1994 on permanent plots, four times per growing season. Autoregression models were used to compare regeneration of tree species in the burned forest with regeneration in the burnt clear-cut area, and to study the effect of distance from nearest seed source to regeneration.

The average number of seedlings germinating in 1993 was higher than in 1994, probably because of differences between these consecutive years in regard to the amount of seed rain and weather conditions. The number of Norway spruce and rowan seedling was higher inside the forest area than in the clear-cut area. The distance to the bordering forest and to the closest seed tree did not explain the result. It is suggested that the more stable microclimatic conditions under the shade of dead tree promote germination and seedling establishment in the forest area. As rowan is a bird-dispersed species, it is likely that dead trees help the dispersal of rowan seed by providing birds place to sit and defecate. The shade provided by dead trees may influence the further succession of the tree stand and vegetation composition and diversity.

  • Vanha-Majamaa, E-mail: iv@mm.unknown (email)
  • Suominen, E-mail: rs@mm.unknown
  • Tonteri, E-mail: tt@mm.unknown
  • Tuittila, E-mail: et@mm.unknown
article id 5571, category Article
Katarina Lindgren, Dag Lindgren. (1996). Germinability of Norway spruce and Scots pine pollen exposed to open air. Silva Fennica vol. 30 no. 1 article id 5571. https://doi.org/10.14214/sf.a9216
Keywords: Pinus sylvestris; Picea abies; gene flow; pollen; pollen viability; conifer pollen
Abstract | View details | Full text in PDF | Author Info

Germination of Norway spruce (Picea abies (L.) H. Karst.) and Scots pine (Pinus sylvestris L.) pollen decreased during exposure to open air conditions. Usually more than half of the pollen remained germinative after a few days outdoors, but following more than four days outdoors the germination became very low. This study supports the opinion that pollen in the atmosphere remains viable long enough to allow for long-distance gene flow by pollen migration, as an important factor in genetic management of conifers and in evolution, maintaining diversity and potential for adaptation.

  • Lindgren, E-mail: kl@mm.unknown (email)
  • Lindgren, E-mail: dl@mm.unknown
article id 5562, category Article
Janne Uuttera, Matti Maltamo. (1995). Impact of regeneration method on stand structure prior to first thinning. Comparative study North Karelia, Finland vs. Republic of Karelia, Russian Federation. Silva Fennica vol. 29 no. 4 article id 5562. https://doi.org/10.14214/sf.a9213
Keywords: natural regeneration; stand structure; artificial regeneration; potential biodiversity; regeneration strategy
Abstract | View details | Full text in PDF | Author Info

Comparisons were made between artificially and naturally regenerated stands in the south-eastern part of North Karelia, Finland, and naturally regenerated stands in the western parts of the Republic of Karelia, Russian Federation. The effect of soil fertility and silvicultural operations on the stand structure was also investigated.

The results of the study show clearly that when forests are artificially regenerated the stand structure includes less variation when compared with the stands naturally regenerated. Differences between the regeneration methods are clearer the more fertile the forest site is. Within the regeneration method there is also a clear trend in stand structure, with the variation decreasing the poorer the site. The effect of silvicultural operations, i.e. the cleaning of the sapling stand, has disappeared by the time of first thinning, although it appears to have a permanent effect on the dynamics of the tree species within a stand.

The variation of the stand structure can be regarded as an essential factor for the potential biodiversity of the stand also at its young vegetation succession stage. This capacity for maintaining the forest biodiversity, developed at the young vegetation succession stage, becomes increasingly important in subsequent vegetation succession stages. Natural regeneration provides improved possibilities for the operations preserving forest biodiversity, as it generates more dense stands with a wider variation in stand structure, compared to artificial regeneration.

  • Uuttera, E-mail: ju@mm.unknown (email)
  • Maltamo, E-mail: mm@mm.unknown
article id 5557, category Article
Klaus Silfverberg. (1995). Forest regeneration on nutrient-poor peatlands: Effects of fertilization, mounding and sowing. Silva Fennica vol. 29 no. 3 article id 5557. https://doi.org/10.14214/sf.a9208
Keywords: Pinus sylvestris; Betula; soil preparation; natural regeneration; drained peatlands; mounding; ground vegetation; sowing; seedling number
Abstract | View details | Full text in PDF | Author Info

The effects of wood ash and PK fertilization on natural regeneration and sowing of Scots pine (Pinus sylvestris L.) were studied in field experiments on nitrogen-poor (Ntot 0.87–1.26%) peat substrates. The study material was derived from three drained, nutrient-poor pine mires (64°52’ N, 25°08’ E) at Muhos, near Oulu, Finland. The experimental fields were laid out in 1985 as a split-split-plot design including the following treatments; mounding, natural regeneration and sowing and fertilization; PK (400 kg ha-1) and wood ash (5,000 kg ha-1). The seedlings were inventoried in circles in July–August 1991.

Changes in the vegetation were small and there were no statistical differences due to the fertilization treatments in the ground vegetation. PK or ash fertilization did not cause vegetation changes harmful to Scots pine regeneration on nitrogen-poor peatlands. Both sowing and fertilization significantly increased the number of pine seedlings, but not their height. Wood ash increased seedling number more than PK fertilizer. The number of seedlings varied from 7,963 (control) to 42,781 ha-1 (mounding + sowing + ash). The seedling number was adequate for successful regeneration even on non-mounded, non-fertilized naturally regenerated plots.

The number of birch seedlings varied more than that of pine (370–25,927 ha-1). Mounding especially increased the number of birches. The difference between PK fertiliser and ash was less pronounced than that for pine. In addition, to the field studies the effects of ash and PK fertilizer on the germination of Scots pine seeds was studied in a greenhouse experiment. Soaking in ash solutions strongly reduced seed germination, while the PK solution was less harmful.

  • Silfverberg, E-mail: ks@mm.unknown (email)
article id 5552, category Article
Liisa Saarenmaa, Timo Leppälä. (1995). Fill-in seedlings in constituting the stocking of Scots pine stands in northern Finland. Silva Fennica vol. 29 no. 2 article id 5552. https://doi.org/10.14214/sf.a9203
Keywords: Pinus sylvestris; planting; spatial pattern; artificial regeneration; fill planting; survival of seedlings
Abstract | View details | Full text in PDF | Author Info

Fill planting is a common procedure following reforestation in Finland. In 1990, 13% of the total of seedlings planted was used for fill planting. The objective of this study is (i) to survey the survival of fill-in seedlings and (ii) to estimate the spatial pattern of stands to evaluate the importance of fill-in seedlings in constituting the stocking of Scots pine (Pinus sylvestris L.) stands in Central and Northern Finland.

A survey of 63 artificially regenerated Scots pine stands was conducted in 1990. Stand densities varied from 950 to 3,925 seedlings/ha. The mean densities of originally planted, fill planted and naturally regenerated seedlings were 863, 639 and 791/ha, respectively. The survival of originally planted seedlings was 36% and that of fill-in seedlings 48%. Death rate of fill-in seedlings of Scots pine increased with longer times between original and fill planting. The survival rate of Norway spruce (Picea abies) seedlings was correlated with temperature sum. Height of the fill-in seedlings was less than that of the originally planted ones. Most stands had an even spatial distribution with the exception of sparsely populated stands, which were somewhat clustered. This indicates that dying of seedlings is not randomly spread. Because of poor survival, fill planting seems to be a risky business in most cases.

  • Saarenmaa, E-mail: ls@mm.unknown (email)
  • Leppälä, E-mail: tl@mm.unknown
article id 5546, category Article
Oliver Schabenberger, Timothy G. Gregoire. (1995). A conspectus on Estimating Function theory and its applicability to recurrent modeling issues in forest biometry. Silva Fennica vol. 29 no. 1 article id 5546. https://doi.org/10.14214/sf.a9197
Keywords: modelling; mixed models; statistical methods; longitudinal data; generalized linear models; optimality
Abstract | View details | Full text in PDF | Author Info

Much of forestry data is characterized by a longitudinal or repeated measures structure where multiple observations taken on some units of interest are correlated. Such dependencies are often ignored in favour of an apparently simpler analysis at the cost of invalid inferences. The last decade has brought to light many new statistical techniques that enable one to successfully deal with dependent observations. Although apparently distinct at first, the theory of Estimating Functions provides a natural extension of classical estimation that encompasses many of these new approaches. This contribution introduces Estimating Function Theory as a principle with potential for unification and presents examples covering a variety of modelling issues to demonstrate its applicability.

  • Schabenberger, E-mail: os@mm.unknown (email)
  • Gregoire, E-mail: tg@mm.unknown
article id 5539, category Article
Tore Skrøppa. (1994). Impacts of tree improvement on genetic structure and diversity of planted forests. Silva Fennica vol. 28 no. 4 article id 5539. https://doi.org/10.14214/sf.a9179
Keywords: diversity; tree breeding; seed orchards; production; adaptability; genetic factors; trees
Abstract | View details | Full text in PDF | Author Info

After a presentation of basic biodiversity concepts, reviews are made of studies reporting genetic implications of tree improvement activities: seed treatments, seedling production, provenance transfers, plus tree selection, seed production in seed orchards and progeny testing.

Several of the activities may influence the genetic structure and diversity of the planted forests. The general conclusion is, however, that planted forests are at least as genetically diverse as the natural stands that they replace. The diversity in forest management and use is best assurance for the future adaptability of the forests.

  • Skrøppa, E-mail: ts@mm.unknown (email)
article id 5537, category Article
Anu Mattila, Anne Pakkanen, Juha Raisio, Pekka Vakkari. (1994). Genetic variation in English oak (Quercus robur) in Finland. Silva Fennica vol. 28 no. 4 article id 5537. https://doi.org/10.14214/sf.a9177
Keywords: Quercus robur; allozymes; populations; genetic differences; heterozygosity
Abstract | View details | Full text in PDF | Author Info

Genetic variation in 5 natural stands of Quercus robur L. in Finland was analysed electrophoretically for 13 isozyme loci. Stands were on average polymorphic at 49.2% of the loci, with 2.1 alleles per locus. Observed heterozygosities, ranging from 13.6% to 16.9%, were slightly lower than estimates reported for German stands. The majority of the species’ genetic variation was found within each studied stand, and only 5.5% was between stands. Mean genetic differentiation (∂) was the same as that found in the primary range of the species, but the differentiation estimates (D) for single Finnish population were more variable.

  • Mattila, E-mail: am@mm.unknown (email)
  • Pakkanen, E-mail: ap@mm.unknown
  • Raisio, E-mail: jr@mm.unknown
  • Vakkari, E-mail: pv@mm.unknown
article id 5534, category Article
Hyun Kang, Inger Ekberg, Gösta Eriksson, Johan Ununger. (1994). Second and third growth period responses of Picea abies families to first growth period photoperiodic, light intensity and temperature treatments. Silva Fennica vol. 28 no. 4 article id 5534. https://doi.org/10.14214/sf.a9174
Keywords: Picea abies; growth; maturation; temperature; seedlings; photoperiod; environmental effects; genetic effects; juvenility
Abstract | View details | Full text in PDF | Author Info

Seedlings of Picea abies (L.) H. Karst. full-sib families of contrasting origins were cultivated in a phytotron under different photoperiodic, light-intensity and temperature treatments during their first growth period. The effects of the treatments on juvenile growth traits – whether enhanced or delayed maturation was induces – were observed during the two subsequent growth periods. The following hypotheses were tested: (A) Enhanced maturation can be induced in the first growth period from sowing with (i) a long period of continuous light during active growth (24 weeks vs. 8 weeks); (ii) a shorter night during bud maturation (12 h vs. 16 h); high temperature (25°C vs. 20°C) during (iii) active growth, growth cessation and bud maturation; and during (iv) the latter part of growth cessation and bud maturation only. (B) Delayed maturation can be induced after (i) low light intensity during growth cessation and bud maturation (114 μmol m-2 s-1 vs. 340 μmol m-2 s-1); low temperature (15°C vs. 20°C) during (ii) active growth, growth cessation and bud maturation; and during (iii) the latter part of growth cessation and bud maturation only.

The most dramatic effect was observed after 24 weeks of continuous light during active growth. All traits showed a significantly more mature performance in the second growth period compared with the control. The effect for all but one trait was carried over to the third growth period. This is in accordance with the hypothesis that the activity of apical shoot meristems controls the maturation process. For the other treatments there was only weak or no support for the hypothesis of induction of enhanced or delayed maturation. Strong family effects were observed for all traits. Differential responses of the various latitudinal families were observed, suggesting that family effects must be considered to predict and interpret correctly how plants will respond to environmental effects.

  • Kang, E-mail: hk@mm.unknown (email)
  • Ekberg, E-mail: ie@mm.unknown
  • Eriksson, E-mail: ge@mm.unknown
  • Ununger, E-mail: ju@mm.unknown
article id 5400, category Article
Kari J. Jokinen, Don J. Durzan. (1994). Properties of rescued embryonal suspensor masses of Norway spruce determined by the genotype and the environment in vitro. Silva Fennica vol. 28 no. 2 article id 5400. https://doi.org/10.14214/sf.a9165
Keywords: Norway spruce; Picea abies; growth; somatic embryogenesis; callus
Abstract | View details | Full text in PDF | Author Info

53 genotypes of embryonal suspensor masses (ESMs) rescued from mature seeds of Norway spruce (Picea abies (L.) H. Karst.) were examined for their pattern of growth and development under standardized culture conditions in vitro. Patterns were classified according to the colour of the colonies grown in darkness, clarity of cell masses and proembryos in the mucilaginous ESM, surface boundary topology of colonies, structure of suspensors, growth rate of the ESM, and recovery of mature embryos.

Five distinctive major growth patterns were observed among ESM colonies under standardized culture conditions. The multiplication of proembryos and early embryos by cleavage and budding polyembryony was the main factor contributing to proliferation and colony growth and further determined the morphology of the colonies. Callus and teratological structures were induced from early embryos by changing the standardized culture conditions i.e. inadequate subculture, excessive dose of 2,4-D in the medium and premature exposure of the colonies to light. Results enable the selection of ESM genotypes for the predictable recovery of mature somatic embryos.

  • Jokinen, E-mail: kj@mm.unknown (email)
  • Durzan, E-mail: dd@mm.unknown
article id 5466, category Article
Hely Häggman. (1991). Application of biotechnology to forest tree breeding. Silva Fennica vol. 25 no. 4 article id 5466. https://doi.org/10.14214/sf.a15624
Keywords: tree breeding; micropropagation; vegetative propagation; somatic embryogenesis; conventional breeding; somaclonal variation; biotechnology; gene transfer
Abstract | View details | Full text in PDF | Author Info

Forest tree breeding involves manipulation of genetic composition of populations and individuals, and biotechnology focuses on selected individuals. The new techniques cannot replace the conventional breeding techniques but both need effective cooperation of each other. Thus, the distinction between conventional breeding and biotechnology is artificial. The biotechnology methods are new and fast developing and the future with field and progeny testing will show which techniques will be permanently adopted into tree breeding. For instance, the earlier hope of the use of somaclonal variation as a new source of variability and a powerful tool for the breeder seem today quite the opposite. Somaclonal variation constituting a major problem in present-day micropropagation is due to the unpredictable variation. Based on knowledge of today, especially micropropagation via somatic embryos, transgenic trees and the identification of major genes seem to be good candidates to be permanently adopted into tree breeding.

  • Häggman, E-mail: hh@mm.unknown (email)
article id 5460, category Article
R. C. Schmidtling. (1991). Effect of rootstock and scion on flowering, growth and foliar nutrients of loblolly pine (Pinus taeda) grafts. Silva Fennica vol. 25 no. 4 article id 5460. https://doi.org/10.14214/sf.a15618
Keywords: seed orchards; Pinus taeda; grafting; incompatibility; genetics
Abstract | View details | Full text in PDF | Author Info

In two separate studies, seedlings from 20 loblolly pine (Pinus taeda L.) families and Virginian pine (Pinus virginiana Mill.) were used as root stocks for grafting loblolly pine seed orchard clones. The rootstocks were open-pollinated seedlings from orchard clones chosen to represent a wide range of flowering and survival capabilities, based on their performance in a first-generation seed orchard. Scions were derived from the same 20 loblolly clones. The effects of scion clone were significant and large for nearly all measured traits. Rootstock significantly affected survival, growth, flowering and foliar nutrients of the grafted ramets. Neither survival nor growth of the crafts was related to survival of growth of the orchard clones from which their rootstocks were derived, however. Survival of incompatible clones was enhanced by grafting on genetically related rootstocks.

The PDF includes an abstract in Finnish.

  • Schmidtling, E-mail: rs@mm.unknown (email)
article id 5458, category Article
Jaroslaw Burczyk, Grzegorz Kosiński, Andrzej Lewandowski. (1991).  . Silva Fennica vol. 25 no. 4 article id 5458. https://doi.org/10.14214/sf.a15616
English title: Mating pattern and empty seed formation in relation to crown level of Larix decidua (Mill.) clones.
Keywords: seeds; crown; Larix decidua; mating systems; genetic load; grafting
Abstract | View details | Full text in PDF | Author Info

The mating system was analysed in the upper and lower crown of two groups of European larch (Larix decidua Mill.) clones divided according to the percentage of full seeds in the upper and lower crown parts. The overall multilocus estimate of outcrossing rate (t) was calculated to be 0.929. The differences of outcrossing rates between crown levels and clonal groups respectively were not statistically significant. The t estimates were greater for the upper crown level and for clones with higher percentage of full seeds in the upper crown level. However, among all observations there was no correlation between outcrossing rates and percentages of full seeds for particular crown levels and groups of clones. Observed similarity of outcrossing among grafts of the same clones may indicate genetic control of self-fertilization rate in individual European larch. 

The PDF includes an abstract in Finnish.

  • Burczyk, E-mail: jb@mm.unknown (email)
  • Kosiński, E-mail: gk@mm.unknown
  • Lewandowski, E-mail: al@mm.unknown
article id 5445, category Article
Taneli Kolström. (1991). Kuusen kylvö- ja istutuskoe viljavilla kivennäismailla Pohjois-Karjalassa. Silva Fennica vol. 25 no. 2 article id 5445. https://doi.org/10.14214/sf.a15599
English title: Results from the sowing and planting experiment of Norway spruce (Picea abies) on fertile sites in North Karelia, Finland.
Original keywords: kuusi; kylvö; istutus; metsänuudistaminen; heinäntorjunta; kuolleisuus; pituuskasvu
English keywords: Picea abies; height growth; planting; mortality; sowing; artificial regeneration; survival rate; control of ground vegetation
Abstract | View details | Full text in PDF | Author Info

Different methods of sowing and planting of Norway spruce (Picea abies (L.) H. Karst.) were compared on fertile sites in North Karelia (62°20’N, 29°35’E, 85–120 m a.s.l.). The planting material were 4-year-old bare-rooted transplants, 2-year-old bare-rooted seedlings, and 2-year-old containerized seedlings raised in plastic greenhouse. The sowing methods were band sowing and shelter sowing. Ground vegetation was controlled during the first growing season mechanically or chemically, or the control was omitted totally.

Planting of spruce gave better results than sowing. After eight growing seasons there were sowed seedlings left in 30% of the sowing pots. The average height of them was 35 cm. Seedling survival was best with large bare-rooted transplants (91%). Survival of containerized seedlings was 79% and of small bare-rooted transplants 71%. The average height of large bare-rooted transplants was 131 cm, of containerized seedlings 86 cm and small bare-rooted seedlings 68 cm.

Sowing is not an advisable method for regeneration of spruce due to the small survival rate and slow initial development when ground vegetation is controlled only once. Also 2-year-old seedlings gave a satisfactory result in regeneration. Seedlings raised in greenhouse were more sensitive to frost damage than seedlings grown on open ground.

The PDF includes an abstract in English.

  • Kolström, E-mail: tk@mm.unknown (email)
article id 5420, category Article
Toyohiro Miyazava, Jukka Laine. (1990). Effect of macroclimate on the development of Scots pine seedling stands on drained oligotrophic pine mires. Silva Fennica vol. 24 no. 2 article id 5420. https://doi.org/10.14214/sf.a15574
Keywords: Pinus sylvestris; fertilization; regeneration; drained peatlands; plantations; ditch spacing; effective temperature sum; microtopography
Abstract | View details | Full text in PDF | Author Info

The influence of different fertilization treatments and ditch spacings on the height growth of young Scots pine (Pinus sylvestris L.) seedling stands growing under various climatic regimes were determined. Comparisons were made between naturally regenerated and planted seedling stands. The effective temperature sum had a stronger effect on the height growth of planted seedlings, and in Northern Finland the planted seedlings seemed to be influenced to a greater degree by the adverse climatic conditions. The heavier the dose of fertilizer that had been applied, the greater the difference in growth caused by macroclimate. A considerably larger proportion of natural seedlings were located on hummocks compared with that of planted seedlings, irrespective of the region. On plots with wider ditch spacings, seedlings growing on hummocks were superior in height growth to those on flat surfaces.

The PDF includes an abstract in Finnish.

  • Miyazava, E-mail: tm@mm.unknown (email)
  • Laine, E-mail: jl@mm.unknown
article id 5415, category Article
Kim von Weissenberg. (1990). Host-parasite relationships in forest ecosystems: A review. Silva Fennica vol. 24 no. 1 article id 5415. https://doi.org/10.14214/sf.a15567
Keywords: Pinus; resistance; Ophiostoma; rust diseases; Cronartium; Cryphonectria; Melampsora; polygenic; oligogenic; gene-for-gene
Abstract | View details | Full text in PDF | Author Info

Based on a survey of world literature it is concluded that 1) the better -researched epidemic forest pathosystems are caused by anthropogenic factors, 2) the systems most likely have a polygenic background, and 3) resistance breeding should maintain polygenic resistance with restrictive incorporation of oligogenic resistance. Corresponding objectives are valid in breeding programs of presently balanced pathosystems, which may turn epidemic if man causes changes in the gene pool and alters critical environmental conditions.

The PDF includes an abstract in English.

  • Weissenberg, E-mail: kw@mm.unknown (email)
article id 5413, category Article
Eljas Pohtila. (1990). Metsän uudistuminen Kivalon vanhoilla kaistalehakkuualoilla. Silva Fennica vol. 24 no. 1 article id 5413. https://doi.org/10.14214/sf.a15565
English title: Forest regeneration of old strip cutting areas in Kivalo.
Original keywords: kuusi; metsänhoito; luontainen uudistaminen; Lappi; kaistalehakkuu
English keywords: Picea abies; natural regeneration; northern Finland; strip cutting; spruce stands
Abstract | View details | Full text in PDF | Author Info

The results of regenerating 54 and 36–38-year old strip cuttings were surveyed in the Kivalo Research Forest (N 66°20’, E 26°40’). 15 measurement plots were placed on each strip. The most common forest type was Hylocomnium-Myrtillus type. Regeneration of the strips proved to be slow. Most of the spruces growing on the strips probably originated from the time before the cutting. The average number of stems was 1,155 per ha, of which one third consisted of broadleaved trees. The average volume increment of stem wood after cutting had been about 1 m3/ha/yr, but it was increasing at the time of the inventory. Both the reforestation of the strips and the development of the emergent stands were dependent on elevation and site fertility. Site fertility was indicated by the abundance of Vaccinium myrtillus.

The PDF includes an abstract in English.

  • Pohtila, E-mail: ep@mm.unknown (email)
article id 5412, category Article
Jari Parviainen. (1990). Metsäpuiden paakkutaimituotannon nykynäkymät. Silva Fennica vol. 24 no. 1 article id 5412. https://doi.org/10.14214/sf.a15564
English title: Future trends for containerized tree seedling production: A literature review.
Original keywords: metsänuudistaminen; juuristo; kustannukset; paakkutaimituotanto; taimilaatu; epämuodostumat
English keywords: regeneration; containerized seedlings; costs; spacing; quality; tree nurseries; root systems; deformations
Abstract | View details | Full text in PDF | Author Info

Containerized tree seedlings will be used on an increasing scale in the future in different parts of the world. There are number of techniques for the production of small one-year-old seedlings but it has not been possible to develop a completely satisfactory methods for large containerized seedlings production. In the long-term development of pine plantations established with containerized seedlings the greatest problem has been deformation of the root system. With a new method, based on a sheet of peat and root pruning, it has been possible to produce conifer seedlings with a good root regeneration potential and favourable morphological root system development. The use of small containerized seedlings allows an increase in planting density without any marked increase in regeneration costs.

The PDF includes an abstract in English.

  • Parviainen, E-mail: jp@mm.unknown (email)
article id 5379, category Article
Timo Kuuluvainen, Timo Pukkala. (1989). Effect of Scots pine seed trees on the density of ground vegetation and tree seedlings. Silva Fennica vol. 23 no. 2 article id 5379. https://doi.org/10.14214/sf.a15536
Keywords: Pinus sylvestris; natural regeneration; spatial variation; ground vegetation; seedlings; seed trees; regeneration models; ecological fields; resource consumption; competitive interface
Abstract | View details | Full text in PDF | Author Info

The study uses the methodology of ecological field theory to model the effect of Scots pine (Pinus sylvestris L.) seed trees on the density of tree seedlings and other plants in the field layer. The seed trees had a clear effect on the expected value of the amount and distribution of the ground vegetation. The vicinity of seed trees had an adverse effect on the growth of grasses, herbs and seedlings, while mosses were most abundant near the trees. Models based on the ecological field approach were derived to describe the effect of seed trees on the ground vegetation.

The PDF includes an abstract in Finnish.

  • Kuuluvainen, E-mail: tk@mm.unknown (email)
  • Pukkala, E-mail: tp@mm.unknown
article id 5365, category Article
Pasi Puttonen. (1988). Metsänuudistamistutkimuksen arvioinnin seminaari 28.4.1988. Silva Fennica vol. 22 no. 4 article id 5365. https://doi.org/10.14214/sf.a15522
English title: Presentation report of evaluation group on forest regeneration research in Finland.
Original keywords: metsäntutkimus; metsänuudistaminen; Suomen Akatemia; arviointi
English keywords: evaluation; forest research; forest regeneration research; Academy of Finland
Abstract | View details | Full text in PDF | Author Info

The Academy of Finland commissioned a review of forest regeneration research. This paper summarises the evaluation. The report deals with the individual research projects and, in addition, the international evaluation group gives several recommendations to develop the forest regeneration research further. The recommendations concern education and training, publications, peer review, experimental design, routine work and extension service, and technology transfer. The paper also includes comments of the Finnish Forest Institute, the Finnish Society of Forest Science and a research scientist on the results of the evaluation.

The PDF includes a summary in English.

  • Puttonen, E-mail:
article id 5350, category Article
Heikki Hänninen, Paavo Pelkonen. (1988). Frost hardiness and over-wintering of forest trees. Silva Fennica vol. 22 no. 3 article id 5350. https://doi.org/10.14214/sf.a15507
Keywords: frost hardiness; breeding; frost damage; genetic variation; dormancy; annual cycle of development; nursery management
Abstract | View details | Full text in PDF | Author Info

This issue of Silva Fennica consists of eight articles, which are based on a co-nordic conference ”Frost hardiness and over-wintering in forest tree seedlings”, held in Joensuu, Finland, during December 1–3, 1986. The whole annual cycle of the trees is considered. Emphasis is given on methods for the study of frost hardiness, genetic variation in frost hardiness, nitrogen metabolism, bud dormancy release, and joint effect of natural and anthropogenic stress factors in the winter damage of forest trees. Practical implications for tree breeding and nursery management are discussed.

The PDF includes an abstract in Finnish.

  • Hänninen, E-mail:
  • Pelkonen, E-mail:
article id 5311, category Article
Timo Pukkala. (1987). Kuusen ja männyn siemensadon ennustemalli. Silva Fennica vol. 21 no. 2 article id 5311. https://doi.org/10.14214/sf.a15468
English title: Model for predicting the seed crop of Picea abies and Pinus sylvestris.
Original keywords: kuusi; mänty; luontainen uudistaminen; Lappi; siemensato; lämpötila; kukinnan aloitus; ennustemalli
English keywords: Pinus sylvestris; Norway spruce; Picea abies; natural regeneration; Scots pine; Finland; Lapland; predictive modelling; seed crop; flowering initiation; influence of temperature
Abstract | View details | Full text in PDF | Author Info

The seed crop of Norway spruce (Picea abies (L.) H. Karst.) and Scots pine (Pinus sylvestris L.) is predicted with the help of mean monthly temperatures during May–August one and two years before the flowering year. The prediction models were made separately for Lapland and for the rest of Finland. The models are based on 10-year periods of seed crop measurements and climatic data. The total number of time series was 59.

In Lapland, Norway spruce flowered abundantly and produced an abundant seed crop after warm July–August and two years after cool July–August. In other parts of Finland, warm June and July produced a good flowering year, especially if these months were cool two years before the flowering year.

In Lapland, Scots pine flowered abundantly if the whole previous growing season was warm. Elsewhere in Finland, a cool June preceded prolific flowering in the coming year if the rest of the growing season was considerably warmer than the average.

The prediction models explained 37–49 % of the variation in the size of the seed crop. The occurrence of good and poor seed years was usually predicted correctly. Using the presented models, the prediction of the seed crop is obtainable 1.5 year for Norway spruce and 2.5 year for Scots pine before the year of seed fall.

The PDF includes an abstract in English.

  • Pukkala, E-mail: tp@mm.unknown (email)
article id 5302, category Article
Seppo Kellomäki, Heikki Hänninen, Taneli Kolström, Ahti Kotisaari, Timo Pukkala. (1987). A tentative model for describing the effects of some regenerative process on the properties of natural seedling stands. Silva Fennica vol. 21 no. 1 article id 5302. https://doi.org/10.14214/sf.a15459
Keywords: Pinus sylvestris; natural regeneration; Scots pine; modelling; stand density; seed crop; seed dispersal; parent tree
Abstract | View details | Full text in PDF | Author Info

The effect of the size of seed crop, dispersal of seeds and the early development of seedlings on the density and spatial distribution of young Scots pine (Pinus sylvestris L.) stands are evaluated on the basis of theoretical models. The models include (i) number and spatial distribution of parent trees on the regeneration area, (ii) size of annual seed crop, (iii) seed dispersal from a particular parent tree, (iv) germination of the seeds (germination percentage), (v) death of ageing seedlings after the establishment process, and (vi) height growth of the seedlings.

As expected, stand density and spatial distribution varied within a large range in relation to the density of the parent trees and the distance from them. The simulations also showed that natural seedling stands can be expected to be heterogenous due to the geometry of seed dispersal, emphasizing the frequency of young and small trees. The properties of the seedling stands were, however, greatly dependent on the density of the parent trees and the length of the regeneration period.

The PDF includes an abstract in Finnish.

  • Kellomäki, E-mail: sk@mm.unknown (email)
  • Hänninen, E-mail: hh@mm.unknown
  • Kolström, E-mail: tk@mm.unknown
  • Kotisaari, E-mail: ak@mm.unknown
  • Pukkala, E-mail: tp@mm.unknown
article id 5243, category Article
Eljas Pohtila, Tapani Pohjola. (1985). Maan kunnostus männyn viljelyssä Lapissa. Silva Fennica vol. 19 no. 3 article id 5243. https://doi.org/10.14214/sf.a15422
English title: Soil preparation in reforestation of Scots pine in Lapland.
Original keywords: mänty; istutus; kulotus; maanmuokkaus; Lappi; laikutus; lautasauraus; uudistustavat; hajakylvö
English keywords: Pinus sylvestris; soil preparation; Scots pine; planting; regeneration methods; scalping; prescribed burning; disc ploughing; broadcast sowing
Abstract | View details | Full text in PDF | Author Info

The study deals with the interaction of various soil preparation and reforestation methods. The most favourable time of the year for broadcast sowing and the effect of stabilization after soil preparation on restocking were studied as special problems.

Prescribed burning, scalping and disc ploughing made a better combination with sowing than planting, and ploughing better combination with planting than sowing. The longer the period was between sowing and germination the fewer seedlings emerged. The best stocking was clearly resulted with sowing in June. Stabilization of soil after preparation had a negative effect on reforestation results.

The PDF includes a summary in English.

  • Pohtila, E-mail: ep@mm.unknown (email)
  • Pohjola, E-mail: tp@mm.unknown
article id 5227, category Article
Harri Vasander, Tapio Lindholm. (1985). Tulen voimakkuus ja maanpinnan lämpötila kulotuksen aikana. Silva Fennica vol. 19 no. 1 article id 5227. https://doi.org/10.14214/sf.a15406
English title: Fire intensities and surface temperatures during prescribed burning.
Keywords: regeneration; slash; prescribed burning; logging waste; controlled burning; surface temperatures; fire intensity
Abstract | View details | Full text in PDF | Author Info

Surface temperature during two prescribed burnings were measured in 1983 in Evo, Southern Finland. Surface temperatures in relation to the amount of slash burned, energy released during the fires, and the fire intensities were studied. The fire intensity was also measured during a third burn. The Lake Nimetön site was burned int the end of May. Due to the uneven distribution of slash, colonization by Calamagrostis arundinacea and the spring moisture, the burning was very uneven. Surface temperatures varied between 410–809°C and the intensity of fire was low (range 0–900 kW/m).

The fire intensity on the other sites burned in May was also low (880 kW/m). During the burn in August the surface temperatures varied between 701–869°C and the intensity of fire was moderate (1,170 kW/m). Slash was burned more evenly and more thoroughly due to the dryness of the site and slash and the fact that grasses and other herbs were not abundant.

The PDF includes a summary in Finnish.

  • Vasander, E-mail: hv@mm.unknown (email)
  • Lindholm, E-mail: tl@mm.unknown
article id 5218, category Article
Jouni Suoheimo. (1984). Isokorvakärsäkkään aikuisten esiintyminen ja merkitys männyn luontaiselle uudistamiselle Pohjois-Lapissa. Silva Fennica vol. 18 no. 3 article id 5218. https://doi.org/10.14214/sf.a15397
English title: The occurrence of Otiorrhynchus nodosus and its significance for the natural regeneration of Scots pine in Lapland.
Original keywords: hyönteistuhot; mänty; luontainen uudistaminen; taimituhot; Pohjois-Lappi; isokorvakärsäkäs
English keywords: Pinus sylvestris; natural regeneration; Scots pine; insect damage; seedling damages; weevils; occurrence
Abstract | View details | Full text in PDF | Author Info

The aim of the present study was to survey the occurrence of Otiorrhynchus nodosus Müller weevils and their significance for the natural regeneration of Scots pine (Pinus sylvestris L.). The study was carried out during summer 1982 at Inari in northern Lapland.

There were two sample plots, one situated in a Scots pine seed-tree area and the other, the control sample plot, in an area with a coverage of mountain birch (Betula pubescens subsp. tortuosa, now subsp. czerepanovii). A total of 177 Otiorhynchus weevils were caught. Movement of the weevils reached its climax in July. There were 86% more individuals in the seed-tree area than in the mountain birch area. No damage to the pine germlings or seedlings was not observed, although the situation could be different during the peaks of the veewil populations.

The PDF includes a summary in English.

  • Suoheimo, E-mail: js@mm.unknown (email)
article id 5198, category Article
Leo Heikurainen, Jukka Laine, Jarmo Lepola. (1983). Lannoitus- ja sarkaleveyskokeita karujen rämeiden uudistamisessa ja taimikoiden kasvatuksessa. Silva Fennica vol. 17 no. 4 article id 5198. https://doi.org/10.14214/sf.a15181
English title: Fertilization and ditch spacing experiments concerned with regeneration and growth of young Scots pine stands on nutrient poor pine bogs.
Original keywords: mänty; luontainen uudistaminen; istutus; uudistaminen; lannoitus; ojitetut suot; taimikot; pituuskasvu; sarkaleveys
English keywords: Pinus sylvestris; fertilization; regeneration; natural regeneration; drained peatlands; Scots pine; height growth; planting; ditch spacing
Abstract | View details | Full text in PDF | Author Info

The effects of variations in the intensity of drainage and NPK fertilization on the natural regeneration and planting results and the subsequent development of seedling stands under various climatic conditions on drained nutrient poor pine bogs was investigated in a 16-year-old study.

Comparison of height development of Scots pine (Pinus sylvestris L.) stands on drained peatlands to that of pine stands growing in mineral soil sites show that in Southern Finland the most efficient forest improvement measures (10 m ditch spacing and 1,000 kg/ha NPK-fertilization) resulted in growth that corresponds a to a height index of a stand in a Vaccinium type site. Less efficient treatment (30 m ditch spacing and no fertilizer) resulted in growth corresponding the development of young stand in a Calluna type site. In Northern Finland the effect of fertilization on height growth was almost negligible. This is possibly due to a decrease in the nitrogen mobilization from south to north of Finland. Thus, it seems evident that fertilization of young Scots pine stands on nutrient poor drained peatlands can be recommended only in the southern part of the country.

The effect of ditch spacing is same in the whole country. The narrower the spacing the better the height growth. In the south planted stands thrive better than naturally regenerated stands, but the situation is reversed in the north.

The PDF includes a summary in English.

  • Heikurainen, E-mail: lh@mm.unknown (email)
  • Laine, E-mail: jl@mm.unknown
  • Lepola, E-mail: jl@mm.unknown
article id 5191, category Article
Juha Lappi. (1983). Metsänuudistamisen vaatiman ajan merkitys uudistamispäätöksissä. Silva Fennica vol. 17 no. 3 article id 5191. https://doi.org/10.14214/sf.a15174
English title: Elevation of the time factor in reforestation decisions.
Original keywords: metsänuudistaminen; metsäsuunnittelu; kustannukset; uudistamisen viivästyminen; uudistumisaika; normaalimetsälaskelma; nollakorkomenetelmä; päätehakkuuikä; nykyarvomenetelmä
English keywords: forest planning; net present value; forest regeneration; cost of time delay; regeneration costs; final cutting age; regeneration method; zero percent interest rate
Abstract | View details | Full text in PDF | Author Info

Length of the regeneration period is a criterion commonly used for comparing different reforestation methods. The time factor should be evaluated using a realistic system for long-term planning. In this paper the preliminary evaluation is made by simplified calculations based on the development series. The slow regeneration method is assumed to be otherwise equal to the rapid one but it has a 5- or 10-years delay at the beginning, and the rotation is thus the final cutting age plus 5- or 10-years delay. Cost of the time delay is taken to be the difference in reforestation costs that makes the rapid and the slow methods equivalent. Calculations are made using zero costs for the slow method; but if the cost of the slow method increases, the critical cost difference decreases very slowly. The final cutting age and the regeneration method must be decided simultaneously. Therefore, the cost of the time delay is presented as a function of final cutting age. By maximizing the average annual revenue, rotation can be even increased if more rapid but more expensive regeneration method is used.

The PDF includes a summary in English.

  • Lappi, E-mail: jl@mm.unknown (email)
article id 5188, category Article
Eljas Pohtila, Tapani Pohjola. (1983). Vuosina 1970-1972 Lappiin perustetun aurattujen alueiden viljelykokeen tulokset. Silva Fennica vol. 17 no. 3 article id 5188. https://doi.org/10.14214/sf.a15171
English title: Results from the reforestation experiment on ploughed sites established in Finnish Lapland during 1970–1972.
Original keywords: kuusi; kylvö; mänty; istutus; maanmuokkaus; Lappi; uudistaminen; kuolleisuus; rauduskoivu; siperianlehtikuusi; auraus
English keywords: Pinus sylvestris; Norway spruce; Betula pendula; Picea abies; regeneration; soil preparation; Scots pine; silver birch; planting; ploughing; Lapland; mortality; Siberian larch; sowing; Larix sibirica
Abstract | View details | Full text in PDF | Author Info

The objective of the study was to compare different reforestation methods on ploughed areas in Finnish Lapland. Four species were compared: Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies (L.) H. Karst.), silver birch (Betula pendula Roth) and Siberian larch (Larix sibirica Ledeb.). The experiments were established in different parts of Lapland on different types of sites in 1970–72.

In Scots pine there was a difference of 15 percentage points in survival of seedlings between the best and worst methods of regeneration. Containerized seedlings and paper pot seedlings had the best survival rates. In Norway spruce the respective difference between sowing and planting was about 20 percentage points. In favour of planting. The survival rate can be increased by about 20 percentage points by selecting the right tree species. The average height varied from 25 cm (the sowed Norway spruce) to 179 cm (the planted silver birch) after 10 growing seasons. The birch was planted at the most fertile sites only. The longer time passed from the afforestation the clearer was the effect of the local growing conditions on the development of the seedlings. The elevation of the site was one factor seemed to influence the success of the seedlings.

The PDF includes a summary in English.

  • Pohtila, E-mail: ep@mm.unknown (email)
  • Pohjola, E-mail: tp@mm.unknown
article id 5176, category Article
J. Lappi, A. Kotisaari, H. Smolander. (1983). Height relascope for regeneration surveys. Silva Fennica vol. 17 no. 1 article id 5176. https://doi.org/10.14214/sf.a15091
Keywords: sampling; forest mensuration; seedling stands; measuring instruments; regeneratoin surveys; seedling height; relascope
Abstract | View details | Full text in PDF | Author Info

The choice of sampling method is of prime importance when seeking relevant information about the height distribution and spatial arrangement of seedlings in the regeneration surveys. It is suggested that the size of sampling plots should depend on the height of the seedlings. Tall seedlings should be sampled from a larger area than short ones since tall seedlings are more important for the future development of the stand. We suggest principles for a technical development task to construct a device which is easy to use in practical regeneration surveys and by which sampling can be made proportional to plant height or any desired function of height.

The PDF includes a summary in Finnish.

  • Lappi, E-mail: jl@mm.unknown (email)
  • Kotisaari, E-mail: ak@mm.unknown
  • Smolander, E-mail: hs@mm.unknown
article id 5160, category Article
P. M. A. Tigerstedt. (1982). Metsäpuiden populaatiogenetiikka. Helsingissä 1981 pidetyn symposion tutkimusraportit. Silva Fennica vol. 16 no. 2 article id 5160. https://doi.org/10.14214/sf.a15075
English title: Population genetics of forest trees.
Original keywords: metsäntutkimus; metsänjalostus; kokoukset; siemenviljelmät; metsägenetiikka; populaatiogenetiikka; populaatiorakenne; sopeutuminen ympäristöolosuhteisiin; ekologinen erilaistuminen
English keywords: adaptation; seed orchards; population structure; population genetics; forest research; forest tree breeding; forest genetics; conference; ecological variation
Abstract | View details | Full text in PDF | Author Info

The publication comprises proceedings of a conference held in Helsinki in 1981. Forest tree populations are investigated for population genetic structure, mating systems, mechanisms of genetic adaptation and ecological adaptation. Methods and techniques used in population genetic research of forest trees are presented. Much concern is given to applications by means of forest tree breeding, particularly the seed orchard breeding technique. Generally, the application of population genetics in cultivated forests is discussed.

The PDF includes a preface and the presentations of the conference (25 short papers) in English, and a comprehensive summary of the themes of the conference in Finnish.

  • Tigerstedt, E-mail:
article id 5127, category Article
Simo Kaila, Juhani Päivänen. (1981). Metsämaanmuokkauksen suoritemäärät ja konekalusto vuosina 1976-1979. Silva Fennica vol. 15 no. 3 article id 5127. https://doi.org/10.14214/sf.a15068
English title: Forest soil cultivation areas and machinery in 1976–1979 in Finland.
Original keywords: maanmuokkaus; uudistaminen; metsäkoneet; lautasauraus; pyöräkoneet; telakoneet; auraus
English keywords: regeneration; soil preparation; disc plow; plowing; ploughing equipment; wheel tractors; crawler tractors
Abstract | View details | Full text in PDF | Author Info

The areas of soil cultivation in regeneration sites using different methods in 1976–1979 and the cultivation units in use in Finland in 1979 were studied by means of a questionnaire. Application of soil cultivation methods appears to have become strongly polarized: in Southern Finland disc plowing with wheel tractors are used, whilst in Northern Finland wing plowing with crawler tractors is the most popular method, each being practically the only alternative in the respective area.

The PDF includes a summary in English.

  • Kaila, E-mail: sk@mm.unknown (email)
  • Päivänen, E-mail: jp@mm.unknown
article id 5068, category Article
S. A. Petrov. (1980). Quantitative analysis of the effect of genotype and environment in forest tree populations. Silva Fennica vol. 14 no. 1 article id 5068. https://doi.org/10.14214/sf.a15009
Keywords: heritability; population structure; population genetics; genotype; Symposiums; intrapopulation variability; environment; quantitative characteristics
Abstract | View details | Full text in PDF | Author Info

The paper discusses the theoretical basis of quantitative analysis of the effect of genotype and environment in forest trees. Perhaps the main problem in the understanding of the laws of intrapopulation variability of the species of woody forest plants is the study of the structure of their populations. It may be characterized by a number of parameters. The intrapopulation variability of quantitative characteristics appears as a result of environmental and genetic factors, but to determine the relative weight of these factors in a concrete case is not easy. The study of the structure of a population by its quantitative characteristics has a wider task: to establish the relevance of the hereditary differences of the individuals of a population. Also, the differences caused by diverse growth conditions and how they are reflected in the level of general phenotypic variability of the quantitative characteristics in a given population has to be identified. The author gives examples of assessment of heritability in forest trees.

The PDF includes a summary in Finnish.

  • Petrov, E-mail: sp@mm.unknown (email)
article id 5058, category Article
Max Hagman. (1980). Suomalais-neuvostoliittolainen metsägenetiikan ja siemenhuollon symposiumi Punkaharjulla 17.–18.8.1978. Silva Fennica vol. 14 no. 1 article id 5058. https://doi.org/10.14214/sf.a14999
English title: Reports of the Finnish-Soviet symposium on forest genetics and forest tree seed production, Punkaharju, Finland, August 17-18. 1978.
Original keywords: metsänjalostus; Suomi; kokoukset; siemenviljely; Neuvostoliitto; seminaarit; genetiikka
English keywords: Finland; seed orchards; forest tree breeding; Soviet Union; USSR; symposium; forest genetics
Abstract | View details | Full text in PDF | Author Info

As a part of the scientific and technical cooperation between Finland and the USSR a symposium and an excursion on forest genetics and seed production was organized in Finland in August 1978. The symposium paper presented at Punkaharju are published here in order to bring them available for a wider audience.

The 12 symposium reports deal with the following subjects: Practical application of forest genetic research (A.I. Novoselceva), geographical variability and provenance transfer (E.P. Prokazin, M. Hagman, I. Etverk), variation in wood density (P. Velling), variation of flowering and seed crops in seed orchards (Y.P. Efimov), and natural stands (V. Koski), vegetative propagation (J. Niiranen), seed size effects and early test problems (J. Mikola), quantitative analysis of genotypic and environmental effects (S.A. Petrov), hormonal induction of flowering (O. Luukkanen) and x-ray photography analysis of the ageing of seeds during storage (M. Ryynänen).

The PDF includes a summary in Finnish.

  • Hagman, E-mail: mh@mm.unknown (email)
article id 5026, category Article
Alpo Enivaara. (1979). Pyynikin harjumännikön uudistaminen ja elvyttäminen. Silva Fennica vol. 13 no. 2 article id 5026. https://doi.org/10.14214/sf.a14881
English title: Regeneration and improvement of the Scots pine forest of Pyynikki esker in Southern Finland.
Original keywords: mänty; uudistaminen; seminaarit; maankäyttö; harju; Pyynikki; taajamametsät
English keywords: Pinus sylvestris; regeneration; Scots pine; urban forests; seminar; MAB; natural park
Abstract | View details | Full text in PDF | Author Info

Pyynikki is a unique natural park in the centre of the city of Tampere in Southern Finland. Currently, the forest of Pyynikki is losing its vigour because of aging and environmental factors. In 1977 a policy was designed for the regrowth of Scots pine (Pinus sylvestris L.) forest and improvement of the condition of the trees to avoid the situation that the whole pine forest (34 ha) would be regrown at one time. The period of regrowth of pine forest is 60 years. It is accomplished by cutting small openings, on which 4-year old potted pine saplings are planted. Birch will be planted only on two spots. The aim is that the forest scenery will be maintained as undamaged as possible. The condition of the growing forest will be improved by fertilization and thinning.

This paper was presented in the ‘Man and the Biosphere’ programme project 2 seminar held on August 24–25 1978 in Hyytiälä research station of University of Helsinki.

The PDF includes a summary in English.

  • Enivaara, E-mail: ae@mm.unknown (email)
article id 5003, category Article
Hannu Saarenmaa. (1978). The occurrence of bark beetles (Col., Scolytidae) in a dead spruce stand flooded by beavers (Castor canadensis Kuhl.) . Silva Fennica vol. 12 no. 3 article id 5003. https://doi.org/10.14214/sf.a14857
Original keywords: kuusi; hyönteistuhot; tulva-alueet; kaarnakuoriaiset; kanadanmajava
English keywords: bark beetles; Norway spruce; Picea abies; Castor canadensis; insect damages; Trypodendron lineatum; flooded areas; Pityogenes chalcographus; Hylurgops palliatus; Ips typographus; Dryocetes autographus
Abstract | View details | Full text in PDF | Author Info

 

The aim of the study was to determine which kinds of insects had infected the Norway spruce (Picea abies (L.) H. Karst.) in different stands killed by flooding caused by beavers (Castor canadensis Kuhl), and if there was any danger that they would subsequently cause damage in the surrounding forests. The effect of tree diameter and certain stand characteristics on the fauna of dead trees are discussed. The occurrence of different insect combinations and qualifications for their coexistence were studied.

Pityogenes chalcographus L., Trypodendron lineatum O., Hylurgops palliatus Gyll. and Dryocetes autographus Ratz. occurred most abundantly. 20 phloem or wood boring species were observed in 5 regular succession types. Secondary species occurred in a virgin stand while Ips typographus L. was found at the edge of a felling area. Owing to the flooding, species preferring moist conditions were abundant. In this case damages had not spread to the surrounding forests which, however, might be possible under certain conditions.

The PDF includes a summary in English.

  • Saarenmaa, E-mail: hs@mm.unknown (email)
article id 4984, category Article
Jyrki Raulo. (1978). Forestation chain for birch (Betula pendula Roth) in Finland. Silva Fennica vol. 12 no. 1 article id 4984. https://doi.org/10.14214/sf.a14838
Keywords: forest management; Betula pendula; regeneration; silver birch; Finland
Abstract | View details | Full text in PDF | Author Info

During the 1970’s an average of 4 million Betula pendula (Roth) seedlings have been planted annually in Finland. The activities connected with the planting of this tree species, the selection of forestation sites, site preparation, planting out the seedlings and follow-up work on the forestation sites are briefly reviewed in the article. The manuscript is based on the studies into the breeding, seedling production and planting techniques of B. pendula started by the Finnish Forest Research Institute already in 1960’s, as well as on practical observations made at the planting sites. A list of some of the Finnish studies concerning B. pendula which have been published in English and studies with a summary in English is included.

The PDF includes a summary in Finnish.

  • Raulo, E-mail: jr@mm.unknown (email)
article id 4962, category Article
Matti Keltikangas, Kustaa Seppälä. (1977). Ojitusalueiden hieskoivikoiden kasvatus taloudellisena vaihtoehtona. Silva Fennica vol. 11 no. 1 article id 4962. https://doi.org/10.14214/sf.a14813
English title: The economics of growing downy birch stands on drained peatlands in Finland.
Original keywords: kannattavuus; mänty; metsän uudistaminen; ojitetut suot; hieskoivu; uudistamiskustannukset
English keywords: Pinus sylvestris; regeneration; Betula pubescens; Scots pine; downy birch; profitability
Abstract | View details | Full text in PDF | Author Info

The aim of this study was to determine under what conditions and with what premises the growing of Betula pubescens Ehrh. is an economically competitive alternative to the growing of Scots pine (Pinus sylvestris L.) in drained peatlands. The basic material consisted of all drainage projects in Ostrobothnia in Western Finland in 1937–38 and 1957–59, according to the archives of the Central Board of Forestry Tapio, including such areas that were at least moderately fertile and had birch dominated young stands or no tree cover. A total of 202 sample plots were measured.

According to the results, the discounted timber yield of the thinned B. pubescens stands is about 10% greater than that of untreated stands. The removing of birch seedlings and the subsequent growing of fully stocked Scots pine is more profitable than growing B. pubescens stands only if the establishment and subsequent development of the pine stand involve no costs. If the site in question is a fertile open drained peatland, establishment of a pine stand is obviously a better financial proposition than a naturally regenerated birch stand. However, if there is already a fully stocked young birch stand on the site, it is more economical to let it grow using a shortish rotation time.

The PDF includes a summary in English.

  • Keltikangas, E-mail: mk@mm.unknown (email)
  • Seppälä, E-mail: ks@mm.unknown
article id 4888, category Article
Olavi Luukkanen. (1973). Havaintoja kuusen vapaapölytysjälkeläistöjen ja männyn metsikköalkuperien CO2-aineenvaihdunnasta. Silva Fennica vol. 7 no. 4 article id 4888. https://doi.org/10.14214/sf.a14726
English title: Observations on CO2 exchange in open pollinated progenies of Norway spruce and provenances of Scots pine.
Original keywords: kuusi; mänty; metsänjalostus; pluspuu; fotosynteesi; geneettinen muuntelu; pimeähengitys; yhteyttämisteho
English keywords: Pinus sylvestris; Norway spruce; Picea abies; Scots pine; tree breeding; photosynthesis capacity; genetic variance; dark respiration
Abstract | View details | Full text in PDF | Author Info

Photosynthesis and dark respiration in five families of autochtonous Norway spruce (Picea abies (L.) H. Karst.) and in seedlings from twenty Finnish stands of Scots pine (Pinus sylvestris L.) were investigated in constant environmental conditions. Values of CO2 exchange were compared with the height growth and weight of seedlings in Norway spruce and with the weight alone in Scots pine. No statistically significant differences were found in CO2 exchange among progenies or stands. Photosynthetic efficiency and photosynthetic capacity showed a positive correlation both in spruce and in pine. Growth and net photosynthetic capacity were linearly and positively correlated in pine. Spruce and a higher light compensation point than pine. The use of an open IRGA system with several simultaneous measurements and the trap-type cuvette construction in genetic work are discussed.

The PDF includes a summary in English.

  • Luukkanen, E-mail: ol@mm.unknown (email)
article id 4876, category Article
Tapio Lehtiniemi, Juhani Sarasto. (1973). Kokemuksia rauduksen istutuksesta ojitetuille soille. Silva Fennica vol. 7 no. 1 article id 4876. https://doi.org/10.14214/sf.a14706
English title: Betula verrucosa plantations on peat.
Original keywords: taimet; turvemaat; uudistaminen; ojitetut suot; kuolleisuus; rauduskoivu; ravinnetasapaino
English keywords: Betula pendula; regeneration; drained peatlands; planting; nutrient deficiency; mortality; seedlings; Betula verrucosa; Godronia multispora
Abstract | View details | Full text in PDF | Author Info

The aim of the present study was to assess whether two-year old Betula verrucosa Ehrh. (now Betula pendula Roth.) transplants can be used in afforestation of drained peatlands and what factors affect the development of the young trees. The seedlings were planted in 1967. The site was repair planted next spring due to mortality caused by a undefined fungal disease, and the plantations were fertilized with NPK fertilizer (soil application. The seedlings were measured twice a year until the autumn 1970.

Only 28% of the original transplants, and 73.4% of the repair plantations were alive in 1970. In some cases, fertilization improved the results, while in others it was detrimental to the trees or had no effect on survival. According to peat analysis, the poor survival and development of the plants could be due to the too high ratios of N/Ca and N/P. Stunted or dead trees displayed often necrosis caused by Godronia multispora. According to the experiences, Betula verrucosa plantations are inferior to those obtained with Scots pine (Pinus sylvestris L.). In addition, the results indicate that in old draining areas calcium and phosphorus are often too low in comparison to nitrogen.

The PDF includes a summary in English.

  • Lehtiniemi, E-mail: tl@mm.unknown (email)
  • Sarasto, E-mail: js@mm.unknown
article id 4869, category Article
Ole Oskarsson, P. M. A. Tigerstedt. (1972). Metsänjalostuksen mahdollisuudet II. Silva Fennica vol. 6 no. 3 article id 4869. https://doi.org/10.14214/sf.a14672
English title: The possibilities in forest tree breeding II. Selection differences and genetic gains in selected seed stands of Scots pine.
Original keywords: mänty; metsänjalostus; tilavuuskasvu; valintahyöty; siemenkeräysmetsät; fenotyyppi; kuutiokasvu
English keywords: Pinus sylvestris; Scots pine; volume growth; genetic gain; forest tree breeding; seed stands; phenotype
Abstract | View details | Full text in PDF | Author Info

The aim of this study was to estimate the genetic gain of volume growth in Scots pine (Pinus sylvestris L.) selected seed stands. To obtain highest possible accuracy, the estimations are based on a large statistical material comprising 197 separate seed stands. It is concluded that the genetic gain of volume growth ranges between 7.4–15.0%. Unwanted pollen contaminations may, however, in the worst case halve this genetic gain.

The PDF includes a summary in English.

  • Oskarsson, E-mail: oo@mm.unknown (email)
  • Tigerstedt, E-mail: pt@mm.unknown
article id 4868, category Article
Mikko Tormilainen. (1972). Havaintoja Oitin siemenviljelyksen mäntykloonien kukinnasta ja käpysadosta. Silva Fennica vol. 6 no. 2 article id 4868. https://doi.org/10.14214/sf.a14671
English title: Studies of flowering and cone crop in a seed orchard of Scots pine.
Original keywords: Etelä-Suomi; mänty; kloonit; siemenviljelmä; siemenviljely; kukinta; käpysato; geneettinen vaihtelu
English keywords: Pinus sylvestris; Scots pine; flowering; genetic variation; seed orchards; clones; Southern Finland; cone production
Abstract | View details | Full text in PDF | Author Info

Male and female flowering, cone crop, and some vegetative characteristics were studied in grafts 10 to 16 years of age in a clonal seed orchard of Scots pine (Pinus sylvestris L.). Genetic variation was found between clones in flowering as well as in cone production. Clone evaluation resulted in similar classifications of clones in different years. A regression analysis showed that crown size clearly increased but previous height growth slightly decreased flowering and cone production. The percentage of pollinated female strobili did not differ between clones.

The PDF includes a summary in English.

  • Tormilainen, E-mail: mt@mm.unknown (email)
article id 4863, category Article
Olavi Luukkanen. (1972). Metsäpuiden fotosynteesin geneettinen vaihtelu. Silva Fennica vol. 6 no. 2 article id 4863. https://doi.org/10.14214/sf.a14666
English title: Genetic variation of photosynthesis in forest trees.
Original keywords: fotosynteesi; geneettinen muuntelu; hengitys; fotorespiraatio; poppeli; hiilidioksidi; kaasuaineenvaihdunta
English keywords: photosynthesis; carbon dioxide; photorespiration; CO2 exhange
Abstract | View details | Full text in PDF | Author Info

In the literature review the current status of information on the genetic variation of CO2 exchange and some reviews and investigations on this subject are listed. Photorespiration is separately discussed and unpublished data of an electron microscope study of poplar leaf microbodies are presented.

Considerable genetic inter- and intraspecific variation is found in several characteristics that affect CO2 exchange in trees. Photosynthesis in young trees does not correlate well with growth through the whole rotation cycle. A special interest has been shown to marginal environmental conditions (e.g. water deficit, low temperature, and low light intensity), as opposed to optimal conditions often employed in laboratory studies of CO2 exchange in trees.

In an unpublished poplar studies by the author et.al. a preliminary experiment with poplar clones showed variation in the CO2 competition point. This variation was negatively correlated with the photosynthesis efficiency of these clones.

The PDF includes a summary in English.

  • Luukkanen, E-mail: ol@mm.unknown (email)
article id 4832, category Article
Paavo Yli-Vakkuri, Pentti K. Räsänen. (1971). Siementen peittämisen ja kylvökohdan polkaisun vaikutus männyn ruutukylvön tulokseen. Silva Fennica vol. 5 no. 1 article id 4832. https://doi.org/10.14214/sf.a14621
English title: The influence of covering and tramping the seeds into the soil on the success of spot sowing of Scots pine.
Original keywords: kylvö; mänty; metsänviljely; taimettuminen; itäminen; kylvösyvyys; siementen suojaus; ruutukylvö
English keywords: Pinus sylvestris; germination; Scots pine; sowing; artificial regeneration; covering of the seeds; stocking with seedlings
Abstract | View details | Full text in PDF | Author Info

This paper reports spot sowing experiments of Scots pine (Pinus sylvestris L.). The seeds were either covered with coarse sand, tramped in the substrate or sowed without any covering, 30 seeds in each treatment in 70 replications. The site was of Vaccinium type with sandy soil. The germination percentage was 81 and 91 on the respective years. The development of seedlings was observed for 3–4 years.

The results indicate that both tramping and covering the seeds to some extent increased the number of seedlings and improved the early development. The highest numbers of seedlings were recorded in the first growing season, after which there was 23 seedlings/100 seeds in the uncovered spots, 27 seedlings in the covered spots and 31 seedlings in the tramped spots in the experiment sowed in 1965.

Mortality of the seedlings was highest between the first and second growing season, and empty spots increased with the time. There was no difference in mortality between the sowing methods, but the number of seedlings after first growing season affected the result. Under favourable conditions four seedlings per spot seemed enough to secure the survival of minimum one seedling per spot during the three first growing seasons. In poor conditions seven seedlings was needed.

The PDF includes a summary in English.

  • Yli-Vakkuri, E-mail: py@mm.unknown (email)
  • Räsänen, E-mail: pr@mm.unknown
article id 4816, category Article
P. M. A. Tigerstedt, Eero Malmivaara. (1970). Metsänjalostuksen mahdollisuudet I.Pluspuiden valintaero ja siemenviljelysten valintahyöty. Silva Fennica vol. 4 no. 2 article id 4816. https://doi.org/10.14214/sf.a14605
English title: The possibilities in forest tree breeding. I Selection differential of plus trees and genetic gain in seed orchards in Finland.
Original keywords: mänty; metsänjalostus; siemenviljelmät; pluspuut; valintahyöty; periytyvyys; vieraspölytys
English keywords: Pinus sylvestris; Scots pine; tree breeding; seed orchards; genetic gain; heredity; selection differential; foreign pollen sources
Abstract | View details | Full text in PDF | Author Info

An attempt was made to estimate critically the genetic gain in clonal seed orchards of Scots pine (Pinus sylvestris L.) in Finland. The selection differential of Scots pine and Norway spruce (Picea abies (L.) H. Karst.) is calculated on the basis of filed information on selected plus trees which has been kept by the genetic register at the Finnish Forest Research Institute. The differentials were computed as realized differences in height between plus-trees and normal stand characteristics on respectively site class and as a function of age.

The genetic gain in height growth of Pinus sylvestris was computed on the basis of information on selection differential and heritability. This genetic gain is between 2.6–4.4% provided there is no pollen contamination from unknown sources outside the seed orchard. The genetic gain of volume growth in Scots pine is about 7–15%, provided there is no pollen contamination in the seed orchard. However, according to investigations, there is invariably some pollen contamination in this kind of seed orchards. The contamination decreases to about 30–50% as the orchard matures and starts to produce endemic pollen. If the pollination would be entirely due to foreign pollen sources, the mathematically calculated genetic gain would be 3.5–7.5%.

The PDF includes a summary in English.

  • Tigerstedt, E-mail: pt@mm.unknown (email)
  • Malmivaara, E-mail: em@mm.unknown
article id 4784, category Article
Matti Leikola, Pentti Pylkkö. (1969). Verhopuuston tiheyden vaikutus metsikön minimilämpötiloihin hallaöinä. Silva Fennica vol. 3 no. 1 article id 4784. https://doi.org/10.14214/sf.a14570
English title: Influence of stand density on the minimum temperatures during frost nights.
Original keywords: kuusi; tiheys; koivu; uudistuminen; lämpötila; suojapuusto; halla
English keywords: Norway spruce; Picea abies; regeneration; Betula pubescens; temperature; stand density; white birch; shelter stand; frost
Abstract | View details | Full text in PDF | Author Info

The objective of this investigation was to study the influence of stand density of white birch (Betula pubescens Ehrl.) on the minimum temperatures in the stand during the growing season, and the actual minimum temperatures of the leading shoot of Norway spruce (Picea abies (L.) H. Karst.) seedlings growing in the open. The 40-year-old uniform white birch stand was situated in 142 m above the sea level in Southern Finland. The stand was treated with thinnings of three different densities in 1961.

Air temperature was recorded in four sample plots at heights of 0.1 m, 0.5 m, 1.0 m, 2 m and 4 m. In the stand of moderate density, temperatures were measured at heights of 6.0 m, and in the stand of full density at 6.0 m, 8.0 m and 10.0 m.

The temperature differences between stands of various densities proved to be rather small. Especially the thinnest stand differed very little from the open area. The soil surface has in all cases been warm compared with the higher air layers indicating meadow-fog-type by Geier (1965). On cloudy or windy weather all the temperature profiles in the various stands resembled each other. The difference between the air temperature and temperature of the spruce shoot was greatest at midnight and decreased steadily thereafter.

The problem in using shelter stands for spruce regeneration areas is that optimum shelter stand density is difficult to define. Already a thin shelter stand causes drawbacks to the young seedlings, but in order to be effective enough against early frosts, the shelter stand should be comparatively dense.

The PDF includes a summary in English.

  • Leikola, E-mail: ml@mm.unknown (email)
  • Pylkkö, E-mail: pp@mm.unknown
article id 4769, category Article
Paavo Yli-Vakkuri, Aimo Autio, Juhani Vehkaoja. (1968). Päätehakkuun ja metsänviljelyn välinen aika. Silva Fennica vol. 2 no. 3 article id 4769. https://doi.org/10.14214/sf.a14554
English title: Time lag between final cutting and regeneration.
Original keywords: kylvö; metsänviljely; istutus; uudistaminen; uudistusaika; istutusrästit; viivästys
English keywords: planting; final felling; sowing; artificial regeneration; seeding; time lag
Abstract | View details | Full text in PDF | Author Info

The time interval between final felling and plantation means a waste of time and thus a production loss, and may lead to lush growth of ground cover and hardwood sprouts, which increases expenses in forest management. The objective of this study was to determine the length of time between final felling and artificial regeneration in private forests in the forest districts of Uusimaa-Häme in Southern Finland and Pohjois-Häme in Central Finland. The material consists of a sample of 150 plans of the 952 cutting and regeneration plans in the district of Uusimaa-Häme and a sample of 140 plans of the 1,102 plans in Pohjois-Häme.

The time lag between final cutting and seeding or planting was on average 1.4 years in Pohjois-Häme district and 0.7 years in Uusimaa-Häme. In the latter district, 56% of the logged area was regenerated in the spring immediately following the cutting, and 84% not later than in the second spring. In Pohjois-Häme, 29% of the harvested area was regenerated immediately in the first spring following cutting, and 79% not later than in the third spring following cutting.

In Pohjois-Häme, the interval was shortest in the smallest forest holdings, and longest in the largest holdings with the largest regeneration areas. The length seems to depend mainly on the size of the regeneration area. In Uusimaa-Häme district, the interval was shortest in the smallest holdings, rather short in the largest, and longest in the intermediate-size forest properties. Seeding with Scots pine (Pinus sylvestris L.) was used in almost all regeneration areas.

The forest owners had mainly carried out the regeneration work themselves. In the Pohjois-Häme area, the interval was shorter when the district forestry board regenerated the area. 35% of the regenerated areas had required supplementary planting in Pohjois-Häme, and 47% in the Uusimaa-Häme area. Supplementary planting was more common in areas regenerated later after the cutting. In Pohjois-Häme, according to the reports of the forest owners, 75% of the regenerated areas required tending during the first three years, in Uusimaa-Häme, 80%.

The PDF includes a summary in English.

  • Yli-Vakkuri, E-mail: py@mm.unknown (email)
  • Autio, E-mail: aa@mm.unknown
  • Vehkaoja, E-mail: jv@mm.unknown
article id 4731, category Article
Leo Heikurainen, Juhani Päivänen, Kustaa Seppälä. (1966). Koetuloksia männyn kylvöstä ja istutuksesta ojitetuilla soilla. Silva Fennica vol. no. 119 article id 4731. https://doi.org/10.14214/sf.a14289
English title: Scots pine seeding and planting on drained peat soils.
Original keywords: kylvö; mänty; istutus; uudistaminen; lannoitus; turvekankaat; ojitetut suot
English keywords: Pinus sylvestris; fertilization; regeneration; Scots pine; planting; sowing; artificial regeneration; seeding
Abstract | View details | Full text in PDF | Author Info

This paper describes the preliminary results of Scots pine (Pinus sylvestris L.) seeding and planting trials on drained peat soils.

The results showed that a perpared peat surface was a better surface for seeding than the unprepared one. Planting of 2+1-year seedlings succeeded better than planting 1-year seedlings. Planting on the turf gave better survival than planting on the unprepared soil surface. The whole growing season was suitable time for planting Scots pine seedlings except May when the peat soil under the surface was still frozen.

Using fertilizers in connection with planting was surveyed in two ways. Mortality of seedlings increased when they were top-dressed with NPK fertilizer. Using a so-called spot fertilizing with several combinations of fertilizers resulted in K and N tending to increase the mortality of seedlings, but P decreasing mortality.

The PDF includes a summary in English.

  • Heikurainen, E-mail: lh@mm.unknown (email)
  • Päivänen, E-mail: jp@mm.unknown
  • Seppälä, E-mail: ks@mm.unknown
article id 4723, category Article
Toivo Matilainen. (1965). Kasvutappiosta ja sen määrittämisestä Etelä-Suomen talousmetsissä. Silva Fennica vol. no. 117 article id 4723. https://doi.org/10.14214/sf.a14281
English title: The increment loss and its determination in economically exploited forests in Southern Finland.
Original keywords: Etelä-Suomi; menetelmät; vajaatuottoiset metsät; kasvutappiot; uudistaminen
English keywords: regeneration; Southern Finland; methods; under-productive forests; deficient stock volume; increment loss
Abstract | View details | Full text in PDF | Author Info

The present study is an attempt to clarify the decrease in growth, or the increment loss, caused by sudden reduction of growth of the growing stock below a certain level, and to find a method for its determination. Increment loss is defined as a decrease in growth during the rotation due to a deficient stock volume. The material consists of Koivisto’s yield tables for repeatedly thinned stands in Southern Finland, and the results of the Third National Forest Inventory concerning the mean volume and increment in the productive sites.

For the calculation of increment loss three formulae were constructed where the increment loss is calculated 1) as the difference between the removal by thinnings in normally developed stands during a time equal with the period of deficient stock and the suddenly removed stock, 2) according to the compound interest calculation principle as the sum of the differences which are obtained by subtracting from the removal in each thinning during the period of deficient stock its initial value, and 3) as the straight interest of the stock deficiency during the period of deficient stock.

According to the calculations, the increment loss is greatest in stands to be grown, viz. 50 m3 solid measure excluding bark per hectare tended Norway spruce stands on Oxalis-Myrtillus type sites at 40% deficiency below minimum stock. In stands to be regenerated the losses are, too, greatest in the similar stands. It exceeds 200 m3/ha when stands younger than 50 years have to been regenerated and the removal amounts to 50% of the stock. In stands to be regenerated the increment loss for spruce, due to the slow initial development by the species, is greater than for Scots pine and birch. The loss is the same at different period of age if the relative deficiency of the stock is of equal size.

According to the study, each stand has a characteristic variation in the increment loss which depends mainly on the relative degree of deficiency from the minimum stock. The formulae and methods can be used to determine the increment loss in average and better stands in Southern Finland when the stock suddenly decreases.

The PDF includes a summary in English.

  • Matilainen, E-mail: tm@mm.unknown (email)
article id 4669, category Article
V. A. Kolehmainen. (1957). Vehkatallinmaa. Silva Fennica vol. no. 90 article id 4669. https://doi.org/10.14214/sf.a9116
English title: Vehkatallinmaa, a successful reforestation area in Central Finland.
Original keywords: luontainen uudistuminen; kylvö; metsitys; metsäpalot
English keywords: regeneration; natural regeneration; forest fire; sowing
Abstract | View details | Full text in PDF | Author Info

In 1933, forest fire caused by locomotive sparkle burned about 600 hectares of forest in a forest district named Vehkatallinmaa, in Central Finland. In 1934–36, the burned area was reforested, using different sowing and planting methods. At the same time, areas with poor runoff were drained. The results from reforestation throughout the area have been good. Also, natural regeneration of coniferous trees, especially Scots pine (Pinus sylvestris L.) has occurred. Even deciduous trees, especially birch (Betula sp.), have regenerated naturally in the area. The forests are an evidence of the adaptability of broadcast sowing on snow crust as a method of reproduction.

The PDF includes a summary in English.

  • Kolehmainen, E-mail: vk@mm.unknown (email)
article id 4644, category Article
Vilho Antero Kolehmainen. (1955). Havaintoja kulotuksen merkityksestä metsiemme uudistamisessa. Silva Fennica vol. no. 85 article id 4644. https://doi.org/10.14214/sf.a9106
English title: Effect of prescribed burning in the forest regeneration.
Original keywords: luontainen uudistaminen; kulotus; metsänuudistaminen; taimettuminen; siemenpuu
English keywords: Pinus sylvestris; Norway spruce; Picea abies; natural regeneration; Scots pine; Betula sp.; seed trees; prescribed burning
Abstract | View details | Full text in PDF | Author Info

Prescribed burning has reported to avail forest regeneration, for instance, by releasing nutrients for the use of seedlings, changing the pH of the soil and decreasing competition of ground vegetation. The aim of the study was to find out if the effects could be verified. Sample plots were measured in the experimental area of Tuomarniemi, in Central Finland, both in previously burned and untreated seedling stands and young forests. The main species in the sample plots was Scots pine (Pinus sylvestris L.).

According to the results, prescribed burning prepares the soil for regeneration. Germination percentage of the seeds is higher on the burned soil. All the species, Scots pine, Norway spruce (Picea abies (L.) Karst.) and birch species (Betula sp.) grow faster. Prescribed burning increases the amount of birch seedlings by improving its regeneration compared to unburned sites. The seed trees survive burning better if they are tall and have short crown, and have thick bark. In general, prescribed burning improves regeneration in seed tree stands.

The article includes a summary in German.

  • Kolehmainen, E-mail: vk@mm.unknown (email)
article id 4638, category Article
Risto Sarvas. (1953). Ohjeita pluspuiden valitsemista ja ilmoittamista varten. Silva Fennica vol. no. 80 article id 4638. https://doi.org/10.14214/sf.a14054
English title: Instruction for the selection and registration of plus trees.
Original keywords: metsäopetus; jatkokoulutus; metsänuudistaminen; siementuotanto; metsänjalostus; pluspuu
English keywords: regeneration; selection; breeding; seed production; forest education
Abstract | View details | Full text in PDF | Author Info

Silva Fennica Issue 80 includes presentations held in 1952 in the 7th professional development courses, arranged for foresters working in the Forest Service. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.

This presentation describes principles of breeding of forest trees and the selection of fast growing, healthy and qualitatively good trees, named plus trees, for seed production in Finland.

  • Sarvas, E-mail: rs@mm.unknown (email)
article id 4637, category Article
Eero Paavola. (1953). Selostus kenttäkokeista, jotka koskivat eräiden kemiallisten aineiden käyttömahdollisuuksia metsänkylvössä. Silva Fennica vol. no. 80 article id 4637. https://doi.org/10.14214/sf.a14053
English title: Review of field experiments on the fitness of certain chemicals for use at seeding.
Original keywords: metsäopetus; kylvö; metsänviljely; jatkokoulutus; metsänuudistaminen; heinäntorjunta; herbisidit; torjunta-aineet
English keywords: ground vegetation; herbicides; forest regeneration; forest education; seeding
Abstract | View details | Full text in PDF | Author Info

Silva Fennica Issue 80 includes presentations held in 1952 in the 7th professional development courses, arranged for foresters working in the Forest Service. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.

This presentation describes a field experiment on the use of herbicides to prevent growth of ground vegetation in cutting areas before sowing. The study suggests that the chemicals used in the experiment were not effective enough to prevent growth of ground vegetation in the more fertile lands, but were effective in poorer lands. However, the treatment affected also growth of tree seedlings.

  • Paavola, E-mail: ep@mm.unknown (email)
article id 4636, category Article
Tauno Hautamäki. (1953). Metsämaan laikutuksen koneellistamiskokeiluista. Silva Fennica vol. no. 80 article id 4636. https://doi.org/10.14214/sf.a14052
English title: Mechanization of ground preparation.
Original keywords: metsäopetus; jatkokoulutus; maanmuokkaus; metsänuudistaminen; koneellistaminen; laikutus
English keywords: regeneration; soil preparation; mechanization; scalping; forest education
Abstract | View details | Full text in PDF | Author Info

Silva Fennica Issue 80 includes presentations held in 1952 in the 7th professional development courses, arranged for foresters working in the Forest Service. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.

Soil preparation improves formation of seedlings in natural regeneration sites. This presentation describes mechanization of the work using four different machines. According to the study, the effectiveness and quality of the work has improved compared to the earlier machines, and the method is becoming a competitive alternative to slash burning.

  • Hautamäki, E-mail: th@mm.unknown (email)
article id 4628, category Article
Gustaf Sirén. (1952). Havaintoja Peräpohjolan valtion mailla vuosina 1948-50 suoritetuista männyn kylvöistä. Silva Fennica vol. no. 78 article id 4628. https://doi.org/10.14214/sf.a9100
English title: Observations on stands of Scots pine sown in state forests in Peräpohjola in northern Finland in 1948-50.
Original keywords: Pohjois-Suomi; valtionmetsät; kylvö; mänty; metsänviljely; Peräpohjola; uudistuminen
English keywords: Pinus sylvestris; regeneration; Scots pine; northern Finland; state forests; sowing
Abstract | View details | Full text in PDF | Author Info

The most common way to artificially regenerate cutting areas in Northern Finland has been sowing, which has, however, often given poor results. The aim of the study was to assess the success rate of sowing and study the causes of poor regeneration.

An inventory was made of 28 areas on Empetrum-Myrtillus and Hylocomnium-Myrtillus type sites sown with Scots pine (Pinus sylvestris L.) seeds in 1948-1950. In addition, to study the effect of substrata, a sowing experiment was established. According to the inventory, regeneration failed completely in 8 and partially in 6 of the 28 sown areas. Factors that explained the poor regeneration included unfavourable weather conditions in 1948-1952, insufficient clearing of sowing spot especially when the humus layer was thick, and insect damage by Otiorynchus dubius weevil.

The article includes a summary in English.

  • Sirén, E-mail: gs@mm.unknown (email)
article id 4578, category Article
Sakari Saarnijoki. (1942). Jalavan esiintymisestä Pyhäjärven, Kokemäenjoen vesistön keskusjärven tulvarannoilla. Silva Fennica vol. no. 58 article id 4578. https://doi.org/10.14214/sf.a9082
English title: Distribution of European white elm in flooded shores of the central lake of Pyhäjärvi and Kokemäenjoki water system.
Original keywords: levinneisyys; jalava; kynäjalava; vuorijalava; rantametsä; tulva-alueet; Ulmus glabra; Ulmus laevis; Ulmus montana; uudistuminen
English keywords: regeneration; distribution; elm; European white elm; coastal forests; flood plains
Abstract | View details | Full text in PDF | Author Info

Distribution of European white elm (Ulmus laevis Pallas) is on its northernmost border in Pyhäjärvi and Kokemäenjoki area. This survey describes distribution of European white elm in the flooded shores of the central lake of Pyhäjärvi and Kokemäenjoki river water system.

Both Ulmus laevis and U. montana (now U. Glabra Hudson) can be found in the area, but most of the elms qrowing in thea area are U. laevis. U. laevis occurs around the lake in two separate areas, almost entirely in flooded shores of the lake. Regeneration of elm from seeds was limited on a narrow belt on the higher part of the flooded shore. Consequently, U. laevis can be found as zones around the lake, created by the changes in water level of the lake. The trees are judged to be native for the area.

The article includes an abstract in German.

  • Saarnijoki, E-mail: ss@mm.unknown (email)
article id 4568, category Article
Atri S. Arimo. (1939). Paloalojen metsittämisestä. Silva Fennica vol. no. 52 article id 4568. https://doi.org/10.14214/sf.a13975
English title: Forest regenration of burned areas.
Original keywords: metsäopetus; kylvö; metsänviljely; metsänhoitajien jatkokurssit; kulotus; metsäpalo; jatkokoulutus; metsänuudistaminen; kulo
English keywords: regeneration; planting; forest fire; forest education; sowing; professional development courses
Abstract | View details | Full text in PDF | Author Info

Silva Fennica issue 52 includes presentations held in professional development courses, arranged for foresters working in public administration in 1938. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.

This presentation describes preparation and forest regeneration methods of a burned area.

  • Arimo, E-mail: aa@mm.unknown (email)
article id 4559, category Article
A. A. Räsänen. (1939). Metsien uudistamisesta Perä-Pohjolassa. Silva Fennica vol. no. 52 article id 4559. https://doi.org/10.14214/sf.a13966
English title: Forest regeneration in Northern Finland.
Original keywords: hakkuut; metsäopetus; metsänviljely; metsänhoitajien jatkokurssit; jatkokoulutus; maanmuokkaus; metsänuudistaminen; Lappi; yli-ikäiset metsät
English keywords: forest management; Norway spruce; Picea abies; regeneration; fellings; forest education; professional development courses
Abstract | View details | Full text in PDF | Author Info

Silva Fennica issue 52 includes presentations held in professional development courses, arranged for foresters working in public administration in 1938. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.

This presentation describes the state of forests in Northern Finland, and suggests that fellings are targeted at the oldest forests until the over-presented over-aged forest are harvested. The preferred regeneration method is natural regeneration in most forest types.

  • Räsänen, E-mail: ar@mm.unknown (email)
article id 4558, category Article
Olli Heikinheimo. (1939). Kokemuksia paksusammaltyypin metsien käsittelystä. Silva Fennica vol. no. 52 article id 4558. https://doi.org/10.14214/sf.a13965
English title: Experiences in forest management of Hylocomnium-Myrtillus type forests in Northern Finland.
Original keywords: hakkuut; metsänhoito; metsäopetus; metsänhoitajien jatkokurssit; jatkokoulutus; metsänuudistaminen; paksusammalkuusikko; Peräpohjola; Lappi; siementuotanto
English keywords: Norway spruce; Picea abies; regeneration; Lapland; seed production; fellings; forest education; professional development courses; Hylocomnium-Myrtillus type
Abstract | View details | Full text in PDF | Author Info

Silva Fennica issue 52 includes presentations held in professional development courses, arranged for foresters working in public administration in 1938. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.

This presentation describes a study of forest management practices suitable for in Hylocomnium-Myrtillus site type forests, typical for Northern Finland. A special problem for the forest site type is poor regeneration due weak seed production of Norway spruce (Picea abies L. Karst.) and thick moss layer.

  • Heikinheimo, E-mail: oh@mm.unknown (email)
article id 4571, category Article
Tauno Lampimäki. (1939). Nautakarjan laiduntamisesta metsämailla. Silva Fennica vol. no. 50 article id 4571. https://doi.org/10.14214/sf.a9074
English title: Grazing of cattle in forest land.
Original keywords: metsätalous; taimettuminen; metsänuudistuminen; laiduntaminen; laitumet; karja; karjatalous; pintakasvillisuus; taimet; taimituhot; metsälaidun
English keywords: regeneration; forestry; ground vegetation; tree seedlings; grazing; cattle; animal husbandry; woodland pasture
Abstract | View details | Full text in PDF | Author Info

Finland has a long tradition of grazing cattle in the forests and common land. There are also reports of degradation of forests by grazing already in 1600th century. The aim of the survey was to study which positive and negative effects grazing has in forests.

The study concludes that grazing has caused considerable economic losses through damages to forests. In addition, woodland pastures cannot give the yields required in modern animal husbandry. The quality of woodland pastures have decreased after the woodlands used in slash and burn culture have become wooded.

Grazing has also some positive effects to forests. It increases the diversity of vegetation in the woodland pastures and spreads species to new areas. This is supported by the lists of species found in different woodland pastures. Cattle destroy large grasses like Calamagrostis, which may avail growth of tree seedlings in the pastures. Grazing can also prepare the site for tree seedlings. On the other hand, prolonged grazing destroys tree seedlings and prevents regeneration.

The article includes a German summary.

  • Lampimäki, E-mail: tl@mm.unknown (email)
article id 4543, category Article
Erkki Laitakari. (1938). Metsän uudistamisesta laihoilla kangasmailla. Silva Fennica vol. no. 46 article id 4543. https://doi.org/10.14214/sf.a13952
English title: Forest regeneration on poor forest sites.
Original keywords: luontainen uudistuminen; kuiva kangas; metsäopetus; metsänhoitajien jatkokurssit; kulotus; kuivahko kangas; kasvupaikka; metsänuudistuminen; uudistushakkuu
English keywords: natural regeneration; forest regeneration; forest education; professional development courses; broadcast burning; dry upland forest site; dryish upland forest site; regeneration felling
Abstract | View details | Full text in PDF | Author Info

Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service. 

This presentation describes forest regeneration on poor forest sites.

  • Laitakari, E-mail: el@mm.unknown (email)
article id 4542, category Article
Erkki Laitakari. (1938). Eräitä metsän uudistamisessa huomioon otettavia näkökohtia. Silva Fennica vol. no. 46 article id 4542. https://doi.org/10.14214/sf.a13951
English title: Forest regeneration.
Original keywords: metsäopetus; metsänviljely; metsänhoitajien jatkokurssit; luontainen uudistaminen; metsänuudistaminen
English keywords: natural regeneration; planting; forest education; professional development courses; articifial regeneration
Abstract | View details | Full text in PDF | Author Info

Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service. 

This presentation describes factors that should be taken into account in forest regeneration. 

  • Laitakari, E-mail: el@mm.unknown (email)
article id 4536, category Article
V. K. Ahola. (1938). Metsän uudistumisen tarkkailusta kovilla metsämailla ja ojitetuilla soilla sekä ojien tarkkailusta. Silva Fennica vol. no. 46 article id 4536. https://doi.org/10.14214/sf.a13945
English title: Inspection of regeneration of forests in mineral soil forests and drained peatlands and checking of ditches.
Original keywords: valtionmetsät; ojitus; metsäopetus; metsänviljely; metsänhoitajien jatkokurssit; luontainen uudistaminen; suot; hoitoalueet; taimettuminen; taimikko; ojitusalueet
English keywords: natural regeneration; drained peatlands; peatlands; seedling stands; forest education; state forests; artificial regeneration; professional development courses; districts
Abstract | View details | Full text in PDF | Author Info

Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service. 

This presentation describes inspection of forest regeneration of mineral soil forest types and drained peatlands, and inspection of ditches. 

  • Ahola, E-mail: va@mm.unknown (email)
article id 4534, category Article
Martti Tertti. (1938). Hakkausalan raivaamisesta. Silva Fennica vol. no. 46 article id 4534. https://doi.org/10.14214/sf.a13943
English title: Clearing of a felling area.
Original keywords: hakkuut; metsäopetus; metsänhoitajien jatkokurssit; metsänraivaus; hakkuuala; metsänuudistaminen; raivaus; alikasvos
English keywords: regeneration; fellings; forest education; professional development courses; cutting area; clearing of felling areas; undergrowth
Abstract | View details | Full text in PDF | Author Info

Silva Fennica issue 46 includes presentations held in professional development coursesarranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administrationespecially in regional levelThe education was arranged by Forest Service. 
This presentation describes clearing of felling areas. 

  • Tertti, E-mail: mt@mm.unknown (email)
article id 4530, category Article
O. J. Lukkala. (1938). Ojitettujen soiden metsittämisestä. Silva Fennica vol. no. 46 article id 4530. https://doi.org/10.14214/sf.a13939
English title: Afforestation of drained peatlands.
Original keywords: valtionmetsät; Metsähallitus; ojitus; metsäopetus; metsänviljely; metsänhoitajien jatkokurssit; luontainen uudistaminen; metsitys; suot
English keywords: afforestation; natural regeneration; drained peatlands; peatlands; forest regeneration; Forest Service; forest education; state forests; professional development courses
Abstract | View details | Full text in PDF | Author Info

Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service. 

This presentation describes afforestation of drained peatlands. 

  • Lukkala, E-mail: ol@mm.unknown (email)
article id 4529, category Article
V. T. Aaltonen. (1938). Maa ja metsän uudistuminen. Silva Fennica vol. no. 46 article id 4529. https://doi.org/10.14214/sf.a13938
English title: Forest regeneration and soil.
Original keywords: metsäopetus; metsänhoitajien jatkokurssit; luontainen uudistaminen; kulotus; maanmuokkaus; metsämaa; maaperä
English keywords: regeneration; soil preparation; natural regeneration; soil; forest education; professional development courses; broadcast burning
Abstract | View details | Full text in PDF | Author Info

Silva Fennica issue 46 includes presentations held in professional development courses, arranged for foresters working in public administration in 1937. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service. 

This presentation describes the effect of properties of soil on forest regeneration. 

  • Aaltonen, E-mail: va@mm.unknown (email)
article id 4492, category Article
V. K. Ahola. (1937). Metsän keinollisesta uudistamisesta. Silva Fennica vol. no. 39 article id 4492. https://doi.org/10.14214/sf.a13902
English title: Artificial regeneration of forests.
Original keywords: metsäopetus; kylvö; metsänhoitajien jatkokurssit; metsän uudistaminen; istutus
English keywords: planting; forest regeneration; forest education; sowing; professional development courses
Abstract | View details | Full text in PDF | Author Info

Silva Fennica Issue 39 includes presentations held in professional development courses in 1935 that were arranged for foresters working in public administration. The presentations focus on practical issues in forest management and administration, especially in regional level.

 This presentation describes the methods of artificial regeneration of forests.

  • Ahola, E-mail: va@mm.unknown (email)
article id 4486, category Article
Martti Tertti. (1937). Metsien luontaisen uudistumisen edistämisestä. Silva Fennica vol. no. 39 article id 4486. https://doi.org/10.14214/sf.a13896
English title: Facilitating natural regeneration.
Original keywords: metsänhoito; metsäopetus; kylvö; metsänhoitajien jatkokurssit; luontainen uudistaminen; taimetuminen
English keywords: natural regeneration; silviculture; forest education; sowing; professional development courses
Abstract | View details | Full text in PDF | Author Info

Silva Fennica Issue 39 includes presentations held in professional development courses in 1935 that were arranged for foresters working in public administration. The presentations focus on practical issues in forest management and administration, especially in regional level. The education was arranged by Forest Service.

This presentation describes natural regeneration in forests and factors affecting the success of regeneration.

  • Tertti, E-mail: mt@mm.unknown (email)
article id 4479, category Article
L. E. T. Borg. (1936). Hankikylvöt Tuomarniemen hoitoalueessa vv. 1913-1930. Silva Fennica vol. no. 38 article id 4479. https://doi.org/10.14214/sf.a9065
English title: Areas broadcast sown on snow in Tuomarniemi district in Finland in 1913-1930.
Original keywords: kuusi; Picea abies; kylvö; euroopanlehtikuusi; kylvömenetelmät; hankikylvö; Larix decidua; mänty; metsänviljely; Pinus sylvestris
English keywords: Norway spruce; sowing; broadcast sowing on snow; Socts pine; artificial regeneration
Abstract | View details | Full text in PDF | Author Info

Broadcast sowing on snow was relatively new method in the beginning of the 20th century in Finland, and the experiences of regeneration were diverse. The aim of the survey was to study the success rate of regeneration in the oldest and largest areas regenerated with this sowing method in Tuomarniemi district. Scots pine (Pinus sylvestris L.) was the most common tree species, but also Norway spruce (Picea abies (L.) H. Karst.) and European larch (Larix decidua Mill.) were used in broadcast sowing on snow.

According to the study, the success of broadcast sowing on snow was as good as patch sowing and sowing in furrows in the sites typical for Tuomarniemi. The regeneration areas were often drained peatlands or paludified lands. When sowing is done using Norway spruce seeds, site preparation either by broadcast burning or scalping with hoe is recommended. Mixed sowing with pine and spruce seldom succeeded due to the differences in site requirements of the species and growth of seedlings. Sowing of Scots pine succeeded well on the drained peatlands. Sowing should be done some years after draining to let the peat dry and sink. Site preparation is needed in sites growing Polytrichum-moss. Broadcast burned areas larger than 10 hectares seemed to regenerate poorer than sites in average, possibly due to dryness of the sites. Trials with European larch were successful, and the growth of the seedlings acceptable despite the sites being relatively poor for the species.

The PDF includes a summary in German.

  • Borg, E-mail: lb@mm.unknown (email)
article id 4451, category Article
Pekka Urho Sakari Tikka. (1928). Havaintoja kuusen esiintymisestä ja kehityksestä Pohjois-Suomen kuivissa kangasmetsissä. Silva Fennica vol. 0 no. 10 article id 4451. https://doi.org/10.14214/sf.a8399
English title: Observations on Norway spruce growth and development in dry mineral soil sites in Northern Finland.
Original keywords: Pohjois-Suomi; luontainen uudistuminen; kuiva kangas; kuusi; Picea abies; taimi
English keywords: Norway spruce; natural regeneration; seedling; northern Finland; dry upland site
Abstract | View details | Full text in PDF | Author Info

Norway spruce (Picea abies (L.) H. Karst.) is rarely the dominant species on dry mineral soil sites in Northern Finland. These sites are, in general, too poor and dry for spruce, and suit better for Scots pine (Pinus sylvestris L.). According to the study, the natural regeneration of spruce is in Northern Finland poor. In the sample plots, cones could be found in 35% of spruce trees in the stands in natural state and 46% in the harvested stands. Compared to the spruce areas in Northern Finland, or fresh mineral soil sites in Southern Finland, cone and seed production of Norway spruce was in dry mineral soil sites very low due to scarcity of seed trees and their low cone number. There were few spruce seedlings in the sample plots, but according to the observations, spruce is able to regenerate on lichen and heath covered sites. The seedling growth was, however, poor on dry sites. Spruce seedlings were often found near fallen trees and stumps. The growing trees prevent growth of seedlings of all species. Norway spruce seems, however, to be able to spread also to the poor sites. The success depends on the vegetation and dryness of the site. For instance, spruce can spread to dry mineral soil sites from seed trees of nearby peatlands.

The PDF includes a summary in German.

  • Tikka, E-mail: pt@mm.unknown (email)

Category: Article

article id 7181, category Article
Eino Oinonen. (1967). Keltalieon (Lycopodium complanatum L.) itiöllinen uudistuminen Etelä-Suomessa kloonien laajuutta ja ikää koskevan tutkimuksen valossa. Silva Fennica vol. 83 no. 3 article id 7181. https://doi.org/10.14214/aff.7181
English title: Sporal regeneration of ground pine (Lycopodium complanatum L.) in Southern Finland in the light of the dimensions and the age of its clones.
Keywords: regeneration; forest fire; Pteridium aquilinum; bracken fern; ground pine; Lycopodium complanatum; Diphasiastrum complanatum; sporal regeneration
Abstract | View details | Full text in PDF | Author Info

In an earlier paper of the author it was established that sporal regeneration of bracken fern (Pteridium aquilinum (L.) Kuhn.) is almost entirely connected with fire, and that the size of the bracken fern clones, distinguished by their characteristics, was related to the time of the fire. In previous studies has been found that also sporal regeneration of ground pine (Lycopodium complanatum L., now Diphasiastrum complanatum) is rare. The occurrence of several plant species were studied in relation to bracken fern in the earlier investigation. This paper reports findings concerning ground pine.

When the size-age problem of bracken fern was solved, parallel measurements of ground pine stand on the same site led to the solution of the size-age problem and sporal regeneration of ground pine. The linkage is also valid when the size of ground pine stands was compared to the dates of fires. The ground pine stands are very long-lasting. The stands are fragmented more easily than bracken fern by environmental factors, such as fires, and tend to form large patch clusters with time. Large individual stands reveal the rarity of sporal regeneration of ground pine. The resemblance with bracken fern clones indicate a common factor of regeneration, fire, and a very even spreading rate. Though considerable variation of the colour and structure of ground pine was observed, the circular stands were identical patch by patch.

The PDF includes a summary in English.

  • Oinonen, E-mail: eo@mm.unknown (email)
article id 7180, category Article
Eino Oinonen. (1967). The correlation between the size of Finnish bracken (Pteridium aquilinum (L.) Kuhn.) clones and certain periods of site history. Silva Fennica vol. 83 no. 2 article id 7180. https://doi.org/10.14214/aff.7180
Keywords: regeneration; distribution; forest fire; Pteridium aquilinum; bracken fern; wars; ground fire
Abstract | View details | Full text in PDF | Author Info

In an earlier paper of the author it was established that bracken fern (Pteridium aquilinum (L.) Kuhn.) that the sporal regeneration of the species is almost entirely connected with fire, and that the size of the bracken fern clones, distinguished by their characteristics, was related to the time of the fire. The time of the fire was determined by the samples taken with auger from the trees on the sample plots. A large data on the occurrence of bracken fern was collected around Finland, but only part of it could be used in the previous study, as the trees of the site provided no means to find out it the site had been burned earlier.

In earlier studies, a mass occurrence of sporelings of bracken fern in England has been connected to cities bombed and burned during the World War II. In this study the data of the clones is connected to historical sources and the time of wars in Finland. The comparisons in this study have confirmed the results achieved by comparing the dimensions of the bracken clones to the fire dates. It was found that the wars have caused an increase in the regeneration frequency of bracken fern.

  • Oinonen, E-mail: eo@mm.unknown (email)
article id 7179, category Article
Eino Oinonen. (1967). Sporal regeneration of bracken (Pteridium aquilinum (L.) Kuhn.) in Finland in the light of the dimensions and the age of its clones. Silva Fennica vol. 83 no. 1 article id 7179. https://doi.org/10.14214/aff.7179
Keywords: regeneration; distribution; Pteridium aquilinum; bracken fern
Abstract | View details | Full text in PDF | Author Info

The objective of the present study was to obtain information on the regeneration by spores of bracken fern (Pteridium aquilinum (L.) Kuhn.) as well as the permanence and vegetative spreading of the clones. Prevention of bracken fern, which can hinder forest regeneration, is possible only when the basic facts of the life cycle is known. It would be useful to know if it is probable that bracken fern can become prevalent by spore regeneration, and to what extent it invades forest land by vegetative spreading.

No bracken fern sporelings were found, and the experiments of regeneration by spores were unsuccessful, therefore, the studies were concentrated on the clones. Due to the fire tolerance of the plant, it was expected that the clone material would show gradation according to the periods of fire. The first part of the work concerns individual variation in bracken, the second part concentrates on the data collected from sample plots, which latest fire could be dated from the trees.

The results of the studied clones confirm the correlation between the stands interpreted as clones and the length of the post-fire period. The stands were delimited to clones by their individual characteristics. The spreading on average sites (Vaccinium and Myrtillus type forests) was in average 35.8±2 cm annually. It is concluded that bracken fern regeneration by spores is very rare in Finland and that it has been rarer during the last 50 years compared to the preceding period. This is probably caused by the end of slash-and-burn cultivation and decrease of forest fires in Finland. Bracken fern clones often survive the fires through their rhizomes, and fragments of old clones originating before the fire could be found in the study areas. These clones may be even 300 years old. Especially in its northern areas of distribution, bracken fern is connected with settlement and old slash and burn sites. Regeneration by spores in the northern distribution areas seem to be less than in the southern areas.

  • Oinonen, E-mail: eo@mm.unknown (email)
article id 7178, category Article
Matti Keltikangas, Pekka Tiililä. (1968). Koivun ja kuusen istutuksen keskinäinen edullisuusjärjestys käenkaali-mustikkatyypin metsämailla. Silva Fennica vol. 82 no. 5 article id 7178. https://doi.org/10.14214/aff.7178
English title: The economic sequence of silver birch (Betula pendula) and Norway spruce (Picea abies) when planting Oxalis-Myrtillus type forest land.
Original keywords: kuusi; kannattavuus; koivu; istutusmetsikkö; puulajivalinta; tuottotaulukot
English keywords: Norway spruce; Betula pendula; Picea abies; silver birch; planting; profitability; artificial regeneration; yield tables; choise of tree species
Abstract | View details | Full text in PDF | Author Info

The present study proposes to calculate the economic sequence of two of Finland’s three main tree species, Norway spruce (Picea abies (L.) H. Karst.) and silver birch (Betula pendula Roth) when planted on Oxalias-Myrtillus type sites where both species are equally suitable, on biological grounds. In addition, the accuracy and applicability of the present Finnish yield tables to an economic comparison is tested. Benefit/cost ratio was selected as criterion of profitableness. All future net incomes and costs were discounted into the planting time and added together. The ratio between the discounted net revenues and the discounted investment costs (later called profit ratio) was the criterion. There is no reliable method to forecast the future wood prices, therefore two price ratios, birch veneer timber to spruce pulpwood and birch cordwood to spruce pulpwood, were chosen as free variables. The economic sequence of the tree species was determined as the function of these variables.

The main conclusions are, first, that under the present price ratios spruce appears to be the better choice for the forest owner, and the most promising policy for changing the situation seems to decrease the production costs of plants in birch nurseries. Second, the present Finnish yield tables are not consistent or accurate enough to enable any sufficiently reliable economic comparisons of tree species in artificial regeneration. The possible error of difference between two rather uncertain estimates is big. More work is needed to construct a uniform system of yield tables covering all main tree species, all site types, all macro climate conditions and all types of regeneration.

The PDF includes a summary in English.

  • Keltikangas, E-mail: mk@mm.unknown (email)
  • Tiililä, E-mail: pt@mm.unknown
article id 7135, category Article
Paavo Yli-Vakkuri. (1961). Kokeellisia tutkimuksia taimien syntymisestä ja ensi kehityksestä kuusikoissa ja männiköissä. Silva Fennica vol. 75 no. 1 article id 7135. https://doi.org/10.14214/aff.7135
English title: Experimental studies on the emergence and development of tree seedlings in spruce and pine stands.
Original keywords: kuusi; kylvö; mänty; taimettuminen; uudistuminen; uudistaminen; itäminen
English keywords: Pinus sylvestris; Norway spruce; Picea abies; germination; regeneration; Scots pine; seedlings; sowing; seeding; restocking with seedlings
Abstract | View details | Full text in PDF | Author Info

This paper aims at studying regeneration of Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) by sowing and natural regeneration of birch (Betula sp.) in Western Finland.

Germination of spruce and pine seeds may be prevented by dryness and temperatures below the optimum for germination. In natural conditions, when temperature and moisture is insufficient for germination, the type of seedbed generally has en effect on germination result. Trenching of the seeding spots showed that root competition during the early stage of regeneration was not of decissive importance. It seemed to, however, improve the preservation of the seedlings later. It is common that it can take long before the seeds germinate, and during that time the number of viable seeds decrease strongly.

Also, the seedling stock quickly began to decrease in number after germination, especially during the first growing season and the following winter. The decrease was larger in intact vegetation than on mineral soil or in the humus layer. The emerging seedlings were destroyed by drought very easily, but their tolerance to drought improved later on.

The PDF includes a summary in English.

  • Yli-Vakkuri, E-mail: py@mm.unknown (email)
article id 7130, category Article
Paavo Yli-Vakkuri. (1961). Tutkimuksia männyn kylvöalojen metsittymisvaiheesta. Silva Fennica vol. 74 no. 3 article id 7130. https://doi.org/10.14214/aff.7130
English title: Studies on the development of young sown pine stands in Central Finland.
Original keywords: kylvö; mänty; taimikko; uudistaminen
English keywords: Pinus sylvestris; regeneration; Scots pine; seedling stand; sowing; seeding
Abstract | View details | Full text in PDF | Author Info

In this paper the development of sown Scots pine (Pinus sylvestris L.) seedling stands into forests is studied. The material was collected in stands sown in 1930–1940 in private forests in the Forestry Board districts of Central and Northern Ostrobothnia. The 119 areas, including both burned and other areas, were studied in 1955.

Most seedings had been carried out on relatively poor soils, mostly representing Vaccinium and Calluna type forests. 71% of the areas consisted of large forest fires, mostly from 1933. The most burned areas did not have seed producing trees nearby. The other sown areas were in general small, 1–2 ha, and near forests capable of producing seeds. The species of previous tree generation, in the older areas mostly pine and in the younger areas Norway spruce, affected tree species composition of the new tree generation.

Over 90% of the burned areas were in silviculturally good or satisfactory condition, while the main part of the other sown stands was in fair or poor condition. Weeding and thinning had been done only in the oldest stands. Most stands had been left untended. Natural new trees often competed with the sown pines, and cull-trees and border forest increased natural regeneration in the areas. In Calluna type the poor soil limited regeneration and growth of broadleaf trees. The worst competitors were naturally regenerated pine seedlings both on Calluna and Vaccinium type. On Vaccinium type also birch and sometimes also aspen (Populus tremula L.) competed with sown pine. On better sites and paludified areas competition by broadleaf trees increased. The rhythm of development of broadleaved trees is so different from pine that only those broadleaved trees that are formed in the stand when the pine seedlings are larger can develop harmoniously with pine. Due to the harmful competition, the seedling stands should be tended early on. In addition, it may be advisable to abandon the practise to leave trees on sowing areas.

The PDF includes a summary in Finnish.

  • Yli-Vakkuri, E-mail: py@mm.unknown (email)
article id 7129, category Article
Erkki K. Kalela. (1961). Natural regeneration of agricultural lands in so-called Porkkala-area of leased land. Silva Fennica vol. 74 no. 2 article id 7129. https://doi.org/10.14214/aff.7129
Keywords: natural regeneration; agricultural land; dominant species
Abstract | View details | Full text in PDF | Author Info

The size of the field and the ditching on it as well as the condition of the field at the time of the surrender of Porkkala area (from Finland to Russia) play an important role on how far the natural regeneration of the fields has progressed.

Larger open fields have naturally regenerated only on sides where the surrounding forest can spread the seeds or the thicket of saplings has reached, whereas small parcels of fields have normally been fully forested. Most important species have been e.g. silver birch and pubescent birch, grey and common alders and European aspen as well as pine and spruce. The broad leaved species are dominant.     

The PDF contains a summary in Finnish.

 

  • Kalela, E-mail: ek@mm.unknown (email)
article id 7128, category Article
Paavo Yli-Vakkuri. (1961). Emergence and initial development of tree seedlings on burnt-over forest land. Silva Fennica vol. 74 no. 1 article id 7128. https://doi.org/10.14214/aff.7128
Keywords: Pinus sylvestris; Norway spruce; Picea abies; germination; regeneration; Scots pine; seeds; prescribed burning; sowing; seeding
Abstract | View details | Full text in PDF | Author Info

Prescribed burning has been used in regeneration areas in Finland as a method to treat the humus layer and creating more favourable chemical, physical and biological conditions for the seedlings. At the same time, fire clears away seedlings and shoots of unwanted trees and other vegetation. Direct sowing or planting, mostly Scots pine (Pinus sylvestris L.), seldom natural regeneration, is used. In this paper, the initial stages of the formation of a new tree generation of Scots pine and Norway spruce (Picea abies (L.) Karst.) on prescribed burned areas is studied in Central Finland in 1956–1960.

The burned area remains almost without vegetation for about two growing seasons. Conditions on a burned area which has not been tiled are very unfavourable for germination of seeds of coniferous and deciduous trees. On the other hand, shoots of deciduous trees occur soon after burning. Conditions for regeneration were found to be better 3–5 years after burning. Removal of humus layer in spots improved regeneration. However, the patches facilitated also natural regeneration of Norway spruce and especially birch (Betula sp.), which compete with Scots pine seedlings.

Continuous rainy periods improved the germination of Scots pine and Norway spruce seeds sown on the humus layer. Pine and spruce developed more rapidly on the exposed soil, however, young seedlings were easily destroyed. Seed eaters destroyed the pine and spruce seeds sown on the humus layer of newly burned areas completely or almost completely. The viability of pine seeds sown on the burned humus layer did not decrease for three weeks, but the viability greatly weakened after six or more weeks. Spruce seeds lost their viability faster than pine seeds.

The PDF includes a summary in Finnish.

  • Yli-Vakkuri, E-mail: py@mm.unknown (email)
article id 7478, category Article
Paavo Yli-Vakkuri. (1958). Tutkimuksia ojitettujen turvemaiden kulotuksesta. Silva Fennica vol. 67 no. 4 article id 7478. https://doi.org/10.14214/aff.7478
English title: Studies on prescribed burning of drained peatlands.
Original keywords: mänty; luontainen uudistaminen; kulotus; suot; pintakasvillisuus; turvemaat; uudistaminen; turvekankaat; vesaikko
English keywords: Pinus sylvestris; regeneration; natural regeneration; drained peatlands; Scots pine; coppice; peatlands; ground vegetation; prescribed burning
Abstract | View details | Full text in PDF | Author Info

Prescribed burning has been used to treat the mineral soil sites, but the method has been little used in drained peatlands. The course and methods of prescribed burning in drained peatlands, and the effect of burning on sprouting of broadleaved trees, growth of ground vegetation and regeneration of Scots pine (Pinus sylvestris L.) by sowing was studied in drained pine bogs in Southern Finland. The top layer of the peat was mostly Sphagnum peat. The material included a prescribed burned 12 ha drained peatland area in Tuomarniemi district, in addition to which ten previously burned areas were investigated.

The burning had succeeded mostly well, but also unsuccessfully burned sites were observed. Estinguishing of the fire was easy, and no peat fires occurred. The fire burned only the logging residue, ground vegetation and the dry top layer of the peat. The roots of brushwood and grasses survived in the peat that insulated the top layer from the heat. For instance, the abundance of cloudberry (Rubus chamaemorus L.) increased after the fire. Similarly, burning did not affect sprouting of the stumps of downy birch (Betula pubescens Ehrh.). It cannot thus be used as a method to restrict the growth of coppice in regenerated areas. The seeds of Scots pine germinated well on the burned surface. 46% of the seeds developed to seedlings on sphagnum-shrub vegetation and 16% in feathermoss-shrub vegetation.

The PDF includes a summary in German.

  • Yli-Vakkuri, E-mail: py@mm.unknown (email)
article id 7472, category Article
Jaakko Lehto. (1956). Tutkimuksia männyn luontaisesta uudistumisesta Etelä-Suomen kangasmailla. Silva Fennica vol. 66 no. 2 article id 7472. https://doi.org/10.14214/aff.7472
English title: Studies on the natural regeneration of Scots pine on the upland soils of Southern Finland.
Original keywords: mänty; luontainen uudistaminen; taimettuminen; uudistuminen
English keywords: Pinus sylvestris; regeneration; natural regeneration; Scots pine; stocking with seedlings
Abstract | View details | Full text in PDF | Author Info

Natural regeneration of Scots pine (Pinus sylvestris L.) by leaving a seed tree stand on a cutting area has long been the most popular regeneration method in Finland. Results of the method have, however, been unsatisfactory. The aim of the investigation was to study the basic problems of natural regeneration of Scots pine. Regeneration success was studied in 144 sample plots in pine stands at different stages of regeneration in Southern Finland. In addition, the data included information of 42 previously investigated areas.

According to the results, Scots pine can be successfully regenerated naturally on sandy and gravelly soils in Southern Finland. Preparing the ground surface by breaking or burning considerably facilitates the establishment of a seedling stand. The number of seedlings was considerably lower in the ground vegetation than in the mineral soil. Considering growth of the seedlings, root competition of the mother trees was heavy in dense stands, but insignificant in thin stands. The stand density did not affect germination of the seeds. In regeneration areas proper, where the density of mother trees usually is under 50 per hectare, there was in average 4,700 seedlings per hectare in Calluna type forests and 5,200 in Vaccinium type forests.

The PDF includes a summary in English.

  • Lehto, E-mail: jl@mm.unknown (email)
article id 7453, category Article
Gustaf Sirén. (1955). The development of spruce forest on raw humus sites in Northern Finland and its ecology. Silva Fennica vol. 62 no. 4 article id 7453. https://doi.org/10.14214/aff.7453
Keywords: Norway spruce; regeneration; succession; northern Finland
Abstract | View details | Full text in PDF | Author Info

According to the National Forest Inventory of Finland, the age structure of forests in Northern Finland require large-scale fellings. One of the problems is regeneration of low-productive Norway spruce (Picea abies (L.) Karst.) forests in the area. The objectives of this investigation were to study the natural development of Hylocomnium-Myrtillus (HMT) type Norway spruce forests in Northern Finland and the ecological causes of the development. A total of 83 sample plots were studied in 1950-52. The stands were chosen to have a total effective temperature during growing season between 600-800 ºC.

According to the results, the HMT type forests represent secondary developmental stages of the northern Myrtillus type, characterized by tree stands of poor quality. By returning the habitats that have reached their secondary stages to their primary stage, e.g. by means of fires, the potential site quality is restored. The climax theory is inapplicable to the Northern Finland’s spruce stands on fresh site types. The natural development of HMT follows not the climax theory, i.e. a return in each succession to the same type stage expressing the same site quality. One reason is accumulation of the thick humus layer, caused by incomplete decomposition of litter. The humus binds nutrients inaccessible to the trees, and the substratum becomes cold, more acid, and moist.

These forests should be managed by clear-cutting, burning-over and artificial regeneration. When thinning the stands, at least a slight mixture of deciduous trees must be left in the stand.

The article is divided into two separate PDFs. The second PDF is in a supplementary file and includes a summary in Finnish.

  • Sirén, E-mail: gs@mm.unknown (email)
article id 7440, category Article
Erkki K. Kalela. (1954). Mäntysiemenpuiden ja -puustojen juurisuhteista. Silva Fennica vol. 61 no. 28 article id 7440. https://doi.org/10.14214/aff.7440
English title: Root systems of Scots pine seed trees and stands.
Original keywords: mänty; taimettuminen; juuristo; uudistuminen; siemenpuut; juurikilpailu
English keywords: Pinus sylvestris; regeneration; natural regeneration; Scots pine; seedlings; seed trees; root system; root competition
Abstract | View details | Full text in PDF | Author Info

Root systems of a Scots pine (Pinus sylvestris L.) stands of seed trees on a Vaccinium sites in Southern Finland were studied by taking soil samples around the seed trees. The results show that root system of an old Scots pine spreads relatively evenly around the tree up to at least 10 meters from the stem. The densest part of the root system is near the stem, which part is often acentric. This is probably due to root competition in the early stages of growth of the tree.

Root systems of the seed trees affect stocking of the site with seedlings and the growth of the seedlings. The root competition can cause, for instance, uneven grouping of the seedlings. It seems that the largest trees of a stand have the most even root system. It is therefore recommended to choose the strongest trees of the stand as seed trees, to ensure even distribution of seedlings.

The Acta Forestalia Fennica issue 61 was published in honour of professor Eino Saari’s 60th birthday.

The PDF includes a summary in German.

  • Kalela, E-mail: ek@mm.unknown (email)
article id 7439, category Article
Leo Heikurainen. (1954). Rämemänniköiden uudistamisesta paljaaksihakkausta käyttäen. Silva Fennica vol. 61 no. 27 article id 7439. https://doi.org/10.14214/aff.7439
English title: Regeneration of Scots pine stands of pine swamps through clear cutting.
Original keywords: männikkö; luontainen uudistuminen; mänty; suot; alikasvos; uudistaminen; räme; turvekangas; avohakkuu
English keywords: Pinus sylvestris; regeneration; natural regeneration; drained peatlands; Scots pine; peatlands; seedlings; fellings; undergrowth; pine swamps; clear cutting
Abstract | View details | Full text in PDF | Author Info

Pine swamps are easily regenerated by natural regeneration of Scots pine (Pinus sylvestris L.). Usually seeding felling is used, but also strip system or clear cutting and regeneration along stand edge has been suggested. This article discusses the regeneration by clear cutting and sparing the existing undergrowth. The article focuses on pine swamps to be drained and the ones in natural state.

Pine swamps in natural state usually have plenty of trees of smaller diameter classes, that can be trusted to form the future tree generation after the felling. This shortens the rotation by 20-30 years. The undergrowth has been shown to recover quickly. The method suits for regeneration of drained peatlands but could fit also for regeneration of pine swamps in natural state.

The seedlings in the pine swamps are mainly 1-5 years old, and the stock is changing. It seems that larger trees produce a wider selection of age groups, but the seedlings survive longer under smaller mother trees. Part of the younger generations of seedlings seem to be destroyed when the peatland is drained. Further studies are needed to investigate how the draining and felling are to be performed to spare the young seedlings.

The Acta Forestalia Fennica issue 61 was published in honour of professor Eino Saari’s 60th birthday.

The PDF includes a summary in German.

  • Heikurainen, E-mail: lh@mm.unknown (email)
article id 7402, category Article
Gustaf Sirén. (1950). Alikasvoskuusten biologiaa. Silva Fennica vol. 58 no. 2 article id 7402. https://doi.org/10.14214/aff.7402
English title: On the biology of undergrown Norway spruce.
Original keywords: kuusi; alikasvos; juuristo; uudistuminen; iänmääritys; latvus; toipumiskyky
English keywords: Norway spruce; Picea abies; regeneration; natural regeneration; age; crown; undergrowth; root system
Abstract | View details | Full text in PDF | Author Info

Norway spruce (Picea abies (L.) Karst.) invading sites is common in Finland. The species tends to establish itself as undergrowth, and takes over when it gets space to grow. To determine whether the undergrowth is suitable as the new generation requires knowledge on the biology of spruce undergrowth. One of the issues is determining the age of the stunted trees. In this investigation, 100 undergrown spruce trees, their crown and their root systems were studied. A method was developed to determine the age of the trees.

The root system of all trees in Vaccinium sites and of stunted trees in Myrtillius sites were superficial. The root systems of older spruces were purely of adventitious origin. The longer the period of stunting growth, the younger is the root system. In addition to acropetal and general adventitious ramification there is often adventitious branching of the roots of pathological causes. Mortality among the long roots is frequent.

A stunted tree has not the same ability as a viable tree to make use of already existing branches for building assimilating surface. When comparing trees with equally large assimilating surface, a stunted tree had greater sum of roots compared to a viable tree. The root system of a stunted undergrown spruce was very superficial compared with the other trees.

The PDF includes a summary in English.

  • Sirén, E-mail: gs@mm.unknown (email)
article id 7377, category Article
S. E. Multamäki. (1942). Kuusen taimien paleltuminen ja sen vaikutus ojitettujen soiden metsittymiseen. Silva Fennica vol. 51 no. 1 article id 7377. https://doi.org/10.14214/aff.7377
English title: Frost injuries of Norway spruce seedlings and their effect on afforestation of drained peatlands .
Original keywords: kuusi; ojitus; uudistuminen; kylmänkestävyys; turvekangas; pakkasvaurio; keväthalla; ojitettu suo; suojametsä
English keywords: Norway spruce; Picea abies; regeneration; drained peatlands; frost damage; peatlands; frost resistance; spring frost; prorective forest
Abstract | View details | Full text in PDF | Author Info

The aim of the investigation was to study natural regeneration of Norway spruce (Picea abies (L.) Karst.) in drained peatlands and frost injuries in seedlings, and to compare microclimates of the regeneration areas. The experiments included peatlands in Satakunta in Western Finland. Restocking of the areas with seedlings and their survival was followed in 1935-40 at sample plots that were mainly 1 are large.

Susceptibility to freezing was shown to be dependent on the stage of development of the shoots. Shoots that have just begun to grow contain little water, and withstand better freezing temperatures than shoots in later stages of growth. Damages to the seedlings were observed when the temperatures decreased to -2.8–-4.3 °C. The most severe damage to a seedling was caused by the death of the leading shoot by spring frost.

Norway spruce regenerates easily on moist peatlands, but peatlands with dry surface tend to have little or no seedlings. The species regenerated better in marshy sites than correspondingly fertile mineral soil sites. However, it needs shelter to avoid frost damage. On clear cut spruce swamp the undergrowth spruce seedlings that were left in the site got severe frost damage. If the site had birch (Betula sp.) coppice or undergrowth, spruce seedlings survived in their shelter depending on the height and density of the birch trees. To be effective, the protective forest should have relatively even crown cover. Young spruce seedlings could grow well even under relatively dense birch stand.

The PDF includes a summary in German.

  • Multamäki, E-mail: sm@mm.unknown (email)
article id 7356, category Article
Peitsa Mikola. (1942). Koivun vesomisesta ja sen metsänhoidollisesta merkityksestä. Silva Fennica vol. 50 no. 3 article id 7356. https://doi.org/10.14214/aff.7356
English title: Silvicultural usefulness of sprouting of birch.
Original keywords: Etelä-Suomi; Pohjois-Suomi; koivu; uudistuminen; rauduskoivu; hieskoivu; vesominen; vesatalous; juurivesa
English keywords: birch; Betula pendula; coppicing; regeneration; silver birch; sprouting; downy birch; decay; Betula verrucosa
Abstract | View details | Full text in PDF | Author Info

Birches’ (Betula sp.) ability to grow sprouts is low. The stump grows root collar and stump shoots, but the stump shoots are not proper stump shoots that will grow from the space between wood and bark. The buds are situated very low in the base, even under the ground. In this study, no actual root shoots could be found. Also the bushy alpine birches seem to be formed from stump and root collar shoots.

In Southern Finland silver birch (Betula pendula Roth) is more common than downy birch (Betula pubescens Ehrh.) in dry upland forest sites, while downy birch is common in fresh mineral soil forests and peatlands. In Northern Finland downy birch is the dominant birch species. Of the two species downy birch has markedly better capacity to form stump and root collar shoots both in Northern and Southern Finland. In general, birches grow sprouts much more strongly in Northern Finland.

Growth of the shoots is fastest during the first year after the felling of the parent tree and slows down gradually. The stump shoots may get separated from the stump when the stump decays, and the decay may also spread to the shoots. It is common that the shoots have no own roots, and die along with the stump. The shoots may have own root system or use roots of the parent tree that have stayed alive, in the latter case decay spreads almost always from the stump to the shoot. Whether the tree was felled with axe or saw had no effect on sprouting, probably because the sprouting buds are situated in the base of the tree. The larger stumps had usually fewer sprouts than smaller stumps. The fertility of the site seemed to have little effect on sprouting, but more moist sites formed more sprouts.

Forest regeneration using sprouts may be possible in peatlands for firewood production. on mineral soil sites birch does not suit for coppicing. The proportion of trees originating from sprouts decreases strongly by the time. Consequently, in Southern Finland sprouts have little effect on regeneration of birch. In Northern Finland sprouting is the most important way of regeneration.

The PDF includes a summary in German.

  • Mikola, E-mail: pm@mm.unknown (email)
article id 7351, category Article
Esko Kangas. (1940). Tuloksia Pohjankankaan ja Hämeenkankaan metsänviljelyksistä. Silva Fennica vol. 49 no. 4 article id 7351. https://doi.org/10.14214/aff.7351
English title: Studies on artificial regeneration in Pohjankangas and Hämeenkangas in Southern Finland.
Original keywords: kylvö; mänty; metsäpalo; metsänuudistaminen; taimikko; kulo; hyönteistuho; sienituho
English keywords: Pinus sylvestris; regeneration; Scots pine; seedling damage; ; seedling stands; insect damage
Abstract | View details | Full text in PDF | Author Info

The regeneration of forests in Hämeenkangas area in Southern Finland has been difficult due to various damages from the middle of the 1800s. Few seed trees were left in the area, and artificial regeneration has been used since 1880s. The area became an experimental area of the Forest Research Institute in 1924. The aim of the study was to survey the area before it was transferred to the Finnish Defense Forces.

The original Scots pine (Pinus sylvestris L.) forest of the esker area suffered from many forest fires. The total area is 13,000-14,000 ha, of which the experimental forests of Forest Research Institute cover 6,000 ha. The area is dry upland forest, and drought affects the survival of germlings. Soil frost is a major cause of loss of young seedlings. Sowing method affects the early development of the seedlings. Band sowing proved to be the best method regarding the soil frost. A total of 39 different harmful insect species, 8 pathogen species and 7 other causes of damages have been detected in the area.

The development of seedling stands follow a certain pattern, reported also in other studies. Many of the pine seedling stands develop well until they reach a certain height. After that seedlings begin to suffer from damages, but after reaching another stage develop normally. The damages affect the height growth of the seedlings. Some common damages are caused by Pissoides weevils, needle damages caused by certain beetles, shoot damages by Evetria resinella, and pine blister rust (Peridermium pini and Cronartium flaccidum).

The PDF includes a summary in German
  • Kangas, E-mail: ek@mm.unknown (email)
article id 7350, category Article
Ilmari Paasio. (1941). Plant sociological principles of open bog types. Silva Fennica vol. 49 no. 3 article id 7350. https://doi.org/10.14214/aff.7350
Keywords: population; classification; treeless bog; open bog; plant sociology; plant stand; phytogenesis
Abstract | View details | Full text in PDF | Author Info

The plant populations of Finnish open bogs are typically formed of two layers. The layers normally consist of one or rarely two species. The structure of plant populations in open bogs is a consequence of the development where determining factors are different site requirements of the species, and the differences in the biotic vitality and capacity for reproduction.

Phytogenesis should be taken as a basic unit for describing the plant societies or vegetation of treeless bogs. However, acknowledging the sub-populations may be of advantage for describing the ecological, genetic and regional characters of open bogs.

The basic classification of open bogs must be done based on the ground layer. The more detailed classification follows mostly based on field layer, partly also based on the ground layer.

The PDF contains a summary in Finnish. 

  • Paasio, E-mail: ip@mm.unknown (email)
article id 7300, category Article
Olli Heikinheimo. (1934). Metsänviljelysmenetelmiä koskevista tutkimuksista. Silva Fennica vol. 40 no. 23 article id 7300. https://doi.org/10.14214/aff.7300
English title: Research on forest regeneration methods.
Original keywords: metsäntutkimus; metsänviljely; metsän uudistaminen; viljelykoe
English keywords: articifial regeneration; forest researc
Abstract | View details | Full text in PDF | Author Info

The article is a review on the research about forest regeneration in Finland, executed in the Forest Research Institute. The climate affects the results of different regeneration methods. Thus, sample plots have been established in different parts of the country. In 1933 there was a total of 386 sample plots around the country. To compare the effect of weather conditions in the regeneration, sample plots have been established in successive years. Other conditions that affect forest regeneration are soil, forest site type, tree species, time from the felling of the stand, and tending of the seedling stand. Of the 386 sample plots 245 were planting experiments and 141 sowing experiments.

The PDF includes a summary in German.

  • Heikinheimo, E-mail: oh@mm.unknown (email)
article id 7294, category Article
Martti Hertz. (1934). Tutkimuksia karjan vaikutuksesta hakkausalojen kasvillisuuteen. Silva Fennica vol. 40 no. 17 article id 7294. https://doi.org/10.14214/aff.7294
English title: Studies on influence of grazing on the vegetation of cutting areas.
Original keywords: luontainen uudistuminen; kuusi; karja; laidunnus; mesälaidun; hakkuualue
English keywords: Picea abies; natural regeneration; spruce; cattle; forest pasture
Abstract | View details | Full text in PDF | Author Info

The aim of the study was to determine the effect of grazing of the cutting areas to the ground vegetation and regeneration of spruce. The cutting areas could be divided into two kinds of areas based on the vegetation. Hillocks were drier and poorer than other parts of the cutting areas. Their vegetation did suffer less from grazing than the other parts of the cutting areas. The shade-loving plant species decreased, but as the poorer sites have less edible plants, cattle caused less damage than in the better sites. The even spaces between the hillocks had both positive and negative changes. Cattle transport seeds, tile and fertilize the soil, promote paludification, and decrease competition by the primary species like large grasses. This is beneficial to new species. Grazing is directed to large grass species like Calamagrostis. Those species that cattle reject, become more abundant. Stamping damages especially shallow rooted species and perennial species, like Norway spruce (Picea abies (L.) H. Karst.). Larger tree seedlings may get injuries in the stem.

The PDF includes a summary in German.

  • Hertz, E-mail: mh@mm.unknown (email)
article id 7256, category Article
V. Pöntynen. (1929). Tutkimuksia kuusen esiintymisestä alikasvoksina Raja-Karjalan valtionmailla. Silva Fennica vol. 35 no. 1 article id 7256. https://doi.org/10.14214/aff.7256
English title: Studies on Norway spruce undergrowth in state forests in Karelia.
Original keywords: kuusi; Picea abies; valtion metsät; metsätyyppi; alikasvos; Karjala
English keywords: natural regeneration; forest site type
Abstract | View details | Full text in PDF | Author Info

The abundance of Norway spruce (Picae abies (L.) H. Karst.) undergrowth is common for the state forests in Karelia near the Russian border, in Finland. In the survey, the occurrence of the undergrowth was studied. The article includes a review on the ownership of the forest, forest soils in the area, and the state of forests in the area. Scots pine (Pinus sylvestris L.) is the dominative species in 67%, Norway spruce in 27% and Betula sp. 6% of the state forests. Only 2% of the forests are 1‒20 years of age. Stands in the age group of 61‒80 years are the most common (25%). Norway spruce undergrowth is most abundant in the municipality of Salmi. The forests are typically moist forest site types or grass-herb site types. If the stands are allowed to develop naturally, even the Vaccinium sites become Norway spruce dominated. Spruce undergrowth is formed seldom under a spruce forest unless the stand is thin or has openings. Because Norway spruce is often rare in the mineral soil sites, the undergrowth is often regenerated from seeds that spread from spruce swamps. Earlier practiced shifting cultivation and its frequent fires prevented regeneration of spruce undergrowth. Similarly, the common felling method used, clear felling in strips, does not promote spruce undergrowth. Consequently, their occurrence is likely to decrease in the future.

The PDF includes a summary in German.

  • Pöntynen, E-mail: vp@mm.unknown (email)
article id 7232, category Article
Edvard Wibeck. (1929). Discussions on the choice of cultivation method in coniferous forests in Sweden. Silva Fennica vol. 34 no. 19 article id 7232. https://doi.org/10.14214/aff.7232
Keywords: regeneration; afforestation; cultivation; Sweden; forest type
Abstract | View details | Full text in PDF | Author Info

In Sweden lot of state owned forests or earlier mining districts or districts of ironworks have been afforested during the last decade. The amount of afforested areas sinks from south to north. Afforestation has taken place also in privately owned forests.

The article discusses the common economic questions related to afforestation work and the biological viewpoints related to it. The best cultivation methods are presented for several common forest types, such as herb-rich forest types, moss-grown forests types, swampy forests with heavy raw humus and barren pine forest sites (lichen type).

The volume 34 of Acta Forestalia Fennica is a jubileum publication of professor Aimo Kaarlo Cajander.

  • Wibeck, E-mail: ew@mm.unknown (email)
article id 7228, category Article
Agnar Barth. (1929). Skjermforyngelsen i produksjonsökonomisk belysning. Silva Fennica vol. 34 no. 15 article id 7228. https://doi.org/10.14214/aff.7228
English title: Natural regeneration from the production economic point of view.
Keywords: yield; natural regeneration; Norway; volume growth; seed tree stands; economic result
Abstract | View details | Full text in PDF | Author Info

Natural regeneration has been studied so far mostly on the perspective of regeneration, while the production capacity of the seed crop stand has been of little interest in the earlier studies. This paper studied volume growth of the seed trees and the economic impact of this regeneration method both in a literature review and measuring sample stands consisting of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) H. Karst.) dominated stands and mixed forests in Norway.

It is concluded that in the most cases the seed tree stands give better pulpwood and timber yield than the stands in the average. The reason might be that the elite stems can better use their production capacity, and, thus, reach bigger and more valuable dimensions. Even though the growing stock diminishes in the seeding felling, the growth of the stand does not decrease or decreases only little. At the same time, the quality growth improves. In addition, the new tree regeneration is achieved usually without extra costs, and there is no unproductive time period in the stand.

  • Barth, E-mail: ab@mm.unknown (email)
article id 7227, category Article
D. Fehér. (1929). Biology of forest soil and its physiological meaning in the life of forest. Silva Fennica vol. 34 no. 14 article id 7227. https://doi.org/10.14214/aff.7227
Keywords: forest management; regeneration; forest soil; soil physiology
Abstract | View details | Full text in PDF | Author Info

The article presents three studies about different aspects of the bio-chemical functions of forest soil. The three studies are i) research on microflora and microfauna of forest soils; ii) study on carbon nutrition of forests in relation to microbial functions of soil and effecting site factors and iii) study on nitrogen metabolism of forest soil. The results of the studies are summarized by every study.

The article discusses the meaning of the results for forest management in practice. The good biological and physiological condition of forest soil is important for the forest growth and it needs to be taken care in regeneration and other forest management. The natural regeneration seems to be better for soil functions. Favoring broadleaved trees as undergrowth enhances the biological processes of forest soil.   

The volume 34 of Acta Forestalia Fennica is a jubileum publication of professor Aimo Kaarlo Cajander.

  • Fehér, E-mail: df@mm.unknown (email)
article id 7195, category Article
Martti Hertz. (1925). Niinipuun uudistumisesta Suomessa. Silva Fennica vol. 29 no. 5 article id 7195. https://doi.org/10.14214/aff.7195
English title: Regeneration of small leaved lime in Finland.
Original keywords: luontainen uudistuminen; siementuotanto; levinneisyys; lehmus; pohjoisraja; niinipuu; Tilia cordata
English keywords: regeneration; natural regeneration; distribution; seed production; northern limit
Abstract | View details | Full text in PDF | Author Info

The northern range of small leaved lime (Tilia cordata Mill.) in Finland has remained the same since the end of 1800s, but according to the studies of subfossiles of lime found in peatlands, the northern limit has once been higher than at the present. The northernmost natural specimens of the species in Reisjärvi do not produce seed, and are therefore probably a relict. The article includes a review on the distribution of the species in Finland and its capacity to regenerate. The natural regeneration of lime in Finland is at present very rare. The species has lost its ability to sexual reproduction, but reproduces readily vegetatively.

The PDF includes a summary in German.

  • Hertz, E-mail: mh@mm.unknown (email)
article id 7036, category Article
I. Lassila. (1920). Tutkimuksia mäntymetsien synnystä ja kehityksestä pohjoisen napapiirin pohjoispuolella. Silva Fennica vol. 14 no. 3 article id 7036. https://doi.org/10.14214/aff.7036
English title: Studies on the regeneration and development of Scots pine forests north of the arctic circle.
Original keywords: luontainen uudistuminen; mänty; Pinus sylvestris; siemensato; käpy
English keywords: natural regeneration; cone; seed yield
Abstract | View details | Full text in PDF | Author Info

The frequency of years when Scots pine (Pinus sylvestris L.) produces cones and seeds affects its reproduction in the north. The study area covered most of the pine lands in Northern Finland. Scots pine seems to be able to produce cones relatively often in the north. The amount of seeds produced in one year was, however, not sufficient to produce a dense seedling stand. Thus, the natural pine stands contain usually trees in different ages-classes, which have germinated in different years. The cone production is highest in 150-170 years old trees. Pine also needs warm summers to produce viable seeds. Brush fires avail the growth of seedlings, because they clear of ground vegetation that hinders germination of seeds. The seedlings need also moisture to survive; good regeneration years have often had rainy summers.

The PDF includes a summary in German.

  • Lassila, E-mail: il@mm.unknown (email)
article id 7035, category Article
V. T. Aaltonen. (1920). Use of water of the trees and the moisture conditions in the soil. Silva Fennica vol. 14 no. 2 article id 7035. https://doi.org/10.14214/aff.7035
Keywords: regeneration; forest soil; water consumption; use of water; moisture conditions; heathy forests type
Abstract | View details | Full text in PDF | Author Info

The data has been collected in the dry heathy forests in Harjavalta commune in south-west Finland and in Sodankylä commune in Lapland in summer time 1918 and 1919. The aim to the study was to find out how does the water consumption of the trees affect the moisture conditions of the soil and how they are linked to the regeneration of the forest.

Trees in one age class have a certain spatial distribution: the greater the distance between trees the older the trees and the smaller the distance, the younger the trees. This seem to rather be due to the development of the root system and the nutrition intake of a tree than the competition for light. The moisture content of the upper soil layers is higher in the open areas than in the closed canopy stands. Hence there are more seedlings growing in open areas.   However, it is not clear whether the results apply for other forest site types as well, and more research is needed.     

  • Aaltonen, E-mail: va@mm.unknown (email)
article id 7024, category Article
August Renvall. (1919). Suojametsäkysymyksestä IV. Poronlaidunnan järjestely suojametsäalueella. Silva Fennica vol. 11 no. 4 article id 7024. https://doi.org/10.14214/aff.7024
English title: Protection forests IV.
Original keywords: luontainen uudistuminen; mänty; Pinus sylvestris; metsäraja; poronhoito
English keywords: natural regeneration; Scots pine; grazing; timber line; reindeer
Abstract | View details | Full text in PDF | Author Info

This fourth part of the six-article series about protection forest in the Northern Finland is a proposal for organizing reindeer grazing to enable scots pine (Pinus sylvestris L.) regeneration near the timber line. Protection forests in the Northern Finland cover third of the natural pastures of reindeer in the area. In these areas reindeer grazing can harm the young pine seedlings. The proposal suggests temporary restrictions in grazing in the coniferous forests. Also, in the northernmost parts of Lapland the pine timberline area would be used only as winter pastures. Regional limits should be set for the number of reindeer. Also the ownership of reindeer herds and herding coopearatives included problematic issues that should be solved.

The article is divided in six parts. A German summary is in a separate PDF.

  • Renvall, E-mail: ar@mm.unknown (email)
article id 7022, category Article
August Renvall. (1919). Suojametsäkysymyksestä II. Suojametsäjärjestelmän tarkoitusperät ja edellytykset. Silva Fennica vol. 11 no. 2-3 article id 7022. https://doi.org/10.14214/aff.7022
English title: Protection forests II.
Original keywords: luontainen uudistuminen; mänty; Pinus sylvestris; metsäraja
English keywords: natural regeneration; Scots pine; timber line
Abstract | View details | Full text in PDF | Author Info

The second part of the six-article series about protection forest in Northern Finland outlines the principles of protecting Scots pine (Pinus sylvestris L.) timber line. The protected areas should include at least all of the viable pine stands and those birch (Betula sp.) lands that are necessary to safeguard fuel wood supply in the pine forest zone. Outside the actual pine forest zone also those birch stands that are near populated regions should be protected to secure sustainable fuel wood supply. Third, south and down of the pine forest zone should be protected pine forest stands and adjacent open fell areas where forests are especially vulnerable. A new protection forest commission should be established to execute the protection forest legislation.

The article is divided in six parts. The parts II and III of the article series are included in the same PDF. A German summary is in a separate PDF.

  • Renvall, E-mail: ar@mm.unknown (email)
article id 7088, category Article
Uunio Saalas. (1919). Kaarnakuoriaisista ja niiden aiheuttamista vahingoista Suomen metsissä. Silva Fennica vol. 10 no. 1 article id 7088. https://doi.org/10.14214/aff.7088
English title: Bark beetles and insect damages in Finnish forest.
Original keywords: kaarnakuoriaiset; hyönteistuho; entomologia
English keywords: forest pests; Tomicus piniperda; insect damage; Pityogenes chalcographus; Ips typographus; entomology; Scolytus Ratzburgi
Abstract | View details | Full text in PDF | Author Info

The aim of the study was to investigate the abundance of bark beetle species and their damage in Finland. The bark beetle populations were studied in several areas in Finland, both in sites with known beetle damage and without. Two-meter wide lines were measured in the sample plots, where all trees were studied for bark beetle damage in the stem and the crown of the trees. The abundance of bark beetle species and the beetle damages in 25 study areas, and 52 different species are discussed in detail.

Divided by their lifestyle, the most important groups of bark beetles are the pine shoot beetles (Tomicus sp.), beetles reproducing in Scots pine (Pinus sylvestris L.) stems and branches under the bark, beetles reproducing in Norway spruce (Picea abies (L.) H. Karst.) stems and branches under the bark, beetles living in the other coniferous trees, beetles reproducing in roots, beetles reproducing under the Betula sp. bark, beetles reproducing in other deciduous trees and beetles reproducing inside of the stems. Of individual species, Blastophagus piniperda (now Tomicus piniperda) and B. minor cause worst damage to pine and Ips typographus (L.) and Pityogenes chalcographus (L.) to spruce. Serious damage is caused also by Xyloterus lineatus to coniferous trees, Polygraphus polygraphus and P. subopacus to Norway spruce and Scolytus Ratzburgi to Betula sp.

The PDF includes a summary in German.
  • Saalas, E-mail: us@mm.unknown (email)
article id 7018, category Article
Olli Heikinheimo. (1917). Metsänhävityksen ja polton vaikutuksesta metsämaahan. Silva Fennica vol. 8 no. 3 article id 7018. https://doi.org/10.14214/aff.7018
English title: Effects of forest devastation and shifting cultivation on forest soil.
Original keywords: maaperä; metsittyminen; kaskeaminen
English keywords: regeneration; soil; shifting cultivation
Abstract | View details | Full text in PDF | Author Info

Soil of open and forest covered lands have different properties. Forest devastation has negative effect on forest soil, for instance, through leaching of nutrients. There are different views on how forest fire influences chemical properties of the soil. Burning of forests can have both positive and negative consequences.

The article publishes partial results from a study that aimed to find out how shifting cultivation affects the soil, and how it may change natural regeneration and forest growth of the stands. Soil samples were collected from four burnt-over sites. Of the samples, size classes of the soil particles, volatile solids and nutrient contents were determined. According to the results, the burning increased soluble nutrients, but the effect was not long-lasting. However, more sample sites would be needed to give definitive results of the differences.

  • Heikinheimo, E-mail: oh@mm.unknown (email)
article id 7013, category Article
Lauri Ilvessalo. (1917). Tutkimuksia mäntymetsien uudistumisvuosista Etelä- ja Keski-Suomessa. Silva Fennica vol. 6 no. 2 article id 7013. https://doi.org/10.14214/aff.7013
English title: Studies on the regeneration years of Scots pine forests in Southern and Central Finland.
Original keywords: luontainen uudistuminen; uudistuminen; siemensato; siemenvuosi
English keywords: natural regeneration; seed production; seed year
Abstract | View details | Full text in PDF | Author Info

The success of natural regeneration of Scots pine (Pinus sylvestris L.) is dependent on the amount and quality of seeds produced in each year. While seed production of Scots pine in Northern Finland is well known, there is little information about frequency of good seed years in Southern and Central Finland. The success of regeneration of Scots pine was studied by defining from the growth rings of felled sample trees in which year they started to grow. The aim was to choose sample trees so that they would form a continual series from seedlings to 100 years old trees. In the material there was too little sample trees from certain years. However, within the period of 1827-1910 it could be identified 13 good regeneration years of pine in Southern Finland. According to the study, pine has regenerated very well every 6th year and at least moderately well every 4th year. When compared to the previous studies made in Northern Finland, part of the good regeneration years seems to be same both in Northern and Southern Finland. Regeneration of Scots pine is affected also by the type and condition of the site.

The PDF includes a summary in German.

  • Ilvessalo, E-mail: li@mm.unknown (email)
article id 7512, category Article
Mauno Pesonen, Arto Kettunen, Petri Räsänen. (1995). Non-industrial private forest landowners’ choices of timber management strategies. Silva Fennica vol. 0 no. 250 article id 7512. https://doi.org/10.14214/aff.7512
Keywords: forest owners; timber management strategies; genetic algorithm; strategic decision making
Abstract | View details | Full text in PDF | Author Info

The factors affecting the non-industrial, private forest owners’ (NIPF) strategic decisions in management planning are studied. A genetic algorithm is used to induce a set of rules predicting potential cut of the forest owners’ choices of preferred timber management strategies. The rules are based on variables describing the characteristics of the landowners and their forest holdings. The predictive ability of a genetic algorithm is compared to linear regression analysis using identical data sets. The data are cross-validated seven times applying both genetic algorithm and regression analyses in order to examine the data-sensitivity and robustness of the generated models.

The optimal rule set derived from genetic algorithm analyses included the following variables: mean initial volume, forest owner’s positive price expectations for the next eight years, forest owner being classified as farmer, and preference for the recreational use of forest property. When tested with previously unseen test data, the optimal rule set resulted in a relative root mean square error of 0.40.

In the regression analyses, the optimal regression equation consisted of the following variables: mean initial volume, proportion of forestry income, intention to cut extensively in future, and positive price expectations for the next two years. The R2 of the optimal regression equation was 0.3 and the relative root mean square error from the test data 0.38.

In both models, mean initial volume and positive stumpage price expectations were entered as significant predictors of potential cut of preferred timber management strategy. When tested with complete data set of 201 observations, both the optimal rule set and the optimal regression model achieved the same level of accuracy.

  • Pesonen, E-mail: mp@mm.unknown (email)
  • Kettunen, E-mail: ak@mm.unknown
  • Räsänen, E-mail: pr@mm.unknown
article id 7506, category Article
Eero Kubin, Lauri Kemppainen. (1994). Effect of soil preparation of boreal spruce forest on air and soil temperature conditions in forest regeneration areas. Silva Fennica vol. 0 no. 244 article id 7506. https://doi.org/10.14214/aff.7506
Keywords: soil preparation; soil temperature; forest regeneration; air temperature; boreal spruce forest
Abstract | View details | Full text in PDF | Author Info

The effect of scarification, ploughing and cross-directional ploughing on temperature conditions in the soil and adjacent air layer have been studied during 11 growth periods by using an unprepared clear-cut area as a control site. The development of seedling stand was followed to determine its shading effect on the soil surface.

Soil preparation decreased the daily temperature amplitude of the air at the height of 10 cm. The maximum temperatures on sunny days were lower in the tilts of the ploughed and in the humps of the cross-directional ploughed sites compared with the unprepared area. Correspondingly, the night temperatures were higher and so the soil preparation reduced the risk of night frost. In the soil at the depth of 5 cm, soil preparation increased daytime temperatures and reduced night temperatures compared with unprepared area. The maximum increase in monthly mean temperatures was almost 5°C, and the daily variation in the surface parts of the tilts and humps increased so that excessively high temperatures for the optimal growth of the root systems were measured from time to time. The temperature also rose at the depths of 50 and 100 cm.

Soil preparation also increased the cumulative temperature sum. The highest sums accumulated during the summer months were recorded at the depth of 5 cm in the humps of cross-directional ploughed area (1,127 dd.) and in the tilts of the ploughed area (1,106 dd.), while the corresponding figure in the unprepared soil was 718 dd. At the height of 10 cm the highest temperature sum was 1,020 dd. in the hump, and 925 dd. in the unprepared area.

The incidence of high temperature amplitudes and frequency of high temperatures at the depth of 5 cm decreased most rapidly in the humps of cross-sectional ploughed area and the ploughing tilts towards the end of the study period. The decrease was attributed principally to the compressing of tilts, the ground vegetation succession and the growth of seedlings. The difference between the prepared and unprepared area did not diminish. The increase in temperature due to soil preparation, thus, lasted at least over 10 years.

  • Kubin, E-mail: ek@mm.unknown (email)
  • Kemppainen, E-mail: lk@mm.unknown
article id 7656, category Article
Kari Leinonen, Matti Leikola, Antti Peltonen, Pentti Räsä­nen. (1989). Kuusen luontainen uudistaminen Pirkka-­Hämeen metsälautakunnassa. Silva Fennica vol. 0 no. 209 article id 7656. https://doi.org/10.14214/aff.7656
English title: Natural regeneration of Norway spruce in Pirkka-Häme Forestry Board District, Southern Finland.
Original keywords: kuusi; luontainen uudistaminen; taimettuminen; suojuspuumenetelmä
English keywords: Picea abies; natural regeneration; shelterwood method; restocking
Abstract | View details | Full text in PDF | Author Info

The aim of the present study was to evaluate and develop the use of natural regeneration of Norway spruce (Picea abies (L.) H. Karst.) in private forestry. The study was carried out using a line-plot survey with permanent circular sample plots. In total 40 regeneration sites were measured. The study includes results from three successive inventories: prior to the shelterwood cutting, in the summer after the cutting, and one year after the cutting. Regression and logistic regression analyses were used to construct models describing the effect of various factors on the restocking of the stands.

The standing volume prior to the shelterwood cutting was on average 236 m3/ha (ranging from 80 to 428 m3/ha) and after the cutting 120 m3/ha (39–220 m3/ha). The average number of stems per hectare decreased from 435 to 186. Prior to the shelterwood cutting 22% of the stands were satisfactorily restocked. After the cutting and one year later these percentages were 6 and 29%, respectively. Prior to the shelterwood cutting the number of acceptable seedlings was 1,440/ha, in the summer and year later 1,308/ha and 1,546/ha, respectively. Prior to the shelterwood cutting the characteristics of the mother stands did not correlate well with the number of seedlings. The change in the number of seedlings during the initial stage of shelterwood method depended on height of the seedling stand, amount of logging waste and number of germlings prior to the cutting. The risk to fail in regeneration was highest in the poorly restocked, sparse shelterwood stands, where a fast expansion of grass vegetation took place.

The PDF includes a summary in English.

  • Leinonen, E-mail: kl@mm.unknown (email)
  • Leikola, E-mail: ml@mm.unknown
  • Peltonen, E-mail: ap@mm.unknown
  • Räsä­nen, E-mail: pr@mm.unknown
article id 7647, category Article
Shikui Peng. (1987). On the combination of multitemporal satellite and field data for forest inventories. Silva Fennica vol. 0 no. 200 article id 7647. https://doi.org/10.14214/aff.7647
Keywords: stratification; SPR; two phase sampling; generalized least squares; multitemporal
Abstract | View details | Full text in PDF | Author Info

The study concerns with the methodology and efficiency of integrating multitemporal satellite image data with permanent sample plots in a continuous forest inventory and compartment wise estimation. The generalized least squares estimation for the population, and the unsupervised stratification for compartments, are the main estimation methods employed. The experimental material is composed of real and simulated multitemporal images (Landsat TM) and field data. Precision analyses for the estimation and comparisons for a number of estimation and updating methods, as well as statistical options are given.

The PDF includes a summary in Finnish.

  • Peng, E-mail: sp@mm.unknown (email)
article id 7642, category Article
Alpo Luomajoki. (1986). Timing of microsporogenesis in trees with reference to climatic adaptation: a review. Silva Fennica vol. 0 no. 196 article id 7642. https://doi.org/10.14214/aff.7642
Keywords: photoperiod; annual cycle; heat sum; microsporogenesis; meiosis; tetrad
Abstract | View details | Full text in PDF | Author Info

The timing of the tetrad phase of microsporogenesis in sixteen tree species, belonging to the genera of Abies, Larix, Picea, Pinus, Alnus, Betula, Corylus and Populus, was studied. The tetrad phase of microsporogenesis in conifers and in Populus tremula L. was reached from late March to early June including the yearly and latitudinal variation. The tetrad phase in Betulaceae was reached in late July to mid-August. The microsporogenesis in Betulaceae species differed in ecophysiological terms from the other species studied in that the timing in Betulaceae was rather day-length dependent than heat sum-correlated. In conifers and in Populus the timing of tetrad phase correlated with heat sums accumulated and did not correlate with day length or any kind of thermal threshold. This difference was, however, judged to be associated to seasonal adaptive strategies rather than taxonomic relationships.

The PDF includes a summary in Finnish.

  • Luomajoki, E-mail: al@mm.unknown (email)
article id 7625, category Article
Antti Turakka, Olavi Luukkanen, Suree Bhumibhamon. (1982). Notes on Pinus kesiya and P. merkusii and their natural regeneration in watershed areas of northern Thailand. Silva Fennica vol. 0 no. 178 article id 7625. https://doi.org/10.14214/aff.7625
Keywords: forest fires; natural regeneration; Pinus kesiya; Thailand; Pinus mercusii
Abstract | View details | Full text in PDF | Author Info

The aim of the present work was to clarify the structure of selected natural Pinus kesiya Royale ex Gord. And P. merkusii Jungh. et de Vriese stands, with particular attention on the amount and quality of naturally-occurring seedlings and young trees. Furthermore, basic information required in the management of pine stands in watershed areas of northern Thailand was obtained.

Large number of pine seedlings were found in P. kesiyana stands only. Two of the examined four localities where this species occurred also included various intermediary height classes under the mature trees. In dense stands the number of seedlings was generally smaller as compared with lower seed tree densities. Neither one of the two localities where P. Merkusii was investigated indicated sufficient natural regeneration. Young pines (over 5 m height) seemed to survive the frequent ground fires quite well, whereas younger seedlings were destroyed and the ground layer vigour was lowered in these cases.

In situ sowing experiments at the beginning of the dry season indicated a faster development and better survival of emerging P. Kesiya seedlings as compared to P. Merkusii. Soil preparation and exposure to sun decreased the survival of seedlings in both species. Utilization of natural regeneration and direct sowing of pines as a silvicultural method as well as the general significance of the two pine species in the succession of plant communities under the influence of forest fires is also discussed.

The PDF includes a summary in Finnish.

  • Turakka, E-mail: at@mm.unknown (email)
  • Luukkanen, E-mail: ol@mm.unknown
  • Bhumibhamon, E-mail: sb@mm.unknown
article id 7620, category Article
Min-Sup Chung. (1981). Biochemical methods for determining population structure in Pinus sylvestris L. Silva Fennica vol. 0 no. 173 article id 7620. https://doi.org/10.14214/aff.7620
Keywords: Scots pine; tree breeding; Finland; genetic variation; population structure; plus trees; self-fertilization
Abstract | View details | Full text in PDF | Author Info

Studies on Finnish Scots pine (Pinus sylvestris L.) plus tree clones by monoterpene and isozyme analyses was undertaken to further investigate mating system, population structure and pollination. Six allozyme systems (3 GOT, 1 GDH and 2 LAP) were properly analysed on the basis of segregation. Monoterpenes were analysed from needle material and segregation in high and low 3-carene content was found to depend on two alleles C and c. Thus, six allozyme systems and one monoterpene system were used as markers in this study.

It was shown that the northern clonal group maintains a much genetic variation as the central or southern clonal groups. The conditional probability of self-fertilization in about 20-year old clones estimated by the multilocus model was 14.1%, of which 8% originate from mating between trees that carry the same alleles to one of the maternal parent at some loci and 6% through self-fertilization.

There was no prominent difference in allele frequency of male gametes that pollinated the very early or very late flowering clones. The northern clonal group has higher a lower frequency of alleles GOT B2 and B3 respectively than of the southern clonal groups. The artificial plus tree selection, particularly in northern Finland, appears to favour heterozygous genotypes for the alleles that control 3-carene content n Scots pine.

The PDF includes a summary in Finnish.

  • Chung, E-mail: mc@mm.unknown (email)
article id 7575, category Article
Robert T Brown, Peitsa Mikola. (1974). The influence of fruticose soil lichens upon the mycorrhizae and seedling growth of forest trees. Silva Fennica vol. 0 no. 141 article id 7575. https://doi.org/10.14214/aff.7575
Keywords: Pinus sylvestris; Norway spruce; Betula pendula; Picea abies; regeneration; Scots pine; lichens; Cladonia alpestris; Cladonia rangiferina; Cladonia arbuscula; Cladonia pleurota; Cetraria islandica; Stereocaulon paschale; growth inhibition
Abstract | View details | Full text in PDF | Author Info

Water extracts of six common soil lichens, Cladonia alpestris, C. rangiferina, C. arbuscula (sylvatica), C. pleurota, Cetraria islandica, Stereocaulon paschale, inhibited growth of ectomycorrhizae of Pinus sylvestris (L.). Of 17 fungi (12 mycorrhizal) tested, many were inhibited while others were scarcely influenced or even occasionally stimulated by the extracts. Cladonia alpestris extract inhibited most fungi while C. rangiferina showed much less influence.

In pure culture synthesis experiments, 32P uptake of Pinus sylvestris was significantly reduced by C. alpestris extract. Different species of fungi showed widely variant abilities to pick up 32 P. in nursery experiments, much more vigorous growth of P. sylvestris and Picea abies (L.) H. Karst. was obtained on plots without C. alpestris than on paired plots covered with it. Betula verrucosa (B. pendula Roth) showed no difference. Under natural forest conditions, P. sylvestris seedlings grow much more rapidly where C. alpestris had been eliminated by road building or reindeer grazing than do seedlings only one meter distant under undisturbed C. alpestris cover. It is suggested that by properly controlled reindeer grazing, establishment and early growth of P. sylvestris on Cladonia sites can be much enhanced. By the time that C. alpestris could become re-established the pine seedlings would have grown large enough to suffer little from reindeer grazing. This study shows the continuity of the major components of the forest tundra biome – the dependence of pines, mycorrhizae, lichens, and reindeer and their predators (human or otherwise) upon each other for a healthy existence.

The PDF includes a summary in Finnish.

  • Brown, E-mail: rb@mm.unknown (email)
  • Mikola, E-mail: pm@mm.unknown
article id 7561, category Article
Kalervo Salonen. (1973). On the life cycle, especially on the reproduction biology of Blastophagus piniperda L. ( Col., Scolytidae). Silva Fennica vol. 0 no. 127 article id 7561. https://doi.org/10.14214/aff.7561
Keywords: regeneration; Tomicus piniperda; swarming; egg galleries; Blastophagus piniperda
Abstract | View details | Full text in PDF | Author Info

Blastophagus piniperda L. (now Tomicus piniperda) life cycle, especially temperature effects at various stages, has been studied in laboratory, outdoor development experiments and in natural conditions in the field in Finland, 1961–1970.

It was established that when a tree with overwintering beetles in its base is felled during the winter, the overwintered insects can establish an egg gallery into the stump without leaving for swarming flight. The egg gallery can be established in this case already about a month before swarming starts. Swarming starts after temperature in the shade exceeds 10° C. It is at its maximum at a temperature over 12° C and in conditions of no wind. The swarming starts in Southern Finland generally in the latter half of April, and in Lapland two to four weeks later than in Southern Finland, and it lasts from a few days to a few weeks. As the temperature rises, the rate of egg gallery construction and the rate of egg laying increase. The characteristics of the galleries and pupal stage are described in detail.

In Finnish conditions, B. Piniperda does not have a diapause. Movement into overwintering sites starts after the minimum daily temperature goes below 0° C. About 10% of the insects found in the overwintering sites in spring were estimated to have died during the winter. With increasing density, the number of new beetles which developed from each egg gallery decreased.

The PDF includes a summary in Finnish.

  • Salonen, E-mail: ks@mm.unknown (email)
article id 7545, category Article
Lalli Laine, Matti Nuorteva. (1970). On the antagonistic influence of insect pathogenic fungi Beauveria bassiana (Bals.) Vuill. and B. tenella (Delacr.) Siem. on root rot disease spongy saprot (Fomes annosus (Fr.) Cooke). Silva Fennica vol. 0 no. 111 article id 7545. https://doi.org/10.14214/aff.7545
Keywords: antagonism; root rot; Fomitopsis annosa; insect pathogenic
Abstract | View details | Full text in PDF | Author Info

The article presents the studies about antagonistic influence of insect pathogenic fungi Beauveria bassiana (Bals.) Vuill. and B. tenella (Delacr.) Siem. against root rot diseases. The experiments were conducted in laboratory where the fungi were grown in Petri dishes. The results show that these fungi are antagonistic with each other. The used stem of B. bassiana was proved as strongest antagonist against all studied F.annosus.

The PDF contains a summary in German. 

  • Laine, E-mail: ll@mm.unknown (email)
  • Nuorteva, E-mail: mn@mm.unknown
article id 7607, category Article
Eino Oinonen. (1969). The time table of vegetative spreading of the lily-of-the-valley (Convallaria majalis L.) and the wood small-reed (Calamagrostis epigeios (L.) Roth) in southern Finland. Silva Fennica vol. 0 no. 97 article id 7607. https://doi.org/10.14214/aff.7607
Keywords: regeneration; forest fire; Calamagrostis epigeios; lily-of-the-valley; Convallaria majalis; wood small-reed
Abstract | View details | Full text in PDF | Author Info

The purpose of this study has been to compile a time-table for the vegetative spreading o the lily-of-the-valley and the wood small-reed. The diameters of the mainly solitary stands of these species have been compared to stand diameters of bracken (Lycopodium clavatum, L. annotium, and L. complanatum), to tree age determined by basal borings, and to times of fires in the site.

Regeneration of the lily-of-the-valley (Convallaria Majalis L.) and the wood small-reed (Calamagrostis epigeios (L.) Roth) from seed is relatively infrequent, and distance between stands may be considerable. Regeneration is more common after fire. The stands of the species are relatively fire resistant, especially on heathlands, where the subsurface parts are safe in the mineral soil. Large stands are generally due to vegetative spreading.

The rate of vegetative spreading of both of the species is practically constant, and about equal, 12.5 cm/year in diameter and 6.2 cm/year in the radial direction. When the actual regeneration time after the fire is not known, the rate of the spreading can be lower than this number.

  • Oinonen, E-mail: eo@mm.unknown (email)
article id 7187, category Article
Eino Oinonen. (1968). Lycopodium clavatum L.- ja L. annotium L.-kasvustojen laajuus rinnastettuna samanpaikkaisiin L. complanatum L.- ja Pteridium aquilinum (L.) Kuhn -esiintymiin sekä puuston ikään ja paloaikoihin. Silva Fennica vol. 0 no. 87 article id 7187. https://doi.org/10.14214/aff.7187
English title: The size of Lycopodium clavatum and L. Annotinum stands as compared to that of L. Complanatum and Pteridium aquilinum stands, the age of the stand and the dates of fire, on the site.
Original keywords: metsäpalo; uudistuminen; kloonit; katinlieko; riidenlieko; lisääntyminen
English keywords: regeneration; forest fire; clones; Pteridium aquilinum; Lycopodium complanatum; Lycopodium clavatum; Lycopodium annotium
Abstract | View details | Full text in PDF | Author Info

The present study is a part of larger project into the size and age of certain forest plants. This study seeks to confirm the size-age relationship of Lycopodium clavatum L. and L. annotium L. stands, and the time of sporal regeneration. The stand dimensions were plotted against the size of bracken (Pteridium aquilinum (L.) Kuhn) and ground pine (Lycopodium complanatum L.) stands growing on the same site, and against the age of the timber and the time of fire on the site.

The method based on parallel measurements carried out in burned forests has proved to be suitable for establishing the rate of spreading of stands and their age. The three club-moss species proved to be much alike in their sporal regeneration. They, as well as bracken, regenerate under the condition created by fire. The largest detached patches are often clones. Both Lycopodium clavatum and L. annotium survive fire poorly. Thus, the maximum size of the clones is usually connected with the last forest fire. Pieces of the shoots may survive the fires.

Sporal regeneration of these species also occur without the aid of fire. As also variation in the annual growth of the shoots can be considerable, the stand size of these species is not as good indicator of the date of fire as with Pteridium aquilinum and L. complanatum. The winding appearance of the shoots increases the error in determination of stand age by the stand size further.

The results emphasize the importance of taking into account the time and spread of the plants in the traditional vegetation analysis. A central question is: what is an individual.

The PDF includes a summary in English.

  • Oinonen, E-mail: eo@mm.unknown (email)

Category: Research article

article id 10544, category Research article
Lars Sprengel, Heinrich Spiecker, Shuirong Wu. (2022). Two subject specific modelling approaches to construct base-age invariant polymorphic site index curves with varying asymptotes. Silva Fennica vol. 56 no. 1 article id 10544. https://doi.org/10.14214/sf.10544
Keywords: generalized algebraic difference approach; stem analysis; dummy variable approach; gnls; nonlinear mixed-effects models; nlme; Zhongtiaoshan forest region
Highlights: Base-age invariant families of height growth curves with polymorphism and varying asymptotes are presented for the seven economically most important tree species in Zhongtiaoshan forest region, China; The nonlinear fixed-effects approach outperforms the nonlinear mixed-effects approach according to the AIC, but according to RMSE and bias these results are not fully supported.
Abstract | Full text in HTML | Full text in PDF | Author Info

For constructing growth and yield models the concept of site index as measure of productivity is crucial. Here, we use nonlinear mixed-effects models (NLME) with random individual effects and nonlinear models with dummy variables as fixed individual effects (NLFE) to fit mechanistic growth functions to stem analysis data of the economically most important tree species in Zhongtiaoshan forest region, China. The Richards and Lundqvist function are formulated into five dynamic equations (R1, R2, L1, L2 and L3) applying the generalized algebraic difference approach (GADA), which inherit polymorphism, varying asymptotes and base-age invariance. According to Akaike information criterion the R1 model as NLFE fits height growth data of Pinus tabuliformis Carrière, Pinus armandii Franch., Quercus liaotungensis Koidz., Quercus aliena Blume and Betula platyphylla Sukaczev best, while for Quercus variabilis Blume R2 as NLFE fits height growth data best. For Larix principis-rupprechtii Mayr L1 as NLME has been selected as best model, as R1 and R2 both as NLFE and NLME are not extrapolating the comparably short length of height growth data well enough. However, according to the root mean square error and bias differences between model fits of both the selected equation and the chosen model fitting approach are not so clear. Presented families of height growth curves serve as planning tools to identify site index and therefore assess productivity of forest stands in the studied region. A direct comparison of the productivity of forest stands of the same tree species is possible due to base-age invariance of the selected models.

  • Sprengel, Chair of Forest Growth and Dendroecology, Albert-Ludwigs-University Freiburg, Tennenbacher Str. 4, 79106 Freiburg, Germany ORCID https://orcid.org/0000-0002-6332-7911 E-mail: lars.sprengel@iww.uni-freiburg.de (email)
  • Spiecker, Chair of Forest Growth and Dendroecology, Albert-Ludwigs-University Freiburg, Tennenbacher Str. 4, 79106 Freiburg, Germany E-mail: instww@uni-freiburg.de
  • Wu, Research Institute of Forest Policy and Information, Chinese Academy of Forestry, No. 1 Dongxiaofu, Haidian District, Beijing 100091, China E-mail: shuirongwu@126.com
article id 10512, category Research article
Mateusz Liziniewicz, Ignacio Barbeito, Andis Zvirgzdins, Lars-Göran Stener, Pentti Niemistö, Nils Fahlvik, Ulf Johansson, Bo Karlsson, Urban Nilsson. (2022). Production of genetically improved silver birch plantations in southern and central Sweden. Silva Fennica vol. 56 no. 1 article id 10512. https://doi.org/10.14214/sf.10512
Keywords: Betula pendula; planting; generalized algebraic difference approach; genetic gain; stand basal area starting function
Highlights: The basal area development of genetically improved birch in Sweden was modeled using a generalized algebraic difference approach; The best model fit, both graphically and statistically was delivered by the Korf base model; The analysis of realized gain trial showed a stability of relative differences in basal area between tested genotypes.
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Investing in planting genetically improved silver birch (Betula pendula Roth) in Swedish plantations requires understanding how birch stands will develop over their entire rotation. Previous studies have indicated relatively low production of birch compared to Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.). This could result from using unrepresentative basic data, collected from unimproved, naturally-regenerated birch (Betula spp.) growing on inventory plots often located in coniferous stands. The objective of this study was to develop a basal area development function of improved silver birch and evaluate production over a full rotation period. We used data from 52 experiments including planted silver birch of different genetic breeding levels in southern and central Sweden. The experimental plots were established on fertile forest sites and on former agricultural lands, and were managed with different numbers of thinnings and basal area removal regimes. The model best describing total stand basal area development was a dynamic equation derived from the Korf base model. The analysis of the realized gain trial for birch showed a good stability of the early calculated relative differences in basal area between tested genotypes over time. Thus, the relative difference in basal area might be with cautious used as representation of the realized genetic gain. On average forest sites in southern Sweden, improved and planted silver birch could produce between 6–10.5 m3 ha–1 year–1, while on fertile agriculture land the average productivity might be higher, especially with material coming from the improvement program. The performed analysis provided a first step toward predicting the effects of genetic improvement on total volume production and profitability of silver birch. However, more experiments are needed to set up the relative differences between different improved material.

  • Liziniewicz, The Forestry Research Institute of Sweden, Ekebo, SE-268 90 Svalöv, Sweden E-mail: mateusz.liziniewicz@skogforsk.se
  • Barbeito, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, Alnarp, Sweden; Université de Lorraine, AgroParisTech, INRAE, UMR Silva, Nancy, France E-mail: ignacio.barbeito@slu.se
  • Zvirgzdins, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, Box 49, 23053 Alnarp, Sweden E-mail: andis.zvirgzdins@slu.se (email)
  • Stener, The Forestry Research Institute of Sweden, Ekebo, SE-268 90 Svalöv, Sweden E-mail: lg.stener@telia.com
  • Niemistö, Natural Resources In-stitute Finland (Luke), Natural resources, Seinäjoki, Finland E-mail: pentti.niemisto@luke.fi
  • Fahlvik, The Forestry Research Institute of Sweden, Ekebo, SE-268 90 Svalöv, Sweden E-mail: nils.fahlvik@skogforsk.se
  • Johansson, Tönnersjöheden Experimental Forest, SLU, Simlångsdalen, Sweden E-mail: ulf.johansson@slu.se
  • Karlsson, The Forestry Research Institute of Sweden, Ekebo, SE-268 90 Svalöv, Sweden E-mail: curly.birch@gmail.com
  • Nilsson, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, Box 49, 23053 Alnarp, Sweden E-mail: urban.nilsson@slu.se
article id 10528, category Research article
Jaana Luoranen, Heli Viiri. (2021). Comparison of the planting success and risks of pine weevil damage on mineral soil and drained peatland sites three years after planting. Silva Fennica vol. 55 no. 4 article id 10528. https://doi.org/10.14214/sf.10528
Keywords: survival; planting; mechanical site preparation; ground vegetation; forest regeneration; Hylobius abietis
Highlights: The planting success was poorer on peatland sites than on mineral soil; Dense ground vegetation cover is more probable on peatland than on mineral soil; No differences in pine weevil feeding damage on mineral soil and peatland were found; Cultivated mineral soil reduced the vegetation cover, feeding damage and seedling mortality.
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Over 20% of regeneration operations will be on drained peatland in the next decade in Finland. There are only a few studies comparing the planting success and the risk of pine weevil (Hylobius abetis (L.) feeding damage on mineral soil and drained peatland. Thirty sites planted with Norway spruce (Picea abies (L.) H. Karst.) container seedlings in 2009 in Southern and Central Finland were inventoried three growing seasons after planting. Prediction models for the probability of survival, pine weevil damage and the presence of ground vegetation cover were done separately for peatland and mineral soil sites. The planting success was 17% lower on peatland sites (1379 surviving seedlings ha–1) than on mineral soil (1654 seedlings ha–1). The factors explaining the survival were the ground vegetation cover and type of the planting spot on the peatland, and the ground vegetation cover on mineral soil. On mineral soil, 76% of the planting spots were on cultivated mineral soil while on peatland only 28% of the seedlings were planted on similar spots. There were also fewer seedlings that were surrounded by dense ground vegetation on mineral soil (4%) than on peatland (14%). Pine weevil feeding damage did not differ significantly on peatland (23%) or mineral soil (18%). The more time there was from clear-cutting, the more the probability of pine weevil feeding damage was reduced on both soil classes. Additionally, cover vegetation in the vicinity of the seedlings increased on mineral soil. Cultivated planting spots, especially those covered by mineral soil, prevented pine weevil feeding and reduced the harmful effects of vegetation on the seedlings both on mineral soil and peatland.

article id 10485, category Research article
Felicia Dahlgren Lidman, Emma Holmström, Tomas Lundmark, Nils Fahlvik. (2021). Management of spontaneously regenerated mixed stands of birch and Norway spruce in Sweden. Silva Fennica vol. 55 no. 4 article id 10485. https://doi.org/10.14214/sf.10485
Keywords: Betula pendula; Picea abies; Betula pubescens; natural regeneration; mixed forest; land expectation value
Highlights: The absence of forest management does not always mean economic loss; With dense spontaneous regeneration of birch and Norway spruce, the first competition release can have a high impact on future stem development; Significantly different effects on stand volume production and diameter development of Norway spruce can be expected with different biomass harvest strategies.
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Timber production and profitability were evaluated for spontaneously-regenerated mixtures on two formerly clearcut areas. The abandoned areas developed into birch-dominated (Betula pendula Roth and Betula pubescens Ehrh.) stands with successional ingrowth of Norway spruce (Picea abies (L.) H. Karst.). An experiment with randomized treatments within blocks was established, using three management strategies and one unthinned control, resulting in variation in optimal rotation age, merchantable volume and species composition. The management strategies were evaluated based on total production (volume) by using measured growth data 42 years after clearcutting and the modelled future stand development. The long-term effects of spontaneous regeneration and management strategies were evaluated based on land expectation value (LEV) and compared with a fifth management strategy using artificial regeneration and intense thinnings. 12 years after treatment, at a stand age of 42 years, the unthinned control had produced the highest total stem volume. At interest rates of 2% or higher, the unmanaged forest was an economically viable strategy, even compared to an intensive management strategy with a preferred merchantable timber species. Interest rates clearly impacted the profitability of the different management strategies. This study shows that when spontaneous regeneration is successful and dense, the first competition release can have a high impact on the development of future crop trees and on the species mixture.

article id 10506, category Research article
Mikko Tikkinen, Terhi Latvala, Tuija Aronen. (2021). Interest in vegetatively propagated Norway spruce materials – a survey among Finnish forest owners and professionals. Silva Fennica vol. 55 no. 3 article id 10506. https://doi.org/10.14214/sf.10506
Keywords: Picea abies; desired traits; intention of adoption; somatic embryogenesis; willingness to pay
Highlights: Survey among forest owners and professionals’ perceptions towards tree breeding and vegetative propagation yielded over 3000 responses; Most valued features in reforestation material were: improved resilience in changing climate, improved pest and pathogen resistance, and securing the species’ gene pool; Majority of respondents accepted reforestation with vegetatively propagated material to some extent; Willingness to pay for improved features was indicated.
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Forests and forestry will encounter several changes of unknown magnitude within the coming decades. In the Nordic, long rotations complicate any anticipation to the upcoming changes. Tree breeding can contribute to coping with these changes. The time span of implementing breeding results in practice may be shortened through vegetative propagation. Introducing vegetative propagation to forest regeneration may phase several challenges before adopted by the industry, some of which are related to perceptions about new technology. Firstly, private forest owners are in a key role in implementing the technology in practice; although they do not represent the overall public, they are the decision makers in their own estates regarding forestry and forest regeneration. Secondly, the professionals related to the production of forest regeneration material and plants from forest species are in a key role when it comes to practically introducing the new technology to the forest owners. In this survey, perceptions of forest owners and professionals towards tree breeding and vegetative propagation were investigated. Additionally, the respondents were asked which traits they considered important to be improved by breeding, and their willingness to pay for these improved traits. The respondents valued the most: improved pest and pathogen resistance, improved resilience of forest in changing climate, and securing the species’ gene pool. Responses indicated that forest owners would be willing to pay more for the improved traits in forest regeneration material. The current novel study provides a foundation to concern public awareness regarding tree breeding and vegetative propagation in the future.

  • Tikkinen, Natural Resources Institute Finland ORCID https://orcid.org/0000-0002-2368-8042 E-mail: mikko.tikkinen@luke.fi (email)
  • Latvala, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: terhi.latvala@luke.fi
  • Aronen, Natural Resources Institute Finland (Luke), Production systems, Vipusenkuja 5, FI-57200 Savonlinna, Finland E-mail: tuija.aronen@luke.fi
article id 10369, category Research article
Tiina Laine, Ville Kankaanhuhta, Juho Rantala, Timo Saksa. (2020). Effects of spot mounding and inverting on growth of conifers, exposed mineral soil and natural birch regeneration. Silva Fennica vol. 54 no. 5 article id 10369. https://doi.org/10.14214/sf.10369
Keywords: boreal forest; regeneration; site preparation; mounding; young stand management
Highlights: There were 21% fewer naturally regenerated birches ha–1 in inverted areas (22 951) compared to spot mounded areas (29 086); Spot mounding exposed slightly, but not statistically significantly, larger area of mineral soil than inverting; There was no difference in seedling mortality or growth between the site preparation methods.
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In Nordic forests, consistent evidence about better seedling survival rate and increased growth due to site preparation have been obtained in numerous studies. Proper site preparation method can reduce costs of the whole regeneration chain through its effects on survival of planted seedlings, abundance of natural regeneration and competition in early stand development. This study compared the natural regeneration of birches (silver birch (Betula pendula Roth) and downy birch (B. pubescens Ehrh.)), amount of exposed mineral soil, and growth of planted seedlings between spot mounding and inverting site preparation methods. Present study was conducted in eight forest stands established in 2012 or 2015. Even though difference was not statistically significant, inverting exposed less mineral soil than spot mounding and thus reduced the natural regeneration of birch seedlings by 6135 seedlings ha–1 compared to spot mounding. However, the variation between regeneration areas was remarkable. There was no difference in seedling mortality or growth between the site preparation methods. In order to achieve high growth of conifers, moderate amount of exposed mineral soil and thus less naturally regenerated birch, inverting should be favored over spot mounding.

  • Laine, Metsä Group, P.O. Box 208, FI-70101 Kuopio, Finland E-mail: tiina.laine@metsagroup.com (email)
  • Kankaanhuhta, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: ville.kankaanhuhta@luke.fi
  • Rantala, Metsä Group, P.O. Box 10, FI-02020 METSÄ, Finland E-mail: juho.rantala@metsagroup.com
  • Saksa, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@luke.fi
article id 10370, category Research article
Juha Lappi, Timo Pukkala. (2020). Analyzing ingrowth using zero-inflated negative binomial models. Silva Fennica vol. 54 no. 4 article id 10370. https://doi.org/10.14214/sf.10370
Keywords: regeneration; continuous cover forestry; count data; generalized linear model; overdispersion; right-censoring
Highlights: Models were developed to describe ingrowth in national forest inventory data; The data were more dispersed than Poisson data and included many zeros; Fixed-effects models had larger zero-inflation probability and overdispersion parameter than mixed-effect models; Mixed-effects models had larger likelihood than fixed-effects models but provided biased predictions; Prediction of right-censored ingrowth may be useful owing to large overdispersion.
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Ingrowth is an important element of stand dynamics in several silvicultural systems, especially in continuous cover forestry. Earlier predictive models for ingrowth in Finnish forests are few and not based on up-to-date statistical methods. Ingrowth is here defined as the number of trees over 1.3 m entering a plot. This study developed new ingrowth models for Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies (L.) H. Karst.) and birch (Betula pendula Roth and B. pubescens Ehrh.) using data from the permanent sample plots of the Finnish national forest inventory. The data were over-dispersed compared to a Poisson process and had many zeros. Therefore, a zero-inflated negative binomial model was used. The total and species-specific stand basal areas, temperature sum and fertility class were used as predictors in the ingrowth models. Both fixed-effects and mixed-effects models were fitted. The mixed-effects model versions included random plot effects. The mixed-effects models had larger likelihoods but provided biased predictions. Also censored prediction was considered where only a certain maximum number of ingrowth trees were accepted for a plot. The models predicted most pine ingrowth in pine-dominated stands on sub-xeric and xeric sites where stand basal area was low. The predicted amount of spruce ingrowth was maximized when the basal area of spruce was 13 m2 ha–1. Increasing temperature sum increased spruce ingrowth. Predicted birch ingrowth decreased with increasing stand basal area and towards low fertility classes. An admixture of pine increased the predicted amount of spruce ingrowth.

  • Lappi, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: juha.lappi.sjk@gmail.com (email)
  • Pukkala, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: timo.pukkala@uef.fi
article id 10220, category Research article
Arnis Gailis, Pauls Zeltiņš, Andis Purviņš, Juris Augustovs, Valts Vīndedzis, Inga Zariņa, Āris Jansons. (2020). Genetic parameters of growth and quality traits in open-pollinated silver birch progeny tests. Silva Fennica vol. 54 no. 2 article id 10220. https://doi.org/10.14214/sf.10220
Keywords: Betula pendula; tree breeding; genetic gain; genetic parameters
Highlights: Growth and stem quality traits were under strong genetic control; Weak genetic correlations between tree growth and stem quality were found; Strong age-age and type-B correlations suggest robust improvement over time and different environments; Simultaneous improvement of growth and stem quality might be applicable.
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Genetic parameters of growth and stem quality traits were estimated for open-pollinated silver birch Betula pendula Roth progenies in Latvia at the age of 10 and 14 years. Tree height and stem volume were found to be under strong genetic control at both inventories (narrow-sense heritabilities varied from 0.41 to 0.66). Mainly low heritabilities were found for stem defects, yet genetic control of branch diameter, stem straightness and overall stem quality varied from low to high depending on study site. High additive genetic coefficient of variation was found for stem volume (25.3–32.5%). Genetic correlations among growth traits were strong and positive (0.90–0.99). Mainly weak genetic correlations between growth and quality traits implied simultaneous improvement. Still, strong negative correlations between branch angle and stem straightness might result in enlarged knot size for straighter logs. The genetic age-age correlations were strong. Weak genotype by environment interaction and stability of best genotypes over different sites was indicated by strong genetic correlations between trials. Each growth or quality trait alone showed substantial improvement in terms of estimated genetic gain (up to 62% over trial mean for stem volume). Therefore, selection index combining both growth and stem quality may be developed.

  • Gailis, Latvian State Forest Research Institute Silava, 111 Rigas street, Salaspils LV-2169, Latvia E-mail: arnis.gailis@silava.lv
  • Zeltiņš, Latvian State Forest Research Institute Silava, 111 Rigas street, Salaspils LV-2169, Latvia ORCID https://orcid.org/0000-0002-6286-5814 E-mail: pauls.zeltins@silava.lv (email)
  • Purviņš, Latvian State Forest Research Institute Silava, 111 Rigas street, Salaspils LV-2169, Latvia E-mail: andis.purvins@silava.lv
  • Augustovs, Latvian State Forest Research Institute Silava, 111 Rigas street, Salaspils LV-2169, Latvia E-mail: juris.augustovs@silava.lv
  • Vīndedzis, Latvian State Forest Research Institute Silava, 111 Rigas street, Salaspils LV-2169, Latvia E-mail: valts.vindedzis@silava.lv
  • Zariņa, Latvian State Forest Research Institute Silava, 111 Rigas street, Salaspils LV-2169, Latvia E-mail: inga.zarina@silava.lv
  • Jansons, Latvian State Forest Research Institute Silava, 111 Rigas street, Salaspils LV-2169, Latvia E-mail: aris.jansons@silava.lv
article id 10243, category Research article
Laura Pikkarainen, Jaana Luoranen, Antti Kilpeläinen, Teppo Oijala, Heli Peltola. (2020). Comparison of planting success in one-year-old spring, summer and autumn plantings of Norway spruce and Scots pine under boreal conditions. Silva Fennica vol. 54 no. 1 article id 10243. https://doi.org/10.14214/sf.10243
Keywords: Pinus sylvestris; Picea abies; seedling damage; forest regeneration; field performance; planting season
Highlights: In Norway spruce, 84% of all plantings were successful, whereas in Scots pine, the corresponding number was 52%; The major reason for poor planting results was poor work quality; An extended planting season is possible for Norway spruce in southern and central Finland; In Scots pine, there are still large uncertainties in the success of summer and autumn plantings.
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In Nordic countries, tree planting of seedlings is mainly performed during spring and early summer. Interest has increased in extending the planting window throughout the unfrozen growing season. This study compared the success of one-year-old spring, summer and autumn plantings in practical forestry in Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) in southern and central Finland. Planting success was based on the number of viable seedlings per hectare relative to a species-specific target density. The influence of different factors to poor planting results were determined, including quality of site preparation and planting, and sources of natural damage. Overall, in Norway spruce, 85, 69 and 84% and in Scots pine 53, 55 and 40% of spring, summer and autumn plantings succeeded. In Norway spruce, the planting results were consistent between the southern and central regions, whereas in Scots pine, the success was slightly lower in the south. The poor work quality and a low density of appropriate planting spots, contributed to poor planting results, regardless of planting season, region or tree species. Considering different damages, especially mammal damage contributed to the failure of Scots pine spring plantings, whereas in summer plantings, corresponding single failure reason could not be identified. Based on our findings, extending the planting season of Norway spruce could be recommended in both regions. For Scots pine, there is still significant uncertainty about the success of summer and autumn plantings, partially due to the limited number of plantings available for analyses.

  • Pikkarainen, School of Forest Sciences, University of Eastern Finland, Yliopistonkatu 7, FI-80100 Joensuu, Finland E-mail: laura.pikkarainen@uef.fi (email)
  • Luoranen, Natural Resources Institute Finland (Luke), Production Systems, Neulaniementie 5, FI-70210 Kuopio, Finland E-mail: jaana.luoranen@luke.fi
  • Kilpeläinen, School of Forest Sciences, University of Eastern Finland, Yliopistonkatu 7, FI-80100 Joensuu, Finland E-mail: antti.kilpelainen@uef.fi
  • Oijala, Metsä Group, Metsä Forest, Kuormaajantie 7, FI-40320 Jyväskylä, Finland E-mail: teppo.oijala@metsagroup.com
  • Peltola, School of Forest Sciences, University of Eastern Finland, Yliopistonkatu 7, FI-80100 Joensuu, Finland E-mail: heli.peltola@uef.fi
article id 10019, category Research article
Junyan Liu, Junfeng Tang, Si-Chong Chen, Wenbao Ma, Zheng Zheng, Tingfa Dong. (2019). Do tree cavity density and characteristics vary across topographical habitats in the tropics? A case study from Xishuangbanna, southwest China. Silva Fennica vol. 53 no. 1 article id 10019. https://doi.org/10.14214/sf.10019
Keywords: heterogeneity; cavity-dependant animals; tropical rainforest; biodiversity conservation
Highlights: Cavities were significantly more abundant in high- and low-slope than high-plateau habitats; There are more “butt hollow” cavities in high-slope habitat and they occurred at a lower height; More “crack” cavities in low-slope habitat and they had a narrower entrance diameter; Certain types of cavities are concentrated in specific habitats, which provide opportunities for forest management and biodiversity conservation.
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Despite the influence of cavities on the survival and distribution of cavity-dependent fauna, the variation in the density and characteristics of tree cavities across different habitat types in tropical forests is unknown. In this study, we surveyed 26 312 living trees from 376 species and compared cavity density and characteristics (height, size, type, and orientation) across five habitat types (valley, low-slope, high-slope, high-gully, and high-plateau) in a 20-hectare tropical rainforest in southwest China. From a total of 2047 cavities, we found that cavity density was mainly driven by habitat rather than tree species richness or diameter at breast height (DBH), and the characteristics of cavities were not uniformly distributed across habitats. Cavities were significantly more abundant in high- and low-slope than high-plateau habitats. Compared with other habitats, more “butt hollow” cavity types were found in high-slope habitat and they occurred at a lower tree height, whereas more “crack” cavities were found in low-slope habitat and they had a narrower entrance diameter. Although the mean orientation of cavities faced towards the northeast, cavity orientation varied significantly across habitat types. Our results indicate that certain types of cavities are concentrated in specific habitat types, which can provide avenues for forest management and biodiversity conservation. We highlight the importance of habitat heterogeneity in providing resources for cavity nesters.

  • Liu, Key Laboratory of Southwest China Wildlife Resources Conservation of Ministry of Education, Key Laboratory of Environmental Science and Biodiversity Conservation (Sichuan Province) and Institute of Plant Adaptation and Utilization in Southwest Mountains, China West Normal University, Nanchong, Sichuan 637009, China; Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla, Yunnan 666303, China E-mail: liujunyan2300@163.com
  • Tang, Key Laboratory of Southwest China Wildlife Resources Conservation of Ministry of Education, Key Laboratory of Environmental Science and Biodiversity Conservation (Sichuan Province) and Institute of Plant Adaptation and Utilization in Southwest Mountains, China West Normal University, Nanchong, Sichuan 637009, China E-mail: jft@nn.ch
  • Chen, Royal Botanic Gardens, Kew, Wakehurst Place, West Sussex RH17 6TN, UK; Mitrani Department of Desert Ecology, Swiss Institute for Dryland Environmental and Energy Research, Jacob Blaustein Institutes for Desert Research, Ben-Gurion University of the Negev, Midreshet Ben-Gurion, Beer-Sheva 8499000, Israel E-mail: chensichong0528@gmail.com
  • Ma, Ecological Restoration and Conservation of Forests and Wetlands Key Laboratory of Sichuan Province, Sichuan Academy of Forestry, Chengdu 610081, China E-mail: mawenbao_2000@126.com
  • Zheng, Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla, Yunnan 666303, China E-mail: dioeco@outlook.com
  • Dong, Key Laboratory of Southwest China Wildlife Resources Conservation of Ministry of Education, Key Laboratory of Environmental Science and Biodiversity Conservation (Sichuan Province) and Institute of Plant Adaptation and Utilization in Southwest Mountains, China West Normal University, Nanchong, Sichuan 637009, China E-mail: dongtf@aliyun.com (email)
article id 10036, category Research article
Karin Hjelm, Lars Rytter. (2018). The demand of hybrid aspen (Populus tremula × P. tremuloides) on site conditions for a successful establishment on forest land. Silva Fennica vol. 52 no. 5 article id 10036. https://doi.org/10.14214/sf.10036
Keywords: regeneration; site preparation; clone effects; poplars; soil acidity
Highlights: Low pH (below 3.5) reduced growth, but not survival, in a greenhouse study; Site preparation methods did not affect survival in field, but differences were found for growth; Mounding had generally the best effect on growth; Clonal differences were found that could be useful for improving establishment and growth.
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Hybrid aspen (Populus tremula L. × P. tremuloides Michx.) is a deciduous tree species suitable for producing large amounts of renewable biomass during short rotations. Its potential under North European conditions could be largely extended if not only agricultural land but also forest land was used for cultivation. Unfortunately, the knowledge of appropriate forest site conditions and effects of site preparation methods on hybrid aspen establishment is limited. In this paper, two studies that explore these questions are presented. In the first study, the sensitivity to acid soils was tested under greenhouse conditions in two type of soils: a) peat soil limed to certain pH levels (3.4–5.7) and b) collected forest soils where pH varied from 3.9 to 5.3. The lowest pH level resulted in reduced growth, elsewhere no significant differences were found. The second study was applied in the field and investigated the effect of four site preparation methods on survival and growth. The methods were: 1) control with no site preparation, 2) patch scarification, 3) mounding and 4) soil inversion. While no differences were found for survival, mounding was generally the method with the highest growth and patch scarification was least successful. The result was probably an effect of good soil aeration and less competition from vegetation after mounding. The field study also revealed clonal differences in growth performance, which stresses the importance of clone selection prior to planting. The results of these studies indicate that hybrid aspen is less sensitive to variation in pH and site preparation methods compared with other poplar species, as have been found in similar studies.

  • Hjelm, Skogforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden E-mail: karin.hjelm@skogforsk.se (email)
  • Rytter, Skogforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden ORCID http://orcid.org/0000-0001-6183-4832 E-mail: lars.rytter@skogforsk.se
article id 10000, category Research article
Hamed Yousefzadeh, Rasta Rajaei, Anna Jasińska, Łukasz Walas, Yann Fragnière, Gregor Kozlowski. (2018). Genetic diversity and differentiation of the riparian relict tree Pterocarya fraxinifolia (Juglandaceae) along altitudinal gradients in the Hyrcanian forest (Iran). Silva Fennica vol. 52 no. 5 article id 10000. https://doi.org/10.14214/sf.10000
Keywords: plant variation; Caucasian wingnut; population genetics; refugium; riparian forest; river system
Highlights: The Caucasian wingnut (Pterocarya fraxinifolia Spach) is an emblematic and relict riparian tree with limited distribution in Hyrcanian forest which investigating its genetic population structure and diversity along altitudinal gradients, and migration patterns are novel; We concluded that rivers are the main seed dispersal vector among P. fraxinifolia populations and there was no trend from upstream to downstream; The high level of gene flow and uniform genetic diversity along each river system suggest the “classical” metapopulation structure of the species.
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Riparian trees, especially relict trees, are attractive and important for research to understand both past and recent biogeographical and evolutionary processes. Our work is the first study to elucidate the genetic diversity and spatial genetic structure of the canopy-dominating riparian Pterocarya fraxinifolia (Juglandaceae) along two altitudinal gradients in different river systems of the Hyrcanian forest, which is one of the most important refugium of relict trees in Western Eurasia. Altitudinal gradients were chosen along two river systems at 100, 400 and 900 m a.s.l. Leaf samples were collected from 116 trees, and the genetic diversity was evaluated with eight SSR markers. Overall, 39 alleles were identified for all of the populations studied. The observed heterozygosity (Ho) varied from 0.79 to 0.87 (with a mean of 0.83). The results of the AMOVA analysis indicated that the variation within populations was 88%, whereas the variation among populations was 12% for all of the gradients. A structure analysis indicated that 93% of the trees were grouped in the same gradient. The genetic distance based on Fst confirmed the structure result and indicated a high rate of gene flow among the investigated populations. Based on high gene flow (low differentiation of the population along the same river) and the clearly distinct genetic structure of the investigated gradients, it can be concluded that rivers are the main seed dispersal vector among P. fraxinifolia populations. The genetic diversity of P. fraxinifolia did not show any trend from upstream to downstream. The high level of gene flow and uniform genetic diversity along each river suggest the “classical” metapopulation structure of the species.

  • Yousefzadeh, Department of Environmental Science, Faculty of Natural Resources and Marine Sciences, Tarbiat Modares University, Noor, Mazandaran, Iran E-mail: h.yousefzadeh@modares.ac.ir (email)
  • Rajaei, Department of Forestry, Faculty of Natural Resources and Marine Sciences, Tarbiat Modares University, Noor, Mazandaran, Iran E-mail: r.rajaei@modares.ac.ir
  • Jasińska, Laboratory of Systematics and Geography, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, PL-62-035 Kornik, Poland E-mail: jasiak9@wp.pl
  • Walas, Laboratory of Systematics and Geography, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, PL-62-035 Kornik, Poland E-mail: lukaswalas@wp.pl
  • Fragnière, Department of Biology and Botanic Garden, University of Fribourg, Chemin du Musée 10, CH-1700 Fribourg, Switzerland E-mail: yann.fragniere@unifr.ch
  • Kozlowski, Department of Biology and Botanic Garden, University of Fribourg, Chemin du Musée 10, CH-1700 Fribourg, Switzerland; Natural History Museum Fribourg, Chemin du Musée 6, CH-1700 Fribourg, Switzerland E-mail: gregor.kozlowski@unifr.ch
article id 9987, category Research article
Monika Dering, Katarzyna Sękiewicz, Grzegorz Iszkuło, Aleksandra Chojnacka, Dominik Tomaszewski, Emilia Pers-Kamczyc, Piotr Karolewski. (2018). Spatial genetic structure and clonal structure of Prunus serotina during invasive spread. Silva Fennica vol. 52 no. 3 article id 9987. https://doi.org/10.14214/sf.9987
Keywords: genetic structure; invasiveness; black cherry; colonization; clonal structure
Highlights: The spread of Prunus serotina in invaded forests is facilitated by high propagule pressure; The seed shadow overlap prevents strong spatial genetic structure; During colonization of isolated site, a strong spatial genetic structure is produced due to founder effect; Overall clonality in P. serotina was low but may efficiently support seedling bank thus contributing to species invasiveness.
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Prunus serotina Ehrh. (black cherry) is one of the most important invaders in the European forests, but existing studies have given limited insight into demo-genetic factors underpinning the process of species invasion. Fine-scale genetic structure (FSGS) may deliver important knowledge on genetics of invasion contributing to efficient management of the alien species. Using eight microsatellites we investigated FSGS, clonal structure and relatedness in four black cherry populations which represented different stages of the invasive spread into Scots pine forests. Three populations were in a continuous forest complex and represented the colonization (Z_1) and established stages (Z_2 and Z_3). To investigate how colonization ability of the species is modified by landscape features, we analyzed an isolated population at colonization stage located in limited forest patch located in an agricultural landscape (R). Populations from continuous forest showed low yet significant positive FSGS with Sp = 0.0068 in Z_1, 0.0054 in Z_2, and 0.0066 in Z_3, while in R spatial structure was the strongest (0.0145). Considerable relatedness noted in population R suggests a dominance of within-population mating and recruitments, low immigration rate and limited seed dispersal, all of which led to the observed strong FSGS. Also, we presume that a founder effect likely involved during colonization of isolated forest patch R led to strong FSGS. In contrary, the seed shadow overlap in the populations from continuous forest prevented strong FSGS and facilitated colonization. Despite of low level of clonality, we argue that it may efficiently support black cherry seedling bank contributing to species invasiveness.

  • Dering, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland E-mail: mdering@man.poznan.pl (email)
  • Sękiewicz, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland E-mail: ksekiewicz@man.poznan.pl
  • Iszkuło, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland; Faculty of Biological Sciences, University of Zielona Góra, Prof. Z. Szafrana 1, 65-516 Zielona Góra, Poland E-mail: iszkulo@man.poznan.pl
  • Chojnacka, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland E-mail: jagoda900@gmail.com
  • Tomaszewski, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland E-mail: dominito@man.poznan.pl
  • Pers-Kamczyc, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland E-mail: epk@man.poznan.pl
  • Karolewski, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland E-mail: pkarolew@man.poznan.pl
article id 9946, category Research article
Frauke Fedderwitz, Niklas Björklund, Velemir Ninkovic, Göran Nordlander. (2018). Does the pine weevil (Hylobius abietis) prefer conifer seedlings over other main food sources? Silva Fennica vol. 52 no. 3 article id 9946. https://doi.org/10.14214/sf.9946
Keywords: Pinus sylvestris; Picea abies; forest regeneration; feeding preference; herbivore; plant tissues
Highlights: Adult pine weevils feed on seedlings and mature conifers, but cause economic damage only on seedlings; Their feeding preferences for branches and roots over seedlings were tested in a laboratory experiment; The only clear preference was for Norway spruce roots; Results support new approaches of seedling protection attempting to redirect pine weevils from planted seedlings to other food sources.
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Adult pine weevils (Hylobius abietis (L.)) feed on the tender bark of branches and roots of mature conifer trees and on the stem bark of conifer seedlings. Their feeding on mature trees does not cause any economic damage, but their feeding on planted seedlings is so devastating that the pine weevil is considered one of the most important forest pest insects in Europe. We asked whether the pine weevil prefers seedlings over other regularly utilized food sources. This question is of particular interest because new approaches to seedling protection are based on decreasing any preference for seedlings by using less palatable plants or by enhancing their defence (by genetic selection or by methyl jasmonate treatment). In a laboratory choice experiment we tested pine weevil feeding preferences for seedlings compared with branches and roots from mature trees (separately for Norway spruce and Scots pine). Pine weevils preferred roots, but not branches, of Norway spruce over seedlings of the same species. With Scots pine there were no clear preferences, but the weevils showed a tendency to prefer roots over seedlings. These results provide support for seedling protection approaches that attempt to redirect pine feeding from planted seedlings to other food sources.

  • Fedderwitz, Department of Ecology, Swedish University of Agricultural Sciences, Box 7044, SE-750 07 Uppsala, Sweden E-mail: frauke.fedderwitz@slu.se (email)
  • Björklund, Department of Ecology, Swedish University of Agricultural Sciences, Box 7044, SE-750 07 Uppsala, Sweden E-mail: niklas.bjorklund@slu.se
  • Ninkovic, Department of Ecology, Swedish University of Agricultural Sciences, Box 7044, SE-750 07 Uppsala, Sweden E-mail: velemir.ninkovic@slu.se
  • Nordlander, Department of Ecology, Swedish University of Agricultural Sciences, Box 7044, SE-750 07 Uppsala, Sweden E-mail: goran.nordlander@slu.se
article id 9933, category Research article
Timo Saksa, Jari Miina, Hilkka Haatainen, Kauko Kärkkäinen. (2018). Quality of spot mounding performed by continuously advancing mounders. Silva Fennica vol. 52 no. 2 article id 9933. https://doi.org/10.14214/sf.9933
Keywords: soil preparation; slash; logging residues; forest regeneration; mixed modelling
Highlights: The number and quality of mounds varied considerably according to the operating conditions;The main factors reducing the quality of spot mounding were steep terrain, a thick humus layer, fresh logging residues, stoniness and soil texture;With careful selection of timing and conditions for mounding, the quality obtained by continuously advancing mounders can be improved.
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Operating conditions affecting the quality of spot mounding by Bracke continuously advancing mounders were investigated on 66 regeneration areas (124 ha) in eastern Finland. The quality of mounds was classified as suitable (good or acceptable after additional compression) or unsuitable for planting. Models were constructed for the number of suitable planting spots obtained per hectare (good and acceptable mounds), the probability of successful mounding (≥1600 planting spots ha–1) and the probability of creating a suitable mound as a function of terrain, site and soil characteristics, as well as slash conditions (removed, fresh or dry logging residues). The average number of mounds created was 1892 ± 290 mounds ha–1, of which 1398 ± 325 mounds ha–1 (74%) were classified as suitable for planting. The quality of spot mounding was reduced by steep terrain, a thick humus layer and fresh logging residues. Stoniness and soil texture also affected the number of planting spots created. Mounding after logging residues had dried increased the number of planting spots by 191 spots ha–1 compared with mounding in the presence of fresh residues. Removing residues did not significantly increase the number of planting spots compared with mounding amongst dry residues. A thick humus layer, very stony soil, steep slopes and valley terrain decreased the number of planting spots by 150–450 spots ha–1. The number and quality of mounds varied considerably according to the operating conditions, but with careful selection of timing and sites the quality obtained by a continuously advancing mounder can be improved.

  • Saksa, Natural Resources Institute Finland (Luke), Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@luke.fi (email)
  • Miina, Natural Resources Institute Finland (Luke), Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: jari.miina@luke.fi
  • Haatainen, Faculty of Science and Forestry, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: hilkka.haatainen@storaenso.com
  • Kärkkäinen, Tornator Oyj, Muuntamontie 2, FI-80100 Joensuu, Finland E-mail: kauko.karkkainen@tornator.fi
article id 7738, category Research article
Samuel Egbäck, Bo Karlsson, Karl-Anders Högberg, Kenneth Nyström, Mateusz Liziniewicz, Urban Nilsson. (2018). Effects of phenotypic selection on height-diameter ratio of Norway spruce and Scots pine in Sweden. Silva Fennica vol. 52 no. 2 article id 7738. https://doi.org/10.14214/sf.7738
Keywords: Pinus sylvestris; Picea abies; genetic correlations; heritability; Genetic selection; slenderness
Highlights: Swedish plus-tree selection promoted less slender Norway spruce trees and more slender Scots pine trees compared to neighboring trees; Similar results were also found for progeny trials which indicated that genetics played a prominent role in phenotypic appearance.
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Genetically improved Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) are extensively used in operational Swedish forestry plantations. However, relatively little is known about the stem slenderness (height-diameter ratio) of genetically improved material. Thus, in this study we investigated effects of plus-tree selection on stem slenderness of Norway spruce and Scots pine in Sweden by evaluating both the plus-tree selection and a large number of progeny trials. Species-specific models for predicting the height-diameter ratio were estimated using regression and mixed model approach. Our results show that phenotypic plus-tree selection promoted less slender Norway spruce trees and more slender Scots pine trees compared to neighboring trees. Similar results were also found for the progeny trials which indicated that genetics played a prominent role in the phenotypic appearance. Compared to the progeny of neighboring trees, Norway spruce plus-tree progenies had a 5.3% lower height-diameter ratio, while Scots pine plus-tree progenies had a 1.5% greater height-diameter ratio. The narrow sense heritability for height-diameter ratio was 0.19 for Norway spruce and 0.11 for Scots pine, indicating that it is possible to modify the height-diameter ratio by breeding. Correlation coefficients between breeding values for height-diameter ratio and diameter were negative for Scots pine (–0.71) and Norway spruce (–0.85), indicating that selection for diameter only would result in less slender stems of both species. Similar correlations were also found between breeding values for height-diameter ratio and height of Scots pine (–0.34) and Norway spruce (–0.74).

  • Egbäck, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, 230 53 Alnarp, Sweden E-mail: samuel.egback@slu.se (email)
  • Karlsson, Skogforsk, Ekebo, 268 90 Svalöv, Sweden E-mail: bo.karlsson@skogforsk.se
  • Högberg, Skogforsk, Ekebo, 268 90 Svalöv, Sweden E-mail: karl-anders.hogberg@skogforsk.se
  • Nyström, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd, 901 83 Umeå, Sweden E-mail: kenneth.nystrom@slu.se
  • Liziniewicz, Skogforsk, Ekebo, 268 90 Svalöv, Sweden E-mail: Mateusz.Liziniewicz@skogforsk.se
  • Nilsson, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, 230 53 Alnarp, Sweden E-mail: urban.nilsson@slu.se
article id 1665, category Research article
Lauri Haataja, Ville Kankaanhuhta, Timo Saksa. (2018). Reliability of self-control method in the management of non-industrial private forests. Silva Fennica vol. 52 no. 1 article id 1665. https://doi.org/10.14214/sf.1665
Keywords: forest management; soil preparation; quality control; young stand management; forest regeneration; boreal silviculture; forest planting
Highlights: Self-control method was found reliable at the main stages of the forest regeneration process; Only slight overestimation was found in self-control results of soil preparation and planting and small underestimation in self-control of young stand management; Diverse utilizing of self-control data is possible in support of service providers operations.
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This study seeks to determine the extent to which self-control data can be relied upon in the management of private forests. Self-control (SC) requires the forest workers to evaluate their own work quality to ensure the clients’ needs are met in terms of soil preparation, planting and young stand management. Self-control data were compared to an independent evaluation of the same worksites. Each dataset had a hierarchical structure (e.g., sample plot, regeneration area and contractor), and key quality indicators (i.e., number of prepared mounds, planted seedlings or crop trees) were measured for each plot.  Self-control and independent-assessments (IA) were analyzed by fitting a multi-level multivariate model containing explanatory variables. No significant differences were observed in terms of soil preparation (number of mounds) or young stand management (number of crop trees) between self-control and independent-assessments. However, the self-control planting data included a slight but significant overestimation of the number of planted seedlings. Discrepancies are discussed in terms of sampling error and other explanatory factors. According to overall results, self-control methods are reliable at every stage of the forest regeneration process. As such, the diverse utilizing of self-control data is possible in support of service providers operations.
  • Haataja, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: lauritapiohaataja@gmail.com (email)
  • Kankaanhuhta, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: ville.kankaanhuhta@luke.fi
  • Saksa, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@luke.fi
article id 1760, category Research article
Marek Fajstavr, Kyriaki Giagli, Hanuš Vavrčík, Vladimír Gryc, Josef Urban. (2017). The effect of stem girdling on xylem and phloem formation in Scots pine. Silva Fennica vol. 51 no. 4 article id 1760. https://doi.org/10.14214/sf.1760
Keywords: Pinus sylvestris; radial increment; cambial activity; cell differentiation; phloemogenesis; xylogenesis
Highlights: Stem girdling ceased the cambial activity, below the girdled area, immediately after the removal of the bark strip; Pinus sylvestris survived for up to two years after stem girdling; The girdled trees formed phloem cells above the girdled area but failed to form latewood cells in the next growing season.
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Scots pine (Pinus sylvestris L.) is a resilient, wide spread species. This paper reports on the xylem and phloem cell formation process, before and after, the species was put under artificial stress by stem girdling. Microcore method was applied to a healthy control group and a standing group of girdled trees within an 80-year-old pine forest for two consecutive growing seasons (2013 and 2014). The stem girdling was applied in the middle of the first growing season (July 2013). Cambial activity timings (onset and cessation of cell division), cell formation intensity, cell differentiation, and the dynamics of the annual radial increment in the stem were analyzed. Cambial activity was inhibited and eventually ceased below the stem girdling immediately after the removal of the strip. Therefore, no latewood tracheids were formed. However, above the stem girdling and in the control trees, cell formation and tissue differentiation continued until the end of the growing season, with the girdled trees moving at a less intensive pace but for a longer period of time. During the following growing season (2014), the cambial zone was reactivated only above the stem girdling, not below, and eventually the girdled trees died. In 2014, the onset of the cambial activity was delayed and the division rate of the cells was slower in the girdled trees. Furthermore, the girdled trees formed less phloem cells than the control trees.

  • Fajstavr, Department of Wood Science, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemedelska 3, 613 00 Brno, Czech Republic E-mail: fajstavr.marek@seznam.cz (email)
  • Giagli, Department of Wood Science, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemedelska 3, 613 00 Brno, Czech Republic E-mail: kyriaki.giagli@mendelu.cz
  • Vavrčík, Department of Wood Science, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemedelska 3, 613 00 Brno, Czech Republic E-mail: vavrcik@mendelu.cz
  • Gryc, Department of Wood Science, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemedelska 3, 613 00 Brno, Czech Republic E-mail: gryc@mendelu.cz
  • Urban, Department of Forest Botany, Dendrology and Geobiocenology, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemedelska 3, 613 00 Brno, Czech Republic;  Siberian Federal University, Svobodnyj Prospect 79, Krasnoyarsk, 660041 Krasnoyarsk, Russia E-mail: josef.urban@email.cz
article id 1691, category Research article
Enéas Ricardo Konzen, Raquel Peron, Márcio Akira Ito, Gilvano Ebling Brondani, Siu Mui Tsai. (2017). Molecular identification of bamboo genera and species based on RAPD-RFLP markers. Silva Fennica vol. 51 no. 4 article id 1691. https://doi.org/10.14214/sf.1691
Keywords: molecular markers; molecular screening; restriction enzymes; genetic divergence; Dendrocalamus; Bambusa
Highlights: We investigated the potential of RAPD-RFLP (Random Amplified Polymorphic DNA, Restriction Fragment Length Polymorphism) in detecting genetic relationships among bamboo genera and species; RFLP profiles resolved in acrylamide gels revealed high number of markers, which accurately differentiated species and genera, based on cophenetic correlation coefficients; We recommend RAPD-RFLP for analyses of genetic diversity and divergence among bamboo genera, species and varieties.
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Bamboo species have a very significant ecological and economic impact. Determining morphological and genetic differences among bamboo genera and species are crucial to explore desirable traits for breeding purposes. Several advances have been made in the taxonomy of bamboos by using molecular fingerprinting tools and next generation sequencing technologies. Nevertheless, classical molecular markers such as RAPD (Random Amplified Polymorphic DNA), AFLP (Amplified Fragment Length Polymorphism) and ISSR (Inter Simple Sequence Repeats) also provide an accurate discrimination among genera and species. Moreover, the RAPD-RFLP (Random Amplified Polymorphic DNA, Restriction Fragment Length Polymorphism) method, in which amplification products from RAPD are digested with restriction enzymes, is a reliable, fast and cost-effective method for fingerprinting. RAPD-RFLP has been scarcely used in the literature and no report regarding bamboo taxonomy is available with this method. Here we explored the molecular (RAPD, RAPD-RFLP) variation among genera (Bambusa, Dendrocalamus, Guadua and Phyllostachys) and species of bamboo cultivated in Brazil. Both molecular markers allowed clear distinction among the genera studied. Moreover, high cophenetic correlation values in UPGMA clusters indicated their potential for discriminating bamboo species. The digestion of RAPD products (RFLP) resulted in high number of polymorphic bands and produced very characteristic profiles for each genus with three enzyme combinations (HindIII/HaeIII, HinfI/RsaI, and single digestion with MspI). We recommend RAPD-RFLP as a reproducible and informative method for screening differences among genera, species and varieties of bamboos. Providing a cost-effective and accurate method for species identification and characterization is straightforward for bamboo conservation, management and breeding.

  • Konzen, Cell and Molecular Biology Laboratory, Center for Nuclear Energy in Agriculture (CENA), University of Sao Paulo (USP), Centenário Av., 303, Piracicaba, SP, P.O. Box 96, Brazil; Forest Sciences, Federal University of Lavras (UFLA), Lavras, MG, P.O. Box 3037, Brazil E-mail: erkonzen@gmail.com (email)
  • Peron, Cell and Molecular Biology Laboratory, Center for Nuclear Energy in Agriculture (CENA), University of Sao Paulo (USP), Centenário Av., 303, Piracicaba, SP, P.O. Box 96, Brazil E-mail: rperon@purdue.edu
  • Ito, Embrapa Western Agriculture, Brazilian Agricultural Research Corporation (EMBRAPA), BR 163 Rd., km 253, Dourados, MS, P.O. Box 449, Brazil E-mail: marcio.ito@embrapa.br
  • Brondani, Forest Sciences, Federal University of Lavras (UFLA), Lavras, MG, P.O. Box 3037, Brazil E-mail: gebrondani@gmail.com
  • Tsai, Cell and Molecular Biology Laboratory, Center for Nuclear Energy in Agriculture (CENA), University of Sao Paulo (USP), Centenário Av., 303, Piracicaba, SP, P.O. Box 96, Brazil E-mail: tsai@cena.usp.br
article id 7001, category Research article
Fei-Fei Zhai, Jun-Xiang Liu, Zhen-Jian Li, Jin-Mei Mao, Yong-Qiang Qian, Lei Han, Zhen-Yuan Sun. (2017). Assessing genetic diversity and population structure of Salix viminalis across Ergun and West Liao basin. Silva Fennica vol. 51 no. 3 article id 7001. https://doi.org/10.14214/sf.7001
Keywords: genetic variation; microsatellites; geographic barrier; basket willow; common osier
Highlights: High genetic diversity and moderate population differentiation were detected in twelve Salix viminalis populations; Populations from Ergun basin presented higher genetic diversity than populations from West Liao basin; Populations with short distances presented low differentiation.
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Salix viminalis L. is an important shrub that has potential for use as a bioenergy crop, for phytoremediation of heavy metal contaminated soil and sewage sludge treatment. It is mainly distributed in the northeast of China, but the species has not yet been used a resource here. We examined the genetic diversity and population structure of populations from the Ergun basin and West Liao basin using 20 microsatellite markers. A high level of genetic diversity (Na = 16.45, He = 0.742) was detected for S. viminalis, and populations from the Ergun basin exhibited higher genetic diversity and private alleles numbers than the West Liao basin. The 12 populations could be divided into two clusters by both Bayesian analysis and UPGMA clustering which were consistent with the populations derived from the two basins. Moderate population differentiation (FST = 0.076) was shown in S. viminalis, and AMOVA analysis confirmed that most of the genetic variation (86.13%) was attributed to individual differences within populations, while 11.49% was attributed to differences between basins and 2.38% to differences within each basin. Significant correlations of FST/(1–FST) with log (geographic distance) among 12 populations (r = 0.634, p < 0.00) and 10 populations within the Ergun basin (r = 0.482, p = 0.0002) indicated that geographical distance was the principal factor influencing genetic structure. As most of genetic variation exist within populations, so protection measures should be focused on populations with higher genetic diversity and unique alleles, such as Tuli, Mordaga downstream, Zhadun1 and Genhe.

  • Zhai, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China; School of Architectural and Artistic Design, Henan Polytechnic University, Century Avenue, Jiaozuo, Henan, 454000, PR China E-mail: lkyzff@163.com
  • Liu, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China E-mail: liu-jx295@163.com
  • Li, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China E-mail: zhenjianli@163.com
  • Mao, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China; Research Institute of Economic Forest, Xinjiang Academy of Forestry, Anjunanlu, Urumqi, Xinjiang, PR China E-mail: 350512173@qq.com
  • Qian, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China E-mail: qianyq@caf.ac.cn
  • Han, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China E-mail: hdd@caf.ac.cn
  • Sun, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China E-mail: lkyszy@126.com (email)
article id 5662, category Research article
Samuel Egbäck, Urban Nilsson, Kenneth Nyström, Karl-Anders Högberg, Nils Fahlvik. (2017). Modeling early height growth in trials of genetically improved Norway spruce and Scots pine in southern Sweden. Silva Fennica vol. 51 no. 3 article id 5662. https://doi.org/10.14214/sf.5662
Keywords: Pinus sylvestris; Picea abies; individual tree growth model; genetic component; genetic multiplier; unimproved material; improved material
Highlights: The developed height growth model based on unimproved material predicted the development relatively well for genetically improved Norway spruce; For genetically improved Scots pine, however, the model needed to be modified; By incorporating a genetic component into the Scots pine model, the prediction errors were reduced.
Abstract | Full text in HTML | Full text in PDF | Author Info

Genetically improved Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) are used extensively in operational Swedish forestry plantations to increase production. Depending on the genetic status of the plant material, the current estimated genetic gain in growth is in the range 10–20% for these species and this is expected to increase further in the near future. However, growth models derived solely from data relating to genetically improved material in Sweden are still lacking. In this study we investigated whether an individual tree growth model based on data from unimproved material could be used to predict the height increment in young trials of genetically improved Norway spruce and Scots pine. Data from 11 genetic experiments with large genetic variation, ranging from offspring of plus-trees selected in the late 1940s to highly improved clonal materials selected from well performing provenances were used. The data set included initial heights at the age of 7–15 years and 5-year increments for almost 2000 genetic entries and more than 20 000 trees. The evaluation indicated that the model based on unimproved trees predicted height development relatively well for genetically improved Norway spruce and there was no need to incorporate a genetic component. However, for Scots pine, the model needed to be modified. A genetic component was developed based on the genetic difference recorded within each trial, using mixed linear models and methods from quantitative genetics. By incorporating the genetic component, the prediction errors were significantly reduced for Scots pine. This study provides