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Category : Climate resilient and sustainable forest management – Editorial

article id 25024, category Climate resilient and sustainable forest management – Editorial
Annamari Laurén. (2025). Forest management is rocket science. Silva Fennica vol. 59 no. 1 article id 25024. https://doi.org/10.14214/sf.25024
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  • Laurén, Department of Forest Sciences, University of Helsinki, P.O. Box 27, 00014 University of Helsinki, Finland; Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland ORCID https://orcid.org/0000-0002-6835-9568 E-mail: annamari.lauren@helsinki.fi (email)

Category : Editorial

article id 25026, category Editorial
Pekka Nygren. (2025). Changes in the editorial staff of Silva Fennica. Silva Fennica vol. 59 no. 1 article id 25026. https://doi.org/10.14214/sf.25026
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article id 24069, category Editorial
Matti Maltamo. (2024). “The Final Cut”. Silva Fennica vol. 58 no. 5 article id 24069. https://doi.org/10.14214/sf.24069
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article id 24058, category Editorial
Henrik Heräjärvi. (2024). Wood is much more than just carbon. Silva Fennica vol. 58 no. 4 article id 24058. https://doi.org/10.14214/sf.24058
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article id 24034, category Editorial
Hanne K. Sjølie. (2024). Normative bias in research. Silva Fennica vol. 58 no. 3 article id 24034. https://doi.org/10.14214/sf.24034
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  • Sjølie, Inland Norway University of Applied Sciences, Faculty of Applied Ecology, Agricultural Sciences and Biotechnology, Department of Forestry and Wildlife Management, Campus Evenstad, Postboks 400, 2418 Elverum, Norway E-mail: sjoliehannek3@gmail.com
article id 24020, category Editorial
Matti Maltamo. (2024). What we pay attention to when we are in the forest? Silva Fennica vol. 58 no. 2 article id 24020. https://doi.org/10.14214/sf.24020
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article id 24016, category Editorial
Pekka Nygren. (2024). Advancing the Everyone’s Right to Forest Science: opening research data and computational codes in Silva Fennica. Silva Fennica vol. 58 no. 1 article id 24016. https://doi.org/10.14214/sf.24016
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article id 23074, category Editorial
Christian Kanzian. (2023). Are productivity studies in forest operations old fashioned and no more publishable? Silva Fennica vol. 57 no. 3 article id 23074. https://doi.org/10.14214/sf.23074
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  • Kanzian, Department of Forest and Soil Sciences, Institute of Forest Engineering, University of Natural Resources and Life Sciences, Vienna (BOKU), Peter-Jordan-Strasse 82, 1190 Vienna, Austria ORCID https://orcid.org/0000-0002-1198-9788 E-mail: christian.kanzian@boku.ac.at
article id 23043, category Editorial
Anna Lintunen. (2023). Passion for science. Silva Fennica vol. 57 no. 2 article id 23043. https://doi.org/10.14214/sf.23043
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  • Lintunen, Institute for Atmospheric and Earth System Research / Forest Sciences, University of Helsinki, Viikinkaari 1, Biokeskus 3, 00790 Helsinki, Finland ORCID https://orcid.org/0000-0002-1077-0784 E-mail: anna.lintunen@helsinki.fi
article id 23011, category Editorial
Sergio de Miguel. (2023). Artificial intelligence-driven disruption in science production ahead. Silva Fennica vol. 57 no. 1 article id 23011. https://doi.org/10.14214/sf.23011
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  • de Miguel, University of Lleida, Av. Alcalde Rovira Roure 191, 25198 Lleida, Spain; Joint Research Unit CTFC – AGROTECNIO – CERCA, Ctra de Sant Llorenç de Morunys, km 2, 25280 Solsona, Spain ORCID https://orcid.org/0000-0002-9738-0657 E-mail: sergio.demiguel@udl.cat
article id 23005, category Editorial
Matti Maltamo. (2023). What does it actually mean to measure a sample plot in forest? Silva Fennica vol. 56 no. 4 article id 23005. https://doi.org/10.14214/sf.23005
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  • Maltamo, University of Eastern Finland, Faculty of Science, Forestry and Technology, Joensuu ORCID 0000-0002-9904-3371 E-mail: matti.maltamo@uef.fi
article id 10790, category Editorial
Henrik Heräjärvi. (2022). What do engineered wood products and Ford Model T have in common? Silva Fennica vol. 56 no. 3 article id 10790. https://doi.org/10.14214/sf.10790
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  • Heräjärvi, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: henrik.herajarvi@luke.fi (email)
article id 10763, category Editorial
Matti Maltamo. (2022). Silva Fennica has improved publishing services by changing manuscript handling system. Silva Fennica vol. 56 no. 2 article id 10763. https://doi.org/10.14214/sf.10763
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  • Maltamo, University of Eastern Finland, School of Forest Sciences, Joensuu E-mail: matti.maltamo@uef.fi (email)
article id 10711, category Editorial
Matti Maltamo. (2022). The persistently developing role of remote sensing in forest sciences. Silva Fennica vol. 56 no. 1 article id 10711. https://doi.org/10.14214/sf.10711
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  • Maltamo, University of Eastern Finland, School of Forest Sciences, Joensuu E-mail: matti.maltamo@uef.fi (email)
article id 10682, category Editorial
Pekka Nygren. (2021). What is similarity check? Silva Fennica vol. 55 no. 5 article id 10682. https://doi.org/10.14214/sf.10682
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  • Nygren, Finnish Society of Forest Science, Viikinkaari 6, FI-00790 Helsinki, Finland E-mail: pekka.nygren@metsatiede.org (email)
article id 10643, category Editorial
Matti Maltamo. (2021). 100 years of national forest inventories. Silva Fennica vol. 55 no. 4 article id 10643. https://doi.org/10.14214/sf.10643
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  • Maltamo, University of Eastern Finland, School of Forest Sciences, Joensuu E-mail: matti.maltamo@uef.fi (email)
article id 10604, category Editorial
Samuel F. Bartels. (2021). The deciduous confusion: it is time to get the contrast right. Silva Fennica vol. 55 no. 3 article id 10604. https://doi.org/10.14214/sf.10604
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  • Bartels, Department of Ecosystem Science and Management, University of Norther British Columbia, 3333 University Way, Prince George, British Columbia, Canada V2N 4Z9 E-mail: samuel.bartels@unbc.ca (email)
article id 10566, category Editorial
Henrik Heräjärvi. (2021). Science as a decision-support tool in forest policies. Silva Fennica vol. 55 no. 2 article id 10566. https://doi.org/10.14214/sf.10566
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  • Heräjärvi, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: henrik.herajarvi@luke.fi (email)
article id 10542, category Editorial
Jussi Manner. (2021). What is (not) an operator effect in forest work science? Silva Fennica vol. 55 no. 1 article id 10542. https://doi.org/10.14214/sf.10542
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article id 10501, category Editorial
Hanne K. Sjølie. (2020). Why are there still too few women in forestry and science? Silva Fennica vol. 54 no. 5 article id 10501. https://doi.org/10.14214/sf.10501
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  • Sjølie, Inland Norway University of Applied Sciences, Postboks 400, 2418 Elverum, Norway E-mail: hanne.sjolie@inn.no (email)
article id 10452, category Editorial
Matti Maltamo. (2020). Re-searching the forests. Silva Fennica vol. 54 no. 4 article id 10452. https://doi.org/10.14214/sf.10452
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  • Maltamo, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: matti.maltamo@uef.fi (email)
article id 10411, category Editorial
Pekka Nygren. (2020). Improvement of reproducibility and research transparency in Silva Fennica. Silva Fennica vol. 54 no. 3 article id 10411. https://doi.org/10.14214/sf.10411
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  • Nygren, Finnish Society of Forest Science, Viikinkaari 6, FI-00790 Helsinki, Finland E-mail: pekka.nygren@metsatiede.org (email)
article id 10362, category Editorial
Svetlana Saarela. (2020). On being a female supervisor in forest research education. Silva Fennica vol. 54 no. 2 article id 10362. https://doi.org/10.14214/sf.10362
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  • Saarela, Department of Forest Resource Management, Swedish University of Agricultural Sciences, Box 7044, SE-750 07 Uppsala, Sweden ORCID https://orcid.org/0000-0002-9044-7249 E-mail: svetlana.saarela@slu.se (email)
article id 10333, category Editorial
Matti Maltamo. (2020). Change of the Subject Editor in Silva Fennica. Silva Fennica vol. 54 no. 1 article id 10333. https://doi.org/10.14214/sf.10333
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  • Maltamo, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: matti.maltamo@uef.fi (email)
article id 10329, category Editorial
Lars Eliasson. (2020). Forest research and the needs of our stakeholders. Silva Fennica vol. 54 no. 1 article id 10329. https://doi.org/10.14214/sf.10329
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article id 10285, category Editorial
Nicole J. Fenton. (2019). The introduction, the link between applied research and a successful publication. Silva Fennica vol. 53 no. 4 article id 10285. https://doi.org/10.14214/sf.10285
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  • Fenton, Université du Québec en Abitibi-Temiscamingue, Institute for Forest Research (IFR) at UQAT, Rouyn-Noranda (Québec), Canada E-mail: nicole.fenton@uqat.ca (email)
article id 10257, category Editorial
Lauri Mehtätalo. (2019). Reporting modern statistical analyses: reproducible and transparent. Silva Fennica vol. 53 no. 3 article id 10257. https://doi.org/10.14214/sf.10257
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  • Mehtätalo, University of Eastern Finland, Faculty of Science and Forestry, School of Computing, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: lauri.mehtatalo@uef.fi (email)
article id 10216, category Editorial
Henrik Heräjärvi. (2019). New age of discovery in wood science. Silva Fennica vol. 53 no. 2 article id 10216. https://doi.org/10.14214/sf.10216
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  • Heräjärvi, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: henrik.herajarvi@luke.fi (email)
article id 10164, category Editorial
Matti Maltamo. (2019). Silva Fennica in 2019. Silva Fennica vol. 53 no. 1 article id 10164. https://doi.org/10.14214/sf.10164
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  • Maltamo, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: matti.maltamo@uef.fi (email)
article id 10122, category Editorial
Eeva Korpilahti. (2018). Silva Fennica’s latest quarter centennial publishing. Silva Fennica vol. 52 no. 5 article id 10122. https://doi.org/10.14214/sf.10122
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  • Korpilahti, Finnish Society of Forest Science, P.O. Box 65, FI-00790 Helsinki, Finland E-mail: eeva.korpilahti@metsatiede.org (email)
article id 443, category Editorial
Eeva Korpilahti. (2012). From the Editor. Silva Fennica vol. 46 no. 1 article id 443. https://doi.org/10.14214/sf.443
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  • Korpilahti, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: eeva.korpilahti@luke.fi (email)

Category : Special section

article id 472, category Special section
Raisa Mäkipää, Jari Liski, Mats Olsson, Pete Smith, Esther Thürig. (2007). Workshop on Development of Models and Forest Soil Surveys for Monitoring of Soil Carbon. Silva Fennica vol. 41 no. 3 article id 472. https://doi.org/10.14214/sf.472
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Selected Papers of the Workshop on Development of Models and Forest Soil Surveys for Monitoring of Soil Carbon.
  • Mäkipää, Finnish Forest Research Institute E-mail: rm@nn.fi (email)
  • Liski, Finnish Environment Institute, Finland E-mail: jl@nn.fi
  • Olsson, Swedish University of Agricultural Sciences, Sweden E-mail: mo@nn.se
  • Smith, University of Aberdeen, UK E-mail: ps@nn.uk
  • Thürig, Swiss Federal Research Institute WSL, Switzerland E-mail: et@nn.ch
article id 290, category Special section
Mikko Peltoniemi, Esther Thürig, Stephen Ogle, Taru Palosuo, Marion Schrumpf, Thomas Wutzler, Klaus Butterbach-Bahl, Oleg Chertov, Alexander Komarov, Aleksey Mikhailov, Annemieke Gärdenäs, Charles Perry, Jari Liski, Pete Smith, Raisa Mäkipää. (2007). Models in country scale carbon accounting of forest soils. Silva Fennica vol. 41 no. 3 article id 290. https://doi.org/10.14214/sf.290
Keywords: National Forest Inventory; soil carbon; greenhouse gas inventory; decomposition; IPCC; regional and national modeling; soil model
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Countries need to assess changes in the carbon stocks of forest soils as a part of national greenhouse gas (GHG) inventories under the United Nations Framework Convention on Climate Change (UNFCCC) and the Kyoto Protocol (KP). Since measuring these changes is expensive, it is likely that many countries will use alternative methods to prepare these estimates. We reviewed seven well-known soil carbon models from the point of view of preparing country-scale soil C change estimates. We first introduced the models and explained how they incorporated the most important input variables. Second, we evaluated their applicability at regional scale considering commonly available data sources. Third, we compiled references to data that exist for evaluation of model performance in forest soils. A range of process-based soil carbon models differing in input data requirements exist, allowing some flexibility to forest soil C accounting. Simple models may be the only reasonable option to estimate soil C changes if available resources are limited. More complex models may be used as integral parts of sophisticated inventories assimilating several data sources. Currently, measurement data for model evaluation are common for agricultural soils, but less data have been collected in forest soils. Definitions of model and measured soil pools often differ, ancillary model inputs require scaling of data, and soil C measurements are uncertain. These issues complicate the preparation of model estimates and their evaluation with empirical data, at large scale. Assessment of uncertainties that accounts for the effect of model choice is important part of inventories estimating large-scale soil C changes. Joint development of models and large-scale soil measurement campaigns could reduce the inconsistencies between models and empirical data, and eventually also the uncertainties of model predictions.
  • Peltoniemi, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: mikko.peltoniemi@metla.fi (email)
  • Thürig, Swiss Federal Institute for Forest, Snow and Landscape Research (WSL), Birmensdorf, Switzerland; European Forest Institute, Joensuu, Finland E-mail: et@nn.ch
  • Ogle, Natural Resources Ecology Laboratory, Colorado State University, Fort Collins, USA E-mail: so@nn.us
  • Palosuo, European Forest Institute, Joensuu, Finland E-mail: tp@nn.fi
  • Schrumpf, Max-Planck-Institute for Biogeochemistry, Jena, Germany E-mail: ms@nn.de
  • Wutzler, Max-Planck-Institute for Biogeochemistry, Jena, Germany E-mail: tw@nn.de
  • Butterbach-Bahl, Institute for Meteorology and Climate Research, Forschungszentrum Karlsruhe GmbH, Garmisch-Partenkirchen, Germany E-mail: kbb@nn.de
  • Chertov, St. Petersburg State University, St. Petersburg-Peterhof, Russia E-mail: oc@nn.ru
  • Komarov, Institute of Physicochemical and Biological Problems in Soil Science of Russian Academy of Sciences, Pushchino, Russia E-mail: ak@nn.ru
  • Mikhailov, Institute of Physicochemical and Biological Problems in Soil Science of Russian Academy of Sciences, Pushchino, Russia E-mail: am@nn.ru
  • Gärdenäs, Dept. of Soil Sciences, Swedish University of Agricultural Sciences, Uppsala, Sweden E-mail: ag@nn.se
  • Perry, USDA Forest Service, Northern Research Station, St. Paul, MN USA E-mail: cp@nn.us
  • Liski, Finnish Environment Institute, Helsinki, Finland E-mail: jl@nn.fi
  • Smith, School of Biological Sciences, University of Aberdeen, Aberdeen, UK E-mail: ps@nn.uk
  • Mäkipää, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: raisa.makipaa@metla.fi
article id 289, category Special section
Thomas Wutzler, Martina Mund. (2007). Modelling mean above and below ground litter production based on yield tables. Silva Fennica vol. 41 no. 3 article id 289. https://doi.org/10.14214/sf.289
Keywords: uncertainty; thinnings; soil carbon; biomass turnover; carbon accounting; harvest residues; sensitivity analysis
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Estimates of litter production are a prerequisite for modeling soil carbon stocks and its changes at regional to national scale. However, the required data on biomass removal is often available only for the recent past. In this study we used yield tables as a source of probable past forest management to drive a single tree based stand growth model. Next, simulated growth and timber volume was converted to tree compartment carbon stocks and biomass turnover. The study explicitly accounted for differences in site quality between stands. In addition we performed a Monte Carlo uncertainty and sensitivity analysis. We exemplify the approach by calculating long-term means of past litter production for 10 species by using yield tables that have been applied in Central Germany during the last century. We found that litter production resulting from harvest residues was almost as large as the one from biomass turnover. Differences in site quality caused large differences in litter production. At a given site quality, the uncertainty in soil carbon inputs were 14%, 17%, and 25% for beech, spruce, and pine stands, respectively. The sensitivity analysis showed that the most influential parameters were associated with foliage biomass and turnover. We conclude that rates of mean past litter production and their uncertainties can reliably be modeled on the basis of yield tables if the model accounts for 1) full rotation length including thinning and final harvest, 2) differences in site quality, and 3) environmental dependency of foliage biomass and foliage turnover.
  • Wutzler, Max Planck Institute for Biogeochemistry, Jena, Germany E-mail: tm@nn.de (email)
  • Mund, Max Planck Institute for Biogeochemistry, Jena, Germany E-mail: mm@nn.de
article id 288, category Special section
Björn Berg, Per Gundersen, Cecilia Akselsson, Maj-Britt Johansson, Åke Nilsson, Lars Vesterdal. (2007). Carbon sequestration rates in Swedish forest soils – a comparison of three approaches. Silva Fennica vol. 41 no. 3 article id 288. https://doi.org/10.14214/sf.288
Keywords: carbon sequestration; stable humus; forest floor C; litter decomposition; limit value
Abstract | View details | Full text in PDF | Author Info

Carbon sequestration rates in forest soil can be estimated using the concept of calculable stable remains in decomposing litter. In a case study of Swedish forest land we estimated C-sequestration rates for the two dominant tree species in the forest floor on top of the mineral soil. Carbon sequestration rates were upscaled to the forested land of Sweden with 23 x 106 ha with Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (Karst.) L.). Two different theoretical approaches, based on limit-value for litter decomposition and N-balance for vegetation and SOM gave rates of the same magnitude. For the upscaling, using these methods, 17 000 grids of 5 x 5 km were used.

The ‘limit-value approach’ gave a sequestration of 4.8   106 tons of C, annually sequestered in the forest floor, with an average of 180 kg C ha–1 yr–1 and a range from 40 to 410 kg C ha–1 yr–1. The ‘N-balance approach’ gave an average value of c. 96 kg ha–1 yr–1 and a range from –60 to 360 kg ha–1 yr–1. A method based on direct measurements of changes in humus depth over 40 years, combined with C analyses gave an average rate that was not very different from the calculated rates, viz. c. 180 kg ha–1 yr–1 and a range from –20 to 730 kg ha–1 yr–1. These values agree with forest floor C sequestration rate based on e.g. sampling of chronsequences but differ from CO2 balance measurements.

The three approaches showed different patterns over the country and regions with high and low carbon sequestration rates that were not always directly related to climate.

  • Berg, Dept. of Forest Ecology, University of Helsinki, Finland (present address: Dipartimento Biologia Strutturale e Funzionale, Complesso Universitario, Monte S. Angelo, Napoli, Italy E-mail: bjorn.berg@helsinki.fi (email)
  • Gundersen, Forest & Landscape Denmark, University of Copenhagen, Denmark E-mail: pg@nn.dk
  • Akselsson, Swedish Environmental Research Institute, IVL, Gothenburg, Sweden E-mail: ca@nn.se
  • Johansson, Department of Forest Soils, SLU, Uppsala, Sweden E-mail: mbj@nn.se
  • Nilsson, Department of Forest Soils, SLU, Uppsala, Sweden E-mail: an@nn.se
  • Vesterdal, Forest & Landscape Denmark, University of Copenhagen, Denmark E-mail: lv@nn.dk
article id 287, category Special section
Mikko Peltoniemi, Juha Heikkinen, Raisa Mäkipää. (2007). Stratification of regional sampling by model-predicted changes of carbon stocks in forested mineral soils. Silva Fennica vol. 41 no. 3 article id 287. https://doi.org/10.14214/sf.287
Keywords: uncertainty; soil carbon; anticipated variance; forest soil; monitoring; repeated measurement; soil survey; stratified sampling
Abstract | View details | Full text in PDF | Author Info
Monitoring changes in soil C has recently received interest due to reporting under the Kyoto Protocol. Model-based approaches to estimate changes in soil C stocks exist, but they cannot fully replace repeated measurements. Measuring changes in soil C is laborious due to small expected changes and large spatial variation. Stratification of soil sampling allows the reduction of sample size without reducing precision. If there are no previous measurements, the stratification can be made with model-predictions of target variable. Our aim was to present a simulation-based stratification method, and to estimate how much stratification of inventory plots could improve the efficiency of the sampling. The effect of large uncertainties related to soil C change measurements and simulated predictions was targeted since they may considerably decrease the efficiency of stratification. According to our simulations, stratification can be useful with a feasible soil sample number if other uncertainties (simulated predictions and forecasted forest management) can be controlled. For example, the optimal (Neyman) allocation of plots to 4 strata with 10 soil samples from each plot (unpaired repeated sampling) reduced the standard error (SE) of the stratified mean by 9–34% from that of simple random sampling, depending on the assumptions of uncertainties. When the uncertainties of measurements and simulations were not accounted for in the division to strata, the decreases of SEs were 2–9 units less. Stratified sampling scheme that accounts for the uncertainties in measured material and in the correlates (simulated predictions) is recommended for the sampling design of soil C stock changes.
  • Peltoniemi, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: mikko.peltoniemi@metla.fi (email)
  • Heikkinen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: jh@nn.fi
  • Mäkipää, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: raisa.makipaa@metla.fi

Category : Climate resilient and sustainable forest management – Research article

article id 23066, category Climate resilient and sustainable forest management – Research article
A. Maarit I. Kallio, Victor F. Strîmbu, Helle Gobakken, Terje Gobakken. (2025). Climate change mitigation through alternative uses of Norwegian forest resources. Silva Fennica vol. 59 no. 1 article id 23066. https://doi.org/10.14214/sf.23066
Keywords: harvested wood products; forest sector; biofuels; greenhouse gas emissions; bioeconomy; carbon sink
Highlights: Climate effects of Norwegian forests under various policies were examined under parametric uncertainty; Shifting some of wood use to high-emission-saving wood products, climate benefits from Norwegian forests can be increased without increasing harvests; Only in the short-term it is possible to increase the climate benefits by increasing harvest; Investments to bioethanol production likely decrease climate benefits of forests.
Abstract | Full text in HTML | Full text in PDF | Author Info

The carbon substitution and storage effects related to Norwegian forests and the forest sector were compared under three potential roundwood harvest scenarios: maintaining harvests at 2021 levels, increasing harvests by 20% due to policies aimed at maximizing economic benefits from the forest sector, and reducing harvests by 20% due to biodiversity concerns. For harvested wood products, both the current product structure and hypothetical alternatives were considered. The carbon stock development in forests was projected using a forestry simulation tool for Norway. Many uncertainties in carbon storage, substitution parameters, and data have been addressed using Monte Carlo simulations. Shifting a portion of pulpwood use to produce wood-based insulation materials and textile fibres was found to increase the climate benefits from the Norwegian forest sector. In contrast, the shift to bioethanol production had only a marginal effect compared to the current production structure. The analysis spanned the next two decades, which is a period relevant to the investment and operational lifespan of industrial facilities. The results suggest that during this time, smarter use of harvested roundwood for HWPs with high carbon substitution benefits can be an effective means of climate change mitigation. However, in the long term, enhancing forest carbon sinks by reducing harvests may be more beneficial for the climate, provided that global efforts to reduce emissions from energy production are successful and lead to a decrease in emissions associated with the production of various materials.

  • Kallio, Norwegian University of Life Sciences, Faculty of Environmental Sciences and Natural Resource Management, P.O.Box 5003, NO-1432 Ås, Norway ORCID https://orcid.org/0000-0002-5393-761X E-mail: maarit.kallio@nmbu.no (email)
  • Strîmbu, Norwegian University of Life Sciences, Faculty of Environmental Sciences and Natural Resource Management, P.O.Box 5003, NO-1432 Ås, Norway ORCID https://orcid.org/0000-0002-0588-2036 E-mail: victor.strimbu@nmbu.no
  • Gobakken, Norwegian University of Life Sciences, Faculty of Environmental Sciences and Natural Resource Management, P.O.Box 5003, NO-1432 Ås, Norway E-mail: helle.ross.gobakken@nmbu.no
  • Gobakken, Norwegian University of Life Sciences, Faculty of Environmental Sciences and Natural Resource Management, P.O.Box 5003, NO-1432 Ås, Norway ORCID https://orcid.org/0000-0001-5534-049X E-mail: terje.gobakken@nmbu.no
article id 23067, category Climate resilient and sustainable forest management – Research article
Lucas N. López, Hanne K. Sjølie, Abbas Nabhani, Francisco X. Aguilar. (2024). Impacts of biodiversity and carbon policies on the management of Norwegian forest and its ecosystem services. Silva Fennica vol. 58 no. 4 article id 23067. https://doi.org/10.14214/sf.23067
Keywords: simulation; optimization; site productivity; bio-economic modeling; NorFor; regional impacts; TreeSim
Highlights: National-level biodiversity and carbon forest sector policies modelled in a simulation-optimization framework; Impacts of policies on management along site productivity gradients estimated; Policies vary in impact across productivity gradients with regional implications.
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Measures to enhance boreal forests’ biodiversity and climate change mitigation potential are high on the policy agenda. Site productivity influences management, ecological attributes, and economic outcomes. However, national-level analyses of management implementation in response to policies considering site productivity are lacking. We analyzed impacts of a carbon policy (Carb), a biodiversity policy (Bio) and a combined biodiversity and carbon policy (BioCarb) in Norway using a simulation-optimization framework, assessing impacts on forest management, timber harvest, ecological attributes, and carbon fluxes until year 2140. Management alternatives were simulated in the single-tree simulator TreeSim before being fed into a market model NorFor to compare policy outcomes to a business-as-usual (BAU) scenario. All policies led to decreased harvests. Old forests expanded from the current 3% to cover 21% or more of the productive forest area in all scenarios. Impacts of policies depended on site productivity. On low-productive land, management under Bio mirrored BAU, while the Carb and BioCarb policies yielded more set-asides. On high-productive land, management intensity under the Carb policy was similar to BAU but the Bio and BioCarb policies resulted in more set-asides and more old forest.  Thus, on low-productive land, the carbon policy showed to have the strongest impact on forest management, while on high-productive land, the biodiversity policy had the strongest impact. With geographical site-productivity gradients, the two policies exhibited different regional effects. The study shows that ex-ante analyses with appropriate tools can provide relevant information of multiple consequences beyond the stated aims which should be considered in policy design.

  • López, Inland Norway University of Applied Sciences, Faculty of Applied Ecology, Agricultural Sciences and Biotechnology, P.O. Box 2400, Koppang, Norway ORCID https://orcid.org/0009-0006-6860-3408 E-mail: lucas.lopez@inn.no (email)
  • Sjølie, Inland Norway University of Applied Sciences, Faculty of Applied Ecology, Agricultural Sciences and Biotechnology, P.O. Box 2400, Koppang, Norway ORCID https://orcid.org/0000-0001-8099-3521 E-mail: hanne.sjolie@inn.no
  • Nabhani, Inland Norway University of Applied Sciences, Faculty of Applied Ecology, Agricultural Sciences and Biotechnology, P.O. Box 2400, Koppang, Norway E-mail: abbas.nabhani@inn.no
  • Aguilar, Swedish University of Agricultural Sciences, Department of Forest Economics, SE-901 83 Umeå, Sweden E-mail: francisco.aguilar@slu.se
article id 23075, category Climate resilient and sustainable forest management – Research article
Lauri Männistö, Jari Miina, Saija Huuskonen. (2024). How to utilize natural regeneration of birch to establish mixed spruce-birch forests in Finland? Silva Fennica vol. 58 no. 3 article id 23075. https://doi.org/10.14214/sf.23075
Keywords: Betula pendula; Picea abies; Betula pubescens; mixed model; Chapman-Richards; mixed stands
Highlights: Naturally emerging birch admixture provides a sufficient starting point for mixed forest in young, planted spruce stands; Creating mixed forest is a time sensitive process, as different growth patterns among tree species might cause a situation where one tree species suppresses the other, leading to monoculture or two-storied mixed forest; Birch admixture should be retained during early cleaning.
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Mixed forests are known for their ability to provide a wide range of ecosystem services. Such forests have higher biodiversity compared to monocultures, are resilient against disturbances and may mitigate the effects of climate change. Despite well-known benefits, there is still little information on how these forests should be established and managed. The aim of this study was to describe the early growth dynamics of current boreal young mixed stands of planted Norway spruces (Picea abies (L.) Karst.) and naturally regenerated birches (Betula spp.). We collected data from 9  stands planted for spruce 8–14 years ago in Southern and Central Finland. Stem analysis was conducted to 144 spruces and to 144 birches to determine previous growth. We modelled the height and diameter development of individual trees in relation to tree age at stump height using non-linear mixed Chapman-Richards model. There were no significant differences between spruce and seed-origin birch in diameter growth at stump height, but the initial height increments of natural birches were larger than those of planted spruces. However, planted spruces were able to keep up with the height development of birches, if spruces received a head start over naturally regenerated seed-origin birch for two growing seasons. Thus, naturally regenerated birch admixture can be utilized to establish single-storied spruce-birch mixtures, and the admixture should be retained during the early cleaning of planted spruce stands.

  • Männistö, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID https://orcid.org/0009-0009-0213-1972 E-mail: lauri.mannisto@luke.fi (email)
  • Miina, Natural Resources Institute Finland (Luke), Natural resources, Yliopistokatu 6 B, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0002-8639-4383 E-mail: jari.miina@luke.fi
  • Huuskonen, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0001-8630-3982 E-mail: saija.huuskonen@luke.fi
article id 23069, category Climate resilient and sustainable forest management – Research article
John Alexander Pulgarin Diaz, Markus Melin, Tiina Ylioja, Päivi Lyytikäinen-Saarenmaa, Heli Peltola, Olli-Pekka Tikkanen. (2024). Relationship between stand and landscape attributes and Ips typographus salvage loggings in Finland. Silva Fennica vol. 58 no. 3 article id 23069. https://doi.org/10.14214/sf.23069
Keywords: Picea abies; wind damage; clear-cuts; European spruce bark beetle; stand attributes
Highlights: In Finland, the European spruce bark beetle (SBB) prefers mature stands (high age and mean diameter at breast hight), herb-rich heath forest sites and semi-coarse or coarse heath forest soil type, as well as a short distance to the closest wind damage from the previous-year, SBB damage from previous-year and particularly to clear-cuts; These stand types should be prioritised for monitoring SBB damage.
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Disturbances caused by the European spruce bark beetle (SBB; Ips typographus L.) on Norway spruce (Picea abies (L.) H. Karst.), have increased immensely across Central and Northern Europe, and are expected to increase further as a result of climate change. While this trend has been noted in Finland, so far limited research has been published. To support proper SBB risk management in Finland, we compared stand properties between salvage loggings due to SBB damage during 2012–2020 (4691 cases) and spruce stands free of SBB damage. Also, we explored the role of landscape attributes as drivers of SBB damage. We considered the forest stand attributes of site fertility class, stand development class, soil type, stand mean diameter at breast height and mean stand age. Considered forest landscape attributes were the distance from SBB-damaged stands to the closest clear-cut, to previous-year SBB-damaged stands and to the previous-year wind-damaged stand. We used nationwide forest logging and forest stock data, and analysed forest stand attributes using chi-squared and Mann-Whitney U tests and landscape attributes using generalised linear mixed models. Based on our findings, the SBB didn’t damage stands randomly, but prevailed in mature stands (high age and high mean diameter at breast height), in herb-rich heath forest site types and in semi-coarse or coarse heath forest soil soils. We found correlation between the landscape variables and the number of salvage loggings, with a higher number of loggings due to SBB damage close to clear-cuts. Our results help to find risk areas of SBB damage.

  • Pulgarin Diaz, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0003-0554-8254 E-mail: alexander.pulgarin.diaz@uef.fi (email)
  • Melin, Natural Resources Institute Finland (Luke), P.O. Box 68, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0001-7290-9203 E-mail: markus.melin@luke.fi
  • Ylioja, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-0079 Helsinki, Finland ORCID https://orcid.org/0000-0002-8840-7504 E-mail: tiina.ylioja@luke.fi
  • Lyytikäinen-Saarenmaa, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0003-1884-3084 E-mail: paivi.lyytikainen-saarenmaa@ef.fi
  • Peltola, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: heli.peltola@uef.fi
  • Tikkanen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0002-3875-2772 E-mail: olli-pekka.tikkanen@uef.fi
article id 23061, category Climate resilient and sustainable forest management – Research article
Noora Tienaho, Ninni Saarinen, Tuomas Yrttimaa, Ville Kankare, Mikko Vastaranta. (2024). Quantifying fire-induced changes in ground vegetation using bitemporal terrestrial laser scanning. Silva Fennica vol. 58 no. 3 article id 23061. https://doi.org/10.14214/sf.23061
Keywords: forest fires; biomass; boreal forest; LIDAR; controlled burning; surface differencing; surface fires
Highlights: Bitemporal terrestrial laser scanning provided a means for identifying surface areas exposed to fire by utilizing a surface differencing method developed in this study; The developed method allowed for the quantification of fire-induced volumetric changes in ground vegetation at high resolution, facilitating the assessment of the impact of surface fires on forest ecosystems.
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Forest fires pose a significant threat to forest carbon storage and sinks, yet they also play a crucial role in the natural dynamics of boreal forests. Accurate quantification of biomass changes resulting from forest fires is essential for damage assessment and controlled burning evaluation. This study utilized terrestrial laser scanning (TLS) to quantify changes in ground vegetation resulting from low-intensity surface fires. TLS data were collected before and after controlled burnings at eight one-hectare test sites in Scots pine (Pinus sylvestris L.) dominated boreal forests in Finland. A surface differencing-based method was developed to identify areas exposed to fire. Validation, based on visual interpretation of 1 × 1 m surface patches (n = 320), showed a recall, precision, and F1-score of 0.9 for the accuracy of identifying burned surfaces. The developed method allowed the assessment of the magnitude of fire-induced vegetation changes within the test sites. The proportions of burned 1 × 1 m areas within the test sites varied between 51–96%. Total volumetric change in ground vegetation was on average –1200 m³ ha-1, with burning reducing the vegetation volume by 1700 m³ ha-1 and vegetation growth increasing it by 500 m³ ha-1. Substantial variations in the volumetric changes within and between the test sites were detected, highlighting the complex dynamics of surface fires, and emphasizing the importance of having observations from multiple sites. This study demonstrates that bitemporal TLS measurements provide a robust means for characterizing fire-induced changes, facilitating the assessment of the impact of surface fires on forest ecosystems.

article id 23042, category Climate resilient and sustainable forest management – Research article
Johanna Jääskeläinen, Lauri Korhonen, Mikko Kukkonen, Petteri Packalen, Matti Maltamo. (2024). Individual tree inventory based on uncrewed aerial vehicle data: how to utilise stand-wise field measurements of diameter for calibration? Silva Fennica vol. 58 no. 3 article id 23042. https://doi.org/10.14214/sf.23042
Keywords: laser scanning; calibration; mixed-effects model; single-tree detection
Highlights: A practical scheme to improve the accuracy of predicted tree and stand attributes in an uncrewed aerial vehicle based individual tree inventory; Accuracy was considerably improved with data from 2–4 sample trees from the target stand; Calibrated existing models and the construction of local models performed equally well; The laborious task of constructing a local model can be avoided by using a calibrated transferred model.
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Uncrewed aerial vehicles (UAV) have great potential for use in forest inventories, but in practice they can be expensive for relatively small inventory areas as a large number of field measurements are needed for model construction. One proposed solution is to transfer previously constructed models to a new inventory area and to calibrate these with a small number of local field measurements. Our objective was to compare calibration of general models and the construction of new models to determine the best approach for UAV-based forest inventories. Our material included field measurements and UAV-based laser scanning data, from which individual trees were automatically identified. A general mixed-effects model for diameter at breast height (DBH) had been formulated earlier based on data from a geographically wider area. It was calibrated to the study area with field measurements from 2–10 randomly selected calibration trees. The calibrated diameters were used to calculate the diameter of a basal area median tree (DGM), tree volumes, and the volume of all trees at plot-level. Next, new DBH-models were formulated based on the 2–10 randomly selected trees and calibrated with plot-level random effects estimated during model construction. Finally, plot-specific height-diameter regression models were formulated by randomly selecting 10 trees from each plot. Calibration reduced the prediction errors of all variables. An increase in the number of calibration trees decreased error rates by 1–6% depending on the variable. Calibrated predictions from the general mixed-effects model were similar to the separately formulated mixed-effects models and plot-specific regression models.

Category : Research article

article id 25013, category Research article
Binod Kafle, Ville Kankare, Harri Kaartinen, Kari Väätäinen, Heikki Hyyti, Tamas Faitli, Juha Hyyppä, Antero Kukko, Kalle Kärhä. (2025). Assessing the consistency of low vegetation characteristics estimated using harvester, handheld, and drone light detection and ranging (LiDAR) systems. Silva Fennica vol. 59 no. 2 article id 25013. https://doi.org/10.14214/sf.25013
Keywords: biodiversity; harvester; wood harvesting; dense area for games; drone laser scanning (DLS); handheld mobile laser scanning (HMLS); point cloud
Highlights: Harvester-mounted LiDAR consistently estimated low vegetation height and volume comparable to handheld and drone LiDAR; Enhancing LiDAR range could improve harvester LiDAR efficiency, reducing processing time and increasing accuracy beyond 20 m.
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Evaluating the potential of a harvester-mounted LiDAR system in monitoring biodiversity indicators such as low vegetation during forest harvesting could enhance sustainable forest management and habitat conservation including dense forest areas for game. However, there is a lack of understanding on the capabilities and limitations of these systems to detect low vegetation characteristics. To address this knowledge gap, this study investigated the performance of a harvester-mounted LiDAR system for measuring low vegetation (height <5 m) attributes in a boreal forest in Finland, by comparing it with handheld mobile laser scanning (HMLS) and drone laser scanning (DLS) systems. LiDAR point cloud data was collected in September 2023 to quantify the low vegetation height (maximum, mean, and percentiles), volume (voxel-based and mean height-based) and cover (grid method). Depending on the system, LiDAR point cloud data was collected either before (HMLS and DLS), during (harvester LiDAR) or after (HMLS and DLS) harvesting operations. A total of 46 fixed-sized (5 m × 5 m) grid cells were studied and analyzed. Results showed harvester-mounted LiDAR provided consistent estimates with HMLS and DLS for maximum height, 99th height percentile, and volume across various grids (5 cm, 10 cm, 20 cm) and voxel (20 cm) sizes. High correlation was observed between the systems used for these attributes. This study demonstrated that harvester-mounted LiDAR is comparable to HMLS and DLS for assessing low vegetation height and volume. The findings could assist forest harvester operators in identifying potential low vegetation and dense areas for conservation and game management.

  • Kafle, School of Forest Sciences, University of Eastern Finland (UEF), Yliopistokatu 7, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0003-0744-3480 E-mail: binod.kafle@uef.fi (email)
  • Kankare, Department of Geography and Geology, University of Turku, FI-20014 Turun yliopisto, Finland ORCID https://orcid.org/0000-0001-6038-1579 E-mail: viveka@utu.fi
  • Kaartinen, Department of Remote Sensing and Photogrammetry, Finnish Geospatial Research Institute (FGI), Opastinsilta 12 C, FI-00520 Helsinki, Finland ORCID https://orcid.org/0000-0002-4796-3942 E-mail: harri.kaartinen@nls.fi
  • Väätäinen, Natural Resources Institute Finland (Luke), Yliopistokatu 6 B, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0002-6886-0432 E-mail: kari.vaatainen@luke.fi
  • Hyyti, Department of Remote Sensing and Photogrammetry, Finnish Geospatial Research Institute (FGI), Opastinsilta 12 C, FI-00520 Helsinki, Finland ORCID https://orcid.org/0000-0003-4664-6221 E-mail: heikki.hyyti@nls.fi
  • Faitli, Department of Remote Sensing and Photogrammetry, Finnish Geospatial Research Institute (FGI), Opastinsilta 12 C, FI-00520 Helsinki, Finland ORCID https://orcid.org/0000-0001-5334-5537 E-mail: tamas.faitli@nls.fi
  • Hyyppä, Department of Remote Sensing and Photogrammetry, Finnish Geospatial Research Institute (FGI), Opastinsilta 12 C, FI-00520 Helsinki, Finland E-mail: juha.coelasr@gmail.com
  • Kukko, Department of Remote Sensing and Photogrammetry, Finnish Geospatial Research Institute (FGI), Opastinsilta 12 C, FI-00520 Helsinki, Finland ORCID https://orcid.org/0000-0002-3841-6533 E-mail: antero.kukko@nls.fi
  • Kärhä, School of Forest Sciences, University of Eastern Finland (UEF), Yliopistokatu 7, FI-80101 Joensuu, Finland E-mail: kalle.karha@uef.fi
article id 25003, category Research article
Lennart Noordermeer, Terje Gobakken, Johannes Breidenbach, Rune Eriksen, Erik Næsset, Hans Ole Ørka, Ole M. Bollandsås. (2025). Effects of sample tree selection and calculation methods on the accuracy of field plot values in area-based forest inventories. Silva Fennica vol. 59 no. 2 article id 25003. https://doi.org/10.14214/sf.25003
Keywords: forest inventory methods; field plot accuracy; height-diameter modeling; sample tree selection
Highlights: Retaining field-measured heights of sample trees improved accuracies of plot values; Selecting sample trees with probability proportional to basal area was recommended; The number of sample trees and sample tree selection method impacted accuracies; The choice of calculation method had a strong influence on accuracies of plot values.
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Accurate field plot data on forest attributes are crucial in area-based forest inventories assisted by airborne laser scanning, providing an essential reference for calibrating predictive models. This study assessed how sample tree selection methods and plot data calculation methods affect the accuracy of field plot values of timber volume, Lorey’s mean height, and dominant height. We used data obtained from 12 420 circular sample plots of 250 m2, measured as part of the Norwegian national forest inventory and 45 local forest management inventories. We applied Monte Carlo simulations by which we tested various numbers of sample trees, methods to select sample trees, and methods to calculate plot-level values from tree-level measurements. Accuracies of plot values were statistically significantly affected by the number of sample trees, sample tree selection method, and calculation method. Obtained values of root mean square error ranged from 5% to 16% relative to the mean observed values, across the factors studied. Accuracy improved with increasing numbers of sample trees for all forest attributes. We obtained greatest accuracies by selecting sample trees with a probability proportional to basal area, and by retaining field-measured heights for sample trees and using heights predicted with a height-diameter model for non-sample trees. This study highlights the importance of appropriate sample tree selection methods and calculation methods in obtaining accurate field plot data in area-based forest inventories.

  • Noordermeer, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, NMBU, P.O. Box 5003, NO-1432 Ås, Norway ORCID https://orcid.org/0000-0002-8840-0345 E-mail: lennart.noordermeer@nmbu.no (email)
  • Gobakken, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, NMBU, P.O. Box 5003, NO-1432 Ås, Norway ORCID https://orcid.org/0000-0001-5534-049X E-mail: terje.gobakken@nmbu.no
  • Breidenbach, Division of Forest and Forest Resources, Norwegian Institute of Bioeconomy Research (NIBIO), P.O. Box 115, NO-1431 Ås, Norway E-mail: johannes.breidenbach@nibio.no
  • Eriksen, Division of Forest and Forest Resources, Norwegian Institute of Bioeconomy Research (NIBIO), P.O. Box 115, NO-1431 Ås, Norway E-mail: rune.eriksen@nibio.no
  • Næsset, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, NMBU, P.O. Box 5003, NO-1432 Ås, Norway E-mail: erik.naesset@nmbu.no
  • Ørka, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, NMBU, P.O. Box 5003, NO-1432 Ås, Norway E-mail: hans-ole.orka@nmbu.no
  • Bollandsås, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, NMBU, P.O. Box 5003, NO-1432 Ås, Norway ORCID https://orcid.org/0000-0002-1231-7692 E-mail: ole.martin.bollandsas@nmbu.no
article id 25004, category Research article
Kalle Kemppainen, Kalle Kärhä, Juha Laitila, Antti Sairanen, Ville Kankaanhuhta, Heli Viiri, Heli Peltola. (2025). Evaluation of the productivity and costs of excavator-based mechanized tree planting in Finland based on automated data collection. Silva Fennica vol. 59 no. 1 article id 25004. https://doi.org/10.14214/sf.25004
Keywords: site preparation; cost-efficiency; mechanization; forest regeneration; planting machine; site selection
Highlights: With proper site selection and competent machine operators, it is possible to achieve a higher productivity than 200 seedlings per operating hour in excavator-based mechanized planting; The hectare-based regeneration costs from excavator-based mechanized planting can be 3% lower than those of conventional manual planting chains; The most cost-efficient planting chain was a continuously advancing mounder combined with manual planting.
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The poor cost-effectiveness of mechanized planting (MECP) is the main reason for the low mechanization rate of planting. In this study, we investigated the productivity of the mechanized excavator-based planting of Norway spruce (Picea abies [L.] H. Karst.) seedlings based on data collected by the Risutec Asta documentation system. We also compared the costs of a MECP chain with two different manual planting (MAP) chains, where mounding was carried out by a crawler excavator (EXC) or a continuously advancing mounder (CONT). The MECP of seedlings was carried out using an EXC equipped with a Risutec PM-160 planting device. Generally, the nine study sites in western Finland contained few surface obstacles (e.g., the logging residues had mainly been harvested), which made the conditions very suitable for MECP. The average production time taken by the MECP was 9 h ha–1. The operating hour (G15-h) productivity averaged 215 seedlings G15-h−1, with the mean planting time being 13.8 s seedling−1. Loading 160 seedlings into the seedling cassette took approximately 10 min (3.8 s seedling−1). Overall, the cost of the MECP was about 3% lower than for the EXC + MAP. However, when productivity was set at <210 seedlings G15-h−1, the cost of the MECP was higher than that of the EXC + MAP. Based on our findings, the most cost-efficient planting chain was CONT + MAP. However, based on our results, the required level of productivity can be achieved if the sites are suitable for MECP and the machine operators are skilled.

  • Kemppainen, School of Forest Sciences, University of Eastern Finland (UEF), P.O. Box 111, FI-80101 Joensuu, Finland; Natural Resources Institute Finland (Luke), Yliopistokatu 6, FI-80100 Joensuu, Finland ORCID https://orcid.org/0009-0000-6184-8812 E-mail: kalle.kemppainen@uef.fi (email)
  • Kärhä, School of Forest Sciences, University of Eastern Finland (UEF), P.O. Box 111, FI-80101 Joensuu, Finland E-mail: kalle.karha@uef.fi
  • Laitila, Natural Resources Institute Finland (Luke), Yliopistokatu 6, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0003-4431-3319 E-mail: juha.laitila@luke.fi
  • Sairanen, School of Forest Sciences, University of Eastern Finland (UEF), P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0009-0008-8632-3797 E-mail: anttsair@student.uef.fi
  • Kankaanhuhta, Natural Resources Institute Finland (Luke), Yliopistokatu 6, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0001-5785-5972 E-mail: ville.kankaanhuhta@luke.fi
  • Viiri, UPM-Kymmene Plc, UPM Forest, Peltokatu 26 C 4, FI-33100 Tampere, Finland ORCID https://orcid.org/0000-0003-3952-9481 E-mail: heli.viiri@upm.com
  • Peltola, School of Forest Sciences, University of Eastern Finland (UEF), P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0003-1384-9153 E-mail: heli.peltola@uef.fi
article id 24046, category Research article
Līga Liepa, Zigmārs Rendenieks, Edgars Dubrovskis, Lāsma Freimane, Inga Straupe, Āris Jansons. (2025). Patterns of short-term vegetation recovery after a fire in protected Scots pine forests of hemiboreal Latvia. Silva Fennica vol. 59 no. 1 article id 24046. https://doi.org/10.14214/sf.24046
Keywords: succession; fire disturbance; plant functional traits; post-fire regeneration; protected forests
Highlights: We observed more rapid vegetation recovery at the ground level, but the dominance increased more steadily at herb and tree layers; We found the highest species diversity at the herb layer during the third (middle-aged stands) and fourth (over-mature stands) years after fire disturbance; From regenerating tree species, only Populus tremula in over-mature stands showed a decline in projective cover during the four studied years.
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Wildfires as natural disturbances have had important impacts on terrestrial ecosystems, including forests. We studied patterns of short-term vegetation recovery after surface fire in protected hemiboreal Pinus sylvestris L.-dominated forest. Our study was carried out near Stikli village in Western Latvia. Seven forest stands – middle-age and over-mature were sampled on nutrient-poor and mesic soils. Forest fire occurred in the summer of 2018 and covered 1440 ha of forested area. In each stand we established 16 sample plots (1 m × 1 m) in a radial pattern from the center. Every summer from 2019 till 2022 we surveyed these sample plots – recorded projective cover (%) and identified Ellenberg indicator values and species traits – plant strategy groups (C-S-R after Grime), Raunkiær life history forms and habitat types. Additionally, the occurrence of specialized fire-adapted plants was recorded. In total we identified 15 species in the ground layer, 47 species in the herbaceous layer, and 9 regenerating tree species. The colonization at the ground layer was the most rapid (projective cover increased overall by 67% in middle-aged stands and by 82% in over-mature stands). Species diversity was the highest at the herb layer during the third (middle-aged stands) and fourth (over-mature stands) after fire disturbance but showed overall declining trends. Betula spp. and Populus tremula L.-dominated regenerating tree species. The dominance of fire-adapted species declined rapidly after the fire except for moss Polytrichum spp. Overall, hemiboreal over-mature stands demonstrated higher vegetation cover and more rapid rate of initial colonization compared to middle-aged stands.

  • Liepa, Latvian State Forest Research Institute “Silava”, Rīgas street 111, LV-2169, Salaspils, Latvia; Latvian University of Life Sciences and Technologies, Faculty of Forest and Environmental Sciences, Akadēmijas street 11, LV-3001, Jelgava, Latvia ORCID https://orcid.org/0000-0002-8270-6722 E-mail: liga.liepa@outlook.com (email)
  • Rendenieks, Latvian State Forest Research Institute “Silava”, Rīgas street 111, LV-2169, Salaspils, Latvia ORCID https://orcid.org/0000-0002-3511-1486 E-mail: zigmars.rendenieks@silava.lv
  • Dubrovskis, Latvian University of Life Sciences and Technologies, Faculty of Forest and Environmental Sciences, Akadēmijas street 11, LV-3001, Jelgava, Latvia ORCID https://orcid.org/0000-0003-0810-5651 E-mail: edgars.dubrovskis@lbtu.lv
  • Freimane, Latvian University of Life Sciences and Technologies, Faculty of Forest and Environmental Sciences, Akadēmijas street 11, LV-3001, Jelgava, Latvia E-mail: lasma.freimane@lbtu.lv
  • Straupe, Latvian University of Life Sciences and Technologies, Faculty of Forest and Environmental Sciences, Akadēmijas street 11, LV-3001, Jelgava, Latvia ORCID https://orcid.org/0000-0002-2098-7194 E-mail: inga.straupe@lbtu.lv
  • Jansons, Latvian State Forest Research Institute “Silava”, Rīgas street 111, LV-2169, Salaspils, Latvia ORCID https://orcid.org/0000-0001-7981-4346 E-mail: aris.jansons@silava.lv
article id 24041, category Research article
Johanna Jetsonen, Annamari Laurén, Heli Peltola, Katariina Laurén, Samuli Launiainen, Marjo Palviainen. (2025). Volume growth responses of Scots pine and Norway spruce to nitrogen fertilization: quantitative synthesis of fertilization experiments in Finland. Silva Fennica vol. 59 no. 1 article id 24041. https://doi.org/10.14214/sf.24041
Keywords: precipitation; growth response; mineral soils; site fertility
Highlights: Volume growth responses increased linearly with the applied nitrogen (N) dose up to 200 kg N ha–1; Precipitation, site fertility, and time since fertilization affected the growth responses; Volume growth response to nitrogen fertilization was larger for Scots pine than for Norway spruce; Results can support planning of environmentally responsible and economically viable precision fertilization.
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The effects of nitrogen (N) fertilization on tree growth have been studied widely in boreal forests in Finland, but a quantitative synthesis is still lacking. We performed a quantitative synthesis on volume growth responses to N fertilization in Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.) stands in experiments established on mineral soils across Finland. Our study employed findings of 9 published studies including 108 Scots pine and 57 Norway spruce observations covering a wide range of N fertilization treatments, as well as forest stand- and climatic conditions. Based on these observations, we built linear mixed models to describe the N fertilization-induced annual volume growth response of Scots pine and Norway spruce stands. Our models showed that the N dose was the best predictor for volume growth response, and the growth response increased linearly with increasing N dose for both tree species. The volume growth responses also increased along with an increase in mean annual precipitation. The annual volume growth response decreased with the time since fertilization. For Scots pine, the best model also contained site fertility; increase in site fertility increased the volume growth response. These findings emphasize the need for site-specific precision fertilization schemes to sustainably improve growth and carbon sequestration of boreal forests.

  • Jetsonen, Department of Forest Sciences, University of Helsinki, P.O. Box 27, 00014 University of Helsinki, Finland ORCID https://orcid.org/0009-0006-4878-8951 E-mail: johanna.jetsonen@helsinki.fi (email)
  • Laurén, Department of Forest Sciences, University of Helsinki, P.O. Box 27, 00014 University of Helsinki, Finland; Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland ORCID https://orcid.org/0000-0002-6835-9568 E-mail: annamari.lauren@helsinki.fi
  • Peltola, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland ORCID https://orcid.org/0000-0003-1384-9153 E-mail: heli.peltola@uef.fi
  • Laurén, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland; University of Helsinki, Faculty of Medicine, Haartmaninkatu 8, 00290 Helsinki, Finland ORCID https://orcid.org/0009-0009-4677-9826 E-mail: katariina.lauren@helsinki.fi
  • Launiainen, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, 00790 Helsinki, Finland ORCID https://orcid.org/0000-0001-6611-6573 E-mail: samuli.launiainen@luke.fi
  • Palviainen, Department of Forest Sciences, University of Helsinki, P.O. Box 27, 00014 University of Helsinki, Finland ORCID https://orcid.org/0000-0001-9963-4748 E-mail: marjo.palviainen@helsinki.fi
article id 24026, category Research article
Olli Muhonen, Heli Peltola, Annamari Laurén, Veli-Pekka Ikonen, Juha Nevalainen, Laura Pikkarainen, Antti Kilpeläinen, Samuli Launiainen, Marjo Palviainen. (2025). Spatial evenness of fertilization and short-term volume growth responses of Scots pine and Norway spruce to fertilization intensity. Silva Fennica vol. 59 no. 1 article id 24026. https://doi.org/10.14214/sf.24026
Keywords: Pinus sylvestris; boreal forest; Picea abies; stand growth; forest fertilization; upland forests; nitrogen fertilizer dose
Highlights: Differences between the target and the measured fertilizer doses were in a range of 3–22% for fertilization treatments in Scots pine and Norway spruce study sites; The volume growth response was affected by the initial basal area and fertilization dose; The fertilization dose correlated with nitrogen concentrations in needles and soil organic (humus) layer and volume growth.
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We studied the spatial evenness of nitrogen (N) fertilizer and the effects of fertilization intensity on the short-term volume growth responses in two ground-fertilized Scots pine and two airborne-fertilized Norway spruce study sites on mesic (Myrtillus-type) upland forests in Eastern Finland. We also studied the relationships between measured fertilizer dose, N concentrations in the needles and soil organic (humus) layer, and volume growth of the trees. In each study site, we established three replicate 1 ha blocks for each fertilization treatment (0, 150 and 200 kg N ha–1). Each block contained three 200 m2 circular plots. The spatial evenness of the fertilizer was measured using textile funnels. The height, breast height diameter and vitality of the trees were measured annually. The nutrient concentrations in the needles and humus layer were measured once. Differences between the target and measured fertilizer doses were 3–10% for Scots pine and 11–22% for Norway spruce. At the Scots pine and Norway spruce sites, the volume growth was 0.4–2.1 and 1.8–2.6 m3 ha–1 a–1 higher with fertilization, respectively. The fertilizer dose correlated with the N concentrations in the needles and humus layer, and volume growth. Significant volume growth responses to the fertilization (p < 0.05) were found only in the Scots pine at Ilomantsi, possibly due to variations in the initial volume and the fertilizer dose realized between and within treatments. We expect that the differences in volume growth responses between fertilization treatments will increase over time.

  • Muhonen, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, FI-80101 Joensuu, Finland ORCID https://orcid.org/0009-0007-4051-8567 E-mail: ollmu@uef.fi (email)
  • Peltola, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0003-1384-9153 E-mail: heli.peltola@uef.fi
  • Laurén, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, FI-80101 Joensuu, Finland; University of Helsinki, Department of Forest Sciences, Latokartanonkaari 7, FI-00014 Helsinki, Finland ORCID https://orcid.org/0000-0002-6835-9568 E-mail: annamari.lauren@helsinki.fi
  • Ikonen, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0003-1732-2922 E-mail: veli-pekka.ikonen@uef.fi
  • Nevalainen, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, FI-80101 Joensuu, Finland ORCID https://orcid.org/0009-0000-2972-4385 E-mail: juha.hs.nevalainen@gmail.com
  • Pikkarainen, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0001-5301-3639 E-mail: laura.pikkarainen@uef.fi
  • Kilpeläinen, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0003-4299-0578 E-mail: antti.kilpelainen@uef.fi
  • Launiainen, Natural Resources Institute Finland, Bioeconomy and Environment, Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0001-6611-6573 E-mail: samuli.launiainen@luke.fi
  • Palviainen, University of Helsinki, Department of Forest Sciences, Latokartanonkaari 7, FI-00014 Helsinki, Finland ORCID https://orcid.org/0000-0001-9963-4748 E-mail: marjo.palviainen@helsinki.fi
article id 24068, category Research article
Kari T. Korhonen. (2024). Corrigendum: Forests of Finland 2019–2023 and their development 1921–2023. Silva Fennica vol. 58 no. 5 article id 24068. https://doi.org/10.14214/sf.24068
Abstract | Full text in HTML | Full text in PDF | Author Info
Corrigendum
article id 24047, category Research article
Kalle Kemppainen, Kalle Kärhä, Juha Laitila, Antti Sairanen, Ville Kankaanhuhta, Heli Viiri, Heli Peltola. (2024). RETRACTED: Evaluation of the productivity and costs of excavator-based mechanized tree planting in Finland based on automated data collection. Silva Fennica vol. 58 no. 5 article id 24047. https://doi.org/10.14214/sf.24047
Keywords: site preparation; cost-efficiency; mechanization; forest regeneration; planting machine; site selection
Highlights: With proper site selection and competent machine operators, it is possible to achieve a higher productivity than 200 seedlings per operating hour in excavator-based mechanized planting; The hectare-based regeneration costs from excavator-based mechanized planting can be 5% lower than those of conventional manual planting chains; The most cost-efficient planting chain was a continuously advancing mounder combined with manual planting.
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The poor cost-effectiveness of mechanized planting (MECP) is the main reason for the low mechanization rate of planting. In this study, we investigated the productivity of the mechanized excavator-based planting of Norway spruce (Picea abies [L.] H. Karst.) seedlings based on data collected by the Risutec Asta documentation system. We also compared the costs of a MECP chain with two different manual planting (MAP) chains, where mounding was carried out by a crawler excavator (EXC) or a continuously advancing mounder (CONT). The MECP of seedlings was carried out using an EXC equipped with a Risutec PM-160 planting device. Generally, the nine study sites in western Finland contained few surface obstacles (e.g., the logging residues had mainly been harvested), which made the conditions very suitable for MECP. The average production time taken by the MECP was 9 h ha-1. The operating hour (G15-h) productivity averaged 215 seedlings G15-h-1 , with the mean planting time being 13.8 s seedling-1. Loading 160 seedlings into the seedling cassette took approximately 10 min (3.8 s seedling-1). Overall, the cost of the MECP was about 5% lower than for the EXC + MAP. However, when productivity was set at <200 seedlings G15-h-1 , the cost of the MECP was higher than that of the EXC + MAP. Based on our findings, the most cost-efficient planting chain was CONT + MAP. However, based on our results, the required level of productivity can be achieved if the sites are suitable for MECP and the machine operators are skilled.

  • Kemppainen, School of Forest Sciences, University of Eastern Finland (UEF), P.O. Box 111, FI-80101 Joensuu, Finland; Natural Resources Institute Finland (Luke), Yliopistokatu 6, FI-80100 Joensuu, Finland ORCID https://orcid.org/0009-0000-6184-8812 E-mail: kalle.kemppainen@uef.fi (email)
  • Kärhä, School of Forest Sciences, University of Eastern Finland (UEF), P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0002-8455-2974 E-mail: kalle.karha@uef.fi
  • Laitila, Natural Resources Institute Finland (Luke), Yliopistokatu 6, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0003-4431-3319 E-mail: juha.laitila@luke.fi
  • Sairanen, School of Forest Sciences, University of Eastern Finland (UEF), P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0009-0008-8632-3797 E-mail: anttsair@student.uef.fi
  • Kankaanhuhta, Natural Resources Institute Finland (Luke), Yliopistokatu 6, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0001-5785-5972 E-mail: ville.kankaanhuhta@luke.fi
  • Viiri, UPM-Kymmene Plc, UPM Forest, Peltokatu 26 C 4, FI-33100 Tampere, Finland ORCID https://orcid.org/0000-0003-3952-9481 E-mail: heli.viiri@upm.com
  • Peltola, School of Forest Sciences, University of Eastern Finland (UEF), P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0003-1384-9153 E-mail: heli.peltola@uef.fi
article id 24045, category Research article
Kari T. Korhonen, Minna Räty, Helena Haakana, Juha Heikkinen, Juha-Pekka Hotanen, Mikko Kuronen, Juho Pitkänen. (2024). Forests of Finland 2019–2023 and their development 1921–2023. Silva Fennica vol. 58 no. 5 article id 24045. https://doi.org/10.14214/sf.24045
Keywords: forest management; National Forest Inventory; growing stock; forest resources; forest damage; increment; monitoring indicators of biodiversity
Highlights: The latest Finnish National Forest Inventory is presented; Volume of growing stock has almost doubled since the 1920s and has continued to increase since the previous inventory; Volume increment is more than double the increment 100 years ago but has declined recently; Mortality is increasing at alarming rate; Amount of dead wood has now increased also in North Finland.
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In 2019–2023 the 13th Finnish National Forest Inventory (NFI) was implemented by measuring a total of 62 266 sample plots across the country. The methodology of the sampling and measurements was similar as in the previous inventory, but the proportion and number of remeasured permanent plots was increased to improve the monitoring of annual increment and other changes in the forests. Only 6.2 M ha (14%) of Finland’s total land area (30.4 M ha) is other land than forestry land. Productive and poorly productive forests cover 22.9 M ha (75%) of the total land area.  The forest area has remained stable in recent decades but the forest area available for wood supply (FAWS) has decreased due to increased forest protection – 23% of the forestry land and 10% of the productive forest are not available for wood supply. Compared to the previous inventory, forest resources have continued to increase but the average annual increment has declined from 107.8 M m3 to 103.0 M m3. The quality of forests from the timber production point of view has remained relatively good or improved slightly. The area of observed forest damage on FAWS is 8.4 M ha (46% of FAWS area), half of these minor damages with no impact on stand quality. Although the area of forest damage has not increased, the amount of mortality has continued to increase, and is now 8.8 M m3 year–1. The amount of dead wood has continued to increase in South Finland, while in North Finland the declining trend has turned into a slight increase. Since the 1920s, the area of forestry land has remained stable, but the area of productive forest has increased due to the drainage of poorly productive or treeless peatlands. The total volume of growing stock has increased by 84% and annual increment has more than doubled.

  • Korhonen, Natural Resources Institute Finland (Luke), P.O.Box 68, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0002-6198-853X E-mail: kari.t.korhonen@luke.fi (email)
  • Räty, Natural Resources Institute Finland (Luke), P.O.Box 2, FI-00790, Helsinki, Finland ORCID https://orcid.org/0000-0001-9898-8712 E-mail: minna.raty@luke.fi
  • Haakana, Natural Resources Institute Finland (Luke), P.O.Box 2, FI-00790, Helsinki, Finland E-mail: helena.haakana@luke.fi
  • Heikkinen, Natural Resources Institute Finland (Luke), P.O.Box 2, FI-00790, Helsinki, Finland ORCID https://orcid.org/0000-0003-3527-774X E-mail: juha.heikkinen@luke.fi
  • Hotanen, Natural Resources Institute Finland (Luke), P.O.Box 68, FI-80100 Joensuu, Finland E-mail: juha-pekka.hotanen@luke.fi
  • Kuronen, Natural Resources Institute Finland (Luke), P.O.Box 2, FI-00790, Helsinki, Finland ORCID https://orcid.org/0000-0002-8089-7895 E-mail: mikko.kuronen@luke.fi
  • Pitkänen, Natural Resources Institute Finland (Luke), P.O.Box 68, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0002-7583-6297 E-mail: juho.pitkanen@luke.fi
article id 24044, category Research article
Shaohui Zhang, Lauri Korhonen, Timo Nummenmaa, Simone Bianchi, Matti Maltamo. (2024). How to implement the data collection of leaf area index by means of citizen science and forest gamification? Silva Fennica vol. 58 no. 5 article id 24044. https://doi.org/10.14214/sf.24044
Keywords: forest canopy; crowdsourcing; hinge angle; plant area index; smartphones
Highlights: Citizen science and gamification are proposed for collecting in situ forest leaf area index data; LAI can be estimated by taking smartphone images of forest canopies at 57° zenith angle; Twenty smartphone images per plot are enough to obtain accurate LAI measurements; Additional images may be required in forests with dense or uneven canopy structure.
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Leaf area index (LAI) is a critical parameter that influences many biophysical processes within forest ecosystems. Collecting in situ LAI measurements by forest canopy hemispherical photography is however costly and laborious. As a result, there is a lack of LAI data for calibration of forest ecosystem models. Citizen science has previously been tested as a solution to obtain LAI measurements from large areas, but simply asking citizen scientists to collect forest canopy images does not stimulate enough interest. As a response, this study investigates how gamified citizen science projects could be implemented with a less laborious data collection scheme. Citizen scientists usually have only mobile phones available for LAI image collection instead of cameras suitable for taking hemispherical canopy images. Our simulation results suggest that twenty directional canopy images per plot can provide LAI estimates that have an accuracy comparable to conventional hemispherical photography with twelve images per plot. To achieve this result, the mobile phone images must be taken at the 57° hinge angle, with four images taken at 90° azimuth intervals at five spread-out locations. However, more images may be needed in forests with large LAI or uneven canopy structure to avoid large errors. Based on these findings, we propose a gamified solution that could guide citizen scientists to collect canopy images according to the proposed scheme.

article id 24042, category Research article
Han Ren, Xuezhen Chen, Xiaohong Qin, Suyang Zhang, Chengqun Lv, Jiangmin Zhou, Hualin Chen. (2024). Bacterial fertilizer and filtered sludge enhance soil fertility and plant growth in a eucalyptus plantation. Silva Fennica vol. 58 no. 5 article id 24042. https://doi.org/10.14214/sf.24042
Keywords: nutrient content; Bacillus megaterium; enzyme activity; PGPR; plant yield
Highlights: Filtered sludge significantly increased eucalyptus growth, particularly after six months of application; Bacterial fertilizer significantly improved soil enzyme activities, specifically urease and sucrase; Bacterial fertilizer influenced overall soil fertility, promoting sustainable eucalyptus cultivation practices; The study indicates the potential of biofertilizers as environmentally friendly alternatives to chemical fertilizers.
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Plant growth-promoting rhizobacteria (PGPR) and filtered sludge are widely used to improve soil fertility and plant yields. In this study, we evaluated the impact of sludge and/or PGPR application on the nutrient contents and enzyme activities of the soil as well as on plant growth. We planted bare-root eucalyptus seedlings in (1) soil amended with filtered sludge from Nanning sugar factory (FS), (2) soil amended with filtered sludge + PGPR (BF), and (3) non-amended soil (control). Soil fertility and eucalyptus growth were determined after 3, 6, 9, and 12 months. Results demonstrated that FS treatment significantly increased eucalyptus growth compared to the control, particularly after six months. Bacterial fertilizer (BF) also increased soil urease and sucrase activities, although differences diminished over the study period. Our findings suggest that the integration of bacterial fertilizers and filtered sludge can serve as an effective and environmentally friendly strategy to improve soil health and promote sustainable eucalyptus cultivation. This research contributes to the growing body of evidence supporting the use of bio-fertilizers in forestry practices, highlighting their potential to reduce or replace the use of chemical fertilizers while increasing plant productivity.

  • Ren, College of Life and Environmental Sciences, Wenzhou University, Wenzhou, Zhejiang, 325035 China; Forestry College, Guangxi University, Daxue E Rd, Xixiangtang District, Nanning, Guangxi, 530004 China ORCID https://orcid.org/0000-0002-0156-0726 E-mail: renhan1225@163.com (email)
  • Chen, College of Life and Environmental Sciences, Wenzhou University, Wenzhou, Zhejiang, 325035 China E-mail: 21211270104@stu.wzu.edu.cn
  • Qin, Forestry College, Guangxi University, Daxue E Rd, Xixiangtang District, Nanning, Guangxi, 530004 China E-mail: qinxiaohong186@163.com
  • Zhang, College of Life and Environmental Sciences, Wenzhou University, Wenzhou, Zhejiang, 325035 China E-mail: 00811091@wzu.edu.cn
  • Lv, Forestry College, Guangxi University, Daxue E Rd, Xixiangtang District, Nanning, Guangxi, 530004 China E-mail: lvchengqun8@163.com
  • Zhou, College of Life and Environmental Sciences, Wenzhou University, Wenzhou, Zhejiang, 325035 China E-mail: rosechl@wzu.edu.cn
  • Chen, College of Life and Environmental Sciences, Wenzhou University, Wenzhou, Zhejiang, 325035 China ORCID https://orcid.org/0000-0002-3043-6919 E-mail: hualin2100@wzu.edu.cn (email)
article id 24031, category Research article
Jaakko Repola, Jaana Luoranen, Saija Huuskonen, Mikko Peltoniemi, Päivi Väänänen, Karri Uotila. (2024). Biomass models for young planted Norway spruce and naturally regenerated silver birch, aspen and rowan trees. Silva Fennica vol. 58 no. 5 article id 24031. https://doi.org/10.14214/sf.24031
Keywords: deciduous trees; Picea abies; tree biomass; model; multi-response
Highlights: Needle biomass was the greatest tree component of young spruce trees, and its proportion of whole biomass was 40-50%; The proportion of foliage biomass showed a decreasing tendency with tree height for all tree species; The existing models (Repola 2008, 2009) were not suitable for predicting spruce and birch biomass growing in young planted stands allocating too much biomass to roots and too little to crown.
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We developed tree level biomass (dry weight) models for Norway spruce (Picea abies [L.] H. Karst.), silver birch (Betula pendula Roth), rowan (Sorbus aucuparia L.) and aspen (Populus tremula L.) growing in young spruce dominated seedling stands with high mixture of broadleaves. The study material was collected from three planted Norway spruce seedling stands located on mineral soil in southern Finland. Biomass models were estimated by individual tree component (stem, living branches, foliage, stump, and roots with diameter of 2 mm) by using a multi-response approach (seemingly unrelated regression), which estimated the parameters of the sub-models (tree component) simultaneously. Even though the application and generalization of the developed models can be restricted by the limited material, they provide new information of seedling biomass allocation and more reliable biomass predictions for spruce and birch growing in young seedling stand compared with those of the commonly applied biomass models (Repola 2008, 2009) in Finland. Repola’s models (2008, 2009) tended to produce biased predictions for crown and below-ground biomasses of seedlings by allocating too much biomass to roots and too little to needle and branches. In addition, this study provides biomass models for aspen and rowan, which were not previously available.

  • Repola, Natural Resources Institute Finland (Luke), Ounasjoentie 6, FI-96200 Rovaniemi, Finland ORCID https://orcid.org/0000-0001-7086-0549 E-mail: jaakko.repola@luke.fi (email)
  • Luoranen, Natural Resources Institute Finland (Luke), Juntintie 154, FI-77600 Suonenjoki, Finland ORCID https://orcid.org/0000-0002-6970-2030 E-mail: jaana.luoranen@luke.fi
  • Huuskonen, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0001-8630-3982 E-mail: saija.huuskonen@luke.fi
  • Peltoniemi, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0003-2028-6969 E-mail: mikko.peltoniemi@luke.fi
  • Väänänen, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: paivi.vaananen@luke.fi
  • Uotila, Natural Resources Institute Finland (Luke), Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: karri.uotila@luke.fi
article id 23080, category Research article
Irene Kuhmonen, Annukka Näyhä, Miisa Solaranta, Janne Keränen. (2024). Can small and medium-sized companies increase the value added from wood-based side streams? Silva Fennica vol. 58 no. 5 article id 23080. https://doi.org/10.14214/sf.23080
Keywords: circular bioeconomy; dynamic capabilities; forest-based sector; strategic niche management; sustainability transition; wood-based SMEs
Highlights: Upgrading the use of wood-based side streams is necessary for circular bioeconomy; Small and medium-sized companies offer innovative solutions for using wood-based side streams; Companies utilizing the side streams host a variety of dynamic capabilities; Strong technological orientation reduces horizontal networking among innovative small and medium-sized companies; The forest-based regime resists the spread of innovations aiming for higher value added.
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One of the central goals of circular bioeconomy in the Finnish forest-based sector is upgrading the use of wood-based materials, especially wood-based side streams, to higher value-added products. However, despite these ambitions, most wood-based side streams are used in energy production. Within the forest-based sector, innovative solutions for higher value-added production of wood-based side streams are being developed within small and medium-sized companies (SMEs). Therefore, to promote the process of upscaling these solutions, understanding the success of these companies is pivotal. For this end, we conducted a qualitative study with 10 forest-based SMEs utilizing wood-based side streams to understand both the internal and external factors affecting their ability to scale up their business models. By applying the dynamic capabilities approach from management research and the strategic niche management approach from sociotechnical transition studies, we found that even though the companies are internally well positioned to succeed in their growth aspirations, they face barriers from the dominant forest-based regime. The studied SMEs are facing a mismatch between their own business models and the rules and operating principles of the forest-based sector based on linear economy. Overcoming these barriers and challenging the dominant structures within the Finnish forest-based regime would require joint efforts from the companies. However, the companies have a strong technological orientation, which makes them hesitant with regard to horizontal networking. They also operate in diverse markets, making it difficult for them to find common ground. As a result, the pressure for systemic transformation within the forest-based sector remains nominal.

  • Kuhmonen, University of Jyväskylä, School of Business and Economics, P.O. Box 35, FI-40014 University of Jyväskylä, Finland ORCID https://orcid.org/0000-0002-1407-8349 E-mail: irene.a.kuhmonen@jyu.fi (email)
  • Näyhä, University of Jyväskylä, School of Business and Economics, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: annukka.nayha@jyu.fi
  • Solaranta, University of Jyväskylä, School of Business and Economics, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: miisa.solaranta@gmail.com
  • Keränen, VTT Technical Research Centre of Finland Ltd, P.O. Box 1603, FI-40101 Jyväskylä, Finland ORCID https://orcid.org/0000-0002-5626-2929 E-mail: janne.keranen@vtt.fi
article id 23035, category Research article
Venla Wallius, Janni Kunttu, Elias Hurmekoski, Teppo Hujala, Anders Q. Nyrud, Hans F. Hoen. (2024). Attractiveness of wood-frame multi-storey buildings in seven European countries: consumer segmentation and the effect of fire safety information. Silva Fennica vol. 58 no. 5 article id 23035. https://doi.org/10.14214/sf.23035
Keywords: wood construction; consumer perceptions; consumer segmentation; fire safety
Highlights: There are significant differences in the perceived attractiveness of wooden multi-storey construction between consumer segments; According to the survey results, young consumers living in urban areas have the most interest in wooden multi-storey construction; Targeted marketing efforts are needed to inform potential consumers of wooden multi-storey construction.
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Wooden construction has the potential to contribute to climate change mitigation, and it is being promoted by the EU and national governments. However, several market barriers to wood-frame multi-storey building (WMSB), have been recognized, including obstacles in national building codes, lack of expertise in wood construction, and material durability concerns among end-users as well as other technical aspects. Given that increased wood construction is a target, understanding consumer perceptions of WMSB is crucial. In this study, consumer attitudes on WMSB were studied through consumer segmentation relying on demographic attributes. Further, the effect of providing fire safety information was explored. To this end, an online survey was deployed in seven European countries, with 7007 responses. The results show that in general, the awareness and attractiveness of WMSB is low amongst European consumers. Out of all respondents, 46% had not heard of WMSB before and only 12% stated that they are interested in the subject and know something about it, showing a clear lack of information and awareness within the general public. Significant differences in the perceived attractiveness of wooden multi-storey construction between consumer segments exist, with younger consumers and urban consumers being more attracted to living in WMSB than older or rural consumers. Fire safety was an important attribute affecting overall attractiveness, yet updated information regarding fire safety and control in WMSBs had a small but statistically significant negative effect on the perceived attractiveness. The results indicate that targeted marketing efforts are needed to inform potential consumers of WMSB and aspects related to fire safety effectively.

  • Wallius, European Forest Institute, Yliopistokatu 6 B, FI-80100, Joensuu, Finland; Jyväskylä University School of Business and Economics, P.O. Box 35, FI-40014 University of Jyväskylä, Finland ORCID https://orcid.org/0000-0003-2576-9994 E-mail: venla.j.wallius@jyu.fi (email)
  • Kunttu, University of Helsinki, Department of Forest Sciences, P.O. Box 4, FI-00014 University of Helsinki, Finland ORCID https://orcid.org/0000-0002-7298-3363 E-mail: janni.kunttu@helsinki.fi
  • Hurmekoski, University of Helsinki, Department of Forest Sciences, P.O. Box 4, FI-00014 University of Helsinki, Finland ORCID https://orcid.org/0000-0001-8717-7287 E-mail: elias.hurmekoski@helsinki.fi
  • Hujala, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0002-7905-7602 E-mail: teppo.hujala@uef.fi
  • Nyrud, Norwegian University of Life Sciences, Faculty of Environmental Sciences and Natural Resource Management, P.O. Box 5003, NO-1432 Ås, Norway E-mail: anders.qvale.nyrud@nmbu.no
  • Hoen, Norwegian University of Life Sciences, Faculty of Environmental Sciences and Natural Resource Management, P.O. Box 5003, NO-1432 Ås, Norway E-mail: hans.hoen@nmbu.no
article id 24011, category Research article
Alexander Kaulen, Benjamin Engler, Thomas Purfürst. (2024). Net carbon storage of supplied timber in highly mechanized timber harvest. Silva Fennica vol. 58 no. 4 article id 24011. https://doi.org/10.14214/sf.24011
Keywords: carbon balance; emissions; forest machine data; forestry technique; timber supply chain; traceability
Highlights: CO2 emissions from timber supply comprise only 1.5–5% of the CO2 stored in wood; Distance to the mill has the greatest influence on CO2 emissions; Successful application by combining a few variable parameters from forest machine data with constant parameters to create a net carbon storage of the supplied timber; Simple presentation of the net CO2 storage capacity of wood can influence policy positively.
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Highly mechanized timber harvesting and timber logistics emit CO2. In turn, the provided timber stores CO2 from the atmosphere as biogenic carbon. This basic assumption resulted in the calculation of net carbon storage of supplied timber. For this, we first developed a formula that represents the carbon content of freshly harvested timber. Coniferous wood contains about 734 kg CO2 m-3 and deciduous wood about 1000 CO2 m-3. Contrary to this, CO2 emissions from trucks, harvesters, and forwarders were calculated using the variable parameters for actual diesel consumption and the distance to the sawmill and constant parameters for the transport of the machine to the stand, lubricants, transport of operators, loading, and fabrication, supply, and maintenance. The method was tested on an actual harvest. The principal findings are that the method is practical, the net carbon storage of the supplied timber is reduced by 1.5% to 5% by harvesting and transport activities, and timber logistics is the largest contributor to emissions. The CO2 emissions for harvesters and forwarders are about 4 kg CO2 m-3, and for downstream timber logistics across all assortments and distances is 11 kg CO2 m-3. We conclude that the emissions are low, vis-a-vis the storage capacity. Emissions and a standardized calculation model are imperative. The model developed here for mapping the net carbon storage of roundwood highlights the climate protection performance of timber and contributes to optimizing climate-friendly timber supply chains.

  • Kaulen, KWF - Kuratorium für Waldarbeit und Forsttechnik e.V., Spremberger Straße 1, 64823 Groß-Umstadt, Germany; University of Freiburg, Chair of Forest Operations, Werthmannstr. 6, 79085 Freiburg, Germany ORCID https://orcid.org/0009-0006-2633-8132 E-mail: alexander.kaulen@kwf-online.de (email)
  • Engler, University of Freiburg, Chair of Forest Operations, Werthmannstr. 6, 79085 Freiburg, Germany ORCID https://orcid.org/0000-0003-2104-8209 E-mail: benjamin.engler@foresteng.uni-freiburg.de
  • Purfürst, University of Freiburg, Chair of Forest Operations, Werthmannstr. 6, 79085 Freiburg, Germany ORCID https://orcid.org/0000-0001-9661-0193 E-mail: thomas.purfuerst@foresteng.uni-freiburg.de
article id 24007, category Research article
Heikki Manninen, Hanna Lehtimäki, Riitta Kilpeläinen, Eila Lautanen, Kalle Kärhä. (2024). The qualifications and competence in supervisory and management skills among recently graduated Finnish forestry professionals. Silva Fennica vol. 58 no. 4 article id 24007. https://doi.org/10.14214/sf.24007
Keywords: competence gap; forestry organizations; generic skills; higher forestry education; working life orientation; work requirements
Highlights: A remarkably large share of forestry professionals has supervisory or managerial duties soon after their graduation; Supervisory skills were considered important for all forestry professionals regardless of their position; The respondents recognized notable gaps in their competence in supervisory and management skills relative to work requirements, which higher forestry education must react to.
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In the past decade, research and several surveys have indicated that the competence of higher forestry education graduates does not meet the requirements of working life regarding supervisory and management skills. The aim of this study was to discover what kind of supervisory and management competence is required in the daily tasks of early-career forestry professionals, and to what extent the teaching of these skills would be advisable to include in higher forestry education. The study was implemented through a Webropol survey of the Master’s of Forestry graduates and forestry engineers graduated between 2018–2021, which mapped their supervisory and managerial duties and skills. The study population was 1046 people, of which 30.4% responded to the survey. The principal finding was of an apparent extensive need for the abovementioned skills, as supervisory duties are typical in the work of forestry professionals soon after their graduation. In this study, approximately one fifth of the employed respondents worked in a supervisory position, one third had supervisory or managerial duties, and half had an indirect network of subordinates. Supervisory skills were seen important for all forestry professionals regardless of their position. Moreover, self-management, team management, and well-being and occupational health management were regarded as important skills. However, there appeared to be notable gaps in all these skills. The study results contribute to improving working life orientation in higher forestry education. Based on these findings, we recommend that teaching of these subjects should be implemented by integrating the topics into field-specific courses.

  • Manninen, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland; South-Eastern  Finland University of Applied Sciences, P.O. Box 68, FI-50101 Mikkeli, Finland ORCID https://orcid.org/0009-0001-4234-7649 E-mail: heikki.manninen@xamk.fi (email)
  • Lehtimäki, University of Eastern Finland, P.O. Box 1627, FI-70211 Kuopio, Finland ORCID https://orcid.org/0000-0002-9061-8984 E-mail: hanna.lehtimaki@uef.fi
  • Kilpeläinen, TTS Työtehoseura, P.O. Box 5, FI-05201 Rajamäki, Finland E-mail: riitta.kilpelainen@tts.fi
  • Lautanen, TTS Työtehoseura, P.O. Box 5, FI-05201 Rajamäki, Finland E-mail: eila.lautanen@tts.fi
  • Kärhä, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0002-8455-2974 E-mail: kalle.karha@uef.fi
article id 23077, category Research article
Timo Domisch, Saija Huuskonen, Juho Matala, Ari Nikula. (2024). Interactive effects of moose browsing and stand composition on the development of mixed species seedling stands. Silva Fennica vol. 58 no. 4 article id 23077. https://doi.org/10.14214/sf.23077
Keywords: boreal forest; tree species; Alces alces; ungulates; moose damage; mixed stands; meta-analysis
Highlights: We conducted meta-analyses assessing effect sizes of moose exclusion on seedling density and height, as well as regressions between stand density and deciduous seedling proportion and effect sizes; The more deciduous trees in a seedling stand, the smaller was the fencing effect, but at some point, the conifer seedling growth was impaired by too many deciduous seedlings. At the same time, the protective fencing effect increased with stand density; An appropriate deciduous admixture in conifer-dominated mixed seedling stands seems to improve moose damage tolerance.
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The moose (Alces alces L.), a common large herbivore in the boreal region, impairs forest regeneration by browsing on tree seedlings and saplings. Moose prefer deciduous species, but during winter more coniferous seedlings are used. We used meta-analyses, separately for deciduous and coniferous seedlings, for evaluating whether excluding moose browsing affected seedling density and height. In addition, we compared (1) deciduous seedling proportion, (2) stand density, (3) elapsed time from fencing and (4) estimated moose density with moose exclusion effect sizes. Fencing had a positive effect on coniferous seedling height. With more deciduous trees in a seedling stand, the fencing effect for both seedling height and density of coniferous seedlings decreased. On the other hand, the fencing effects increased with denser stands. At some point effect sizes turned to negative, and conifer species varied in their response to browsing. This implies that deciduous seedlings can protect conifers from browsing by moose up to some mixing ratio, but when deciduous seedling densities are too high, their negative effect increases, presumably through increased competition. Our results suggest that a moderate deciduous admixture in conifer-dominated mixed seedling stands can decrease moose damage but also underline the significance of timely silvicultural measures to minimize the negative effects of excessive deciduous seedlings and too dense stands. Due to differences in coniferous and deciduous species, as well as their compositions and amounts in studied experiments, more studies adjusted to local conditions are still needed to give exact measures for silvicultural recommendations.

  • Domisch, Natural Resources Institute Finland (Luke), Natural Resources, Yliopistokatu 6B, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0001-7026-1087 E-mail: timo.domisch@luke.fi (email)
  • Huuskonen, Natural Resources Institute Finland (Luke), Natural Resources, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: saija.huuskonen@luke.fi
  • Matala, Natural Resources Institute Finland (Luke), Natural Resources, Yliopistokatu 6B, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0002-5867-5057 E-mail: juho.matala@luke.fi
  • Nikula, Natural Resources Institute Finland (Luke), Natural Resources, Ounasjoentie 6, FI-96200 Rovaniemi, Finland ORCID https://orcid.org/0000-0001-8372-8440 E-mail: ari.nikula@luke.fi
article id 23068, category Research article
Anne Viljanen, Mikko Kurttila, Anne Toppinen. (2024). Retrofitting urban areas with wood: the origin of new projects in an emerging business ecosystem. Silva Fennica vol. 58 no. 4 article id 23068. https://doi.org/10.14214/sf.23068
Keywords: decision-making; business ecosystem; dynamic capabilities; project initiation; vertical building; wooden retrofits
Highlights: Decarbonizing building stock requires businesses to create new solutions for markets; Wood-material based retrofits offer new possibilities but are yet rare in urban context; The city-owned housing companies are found to be most aware of seizing opportunities of climate-benefits and facelifts in the less appealing suburban areas; Actors use dynamic capabilities in niche-level projects, yet the reconfiguring stage of business models is not visible.
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Nurturing a low-carbon residential building stock requires businesses to create new solutions for markets. Wood material-based retrofits would be one solution but have remained rare in the urban context. Our study explores the structure and dynamics of an emerging business ecosystem (BE) of wooden retrofits in Finland. We study wooden retrofit projects, from the perspective of the initial steps of local-level development. By applying the concepts of BE and dynamic capabilities, we aim to shed light on the role of actors and their early-stage decision-making and use actor mapping and qualitative analysis of 27 thematic interviews with retrofit businesses and municipal actors. Results show that project initiation has relied on individual champions seizing opportunities as a main base in building dynamic capabilities. Builders and customers are key actors of wooden retrofit projects in local BEs in Finland. The identified actors are further involved with future plans for wooden retrofits, some with ambitious deep renovation with wood. Motivations for projects are driven from urban densification strategies, improvement of suburban neighborhood attractiveness and from the efficiency of space utilization. Results further elaborate a certain degree of difficulty in the early-stage decision-making. This study contributes to the scientific knowledge of both the BE and the dynamic capability perspective by exploring a path to material-driven sustainable construction in the Finnish context. We provide new information on this emerging retrofit construction business with potentially significant international implications if scaled up more widely.

  • Viljanen, University of Helsinki, Faculty of Agriculture and Forestry, Latokartanonkaari 7, 00014 Helsingin yliopisto, Finland ORCID https://orcid.org/0000-0001-7166-8022 E-mail: anne.viljanen@helsinki.fi (email)
  • Kurttila, Natural Resources Institute Finland (Luke), Research and Customer Relationships, Yliopistokatu 6, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0001-5290-4771 E-mail: mikko.kurttila@luke.fi
  • Toppinen, University of Helsinki, Faculty of Agriculture and Forestry, Latokartanonkaari 7, 00014 Helsingin yliopisto, Finland ORCID https://orcid.org/0000-0003-0910-1505 E-mail: anne.toppinen@helsinki.fi
article id 23044, category Research article
Kyle Eyvindson, Annika Kangas, Olha Nahorna, Juliette Hunault-Fontbonne, Maria Potterf. (2024). Integrating wind disturbances into forest planning: a stochastic programming approach. Silva Fennica vol. 58 no. 4 article id 23044. https://doi.org/10.14214/sf.23044
Keywords: forest planning; wind damage; risk mitigation; stochastic optimization
Highlights: Assessing risk should focus on the objectives of the decision maker, not simply to minimize wind damage; We explored timber income-oriented objectives, maximizing net profit and maintaining a high even-flow of timber related income; Integrating wind disturbances had limited advantages when prioritizing net profits, however, the impact was dramatic when striving to maintain a high even-flow of timber.
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Forest disturbances challenge our ability to carefully plan for sustainable use of forest resources. As forest disturbances are stochastic, we cannot plan for the disturbance at any specific time or location. However, we can prepare for the possibility of a disturbance by integrating its potential intensity range and frequency when developing forest management plans. This study uses stochastic programming to integrate wind intensity (wind speed) and wind event frequency (number of occurrences) into the forest planning process on a small coastal Finnish forest landscape. We used a mechanistic model to quantify the critical wind speed for tree felling, with a Monte Carlo approach to include wind damage and salvage logging into forest management alternatives. We apply a stochastic programming model to explore two objectives: maximizing the expected forest net present value or maximizing the even-flow of income. To assess the effects of improper wind risk assumptions in planning, we compare the results when optimizing for correct versus incorrect wind intensity and frequency assumptions. When maximizing for net present value, the impacts of misidentifying wind intensity and frequency are minor, likely due to harvests planned immediately as trees reach maturity. For the case when maximizing even-flow of income, incorrectly identifying wind intensity and frequency severely impacts the ability to meet the required harvest targets and reduces the expected net present value. The specific utility of risk mitigation therefore depends on the planning problem. Overall, we show that incorporating wind disturbances into forest planning can inform forest owners about how they can manage wind risk based on their specific risk preferences.

  • Eyvindson, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, Campus Ås, Norway; Natural Resource Institute Finland (Luke), Bioeconomy and Environment, Laatokartanonkaari 9, 00790 Helsinki, Finland ORCID https://orcid.org/0000-0003-0647-1594 E-mail: kyle.eyvindson@nmbu.no (email)
  • Kangas, Natural Resources Institute Finland (Luke), Bioeconomy and Environment, Yliopistokatu 6, 80100 Joensuu, Finland ORCID https://orcid.org/0000-0002-8637-5668 E-mail: annika.kangas@luke.fi
  • Nahorna, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, Campus Ås, Norway ORCID https://orcid.org/0000-0002-5497-0315 E-mail: olha.nahorna@nmbu.no
  • Hunault-Fontbonne, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, Campus Ås, Norway ORCID https://orcid.org/0009-0004-1864-5162 E-mail: juliette.hunault@nmbu.no
  • Potterf, Ecosystem Dynamics and Forest Management Group, Technical University of Munich, Hans Carl-von-Carlowitz-Platz 2, 85354 Freising, Germany ORCID https://orcid.org/0000-0001-6763-1948 E-mail: maria.potterf@tum.de
article id 23073, category Research article
Curt Almqvist. (2024). High stand density improves seed production in seed orchards of the masting species Picea abies. Silva Fennica vol. 58 no. 3 article id 23073. https://doi.org/10.14214/sf.23073
Keywords: Norway spruce; thinning; cone production
Highlights: Cone production per tree in a Picea abies seed orchard is independent of stand density up to at least 800 stems ha–1; The higher the stand density, the higher per hectare cone production will be in the seed orchard.
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Reproduction in masting species is characterised by long intervals between good cone and seed production years, and only sparse reproduction between mast years. The physiological mechanisms behind masting, and how these are linked to internal resource status and external weather factors, is still a subject of scientific exploration and debate, as is the effect of climate change on masting. This study investigates cone production in one operational seed orchard in Sweden which was established with two different spacings and has since been subject to three tree thinning experiments. The spacings before thinning varied between 800 and 400 stems ha–1, and then thinning reduced the stand density in all trials to half, i.e. between 400 and 200 stems ha–1. In all three experiments cone production per tree was equal in un-thinned and thinned treatments, both in mast years and in non-mast years. Thus, the cone production per unit area was twice as high in the un-thinned areas. The conclusion from these experiments is that the establishment of Picea abies (L.) H. Karst. seed orchards with wide tree spacing is both a misuse of good orchard locations and bad economics.

article id 23030, category Research article
Harri Silvennoinen, Laura Pikkarainen, Heini Nakola, Matti Koivula, Liisa Tyrväinen, Jukka Tikkanen, Philip Chambers, Heli Peltola. (2024). Consistency of video and photo surveys in measuring attractiveness of forest stands managed with varying intensities. Silva Fennica vol. 58 no. 3 article id 23030. https://doi.org/10.14214/sf.23030
Keywords: environment; assessment; forest treatment; preference; visualization
Highlights: The more intensely a forest is treated silviculturally, the less it corresponds to people’s wishes and expectations for nature; It is possible to assess the attractiveness of nature from both photographs and videos; Evaluations from videos and images differed slightly, but these differences do not affect how treatments are ranked; Photo and video assessments are not easy for everyone.
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We evaluated the consistency of video, ordinary photo, and panoramic photo surveys in measuring the attractiveness (recreational use, scenic values etc.) of forest stands managed with varying intensities. We also evaluated possible effects on the results caused by the personal background of citizen respondents and how the respondents experienced the evaluation events. Our experimental sites were in mature Scots pine (Pinus sylvestris L.) forests in eastern Finland and included two replicate sites which were unharvested (control, basal area 26 m2 ha–1), a selective cutting site (basal area 18 m2 ha–1), small openings sites (gap cut) with 5 and 20% retained trees, respectively, and one site which was clear cut with 3% retained trees. In our study, 71 volunteer forestry students evaluated the attractiveness of these sites from an ordinary photo, a panoramic photo, and a video, with a 0–10 scale. Based on this study, the unharvested forest was the most attractive and clear cutting was the least attractive, regardless of the evaluation method. This result was in line with a previous study using on-site evaluations of the same sites. The differences of respondents considering in how easy they felt to assess the attractiveness of the environment as a whole and in using different visualisation methods affected the result, unlike background variables of the respondents. The results of forest attractiveness were consistent between panoramic and ordinary photos, and the attractiveness scoring was slightly higher for them than for the video. We conclude that all the compared visualisation methods seem to be suitable for assessment of the attractiveness of forest views.

  • Silvennoinen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0002-9095-7986 E-mail: harri.silvennoinen@uef.fi (email)
  • Pikkarainen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0001-5301-3639 E-mail: laura.pikkarainen@uef.fi
  • Nakola, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: heini.nakola@gmail.com
  • Koivula, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0001-6415-4904 E-mail: matti.koivula@luke.fi
  • Tyrväinen, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0001-5144-7150 E-mail: liisa.tyrvainen@luke.fi
  • Tikkanen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0002-9693-8209 E-mail: jukka.tikkanen@uef.fi
  • Chambers, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0002-0586-3142 E-mail: philip.chambers@uef.fi
  • Peltola, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0003-1384-9153 E-mail: heli.peltola@uef.fi
article id 23018, category Research article
Anssi Ahtikoski, Kari Väätäinen, Perttu Anttila, Juha Laitila, Antti Mutanen, Jari Lindblad, Lauri Sikanen, Johanna Routa. (2024). The effects of the EU’s forest-related policies on harvesting costs in Finland. Silva Fennica vol. 58 no. 3 article id 23018. https://doi.org/10.14214/sf.23018
Keywords: forests; scenario approach; harvesting costs; directives and regulations; EU legislation; structural systems analysis
Highlights: The EU’s forest-related policies set direct and indirect limitations on the harvesting operations and operating environment; The effect on harvesting costs of Directives and Regulations depends on the magnitude of policy implementation; In a high-impact scenario, harvesting costs could increase by as much as 18% from the current level of costs.
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The EU’s influence on national forest policies is growing, and the implementation of forest-related policies proposed by the Commission will affect the practice of forestry in Europe. For instance, the Nature Restoration Law sets concrete areal goals for restoring forest ecosystems and for conservation, the Deforestation Regulation requires meticulous tracking of wood’s origin, and the renewed Renewable Energy Directive (RED III) sets new criteria to sustainable forest biomass procurement. So far there have been no studies that have looked into the impacts from the economic and operational point of view. In this study, structural systems analysis was first performed to discover the relevant variables (and their functioning) associated with the roundwood harvesting operations and the operating environment. A scenario approach was then applied to capture the potential levels of implementation of the EU’s forest-related policies. Finally, using different scenarios (low-, moderate- and high-impact) and a systems analysis framework, the impact of alternative levels of implementation was quantified in terms of harvesting costs, measured in € m–3. The results indicate that with the low- and moderate-impact scenarios the harvesting costs would increase by less than 10% from the current levels in three different regions in Finland. Such an increase (less than 10%) could be tolerated over a period of a few years, but a sudden increase is likely to lead to challenges to the running of businesses. With the high-impact scenario the harvesting costs would increase by between 15% and 18%, depending on the region. This magnitude of increase (of approximately a sixth) corresponds to a severe change in the roundwood harvesting operations and operating environment.

  • Ahtikoski, Natural Resources Institute Finland (Luke), Tekniikankatu 1, FI-33720 Tampere, Finland ORCID https://orcid.org/0000-0003-1658-3813 E-mail: anssi.ahtikoski@luke.fi (email)
  • Väätäinen, Natural Resources Institute Finland (Luke), Yliopistonkatu 6, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0002-6886-0432 E-mail: kari.vaatainen@luke.fi
  • Anttila, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0002-6131-392X E-mail: perttu.anttila@luke.fi
  • Laitila, Natural Resources Institute Finland (Luke), Yliopistonkatu 6, FI-80100 Joensuu, Finland E-mail: juha.laitila@luke.fi
  • Mutanen, Natural Resources Institute Finland (Luke), Yliopistonkatu 6, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0003-0533-9356 E-mail: antti.mutanen@luke.fi
  • Lindblad, Natural Resources Institute Finland (Luke), Yliopistonkatu 6, FI-80100 Joensuu, Finland ORCID https://orcid.org/0009-0003-6766-6587 E-mail: jari.lindblad@luke.fi
  • Sikanen, Natural Resources Institute Finland (Luke), Yliopistonkatu 6, FI-80100 Joensuu, Finland E-mail: lauri.sikanen@luke.fi
  • Routa, Natural Resources Institute Finland (Luke), Yliopistonkatu 6, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0001-7225-1798 E-mail: johanna.routa@luke.fi
article id 23065, category Research article
Pentti Niemistö, Heikki Korpunen, Yrjö Nuutinen. (2024). Harvesting efficiency and understory damage with different cutting methods on two storied stands of silver birch and Norway spruce. Silva Fennica vol. 58 no. 2 article id 23065. https://doi.org/10.14214/sf.23065
Keywords: productivity; thinning; costs; harvesting damage; mixed stand; continuous cover; multistorey forest
Highlights: With careful, well-timed harvesting of overstory birch, it is possible to successfully use initial understory spruce in producing the next tree generation; Preserving understory spruce when cutting overstory birch reduced harvesting productivity by 30% in comparison to unpreserved harvesting; Removal of overstory birch had a positive correlation in cutting productivity but negative correlation in the quality of remaining understory spruce.
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Uneven-aged forests set certain challenges for cut-to-length harvesting work. It is a challenge to cost-effectively remove larger trees while leaving a healthy understory for regrowth. The study’s aim was to evaluate productivity and costs of harvesting two-storied Silver birch (Betula pendula Roth) and Norway spruce (Picea abies (L.) H. Karst.) stands by creating time consumption models for cutting, and using existing models for forwarding. Damage to the remaining understory spruce was also examined. Four different harvesting methods were used: 1) all dominant birches were cut; 2) half of them thinned and understory was preserved; compared to 3) normal thinning of birch stand without understory; and 4) clear cutting of two-storied stand. Results showed the time needed for birch cutting was 26–30% lower when the understory was not preserved. Pulpwood harvesting of small sized spruces that prevent birch cutting was expensive, especially because of forwarding of small amounts with low timber density on the strip roads. Generally, when taking the cutting and forwarding into account, the unit cost at clear cuttings was lowest, due to lesser limitations on work. It was noted that with increasing removal from 100 to 300 m3 ha–1, the relative share of initial undamaged spruces after the harvest decreased from 65 to 50% when the aim was to preserve them.  During summertime harvesting, the amount of stem damage was bigger than during winter. In conclusion, two-storied stands are possible to transit to spruce stands by accepting some losses in harvesting productivity and damages on remaining trees.

  • Niemistö, Natural Resources Institute Finland (Luke), Natural resources, Kampusranta 9 C, FI-60320 Seinäjoki, Finland ORCID https://orcid.org/0000-0002-9152-2108 E-mail: ext.pentti.niemisto@luke.fi
  • Korpunen, Norsk institutt for bioøkonomi (NIBIO), Division of Forest and Forest Resources, Department of Forest operations and digitalization, Divisjon for skog og utmark, Avdeling for Driftsteknikk og digitalisering, Høgskoleveien 8, 1433 Ås, Norway ORCID https://orcid.org/0000-0001-9749-5684 E-mail: heikki.korpunen@nibio.no
  • Nuutinen, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6 B, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0003-3360-4444 E-mail: yrjo.nuutinen@luke.fi (email)
article id 23045, category Research article
Laura Pikkarainen, Harri Strandman, Eerik Vento, Aaron Petty, Olli-Pekka Tikkanen, Antti Kilpeläinen, Heli Peltola. (2024). Effects of forest conservation and management on timber, ecosystem carbon, dead wood and habitat suitability area in a boreal forest under climate change. Silva Fennica vol. 58 no. 2 article id 23045. https://doi.org/10.14214/sf.23045
Keywords: saproxylic species; simulation; carbon storage; forest fertilization; Fennoscandia; forest ecosystems services; improved regeneration material
Highlights: Increasing forest conservation areas increased all other ecosystem services except timber yield. Intensive forest management enhanced this increase; Increased conservation area decreased timber yield, but intensive forest management reduced this effect and even overcompensated it at the end of simulation period in the 10% conservation scenario with intensified forest management; Climate change increased all other ecosystem services, except carbon stocks.
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We used forest ecosystem model simulations to study how forest conservation and management intensity affected timber yield, ecosystem carbon stocks, amount of dead wood, and habitat suitability area in a middle boreal forest region of Finland under changing climate over a 90-year simulation period. We used the following forest conservation and management scenarios: baseline forest management (BM), BM with 10 or 20% increase of conservation area with or without intensified forest management (i.e. improved forest regeneration material and forest fertilization). The simulations were done under current climate (reference period of 1981–2010), and Representative Concentration Pathway (RCP) climate change projections under the RCP2.6 and RCP4.5 forcing scenarios. Overall, increasing the forest conservation area decreased timber yield and increased the ecosystem carbon stock, the amount of dead wood and consequently the area of suitable habitat for saproxylic species. The use of intensified forest management reduced the loss of timber yield, increased ecosystem carbon stock, the amount of dead wood and area of suitable habitat for saproxylic species. At the end of simulation period, the use of intensified forest management even overcompensated (4–6% higher) the timber loss from 10% increase of conservation area. Under changing climate, timber yield, the amount of dead wood and the area of suitable habitats for saproxylic species increased. To conclude, with intensified forest management it is possible, in the short term, to decrease the loss of timber yield through increased forest conservation area and in the long term maintain or even increase it compared to baseline forest management.

  • Pikkarainen, School of Forest Sciences, University of Eastern Finland, Yliopistokatu 7, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0001-5301-3639 E-mail: laura.pikkarainen@uef.fi (email)
  • Strandman, School of Forest Sciences, University of Eastern Finland, Yliopistokatu 7, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0002-9400-6424 E-mail: harri.strandman@uef.fi
  • Vento, School of Forest Sciences, University of Eastern Finland, Yliopistokatu 7, FI-80100 Joensuu, Finland E-mail: eerik.vento@gmail.com
  • Petty, School of Forest Sciences, University of Eastern Finland, Yliopistokatu 7, FI-80100 Joensuu, Finland ORCID https://orcid.org/0009-0006-6595-1386 E-mail: aaron.petty@uef.fi
  • Tikkanen, School of Forest Sciences, University of Eastern Finland, Yliopistokatu 7, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0002-3875-2772 E-mail: olli-pekka.tikkanen@uef.fi
  • Kilpeläinen, School of Forest Sciences, University of Eastern Finland, Yliopistokatu 7, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0003-4299-0578 E-mail: antti.kilpelainen@uef.fi
  • Peltola, School of Forest Sciences, University of Eastern Finland, Yliopistokatu 7, FI-80100 Joensuu, Finland E-mail: heli.peltola@uef.fi
article id 23029, category Research article
Amira Fetouab, Nicole J. Fenton, Nelson Thiffault, Martin Barrette. (2024). Planting density and mechanical site preparation effects on understory composition, functional diversity and planted black spruce growth in boreal forests. Silva Fennica vol. 58 no. 2 article id 23029. https://doi.org/10.14214/sf.23029
Keywords: silviculture; paludification; reforestation; plantation establishment; community ecology; functional ecology
Highlights: Mechanical site preparation and planting density interacted to influence understory diversity and composition in plowed plots; Careful logging alone resulted in higher density of naturally established conifers; Planting density and site preparation alone had no effect on tree height and diameter; Findings provide insights for managing paludified forests and enhancing forest ecosystem functions.
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Mechanical site preparation (MSP) is used prior to planting to control competing vegetation and enhance soil conditions, particularly in areas prone to paludification. Tree planting density can be adapted to the management context and objectives, as it influences yield and wood quality. However, the combined effects of MSP and planting density on understory vegetation composition, functional traits, and diversity remain uncertain. We thus conducted a study in the Clay Belt region of northwestern Quebec, Canada. After careful logging, the study area was divided into nine sites, each receiving one of three treatments: plowing, disc trenching, or no preparation. Sites were further divided into two, with black spruce (Picea mariana [Mill.] Britton, Sterns & Poggenb.) seedlings planted at either a low planting density of 1100 seedlings ha-1 or a high planting density of 2500 seedlings ha-1. After nine years, we assessed understory composition, diversity, key functional traits, sapling density and growth of planted trees. Careful logging alone led to a higher density of naturally established conifers compared to plowing or disc trenching. The interaction between planting density and MSP significantly influenced understory diversity and composition in plowed plots. Understory composition was affected by the soil C/N ratio, coniferous species, and deciduous species density. The growth of black spruce was notably enhanced with higher planting density in the plow treatment only. Neither planting density nor MSP alone affected tree height and diameter. Our results suggest that combining plowing with high-density planting can enhance stand growth and improve forest productivity. These findings guide future research on paludified forests.

  • Fetouab, Institute for Forest Research and Centre for Forest Research, Université du Québec en Abitibi-Témiscamingue, 445 boul. de l’Université, Rouyn-Noranda, QC J9X 5E4, Canada E-mail: amira.fetouab@uqat.ca
  • Fenton, Institute for Forest Research and Centre for Forest Research, Université du Québec en Abitibi-Témiscamingue, 445 boul. de l’Université, Rouyn-Noranda, QC J9X 5E4, Canada ORCID https://orcid.org/0000-0002-3782-2361 E-mail: nicole.fenton@uqat.ca
  • Thiffault, Institute for Forest Research and Centre for Forest Research, Université du Québec en Abitibi-Témiscamingue, 445 boul. de l’Université, Rouyn-Noranda, QC J9X 5E4, Canada; Canadian Wood Fibre Centre, Natural Resources Canada, 1055 du P.E.P.S, P.O. Box 10380, Sainte-Foy Stn, Québec, QC G1V 4C7, Canada ORCID https://orcid.org/0000-0003-2017-6890 E-mail: nelson.thiffault@canada.ca (email)
  • Barrette, Institute for Forest Research and Centre for Forest Research, Université du Québec en Abitibi-Témiscamingue, 445 boul. de l’Université, Rouyn-Noranda, QC J9X 5E4, Canada; Direction de la recherche forestière, Ministère des Ressources naturelles et des Forêts, 2700 rue Einstein, Québec, QC G1P 3W8, Canada ORCID https://orcid.org/0000-0001-5937-382X E-mail: martin.barrette@mffp.gouv.qc.ca
article id 23025, category Research article
Gundega Done, Jānis Ozoliņš, Guna Bagrade, Jurģis Jansons, Jānis Baumanis, Alekss Vecvanags, Dainis Jakovels. (2024). A case study for best suitable methods of monitoring demographic structure in cervid populations to predict increasing forest damages. Silva Fennica vol. 58 no. 2 article id 23025. https://doi.org/10.14214/sf.23025
Keywords: age structure; Alces alces; Cervus elaphus; camera trapping; damage to trees; pellet group counts; sex structure
Highlights: The study examines the interaction between sex-age structure of deer populations and forest damage throughout the country (Latvia); In pine stands, a higher female biased pellet group density index correlates with larger damage to pine trees through the browsing or debarking; Counting pellet groups and setting trail cameras both provide useful data on sex and age structure in moose and red deer populations that can be used for population monitoring.
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The overall increase of ungulate populations in modern Europe has contributed to conflicts in national economies, particularly between game management and the forestry sector. This study assessed damage risks to young pine (Pinus sylvestris L.), spruce (Picea abies (L.) H. Karst.) and aspen (Populus tremula L.) stands at two spatial scales. One level assessed the interaction between sex-age structure of cervid populations, measured by pellet group density, and forest damages, measured as the percentage of heavily browsed trees in 2040 stand surveys distributed proportionally throughout the country. The second level compared pellet counts and trail-camera-based records of moose (Alces alces L.) and red deer (Cervus elaphus L.) presence in a pilot study area. We examined whether 1) there is a correlation between damage amount and ungulate population structure and 2) are the data from trail cameras suitable for wider use in monitoring ungulate population structure. The study confirmed significantly higher moose pellet group densities in pine than in spruce and aspen stands. Pine damages were greater in stands with higher moose pellet group density, especially with female moose prevailing over male moose density index. The red deer pellet group densities were significantly higher in heavily damaged pine and spruce stands, regardless of sex-age structure. In most cases, there were no statistically significant differences between the two survey methods of ungulate population structure by using pellet count transects and trail camera fixations. Both methods provide comparable data on sex-age structure in moose and red deer populations if seasonal and habitat-predicted biases are considered. However, trail cameras are more widely applicable and easier to use by hunters than pellet counts.

  • Done, Latvian State Forest Research Institute ‘Silava’, Riga street 111, Salaspils, LV-2169, Latvia ORCID https://orcid.org/0000-0002-2122-7154 E-mail: gundega.done@silava.lv (email)
  • Ozoliņš, Latvian State Forest Research Institute ‘Silava’, Riga street 111, Salaspils, LV-2169, Latvia ORCID https://orcid.org/0000-0002-6647-9128 E-mail: janis.ozolins@silava.lv
  • Bagrade, Latvian State Forest Research Institute ‘Silava’, Riga street 111, Salaspils, LV-2169, Latvia ORCID https://orcid.org/0000-0002-1031-0665 E-mail: guna.bagrade@gmail.com
  • Jansons, Latvian State Forest Research Institute ‘Silava’, Riga street 111, Salaspils, LV-2169, Latvia E-mail: jurgis.jansons@silava.lv
  • Baumanis, Latvian State Forest Research Institute ‘Silava’, Riga street 111, Salaspils, LV-2169, Latvia E-mail: jbaumanis@inbox.lv
  • Vecvanags, Institute for Environmental Solutions ‘Lidlauks’, Cēsis, LV-4126, Latvia ORCID https://orcid.org/0000-0003-1233-764X E-mail: alekss.vecvanags@vri.lv
  • Jakovels, Institute for Environmental Solutions ‘Lidlauks’, Cēsis, LV-4126, Latvia ORCID https://orcid.org/0000-0002-2969-5972 E-mail: dainis.jakovels@vri.lv
article id 23014, category Research article
Hao Xiong, Yong Pang, Wen Jia, Yu Bai. (2024). Forest stand delineation using airborne LiDAR and hyperspectral data. Silva Fennica vol. 58 no. 2 article id 23014. https://doi.org/10.14214/sf.23014
Keywords: canopy height model; automatic delineation; merge rule; over-segmentation
Highlights: Delineate forest stands by the fusion of airborne LiDAR and hyperspectral data automatically; The forest height, canopy closure, and species information were taken into account during the delineation process, aligning with forest management in reality; The delineation accuracy was verified through comparison with three reference data sources commonly used in forest management.
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Forest stands, crucial for inventory, planning, and management, traditionally rely on time-consuming visual analysis by forest managers. To enhance efficiency, there is a growing need for automated methods that take into account essential forest attributes. In response, we propose a novel approach utilizing airborne Light Detection and Ranging (LiDAR) and hyperspectral data for automated forest stand delineation. Our approach initiates with over-segmentation of the Canopy Height Model (CHM), followed by attribute calculation for each segment using both CHM and hyperspectral data. Two rules are applied to merge homogeneous segments and eliminate others based on calculated attributes. The effectiveness of our method was validated using three types of reference forest stands with two indices: the explained variance (R2) and Intersection over Union (IoU). Results from our study demonstrated notable accuracy, with a R2 of 97.35% and 97.86% for mean tree height and mean diameter at breast height (DBH), respectively. The R2 for mean canopy height is 81.80%, outperforming manual delineation by 7.31% and multi-scale segmentation results by 2.13%. Furthermore, our approach achieved high IoU values, which indicates a strong spatial agreement with manually delineated forest stands and leading to fewer manual adjustments when applied directly to forest management. In conclusion, our forest stand delineation method enhances both internal consistency and spatial accuracy. This method contributes to improving practical performance and forest management efficiency.

  • Xiong, Institute of Forest Resource Information Techniques, Chinese Academy of Forestry, Beijing 100091, China; Key Laboratory of Forestry Remote Sensing and Information System, National Forestry and Grassland Administration, Beijing 100091, China; School of Geospatial Engineering and Science, Sun Yat-sen University, Zhuhai 519082, China ORCID https://orcid.org/0000-0003-4432-2485 E-mail: xiongh29@mail2.sysu.edu.cn
  • Pang, Institute of Forest Resource Information Techniques, Chinese Academy of Forestry, Beijing 100091, China; Key Laboratory of Forestry Remote Sensing and Information System, National Forestry and Grassland Administration, Beijing 100091, China ORCID https://orcid.org/0000-0002-9760-6580 E-mail: pangy@ifrit.ac.cn (email)
  • Jia, Institute of Forest Resource Information Techniques, Chinese Academy of Forestry, Beijing 100091, China; Key Laboratory of Forestry Remote Sensing and Information System, National Forestry and Grassland Administration, Beijing 100091, China E-mail: jiawen@ifrit.ac.cn
  • Bai, Institute of Forest Resource Information Techniques, Chinese Academy of Forestry, Beijing 100091, China; Key Laboratory of Forestry Remote Sensing and Information System, National Forestry and Grassland Administration, Beijing 100091, China E-mail: baiyu9224@163.com
article id 23072, category Research article
Matti Haapanen. (2024). Realised genetic gains from past Finnish silver birch seed orchards. Silva Fennica vol. 58 no. 1 article id 23072. https://doi.org/10.14214/sf.23072
Keywords: Betula pendula; seed orchards; genetic gain; forest reproductive materials
Highlights: Seed orchard materials generally outperformed unimproved trees in all growth and quality traits, but individual seed orchards showed substantial performance variability; Realised gains in stem volume and forking showed an increasing temporal trend, whereas gains in branch quality were positive and steady; Gains in stem volume and branch quality trade off slightly; The bi-clonal seed orchard variety “JR-2” emerged as the best overall performer across all the seed orchards, whereas the older variety “JR-1” was mediocre; An alternative statistical concept, D-value, was introduced to compare levels of genetic gains for different scaled, normally distributed traits.
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Genetic gains realised through silver birch (Betula pendula Roth) seed orchards were studied using data from common-garden trials established at 34 sites in southern and central Finland. The test materials include seedlots representing 19 commercial seed orchards that operated between 1972 and 2009, and 21 natural stands. All the trials were assessed for several growth and quality traits between the ages of 9 and 24. Realised gains were estimated based on univariate linear mixed models with corrections for latitudinal seed transfer effects. Overall, seed orchard materials outperformed unimproved reference materials in all the traits but results for individual seed orchards varied substantially. Stem volume gains ranged from 1.0% to 31.1%. Improved trees had, on average, 6.8% (up to 26.7%) fewer ramicorn branches and 16.2% (up to 57.6%) fewer forks than unimproved trees. Branch and overall quality showed consistently positive gains. More recently established seed orchards performed better than older ones, and seed orchards with fewer clones outperformed those with dozens of clones. "JR-1" and "JR-2" bi-clonal seed orchards fared differently, with "JR-1" showing modest genetic gains and "JR-2" emerging as the top overall performer across all seed orchards. An alternative statistical concept, the D-value, was utilised to assess the magnitude of genetic gain for different scaled, normally distributed traits. Average D-values implied a similar level of improvement for stem volume, branch quality, and forking, and a smaller gain for stem slenderness and the number of ramicorn branches. The results for individual seed orchards suggest a slight trade-off between stem volume growth and branch quality.

article id 23062, category Research article
Tobias Semberg, Anders Nilsson, Rolf Björheden, Linnea Hansson. (2024). Real-time target point identification and automated log grasping by a forwarder, using a single stereo camera for both object detection and boom-tip control. Silva Fennica vol. 58 no. 1 article id 23062. https://doi.org/10.14214/sf.23062
Keywords: vector analysis; forwarder; forest operations; loading; boom automation; object detection; stereo camera
Highlights: Simple target-point detection in real time using only a stereo camera; Sturdiness ensured through the simple feedback system based on the same camera; Automated boom-tip control and log grasping successfully tested on full-sized forwarder; A step toward semi-automation (operator support) or autonomous forwarding.
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The forest industry is constantly striving to increase productivity and cut costs, and many research and innovation projects are currently focusing on semi-automated or autonomous systems. A key element, with several possible solutions, is automated log grasping, where researchers and manufacturers are looking for efficient and sturdy ways to solve the task in real-time forwarding operations. This study presents a simple method for automated log grasping using only a single stereo camera for object detection (log and grapple) and a simple controller moving the boom, with feedback from the camera as boom-tip control. The accuracy, precision, and repeatability of the method was tested on a full-scale forwarder. Boom movements were examined from two different start positions in relation to the target position, with the log placed at three different angles. The overall log-grasping success was also evaluated. The tests were performed in a full-scale, real-time operation, without hand-eye calibration or other sensor data from the machine. The method was precise, with high repeatability, but the grasping point showed a minor systematic offset, depending on log angle. However, the deviation in accuracy was too small to affect the success rate. In practice, the most difficult log angles can be avoided by moving the machine slightly. The log grasping method may become part of an autonomous forwarding system or could provide operator support in semi-automated systems.

  • Semberg, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, 751 83 Uppsala, Sweden E-mail: tobias.semberg@skogforsk.se
  • Nilsson, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, 751 83 Uppsala, Sweden E-mail: anders.nilsson@skogforsk.se
  • Björheden, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, 751 83 Uppsala, Sweden ORCID https://orcid.org/0000-0002-4158-102X E-mail: rolf.bjorheden@skogforsk.se
  • Hansson, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, 751 83 Uppsala, Sweden ORCID https://orcid.org/0000-0002-9788-1734 E-mail: linnea.hansson@skogforsk.se (email)
article id 23058, category Research article
Johanna Jetsonen, Annamari Laurén, Heli Peltola, Olli Muhonen, Juha Nevalainen, Veli-Pekka Ikonen, Antti Kilpeläinen, Eeva-Stiina Tuittila, Elisa Männistö, Nicola Kokkonen, Marjo Palviainen. (2024). Effects of nitrogen fertilization on the ground vegetation cover and soil chemical properties in Scots pine and Norway spruce stands. Silva Fennica vol. 58 no. 1 article id 23058. https://doi.org/10.14214/sf.23058
Keywords: boreal forest; competition index; forest fertilization; upland forests; vegetation cover
Highlights: Nitrogen (N) fertilization decreased the total cover of ground vegetation; N fertilization decreased the cover of mosses and dwarf shrubs and increased the cover of herbaceous plants; N concentration of the mor humus layer increased with the N fertilization; The magnitude of these changes depended on the intensity of the N fertilization.
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The aim of this study was to investigate the short-term effects of nitrogen (N) fertilization intensity on the ground vegetation cover and soil chemical properties  in two Scots pine (Pinus sylvestris L.)  and two Norway spruce (Picea abies (L.) Karst.) dominated stands on upland forest sites in Eastern Finland. The fertilizer was applied using a helicopter in the spruce stands and a forwarder in the pine stands. The distribution and the amount of fertilizer was measured with funnel traps. Cover of each species of ground vegetation was estimated  before fertilization and 3–4 years after it in pine and 2–3 years after it in spruce stands. Further, the cover observations were aggregated by plant types. Based on measurements, we analyzed the effects of the funnel-trap-observed amount of N fertilizer on the cover and plant type composition of ground vegetation and soil N and C concentration. In addition, we analyzed  the impacts of competition caused by trees on the ground vegetation cover based on competition indices. N fertilization increased the cover of herbaceous plants and decreased the cover of mosses and dwarf shrubs, and the total cover of ground vegetation. Further, it increased the N concentration of the mor humus layer. The magnitude of the changes increased with the intensity of the N fertilization. The competition caused by trees did not affect the cover of ground vegetation.

  • Jetsonen, Department of Forest Sciences, University of Helsinki, P.O. Box 27, 00014 University of Helsinki, Finland E-mail: johanna.jetsonen@helsinki.fi (email)
  • Laurén, Department of Forest Sciences, University of Helsinki, P.O. Box 27, 00014 University of Helsinki, Finland; Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland ORCID https://orcid.org/0000-0002-6835-9568 E-mail: annamari.lauren@helsinki.fi
  • Peltola, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland E-mail: heli.peltola@uef.fi
  • Muhonen, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland ORCID https://orcid.org/0009-0007-4051-8567 E-mail: olli.muhonen@forestvital.com
  • Nevalainen, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland ORCID https://orcid.org/0009-0000-2972-4385 E-mail: juha.hs.nevalainen@gmail.com
  • Ikonen, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland ORCID https://orcid.org/0000-0003-1732-2922 E-mail: veli-pekka.ikonen@uef.fi
  • Kilpeläinen, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland ORCID https://orcid.org/0000-0003-4299-0578 E-mail: antti.kilpelainen@uef.fi
  • Tuittila, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland E-mail: eeva-stiina.tuittila@uef.fi
  • Männistö, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland ORCID https://orcid.org/0000-0003-3869-6739 E-mail: elisa.mannisto@uef.fi
  • Kokkonen, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, 80101 Joensuu, Finland ORCID https://orcid.org/0000-0003-0197-2672 E-mail: nicola.kokkonen@uef.fi
  • Palviainen, Department of Forest Sciences, University of Helsinki, P.O. Box 27, 00014 University of Helsinki, Finland E-mail: marjo.palviainen@helsinki.fi
article id 23032, category Research article
Maria Anna Gartner, Matthias Kaltenbrunner, Manfred Gronalt. (2024). Dynamic box assignment planning in log yards. Silva Fennica vol. 58 no. 1 article id 23032. https://doi.org/10.14214/sf.23032
Keywords: storage assignment; seasonality; binary integer program; optimisation model; sawmill storage
Highlights: Seasonal change of assortments calls for dynamic box assignment planning in log yards; Multi-period planning better suited for dynamic problem, however period per period planning improves with decreasing capacity on the log yard; Rearrangement of assortment amounts to 8–11% of total transportation distance (loaded travelled distances of transportation vehicle); Considering separate box allocation (storage and ejection), which results in double stage planning of box allocation, benefits most if 10% additional volume may be cut in to clear the box.
Abstract | Full text in HTML | Full text in PDF | Author Info

The situation on the log yard changes seasonally and also over the years. The quantities of assortments to be stored, their number and also the type of wood can change. To respond to this, we have developed a dynamic log yard planning model for assigning roundwood to specific ejection boxes and storage areas in order to minimise the overall transport distances of the loaded transportation vehicles on the log yard, including any possible re-allocation of assortments. The study centres on the log yard of a medium-sized hardwood sawmill in Europe, with actual cutting data from a six-month period. We are comparing a multi-period binary integer program with a model that operates on a period per period basis and a solution approach that splits the problem into two subproblems and solves them sequentially. The models undergo testing with decreasing space capacities at the storage boxes on the log yard and are compared. If capacity is continuously decreasing from 100% to 80%, then period per period planning is on average 13% worse than multi-period planning. We also investigate how the solutions change when twice as many or half as many assortments are stored at the log yard. In addition, we study how much the solutions improve when logs can be removed from the storage boxes to clear them and release them for other material in the following period.

  • Gartner, University of Natural Resources and Life Sciences, Department of Economics and Social Sciences, Institute of Production and Logistics, Feistmantelstrasse 4, 1180 Vienna, Austria ORCID https://orcid.org/0000-0001-8547-718X E-mail: maria.gartner@boku.ac.at (email)
  • Kaltenbrunner, improvem GmbH, Holzinnovationszentrum 1a, 8740 Zeltweg, Austria ORCID https://orcid.org/0000-0002-1178-0087 E-mail: matthias.kaltenbrunner@improvem.at
  • Gronalt, University of Natural Resources and Life Sciences, Department of Economics and Social Sciences, Institute of Production and Logistics, Feistmantelstrasse 4, 1180 Vienna, Austria ORCID https://orcid.org/0000-0003-0944-4911 E-mail: manfred.gronalt@boku.ac.at
article id 22017, category Research article
Andis Lazdiņš, Ainārs Lupiķis, Kaspars Polmanis, Arta Bārdule, Aldis Butlers, Santa Kalēja. (2024). Carbon stock changes of drained nutrient-rich organic forest soils in Latvia. Silva Fennica vol. 58 no. 1 article id 22017. https://doi.org/10.14214/sf.22017
Keywords: drainage; hemiboreal forests; organic soil; peat compaction; peatland forests; subsidence of the peat layer
Highlights: In moderate nutrient-rich forest site type (Myrtillosa turf. mel.), a significant subsidence of peat layer after drainage is associated with compaction rather than decomposition of peat; In nutrient-rich forest site type (Oxalidosa turf. mel.), a contribution of soil C stock losses to subsidence of the peat layer is significant; In moderate nutrient-rich forest site type (Myrtillosa turf. mel.), type of dominant tree species has higher impact on changes in soil C stock after drainage than in nutrient-rich forest site type (Oxalidosa turf. mel.); Distribution of different forest site types involving soil nutrient status has to be taken into account when CO2 emissions from drained organic soil in forest land are estimated at national level.
Abstract | Full text in HTML | Full text in PDF | Author Info

Impact of drainage of organic soils in forest land on soil carbon (C) stock changes is of high interest not only to accurately estimate soil C stock changes, but also to provide scientifically based recommendations for forest land management in context of climate change mitigation. To improve knowledge about long-term impact of drainage on nutrient-rich organic soils in hemiboreal forests in Latvia, 50 research sites representing drained conditions (Oxalidosa turf. mel. (Kp) and Myrtillosa turf. mel. (Ks) forest site types) and undrained conditions as control areas (Caricoso-phragmitosa, Dryopterioso-caricosa and Filipendulosa forest site types) were selected. Soil C stock changes after drainage was evaluated by comparing current C stock in drained organic soils to theoretical C stock before drainage considering impact of soil subsidence. During the 53-years period after drainage, the peat subsidence was higher in nutrient-rich Kp forest site type compared to moderate nutrient-rich Ks forest site type (peat subsided by 37.0 ± 4.8 and 23.3 ± 4.8 cm, respectively). In nutrient-rich Kp forest site type, soil C stock decreased by 4.98 ± 1.58 Mg C ha-1 yr-1 after drainage, while no statistically significant changes in soil C stock (0.19 ± 1.31 Mg C ha-1 yr-1) were observed in moderate nutrient-rich soils in Ks forest site type. Thus, in Ks forest site type, the main driver of the peat subsidence was the physical compaction, while in Kp forest site type contribution of organic matter decomposition and consequent soil C losses to subsidence of the peat was significant.

  • Lazdiņš, Latvian State Forest Research Institute ‘Silava’ (LSFRI Silava), Rigas str. 111, Salaspils, LV-2169, Latvia ORCID https://orcid.org/0000-0002-7169-2011 E-mail: andis.lazdins@silava.lv
  • Lupiķis, Latvian State Forest Research Institute ‘Silava’ (LSFRI Silava), Rigas str. 111, Salaspils, LV-2169, Latvia E-mail: ainars.lupikis@inbox.lv
  • Polmanis, Latvian State Forest Research Institute ‘Silava’ (LSFRI Silava), Rigas str. 111, Salaspils, LV-2169, Latvia ORCID https://orcid.org/0000-0003-2579-353X E-mail: kaspars.polmanis@silava.lv (email)
  • Bārdule, Latvian State Forest Research Institute ‘Silava’ (LSFRI Silava), Rigas str. 111, Salaspils, LV-2169, Latvia ORCID https://orcid.org/0000-0003-0961-5119 E-mail: arta.bardule@silava.lv
  • Butlers, Latvian State Forest Research Institute ‘Silava’ (LSFRI Silava), Rigas str. 111, Salaspils, LV-2169, Latvia ORCID https://orcid.org/0000-0003-3118-1716 E-mail: aldis.butlers@silava.lv
  • Kalēja, Latvian State Forest Research Institute ‘Silava’ (LSFRI Silava), Rigas str. 111, Salaspils, LV-2169, Latvia E-mail: santa.kaleja@silava.lv
article id 23054, category Research article
Stephan Böhm, Peter Baier, Thomas Kirisits, Christian Kanzian. (2023). Blue-stain development on Norway spruce logs under alpine conditions. Silva Fennica vol. 57 no. 3 article id 23054. https://doi.org/10.14214/sf.23054
Keywords: Picea abies; moisture content; temperature; weather data; bark beetle; predictive model; sap-stain
Highlights: A nonlinear model was developed to predict the temperature-dependent spread rate of blue-stain in Norway spruce logs in alpine areas in Austria; The influence of temperature sum on the development of blue-stain was confirmed; The effect of harvesting season on the development and amount of sap-stain (faster and more extensive in summer than in spring) was observed.
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Discoloration of the sapwood caused by blue-stain fungi on conifer logs during interim storage causes significant loss to the forest industry. The fungal infection is often associated with bark beetle attacks because the spores are transmitted by the beetles. They can also be disseminated by rain-splash and moist air. While there are methods to protect logs from sap-stain in wood yards, this is often not possible in the forest for practical and regulatory reasons. Timing of harvesting and timely transportation are often the only ways to prevent blue-stain. To estimate the urgency of transportation, knowledge of the growth of blue-stain fungi and its dependence on weather conditions is of great interest.   The proportion of discolored sapwood on Norway spruce logs was recorded along a time series, together with weather data in two field experiments conducted in spring and summer at two alpine sites in Austria. A predictive model was developed to estimate the proportion of blue-stained sapwood based on the temperature sum to which the logs were exposed. After harvest in March, there was a time lag of 82 and 97 days at the two respective sites, caused by initially low temperatures, before discoloration started. In contrast, sap-stain occurred 14 days after the harvest in June, when warm conditions prevailed from the start. The nonlinear least square regression model can help to estimate a window of opportunity to transport wood before it loses its value and serves as a sub model for lead time estimation within logistic decision support systems.

  • Böhm, Department of Forest and Soil Sciences, Institute of Forest Engineering, University of Natural Resources and Life Sciences, Vienna (BOKU), Peter-Jordan-Strasse 82, 1190 Vienna, Austria ORCID https://orcid.org/0000-0001-7803-6618 E-mail: stephan.boehm@boku.ac.at (email)
  • Baier, Department of Forest and Soil Sciences, Institute of Forest Entomology, Forest Pathology and Forest Protection, University of Natural Resources and Life Sciences, Vienna (BOKU), Peter-Jordan-Strasse 82, 1190 Vienna, Austria ORCID https://orcid.org/0000-0002-1029-5637 E-mail: peter.baier@boku.ac.at
  • Kirisits, Department of Forest and Soil Sciences, Institute of Forest Entomology, Forest Pathology and Forest Protection, University of Natural Resources and Life Sciences, Vienna (BOKU), Peter-Jordan-Strasse 82, 1190 Vienna, Austria ORCID https://orcid.org/0000-0002-9918-3593 E-mail: thomas.kirisits@boku.ac.at
  • Kanzian, Department of Forest and Soil Sciences, Institute of Forest Engineering, University of Natural Resources and Life Sciences, Vienna (BOKU), Peter-Jordan-Strasse 82, 1190 Vienna, Austria ORCID https://orcid.org/0000-0002-1198-9788 E-mail: christian.kanzian@boku.ac.at
article id 23040, category Research article
Jānis Liepiņš, Ieva Jaunslaviete, Kaspars Liepiņš, Līga Jansone, Roberts Matisons, Andis Lazdiņš, Āris Jansons. (2023). Effect of stem rot on wood basic density, carbon, and nitrogen content of living deciduous trees in hemiboreal forests. Silva Fennica vol. 57 no. 3 article id 23040. https://doi.org/10.14214/sf.23040
Keywords: wood specific gravity; birch; climate change mitigation; biomass estimation; alder; aspen; wood decay
Highlights: Stem rot significantly reduces the basic density of wood and increases its nitrogen content in living deciduous trees, while the carbon content appears irresponsive; The effect of the distance from the pith on the basic density and nitrogen content of wood varies, depending on presence of discoloration or decomposition in the wood.
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While numerous studies have focused on analyzing various aspects of the carbon (C) budget in forests, there appears to be a lack of comprehensive assessments specifically addressing the impact of stem rot on the C budget of broadleaf tree species, especially in old-growth forests where stem rot is prevalent. One of the main challenges in accurately quantifying C losses caused by stem rot is the lack of precise data on the basic density and C content of decayed wood, which are crucial for converting decayed wood volume into biomass and C stocks. Using linear mixed-effects models, we examine the variability of wood basic density, C content, and nitrogen (N) content. Discolored and decomposed wood was collected from the stems of 136 living deciduous trees common in hemiboreal forests in Latvia. Our research indicates a noticeable reduction in the wood basic density, coupled with an increase in the N content within the stem wood throughout the decomposition process in birch (Betula spp.), European aspen (Populus tremula L.), grey alder (Alnus incana (L.) Moench), and common alder (Alnus glutinosa (L.) Gaertn.). While aspen wood showed a decreasing trend in C content as decay progressed, a pairwise comparison test revealed no significant differences in C content between discolored and decomposed wood for the studied species, unlike the findings for basic density and N content. This study emphasizes the need to account for stem rot in old-growth forest carbon budgets, especially in broadleaf species, and calls for more research on stem rot-induced carbon losses.

  • Liepiņš, Latvian State Forest Research Institute “Silava,” Rigas Street 111, LV-2169 Salaspils, Latvia ORCID https://orcid.org/0000-0003-3030-1122 E-mail: janis.liepins@silava.lv (email)
  • Jaunslaviete, Latvian State Forest Research Institute “Silava,” Rigas Street 111, LV-2169 Salaspils, Latvia ORCID https://orcid.org/0009-0000-7322-2729 E-mail: ieva.jaunslaviete@silava.lv
  • Liepiņš, Latvian State Forest Research Institute “Silava,” Rigas Street 111, LV-2169 Salaspils, Latvia ORCID https://orcid.org/0000-0002-1179-8586 E-mail: kaspars.liepins@silava.lv
  • Jansone, Latvian State Forest Research Institute “Silava,” Rigas Street 111, LV-2169 Salaspils, Latvia ORCID https://orcid.org/0000-0003-2748-3797 E-mail: liga.jansone@silava.lv
  • Matisons, Latvian State Forest Research Institute “Silava,” Rigas Street 111, LV-2169 Salaspils, Latvia E-mail: roberts.matisons@silava.lv
  • Lazdiņš, Latvian State Forest Research Institute “Silava,” Rigas Street 111, LV-2169 Salaspils, Latvia ORCID https://orcid.org/0000-0002-7169-2011 E-mail: andis.lazdins@silava.lv
  • Jansons, Latvian State Forest Research Institute “Silava,” Rigas Street 111, LV-2169 Salaspils, Latvia ORCID https://orcid.org/0000-0001-7981-4346 E-mail: aris.jansons@silava.lv
article id 23024, category Research article
Anu Laakkonen, Katri Rusanen, Teppo Hujala, Mika Gabrielsson, Jouni Pykäläinen. (2023). Implications of the sustainability transition on the industry value creation logic – case of Finnish pulp and paper industry. Silva Fennica vol. 57 no. 3 article id 23024. https://doi.org/10.14214/sf.23024
Keywords: forest industry; business transformation; circular bioeconomy; sustainable business
Highlights: Global change drivers are inducing a sustainability transition to a circular bioeconomy; Established industries need to reconfigure their value creation logic; A document analysis investigated pulp and paper industry’s communicated value creation logic; An interdisciplinary approach helps to understand a changing business environment; Recognition of a socio-economic-technological-environmental system is needed.
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Global economic, social and environmental change drivers have tremendous effects on the dynamic and nested business environment calling for a sustainability transition to a circular bioeconomy. The transition will pressurise established industries to alter their value creation logic to consider sustainability holistically. The study follows a case study research strategy and investigates how an established Finnish pulp and paper industry reconfigures its communicated value creation logic. The findings of a qualitative document analysis suggest that the pulp and paper industry has started to explore new sustainable path-breaking innovations and create a common development agenda, which has resulted in incremental adaptations in the value creation logic. However, the industry’s narrative of already being sustainable has hampered the reconfiguration and stabilisation of the adapted value creation logic. From a theoretical perspective, adopting an interdisciplinary and systemic perspective is necessary to understand the changing business environment. From a managerial perspective, cross-sectoral collaboration and including perspectives of different actors can help in creating a holistically sustainable value creation logic.

  • Laakkonen, School of Forest Sciences, Faculty of Science, Forestry and Technology, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland; Department of Forest Sciences, Faculty of Agriculture and Forestry, University of Helsinki, P.O. Box 27, 00014 University of Helsinki, Finland ORCID https://orcid.org/0000-0002-6384-7773 E-mail: anu.laakkonen@uef.fi (email)
  • Rusanen, School of Forest Sciences, Faculty of Science, Forestry and Technology, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0003-1705-5561 E-mail: katri.rusanen@uef.fi
  • Hujala, School of Forest Sciences, Faculty of Science, Forestry and Technology, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0002-7905-7602 E-mail: teppo.hujala@uef.fi
  • Gabrielsson, Department of Marketing, Hanken School of Economics, P.O. Box 479, FI-00101, Helsinki, Finland; UEF Business School, Faculty of Social Sciences and Business Studies, University of Eastern Finland, P.O. Box 1627, FI-70211 Kuopio, Finland ORCID https://orcid.org/0000-0002-0633-6918 E-mail: mika.gabrielsson@uef.fi
  • Pykäläinen, School of Forest Sciences, Faculty of Science, Forestry and Technology, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: jouni.pykalainen@uef.fi
article id 23023, category Research article
Lennart Noordermeer, Hans Ole Ørka, Terje Gobakken. (2023). Imputing stem frequency distributions using harvester and airborne laser scanner data: a comparison of inventory approaches. Silva Fennica vol. 57 no. 3 article id 23023. https://doi.org/10.14214/sf.23023
Keywords: forest inventory; airborne laser scanning; harvester data; inventory approaches
Highlights: We imputed stem frequency distributions using harvester reference data and predictor variables computed from airborne laser scanner data.; Stand-level distributions of stem diameter, tree height, volume, and sawn wood volume; (Enhanced) area-based and semi-individual tree crown approaches outperformed the individual tree crown method.
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Stem frequency distributions provide useful information for pre-harvest planning. We compared four inventory approaches for imputing stem frequency distributions using harvester data as reference data and predictor variables computed from airborne laser scanner (ALS) data. We imputed distributions and stand mean values of stem diameter, tree height, volume, and sawn wood volume using the k-nearest neighbor technique. We compared the inventory approaches: (1) individual tree crown (ITC), semi-ITC, area-based (ABA) and enhanced ABA (EABA). We assessed the accuracies of imputed distributions using a variant of the Reynold’s error index, obtaining the best mean accuracies of 0.13, 0.13, 0.10 and 0.10 for distributions of stem diameter, tree height, volume and sawn wood volume, respectively. Accuracies obtained using the semi-ITC, ABA and EABA inventory approaches were significantly better than accuracies obtained using the ITC approach. The forest attribute, inventory approach, stand size and the laser pulse density had significant effects on the accuracies of imputed frequency distributions, however the ALS delay and percentage of deciduous trees did not. This study highlights the utility of harvester and ALS data for imputing stem frequency distributions in pre-harvest inventories.

  • Noordermeer, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway ORCID https://orcid.org/0000-0002-8840-0345 E-mail: lennart.noordermeer@nmbu.no (email)
  • Ørka, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway ORCID https://orcid.org/0000-0002-7492-8608 E-mail: hans-ole.orka@nmbu.no
  • Gobakken, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: terje.gobakken@nmbu.no
article id 23021, category Research article
Virpi Stenman, Annika Kangas, Markus Holopainen. (2023). Upper stem diameter and volume prediction strategies in the National Forest Inventory of Finland. Silva Fennica vol. 57 no. 3 article id 23021. https://doi.org/10.14214/sf.23021
Keywords: forest inventory; measurement errors; accuracy; Bland-Altman plot
Highlights: National Forest Inventory specific methods were applied with a number of measurement instruments, including a laser-based dendrometer, to collect tree stem diameter measurements; Bland-Altman plots and measurement error variances were used to determine measurement precision and accuracy; The laser-based dendrometer did not perform better than the other instruments in the study.
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In forest inventories, field data are needed for the prediction of tree volumes. However, gathering field data requires resources, such as labour, equipment, and data management operations. This means that time and budget, as well as quality, must be carefully considered when National Forest Inventory (NFI) field measurement activities are planned. Therefore, the development of cost efficient, simple, safe and reliable measurement methods and tools are of great interest. To date, upper stem diameter (d6), which provides a more reliable estimation of tree stem volume, has typically been measured with a parabolic calliper. In this study, the performance of the Criterion laser-based dendrometer was examined for d6 measurements. A total of 326 sample trees were measured multiple times with three different measurement instruments. These instruments were used to measure diameter at breast height (dbh) as well as d6 measurements. Bland-Altman plots and measurement error variances were used to determine measurement instrument reliability. For all trees, the standard deviation for the laser based dendrometer was 18.73 mm at dbh and 15.36 mm for the d6 measurements. When the performance of Criterion was analysed with reference to the mean value of repeated measurements, the standard deviation in the dbh measurements was 12.21 mm, and 8.88 mm in the d6 measurements.

  • Stenman, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID https://orcid.org/0000-0003-1176-7840 E-mail: virpi.stenman@helsinki.fi (email)
  • Kangas, Natural Resources Institute Finland (Luke), Bio­economy and Environment, P.O. Box 68, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0002-8637-5668 E-mail: annika.kangas@luke.fi
  • Holopainen, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: markus.holopainen@helsinki.fi
article id 23009, category Research article
Abubakari H. Munna, Nyambilila A. Amuri, Proches Hieronimo, Dino A. Woiso. (2023). Modelling ecological niches of Sclerocarya birrea subspecies in Tanzania under the current and future climates. Silva Fennica vol. 57 no. 3 article id 23009. https://doi.org/10.14214/sf.23009
Keywords: climate change; conservation; GIS; agroforestry; domestication; MaxEnt; protected areas network
Highlights: Tanzania harbors ecological niches of Sclerocarya birrea (S. birrea) subsp. caffra, multifoliata and birrea in the eastern, southern-central-northern, and northeastern part of the country, covering 184 814 km2, 139 918 km2 and 28 446 km2 of Tanzania’s land area, respectively; Ecological niches will contract under future warming climates; Currently, significant parts of ecological niches for Sclerocarya birrea subspecies are beyond Tanzania’s protected areas network.
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The information on ecological niches of the Marula tree, Sclerocarya birrea (A. Rich.) Horchst. subspecies are needed for sustainable management of this tree, considering its nutritional, economic, and ecological benefits. However, despite Tanzania being regarded as a global genetic center of diversity of S. birrea, information on the subspecies ecological niches is lacking. We aimed to model ecological niches of S. birrea subspecies in Tanzania under the current and future climates. Ecological niches under the current climate were modelled by using ecological niche models in MaxEnt using climatic, edaphic, and topographical variables, and subspecies occurrence data. The Hadley Climate Center and National Center for Atmospheric Research's Earth System Models were used to predict ecological niches under the medium and high greenhouse gases emission scenarios for the years 2050 and 2080. Area under the curves (AUCs) were used to assess the accuracy of the models. The results show that the models were robust, with AUCs of 0.85–0.95. Annual and seasonal precipitation, elevation, and soil cation exchange capacity are the key environmental factors that define the ecological niches of the S. birrea subspecies. Ecological niches of subsp. caffra, multifoliata, and birrea are currently found in 30, 22, and 21 regions, and occupy 184 814 km2, 139 918 km2, and 28 446 km2 of Tanzania's land area respectively, which will contract by 0.4–44% due to climate change. Currently, 31–51% of ecological niches are under Tanzania’s protected areas network. The findings are important in guiding the development of conservation and domestication strategies for the S. birrea subspecies in Tanzania.

  • Munna, Department of Soil and Geological Sciences, Sokoine University of Agriculture, P.O. Box 3008, Morogoro, Tanzania ORCID https://orcid.org/0000-0001-8858-0457 E-mail: amabmunna81@gmail.com (email)
  • Amuri, Department of Soil and Geological Sciences, Sokoine University of Agriculture, P.O. Box 3008, Morogoro, Tanzania ORCID https://orcid.org/0000-0003-3092-3458 E-mail: namuri@sua.ac.tz
  • Hieronimo, Department of Agricultural Engineering, Sokoine University of Agriculture, P.O. Box 3003 Morogoro, Tanzania ORCID https://orcid.org/0000-0002-4450-5073 E-mail: phmusigula@gmail.com
  • Woiso, Department of Biosciences, Sokoine University of Agriculture, P.O. Box 3038 Morogoro, Tanzania E-mail: dino@sua.ac.tz
article id 23004, category Research article
Per Nordin, Erika Olofsson, Karin Hjelm. (2023). Within-site adaptation: growth and mortality of Norway spruce, Scots pine and silver birch seedlings in different planting positions across a soil moisture gradient. Silva Fennica vol. 57 no. 3 article id 23004. https://doi.org/10.14214/sf.23004
Keywords: regeneration; microsite; decision making; mounding; planting; depth-to-water
Highlights: A soil moisture map could be used to support the choice of planting position for different soil moisture conditions; Mounds reduced mortality rates for conifers when conditions were wet, but at drier conditions differences between planting positions were small; Contradictory, silver birch had higher survival in lower planting positions compared with mounds; Height and diameter were higher in mounds for conifers, but only small differences occurred between planting positions for silver birch.
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Adapting to site conditions is a central part of forest regeneration and can be done through selection of different planting positions. Requirements are tree species specific, and the use of soil moisture maps could be a way to support decision making in forest regeneration planning. At two experimental sites with varying soil moisture conditions in southern Sweden Norway spruce (Picea abies (L.) Karst.), Scots pine (Pinus sylvestris L.), and silver birch (Betula pendula Roth) seedlings were planted in four different planting positions following mounding site preparation; Depression, Hinge, Mound and Unscarified. Soil moisture estimates were obtained from a high-resolution depth-to-water raster for each planting spot. The effect of soil moisture, planting position and their interactions on mortality, height and diameter was evaluated for each tree species. In wet conditions mounds proved to be the best option to minimize seedling mortality for conifers, but with decreasing soil moisture, differences between the planting positions decreased. Birch on the other hand had the greatest survival in the hinge. The coniferous species displayed increased height and diameter when planted in mounds independent of the soil moisture conditions, whereas silver birch was less dependent on a specific planting position. Results from this study shows that a soil moisture map can explain mortality, height and diameter and thus can be a useful tool when choosing planting position in different soil moisture conditions.

article id 23003, category Research article
Olli-Pekka Tikkanen, Ilari Lehtonen. (2023). Changing climatic drivers of European spruce bark beetle outbreaks: a comparison of locations around the Northern Baltic Sea. Silva Fennica vol. 57 no. 3 article id 23003. https://doi.org/10.14214/sf.23003
Keywords: climate change; temperature sum; precipitation; Ips typographus; hydrothermic index
Highlights: Temperatures in the Northern Baltic Sea area will reach new and higher levels in all studied climate change scenarios, speeding up the development of the European spruce bark beetle; Unless greenhouse gas emissions are reduced rapidly, more frequent droughts will facilitate spruce bark beetle outbreaks, especially in Sweden and, to a lesser extent, in Estonia and in Southern Finland.
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European spruce bark beetle (Ips typographus [L.]; SBB) damage has reached extreme and unprecedented levels in East Central Sweden, likely driven by increasing temperatures and severe drought due to climate change. However, SBB outbreaks have been less severe on the eastern side of the Baltic Sea, in Estonia and Finland, than in Sweden. This study investigated how precipitation, temperature sum, and droughts (hydrothermic index) have varied in land areas surrounding the Baltic Sea. We studied past meteorological observations from 1950 to 1999. We modeled the effect of climate change on precipitation and temperature using three representative concentration pathway (RCP) scenarios for greenhouse gas emissions (RCP2.6, RCP4.5, and RCP8.5) and multiple (17–23) climate models. Future climate projections (up to 2100) were made for Southeastern Estonia, Southern Finland, and East Central Sweden. Weather data showed that temperature sums had been high and droughts severe in the 2010s, particularly in East Central Sweden, where SBB outbreaks have been a more significant problem than on the eastern shores of the Baltic Sea. Future climate projections suggest that increases in temperature sum will further enhance SBB reproduction, especially in the RCP4.5 and RCP8.5 scenarios. In all climate change scenarios, drought continues to be a problem in East Central Sweden, potentially facilitating SBB outbreaks. In addition, moderate and severe droughts may become more frequent in Southeastern Estonia and Southern Finland if climate change proceeds as predicted in the RCP4.5 or RCP8.5 scenarios.

article id 23019, category Research article
Victor F. Strîmbu, Tron Eid, Terje Gobakken. (2023). A stand level scenario model for the Norwegian forestry – a case study on forest management under climate change. Silva Fennica vol. 57 no. 2 article id 23019. https://doi.org/10.14214/sf.23019
Keywords: forest planning; carbon balance; climate change mitigation; forest stand simulator
Highlights: GAYA 2.0: a new scenario analysis model focusing on forest carbon fluxes; Carbon sequestration potential estimated at regional level; GAYA 2.0 may be used to estimate the costs of obtaining carbon benefits by adapting the forest management.
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Carbon sequestration and income generation are competing objectives in modern forest management. The climate commitments of many countries depend on forests as carbon sinks which must be quantified, monitored, and projected into the future. For projections we need tools to model forest development and perform scenario analyses to assess future carbon sequestration potentials under different management regimes, the expected net present value of such regimes, and possible impacts of climate change. We propose a scenario analysis software tool (GAYA 2.0) that can assist in answering these types of questions using stand level simulations, detailed carbon flow models and an optimizer. This paper has two objectives: (1) to describe GAYA 2.0, and (2) demonstrate its potential in a case study where we analyze the forest carbon balance over a region in Norway based on national forest inventory sample plots. The tool was used to map the optimality front between the carbon benefit and net present value. We observed changes in net present value for different levels of carbon benefit as well as changes in optimal management strategies. We predicted future changes in several forest carbon pools as well as albedo and illustrated the impact of gradual increase in forest productivity (i.e., due to climate warming). Having been updated and modernized from its previous version with increased attention to forest carbon and energy fluxes, GAYA 2.0 is an effective tool that offers multiple opportunities to perform various types of scenario analyses in forest management.

  • Strîmbu, Norwegian University of Life Sciences, Faculty of Environmental Sciences and Natural Resource Management, NO-1432 Ås, Norway ORCID https://orcid.org/0000-0002-0588-2036 E-mail: victor.strimbu@nmbu.no (email)
  • Eid, Norwegian University of Life Sciences, Faculty of Environmental Sciences and Natural Resource Management, NO-1432 Ås, Norway E-mail: tron.eid@nmbu.no
  • Gobakken, Norwegian University of Life Sciences, Faculty of Environmental Sciences and Natural Resource Management, NO-1432 Ås, Norway ORCID https://orcid.org/0000-0001-5534-049X E-mail: terje.gobakken@nmbu.no
article id 23012, category Research article
Jari Vauhkonen, Juho Matala, Ari Nikula. (2023). Future browsing damage in seedling stands according to projected forest resources and moose population density. Silva Fennica vol. 57 no. 2 article id 23012. https://doi.org/10.14214/sf.23012
Keywords: forest management planning; forest development simulation; forest projection; Markov chain; scenario analysis; transition matrix model
Highlights: Projections of forest resources and seedling stands damaged by moose browsing; Damaged seedling stand area modelled by moose population and forest characteristics; Moose damage predicted by the age class structure of simulated future forests.
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An important modifier of forests and forestry practices is browsing by cervids. As high populations of moose (Alces alces L.) cause extensive forest damage in the Fennoscandian boreal forests, models should be able to predict the susceptibility of projected forest structures to browse damage. We augmented the European Forestry Dynamics Model (EFDM) for the area of seedling stands damaged by moose. The augmented model was tested in projecting both forest resources and moose damage for 18 million hectares of forest land in Finland, based on input data from the National Forest Inventory (NFI). Modeling the area of seedling stands damaged as a function of moose population density, forest characteristics, and region-specific interactions of these variables was found to work realistically for 30 years, predicting that the area of seedling stands damaged by moose would increase by up to a third from the last NFI observation. Our work lays the groundwork for modeling consequential, large-scale ecological and socio-economic effects of moose browsing.

  • Vauhkonen, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, FI-80101 Joensuu, Finland; University of Helsinki, Department of Forest Sciences, Latokartanonkaari 7, FI-00014 Helsingin yliopisto, Finland E-mail: jari.vauhkonen@uef.fi (email)
  • Matala, Natural Resources Institute Finland (Luke), Natural resources, Yliopistokatu 6 B, FI-80100 Joensuu, Finland E-mail: juho.matala@luke.fi
  • Nikula, Natural Resources Institute Finland (Luke), Natural resources, Ounasjoentie 6, FI-96200 Rovaniemi, Finland ORCID https://orcid.org/0000-0001-8372-8440 E-mail: ari.nikula@luke.fi
article id 22028, category Research article
Eelis Halme, Matti Mõttus. (2023). Improved parametrisation of a physically-based forest reflectance model for retrieval of boreal forest structural properties. Silva Fennica vol. 57 no. 2 article id 22028. https://doi.org/10.14214/sf.22028
Keywords: forest structure; Sentinel-2; reflectance; hyperspectral; tree distribution
Highlights: Spatial distribution of trees is a key driver for forest reflectance; Knowledge of the ratio of branch to leaf area improves forest reflectance simulation substantially; Different optical properties of the two leaf sides have a notable effect on forest reflectance.
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Physically-based reflectance models offer a robust and transferable method to assess biophysical characteristics of vegetation in remote sensing. Forests exhibit explicit structure at many scales, from shoots and branches to landscape patches, and hence present a specific challenge to vegetation reflectance modellers. To relate forest reflectance with its structure, the complexity must be parametrised leading to an increase in the number of reflectance model inputs. The parametrisations link reflectance simulations to measurable forest variables, but at the same time rely on abstractions (e.g. a geometric surface forming a tree crown) and physically-based simplifications that are difficult to quantify robustly. As high-quality data on basic forest structure (e.g. tree height and stand density) and optical properties (e.g. leaf and forest floor reflectance) are becoming increasingly available, we used the well-validated forest reflectance and transmittance model FRT to investigate the effect of the values of the “uncertain” input parameters on the accuracy of modelled forest reflectance. With the state-of-the-art structural and spectral forest information, and Sentinel-2 Multispectral Instrument imagery, we identified that the input parameters influencing the most the modelled reflectance, given that the basic forestry variables are set to their true values and leaf mass is determined from reliable allometric models, are the regularity of the tree distribution and the amount of woody elements. When these parameters were set to their new adjusted values, the model performance improved considerably, reaching in the near infrared spectral region (740–950 nm) nearly zero bias, a relative RMSE of 13% and a correlation coefficient of 0.81. In the visible part of the spectrum, the model performance was not as consistent indicating room for improvement.

article id 22027, category Research article
Timo Saksa, Jari Miina, Olli Luukkonen. (2023). Uprooting as a pre-commercial thinning operation in young Norway spruce stands. Silva Fennica vol. 57 no. 2 article id 22027. https://doi.org/10.14214/sf.22027
Keywords: Picea abies; simulation; precommercial thinning; mechanical young stand management
Highlights: The number of competing birches varied considerably according to the operating conditions; The main factors reducing the quality of uprooting were early operation timing, soil moisture and site fertility; With careful selection of the timing and conditions for uprooting, the quality obtained by uprooting can be improved.
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Uprooting, as a mechanical pre-commercial thinning operation (PCT), removes competitive, undesired trees with roots to prevent or reduce the need for a second PCT. The state of 66 young Norway spruce (Picea abies (L.) H. Karst.) stands was explored 3–5 years after uprooting to assess the quality of uprooting in terms of the number of competing birches (Betula spp.) and the probability of freely growing crop tree spruce (no competing trees taller than 2/3 of the crop tree’s height). In the uprooted spruce stands, the number of birches was, on average, about 3800 stems ha−1, and in every fifth stand, it was less than 2000 stems ha−1. The number of birches increased with increasing site wetness (TWI), site fertility and thickness of the humus layer. In 80% of the stands, there were at least 1000 freely growing spruces ha−1, which is approximately the stem number to be left growing after first thinning in a spruce stand. If the height of crop tree spruces was 125 cm at uprooting, the probability of freely growing stems varied from 0.6 to 0.8 in typical cases. According to simulations at the first thinning stage, in most stands, birch was less than 10% taller than spruce, and the volume of birches was low enough that, in the future, spruce and birch would grow in the same crown layer as a mixed spruce–birch stand. The results showed that uprooting can serve as the only PCT operation in a planted spruce stand. The quality obtained by uprooting can be improved with careful selection of timing and conditions for uprooting; the optimal uprooting time is when crop tree spruces have reached about breast height.

  • Saksa, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, 77600 Suonenjoki, Finland ORCID https://orcid.org/0000-0002-1776-2357 E-mail: ext.timo.saksa@luke.fi (email)
  • Miina, Natural Resources Institute Finland (Luke), Natural resources, Yliopistokatu 6 B, 80100 Joensuu, Finland ORCID https://orcid.org/0000-0002-8639-4383 E-mail: jari.miina@luke.fi
  • Luukkonen, Metsäsydän Oy, Junninmäentie 53, 52200 Puumala, Finland E-mail: olli.p.j.luukkonen@gmail.com
article id 22026, category Research article
Annika Kangas, Mari Myllymäki, Lauri Mehtätalo. (2023). Understanding uncertainty in forest resources maps. Silva Fennica vol. 57 no. 2 article id 22026. https://doi.org/10.14214/sf.22026
Keywords: autocorrelation; ensemble modelling; kriging; quantile; random forest; sequential Gaussian simulation
Highlights: Forest resources maps without uncertainty assessment may lead to false impression of precision; Suitable tools for visualization of map products are lacking; Kriging method provided accurate uncertainty assessment for pixel-level predictions; Quantile random forest algorithm slightly underestimated the pixel-level uncertainties; With simulation it is possible to assess the uncertainty also for landscape-level characteristics.
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Maps of forest resources and other ecosystem services are needed for decision making at different levels. However, such maps are typically presented without addressing the uncertainties. Thus, the users of the maps have vague or no understanding of the uncertainties and can easily make wrong conclusions. Attempts to visualize the uncertainties are also rare, even though the visualization would be highly likely to improve understanding. One complication is that it has been difficult to address the predictions and their uncertainties simultaneously. In this article, the methods for addressing the map uncertainty and visualize them are first reviewed. Then, the methods are tested using laser scanning data with simulated response variable values to illustrate their possibilities. Analytical kriging approach captured the uncertainty of predictions at pixel level in our test case, where the estimated models had similar log-linear shape than the true model. Ensemble modelling with random forest led to slight underestimation of the uncertainties. Simulation is needed when uncertainty estimates are required for landscape level features more complicated than small areas.

  • Kangas, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Yliopistokatu 6, 80101 Joensuu, Finland ORCID https://orcid.org/0000-0002-8637-5668 E-mail: annika.kangas@luke.fi (email)
  • Myllymäki, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0002-2713-7088 E-mail: mari.myllymaki@luke.fi
  • Mehtätalo, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Yliopistokatu 6, 80101 Joensuu, Finland ORCID https://orcid.org/0000-0002-8128-0598 E-mail: lauri.mehtatalo@luke.fi
article id 22013, category Research article
Bodil Häggström, Reimo Lutter, Tomas Lundmark, Fredrik Sjödin, Annika Nordin. (2023). Effect of arginine-phosphate addition on early survival and growth of Scots pine, Norway spruce and silver birch. Silva Fennica vol. 57 no. 2 article id 22013. https://doi.org/10.14214/sf.22013
Keywords: Pinus sylvestris; Betula pendula; Picea abies; forest regeneration; seedling growth; seedling survival; arginine
Highlights: Arginine-phosphate addition (APA) represents a potential tool to aid regeneration of planted trees, especially to increase survival of Scots pine seedlings on sites where susceptible to pests;Effects of APA however varies between different sites.
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Applying arginine-phosphate (AP) to tree seedlings at planting is a novel silvicultural practice in Northern Europe to improve the success of forest regeneration. We present three case-studies of the potential advantages of adding AP at planting on the establishment and damage susceptibility of seedlings in pure and mixed plantings of Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies (L.) H. Karst. ) and silver birch (Betula pendula Roth) over two years in the field. Location of study sites were in southern (S), northeastern (NE) and northwestern (NW) Sweden. The main agents of damage were pine weevil (Hylobius abietis L.) on conifers at the south site, browsing of birch at all sites and browsing/other top damage to conifers at the north sites. The effect of adding AP varied between the sites. It was positive for survival of pine at site S, despite considerable damage by pine weevil. However, at the S site more of the surviving spruce and birch were browsed when treated with AP. At the NE site AP-treatment had positive effects on conifer growth. At the NW site adding AP positively affected survival and growth of all three species, and AP-treated seedlings of all species were less browsed than untreated seedlings. AP treatment presents a potential tool to improve the success of forest regeneration, especially when establishing pine stands in south Sweden.

  • Häggström, Umeå Plant Science Centre, Department of Forest Genetics and Plant Physiology, Swedish University of Agricultural Sciences, 90183 Umeå, Sweden ORCID https://orcid.org/0000-0002-7738-5493 E-mail: bodil.haggstrom@slu.se (email)
  • Lutter, Institute of Forestry and Engineering, Estonian University of Life Sciences, Kreutzwaldi 5, Tartu 51006, Estonia ORCID https://orcid.org/0000-0001-5847-4282 E-mail: reimo.lutter@emu.ee
  • Lundmark, Department of Forest Ecology and Management, Swedish University of Agricultural Sciences, 90183 Umeå, Sweden ORCID https://orcid.org/0000-0003-2271-3469 E-mail: tomas.lundmark@slu.se
  • Sjödin, Unit for field-based forest research, Swedish University of Agricultural Sciences, 90183 Umeå, Sweden E-mail: fredrik.sjodin@slu.se
  • Nordin, Umeå Plant Science Centre, Department of Forest Genetics and Plant Physiology, Swedish University of Agricultural Sciences, 90183 Umeå, Sweden ORCID https://orcid.org/0000-0002-5765-3550 E-mail: annika.nordin@slu.se
article id 10764, category Research article
Sabiha Acer, Zeynel Arslangündoğdu, Asko Lehtijärvi. (2023). The occurrence and pathogenicity of fungi associated with Orthotomicus erosus on Pinus brutia in the Southern Marmara, Türkiye. Silva Fennica vol. 57 no. 2 article id 10764. https://doi.org/10.14214/sf.10764
Keywords: Ophiostoma; pathogenicity; Graphilbum; Leptographium; Mediterranean pine beetle; Turkish pine; Southern Marmara
Highlights: Three ophiostomatoid fungi species are recorded, associated with Orthotomicus erosus on Pinus brutia for the first time in Türkiye; Ceratocystis ips has the highest frequency. The association between Leptographium wingfieldiiand Orthotomicus erosus occurred with high frequency; While all three fungal species are severely pathogenic for pines in some regions, they are weak pathogens for Turkish pine in Türkiye.
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Fungal pathogens associated with bark beetles constitute one of the most significant problems to forest health. The Turkish pine (Pinus brutia Ten.) is a native species in the forests of Türkiye and occurs in the Mediterranean-type climate. The Southern Marmara is a natural occurrence area of Turkish pine in the Marmara Region. In the present study, trap logs were set up in pure Pinus brutia forests to investigate fungi associated with Orthotomicus erosus (Wollaston) (Mediterranean pine beetle) throughout Southern Marmara. Orthotomicus erosus adults, larvae, and their galleries were sampled and individually cultured on a 1% CSMA (cycloheximide–streptomycin malt agar) medium. Three ophiostomatoid fungi were identified using morphological characteristics and molecular genetic analyses: Ceratocystis (syn. Ophiostoma) ips (Rumbold) C. Moreau, Graphilbum sp. H.P. Upadhyay & W.B. Kendr., and Leptographium wingfieldii M. Morelet. All three species were new in records of the fungal flora of Türkiye. The most dominant of these species, Ceratocystis ips was isolated 69%. Unexpectedly, L. wingfieldii had a high-frequency association with O. erosus (27%). The pathogenicity tests showed that all three species could cause lesions on branches of Turkish pine but were non-pathogenic or weak pathogenic.

  • Acer, Istanbul University-Cerrahpaşa, Department of Forest Entomology and Protection, Faculty of Forestry, 34473, Istanbul, Türkiye E-mail: sacer@iuc.edu.tr (email)
  • Arslangündoğdu, Istanbul University-Cerrahpaşa, Department of Forest Entomology and Protection, Faculty of Forestry, 34473, Istanbul, Türkiye E-mail: zeynel@iuc.edu.tr
  • Lehtijärvi, Isparta University of Applied Sciences, Sütçüler Prof. Dr. Hasan Gürbüz Vocational School, 32950, Isparta, Türkiye E-mail: askolehtijarvi@isparta.edu.tr
article id 23001, category Research article
Eirik Ogner Jåstad, Niels Oliver Nagel, Junhui Hu, Per Kristian Rørstad. (2023). The location and capacity-dependent price impacts of biofuel production and its effect on the forest industry. Silva Fennica vol. 57 no. 1 article id 23001. https://doi.org/10.14214/sf.23001
Keywords: partial equilibrium model; biofuel location policy; biomass supply; forest-based biofuel; nordic forest sector
Highlights: Large biofuel units prefer regions close to transportation facilities; Forest owners are the main winners if large-scale biofuel production is established; The first production units reduce export, hence should be located at an exporting hub; Biofuel production will reduce the Norwegian export of roundwood to Sweden; Biofuel production increases the local demand and pulpwood prices.
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Forest-based biofuel stands out as a promising solution to avoid fossil emissions in parts of the transport sector. Biofuel production will need large amounts of forest biomass, collected from a large area. Roundwood is costly to transport compared with other goods. Therefore, the location of forest-based biofuel production is a crucial part of an investment decision. This study analyses the optimal location of biofuel plants in Norway and the implications for the traditional forest sector in the Nordic countries. We test different numbers of production units, different sizes of the units, and various raw materials. The study applies a partial equilibrium model that covers the Norwegian and Nordic forest sectors, with 356 regions in Norway. The results indicate that small biofuel plants have the potential to turn exporting regions into importing regions. Larger biofuel plants are suitable for areas with large harvest activity today or regions with access to harbour or timber terminals along railways. We find that forest owners close to a biofuel plant will profit the most from biofuel production. Policymakers and investors should take into account that different locations and production capacities have different impacts on the forest sectors.

  • Jåstad, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway ORCID https://orcid.org/0000-0002-1089-0284 E-mail: eirik.jastad@nmbu.no (email)
  • Nagel, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway ORCID https://orcid.org/0000-0002-3171-0262 E-mail: niels.oliver.nagel@nmbu.no
  • Hu, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway ORCID https://orcid.org/0000-0003-0001-5993 E-mail: junhui.hu@nmbu.no
  • Rørstad, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: per.kristian.rorstad@nmbu.no
article id 22019, category Research article
Aleksi Nirhamo, Juha Pykälä, Kimmo Jääskeläinen, Jari Kouki. (2023). Habitat associations of red-listed epiphytic lichens in Finland. Silva Fennica vol. 57 no. 1 article id 22019. https://doi.org/10.14214/sf.22019
Keywords: boreal forests; deciduous trees; biodiversity; conservation; threatened species
Highlights: We analyzed the habitat associations of 231 nationally red-listed epiphytic lichen species in Finland; Their habitat associations were varying, but deciduous trees, old forests and trees, and microclimates with intermediate or high light availability and humidity were particularly important; The maintenance of the habitats of many red-listed epiphytic lichens is difficult if not impossible to combine with intensive forest management.
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The Finnish red list shows that the epiphytic lichen flora of Finnish forests is highly threatened and declining steeply. Red lists provide limited information on the habitat associations of threatened species, which could be relevant in informing management and conservation measures. We used documented empirical data and expert assessments to determine for each red-listed (IUCN categories Near Threatened, NT; Vulnerable, VU; Endangered, EN; Critically Endangered, CR; Regionally Extinct, RE) epiphytic lichen species of Finland the following key habitat associations: host tree species, substrate type, habitat type, geographical distribution, preferred microclimate, and minimum required forest and tree age. The most important host tree species were Picea abies (L.) H. Karst. and Populus tremula L. Other tree species of high importance included Sorbus aucuparia L. and Salix caprea L. One fourth of red-listed epiphytic lichens were primarily lignicolous. Most species required old-growth forests (required by 41% of species) or old trees (52%), but many species required only mature forests (36%) or trees (35%). The microclimatic preferences of most red-listed epiphytic lichens consisted of high or intermediate light availability and humidity. Most species whose status had deteriorated were dependent on deciduous trees. The continuous availability of old deciduous trees (especially Populus, Salix and Sorbus) requires special attention in both managed and protected forests. Red-listed epiphytic lichens would be aided by increased forest protection or transitioning to less intensive management regimes.

article id 22014, category Research article
Nea Kuusinen, Aarne Hovi, Miina Rautiainen. (2023). Estimation of boreal forest floor lichen cover using hyperspectral airborne and field data. Silva Fennica vol. 57 no. 1 article id 22014. https://doi.org/10.14214/sf.22014
Keywords: remote sensing; Cladonia; spectroscopy
Highlights: A pilot study on estimating forest floor lichen cover from hyperspectral data; Multiple endmember spectral mixture analysis applied to field and airborne data; Accuracy of lichen cover estimates was good; Tree cover and presence of dwarf shrubs may influence lichen cover estimation.
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Lichens are sensitive to competition from vascular plants, intensive silviculture, pollution and reindeer and caribou grazing, and can therefore serve as indicators of environmental changes. Hyperspectral remote sensing data has been proved promising for estimation of plant diversity, but its potential for forest floor lichen cover estimation has not yet been studied. In this study, we investigated the use of hyperspectral data in estimating ground lichen cover in boreal forest stands in Finland. We acquired airborne and in situ hyperspectral data of lichen-covered forest plots, and applied multiple endmember spectral mixture analysis to estimate the fractional cover of ground lichens in these plots. Estimation of lichen cover based on in situ spectral data was very accurate (coefficient of determination (r2) 0.95, root mean square error (RMSE) 6.2). Estimation of lichen cover based on airborne data, on the other hand, was fairly good (r2 0.77, RMSE 11.7), but depended on the choice of spectral bands. When the hyperspectral data were resampled to the spectral resolution of Sentinel-2, slightly weaker results were obtained. Tree canopy cover near the flight plots was weakly related to the difference between estimated and measured lichen cover. The results also implied that the presence of dwarf shrubs could influence the lichen cover estimates.

  • Kuusinen, Department of Built Environment, School of Engineering, Aalto University, P.O. Box 14100, FI-00076 Aalto, Finland ORCID https://orcid.org/0000-0002-8063-1739 E-mail: nea.kuusinen@aalto.fi (email)
  • Hovi, Department of Built Environment, School of Engineering, Aalto University, P.O. Box 14100, FI-00076 Aalto, Finland ORCID https://orcid.org/0000-0002-4384-5279 E-mail: aarne.hovi@aalto.fi
  • Rautiainen, Department of Built Environment, School of Engineering, Aalto University, P.O. Box 14100, FI-00076 Aalto, Finland ORCID https://orcid.org/0000-0002-6568-3258 E-mail: miina.a.rautiainen@aalto.fi
article id 10767, category Research article
Semo Mogeia, Alberto A. Manhiça, Andrade F. Egas. (2023). Wood ash content variation in Eucalyptus grandis clones in Mozambique. Silva Fennica vol. 57 no. 1 article id 10767. https://doi.org/10.14214/sf.10767
Keywords: sapwood; heartwood; base-top; bioenergy quality; woody fuel
Highlights: Eucalyptus grandis heartwood produces better fuel than sapwood, if assuming ash content as energy quality parameter; Younger individuals have higher ash content in sapwood, and older individuals in the heartwood; There was not significant stem end variation of ash content in heartwood and sapwood.
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The sustainability of native forests in Sub-Saharan Africa depends on the diversification of sources to generate bioenergy, and Eucalyptus spp. wood has been highlighted. However, the determination of energy quality parameters has been a challenge to enable plantation wood to generate energy. The research assessed the ash content of radial and longitudinal samples of Eucalyptus grandis (Hill) clone with different ages and growth sites. Samples were collected in three pre-established plots in the center of Mozambique. Five trees were cut down in each plot and six discs were removed from each tree. Grinded samples with <0.5 mm particle size were generated from the heartwood and sapwood of each disk to determine the ash content. Wood from 7-year-olds had a higher ash content compared to 9-year-olds. The two sample plots differed from each other in terms of wood ash content. Heartwood samples had smaller ash content than sapwood samples. In general, the ash content of the intermediate positions was lower than those from the base and top of the stem, for both radial sections. No conclusive differences were found between samples from the base and the top of the trees, indicating that the material from the top of the trees can also be used as wood fuel. Ash content can be a considerable parameter to assess the quality of the wood of Eucalyptus spp. as a fuel.

  • Mogeia, Universidade Lúrio, Faculdade de Ciências Agrárias, Departamento de Silvicultura e Maneio [Lurio University, Faculty of Agricultural Sciences, Department of Forestry and Management], Campus de Wanaango, EN733, Km 42, Unango, Niassa, Mozambique E-mail: smogeia@unilurio.ac.mz (email)
  • Manhiça, Centro de Investigação Florestal, [Forestry Research Center], Marracuene, EN1, Maputo província, Mozambique E-mail: albertomanhica@gmail.com
  • Egas, Universidade Eduardo Mondlane, Faculdade de Agronomia e Engenharia Florestal, Departamento de Florestas, [Eduardo Mondlane University, Faculty of Agronomy and Forestry Engineering, Department of Forests], Av. Julius Nyerere, Maputo cidade, Mozambique E-mail: aegas8@gmail.com
article id 10752, category Research article
Oiva Hiltunen, Ville Hallikainen, Teijo Palander. (2023). Analysing the groundwater level and its determinants in a drained peatland forest: a case study in South Lapland, Finland. Silva Fennica vol. 57 no. 1 article id 10752. https://doi.org/10.14214/sf.10752
Keywords: peatland; groundwater level; load-bearing capacity; subsoil
Highlights: Mineral subsoils under a peat layer (less than 1 m) affect the groundwater level; During average precipitation, silty subsoil often makes groundwater level remain too high for timber haulage; If the subsoil is sandy and the peat layer is thin, the groundwater level gets lower; The amount of stand evapotranspiration alone is not enough to keep the groundwater level low enough.
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In southern Lapland, 70% of drained peatland forests have a peat layer thickness of less than one metre. On these sites, the question is how the subsoil under the peat affects groundwater level and thus timber harvesting. The aim of this study was to investigate the effect of the peat layer (<1 m) and subsoil on the groundwater level and its variation during the growing season (non-frost) by modelling the factors affecting water level. In sandy soils, the groundwater level rose by 20 cm when the peat layer thickness increased from 20 to 70 cm. In silty soils the effect of the peat thickness on groundwater remained minor. When the subsoil was sand or coarser, the groundwater level was usually deeper than when it was silty or finer. The effect of stand volume (m–3 ha–1) on the groundwater level was rather weak albeit significant. The model explained a significant part of the groundwater surface variation, with a marginal coefficient of determination (R2) of 68%. It seems that the rutting of roads could be avoided in late summer if the precipitation is remarkably lower during that period, or if the subsoil is sandy with thin peat layer on top of it. Because the groundwater level affects the load-bearing capacity of timber-harvesting machinery, it is important to study this issue in more detail in the future.

  • Hiltunen, Lapland University of Applied Sciences, Jokiväylä 11, FI-96300 Rovaniemi, Finland E-mail: oivah@student.uef.fi (email)
  • Hallikainen, Natural Resources Institute Finland, Ounasjoentie 6, FI-96300 Rovaniemi, Finland E-mail: ville.hallikainen@luke.fi
  • Palander, The University of Eastern Finland, Faculty of Science, Forestry and Technology, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0002-9284-5443 E-mail: teijo.s.palander@uef.fi
article id 10648, category Research article
Antti-Jussi Lindroos, Hannu Ilvesniemi. (2023). Weathering rates of Ca and Mg related to granitic and gabbro mineralogy in boreal forest soils and the effect of mechanical soil disturbance on weathering release. Silva Fennica vol. 57 no. 1 article id 10648. https://doi.org/10.14214/sf.10648
Keywords: ploughing; base cations; rotation period; mineral soil weathering
Highlights: Weathering rates were higher in the gabbro than the granitic areas; Weathering was an important Ca and Mg source to forest trees, and it increased after mechanical soil disturbance.
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This study’s aim was to calculate the weathering rates of Ca and Mg for five boreal forest soils in southern Finland on granitic and gabbro containing bedrock. The effect of mineralogy on the total concentrations of Ca and Mg in soil and weathering rates was evaluated. The aim was also to estimate the effect of mechanical soil disturbance related to ploughing on the weathering in the gabbro area. The total concentrations of SiO2, CaO, MgO, and Zr were determined by XRF, and weathering rates of Ca and Mg were determined based on the changes in the CaO, MgO, and Zr concentrations. The weathering rates of Ca+Mg varied 5–38 mmolc m–2 year–1 in the E+B/BC horizons among the plots. Soil disturbance related to ploughing increased the weathering of Ca and Mg largely in the disturbed part of the topmost mineral soil as indicated by the decreasing concentrations of Ca and Mg after mechanical soil disturbance. The weathering input of Ca in the undisturbed soil did not fully replace the Ca output in final whole-tree cutting. The weathering input of Mg in the undisturbed soil was sufficient to replace the lost Mg in stemwood harvesting but not on all the plots the lost Mg in whole-tree harvesting. Weathering rates were higher in the gabbro than the granitic areas.

  • Lindroos, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: antti.lindroos@luke.fi (email)
  • Ilvesniemi, Natural Resources Institute Finland (Luke), Production systems, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: hannu.ilvesniemi@luke.fi
article id 22007, category Research article
Ilkka Korpela, Antti Polvivaara, Saija Papunen, Laura Jaakkola, Noora Tienaho, Johannes Uotila, Tuomas Puputti, Aleksi Flyktman. (2023). Airborne dual-wavelength waveform LiDAR improves species classification accuracy of boreal broadleaved and coniferous trees. Silva Fennica vol. 56 no. 4 article id 22007. https://doi.org/10.14214/sf.22007
Keywords: crown modeling; laser scanning; photogrammetry; individual tree detection; Scandinavia
Highlights: First study to assess dual-wavelength waveform data in tree species identification; New findings regarding waveform features of previously unstudied species; Waveform features correlated with tree size displaying wavelength- and species-specific differences linked to bark reflectance, height growth rate and foliage density; Effects by pulse length and beam divergence are highlighted.
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Tree species identification constitutes a bottleneck in remote sensing applications. Waveform LiDAR has been shown to offer potential over discrete-return observations, and we assessed if the combination of two-wavelength waveform data can lead to further improvements. A total of 2532 trees representing seven living and dead conifer and deciduous species classes found in Hyytiälä forests in southern Finland were included in the experiments. LiDAR data was acquired by two single-wavelength sensors. The 1064-nm and 1550-nm data were radiometrically corrected to enable range-normalization using the radar equation. Pulses were traced through the canopy, and by applying 3D crown models, the return waveforms were assigned to individual trees. Crown models and a terrain model enabled a further split of the waveforms to strata representing the crown, understory and ground segments. Different geometric and radiometric waveform attributes were extracted per return pulse and aggregated to tree-level mean and standard deviation features. We analyzed the effect of tree size on the features, the correlation between features and the between-species differences of the waveform features. Feature importance for species classification was derived using F-test and the Random Forest algorithm. Classification tests showed significant improvement in overall accuracy (74→83% with 7 classes, 88→91% with 4 classes) when the 1064-nm and 1550-nm features were merged. Most features were not invariant to tree size, and the dependencies differed between species and LiDAR wavelength. The differences were likely driven by factors such as bark reflectance, height growth induced structural changes near the treetop as well as foliage density in old trees.

  • Korpela, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID 0000-0002-1665-3984 E-mail: ilkka.korpela@helsinki.fi (email)
  • Polvivaara, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail:
  • Papunen, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID 0000-0001-5383-4314 E-mail: saija.papunen@outlook.com
  • Jaakkola, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: laura.jaakkola@helsinki.fi
  • Tienaho, University of Eastern Finland, Faculty of Science and Forestry, P.O. Box 111, FI-80101 Joensuu, Finland ORCID 0000-0002-6574-5797 E-mail: noora.tienaho@uef.fi
  • Uotila, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: johannes.uotila@helsinki.fi
  • Puputti, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID 0000-0003-1972-1636 E-mail: tuomas.puputti@helsinki.fi
  • Flyktman, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID 0000-0002-5235-317X E-mail: aleksi.flyktman@helsinki.fi
article id 10798, category Research article
Perttu Anttila, Johannes Ojala, Teijo Palander, Kari Väätäinen. (2023). The effect of road characteristics on timber truck driving speed and fuel consumption based on visual interpretation of road database and data from fleet management system. Silva Fennica vol. 56 no. 4 article id 10798. https://doi.org/10.14214/sf.10798
Keywords: fuel consumption; forest roads; CAN bus; forest logistics; greenhouse gas emissions; log truck; road classes
Highlights: Finnish road and pavement classes explain driving speed and fuel consumption of a timber truck; Other significant explanatory variables include the number of road crossings, season, proportion of distance travelled with a loader, and total laden mass of a truck; In the future, higher-resolution tracking data is needed to construct generalisable models for 76-tonne vehicles.
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Road transport produces 90% of greenhouse gas emissions in timber transport in Finland. It is therefore necessary to understand the factors that affect driving speed, fuel consumption, and ultimately, emissions. The objective of this study was to assess the effect of road characteristics on timber truck driving speed and fuel consumption. Data from the fleet management and transport management systems of two timber trucks were collected over a year. A sample of 104 trips was drawn, and the tracking points were overlaid on the road data in a geographical information system. Thereafter, work phases were determined for the points, and they were visually classified into road and pavement classes. Subsequently, the data of 80 trips were utilised in regression analysis to further study the effects of the visually interpreted variables on driving speed and fuel consumption. Fuel consumption was explained by the proportion of forest roads and distance travelled with a loader, and the number of crossings and season when driving without a load. When driving with a load, both asphalt and gravel pavements decreased consumption, in contrast to an unpaved road. Crossings increased fuel consumption, as did the winter and spring months, and the total laden mass of the truck. In conclusion, the study showed that the functional Finnish road and pavement classes can be used to predict driving speed and fuel consumption.

  • Anttila, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0002-6131-392X E-mail: perttu.anttila@luke.fi (email)
  • Ojala, UPM Metsä, Sirkkalantie 13 b, FI-80100 Joensuu, Finland E-mail: johannes.ojala@upm.com
  • Palander, School of Forest Sciences, University of Eastern Finland (UEF), Yliopistokatu 7, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0002-9284-5443 E-mail: teijo.s.palander@uef.fi
  • Väätäinen, Natural Resources Institute Finland (Luke), Yliopistokatu 6, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0002-6886-0432 E-mail: kari.vaatainen@luke.fi
article id 10769, category Research article
Harri Mäkinen, Pekka Nöjd, Samuli Helama. (2022). Recent unexpected decline of forest growth in North Finland: examining tree-ring, climatic and reproduction data. Silva Fennica vol. 56 no. 4 article id 10769. https://doi.org/10.14214/sf.10769
Keywords: Pinus sylvestris; Picea abies; precipitation; temperature; seed production; tree-ring width; growth variation
Highlights: Tree-ring indices of Scots pine showed decadal variations and a prolonged reduction both on mineral soil sites and peatlands after the mid 2000s; The indices of Norway spruce had less pronounced decadal variations and no trend-like reduction over the last 15 years; Temperature and drought explain some part of the observed growth variability.
Abstract | Full text in HTML | Full text in PDF | Author Info

After a decades-long increasing trend, the recent results of the National Forest Inventory (NFI) reported a decline of forest growth in North Finland. The aim of this study was to assess climatic and reproduction influences behind the growth decline. We used tree-ring data that had been collected by NFI using systematic sampling. The tree-ring width series were detrended using the regional curve standardisation (RCS) removing age-related trends. The resulting tree-ring indices of Scots pine (Pinus sylvestris L.) showed decadal variations with low increment in the 1990s, and high increment in the 1980s and the early years of the current century. Thereafter, a prolonged growth reduction for pine started both on the mineral soil sites and peatlands. The tree-ring indices of Norway spruce (Picea abies (L.) Karst.) had less pronounced decadal variations and no trend-like reduction over the last 15 years. High spring and summer temperatures were found to enhance radial growth, but high winter temperatures were related to low growth for pine and spruce in the following summer. Temperature variation, accompanied by variables indicating years of drought and intensive flowering, accounted for 34% annual growth variance of pine and 21–44% for spruce. Thus, the results imply that climatic factors may have to some extent contributed to the recent growth reduction of pine. Due to its ecological and economic consequences growth decline needs to be further monitored and investigated. Moreover, analyses of stand and age structure, potentially affecting the growth decline, were beyond the scope of this paper, but also warrant further investigation.

  • Mäkinen, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, 00790 Helsinki ORCID https://orcid.org/0000-0002-1820-6264 E-mail: harri.makinen@luke.fi (email)
  • Nöjd, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, 00790 Helsinki E-mail: pekka.nojd@luke.fi
  • Helama, Natural Resources Institute Finland (Luke), Ounasjoentie 6, 96200 Rovaniemi, Finland ORCID https://orcid.org/0000-0002-9777-3354 E-mail: samuli.helama@luke.fi
article id 10762, category Research article
Karri Uotila, Jaana Luoranen, Timo Saksa, Tiina Laine, Juha Heiskanen. (2022). Long-term growth response of Norway spruce in different mounding and vegetation control treatments on fine-textured soils. Silva Fennica vol. 56 no. 4 article id 10762. https://doi.org/10.14214/sf.10762
Keywords: boreal forest; establishment; regeneration; site preparation; plantation; tree growth; scarification; fine-textured soils; mechanical vegetation control
Highlights: Spot and ditch mounding methods favoured spruce sapling development on fine-textured soils; Inverted and unprepared plots showed the weakest growth; Vegetation control suppressed the growth differences between site preparation methods; Vegetation control reduced the density of resprouts after early cleaning.
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Mechanical site preparation (MSP) is a common practice that precedes the planting of Norway spruce (Picea abies (L.) H. Karst.) in Nordic forests. Mounding has become the most used method in spruce planting in recent years. This study examined the effects of different mounding treatments (spot and ditch mounding, inversion, unprepared control with or without herbicide application) and a mechanical vegetation control (MVC) treatment done 3–4 years after planting on the post-planting growth of spruce container seedlings and their development to saplings during the first 11–13 years on two forest till soils in central Finland, one on flat terrain and other on a southwest slope. On these fine-textured soils the spot and ditch mounding methods favoured spruce saplings development. Inversion and unprepared plots showed weakest growth. On the site with flat terrain, 11 years post planting, spruce saplings were 78–144 cm (38–80%) taller and their breast height diameters were 11–13 mm (60–74%) thicker for ditch or spot mounding than for inversion or herbicide treatment. On the site with sloped terrain the differences were minor between the MSP treatments. MVC improved spruce height growth on sites which did not have intensive MSP, especially on control saplings planted on unprepared soil in herbicide and inversion treatments. On the flat terrain, MVC reduced the density of resprouts to be removed later in pre-commercial thinning. As a conclusion, spot or ditch mounding favoured the growth of spruce over inversion especially on flat terrain with fine-textured soil.

  • Uotila, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: karri.uotila@luke.fi (email)
  • Luoranen, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID https://orcid.org/0000-0002-6970-2030 E-mail: jaana.luoranen@luke.fi
  • Saksa, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID https://orcid.org/0000-0002-1776-2357 E-mail: timo.saksa@luke.fi
  • Laine, Metsä Forest, Revontulenpuisto 2, FI-02100 Espoo, Finland ORCID https://orcid.org/0000-0002-6448-8274 E-mail: tiina.laine@metsagroup.com
  • Heiskanen, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland ORCID https://orcid.org/0000-0002-0549-835X E-mail: juha.heiskanen@luke.fi
article id 10757, category Research article
Silvana M.J. Sione, Silvia G. Ledesma, Pablo G. Aceñolaza, Marcelo G. Wilson. (2022). Biomass Carbon and Nitrogen allocation in different tree species: do tree compartments and size affect C:N relationship? Silva Fennica vol. 56 no. 4 article id 10757. https://doi.org/10.14214/sf.10757
Keywords: Neltuma nigra; Neltuma affinis; Vachellia caven; biomass compartments; C:N ratio; native forests
Highlights: C:N relationship variations in biomass compartments were positively correlated with N concentration and did not exhibit significant association with C; C:N ratios differed significantly among species and compartments; Only in Neltuma affinis and Vachellia caven stems C:N relationship differed among tree size.
Abstract | Full text in HTML | Full text in PDF | Author Info

Tree carbon (C) and nitrogen (N) concentrations and C:N ratio are critical for understanding the elemental compositions of forests, N use efficiency, productivity and the biogeochemical cycles. We evaluate differences in C and N allocation among biomass compartments of three N‑fixing tree species of Espinal Argentine eco-region; the scaling relationship between C and N and the C:N ratio variation among compartments and tree size. Neltuma affinis (Spreng.) C.E. Hughes & G.P. Lewis, Neltuma nigra (Griseb.) C.E. Hughes & G.P. Lewis and Vachellia caven (Molina) Seigler & Ebinger plants (n = 30 for each species) were felled, grouped by stem basal diameter-based size classes and partitioned into 3 biomass compartments: stem (st), large branches (lb) and small branches + leaves, flowers and fruits (sbl). C and N concentrations were markedly influenced by species and biomass compartments. In general, sbl compartment presented more N than the st and lb, while C concentrations in Neltuma stems were the highest. Overall, no isometric C–N scaling relationships were found in different compartments. C:N variations in compartments were positively correlated with N concentrations but did not exhibit any significant association with C concentrations. C:N ratios differed significantly among species and biomass compartments. The C:N ratio for compartments ranked in an order of st > lb > sbl. C:N ratio variability in sbl was the least. Only in N. affinis and V. caven stems C:N relationship differed among tree size. Our results provide evidence of the importance of using in situ C and N concentration per main tree species and biomass compartments, to more accurate estimates of C and N stocks.

  • Sione, National University of Entre Ríos, Faculty of Agricultural Sciences, Ruta 11 Km 10,5 (3101) Oro Verde, Entre Ríos, Argentina; CICyTTP-CONICET (National Research and Technological Investigation Council), Materi y España (3105) Diamante, Entre Ríos, Argentina ORCID https://orcid.org/0000-0002-2119-0363 E-mail: silvana.sione@fca.uner.edu.ar (email)
  • Ledesma, National University of Entre Ríos, Faculty of Agricultural Sciences, Ruta 11 Km 10,5 (3101) Oro Verde, Entre Ríos, Argentina ORCID https://orcid.org/0000-0002-9264-1766 E-mail: silvia.ledesma@fca.uner.edu.ar
  • Aceñolaza, National University of Entre Ríos, Faculty of Agricultural Sciences, Ruta 11 Km 10,5 (3101) Oro Verde, Entre Ríos, Argentina; CICyTTP-CONICET (National Research and Technological Investigation Council), Materi y España (3105) Diamante, Entre Ríos, Argentina ORCID https://orcid.org/0000-0002-5033-3466 E-mail: pablo.acenolaza@fca.uner.edu.ar
  • Wilson, National University of Entre Ríos, Faculty of Agricultural Sciences, Ruta 11 Km 10,5 (3101) Oro Verde, Entre Ríos, Argentina ORCID https://orcid.org/0000-0002-1214-1041 E-mail: wilson.marcelo@inta.gob.ar
article id 10676, category Research article
Eva Ring, Fredrik Johansson, Claudia von Brömssen, Isabelle Bergkvist. (2022). A snapshot of forest buffers near streams, ditches, and lakes on forest land in Sweden – lessons learned. Silva Fennica vol. 56 no. 4 article id 10676. https://doi.org/10.14214/sf.10676
Keywords: forestry; conifer; harvest; lake; riparian; stream; watercourse
Highlights: Forest buffers were inventoried on 174 harvested and site-prepared compartments bordering surface water in Sweden; Buffers with 100% shoreline coverage were present beside all 16 lakes and 55% of the natural or modified stream reaches; Judging streams´ character from field inspection of individual reaches alone proved difficult on forest land affected by historic drainage activities.
Abstract | Full text in HTML | Full text in PDF | Author Info

Forest buffers beside surface water can mitigate negative effects of logging. To gain more information on buffer implementation in operational forestry, forest buffers were inventoried during 2018 on 174 harvested and site-prepared compartments traversed by or bordering streams, ditches and lakes in three regions across Sweden 2–4 years after clearcutting. Most of the inventoried stream and ditch reaches were ≤5 m wide. The water reaches were categorized as lakes (n = 16), natural streams (n = 50), modified streams (n = 21) or ditches (n = 87). Forest buffers with 100% shoreline coverage were present along all lake reaches and 55% and 10% of the natural or modified stream and ditch reaches, respectively. Buffers were absent beside 14% of the natural or modified stream reaches and 61% of the ditch reaches. Lake reaches had significantly wider buffers on average than ditch reaches and natural or modified stream reaches. The mean (SE) buffer widths beside lakes, natural or modified stream reaches and ditch reaches across all three regions and shoreline coverage classes were 12 (1.1), 6.6 (0.6) and 1.5 (0.5) m, respectively. The character of the local stream networks (natural or modified streams or ditches) containing each inventoried reach, were assessed using map information and the reaches´ field classifications. This illustrated the difficulty of judging a streams´ character based solely on field inspections of individual reaches on forest land where historic drainage activities have been performed. We recommend that also upstream and downstream conditions should be considered when planning environmental measures to protect surface water bodies.

  • Ring, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, 751 83 Uppsala, Sweden ORCID https://orcid.org/0000-0002-8962-9811 E-mail: eva.ring@skogforsk.se (email)
  • Johansson, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, 751 83 Uppsala, Sweden E-mail: fredrik.johansson@skogforsk.se
  • von Brömssen, Department of energy and technology, Division of applied statistics and mathematics, Swedish University of Agricultural Sciences, 750 07 Uppsala, Sweden ORCID https://orcid.org/0000-0002-1452-8696 E-mail: Claudia.von.Bromssen@slu.se
  • Bergkvist, Mellanskog, Uppsala Science Park, Box 127, 751 04 Uppsala, Sweden E-mail: isabelle.bergkvist@mellanskog.se
article id 10755, category Research article
Rikard Jonsson, Lotta Woxblom, Rolf Björheden, Eva-Maria Nordström, Bosko Blagojevic, Ola Lindroos. (2022). Analysis of decision-making processes for strategic technology investments in Swedish large-scale forestry. Silva Fennica vol. 56 no. 3 article id 10755. https://doi.org/10.14214/sf.10755
Keywords: information needs; qualitative analysis; harwarder; forest technology development; semi-structured interviews; unstructured decision processes
Highlights: When making development decisions, respondents representing six relatively large users of forest technology aimed to maximize economic criteria without falling below threshold values for criteria such as operator well-being, soil rutting, and wood value; Collaboration between users, manufacturers, and researchers was found to be important; Decision-making could be improved by using tools such as problem-structuring methods, simulations, and optimization.
Abstract | Full text in HTML | Full text in PDF | Author Info

Technological development gives forest companies opportunities to maintain competitiveness in the highly cost-sensitive market for forest products. However, no previous studies have examined the technological development decisions made by forest companies or the support tools used when making them. We therefore aimed to describe and analyze 1) the processes used when making such decisions, 2) the associated decision situations, and 3) the use of and need for decision support tools in these processes, with a harwarder concept as case. Semi-structured interviews were conducted with respondents from six forestry organizations. Two theoretical frameworks were used to analyze the interviews, one for unstructured decision processes and one for decision situations. The respondents’ descriptions of their decision processes were consistent with those observed in other industries, and it was shown that decision-making could potentially be improved by investing more resources into diagnosing the problem at hand. The main objective in decision-making was to maximize economic criteria while satisfying threshold requirements relating to criteria such as operator well-being, soil rutting, and wood value. When facing large uncertainties, interviewees preferred to gather data through operational trials and/or scientific studies. If confronted with large uncertainties that could not be reduced, they proceeded with development only if the potential gains exceeded the estimated uncertainties, and implemented innovations in a stepwise manner. These results indicate a need for greater use of existing decision-support tools such as problem-structuring methods to enable more precise diagnoses, simulations to better understand new innovations, and optimization to better evaluate their theoretical large-scale potential.

  • Jonsson, The Forestry Research Institute of Sweden (Skogforsk), Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: rikard.jonsson@skogforsk.se (email)
  • Woxblom, The Forestry Research Institute of Sweden (Skogforsk), Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: lotta.woxblom@skogforsk.se
  • Björheden, The Forestry Research Institute of Sweden (Skogforsk), Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: rolf.bjorheden@skogforsk.se
  • Nordström, Swedish University of Agricultural Sciences, Department of Forest Resource Management, SE-901 83 Umeå, Sweden E-mail: eva-maria.nordstrom@slu.se
  • Blagojevic, University of Novi Sad, Department of Water Management, Trg D. Obradovica 8, 21000 Novi Sad, Serbia E-mail: bosko.blagojevic@polj.edu.rs
  • Lindroos, Swedish University of Agricultural Sciences, Department of Forest Biomaterials and Technology, SE-901 83 Umeå, Sweden E-mail: ola.lindroos@slu.se
article id 10754, category Research article
Robert E. Tatina, Brice B. Hanberry. (2022). Historical forests of the Black Hills, South Dakota, USA, determined using General Land Office surveys. Silva Fennica vol. 56 no. 3 article id 10754. https://doi.org/10.14214/sf.10754
Keywords: fire; Ponderosa pine; settlement; ecological reference; Government Land Office; GLO; range of variation
Highlights: Based on the U.S. General Land Office survey, the historical (1878–1915) Black Hills landscape in southwestern South Dakota, USA, was relatively open; Historical survey points with trees had lesser tree densities than current forest plots; Ponderosa pine has become less dominant as fire-sensitive species have increased; This study provides another line of evidence of a changed forest structure.
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Forests in the western United States generally have increased in tree density since Euro-American settlement, particularly through increases in fire-sensitive species, such as spruces, firs, and junipers. Like most areas, the Black Hills region in western South Dakota and eastern Wyoming was logged for forest products and underwent agricultural conversion before historical forests were documented. To supplement historical reconstructions and accounts, we compared tree composition and densities (diameters ≥12.7 cm at 1.37 m above ground height) from historical General Land Office (GLO) records (years 1878 to 1915) and current Forest Inventory and Analysis (FIA) tree surveys (years 2011 to 2016) in the Black Hills Highlands of South Dakota. For composition, ponderosa pine (Pinus ponderosa P. Lawson & C. Lawson) decreased from 95% to 86% of all trees, with a consequent increase specifically of white spruce (Picea glauca (Moench) Voss) from 1.5% to 6.7% of all trees. Ponderosa pine currently is smaller in mean diameter by 7.4 cm, while white spruce is larger in mean diameter by 2.4 cm than historically. When the 35% of historical survey points without recorded trees were excluded, historical tree densities indicated an overall forested structure of savannas and open woodlands with tree densities ranging from 66 trees ha–1 to 162 trees ha–1. However, historical forests of the Black Hills incorporated dense stands. Tree densities have increased two- to more than four-fold, to 311 trees ha–1 currently. These comparisons provide another source of information, paralleling changes documented in surface fire-dependent pine and oak forests throughout the United States, of transitions in forest composition and structure since Euro-American settlement.

  • Tatina, Department of Biological Sciences, Dakota Wesleyan University, Mitchell, SD 57301 USA E-mail: rotatina@dwu.edu (email)
  • Hanberry, USDA Forest Service, Rocky Mountain Research Station, Rapid City, SD 57702 USA E-mail: brice.hanberry@usda.gov
article id 10732, category Research article
Ana Aza, A. Maarit I. Kallio, Timo Pukkala, Ari Hietala, Terje Gobakken, Rasmus Astrup. (2022). Species selection in areas subjected to risk of root and butt rot: applying Precision forestry in Norway. Silva Fennica vol. 56 no. 3 article id 10732. https://doi.org/10.14214/sf.10732
Keywords: Norway spruce; Scots pine; growth modelling; precision forestry; root and butt rot severity; tree species selection
Highlights: We present the best species to plant on previously spruce-dominated sites with different site indexes and rot levels; We recommend planting Norway spruce on low-rot sites, Scots pine on higher-rot sites, and allowing natural regeneration on low site indexes; We demonstrate the Precision forestry method for determining the optimal tree species in heterogenous stands; In the case study, the method increased net present value by approximately 6% on average.
Abstract | Full text in HTML | Full text in PDF | Author Info

Norway’s most common tree species, Picea abies (L.) Karst. (Norway spruce), is often infected with Heterobasidion parviporum Niemelä & Korhonen and Heterobasidion annosum (Fr.) Bref.. Because Pinus sylvestris L. (Scots pine) is less susceptible to rot, it is worth considering if converting rot-infested spruce stands to pine improves economic performance. We examined the economically optimal choice between planting Norway spruce and Scots pine for previously spruce-dominated clear-cut sites of different site indexes with initial rot levels varying from 0% to 100% of stumps on the site. While it is optimal to continue to plant Norway spruce in regions with low rot levels, shifting to Scots pine pays off when rot levels get higher. The threshold rot level for changing from Norway spruce to Scots pine increases with the site index. We present a case study demonstrating a practical method (“Precision forestry”) for determining the tree species in a stand at the pixel level when the stand is heterogeneous both in site indexes and rot levels. This method is consistent with the concept of Precision forestry, which aims to plan and execute site-specific forest management activities to improve the quality of wood products while minimising waste, increasing profits, and maintaining environmental quality. The material for the study includes data on rot levels and site indexes in 71 clear-cut stands. Compared to planting the entire stand with a single species, pixel-level optimised species selection increases the net present value in almost every stand, with average increase of approximately 6%.

  • Aza, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, PO Box 5003, NO-1432, Ås, Norway ORCID https://orcid.org/0000-0002-6416-6697 E-mail: anfe@nmbu.no (email)
  • Kallio, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, PO Box 5003, NO-1432, Ås, Norway E-mail: maarit.kallio@nmbu.no
  • Pukkala, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: timo.pukkala@uef.fi
  • Hietala, Norwegian Institute of Bioeconomy Research, PO Box 115, NO-1431 Ås, Norway E-mail: ari.hietala@nibio.no
  • Gobakken, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, PO Box 5003, NO-1432, Ås, Norway E-mail: terje.gobakken@nmbu.no
  • Astrup, Norwegian Institute of Bioeconomy Research, PO Box 115, NO-1431 Ås, Norway E-mail: rasmus.astrup@nibio.no
article id 10712, category Research article
Kenneth Olofsson, Johan Holmgren. (2022). Co-registration of single tree maps and data captured by a moving sensor using stem diameter weighted linking. Silva Fennica vol. 56 no. 3 article id 10712. https://doi.org/10.14214/sf.10712
Keywords: airborne laser scanning; terrestrial laser scanning; field plot; mobile laser scanning; simultaneous location and mapping; stem map
Highlights: A stem diameter weighted linking algorithm for tree maps was introduced which improves linking accuracy; A new simultaneous location and mapping-based co-registration method for stem maps measured with moving sensors was introduced that operates with high linking accuracy.
Abstract | Full text in HTML | Full text in PDF | Author Info

A new method for the co-registration of single tree data in forest stands and forest plots applicable to static as well as dynamic data capture is presented. This method consists of a stem diameter weighted linking algorithm that improves the linking accuracy when operating on diverse diameter stands with stem position errors in the single tree detectors. A co-registration quality metric threshold, QT, is also introduced which makes it possible to discriminate between correct and incorrect stem map co-registrations with high probability (>99%). These two features are combined to a simultaneous location and mapping-based co-registration method that operates with high linking accuracy and that can handle sensors with drifting errors and signal bias. A test with simulated data shows that the method has an 89.35% detection rate. The statistics of different settings in a simulation study are presented, where the effect of stem density and position errors were investigated. A test case with real sensor data from a forest stand shows that the average nearest neighbor distances decreased from 1.90 m to 0.51 m, which indicates the feasibility of this method.

  • Olofsson, Section of Forest Remote Sensing, Department of Forest Resource Management, Swedish University of Agricultural Sciences, Umeå, Sweden ORCID https://orcid.org/0000-0002-2836-2316 E-mail: kenneth.olofsson@slu.se (email)
  • Holmgren, Section of Forest Remote Sensing, Department of Forest Resource Management, Swedish University of Agricultural Sciences, Umeå, Sweden ORCID https://orcid.org/0000-0002-7112-8015 E-mail: johan.holmgren@slu.se
article id 10698, category Research article
Constança Camilo-Alves, Jose Antonio Nunes, Ana Patricia Poeiras, Joao Ribeiro, Cati Dinis, Joao M. Barroso, Margarida Vaz, Nuno Almeida-Ribeiro. (2022). Influence of water and nutrients on cork oak radial growth – looking for an efficient fertirrigation regime. Silva Fennica vol. 56 no. 3 article id 10698. https://doi.org/10.14214/sf.10698
Keywords: irrigation; mixed models; Quercus suber; climate–growth relationship; increments; stem radius
Highlights: Different summer fertirrigation treatments were tested on cork oaks over four years in a 1 ha plot; Radial growth, meteorological parameters and fertirrigation volume were measured every 15–30 days; During summer fertirrigated trees grew significantly more, independently of air vapor pressure deficit; Increments were linearly related with fertirrigation volume up to 140 m3 week–1.
Abstract | Full text in HTML | Full text in PDF | Author Info

The widespread cork oak (Quercus suber L.) mortality and reduced afforestation /regeneration are causing an overall reduction in cork production. To enhance trees’ growth and vitality, afforestation techniques using fertirrigation were tested. The main objective was the promotion of trees’ growth on new dense plantations using minimum water requirements until reaching productive forests. The experimental plot – Irricork – was installed in 2017 in a ≈1 ha stand with 14 years’ age cork oaks summer-fertirrigated since plantation. Four fertirrigation treatments were applied during fertirrigation campaigns. Radial growth, meteorological parameters and fertirrigation volume were measured every 15–30 days over four years. It was observed that weather, tree size, debarking and trees’ intra-competition had a significant effect on radial increments. Fertirrigation significantly enhanced growth during summer drought and decoupled increments from air vapor pressure deficit constraints. There was a linear relationship between trees’ radial increments and fertirrigation volume up to 140 m3 week–1. Above this value, increments were smoother. In conclusion, summer fertirrigation of 140 m3 week–1 efficiently enhanced the radial growth of trees with 50–75 circumference at breast height, under the particular edaphoclimatic conditions of the stand. This study showed to be, therefore, promising in the use of efficient fertirrigation the enhance cork oaks’ radial growth.

  • Camilo-Alves, MED – Mediterranean Institute for Agriculture, Environment and Development & CHANGE – Global Change and Sustainability Institute, Institute for Advanced Studies and Research, University of Evora, Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal ORCID https://orcid.org/0000-0001-5156-172X E-mail: calves@uevora.pt (email)
  • Nunes, Department of Plant Science, School of Science and Technology, University of Evora, Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal ORCID https://orcid.org/0000-0002-6144-3484 E-mail: jain@uevora.pt
  • Poeiras, MED – Mediterranean Institute for Agriculture, Environment and Development & CHANGE – Global Change and Sustainability Institute, Institute for Advanced Studies and Research, University of Evora, Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal ORCID https://orcid.org/0000-0002-6049-807X E-mail: apcp@uevora.pt
  • Ribeiro, Department of Plant Science, School of Science and Technology, University of Evora, Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal ORCID https://orcid.org/0000-0001-7242-5866 E-mail: jmrpr@uevora.pt
  • Dinis, ORCID https://orcid.org/0000-0001-6984-1033 E-mail: dinis.cati@gmail.com
  • Barroso, MED – Mediterranean Institute for Agriculture, Environment and Development & CHANGE – Global Change and Sustainability Institute, and Department of Plant Science, School of Science and Technology, University of Evora, Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal ORCID https://orcid.org/0000-0002-0160-3845 E-mail: jmmb@uevora.pt
  • Vaz, MED – Mediterranean Institute for Agriculture, Environment and Development & CHANGE – Global Change and Sustainability Institute, and Department of Biology, School of Science and Technology, University of Evora. Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal ORCID https://orcid.org/0000-0002-3368-757X E-mail: mvaz@uevora.pt
  • Almeida-Ribeiro, ICT – Institute of Earth Sciences and Department of Plant Science, School of Science and Technology, University of Evora, Pólo da Mitra, Ap. 94, 7006-554 Évora, Portugal ORCID https://orcid.org/0000-0002-0160-3845 E-mail: nmcar@uevora.pt
article id 10660, category Research article
Åsa Gustafsson, Jimmy Johansson. (2022). Identifying present drivers of product development and describing roles of identified actors primarily affecting the development of harvesters: a multiple-case study. Silva Fennica vol. 56 no. 3 article id 10660. https://doi.org/10.14214/sf.10660
Keywords: forest operations; logging contractor; industry driver; interview; softwood lumber supply chain
Highlights: Legislators, logging contractors, and expert and research organizations are present drivers of product development of harvesters; They appear to prioritize meeting legal regulations and lowering costs for logging contractors as they outline requirements for manufacturing harvesters.
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Forest operations involve several different actors. Each actor imposes their own requirements on the harvester in relation to their differing roles in the industry, whether they are concerned with the harvester itself, information, environmental concerns, etc. The manufacturers of harvesters need to meet the requirements imposed by multiple actors, among them logging contractors, whose survival depends on their harvesters. This paper aims to identify the present drivers of product development and describe the roles of the actors who have been identified as those currently affecting the development of harvesters. A multiple-case study of harvester manufacturers was conducted. In total, 4 cases were studied. Each case was comprised of five interviewees: two from each harvesting manufacturer, two logging contractors, and one dealer. Following 20 interviews and 3 validation interviews (with experts from both the industry and academia), the paper concludes that the present drivers of product development of harvesters are legislators, logging contractors, and expert and research organizations. Harvester manufacturers appear to develop harvesters aligned with requirements coming from both logging contractors and legislators. Logging contractors are the primary customers, and they prioritize requirements that reduce cost and improve work environments. Legislators, and expert and research organizations are supporting development in relation to current regulations.

  • Gustafsson, Department of Accounting and Logistics, Linnaeus University, Universitetsplatsen 1, 352 52 Växjö, Sweden E-mail: asa.gustafsson@lnu.se (email)
  • Johansson, Department of Forestry and Wood Technology, Linnaeus University, 391 82 Kalmar, Sweden E-mail: jimmy.johansson@lnu.se
article id 10707, category Research article
Martin Goude, Urban Nilsson, Euan Mason, Giulia Vico. (2022). Comparing basal area growth models for Norway spruce and Scots pine dominated stands. Silva Fennica vol. 56 no. 2 article id 10707. https://doi.org/10.14214/sf.10707
Keywords: Pinus sylvestris; basal area; Picea abies; National Forest Inventory; regression; difference equation; long-term experiment
Highlights: Models were developed that predict basal area growth for Scot pine and Norway spruce stands in Sweden; There were no apparent differences in the ability to predict basal area development between a linear regression model for basal area growth or a compatible growth and yields model for basal area; The model based on data from the 80s had similar performance as the models with data from the 2000s, showing that both can reliably be used to predict forest development.
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Models that predict forest development are essential for sustainable forest management. Constructing growth models via regression analysis or fitting a family of sigmoid equations to construct compatible growth and yield models are two ways these models can be developed. In this study, four species-specific models were developed and compared. A compatible growth and yield stand basal area model and a five-year stand basal area growth model were developed for Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.). The models were developed using data from permanent inventory plots from the Swedish national forest inventory and long-term experiments. The species-specific models were compared, using independent data from long-term experiments, with a stand basal area growth model currently used in the Swedish forest planning system Heureka (Elfving model). All new models had a good, relatively unbiased fit. There were no apparent differences between the models in their ability to predict basal area development, except for the slightly worse predictions for the Norway spruce growth model. The lack of difference in the model comparison showed that despite the simplicity of the compatible growth and yield models, these models could be recommended, especially when data availability is limited. Also, despite using more and newer data for model development in this study, the currently used Elfving model was equally good at predicting basal area. The lack of model difference indicate that future studies should instead focus on model development for heterogeneous forests which are common but lack in growth and yield modelling research.

  • Goude, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, SE-230 53 Alnarp, Sweden ORCID https://orcid.org/0000-0002-2179-292X E-mail: martin.goude@slu.se (email)
  • Nilsson, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, SE-230 53 Alnarp, Sweden E-mail: urban.nilsson@slu.se
  • Mason, School of Forestry, University of Canterbury, Private Bag 4800, Christchurch, New Zealand E-mail: euan.mason@canterbury.ac.nz
  • Vico, Department of Crop Production Ecology, Swedish University of Agricultural Sciences, SE-750 07 Uppsala, Sweden E-mail: giulia.vico@slu.se
article id 10695, category Research article
Ana de Lera Garrido, Terje Gobakken, Hans Ole Ørka, Erik Næsset, Ole M. Bollandsås. (2022). Estimating forest attributes in airborne laser scanning based inventory using calibrated predictions from external models. Silva Fennica vol. 56 no. 2 article id 10695. https://doi.org/10.14214/sf.10695
Keywords: forest inventory; LIDAR; calibration; area-based approach; spatial transferability; temporal transferability
Highlights: Three approaches to calibrate temporal and spatial external models using field observations from different numbers of local plots are presented; Calibration produced satisfactory results, reducing the mean difference between estimated and observed values in 89% of all trials; Using few calibration plots, ratio-calibration provided the lowest mean difference; Calibration using 20 plots gave comparable results to a local forest inventory.
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Forest management inventories assisted by airborne laser scanner data rely on predictive models traditionally constructed and applied based on data from the same area of interest. However, forest attributes can also be predicted using models constructed with data external to where the model is applied, both temporal and geographically. When external models are used, many factors influence the predictions’ accuracy and may cause systematic errors. In this study, volume, stem number, and dominant height were estimated using external model predictions calibrated using a reduced number of up-to-date local field plots or using predictions from reparametrized models. We assessed and compared the performance of three different calibration approaches for both temporally and spatially external models. Each of the three approaches was applied with different numbers of calibration plots in a simulation, and the accuracy was assessed using independent validation data. The primary findings were that local calibration reduced the relative mean difference in 89% of the cases, and the relative root mean squared error in 56% of the cases. Differences between application of temporally or spatially external models were minor, and when the number of local plots was small, calibration approaches based on the observed prediction errors on the up-to-date local field plots were better than using the reparametrized models. The results showed that the estimates resulting from calibrating external models with 20 plots were at the same level of accuracy as those resulting from a new inventory.

  • de Lera Garrido, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: ana.de.lera@nmbu.no (email)
  • Gobakken, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: terje.gobakken@nmbu.no
  • Ørka, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: hans-ole.orka@nmbu.no
  • Næsset, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: erik.naesset@nmbu.no
  • Bollandsås, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: ole.martin.bollandsas@nmbu.no
article id 10663, category Research article
Back Tomas Ersson, Lars-Göran Sundblad, Jussi Manner. (2022). Cost analysis of seedling supply systems adapted for mechanized tree planting: a case study from southern Sweden. Silva Fennica vol. 56 no. 2 article id 10663. https://doi.org/10.14214/sf.10663
Keywords: logistics; silviculture; reforestation; tree planting machine; containerized seedling; seedling handling; system analysis
Highlights: The total cost of cardboard box concepts that increase the productivity of tree planting machines is higher than of the cultivation tray system (5–49% in the basic scenario); Increasing the boxes’ packing densities and/or the planting machines’ hourly cost increases the boxes’ cost-competitiveness; Packing density is a key factor in achieving highly cost-efficient seedling supply systems for mechanized tree planting.
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Because today’s tree planting machines do a good job silviculturally, the Nordic forest sector is interested in finding ways to increase the planting machines’ productivity. Faster seedling reloading increases machine productivity, but that solution might require investments in specially designed seedling packaging. The objective of our study was to compare the cost-efficiency of cardboard box concepts that increase the productivity of tree planting machines with that of today’s two most common seedling packaging systems in southern Sweden. We modelled the total cost of these five different seedling packaging systems using data from numerous sources including manufacturers, nurseries, contractors, and forest companies. Under these southern Swedish conditions, the total cost of cardboard box concepts that increase the productivity of intermittently advancing tree planting machines was higher than the cost of the cultivation tray system (5–49% in the basic scenario). However, the conceptual packaging system named ManBox_fast did show promise, especially with increasing primary transport distances and increased planting machine productivities and hourly costs. Thus, our results show that high seedling packing density is of fundamental importance for cost-efficiency of cardboard box systems designed for mechanized tree planting. Our results also illustrate how different factors in the seedling supply chain affect the cost-efficiency of tree planting machines. Consequently, our results underscore that the key development factor for mechanized tree planting in the Nordic countries is the development of cost-efficient seedling handling systems between nurseries and planting machines.

  • Ersson, SLU, School of Forest Management, SE-739 21 Skinnskatteberg, Sweden ORCID https://orcid.org/0000-0003-2442-7482 E-mail: back.tomas.ersson@slu.se (email)
  • Sundblad, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: lars-goran.sundblad@skogforsk.se
  • Manner, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID https://orcid.org/0000-0002-4982-3855 E-mail: jussi.manner@skogforsk.se
article id 10620, category Research article
Tapani Repo, Virva Volanen, Pertti Pulkkinen. (2022). No difference in the maximum frost hardiness of different pedunculate oak populations in Finland. Silva Fennica vol. 56 no. 2 article id 10620. https://doi.org/10.14214/sf.10620
Keywords: climate change; differential thermal analysis; electrolyte leakage; low-temperature exotherm; species distribution; visual damage scoring
Highlights: Four pedunculate oak populations were compared for their maximum frost hardiness (FHmax) at two sampling times in midwinter; Based on the initiation of the low temperature exotherm (LTE), FHmax was an average of –41 °C; Based on the relative electrolyte leakage method, FHmax was an average of –46 °C and –41 °C in the first and second sampling time respectively; No significant differences were observed among the populations; Within-population variation in the LTE was high, providing potential for breeding; In extreme winters, FHmax may be critical for the growth and survival of oak in central and northern Finland.
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The natural northern distribution limit for pedunculate oak (Quercus robur L.) is in southern Finland. We hypothesized that the maximum frost hardiness (FHmax) in the winter limited the cultivation of oaks in northern latitudes. We tested the hypothesis with controlled freezing tests in midwinter. The acorns for the experiment were collected from the four main oak populations in southernmost Finland. The seedlings were raised in the nursery, frost hardened in field conditions, and then moved to a growth chamber at –2 °C on two occasions in winter and tested for FHmax in controlled freezing tests. Frost hardiness was assessed by differential thermal analysis (DTA) based on the low temperature exotherm (LTE) and relative electrolyte leakage (REL) of the stem, and visual damage scoring (VD) of the buds and stem. The initiation and peak of the LTE took place at an average of –41 °C and –43 °C respectively, without differences among the populations. The variation in the initiation and peak of the LTE was high, ranging from –34.6 °C to –45.5 °C and from –37.1 °C to –46.9 °C respectively. According to the REL method, the frost hardiness of the populations ranged from –44.0 °C to –46.4 °C in February and from –40.6 °C to –41.6 °C in March, without significant differences among the populations. According to VD, the bud was the least frost hardy organ, with FH between –19 °C and –33 °C, depending on population and assessment time. We conclude that the maximum hardiness may set the limit for the distribution of pedunculate oak northwards, but the high within-population variation offers potential to breed more frost hardy genotypes.

  • Repo, Natural Resources Institute Finland (Luke), Natural Resources, Yliopistokatu 6b, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0002-7443-6275 E-mail: tapani.repo@luke.fi (email)
  • Volanen, Kalevankatu 4b B21, FI-80110 Joensuu, Finland E-mail: virva.volanen@siunsote.fi
  • Pulkkinen, Natural Resources Institute Finland (Luke), Production systems, Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0002-1643-7691 E-mail: pertti.pulkkinen@luke.fi
article id 10617, category Research article
Yao Anicet Gervais Kouamé, Mathieu Millan, Aya Brigitte N'Dri, Tristan Charles-Dominique, Marcel Konan, Adama Bakayoko, Jacques Gignoux. (2022). Multispecies allometric equations for shrubs and trees biomass prediction in a Guinean savanna (West Africa). Silva Fennica vol. 56 no. 2 article id 10617. https://doi.org/10.14214/sf.10617
Keywords: carbon stocks; allometric equations; shrubs; trees; aboveground and belowground biomass; Guinean savannas
Highlights: New allometric equations were developed for predicting aboveground and belowground biomass (AGB and BGB) of trees and multi-stemmed shrubs in the Guinean savannas based on field measurements, providing information for West African mesic savannas and filling a critical knowledge gap; AGB and BGB of trees were better predicted from the quantity ρDb2H (with ρ the specific wood density in g cm–3, Db the stem basal diameter in cm, and H the tree height in m); Obtaining accurate estimates of AGB and BGB in multi-stemmed shrubs required additional consideration of the total number of stems; The root/shoot biomass ratio decreased with increasing of the stem size (measured by Db) for trees but remains relatively unchanged for shrubs.
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Currently, tools to predict the aboveground and belowground biomass (AGB and BGB) of woody species in Guinean savannas (and the data to calibrate them) are still lacking. Multispecies allometric equations calibrated from direct measurements can provide accurate estimates of plant biomass in local ecosystems and can be used to extrapolate local estimates of carbon stocks to the biome scale. We developed multispecies models to estimate AGB and BGB of trees and multi-stemmed shrubs in a Guinean savanna of Côte d’Ivoire. The five dominant species of the area were included in the study. We sampled a total of 100 trees and 90 shrubs destructively by harvesting their biometric data (basal stem diameter Db, total stem height H, stump area SS, as well as total number of stems n for shrubs), and then measured their dry AGB and BGB. We fitted log-log linear models to predict AGB and BGB from the biometric measurements. The most relevant model for predicting AGB in trees was fitted as follows: AGB = 0.0471 (ρDb2H)0.915 (with AGB in kg and ρDb2H in g cm–1 m). This model had a bias of 19%, while a reference model for comparison (fitted from tree measurements in a similar savanna ecosystem, Ifo et al. 2018) overestimated the AGB of trees of our test savannas by 132%. The BGB of trees was also better predicted from ρDb2H as follows: BGB = 0.0125 (ρDb2H)0.6899 (BGB in kg and ρDb2H in g cm–1 m), with 6% bias, while the reference model had about 3% bias. In shrubs, AGB and BGB were better predicted from ρDb2H together with the total number of stems (n). The best fitted allometric equation for predicting AGB in shrubs was as follows: AGB = 0.0191 (ρDb2H)0.6227 n0.9271. This model had about 1.5% bias, while the reference model overestimated the AGB of shrubs of Lamto savannas by about 79%. The equation for predicting BGB of shrubs is: BGB = 0.0228 (ρDb2H)0.7205 n0.992 that overestimated the BGB of the shrubs of Lamto savannas with about 3% bias, while the reference model underestimated the BGB by about 14%. The reference model misses an important feature of fire-prone savannas, namely the strong imbalance of the BGB/AGB ratio between trees and multi-stemmed shrubs, which our models predict. The allometric equations we developed here are therefore relevant for C stocks inventories in trees and shrubs communities of Guinean savannas.

  • Kouamé, UFR Sciences de la Nature, UFR-SN/ Station d’Ecologie de Lamto (CRE), Pôle de recherche Environnement et Développement Durable, Université NANGUI ABROGOUA, 02 BP 801 Abidjan 02, Côte d’Ivoire); Institute of Ecology and Environmental Sciences IEES-Paris (Sorbonne Université, CNRS, Université Paris Diderot, IRD, UPEC, INRA), 4 Place Jussieu, 75005, Paris, France ORCID https://orcid.org/0000-0002-0847-2569 E-mail: kouameyag@gmail.com (email)
  • Millan, Centre for African Ecology, School of Animal, Plant and Environmental Sciences, University of the Witwatersrand, Private Bag 3, Johannesburg, South Africa; Global Change Biology Group, Department of Botany and Zoology, Stellenbosch University, Private Bag X1, Matieland 7602, South Africa; Institute of Botany of the Czech Academy of Sciences, v.v.i, Dukelská 135, Třeboň, 379 01, Czech Republic ORCID https://orcid.org/0000-0002-0151-6055 E-mail: mathieu.millan@gmail.com
  • N'Dri, UFR Sciences de la Nature, UFR-SN/ Station d’Ecologie de Lamto (CRE), Pôle de recherche Environnement et Développement Durable, Université NANGUI ABROGOUA, 02 BP 801 Abidjan 02, Côte d’Ivoire ORCID https://orcid.org/0000-0002-6333-6279 E-mail: brigitte.aya@gmail.com
  • Charles-Dominique, Institute of Ecology and Environmental Sciences IEES-Paris (Sorbonne Université, CNRS, Université Paris Diderot, IRD, UPEC, INRA), 4 Place Jussieu, 75005, Paris, France ORCID https://orcid.org/0000-0002-5767-0406 E-mail: tristan.charles-dominique@sorbonne-universite.fr
  • Konan, UFR Sciences de la Nature, UFR-SN/ Station d’Ecologie de Lamto (CRE), Pôle de recherche Environnement et Développement Durable, Université NANGUI ABROGOUA, 02 BP 801 Abidjan 02, Côte d’Ivoire E-mail: marcelkonan.lamto@gmail.com
  • Bakayoko, UFR Sciences de la Nature, UFR-SN/ Station d’Ecologie de Lamto (CRE), Pôle de recherche Environnement et Développement Durable, Université NANGUI ABROGOUA, 02 BP 801 Abidjan 02, Côte d’Ivoire E-mail: bakadamaci@yahoo.fr
  • Gignoux, Institute of Ecology and Environmental Sciences IEES-Paris (Sorbonne Université, CNRS, Université Paris Diderot, IRD, UPEC, INRA), 4 Place Jussieu, 75005, Paris, France ORCID https://orcid.org/0000-0003-3853-9282 E-mail: jacques.gignoux@upmc.fr
article id 10606, category Research article
Benjamin Allen, Michele Dalponte, Ari M. Hietala, Hans Ole Ørka, Erik Næsset, Terje Gobakken. (2022). Detection of Root, Butt, and Stem Rot presence in Norway spruce with hyperspectral imagery. Silva Fennica vol. 56 no. 2 article id 10606. https://doi.org/10.14214/sf.10606
Keywords: Picea abies; Heterobasidion; remote sensing; root rot; hyperspectral imagery; forest pathology
Highlights: Hyperspectral imagery can be used to detect Root, Butt, and Stem Rot in Picea abies with moderate accuracy; Spectral derivatives improved classification accuracy; Bands around 540, 700, and 1650 nm tended to be the most important for classification models.
Abstract | Full text in HTML | Full text in PDF | Author Info

Pathogenic wood decay fungi such as species of Heterobasidion are some of the most serious forest pathogens in Europe, causing rot of tree boles and loss of growth, with estimated economic losses of eight hundred million euros per year. In conifers with low resinous heartwood such as species of Picea and Abies, these fungi are commonly confined to heartwood and thus external infection signs on the bark or foliage of trees are normally absent. Consequently, determining the extent of disease presence in a forest stand with field surveys is not practical for guiding forest management decisions such as optimal rotation time. Remote sensing technologies such as airborne laser scanning and aerial imagery are already used to reduce the reliance on fieldwork in forest inventories. This study aimed to use remote sensing to detect rot in spruce (Picea abies L. Karst.) forests in Norway. An airborne hyperspectral imager provided information for classifying the presence or absence of rot in a single-tree-based framework. Ground reference data showing the presence of rot were collected by harvest machine operators during the harvest of forest stands. Random forest and support vector machine algorithms were used to classify the presence and absence of rot. Results indicate a 64% overall classification accuracy for presence-absence classification of rot, although additional work remains to make the classifications usable for practical forest management.

  • Allen, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: benjamin.allen@nmbu.no (email)
  • Dalponte, Department of Sustainable Agro-ecosystems and Bioresources, Research and Innovation Centre, Fondazione E. Mach, Via E. Mach 1, 38098 San Michele all’Adige (TN), Italy E-mail: michele.dalponte@fmach.it
  • Hietala, Norwegian Institute of Bioeconomy Research, Innocamp Steinkjer, Skolegata 22, NO-7713 Steinkjer, Norway E-mail: Ari.Hietala@nibio.no
  • Ørka, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: hans-ole.orka@nmbu.no
  • Næsset, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: erik.naesset@nmbu.no
  • Gobakken, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: terje.gobakken@nmbu.no
article id 10599, category Research article
Anu Laakkonen, Teppo Hujala, Jouni Pykäläinen. (2022). Defining the systemic development of the Finnish pulp and paper industry’s business network. Silva Fennica vol. 56 no. 2 article id 10599. https://doi.org/10.14214/sf.10599
Keywords: forest cluster; competence; Actors-Resources-Activities framework; coopetition; historical pathways
Highlights: Systemic view helps to understand the phenomena reshaping business field networks; Forest sector companies operate in complex, dynamic, and international environment; Actors-Resources-Activities framework -based analysis of Finnish pulp and paper industry’s network development; The role of actors, resources, and activities have varied between different phases; Network structure altered due to radical changes in the operating environment.
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Companies operate in a nested and complex system where global challenges shape their environments and put pressure on business activities. Systemic understanding of the past and ongoing changes within a national industry help to analyze the global influences and identify phenomena that reshape business collaborations. To address this issue in the case of a forest sector, this study constructs a systemic picture of the historical development of the Finnish pulp and paper industry’s business network and analyzes it qualitatively through the Actors-Resources-Activities framework. Books discussing the history of the Finnish forest industry were used as secondary data, which were analyzed with a theory-based content analysis method. The analysis revealed four development phases during which the network has evolved from rather simple one emphasizing cooperation organizations (1st) to a more complex one with stronger roles of the state and individual influencers (2nd), and then emphasizing export and advocacy associations (3rd), before returning to be rather simple, based around three large multinationals and the EU playing an important role (4th). The industry is concerned about securing its key resources, with varying foci. Research and technological innovation activities play an important role together with cooperative interactions. Overall, actors favor a business-as-usual strategy, which is overruled only by a radical change in the operating environment, leading to notable changes in the network. Thus, a suggestion for all actors within the forest sector is that actively detecting and interpreting change signals in the whole environment can help actors in pursuing sustainable activities.

  • Laakkonen, School of Forest Sciences, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0002-6384-7773 E-mail: anu.laakkonen@uef.fi (email)
  • Hujala, School of Forest Sciences, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0002-7905-7602 E-mail: teppo.hujala@uef.fi
  • Pykäläinen, School of Forest Sciences, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: jouni.pykalainen@uef.fi
article id 10627, category Research article
Christian Kuehne, J. Paul McLean, Kobra Maleki, Clara Antón-Fernández, Rasmus Astrup. (2022). A stand-level growth and yield model for thinned and unthinned even-aged Scots pine forests in Norway. Silva Fennica vol. 56 no. 1 article id 10627. https://doi.org/10.14214/sf.10627
Keywords: Pinus sylvestris; mortality; volume growth; seemingly unrelated regression; production forestry; system of equations
Highlights: The presented growth and yield model consists of component equations for dominant height, stem density, total basal area, and total stem volume; The component equations were fitted simultaneously using seemingly unrelated regression; The model is capable to forecast and compare outcomes of varying thinning regimes; The new component equations better represent the improved growing conditions for Scots pine in Norway.
Abstract | Full text in HTML | Full text in PDF | Author Info

Management of Scots pine (Pinus sylvestris L.) in Norway requires a forest growth and yield model suitable for describing stand dynamics of even-aged forests under contemporary climatic conditions with and without the effects of silvicultural thinning. A system of equations forming such a stand-level growth and yield model fitted to long-term experimental data is presented here. The growth and yield model consists of component equations for (i) dominant height, (ii) stem density (number of stems per hectare), (iii) total basal area, (iv) and total stem volume fitted simultaneously using seemingly unrelated regression. The component equations for stem density, basal area, and volume include a thinning modifier to forecast stand dynamics in thinned stands. It was shown that thinning significantly increased basal area and volume growth while reducing competition related mortality. No significant effect of thinning was found on dominant height. Model examination by means of various fit statistics indicated no obvious bias and improvement in prediction accuracy in comparison to existing models in general. An application of the developed stand-level model comparing different management scenarios exhibited plausible long-term behavior and we propose this is therefore suitable for national deployment.

  • Kuehne, Norwegian Institute of Bioeconomy Research, Division of Forestry and Forest Resources, P.O. Box 115, NO-1431 Ås, Norway E-mail: christian.kuehne@nibio.no (email)
  • McLean, Norwegian Institute of Bioeconomy Research, Division of Forestry and Forest Resources, P.O. Box 115, NO-1431 Ås, Norway E-mail: paul.mclean@nibio.no
  • Maleki, Norwegian Institute of Bioeconomy Research, Division of Forestry and Forest Resources, P.O. Box 115, NO-1431 Ås, Norway E-mail: kobra.maleki@nibio.no
  • Antón-Fernández, Norwegian Institute of Bioeconomy Research, Division of Forestry and Forest Resources, P.O. Box 115, NO-1431 Ås, Norway E-mail: clara.anton.fernandez@nibio.no
  • Astrup, Norwegian Institute of Bioeconomy Research, Division of Forestry and Forest Resources, P.O. Box 115, NO-1431 Ås, Norway E-mail: rasmus.astrup@nibio.no
article id 10608, category Research article
Lennart Noordermeer, Erik Næsset, Terje Gobakken. (2022). Effects of harvester positioning errors on merchantable timber volume predicted and estimated from airborne laser scanner data in mature Norway spruce forests. Silva Fennica vol. 56 no. 1 article id 10608. https://doi.org/10.14214/sf.10608
Keywords: forest inventory; ALS; forest harvester; GNSS; precision forestry
Highlights: Timber volume was estimated using harvester and airborne laser scanner (ALS) data acquired with different scanners over eight years; The year of ALS acquisition did not have a significant effect on errors in timber volume estimates; Accuracies of timber volume estimates decreased significantly with increasing levels of positioning error; When using inaccurately positioned harvester data, larger grid cells are beneficial.
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Newly developed positioning systems in cut-to-length harvesters enable georeferencing of individual trees with submeter accuracy. Together with detailed tree measurements recorded during processing of the tree, georeferenced harvester data are emerging as a valuable tool for forest inventory. Previous studies have shown that harvester data can be linked to airborne laser scanner (ALS) data to estimate a range of forest attributes. However, there is little empirical evidence of the benefits of improved positioning accuracy of harvester data. The two objectives of this study were to (1) assess the accuracy of timber volume estimation using harvester data and ALS data acquired with different scanners over multiple years and (2) assess how harvester positioning errors affect merchantable timber volume predicted and estimated from ALS data. We used harvester data from 33 commercial logging operations, comprising 93 731 harvested stems georeferenced with sub-meter accuracy, as plot-level training data in an enhanced area-based inventory approach. By randomly altering the tree positions in Monte Carlo simulations, we assessed how prediction and estimation errors were influenced by different combinations of simulated positioning errors and grid cell sizes. We simulated positioning errors of 1, 2, …, 15 m and used grid cells of 100, 200, 300 and 400 m2. Values of root mean square errors obtained for cell-level predictions of timber volume differed significantly for the different grid cell sizes. The use of larger grid cells resulted in a greater accuracy of timber volume predictions, which were also less affected by positioning errors. Accuracies of timber volume estimates at logging operation level decreased significantly with increasing levels of positioning error. The results highlight the benefit of accurate positioning of harvester data in forest inventory applications. Further, the results indicate that when estimating timber volume from ALS data and inaccurately positioned harvester data, larger grid cells are beneficial.

  • Noordermeer, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, NMBU, P.O. Box 5003, NO-1432 Ås, Norway E-mail: lennart.noordermeer@nmbu.no (email)
  • Næsset, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, NMBU, P.O. Box 5003, NO-1432 Ås, Norway E-mail: erik.naesset@nmbu.no
  • Gobakken, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, NMBU, P.O. Box 5003, NO-1432 Ås, Norway E-mail: terje.gobakken@nmbu.no
article id 10598, category Research article
Argo Orumaa, Kajar Köster, Arvo Tullus, Tea Tullus, Marek Metslaid. (2022). Forest fires have long-term effects on the composition of vascular plants and bryophytes in Scots pine forests of hemiboreal Estonia. Silva Fennica vol. 56 no. 1 article id 10598. https://doi.org/10.14214/sf.10598
Keywords: disturbance; bryophytes; understorey vegetation; vascular plants; hemiboreal forest; fire chronosequence; wildfire
Highlights: We recorded 31 vascular plant and 39 bryophyte species in a chronosequence of Scots pine stands with 12–183 years since fire; Time since fire affected the compositional patterns of vascular plants and bryophytes; The richness of liverworts was higher in recently burned stands due to the presence of Cephaloziella spp.; The richness of dwarf-shrubs increased with longer period since fire.
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Since fire frequency is expected to increase globally due to climate change, it is important to understand its effects on forest ecosystems. We studied the long-term patterns in species diversity, cover and composition of vascular plants and bryophytes after forest fire and the site-related factors behind them. Research was carried out in northwestern Estonia, using a chronosequence of Scots pine (Pinus sylvestris L.) stands, located on nutrient poor sandy soils, where fires had occurred 12, 23, 38, 69, 80 and 183 years ago. In every stand three 100 m2 vegetation plots were established to collect floristic and environmental information. The effects on floristic characteristics of time since fire, light, and soil variables were evaluated with linear mixed models, followed by backward variable selection. Compositional variation was analysed with non-metric multidimensional scaling, Multi-response Permutation Procedures, and Indicator Species Analysis. Altogether, 31 vascular plant and 39 bryophyte species were found in vegetation plots. The cover of the vascular plant and bryophyte layers increased with a longer time since fire. Soil and light variables impacted the richness of several vascular plant and bryophyte groups, whereas only the richness of liverworts and dwarf-shrubs correlated with time since fire. Considerable compositional differences were observed in vascular plant and bryophyte assemblages between recently vs. long-time ago burned stands. To conclude, time since fire significantly impacted compositional patterns of vascular plants and bryophytes in pine forests on nutrient poor soils, although time-related trends in species richness were less evident.

  • Orumaa, Institute of Forestry and Engineering, Estonian University of Life Sciences, Kreutzwaldi 5, 51006, Tartu, Estonia E-mail: argo.orumaa@emu.ee (email)
  • Köster, Department of Environmental and Biological Sciences, University of Eastern Finland, PO Box 111 (Yliopistokatu 7), 80130, Joensuu, Finland E-mail: kajar.koster@helsinki.fi
  • Tullus, Department of Botany, Institute of Ecology and Earth Sciences, University of Tartu, Lai 40, Tartu, 51003, Estonia E-mail: arvo.tullus@ut.ee
  • Tullus, Institute of Forestry and Engineering, Estonian University of Life Sciences, Kreutzwaldi 5, 51006, Tartu, Estonia E-mail: tea.tullus@emu.ee
  • Metslaid, Institute of Forestry and Engineering, Estonian University of Life Sciences, Kreutzwaldi 5, 51006, Tartu, Estonia E-mail: marek.metslaid@emu.ee
article id 10550, category Research article
Miro Demol, Phil Wilkes, Pasi Raumonen, Sruthi M. Krishna Moorthy, Kim Calders, Bert Gielen, Hans Verbeeck. (2022). Volumetric overestimation of small branches in 3D reconstructions of Fraxinus excelsior. Silva Fennica vol. 56 no. 1 article id 10550. https://doi.org/10.14214/sf.10550
Keywords: aboveground biomass; crown architecture; LIDAR; quantitative structure models; common ash; woody tree volume
Highlights: We compare branch diameter and tree woody volume estimates from terrestrial laser scanning data with manual measurements of two Fraxinus excelsior trees; Smaller branch diameters are generally overestimated due to scattering and misalignment errors in the point cloud; Consequently, tree woody volume is overestimated by 38% to 52%; Filtering by reflectance and improved alignment partly mitigate this effect.
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Terrestrial laser scanning (TLS) has been applied to estimate forest wood volume based on detailed 3D tree reconstructions from point cloud data. However, sources of uncertainties in the point cloud data (alignment and scattering errors, occlusion, foliage...) and the reconstruction algorithm type and parameterisation are known to affect the reconstruction, especially around finer branches. To better understand the impacts of these uncertainties on the accuracy of TLS-derived woody volume, high-quality TLS scans were collected in leaf-off conditions prior to destructive harvesting of two forest-grown common ash trees (Fraxinus excelsior L.; diameter at breast height ~28 cm, woody volume of 732 and 868 L). We manually measured branch diameters at 265 locations in these trees. Estimates of branch diameters and tree volume from Quantitative Structure Models (QSM) were compared with these manual measurements. The accuracy of QSM branch diameter estimates decreased with smaller branch diameters. Tree woody volume was overestimated (+336 L and +392 L) in both trees. Branches measuring < 5 cm in diameter accounted for 80% and 83% of this overestimation respectively. Filtering for scattering errors or improved coregistration approximately halved the overestimation. Range filtering and modified scanning layouts had mixed effects. The small branch overestimations originated primarily in limitations in scanner characteristics and coregistration errors rather than suboptimal QSM parameterisation. For TLS-derived estimates of tree volume, a higher quality point cloud allows smaller branches to be accurately reconstructed. Additional experiments need to elucidate if these results can be generalised beyond the setup of this study.

  • Demol, CAVElab – Computational and Applied Vegetation Ecology, Department of Environment, Faculty of Bioscience Engineering, Ghent University, Coupure Links 653, B-9000 Ghent, Belgium; PLECO – Plants and Ecosystems, Faculty of Science, Antwerp University, Universiteitsplein 1, B-2610 Wilrijk, Belgium ORCID https://orcid.org/0000-0002-5492-2874 E-mail: miro.demol@ugent.be (email)
  • Wilkes, UCL Department of Geography, Gower Street, London WC1E 6BT, UK; NERC National Centre for Earth Observation (NCEO), UK ORCID https://orcid.org/0000-0001-6048-536X E-mail: p.wilkes@ucl.ac.uk
  • Raumonen, Mathematics, Tampere University, FI-33101 Tampere, Finland ORCID https://orcid.org/0000-0001-5471-0970 E-mail: pasi.raumonen@tuni.fi
  • Krishna Moorthy, CAVElab – Computational and Applied Vegetation Ecology, Department of Environment, Faculty of Bioscience Engineering, Ghent University, Coupure Links 653, B-9000 Ghent, Belgium ORCID https://orcid.org/0000-0002-6838-2880 E-mail: Sruthi.KrishnaMoorthyParvathi@ugent.be
  • Calders, CAVElab – Computational and Applied Vegetation Ecology, Department of Environment, Faculty of Bioscience Engineering, Ghent University, Coupure Links 653, B-9000 Ghent, Belgium ORCID https://orcid.org/0000-0002-4562-2538 E-mail: kim.calders@ugent.be
  • Gielen, PLECO – Plants and Ecosystems, Faculty of Science, Antwerp University, Universiteitsplein 1, B-2610 Wilrijk, Belgium ORCID https://orcid.org/0000-0002-4890-3060 E-mail: bert.gielen@uantwerpen.be
  • Verbeeck, CAVElab – Computational and Applied Vegetation Ecology, Department of Environment, Faculty of Bioscience Engineering, Ghent University, Coupure Links 653, B-9000 Ghent, Belgium ORCID https://orcid.org/0000-0003-1490-0168 E-mail: hans.verbeeck@ugent.be
article id 10544, category Research article
Lars Sprengel, Heinrich Spiecker, Shuirong Wu. (2022). Two subject specific modelling approaches to construct base-age invariant polymorphic site index curves with varying asymptotes. Silva Fennica vol. 56 no. 1 article id 10544. https://doi.org/10.14214/sf.10544
Keywords: generalized algebraic difference approach; stem analysis; dummy variable approach; gnls; nonlinear mixed-effects models; nlme; Zhongtiaoshan forest region
Highlights: Base-age invariant families of height growth curves with polymorphism and varying asymptotes are presented for the seven economically most important tree species in Zhongtiaoshan forest region, China; The nonlinear fixed-effects approach outperforms the nonlinear mixed-effects approach according to the AIC, but according to RMSE and bias these results are not fully supported.
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For constructing growth and yield models the concept of site index as measure of productivity is crucial. Here, we use nonlinear mixed-effects models (NLME) with random individual effects and nonlinear models with dummy variables as fixed individual effects (NLFE) to fit mechanistic growth functions to stem analysis data of the economically most important tree species in Zhongtiaoshan forest region, China. The Richards and Lundqvist function are formulated into five dynamic equations (R1, R2, L1, L2 and L3) applying the generalized algebraic difference approach (GADA), which inherit polymorphism, varying asymptotes and base-age invariance. According to Akaike information criterion the R1 model as NLFE fits height growth data of Pinus tabuliformis Carrière, Pinus armandii Franch., Quercus liaotungensis Koidz., Quercus aliena Blume and Betula platyphylla Sukaczev best, while for Quercus variabilis Blume R2 as NLFE fits height growth data best. For Larix principis-rupprechtii Mayr L1 as NLME has been selected as best model, as R1 and R2 both as NLFE and NLME are not extrapolating the comparably short length of height growth data well enough. However, according to the root mean square error and bias differences between model fits of both the selected equation and the chosen model fitting approach are not so clear. Presented families of height growth curves serve as planning tools to identify site index and therefore assess productivity of forest stands in the studied region. A direct comparison of the productivity of forest stands of the same tree species is possible due to base-age invariance of the selected models.

  • Sprengel, Chair of Forest Growth and Dendroecology, Albert-Ludwigs-University Freiburg, Tennenbacher Str. 4, 79106 Freiburg, Germany ORCID https://orcid.org/0000-0002-6332-7911 E-mail: lars.sprengel@iww.uni-freiburg.de (email)
  • Spiecker, Chair of Forest Growth and Dendroecology, Albert-Ludwigs-University Freiburg, Tennenbacher Str. 4, 79106 Freiburg, Germany E-mail: instww@uni-freiburg.de
  • Wu, Research Institute of Forest Policy and Information, Chinese Academy of Forestry, No. 1 Dongxiaofu, Haidian District, Beijing 100091, China E-mail: shuirongwu@126.com
article id 10520, category Research article
Shaoqin Yang, Lita Yi, Nuonan Ye, Mengyuan Wu, Meihua Liu. (2022). Spatial pattern dynamics of Cyclobalanopsis myrsinifolia in mixed broad-leaved forests on Tianmu Mountain, eastern China, 1996–2012. Silva Fennica vol. 56 no. 1 article id 10520. https://doi.org/10.14214/sf.10520
Keywords: forest dynamics; spatial distribution; East Asia; evergreen and deciduous broad-leaved mixed forests
Highlights: Spatial distribution pattern monitoring of Cyclobalanopsis myrsinifolia was performed over 16 years in a 1 ha plot; The importance value of C. myrsinifolia decreased between 1996 and 2012; The spatial distribution pattern changed at a spatial scale of 0–25 m; The drivers of the variation in spatial distribution were intra- and interspecific mutual relationships.
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Studies of the spatial patterns of dominant plant species may provide significant insights into processes and mechanisms that maintain stand stability. This study was performed in a permanent 1 ha plot in evergreen and deciduous broad-leaved mixed forests on Tianmu Mountain. Based on two surveys (1996 and 2012), the dynamics of the spatial distribution pattern of the dominant population (Cyclobalanopsis myrsinifolia (Blume) Oersted) and the intra- and interspecific relationships between C. myrsinifolia and other dominant species populations were analyzed using Ripley’s K(r) function. We identified the importance value of a species in a community, which is the sum of the relative density, relative frequency, and relative dominance. The drivers of spatial distribution variation and the maintenance mechanisms of the forest were discussed. The results showed that the importance value of C. myrsinifolia within the community decreased over the past 16 years. The C. myrsinifolia population exhibited a significantly aggregated distribution within a spatial scale of 0–25 m in 1996 whereas it changed to a random distribution at scales larger than 5.5 m in 2012. From 1996 to 2012, the spatial distribution patterns between C. myrsinifolia and Cyclocarya paliurus (Batal.) Iljinsk. and between C. myrsinifolia and Cunninghamia lanceolata (Lamb.) Hook did not change significantly. In 1996, C. myrsinifolia and Daphniphyllum macropodum Miq. were positively associated at the scale of 0–25 m; this relationship was strongly significant at the scale of 6–10 m. However, there was no association between the populations of two species in terms of the spatial distribution at the scale of 0–25 m in 2012. Our findings indicate that the drivers of variation in the spatial distribution of the C. myrsinifolia population were intra- and interspecific mutual relationships as well the seed-spreading mechanism of this species.

  • Yang, Zhejiang Forest Resources Monitoring Center, Hangzhou 310020, China E-mail: 20080095@zafu.edu.cn
  • Yi, School of Forestry and Biotechnology, Zhejiang A & F University, Lin’an 311300, China E-mail: yilita@zafu.edu.cn
  • Ye, School of Forestry and Biotechnology, Zhejiang A & F University, Lin’an 311300, China E-mail: 542243187@qq.com
  • Wu, School of Forestry and Biotechnology, Zhejiang A & F University, Lin’an 311300, China E-mail: 326585523@qq.com
  • Liu, School of Forestry and Biotechnology, Zhejiang A & F University, Lin’an 311300, China E-mail: mhliu@zafu.edu.cn (email)
article id 10512, category Research article
Mateusz Liziniewicz, Ignacio Barbeito, Andis Zvirgzdins, Lars-Göran Stener, Pentti Niemistö, Nils Fahlvik, Ulf Johansson, Bo Karlsson, Urban Nilsson. (2022). Production of genetically improved silver birch plantations in southern and central Sweden. Silva Fennica vol. 56 no. 1 article id 10512. https://doi.org/10.14214/sf.10512
Keywords: Betula pendula; planting; generalized algebraic difference approach; genetic gain; stand basal area starting function
Highlights: The basal area development of genetically improved birch in Sweden was modeled using a generalized algebraic difference approach; The best model fit, both graphically and statistically was delivered by the Korf base model; The analysis of realized gain trial showed a stability of relative differences in basal area between tested genotypes.
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Investing in planting genetically improved silver birch (Betula pendula Roth) in Swedish plantations requires understanding how birch stands will develop over their entire rotation. Previous studies have indicated relatively low production of birch compared to Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.). This could result from using unrepresentative basic data, collected from unimproved, naturally-regenerated birch (Betula spp.) growing on inventory plots often located in coniferous stands. The objective of this study was to develop a basal area development function of improved silver birch and evaluate production over a full rotation period. We used data from 52 experiments including planted silver birch of different genetic breeding levels in southern and central Sweden. The experimental plots were established on fertile forest sites and on former agricultural lands, and were managed with different numbers of thinnings and basal area removal regimes. The model best describing total stand basal area development was a dynamic equation derived from the Korf base model. The analysis of the realized gain trial for birch showed a good stability of the early calculated relative differences in basal area between tested genotypes over time. Thus, the relative difference in basal area might be with cautious used as representation of the realized genetic gain. On average forest sites in southern Sweden, improved and planted silver birch could produce between 6–10.5 m3 ha–1 year–1, while on fertile agriculture land the average productivity might be higher, especially with material coming from the improvement program. The performed analysis provided a first step toward predicting the effects of genetic improvement on total volume production and profitability of silver birch. However, more experiments are needed to set up the relative differences between different improved material.

  • Liziniewicz, The Forestry Research Institute of Sweden, Ekebo, SE-268 90 Svalöv, Sweden E-mail: mateusz.liziniewicz@skogforsk.se
  • Barbeito, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, Alnarp, Sweden; Université de Lorraine, AgroParisTech, INRAE, UMR Silva, Nancy, France E-mail: ignacio.barbeito@slu.se
  • Zvirgzdins, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, Box 49, 23053 Alnarp, Sweden E-mail: andis.zvirgzdins@slu.se (email)
  • Stener, The Forestry Research Institute of Sweden, Ekebo, SE-268 90 Svalöv, Sweden E-mail: lg.stener@telia.com
  • Niemistö, Natural Resources In-stitute Finland (Luke), Natural resources, Seinäjoki, Finland E-mail: pentti.niemisto@luke.fi
  • Fahlvik, The Forestry Research Institute of Sweden, Ekebo, SE-268 90 Svalöv, Sweden E-mail: nils.fahlvik@skogforsk.se
  • Johansson, Tönnersjöheden Experimental Forest, SLU, Simlångsdalen, Sweden E-mail: ulf.johansson@slu.se
  • Karlsson, The Forestry Research Institute of Sweden, Ekebo, SE-268 90 Svalöv, Sweden E-mail: curly.birch@gmail.com
  • Nilsson, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, Box 49, 23053 Alnarp, Sweden E-mail: urban.nilsson@slu.se
article id 10326, category Research article
Per K. Rørstad, Birger Solberg, Erik Trømborg. (2022). Can we detect regional differences in econometric analyses of the Norwegian timber supply? Silva Fennica vol. 56 no. 1 article id 10326. https://doi.org/10.14214/sf.10326
Keywords: econometric specification test; panel data analysis; price elasticities; volume elasticities
Highlights: The first difference econometric specification yields better overall fit than fixed and random effects models; Using region specific price elasticities improve the fit for fixed and random effects models; Statistically significant different price elasticities are found in 12 out of total 15 pairs of regions; Western Norway has particularly high growing stock volume elasticities and low short-term price elasticities.
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Forestry and forest industries are important for regional income and employment in Norway as well as in most North European countries, but few studies exist about factors affecting the timber supply at regional level. The main objective of this study is to estimate aggregated regional timber supply elasticities for six regions in Norway. Thereby we also test for regional differences, focusing on wood prices, standing stock volume and interest rate as explanatory variables. We have used three different statistical models (fixed and random effects panel models and first difference models) on regional data from the Norwegian forest inventory on standing volume and official statistics on harvested volumes, interest rate and prices of sawlogs and pulpwood for the period 1996–2016. Statistically significant different price elasticities are found in 12 out of total 15 pairs of regions. The price elasticity was lower and the volume elasticity higher in the western region compared to the other regions. The first difference models are best with respect to specification tests. The use of region specific price elasticities gives slightly better fit for the panel data models than using a uniform price parameter. The results show that the econometric specification influence the parameter values, and it is thus complicated to directly compare results in different timber supply studies. Regional differences in timber supply are important to consider.

  • Rørstad, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: per.kristian.rorstad@nmbu.no (email)
  • Solberg, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: birger.solberg@nmbu.no
  • Trømborg, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: erik.tromborg@nmbu.no
article id 10662, category Research article
Kari T. Korhonen, Arto Ahola, Juha Heikkinen, Helena M. Henttonen, Juha-Pekka Hotanen, Antti Ihalainen, Markus Melin, Juho Pitkänen, Minna Räty, Maria Sirviö, Mikael Strandström. (2021). Forests of Finland 2014–2018 and their development 1921–2018. Silva Fennica vol. 55 no. 5 article id 10662. https://doi.org/10.14214/sf.10662
Keywords: biodiversity; National Forest Inventory; growing stock; forest resources; forest damage
Highlights: Current volume of growing stock, 2500 M m3, is 1.7 times the volume in the 1920s; Annual volume increment is 107.8 M m3, which is double the increment estimated in the 1930s; Serious damage is observed on 2% of the forests available for wood supply; The amount of dead wood is on average 5.8 m3 per ha on productive forest.
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We describe the methodology applied in the 12th national forest inventory of Finland (NFI12) and describe the state of Finland’s forests as well as the development of some key parameters since 1920s. According to the NFI12, the area of forestry land (consisting of productive and poorly productive forest, unproductive land, and other forestry land) is 26.2 M ha. The area of forestry land has decreased from 1920s to 1960s due to expansion of agriculture and built-up land. 20% of the forestry land is not available for wood supply and 13% is only partly available for wood supply. The area of peatlands is 8.8 M ha, which is one third of the forestry land. 53% of the current area of peatlands is drained. The volume of growing stock, 2500 M m3, is 1.7 times the volume estimated in NFI1 in the 1920s for the current territory of Finland. The estimated annual volume increment is 107.8 M m3. The increment estimate has doubled since the estimate of NFI2 implemented in late 1930s. The annual mortality is estimated to 7 M m3, which is 0.5 M m3 more than according to the previous inventory. Serious or complete damage was observed on 2% of the productive forest available for wood supply. The amount of dead wood is on average 5.8 m3 ha–1 in productive forests. Since the NFI9 (1996–2003) the amount of dead wood has increased in South Finland and decreased in North Finland both in protected forests and forests available for wood supply (FAWS). The area of natural or almost natural forests on productive forest is 380 000 ha, out of this, 42 000 ha are in FAWS and 340 000 ha in protected forests.

  • Korhonen, Natural Resources Institute Finland (Luke), P.O. Box 68, FI-80100 Joensuu, Finland E-mail: kari.t.korhonen@luke.fi (email)
  • Ahola, Natural Resources Institute Finland (Luke), P.O. Box 2, FI-00790, Helsinki, Finland E-mail: arto.ahola@luke.fi
  • Heikkinen, Natural Resources Institute Finland (Luke), P.O. Box 2, FI-00790, Helsinki, Finland E-mail: juha.heikkinen@luke.fi
  • Henttonen, Natural Resources Institute Finland (Luke), P.O. Box 2, FI-00790, Helsinki, Finland E-mail: helena.henttonen@luke.fi
  • Hotanen, Natural Resources Institute Finland (Luke), P.O. Box 68, FI-80100 Joensuu, Finland E-mail: juha-pekka.hotanen@luke.fi
  • Ihalainen, Natural Resources Institute Finland (Luke), P.O. Box 2, FI-00790, Helsinki, Finland E-mail: anttivj.ihalainen@elisanet.fi
  • Melin, Natural Resources Institute Finland (Luke), P.O. Box 68, FI-80100 Joensuu, Finland E-mail: markus.melin@luke.fi
  • Pitkänen, Natural Resources Institute Finland (Luke), P.O. Box 68, FI-80100 Joensuu, Finland E-mail: juho.pitkanen@luke.fi
  • Räty, Natural Resources Institute Finland (Luke), P.O. Box 2, FI-00790, Helsinki, Finland E-mail: minna.raty@luke.fi
  • Sirviö, Natural Resources Institute Finland (Luke), P.O. Box 2, FI-00790, Helsinki, Finland E-mail: maria.sirvio@uudenmaanliitto.fi
  • Strandström, Natural Resources Institute Finland (Luke), P.O. Box 2, FI-00790, Helsinki, Finland E-mail: mikael.strandstrom@luke.fi
article id 10612, category Research article
Daesung Lee, Jouni Siipilehto, Jari Hynynen. (2021). Models for diameter distribution and tree height in hybrid aspen plantations in southern Finland. Silva Fennica vol. 55 no. 5 article id 10612. https://doi.org/10.14214/sf.10612
Keywords: Näslund’s height curve; Weibull distribution; parameter recovery; Populus tremula × P. tremuloides; clonal plantation; nonlinear mixed-effects model
Highlights: Parameter recovery method for the Weibull function fitted diameter distributions well by means of sum and mean forest stand characteristics for hybrid aspen plantations; Arithmetic and weighted mean diameters performed better for the recovery method than the corresponding median diameters; Two alternative Näslund’s height curve models with stand characteristics and tree dbh predictors provided unbiased tree height predictions.
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Hybrid aspen (Populus tremula L. × P. tremuloides Michx.) is known with outstanding growth rate and some favourable wood characteristics, but models for stand management have not yet been prepared in northern Europe. This study introduces methods and models to predict tree dimensions, diameter at breast height (dbh) and tree height for a hybrid aspen plantation using data from repeatedly measured permanent sample plots established in clonal plantations in southern Finland. Dbh distributions using parameter recovery method for the Weibull function was used with Näslund’s height curve to model tree heights. According to the goodness-of-fit statistics of Kolmogorov-Smirnov and the Error Index, the arithmetic mean diameter (D) and basal area-weighted mean diameter (DG) provided more stable parameter recovery for the Weibull distribution than the median diameter (DM) and basal area-weighted median diameter (DGM), while DG showed the best overall fit. Thus, Näslund’s height curve was modelled using DG with Lorey’s height (HG), age, basal area (BA), and tree dbh (Model 1). Also, Model 2 was tested using all predictors of Model 1 with the number of trees per ha (TPH). All predictors were shown to be significant in both Models, showing slightly different behaviour. Model 1 was sensitive to the mean characteristics, DG and HG, while Model 2 was sensitive to stand density, including both BA and TPH as predictors. Model 1 was considered more reasonable to apply based on our results. Consequently, the parameter recovery method using DG and Näslund’s models were applicable for predicting tree diameter and height.

  • Lee, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0003-1586-9385 E-mail: daesung.lee@luke.fi (email)
  • Siipilehto, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: jouni.siipilehto@luke.fi
  • Hynynen, Natural Resources Institute Finland (Luke), Natural resources, Vipusenkuja 5, FI-57200 Savonlinna, Finland ORCID https://orcid.org/0000-0002-9132-8612 E-mail: jari.hynynen@luke.fi
article id 10605, category Research article
Charlotta Harju, Katja Lähtinen. (2021). Perceptions of wooden interior product quality – insights on sustainability views among Finnish consumers. Silva Fennica vol. 55 no. 5 article id 10605. https://doi.org/10.14214/sf.10605
Keywords: wood products; consumer behavior; perceived quality; sociodemographic variables; sustainable consumption
Highlights: The quality indicators of wooden interior products can be grouped into four factors relating to products’ environmental friendliness, fit with lifestyle and home design, visual and tactile attractiveness, and technical solidity, which are in multiple ways connected with sustainability; The sociodemographic background of the respondents was found to be linked with consumer scores for those factors; Wooden interior products should be designed to meet different types of product quality expectations in the home environment.
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Evaluation of product attributes and the overall quality significantly affect consumer purchasing decisions. Previous studies on wooden products have mostly addressed wood product quality from technical viewpoints, while largely disregarding environmental, social, and economic aspects in the assessments. Therefore, knowledge on how sustainability aspects are evaluated as a feature of wood product quality is narrow. This study investigated consumer perceptions of different quality indicators (i.e., quality cues and attributes) of wooden interior products with a special focus on sustainability and value chain phases. In addition, the connections between consumers’ sociodemographic background and their perceptions of the quality features of wooden interior products were evaluated. The material of the study was based on data gathered in 2018 with a postal survey sent to 1000 people living in Finland with a response rate of 25.6%. As methods of analysis, exploratory factor analysis, the Mann-Whitney U test, and the Kruskall-Wallis test were utilized. The results show that the quality indicators of wooden interior products can be grouped into four factors relating to products’ environmental friendliness, fit with lifestyle and home design, visual and tactile attractiveness, and technical solidity, which are in multiple ways connected with sustainability. The sociodemographic background of the respondents was found to be linked with consumer scores for those factors. Engaging consumers in sustainable consumption choices requires providing them with information on wooden product value chains that meets their individual needs in relation to their existing knowledge of those issues and individual values.
 

  • Harju, University of Vaasa, School of Marketing and Communication, P.O. Box 700, FI-65200 Vaasa, Finland ORCID https://orcid.org/0000-0002-0432-2598 E-mail: charlotta.harju@uwasa.fi (email)
  • Lähtinen, Natural Resources Institute Finland (Luke), Bioeconomy and Environment Unit, P.O. Box 2, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0001-6260-5062 E-mail: katja.lahtinen@luke.fi
article id 10573, category Research article
Jari Miina, Inka Bohlin, Torgny Lind, Jonas Dahlgren, Kari Härkönen, Tuula Packalen, Anne Tolvanen. (2021). Lessons learned from assessing the cover and yield of bilberry and lingonberry using the national forest inventories in Finland and Sweden. Silva Fennica vol. 55 no. 5 article id 10573. https://doi.org/10.14214/sf.10573
Keywords: forest management planning; berry models; field measurements; predictions
Highlights: Model-based predictions of the berry yields of an average crop year are produced using the Finnish National Forest Inventory (NFI); Inventory-based estimates of seasonal berry yields are produced using the Swedish NFI observations; The inventory-based method provides seasonal estimates, whereas models can be utilised to integrate vegetation cover and berry yields in numerical multi-objective forest planning.
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Bilberry (Vaccinium myrtillus L.) and lingonberry (V. vitis-idaea L.) can be a part of healthy diet and are important for many animals. Two approaches are described to assessing their vegetation cover and berry yield via national forest inventory (NFI) observations. The aim was to provide estimates and predictions of the abundance and yield of the species at regional and national levels in Finland and Sweden. In Finland, the model-based predictions are used in evaluating the impacts of cutting intensity on forest berries needed in forest-related decision making. In Sweden, seasonal inventory-based estimates are used to evaluate the annual national and regional berry yields, and in a forecasting system aimed at large public and berry enterprises. Based on the NFI sample plots measured between 2014 and 2018, the total annual yields are estimated to be 208 Mkg of bilberry and 246 Mkg of lingonberry on productive forest land (increment at least 1 m3 ha–1 year–1) in Finland, and 336 and 382 Mkg respectively in Sweden (average of NFI inventories in 2015–2019). The predicted development of berry yields is related to the intensity of cuttings in alternative forest management scenarios: lower removals favoured bilberry, and higher removals lingonberry. The model-based method describes the effects of stand development and management on berry yields, whereas the inventory-based method can calibrate seasonal estimates through field observations. In providing spatially and timely more accurate information concerning seasonal berry yields, an assessment of berry yields should involve the elements of both inventory-based and model-based approaches described in this study.

  • Miina, Natural Resources Institute Finland (Luke), Yliopistokatu 6 B, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0002-8639-4383 E-mail: jari.miina@luke.fi (email)
  • Bohlin, Swedish University of Agricultural sciences (SLU), Department of Forest Resource Management, Skogsmarksgränd, S-901 83 Umeå, Sweden E-mail: inka.bohlin@slu.se
  • Lind, Swedish University of Agricultural sciences (SLU), Department of Forest Resource Management, Skogsmarksgränd, S-901 83 Umeå, Sweden E-mail: torgny.lind@slu.se
  • Dahlgren, Swedish University of Agricultural sciences (SLU), Department of Forest Resource Management, Skogsmarksgränd, S-901 83 Umeå, Sweden E-mail: jonas.dahlgren@slu.se
  • Härkönen, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: kari.harkonen@luke.fi
  • Packalen, Natural Resources Institute Finland (Luke), Yliopistokatu 6 B, FI-80100 Joensuu, Finland; Ministry of Agriculture and Forestry of Finland, P.O. Box 30, FI-00023 Government, Finland E-mail: tuula.packalen@mmm.fi
  • Tolvanen, Natural Resources Institute Finland (Luke), Paavo Havaksentie 3, FI-90014 University of Oulu, Finland E-mail: anne.tolvanen@luke.fi
article id 10568, category Research article
Juha Kaitera, Leena Aarnio, Tiina Ylioja, Jouni Karhu. (2021). Naohidemyces vaccinii sporulates on wild species of ground flora in Finnish Norway spruce seed orchards but Thekopsora areolata does not on other species than Prunus. Silva Fennica vol. 55 no. 5 article id 10568. https://doi.org/10.14214/sf.10568
Keywords: Picea abies; cherry-spruce rust; epidemics; Prunus; blueberry rust; Naohidemyces vaccinii
Highlights: Cherry-spruce rust, Thekopsora areolata, was not found on any of the common species of ground vegetation in Finnish Norway spruce seed orchards; Blueberry rust, Naohidemyces vaccinii, was common on Vaccinium myrtillus and occasional on V. vitis-idaea in all seed orchards; Thekopsora areolata occurs only on Prunus in Finnish Norway spruce seed orchards.
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Thekopsora areolata (Fr.) Magnus is a serious cone pathogen that reduces seed crop of Picea abies (L.) Karst. and other Picea spp. Natural sporulation of T. areolata was investigated in nine Norway spruce seed orchards suffering from severe successive T. areolata epidemics in Finland. Habitats occupied by Vaccinium myrtillus L., V. vitis-idaea L., Empetrum nigrum L. and Calluna vulgaris (L.) Hull, and a number of other wild species belonging to ground flora were investigated for Thekopsora areolata uredinia 9–10 times in May–September 2018–2019. Occurrence of Thekopsora uredinia was estimated in current-year leaves of the plants in ca. 25 sample plots of 1 m2 in each seed orchard. A sample of plant leaves with rust uredinia or necrotic pustules were collected from each plot. No rust fruiting stages of T. areolata were found on any of the test species of ground flora. However, rust uredinia were observed regularly on leaves of V. myrtillus and V. vitis-idaea in all seed orchards between mid-July and the end of September. Rust sporulation started on V. myrtillus in July and on V. vitis-idaea in August. Based on symptoms, uredinia and spore morphology, the rust on both V. myrtillus and V. vitis-idaea was identified as blueberry rust, Naohidemyces vaccinii (Jørst.) S. Sato, Katsuya & Y. Hirats. ex Vanderwegen & Fraiture. The uredinial stage of the rust on Vaccinium spp. were described. No evidence of natural sporulation of T. areolata on wild plant species other than Prunus was observed in Finnish Norway spruce seed orchards.

  • Kaitera, Natural Resources Institute Finland (Luke), Natural Resources and Bioproduction, FI-90570 Oulu, Finland E-mail: juha.kaitera@luke.fi (email)
  • Aarnio, Natural Resources Institute Finland (Luke), Natural Resources and Bioproduction, FI-00790 Helsinki, Finland E-mail: leena.aarnio@luke.fi
  • Ylioja, Natural Resources Institute Finland (Luke), Natural Resources and Bioproduction, FI-00790 Helsinki, Finland E-mail: tiina.ylioja@luke.fi
  • Karhu, Natural Resources Institute Finland (Luke), Natural Resources and Bioproduction, FI-90570 Oulu, Finland E-mail: jouni.karhu@luke.fi
article id 10534, category Research article
Matti Haapanen, Seppo Ruotsalainen. (2021). Adaptive performance of genetically improved and unimproved seedlings of Scots pine. Silva Fennica vol. 55 no. 5 article id 10534. https://doi.org/10.14214/sf.10534
Keywords: Pinus sylvestris; cold hardiness; growth rhythm; seed orchards; bud set; freezing test; reproductive materials
Highlights: We studied the variation in adaptive traits in one-year-old seedlings of Scots pine representing different levels of genetic gain and geographical origins; All the adaptive traits analyzed showed clinal co-variation with the latitude of origin; Differences in adaptive performance between genetically improved and unimproved reproductive materials were mostly small and insignificant when the effect of the latitudinal origin was considered; First-generation seed orchard materials implied slightly poorer autumn frost hardiness compared to other materials, but the results were ambiguous.
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Our main objective was to determine whether various genetically improved reproductive materials of Scots pine (Pinus sylvestris L.) differ in growth rhythm, autumn cold acclimation and resilience from unimproved materials. The study consisted of two successive indoor experiments with Scots pine seedlings representing four levels of genetic gain (unimproved natural stands, first-generation seed orchards, 1.5-generation seed orchards and seed orchards established with freezing-tested parents) and a wide range of geographical origins within Finland. The seedlings were assessed for terminal shoot elongation, growth cessation, bud set, freezing injuries and bud flushing over the first growth period. All the adaptive traits showed a latitudinal trend regardless of the genetic level. Seed orchard progenies and natural stand progenies did not differ significantly in the timing of growth cessation, bud set, and the flushing rate of the frost-injured seedlings, after the trait variation was adjusted to the latitude of origin. The differences in autumn frost hardiness were insignificant, too, except for the somewhat higher injury rate displayed by the first-generation seed orchard materials. The finding was not conclusive due to ambiguous results from the two experiments. Overall, we did not find evidence of alarming compromises in the adaptive performance of genetically improved materials.

  • Haapanen, Natural Resources Institute Finland (Luke), Production systems, Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0003-3294-501X E-mail: matti.haapanen@luke.fi (email)
  • Ruotsalainen, Natural Resources Institute Finland (Luke), Production systems, Vipusenkuja 5, FI-57200 Savonlinna, Finland ORCID https://orcid.org/0000-0002-2547-0282 E-mail: seppo.ruotsalainen@luke.fi
article id 10525, category Research article
Markus Melin, Tiina Ylioja, Leena Aarnio, Katri Hamunen, Seppo Nevalainen, Antti Pouttu, Heli Viiri. (2021). Emergence levels of pine shoot beetles from roundwood piles of Scots pine and the cascading damage in the surrounding forests. Silva Fennica vol. 55 no. 5 article id 10525. https://doi.org/10.14214/sf.10525
Keywords: forest management; forestry; Tomicus piniperda; Tomicus minor; roundwood; legislation; timber storage
Highlights: Emerged pine shoot beetles were counted from piles of harvested Scots pine, and the shoot damage in the surrounding forests was measured; Damage was noticeable up to a distance of 40–60 m, being more severe near large piles; For piles smaller than 50 m3 the level of damage (fallen shoots) was mainly below known thresholds for growth losses; Logs with harvester-damaged bark were significantly less colonized by the beetles.
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Bark beetles are amongst the most aggressive pest agents of coniferous forests. Due to this, many boreal countries have designated laws aiming to lower the risk of bark beetle epidemics. Finland’s forest legislation has pre-emptive measures targeted against bark beetles, and for Scots pine (Pinus sylvestris L.), the law concerns pine shoot beetles (Tomicus spp.). This study used data collected around 25 piles of Scots pine roundwood that were harvested in the winter but left in the forest until the following November. Thus, the pine shoot beetles were able to use the piles for breeding. We assessed the number of emerged insects from the piles and the cascading damage they caused in the surrounding forests. All roundwood piles, regardless of their volume, were used by the beetles for breeding. Highest densities of beetle exit holes were found from the parts of the log with thick and intact bark. If the bark of the log was damaged by the harvester head, the number of beetles decreased significantly. Depending on the volume of the roundwood pile, the cascading damage (fallen shoots) was noticeable up to ca. 40–60 m from the roundwood pile. Storing of piles smaller than 50 m3 did not cause excess damage. The number of fallen shoots per tree was generally below the known thresholds for when growth losses can occur. However, the study was conducted in mature forests, and it can be assumed that the recorded damage levels would severely affect the growth of young pines, raising the question of where to store the roundwood. As with other bark beetles, the role of Tomicus beetles as damage agents may change in the future, but based on this as well as past studies, the species can be viewed as a notable damage agents only around long-term wood storage sites in the current northern conditions.

  • Melin, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Yliopistokatu 6b, FI-80100 Joensuu, Finland E-mail: markus.melin@luke.fi (email)
  • Ylioja, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: tiina.ylioja@luke.fi
  • Aarnio, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: leena.aarnio@luke.fi
  • Hamunen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Yliopistokatu 6b, FI-80100 Joensuu, Finland E-mail: katri.hamunen@luke.fi
  • Nevalainen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Yliopistokatu 6b, FI-80100 Joensuu, Finland E-mail: seppo.nevalainen@gmail.com
  • Pouttu, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: antti.pouttu@kolumbus.fi
  • Viiri, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Yliopistokatu 6b, FI-80100 Joensuu, Finland; UPM-Kymmene Oyj, UPM Forest, Peltokatu 26 C, PL 85, FI-33100, Tampere, Finland E-mail: heli.viiri@upm.com
article id 10619, category Research article
Yrjö Nuutinen. (2021). Corrigendum: Comparing the characteristics of boom-corridor and selectively thinned stands of Scots pine and birch. Silva Fennica vol. 55 no. 4 article id 10619. https://doi.org/10.14214/sf.10619
Abstract | Full text in HTML | Full text in PDF | Author Info

Corrigendum

  • Nuutinen, Natural Resources Institute Finland (Luke), Production systems, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: yrjo.nuutinen@luke.fi (email)
article id 10580, category Research article
Tomi Kaakkurivaara, Nopparat Kaakkurivaara. (2021). Cost-efficiency and ergonomic study of two methods for planting Eucalyptus spp. seedlings in plantation forestry. Silva Fennica vol. 55 no. 4 article id 10580. https://doi.org/10.14214/sf.10580
Keywords: productivity; time study; planting stick; planting tube; REBA; work load; work sampling
Highlights: We present a first assessment of the planting stick method as used in Thailand for planting Eucalyptus spp. seedlings in plantations in terms of time, cost and ergonomic parameters; Parallel analysis shows that the planting tube method may be superior for planting Eucalyptus spp. seedlings
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In Thailand and various other countries tree seedlings are generally planted using simple manual tools, often a ‘planting stick’, but the method requires time-consuming, labour-intensive teamwork. However, use of a ‘planting tube’ allows a single person to perform both the preparation and planting work. Thus, in a classical time study and ergonomic survey we compared the productivity, cost-effectiveness, and ergonomic impact of planting Eucalyptus spp. seedlings using the two tools at the same planting site in Western Thailand. The planting tube method proved to be more productive, more cost-efficient, and less burdensome than the planting stick method (with time and cost requirements of 21 s and €0.0061 per seedling, versus 16.6 s and €0.0463 per seedling, respectively). In conclusion, the planting tube method may be a viable alternative to reduce costs and increase productivity, while maintaining reasonable workloads for the workers, despite the higher purchase price of planting tubes.

  • Kaakkurivaara, Department of Forest Engineering, Faculty of Forestry, Kasetsart University, 50 Phahonyothin Rd, Lat Yao, Chatuchak, Bangkok 10900, Thailand E-mail: tomi.kaakkurivaara@gmail.com (email)
  • Kaakkurivaara, Department of Forest Engineering, Faculty of Forestry, Kasetsart University, 50 Phahonyothin Rd, Lat Yao, Chatuchak, Bangkok 10900, Thailand E-mail: ffornrm@ku.ac.th
article id 10539, category Research article
Jaakko Repola, Juha Heikkinen, Jari Lindblad. (2021). Pulpwood green density prediction models and sampling-based calibration. Silva Fennica vol. 55 no. 4 article id 10539. https://doi.org/10.14214/sf.10539
Keywords: pulpwood; conversion factor; green density
Highlights: The developed models provided a realistic description of the observed seasonal variation in pulpwood green density; The model predictions were more reliable than those obtained with current practices.
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Pulpwood arriving at the mills is mainly measured by weighing. In the loading phase of forwarding and trucking, timber is weighed using scales mounted in the grapple loader. The measured weight of timber is converted into volume using a conversion factor defined as green density (kg m–3). At the mill, the green density factor is determined by sampling measurements, while in connection with weighing with grapple-mounted scales during transportation, fixed green density factors are used. In this study, we developed predictive regression models for the green density of pulpwood. The models were constructed separately by pulpwood assortments: pine (contains mainly Pinus sylvestris L); spruce (mainly Picea abies (L.) Karst.); decayed spruce; birch (mainly Betula pubescens Ehrh. and Betula pendula Roth); and aspen (mainly Populus tremula L.). Study material was composed of the sampling-based measurements at the mills between 2013–2019. The models were specified as linear mixed models with both fixed and random parameters. The fixed effect produced the expected value of green density as a function of delivery week, storage time, and meteorological conditions during storage. The random effects allowed the model calibration by utilizing the previous sampling weight measurements. The model validation showed that the model predictions faithfully reproduced the observed seasonal variation in green density. They were more reliable than those obtained with the current practices. Even the uncalibrated (fixed) predictions had lower relative root mean squared prediction errors than those obtained with the current practices.

  • Repola, Natural Resources Institute Finland (Luke), Natural resources, Ounasjoentie 6, 96200 Rovaniemi, Finland E-mail: jaakko.repola@luke.fi (email)
  • Heikkinen, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: juha.heikkinen@luke.fi
  • Lindblad, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, 00790 Helsinki, Finland E-mail: jari.lindblad@luke.fi
article id 10528, category Research article
Jaana Luoranen, Heli Viiri. (2021). Comparison of the planting success and risks of pine weevil damage on mineral soil and drained peatland sites three years after planting. Silva Fennica vol. 55 no. 4 article id 10528. https://doi.org/10.14214/sf.10528
Keywords: survival; planting; mechanical site preparation; ground vegetation; forest regeneration; Hylobius abietis
Highlights: The planting success was poorer on peatland sites than on mineral soil; Dense ground vegetation cover is more probable on peatland than on mineral soil; No differences in pine weevil feeding damage on mineral soil and peatland were found; Cultivated mineral soil reduced the vegetation cover, feeding damage and seedling mortality.
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Over 20% of regeneration operations will be on drained peatland in the next decade in Finland. There are only a few studies comparing the planting success and the risk of pine weevil (Hylobius abetis (L.) feeding damage on mineral soil and drained peatland. Thirty sites planted with Norway spruce (Picea abies (L.) H. Karst.) container seedlings in 2009 in Southern and Central Finland were inventoried three growing seasons after planting. Prediction models for the probability of survival, pine weevil damage and the presence of ground vegetation cover were done separately for peatland and mineral soil sites. The planting success was 17% lower on peatland sites (1379 surviving seedlings ha–1) than on mineral soil (1654 seedlings ha–1). The factors explaining the survival were the ground vegetation cover and type of the planting spot on the peatland, and the ground vegetation cover on mineral soil. On mineral soil, 76% of the planting spots were on cultivated mineral soil while on peatland only 28% of the seedlings were planted on similar spots. There were also fewer seedlings that were surrounded by dense ground vegetation on mineral soil (4%) than on peatland (14%). Pine weevil feeding damage did not differ significantly on peatland (23%) or mineral soil (18%). The more time there was from clear-cutting, the more the probability of pine weevil feeding damage was reduced on both soil classes. Additionally, cover vegetation in the vicinity of the seedlings increased on mineral soil. Cultivated planting spots, especially those covered by mineral soil, prevented pine weevil feeding and reduced the harmful effects of vegetation on the seedlings both on mineral soil and peatland.

article id 10515, category Research article
Alwin A. Hardenbol, Anton Kuzmin, Lauri Korhonen, Pasi Korpelainen, Timo Kumpula, Matti Maltamo, Jari Kouki. (2021). Detection of aspen in conifer-dominated boreal forests with seasonal multispectral drone image point clouds. Silva Fennica vol. 55 no. 4 article id 10515. https://doi.org/10.14214/sf.10515
Keywords: Populus tremula; deciduous trees; mixed forest; protected areas; tree species classification; unmanned aerial vehicles
Highlights: Four boreal tree species (Scots pine, Norway spruce, birches and European aspen) classified with an overall accuracy of 95%; Presence of European aspen detected with excellent accuracy (UA: 97%, PA: 96%); Late spring is the best time for species classification by remote sensing; Best time to separate aspen from birch was when birch had leaves, but aspen did not.
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Current remote sensing methods can provide detailed tree species classification in boreal forests. However, classification studies have so far focused on the dominant tree species, with few studies on less frequent but ecologically important species. We aimed to separate European aspen (Populus tremula L.), a biodiversity-supporting tree species, from the more common species in European boreal forests (Pinus sylvestris L., Picea abies [L.] Karst., Betula spp.). Using multispectral drone images collected on five dates throughout one thermal growing season (May–September), we tested the optimal season for the acquisition of mono-temporal data. These images were collected from a mature, unmanaged forest. After conversion into photogrammetric point clouds, we segmented crowns manually and automatically and classified the species by linear discriminant analysis. The highest overall classification accuracy (95%) for the four species as well as the highest classification accuracy for aspen specifically (user’s accuracy of 97% and a producer’s accuracy of 96%) were obtained at the beginning of the thermal growing season (13 May) by manual segmentation. On 13 May, aspen had no leaves yet, unlike birches. In contrast, the lowest classification accuracy was achieved on 27 September during the autumn senescence period. This is potentially caused by high intraspecific variation in aspen autumn coloration but may also be related to our date of acquisition. Our findings indicate that multispectral drone images collected in spring can be used to locate and classify less frequent tree species highly accurately. The temporal variation in leaf and canopy appearance can alter the detection accuracy considerably.

  • Hardenbol, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0002-0615-505X E-mail: alwin.hardenbol@uef.fi (email)
  • Kuzmin, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland; University of Eastern Finland, Department of Geographical and Historical Studies, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: anton.kuzmin@uef.fi
  • Korhonen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: lauri.korhonen@uef.fi
  • Korpelainen, University of Eastern Finland, Department of Geographical and Historical Studies, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: pasi.korpelainen@uef.fi
  • Kumpula, University of Eastern Finland, Department of Geographical and Historical Studies, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: timo.kumpula@uef.fi
  • Maltamo, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: matti.maltamo@uef.fi
  • Kouki, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: jari.kouki@uef.fi
article id 10507, category Research article
Karri Uotila, Timo Saksa. (2021). Cost-efficient pre-commercial thinning: effects of method and season of early cleaning. Silva Fennica vol. 55 no. 4 article id 10507. https://doi.org/10.14214/sf.10507
Keywords: productivity; young stand management; cost-effectiveness; release treatment
Highlights: The first stage of multistage pre-commercial thinning, early cleaning, took 27–30% less time when carried out in the spring instead of the summer; The two stage pre-commercial thinning program was 11% less expensive to apply when early cleaning had been applied in the spring instead of the summer.
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This study’s aim was to identify how the application season and the method of early cleaning (EC), the first stage of multistage pre-commercial thinning (PCT), affected the time consumption in EC and in the subsequent second PCT operation. The worktime in EC was recorded in the spring, summer, and autumn in 22 sites, which were either totally cleaned or point cleaned. Later, these sites were measured at the time of the second PCT. Time consumption was estimated in PCT, based on the removal of the sites. The time consumption in EC was 5.3 productive work hours (pwh) ha–1, 7.3 pwh ha–1, and 6.2 pwh ha–1 respectively in the spring, summer, and autumn. EC in the spring instead of the summer saved 27–30% of working time, depending on the cleaning method. Point cleaning was 0.8 pwh ha–1 quicker than total cleaning, but the difference was statistically insignificant. The second stage, PCT, was 1 pwh ha–1 slower to conduct in sites which had been early cleaned in the spring instead of the summer. However, at the entire management program level, EC applied in the spring or autumn instead of the summer saved 11% or 5% respectively of the total discounted costs (3% interest rate) of multistage pre-commercial thinning. Today, the commonest time to conduct EC is in the summer, which was the most expensive of the analyzed management alternatives here. We can expect savings in juvenile stand management in forestry throughout boreal conifer forests by rethinking the seasonal workforce allocation.

  • Uotila, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: karri.uotila@luke.fi (email)
  • Saksa, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@luke.fi
article id 10485, category Research article
Felicia Dahlgren Lidman, Emma Holmström, Tomas Lundmark, Nils Fahlvik. (2021). Management of spontaneously regenerated mixed stands of birch and Norway spruce in Sweden. Silva Fennica vol. 55 no. 4 article id 10485. https://doi.org/10.14214/sf.10485
Keywords: Betula pendula; Picea abies; Betula pubescens; natural regeneration; mixed forest; land expectation value
Highlights: The absence of forest management does not always mean economic loss; With dense spontaneous regeneration of birch and Norway spruce, the first competition release can have a high impact on future stem development; Significantly different effects on stand volume production and diameter development of Norway spruce can be expected with different biomass harvest strategies.
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Timber production and profitability were evaluated for spontaneously-regenerated mixtures on two formerly clearcut areas. The abandoned areas developed into birch-dominated (Betula pendula Roth and Betula pubescens Ehrh.) stands with successional ingrowth of Norway spruce (Picea abies (L.) H. Karst.). An experiment with randomized treatments within blocks was established, using three management strategies and one unthinned control, resulting in variation in optimal rotation age, merchantable volume and species composition. The management strategies were evaluated based on total production (volume) by using measured growth data 42 years after clearcutting and the modelled future stand development. The long-term effects of spontaneous regeneration and management strategies were evaluated based on land expectation value (LEV) and compared with a fifth management strategy using artificial regeneration and intense thinnings. 12 years after treatment, at a stand age of 42 years, the unthinned control had produced the highest total stem volume. At interest rates of 2% or higher, the unmanaged forest was an economically viable strategy, even compared to an intensive management strategy with a preferred merchantable timber species. Interest rates clearly impacted the profitability of the different management strategies. This study shows that when spontaneous regeneration is successful and dense, the first competition release can have a high impact on the development of future crop trees and on the species mixture.

article id 10449, category Research article
Emanuel Strieder, Sonja Vospernik. (2021). Intra-annual diameter growth variation of six common European tree species in pure and mixed stands. Silva Fennica vol. 55 no. 4 article id 10449. https://doi.org/10.14214/sf.10449
Keywords: growing season; climate effects; dendrometer; logistic model; mixing effect; seasonal growth; social tree position
Highlights: 244 intra-annual growth patterns of six tree species on five sites in mixed and corresponding pure stands were analyzed; Humid sites showed a longer growing season than dry sites; Deciduous species showed an earlier growth culmination than conifer species; Mixture effects were both positive and negative and clearly differed between years, indicating that climate alters mixture effects.
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Studies of intra-annual growth are particularly useful for understanding tree growth because of their high temporal resolution. This study was performed in Austria and included hourly band dendrometer data of 244 annual tree recordings from six tree species (Picea abies (L.) Karst., Pinus sylvestris L., Larix decidua Mill., Abies alba Mill., Fagus sylvatica L., Quercus spp. (Quercus petraea (Matt.) Liebl., Quercus robur L.) sampled on five sites with contrasting site conditions in pure and mixed stands and on trees of different social position. Measurements encompassed 1–7 years. Cumulative diameter increment was modelled by logistic mixed-effects models with random effects at the tree and year level. The results showed large differences in seasonal growth patterns between sites, with a clearly shorter growing season at the drier sites. Species specific response on dry sites could be linked to drought characteristics, whereas response on more humid sites was related to light requirements or successional status. The deciduous trees showed earlier growth culmination and shorter growing periods than the evergreen species. Individual tree growth of Quercus spp., P. abies, and F. sylvatica was positively affected by mixture whereas L. decidua, P. sylvestris and A. alba showed no or adverse mixture effects. Mixture effects differed between years and social position. Furthermore, increment culmination was earlier in mixed stands, but shifts were minor. Tree growth differed by social position with dominant trees showing the largest increment and the longest growth duration, with shifts in tree growth patterns due to social position being as large as those between different sites.

  • Strieder, Boku, University of Natural Resources and Life Sciences Vienna, Department of Forest- and Soil Sciences, Institute of Forest Growth, Peter-Jordan-Str. 82, A-1190 Vienna, Austria ORCID https://orcid.org/0000-0001-6398-8536 E-mail: emanuel.strieder@students.boku.ac.at
  • Vospernik, Boku, University of Natural Resources and Life Sciences Vienna, Department of Forest- and Soil Sciences, Institute of Forest Growth, Peter-Jordan-Str. 82, A-1190 Vienna, Austria ORCID http://orcid.org/0000-0002-4201-6444 E-mail: sonja.vospernik@boku.ac.at (email)
article id 10441, category Research article
Dipak Mahatara, Amul K. Acharya, Bishnu P. Dhakal, Dipesh K. Sharma, Sunita Ulak, Prashant Paudel. (2021). Maxent modelling for habitat suitability of vulnerable tree Dalbergia latifolia in Nepal. Silva Fennica vol. 55 no. 4 article id 10441. https://doi.org/10.14214/sf.10441
Keywords: rosewood; satisal; distribution mapping; environmental variables; Maxent modelling
Highlights: Dalbergia latifolia is a vulnerable species of Nepal with very few conservation priorities; Habitat suitability modelling for this species is essential to endorse different conservation interventions; 43 presence locations and different environmental variables were retained for model prediction in Maxent; Province 2 was found most suitable habitat for the growth of D. latifolia, with western most province as unsuitable.
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Dalbergia latifolia Roxb., commonly known as rosewood, is one of the highly valuable tropical timber species of Nepal. The tree species was widely distributed in the past, however, over-exploitation of natural habitat, deforestation, forest conversion for agriculture, illegal logging and the invasion of alien species resulted in the classification of this species as vulnerable by the IUCN (International Union for Conservation of Nature) category. So, the prediction of habitat suitability and potential distribution of the species is required to develop restoration mechanisms and conservation interventions. In this study, we modelled the suitable habitat of D. latifolia over the entire possible range of Nepal using a Maxent model. We compiled 23 environmental variables (19 bioclimatic, 3 topographic and a vegetative layer), however, only 12 least correlated variables along with 43 spatially representative presence locations were retained for model prediction. We used a receiver operating characteristic (ROC) curve to assess the model’s performance and a Jackknife procedure to evaluate the relative importance of predictor variables. The model was statistically significant with an area under the curve (AUC) value of 0.969. The internal Jackknife test indicated that elevation was the most important variable for the model prediction with 71.3% contribution followed by mean temperature of driest quarter (9.8%). The most (>0.6) suitable habitat for the D. latifolia was 235 484 hectares with large sections of area in two provinces whereas, the western most provinces were not suitable for D. latifolia as per Maxent model. The information presented here can provide a framework for nature conservation planning, monitoring and habitat management of this rare and endangered species.

  • Mahatara, Forest Research and Training Centre, Government of Nepal, P.O. Box 3339, Babarmahal, Kathmandu 44600, Nepal E-mail: honeystar73@gmail.com (email)
  • Acharya, Forest Research and Training Centre, Government of Nepal, P.O. Box 3339, Babarmahal, Kathmandu 44600, Nepal E-mail: acharya.amulkumar@gmail.com
  • Dhakal, Forest Research and Training Centre, Government of Nepal, P.O. Box 3339, Babarmahal, Kathmandu 44600, Nepal E-mail: dhakalbp.shorea@gmail.com
  • Sharma, Forest Research and Training Centre, Government of Nepal, P.O. Box 3339, Babarmahal, Kathmandu 44600, Nepal E-mail: dipeshsharmadiyu2015@gmail.com
  • Ulak, Forest Research and Training Centre, Government of Nepal, P.O. Box 3339, Babarmahal, Kathmandu 44600, Nepal E-mail: sunitaulak@gmail.com
  • Paudel, Agriculture Forestry University, P.O. Box 13712 Rampur, Chitwan, Nepal E-mail: prashant.paudel88@gmail.com
article id 10244, category Research article
Hans Ole Ørka, Endre H. Hansen, Michele Dalponte, Terje Gobakken, Erik Næsset. (2021). Large-area inventory of species composition using airborne laser scanning and hyperspectral data. Silva Fennica vol. 55 no. 4 article id 10244. https://doi.org/10.14214/sf.10244
Keywords: airborne laser scanning; Dirichlet regression; hyperspectral; species proportions; species-specific forest inventory
Highlights: A methodology for using hyperspectral data in the area-based approach is presented; Hyperspectral data produced satisfactory results for species composition in 90% of the cases; Parametric Dirichlet regression is an applicable method to predicting species proportions; Normalization and a tree-based selection of pixels provided the overall best results; Both visible to near-infrared and shortwave-infrared sensors gave acceptable results.
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Tree species composition is an essential attribute in stand-level forest management inventories and remotely sensed data might be useful for its estimation. Previous studies on this topic have had several operational drawbacks, e.g., performance studied at a small scale and at a single tree-level with large fieldwork costs. The current study presents the results from a large-area inventory providing species composition following an operational area-based approach. The study utilizes a combination of airborne laser scanning and hyperspectral data and 97 field sample plots of 250 m2 collected over 350 km2 of productive forest in Norway. The results show that, with the availability of hyperspectral data, species-specific volume proportions can be provided in operational forest management inventories with acceptable results in 90% of the cases at the plot level. Dominant species were classified with an overall accuracy of 91% and a kappa-value of 0.73. Species-specific volumes were estimated with relative root mean square differences of 34%, 87%, and 102% for Norway spruce (Picea abies (L.) Karst.), Scots pine (Pinus sylvestris L.), and deciduous species, respectively. A novel tree-based approach for selecting pixels improved the results compared to a traditional approach based on the normalized difference vegetation index.

  • Ørka, Norwegian University of Life Sciences, Faculty of Environmental Sciences and Natural Resource Management, P.O. Box 5003, NO-1432 Ås, Norway ORCID https://orcid.org/0000-0002-7492-8608 E-mail: hans-ole.orka@nmbu.no (email)
  • Hansen, Norwegian University of Life Sciences, Faculty of Environmental Sciences and Natural Resource Management, P.O. Box 5003, NO-1432 Ås, Norway; Norwegian Forest Extension Institute, Honnevegen 60, NO-2836 Biri, Norway ORCID https://orcid.org/0000-0001-5174-4497 E-mail: eh@skogkurs.no
  • Dalponte, Department of Sustainable Agro-ecosystems and Bioresources, Research and Innovation Centre, Fondazione E. Mach, Via E. Mach 1, 38010 San Michele all’Adige, TN, Italy ORCID https://orcid.org/0000-0001-9850-8985 E-mail: michele.dalponte@fmach.it
  • Gobakken, Norwegian University of Life Sciences, Faculty of Environmental Sciences and Natural Resource Management, P.O. Box 5003, NO-1432 Ås, Norway ORCID https://orcid.org/0000-0001-5534-049X E-mail: terje.gobakken@nmbu.no
  • Næsset, Norwegian University of Life Sciences, Faculty of Environmental Sciences and Natural Resource Management, P.O. Box 5003, NO-1432 Ås, Norway E-mail: erik.naesset@nmbu.no
article id 10555, category Research article
Ferréol Berendt, Felix Wolfgramm, Tobias Cremer. (2021). Reliability of photo-optical measurements of log stack gross volume. Silva Fennica vol. 55 no. 3 article id 10555. https://doi.org/10.14214/sf.10555
Keywords: photogrammetry; timber trade; logistic; volume estimation; wood pile
Highlights: Gross volume estimations of larger log stacks showed a smaller deviation compared to those of piles with smaller volumes; Log stack quality affects estimation accuracy; The deviations between the manual section-wise estimations were of similar amplitude as those for the photo-optical estimations.
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In terms of assessing economic impact, one of the most important elements in the wood supply chain is the measurement of round wood. Besides the one-by-one measurement of logs, logs are often measured when stacked at the forest road. The gross stacked volume includes the volume of the wood, bark and airspace and is widely used for industrial wood assortments. The increasing international attention given to photo-optical measurement systems for portable devices is due to their simplicity of use and efficiency. The aim of this study was to compare the gross volumes of hardwood log stacks measured using one widespread photo-optical app with two manual section-wise volume estimations of log stacks based on the German framework agreement for timber trade (RVR). The manual volume estimations were done starting from the left (RVRleft) and right (RVRright) sides of the log stacks. The results showed an average deviation of the photo-optical gross volume estimation in comparison to the manual estimation of –2.09% (RVRleft) and –3.66% (RVRright) while the deviation between RVRleft and RVRright was +2.54%. However, the log stack gross volume had a highly significant effect on the deviation and better accuracy with smaller deviation were reached for larger log stacks. Moreover, results indicated that the gross volume estimations of higher quality log stacks were closer for the three analyzed methods compared to estimations of poor-quality log stacks.

  • Berendt, Department of Forest Utilization and Timber Markets, Eberswalde University for Sustainable Development, 16225 Eberswalde, Germany ORCID https://orcid.org/0000-0002-6285-7590 E-mail: ferreol.berendt@hnee.de (email)
  • Wolfgramm, Landesforst MV Anstalt des öffentlichen Rechts, Forstamt Billenhagen, 18182 Blankenhagen, Germany E-mail: felixwolfgramm@yahoo.de
  • Cremer, Department of Forest Utilization and Timber Markets, Eberswalde University for Sustainable Development, 16225 Eberswalde, Germany E-mail: tobias.cremer@hnee.de
article id 10545, category Research article
Karol Tomczak, Tomczak Arkadiusz, Bartłomiej Naskrent, Tomasz Jelonek. (2021). The radial gradient of moisture content of silver birch wood in different seasons. Silva Fennica vol. 55 no. 3 article id 10545. https://doi.org/10.14214/sf.10545
Keywords: birch; Betula pendula; wood properties; living trees; seasonal changes; trunk cross section
Highlights: Seasonal variation in moisture content is significant, the greatest moisture content of wood was recorded in winter, and the lowest in summer; The greatest moisture content on cross-section was observed near to the pith, and lower values near to the bark; From environmental perspective results of this study may have an impact for log transport planning, weight-scaling systems, lumber drying.
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Silver birch (Betula pendula Roth) is classified in diffuse-porous wood category. In this case structure of wood tissue is quite similar across whole cross-sectional area. The aim of this study was to analyse cross-section variability of moisture content (MC) of growing silver birch wood, significant hardwood species in Polish forests. Investigations were performed on 120 model trees. In the trunk of each model tree, an increment core was collected at breast height. Samples were collected of 30 different trees in four different seasons. The greatest MC was observed during winter, lowest MC in summer. Differences in MC were statistically significant only between winter versus spring, summer, and autumn. Distribution of MC on cross-section was similar in each season. The greatest average value was observed close to pith, then it was decreasing in bark direction. The greatest difference between observed in spring – 19.51% (p < 0.05) and lowest in autumn – 4.66%. Distribution of green density (GD) on cross section was inverse proportional to MC value. Variations in GD and MC are relevant for log transport planning, weight-scaling systems, lumber drying and dynamic assessment of stiffness. Therefore, from an environmental loss perspective, it is important to determine changes in MC and GD across the year.

  • Tomczak, Department of Forest Utilization, Faculty of Forestry and Wood Technology, Poznań University of Life Sciences, Wojska Polskiego 71A, 60-625 Poznań, Poland ORCID https://orcid.org/0000-0001-5192-0294 E-mail: karol.tomczak@up.poznan.pl (email)
  • Arkadiusz, Department of Forest Utilization, Faculty of Forestry and Wood Technology, Poznań University of Life Sciences, Wojska Polskiego 71A, 60-625 Poznań, Poland ORCID https://orcid.org/0000-0003-1140-8282 E-mail: arkadiusz.tomczak@up.poznan.pl
  • Naskrent, Department of Forest Utilization, Faculty of Forestry and Wood Technology, Poznań University of Life Sciences, Wojska Polskiego 71A, 60-625 Poznań, Poland ORCID https://orcid.org/0000-0002-0756-4162 E-mail: bartlomiej.naskrent@up.poznan.pl
  • Jelonek, Department of Forest Utilization, Faculty of Forestry and Wood Technology, Poznań University of Life Sciences, Wojska Polskiego 71A, 60-625 Poznań, Poland ORCID https://orcid.org/0000-0001-9558-9951 E-mail: tomasz.jelonek@up.poznan.pl
article id 10509, category Research article
Chuan Xie, Tianfeng Liu, Song Guo, Jian Peng, Zailiu Li. (2021). Effects of ultra-dry storage on seed germination and seedling growth of Handeliondendron bodinieri. Silva Fennica vol. 55 no. 3 article id 10509. https://doi.org/10.14214/sf.10509
Keywords: Handeliodendron bodinieri; priming treatment seed; ultra-dry treatment
Highlights: By reducing the moisture content of the seeds, Handeliodendron bodinieri seeds showed a strong tolerance; PEG played a protective role in the process of re-wetting ultra-dry seeds; Ultra-dry storage could promote Handeliodendron bodinieri seedling growth and root development.
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Handeliodendron bodinieri (H. Lév.) Rehder is a rare, endangered, and therefore, protected tree species native to China. However, there are serious limitations to the effective protection of the species, including a low seed germination-rate and difficult storage due to a high seed oil-content. Here, we evaluated the feasibility of ultra-dry seed storage and its effects on seedling growth. We used the silica gel method to prepare ultra-dry seeds with different moisture contents to find an optimal moisture content range (2.54%–4.77%). Ultra-dry treatment improved storability of H. bodinieri seeds. Furthermore, seeds with a moisture content of 4.77% stored at room temperature, and seeds with a moisture content of 3.97% stored at 4 °C yielded the best results. Priming with an appropriate concentration of polyethylene glycol had a certain repairing effect on ultra-dry stored seeds and improved seed vigor, with a two-day priming treatment with 20% polyethylene glycol having the best effect. Finally, compared with sand storage at 4 °C, ultra-dry storage promoted seedling growth and root development; furthermore, it alleviated storage damage to H. bodinieri seeds, promoted soluble sugar and soluble protein accumulation, and increased seedling nitrogen, phosphorus, and potassium uptake. Therefore, ultra-dry storage can be effectively used to preserve H. bodinieri seeds. Specifically, low-temperature storage of ultra-dry seeds with a moisture content of 3.97% enhanced H. bodinieri seed vigor, and seedling growth and development.

  • Xie, Forestry College, Guangxi University, Daxue E Rd., Xiangtang District, Nanning 530004, Guangxi, China; Huanggang forestry research institute, Huanggang 438000, Hubei, China E-mail: sherman99c@163.com
  • Liu, Forestry College, Guangxi University, Daxue E Rd., Xiangtang District, Nanning 530004, Guangxi, China E-mail: ltfltfll@163.com
  • Guo, Forestry College, Guangxi University, Daxue E Rd., Xiangtang District, Nanning 530004, Guangxi, China E-mail: guosong@gxu.edu.cn
  • Peng, Sanmengjiang forest farm of Guangxi, Liuzhou 545001, Guangxi, China E-mail: pj75481425@126.com
  • Li, Forestry College, Guangxi University, Daxue E Rd., Xiangtang District, Nanning 530004, Guangxi, China E-mail: lizailiu666@163.com (email)
article id 10506, category Research article
Mikko Tikkinen, Terhi Latvala, Tuija Aronen. (2021). Interest in vegetatively propagated Norway spruce materials – a survey among Finnish forest owners and professionals. Silva Fennica vol. 55 no. 3 article id 10506. https://doi.org/10.14214/sf.10506
Keywords: Picea abies; desired traits; intention of adoption; somatic embryogenesis; willingness to pay
Highlights: Survey among forest owners and professionals’ perceptions towards tree breeding and vegetative propagation yielded over 3000 responses; Most valued features in reforestation material were: improved resilience in changing climate, improved pest and pathogen resistance, and securing the species’ gene pool; Majority of respondents accepted reforestation with vegetatively propagated material to some extent; Willingness to pay for improved features was indicated.
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Forests and forestry will encounter several changes of unknown magnitude within the coming decades. In the Nordic, long rotations complicate any anticipation to the upcoming changes. Tree breeding can contribute to coping with these changes. The time span of implementing breeding results in practice may be shortened through vegetative propagation. Introducing vegetative propagation to forest regeneration may phase several challenges before adopted by the industry, some of which are related to perceptions about new technology. Firstly, private forest owners are in a key role in implementing the technology in practice; although they do not represent the overall public, they are the decision makers in their own estates regarding forestry and forest regeneration. Secondly, the professionals related to the production of forest regeneration material and plants from forest species are in a key role when it comes to practically introducing the new technology to the forest owners. In this survey, perceptions of forest owners and professionals towards tree breeding and vegetative propagation were investigated. Additionally, the respondents were asked which traits they considered important to be improved by breeding, and their willingness to pay for these improved traits. The respondents valued the most: improved pest and pathogen resistance, improved resilience of forest in changing climate, and securing the species’ gene pool. Responses indicated that forest owners would be willing to pay more for the improved traits in forest regeneration material. The current novel study provides a foundation to concern public awareness regarding tree breeding and vegetative propagation in the future.

  • Tikkinen, Natural Resources Institute Finland ORCID https://orcid.org/0000-0002-2368-8042 E-mail: mikko.tikkinen@luke.fi (email)
  • Latvala, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: terhi.latvala@luke.fi
  • Aronen, Natural Resources Institute Finland (Luke), Production systems, Vipusenkuja 5, FI-57200 Savonlinna, Finland E-mail: tuija.aronen@luke.fi
article id 10494, category Research article
Hannu Hökkä, Ari Laurén, Leena Stenberg, Samuli Launiainen, Kersti Leppä, Mika Nieminen. (2021). Defining guidelines for ditch depth in drained Scots pine dominated peatland forests. Silva Fennica vol. 55 no. 3 article id 10494. https://doi.org/10.14214/sf.10494
Keywords: Pinus sylvestris; forest management; peatland; drainage; ditch network maintenance; process model; water table
Highlights: Process-based hydrological model was applied to drained peatland forests representing a wide range of conditions in Finland; Ditch depth keeping the median July–August water table below 0.35 m was defined; Ditch depth depended on climatic conditions, stand volume, peat properties, and ditch spacing; Shallower ditches than recommended in practice proved to be sufficient in most situations.
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We used a process-based hydrological model SUSI to improve guidelines for ditch network maintenance (DNM) operations on drained peatland forests. SUSI takes daily weather data, ditch depth, strip width, peat properties, and forest stand characteristics as input and calculates daily water table depth (WTD) at different distances from ditch. The study focuses on Scots pine (Pinus sylvestris L.) dominated stands which are the most common subjects of DNM. Based on a literature survey, and consideration of the tradeoffs between forest growth and detrimental environmental impacts, long term median July–August WTD of 0.35 m was chosen as a target WTD. The results showed that ditch depths required to reach such WTD depends strongly on climatic locations, stand volume, ditch spacing, and peat thickness and type. In typical ditch cleaning areas in Finland with parallel ditches placed about 40 m apart and tree stand volumes exceeding 45 m3 ha–1, 0.3–0.8 m deep ditches were generally sufficient to lower WTD to the targeted depth of 0.35 m. These are significantly shallower ditch depths than generally recommended in operational forestry. The main collector ditch should be naturally somewhat deeper to permit water outflow. Our study brings a firmer basis on environmentally sound forestry on drained peatlands.

  • Hökkä, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: hannu.hokka@luke.fi (email)
  • Laurén, University of Eastern Finland, Faculty of Science and Forestry, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: ari.lauren@uef.fi
  • Stenberg, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: leena.stenberg@luke.fi
  • Launiainen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: samuli.launiainen@luke.fi
  • Leppä, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: kersti.leppa@luke.fi
  • Nieminen, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: mika.nieminen@luke.fi
article id 10468, category Research article
Arto Haara, Juho Matala, Markus Melin, Janne Miettinen, Kari T. Korhonen, Tuula Packalen, Jari Varjo. (2021). Economic effects of grouse-friendly forest management. Silva Fennica vol. 55 no. 3 article id 10468. https://doi.org/10.14214/sf.10468
Keywords: grouse; simulation; trade-offs; game keeping; interest rate
Highlights: The economic effects of grouse friendly forest management were evaluated by simulating alternative forest management approaches on four large forest holdings in different parts of Finland; The grouse-friendly management of forest holdings was possible with minor effects on the economics in most cases: only in one case was the reduction of NPV more than 5% during a 30-year simulation period; The interest rates had an impact on the differences between the economic effects of the forest management approaches.
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Traditional timber production may have negative effects on other ecosystem services. Therefore, new forest management guidelines have been developed in order to enhance a habitat suitable for wildlife. In Finland, a recent example of this is grouse-friendly forest management (GFFM) which emphasises the preservation of grouse species (Tetronidae) habitats. This study aimed to analyse the economic effects of these guidelines. An analysis was made on how the application of GFFM affected the Net Present Value (NPV) in a 30-year simulation of forest management of four large forest holdings located from south to north in Finland. In the simulations, traditional forest management practices were compared to two levels of GFFM. Five levels of interest rate were used, namely 1, 2, 3, 4, and 5%. In most of the simulations, the NPV was reduced by about 1% or less due to the application of GFFM in comparison to the traditional reference forest management. Only in one case with more intensive GFFM, was the reduction of NPV more than 5%. The interest rates had an impact on the differences between the management approaches. For example, a low interest rate resulted in a higher thinning intensity in GFFM in comparison to traditional forest management, which lead to a higher NPV in GFFM. To sum up, it seems that it would be possible to manage forest holdings in a grouse-friendly manner with minor effects on the economics.

  • Haara, LUKE ORCID https://orcid.org/0000-0002-6895-5300 E-mail: arto.haara@luke.fi (email)
  • Matala, Natural Resources Institute Finland, (Luke), Natural resources, Yliopistokatu 6B, FI-80100 Joensuu, Finland E-mail: juho.matala@luke.fi
  • Melin, Natural Resources Institute Finland, (Luke), Bioeconomy and environment, Yli opistokatu 6B, FI-80100 Joensuu, Finland E-mail: markus.melin@luke.fi
  • Miettinen, Finnish Wildlife Agency, Ratatie 41, FI-91501 Muhos, Finland E-mail: janne.miettinen@riista.fi
  • Korhonen, Natural Resources Institute Finland, (Luke), Bioeconomy and environment, Yli opistokatu 6B, FI-80100 Joensuu, Finland E-mail: kari.t.korhonen@luke.fi
  • Packalen, Ministry of Agriculture and Forestry, P.O. Box 30, FI-00023 GOVERNMENT, Finland E-mail: tuula.packalen@mmm.fi
  • Varjo, Finnish Wildlife Agency, Sompiontie 1, FI-00730 Helsinki, Finland E-mail: jari.varjo@riista.fi
article id 10462, category Research article
Yrjö Nuutinen, Jari Miina, Timo Saksa, Dan Bergström, Johanna Routa. (2021). Comparing the characteristics of boom-corridor and selectively thinned stands of Scots pine and birch. Silva Fennica vol. 55 no. 3 article id 10462. https://doi.org/10.14214/sf.10462
Keywords: biomass; forestry; first thinning; geometrical thinning; systematic thinning
Highlights: After boom-corridor thinning (BCT), the number of stems per hectare was higher than that after selective thinning. The number of future crop trees was at the same level; The removal of the simulated intermediate thinnings and clearcutting in BCT treatments was 10–18% higher than in selective thinning. The saw log volumes were at the same level in both treatments.
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Young, dense forest in Finland and Sweden urgently need to receive first thinning. In such stands, conventional selective thinning methods make the harvester work time consuming and, thus, costly. To make small-sized trees economically competitive as raw material for bioenergy and biorefining, new harvesting technologies and/or thinning methods need to be developed. A potential solution is boom-corridor thinning (BCT), rendering effective cutting work. The aim of this study was to describe and compare the stand structure of two Scots pine stands (Pinus sylvestris L.) and one birch-dominated (Betula pendula Roth with natural downy birch, B. pubescens Ehrh.) stand after BCT and selective thinning at the first thinning phase. Furthermore, simulations were conducted to predict the future stand development after the first thinning treatments. The density of the growing stock was 16–46% higher after BCT treatment than after selective thinning because BCT stands included more small and supressed trees with a dbh < 100 mm. However, the numbers of future crop trees with a dbh > 140 mm per hectare were at the same level in both treatments. The stem volume removal per hectare did not differ between treatments. However, simulation of stand development and intermediate thinning and clearcutting revealed that the total removal volume was 10–18% higher in BCT stands compared to selectively thinned ones. The saw log volumes harvested did, however, not differ between treatments. This study shows that BCT generates stands with higher biodiversity compared to conventional thinning as higher levels of biomass removal can be reached throughout stand rotations.

  • Nuutinen, Natural Resources Institute Finland (Luke), Production systems, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: yrjo.nuutinen@luke.fi (email)
  • Miina, Natural Resources Institute Finland (Luke), Natural resources, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jari.miina@luke.fi
  • Saksa, Natural Resources Institute Finland (Luke), Natural resources, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@luke.fi
  • Bergström, Swedish University of Agricultural Sciences (SLU), Dept of Forest Biomaterials and Technology, Section of Forest Operations, SE-90183 Umeå, Sweden E-mail: dan.bergstrom@slu.se
  • Routa, Natural Resources Institute Finland (Luke), Production systems, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: johanna.routa@luke.fi
article id 10503, category Research article
Katja Lähtinen, Liina Häyrinen, Anders Roos, Anne Toppinen, Francisco X. Aguilar Cabezas, Bo J. Thorsen, Teppo Hujala, Anders Q. Nyrud, Hans F. Hoen. (2021). Consumer housing values and prejudices against living in wooden homes in the Nordic region. Silva Fennica vol. 55 no. 2 article id 10503. https://doi.org/10.14214/sf.10503
Keywords: housing markets; industrial building; structural material; sustainable urbanization; timber structures; urban construction; value expectations
Highlights: Consumers in the Nordic region are similar in their housing value expectations and prejudices against building with wood; Physical properties of houses seem to be less important as constituents of housing value for the consumers compared to intangible factors related to lifestyles and milieus; Urban consumers are the most prejudiced against wood building, and thus supply of homes meeting their value expectations is of a critical importance for sustainable urbanization.
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So far, consumer housing values have not been addressed as factors affecting the market diffusion potential of multi-storey wood building (MSWB). To fill the void, this study addresses different types of consumer housing values in Denmark, Finland, Norway, and Sweden (i.e., Nordic region), and whether they affect the likelihood of prejudices against building with wood in the housing markets. The data collected in 2018 from 2191 consumers in the Nordic region were analyzed with exploratory factor analysis and logistic binary regression analysis. According to the results, consumers’ perceptions on ecological sustainability, material usage and urban lifestyle were similar in all countries, while country-specific differences were detected for perceptions on aesthetics and natural milieus. In all countries, appreciating urban lifestyle and living in attractive neighborhoods with good reputation increased the likelihood of prejudices against wood building, while appreciation of aesthetics and natural milieus decreased the likelihood of prejudices. In strengthening the demand for MSWB and sustainable urbanization through actions in businesses (e.g., branding) and via public policy support (e.g., land zoning), few messages derive from the results. In all, abreast with the already existing knowledge on the supply side factors (e.g., wood building innovations), more customized information is needed on the consumer-driven issues affecting the demand potential of MSWB in the housing markets. This would enable, e.g., both enhancing the supply of wooden homes for consumers appreciating urban lifestyle and neighborhoods and fortifying positive image of wood among consumers especially appreciating good architecture and pleasant environmental milieus.

  • Lähtinen, Vaasan yliopisto/Seinäjoen yliopistokeskus E-mail: katja.lahtinen@luke.fi (email)
  • Häyrinen, Natural Resources Institute Finland (Luke), Bioeconomy and Environment Unit, P.O. Box 2, FI-00790 Helsinki, Finland E-mail: liina.hayrinen@luke.fi
  • Roos, Sveriges lantbruksuniversitet (SLU), Department of Forest Economics, Box 7060, SE-750 07 Uppsala, Sweden E-mail: anders.roos@slu.se
  • Toppinen, University of Helsinki, Department of Forest Sciences/Helsinki Institute of Sustainability Science (HELSUS), P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: anne.toppinen@helsinki.fi
  • Aguilar Cabezas, Sveriges lantbruksuniversitet (SLU), Department of Forest Economics, SE-901 83 Umeå, Sweden E-mail: francisco.aguilar@slu.se
  • Thorsen, University of Copenhagen, Department of Food and Resource Economics, Rolighedsvej 23, DK-1958 Frederiksberg C, DenmarkBo Jellesmark Thorsen E-mail: bjt@ifro.ku.dk
  • Hujala, University of Eastern Finland (UEF), Faculty of Science and Forestry, School of Forest Sciences, P.O. Box 111, 80101 FI-Joensuu, Finland E-mail: teppo.hujala@uef.fi
  • Nyrud, Norwegian University of Life Sciences (NMBU), Faculty of Environmental Sciences and Natural Resource Management, P.O. Box 5003, NO-1432 Ås, Norway E-mail: anders.qvale.nyrud@nmbu.no
  • Hoen, Norwegian University of Life Sciences (NMBU), Faculty of Environmental Sciences and Natural Resource Management, P.O. Box 5003, NO-1432 Ås, Norway E-mail: hans.hoen@nmbu.no
article id 10496, category Research article
Jouni Siipilehto, Harri Mäkinen, Kjell Andreassen, Mikko Peltoniemi. (2021). Models for integrating and identifying the effect of senescence on individual tree survival probability for Norway spruce. Silva Fennica vol. 55 no. 2 article id 10496. https://doi.org/10.14214/sf.10496
Keywords: forest dynamics; model comparison; between-tree competition; mortality model
Highlights: The effect of senescence was integrated into an individual tree survival model; The best model showed good fit for managed, unmanaged and old-growth stands; The probability for a large tree to survive decreased with increasing stand age; The best performed model included an interaction term between stem diameter and stand age and also stand age as a separate independent variable.
Abstract | Full text in HTML | Full text in PDF | Author Info

Ageing and competition reduce trees’ ability to capture resources, which predisposes them to death. In this study, the effect of senescence on the survival probability of Norway spruce (Picea abies (L.) Karst.) was analysed by fitting alternative survival probability models. Different model formulations were compared in the dataset, which comprised managed and unmanaged plots in long-term forest experiments in Finland and Norway, as well as old-growth stands in Finland. Stand total age ranged from 19 to 290 years. Two models were formulated without an age variable, such that the negative coefficient for the squared stem diameter described a decreasing survival probability for the largest trees. One of the models included stand age as a separate independent variable, and three models included an interaction term between stem diameter and stand age. According to the model including stand age and its interaction with stem diameter, the survival probability curves could intersect each other in stands with a similar structure but a different mean age. Models that did not include stand age underestimated the survival rate of the largest trees in the managed stands and overestimated their survival rate in the old-growth stands. Models that included stand age produced more plausible predictions, especially for the largest trees. The results supported the hypothesis that the stand age and senescence of trees decreases the survival probability of trees, and that the ageing effect improves survival probability models for Norway spruce.

  • Siipilehto, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, P.O. Box 2, FI-00790 Helsinki, Finland E-mail: jouni.siipilehto@luke.fi (email)
  • Mäkinen, Natural Resources Institute Finland (Luke), Production systems, Latokartanonkaari 9, P.O. Box 2, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0002-1820-6264 E-mail: harri.makinen@luke.fi
  • Andreassen, Norwegian Institute of Bioeconomy Research (NIBIO), NO-1431 Ås, Norway E-mail: kjellandreassen@gmail.com
  • Peltoniemi, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, P.O. Box 2, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0003-2028-6969 E-mail: mikko.peltoniemi@luke.fi
article id 10483, category Research article
Māra Kitenberga, Guntars Šņepsts, Jānis Vuguls, Didzis Elferts, Ieva Jaunslaviete, Āris Jansons. (2021). Tree- and stand-scale factors shape the probability of wind damage to birch in hemiboreal forests. Silva Fennica vol. 55 no. 2 article id 10483. https://doi.org/10.14214/sf.10483
Keywords: stem breakage; Betula pendula; snapping height; stem damage; stem fracture; tree stability
Highlights: Prior stem and root damage significantly increased the probability of birch being wind-damaged; Trees with a high slenderness ratio had a lower relative snapping height; The snapping height was significantly negatively affected by the prior stem and root damage.
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Strong wind is the major natural disturbance in European forests, that periodically causes tremendous damages to forestry. Yet, factors that affect the probability of wind damage for birch (Betula pendula Roth and B. pubescens Ehrh.), the most common deciduous tree species in hemiboreal forests, are studied scarcely. This study aimed to assess the effects of several tree- and stand-scale variables on the probability of wind damage to birch using data from the Latvian National Forest Inventory (2004–2018), and determine individual tree characteristics that affect the height of the stem breakage. The data analysis was done using the Bayesian binary logistic generalized linear mixed-effects model and a linear mixed-effects model. The probability of wind damage significantly increased by stand age, basal area, and slenderness ratio. Trees with prior damage had a significantly higher probability (odds ratio 4.32) for wind damage. For wind-damaged trees, the snapping height was significantly decreased by an increase in the slenderness ratio (p = 0.03) and prior damage (p = 0.003). Previously damaged trees were more frequently (73%) snapped in the lowest 40% of tree height than trees without prior damage (54%). The probability of wind damage is largely set by factors related to the selection of site, species composition, and rotation. The damage probability could be decreased by management measures that lower competition within the stand with particular regard to preserving intact remaining trees during these manipulations. Factors that reduce the probability of the damage simultaneously increase the snapping height, emphasizing their relevance for mitigation of the wind damages.

  • Kitenberga, Latvian State Forest Research Institute ‘Silava’, Rigas Street 111, LV–2169, Salaspils, Latvia ORCID https://orcid.org/0000-0002-6192-988X E-mail: mara.kitenberga@silava.lv
  • Šņepsts, Latvian State Forest Research Institute ‘Silava’, Rigas Street 111, LV–2169, Salaspils, Latvia E-mail: guntars.snepsts@silava.lv
  • Vuguls, Latvian State Forest Research Institute ‘Silava’, Rigas Street 111, LV–2169, Salaspils, Latvia E-mail: janis.vuguls@silava.lv
  • Elferts, Latvian State Forest Research Institute ‘Silava’, Rigas Street 111, LV–2169, Salaspils, Latvia; Faculty of Biology, University of Latvia, Jelgavas Street 1, LV–1004, Rīga, Latvia ORCID https://orcid.org/0000-0002-9401-1231 E-mail: didzis.elferts@lu.lv
  • Jaunslaviete, Latvian State Forest Research Institute ‘Silava’, Rigas Street 111, LV–2169, Salaspils, Latvia E-mail: ieva.jaunslaviete@silava.lv
  • Jansons, Latvian State Forest Research Institute ‘Silava’, Rigas Street 111, LV–2169, Salaspils, Latvia ORCID https://orcid.org/0000-0001-7981-4346 E-mail: aris.jansons@silava.lv (email)
article id 10447, category Research article
Marian Schönauer, Stephan Hoffmann, Martin Nolte, Dirk Jaeger. (2021). Evaluation of a new pruning and tending system for young stands of Douglas fir. Silva Fennica vol. 55 no. 2 article id 10447. https://doi.org/10.14214/sf.10447
Keywords: thinning; time study; Pseudotsuga menziesii; body posture; heart rate; OWAS; physical workload; young stand
Highlights: Electric pruning shears combined with workflow re-organization reduced time demand for pruning; Thinning by chainsaw was more productive than using the clearing saw Husqvarna 535FBX, which on the other hand showed lower physical workload; Overall cost savings through the use of the new system were quantified at 6%.
Abstract | Full text in HTML | Full text in PDF | Author Info

In Germany, management restrictions for Norway spruce (Picea abies (L.) H. Karst.) due to climate change lead to increasing interest in Douglas fir (Pseudotsuga menziesii (Mirb.) Franco) as a potential substituting species. However, Douglas fir requires cost-intensive silvicultural treatments, such as periodic thinnings and, in particular, pruning. In order to improve the efficiency of such treatments, a new tending system with an adapted two-step work system was analyzed. The new system, using electric pruning shears and the backpack clearing saw Husqvarna 535FBX ‘Spacer’, was compared to the conventional three-step work system, using handsaw and chainsaw and characterized by tree selection previously conducted as an independent work step. Time and motion studies to determine productivity and costs, as well as ergonomic analysis through heart rate measurements and posture analysis were conducted. Overall, the new system was found to be more productive and to have lower costs, with 8.9 trees per scheduled system hour (4.17 € tree–1), compared to the conventional system with 8.1 trees per scheduled system hour (4.44 € tree–1). Ergonomic improvements with the new system could be mainly observed during the felling of competing trees, when the level of heart rate reserve was reduced by 9.3 percent points, compared to the conventional system. However, significant advantages in reducing unfavorable body postures expected for the ‘Spacer’ could not be confirmed. Since time savings within the new system were mainly attributed to the adaptation of workflow and the use of the electric shears during pruning, it should be considered to replace the ‘Spacer’ within the new system by light chainsaws for best results under the conditions investigated.

  • Schönauer, Department of Forest Work Science and Engineering, University of Göttingen, Göttingen/Germany E-mail: marian.schoenauer@uni-goettingen.de (email)
  • Hoffmann, Department of Forest Work Science and Engineering, University of Göttingen, Göttingen/Germany; School of Forestry, University of Canterbury, Christchurch/New Zealand ORCID https://orcid.org/0000-0002-8077-967X E-mail: stephan.hoffmann@uni-goettingen.de
  • Nolte, Forest Education Center FBZ / State Enterprise Forestry and Timber NRW, Arnsberg/Germany E-mail: Martin.Nolte@wald-und-holz.nrw.de
  • Jaeger, Department of Forest Work Science and Engineering, University of Göttingen, Göttingen/Germany E-mail: dirk.jaeger@uni-goettingen.de
article id 10446, category Research article
Juha Kaitera, Tuomas Kauppila, Jarkko Hantula. (2021). Assessment of the potential of Norway-spruce-seed-orchard associated plants to serve as alternate hosts of Thekopsora areolata. Silva Fennica vol. 55 no. 2 article id 10446. https://doi.org/10.14214/sf.10446
Keywords: Picea abies; cherry-spruce rust; epidemics; rust diseases; Prunus; heteroecism
Highlights: Prunus is the only alternate host of cherry-spruce rust identified in this study; Cherry-spruce rust does not infect plants that belong to common ground vegetation flora in Finnish Norway spruce seed orchards; Spores from different Finnish cherry-spruce rust populations are similarly pathogenic on Prunus; Disease control should be focused only on Prunus in seed orchards.
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The alternate host range of cherry-spruce rust is poorly studied although such information could be important in protecting spruce seed orchards from infections. Pathogenicity of cherry-spruce rust, Thekopsora areolata (Fr.) Magnus, was investigated on potential alternate host species in a greenhouse and in a laboratory in Finland. Five common species of Ericaceae, Vaccinium myrtillus L., V. uliginosum L., V. vitis-idaea L., Empetrum nigrum L. and Arctostaphylos uva-ursi (L.) Spreng, were inoculated in the greenhouse using aeciospores from seven Norway spruce [Picea abies (L.) H. Karst.] seed orchards suffering from T. areolata in 2018. In addition, young detached leaves of Vaccinium spp. and 17 other plant species of ground vegetation from eight Norway spruce seed orchards were inoculated with aeciospores from six seed orchards in the laboratory in 2019. Also, young leaves of Prunus padus L. trees growing within the seed orchards or close to them were inoculated as controls. None of the inoculated leaves of the potential alternate hosts formed uredinia either in the greenhouse or in the laboratory. In contrast, leaves of P. padus from the seed orchards were infected by the six spore sources from six seed orchards and produced uredinia. As T. areolata spores were able to infect only P. padus, but not the other tested species belonging to ground flora, it was concluded that T. areolata disperses only via Prunus spp. in Finnish seed orchards.

  • Kaitera, Natural Resources Institute Finland (Luke), Natural Resources, FI-90570 Oulu, Finland E-mail: juha.kaitera@luke.fi (email)
  • Kauppila, Botanical Gardens, University of Oulu, FI-90014 Oulu, Finland E-mail: tuomas.kauppila@oulu.fi
  • Hantula, Natural Resources Institute Finland (Luke), Natural Resources, FI-00790 Helsinki, Finland E-mail: jarkko.hantula@luke.fi
article id 10415, category Research article
Lele Lu, Sophan Chhin, Jianguo Zhang, Xiongqing Zhang. (2021). Modelling tree height-diameter allometry of Chinese fir in relation to stand and climate variables through Bayesian model averaging approach. Silva Fennica vol. 55 no. 2 article id 10415. https://doi.org/10.14214/sf.10415
Keywords: Cunninghamia lanceolata; Bayesian model averaging; height-diameter allometry; stand and climate variables; stepwise regression
Highlights: Bayesian model averaging (BMA) and stepwise regression (SR) were compared for modelling tree height-diameter allometry; The model acquired by SR was equal to the model with the third highest posterior probability of the BMA models; BMA produced estimates of the model parameters with slightly narrower ranges around the estimate of the population parameter; Temperature was the dominant climate variable shaping the allometry.
Abstract | Full text in HTML | Full text in PDF | Author Info

Tree height-diameter allometry reflects the response of specific species to above and belowground resource allocation patterns. However, traditional methods (e.g. stepwise regression (SR)) may ignore model uncertainty during the variable selection process. In this study, 450 trees of Chinese fir (Cunninghamia lanceolata (Lamb.) Hook.) grown at five spacings were used. We explored the height-diameter allometry in relation to stand and climate variables through Bayesian model averaging (BMA) and identifying the contributions of these variables to the allometry, as well as comparing with the SR method. Results showed the SR model was equal to the model with the third highest posterior probability of the BMA models. Although parameter estimates from the SR method were similar to BMA, BMA produced estimates with slightly narrower 95% intervals. Heights increased with increasing planting density, dominant height, and mean annual temperature, but decreased with increasing stand basal area and summer mean maximum temperature. The results indicated that temperature was the dominant climate variable shaping the height-diameter allometry for Chinese fir plantations. While the SR model included the mean coldest month temperature and winter mean minimum temperature, these variables were excluded in BMA, which indicated that redundant variables can be removed through BMA.

  • Lu, Key Laboratory of Tree Breeding and Cultivation of the National Forestry and Grassland Administration, Research Institute of Forestry, Chinese Academy of Forestry, Beijing 100091, P. R. China; Collaborative Innovation Center of Sustainable Forestry in Southern China, Nanjing Forestry University, Nanjing, 210037, P. R. China E-mail: 18556439861@163.com
  • Chhin, Division of Forestry and Natural Resources, West Virginia University, 322 Percival Hall, 1145 Evansdale Dr, Morgantown, West Virginia, 26506, USA E-mail: steve.chhin@mail.wvu.edu
  • Zhang, Key Laboratory of Tree Breeding and Cultivation of the National Forestry and Grassland Administration, Research Institute of Forestry, Chinese Academy of Forestry, Beijing 100091, P. R. China E-mail: xqzhang85@caf.ac.cn
  • Zhang, Key Laboratory of Tree Breeding and Cultivation of the National Forestry and Grassland Administration, Research Institute of Forestry, Chinese Academy of Forestry, Beijing 100091, P. R. China; Collaborative Innovation Center of Sustainable Forestry in Southern China, Nanjing Forestry University, Nanjing, 210037, P. R. China E-mail: xqzhang85@yahoo.com (email)
article id 10409, category Research article
Noé Dumas, Mathieu Dassot, Jonathan Pitaud, Jérôme Piat, Lucie Arnaudet, Claudine Richter, Catherine Collet. (2021). Four-year-performance of oak and pine seedlings following mechanical site preparation with lightweight excavators. Silva Fennica vol. 55 no. 2 article id 10409. https://doi.org/10.14214/sf.10409
Keywords: vegetation control; reforestation; seedling growth; seedling survival
Highlights: Mechanical site preparation (MSP) with lightweight excavators controls highly competitive plant species (Molinia caerulea, Pteridium aquilinum) much more efficiently than MSP with conventional methods; This MSP approach improves four-year survival and growth of pine seedlings, but it is less evident for oak seedlings.
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Mechanical site preparation methods that used tools mounted on lightweight excavators and that provided localised intensive preparation were tested in eight experimental sites across France where the vegetation was dominated either by Molinia caerulea (L.) Moench or Pteridium aquilinum (L.) Kuhn. Two lightweight tools (Deep Scarifier: DS; Deep Scarifier followed by Multifunction Subsoiler: DS+MS) were tested in pine (Pinus sylvestris L., Pinus nigra var. corsicana (Loudon) Hyl. or Pinus pinaster Aiton) and oak (Quercus petraea (Matt.) Liebl. or Quercus robur L.) plantations. Regional methods commonly used locally (herbicide, disk harrow, mouldboard plow) and experimental methods (repeated herbicide application; untreated control) were used as references in the experiments. Neighbouring vegetation cover, seedling survival, height and basal diameter were assessed over three to five years after plantation. For pines growing in M. caerulea, seedling diameter after four years was 37% and 98% greater in DS and DS+MS, respectively, than in the untreated control. For pines growing in P. aquilinum, it was 62% and 107% greater in the same treatments. For oak, diameter was only 4% and 15% greater in M. caerulea, and 13% and 25% greater in P. aquilinum, in the same treatments. For pines, the survival rate after four years was 26% and 32% higher in M. caerulea and 64% and 70% higher in P. aquilinum, in the same treatments. For oak, it was 3% and 29% higher in M. caerulea and 37% and 31% higher in P. aquilinum. Herbicide, when applied for three or four years after planting, provided the best growth performances for pines growing in M. caerulea and P. aquilinum and for oaks growing in P. aquilinum. For these species and site combinations, DS+MS and DS treatments reduced the neighbouring vegetation cover for one to four years following site preparation.

  • Dumas, Université de Lorraine, AgroParisTech, INRAE, UMR Silva, 54000 Nancy, France E-mail: noe.dumas@inrae.fr
  • Dassot, EcoSustain, Environmental Engineering Office, Research and Development, 31, rue de Volmerange, 57330 Kanfen, France; Institut National de l’Information Géographique et Forestière, 1 rue des Blanches Terres, 54250 Champigneulles, France E-mail: mathieu.dassot@ign.fr
  • Pitaud, Office National des Forêts, Département Recherche Développement et Innovation, route d’Amance, 54280 Champenoux, France E-mail: jonathan.pitaud@onf.fr
  • Piat, Office National des Forêts, Département Recherche Développement et Innovation, 3 rue du petit château, 60200 Compiègne, France E-mail: jerome.piat@onf.fr
  • Arnaudet, Office National des Forêts, Département Recherche Développement et Innovation, 100 boulevard de la Salle, 45760 Boigny-sur-Bionne, France E-mail: lucie.arnaudet@onf.fr
  • Richter, Office National des Forêts, Département Recherche Développement et Innovation, Boulevard de Constance, 77300 Fontainebleau, France E-mail: claudine.richter@onf.fr
  • Collet, Université de Lorraine, AgroParisTech, INRAE, UMR Silva, 54000 Nancy, France ORCID https://orcid.org/0000-0003-0861-7796 E-mail: catherine.collet@inrae.fr (email)
article id 10460, category Research article
Hanna Lundmark, Lars Östlund, Torbjörn Josefsson. (2021). Continuity forest or second-generation forest? Historic aerial photos provide evidence of early clear-cutting in northern Sweden. Silva Fennica vol. 55 no. 1 article id 10460. https://doi.org/10.14214/sf.10460
Keywords: continuous cover forestry; forest history; aerial photography; satellite image; clear-cutting; clear-felling
Highlights: In the early 1900s clear-cutting had been applied to 10% of the forest land in the study area situated in Västernorrland province, northern Sweden; By the end of the 1940s 40% of the study area had been clear-cut and constituted second-generation forest; 50–70 years is too short of a time frame for assessing the continuity of a forest in the study area.
Abstract | Full text in HTML | Full text in PDF | Author Info

Modern forestry, which mainly consists of clear-cutting, is one of the most important factors influencing today’s boreal forests. In Sweden, the breaking point for modern forestry is generally considered to be around 1950. Recently, our common knowledge of the implementation of clear-cutting in Sweden has increased, and new research indicates that clear-cutting systems were already applied before the 1950s. In this case study, we used aerial photographs from the 1940s to analyze the extent of contemporaneous clear-cuts and even-aged young forests in an area in northern Sweden. Our results show that almost 40% of the study area had already been clear-cut by the end of the 1940s, but also that clear-cutting had been applied to 10% of the forest land in the early 1900s. This implies that the historical development of forestry in northern Sweden is more complex than previously thought, and that certain proportions of the forest land were already second-generation forests in the 1950s. Our results have implications for the use of concepts such as “continuity forest”, suggesting that this concept should employ a time frame of at least 100 years.

  • Lundmark, Department of Forest Ecology and Management, Swedish University of Agricultural Sciences, S-901 83 Umeå, Sweden ORCID https://orcid.org/0000-0001-8402-7152 E-mail: hanna.lundmark@slu.se
  • Östlund, Department of Forest Ecology and Management, Swedish University of Agricultural Sciences, S-901 83 Umeå, Sweden ORCID https://orcid.org/0000-0002-7902-3672 E-mail: lars.ostlund@slu.se
  • Josefsson, Department of Forest Ecology and Management, Swedish University of Agricultural Sciences, S-901 83 Umeå, Sweden ORCID https://orcid.org/0000-0002-8734-5778 E-mail: torbjorn.josefsson@slu.se (email)
article id 10444, category Research article
Risto Kalliola, Timo Saarinen, Niko Tanski. (2021). Seasonal variations of foliar element distributions of silver birch in different habitats. Silva Fennica vol. 55 no. 1 article id 10444. https://doi.org/10.14214/sf.10444
Keywords: foliar nutrients; Betula pendula; deciduous tree; drought injury; element mapping; X-ray fluorescence
Highlights: Detailed mapping using micro X-ray fluorescence showed element variations in a seasonal cohort of silver birch leaves in six different habitats; Seasonal changes occurred in most of the ten studied nutrient elements; Different habitats showed unique seasonal development patterns in specific elements; Element distributions within individual leaf blades were variable; Phenotypic plasticity of foliar nutrients supports adaptation to patchy environments.
Abstract | Full text in HTML | Full text in PDF | Author Info

Novel information on silver birch (Betula pendula Roth) foliar element contents and their seasonal, between-habitat and leaf level variations are provided by applying fine-scaled element mapping with micro X-ray fluorescence. In the monthly leaf samples collected from May to October from six different habitats, pairwise scatter plots and Spearman’s rank correlations showed statistically significant positive correlations between Si, Al and Fe, and covariations between also many other pairs of elements. Of the ten elements studied, seven showed statistically significant changes in their average levels between May and June. The contents of P, S and K decreased in most habitats during the later season, whereas Ca and in some habitats also Mn and Zn increased. Comparing habitats, trees in the limestone habitat had relatively low content of Mg, strongly increasing levels of P until the late season, and high content of Ca and Fe. Other habitats also revealed distinctive particularities in their foliar elements, such as a high relative content of S and a low content of Ca at the seashore. Mn was high in three habitats, possibly due to bedrock characteristics. Except for P, the contents of all elements diverged between the midrib and other leaf areas. Zn content was particularly high in the leaf veins. Mn levels were highest at the leaf margins, indicating a possible sequestration mechanism for this potentially harmful element. Si may help to alleviate the metallic toxicities of Al and Fe. Because the growing season studied was dry, some trees developed symptoms of drought stress. The injured leaf parts had reduced levels of P, S and K, suggesting translocation of these nutrients before permanent damage.

  • Kalliola, Department of geography and geology, University of Turku, FI-20014 Turku, Finland ORCID https://orcid.org/0000-0003-2454-8217 E-mail: risto.kalliola@utu.fi (email)
  • Saarinen, Department of geography and geology, University of Turku, FI-20014 Turku, Finland E-mail: tijusa@utu.fi
  • Tanski, Department of geography and geology, University of Turku, FI-20014 Turku, Finland E-mail: niko.tanski@utu.fi
article id 10422, category Research article
Juha Kaitera, Jouni Karhu. (2020). Temperature range for germination of Thekopsora areolata aeciospores from Finnish Norway spruce seed orchards. Silva Fennica vol. 55 no. 1 article id 10422. https://doi.org/10.14214/sf.10422
Keywords: Picea abies; cherry-spruce rust; cones; epidemics; rust diseases
Highlights: Cherry-spruce rust was able to germinate at very low temperature (6 °C) close to the temperature when thermal growth starts; Optimum spore germination occurred between 15–24 °C; High temperature of 30 °C drastically reduced germination; Temperature had the most significant effect on germination, while spore source had less significant effect and agar media a non-significant effect on germination in a Fixed Effects model.
Abstract | Full text in HTML | Full text in PDF | Author Info

Cherry-spruce rust caused by Thekopsora areolata (Fr.) Magnus is a serious cone pathogen of Norway spruce [Picea abies (L.) Karst.]. The rust causes great economical losses in seed orchards specialized in the production of high quality seeds. Germination range of T. areolata aeciospores from rust populations (spore sources) in seven Finnish Norway spruce seed orchards was tested on water agar and malt agar at nine temperatures varying between 6–30 °C. The temperature range of spore germination was high varying between 6 °C and 27 °C, while germination was retarded at 30 °C. The peak in germination rate of all spore sources occurred between 15–24 °C. In a model with fixed effects of agar media, temperature and spore source, temperature had the most significant effect on germination. Spore source had a less significant effect, while agar media had a non-significant effect on germination. The rust was able to germinate at low temperatures corresponding to temperatures when the thermal growing season starts at 5 °C in the spring. As spores from cones from both the spruce canopy and the ground showed very similar germination ranges, it indicated the great capacity of all spores of the rust to germinate early in the spring. Hot temperatures with over 30 °C drastically reduced germination of the rust.

  • Kaitera, Natural Resources Institute Finland (Luke), Natural resources and bioproduction, Paavo Havaksentie 3, FI-90014 University of Oulu, Finland E-mail: juha.kaitera@luke.fi (email)
  • Karhu, Natural Resources Institute Finland (Luke), Natural resources and bioproduction, Paavo Havaksentie 3, FI-90014 University of Oulu, Finland E-mail: jouni.karhu@luke.fi
article id 10420, category Research article
Eeva-Liisa Terhonen, Jumoke Babalola, Risto Kasanen, Risto Jalkanen, Kathrin Blumenstein. (2021). Sphaeropsis sapinea found as symptomless endophyte in Finland. Silva Fennica vol. 55 no. 1 article id 10420. https://doi.org/10.14214/sf.10420
Keywords: Pinus sylvestris; Scots pine; Diplodia sapinea; Diplodia tip blight
Highlights: Sphaeropsis sapinea was found for the first time as an endophyte in healthy Scots pine in Finland; This finding confirms that S. sapinea can proliferate in a symptomless stage in Scots pine in Finland.
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The aim of this study was to determine if the ascomycete fungus Sphaeropsis sapinea (Fr.) Dyko & B. Sutton (syn. Diplodia sapinea (Fr.) Fuckel) could be cultured from surface sterilized Scots pine twigs presenting the endophytic stage of this fungus. This fungus causes the disease called Diplodia tip blight in conifers. Symptoms become visible when trees have been weakened by abiotic stressors related to temperature, drought and hailstorms. The disease is rapidly increasing and is observed regularly in Scots pine (Pinus sylvestris L.) forests in Europe. Changes in climatic conditions will gradually increase the damage of this pathogen, because it is favored by elevated temperatures and additionally the host trees will be more susceptible due to related environmental stress. Diplodia tip blight is emerging towards Northern latitudes, thus, actions to monitor the spread of S. sapinea in pine-dominated forests should be undertaken in Finland. Our aim was to search for S. sapinea in Scots pine along a transect in Finland. Branch samples were collected from healthy Scots pine, fungal endophytes were isolated and morphologically identified. Sixteen S. sapinea strains were found from four Scots pine trees from two locations. This finding confirms that S. sapinea is found as an endophyte in healthy Scots pine in Finland.

  • Terhonen, Forest Pathology Research Group, Department of Forest Botany and Tree Physiology, Faculty of Forest Sciences and Forest Ecology, University of Goettingen, Büsgen-Institute, Büsgenweg 2, D-37077 Göttingen, Germany E-mail: terhonen@uni-goettingen.de (email)
  • Babalola, Forest Pathology Research Group, Department of Forest Botany and Tree Physiology, Faculty of Forest Sciences and Forest Ecology, University of Goettingen, Büsgen-Institute, Büsgenweg 2, D-37077 Göttingen, Germany E-mail: j.babalola@stud.uni-goettingen.de
  • Kasanen, Forest Pathology Lab, Department of Forest Sciences, University of Helsinki, Latokartanonkaari 7, FI-00014 University of Helsinki, Finland E-mail: risto.kasanen@helsinki.fi
  • Jalkanen, Rovaniemi Research Unit, Natural Resources Institute Finland (Luke), Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: ristjal@gmail.com
  • Blumenstein, Forest Pathology Research Group, Department of Forest Botany and Tree Physiology, Faculty of Forest Sciences and Forest Ecology, University of Goettingen, Büsgen-Institute, Büsgenweg 2, D-37077 Göttingen, Germany E-mail: kathrin.blumenstein@uni-goettingen.de
article id 10391, category Research article
Victoria A.M. Poljatschenko, Lauri T. Valsta. (2021). Carbon emissions displacement effect of Finnish mechanical wood products by dominant tree species in a set of wood use scenarios. Silva Fennica vol. 55 no. 1 article id 10391. https://doi.org/10.14214/sf.10391
Keywords: avoided emissions; mechanical forest industry; substitution effect; wooden construction
Highlights: Wood product consumption patterns are combined with production-based displacement factors to estimate displacement factors for logs by tree species; The main tree species in Finland have somewhat unequal displacement factors; Historical and projected wood use scenarios show large variation in the levels of avoided emissions between individual years and scenarios.
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The carbon emissions displacement effect of Finnish logs for mechanical wood products by dominant tree species (Scots pine, Pinus sylvestris L.; Norway spruce, Picea abies (L.) H. Karst.; Birch, Betula spp.) was assessed by combining information from previous studies of current consumption patterns with displacement factors (DF) for structural construction, non-structural construction, and energy usage. We did not conduct additional life cycle analyses compared to the current literature. Our aim was to identify the factors that most extensively influence the displacement effect and to estimate the overall climate effect of Finnish logs in light of current production levels of mechanical forest industry. The analyses were based on information from both statistics and proprietary sources. Contrary to previous studies, we provide DFs by main tree species in Finland, which has been an unidentified area of research to date. Additionally, we apply a more detailed classification of structural and non-structural wood products. This study did not include effects on the forest carbon sink, as they depend case-wise on forest resources and forest management. According to our results, with current production and consumption trends, the average displacement effects for domestic Scots pine, Norway spruce, and birch logs were 1.28, 1.16, and 1.43 Mg C/Mg C, respectively. The corresponding overall annual displacement effect caused by the current production of sawn wood and wood-based panels was 12.3 Tg CO2 for Finland for the BAU scenario and varied between 8.6 and 16.3 Tg CO2 depending on the wood use scenario.

  • Poljatschenko, Simosol Oy, Hämeenkatu 10, FI-11100 Riihimäki, Finland E-mail: victoria.poljatschenko@simosol.fi (email)
  • Valsta, Department of Forest Sciences, University of Helsinki, Latokartanonkaari 7, FI-00014 University of Helsinki, Finland E-mail: lauri.valsta@helsinki.fi
article id 10371, category Research article
Katalin Waga, Jukka Malinen, Timo Tokola. (2021). Locally invariant analysis of forest road quality using two different pulse density airborne laser scanning datasets. Silva Fennica vol. 55 no. 1 article id 10371. https://doi.org/10.14214/sf.10371
Keywords: ALS; DEM; forest road quality; reference DEM; unpaved forest road
Highlights: Airborne laser scanning is used to assess forest road quality; High-pulse data analysis classified roads with good performance; Two-step classification further improved the accuracy; A reference surface improved the classification results of the low pulse data; 66–75% of the roads were correctly classified using the reference surface.
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Two different pulse density airborne laser scanning datasets were used to develop a quality assessment methodology to determine how airborne laser scanning derived variables with the use of reference surface can determine forest road quality. The concept of a reference DEM (Digital Elevation Model) was used to guarantee locally invariant topographic analysis of road roughness. Structural condition, surface wear and flatness were assessed at two test sites in Eastern Finland, calculating surface indices with and without the reference DEM. The high pulse density dataset (12 pulses m–2) gave better classification results (77% accuracy of the correctly classified road sections) than the low pulse density dataset (1 pulse m–2). The use of a reference DEM increased the precision of the road quality classification with the low pulse density dataset when the classification was performed in two-steps. Four interpolation techniques (Inverse Weighted Distance, Kriging, Natural Neighbour and Spline) were compared, and spline interpolation provided the best classification. The work shows that applying a spline reference DEM it is possible to identify 66% of the poor quality road sections and 78% of the good ones. Locating these roads is essential for road maintenance.

  • Waga, Faculty of Science and Forestry, University of Eastern Finland, Yliopistokatu 7, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0003-1496-7012 E-mail: katalin.waga@uef.fi (email)
  • Malinen, Faculty of Science and Forestry, University of Eastern Finland, Yliopistokatu 7, FI-80100 Joensuu, Finland; Metsäteho Ltd., Vernissakatu 1, FI-01300 Vantaa, Finland ORCID https://orcid.org/0000-0002-5023-1056 E-mail: jukka.malinen@uef.fi
  • Tokola, Faculty of Science and Forestry, University of Eastern Finland, Yliopistokatu 7, FI-80100 Joensuu, Finland E-mail: timo.tokola@uef.fi
article id 10360, category Research article
Mikko Kukkonen, Eetu Kotivuori, Matti Maltamo, Lauri Korhonen, Petteri Packalen. (2021). Volumes by tree species can be predicted using photogrammetric UAS data, Sentinel-2 images and prior field measurements. Silva Fennica vol. 55 no. 1 article id 10360. https://doi.org/10.14214/sf.10360
Keywords: forest inventory; satellite image; open data; drone; stereo matching; unmanned aircraft system
Highlights: A UAS-based species-specific forest inventory approach that avoids new field measurements is presented; Models were constructed using previously measured training plots and remotely sensed data; Bi-seasonal Sentinel-2 data were beneficial in the prediction of species-specific volumes; RMSE values associated with the prediction of volumes by tree species and total volume at the validation plot level were 33.4–62.6% and 9.0%, respectively.
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Photogrammetric point clouds obtained with unmanned aircraft systems (UAS) have emerged as an alternative source of remotely sensed data for small area forest management inventories (FMI). Nonetheless, it is often overlooked that small area FMI require considerable field data in addition to UAS data, to support the modelling of forest attributes. In this study, we propose a method whereby tree volumes by species are predicted with photogrammetric UAS data and Sentinel-2 images, using models fitted with airborne laser scanning data. The study area is in a managed boreal forest area in Eastern Finland. First, we predicted total volume with UAS point cloud metrics using a prior regression model fitted in another area with ALS data. Tree species proportions were then predicted by k nearest neighbor (k-NN) imputation based on bi-seasonal Sentinel-2 images without measuring new field plot data. Species-specific volumes were then obtained by multiplying the total volume by species proportions. The relative root mean square error (RMSE) values for total and species-specific volume predictions at the validation plot level (30 m × 30 m) were 9.0%, and 33.4–62.6%, respectively. Our approach appears promising for species-specific small area FMI in Finland and in comparable forest conditions in which suitable field plots are available.

  • Kukkonen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: mikko.kukkonen@uef.fi (email)
  • Kotivuori, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: eetu.kotivuori@uef.fi
  • Maltamo, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: matti.maltamo@uef.fi
  • Korhonen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: lauri.korhonen@uef.fi
  • Packalen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: petteri.packalen@uef.fi
article id 10414, category Research article
Jouni Siipilehto, Micky Allen, Urban Nilsson, Andreas Brunner, Saija Huuskonen, Soili Haikarainen, Narayanan Subramanian, Clara Antón-Fernández, Emma Holmström, Kjell Andreassen, Jari Hynynen. (2020). Stand-level mortality models for Nordic boreal forests. Silva Fennica vol. 54 no. 5 article id 10414. https://doi.org/10.14214/sf.10414
Keywords: Norway spruce; Scots pine; simulation; broadleaved species; logistic function; period length; plot size
Highlights: Models were developed for predicting stand-level mortality from a large representative NFI data set; The logistic function was used for modelling the probability of no mortality and the proportion of basal area in surviving trees; The models take into account the variation in prediction period length and in plot size; The models showed good fit with respect to stand density, developmental stage and species structure, and showed satisfying fit in the independent data set of unmanaged spruce stands.
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New mortality models were developed for the purpose of improving long-term growth and yield simulations in Finland, Norway, and Sweden and were based on permanent national forest inventory plots from Sweden and Norway. Mortality was modelled in two steps. The first model predicts the probability of survival, while the second model predicts the proportion of basal area in surviving trees for plots where mortality has occurred. In both models, the logistic function was used. The models incorporate the variation in prediction period length and in plot size. Validation of both models indicated unbiased mortality rates with respect to various stand characteristics such as stand density, average tree diameter, stand age, and the proportion of different tree species, Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies (L.) Karst.), and broadleaves. When testing against an independent dataset of unmanaged spruce-dominated stands in Finland, the models provided unbiased prediction with respect to stand age.

  • Siipilehto, Natural Resources Institute Finland (Luke), Natural resources, P.O. Box 2, FI-00790 Helsinki, Finland E-mail: jouni.siipilehto@luke.fi (email)
  • Allen, Norwegian Institute of Bioeconomy Research (NIBIO), Division of Forest and Forest Products, NO-1431 Ås, Norway; Larson and McGowin Inc., Mobile, AL 36607, USA ORCID https://orcid.org/0000-0002-7824-2849 E-mail: micky.allen@nibio.no
  • Nilsson, Swedish University of Agricultural Sciences (SLU), Southern Swedish Forest Research Centre, SE-23053 Alnarp, Sweden ORCID https://orcid.org/0000-0002-7624-4031 E-mail: urban.nilsson@slu.se
  • Brunner, Norwegian University of Life Sciences (NMBU), Faculty of Environmental Sciences and Natural Resource Management, NO-1432 Ås, Norway ORCID https://orcid.org/0000-0003-1668-9714 E-mail: andreas.brunner@nmbu.no
  • Huuskonen, Natural Resources Institute Finland (Luke), Natural resources, P.O. Box 2, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0001-8630-3982 E-mail: saija.huuskonen@luke.fi
  • Haikarainen, Natural Resources Institute Finland (Luke), Natural resources, P.O. Box 2, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0001-8703-3689 E-mail: soili.haikarainen@luke.fi
  • Subramanian, Swedish University of Agricultural Sciences (SLU), Southern Swedish Forest Research Centre, SE-23053 Alnarp, Sweden ORCID https://orcid.org/0000-0003-2777-3241 E-mail: narayanan.subramanian@slu.se
  • Antón-Fernández, Norwegian Institute of Bioeconomy Research (NIBIO), Division of Forest and Forest Products, NO-1431 Ås, Norway ORCID https://orcid.org/0000-0001-5545-3320 E-mail: clara.anton.fernandez@nibio.no
  • Holmström, Swedish University of Agricultural Sciences (SLU), Southern Swedish Forest Research Centre, SE-23053 Alnarp, Sweden ORCID https://orcid.org/0000-0003-2025-1942 E-mail: emma.holmstrom@slu.se
  • Andreassen, Norwegian Institute of Bioeconomy Research (NIBIO), Division of Forest and Forest Products, NO-1431 Ås, Norway ORCID https://orcid.org/0000-0003-4272-3744 E-mail: kjellandreassen@gmail.com
  • Hynynen, Natural Resources Institute Finland (Luke), Natural resources, P.O. Box 2, FI-00790 Helsinki, Finland E-mail: jari.hynynen@luke.fi
article id 10396, category Research article
Agnese Gailīte, Anita Gaile, Dainis E. Ruņģis. (2020). Genetic diversity and structure of wild Vaccinium populations - V. myrtillus, V. vitis-idaea and V. uliginosum in the Baltic States. Silva Fennica vol. 54 no. 5 article id 10396. https://doi.org/10.14214/sf.10396
Keywords: molecular markers; bilberries; bog bilberries; chloroplast SSR; in situ conservation; lingonberries; nuclear SSR
Highlights: Wild Vaccinium species were studied using EST-SSR and chloroplast SSR markers; Populations were moderately genetically differentiated, but without higher order clustering of groups of populations; Genetic diversity of populations growing under different management regimes was similar; Selection of populations for in situ conservation should focus on rare genotypes, more differentiated populations and geographic coverage.
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Vaccinium myrtillus L., V. vitis-idaea L. and V. uliginosum L. belong to the genus Vaccinium. These wild species are widely distributed and ecologically important within the Baltic countries but they have not been extensively studied using molecular markers. EST-SSR and cpSSR markers were used to investigate the population structure and genetic diversity of these species to obtain information useful for the development of in situ conservation strategies. Wild Vaccinium species populations are moderately genetically differentiated, with some populations more highly differentiated, but without higher order clustering of groups of populations, indicating that there are no dispersal barriers for these species within the Baltic countries. Genetic diversity of populations growing in protected areas, managed forests and intensively utilised public recreational areas is similar. The results from this study can be utilised for the selection of populations for the in situ conservation of the studied Vaccinium species. In addition, complementary ex situ conservation strategies can be used for the preservation of rare varieties (e.g. V. myrtillus var. leucocarpum).

  • Gailīte, Genetic Resource Centre, Latvian State Forest Research Institute “Silava”, 111 Rigas Street, Salaspils, Latvia, LV-2169 E-mail: agnese.gailite@silava.lv (email)
  • Gaile, Genetic Resource Centre, Latvian State Forest Research Institute “Silava”, 111 Rigas Street, Salaspils, Latvia, LV-2169 E-mail: anita.gaile@silava.lv
  • Ruņģis, Genetic Resource Centre, Latvian State Forest Research Institute “Silava”, 111 Rigas Street, Salaspils, Latvia, LV-2169 ORCID https://orcid.org/0000-0001-5173-2912 E-mail: dainis.rungis@silava.lv
article id 10379, category Research article
Juha Laitila, Kari Väätäinen. (2020). Productivity of harvesting and clearing of brushwood alongside forest roads. Silva Fennica vol. 54 no. 5 article id 10379. https://doi.org/10.14214/sf.10379
Keywords: logging; energy wood; time study; harwarder; spiral cutter; whole-trees
Highlights: The results can be used as a basis to determine in what kinds of cases brushwood biomass should be recovered and where it should be left to decay; The average volume of harvested brushwood and forwarding distance are the key elements to harvesting productivity with a harwarder; Stump diameter has a strong impact on clearing productivity of brushwood.
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Expertise in the cost-efficient utilization and treatment of brushwood on forest roadside sites is limited. In the present study, the productivity of brushwood clearing and harvesting on forest roadside sites was defined by creating time-consumption models or parameters for the aforementioned working methods. Compiled time consumption models and parameters for the brushwood clearing and harvesting can be used as a basis for evaluating alternative management practices and to determine when brushwood biomass should be harvested and when it should be left to decay. The harvesting of brushwood was based on the harwarder system and the clearing of brushwood was done with a spiral cutter, which is a novel accessory for cutting roadside vegetation. Based on the study results, the average volume of harvested brushwood and forwarding distance are the key elements that have an effect on harvesting productivity with harwarders. Correspondingly, stump diameter has a strong impact on the clearing productivity of brushwood. The plot-wise productivity of the spiral cutter in brushwood clearings varied in the range of 0.19–0.61 ha per PMh. An increase in stump diameter slowed down the clearing productivity of the spiral cutter and there was a clear step downward in clearing productivity as the average diameter increased from 30 mm to 40 mm. The machinery under study operated well and there were no interruptions due to machine breakdowns.

  • Laitila, Natural Resources Institute Finland (Luke), Production systems, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: juha.laitila@luke.fi (email)
  • Väätäinen, Natural Resources Institute Finland (Luke), Production systems, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: kari.vaatainen@luke.fi
article id 10369, category Research article
Tiina Laine, Ville Kankaanhuhta, Juho Rantala, Timo Saksa. (2020). Effects of spot mounding and inverting on growth of conifers, exposed mineral soil and natural birch regeneration. Silva Fennica vol. 54 no. 5 article id 10369. https://doi.org/10.14214/sf.10369
Keywords: boreal forest; regeneration; site preparation; mounding; young stand management
Highlights: There were 21% fewer naturally regenerated birches ha–1 in inverted areas (22 951) compared to spot mounded areas (29 086); Spot mounding exposed slightly, but not statistically significantly, larger area of mineral soil than inverting; There was no difference in seedling mortality or growth between the site preparation methods.
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In Nordic forests, consistent evidence about better seedling survival rate and increased growth due to site preparation have been obtained in numerous studies. Proper site preparation method can reduce costs of the whole regeneration chain through its effects on survival of planted seedlings, abundance of natural regeneration and competition in early stand development. This study compared the natural regeneration of birches (silver birch (Betula pendula Roth) and downy birch (B. pubescens Ehrh.)), amount of exposed mineral soil, and growth of planted seedlings between spot mounding and inverting site preparation methods. Present study was conducted in eight forest stands established in 2012 or 2015. Even though difference was not statistically significant, inverting exposed less mineral soil than spot mounding and thus reduced the natural regeneration of birch seedlings by 6135 seedlings ha–1 compared to spot mounding. However, the variation between regeneration areas was remarkable. There was no difference in seedling mortality or growth between the site preparation methods. In order to achieve high growth of conifers, moderate amount of exposed mineral soil and thus less naturally regenerated birch, inverting should be favored over spot mounding.

  • Laine, Metsä Group, P.O. Box 208, FI-70101 Kuopio, Finland E-mail: tiina.laine@metsagroup.com (email)
  • Kankaanhuhta, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: ville.kankaanhuhta@luke.fi
  • Rantala, Metsä Group, P.O. Box 10, FI-02020 METSÄ, Finland E-mail: juho.rantala@metsagroup.com
  • Saksa, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@luke.fi
article id 10351, category Research article
Karol Bronisz, Michał Zasada. (2020). Taper models for black locust in west Poland. Silva Fennica vol. 54 no. 5 article id 10351. https://doi.org/10.14214/sf.10351
Keywords: taper; Robinia pseudoacacia; mixed-effects models; section diameter over and under bark; volume prediction
Highlights: Seven taper models with different numbers of estimated parameters were analysed; Section diameter and volume was modelled using fixed and mixed-effects modelling approaches; The variable-form taper model with eight estimated parameters fitted the data the best; The lowest error for volume prediction was achieved for the fixed-effects taper model.
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The diameter at any point on a stem and tree volume are some of the most important types of information used in forest management planning. One of the methods to predict the diameter at any point on a stem is to develop taper models. Black locust (Robinia pseudoacacia L.) occurs in almost all forests in Poland, with the largest concentration in the western part of the country. Using empirical data obtained from 13 black locust stands (48 felled trees), seven taper models with different numbers of estimated parameters were analysed for section diameters both over and under bark using fixed and mixed-effects modelling approaches. Assuming a lack of additional measurements, the best fitted taper models were used for the prediction of over bark volume using both methods. The predicted volume was compared with the results from different volume equations available for black locust. The variable-form taper model with eight estimated parameters fitted the data the best. The lowest root mean square error for volume prediction was achieved for the elaborated fixed-effects taper model (0.0476), followed by the mixed-effects taper model (0.0489). At the same time, the difference between the volume relative errors achieved based on the taper models does not differ significantly from the results obtained using the volume equations already available for black locust (two of the three analysed).

  • Bronisz, Institute of Forest Sciences, Warsaw University of Life Sciences-SGGW, Nowoursynowska 159, PL 02-776 Warsaw, Poland E-mail: karol.bronisz@wl.sggw.pl (email)
  • Zasada, Institute of Forest Sciences, Warsaw University of Life Sciences-SGGW, Nowoursynowska 159, PL 02-776 Warsaw, Poland ORCID https://orcid.org/0000-0002-4881-296X E-mail: Michal.Zasada@wl.sggw.pl
article id 10334, category Research article
Annija Kārkliņa, Guntis Brūmelis, Iluta Dauškane, Didzis Elferts, Lāsma Freimane, Māra Kitenberga, Zane Lībiete, Roberts Matisons, Āris Jansons. (2020). Effect of salvage-logging on post-fire tree establishment and ground cover vegetation in semi-natural hemiboreal forests. Silva Fennica vol. 54 no. 5 article id 10334. https://doi.org/10.14214/sf.10334
Keywords: management; forest fire; natural disturbance; legacies; understory vegetation
Highlights: Effect of salvage logging on post-fire understory vegetation was assessed; Effect of salvage logging differed depending on forest types; In dry-poor stands, Calluna vulgaris was hindering other plant species; In wet stands, logging had positive effect on understory vegetation diversity; Salvage logging enhanced the effect of natural disturbance in dry-rich stands.
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Fire is a common disturbance in boreal forests causing changes in biological diversity at various spatial scales. In the past 100 years, forest management has limited fire outbreaks, but in the future, the fire-affected forest area is expected to increase in many regions due to climate change. Burned forests are typically salvage-logged, but the effect of this type of management versus natural regeneration on biological diversity is not well understood, particularly the mid-term effect to tree establishment and understory vegetation composition and diversity. Various management methods were used after a large fire in 1992 in a peatland-forest complex and neighbouring managed forests, which created an experimental setup for study of the effect of management after fire in the Sliteres National park, northwestern Latvia. Understory vegetation was described in plots using a design of four forest and three management types: natural regeneration (unmanaged) and managed sites with salvage logging followed by no further human intervention and salvage logging with planting. Post-fire management had different effect in each forest type. Species richness was higher in forest types with salvage logging than in natural regenerated sites on rich wet and rich dry forest types, but not for the poor forest types. Tree regeneration was generally greater in salvage-logged stands, but differed between forest types. Species composition was related to tree regeneration and canopy openness. In contrast to other studies, salvage logging had a positive mid-term effect to ground vegetation diversity and tree establishment in the studied stands, implying potential for concomitant management and conservation of ground cover vegetation in semi-natural stands.

  • Kārkliņa, Latvian State Forest Research Institute “Silava”, 111 Rigas Street, LV-2169, Salaspils, Latvia E-mail: annija.karklina@silava.lv (email)
  • Brūmelis, University of Latvia, Faculty of Biology, Jelgavas street 1, LV-1004, Riga, Latvia E-mail: guntis.brumelis@lu.lv
  • Dauškane, University of Latvia, Faculty of Biology, Jelgavas street 1, LV-1004, Riga, Latvia E-mail: iluta.dauskane@lu.lv
  • Elferts, Latvian State Forest Research Institute “Silava”, 111 Rigas Street, LV-2169, Salaspils, Latvia; University of Latvia, Faculty of Biology, Jelgavas street 1, LV-1004, Riga, Latvia E-mail: didzis.elferts@lu.lv
  • Freimane, Latvian State Forest Research Institute “Silava”, 111 Rigas Street, LV-2169, Salaspils, Latvia E-mail: lasma.freimane@silava.lv
  • Kitenberga, Latvian State Forest Research Institute “Silava”, 111 Rigas Street, LV-2169, Salaspils, Latvia E-mail: mara.kitenberga@silava.lv
  • Lībiete, Latvian State Forest Research Institute “Silava”, 111 Rigas Street, LV-2169, Salaspils, Latvia E-mail: zane.libiete@silava.lv
  • Matisons, Latvian State Forest Research Institute “Silava”, 111 Rigas Street, LV-2169, Salaspils, Latvia E-mail: robism@inbox.lv
  • Jansons, Latvian State Forest Research Institute “Silava”, 111 Rigas Street, LV-2169, Salaspils, Latvia E-mail: aris.jansons@silava.lv
article id 10309, category Research article
Petteri Seppänen, Antti Mäkinen. (2020). Comprehensive yield model for plantation teak in Panama. Silva Fennica vol. 54 no. 5 article id 10309. https://doi.org/10.14214/sf.10309
Keywords: simulation; teak; decision support system; Tectona grandis; Panama; taper curve; volume equation; yield model
Highlights: Tree level teak stem volume models, taper model and three sets of stand level yield models were developed using large empirical datasets; Tree volume models were satisfactorily validated against independent measurement data and other published models; Tree height as input parameter improved the stem volume model marginally; Stand level yield models produced comparable harvest volumes with models published in the literature; Stand level timber product outputs were found like actual harvests with an exception that the models marginally underestimate the share of logs in very large diameter classes.
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The purpose of this study was to prepare a comprehensive, computerized teak (Tectona grandis L.f) plantation yield model system that can be used to describe the forest dynamics, predict growth and yield and support forest planning and decision-making. Extensive individual tree and permanent sample plot data were used to develop tree-level volume models, taper curve models and stand-level yield models for teak plantations in Panama. Tree volume models were satisfactorily validated against independent measurement data and other published models. Tree height as input parameter improved the stem volume model marginally. Stand level yield models produced comparable harvest volumes with models published in the literature. Stand level volume product outputs were found like actual harvests with an exception that the models marginally underestimate the share of logs in very large diameter classes. The kind of comprehensive model developed in this study and implemented in an easy to use software package provides a very powerful decision support tool. Optimal forest management regimes can be found by simulating different planting densities, thinning regimes and final harvest ages. Forest practitioners can apply growth and yield models in the appropriate stand level inventory data and perform long term harvest scheduling at property level or even at an entire timberland portfolio level. Harvest schedules can be optimized using the applicable financial parameters (silviculture costs, harvesting costs, wood prices and discount rates) and constraints (market size and operational capacity).

  • Seppänen, Verdas Oy, Kihlinkuja 7, FI-50600 Mikkeli, Finland E-mail: petteri@verdas.fi (email)
  • Mäkinen,  Simosol Oy. Hämeenkatu 10, FI-11100 Riihimäki, Finland E-mail: antti.makinen@simosol.fi
article id 10291, category Research article
Sakari Tuominen, Andras Balazs, Annika Kangas. (2020). Comparison of photogrammetric canopy models from archived and made-to-order aerial imagery in forest inventory. Silva Fennica vol. 54 no. 5 article id 10291. https://doi.org/10.14214/sf.10291
Keywords: distribution; prediction; forest resources; mapping; aerial imaging; digital stereo-photogrammetry
Highlights: Two photogrammetric canopy models were tested in forest inventory: one based on archived standard aerial imagery acquired for ortho-mosaic production and another based on stereo-photogrammetrically oriented aerial imaging adjusted for stereo-photogrammetric canopy modelling; Both data sets were tested in the estimation of forest variables; Despite the differences in imaging parameters, there was little difference in their performance in predicting the forest inventory variables.
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In remote sensing-based forest inventories 3D point cloud data, such as acquired from airborne laser scanning, are well suited for estimating the volume of growing stock and stand height, but tree species recognition often requires additional optical imagery. A combination of 3D data and optical imagery can be acquired based on aerial imaging only, by using stereo photogrammetric 3D canopy modeling. The use of aerial imagery is well suited for large-area forest inventories, due to low costs, good area coverage and temporally rapid cycle of data acquisition. Stereo-photogrammetric canopy modeling can also be applied to previously acquired imagery, such as for aerial ortho-mosaic production, assuming that the imagery has sufficient stereo overlap. In this study we compared two stereo-photogrammetric canopy models combined with contemporary satellite imagery in forest inventory. One canopy model was based on standard archived imagery acquired primarily for ortho-mosaic production, and another was based on aerial imagery whose acquisition parameters were better oriented for stereo-photogrammetric canopy modeling, including higher imaging resolution and greater stereo-coverage. Aerial and satellite data were tested in the estimation of growing stock volume, volumes of main tree species, basal area and diameter and height. Despite the better quality of the latter canopy model, the difference of the accuracy of the forest estimates based on the two different data sets was relatively small for most variables (differences in RMSEs were 0–20%, depending on variable). However, the estimates based on stereo-photogrammetrically oriented aerial data retained better the original variation of the forest variables present in the study area.

  • Tuominen, Natural Resources Institute Finland (Luke), Bioeconomy and Environment, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: sakari.tuominen@luke.fi (email)
  • Balazs, Natural Resources Institute Finland (Luke), Bioeconomy and Environment, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: andras.balazs@luke.fi
  • Kangas, Natural Resources Institute Finland (Luke), Bioeconomy and Environment, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: annika.kangas@luke.fi
article id 10272, category Research article
Ana de Lera Garrido, Terje Gobakken, Hans Ole Ørka, Erik Næsset, Ole M. Bollandsås. (2020). Reuse of field data in ALS-assisted forest inventory. Silva Fennica vol. 54 no. 5 article id 10272. https://doi.org/10.14214/sf.10272
Keywords: airborne laser scanning; data reuse; temporal model transferability
Highlights: Six biophysical forest attributes were estimated for small stands without using up-to-date field data; The approaches included reused model relationships and forecasted field data; The accuracy of height estimates was comparable with the accuracy of an ordinary forest inventory with up-to-date field- and ALS data; Both approaches tended to produce estimates systematically different from the ground reference.
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Forest inventories assisted by wall-to-wall airborne laser scanning (ALS), have become common practice in many countries. One major cost component in these inventories is the measurement of field sample plots used for constructing models relating biophysical forest attributes to metrics derived from ALS data. In areas where ALS-assisted forest inventories are planned, and in which the previous inventories were performed with the same method, reusing previously acquired field data can potentially reduce costs, either by (1) temporally transferring previously constructed models or (2) projecting field reference data using growth models that can serve as field reference data for model construction with up-to-date ALS data. In this study, we analyzed these two approaches of reusing field data acquired 15 years prior to the current ALS acquisition to estimate six up-to-date forest attributes (dominant tree height, mean tree height, stem number, stand basal area, volume, and aboveground biomass). Both approaches were evaluated within small stands with sizes of approximately 0.37 ha, assessing differences between estimates and ground reference values. The estimates were also compared to results from an up-to-date forest inventory relying on concurrent field- and ALS data. The results showed that even though the reuse of historical information has some potential and could be beneficial for forest inventories, systematic errors may appear prominent and need to be overcome to use it operationally. Our study showed systematic trends towards the overestimation of lower-range ground references and underestimation of the upper-range ground references.

  • de Lera Garrido, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: ana.maria.lera.garrido@nmbu.no (email)
  • Gobakken, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: terje.gobakken@nmbu.no
  • Ørka, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: hans-ole.orka@nmbu.no
  • Næsset, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: erik.naesset@nmbu.no
  • Bollandsås, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: ole.martin.bollandsas@nmbu.no
article id 10392, category Research article
Paula Jylhä, Pasi Rikkonen, Katri Hamunen. (2020). Size matters – an analysis of business models and the financial performance of Finnish wood-harvesting companies. Silva Fennica vol. 54 no. 4 article id 10392. https://doi.org/10.14214/sf.10392
Keywords: profitability; business model; business model canvas; entrepreneur; success factor; wood-harvesting enterprise
Highlights: Economic success was related to company’s size, Small companies with a turnover of less than 600 000 € a–1 are struggling with profitability; Large enterprises continue to grow and innovate new business concepts; The competitive edge of large companies was reflected in large production capacity, efficient operations, versatile supply of services, and power in negotiations.
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The size of Finnish wood harvesting enterprises has grown, and entrepreneurs have become responsible for various additional tasks, resulting in networking with other harvesting enterprises of various sizes and suppliers of supporting services, but the profitability of the wood harvesting sector has remained low. In the present study, the financial performance of 83 wood harvesting companies in Eastern and Northern Finland was evaluated, based on public final account data from a five-year period between 2013 and 2017. The factors underlying economic success were identified based on 19 semi-structured entrepreneur interviews. The Business Model Canvas framework was applied in the analyses. In particular, the smallest companies (with an annual turnover of less than 600 000 €) struggled with profitability. They showed increasing indebtedness, suffered from poor power in negotiations, had typically short-term contracts, and faced difficulties in retaining skilled operators. Most of the small companies were subcontractors of larger wood-harvesting companies. The better economic success of larger companies was likely based on their capacity to provide wood harvesting services in large volumes and supply versatile services, power in negotiations, and more cost-effective operations. The future development of wood harvesting seems to be polarised: larger enterprises are likely to continue growing, while the size of smaller enterprises has stabilised. Enhancing business management skills and practices is required in enterprises of all size groups.

  • Jylhä, Natural Resources Institute Finland (Luke), Production systems, Teknologiakatu 7, FI-67100 Kokkola, Finland E-mail: paula.jylha@luke.fi (email)
  • Rikkonen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Lönnrotinkatu 7, FI-50100 Mikkeli, Finland E-mail: pasi.rikkonen@luke.fi
  • Hamunen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Yliopistokatu 6B, FI-80100 Joensuu, Finland E-mail: katri.hamunen@luke.fi
article id 10385, category Research article
Perttu Anttila, Tuomas Nummelin, Kari Väätäinen, Juha Laitila. (2020). The effect of winter weather on timber truck tare weights. Silva Fennica vol. 54 no. 4 article id 10385. https://doi.org/10.14214/sf.10385
Keywords: forestry logistics; payload; tare weight; weather data
Highlights: In a one-year follow-up study of eight timber trucks in Finland, up to 6–12% increase in tare weights was observed in wintertime; Compared with summertime tare weights, the maximum increase varied between 1560 kg and 3100 kg; Potential payload loss was explained by temperature, relative humidity and precipitation.
Abstract | Full text in HTML | Full text in PDF | Author Info

In wintertime, the payload capacity of a timber truck is reduced by snow that accumulates on the structures of the truck. The aim of this study was to quantify the potential payload loss due to snow and winter accessories and to predict the loss with weather variables. Tare weights of eight timber trucks were collected at mill receptions in Finland over a one-year period. Monthly and annual loss of potential payload was estimated using the tare measurements in summer months as a reference. Each load was also connected with weather data at the location and time of delivery and payload loss explained by the weather data with the aid of regression models. The maximum loss of payload varied between 1560 kg and 3100 kg. On a monthly basis, the highest losses occurred in January, when the median values varied between 760 kg and 2180 kg. Over the year, the payload loss ranged between the trucks from 0.5% to 1.5% (from 1.9% and 5.1% in January) of the total number of loads in the study. Payload loss was found to increase with decreasing temperature, increasing relative humidity and increasing precipitation. Although the average payload loss was not very high, the biggest losses occur just during the season of highest capacity utilization. Big differences were also found in the tare weights between the trucks. The results of the study give incentive to develop truck and trailer structures that reduce the adherence of snow.

  • Anttila, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0002-6131-392X E-mail: perttu.anttila@luke.fi (email)
  • Nummelin, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: tuomas.nummelin@luke.fi
  • Väätäinen, Natural Resources Institute Finland (Luke), Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: kari.vaatainen@luke.fi
  • Laitila, Natural Resources Institute Finland (Luke), Yliopistokatu 6, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0003-4431-3319 E-mail: juha.laitila@luke.fi
article id 10375, category Research article
Amy Wotherspoon, Nelson Thiffault, Robert L. Bradley. (2020). Resource availability and physiological response of black spruce to scarification in two climatic regions of Québec (Canada). Silva Fennica vol. 54 no. 4 article id 10375. https://doi.org/10.14214/sf.10375
Keywords: Picea mariana; foliar δ18O; foliar δ13C; needle gas exchange; photosynthetic nitrogen use efficiency; water use efficiency
Highlights: Mechanical scarification of clearcut sites in two climatic regions maintained high growth rates of regenerating black spruce 18 years after treatment; In both regions, scarification reduced soil moisture and increased water use efficiency; In the cooler-humid region, scarification improved soil nitrogen mineralization.
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Scarification is a mechanical site preparation technique designed to create microsites that will favor the growth of planted tree seedlings after clearcutting. However, the positive growth response of black spruce (Picea mariana (Mill.) Britton, Sterns & Poggenb.) to scarification varies across different sites. We hypothesized that this was due to different forms of physiological stress induced by different climates or by the severity of competition from ericaceous shrubs. We thus compared the effects of scarification on black spruce needle gas exchange and other foliar properties, as well as on indices of soil water and nitrogen availability, in relatively warm-dry (Abitibi) vs. cool-humid (Côte-Nord) climates in the province of Québec (Canada). We found a similar positive effect of scarification on tree height in Abitibi and Côte-Nord. Scarification reduced soil moisture in both climatic regions, but increased soil N mineralization in Côte-Nord only. Accordingly, scarification increased the instantaneous water use efficiency in both climate regions, but decreased photosynthetic N use efficiency in Côte-Nord only. In both regions, we found a positive relationship between foliar δ18O and δ13C on scarified plots, providing further evidence that increased growth due to scarification depends on a decrease in stomatal conductance. We conclude that scarification increases total evapotranspiration of trees evenly across the east-to-west moisture gradient in the province of Québec, but also improves long-term soil nutritional quality in a cooler-humid climate.

  • Wotherspoon, Département de biologie, Université de Sherbrooke, Sherbrooke, Québec, J1K 2R1, Canada E-mail: amy.wotherspoon@usherbrooke.ca (email)
  • Thiffault, Canadian Wood Fibre Centre, Canadian Forest Service, Natural Resources Canada, 1055 du. P.E.P.S., P.O. Box 10380, Québec, G1V 4C7, Canada E-mail: nelson.thiffault@canada.ca
  • Bradley, Département de biologie, Université de Sherbrooke, Sherbrooke, Québec, J1K 2R1, Canada E-mail: robert.bradley@usherbrooke.ca
article id 10370, category Research article
Juha Lappi, Timo Pukkala. (2020). Analyzing ingrowth using zero-inflated negative binomial models. Silva Fennica vol. 54 no. 4 article id 10370. https://doi.org/10.14214/sf.10370
Keywords: regeneration; continuous cover forestry; count data; generalized linear model; overdispersion; right-censoring
Highlights: Models were developed to describe ingrowth in national forest inventory data; The data were more dispersed than Poisson data and included many zeros; Fixed-effects models had larger zero-inflation probability and overdispersion parameter than mixed-effect models; Mixed-effects models had larger likelihood than fixed-effects models but provided biased predictions; Prediction of right-censored ingrowth may be useful owing to large overdispersion.
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Ingrowth is an important element of stand dynamics in several silvicultural systems, especially in continuous cover forestry. Earlier predictive models for ingrowth in Finnish forests are few and not based on up-to-date statistical methods. Ingrowth is here defined as the number of trees over 1.3 m entering a plot. This study developed new ingrowth models for Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies (L.) H. Karst.) and birch (Betula pendula Roth and B. pubescens Ehrh.) using data from the permanent sample plots of the Finnish national forest inventory. The data were over-dispersed compared to a Poisson process and had many zeros. Therefore, a zero-inflated negative binomial model was used. The total and species-specific stand basal areas, temperature sum and fertility class were used as predictors in the ingrowth models. Both fixed-effects and mixed-effects models were fitted. The mixed-effects model versions included random plot effects. The mixed-effects models had larger likelihoods but provided biased predictions. Also censored prediction was considered where only a certain maximum number of ingrowth trees were accepted for a plot. The models predicted most pine ingrowth in pine-dominated stands on sub-xeric and xeric sites where stand basal area was low. The predicted amount of spruce ingrowth was maximized when the basal area of spruce was 13 m2 ha–1. Increasing temperature sum increased spruce ingrowth. Predicted birch ingrowth decreased with increasing stand basal area and towards low fertility classes. An admixture of pine increased the predicted amount of spruce ingrowth.

  • Lappi, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: juha.lappi.sjk@gmail.com (email)
  • Pukkala, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: timo.pukkala@uef.fi
article id 10347, category Research article
Matti Katila, Tuomas Rajala, Annika Kangas. (2020). Assessing local trends in indicators of ecosystem services with a time series of forest resource maps. Silva Fennica vol. 54 no. 4 article id 10347. https://doi.org/10.14214/sf.10347
Keywords: National Forest Inventory; satellite images; k-nearest neighbour estimation; Landsat; Mann-Kendall test; multitemporal inventory data; temporal trend analysis
Highlights: Untitled Document Contextual Mann-Kendall test detects significant trends in time-series of forest maps; Trends become more consistent as the areal unit size used for test input increases; Changes in different scales reflect different phenomena in forests; Significant trends were detected even after multiple testing correction.
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Since the 1990’s, forest resource maps and small area estimates have been produced by combining national forest inventory (NFI) field plot data, optical satellite images and numerical map data using a non-parametric k-nearest neighbour method. In Finland, thematic maps of forest variables have been produced by the means of multi-source NFI (MS-NFI) for eight to ten times depending on the geographical area, but the resulting time series have not been systematically utilized. The objective of this study was to explore the possibilities of the time series for monitoring the key ecosystem condition indicators for forests. To this end, a contextual Mann-Kendall (CMK) test was applied to detect trends in time-series of two decades of thematic maps. The usefulness of the observed trends may depend both on the scale of the phenomena themselves and the uncertainties involved in the maps. Thus, several spatial scales were tested: the MS-NFI maps at 16 × 16 m2 pixel size and units of 240 × 240 m2, 1200 × 1200 m2 and 12  000 × 12  000 m2 aggregated from the MS-NFI map data. The CMK test detected areas of significant increasing trends of mean volume on both study sites and at various unit sizes except for the original thematic map pixel size. For other variables such as the mean volume of tree species groups, the proportion of broadleaved tree species and the stand age, significant trends were mostly found only for the largest unit size, 12  000 × 12  000 m2. The multiple testing corrections decreased the amount of significant p-values from the CMK test strongly. The study showed that significant trends can be detected enabling indicators of ecosystem services to be monitored from a time-series of satellite image-based thematic forest maps.

  • Katila, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, FI-00790 Helsinki, Finland; ORCID https://orcid.org/0000-0001-6946-5736 E-mail: matti.katila@luke.fi (email)
  • Rajala, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: tuomas.rajala@luke.fi
  • Kangas, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Yliopistokatu 6, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0002-8637-5668 E-mail: annika.kangas@luke.fi
article id 10331, category Research article
Jussi Juola, Aarne Hovi, Miina Rautiainen. (2020). Multiangular spectra of tree bark for common boreal tree species in Europe. Silva Fennica vol. 54 no. 4 article id 10331. https://doi.org/10.14214/sf.10331
Keywords: classification; reflectance; hyperspectral; imaging spectrometer; near-infrared; SVM; visible
Highlights: Novel multiangular measurement set-up for hyperspectral imaging; Multiangular spectra of silver birch (Betula pendula), Scots pine (Pinus sylvestris) and Norway spruce (Picea abies) stem bark samples were collected; Intra- and interspecific variations in reflectance were analyzed; Demonstration of tree species identification based on stem bark spectra; Collected spectra openly available in SPECCHIO Spectral Information System.
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Despite the importance of spectral properties of woody tree structures, they are seldom represented in research related to forests, remote sensing, and reflectance modeling. This study presents a novel imaging multiangular measurement set-up that utilizes a mobile handheld hyperspectral camera (Specim IQ, 400–1000 nm), and can measure stem bark spectra in a controlled laboratory setting. We measured multiangular reflectance spectra of silver birch (Betula pendula Roth), Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.) stem bark, and demonstrated the potential of using bark spectra in identifying tree species using a Support Vector Machine (SVM) based approach. Intraspecific reflectance variability was the lowest in visible (400–700 nm), and the highest in near-infrared (700–1000 nm) wavelength regions. Interspecific variation was the largest in the red, red-edge and near-infrared spectral bands. Spatial variation of reflectance along the tree height and different sides of the stem (north and south) were found. Both birch and pine had increased reflectance in the forward-scattering directions for visible to near-infrared wavelength regions, whilst spruce displayed the same only for the visible wavelength region. In addition, spruce had increased reflectance in the backward-scattering directions. In spite of the intraspecific variations, SVM could identify tree species with 88.8% overall accuracy when using pixel-specific spectra, and with 97.2% overall accuracy when using mean spectra per image. Based on our results it is possible to identify common boreal tree species based on their stem bark spectra using images from mobile hyperspectral cameras.

  • Juola, Aalto University, School of Engineering, Department of Built Environment, P.O. Box 14100, FI-00760 Aalto, Finland ORCID https://orcid.org/0000-0002-6050-7247 E-mail: jussi.juola@aalto.fi (email)
  • Hovi, Aalto University, School of Engineering, Department of Built Environment, P.O. Box 14100, FI-00760 Aalto, Finland ORCID https://orcid.org/0000-0002-4384-5279 E-mail: aarne.hovi@aalto.fi
  • Rautiainen, Aalto University, School of Engineering, Department of Built Environment, P.O. Box 14100, FI-00760 Aalto, Finland; Aalto University, School of Electrical Engineering, Department of Electronics and Nanoengineering, P.O. Box 15500, FI-00760 Aalto, Finland ORCID https://orcid.org/0000-0002-6568-3258 E-mail: miina.a.rautiainen@aalto.fi
article id 10276, category Research article
Adas Marčiulynas, Vaida Sirgedaitė-Šėžienė, Povilas Žemaitis, Āris Jansons, Virgilijus Baliuckas. (2020). Resistance of Scots pine half-sib families to Heterobasidion annosum in progeny field trials. Silva Fennica vol. 54 no. 4 article id 10276. https://doi.org/10.14214/sf.10276
Keywords: Pinus sylvestris; field trials; half-sib families; root rot pathogen; total phenolic compounds; tree susceptibility
Highlights: Large genetic variation was estimated in the resistance of Scots pine half-sib families to root rot in field trials; A strong relationship was observed between family resistance to root rot and phenolic compound concentration in the wood.
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Five Scots pine (Pinus sylvestris L.) progeny field trials, each established in different Lithuanian regions of provenance in 1983, were studied. Each progeny field trial consists of 140 half-sib families from seven populations (20 families from each population). The evaluation was carried out in 2012 and 2018 to assess the families resistance to Heterobasidion annosum (Fr.) Bref. An index of resistance in the infected plots was calculated. To verify the accuracy of the method, total phenolic compounds (TPC) was chosen as key parameter to compare with the plant resistance index. During the six years between the two assessments, the percentage of living Scots pine trees in the progeny field trials decreased up to 20 percentage points (range: 4 p.p. to 20 p.p.). In 2018 the area of H. annosum damaged plots (in percentage from total field trial area) varied from 17 to 27%. Tree mortality in the trial correlates with site soil fertility – more fertile soils were distinguished by higher tree percentage loss and vice versa. Using analysis from combined data of all progeny trials, the family variance component reached 13.3 ± 2.2% and family heritability was 0.81. Family heritability estimates for root rot resistance show possibilities of high breeding effectiveness. The correlations between the trials in family resistance estimates were negligible (ranging from 0 to 0.28). The significant high correlation coefficient was determined between the resistance index and TPC concentration (r = 0.77, p = 0.0003). This allows us to assume that plant resistance is directly linked on TPC synthesis. The results indicate that the chosen methods of chemical resistance for identification of root rot-resistant genotypes are applicable for the selection of Scots pine half-sib families in the field trials with higher resistance to pathogens.

  • Marčiulynas, Institute of Forestry, Lithuanian Research Centre for Agriculture and Forestry, Liepų str. 1, LT-53101 Girionys, Kaunas District, Lithuania E-mail: adas.marciulynas@mi.lt (email)
  • Sirgedaitė-Šėžienė, Institute of Forestry, Lithuanian Research Centre for Agriculture and Forestry, Liepų str. 1, LT-53101 Girionys, Kaunas District, Lithuania E-mail: vaida.seziene@mi.lt
  • Žemaitis, Institute of Forestry, Lithuanian Research Centre for Agriculture and Forestry, Liepų str. 1, LT-53101 Girionys, Kaunas District, Lithuania E-mail: povilas.zemaitis@mi.lt
  • Jansons, Latvian State Forest Research Institute “Silava”, Department of Forest Tree Breeding, Rigas St.t. 111, Salaspils LV-2169, Latvia E-mail: aris.jansons@silava.lv
  • Baliuckas, Institute of Forestry, Lithuanian Research Centre for Agriculture and Forestry, Liepų str. 1, LT-53101 Girionys, Kaunas District, Lithuania E-mail: virgilijus.baliuckas@mi.lt
article id 10269, category Research article
Annika Kangas, Helena M. Henttonen, Timo P. Pitkänen, Sakari Sarkkola, Juha Heikkinen. (2020). Re-calibrating stem volume models – is there change in the tree trunk form from the 1970s to the 2010s in Finland? Silva Fennica vol. 54 no. 4 article id 10269. https://doi.org/10.14214/sf.10269
Keywords: TLS; OLS; regression; terrestrial laser scanning
Highlights: TLS data showed that trunk form has changed in Finland from the 1970s; Significant differences were observed for all tree species; The trees in TLS data are on average more slender than in the old data.
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The tree stem volume models of Norway spruce, Scots pine and silver and downy birch currently used in Finland are based on data collected during 1968–1972. These models include four different formulations of a volume model, with three different combinations of independent variables: 1) diameter at height of 1.3 m above ground (dbh), 2) dbh and tree height (h) and 3) dbh, h and upper diameter at height of 6 m (d6). In recent National Forest Inventories of Finland, a difference in the mean volume prediction between the models with and without the upper diameter as predictor has been observed. To analyze the causes of this difference, terrestrial laser scanning (TLS) was used to acquire a large dataset in Finland during 2017–2018. Field-measured predictors and volumes predicted using spline functions fitted to the TLS data were used to re-calibrate the current volume models. The trunk form is different in these two datasets. The form height is larger in the new data for all diameter classes, which indicates that the tree trunks are more slender than they used to be. One probable reason for this change is the increase in stand densities, which is at least partly due to changed forest management. In models with both dbh and h as predictors, the volume is smaller a given h class in the data new data than in the old data, and vice versa for the diameter classes. The differences between the old and new models were largest with pine and smallest with birch.

  • Kangas, Natural Resources Institute Finland (Luke), Yliopistokatu 6 B, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0002-8637-5668 E-mail: annika.kangas@luke.fi (email)
  • Henttonen, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: helena.henttonen@luke.fi
  • Pitkänen, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0001-5389-8713 E-mail: timo.p.pitkanen@luke.fi
  • Sarkkola, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: sakari.sarkkola@luke.fi
  • Heikkinen, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0003-3527-774X E-mail: juha.heikkinen@luke.fi
article id 10389, category Research article
Juho Matala, Harri Kilpeläinen, Henrik Heräjärvi, Tapio Wall, Erkki Verkasalo. (2020). Sawlog quality and tree dimensions of Scots pine 34 years after artificial moose browsing damage. Silva Fennica vol. 54 no. 3 article id 10389. https://doi.org/10.14214/sf.10389
Keywords: Pinus sylvestris; growth; timber quality; Alces alces; defect
Highlights: The first controlled, long-term, experimental study on the tree dimensions and sawlog quality after moose damage; The trees damaged at the seedling stage had a smaller diameter, height, and tree volume at the end of the experiment; The heavier the clipping treatment, the more likely the stem form deteriorated; Deteriorated stem form and vertical branches were the most typical defects.
Abstract | Full text in HTML | Full text in PDF | Author Info

Moose (Alces alces L.) browsing causes severe damage in Scots pine (Pinus sylvestris L.) seedling stands. The effects of this damage on the quality of sawlogs were studied in a long-term controlled experiment. This article reports the stem size and external quality characteristics of Scots pine stems 34 years after artificial moose browsing damage. Damaging the trees by clipping the main stem at the seedling stage reduced the diameter, height, and tree volume of the trees at the end of the experiment. The tree growth reduction was dependent on the severity of clipping. The differences between the damaged and the control trees were more obvious in diameter than in height at the time of final felling. Stem form defects and vertical branches were the most typical externally detectable defects caused by clipping. Defects in the butt logs were detected in 71–89% of the damaged trees, depending on the clipping treatment severity. The stronger the clipping treatment, the more likely the stem form was defected and the more commonly were vertical branches and crooks detected in the stems. The results indicate that both tree dimensions and stem quality suffer from moose browsing. The findings of this controlled experiment more likely underestimate than overestimate the damage in comparison to real moose browsing. Further analyses are required to assess the effects of browsing damage on the internal quality of sawlogs and subsequent economic outcomes.

  • Matala, Natural Resources Institute Finland (Luke), Natural resources, Yliopistokatu 6B, FI-80101 Joensuu, Finland E-mail: juho.matala@luke.fi (email)
  • Kilpeläinen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Yliopistokatu 6B, FI-80101 Joensuu, Finland E-mail: harri.kilpelainen@luke.fi
  • Heräjärvi, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6B, FI-80101 Joensuu, Finland E-mail: henrik.herajarvi@luke.fi
  • Wall, Natural Resources Institute Finland (Luke), Research infrastructure services, Yliopistokatu 6B, FI-80101 Joensuu, Finland E-mail: tapio.wall@luke.fi
  • Verkasalo, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6B, FI-80101 Joensuu, Finland E-mail: erkki.verkasalo@luke.fi
article id 10373, category Research article
Chintan Sheth, Aparajita Datta, Devathi Parashuram. (2020). Persistent loss of biologically-rich tropical forests in the Indian Eastern Himalaya. Silva Fennica vol. 54 no. 3 article id 10373. https://doi.org/10.14214/sf.10373
Keywords: deforestation; Arunachal Pradesh; biodiversity hotspot; Buceros bicornis; hornbill nest trees; illegal logging; north-east India
Highlights: We found a high rate of deforestation occurring in a state managed reserve forest in Indian Eastern Himalaya; Fine-scale analysis showed considerable forest loss around nesting trees for hornbills; Forest monitoring, protection and honest governance are required to effectively protect forests in the Eastern Himalaya biodiversity hotspot.
Abstract | Full text in HTML | Full text in PDF | Author Info

Using fine-resolution satellite imagery from multiple satellite data products, we assessed the change in forest cover of a state-managed Reserve Forest (RF) located in India’s Eastern Himalaya biodiversity hot-spot. 4.6% of forest cover was lost from Papum RF between 2013 and 2017 at the rate of 8.2 km2 year–1. Three species of hornbills: Great Hornbill Buceros bicornis Linnaeus, 1758, Wreathed Hornbill Rhyticeros undulatus (Shaw, 1811) and Oriental Pied Hornbill Anthracoceros albirostris (Shaw, 1808), that are functionally important are found here with nesting habitat in the areas affected by illegal logging. Therefore, we assessed the habitat loss within a 1 km radius around 29 nest trees. From 2011 to 2019, forest cover declined from 38.55 km2 to 21.94 km2 around these hornbill nest trees. Illegal logging is the main driver that is depleting forest cover within this important bird area. Our results highlight the ongoing threats to biologically-rich forests and the need for urgent measures to halt this loss. We suggest that this study has practical implications for the monitoring and governance of state-managed forests in Arunachal Pradesh.

  • Sheth, #4 Ananda Nilaya, 4th Main Road, Siddhivinayaka Layout, Bengaluru 560097, Karnataka, India E-mail: chintz604@gmail.com
  • Datta, Nature Conservation Foundation, 1311,“Amritha”, 12th Main, Vijayanagar 1st Stage, Mysore 570017, Karnataka, India E-mail: aparajita@ncf-india.org (email)
  • Parashuram, Nature Conservation Foundation, 1311,“Amritha”, 12th Main, Vijayanagar 1st Stage, Mysore 570017, Karnataka, India; Department of Geography, University of Cambridge, CB2 3EN, United Kingdom E-mail: dp608@cam.ac.uk
article id 10355, category Research article
Grzegorz Szewczyk, Raffaele Spinelli, Natascia Magagnotti, Paweł Tylek, Janusz M. Sowa, Piotr Rudy, Dominika Gaj-Gielarowiec. (2020). The mental workload of harvester operators working in steep terrain conditions. Silva Fennica vol. 54 no. 3 article id 10355. https://doi.org/10.14214/sf.10355
Keywords: logging; ergonomics; eye tracking; fatigue; gaze behaviour
Highlights: We analysed a harvester operator’s perception of his work environment as slope gradient increased; Increasingly difficult work conditions (steeper terrain) were reflected by increased eyeball activity; Fixation duration increased with slope gradient, while the duration of saccades was shorter; Variation of the eyeball movement cycles was also related to work difficulty caused by slope gradient.
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The use of modern multi-functional forestry machines has already been associated with central nervous system fatigue induced by high mental workload. As these machines are being used under increasingly difficult terrain conditions, further knowledge is required on the expected aggravation of operators’ mental workload, so that suitable work/rest schedules can be developed. Within such a context, the aim of this study was to gauge aggravations of mental workload derived from increasing slope gradient. Measurements of eye activity were obtained from a representative harvester operator working in corridors with the following mean inclinations: 9%, 23% and 47%. The duration, frequency and trajectory of eye movements were used to determine the harvester operator’s mental workload, on the assumption that worsening work conditions would be reflected by increased eyeball activity. The number of fixations during the performance of all tasks increased with the increasing slope gradient. Similarly, fixation duration increased with slope gradient. The mean duration of saccades when working on a 23% slope was 5% shorter compared to work under a 9% gradient. A further significant shortening of saccade duration (~22%) occurred when working on a 47% slope. The good match between eye activity cycles and work cycles, visible especially on steep slopes, indicates that mental workload is related to work conditions. Overall, operating a forest harvester on steep slopes results in a greatly increased mental workload and calls for suitable rest schedules.

  • Szewczyk, University of Agriculture in Krakow; al. 29 Listopada 46, 31-425 Kraków, Poland E-mail: rlszewcz@cyf-kr.edu.pl (email)
  • Spinelli, National Research Council of Italy, Firenze, via Madonna del Piano, 10 50019 – Sesto Fiorentino, Italy E-mail: spinelli@ivalsa.cnr.it
  • Magagnotti, National Research Council of Italy, Firenze, via Madonna del Piano, 10 50019 – Sesto Fiorentino, Italy E-mail: natascia.magagnotti@ibe.cnr.it
  • Tylek, University of Agriculture in Krakow; al. 29 Listopada 46, 31-425 Kraków, Poland E-mail: rltylek@cyf-kr.edu.pl
  • Sowa, University of Agriculture in Krakow; al. 29 Listopada 46, 31-425 Kraków, Poland E-mail: janusz.sowa@urk.edu.pl
  • Rudy, State Forests, Biłgoraj Forest District, ul. Zamojska 96, 23-400 Biłgoraj, Poland E-mail: rudy.piotrek@o2.pl
  • Gaj-Gielarowiec, State Forests, Barycz Forest District, Barycz 69, 26-200 Końskie, Poland E-mail: dominika.gaj@gmail.com
article id 10341, category Research article
Arta Bārdule, Edgars Jūrmalis, Zane Lībiete, Ilze Pauliņa, Jānis Donis, Agita Treimane. (2020). Use of retail market data to assess prices and flows of non-wood forest products in Latvia. Silva Fennica vol. 54 no. 3 article id 10341. https://doi.org/10.14214/sf.10341
Keywords: forest berries; mushrooms; provisional ecosystem services; retail prices; spatial flows of NWFPs; tree sap
Highlights: Retail prices of non-wood forest products (NWFPs) may be used to study lifestyle-related consumption patterns; While retail sales of NWFPs may increase household budgets, this source of income is highly variable due to varying meteorological conditions; NWFP retail price analysis illustrates aspect of household economies not recorded in official statistics and cash flows of declared income.
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In northern Europe, largest part of non-wood forest products (NWFPs) are gathered for recreational purposes and household consumption, but considerable amount of forest berries and mushrooms are sold as well. Retail market, largely invisible for the official statistics, reveals the lifestyle-related aspects of NWFP trade and may help to understand the flows of this ecosystem service when information on wholesale trade is inaccessible. The prices and flows of most common NWFPs – edible berries, mushrooms and tree sap – in the retail market in Latvia in 2017 and 2018 were analysed based on direct interviews with the sellers in marketplaces and telephone interviews with online retailers. The mean retail prices of NWFPs were compared between statistical regions and years and correlated with socio-economic data and forest characteristics. The directions of the NWFP flows were analysed according to the place of origin and place of retail sales. The highest prices were recorded for stinkhorn (Phallus impudicus Pers.) and Boletes spp. among mushrooms, for wild strawberries (Fragaria vesca L.) among berries and for maple (Acer platanoides L.) sap in the product group of tree sap. The retail price of the same products differed between years, most likely due to the product availability, largely caused by meteorological conditions. In more than half of the cases of recorded sales, NWFPs were consumed in the same region as they were gathered. For other cases of sales, the capital, Rīga, was the main service benefitting area of NWFP retail trade, and the largest part of the products originated from the two closest statistical regions.

  • Bārdule, Latvian State Forest Research Institute “Silava”, Rigas str. 111, Salaspils, Latvia, LV-2169 E-mail: arta.bardule@silava.lv (email)
  • Jūrmalis, Latvian State Forest Research Institute “Silava”, Rigas str. 111, Salaspils, Latvia, LV-2169 E-mail: edgars.jurmalis@silava.lv
  • Lībiete, Latvian State Forest Research Institute “Silava”, Rigas str. 111, Salaspils, Latvia, LV-2169 E-mail: zane.libiete@silava.lv
  • Pauliņa, Latvian State Forest Research Institute “Silava”, Rigas str. 111, Salaspils, Latvia, LV-2169 E-mail: paulina.ilze@gmail.com
  • Donis, Latvian State Forest Research Institute “Silava”, Rigas str. 111, Salaspils, Latvia, LV-2169 E-mail: janis.donis@silava.lv
  • Treimane, Latvian State Forest Research Institute “Silava”, Rigas str. 111, Salaspils, Latvia, LV-2169; University of Latvia, Jelgavas str. 1, Riga, Latvia, LV-1004 E-mail: agita.treimane@silava.lv
article id 10284, category Research article
Togtokhbayar Erdene-Ochir , Futoshi Ishiguri, Ikumi Nezu, Bayasaa Tumenjargal, Bayartsesteg Baasan, Ganbaatar Chultem, Jyunichi Ohshima, Shinso Yokota. (2020). Utilization potential of naturally regenerated Mongolian Betula platyphylla wood based on growth characteristics and wood properties. Silva Fennica vol. 54 no. 3 article id 10284. https://doi.org/10.14214/sf.10284
Keywords: basic density; stress-wave velocity; compressive strength; early evaluation
Highlights: Wood with high density and high compressive strength can be obtained near to the bark of Betula platyphylla trees; Basic density of B. platyphylla wood may be predicted by the stress-wave velocity of stems; Growth characteristics are positively correlated with stress-wave velocity and basic density; Early evaluation of basic density is possible when using the wood at 2 cm from the pith: A significant between-site variation was found in the basic density at the position from the 1st to the 15th annual ring from the pith; Wood from B. platyphylla trees grown in Mongolia may be used for industrial products equally to the same species in other countries.
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To preliminary evaluate the potential wood utilization of Betula platyphylla Sukaczev trees naturally regenerated in Mongolia, growth characteristics (stem diameter and tree height), wood properties (annual ring width, basic density, and compressive strength parallel to grain at the green condition) of core samples, and stress-wave velocity in stems were investigated for Betula platyphylla trees grown naturally in three different sites in Selenge, Mongolia. Betula platyphylla trees, naturally grown in Nikko, Japan, were also examined to compare wood properties between the two regions. The mean values of stem diameter, tree height, stress-wave velocity of stems, annual ring width, basic density, and compressive strength parallel to grain at green condition in Mongolian B. platyphylla were 17.6 cm, 14.1 m, 3.50 km s–1, 1.27 mm, 0.51 g cm–3, and 20.4 MPa, respectively. Basic density and compressive strength were decreased first from the pith, and then gradually increased toward the bark. The wood properties of B. platyphylla trees grown naturally in Mongolia were similar to those in B. platyphylla trees grown in Japan. Growth characteristics, especially stem diameter, were positively correlated with the stress-wave velocity of stems and basic density. Early evaluation of basic density in B. platyphylla trees is possible by using wood located 2 cm from the pith. Basic density at the position from the 1st to the 15th annual ring from the pith showed significant between-site differences in Mongolian B. platyphylla. Based on the results, it is concluded that the wood of B. platyphylla trees grown in Mongolia may be used for industrial products as well as those from similar species in other countries.

  • Erdene-Ochir , School of Agriculture, Utsunomiya University, Utsunomiya, Tochigi 321-8505, Japan; United Graduate School of Agricultural Science, Tokyo University of Agriculture and Technology, Fuchu, Tokyo 183-8509, Japan; Training and Research Institute of Forestry and Wood Industry, Mongolian University of Science and Technology, Ulaanbaatar 14191, Mongolia E-mail: togtokhbayarerdeneochir@gmail.com
  • Ishiguri, School of Agriculture, Utsunomiya University, Utsunomiya, Tochigi 321-8505, Japan E-mail: ishiguri@cc.utsunomiya-u.ac.jp (email)
  • Nezu, School of Agriculture, Utsunomiya University, Utsunomiya, Tochigi 321-8505, Japan E-mail: zoo-sk3.primo@outlook.jp
  • Tumenjargal, School of Agriculture, Utsunomiya University, Utsunomiya, Tochigi 321-8505, Japan; United Graduate School of Agricultural Science, Tokyo University of Agriculture and Technology, Fuchu, Tokyo 183-8509, Japan; Training and Research Institute of Forestry and Wood Industry, Mongolian University of Science and Technology, Ulaanbaatar 14191, Mongolia E-mail: t_bayasaa88@yahoo.com
  • Baasan, Training and Research Institute of Forestry and Wood Industry, Mongolian University of Science and Technology, Ulaanbaatar 14191, Mongolia E-mail: bayartsetseg@must.edu.mn
  • Chultem, Training and Research Institute of Forestry and Wood Industry, Mongolian University of Science and Technology, Ulaanbaatar 14191, Mongolia E-mail: ganbaatar_ch@must.edu.mn
  • Ohshima, School of Agriculture, Utsunomiya University, Utsunomiya, Tochigi 321-8505, Japan E-mail: joshima@cc.utsunomiya-u.ac.jp
  • Yokota, School of Agriculture, Utsunomiya University, Utsunomiya, Tochigi 321-8505, Japan E-mail: yokotas@cc.utsunomiya-u.ac.jp
article id 10273, category Research article
Lasse Aro, Anssi Ahtikoski, Jyrki Hytönen. (2020). Profitability of growing Scots pine on cutaway peatlands. Silva Fennica vol. 54 no. 3 article id 10273. https://doi.org/10.14214/sf.10273
Keywords: Pinus sylvestris; afforestation; bare land value; financial performance; peat cutaway area; stand-level optimization
Highlights: The potential wood production on cutaway peatlands can be high; Afforestation with seeding or planting was profitable at 3% rate of interest; In northern Finland, the afforestation method, ditch spacing and fertilization affected the bare land value.
Abstract | Full text in HTML | Full text in PDF | Author Info

A major after-use option for former peat harvesting areas has been afforestation. The profitability of afforestation with Scots pine trees (Pinus sylvestris L.) was studied in two 31–32-year old experiments in southern and northern Finland. The stands were established by seeding and planting, and various fertilization treatments and drainage intensities were tested. The financial performance for each plot was assessed in three steps. First, the costs occurred during the measurement time were summed up according to their present value. Then, for the rest of the rotation (i.e., from the age of 31/32 onwards) the stand management was optimized in order to maximize the net present value (MaxNPV). Finally, bare land values (BLVs) were calculated by summing up the present value of costs and the MaxNPV and converting the sum of the series into infinity. The afforestation method did not affect the mean annual increment (MAI; 9.2–9.5 m3 ha–1 a–1) in the southern experiment. In the northern experiment the afforestation method, ditch spacing and fertilization had significant effects on the MAI of the stands. The average MAI of the planted pines was 8.9 m3 ha–1 a–1, and for seeded pines it was 7.5 m3 ha–1 a–1. The BLV at an interest rate of 3% was positive for all stands in both regions. In the northern region afforestation method, ditch spacing and fertilization also had a significant effect on the BLV. When the interest rate was 5%, almost two thirds of the stands had a negative BLV in both regions.

  • Aro, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Itäinen Pitkäkatu 4A, FI-20520 Turku, Finland E-mail: lasse.aro@luke.fi (email)
  • Ahtikoski, Natural Resources Institute Finland (Luke), Natural resources, Paavo Havaksentie 3, FI-90570 Oulu, Finland ORCID https://orcid.org/0000-0003-1658-3813 E-mail: anssi.ahtikoski@luke.fi
  • Hytönen, Natural Resources Institute Finland (Luke), Natural resources, Teknologiakatu 7, FI-67100 Kokkola, Finland ORCID http://orcid.org/0000-0001-8475-3568 E-mail: jyrki.hytonen@luke.fi
article id 10205, category Research article
Michel Soucy, Martin Béland. (2020). A crop tree release variant of precommercial thinning using a backpack mounted chain saw. Silva Fennica vol. 54 no. 3 article id 10205. https://doi.org/10.14214/sf.10205
Keywords: thinning; hardwood; spacing; time study; brushsaw; motor-manual; work performance
Highlights: Backpack mounted chain saws offer an opportunity to effectively cut larger stems in precommercial thinning treatments compared to conventional circular blade brushsaws; Productivity in performing precommercial crop tree release (PCTR) in a hardwood stand using a backpack mounted chain saw varied between 0.22 to 0.47 ha h–1.
Abstract | Full text in HTML | Full text in PDF | Author Info

Although crop tree release (CTR) in hardwood stands is an accepted variant of precommercial thinning (PCT), the lack of an affordable and feasible method hinders its adoption. CTR implies selecting between 150 and 500 trees ha–1 when trees are between 7 and 12 m high and cutting only stems competing with the target crop trees. We performed a field trial of a CTR variant of PCT in a 27.8 ha hardwood stand using a backpack mounted chain saw. A detailed time study was performed to document the trial over 13 days. Compared to conventional PCT performed earlier in the life of a stand, precommercial crop tree release required cutting larger stems, which showed to be feasible and productive using a backpack mounted chain saw. Productivity varied between 0.22 to 0.47 ha h–1 during the trial, Although productivity could vary with stand characteristics and worker, this proof of concept trial demonstrates some of the potential uses that this new saw configuration offers and sets the basis for an eventual larger scale deployment of this treatment.

article id 10270, category Research article
Aarne Hovi, Matti Mõttus, Jussi Juola, Farshid Manoocheri, Erkki Ikonen, Miina Rautiainen. (2020). Evaluating the performance of a double integrating sphere in measurement of reflectance, transmittance, and albedo of coniferous needles. Silva Fennica vol. 54 no. 2 article id 10270. https://doi.org/10.14214/sf.10270
Keywords: vegetation; albedo; reflectance; transmittance; needle carrier; spectra
Highlights: Adaptation of a compact double integrating sphere for spectral measurements of coniferous needles; Double integrating sphere is fast to operate and suitable for monitoring purposes and collection of large spectral databases; Measured spectra showed negative bias, which could potentially be reduced by building an optimized measurement setup.
Abstract | Full text in HTML | Full text in PDF | Author Info

Leaf reflectance and transmittance spectra are essential information in many applications such as developing remote sensing methods, computing shortwave energy balance (albedo) of forest canopies, and monitoring health or stress of trees. Measurement of coniferous needle spectra has usually been carried out with single integrating spheres, which has involved a lot of tedious manual work. A small double integrating sphere would make the measurements considerably faster, because of its ease of operation and small sample sizes required. Here we applied a compact double integrating sphere setup, used previously for measurement of broad leaves, for measurement of coniferous needles. Test measurements with the double integrating sphere showed relative underestimation of needle albedo by 5–39% compared to a well-established single integrating sphere setup. A small part of the bias can be explained by the bias of the single sphere. Yet the observed bias is quite significant if absolute accuracy of measurements is required. For relative measurements, e.g. for monitoring development of needle spectra over time, the double sphere system provides notable improvement. Furthermore, it might be possible to reduce the bias by building an optimized measurement setup that minimizes absorption losses in the sample port. Our study indicates that double spheres, after some technical improvement, may provide a new and fast way to collect extensive spectral libraries of tree species.

  • Hovi, Aalto University, School of Engineering, Department of Built Environment, P.O.Box 14100, FI-00760 Aalto, Finland ORCID https://orcid.org/0000-0002-4384-5279 E-mail: aarne.hovi@aalto.fi (email)
  • Mõttus, VTT Technical Research Centre Finland, P.O. Box 1000, FI-02044 VTT, Finland ORCID https://orcid.org/0000-0002-2745-1966 E-mail: matti.mottus@gmail.com
  • Juola, Aalto University, School of Engineering, Department of Built Environment, P.O.Box 14100, FI-00760 Aalto, Finland E-mail: jussi.juola@aalto.fi
  • Manoocheri, Aalto University, School of Electrical Engineering, Metrology Research Institute, Maarintie 8, FI-02150 Espoo, Finland ORCID https://orcid.org/0000-0003-3935-3930 E-mail: farshid.manoocheri@aalto.fi
  • Ikonen, VTT Technical Research Centre Finland, P.O. Box 1000, FI-02044 VTT, Finland; Aalto University, School of Electrical Engineering, Metrology Research Institute, Maarintie 8, FI-02150 Espoo, Finland ORCID https://orcid.org/0000-0001-6444-5330 E-mail: erkki.ikonen@aalto.fi
  • Rautiainen, Aalto University, School of Engineering, Department of Built Environment, P.O.Box 14100, FI-00760 Aalto, Finland; Aalto University, School of Electrical Engineering, Department of Electronics and Nanoengineering, P.O. Box 15500, FI-00760 Aalto, Finland ORCID https://orcid.org/0000-0002-6568-3258 E-mail: miina.a.rautiainen@aalto.fi
article id 10259, category Research article
Jyrki Hytönen, Hannu Hökkä. (2020). Comparison of granulated and loose ash in fertilisation of Scots pine on peatland. Silva Fennica vol. 54 no. 2 article id 10259. https://doi.org/10.14214/sf.10259
Keywords: Pinus sylvestris; wood ash; nutrition; fertilisation; granulation; PK fertiliser
Highlights: Granulated ash and commercial PK fertilizer increased stand growth in similar way during 15-year study period; Loose ash gave stronger and faster response than granulated ash.
Abstract | Full text in HTML | Full text in PDF | Author Info

The effects of wood ash fertilisation on tree nutrition and growth on forested peatlands has been studied using loose ash, but in practice, ash fertilisation is done almost exclusively with granulated ash. In this study, the effects of granulated ash and loose ash (both 5 Mg ha–1) on the growth and nutrition of Scots pine (Pinus sylvestris L.) stands were compared between a nitrogen-poor and a nitrogen-rich site over 15 years. On the nitrogen-rich site, wood ash application was also compared with commercial PK fertilisation. On the nitrogen-rich site, mean stand volume growth increase over unfertilised control treatment during the 15 year study period using granulated ash and commercial PK fertiliser was of the same magnitude (on average, 2.2–2.3 m3 ha–1 a–1). However, when loose ash was used growth increase over control was higher (3.7 m3 ha–1 a–1). On the nitrogen-poor site, the mean growth increase gained by loose or granulated ash (1.4–1.5 m3 ha–1 a–1) over the unfertilised control treatment was not significant. Fertilisation with loose ash or PK increased foliar P, K and B concentrations already in the first or second growing season, following fertilisation on both sites. Granulated ash increased foliar P concentrations on the nitrogen-rich site less than loose ash. After an initial increase, foliar P, K and B concentrations decreased at the end of study period. On the nitrogen-poor site, foliar P concentrations were below the deficiency limit by the end of the study period.

  • Hytönen, Natural Resources Institute Finland, Natural Resources, Teknologiakatu 7, FI-67100 Kokkola, Finland ORCID https://orcid.org/0000-0001-8475-3568 E-mail: jyrki.hytonen@luke.fi (email)
  • Hökkä, Natural Resources Institute Finland, Natural Resources, Paavo Havaksentie 3, FI-90570 Oulu, Finland E-mail: hannu.hokka@luke.fi
article id 10247, category Research article
Agnese Marcelli, Walter Mattioli, Nicola Puletti, Francesco Chianucci, Damiano Gianelle, Mirko Grotti, Gherardo Chirici, Giovanni D' Amico, Saverio Francini, Davide Travaglini, Lorenzo Fattorini, Piermaria Corona. (2020). Large-scale two-phase estimation of wood production by poplar plantations exploiting Sentinel-2 data as auxiliary information. Silva Fennica vol. 54 no. 2 article id 10247. https://doi.org/10.14214/sf.10247
Keywords: national forest inventories; Sentinel-2; design-based inference; first-phase tessellation stratified sampling; regression estimator; second-phase stratified sampling; simulation study
Highlights: A two-phase sampling for large-scale assessment of fast-growing forest crops is developed; Vegetation indices from Sentinel-2 are exploited in a linear regression estimator; The linear regression estimator turns out to be better than the estimator based on the sole sample information; The approach represents a reference for supporting outside-forest resource monitoring and assessment.
Abstract | Full text in HTML | Full text in PDF | Author Info

Growing demand for wood products, combined with efforts to conserve natural forests, have supported a steady increase in the global extent of planted forests. Here, a two-phase sampling strategy for large-scale assessment of the total area and the total wood volume of fast-growing forest tree crops within agricultural land is presented. The first phase is performed using tessellation stratified sampling on high-resolution remotely sensed imagery and is sufficient for estimating the total area of plantations by means of a Monte Carlo integration estimator. The second phase is performed using stratified sampling of the plantations selected in the first phase and is aimed at estimating total wood volume by means of an approximation of the first-phase Horvitz-Thompson estimator. Vegetation indices from Sentinel-2 are exploited as freely available auxiliary information in a linear regression estimator to improve the design-based precision of the estimator based on the sole sample data. Estimators of the totals and of the design-based variances of total estimators are presented. A simulation study is developed in order to check the design-based performance of the two alternative estimators under several artificial distributions supposed for poplar plantations (random, clustered, spatially trended). An application in Northern Italy is also reported. The regression estimator turns out to be invariably better than that based on the sole sample information. Possible integrations of the proposed sampling scheme with conventional national forest inventories adopting tessellation stratified sampling in the first phase are discussed.

  • Marcelli, University of Tuscia, Department for Innovation in Biological, Agro-food and Forest systems, Viterbo, Italy; Fondazione Edmund Mach, Department of Sustainable Agro-Ecosystems and Bioresources, Research and Innovation Centre, San Michele all’Adige, Italy E-mail: agnese.marcelli@student.unisi.it (email)
  • Mattioli, University of Tuscia, Department for Innovation in Biological, Agro-food and Forest systems, Viterbo, Italy; CREA, Research Centre for Forestry and Wood, Arezzo, Italy E-mail: walter.mattioli@crea.gov.it
  • Puletti, CREA, Research Centre for Forestry and Wood, Arezzo, Italy E-mail: nicola.puletti@crea.gov.it
  • Chianucci, CREA, Research Centre for Forestry and Wood, Arezzo, Italy E-mail: fchianucci@gmail.com
  • Gianelle, Fondazione Edmund Mach, Department of Sustainable Agro-Ecosystems and Bioresources, Research and Innovation Centre, San Michele all’Adige, Italy E-mail: damiano.gianelle@fmach.it
  • Grotti, CREA, Research Centre for Forestry and Wood, Arezzo, Italy; University of Roma La Sapienza, Department of Architecture and Design, Rome, Italy E-mail: mirkogrotti@gmail.com
  • Chirici, University of Firenze, Department of Agriculture, Food, Environment and Forestry, Florence, Italy E-mail: gherardo.chirici@unifi.it
  • D' Amico, University of Firenze, Department of Agriculture, Food, Environment and Forestry, Florence, Italy E-mail: giovanni.damico@unifi.it
  • Francini, University of Firenze, Department of Agriculture, Food, Environment and Forestry, Florence, Italy; University of Molise, Department of Agricultural, Environmental and Food Sciences, Campobasso, Italy E-mail: saverio.francini@gmail.com
  • Travaglini, University of Firenze, Department of Agriculture, Food, Environment and Forestry, Florence, Italy E-mail: davide.travaglini@unifi.it
  • Fattorini, University of Siena, Department of Economics and Statistics, Siena, Italy E-mail: lorenzo.fattorini@unisi.it
  • Corona, CREA, Research Centre for Forestry and Wood, Arezzo, Italy E-mail: piermaria.corona@crea.gov.it
article id 10220, category Research article
Arnis Gailis, Pauls Zeltiņš, Andis Purviņš, Juris Augustovs, Valts Vīndedzis, Inga Zariņa, Āris Jansons. (2020). Genetic parameters of growth and quality traits in open-pollinated silver birch progeny tests. Silva Fennica vol. 54 no. 2 article id 10220. https://doi.org/10.14214/sf.10220
Keywords: Betula pendula; tree breeding; genetic gain; genetic parameters
Highlights: Growth and stem quality traits were under strong genetic control; Weak genetic correlations between tree growth and stem quality were found; Strong age-age and type-B correlations suggest robust improvement over time and different environments; Simultaneous improvement of growth and stem quality might be applicable.
Abstract | Full text in HTML | Full text in PDF | Author Info

Genetic parameters of growth and stem quality traits were estimated for open-pollinated silver birch Betula pendula Roth progenies in Latvia at the age of 10 and 14 years. Tree height and stem volume were found to be under strong genetic control at both inventories (narrow-sense heritabilities varied from 0.41 to 0.66). Mainly low heritabilities were found for stem defects, yet genetic control of branch diameter, stem straightness and overall stem quality varied from low to high depending on study site. High additive genetic coefficient of variation was found for stem volume (25.3–32.5%). Genetic correlations among growth traits were strong and positive (0.90–0.99). Mainly weak genetic correlations between growth and quality traits implied simultaneous improvement. Still, strong negative correlations between branch angle and stem straightness might result in enlarged knot size for straighter logs. The genetic age-age correlations were strong. Weak genotype by environment interaction and stability of best genotypes over different sites was indicated by strong genetic correlations between trials. Each growth or quality trait alone showed substantial improvement in terms of estimated genetic gain (up to 62% over trial mean for stem volume). Therefore, selection index combining both growth and stem quality may be developed.

  • Gailis, Latvian State Forest Research Institute Silava, 111 Rigas street, Salaspils LV-2169, Latvia E-mail: arnis.gailis@silava.lv
  • Zeltiņš, Latvian State Forest Research Institute Silava, 111 Rigas street, Salaspils LV-2169, Latvia ORCID https://orcid.org/0000-0002-6286-5814 E-mail: pauls.zeltins@silava.lv (email)
  • Purviņš, Latvian State Forest Research Institute Silava, 111 Rigas street, Salaspils LV-2169, Latvia E-mail: andis.purvins@silava.lv
  • Augustovs, Latvian State Forest Research Institute Silava, 111 Rigas street, Salaspils LV-2169, Latvia E-mail: juris.augustovs@silava.lv
  • Vīndedzis, Latvian State Forest Research Institute Silava, 111 Rigas street, Salaspils LV-2169, Latvia E-mail: valts.vindedzis@silava.lv
  • Zariņa, Latvian State Forest Research Institute Silava, 111 Rigas street, Salaspils LV-2169, Latvia E-mail: inga.zarina@silava.lv
  • Jansons, Latvian State Forest Research Institute Silava, 111 Rigas street, Salaspils LV-2169, Latvia E-mail: aris.jansons@silava.lv
article id 10211, category Research article
Gernot Erber, Raffaele Spinelli. (2020). Timber extraction by cable yarding on flat and wet terrain: a survey of cable yarder manufacturer’s experience. Silva Fennica vol. 54 no. 2 article id 10211. https://doi.org/10.14214/sf.10211
Keywords: forest soils; soil compaction; logging equipment; sensitive soils
Highlights: Survey of all European cable yarder manufacturers on flat-terrain yarding; Manufacturers are frequently contacted concerning flat-terrain yarding; Forest resource inaccessibility, regulatory and environmental considerations are most important motivations; Lack of clearance, tree stability and installation costs are major challenges; Mobile, self-anchoring tail spar is considered a chief adaptation; Cost-competitiveness with ground-based systems cannot be achieved without subsidies; Increasing environmental awareness and climate change present opportunity to expand flat-terrain cable yarding.
Abstract | Full text in HTML | Full text in PDF | Author Info

Cable yarding is a general solution for load handling on sites not accessible to ground-based machinery, and is typically associated with steep terrain. On flat terrain, such conditions can primarily be found on soft or wet soils, most frequently encountered in Central and Northern European countries. Today, changed environmental and market conditions may offer an unprecedented opportunity to the actual implementation of cable yarding on flat terrain in commercial operations. The study goal was to collect cable yarder manufacturers experience regarding the use and adaption of cable yarding technology on flat terrain. European manufacturers of cable yarding technology were interviewed about customer experience, particular challenges, adaptation potential, future potential and main hurdles for the expansion of cable yarding on flat terrain. Almost all manufacturers have received requests for flat-terrain yarding technology solutions, primarily from Germany. Temporal or permanent inaccessibility, regulatory or environmental reasons were the most frequent motivation for considering cable yarding technology. Installation was considered particularly challenging (clearance, stable anchoring). Potential adaptations included higher towers, artificial anchors, mechanized bunching before extraction and un-guyed yarder-systems. An artificial, highly mobile, self-anchoring tail spar was considered the most useful adaptation. While concerned about limited profitability and qualified labour shortage, most manufacturers demonstrated a positive or neutral view concerning the expansion of cable yarding on flat terrain. However, cable yarding is not considered to be cost-competitive wherever ground-based systems can be employed and cable yarding is not subsidized.

  • Erber, University of Natural Resources and Life Sciences Vienna, Department of Forest and Soil Sciences, Institute of Forest Engineering, Peter Jordan Strasse 82, 1190 Vienna, Austria ORCID https://orcid.org/0000-0003-1606-5258 E-mail: gernot.erber@boku.ac.at (email)
  • Spinelli, CNR-IBE Consiglio Nazionale delle Ricerche-Istituto per la BioEconomia, Via Madonna del Piano 10, Sesto Fiorentino, Firenze, I-50019, Italy; AFORA, University of the Sunshine Coast, Locked Bag 4, Maroochydore, QLD, Australia ORCID https://orcid.org/0000-0001-9545-1004 E-mail: spinelli@ivalsa.cnr.it
article id 10167, category Research article
Cheng Bai, Shixue You, Weipeng Ku, Qilin Dai, Zhengyi Wang, Mingshui Zhao, Shuquan Yu. (2020). Life form dynamics of the tree layer in evergreen and deciduous broad-leaved mixed forest during 1996–2017 in Tianmu Mountains, eastern China. Silva Fennica vol. 54 no. 2 article id 10167. https://doi.org/10.14214/sf.10167
Keywords: climate change; biodiversity; long-term monitoring; vegetation dynamics
Highlights: Biodiversity monitoring was performed over twenty one years in a 1 ha plot; Life form composition changed significantly between 1996 and 2017; Evergreen trees in the understory expanded upwards; Forests were currently dominated by evergreen trees.
Abstract | Full text in HTML | Full text in PDF | Author Info

In the forest areas of eastern China, there is a change from forest dominated by deciduous broad-leaved trees to forest dominated by evergreen broad-leaved trees as the latitude or altitude decreases. Different life forms have different survival strategies to deal with climate change, and studying the life form dynamics of the tree layers in the mixed forest in eastern China, with increasing temperature, can help us understand how the forest responds. This study was performed in a 1 ha plot in evergreen and deciduous broad-leaved mixed forest in Tianmu Mountain National Nature Reserve. Based on the data from two surveys (1996 and 2017), the changes in life form composition and biodiversity over the past 21 years were analyzed. We obtained the following results: (1) The proportion of evergreen trees increased from 55.0% in 1996 to 67.5% in 2017, and the dominance of evergreen species was enhanced. (2) The diversity of both life forms increased, and the tree species were more abundant. (3) The average annual recruitment rate of the evergreen species was 2.1% greater than their mortality rate, and the average annual recruitment rate of the deciduous species was 0.5% less than their mortality rate. (4) The competition among the trees in the small-diameter class (10 cm ≤ DBH < 20 cm) was fierce for many tree species. The proportion of the evergreen species in the small-diameter class was high. The life forms making up the mixed climax forest community has changed over the past 21 years, with the proportion and dominance of evergreen trees increasing significantly.

  • Bai, State Key Laboratory of Subtropical Silviculture, Zhejiang A&F University, Hangzhou 311300, China E-mail: baicheng111@gmail.com
  • You, College of Environmental and Resource Sciences, Zhejiang University, Hangzhou 310011, China E-mail: sxyou@zju.edu.cn
  • Ku, State Key Laboratory of Subtropical Silviculture, Zhejiang A&F University, Hangzhou 311300, China E-mail: 2732684475@qq.com
  • Dai, State Key Laboratory of Subtropical Silviculture, Zhejiang A&F University, Hangzhou 311300, China E-mail: 757692949@qq.com
  • Wang, State Key Laboratory of Subtropical Silviculture, Zhejiang A&F University, Hangzhou 311300, China E-mail: 237600341@qq.com
  • Zhao, Management Bureau of Tianmu Mountain National Nature Reserve, Hangzhou 311311, China E-mail: 973659738@qq.com
  • Yu, State Key Laboratory of Subtropical Silviculture, Zhejiang A&F University, Hangzhou 311300, China E-mail: yushq@zafu.edu.cn (email)
article id 10143, category Research article
Olga Grigorieva, Olga Brovkina, Alisher Saidov. (2020). An original method for tree species classification using multitemporal multispectral and hyperspectral satellite data. Silva Fennica vol. 54 no. 2 article id 10143. https://doi.org/10.14214/sf.10143
Keywords: boreal forest; phenological period; space spectroscopy; spectral signature
Highlights: Differences between spectral reflectance of tree species are statistically significant in the sub-seasons of spring, first half of summer, and main autumn; Classification using multitemporal multispectral data is more productive than is classification using a single hyperspectral image; the method improves recent forest mapping in the study regions.
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This study proposes an original method for tree species classification by satellite remote sensing. The method uses multitemporal multispectral (Landsat OLI) and hyperspectral (Resurs-P) data acquired from determined vegetation periods. The method is based on an original database of spectral features taking into account seasonal variations of tree species spectra. Changes in the spectral signatures of forest classes are analyzed and new spectral–temporal features are created for the classification. Study sites are located in the Czech Republic and northwest (NW) Russia. The differences in spectral reflectance between tree species are shown as statistically significant in the sub-seasons of spring, first half of summer, and main autumn for both study sites. Most of the errors are related to the classification of deciduous species and misclassification of birch as pine (NW Russia site), pine as mixture of pine and spruce, and pine as mixture of spruce and beech (Czech site). Forest species are mapped with accuracy as high as 80% (NW Russia site) and 81% (Czech site). The classification using multitemporal multispectral data has a kappa coefficient 1.7 times higher than does that of classification using a single multispectral image and 1.3 times greater than that of the classification using single hyperspectral images. Potentially, classification accuracy can be improved by the method when applying multitemporal satellite hyperspectral data, such as in using new, near-future products EnMap and/or HyspIRI with high revisit time.

  • Grigorieva, A.F. Mozhaysky’s Military-Space Academy, Krasnogo Kursanta Street 19a, 197198, Saint Petersburg, Russia E-mail: alenka12003@gmail.com
  • Brovkina, Global Change Research Institute CAS, Bělidla 986/4a, 603 00, Brno, Czech Republic E-mail: brovkina.o@czechglobe.cz (email)
  • Saidov, A.F. Mozhaysky’s Military-Space Academy, Krasnogo Kursanta Street 19a, 197198, Saint Petersburg, Russia E-mail: celestial.azura@gmail.com
article id 10243, category Research article
Laura Pikkarainen, Jaana Luoranen, Antti Kilpeläinen, Teppo Oijala, Heli Peltola. (2020). Comparison of planting success in one-year-old spring, summer and autumn plantings of Norway spruce and Scots pine under boreal conditions. Silva Fennica vol. 54 no. 1 article id 10243. https://doi.org/10.14214/sf.10243
Keywords: Pinus sylvestris; Picea abies; seedling damage; forest regeneration; field performance; planting season
Highlights: In Norway spruce, 84% of all plantings were successful, whereas in Scots pine, the corresponding number was 52%; The major reason for poor planting results was poor work quality; An extended planting season is possible for Norway spruce in southern and central Finland; In Scots pine, there are still large uncertainties in the success of summer and autumn plantings.
Abstract | Full text in HTML | Full text in PDF | Author Info

In Nordic countries, tree planting of seedlings is mainly performed during spring and early summer. Interest has increased in extending the planting window throughout the unfrozen growing season. This study compared the success of one-year-old spring, summer and autumn plantings in practical forestry in Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) in southern and central Finland. Planting success was based on the number of viable seedlings per hectare relative to a species-specific target density. The influence of different factors to poor planting results were determined, including quality of site preparation and planting, and sources of natural damage. Overall, in Norway spruce, 85, 69 and 84% and in Scots pine 53, 55 and 40% of spring, summer and autumn plantings succeeded. In Norway spruce, the planting results were consistent between the southern and central regions, whereas in Scots pine, the success was slightly lower in the south. The poor work quality and a low density of appropriate planting spots, contributed to poor planting results, regardless of planting season, region or tree species. Considering different damages, especially mammal damage contributed to the failure of Scots pine spring plantings, whereas in summer plantings, corresponding single failure reason could not be identified. Based on our findings, extending the planting season of Norway spruce could be recommended in both regions. For Scots pine, there is still significant uncertainty about the success of summer and autumn plantings, partially due to the limited number of plantings available for analyses.

  • Pikkarainen, School of Forest Sciences, University of Eastern Finland, Yliopistonkatu 7, FI-80100 Joensuu, Finland E-mail: laura.pikkarainen@uef.fi (email)
  • Luoranen, Natural Resources Institute Finland (Luke), Production Systems, Neulaniementie 5, FI-70210 Kuopio, Finland E-mail: jaana.luoranen@luke.fi
  • Kilpeläinen, School of Forest Sciences, University of Eastern Finland, Yliopistonkatu 7, FI-80100 Joensuu, Finland E-mail: antti.kilpelainen@uef.fi
  • Oijala, Metsä Group, Metsä Forest, Kuormaajantie 7, FI-40320 Jyväskylä, Finland E-mail: teppo.oijala@metsagroup.com
  • Peltola, School of Forest Sciences, University of Eastern Finland, Yliopistonkatu 7, FI-80100 Joensuu, Finland E-mail: heli.peltola@uef.fi
article id 10242, category Research article
Shisheng Long, Siqi Zeng, Falin Liu, Guangxing Wang. (2020). Influence of slope, aspect and competition index on the height-diameter relationship of Cyclobalanopsis glauca trees for improving prediction of height in mixed forests. Silva Fennica vol. 54 no. 1 article id 10242. https://doi.org/10.14214/sf.10242
Keywords: secondary forest; effect; improved Hegyi_I; topographic feature; tree height estimation
Highlights: In this study, the effects of slope, aspect and competition index (CI) on the H-DBH relationship were explored and an improved CI was developed and included to improve predictions of Cyclobalanopsis glauca tree height; It was found that the effects were statistically significant and considering slope, aspect and CI for developing the H-DBH models significantly increased the H prediction accuracy.
Abstract | Full text in HTML | Full text in PDF | Author Info

Diameter at breast height (DBH) and height (H) of trees are two important variables used in forest management plans. However, collecting the measurements of H is time-consuming and costly. Instead, the H-DBH relationship is modeled and used to estimate H. But, ignoring the effects of slope, aspect and tree competition on the H-DBH relationship often impedes the improvement of H predictions. In this study, to improve predictions of Cyclobalanopsis glauca (Thunb.) Oerst. tree H in mixed forests, we compared eleven H-DBH models and examined the influence of slope and aspect on the H-DBH relationship using 426 trees. We then improved Hegyi competition index and explored its effect on the H predictions by including it in the selected models. Results showed 1) There were statistically significant effects of slope and aspect on the H-DBH relationship; 2) The log transformation and exponential model performed best for sunny- and shady-steep, respectively, and the Gompertz’s model was optimal for both sunny- and shady-gentle; 3) Compared with the whole dataset, the division of the data into the slope and aspect sub-datasets significantly reduced the RMSE of H predictions; 4) Compared with the selected models without competition index, adding the original Hegyi and improved Hegyi_I into the models improved the H predictions but only the models containing the improved Hegyi_I significantly increased the prediction accuracy at the significant level of 0.1. This study implied that modeling the H-DBH relationship under different slopes and aspects and including the improved Hegyi_I provided the great potential to improve the H predictions.

  • Long, Faculty of Forestry, Central South University of Forestry and Technology, Changsha, Hunan 410004, China E-mail: shisheng3604@21cn.com
  • Zeng, Faculty of Forestry, Central South University of Forestry and Technology, Changsha, Hunan 410004, China E-mail: zengsiqi@21cn.com
  • Liu, Faculty of Forestry, Central South University of Forestry and Technology, Changsha, Hunan 410004, China E-mail: liufl680@126.com
  • Wang, Research Center of Forestry Remote Sensing & Information Engineering, Central South University of Forestry and Technology, Changsha 410004, China; Department of Geography and Environmental Resources, Southern Illinois University, Carbondale, IL 62901, USA E-mail: gxwang@siu.edu (email)
article id 10215, category Research article
Korotimi Ouédraogo, Kangbéni Dimobe, Adjima Thiombiano. (2020). Allometric models for estimating aboveground biomass and carbon stock for Diospyros mespiliformis in West Africa. Silva Fennica vol. 54 no. 1 article id 10215. https://doi.org/10.14214/sf.10215
Keywords: Burkina Faso; biometric variables; African ebony; biomass estimation equations; carbon storages; jackalberry; Sudanian savanna
Highlights: Biomass estimation models developed for Diospyros mespiliformis; Models based on DBH alone predicted aboveground biomass with 97.11% accuracy; Published models had relative error between –72% and +98%; Models for branch and stem biomass were more accurate than those for leaf biomass.
Abstract | Full text in HTML | Full text in PDF | Author Info

Accurate estimates of aboveground biomass (AGB) strongly depend on the suitability and precision of allometric models. Diospyros mespiliformis Hochst. ex A. DC. is a key component of most sub-Sahara agroforestry systems and, one of the most economically important trees in Africa. Despite its importance, very few scientific information exists regarding its biomass and carbon storage potential. In this study direct method was used to develop site-specific biomass models for D. mespiliformis tree components in Burkina Faso. Allometric models were developed for stem, branch and leaf biomass using data from 39 tree harvested in Sudanian savannas of Burkina Faso. Diameter at breast height (DBH), tree height, crown diameter (CD) and basal diameter (D20) were regressed on biomass component using non-linear models with DBH alone, and DBH in combination with height and/or CD as predictor variables. Carbon content was estimated for each tree component using the ash method. Allometric models differed between the experimental sites, except for branch biomass models. Site-specific models developed in this study exhibited good model fit and performance, with explained variance of 81–98%. Using models developed from other areas would have underestimated or overestimated biomass by between –72% and +98%. Carbon content in aboveground components of D. mespiliformis in Tiogo, Boulon and Tapoa-Boopo was 55.40% ± 1.50, 55.52% ± 1.06 and 55.63% ± 1.00, respectively, and did not vary significantly (P-value = 0.909). Site-specific models developed in this study are useful tool for estimating carbon stocks and can be used to accurately estimate tree components biomass in vegetation growing under similar conditions.

  • Ouédraogo, University Joseph Ki-Zerbo, UFR/SVT, Laboratory of Plant Biology and Ecology, 03 B.P. 7021 Ouagadougou 03, Burkina Faso E-mail: okorotimi@yahoo.fr (email)
  • Dimobe, University Joseph Ki-Zerbo, UFR/SVT, Laboratory of Plant Biology and Ecology, 03 B.P. 7021 Ouagadougou 03, Burkina Faso; University of Dédougou, Institut des Sciences de l’Environnement et du Développement Rural (ISEDR), BP 139 Dédougou, Burkina Faso; West African Science Service Center on Climate Change and Adapted Land Use, Competence Center, Avenue Muamar Ghadhafi, Ouagadougou, BP 9507, Burkina Faso ORCID https://orcid.org/0000-0001-5536-9700 E-mail: kangbenidimobe@gmail.com
  • Thiombiano, University Joseph Ki-Zerbo, UFR/SVT, Laboratory of Plant Biology and Ecology, 03 B.P. 7021 Ouagadougou 03, Burkina Faso E-mail: adjima_thiombiano@yahoo.fr
article id 10196, category Research article
Karri Uotila, Jari Miina, Timo Saksa, Ron Store, Kauko Kärkkäinen, Mika Härkönen. (2020). Low cost prediction of time consumption for pre-commercial thinning in Finland. Silva Fennica vol. 54 no. 1 article id 10196. https://doi.org/10.14214/sf.10196
Keywords: forest vegetation management; early cleaning; release treatment; linear mixed model; topographic wetness index; work productivity
Highlights: Time consumption (TC) in pre-commercial thinning (PCT) can be predicted by variables describing site and stands conditions and previous silvicultural management; Applying variables available in forest resources data the field-assessment of worksite difficulty factors is not needed; The TC model could facilitate the predictions of the labour costs of PCT in forest information systems.
Abstract | Full text in HTML | Full text in PDF | Author Info

The time consumption (TC) of pre-commercial thinning (PCT) varies greatly among sites, stands and forest workers. The TC in PCT is usually estimated by field-assessed work difficulty factors. In this study, a linear mixed model for the TC in PCT was prepared by utilizing forest resources data (FRD). The modelling data included 11 848 and validation data included 3035 worksites with TC information recorded by forest workers within the period of 2008–2018. The worksites represented a range of site and stand conditions across a broad geographical area in Finland. Site and stand characteristics and previous management logically explained the TC in PCT. The more fertile the site, the more working time was needed in PCT. On sites of medium fertility, TC in the initial PCT increased with stand age by 0.5 h ha–1 yr–1. Site wetness increased the TC. PCT in summer was more time consuming than in spring. Small areas were more time consuming to PCT per hectare than larger ones. The between-forest worker variation involved in the TC was as high as 35% of the variation unexplained by the TC model. The coefficient of determination in validation data was 19.3%, RMSE 4.75 h ha–1 and bias –1.6%. The TC model based on FRD was slightly less precise than the one based on field-assessed work difficulty factors (removal quantity and type and terrain difficulty): RMSE 4.9 h ha–1 vs. 4.1 h ha–1 (52% vs. 43%). The TC model could be connected to forest information systems where it would facilitate the predictions of the labour costs of PCT without field-assessing work difficulty factors.

  • Uotila, Natural Resources Institute Finland (Luke), Natural resources, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: karri.uotila@luke.fi (email)
  • Miina, Natural Resources Institute Finland (Luke), Natural resources, Yliopistokatu 6 B, FI-80100 Joensuu, Finland E-mail: jari.miina@luke.fi
  • Saksa, Natural Resources Institute Finland (Luke), Natural resources, Survontie 9, FI-40500 Jyväskylä, Finland E-mail: timo.saksa@luke.fi
  • Store, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Teknologiakatu 7, FI-67100 Kokkola, Finland E-mail: ron.store@luke.fi
  • Kärkkäinen, E-mail: kauko.karkkainen@gmail.com
  • Härkönen, Tornator Oyj, Pielisentie 2–6, FI-81700 Lieksa, Finland E-mail: mika.harkonen@tornator.fi
article id 10163, category Research article
Uttam Thangjam, Uttam K. Sahoo, Pentile Thong. (2020). Characterization of morphometric, reproductive and seedling traits of Parkia timoriana in northeast India. Silva Fennica vol. 54 no. 1 article id 10163. https://doi.org/10.14214/sf.10163
Keywords: seed source; heritability; genotype; tree bean; variations
Highlights: Among provenances, Pherema (P1) Serchip (P10) and Jiribam (P7) were associated with stress tolerant and better quality seedlings.; P1 gave the best result for seed traits including high germinability, P10 showed highest seedling vigour, while P7 corresponds to high-quality pod related traits; Variations in morphological, germinative and seedling growth characters of seeds across provenance further illustrate the importance of site selection for the production of better quality trees.
Abstract | Full text in HTML | Full text in PDF | Author Info

We studied variations on different traits of Parkia timoriana (D.C.) Merr. in twelve provenances systematically from their source of origin to a controlled environment where representative seedlings were grown. Among the provenances, P1 gave the best result for seed traits including germination traits, P7 for pod traits and P10 for seedling vigour. Effects of seasonal distribution of rainfall and temperature on seed and pod traits were also determined by computing multiple regression analysis. The results displayed winter rainfall and summer temperature as the most important factor determining pod and seed traits. Latitude also significantly (P < 0.001) affected PWT (r = 0.52), SWP (r = 0.46) and SW (r = 0.50). A common garden study for germination and seedling growth indicated P1 and P10 provenance as the best among all. Seeds drawn from P10 gave the highest seedling vigour with an average growth rate of 0.61 cm/day from 90th to 180th day. Highest broad-sense heritability values (h2) were observed in germination traits, followed by seedling collar diameter. The lowest h2 was observed for seedling height.

  • Thangjam, Department of Forestry, School of Earth Sciences & Natural Resource Management, Mizoram University, Aizawl-796004, Mizoram, India ORCID https://orcid.org/0000-0003-1263-5348 E-mail: thangjam1987@gmail.com
  • Sahoo, Department of Forestry, School of Earth Sciences & Natural Resource Management, Mizoram University, Aizawl-796004, Mizoram, India ORCID https://orcid.org/0000-0002-6524-1775 E-mail: uksahoo_2003@rediffmail.com (email)
  • Thong, Department of Forestry, School of Earth Sciences & Natural Resource Management, Mizoram University, Aizawl-796004, Mizoram, India ORCID https://orcid.org/0000-0002-5358-181X E-mail: pentilethong@gmail.com
article id 10084, category Research article
Mihails Čugunovs, Eeva-Stiina Tuittila, Jari Kouki. (2020). Proximity to charred logs in burned forests likely affects decomposition processes in the soil. Silva Fennica vol. 54 no. 1 article id 10084. https://doi.org/10.14214/sf.10084
Keywords: coarse woody debris; forest fire; prescribed burning; cellulose decomposition; tea-bag method
Highlights: Standardised organic substrate decomposition was tentatively observed to be faster adjacent to non-charred downed logs than away from the logs or adjacent to charred logs; A spatial linkage was observed between non-charred logs and decomposition in the soil in burned boreal forests; Proximity to a charred log may provide a micro-environment where decomposition rates differ from the surrounding forest soil.
Abstract | Full text in HTML | Full text in PDF | Author Info

We studied the spatial decomposition rates of standardised organic substrates in soils (burned boreal pine-dominated sub-xeric forests in eastern Finland), with respect to charred and non-charred coarse woody debris (CWD). Decomposition rates of rooibos plant litter inside teabags (C:N = 42.870 ± 1.841) and pressed-sheet Nordic hardwood pulp (consisting of mainly alpha-cellulose) were measured at 0.2 m distance from 20 charred (LC0.2) and 40 non-charred logs (LNC0.2). We also measured decomposition at 60 plots located 3–10 m away from downed logs (L3,10). The rooibos decomposition rate constant ‘k’ was 8.4% greater at the LNC0.2 logs than at the L3,10 or LC0.2 logs. Cellulose decomposed more completely in 1 micron mesh bags at LNC0.2 (44% of buried bags had leftover material) than at LC0.2 (76%) or L3,10 (70%). Decomposition of cellulose material was rapid but varied greatly between sampling plots. Our results indicate that decomposition of the standardised organic matter was more rapid close to CWD pieces than further away. However, only the plots located near non-charred logs (LNC0.2) exhibited high decomposition rates, with no corresponding increase observed at the charred logs (LC0.2). This suggests a possible noteworthy indirect effect of forest burning on soil organic matter (SOM) decomposition rates close to charred CWD after forest fires. We urge for more studies on this tentative observation as it may affect the estimates on how fires affect carbon cycling in forests.

  • Čugunovs, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: mihails.cugunovs@gmail.com (email)
  • Tuittila, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0001-8861-3167 E-mail: eeva-stiina.tuittila@uef.fi
  • Kouki, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0003-2624-8592 E-mail: jari.kouki@uef.fi
article id 10051, category Research article
Anna-Kaisa Kosenius, Artti Juutinen, Liisa Tyrväinen. (2020). The role of state-owned commercial forests and firm features in nature-based tourism business performance. Silva Fennica vol. 54 no. 1 article id 10051. https://doi.org/10.14214/sf.10051
Keywords: business performance; entrepreneurial attitude; forest-based tourism business; forest management practices; landscape trading; recreational value trading
Highlights: State-owned commercial forests provide an important part of nature-based tourism enterprises’ service scope; Increase in turnover during five years vary depending on provided services; Entrepreneurial attitude associates with willingness to participate in a landscape and recreational value trading scheme; Increasing international demand for nature-based tourism promotes business possibilities, needs attention to forest environments and smooth co-operation between actors.
Abstract | Full text in HTML | Full text in PDF | Author Info

Nature-based tourism (NBT) is a growing industry within regions rich in natural amenities worldwide. An important feature of NBT business is the dependence on the quality of surrounding environment. This paper addresses the role of the management of commercial forests owned by the state in Finnish Lapland. The paper explores the NBT entrepreneurs’ willingness to participate in a proposed new landscape and recreational value trading (LRVT) and elaborates the effect of entrepreneur and enterprise characteristics, such as entrepreneurial attitude, venture size, and a variety of services offered to customers, on the experienced and expected growth of NBT enterprise. The survey data on NBT enterprises were analyzed with ordered and binary logit models. The willingness of enterprises to participate in LRVT depended on the venture size, entrepreneurial attitude, and type of activities offered to customers. The results show that relatively young and small-sized enterprises have faced difficulties in developing their business. Entrepreneurial experience, risk-taking and intention to develop new business associate positively with expected increase in turnover.

  • Kosenius, University of Helsinki, Department of Economics and Management, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID https://orcid.org/0000-0002-1528-8145 E-mail: anna-kaisa.kosenius@helsinki.fi (email)
  • Juutinen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Paavo Havaksentie 3, FI-90570 Oulu, Finland ORCID https://orcid.org/0000-0003-4300-5365 E-mail: artti.juutinen@luke.fi
  • Tyrväinen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0001-9941-8228 E-mail: liisa.tyrvainen@luke.fi
article id 10230, category Research article
Mohammed Henneb, Osvaldo Valeria, Nelson Thiffault, Nicole Fenton. (2019). Black spruce seedling growth response in controlled organic and organic-mineral substrates. Silva Fennica vol. 53 no. 4 article id 10230. https://doi.org/10.14214/sf.10230
Keywords: Picea mariana; paludification; seedling growth; seedling nutrition; semi-controlled experiment; substrate
Highlights: Seedling height and diameter were highest on clay and mesic substrates respectively; Foliar nutrients were relatively high in seedlings that were established on mesic substrates; We recommend the application of mechanical soil preparation techniques that promote the creation of organic-mesic substrates to support regeneration establishment.
Abstract | Full text in HTML | Full text in PDF | Author Info

In the boreal forest of eastern Canada, a large proportion of black spruce (Picea mariana [Mill.] Britton, Sterns & Poggenb.) stands are affected by paludification. Edaphic conditions that are created by paludification processes, including an abundance of microsites with high moisture and low nutrient contents, hinder forest regeneration. Disturbance of paludified sites by mechanical soil preparation (MSP) reduces organic layer thickness, while generating a range of substrates for regeneration establishment. Yet, little information is available regarding the effects of these substrates on tree growth. Our objective was to determine the effect of organic, mineral and organo-mineral substrates that are created following MSP of a paludified site on the growth and root development of black spruce seedlings in a semi-controlled environment. We demonstrated that substrate exerted a significant effect on seedling growth and foliar concentrations of N, P and K. Increase in height and diameter were respectively greatest on clay (mineral) and mesic substrates. Substrate effects did not affect total biomass increases or final root biomass. Foliar nutrients (N, P, K) were relatively high in seedlings that were established on mesic substrates and relatively low for those established on clay substrates. To ensure successful seedling establishment, we recommend the application of MSP techniques that expose organic-mesic substrates on sites that are susceptible to paludification.

  • Henneb, Institut de recherche sur les forêts (IRF), Université du Québec en Abitibi-Témiscamingue, 445 boul. de l’Université, Rouyn-Noranda, QC J9X 5E4, Canada ORCID http://orcid.org/0000-0003-4507-1219 E-mail: mohammed.henneb@uqat.ca (email)
  • Valeria, Institut de recherche sur les forêts (IRF), Université du Québec en Abitibi-Témiscamingue, 445 boul. de l’Université, Rouyn-Noranda, QC J9X 5E4, Canada ORCID http://orcid.org/0000-0002-9921-7474 E-mail: osvaldo.valeria@uqat.ca
  • Thiffault, Institut de recherche sur les forêts (IRF), Université du Québec en Abitibi-Témiscamingue, 445 boul. de l’Université, Rouyn-Noranda, QC J9X 5E4, Canada; Natural Resources Canada, Canadian Wood Fibre Centre, 1055 rue du PEPS, P.O. Box 10380, Stn Sainte Foy, Quebec, QC G1V 4C7, Canada ORCID http://orcid.org/0000-0003-2017-6890 E-mail: nelson.thiffault@canada.ca
  • Fenton, Institut de recherche sur les forêts (IRF), Université du Québec en Abitibi-Témiscamingue, 445 boul. de l’Université, Rouyn-Noranda, QC J9X 5E4, Canada ORCID http://orcid.org/0000-0002-3782-2361 E-mail: nicole.fenton@uqat.ca
article id 10217, category Research article
Xingji Jin, Timo Pukkala, Fengri Li, Lihu Dong. (2019). Developing growth models for tree plantations using inadequate data – a case for Korean pine in Northeast China. Silva Fennica vol. 53 no. 4 article id 10217. https://doi.org/10.14214/sf.10217
Keywords: Pinus koraiensis; optimization-based modeling; quantile regression; self-thinning
Highlights: The permanent sample plots of Chinese plantation trees have not been designed for producing data for growth modeling; We used various methods to deal with the inadequacies of sample plot data; Optimization was used to fit diameter increment and survival models using data with varying measurement intervals and tree identification errors; Quantile regression was used to model self-thinning limit.
Abstract | Full text in HTML | Full text in PDF | Author Info

Korean pine (Pinus koraiensis Siebold & Zucc.) is economically the most important tree species in northeast China. Korean pine plantations are established and managed for the production of timber and seeds. Despite the importance of the species, few models have been developed for the comparison of alternative management schedules. Model development is affected by the fact that permanent sample plots and thinning experiments have not been designed and managed for modeling purposes. The permanent sample plots include few non-thinned plots, and weak trees are removed in thinning treatments, leading to low mortality rate. Moreover, the measurement interval is irregular. This study used optimization-based modeling approach in tree-level diameter increment and survival modeling to deal with the above problems. Models for self-thinning limit were developed to alleviate the problem of underestimated mortality arising from the features of the data. In addition, improved site index and individual-tree height models were developed. The model of Lundqvist and Korf was used as the site index model and the model proposed by Schumacher as the height model. Quantile regression was used to model the maximum stand basal area and maximum number of trees as a function of mean tree diameter and site index. Tree diameter, stand basal area, basal area in larger trees and site index were used as the predictors of diameter increment and tree survival. The models developed in this study constitute a model set that is suitable for simulation and optimization studies. The models produced simulation results that correspond to measured stand development.

  • Jin, Key Laboratory of Sustainable Forest Ecosystem Management - Ministry of Education, School of Forestry, Northeast Forestry University, Harbin 150040, Heilongjiang, People’s Republic of China ORCID http://orcid.org/0000-0003-2971-2709 E-mail: xingji_jin@163.com
  • Pukkala, Key Laboratory of Sustainable Forest Ecosystem Management - Ministry of Education, School of Forestry, Northeast Forestry University, Harbin 150040, Heilongjiang, People’s Republic of China; University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: timo.pukkala@uef.fi (email)
  • Li, Key Laboratory of Sustainable Forest Ecosystem Management - Ministry of Education, School of Forestry, Northeast Forestry University, Harbin 150040, Heilongjiang, People’s Republic of China ORCID http://orcid.org/0000-0002-4058-769X E-mail: fengrili@126.com
  • Dong, Key Laboratory of Sustainable Forest Ecosystem Management - Ministry of Education, School of Forestry, Northeast Forestry University, Harbin 150040, Heilongjiang, People’s Republic of China ORCID http://orcid.org/0000-0002-3985-9475 E-mail: ldonglihu2006@163.com
article id 10209, category Research article
Claudie-Maude Canuel, Nelson Thiffault, Michael K. Hoepting, James C.G. Farrell. (2019). Legacy effects of precommercial thinning on the natural regeneration of next rotation balsam fir stands in eastern Canada. Silva Fennica vol. 53 no. 4 article id 10209. https://doi.org/10.14214/sf.10209
Keywords: silviculture; Abies balsamea; PCT; conifer; density management
Highlights: We investigated the potential legacy effects of precommercial thinning in next rotation, dense natural balsam fir stands; Precommercial thinning had few legacy effects on next rotation stands and should not impair their regeneration; Balsam fir dominated the regeneration layer. Other tree species were almost absent.
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The Green River precommercial thinning (PCT) trial was established between 1959–1961 in New Brunswick (Canada) within natural balsam fir (Abies balsamea (L.) Mill.)-dominated stands. Three silviculture scenarios differing only by the increasing nominal spacings of PCT treatments (1.2 m, 1.8 m, 2.4 m) were compared to an unthinned control within randomized replicates that were clearcut harvested in 2008 and treated with herbicide in 2011. During the fourth post-harvest growing season, we assessed regeneration, competing vegetation and coarse woody debris (CWD; differentiated between large woody debris and slash) to assess the legacy effects of PCT on regeneration of next rotation stands. Our results confirmed that silviculture scenarios including PCT significantly increased conifer stocking in treated plots compared to control conditions, but only in the 1.8 m nominal spacing. Considering that treated and untreated stands were fully stocked, we conclude that PCT using the spacing gradient tested has no legacy effect on the regeneration of next rotation natural balsam fir stands. Given the known sensitivity of balsam fir to future climate conditions in this region, we suggest that future treatments should promote tree species diversity to support ecosystem resilience to climate change by favouring more warm-adapted species, such as some hardwoods.

  • Canuel, Faculté de foresterie, géographie et géomatique, Université Laval, Québec, QC G1V 0A6, Canada;  Canadian Wood Fibre Centre, Natural Resources Canada, 1055 du P.E.P.S., P.O. Box 10380, Sainte-Foy Stn., Québec, QC G1V 4C7, Canada E-mail: claudie-maude.canuel.1@ulaval.ca
  • Thiffault, Canadian Wood Fibre Centre, Natural Resources Canada, 1055 du P.E.P.S., P.O. Box 10380, Sainte-Foy Stn., Québec, QC G1V 4C7, Canada ORCID http://orcid.org/0000-0003-2017-6890 E-mail: nelson.thiffault@canada.ca (email)
  • Hoepting, Canadian Wood Fibre Centre, Natural Resources Canada, 1219 Queen St. E., Sault Ste. Marie, ON P6A 2E5, Canada E-mail: michael.hoepting@canada.ca
  • Farrell, Canadian Wood Fibre Centre, Natural Resources Canada, 1350 Regent Street, P.O. Box 4000, Fredericton, NB E3B 5P7, Canada E-mail: jamescg.farrell@canada.ca
article id 10183, category Research article
Tomi Karjalainen, Petteri Packalen, Janne Räty, Matti Maltamo. (2019). Predicting factual sawlog volumes in Scots pine dominated forests using airborne laser scanning data. Silva Fennica vol. 53 no. 4 article id 10183. https://doi.org/10.14214/sf.10183
Keywords: Pinus sylvestris; area based approach; k-NN; sawlog
Highlights: We predicted visually bucked factual sawlog volumes at the 30 × 30 m plot-level with several alternatives; The lowest root mean squared error value of approximately 21% was obtained with a linear mixed-effects model that employed factual sawlog volume as a response variable and airborne laser scanning metrics as predictors; The sawlog reduction model commonly used in Finland performed poorly.
Abstract | Full text in HTML | Full text in PDF | Author Info

The aim in the study was to compare alternatives for the prediction of factual sawlog volumes using airborne laser scanning (ALS) data in Scots pine (Pinus sylvestris L.) dominated forests in eastern Finland. Accurate estimates of factual sawlog volume are desirable to ease the planning of harvesting operations. The factual sawlog volume of pines was derived from visual bucking, i.e. a procedure where the defects were located on each stem during sample plot measurements. For other species, the theoretical sawlog volume was considered also as the factual sawlog volume due to data restrictions. We predicted factual sawlog volume with eight alternatives that were based on either linear mixed-effects models or k-nearest neighbour imputations. An existing sawlog reduction model, commonly used in Finland, was also tested individually and combined with a number of the alternatives, and site type information was also utilised. Model fitting and prediction was implemented at the 15 × 15 m level, but accuracy was assessed at the 30 × 30 m level. The relative root mean squared error (RMSE%) values for the factual sawlog volume predictions varied between 20.9% and 33.5%, and the best accuracy was obtained with a linear mixed-effects model. These results indicate that factual sawlog volumes in Scots pine dominated forests can be predicted with reasonable accuracy with ALS data.

  • Karjalainen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: tomikar@uef.fi (email)
  • Packalen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: petteri.packalen@uef.fi
  • Räty, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: janne.raty@uef.fi
  • Maltamo, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: matti.maltamo@uef.fi
article id 10151, category Research article
Jyri Hietala, Riitta Hänninen, Matleena Kniivilä, Anne Toppinen. (2019). Networks in international opportunity recognition among Finnish wood product industry SMEs. Silva Fennica vol. 53 no. 4 article id 10151. https://doi.org/10.14214/sf.10151
Keywords: wood products; business networks; institutional networks; internationalization; opportunity recognition; social networks
Highlights: In line with earlier literature, we found the networks in our study to positively impact international opportunity recognition; Despite the reliance on various network forms and levels, a strategic stance towards opportunity recognition can be characterized as being more reactive than proactive; Institutional networks represented a more systematic way of recognizing international opportunities among case companies.
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Bioeconomy development will create new opportunities for firms operating in the international wood products markets, and identifying and exploiting these opportunities is emphasized as a key concept to achieving business success. Our study will attempt to address a gap in the literature on sawmill industry business development from the viewpoint of international opportunity recognition. The aim of our study is to provide a holistic description on how small and medium-sized enterprises (SMEs) in the wood products industry recognize and exploit international business opportunities, and how they utilize network perspectives in this context. The subject was examined through Finnish wood product industry SMEs by interviewing 11 managers and industry representatives. The results suggest that SMEs recognize international opportunities reactively per se. Social networks formed in professional forums were an important information channel for identifying international opportunities. Through vertical business networks, such as sales agents, firms have been able to increase their international market presence and free their own resources for other important activities. Horizontal dyadic business networks were seen to facilitate new international opportunities through cooperation, while excessive reliance on vertical networks raised concerns and seemed not to be effective in international opportunity recognition. Institutional networks formed a systematic way of recognizing international opportunities, but more so at the initial market entry stage.

  • Hietala, United Bankers, Aleksanterinkatu 21 A, FI-00100 Helsinki, Finland E-mail: jyri.hietala@unitedbankers.fi (email)
  • Hänninen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: riitta.hanninen@luke.fi
  • Kniivilä, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: matleena.kniivila@luke.fi
  • Toppinen, University of Helsinki, Helsinki Institute of Sustainability Science, Latokartanonkaari 7, P.O. 27, FI-00014 University of Helsinki, Finland E-mail: anne.toppinen@helsinki.fi
article id 10147, category Research article
Mika Aalto, Olli-Jussi Korpinen, Tapio Ranta. (2019). Feedstock availability and moisture content data processing for multi-year simulation of forest biomass supply in energy production. Silva Fennica vol. 53 no. 4 article id 10147. https://doi.org/10.14214/sf.10147
Keywords: bioenergy; simulation; forest resources; data analysis; geographic information system
Highlights: A method for allocating forest biomass availability for a multi-year simulation model was developed; The possibility to take the quality change of feedstock into account by moisture estimations was studied; A method to estimate weather data for moisture estimation equations with fewer parameters was presented.
Abstract | Full text in HTML | Full text in PDF | Author Info

Simulation and modeling have become more common in forest biomass studies. Dynamic simulation has been used to study the supply chain of forest biomass with numerous different models. A robust predictive multi-year model requires biomass availability data, where annual variation is included spatially and temporally. This can be done by using data from enterprises, but in some cases relevant data is not accessible. Another option is to use forest inventory data to estimate biomass availability, but this data must be processed in the correct form to be utilized in the model. This study developed a method for preparing forest inventory data for a multi-year simulation supply model using the theoretical availability of feedstock. Methods for estimating quality changes during roadside storage are also presented, including a possible parameter estimation to decrease the amount of data needed. The methods were tested case by case using the inventory database “Biomass Atlas” and weather data from a weather station in Mikkeli, Finland. The data processing method for biomass allocation produced a reasonable quantity of stands and feedstock, having a realistic annual supply with variation for the demand point. The results of the study indicate that it is possible to estimate moisture content changes using weather data. The estimations decreased the accuracy of the model and, therefore, estimations should be kept minimal. The presented data preparation method can generate a supply of forest biomass for the simulation model, but the validity of the data must be ensured for correct model behavior.

  • Aalto, Lappeenranta-Lahti University of Technology LUT, School of Energy Systems, Laboratory of Bioenergy, Lönnrotinkatu 7, FI-50100 Mikkeli, Finland ORCID https://orcid.org/0000-0002-7768-1145 E-mail: mika.aalto@lut.fi (email)
  • Korpinen, Lappeenranta-Lahti University of Technology LUT, School of Energy Systems, Laboratory of Bioenergy, Lönnrotinkatu 7, FI-50100 Mikkeli, Finland E-mail: olli-jussi.korpinen@lut.fi
  • Ranta, Lappeenranta-Lahti University of Technology LUT, School of Energy Systems, Laboratory of Bioenergy, Lönnrotinkatu 7, FI-50100 Mikkeli, Finland ORCID https://orcid.org/0000-0001-5464-5136 E-mail: tapio.ranta@lut.fi
article id 10016, category Research article
Ivars Kļaviņš, Arta Bārdule, Zane Lībiete, Dagnija Lazdiņa, Andis Lazdiņš. (2019). Impact of biomass harvesting on nitrogen concentration in the soil solution in hemiboreal woody ecosystems. Silva Fennica vol. 53 no. 4 article id 10016. https://doi.org/10.14214/sf.10016
Keywords: nitrogen concentration; stump harvesting; whole-tree harvesting; soil solution; hemiboreal forest; short-rotation coppice
Highlights: Soil solution nitrogen concentrations in whole-tree harvesting sites are higher in sites of medium to high fertility than in sites of low fertility; In whole-tree harvesting and stem-only harvesting sites, soil solution nitrogen concentrations are highest 2 to 3 years after harvesting; The risks of nitrogen leaching immediately after harvesting are higher in traditional forestry systems compared to short-rotation cropping.
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Considering the increasing use of wood biomass for energy and the related intensification of forest management, the impacts of different intensities of biomass harvesting on nutrient leaching risks must be better understood. Different nitrogen forms in the soil solution were monitored for 3 to 6 years after harvesting in hemiboreal forests in Latvia to evaluate the impacts of different biomass harvesting regimes on local nitrogen leaching risks, which potentially increase eutrophication in surface waters. In forestland dominated by Scots pine Pinus sylvestris L. or Norway spruce Picea abies L. (Karst.), the soil solution was sampled in: (i) stem-only harvesting (SOH), (ii) whole‐tree harvesting, with only slash removed (WTH), and (iii) whole‐tree harvesting, with both slash and stumps harvested (WTH + SB), subplots. In agricultural land, sampling was performed in an initially fertilised hybrid aspen (Populus tremula L.× P. tremuloides Michx.) short-rotation coppice (SRC), where above-ground biomass was harvested. In forestland, soil solution N (nitrogen) concentrations were highest in the second and third year after harvesting. Mean annual values in WTH subplots of medium to high fertility sites exceeded the mean values in SOH subplots and control subplots (mature stand where no harvesting was performed) for the entire study period; the opposite trend was observed for the low-fertility site. Biomass harvesting in the hybrid aspen SRC only slightly affected NO3-N (nitrate nitrogen) and NH4+-N (ammonium nitrogen) concentrations in the soil solution within 3 years after harvesting, but a significant decrease in the TN (total nitrogen) concentration in the soil solution was found in plots with additional N fertilisation performed once initially.

  • Kļaviņš, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia; University of Latvia, Raiņa blvd 19-125, LV 1586, Riga, Latvia E-mail: ivars.klavins@silava.lv (email)
  • Bārdule, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia; University of Latvia, Raiņa blvd 19-125, LV 1586, Riga, Latvia E-mail: arta.bardule@silava.lv
  • Lībiete, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: zane.libiete@silava.lv
  • Lazdiņa, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: dagnija.lazdina@silava.lv
  • Lazdiņš, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: andis.lazdins@silava.lv
article id 10010, category Research article
Panu Halme, Jenna Purhonen, Emma-Liina Marjakangas, Atte Komonen, Katja Juutilainen, Nerea Abrego. (2019). Dead wood profile of a semi-natural boreal forest – implications for sampling. Silva Fennica vol. 53 no. 4 article id 10010. https://doi.org/10.14214/sf.10010
Keywords: coarse woody debris; coniferous forest; forest structure; fine woody debris; forest disturbance dynamics; saproxylic
Highlights: We constructed a full dead wood profile of a semi-natural boreal forest; Abundance-diameter distributions were different among tree species; Extensive sampling is needed if focus on large dead wood and rare tree species.
Abstract | Full text in HTML | Full text in PDF | Author Info

Dead wood profile of a forest is a useful tool for describing forest characteristics and assessing forest disturbance history. Nevertheless, there are few studies on dead wood profiles, including both coarse and fine dead wood, and on the effect of sampling intensity on the dead wood estimates. In a semi-natural boreal forest, we measured every dead wood item over 2 cm in diameter from 80 study plots. From eight plots, we further recorded dead wood items below 2 cm in diameter. Based on these data we constructed the full dead wood profile, i.e. the overall number of dead wood items and their distribution among different tree species, volumes of different size and decay stage categories. We discovered that while the number of small dead wood items was immense, their number dropped drastically from the diameter below 1 cm to diameters 2–3 cm. Different tree species had notably different abundance-diameter distribution patterns: spruce dead wood comprised most strikingly the smallest diameter fractions, whereas aspen dead wood comprised a larger share of large-diameter items. Most of the dead wood volume constituted of large pieces (>10 cm in diameter), and 62% of volume was birch. The variation in the dead wood estimates was small for the numerically dominant tree species and smallest diameter categories, but high for the sub-dominant tree species and larger size categories. In conclusion, the more the focus is on rare tree species and large dead wood items, the more comprehensive should the sampling be.

  • Halme, Department of Biological and Environmental Science, University of Jyväskylä, P.O. Box 35, FI-40014 University of Jyväskylä, Finland; School of Resource Wisdom, University of Jyväskylä, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: panu.halme@jyu.fi (email)
  • Purhonen, Department of Biological and Environmental Science, University of Jyväskylä, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: jenna.e.i.purhonen@jyu.fi
  • Marjakangas, Centre for Biodiversity Dynamics, Department of Biology, Norwegian University of Science and Technology, N-7491 Trondheim, Norway E-mail: emma-liina.marjakangas@ntnu.no
  • Komonen, Department of Biological and Environmental Science, University of Jyväskylä, P.O. Box 35, FI-40014 University of Jyväskylä, Finland; School of Resource Wisdom, University of Jyväskylä, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: atte.komonen@jyu.fi
  • Juutilainen, Department of Biological and Environmental Science, University of Jyväskylä, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: kjuutilainen@yahoo.com
  • Abrego, Department of Agricultural Sciences, University of Helsinki, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: nerea.abrego@helsinki.fi
article id 10187, category Research article
Timo Pukkala, Kjersti Holt Hanssen, Kjell Andreassen. (2019). Stem taper and bark functions for Norway spruce in Norway. Silva Fennica vol. 53 no. 3 article id 10187. https://doi.org/10.14214/sf.10187
Keywords: forest management; Picea abies; Kozak model; variable-exponent taper function
Highlights: New variable-exponent stem taper and bark functions were developed for Norway spruce; Both fixed and mixed-effects models were developed; Site index and tree age had statistically significant but small effects on stem taper.
Abstract | Full text in HTML | Full text in PDF | Author Info

Based on data from long-term experimental fields with Norway spruce (Picea abies (L.) H. Karst.), we developed new stem taper and bark functions for Norway. Data was collected from 477 trees in stands across Norway. Three candidate functions which have shown good performance in previous studies (Kozak 02, Kozak 97 and Bi) were fitted to the data as fixed-effects models. The function with the smallest Akaike Information Criterion (AIC) was then chosen for additional analyses, fitting 1) site index-dependent and 2) age-dependent versions of the model, and 3) fitting a mixed-effects model with tree-specific random parameters. Kozak 97 was found to be the function with the smallest AIC, but all three tested taper functions resulted in fairly similar predictions of stem taper. The site index-dependent function reduced AIC and residual standard error and showed that the effect of site index on stem taper is different in small and large trees. The predictions of the age-independent and age-dependent models were very close to each other. Adding tree-specific random parameters to the model clearly reduced AIC and residual variation. However, the results suggest that the mixed-effects model should be used only when it is possible to calibrate it for each tree, otherwise the fixed-effects Kozak 97 model should be used. A model for double bark thickness was also fitted as fixed-effects Kozak 97 model. The model behaved logically, predicting larger relative but smaller absolute bark thickness for small trees.

article id 10179, category Research article
Lauri Korhonen, Jaakko Repola, Tomi Karjalainen, Petteri Packalen, Matti Maltamo. (2019). Transferability and calibration of airborne laser scanning based mixed-effects models to estimate the attributes of sawlog-sized Scots pines. Silva Fennica vol. 53 no. 3 article id 10179. https://doi.org/10.14214/sf.10179
Keywords: Pinus sylvestris; LIDAR; crown base height; hierarchical data; individual tree detection; sawlog quality
Highlights: Attributes of individual sawlog-sized pines estimated by transferring ALS-based models between sites; Mixed effects models were more accurate than k-NN imputation tested earlier; Calibration with a small number of field measured trees improved the accuracy.
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Airborne laser scanning (ALS) data is nowadays often available for forest inventory purposes, but adequate field data for constructing new forest attribute models for each area may be lacking. Thus there is a need to study the transferability of existing ALS-based models among different inventory areas. The objective of our study was to apply ALS-based mixed models to estimate the diameter, height and crown base height of individual sawlog sized Scots pines (Pinus sylvestris L.) at three different inventory sites in eastern Finland. Different ALS sensors and acquisition parameters were used at each site. Multivariate mixed-effects models were fitted at one site and the models were validated at two independent test sites. Validation was carried out by applying the fixed parts of the mixed models as such, and by calibrating them using 1–3 sample trees per plot. The results showed that the relative RMSEs of the predictions were 1.2–6.5 percent points larger at the test sites compared to the training site. Systematic errors of 2.4–6.2 percent points also emerged at the test sites. However, both the RMSEs and the systematic errors decreased with calibration. The results showed that mixed-effects models of individual tree attributes can be successfully transferred and calibrated to other ALS inventory areas in a level of accuracy that appears suitable for practical applications.

  • Korhonen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland ORCID http://orcid.org/0000-0002-9352-0114 E-mail: lauri.korhonen@uef.fi (email)
  • Repola, Natural Resources Institute of Finland (Luke), Natural resources, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: jaakko.repola@luke.fi
  • Karjalainen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: tomikar@uef.fi
  • Packalen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: petteri.packalen@uef.fi
  • Maltamo, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: matti.maltamo@uef.fi
article id 10178, category Research article
Franz Holzleitner, Magdalena Langmaier, Eduard Hochbichler, Bernhardt Obermayer, Karl Stampfer, Christian Kanzian. (2019). Effect of prior tree marking, thinning method and topping diameter on harvester performance in a first thinning operation – a field experiment. Silva Fennica vol. 53 no. 3 article id 10178. https://doi.org/10.14214/sf.10178
Keywords: productivity; time study; harvester; prior tree marking; thinning method; topping diameter; video analysis
Highlights: No effect on harvester performance due to prior tree marking detected; Operator selection versus prior tree marking was assessed; Operator could apply two different thinning methods; Prior tree marking seems to have a positive effect on residual stand damage.
Abstract | Full text in HTML | Full text in PDF | Author Info

The effect of harvester operator tree selection or prior tree marking in thinning operations on satisfactory results and performance has been widely discussed. In harvester operator tree selection, the machine operator decides on the fly which trees are selected to remain or cut. The objective of the study was to analyze the effect of prior tree marking, thinning method and topping diameter on harvester performance in low-diameter thinning operations. The entire thinning operation was captured using video technology. Overall, 2.36 ha divided into 48 plots with 5202 trees were thinned with an average diameter at breast height (dbh) over bark for all plots of between 12.5 and 14.7 cm. In total, 3122 trees were harvested, resulting in 60% removal of stem number over all plots. The harvester achieved a mean productivity of 7.38 m3 PMH0–1 with 1.48 m3 PMH0–1 SEM, with stem volume having the major influence on harvesting productivity. Prior tree marking, topping and thinning method did not significantly affect productivity. Without prior tree marking by the foresters, harvesting removal was shifted toward lower diameters. Within the unmarked plots, 7.0% of the residual trees were damaged compared with 3.2% in marked plots.

  • Holzleitner, University of Natural Resources and Life Sciences Vienna, Department of Forest and Soil Sciences, Institute of Forest Engineering, Peter-Jordan-Straße 82/3, A-1190 Vienna, Austria ORCID https://orcid.org/0000-0001-8489-3050 E-mail: franz.holzleitner@boku.ac.at (email)
  • Langmaier, University of Natural Resources and Life Sciences Vienna, Department of Forest and Soil Sciences, Institute of Silviculture, Peter-Jordan-Straße 82/3, A-1190 Vienna, Austria; Austrian Research Centre for Forests, Department of Forest Growth and Silviculture, Seckendorff Gudent Weg 8, A-1130 Vienna, Austria E-mail: magdalena.langmaier@bfw.gv.at
  • Hochbichler, University of Natural Resources and Life Sciences Vienna, Department of Forest and Soil Sciences, Institute of Silviculture, Peter-Jordan-Straße 82/3, A-1190 Vienna, Austria E-mail: eduard.hochbichler@boku.ac.at
  • Obermayer, University of Natural Resources and Life Sciences Vienna, Department of Forest and Soil Sciences, Institute of Forest Engineering, Peter-Jordan-Straße 82/3, A-1190 Vienna, Austria; Agricultural Technical School of Pyhra, Kyrnbergstraße 4, A-3143 Pyhra, Austria E-mail: bernhardt.obermayer@lfs-pyhra.ac.at
  • Stampfer, University of Natural Resources and Life Sciences Vienna, Department of Forest and Soil Sciences, Institute of Forest Engineering, Peter-Jordan-Straße 82/3, A-1190 Vienna, Austria ORCID http://orcid.org/0000-0001-9350-2859 E-mail: karl.stampfer@boku.ac.at
  • Kanzian, University of Natural Resources and Life Sciences Vienna, Department of Forest and Soil Sciences, Institute of Forest Engineering, Peter-Jordan-Straße 82/3, A-1190 Vienna, Austria ORCID http://orcid.org/0000-0002-1198-9788 E-mail: christian.kanzian@boku.ac.at
article id 10075, category Research article
Matti Maltamo, Marius Hauglin, Erik Naesset, Terje Gobakken. (2019). Estimating stand level stem diameter distribution utilizing harvester data and airborne laser scanning. Silva Fennica vol. 53 no. 3 article id 10075. https://doi.org/10.14214/sf.10075
Keywords: LIDAR; cut-to length harvester; GPS; merchantable volume; tree list
Highlights: Tree level-positioned harvester data were successfully used as plot-level training data for k-nearest neighbor stem diameter distribution modelling applying airborne laser scanning information as predictor variables; Stand-level validation showed that merchantable volume of total tree stock could be estimated with RMSE value of about 9%; The fit of the stem diameter distribution assessed by a variant of Reynold’s error index showed values smaller than 0.2; The most accurate results were obtained for the training plot sizes of 200 m2 and 400 m2.
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Accurately positioned single-tree data obtained from a cut-to-length harvester were used as training harvester plot data for k-nearest neighbor (k-nn) stem diameter distribution modelling applying airborne laser scanning (ALS) information as predictor variables. Part of the same harvester data were also used for stand-level validation where the validation units were stands including all the harvester plots on a systematic grid located within each individual stand. In the validation all harvester plots within a stand and also the neighboring stands located closer than 200 m were excluded from the training data when predicting for plots of a particular stand. We further compared different training harvester plot sizes, namely 200 m2, 400 m2, 900 m2 and 1600 m2. Due to this setup the number of considered stands and the areas within the stands varied between the different harvester plot sizes. Our data were from final fellings in Akershus County in Norway and consisted of altogether 47 stands dominated by Norway spruce. We also had ALS data from the area. We concentrated on estimating characteristics of Norway spruce but due to the k-nn approach, species-wise estimates and stand totals as a sum over species were considered as well. The results showed that in the most accurate cases stand-level merchantable total volume could be estimated with RMSE values smaller than 9% of the mean. This value can be considered as highly accurate. Also the fit of the stem diameter distribution assessed by a variant of Reynold’s error index showed values smaller than 0.2 which are superior to those found in the previous studies. The differences between harvester plot sizes were generally small, showing most accurate results for the training harvester plot sizes 200 m2 and 400 m2.

  • Maltamo, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu E-mail: matti.maltamo@uef.fi (email)
  • Hauglin, Norwegian Institute of Bioeconomy Research, Division of Forest and Forest Resources, P.O. Box 115, 1431 Ås, Norway E-mail: marius.hauglin@nibio.no
  • Naesset, Norwegian University of Life Sciences, Faculty of Environmental Sciences and Natural Resource Management, P.O. Box 5003, 1432 Ås, Norway E-mail: erik.naesset@nmbu.no
  • Gobakken, Norwegian University of Life Sciences, Faculty of Environmental Sciences and Natural Resource Management, P.O. Box 5003, 1432 Ås, Norway E-mail: terje.gobakken@nmbu.no
article id 10074, category Research article
Sebastian Kühle, Alfred Teischinger, Manfred Gronalt. (2019). Optimal location of laminated beech production plants within the solid hardwood supply network in Austria. Silva Fennica vol. 53 no. 3 article id 10074. https://doi.org/10.14214/sf.10074
Keywords: decision support system; facility location; laminated timber products; mixed integer linear programming; supply chain network design
Highlights: This paper provides data to the solid hardwood business and develops a mixed integer linear program model to design a laminated beech wood supply network; It covers the strategic decision where to locate a new production facility within the existing supply network with the lowest supply network cost; Sufficient sawn wood suppliers and potential facility locations are provided.
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Due to changes in forest management in various European countries, hardwood forest areas and amounts will increase. Sustainable and individual utilization concepts have to be developed for the upcoming available resource. Studies conclude that there is low potential for hardwoods in the traditional appearance market thus the application areas have to be extended to new structural innovative products. This paper examines the extension to a future laminated beech wood supply network which would be a combination of already existing and new production facilities. For a better future use of hardwood raw materials it is necessary to consider the entire supply chain. This also better shows a total hardwood value chain. Therefore, this paper provides data to the solid hardwood business and develops a mixed integer linear programming to design a laminated beech wood supply network. The model is applied to Austria as the sample region. It covers the important strategic decisions where to locate a downstream facility within the existing production network with the lowest supply network cost. Fourteen scenarios are developed to examine various future network configurations. Results about optimal material flows and used sawmills as well as downstream production facilities are presented in form of material and financial performances. Two optimal laminated beech production locations are determined by the calculated scenarios results, and the impact of a new sawmill is analyzed which is focused on beech.

  • Kühle, BOKU - University of Natural Resources and Life Sciences, Vienna, Department of Material Science and Process Engineering, and Renewable Institute of Wood Technology Materials, Konrad-Lorenz-Straße 24, 3430 Tulln, Austria E-mail: skuehle@boku.ac.at (email)
  • Teischinger, BOKU - University of Natural Resources and Life Sciences, Vienna, Department of Material Science and Process Engineering, and Renewable Institute of Wood Technology Materials, Konrad-Lorenz-Straße 24, 3430 Tulln, Austria E-mail: alfred.teischinger@boku.ac.at
  • Gronalt, BOKU - University of Natural Resources and Life Sciences, Vienna, Department of Economics and Social Sciences, Institute of Production and Logistics, Feistmantelstraße 4, 1180 Vienna, Austria E-mail: manfred.gronalt@boku.ac.at
article id 10043, category Research article
Outi Manninen, Rainer Peltola. (2019). Continuous picking may increase bilberry yields. Silva Fennica vol. 53 no. 3 article id 10043. https://doi.org/10.14214/sf.10043
Keywords: sustainability; Vaccinium myrtillus; wild forest berries; collectable goods
Highlights: Bilberry fruit production and fruit set increased under continuous picking by rake in three-year study; Bilberry flower number and fruit mass were not affected by picking; Bilberry compensated for biomass loss; The highest relative deciduous species abundance was found in the picking treatment plots at the end of the experiment.
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Accumulated knowledge about the health benefits of bilberry (Vaccinium myrtillus L.) has increased the demand and utilization of wild bilberries. Intensive berry picking by metal rakes is believed to cause damage in bilberry stands in areas under continuous picking pressure, and hence expected to hamper the production of berries in forthcoming years. We conducted an experiment to examine the effect of continuous bilberry picking by metal rake on the number of bilberry flowers and fruits, fruit mass, compensation for biomass loss after picking, and plant functional type abundance in the understorey in northern Finland. Bilberry lost less than 0.5% of its biomass annually during the three-year study period due to rake harvesting. The number of flowers was not significantly affected by damage caused by picking, while both fruit production and fruit set increased without any indication of reduced fruit mass, and biomass loss was fully compensated. Moreover, the relative abundance of plant functional types was not affected by picking during the study. We suggest that the low intensity and timing of damage act as a buffer against the adverse effects of picking on bilberry fruit production. On the basis of this study, it is reasonable to anticipate that there are no indications that current intensive berry picking would not be on a sustainable level.

  • Manninen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Paavo Havaksentie 3, FI-90014 University of Oulu, Finland E-mail: outi.manninen@luke.fi (email)
  • Peltola, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Ounasjoentie 6, FI-96200 Rovaniemi, Finland E-mail: rainer.peltola@luke.fi
article id 10161, category Research article
Jussi Manner, Anders Mörk, Martin Englund. (2019). Comparing forwarder boom-control systems based on an automatically recorded follow-up dataset. Silva Fennica vol. 53 no. 2 article id 10161. https://doi.org/10.14214/sf.10161
Keywords: automation; cut-to-length logging; haulage; crane work; extraction; boom-tip control
Highlights: Boom-tip control saves time compared to the conventional system; This study introduced a field-study design enabling establishment of causal relationships during ordinary forwarding operations; Although the study design requires some extra arrangements, it efficiently combines the representativeness of conventional follow-up datasets with establishment of causal relationships that traditionally have been possible only through observational time studies or standardized experiments.
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Crane work is the most time-consuming work element in forwarding. Hence, assist systems like boom-tip control are of interest. The first commercially available boom-tip control for forwarders was introduced in 2013. In this study we analysed whether replacing conventional boom control (CBC) with John Deere’s version of boom-tip control (named Intelligent Boom Control, IBC), increases crane-work productivity. We used data automatically gathered from 10 final-felling stands, covering typical logging conditions for southern, central and northern Sweden. Two John Deere 1510E and two John Deere 1910G forwarders were operated by seven experienced operators during the follow-up study, covering 1238 loads in total. A split-plot design was applied to isolate effects of the boom-control system being used (CBC, IBC). We found that using IBC for loading work (crane work and driving included) saved 5.2% of productive machine time compared to using CBC (p ≤ 0.05). The corresponding saving when using IBC for unloading work was 7.9% (p ≤ 0.05). Depending on geophysical factors, this corresponds to approximately 4% savings in productive machine time for forwarding as a whole, including pure transport (with and without load). Moreover, the study introduced in cut-to-length context a novel field-study design to collect a large follow-up dataset in the course of ordinary forwarding operations. We found the study design to be a cost-efficient way to combine the representativeness of conventional follow-up datasets with the ability to establish causal relationships. Establishment of causal relationships has traditionally been possible only through observational time studies or standardized experiments.

  • Manner, The Forestry Research Institute of Sweden (Skogforsk), Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID http://orcid.org/0000-0002-4982-3855 E-mail: jussi.manner@skogforsk.se (email)
  • Mörk, The Forestry Research Institute of Sweden (Skogforsk), Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: anders.mork@skogforsk.se
  • Englund, The Forestry Research Institute of Sweden (Skogforsk), Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: martin.englund@skogforsk.se
article id 10150, category Research article
Petri Forsström, Jouni Peltoniemi, Miina Rautiainen. (2019). Seasonal dynamics of lingonberry and blueberry spectra. Silva Fennica vol. 53 no. 2 article id 10150. https://doi.org/10.14214/sf.10150
Keywords: understory; remote sensing; boreal forest vegetation; berries; flowers; goniometer; FIGIFIGO
Highlights: Seasonal series of multiangular spectra for lingonberry (Vaccinium vitis-idaea L.) and blueberry (Vaccinium myrtillus L.); Decidous blueberry has strong seasonal pattern while temporal variations of evergreen lingonberry were linked to phenological stages of flowering and berrying; Detection of flowers and berries from shrub spectra was possible; Collected spectral data are openly available through SPECCHIO Spectral Information System.
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Accurate mapping of the spatial distribution of understory species from spectral images requires ground reference data which represent the prevailing phenological stage at the time of image acquisition. We measured the spectral bidirectional reflectance factors (BRFs, 350–2500 nm) at varying view angles for lingonberry (Vaccinium vitis-idaea L.) and blueberry (Vaccinium myrtillus L.) throughout the growing season of 2017 using Finnish Geospatial Research Institute’s FIGIFIGO field goniometer. Additionally, we measured spectra of leaves and berries of both species, and flowers of lingonberry. Both lingonberry and blueberry showed seasonality in visible and near-infrared spectral regions which was linked to occurrences of leaf growth, flowering, berrying, and leaf senescence. The seasonality of spectra differed between species due to different phenologies (evergreen vs. deciduous). Vegetation indices, normalized difference vegetation index (NDVI), moisture stress index (MSI), plant senescence reflectance index (PSRI), and red-edge inflection point (REIP2), showed characteristic seasonal trends. NDVI and PSRI were sensitive to the presence of flowers and berries of lingonberry, while with blueberry the effects were less evident. Off-nadir observations supported differentiating the dwarf shrub species from each other but showed little improvement for detection of flowers and berries. Lingonberry and blueberry can be identified by their spectral signatures if ground reference data are available over the entire growing season. The spectral data measured in this study are reposited in the publicly open SPECCHIO Spectral Information System.

  • Forsström, Aalto University, School of Engineering, Department of Built Environment, FI-00076 Aalto, Finland ORCID https://orcid.org/0000-0002-2357-2517 E-mail: petri.forsstrom@aalto.fi (email)
  • Peltoniemi, Finnish Geospatial Research Institute (FGI), Department of Geodesy and Geodynamics, Geodeetinrinne 2, FI-02430 Masala, Finland ORCID https://orcid.org/0000-0002-4701-128X E-mail: jouni.peltoniemi@nls.fi
  • Rautiainen, Aalto University, School of Engineering, Department of Built Environment, FI-00076 Aalto, Finland; Aalto University, Department of Electronics and Nanoengineering, FI-00076 Aalto, Finland ORCID https://orcid.org/0000-0002-6568-3258 E-mail: miina.a.rautiainen@aalto.fi
article id 10134, category Research article
Matti Sirén, Jari Ala-Ilomäki, Harri Lindeman, Jori Uusitalo, Kalle E.K. Kiilo, Aura Salmivaara, Ari Ryynänen. (2019). Soil disturbance by cut-to-length machinery on mid-grained soils. Silva Fennica vol. 53 no. 2 article id 10134. https://doi.org/10.14214/sf.10134
Keywords: rut formation; soil compaction; sandy soil; silty soil; harvesting damage
Highlights: The number of machine passes, volumetric water content in the mineral soil and the depth of the organic layer were the controlling factors for rut formation; The harvester rut depth was a good predictor of the forwarder rut formation; Changes in the penetration resistance were highest at depths of 20–40 cm.
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Factors affecting soil disturbance caused by harvester and forwarder were studied on mid-grained soils in Finland. Sample plots were harvested using a one-grip harvester. The harvester operator processed the trees outside the strip roads, and the remaining residues were removed to exclude the covering effect of residues. Thereafter, a loaded forwarder made up to 5 passes over the sample plots. The average rut depth after four machine passes was positively correlated to the volumetric water content at a depth of 0–10 cm in mineral soil, as well as the thickness of the organic layer and the harvester rut depth, and negatively correlated with penetration resistance at depths of both 0–20 cm and 5–40 cm. We present 5 models to predict forwarder rut depth. Four include the cumulative mass driven over a measurement point and combinations of penetration resistance, water content and the depth of organic layer. The fifth model includes harvester rut depth and the cumulative overpassed mass and provided the best fit. Changes in the penetration resistance (PR) were highest at depths of 20–40 cm. Increase in BD and VWC decreased PR, which increased with total overdriven mass. After four to five machine passes PR values started to stabilize.

  • Sirén, Natural Resources Institute Finland (Luke) c/o Aalto University, P.O. Box 15600, FI-00076 Aalto, Finland E-mail: matti.siren@luke.fi (email)
  • Ala-Ilomäki, Natural Resources Institute Finland (Luke) c/o Aalto University, P.O. Box 15600, FI-00076 Aalto, Finland ORCID http://orcid.org/0000-0002-6671-7624 E-mail: jari.ala-ilomaki@luke.fi
  • Lindeman, Natural Resources Institute Finland (Luke), Korkeakoulunkatu 7, FI-33720 Tampere, Finland E-mail: harri.lindeman@luke.fi
  • Uusitalo, Natural Resources Institute Finland (Luke), Korkeakoulunkatu 7, FI-33720 Tampere, Finland ORCID http://orcid.org/0000-0003-3793-1215 E-mail: jori.uusitalo@luke.fi
  • Kiilo, Versowood, Teollisuuskatu 1, FI-11130 Riihimäki, Finland E-mail: kalle.kiilo@versowood.fi
  • Salmivaara, Natural Resources Institute Finland (Luke), P.O. Box 2, FI-00791 Helsinki, Finland E-mail: aura.salmivaara@luke.fi
  • Ryynänen, Natural Resources Institute Finland (Luke), Kaironiementie 15, FI-39700 Parkano, Finland E-mail: ari.ryynanen@luke.fi
article id 10132, category Research article
Yan Liu, Yuan Zhang, Qing Zhou, Jian Wu, Pingdong Zhang. (2019). Colchicine did not affect the viability of induced 2n pollen in Populus tomentosa. Silva Fennica vol. 53 no. 2 article id 10132. https://doi.org/10.14214/sf.10132
Keywords: colchicine-induced 2n pollen; pollen production; pollen viability; pollen germination rate; triploid
Highlights: The number of colchicine injections and the meiotic stage at which they were administered both had a significant effect on the occurrence rate of induced 2n pollen in Populus tomentosa; Treating male buds with 5000 ppm colchicine solution during meiosis led to a significant decrease in pollen production; Colchicine injection could induce P. tomentosa to produce 2n pollen and did not lead to dysfunction of induced diploid pollen.
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Colchicine is widely used as a mutagen to induce production of diploid gametes in plants. However, whether colchicine affects induced pollen viability remains unclear. To clarify whether colchicine affected the viability of induced pollen, we induced production of diploid pollen by colchicine, followed by pollen germination in vitro and crossing induced pollen with normal gametes to produce triploid in Populus tomentosa Carrière. The results showed that the predominant meiotic stages and the number of colchicine injections had significant effects on the occurrence rates of induced 2n pollen. When the colchicine injection was given at diakinesis, a significant decrease in the pollen production per bud was observed (p < 0.001). The morphology of the colchicine-induced 2n pollen was similar to that of the natural 2n pollen in its ectexine structure. The pollen germination experiments revealed that there was also no significant difference in germination rates between the induced diploid pollen and natural 2n pollen grains, and 68 triploids were created by crossing colchicine-induced pollen. Our findings revealed that colchicine injection could induce P. tomentosa to produce 2n pollen and will not lead to dysfunction of induced diploid pollen.

  • Liu, National Engineering Laboratory for Tree Breeding, Beijing Forestry University, Beijing 100083, China; College of Bioscience and Biotechnology, Beijing Forestry University, Beijing 100083, China E-mail: 342767649@qq.com
  • Zhang, National Engineering Laboratory for Tree Breeding, Beijing Forestry University, Beijing 100083, China; College of Bioscience and Biotechnology, Beijing Forestry University, Beijing 100083, China E-mail: 409192881@qq.com
  • Zhou, National Engineering Laboratory for Tree Breeding, Beijing Forestry University, Beijing 100083, China; College of Bioscience and Biotechnology, Beijing Forestry University, Beijing 100083, China E-mail: 876034493@qq.com
  • Wu, National Engineering Laboratory for Tree Breeding, Beijing Forestry University, Beijing 100083, China; College of Bioscience and Biotechnology, Beijing Forestry University, Beijing 100083, China E-mail: 1269485709@qq.com
  • Zhang, National Engineering Laboratory for Tree Breeding, Beijing Forestry University, Beijing 100083, China; College of Bioscience and Biotechnology, Beijing Forestry University, Beijing 100083, China E-mail: zhangpd@bjfu.edu.cn (email)
article id 10089, category Research article
Arto Haara, Annika Kangas, Sakari Tuominen. (2019). Economic losses caused by tree species proportions and site type errors in forest management planning. Silva Fennica vol. 53 no. 2 article id 10089. https://doi.org/10.14214/sf.10089
Keywords: forest inventory; value of information; uncertainty; sub-optimality loss
Highlights: Errors in tree species proportions caused more economic losses for forest owners than site type errors; Economic losses due to sub-optimal treatments were observed from 26.5% to 31.7% of plots, depending on the remote sensing data set used; Even with the most accurate remote sensing data set, namely ALS data set, NPV losses were on average 124.4 € ha–1 with 3% interest rate.
Abstract | Full text in HTML | Full text in PDF | Author Info

The aim of this study was to estimate economic losses, which are caused by forest inventory errors of tree species proportions and site types. Our study data consisted of ground truth data and four sets of erroneous tree species proportions. They reflect the accuracy of tree species proportions in four remote sensing data sets, namely 1) airborne laser scanning (ALS) with 2D aerial image, 2) 2D aerial image, 3) 3D and 2D aerial image data together and 4) satellite data. Furthermore, our study data consisted of one simulated site type data set. We used the erroneous tree species proportions to optimise the timing of forest harvests and compared that to the true optimum obtained with ground truth data. According to the results, the mean losses of Net Present Value (NPV) because of erroneous tree species proportions at an interest rate of 3% varied from 124.4 € ha–1 to 167.7 € ha–1. The smallest losses were observed using tree species proportions predicted using ALS data and largest using satellite data. In those stands, respectively, in which tree species proportion errors actually caused economic losses, they were 468 € ha–1 on average with tree species proportions based on ALS data. In turn, site type errors caused only small losses. Based on this study, accurate tree species identification seems to be very important with respect to operational forest inventory.

  • Haara, Natural Resources Institute Finland (Luke), Bioeconomy and environment, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: arto.haara@luke.fi (email)
  • Kangas, Natural Resources Institute Finland (Luke), Bioeconomy and environment, P.O. Box 68, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0002-8637-5668 E-mail: annika.kangas@luke.fi
  • Tuominen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, P.O. Box 2, FI-00791 Helsinki, Finland ORCID https://orcid.org/0000-0001-5429-3433 E-mail: sakari.tuominen@luke.fi
article id 10088, category Research article
Antti-Jussi Lindroos, Kira Ryhti, Tomi Kaakkurivaara, Jori Uusitalo, Heljä-Sisko Helmisaari. (2019). Leaching of heavy metals and barium from forest roads reinforced with fly ash. Silva Fennica vol. 53 no. 2 article id 10088. https://doi.org/10.14214/sf.10088
Keywords: recycling; lysimeter; fly ash; forest road rehabilitation; environmental impact assessment; low-volume road
Highlights: Heavy metal concentrations were generally low in percolation and ditch water samples of ash roads, but elevated concentrations were found in some parts of ash roads; Risk for heavy metal leaching is negligible if road parts producing high concentrations are rare.
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The aim of this study was to determine the effect of leaching of heavy metals (Cr, As, Cd, Cu, Ni, Pb, Zn, Co, Mo) and earth-alkaline metal, barium (Ba), on the percolation and ditch water quality from the forest roads that contained ash in the road structures. Water quality was studied in the immediate vicinity below the ash layers as well as deeper in the road structure. Water quality was also determined in the drainage water in ditches that crossed the forest roads. A mixture of wood and peat based fly ash was used in the road structures. The treatments were: 1) no ash, 2) a 15 cm layer of ash/gravel mixture, 3) a 20 cm layer of ash/gravel mixture, 4) a 25 cm layer of ash, and 5) a 50 cm layer of ash. Large variation in the concentrations of Cr, As, Cu, Ni, Pb, Mo and Ba in the percolation water, even within the same treatment, caused difficulties to generalize the results. The concentrations of Cr, As, Ni, Pb, Mo and Ba in water samples were high in some treatment plot lysimeters containing ash compared to the control (no ash). On the other hand, many lysimeters had low and similar concentrations in water samples in the treatment plots containing ash compared to concentrations in the control plots. The ash in the roads did not affect the concentrations in the ditches. The leaching is uneven and seems to take place only from some parts of the ash layer. Risk for leaching is minimal if such parts are not widely spread.

  • Lindroos, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: antti.lindroos@luke.fi (email)
  • Ryhti, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: kira.ryhti@helsinki.fi
  • Kaakkurivaara, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: tomi.kaakkurivaara@gmail.com
  • Uusitalo, Natural Resources Institute Finland (Luke), Korkeakoulunkatu 7, FI-33720 Tampere, Finland E-mail: jori.uusitalo@luke.fi
  • Helmisaari, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: helja-sisko.helmisaari@helsinki.fi
article id 10068, category Research article
Lari Melander, Risto Ritala, Markus Strandström. (2019). Classifying soil stoniness based on the excavator boom vibration data in mounding operations. Silva Fennica vol. 53 no. 2 article id 10068. https://doi.org/10.14214/sf.10068
Keywords: spot mounding; activity recognition; stoniness classification; supervised machine learning
Highlights: An excavator was equipped with an inertial measurement unit for taking automatic measurements of soil stoniness during mounding work; Supervised machine-learning classifiers were trained utilizing both the automatically measured data and manual stoniness measurements; The class prediction for the soil stoniness achieved an accuracy of 70% when assigned to constant grid cells.
Abstract | Full text in HTML | Full text in PDF | Author Info

The stoniness index of forest soil describes the stone content in the upper soil layer at depths of 20–30 centimeters. This index is not available in any existing map databases, and traditional measurements for the stoniness of the soil have always necessitated laborious soil-penetration methods. Knowledge of the stone content of a forest site could be of use in a variety of forestry operations. This paper presents a novel approach to obtaining automatic measurements of soil stoniness during an excavator-based mounding operation. The excavator was equipped with only a low-cost inertial measurement unit and a satellite navigation receiver. Using the data from these sensors and manually conducted soil stoniness measurements, supervised machine learning methods were utilized to build a model that is capable of predicting the stoniness class of a given mounding location. This study compares different classifiers and feature selection methods to find the most promising solution for this learning problem. The discussion includes a proposition for a meaningful measurement resolution of the soil’s stoniness, and a practical method for evaluating the variability of the stone content of the soil. The results indicate that it is possible to predict the soil stoniness class with 70% accuracy using only the inertial and location measurements.

  • Melander, Automation Technology and Mechanical Engineering, Tampere University, FI-33014 Tampere University, Finland ORCID http://orcid.org/0000-0003-3662-5187 E-mail: lari.melander@tuni.fi (email)
  • Ritala, Automation Technology and Mechanical Engineering, Tampere University, FI-33014 Tampere University, Finland ORCID http://orcid.org/0000-0003-0721-9948 E-mail: risto.ritala@tuni.fi
  • Strandström, Metsäteho Oy, Vernissakatu 1, FI-01300 Vantaa, Finland E-mail: markus.strandstrom@metsateho.fi
article id 10048, category Research article
Urszula Zajączkowska, Karina Kaczmarczyk, Janusz Liana. (2019). Birch sap exudation: influence of tree position in a forest stand on birch sap production, trunk wood anatomy and radial bending strength. Silva Fennica vol. 53 no. 2 article id 10048. https://doi.org/10.14214/sf.10048
Keywords: biomechanics; wood anatomy; forest edge; xylem sap
Highlights: Birch trees along the forest edge exude more xylem sap but less concentrated than the trees from the interior; Radial bending strength of wood in birch trunk is higher in the trees from forest edge; Trees exhibit higher bending strength in western side of the trunk, where the number of vessels and the wood potential conductivity index are smaller.
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It is commonly accepted that the period of early-spring xylem sap exudation marks a stage during which a positive pressure builds inside the tree trunks. This state changes when leaves appear, initiating water transport within the trunk. It is unknown, however, how the wood anatomical structure and its mechanical resistance influences the sap. We present the results of research on the relationship between exudation of sap from Betula pendula Roth trees from the interior of a forest stand and from its edge, and the anatomical structure of the trunk wood and its bending strength. During the period between March 21 and April 18, we performed five sets of measurements of sap exudation from trees at the edge of the stand and from the forest interior. The resulting radial wood samples were tested for bending strength using a fractometer. We tested the sap for electrolytic conductivity and sugars content. For the anatomical analysis of the wood, we determined the number of vessels per 1 mm2, average vessel lumen area and potential conductivity index. We found that the trees along the edge of the stand exude more sap, but it is less concentrated than the sap from the trees from the interior. Bending strength perpendicular to wood fibres is higher in the trees from the stand edge and in the western side of the trunk, where the number of vessels per 1 mm2 and conductivity index are smaller. Seemingly, this is the result of western winds, which are dominant in Poland.

  • Zajączkowska, Department of Forest Botany, Faculty of Forestry, Warsaw University of Life Sciences, 159 Nowoursynowska St., 02-776 Warsaw, Poland E-mail: urszula.zajaczkowska@wl.sggw.pl (email)
  • Kaczmarczyk, Department of Forest Botany, Faculty of Forestry, Warsaw University of Life Sciences, 159 Nowoursynowska St., 02-776 Warsaw, Poland E-mail: karina.kaczmarczyk@wl.sggw.pl
  • Liana, Department of Forest Botany, Faculty of Forestry, Warsaw University of Life Sciences, 159 Nowoursynowska St., 02-776 Warsaw, Poland E-mail: janusz.liana@wl.sggw.pl
article id 10076, category Research article
Tore Skrøppa, Arne Steffenrem. (2019). Genetic variation in phenology and growth among and within Norway spruce populations from two altitudinal transects in Mid-Norway. Silva Fennica vol. 53 no. 1 article id 10076. https://doi.org/10.14214/sf.10076
Keywords: Picea abies; adaptation; tree breeding; height; clinal variation; populations; families; bud flush
Highlights: Norway spruce populations distributed along each of two altitudinal transects showed strong clinal relationships between the annual mean temperatures of the sites of the populations and height and phenology traits in short term tests and height in field trials; Large variation was present among families within populations for height and phenology traits and with a wider range within than among populations; Correlation patterns among traits were different for provenances and families.
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Progenies from open pollinated cones collected in natural populations of Norway spruce (Picea abies (L.) Karst.) distributed along two altitudinal transects in Mid-Norway were tested in the nursery, in short term tests and in long-term field trials. The populations showed clinal variation related to the mean annual temperatures of the populations, with the earliest bud flush and cessation of shoot elongation and lowest height at age nine years for the high altitude populations. Within population variation was considerable as the narrow sense heritability for these traits was 0.67, 0.31 and 0.09 in one transect and 0.55, 0.18 and 0.14 in the other transect, respectively. Lammas shoots occurred in the short term trials with large variation in frequency between years. There was significant family variation for this trait, but also interactions between populations and year. The variance within populations was considerably larger in the populations from low altitude compared to the high-altitude populations. Significant genetic correlations between height and phenology traits and damage scores indicate that families flushing early and ceasing growth late were taller. Taller families also had higher frequencies of damages. Selection of the top 20% families for height growth in short term tests at age nine years gave a simulated gain of 11% increased height growth at age 18 years in long term trials at altitudes similar to those of origin of the populations. The gain was negative when high altitude populations were selected based on testing in the lowland.

  • Skrøppa, Norwegian Institute of Bioeconomy Research, P.O. Box 115, 1431 Ås, Norway E-mail: tore.skroppa@nibio.no (email)
  • Steffenrem, E-mail: as@nn.no
article id 10062, category Research article
Jouni Siipilehto, Miika Rajala. (2019). Model for diameter distribution from assortments volumes: theoretical formulation and a case application with a sample of timber trade data for clear-cut sections. Silva Fennica vol. 53 no. 1 article id 10062. https://doi.org/10.14214/sf.10062
Keywords: bucking; optimization; simplex method; truncated Weibull function
Highlights: The Weibull distribution was solved successfully from assortment volumes using optimization; The solved distribution provided accurate assortment volume when the input variables were correct; Goodness-of-fit tests indicate the compatibility between the solved distribution and the cut trees, according to harvester data; Timber trade contracts showed overestimated average merchantable tree sizes, which resulted in an underestimation of the number of cut trees; The reason for underestimation seemed to be in the decreasing distributions.
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This study examined a theoretical model for stand structures from the volumes of pulpwood and saw logs of clear-cut stands. The average stem size was used to estimate the number of cut trees. The distribution was solved using nonlinear derivative-free optimization. The truncated 2-parameter Weibull distribution was used to describe the stand structure of the commercial stems. This method was first tested with harvester data collected from seven clear-cut stands in southern Finland. Validation included reliability in the stand characteristics and goodness-of-fit of the species-specific distributions. The distributions provided unbiased estimates for the saw log volume, while the bias in the estimated pulpwood volume was 2%. The standard stand characteristics from the Weibull distributions corresponded notably well with the harvester data. A Kolmogorov-Smirnov (KS) test rejected two distributions out of 21 cases, when the accurate input variables were available for the theoretical model. The results of the study suggest that the presented method is a relevant option for predicting the stand structure. In practice, the reliability of the presented method was dependent on the quality of the information available from the stand prior to cutting. With a timber trade data set, the solution for the distribution for a clear-cut section was found. The goodness-of-fit was dependent on the accuracy of the visually assessed timber trade variables. Especially the average stem size proved difficult to assess due to high number of understorey pulpwood stems. Due to overestimated average stem sizes, the solved number of harvested trees was underestimated. Less than 50% of the distributions predicted for clear-cut sections passed the KS test.

  • Siipilehto, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, P.O. Box 2, FI-00790 Helsinki, Finland E-mail: jouni.siipilehto@luke.fi (email)
  • Rajala, Metsä Group, Revontulenpuisto 2, P.O. Box 10, 02020 METSÄ, FI-02100 Espoo, Finland E-mail: miika.rajala@metsagroup.com
article id 10052, category Research article
Pentti Niemistö, Harri Kilpeläinen, Henrik Heräjärvi. (2019). Effect of pruning season and tool on knot occlusion and stem discolouration in Betula pendula – situation five years after pruning. Silva Fennica vol. 53 no. 1 article id 10052. https://doi.org/10.14214/sf.10052
Keywords: silver birch; pruning saw; discolouration; healing-over; secateurs; stick pruning
Highlights: The occlusion was the fastest in the case of small living branches of fast growing trees pruned in springtime; Occlusion was quicker after saw pruning than after secateurs pruning, due to shorter knot stubs; Branches that were pruned in living state occluded faster than the ones pruned as dead; Dead branches hit down with a stick occluded slowly.
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This paper investigates and models the effects of pruning season and tool on wound occlusion with varying tree and branch characteristics of silver birch (Betula pendula Roth) stems at the pruning height of 0−4 metres. Dates of eight secateurs prunings, three saw prunings and two sticks prunings as well as unpruned control were tested in permanent plots on four sites. Knot occlusion and discolouration in stemwood were measured from about 1600 studied knots of 112 sample trees felled five to six years after pruning in 2010. Knot occlusion rate was modelled according to pruning tool, date, tree growth, and branch characteristics. The occlusion was the fastest in trees pruned in spring or early summer, and the slowest in trees pruned in autumn. Stubs of living branches occluded faster than the dead ones with the same diameter. Saw pruning resulted in clearly better occlusion rates than secateurs pruning, caused by the shorter knot stubs after saw pruning. Hitting dead branches away with a stick resulted in the worst occlusion status. The colour defects spread more often upward from the knot than downward. Discolouration in stemwood was detected more frequently near to the pruned branches than the unpruned ones, and more widely near to the stubs of dead branches than the living ones. Most saw and secateurs pruned branches were completely occluded during the experiment, so these prunings were suitable for all branches under 20 mm in diameter, and for living branches even up to 30 mm in fast-growing trees.

  • Niemistö, Natural Resources Institute Finland (Luke), Natural resources, Kampusranta 9 C, FI-60320 Seinäjoki, Finland E-mail: pentti.niemisto@luke.fi (email)
  • Kilpeläinen, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6 B, FI-80100 Joensuu, Finland E-mail: harri.kilpelainen@luke.fi
  • Heräjärvi, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6 B, FI-80100 Joensuu, Finland E-mail: henrik.herajarvi@luke.fi
article id 10050, category Research article
Jori Uusitalo, Jari Ala-Ilomäki, Harri Lindeman, Jenny Toivio, Matti Siren. (2019). Modelling soil moisture – soil strength relationship of fine-grained upland forest soils. Silva Fennica vol. 53 no. 1 article id 10050. https://doi.org/10.14214/sf.10050
Keywords: cone index; penetration resistance; shear strength; soil bulk density; VWC
Highlights: Penetration resistance (PR) is best predicted with moisture content (MC), bulk density and clay content; In fully saturated silty or clayey soils PR range from 600 to 800 kPa; The models can be linked with mobility models predicting rutting of forest machines.
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The strength of soil is known to be dependent on water content but the relationship is strongly affected by the type of soil. Accurate moisture content – soil strength models will provide forest managers with the improved ability to reduce soil disturbances and increase annual forest machine utilization rates. The aim of this study was to examine soil strength and how it is connected to the physical properties of fine-grained forest soils; and develop models that could be applied in practical forestry to make predictions on rutting induced by forest machines. Field studies were conducted on two separate forests in Southern Finland. The data consisted of parallel measurements of dry soil bulk density (BD), volumetric water content (VWC) and penetration resistance (PR). The model performance was logical, and the results were in harmony with earlier findings. The accuracy of the models created was tested with independent data. The models may be regarded rather trustworthy, since no significant bias was found. Mean absolute error of roughly 20% was found which may be regarded as acceptable taken into account the character of the penetrometer tool. The models can be linked with mobility models predicting either risks of rutting, compaction or rolling resistance.

  • Uusitalo, Natural Resources Institute Finland (Luke), Production systems, Korkeakoulunkatu 7, FI-33720 Tampere, Finland E-mail: jori.uusitalo@luke.fi (email)
  • Ala-Ilomäki, Natural Resources Institute Finland (Luke), Production systems Maarintie 6, FI-02150 Espoo, Finland E-mail: jari.ala-ilomaki@luke.fi
  • Lindeman, Natural Resources Institute Finland (Luke), Production systems, Korkeakoulunkatu 7, FI-33720 Tampere, Finland E-mail: harri.lindeman@luke.fi
  • Toivio, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: toiviojenny@gmail.com
  • Siren, Natural Resources Institute Finland (Luke), Production systems Maarintie 6, FI-02150 Espoo, Finland E-mail: matti.siren@luke.fi
article id 10019, category Research article
Junyan Liu, Junfeng Tang, Si-Chong Chen, Wenbao Ma, Zheng Zheng, Tingfa Dong. (2019). Do tree cavity density and characteristics vary across topographical habitats in the tropics? A case study from Xishuangbanna, southwest China. Silva Fennica vol. 53 no. 1 article id 10019. https://doi.org/10.14214/sf.10019
Keywords: heterogeneity; cavity-dependant animals; tropical rainforest; biodiversity conservation
Highlights: Cavities were significantly more abundant in high- and low-slope than high-plateau habitats; There are more “butt hollow” cavities in high-slope habitat and they occurred at a lower height; More “crack” cavities in low-slope habitat and they had a narrower entrance diameter; Certain types of cavities are concentrated in specific habitats, which provide opportunities for forest management and biodiversity conservation.
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Despite the influence of cavities on the survival and distribution of cavity-dependent fauna, the variation in the density and characteristics of tree cavities across different habitat types in tropical forests is unknown. In this study, we surveyed 26 312 living trees from 376 species and compared cavity density and characteristics (height, size, type, and orientation) across five habitat types (valley, low-slope, high-slope, high-gully, and high-plateau) in a 20-hectare tropical rainforest in southwest China. From a total of 2047 cavities, we found that cavity density was mainly driven by habitat rather than tree species richness or diameter at breast height (DBH), and the characteristics of cavities were not uniformly distributed across habitats. Cavities were significantly more abundant in high- and low-slope than high-plateau habitats. Compared with other habitats, more “butt hollow” cavity types were found in high-slope habitat and they occurred at a lower tree height, whereas more “crack” cavities were found in low-slope habitat and they had a narrower entrance diameter. Although the mean orientation of cavities faced towards the northeast, cavity orientation varied significantly across habitat types. Our results indicate that certain types of cavities are concentrated in specific habitat types, which can provide avenues for forest management and biodiversity conservation. We highlight the importance of habitat heterogeneity in providing resources for cavity nesters.

  • Liu, Key Laboratory of Southwest China Wildlife Resources Conservation of Ministry of Education, Key Laboratory of Environmental Science and Biodiversity Conservation (Sichuan Province) and Institute of Plant Adaptation and Utilization in Southwest Mountains, China West Normal University, Nanchong, Sichuan 637009, China; Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla, Yunnan 666303, China E-mail: liujunyan2300@163.com
  • Tang, Key Laboratory of Southwest China Wildlife Resources Conservation of Ministry of Education, Key Laboratory of Environmental Science and Biodiversity Conservation (Sichuan Province) and Institute of Plant Adaptation and Utilization in Southwest Mountains, China West Normal University, Nanchong, Sichuan 637009, China E-mail: jft@nn.ch
  • Chen, Royal Botanic Gardens, Kew, Wakehurst Place, West Sussex RH17 6TN, UK; Mitrani Department of Desert Ecology, Swiss Institute for Dryland Environmental and Energy Research, Jacob Blaustein Institutes for Desert Research, Ben-Gurion University of the Negev, Midreshet Ben-Gurion, Beer-Sheva 8499000, Israel E-mail: chensichong0528@gmail.com
  • Ma, Ecological Restoration and Conservation of Forests and Wetlands Key Laboratory of Sichuan Province, Sichuan Academy of Forestry, Chengdu 610081, China E-mail: mawenbao_2000@126.com
  • Zheng, Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla, Yunnan 666303, China E-mail: dioeco@outlook.com
  • Dong, Key Laboratory of Southwest China Wildlife Resources Conservation of Ministry of Education, Key Laboratory of Environmental Science and Biodiversity Conservation (Sichuan Province) and Institute of Plant Adaptation and Utilization in Southwest Mountains, China West Normal University, Nanchong, Sichuan 637009, China E-mail: dongtf@aliyun.com (email)
article id 9968, category Research article
Hubert Lachowicz, Anna Bieniasz, Rafał Wojtan. (2019). Variability in the basic density of silver birch wood in Poland. Silva Fennica vol. 53 no. 1 article id 9968. https://doi.org/10.14214/sf.9968
Keywords: Betula pendula; tree age; forest habitat type; thickness of trees; geographical location; physical properties of wood
Highlights: Location, tree age and forest habitat type, and the interactions between those factors, have a statistically significant impact on the basic density of silver birch wood; The average basic density of silver birch wood increases with the age of the tree.
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This work presents the findings of a study concerning variability in the basic density of silver birch (Betula pendula Roth) wood, depending on the geographical location of tree stands, the age and thickness of the trees, the forest habitat type, and interactions between some of these factors. The study was carried out on wood from trees aged approximately 30, 50 and 70 years in 12 forest districts located throughout Poland. In total 4777 wood samples, taken from 306 trees from 51 test plots, were measured. The location, the age of the trees, the thickness of the trees and the forest habitat type, as well as interactions between these factors, proved to have a significant influence on the basic density of silver birch wood. The highest mean values of the basic density of the birch wood were found in Sokołów forest district on the FBF habitat type (549 kg m–3) and in Giżycko forest district on the FMBF habitat type (548 kg m–3). For the entire set of examined material, the average values of the basic density of wood increase with tree age. For the examined material originating in FBF and FMBF habitats the average values of basic density showed no significant differences; however, in the cases of the forest districts of Giżycko, Łobez and Rudziniec, significant differences in the analysed property were observed.

  • Lachowicz, Department of Forest Utilization, Warsaw University of Life Sciences SGGW, Nowoursynowska 159 02-787 Warsaw, Poland E-mail: hubert.lachowicz@wl.sggw.pl
  • Bieniasz, Department of Forest Utilization, Warsaw University of Life Sciences SGGW, Nowoursynowska 159 02-787 Warsaw, Poland E-mail: anna.bieniasz@wl.sggw.pl (email)
  • Wojtan, Laboratory of Dendrometry and Forest Productivity, Warsaw University of Life Sciences SGGW, Nowoursynowska 159 02-787 Warsaw, Poland E-mail: rwojtan@wl.sggw.pl
article id 9918, category Research article
Ari Nikula, Vesa Nivala, Juho Matala, Kari Heliövaara. (2019). Modelling the effect of habitat composition and roads on the occurrence and number of moose damage at multiple scales. Silva Fennica vol. 53 no. 1 article id 9918. https://doi.org/10.14214/sf.9918
Keywords: forestry; Alces alces; damage probability; forest damage; forest plantation; habitat selection; habitat modelling; zero-inflated negative binomial distribution
Highlights: The occurrence and number of moose damage were modelled with a zero-inflated count model; An admixture of mature forests within plantations increased the number of damage; Vicinity of inhabited areas and roads reduced damage; Plantations in landscapes with a large amount of pine-dominated thinning forests had less damage in Lapland; Damage risk assessment should include characteristics specific to each region.
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We modelled the effect of habitat composition and roads on the number and occurrence of moose (Alces alces L.) damage in Ostrobothnia and Lapland using a zero-inflated count model. Models were developed for 1 km2, 25 km2 and 100 km2 landscapes consisting of equilateral rectangular grid cells. Count models predict the number of damage, i.e. the number of plantations and zero models the probability of a landscape being without damage for a given habitat composition. The number of moose damage in neighboring grid cells was a significant predictor in all models. The proportion of mature forest was the most frequent significant variable, and an increasing admixture of mature forests among plantations increased the number and occurrence of damage. The amount of all types of plantations was the second most common significant variable predicting increasing damage along with increasing amount of plantations. An increase in thinning forests as an admixture also increased damage in 1 km2 landscapes in both areas, whereas an increase in pine-dominated thinning forests in Lapland reduced the number of damage in 25 km2 landscapes. An increasing amount of inhabited areas in Ostrobothnia and the length of connecting roads in Lapland reduced the number of damage in 1 and 25 km2 landscapes. Differences in model variables between areas suggest that models of moose damage risk should be adjusted according to characteristics that are specific to the study area.

  • Nikula, Natural Resources Institute Finland (Luke), Bioeconomy and Environment, Ounasjoentie 6, FI-96200 Rovaniemi, Finland E-mail: ari.nikula@luke.fi (email)
  • Nivala, Natural Resources Institute Finland (Luke), Bioeconomy and Environment, Ounasjoentie 6, FI-96200 Rovaniemi, Finland E-mail: vesa.nivala@luke.fi
  • Matala, Natural Resources Institute Finland (Luke), Natural resources, Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: juho.matala@luke.fi
  • Heliövaara, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: kari.heliovaara@helsinki.fi
article id 10055, category Research article
Jaakko Repola, Hannu Hökkä, Hannu Salminen. (2018). Models for diameter and height growth of Scots pine, Norway spruce and pubescent birch in drained peatland sites in Finland. Silva Fennica vol. 52 no. 5 article id 10055. https://doi.org/10.14214/sf.10055
Keywords: Pinus sylvestris; Picea abies; Betula pubescens; drained peatlands; forest drainage; mixed model; tree growth model
Highlights: Tree growth strongly correlated with site drainage status; Between-tree competition had a higher impact on tree diameter growth than on height growth; Growth predicted by the constructed models were calibrated using NFI11 data to ensure generally applicable growth predictions level in whole country.
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The aim of this study was to develop individual-tree diameter and height growth models for Scots pine, Norway spruce, and pubescent birch growing in drained peatlands in Finland. Trees growing in peatland sites have growth patterns that deviate from that of trees growing in mineral soil sites. Five-year growth was explained by tree diameter, different tree and stand level competition measures, management operations and site characteristics. The drainage status of the site was influencing growth directly or in interaction with other variables. Site quality had a direct impact but was also commonly related to current site drainage status (need for ditch maintenance). Recent thinning increased growth of all species and former PK fertilization increased growth of pine and birch. Temperature sum was a significant predictor in all models and altitude for spruce and birch. The data were a subsample of the 7th National Forest Inventory (NFI) sample plots representing northern and southern Finland and followed by repeated measurements for 15–20 yrs. Growth levels predicted by the models were calibrated using NFI11 data to remove bias originating from the sample of the modelling data. The mixed linear models technique was used in model estimation. The models will be incorporated into the MOTTI stand simulator to replace the current peatlands growth models.

  • Repola, Natural Resources Institute Finland (Luke), Natural resources, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: jaakko.repola@luke.fi (email)
  • Hökkä, Natural Resources Institute Finland (Luke), Natural resources, Paavo Havaksen tie 3, FI-90014 University OF Oulu, Finland E-mail: hannu.hokka@luke.fi
  • Salminen, Natural Resources Institute Finland (Luke), Natural resources, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: hannu.salminen@luke.fi
article id 10040, category Research article
Bengt Andersson Gull, Torgny Persson, Aleksey Fedorkov, Tim J. Mullin. (2018). Longitudinal differences in Scots pine shoot elongation. Silva Fennica vol. 52 no. 5 article id 10040. https://doi.org/10.14214/sf.10040
Keywords: Pinus sylvestris; climate change; adaptation; shoot phenology; heritability of phenological traits; growth onset; growth cessation
Highlights: More northerly Scots pine origins exhibit earlier onset and cessation of shoot growth; Continental origins show more northern phenological behaviour; Heat accumulation requirements for onset are not fixed and may be lower when accumulating slower; Scots pine may suffer from spring frost due to earlier growth onset in a warming climate; Phenological traits show potential to adapt to new climate conditions by breeding.
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Phenology can have a profound effect on growth and climatic adaptability of long-lived, northern tree species such as Scots pine (Pinus sylvestris L.), where the onset of growth in the spring is triggered mainly by accumulated heat, while cessation of growth is related to the joint effect of photoperiod and temperature. In this study, the objectives were: (1) to compare shoot phenology of genetic material from Scandinavia (maritime climate origin) and northern Russia (continental climate origin) sources, under field conditions in both Scandinavia and Russia (maritime and continental growth conditions); and (2) to estimate the heritabilities of phenological parameters. The material used was part of a larger provenance test series involving Scots pine populations and open-pollinated plus-tree families from Russia, Sweden and Finland. Terminal shoot elongation was measured on multiple occasions during the seventh growing season from seed at a trial near Bäcksjön (Sweden) and Syktyvkar (northern Russia). We calculated the regression of relative shoot elongation over accumulated heat sum above +5 °C using an exponential expression. Seedlings of Swedish and Russian provenance had similar heat-sum requirements for growth onset and cessation in both trials. More northern provenances started onset and cessation at a lower temperature sum, but heat accumulation requirements for onset were not fixed. Scots pine may suffer from spring frost due to earlier growth onset in a warming climate. Variation and heritability of phenological traits show potential to adapt Scots pine to new climate conditions by breeding.

  • Andersson Gull, The Swedish Forestry Research Institute (Skogforsk), Box 3, SE-918 21 Sävar, Sweden ORCID https://orcid.org/0000-0003-3556-3172 E-mail: bengt.anderssongull@skogforsk.se
  • Persson, The Swedish Forestry Research Institute (Skogforsk), Box 3, SE-918 21 Sävar, Sweden E-mail: torgny.persson@skogforsk.se
  • Fedorkov, The Institute of Biology of Komi Scientific Centre of the Ural Branch of the Russian Academy of Sciences (IB Komi SC UB RAS), Kommunisticheskaya St., 28, Syktyvkar, 167982, Russia ORCID https://orcid.org/0000-0001-7800-7534 E-mail: fedorkov@ib.komisc.ru
  • Mullin, The Swedish Forestry Research Institute (Skogforsk), Box 3, SE-918 21 Sävar, Sweden ORCID https://orcid.org/0000-0003-4924-1836 E-mail: tim.mullin@skogforsk.se (email)
article id 10036, category Research article
Karin Hjelm, Lars Rytter. (2018). The demand of hybrid aspen (Populus tremula × P. tremuloides) on site conditions for a successful establishment on forest land. Silva Fennica vol. 52 no. 5 article id 10036. https://doi.org/10.14214/sf.10036
Keywords: regeneration; site preparation; clone effects; poplars; soil acidity
Highlights: Low pH (below 3.5) reduced growth, but not survival, in a greenhouse study; Site preparation methods did not affect survival in field, but differences were found for growth; Mounding had generally the best effect on growth; Clonal differences were found that could be useful for improving establishment and growth.
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Hybrid aspen (Populus tremula L. × P. tremuloides Michx.) is a deciduous tree species suitable for producing large amounts of renewable biomass during short rotations. Its potential under North European conditions could be largely extended if not only agricultural land but also forest land was used for cultivation. Unfortunately, the knowledge of appropriate forest site conditions and effects of site preparation methods on hybrid aspen establishment is limited. In this paper, two studies that explore these questions are presented. In the first study, the sensitivity to acid soils was tested under greenhouse conditions in two type of soils: a) peat soil limed to certain pH levels (3.4–5.7) and b) collected forest soils where pH varied from 3.9 to 5.3. The lowest pH level resulted in reduced growth, elsewhere no significant differences were found. The second study was applied in the field and investigated the effect of four site preparation methods on survival and growth. The methods were: 1) control with no site preparation, 2) patch scarification, 3) mounding and 4) soil inversion. While no differences were found for survival, mounding was generally the method with the highest growth and patch scarification was least successful. The result was probably an effect of good soil aeration and less competition from vegetation after mounding. The field study also revealed clonal differences in growth performance, which stresses the importance of clone selection prior to planting. The results of these studies indicate that hybrid aspen is less sensitive to variation in pH and site preparation methods compared with other poplar species, as have been found in similar studies.

  • Hjelm, Skogforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden E-mail: karin.hjelm@skogforsk.se (email)
  • Rytter, Skogforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden ORCID http://orcid.org/0000-0001-6183-4832 E-mail: lars.rytter@skogforsk.se
article id 10034, category Research article
Yan Ao, Peter M. Hirst, Guolei Li, Yahui Miao, Runzhe Zhang. (2018). Combined effects of provenance and slow-release fertilizer on nursery and field performance of yellowhorn seedlings. Silva Fennica vol. 52 no. 5 article id 10034. https://doi.org/10.14214/sf.10034
Keywords: nursery response; field performance; Xanthoceras sorbifolium; plant nutrition; short rotation plantation
Highlights: Combining slow-release fertilizer (SRF) and provenance in the nursery has large effects on most seedling characteristics in yellowhorn; Stem and root P contents in the nursery, and height at the end of the second growing season (T3) in the field were mainly affected by provenance; Higher rates of SRF tended to increase root N, stem and root P contents in the nursery, diameter, and biomass at T3; The combination of AQ provenance with 120–200 mg N seedling–1 SRF yielding better nursery and field performance was recommended.
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Yellowhorn (Xanthoceras sorbifolium Bunge) has been widely planted for biodiesel production in China, but has frequently shown poor field performance. Container-grown yellowhorn seedlings originating from three Chinese provenances, Wengniute Qi (WQ), Alukeerqin Qi (AQ), and Shanxian (SX), were fertilized with slow-release fertilizer (SRF) at 40, 80, 120, 160 or 200 mg N seedling–1. Tree growth, survival and nutrient content were measured after one year’s growth in a greenhouse followed by two years in a field site. Plants from AQ and SX tended to have higher stem and root P contents in the nursery. Higher rates of SRF increased root N, and stem and root P contents. After one year in the nursery, there were a number of interactions between provenance and SRF for plant growth responses and nutrient content in the nursery, however after two years of additional growth in the field, plants from the different provenances generally responded similarly to applied SRF in the nursery, with few interactions. Final plant height was approximately 10% lower in trees from provenance SX but was not affected by application of SRF. Conversely, final trunk diameter and stem and root biomass were unaffected by provenance but increased with higher rates of applied SRF. Our results indicate that application of SRF may be a useful tool to nutrient load yellowhorn in the nursery and facilitate transplanting performance in the field. Overall, optimal nursery and field performance of yellowhorn were observed in provenance AQ at 120–200 mg N seedling–1 SRF. We suggest that growers consider a wider range of yellowhorn provenances and SRF rates (above 200 mg N seedling–1) to yield even better growth response.

  • Ao, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing Forestry University, 35 East Qinghua Road, Haidian District, Beijing 100083, China E-mail: aoyan316@163.com (email)
  • Hirst, Department of Horticulture and Landscape Architecture, Purdue University, West Lafayette, IN 47907, USA E-mail: hirst@purdue.edu
  • Li, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing Forestry University, 35 East Qinghua Road, Haidian District, Beijing 100083, China E-mail: glli226@163.com
  • Miao, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing Forestry University, 35 East Qinghua Road, Haidian District, Beijing 100083, China E-mail: 372902610@qq.com
  • Zhang, College of Forestry, Beijing Forestry University, 35 East Qinghua Road, Haidian District, Beijing 100083, China E-mail: 793755837@qq.com
article id 10001, category Research article
Karoliina Hämäläinen, Teemu Tahvanainen, Kaisa Junninen. (2018). Characteristics of boreal and hemiboreal herb-rich forests as habitats for polypore fungi. Silva Fennica vol. 52 no. 5 article id 10001. https://doi.org/10.14214/sf.10001
Keywords: biodiversity; species richness; coarse woody debris; threatened species; wood-decaying fungi
Highlights: Polypore species richness and diversity were affected positively by dead-wood diversity, and negatively by increasing latitude; Red-listed species responded only to the abundance of large-diameter dead wood; Main factor determining composition of polypore assemblages was host-tree species; High proportion of deciduous dead-wood in herb-rich forests provides complementary effect on polypore assemblages in boreal forest landscapes.
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Herb-rich forests are often considered biodiversity hotspots in the boreal zone but their fungal assemblages, particularly those of wood-decaying fungi, remain poorly known. We studied herb-rich forests as habitats for polypores, a distinct group of wood-decaying fungi, and assessed the importance of tree- and stand-scale variables for polypore species richness, abundance, and diversity, including red-listed species. The data include 71 herb-rich forest stands in Finland and 4797 dead wood items, on which we made 2832 observations of 101 polypore species. Dead-wood diversity was the most important variable explaining polypore species richness and diversity, whereas increasing latitude had a negative effect. Red-listed species showed a positive response to the abundance of large-diameter dead wood, which, especially birch, supported also high general abundance of polypores. The composition of polypore assemblages reflected their host-tree species. The red-listed species did not show explicit patterns in the ordination space. Compared to old-growth spruce forests, herb-rich forests seem to host lower polypore species richness and less red-listed species. However, because of high proportion of deciduous trees in the dead wood profile, herb-rich forests have a clear complementary effect on polypore assemblages in boreal forest landscapes.

  • Hämäläinen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: karoham@uef.fi (email)
  • Tahvanainen, Department of Environmental and Biological Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: teemu.tahvanainen@uef.fi
  • Junninen, Metsähallitus Parks & Wildlife Finland, c/o UEF/Borealis, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: kaisa.junninen@metsa.fi
article id 10000, category Research article
Hamed Yousefzadeh, Rasta Rajaei, Anna Jasińska, Łukasz Walas, Yann Fragnière, Gregor Kozlowski. (2018). Genetic diversity and differentiation of the riparian relict tree Pterocarya fraxinifolia (Juglandaceae) along altitudinal gradients in the Hyrcanian forest (Iran). Silva Fennica vol. 52 no. 5 article id 10000. https://doi.org/10.14214/sf.10000
Keywords: plant variation; Caucasian wingnut; population genetics; refugium; riparian forest; river system
Highlights: The Caucasian wingnut (Pterocarya fraxinifolia Spach) is an emblematic and relict riparian tree with limited distribution in Hyrcanian forest which investigating its genetic population structure and diversity along altitudinal gradients, and migration patterns are novel; We concluded that rivers are the main seed dispersal vector among P. fraxinifolia populations and there was no trend from upstream to downstream; The high level of gene flow and uniform genetic diversity along each river system suggest the “classical” metapopulation structure of the species.
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Riparian trees, especially relict trees, are attractive and important for research to understand both past and recent biogeographical and evolutionary processes. Our work is the first study to elucidate the genetic diversity and spatial genetic structure of the canopy-dominating riparian Pterocarya fraxinifolia (Juglandaceae) along two altitudinal gradients in different river systems of the Hyrcanian forest, which is one of the most important refugium of relict trees in Western Eurasia. Altitudinal gradients were chosen along two river systems at 100, 400 and 900 m a.s.l. Leaf samples were collected from 116 trees, and the genetic diversity was evaluated with eight SSR markers. Overall, 39 alleles were identified for all of the populations studied. The observed heterozygosity (Ho) varied from 0.79 to 0.87 (with a mean of 0.83). The results of the AMOVA analysis indicated that the variation within populations was 88%, whereas the variation among populations was 12% for all of the gradients. A structure analysis indicated that 93% of the trees were grouped in the same gradient. The genetic distance based on Fst confirmed the structure result and indicated a high rate of gene flow among the investigated populations. Based on high gene flow (low differentiation of the population along the same river) and the clearly distinct genetic structure of the investigated gradients, it can be concluded that rivers are the main seed dispersal vector among P. fraxinifolia populations. The genetic diversity of P. fraxinifolia did not show any trend from upstream to downstream. The high level of gene flow and uniform genetic diversity along each river suggest the “classical” metapopulation structure of the species.

  • Yousefzadeh, Department of Environmental Science, Faculty of Natural Resources and Marine Sciences, Tarbiat Modares University, Noor, Mazandaran, Iran E-mail: h.yousefzadeh@modares.ac.ir (email)
  • Rajaei, Department of Forestry, Faculty of Natural Resources and Marine Sciences, Tarbiat Modares University, Noor, Mazandaran, Iran E-mail: r.rajaei@modares.ac.ir
  • Jasińska, Laboratory of Systematics and Geography, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, PL-62-035 Kornik, Poland E-mail: jasiak9@wp.pl
  • Walas, Laboratory of Systematics and Geography, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, PL-62-035 Kornik, Poland E-mail: lukaswalas@wp.pl
  • Fragnière, Department of Biology and Botanic Garden, University of Fribourg, Chemin du Musée 10, CH-1700 Fribourg, Switzerland E-mail: yann.fragniere@unifr.ch
  • Kozlowski, Department of Biology and Botanic Garden, University of Fribourg, Chemin du Musée 10, CH-1700 Fribourg, Switzerland; Natural History Museum Fribourg, Chemin du Musée 6, CH-1700 Fribourg, Switzerland E-mail: gregor.kozlowski@unifr.ch
article id 9985, category Research article
Antonín Martiník, Robert Knott, Jan Krejza, Jakub Černý. (2018). Biomass production of Betula pendula stands regenerated in the region of allochthonous Picea abies dieback. Silva Fennica vol. 52 no. 5 article id 9985. https://doi.org/10.14214/sf.9985
Keywords: silver birch; stem biomass; branch biomass; rotation period
Highlights: Biomass equations for individual components of above-ground wood biomass estimation are presented for stands at the age of 4, 8, 17 and 22 years; Peak of the mean annual increment was found at the age from 15 to 20 years and reached over 5.0 t ha–1 y–1 of dry biomass; The share of the stem to the total biomass increased with stand age.
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The paper deals with production of above-ground biomass of silver birch (Betula pendula Roth) stands in the Czech Republic. One-year biomass dynamics was studied within chronosequence of birch stands at the age of 4–5, 8–9, 17–18 and 22–23 years. With the exception of the youngest stand, which was established by seeding, all experimental birch stands were regenerated naturally after the allochthonous spruce stands. Above-ground biomass (AB) was calculated from plot inventory data and biomass equations were parameterized from destructive sampling of biomass component of sampled trees. Results reveal that the peak of the mean annual increment (MAIABtotal) of birch stands can be expected at the age from 15 to 20 years. Additionally, the stand age, the value of basal area (BA) should be considered as a predictor of stand productivity. If the value of BA varied from 25 to 35 m2 ha–1, the MAI of the birch stands reached the range from 5.0 to 6.5 t of dry biomass per ha y–1 at the age ranging between 15 and 25 years. The stem/branch proportion increased with stand age, the stem relative proportion ranging from 75 to 90% of total above-ground biomass. According to the results of this study, birch stand biomass production and utilization is one of the approaches in terms of forest recovery management in large disturbed areas. Although, no silvicultural treatments were occurred in all analysed stands, the pre-commercial thinning method could increase stand productivity and stability as well.

  • Martiník, Department of Silviculture, Faculty of Forestry and Wood Technology, Mendel University in Brno, 613 00 Brno, Czech Republic ORCID https://orcid.org/0000-0002-5906-8830 E-mail: martinik@mendelu.cz (email)
  • Knott, Department of Silviculture, Faculty of Forestry and Wood Technology, Mendel University in Brno, 613 00 Brno, Czech Republic E-mail: robert.knott@mendelu.cz
  • Krejza, Global Change Research Institute CAS, v.v.i., Bělidla 4a, 603 00 Brno, Czech Republic E-mail: krejza.j@czechglobe.cz
  • Černý, The Forestry and Game Management Research Institute, Research Station at Opočno, Na Olivě 550, 517 73 Opočno, Czech Republic E-mail: cerny@vulhmop.cz
article id 6993, category Research article
Lars Fridh, Lars Eliasson, Dan Bergström. (2018). Precision and accuracy in moisture content determination of wood fuel chips using a handheld electric capacitance moisture meter. Silva Fennica vol. 52 no. 5 article id 6993. https://doi.org/10.14214/sf.6993
Keywords: heating value; biofuels; payment
Highlights: The studied capacitance meter can provide accurate estimates of mean moisture content for chips with M < 50% if a large sample is taken; It should be possible to use the capacitance meter to measure moisture content even for calculating payments depending of the needed accuracy; However a calibration function for each assortment is needed.
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According to the Swedish Timber Measurement Act, measurements affecting payments for wood fuels to landowners must be accurate and precise. In this regard, moisture content is an important quality parameter for wood chips which influences the net calorific value as received and thus the economic value. As standard practice moisture content is determined with the oven-drying method, which is cumbersome to use for deliveries to facilities without drying-ovens, which in turn necessitates that samples are taken elsewhere for measurement. An alternative solution is to use a portable moisture meter. Our aim was to evaluate the precision of a handheld capacitance moisture meter. Accuracy and precision of a capacitance meter was determined in the lab and a calibration function was made. Thereafter, the calibrated moisture meter was compared with the standard method for moisture content determination of truckloads of chips. The capacitance meter showed a moderate accuracy by underestimating moisture content by 6.0 percentage points (pp), compared to the reference method, at a precision of ±3.8 pp (CI 95%). For chips with M > 50%, both accuracy and precision decreased. Calibration increased the accuracy in the follow up study by 3 pp for chips with M < 50% but could not be made for wetter chips. The oven-drying method and the capacitance meter can provide equally accurate estimates of mean moisture content for chips with M < 50% if a larger sample is taken with the latter. It should be possible to use capacitance meters to measure moisture content even when used to calculate payments depending of the needed accuracy. A calibration function for each assortment is needed.

  • Fridh, Skogforsk, The Forestry Research Institute of Sweden, Uppsala Science Park, 751 83 Uppsala, Sweden; Skogsägarna Mellanskog, Uppsala Science Park, Box 127, 751 04 Uppsala, Sweden ORCID http://orcid.org/0000-0002-4721-1193 E-mail: lars.fridh@mellanskog.se
  • Eliasson, Skogforsk, The Forestry Research Institute of Sweden, Uppsala Science Park, 751 83 Uppsala, Sweden ORCID http://orcid.org/0000-0002-2038-9864 E-mail: lars.eliasson@skogforsk.se (email)
  • Bergström, Swedish University of Agricultural Sciences, Department of Forest Biomaterials and Technology, S-901 83 Umeå, Sweden E-mail: dan.bergstrom@slu.se
article id 10013, category Research article
Hardo Becker, Jürgen Aosaar, Mats Varik, Gunnar Morozov, Kristiina Aun, Ülo Mander, Kaido Soosaar, Veiko Uri. (2018). Annual net nitrogen mineralization and litter flux in well-drained downy birch, Norway spruce and Scots pine forest ecosystems. Silva Fennica vol. 52 no. 4 article id 10013. https://doi.org/10.14214/sf.10013
Keywords: Pinus sylvestris; Picea abies; Betula pubescens; drained peatland forests; ammonification; effect of tree species; swamp; nitrification; transitional bog
Highlights: The net nitrogen mineralization (NNM) flux in drained peat soils depends largely on the C/N ratio and tree species; The soil NNM process is affected by trees through organic litter input into soil; Pine stand in low-fertility drained transitional bog is dominated by net ammonification; Birch and spruce stands on the fertile drained peat soil with higher pH and N content are dominated by net nitrification.
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The main aim of the current study was to estimate the annual net nitrogen mineralization (NNM) flux in stands of different tree species growing on drained peatlands, as well as to clarify the effect of tree species, soil properties and litter on annual NNM dynamics. Three study sites were set up in May 2014: a downy birch (Betula pubescens Ehrh.) stand and a Norway spruce (Picea abies (L.) Karst.) stand in Oxalis full-drained swamp (ODS) and a Scots pine (Pinus sylvestris L.) stand in Myrtillus full-drained swamp (MDS). The NNM flux was estimated using the in situ method with incubated polyethylene bags. The highest value of NNM was found in stands that were growing on fertile ODS: 127.5 kg N ha–1 yr–1 and 87.7 kg N ha–1 yr–1, in the downy birch stand and in the Norway spruce stand, respectively. A significantly lower annual NNM flux (11.8 kg N ha–1 yr–1) occurred in the Scots pine stand growing in MDS. Nitrification was highest at fertile ODS sites and ammonification was the highest at the low fertility MDS site. For all study sites, positive correlation was found between soil temperature and NNM intensity. The difference in annual NNM between the downy birch stand and the Norway spruce stand growing on similar drained fertile peatlands was due to litter quality. The annual N input into the soil through leaf litter was the highest at the downy birch site where also the C/N ratio of litter was the lowest. The second highest N input into the soil was found in the spruce stand and the lowest in the pine stand.

  • Becker, Chair of Silviculture and Forest Ecology, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Friedrich Reinhold Kreutzwaldi 1, 51014 Tartu, Estonia E-mail: hardo.becker@emu.ee (email)
  • Aosaar, Chair of Silviculture and Forest Ecology, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Friedrich Reinhold Kreutzwaldi 1, 51014 Tartu, Estonia E-mail: jyrgen.aosaar@emu.ee
  • Varik, Chair of Silviculture and Forest Ecology, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Friedrich Reinhold Kreutzwaldi 1, 51014 Tartu, Estonia E-mail: mats.varik@emu.ee
  • Morozov, Chair of Silviculture and Forest Ecology, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Friedrich Reinhold Kreutzwaldi 1, 51014 Tartu, Estonia E-mail: gunnar.morozov@emu.ee
  • Aun, Chair of Silviculture and Forest Ecology, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Friedrich Reinhold Kreutzwaldi 1, 51014 Tartu, Estonia E-mail: kristiina.aun@emu.ee
  • Mander, Department of Geography, Institute of Ecology & Earth Sciences, University of Tartu, Ülikooli 18, 50090 Tartu, Estonia E-mail: ulo.mander@ut.ee
  • Soosaar, Department of Geography, Institute of Ecology & Earth Sciences, University of Tartu, Ülikooli 18, 50090 Tartu, Estonia E-mail: kaido.soosaar@ut.ee
  • Uri, Chair of Silviculture and Forest Ecology, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Friedrich Reinhold Kreutzwaldi 1, 51014 Tartu, Estonia E-mail: veiko.uri@emu.ee
article id 10002, category Research article
Bayasaa Tumenjargal, Futoshi Ishiguri, Haruna Aiso-Sanada, Yusuke Takahashi, Bayartsetseg Baasan, Ganbaatar Chultem, Jyunichi Ohshima, Shinso Yokota. (2018). Geographic variations of wood properties of Larix sibirica naturally grown in Mongolia. Silva Fennica vol. 52 no. 4 article id 10002. https://doi.org/10.14214/sf.10002
Keywords: natural forests; basic density; dynamic Young’s modulus; nondestructive testing; stress-wave velocity
Highlights: Significant differences (p  < 0.01) among five stands were found in tree height, stress-wave velocity of stems, dynamic Young’s modulus of stems and logs, annual ring width, and latewood percentage, suggesting that there was geographic variation of mechanical properties of wood in Larix sibirica (Münchh.) Ledeb. grown in Mongolia; Dynamic Young’s modulus of logs in L. sibirica trees can be predicted by stress-wave velocity of stems; Stem diameter of L. sibirica naturally grown in Mongolia is closely related to radial growth at initial stage of growth, especially within the first twenty years.
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Geographic variations in growth, stress-wave velocity of stem, dynamic Young’s modulus of stems and logs, annual ring width, latewood percentage and basic density were investigated for Larix sibirica (Münchh.) Ledeb. naturally grown in Mongolia. A total of 250 trees with 20 to 30 cm in stem diameter at a height of 1.3 m above ground level were selected from each natural stand in five different provenances in Mongolia. In addition, five trees in each stand were cut for measuring dynamic Young’s modulus of stems and logs, annual ring width, latewood percentage and basic density. Mean values of stress-wave velocity of stems in each stand ranged from 2.92 to 3.41 km s–1, and the mean value of five stands was 3.23 km s–1. Mean values of dynamic Young’s modulus of logs in each stand ranged from 5.17 to 9.72 GPa. A significant correlation (r = 0.798, p < 0.01) was found between stress-wave velocity of stems and dynamic Young’s modulus of logs. Among the five stands, the highest and the lowest values of average annual ring number were 193 and 44, respectively. Mean values of basic density in five trees within each stand were examined and ranged from 0.52 to 0.56 g cm–3. Significant differences among five stands were found in tree height, stress-wave velocity of stem, dynamic Young’s modulus of stems and logs, annual ring width and latewood percentage, suggesting that L. sibirica trees naturally grown in Mongolia have geographic variations in mechanical properties of wood.

  • Tumenjargal, Faculty of Agriculture, Utsunomiya University, Utsunomiya, Tochigi 321-8505, Japan; United Graduate School of Agricultural Science, Tokyo University of Agriculture and Technology, Fuchu, Tokyo 183-8509, Japan; Research and Training Institute of Forestry and Wood Industry, Mongolian University Science and Technology, Ulaanbaatar 14191, Mongolia E-mail: t_bayasaa88@yahoo.com
  • Ishiguri, Faculty of Agriculture, Utsunomiya University, Utsunomiya, Tochigi 321-8505, Japan E-mail: ishiguri@cc.utsunomiya-u.ac.jp (email)
  • Aiso-Sanada, Forestry and Forest Products Research Institute, Tsukuba, Ibaraki 305-8687, Japan E-mail: haiso@ffpri.affrc.go.jp
  • Takahashi, Faculty of Agriculture, Utsunomiya University, Utsunomiya, Tochigi 321-8505, Japan E-mail: zo.r.by0814@gmail.com
  • Baasan, Research and Training Institute of Forestry and Wood Industry, Mongolian University Science and Technology, Ulaanbaatar 14191, Mongolia E-mail: bayartsetseg@must.edu.mn
  • Chultem, Research and Training Institute of Forestry and Wood Industry, Mongolian University Science and Technology, Ulaanbaatar 14191, Mongolia E-mail: ganbaatar_ch@must.edu.mn
  • Ohshima, Faculty of Agriculture, Utsunomiya University, Utsunomiya, Tochigi 321-8505, Japan E-mail: joshima@cc.utsunomiya-u.ac.jp
  • Yokota, Faculty of Agriculture, Utsunomiya University, Utsunomiya, Tochigi 321-8505, Japan E-mail: yokotas@cc.utsunomiya-u.ac.jp
article id 9996, category Research article
Mulualem Tigabu, Mostafa Farhadi, Lars-Göran Stener, Per C. Odén. (2018). Visible + Near Infrared Spectroscopy as taxonomic tool for identifying birch species. Silva Fennica vol. 52 no. 4 article id 9996. https://doi.org/10.14214/sf.9996
Keywords: Betula; silver birch; downy birch; seeds; NIRS; OPLS; multivariate modelling
Highlights: Multivariate modelling of visible + near infrared (NIR) reflectance spectra of single seeds distinguished Betula pubescens and B. pendula with 100% and 99% accuracy, respectively; The results demonstrate the feasibility of NIR spectroscopy as taxonomic tool for classification of species that have morphological resemblance.
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The genus Betula L. is composed of several species, which are difficult to distinguish in the field on the basis of morphological traits. The aim of this study was to evaluate the taxonomic importance of using visible + near infrared (Vis + NIR) spectra of single seeds for differentiating Betula pendula Roth and Betula pubescens Ehrh. Seeds from several families (controlled crossings of known parent trees) of each species were used and Vis + NIR reflectance spectra were obtained from single seeds. Multivariate discriminant models were developed by Orthogonal Projections to Latent Structures – Discriminant Analysis (OPLS-DA). The OPLS-DA model fitted on Vis + NIR spectra recognized B. pubescens with 100% classification accuracy while the prediction accuracy of class membership for B. pendula was 99%. However, the discriminant models fitted on NIR spectra alone resulted in 100% classification accuracies for both species. Absorption bands accounted for distinguishing between birch species were attributed to differences in color and chemical composition, presumably polysaccharides, proteins and fatty acids, of the seeds. In conclusion, the results demonstrate the feasibility of NIR spectroscopy as taxonomic tool for classification of species that have morphological resemblance.

  • Tigabu, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, Box 49, SE-230 52 Alnarp, Sweden E-mail: mulualem.tigabu@slu.se (email)
  • Farhadi, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, Box 49, SE-230 52 Alnarp, Sweden E-mail: mostafa.farhadi@gmail.com
  • Stener, The Forestry Research Institute of Sweden, Ekebo 2250, SE-268 90 Svalöv, Sweden E-mail: lars-goran.stener@skogforsk.se
  • Odén, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, Box 49, SE-230 52 Alnarp, Sweden E-mail: per.christer.oden@slu.se
article id 9972, category Research article
Jukka Malinen, Harri Kilpeläinen, Erkki Verkasalo. (2018). Validating the predicted saw log and pulpwood proportions and gross value of Scots pine and Norway spruce harvest at stand level by Most Similar Neighbour analyses and a stem quality database. Silva Fennica vol. 52 no. 4 article id 9972. https://doi.org/10.14214/sf.9972
Keywords: stem quality; bucking; non-parametric prediction; database; assortment recovery; value recovery
Highlights: Non-parametric prediction together with external stem quality database provides predictions usable for pre-harvest assessment at a stand level; The prediction of Norway spruce assortment recovery and value proved to be more accurate than the predictions for Scots pine; RMSE and bias of unit prices were 3.50 € m–3 and 0.58 € m–3 for pine and 2.60 € m–3 and 0.35 € m–3 for spruce.
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Detailed pre-harvest information about the volumes and properties of growing stocks is needed for increased precision in wood procurement planning for just-in-time wood deliveries by cut-to-length (CTL) harvesters. In the study, the non-parametric Most Similar Neighbour (MSN) methodology was evaluated for predicting external quality of Scots pine and Norway spruce, expressed as stem sections fulfilling the saw log dimension and quality requirements of Finnish forest industry, as they affect the recovery of timber assortments and the value of a pre-harvest stand. Effects of external tree quality were evaluated using saw log recovery and saw log reduction caused by stem defects, as well as total timber value (€) and average unit value (€ m–3) in a stand. Root mean square error (RMSE) of saw log recovery and reduction were 9.12 percentile points (pp) for Scots pine and 6.38 pp for Norway spruce stands. In the unit value considerations, the predictions compared with measurements resulted in the RMSE of 3.50 € m–3 and the bias of 0.58 € m–3 in Scots pine stands and 2.60 € m–3, and 0.35 € m–3 in Norway spruce stands, respectively. The presented MSN based approach together with the utilization of the external stem quality database included in the ARVO software could provide dimension and external quality predictions usable for pre-harvest assessment of timber stock at a stand level. This prediction methodology is usable especially in analyses where timber assortment recoveries, values and unit prices are compared when different bucking objectives are used.

  • Malinen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: jukka.malinen@uef.fi (email)
  • Kilpeläinen, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: harri.kilpelainen@luke.fi
  • Verkasalo, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: erkki.verkasalo@luke.fi
article id 7763, category Research article
Sergei Senko, Mikko Kurttila, Timo Karjalainen. (2018). Prospects for Nordic intensive forest management solutions in the Republic of Karelia. Silva Fennica vol. 52 no. 4 article id 7763. https://doi.org/10.14214/sf.7763
Keywords: forest management; thinning; forestry; Russia; SWOT
Highlights: SWOT (strengths, weaknesses, opportunities and threats) and multi-criteria decision support analysis were combined to examine the potential for Nordic intensive forest management solutions (NIFMS) in Karelia, Russia; NIFMS looks promising for Karelian forestry; Improving quality-and-value of timber and sustained yield are the highly prioritized strengths; Unprepared forestry regulations are the main threat that needs to be taken into account.
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In this study, the prospects for future forest management in Republic of Karelia, Russia were analyzed. Forestry has an important role in the economy of Karelia. However, productivity and profitability in the forestry sector are extremely low, forest stand structure and quality are weak, the commercial forest land of coniferous species has declined and the wood processing industry struggles with a deficit of raw materials. The situation is typical to many forest regions in Russia with extensive forest management cited as one reason for the current situation. In contrast, the Nordic countries have significant experience in intensive and sustainable forest management and the results have been to a large extent positive. The transfer of Nordic intensive forest management solutions (NIFMS) could improve forestry in Karelia. SWOT analysis, combined with the multi-criteria decision support (MCDS) method was used to identify local operational environments and to assign priorities. Major threats included unprepared regulations, poor road infrastructure, insecure investments, low forestry productivity, forest degradation, high investment costs and a negative attitude to intensive forestry. The main opportunities are high forest resource potential in Karelia, favorable authority development programs, proven Nordic expertise, wood-based energy development and availability of new technology. Results also showed that the main weaknesses that might influence the NIFMS in Karelia are slow return on investments, low market demand for energy wood, high costs associated with young forest thinnings, high demand for skilled specialists and a lack of investment in research and development.

  • Senko, University of Eastern Finland (UEF), School of Forest Sciences, Yliopistokatu 7, FI-80111 Joensuu, Finland E-mail: sergei.senko@uef.fi (email)
  • Kurttila, The Natural Resources Institute Finland (Luke), Bioeconomy and environment, Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: mikko.kurttila@luke.fi
  • Karjalainen, &#8224; Deceased E-mail: tjk@nn.fi
article id 10006, category Research article
Matti Maltamo, Tomi Karjalainen, Jaakko Repola, Jari Vauhkonen. (2018). Incorporating tree- and stand-level information on crown base height into multivariate forest management inventories based on airborne laser scanning. Silva Fennica vol. 52 no. 3 article id 10006. https://doi.org/10.14214/sf.10006
Keywords: forest inventory; LIDAR; alpha shape; crown height; nearest neighbor; mixed-effects model
Highlights: The most accurate tree-level alternative is to include crown base height (CBH) to nearest neighbour imputation; Also mixed-effects models can be applied to predict CBH using tree attributes and airborne laser scanning (ALS) metrics; CBH prediction can be included with an accuracy of 1–1.5 m to forest management inventory applications.
Abstract | Full text in HTML | Full text in PDF | Author Info

This study examines the alternatives to include crown base height (CBH) predictions in operational forest inventories based on airborne laser scanning (ALS) data. We studied 265 field sample plots in a strongly pine-dominated area in northeastern Finland. The CBH prediction alternatives used area-based metrics of sparse ALS data to produce this attribute by means of: 1) Tree-level imputation based on the k-nearest neighbor (k-nn) method and full field-measured tree lists including CBH observations as reference data; 2) Tree-level mixed-effects model (LME) prediction based on tree diameter (DBH) and height and ALS metrics as predictors of the models; 3) Plot-level prediction based on analyzing the computational geometry and topology of the ALS point clouds; and 4) Plot-level regression analysis using average CBH observations of the plots for model fitting. The results showed that all of the methods predicted CBH with an accuracy of 1–1.5 m. The plot-level regression model was the most accurate alternative, although alternatives producing tree-level information may be more interesting for inventories aiming at forest management planning. For this purpose, k-nn approach is promising and it only requires that field measurements of CBH is added to the tree lists used as reference data. Alternatively, the LME-approach produced good results especially in the case of dominant trees.

  • Maltamo, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: matti.maltamo@uef.fi (email)
  • Karjalainen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: tomimkarjalainen@gmail.com
  • Repola, Natural Resources Institute of Finland (Luke), Natural resources, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: jaakko.repola@luke.fi
  • Vauhkonen, Natural Resources Institute of Finland (Luke), Bioeconomy and environment, Yliopistokatu 6, 80100 Joensuu, Finland E-mail: jari.vauhkonen@luke.fi
article id 9987, category Research article
Monika Dering, Katarzyna Sękiewicz, Grzegorz Iszkuło, Aleksandra Chojnacka, Dominik Tomaszewski, Emilia Pers-Kamczyc, Piotr Karolewski. (2018). Spatial genetic structure and clonal structure of Prunus serotina during invasive spread. Silva Fennica vol. 52 no. 3 article id 9987. https://doi.org/10.14214/sf.9987
Keywords: genetic structure; invasiveness; black cherry; colonization; clonal structure
Highlights: The spread of Prunus serotina in invaded forests is facilitated by high propagule pressure; The seed shadow overlap prevents strong spatial genetic structure; During colonization of isolated site, a strong spatial genetic structure is produced due to founder effect; Overall clonality in P. serotina was low but may efficiently support seedling bank thus contributing to species invasiveness.
Abstract | Full text in HTML | Full text in PDF | Author Info

Prunus serotina Ehrh. (black cherry) is one of the most important invaders in the European forests, but existing studies have given limited insight into demo-genetic factors underpinning the process of species invasion. Fine-scale genetic structure (FSGS) may deliver important knowledge on genetics of invasion contributing to efficient management of the alien species. Using eight microsatellites we investigated FSGS, clonal structure and relatedness in four black cherry populations which represented different stages of the invasive spread into Scots pine forests. Three populations were in a continuous forest complex and represented the colonization (Z_1) and established stages (Z_2 and Z_3). To investigate how colonization ability of the species is modified by landscape features, we analyzed an isolated population at colonization stage located in limited forest patch located in an agricultural landscape (R). Populations from continuous forest showed low yet significant positive FSGS with Sp = 0.0068 in Z_1, 0.0054 in Z_2, and 0.0066 in Z_3, while in R spatial structure was the strongest (0.0145). Considerable relatedness noted in population R suggests a dominance of within-population mating and recruitments, low immigration rate and limited seed dispersal, all of which led to the observed strong FSGS. Also, we presume that a founder effect likely involved during colonization of isolated forest patch R led to strong FSGS. In contrary, the seed shadow overlap in the populations from continuous forest prevented strong FSGS and facilitated colonization. Despite of low level of clonality, we argue that it may efficiently support black cherry seedling bank contributing to species invasiveness.

  • Dering, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland E-mail: mdering@man.poznan.pl (email)
  • Sękiewicz, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland E-mail: ksekiewicz@man.poznan.pl
  • Iszkuło, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland; Faculty of Biological Sciences, University of Zielona Góra, Prof. Z. Szafrana 1, 65-516 Zielona Góra, Poland E-mail: iszkulo@man.poznan.pl
  • Chojnacka, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland E-mail: jagoda900@gmail.com
  • Tomaszewski, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland E-mail: dominito@man.poznan.pl
  • Pers-Kamczyc, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland E-mail: epk@man.poznan.pl
  • Karolewski, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland E-mail: pkarolew@man.poznan.pl
article id 9983, category Research article
Mika Rekola, Jaakko Nippala, Päivi Tynjälä, Anne Virtanen. (2018). Modelling competences and anticipating the future competence needs in the forest sector. Silva Fennica vol. 52 no. 3 article id 9983. https://doi.org/10.14214/sf.9983
Keywords: forest sector; competence modelling; human resources; work life; future needs
Highlights: The most frequently used practices of modelling competences in Finnish forest sector organisations were superior-subordinate review discussions and quantitative surveys; Competence modelling was used for several human resources functions but surprisingly not for hiring and compensation; The experts interviewed underlined the need for generic competences in the future, especially they highlighted the importance of information processing and personal self-management skills.
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This explorative study examined practices of competence modelling in the forest sector organisations and how organisations anticipate changes in competence needs in the future. Semi-structured in-depth interviews (n=10) were conducted amongst forest sector experts in Finland and data was analysed by thematic analysis. The findings showed that the practices of modelling competences were diverse, most frequently used ones being superior-subordinate review discussions and quantitative competence surveys. In addition to these formal systems, informal modelling, especially on the team level and in smaller companies was also frequent. Organisations used competence modelling for several human resources functions, such as appraisal, motivation and promotion of employees. Surprisingly hiring and compensation functions were not mentioned. Perceptions related to competence modelling were generally speaking positive. The most important challenges were the lack of further actions and sometimes the extraordinary burden to the employees. When anticipating the future, the experts interviewed mentioned several commonly recognised trends, e.g., development of information technology, fragmentation of working life and structural changes in labour markets. All these require more generic competences related to information processing and personal self-management, especially respondents highlighted the importance of self-awareness skills. It is concluded that several useful practices for competence modelling already exist and that present study provides a basis for further quantitative further study.

  • Rekola, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: mika.rekola@helsinki.fi (email)
  • Nippala, Suomen Partiolaiset – Finlands Scouter ry, Töölönkatu 55, FI-00250 Helsinki, Finland E-mail: jaakko.nippala@partio.fi
  • Tynjälä, Finnish Institute for Educational Research, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: paivi.a.tynjala@jyu.fi
  • Virtanen, Finnish Institute for Educational Research, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: anne.virtanen@jyu.fi
article id 9980, category Research article
Eino Levkoev, Lauri Mehtätalo, Katri Luostarinen, Pertti Pulkkinen, Anatoly Zhigunov, Heli Peltola. (2018). Development of height growth and frost hardiness for one-year-old Norway spruce seedlings in greenhouse conditions in response to elevated temperature and atmospheric CO2 concentration. Silva Fennica vol. 52 no. 3 article id 9980. https://doi.org/10.14214/sf.9980
Keywords: climate change; Picea abies; needles; tree growth; seedlings; cold acclimation
Highlights: Elevated temperature resulted in increased height growth, delayed onset and shortened duration of autumn frost hardiness development in Norway spruce seedlings; Elevated temperature increased variation between genotypes in height growth and frost hardiness development; Elevated atmospheric CO2 concentration had no effect on the development of height or autumn frost hardiness in Norway spruce seedlings.
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The mean temperature during the potential growing season (April–September) may increase by 1 °C by 2030, and by 4 °C, or even more, by 2100, accompanied by an increase in atmospheric CO2 concentrations of 536–807 ppm, compared to the current climate of 1981–2010, in which atmospheric CO2 is at about 350 ppm. This may affect both the growth and frost hardiness of boreal trees. In this work, we studied the responses of height and autumn frost hardiness development in 22 half-sib genotypes of one-year-old Norway spruce (Picea abies (L.) Karst.) seedlings to elevated temperatures and atmospheric CO2 concentration under greenhouse conditions. The three climate treatments used were: T+1 °C above ambient and ambient CO2; T+4 °C above ambient and ambient CO2; and T+4 °C above ambient and elevated CO2 (700 ppm). The height growth rate and final height were both higher under T+4 °C compared to T+1 °C. Temperature increase also delayed the onset, and shortened the duration, of autumn frost hardiness development. Elevated CO2 did not affect the development of height or frost hardiness, when compared to the results without CO2 elevation under the same temperature treatment. Higher temperatures resulted in greater variation in height and frost hardiness development among genotypes. Three genotypes with different genetic backgrounds showed superior height growth, regardless of climate treatment; however, none showed a superior development of autumn frost hardiness. In future studies, clonal or full-sib genetic material should be used to study the details of autumn frost hardiness development among different genotypes.

  • Levkoev, University of Eastern Finland, Faculty of Science and Forestry, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: eino.levkoev@uef.fi (email)
  • Mehtätalo, University of Eastern Finland, Faculty of Science and Forestry, School of Computing, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: lauri.mehtatalo@uef.fi
  • Luostarinen, University of Eastern Finland, Faculty of Science and Forestry, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: katri.luostarinen@uef.fi
  • Pulkkinen,  Natural Resources Institute Finland (Luke), Production systems, Haapastensyrjä Breeding Station, FI-16200 Läyliäinen, Finland E-mail: pertti.pulkkinen@luke.fi
  • Zhigunov, Saint-Petersburg State Forest Technical University, Forestry Faculty, RU-194021, Institutskiy per. 5, Saint-Petersburg, Russia E-mail: a.zhigunov@bk.ru
  • Peltola, University of Eastern Finland, Faculty of Science and Forestry, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: heli.peltola@uef.fi
article id 9948, category Research article
Juha Heiskanen, Ville Hallikainen, Jori Uusitalo, Hannu Ilvesniemi. (2018). Co-variation relations of physical soil properties and site characteristics of Finnish upland forests. Silva Fennica vol. 52 no. 3 article id 9948. https://doi.org/10.14214/sf.9948
Keywords: site quality; vegetation; bulk density; trafficability; particle size; humus layer; water retention
Highlights: Atmospheric temperature sum is related to site index H100 as a covariate;Soil pH and water retention at field capacity (FC) are also closely related to H100;Fine fraction is related to water retention at FC, soil layer and site type;Fine fraction co-varies also with temperature sum, H100 and slope.
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Physical soil properties have a marked influence on the quality of forest sites and on the preconditions for forest growth and management. In this study, water retention characteristics (WRC) and related physical soil properties in addition to vegetation coverage and tree stand data were studied at upland forest sites in Finland. Fixed and mixed models between soil and site characteristics were formed to estimate physical and hydrologic soil characteristics and the site quality with indirect co-varying variables. In the present data, the site quality index (H100) shows a high coefficient of determination in respect to the temperature sum. It is also related to soil fine fraction content, topsoil pH and water retention at field capacity. The thickness of the humus layer is predictable from the pH and cover of xeric and mesic plant species. The soil fine fraction content (clay + silt) is closely related to water retention at field capacity, the soil layer and site type, and without WRC to the temperature sum and site index and type, as well as the slope angle. The soil bulk density is related to organic matter, depth (layer) or alternatively to organic matter, slope and field estimated textural class (fine, medium, coarse). Water retention characteristics were found to be best determinable by the fine fraction content, depth and bulk density. Water content and air-filled porosity at field capacity are closely related to the fine fraction. This study provides novel models for further investigations that aim at improved prediction models for forest growth, hydrology and trafficability.

  • Heiskanen, Natural Resources Institute Finland (Luke), Soil ecosystems, Neulaniementie 5, FI-70100 Kuopio, Finland E-mail: juha.heiskanen@luke.fi (email)
  • Hallikainen, Natural Resources Institute Finland (Luke), Applied statistical methods, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: ville.hallikainen@luke.fi
  • Uusitalo, Natural Resources Institute Finland (Luke), Forest technology and logistics, Korkeakoulunkatu 7, FI-33720 Tampere, Finland E-mail: jori.uusitalo@luke.fi
  • Ilvesniemi, Natural Resources Institute Finland (Luke), Biorefinery and bioproducts, Tietotie 2, FI-02150 Espoo, Finland E-mail: hannu.ilvesniemi@luke.fi
article id 9947, category Research article
Eric R. Labelle, Linus Huß. (2018). Creation of value through a harvester on-board bucking optimization system operated in a spruce stand. Silva Fennica vol. 52 no. 3 article id 9947. https://doi.org/10.14214/sf.9947
Keywords: buck-to-value; processing; single-grip-harvester; product recovery; harvesting productivity
Highlights: Use of harvester on-board computer bucking optimization remains highly under-utilized in German forestry; Revenue per tree and harvesting productivity were both statistically higher with automatic bucking as compared to quality bucking during a thinning operation in a spruce dominated stand.
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Tree bucking, defined as the process in which a stem is segmented into shorter logs of varying lengths, has a significant effect on the value adding potential of a forest enterprise. Because of its importance in terms of correct product and length combinations, improper bucking can lead to financial losses. In this study, two treatments (OFF: quality bucking performed by the operator while using hot keys and ON: automatic bucking using the optimized suggestions from the harvester on-board computer; OBC) were tested in a Norway spruce (Picea abies [L.] Karst.) dominated stand located in Germany. Both treatments had the aim to maximize the value of a stem. The research took place in an 80-year old spruce and beech stand under a regenerative cutting. Fully-mechanized harvesting was performed with an 8-wheel Ponsse Bear single-grip harvester equipped with a H8 harvesting head. Results indicated that the product recovery of the two treatments differed by 4% in undamaged trees (no broken tree-tops or stems) to the benefit of manual bucking. However, the revenue of trees subjected to optimized bucking were up to 4% higher (in average 3%) than those of the manual bucking once expressed on a per cubic meter basis. Moreover, the harvesting productivity of the ON treatment was at the maximum 17% higher compared to the OFF treatment. Based on the results from this case study, the use of an optimization software in Norway spruce dominated stands with the aim to maximize the value of single stems showed promising results.

  • Labelle, Assistant Professorship of Forest Operations, Department of Ecology and Ecosystem Management, Technical University of Munich, Hans-Carl-von-Carlowitz-Platz 2, D-85354 Freising, Germany E-mail: eric.labelle@tum.de (email)
  • Huß, Assistant Professorship of Forest Operations, Department of Ecology and Ecosystem Management, Technical University of Munich, Hans-Carl-von-Carlowitz-Platz 2, D-85354 Freising, Germany E-mail: linus.huss@gmx.de
article id 9946, category Research article
Frauke Fedderwitz, Niklas Björklund, Velemir Ninkovic, Göran Nordlander. (2018). Does the pine weevil (Hylobius abietis) prefer conifer seedlings over other main food sources? Silva Fennica vol. 52 no. 3 article id 9946. https://doi.org/10.14214/sf.9946
Keywords: Pinus sylvestris; Picea abies; forest regeneration; feeding preference; herbivore; plant tissues
Highlights: Adult pine weevils feed on seedlings and mature conifers, but cause economic damage only on seedlings; Their feeding preferences for branches and roots over seedlings were tested in a laboratory experiment; The only clear preference was for Norway spruce roots; Results support new approaches of seedling protection attempting to redirect pine weevils from planted seedlings to other food sources.
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Adult pine weevils (Hylobius abietis (L.)) feed on the tender bark of branches and roots of mature conifer trees and on the stem bark of conifer seedlings. Their feeding on mature trees does not cause any economic damage, but their feeding on planted seedlings is so devastating that the pine weevil is considered one of the most important forest pest insects in Europe. We asked whether the pine weevil prefers seedlings over other regularly utilized food sources. This question is of particular interest because new approaches to seedling protection are based on decreasing any preference for seedlings by using less palatable plants or by enhancing their defence (by genetic selection or by methyl jasmonate treatment). In a laboratory choice experiment we tested pine weevil feeding preferences for seedlings compared with branches and roots from mature trees (separately for Norway spruce and Scots pine). Pine weevils preferred roots, but not branches, of Norway spruce over seedlings of the same species. With Scots pine there were no clear preferences, but the weevils showed a tendency to prefer roots over seedlings. These results provide support for seedling protection approaches that attempt to redirect pine feeding from planted seedlings to other food sources.

  • Fedderwitz, Department of Ecology, Swedish University of Agricultural Sciences, Box 7044, SE-750 07 Uppsala, Sweden E-mail: frauke.fedderwitz@slu.se (email)
  • Björklund, Department of Ecology, Swedish University of Agricultural Sciences, Box 7044, SE-750 07 Uppsala, Sweden E-mail: niklas.bjorklund@slu.se
  • Ninkovic, Department of Ecology, Swedish University of Agricultural Sciences, Box 7044, SE-750 07 Uppsala, Sweden E-mail: velemir.ninkovic@slu.se
  • Nordlander, Department of Ecology, Swedish University of Agricultural Sciences, Box 7044, SE-750 07 Uppsala, Sweden E-mail: goran.nordlander@slu.se
article id 9940, category Research article
Esteban Ceriani-Nakamurakare, Sergio Ramos, Carolina A. Robles, María V. Novas, María F. D´Jonsiles, Paola Gonzalez-Audino, Cecilia Carmarán. (2018). Metagenomic approach of associated fungi with Megaplatypus mutatus (Coleoptera: Platypodinae). Silva Fennica vol. 52 no. 3 article id 9940. https://doi.org/10.14214/sf.9940
Keywords: ambrosia; forest pest; metagenomic; fungi; Platypodinae; insect-fungus interactions
Highlights: There were no significant effects of host plant and location on fungal richness; Two fungal species, belonging to Fusarium and Candida genera, were present in all the studied associations; Results suggest that host plant identity would not be crucial to determine the composition of fungal communities associated to Megaplatypus mutatus.
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Megaplatypus mutatus is a major forest pest in Argentina and an emerging pest in Europe. In this study the multitrophic interactions between M. mutatus and associated fungi were assessed with a metagenomics approach (454-pyrosequencing). A total of 270 collection points from insect galleries from three locations in Argentina were pooled for pyrosequencing analyses. Two hosts, Populus deltoides and Casuarina cunninghamiana, were independently evaluated to characterize the fungal communities associated to M. mutatus; compare the culture-independent approach with previous culturing studies, in terms of data recovery related to the fungal community composition, and test the specificity of the fungal communities amongst locations and hosts. A Generalized Linear Mixed Model was performed to compare the fungal richness in each dataset, which showed no significant differences between taxa richness amongst locations. Principal Coordinates Analyses showed a separation between fungal communities within the same host, suggesting that host identity would not be crucial to determine the specificity in fungal communities. Candida insectalens and one Fusarium species, present in all hosts and locations, achieved 37.6% of the total relative frequency per taxa. These results complement the data from culturing methods previously reported, thus improving the accuracy and understanding of the fungal assemblages associated to M. mutatus.

  • Ceriani-Nakamurakare, Universidad de Buenos Aires. Facultad de Ciencias Exactas y Naturales. Depto. Biodiversidad y Biología Experimental. Buenos Aires, Argentina; CONICET- Universidad de Buenos Aires. Instituto de Micología y Botánica (INMIBO). Buenos Aires, (C1428EHA) Argentina E-mail: cerianinaka@gmail.com
  • Ramos, Instituto Nacional de Tecnología Agropecuaria. Estación Experimental Agropecuaria Concordia. Entre Ríos, (E3200) Argentina E-mail: ramos.sergio@inta.gob.ar
  • Robles, Universidad de Buenos Aires. Facultad de Ciencias Exactas y Naturales. Depto. Biodiversidad y Biología Experimental. Buenos Aires, Argentina; CONICET- Universidad de Buenos Aires. Instituto de Micología y Botánica (INMIBO). Buenos Aires, (C1428EHA) Argentina E-mail: carorobles@bg.fcen.uba.ar
  • Novas, Universidad de Buenos Aires. Facultad de Ciencias Exactas y Naturales. Depto. Biodiversidad y Biología Experimental. Buenos Aires, Argentina; CONICET- Universidad de Buenos Aires. Instituto de Micología y Botánica (INMIBO). Buenos Aires, (C1428EHA) Argentina E-mail: vicnovas@bg.fcen.uba.ar
  • D´Jonsiles, Universidad de Buenos Aires. Facultad de Ciencias Exactas y Naturales. Depto. Biodiversidad y Biología Experimental. Buenos Aires, Argentina; CONICET- Universidad de Buenos Aires. Instituto de Micología y Botánica (INMIBO). Buenos Aires, (C1428EHA) Argentina E-mail: lalijonsi@gmail.com
  • Gonzalez-Audino, Centro de Investigaciones de Plagas e Insecticidas (CITEFA-CONICET). Buenos Aires, (B1603ALO) Argentina E-mail: pgonzalezaudino@citedef.gob.ar
  • Carmarán, Centro de Investigaciones de Plagas e Insecticidas (CITEFA-CONICET). Buenos Aires, (B1603ALO) Argentina E-mail: carmaran@bg.fcen.uba.ar (email)
article id 9905, category Research article
Mercedes M. Fernandez, Diana Bezos, Julio J. Diez. (2018). Fungi associated with necrotic galls of Dryocosmus kuriphilus (Hymenoptera: Cynipidae) in northern Spain. Silva Fennica vol. 52 no. 3 article id 9905. https://doi.org/10.14214/sf.9905
Keywords: Asian chestnut gall wasp; Castanea sativa; Fusarium spp.; Gnomoniopsis smithogilvyi; entomopathogens; fungal diversity.
Highlights: Presence of Dryocosmus kuriphilus in Northern Spain; The mycobiota associated to necrotic galls was studied for the first time; 7 fungal species were identified; The entomopathogenic fungi found could be use as potential biological control agents; Gnomomiopsis smithogilvyi, Fusarium oxysporum and F. avenaceum known by their toxicity against the insect, were found.
Abstract | Full text in HTML | Full text in PDF | Author Info

The Asian chestnut gall wasp (ACGW), Dryocosmus kuriphilus Yasumatsu (Hymenoptera: Cynipidae) is one of the most important pests in Castanea species worldwide. In 2012, it was found for the first time in Catalonia (Spain) and a year later, in the north of Spain (Cantabria). Today, it is present in 14 Spanish provinces. In search of biological control against the ACGW, several authors have previously found the relationship between the presence of some Fusarium Link species in necrotic galls and wasp mortality due to the production of different types of wall-degrading enzymes and entomopathogenic mycotoxins. The objective of this study was to investigate the mycobiota associated with necrotic galls to find interesting perspectives for biological control of the ACGW. For this purpose, in 2014, 119 necrotic galls of Castanea sativa Miller were plated to isolate and identify the associated fungi. The fungal isolates were identified by the morphology of the fruiting bodies and DNA analyses. From necrotic galls, 7 species of fungi were identified. Of these, we highlight three species of Fusarium Link as well as the presence of Gnomoniopsis smithogilvyi Shuttlew, Liew & Guest due to its toxic capacity. Further studies are required to verify the effectiveness of these fungal species as biocontrol agents against the ACGW.

  • Fernandez, Dpt. of Agroforestry Sciences, ETSIIAA, University of Valladolid, Av. Madrid 50, 34071 Palencia, Spain; Sustainable Forest Management Research Institute UVa-INIA, ETSIIAA, 34071 Palencia, Spain ORCID http://orcid.org/0000-0002-1646-5027 E-mail: mffernan@agro.uva.es (email)
  • Bezos, Sustainable Forest Management Research Institute UVa-INIA, ETSIIAA, 34071 Palencia, Spain E-mail: dianabezos@yahoo.es
  • Diez, Sustainable Forest Management Research Institute UVa-INIA, ETSIIAA, 34071 Palencia, Spain; Dpt. of Plant Production and Forest Resources, University of Valladolid, Av. Madrid 50, 34071 Palencia, Spain E-mail: jdcasero@pvs.uva.es
article id 7710, category Research article
Pekka Hyvönen, Jaakko Heinonen. (2018). Estimating storm damage with the help of low-altitude photographs and different sampling designs and estimators. Silva Fennica vol. 52 no. 3 article id 7710. https://doi.org/10.14214/sf.7710
Keywords: simulation; simple random sampling estimator; ratio estimator; Lindeberg; flight line
Highlights: Digital photographs taken from low altitudes are usable for monitoring storm damage; Simple random sampling and ratio estimators resulted in similar standard errors; Characteristics of the storm influence the optimal flight plan and which variance estimator should be used; The developed model for simulations can be modified and utilized with future storms.
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Climate change has been estimated to increase the risk of storm damage in forests in Finland. There is a growing need for methods to obtain information on the extent and severity of storm damage after a storm occurrence. The first objective of this study was to test whether digital photographs taken from aircrafts flying at low-altitude can be utilized in locating storm-damaged areas and estimating the need for harvesting of wind-thrown trees. The second objective was to test the performance of selected estimators. Depending on distances between flight lines, plots on lines and the used estimator, the relative standard errors of storm area estimates varied between 7.7 and 48.7%. For the area for harvesting and volume of wind-thrown trees, the relative standard errors of estimates varied between 16.8 and 167.3%. Using forest area information from Multisource National Forest Inventory data improved the accuracy of the estimates. However, performance of a simple random sampling estimator and ratio estimator were quite similar. Lindeberg’s method for variance estimation based on adjacent lines was sensitive to line directions in relation to possible trends in storm-damaged area locations. Our results showed that the tested method could be used in estimating storm-damaged area provided that the network of flight lines and photographs on lines are sufficiently dense. The developed model for simulations can be utilized also with forthcoming storms as model’s parameters can be freely adjusted to meet, e.g., the intensity and extent of the storm.

  • Hyvönen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: pekka.hyvonen@luke.fi (email)
  • Heinonen, E-mail: jaakkoheinonen@gmail.com
article id 9938, category Research article
Jyrki Hytönen, Egbert Beuker, Anneli Viherä-Aarnio. (2018). Clonal variation in basic density, moisture content and heating value of wood, bark and branches in hybrid aspen. Silva Fennica vol. 52 no. 2 article id 9938. https://doi.org/10.14214/sf.9938
Keywords: moisture content; wood; bark; hybrid aspen; basic density; branches; clonal variation; heating value; ash content
Highlights: Hybrid aspen clones differed in their moisture content, ash content, basic density and heating value; Stem wood had lower ash content, basic density and effective heating value than stem bark; There was significant vertical variation in wood and bark along the stem in moisture content and basic density.
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Hybrid aspen (Populus tremula × P. tremuloides) is one of the fastest growing tree species in Finland. During the mid-1990s, a breeding programme was started with the aim of selecting clones that were superior in producing pulpwood. Hybrid aspen can also be grown as a short-rotation crop for bioenergy. To study clonal variation in wood and bark properties, seven clones were selected from a 12-year-old field trial located in southern Finland. From each clone, five trees were harvested and samples were taken from stem wood, stem bark and branches to determine basic density, effective heating value, moisture and ash content. Vertical within-tree variation in moisture content and basic density was also studied. The differences between clones were significant for almost all studied properties. For all studied properties there was a significant difference between wood and bark. Wood had lower ash content (0.5% vs. 3.9%), basic density (378 kg m–3 vs. 450 kg m–3) and effective heating value (18.26 MJ kg–1 vs. 19.24 MJ kg–1), but higher moisture content (55% vs. 49%) than bark. The values for branches were intermediate. These results suggest that the properties of hybrid aspen important for energy use could be improved by clonal selection. However, selecting clones based on fast growth only may be challenging since it may lead to a decrease in hybrid aspen wood density.

  • Hytönen, Natural Resources Institute Finland (Luke), Natural resources, Teknologiakatu 7, FI-67100 Kokkola, Finland E-mail: jyrki.hytonen@luke.fi (email)
  • Beuker, Natural Resources Institute Finland (Luke), Production systems, Vipusenkuja 6, FI-57200 Savonlinna, Finland E-mail: egbert.beuker@luke.fi
  • Viherä-Aarnio, Natural Resources Institute Finland (Luke), Production systems, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: anneli.vihera-aarnio@luke.fi
article id 9933, category Research article
Timo Saksa, Jari Miina, Hilkka Haatainen, Kauko Kärkkäinen. (2018). Quality of spot mounding performed by continuously advancing mounders. Silva Fennica vol. 52 no. 2 article id 9933. https://doi.org/10.14214/sf.9933
Keywords: soil preparation; slash; logging residues; forest regeneration; mixed modelling
Highlights: The number and quality of mounds varied considerably according to the operating conditions;The main factors reducing the quality of spot mounding were steep terrain, a thick humus layer, fresh logging residues, stoniness and soil texture;With careful selection of timing and conditions for mounding, the quality obtained by continuously advancing mounders can be improved.
Abstract | Full text in HTML | Full text in PDF | Author Info

Operating conditions affecting the quality of spot mounding by Bracke continuously advancing mounders were investigated on 66 regeneration areas (124 ha) in eastern Finland. The quality of mounds was classified as suitable (good or acceptable after additional compression) or unsuitable for planting. Models were constructed for the number of suitable planting spots obtained per hectare (good and acceptable mounds), the probability of successful mounding (≥1600 planting spots ha–1) and the probability of creating a suitable mound as a function of terrain, site and soil characteristics, as well as slash conditions (removed, fresh or dry logging residues). The average number of mounds created was 1892 ± 290 mounds ha–1, of which 1398 ± 325 mounds ha–1 (74%) were classified as suitable for planting. The quality of spot mounding was reduced by steep terrain, a thick humus layer and fresh logging residues. Stoniness and soil texture also affected the number of planting spots created. Mounding after logging residues had dried increased the number of planting spots by 191 spots ha–1 compared with mounding in the presence of fresh residues. Removing residues did not significantly increase the number of planting spots compared with mounding amongst dry residues. A thick humus layer, very stony soil, steep slopes and valley terrain decreased the number of planting spots by 150–450 spots ha–1. The number and quality of mounds varied considerably according to the operating conditions, but with careful selection of timing and sites the quality obtained by a continuously advancing mounder can be improved.

  • Saksa, Natural Resources Institute Finland (Luke), Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@luke.fi (email)
  • Miina, Natural Resources Institute Finland (Luke), Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: jari.miina@luke.fi
  • Haatainen, Faculty of Science and Forestry, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: hilkka.haatainen@storaenso.com
  • Kärkkäinen, Tornator Oyj, Muuntamontie 2, FI-80100 Joensuu, Finland E-mail: kauko.karkkainen@tornator.fi
article id 9927, category Research article
Eva Dařenová, Richard A. Crabbe, Robert Knott, Barbora Uherková, Jan Kadavý. (2018). Effect of coppicing, thinning and throughfall reduction on soil water content and soil CO2 efflux in a sessile oak forest. Silva Fennica vol. 52 no. 2 article id 9927. https://doi.org/10.14214/sf.9927
Keywords: soil respiration; coppice; precipitation; soil moisture; Quercus petraea
Highlights: The coppice stand reached similar soil CO2 efflux rates eight years after harvest as the original stand; Thinning increased SWC but did not affect soil CO2 efflux; 30% throughfall reduction decreased soil CO2 efflux by 50.7%.
Abstract | Full text in HTML | Full text in PDF | Author Info

In this study we determined the effect of transformation of a mature sessile oak forest stand into a coppiced forest, and of thinning and throughfall reduction in a coppice stand on soil water content (SWC) and soil CO2 efflux. The precipitation reduction was induced by installing parallel drainage channels in both unthinned and thinned coppice stands. The driving factor for temporal dynamics of soil CO2 efflux in all plots was soil temperature. The other factor was soil water content but only up to about 15%. Above this threshold, there was no more effect on CO2 efflux. We found no clear difference in SWC or soil CO2 efflux between the mature and coppiced stand eight years after harvesting. On the other hand, thinning of the coppice stand resulted in increase in SWC up to 22% in proportion, which we assume to be a result of increased gap fraction of the canopy. However, no effect on soil CO2 efflux was observed two years after the thinning. Installation of the drainage channels in two plots covering 30% of the ground area resulted in decrease in SWC up to a proportional 30.5% and thus contributed up to 50.7% reduction in soil CO2 efflux.

  • Dařenová, Global Change Research Institute CAS, v.v.i., Belidla 4a, 603 00 Brno, Czech Republic E-mail: darenova.e@czechglobe.cz (email)
  • Crabbe, Global Change Research Institute CAS, v.v.i., Belidla 4a, 603 00 Brno, Czech Republic E-mail: crabbe.r@czechglobe.cz
  • Knott, Mendel University in Brno, Zemedelska 3, 613 00 Brno, Czech Republic E-mail: robert.knott@mendelu.cz
  • Uherková, Mendel University in Brno, Zemedelska 3, 613 00 Brno, Czech Republic E-mail: xfedorov@node.mendelu.cz
  • Kadavý, Mendel University in Brno, Zemedelska 3, 613 00 Brno, Czech Republic E-mail: jan.kadavy@mendelu.cz
article id 9914, category Research article
Jun Tanabe, Futoshi Ishiguri, Akira Tamura, Yuya Takashima, Jyunichi Ohshima, Kazuya Iizuka, Shinso Yokota. (2018). Within-tree radial and among-family variations in wood density, microfibril angle, and mechanical properties in Picea glehnii. Silva Fennica vol. 52 no. 2 article id 9914. https://doi.org/10.14214/sf.9914
Keywords: modulus of elasticity; modulus of rupture; early selection; tree breeding for wood quality
Highlights: The modulus of elasticity was affected by both microfibril angle and wood density, whereas the modulus of rupture was mainly affected by wood density in Picea glehnii; A larger degree of among-family variation in wood properties was detected in juvenile wood than in mature wood, indicating that genetic improvements in the mechanical properties may be more effective for juvenile wood.
Abstract | Full text in HTML | Full text in PDF | Author Info

Genetic improvements in the mechanical properties of wood are important in forestry species used for lumber, such as Picea. The within-tree radial and among-family variations for the modulus of elasticity (MOE), modulus of rupture (MOR), and their related traits [i.e., microfibril angle (MFA) of the S2 layer in latewood tracheid and air-dry density (AD)] were evaluated in nine open-pollinated families of Picea glehnii (F. Schmidt) Mast. The radial variation in MOR was mainly affected by AD, whereas MOE was affected by MFA and AD. Higher F-values obtained by analysis of variance and coefficient of variation were observed for all properties at the 6th–15th annual ring, except for AD at the 6th–10th annual ring. This result suggests that the contribution of genetic effect is larger in these highly variable regions. In addition, positive correlation coefficients were obtained between wood properties at the 6th–15th annual ring and mean values of these properties. Therefore, genetic improvements for MOE, MOR, and their related traits in P. glehnii is likely to be more effective in juvenile wood, specifically at the 6th–15th annual ring from the pith.

  • Tanabe, Faculty of Education, Chiba University, Chiba, 263-8522, Japan E-mail: tanabe_j@chiba-u.jp (email)
  • Ishiguri, Faculty of Agriculture, Utsunomiya University, Utsunomiya 321-8505, Japan E-mail: ishiguri@cc.utsunomiya-u.ac.jp
  • Tamura, Forest Tree Breeding Center, Forestry and Forest Products Research Institute, Hitachi, 319-1301, Japan E-mail: akirat@affrc.go.jp
  • Takashima, Forest Tree Breeding Center, Forestry and Forest Products Research Institute, Hitachi, 319-1301, Japan E-mail: ytakashima@ffpri.affrc.go.jp
  • Ohshima, Faculty of Agriculture, Utsunomiya University, Utsunomiya 321-8505, Japan E-mail: joshima@cc.utsunomiya-u.ac.jp
  • Iizuka, Faculty of Agriculture, Utsunomiya University, Utsunomiya 321-8505, Japan E-mail: kiizuka@cc.utsunomiya-u.ac.jp
  • Yokota, Faculty of Agriculture, Utsunomiya University, Utsunomiya 321-8505, Japan E-mail: yokotas@cc.utsunomiya-u.ac.jp
article id 9902, category Research article
Perttu Anttila, Vesa Nivala, Olli Salminen, Markus Hurskainen, Janne Kärki, Tomi J. Lindroos, Antti Asikainen. (2018). Regional balance of forest chip supply and demand in Finland in 2030. Silva Fennica vol. 52 no. 2 article id 9902. https://doi.org/10.14214/sf.9902
Keywords: bioenergy; energy wood; GIS; availability; potential
Highlights: The impact of increasing forest chip demand in 2030 was analyzed in Finland; Demand of small trees may exceed potential at the national level; Surplus potential will remain in logging residues and stumps; Hot spots of demand call for efficient logistical solutions.
Abstract | Full text in HTML | Full text in PDF | Author Info

According to the National Energy and Climate Strategy of Finland in 2016, the demand for forest chips, that is, wood chips made of forest biomass directly for energy use, could even double by 2030 compared to the present situation. A spatially explicit impact analysis of regional supply and demand balances for forest chips was carried out. The balances were calculated as the difference between technical harvesting potentials and demand. First, the technical potentials were estimated based on the national forest inventory data. Secondly, three demand scenarios were defined for 2030 and subsequently deducted from the potentials. The results suggested that there would be increasing competition for feedstock in southern and western Finland, whereas in eastern and northern Finland there would still be surplus potential. Moreover, due to the remarkable deficit of small trees in southern Finland, there might be pressure towards using more pulpwood-sized and/or imported wood in energy production. The results also showed that, in particular, large new plants consuming substantial amounts of forest chips could have a significant effect on the regional availability of forest chips. Moreover, with increasing transport distances, new logistical solutions will be needed.

  • Anttila, Natural Resources Institute Finland (Luke), Yliopistokatu 6, FI-80100 Joensuu, Finland ORCID http://orcid.org/0000-0002-6131-392X E-mail: perttu.anttila@luke.fi (email)
  • Nivala, Natural Resources Institute Finland (Luke), Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: vesa.nivala@luke.fi
  • Salminen, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: olli.salminen@luke.fi
  • Hurskainen, VTT Technical Research Centre of Finland Ltd, Koivurannantie 1, FI-40400 Jyväskylä, Finland E-mail: markus.hurskainen@vtt.fi
  • Kärki, VTT Technical Research Centre of Finland Ltd, Koivurannantie 1, FI-40400 Jyväskylä, Finland E-mail: janne.karki@vtt.fi
  • Lindroos, VTT Technical Research Centre of Finland Ltd, Vuorimiehentie 3 (Espoo), P.O. Box 1000, FI-02044 VTT, Finland E-mail: tomi.j.lindroos@vtt.fi
  • Asikainen, Natural Resources Institute Finland (Luke), Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: antti.asikainen@luke.fi
article id 7830, category Research article
Jari Lindblad, Johanna Routa, Johanna Ruotsalainen, Marja Kolström, Ari Isokangas, Lauri Sikanen. (2018). Weather based moisture content modelling of harvesting residues in the stand. Silva Fennica vol. 52 no. 2 article id 7830. https://doi.org/10.14214/sf.7830
Keywords: logging residues; harvesting residues; energy wood measurement; conversion factor
Highlights: Weather data used for estimating the moisture content of energy wood; The validation of the weather based models performed based on the field data.
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Harvesting residues collected from the final cuttings of boreal forests are an important source of solid biofuel for energy production in Finland and Sweden. In the Finnish supply chain, the measurement of residues is performed by scales integrated in forwarders. The mass of residues is converted to volume by conversion factors. In this study, weather based models for defining the moisture content of residues were developed and validated. Models were also compared with the currently used fixed tables of conversion factors. The change of the moisture content of residues is complex, and an exact estimation was challenging. However, the model predicting moisture change for three hour periods was found to be the most accurate. The main improvement compared to fixed tables was the lack of a systematic error. It can be assumed that weather based models will give more reliable estimates for the moisture in varying climate conditions and the further development of models should be focused on obtaining more appropriate data from varying drying conditions in different geographical and microclimatological locations.

  • Lindblad, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: jari.lindblad@luke.fi (email)
  • Routa, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: johanna.routa@luke.fi
  • Ruotsalainen, Finnish Meteorological Institute, Aviation and Military Weather Services, P.O. Box 1627, FI-70211 Kuopio, Finland E-mail: johanna.ruotsalainen@fmi.fi
  • Kolström, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: marja.kolstrom@uef.fi
  • Isokangas, University of Oulu, Control Engineering, P.O. Box 8000, FI-90014 University of Oulu, Finland E-mail: ari.isokangas@oulu.fi
  • Sikanen, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: lauri.sikanen@luke.fi
article id 7816, category Research article
Pentti Niemistö, Harri Kilpeläinen, Eero Poutiainen. (2018). Effect of first thinning type and age on growth, stem quality and financial performance of a Scots pine stand in Finland. Silva Fennica vol. 52 no. 2 article id 7816. https://doi.org/10.14214/sf.7816
Keywords: bucking; growth and yield; external stem quality; profitability; Pinus silvestris; tree selection; saw log
Highlights: Early commercial first thinning at the top height of 11 m with tree selection targeting high quality of remaining stems was more profitable in a Scots pine stand than thinning from below at similar time or delayed thinning 10 years later; After early first thinning only one intermediate thinning was needed, but a late thinning at age of 60 years was feasible in connection with increased amount of high quality butt logs after artificial pruning; The optimal rotation period for a Scots pine stand using a lower interest rate of 1% was 80–85 years depending on the thinning pattern. With a higher rate of 4%, the optimal rotation took only 60 years.
Abstract | Full text in HTML | Full text in PDF | Author Info

The objective of the study was to ascertain the effects of tree selection (thinning from below, from above and according to stem quality) and timing of first commercial thinning (early and delayed) on the growth, yield and quality of trees in a Scots pine (Pinus sylvestris L.) stand. A long-term field experiment (25 years) was measured in 5-year periods and the further development was simulated with growth and yield models to final cuttings using alternative rotation periods of 55–85 years. The measurements included also the exact location and type of technical defects detected on all trees in the experimental plots. The measured volume increment per unit area during the study period, 25 years after the early thinning stage was the lowest in the plots thinned from below, and the highest in the plots thinned from above or in the delayed thinning plots. However, the largest volume of saw logs during the whole rotation of 80 years was yielded after early first thinning according to the quality. The largest volume of very high-quality butt logs was produced by pruning connected with early thinning from above, and a smaller volume after early thinning according to stem quality but no after thinning from below or delayed first thinning. Without pruning an early quality thinning with one intermediate thinning was the most profitable thinning treatment in the Scots pine stand regardless the rotation length or the interest rate used. By interest rates of 1% and 2%, the optimal rotations were 80–85 years and 70 years respectively. A late thinning at the age of 60 year with long rotation was profitable only for the pruned pine stands with a low interest rate.

  • Niemistö, Natural Resources Institute Finland (Luke), Natural resources, Kampusranta 9 C, FI-60320 Seinäjoki, Finland E-mail: pentti.niemisto@luke.fi (email)
  • Kilpeläinen, Natural Resources Institute Finland (Luke), Production systems, Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: harri.kilpelainen@luke.fi
  • Poutiainen, Oulu E-mail: eero1.poutiainen@dnainternet.net
article id 7813, category Research article
Jaana Luoranen. (2018). Autumn versus spring planting: the initiation of root growth and subsequent field performance of Scots pine and Norway spruce seedlings. Silva Fennica vol. 52 no. 2 article id 7813. https://doi.org/10.14214/sf.7813
Keywords: bud burst; planting date; root growth; shoot growth
Highlights: Conifer seedlings planted after mid-September generally have poor rooting, which causes poor root egress during the following spring; Although Scots pine and Norway spruce seedlings planted in late autumn may have a slightly reduced growth, it is possible to plant them if weather conditions are favorable in late-autumn, without increased mortality.
Abstract | Full text in HTML | Full text in PDF | Author Info

There is a need to extend the planting season of conifer regeneration into periods where the soil remains unfrozen due to a lack of available labor and the mechanization of planting. This study investigated how the summer- (August) and autumn-, especially late autumn (mid-September to mid-October) plantings affect the field performance of Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.) container seedlings. This study examined the timing of root growth just after planting, shoot flush and the start of root growth the following spring, and subsequent field performance. Seedlings of both species were planted in a nursery field trial, and in a clearcut reforestation site from August to October and the following May. The root growth of planted seedlings declined in September and ceased after mid-September. In the following spring, seedlings which were planted in early-autumn started their root growth faster than late-autumn-planted seedlings in both species. There was no difference in the timing of shoot flush for various planting dates. During the initial two years after planting, the shoot growth of spring-planted seedlings was lower, compared to autumn-planted seedlings. In conclusion, it is possible to plant conifer seedlings in the boreal forest zone up to October under non-limiting field conditions.

  • Luoranen, Natural Resources Institute Finland (Luke), Production systems, FI-77600 Suonenjoki, Finland E-mail: jaana.luoranen@luke.fi (email)
article id 7772, category Research article
Curt Almqvist. (2018). Improving floral initiation in potted Picea abies by supplemental light treatment. Silva Fennica vol. 52 no. 2 article id 7772. https://doi.org/10.14214/sf.7772
Keywords: Norway spruce; seed production; flower stimulation; strobili initiation; indoor seed orchard
Highlights: Supplemental light treatment:
  • Increases the proportion of genotypes initiating reproductive buds.
  • Increases floral induction, especially of female floral buds.
  • Facilitates breeding programmes, and seed production of highly improved base material from new selections for vegetative production programmes, to be more efficient.
Abstract | Full text in HTML | Full text in PDF | Author Info

Light is an important environmental factor for all green plants. Its intensity, spectral composition and photoperiod can affect the regulatory pathways in plants that lead to floral initiation. In this report, results are presented from three experiments in which supplemental light with metal halide lamps (250 µmol m–2 s–1, 20 hours day–1, approx. 6 weeks) was tested as a complement to other flowering stimulation treatments (elevated temperature, treatment with gibberellin A4 and A 7 (GA4/7), restricted water supply) applied to potted Picea abies (L.) Karst. in the greenhouse. Flower stimulation in a greenhouse resulted in more floral initiation compared to flower stimulation outdoors. Supplemental light treatment increased floral initiation further, and to a larger extent in female than in male flowers. It also increased the proportion of trees and genotypes that induced reproductive buds. In a practical application of the supplemental light treatment to potted Picea abies breeding material, 90.6% of the clones produced either female or male flowers, or both. A subset of the same material kept outdoors, and thus subjected to natural light and temperatures, produced no flowers despite being treated with GA4/7 and receiving a restricted water supply. In conclusion, supplemental light treatment facilitates breeding programmes, and seed production of highly improved base material from new selections for vegetative production programmes, to be more efficient.

  • Almqvist, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, 751 83 Uppsala, Sweden ORCID http://orcid.org/0000-0001-5739-4854 E-mail: curt.almqvist@skogforsk.se (email)
article id 7740, category Research article
Jonas R. Coussement, Kathy Steppe, Peter Lootens, Isabel Roldán-Ruiz, Tom De Swaef. (2018). A flexible geometric model for leaf shape descriptions with high accuracy. Silva Fennica vol. 52 no. 2 article id 7740. https://doi.org/10.14214/sf.7740
Keywords: image processing; leaf contour; leaf shape; shape function; digitising; tree leaf shape
Highlights: A method for assessing leaf shape for 3D plant models is proposed; The model is highly flexible and fits a large variety of shapes; It allows analysis of shape differences within and between leaf datasets.
Abstract | Full text in HTML | Full text in PDF | Author Info

Accurate assessment of canopy structure is crucial in studying plant-environment interactions. The advancement of functional-structural plant models (FSPM), which incorporate the 3D structure of individual plants, increases the need for a method for accurate mathematical descriptions of leaf shape. A model was developed as an improvement of an existing leaf shape algorithm to describe a large variety of leaf shapes. Modelling accuracy was evaluated using a spatial segmentation method and shape differences were assessed using principal component analysis (PCA) on the optimised parameters. Furthermore, a method is presented to calculate the mean shape of a dataset, intended for obtaining a representative shape for modelling purposes. The presented model is able to accurately capture a large range of single, entire leaf shapes. PCA illustrated the interpretability of the parameter values and allowed evaluation of shape differences. The model parameters allow straightforward digital reconstruction of leaf shapes for modelling purposes such as FSPMs.

  • Coussement, Plant Sciences Unit, Institute for Agricultural and Fisheries Research (ILVO), Caritasstraat 39, B-9090 Melle, Belgium; Laboratory of Plant Ecology, Department of Plants and Crops, Faculty of Bioscience Engineering, Ghent University, Coupure links 653, B-9000 Ghent, Belgium E-mail: jonas.coussement@ilvo.vlaanderen.be
  • Steppe, Laboratory of Plant Ecology, Department of Plants and Crops, Faculty of Bioscience Engineering, Ghent University, Coupure links 653, B-9000 Ghent, Belgium E-mail: kathy.steppe@ugent.be
  • Lootens, Plant Sciences Unit, Institute for Agricultural and Fisheries Research (ILVO), Caritasstraat 39, B-9090 Melle, Belgium E-mail: peter.lootens@ilvo.vlaanderen.be
  • Roldán-Ruiz, Plant Sciences Unit, Institute for Agricultural and Fisheries Research (ILVO), Caritasstraat 39, B-9090 Melle, Belgium; Department of Plant Biotechnology and Bioinformatics, Faculty of Sciences, Ghent University, Technologiepark Zwijnaarde 927, B-9052 Zwijnaarde, Belgium E-mail: isabel.roldan-ruiz@ilvo.vlaanderen.be
  • De Swaef, Plant Sciences Unit, Institute for Agricultural and Fisheries Research (ILVO), Caritasstraat 39, B-9090 Melle, Belgium E-mail: tom.deswaef@ilvo.vlaanderen.be (email)
article id 7738, category Research article
Samuel Egbäck, Bo Karlsson, Karl-Anders Högberg, Kenneth Nyström, Mateusz Liziniewicz, Urban Nilsson. (2018). Effects of phenotypic selection on height-diameter ratio of Norway spruce and Scots pine in Sweden. Silva Fennica vol. 52 no. 2 article id 7738. https://doi.org/10.14214/sf.7738
Keywords: Pinus sylvestris; Picea abies; genetic correlations; heritability; Genetic selection; slenderness
Highlights: Swedish plus-tree selection promoted less slender Norway spruce trees and more slender Scots pine trees compared to neighboring trees; Similar results were also found for progeny trials which indicated that genetics played a prominent role in phenotypic appearance.
Abstract | Full text in HTML | Full text in PDF | Author Info

Genetically improved Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) are extensively used in operational Swedish forestry plantations. However, relatively little is known about the stem slenderness (height-diameter ratio) of genetically improved material. Thus, in this study we investigated effects of plus-tree selection on stem slenderness of Norway spruce and Scots pine in Sweden by evaluating both the plus-tree selection and a large number of progeny trials. Species-specific models for predicting the height-diameter ratio were estimated using regression and mixed model approach. Our results show that phenotypic plus-tree selection promoted less slender Norway spruce trees and more slender Scots pine trees compared to neighboring trees. Similar results were also found for the progeny trials which indicated that genetics played a prominent role in the phenotypic appearance. Compared to the progeny of neighboring trees, Norway spruce plus-tree progenies had a 5.3% lower height-diameter ratio, while Scots pine plus-tree progenies had a 1.5% greater height-diameter ratio. The narrow sense heritability for height-diameter ratio was 0.19 for Norway spruce and 0.11 for Scots pine, indicating that it is possible to modify the height-diameter ratio by breeding. Correlation coefficients between breeding values for height-diameter ratio and diameter were negative for Scots pine (–0.71) and Norway spruce (–0.85), indicating that selection for diameter only would result in less slender stems of both species. Similar correlations were also found between breeding values for height-diameter ratio and height of Scots pine (–0.34) and Norway spruce (–0.74).

  • Egbäck, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, 230 53 Alnarp, Sweden E-mail: samuel.egback@slu.se (email)
  • Karlsson, Skogforsk, Ekebo, 268 90 Svalöv, Sweden E-mail: bo.karlsson@skogforsk.se
  • Högberg, Skogforsk, Ekebo, 268 90 Svalöv, Sweden E-mail: karl-anders.hogberg@skogforsk.se
  • Nyström, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd, 901 83 Umeå, Sweden E-mail: kenneth.nystrom@slu.se
  • Liziniewicz, Skogforsk, Ekebo, 268 90 Svalöv, Sweden E-mail: Mateusz.Liziniewicz@skogforsk.se
  • Nilsson, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, 230 53 Alnarp, Sweden E-mail: urban.nilsson@slu.se
article id 7699, category Research article
Dalytė Matulevičiūtė, Jurga Motiejunaite, Domas Uogintas, Ričardas Taraškevičius, Mindaugas Dagys, Valerijus Rašomavičius. (2018). Decline of a protected coastal pine forest under impact of a colony of great cormorants and the rate of vegetation change under ornithogenic influence. Silva Fennica vol. 52 no. 2 article id 7699. https://doi.org/10.14214/sf.7699
Keywords: coniferous forests; piscivorous birds; Baltic Sea; habitats
Highlights: We studied vegetation and its changes in a pine forest affected by a colony of great cormorants; Vegetation in the colony varied according to the period of bird influence and the stand elevation above sea level; Considerable vegetation changes occur in several years after bird colony establishment; Pine forest ecosystem cease to exist following a decade of bird activity.
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We investigated the impact of a colony of great cormorants on the vegetation of the old growth Pinus sylvestris L. forest on the Curonian Spit peninsula, Lithuania. We studied the characteristics and rates of plant cover changes under varying length and intensity of bird influence. Plant species numbers, as well as the coverage of plants with different ecological requirements, varied according to the period of bird influence, but the resulting vegetation also depended upon the stand elevation above sea level. In our study, the initial increase in plant species richness at the start of bird nesting was not obvious and was of a transient character, due to the weak invasion of non-forest species and the rapid decline of forest plants. The colony area showed obvious and rapid vegetation changes during the investigation period. According to the calculated colony expansion rates, after 6–7 years of impact from birds the tree layer decreased by about four fold; the shrub layer decreased by about two fold; the field layer decreased by about 15 fold; and the dwarf shrub and bottom layers disappeared. The coverage by oligotrophic species decreased by more than four fold, while the coverage by eutrophic species increased by more than 60 fold. After 9–10 years of ornithogenic impact, all the trees were dead, and the protected coniferous forest ecosystem, with its characteristic plant species, had ceased to exist as such.

  • Matulevičiūtė, Institute of Botany, Nature Research Centre, Žaliųjų ežerų Str. 49, LT-08406 Vilnius, Lithuania ORCID http://orcid.org/0000-0001-7550-1860 E-mail: dalyte.matuleviciute@botanika.lt (email)
  • Motiejunaite, Institute of Botany, Nature Research Centre, Žaliųjų ežerų Str. 49, LT-08406 Vilnius, Lithuania ORCID http://orcid.org/0000-0002-6949-1990 E-mail: jurga.motiejunaite@botanika.lt
  • Uogintas, Institute of Botany, Nature Research Centre, Žaliųjų ežerų Str. 49, LT-08406 Vilnius, Lithuania ORCID http://orcid.org/0000-0002-3937-1218 E-mail: domas.uogintas@botanika.lt
  • Taraškevičius, Geology and Geography Institute, Nature Research Centre, Akademijos Str. 2, LT-08412 Vilnius, Lithuania ORCID http://orcid.org/0000-0002-5110-6142 E-mail: taraskevicius@geo.lt
  • Dagys, Institute of Ecology, Nature Research Centre, Akademijos Str. 2, LT-08412 Vilnius, Lithuania ORCID http://orcid.org/0000-0001-9342-3464 E-mail: dagys@ekoi.lt
  • Rašomavičius, Institute of Botany, Nature Research Centre, Žaliųjų ežerų Str. 49, LT-08406 Vilnius, Lithuania ORCID http://orcid.org/0000-0003-1314-4356 E-mail: valerijus.rasomavicius@botanika.lt
article id 9923, category Research article
Annika Kangas, Terje Gobakken, Stefano Puliti, Marius Hauglin, Erik Naesset. (2018). Value of airborne laser scanning and digital aerial photogrammetry data in forest decision making. Silva Fennica vol. 52 no. 1 article id 9923. https://doi.org/10.14214/sf.9923
Keywords: forest inventory; value of information; uncertainty; remote sensing; cost-plus-loss; data quality
Highlights: Airborne laser scanning (ALS) and digital aerial photogrammetry (DAP) are nearly equally valuable for harvest scheduling decisions even though ALS data is more precise; Large underestimates of stand volume are most dangerous errors for forest owner because of missed cutting probabilities; Relative RMSE of stand volume and the mean volume in a test area explain 77% of the variation between the expected losses due to errors in the data in the published studies; Increasing the relative RMSE of volume by 1 unit, increased the losses in average by 4.4 € ha–1.
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Airborne laser scanning (ALS) has been the main method for acquiring data for forest management planning in Finland and Norway in the last decade. Recently, digital aerial photogrammetry (DAP) has provided an interesting alternative, as the accuracy of stand-based estimates has been quite close to that of ALS while the costs are markedly smaller. Thus, it is important to know if the better accuracy of ALS is worth the higher costs for forest owners. In many recent studies, the value of forest inventory information in the harvest scheduling has been examined, for instance through cost-plus-loss analysis. Cost-plus-loss means that the quality of the data is accounted for in monetary terms through calculating the losses due to errors in the data in the forest management planning context. These costs are added to the inventory costs. In the current study, we compared the losses of ALS and DAP at plot level. According to the results, the data produced using DAP are as good as data produced using ALS from a decision making point of view, even though ALS is slightly more accurate. ALS is better than DAP only if the data will be used for more than 15 years before acquiring new data, and even then the difference is quite small. Thus, the increased errors in DAP do not significantly affect the results from a decision making point of view, and ALS and DAP data can be equally well recommended to the forest owners for management planning. The decision of which data to acquire, can thus be made based on the availability of the data on first hand and the costs of acquiring it on the second hand.

  • Kangas, Natural Resources Institute Finland (Luke), Bioeconomy and environment, P.O. Box 68, FI-80170 Joensuu, Finland E-mail: annika.kangas@luke.fi (email)
  • Gobakken, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: terje.gobakken@nmbu.no
  • Puliti, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: stefano.puliti@nibio.no
  • Hauglin, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: marius.hauglin@nmbu.no
  • Naesset, Faculty of Environmental Sciences and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, NO-1432 Ås, Norway E-mail: erik.naesset@nmbu.no
article id 7822, category Research article
Mulualem Tigabu, Annika M. Felton. (2018). Multivariate calibration of near infrared spectra for predicting nutrient concentrations of solid moose rumen contents. Silva Fennica vol. 52 no. 1 article id 7822. https://doi.org/10.14214/sf.7822
Keywords: Alces alces; lignin; NIRS; acid-detergent fiber; crude protein; microbial-N; neutral detergent fiber
Highlights: Multivariate calibrations were established for predicting nutrient concentrations of solid moose rumen contents by near infrared spectroscopy (NIRS); Crude protein, available protein and ash contents were accurately predicted; Prediction of microbial nitrogen, ash, acid-detergent fiber, neutral detergent fiber and lignin were satisfactory; The results demonstrate that NIRS offers quick and inexpensive procedure to quantify nutrient concentrations of solid rumen contents.
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This study aimed at establishing calibrations to predict nutrient concentrations of solid moose (Alces alces L.) rumen content using near infrared spectroscopy (NIRS), as an alternative to expensive chemical analyses. NIR reflectance spectra of 148 dry pulverized samples were recorded. The scanned samples were then analyzed for crude protein, available protein, microbial nitrogen (N), ash, acid-detergent fiber (ADF), neutral detergent fiber (NDF) and lignin contents following standard chemical analysis procedures. The calibration models were derived by Orthogonal Projection to Latent Structure (OPLS) and validated using external prediction sets. The calibration models accurately predicted crude protein, available protein and ash contents (R2 = 0.99, 0.96, and 0.92, prediction error = 0.39, 0.72 and 0.53% dry matter, respectively) while NDF (R2 = 0.92; prediction error = 2.23% dry matter) and ADF (R2 = 0.89; prediction error = 1.94% dry matter) were predicted with sufficient accuracy and that of microbial-N (R2 = 0.81; prediction error = 1.25 mg yeast-RNA g–1 dry matter) and lignin (R2 = 0.84; prediction error = 1.05% dry matter) were acceptable. The ratio of performance to deviation values were > 3.0 for crude protein and available protein, between 3.0 and 2.5 for ADF, NDF and lignin, and 2.32 for microbial-N; attesting the robustness of the calibration models. It can be concluded that NIR spectroscopy offers a quick and inexpensive procedure for prediction of nutrient concentrations of solid rumen contents in wild herbivores.

  • Tigabu, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, Box 49, SE-230 53 Alnarp, Sweden ORCID http://orcid.org/0000-0003-2471-1168 E-mail: mulualem.tigabu@slu.se (email)
  • Felton, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, Box 49, SE-230 53 Alnarp, Sweden E-mail: annika.felton@slu.se
article id 7803, category Research article
Lingbo Dong, Pete Bettinger, Huiyan Qin, Zhaogang Liu. (2018). Reflections on the number of independent solutions for forest spatial harvest scheduling problems: a case of simulated annealing. Silva Fennica vol. 52 no. 1 article id 7803. https://doi.org/10.14214/sf.7803
Keywords: simulated annealing; forest management planning; combinatorial optimization; neighborhood search; adjacency constraint
Highlights: No one particular neighborhood search technique of simulated annealing was found to be universally acceptable; The optimal number of independent solutions necessary for addressing the area restriction harvest scheduling model was described with a negative logarithmic function that was related with the problem size. However, optimal number of independent solutions necessary was not sensitive to the problem size for non-spatial and unit restriction harvest scheduling model problems, which should be somewhat above 250 independent runs; The types of adjacency constraints have moderate effects on the number of independent solutions, but these effects are not significant.
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To assess the quality of results obtained from heuristics through statistical procedures, a number of independently generated solutions to the same problem are required, however the knowledge of how many solutions are necessary for this purpose using a specific heuristic is still not clear. Therefore, the overall aims of this paper are to quantitatively evaluate the effects of the number of independent solutions generated on the forest planning objectives and on the performance of different neighborhood search techniques of simulated annealing (SA) in three increasing difficult forest spatial harvest scheduling problems, namely non-spatial model, area restriction model (ARM) and unit restriction model (URM). The tested neighborhood search techniques included the standard version of SA using the conventional 1-opt moves, SA using the combined strategy that oscillates between the conventional 1-opt moves and the exchange version of 2-opt moves, and SA using the change version of 2-opt moves. The obtained results indicated that the number of independent solutions generated had clear effects on the conclusions of the performances of different neighborhood search techniques of SA, which indicated that no one particular neighborhood search technique of SA was universally acceptable. The optimal number of independent solutions generated for all alternative neighborhood search techniques of SA for ARM problems could be estimated using a negative logarithmic function based on the problem size, however the relationships were not sensitive (i.e., 0.13 < p < 0.78) to the problem size for non-spatial and URM harvest scheduling problems, which should be somewhat above 250 independent runs. The types of adjacency constraints did moderately affect the number of independent solutions necessary, but not significantly. Therefore, determining an optimal number of independent solutions generated is a necessary process prior to employing heuristics in forest management planning practices.

  • Dong, College of Forestry, Northeast Forestry University, Harbin 150040, China E-mail: farrell0503@126.com
  • Bettinger, Warnell School of Forestry and Natural Resources, University of Georgia, Athens 30602, GA, USA E-mail: pbettinger@warnell.uga.edu
  • Qin, College of Economic and Management, Northeast Forestry University, Harbin 150040, China E-mail: huiyanqin@hotmail.com
  • Liu, College of Forestry, Northeast Forestry University, Harbin 150040, China E-mail: lzg19700602@163.com (email)
article id 7791, category Research article
Tadeusz B. Splawinski, Sylvie Gauthier, Nicole J. Fenton, Daniel Houle, Yves Bergeron. (2018). The colonization of young fire initiated stands by the crustose lichen Trapeliopsis granulosa and its potential effect on conifer establishment and stand succession. Silva Fennica vol. 52 no. 1 article id 7791. https://doi.org/10.14214/sf.7791
Keywords: establishment; Pinus banksiana; forest ecology; lichen woodland; stand succession.; Trapeliopsis granulosa
Highlights: T. granulosa is a poor seedbed for jack pine establishment; The presence of extensive T. granulosa cover can limit ongoing tree recruitment, thereby maintaining open lichen woodland; Dry open conditions favor the establishment of T. granulosa; Stands with significant T. granulosa cover may be good candidates for afforestation initiatives due to lower evaporation potential and decreased water stress.
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The resilience of closed-crown coniferous stands within the boreal forest of North America is highly dependent on successful re-establishment of tree species following fire. A shift from closed-crown forest to open lichen woodland is possible following poor natural regeneration during the initial establishment phase, followed by the development of extensive lichen cover, which may hinder ongoing recruitment. We examined the development of the crustose lichen Trapeliopsis granulosa (Hoffm.) 18 to 21 years following fire within six sites in the boreal forest of northwestern Quebec, and explored its potential to affect ongoing recruitment during early successional stages of stand development. Germination and survivorship trials were conducted within the laboratory to determine the establishment rate of Pinus banksiana Lamb. (jack pine) on T. granulosa, mineral soil, and burnt duff under two separate watering frequencies (observed and drought). Survival and establishment rates of jack pine were highest on burnt duff, and poor on both T. granulosa and mineral soil. Under the drought treatment, no seedlings survived on any substrates. In the field, T. granulosa cover had a positive relationship with mineral soil cover, and negative relationships with duff cover, ericaceous shrub cover, organic layer depth, other lichen cover, and Sphagnum moss cover. No discernable relationship was found between T. granulosa and tree density, rock cover, dead wood cover or other moss cover. The development of extensive T. granulosa cover in fire-initiated stands can impede ongoing recruitment of conifer species due to its poor seedbed quality, thereby maintaining open forests.

  • Splawinski, Institut de recherche sur les forêts, Université du Québec en Abitibi-Témiscamingue, 445, boul. de l’Université, Rouyn-Noranda, QC, J9X 5E4, Canada E-mail: tsplawinski@gmail.com (email)
  • Gauthier, Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, 1055 rue du PEPS, P.O. Box 10380, Stn Sainte Foy, QC, G1V 4C7, Canada E-mail: sylvie.gauthier@rncan-nrcan.gc.ca
  • Fenton, Institut de recherche sur les forêts (IRF), Université du Québec en Abitibi-Témiscamingue, 445 boul. de l’Université, Rouyn-Noranda, QC, J9X 5E4, Canada E-mail: nicole.fenton@uqat.ca
  • Houle, Ministère des Forêts, de la Faune et des Parcs, Direction de la recherché forestière, Québec, QC, G1P 3W8, Canada; Ouranos Climate Change Consortium, Montréal, QC, H3A 1B9, Canada E-mail: daniel.houle@mffp.gouv.qc.ca
  • Bergeron, Centre d’étude sur la forêt and Chaire industrielle en aménagement forestier durable, Université du Québec à Montréal, CP 8888 Succursale A, Montréal, QC, H3C 3P8, Canada E-mail: bergeron.yves@uqam.ca
article id 7003, category Research article
Poje Anton, Raffaele Spinelli, Natascia Magagnotti, Matevz Mihelic. (2018). The effect of feedstock, knife wear and work station on the exposure to noise and vibrations in wood chipping operations. Silva Fennica vol. 52 no. 1 article id 7003. https://doi.org/10.14214/sf.7003
Keywords: biomass; fuelwood; ergonomics; safety
Highlights: Exposure to noise and vibration is higher when chipping branches rather than logs; Noise levels are higher in the separate cab, especially at some frequencies; Vibration is higher for the Z axis in the separate cab, for the X axis in the truck; Noise is higher when working with blunt knives, rather than new knives; Knife wear has no significant effect on exposure to whole-body vibration.
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Industrial chipping is becoming increasingly popular, as the result of a growing demand for woody biomass. Industrial chippers are large, powerful machines that generate much noise and vibration. This study explored some factors that may affect exposure to noise and vibration, namely: feedstock type (branches vs. logs), work station characteristics (truck cab vs. separate cab) and knife wear (new knives vs. blunt knives). Exposure to noise was significantly affected by all three factors, and it was higher for branch feedstock, separate cabs and blunt knives. The higher exposure levels recorded for the separate cab were especially insidious, because they were below and above the hearing threshold and would elude immediate perception. Exposure to whole-body vibration (WBV) was significantly higher for branch feedstock and for the separate cab. Knife wear seemed to determine an increase in WBV, but this effect had no statistical significance and the result could not be taken as conclusive. Among the three factors studied, work station characteristics had the strongest effect. Further studies may extend the comparison to a wider range of options, as well as explore the use of exposure variation for machine diagnostics.

  • Anton, University of Ljubljana, Dept. of Forestry and Renewable Resources, Večna pot 83, 1000 Ljubljana, Slovenia E-mail: anton.poje@bf.uni-lj.si
  • Spinelli, CNR IVALSA, Via Madonna del Piano 10, I-50019 Sesto Fiorentino, Italy; AFORA, University of the Sunshine Coast, Locked Bag 4, Maroochydore DC, Queensland, 4558 Australia ORCID http://orcid.org/0000-0001-9545-1004 E-mail: spinelli@ivalsa.cnr.it (email)
  • Magagnotti, CNR IVALSA, Via Madonna del Piano 10, I-50019 Sesto Fiorentino, Italy; AFORA, University of the Sunshine Coast, Locked Bag 4, Maroochydore DC, Queensland, 4558 Australia E-mail: magagnotti@ivalsa.cnr.it
  • Mihelic, University of Ljubljana, Dept. of Forestry and Renewable Resources, Večna pot 83, 1000 Ljubljana, Slovenia E-mail: matevz.mihelic@bf.uni-lj.si
article id 6999, category Research article
Virginia Morales Olmos, Jacek P. Siry. (2018). The Law of One Price in global coniferous sawlog markets. Silva Fennica vol. 52 no. 1 article id 6999. https://doi.org/10.14214/sf.6999
Keywords: cointegration; roundwood prices; international trade
Highlights: The Law of One Price did not hold for North American, European and South American sawlog markets between 1995–2012; There are long-term relationships between some of the analyzed sawlog prices in these markets; The Law of One Price may hold between sawlogs traded in Sweden and Norway, Norway and Finland, and Canada West and US Northwest.
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With progressing globalization of forest production, roundwood prices in different countries may follow similar trends. The Law of One Price (LOP) postulates that the price of a similar product should be the same in different markets when expressed in the same currency. The objectives of this research were (1) to test the LOP in selected coniferous sawlog markets, and (2) to analyze whether a common market – the European Union – leads to the existence of a single sawlog market. The analysis included Brazil, Chile, Finland, Germany, Norway, Poland, Russia Northwest, Sweden, the US South, the US Northwest, Canada East, and Canada West. The results suggest that some of the coniferous sawlog markets were cointegrated which means that they shared a long-term relationship even if in the short-term they do not necessarily adjust to each other. The LOP may hold between coniferous sawlog markets in Sweden and Norway from 1995 through 2012 when sawlog prices were expressed in USD, and in Norway and Finland for 2001–2012 for prices in EUR. Furthermore, the LOP may hold for North American markets in the West for 2004–2012.

  • Morales Olmos, Universidad de la República-Sede Tacuarembó, Ruta 5. km, 386.200, Tacuarembó, CP 45000, Uruguay E-mail: vmolmos@gmail.com (email)
  • Siry, Warnell School of Forestry and Natural Resources, University of Georgia, 180 E. Green Street, Athens, GA 30602-2152, USA E-mail: jsiry@uga.edu
article id 1665, category Research article
Lauri Haataja, Ville Kankaanhuhta, Timo Saksa. (2018). Reliability of self-control method in the management of non-industrial private forests. Silva Fennica vol. 52 no. 1 article id 1665. https://doi.org/10.14214/sf.1665
Keywords: forest management; soil preparation; quality control; young stand management; forest regeneration; boreal silviculture; forest planting
Highlights: Self-control method was found reliable at the main stages of the forest regeneration process; Only slight overestimation was found in self-control results of soil preparation and planting and small underestimation in self-control of young stand management; Diverse utilizing of self-control data is possible in support of service providers operations.
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This study seeks to determine the extent to which self-control data can be relied upon in the management of private forests. Self-control (SC) requires the forest workers to evaluate their own work quality to ensure the clients’ needs are met in terms of soil preparation, planting and young stand management. Self-control data were compared to an independent evaluation of the same worksites. Each dataset had a hierarchical structure (e.g., sample plot, regeneration area and contractor), and key quality indicators (i.e., number of prepared mounds, planted seedlings or crop trees) were measured for each plot.  Self-control and independent-assessments (IA) were analyzed by fitting a multi-level multivariate model containing explanatory variables. No significant differences were observed in terms of soil preparation (number of mounds) or young stand management (number of crop trees) between self-control and independent-assessments. However, the self-control planting data included a slight but significant overestimation of the number of planted seedlings. Discrepancies are discussed in terms of sampling error and other explanatory factors. According to overall results, self-control methods are reliable at every stage of the forest regeneration process. As such, the diverse utilizing of self-control data is possible in support of service providers operations.
  • Haataja, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: lauritapiohaataja@gmail.com (email)
  • Kankaanhuhta, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: ville.kankaanhuhta@luke.fi
  • Saksa, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@luke.fi
article id 7783, category Research article
Markku T. Lehtinen, Pertti Pulkkinen. (2017). Effects of Scots pine paternal genotypes of two contiguous seed orchards on the budset and frost hardening of first-year progeny. Silva Fennica vol. 51 no. 5 article id 7783. https://doi.org/10.14214/sf.7783
Keywords: Pinus sylvestris; provenance; conifer; environmental influence; pollen; genotype effect
Highlights: This environmentally controlled study on Scots pine demonstrated the effect of the paternal genotype on the budset and frost hardening of the progeny; With the applied study design, no significant indication of an environmental influence on the effect of the Scots pine paternal genotype was obtained.
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In Scots pine (Pinus sylvestris L.), it has been shown that the parental conditions have a role in the phenological variation among first-year seedlings. For this reason, it is argued that they should be comprehensively controlled before estimating the parental genotype effects. This controlled-cross study examined the effects of a set of fathers of Scots pines on the timing of budset and autumn frost hardening of first-year seedlings. The paternal genotypes had either a northern or southern provenance, but had spent a period of over 25 years as grafts in a shared climatic environment in two closely located southern orchards. Pollen applied in the crosses was collected from these orchards in one year and all the maternal genotypes were pollinated in only one seed orchard. The results of freeze tests and budset observations of the consequent progeny were analysed and additionally compared with results obtained using seedlings from seed lots of natural forests in order to estimate the ability of northern paternal genotypes to maintain a northern effect under southern conditions. This environmentally controlled study demonstrated a significant effect of the paternal genotype on the budset and autumn frost hardening of first-year seedling of Scots pine. With the applied study design, no significant indication of an environmental influence on the effect of the paternal genotype was obtained. The accuracy of the observations is discussed. It is concluded that the results suggest a minor role of mutability in the effects of Scots pine paternal genotypes.

  • Lehtinen, University of Helsinki, Department of Agricultural Sciences, Latokartanonkaari 5 and 7, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: markku.t.lehtinen@helsinki.fi (email)
  • Pulkkinen, Natural Resources Institute Finland (Luke), Green Technology, Haapastensyrjäntie 34, FI-12600 Läyliäinen, Finland E-mail: pertti.pulkkinen@luke.fi
article id 7759, category Research article
Jaana Luoranen, Sirkka Sutinen. (2017). Reduced height of short day induced bud scale complex may partly explain early bud burst in Norway spruce seedlings. Silva Fennica vol. 51 no. 5 article id 7759. https://doi.org/10.14214/sf.7759
Keywords: Picea abies; elongation; bud scale complex; primordia; shape
Highlights: Short day treatment used in tree seedling nurseries affects the structure of apical buds; Changes in bud structure may partly explain early bud burst and may be a reason for unburst buds of short day treated seedlings.
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Short day (SD) treatment is used as a dormancy induction in forest tree seedling nurseries in the boreal forest zone. However, SD treatment has caused early bud burst in the following spring, which may expose the seedlings to spring frosts. Because the mechanisms affecting earlier bud burst in SD treated seedlings are not fully understood yet, here we have studied the effect of SD treatment on the structure of buds in Norway spruce [Picea abies (L.) Karst.] seedlings. Seedlings were exposed to SD treatments or natural (CTRL) light and photoperiod in July in a nursery in Central Finland. The experiments included two lots of seedlings over two summers and the analyses were done under a stereo microscope. SD treatment advanced initiation of bud scales and formation of needle primordia, and thus the formation period was shorter in CTRL seedlings. In mature buds, no differences in primordial shoots were found between the treatments, whereas notable differences were found in bud scales. The SD buds had fewer and shorter bud scales than the CTRL buds. This led to significantly shorter bud scale complex and, consequently, to shorter buds in SD than in CTRL seedlings. Buds and needles matured earlier in SD treated seedlings. In the following spring, the primordial shoots started to elongate in both treatments around mid-May, when the SD buds started to break down, whereas CTRL buds started to break down in late May. The fewer number and shorter height of protective bud scales may expose buds to harsh winter temperatures and early loss of scales may predispose the SD buds to spring frosts.

  • Luoranen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: jaana.luoranen@luke.fi (email)
  • Sutinen, E-mail: sirusuti@gmail.com
article id 7751, category Research article
Göran Nordlander, Euan G. Mason, Karin Hjelm, Henrik Nordenhem, Claes Hellqvist. (2017). Influence of climate and forest management on damage risk by the pine weevil Hylobius abietis in northern Sweden. Silva Fennica vol. 51 no. 5 article id 7751. https://doi.org/10.14214/sf.7751
Keywords: temperature sum; reforestation; soil scarification; clear-cut age; conifer seedling; damage prediction; warmer climate
Highlights: Analysis of survey data from 292 reforestation areas in northern Sweden show that the probability of pine weevil damage can be predicted with a standard error of 0.12; Three variables are important in the optimal model: proportion of seedlings in mineral soil, age of clear-cut, and temperature sum; Temperature sum in the model can be adjusted to reflect future climate scenarios.
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The pine weevil Hylobius abietis L. is an economically important pest insect that kills high proportions of conifer seedlings in reforestation areas. It is present in conifer forests all over Europe but weevil abundance and risk for damage varies considerably between areas. This study aimed to obtain a useful model for predicting damage risks by analyzing survey data from 292 regular forest plantations in northern Sweden. A model of pine weevil attack was constructed using various site characteristics, including both climatic factors and factors related to forest management activities. The optimal model was rather imprecise but showed that the risk of pine weevil attack can be predicted approximatively with three principal variables: 1) the proportion of seedlings expected to be planted in mineral soil rather than soil covered with duff and debris, 2) age of clear-cut at the time of planting, and 3) calculated temperature sum at the location. The model was constructed using long-run average temperature sums for epoch 2010, and so effects of climate change can be inferred from the model by adjustment to future epochs. Increased damage risks with a warmer climate are strongly indicated by the model. Effects of a warmer climate on the geographical distribution and abundance of the pine weevil are also discussed. The new tool to better estimate the risk of damage should provide a basis for foresters in their choice of countermeasures against pine weevil damage in northern Europe.

  • Nordlander, Swedish University of Agricultural Sciences (SLU), Department of Ecology, P.O. Box 7044, SE-750 07 Uppsala, Sweden E-mail: Goran.Nordlander@slu.se
  • Mason, University of Canterbury, School of Forestry, Private Bag 4800, Christchurch 8140, New Zealand; Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53 Alnarp, Sweden ORCID http://orcid.org/0000-0001-9024-9106 E-mail: euan.mason@canterbury.ac.nz (email)
  • Hjelm, Skogforsk, The Forest Research Institute of Sweden, Ekebo 2250, SE-268 90 Svalöv, Sweden; Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: karin.hjelm@skogforsk.se
  • Nordenhem, Swedish University of Agricultural Sciences (SLU), Department of Ecology, P.O. Box 7044, SE-750 07 Uppsala, Sweden E-mail: h.nordenhem@telia.com
  • Hellqvist, Swedish University of Agricultural Sciences (SLU), Department of Ecology, P.O. Box 7044, SE-750 07 Uppsala, Sweden E-mail: Claes.Hellqvist@slu.se
article id 7748, category Research article
Dominik Bayer, Hans Pretzsch. (2017). Reactions to gap emergence: Norway spruce increases growth while European beech features horizontal space occupation – evidence by repeated 3D TLS measurements. Silva Fennica vol. 51 no. 5 article id 7748. https://doi.org/10.14214/sf.7748
Keywords: Picea abies; gap dynamics; Fagus sylvatica; crown expansion; crown allometry; TLS; growing area competition; growing space efficiency
Highlights: Analysis of the closure dynamics of a Norway spruce, a European beech and a mixed forest gap by repeated TLS measurements; Norway spruce allocated additional resources predominantly into DBH growth and displayed stronger resilience against mechanical crown damage; European beech allocated resources towards space occupation and displayed higher crown plasticity; Species mixture had no significant effect.
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The reach of different tree species’ crowns and the velocity of gap closure during the occupation of canopy gaps resulting from mortality and thinning during stand development determine species-specific competition and productivity within forest stands. However, classical dendrometric methods are rather inaccurate or even incapable of time- and cost-effectively measuring 3D tree structure, crown dynamics and space occupation non-destructively. Therefore, we applied terrestrial laser scanning (TLS) in order to measure the structural dynamics at tree and stand level from gap cutting in 2006 until 2012 in pure and mixed stands of Norway spruce (Picea abies [L.] Karst.) and European beech (Fagus sylvatica L.). In conclusion, our results suggest that Norway spruce invests newly available above-ground resources primarily into DBH as well as biomass growth and indicate a stronger resilience against loss of crown mass induced by mechanical damage. European beech showed a vastly different reaction, investing gains from additional above-ground resources primarily into faster occupation of canopy space. Whether our sample trees were located in pure or mixed groups around the gaps had no significant impact on their behavior during the years after gap cutting.

  • Bayer, Address Technical University of Munich (TUM), Chair for Forest Growth and Yield Science, Hans-Carl-von-Carlowitz-Platz 2, 85354, Freising, Germany ORCID http://orcid.org/0000-0002-2084-3019 E-mail: dominik.bayer@lrz.tu-muenchen.de (email)
  • Pretzsch, Address Technical University of Munich (TUM), Chair for Forest Growth and Yield Science, Hans-Carl-von-Carlowitz-Platz 2, 85354, Freising, Germany E-mail: hans.pretzsch@lrz.tu-muenchen.de
article id 7746, category Research article
Jānis Donis, Māra Kitenberga, Guntars Šņepsts, Roberts Matisons, Juris Zariņš, Āris Jansons. (2017). The forest fire regime in Latvia during 1922–2014. Silva Fennica vol. 51 no. 5 article id 7746. https://doi.org/10.14214/sf.7746
Keywords: forest disturbance; fire regime; hemiboreal zone; spatial analysis
Highlights: Climate effects and human influence on forest fire activity in Latvia was assessed using time-series analysis; Drought conditions during summer season had the strongest effect on fire activity of tested climatic variables; Negative trends and spatial distribution pattern of fire activity suggests of prevailing human influence on forest fire regime over the 20th century.
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Fire as disturbance of forests has an important ecological and economical role in boreal and hemiboreal forests. The occurrence of forest fires is both climatically and anthropogenically determined and shifts in fire regimes are expected due to climate change. Although fire histories have been well documented in boreal regions, there is still insufficient information about fire occurrence in the Baltic States. In this study, spatio-temporal patterns and climatic drivers of forest fires were assessed by means of spatial and time-series analysis. The efficiency of Canadian Fire Weather (FWI) indices as indicators for fire activity was tested. The study was based on data from the literature, archives, and the Latvian State Forest service database. During the period 1922–2014, the occurrence and area affected by forest fires has decreased although the total area of forest land has nearly doubled, suggesting improvement of the fire suppression system as well as changes in socioeconomic situation. The geographical distribution of forest fires revealed two pronounced clusters near the largest cities of Riga and Daugavpils, suggesting dominance of human causes of ignitions. The occurrence of fires was mainly influenced by drought. FWI appeared to be efficient in predicting the fire occurrence: 23–34% of fires occurred on days with a high or extremely high fire danger class, which overall had a relative occurrence of only 4.3–4.6%. During the 20th century, the peak of fire activity shifted from May to April, probably due to global warming and socioeconomic reasons. The results of this study are relevant for forest hazard mitigation and development of fire activity prediction system in Latvia.

  • Donis, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: janis.donis@silava.lv
  • Kitenberga, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: mara.kitenberga@gmail.com (email)
  • Šņepsts, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: guntars.snepsts@silava.lv
  • Matisons, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: robism@inbox.lv
  • Zariņš, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: juris.zarins@silava.lv
  • Jansons, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: aris.jansons@silava.lv
article id 7723, category Research article
Mihails Čugunovs, Eeva-Stiina Tuittila, Ida Sara-Aho, Laura Pekkola, Jari Kouki. (2017). Recovery of boreal forest soil and tree stand characteristics a century after intensive slash-and-burn cultivation. Silva Fennica vol. 51 no. 5 article id 7723. https://doi.org/10.14214/sf.7723
Keywords: restoration; fire disturbance; historical land-use; rewildening
Highlights: Soil organic matter stocks have still not fully recovered after a century of stand succession and passive recovery after slash-and-burn period; Historical slash-and-burn stands feature higher live birch and standing dead wood volume than controls; If passive rewildening is used, Fennoscandian boreal forests need more than a century to regain naturalness.
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Passive rewildening of forest ecosystems is commonly used for rehabilitating degraded habitats closer to their natural origin in addition to costly active restoration measures. However, it is not clear if passive processes are effective and how long the recovery of main ecosystem properties takes. We investigate the recovery of forest soil and tree stand characteristics a century after cessation of slash-and-burn cultivation, a major historical intensive disturbance regime that was applied widely in boreal zone of Finland until late 1800s. We systematically sampled soil and tree stand parameters within former slash-and-burn and nearby control areas. Humus layer thickness and soil organic matter (SOM) stocks were still lower in the historical slash-and-burn than in control areas. Slash-and-burn areas also had a larger volume of live birch trees and a higher standing dead wood volume than control areas. Accordingly, organic matter (humus layer thickness and SOM stocks) correlated negatively with birch standing live tree volume. Combined OM stock in humus and uppermost 10 cm mineral soil layer was positively correlated with lying dead wood volume. Overall, we observed clear recovery of several natural properties but we also found that a century after cessation of frequent anthropogenic burnings, clear legacies of disturbance in the above- and below-ground parts of boreal ecosystem were evident. Our results indicate that if only passive rewildening is applied as a restoration measure, the full recovery of boreal forest is slow and the effects of historical land-use may persist for over hundred years in soil and tree properties.

  • Čugunovs, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: mihails.cugunovs@uef.fi (email)
  • Tuittila, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland ORCID http://orcid.org/0000-0001-8861-3167 E-mail: eeva-stiina.tuittila@uef.fi
  • Sara-Aho, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: ida.sara-aho@mhy.fi
  • Pekkola, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: laura.pekkola@gmail.com
  • Kouki, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland ORCID http://orcid.org/0000-0003-2624-8592 E-mail: jari.kouki@uef.fi
article id 7721, category Research article
Sakari Tuominen, Andras Balazs, Eija Honkavaara, Ilkka Pölönen, Heikki Saari, Teemu Hakala, Niko Viljanen. (2017). Hyperspectral UAV-imagery and photogrammetric canopy height model in estimating forest stand variables. Silva Fennica vol. 51 no. 5 article id 7721. https://doi.org/10.14214/sf.7721
Keywords: forest inventory; digital photogrammetry; aerial imagery; hyperspectral imaging; radiometric calibration; UAVs; stereo-photogrammetric canopy modelling
Highlights: Hyperspectral imagery and photogrammetric 3D point cloud based on RGB imagery were acquired under weather conditions changing from cloudy to sunny; Calibration of hyperspectral imagery was required for compensating the effect of varying weather conditions; The combination of hyperspectral imagery and photogrammetric point cloud data resulted in accurate forest estimates, especially for volumes per tree species.
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Remote sensing using unmanned aerial vehicle (UAV) -borne sensors is currently a highly interesting approach for the estimation of forest characteristics. 3D remote sensing data from airborne laser scanning or digital stereo photogrammetry enable highly accurate estimation of forest variables related to the volume of growing stock and dimension of the trees, whereas recognition of tree species dominance and proportion of different tree species has been a major complication in remote sensing-based estimation of stand variables. In this study the use of UAV-borne hyperspectral imagery was examined in combination with a high-resolution photogrammetric canopy height model in estimating forest variables of 298 sample plots. Data were captured from eleven separate test sites under weather conditions varying from sunny to cloudy and partially cloudy. Both calibrated hyperspectral reflectance images and uncalibrated imagery were tested in combination with a canopy height model based on RGB camera imagery using the k-nearest neighbour estimation method. The results indicate that this data combination allows accurate estimation of stand volume, mean height and diameter: the best relative RMSE values for those variables were 22.7%, 7.4% and 14.7%, respectively. In estimating volume and dimension-related variables, the use of a calibrated image mosaic did not bring significant improvement in the results. In estimating the volumes of individual tree species, the use of calibrated hyperspectral imagery generally brought marked improvement in the estimation accuracy; the best relative RMSE values for the volumes for pine, spruce, larch and broadleaved trees were 34.5%, 57.2%, 45.7% and 42.0%, respectively.

  • Tuominen, Natural Resources Institute Finland (Luke), Economics and society, P.O. Box 2, FI-00791 Helsinki, Finland ORCID http://orcid.org/0000-0001-5429-3433 E-mail: sakari.tuominen@luke.fi (email)
  • Balazs, Natural Resources Institute Finland (Luke), Economics and society, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: andras.balazs@luke.fi
  • Honkavaara, Finnish Geospatial Research Institute, National Land Survey of Finland, Geodeetinrinne 2, FI-02430 Masala, Finland E-mail: eija.honkavaara@nls.fi
  • Pölönen, University of Jyväskylä, Faculty of Information Technology, P.O. Box 35, FI-40014 Jyväskylä, Finland E-mail: ilkka.polonen@jyu.fi
  • Saari, VTT Microelectronics, P.O. Box 1000, FI-02044 VTT, Finland E-mail: heikki.saari@vtt.fi
  • Hakala, Finnish Geospatial Research Institute, National Land Survey of Finland, Geodeetinrinne 2, FI-02430 Masala, Finland E-mail: teemu.hakala@nls.fi
  • Viljanen, Finnish Geospatial Research Institute, National Land Survey of Finland, Geodeetinrinne 2, FI-02430 Masala, Finland E-mail: niko.viljanen@nls.fi
article id 7693, category Research article
Chunyu Zhu, Jiaojun Zhu, Xiao Zheng, Deliang Lu, Xiufen Li. (2017). Comparison of gap formation and distribution pattern induced by wind/snowstorm and flood in a temperate secondary forest ecosystem, Northeast China. Silva Fennica vol. 51 no. 5 article id 7693. https://doi.org/10.14214/sf.7693
Keywords: secondary forests; canopy opening; windstorm; flood; large scale
Highlights: The canopy gaps induced by wind/snowstorm were aggregated in steep slope and high altitude areas, while the gaps formed by flood were gathered in steep slope and low altitude areas; The wind/snowstorm mainly driven the formation of medium gaps, while the flood mainly promoted the percentage of small gaps and vacant lands.
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Canopy gap is the driving force of forest succession. Due to the uncontrollability, however, the influences of natural disturbances on gap formation and gap distribution pattern have been rarely understood in temperate secondary forest ecosystems. We monitored the gap formation and gap distribution pattern using high-resolution remote sensing images before and after two disturbances (wind/snowstorm in 2003 and flood in 2013). The results showed that after wind/snowstorm, the gap nearest neighbor index (GNNI) decreased, the vacant land area did not obviously change while the gap fraction and gaps density (especially medium size) increased. After the flood, GNNI decreased, the number of small gaps increased but larger gaps were in many cases extended to vacant land areas leading to a smaller total number of medium and large gaps but considerable increase in vacant land area. We also found that the gap densities increased with slope and altitude for wind/snowstorm-formed gaps, but they increased with increasing slope and decreasing altitude for flood-formed gaps. These results indicated that gaps were aggregated in steep slope and high altitude areas after wind/snowstorm, but in steep slope and low altitude areas after the flood. Medium gaps were mainly created by the wind/snowstorm due to the individual-level death of dominant tree with the continuous fall of surrounding trees. While, vacant lands were obviously created during the flood because of integral sweeping. Besides, smaller trees were easily damaged by runoff of flood, which induced small gaps. In summary, forest managers may pay more attention to use gaps to accelerate forest succession after wind/snowstorms and to restore vegetation in vacant lands after floods.

  • Zhu, E-mail: Chunyuzhu123@126.com
  • Zhu, E-mail: jiaojunzhu@iae.ac.cn (email)
  • Zheng, E-mail: xiaozheng@iae.ac.cn
  • Lu, E-mail: delianglu14@hotmail.com
  • Li, E-mail: delianglu14@hotmail.com
article id 2018, category Research article
Sima Mohtashami, Lars Eliasson, Gunnar Jansson, Johan Sonesson. (2017). Influence of soil type, cartographic depth-to-water, road reinforcement and traffic intensity on rut formation in logging operations: a survey study in Sweden. Silva Fennica vol. 51 no. 5 article id 2018. https://doi.org/10.14214/sf.2018
Keywords: soil disturbance; forestry; forwarder; decision support tool
Highlights: Soil type and traffic intensity had significant effects on rut formation; Further studies are required to identify all factors affecting rut formation, especially on soils with medium bearing capacity; The cartographic depth-to-water index (DTW) alone did not predict rut formation, but used in combination with other information, e.g. soil type, could be an interesting tool for delineating soil areas that are potentially vulnerable to rut formation in logging operations.
Abstract | Full text in HTML | Full text in PDF | Author Info

Rut formation caused by logging operations has been recognised as a challenge for Swedish forestry. Frequent traffic with heavy machines on extraction roads, together with a warmer climate, is one of the factors that increases the risk of rut formation in forests. One possible way to control this impact of logging operations is to design and apply decision support tools that enable operators to take sensitive areas into account when planning extraction roads. In this study, 16 different logging sites in south-eastern Sweden were surveyed after clear-cut. Information was collected about extraction roads (i.e. traffic intensity, whether the roads had been reinforced with slash) and ruts. Digital maps such as cartographic depth-to-water (DTW) index and soil type were also examined for any connection to rut positions. Soil type and traffic intensity were found to be significant factors in rut formation, while DTW and slash reinforcement were not. However, the DTW map combined with other information, such as soil type, could contribute to decision support tools that improve planning of extraction roads.

  • Mohtashami, The forestry research institute of Sweden, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: sima.mohtashami@skogforsk.se (email)
  • Eliasson, The forestry research institute of Sweden, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID http://orcid.org/0000-0002-2038-9864 E-mail: lars.eliasson@skogforsk.se
  • Jansson, The forestry research institute of Sweden, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID http://orcid.org/0000-0002-3018-9161 E-mail: gunnar.jansson@skogforsk.se
  • Sonesson, The forestry research institute of Sweden, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID http://orcid.org/0000-0002-2018-7496 E-mail: johan.sonesson@skogforsk.se
article id 1740, category Research article
Ram P. Sharma, Zdeněk Vacek, Stanislav Vacek. (2017). Modelling tree crown-to-bole diameter ratio for Norway spruce and European beech. Silva Fennica vol. 51 no. 5 article id 1740. https://doi.org/10.14214/sf.1740
Keywords: Picea abies; Fagus sylvatica; dominant height; exponential decay function; mixed effect model; spatially explicit competition index; species proportion; species mixture effect
Highlights: Modelled crown-to-bole diameter ratio (CDBDR) using tree and stand-level predictors, and sample plot random effects; Spatially explicit mixed-effects model described the largest part of CDBDR variation with no significant trend in the residuals; The CDBDR increased with increasing stand development stage and site quality, but decreased with decreasing proportion of the species of interest, and increasing competition.
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Crown dimensions are correlated to growth of other parts of a tree and often used as predictors in growth models. The crown-to-bole diameter ratio (CDBDR), which is a ratio of maximum crown width to diameter at breast height (DBH), was modelled using data from permanent sample plots located on Norway spruce (Picea abies (L.) Karst.) and European beech (Fagus sylvatica L.) stands in different parts of the Czech Republic. Among various tree and stand-level measures evaluated, DBH, height to crown base (HCB), dominant height (HDOM), basal area of trees larger in diameter than a subject tree (BAL), basal area proportion of the species of interest (BAPOR), and Hegyi’s competition index (CI) were found to be significant predictors in the CDBDR model. Random effects were included using the mixed-effects modelling to describe sample plot-level variation. For each species, the mixed-effects model described a larger part of the variation of the CDBDR than nonlinear ordinary least squares model with no trend in the residuals. The spatially explicit mixed-effects model showed more attractive fit statistics [conditional R2 ≈ 0.73 (spruce), 0.78 (beech)] than its spatially inexplicit counterpart [conditional R2 ≈ 0.71 (spruce), 0.76 (beech)]. The model showed that CDBDR increased with increasing HDOM – a measure that combines the stand development stage and site quality – but decreased with increasing HCB and competition (increasing BAL and CI), and decreasing proportions of the species of interest (increasing BAPOR). For both species, the spatially explicit mixed-effects model should be a preferred choice for a precise prediction of the CDBDR. The CDBDR model will have various management implications such as determination of spacing, stand basal area, stocking, and planning of appropriate species mixture.

  • Sharma, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Kamýcká 129, 16521, Praha 6 – Suchdol, Czech Republic E-mail: sharmar@fld.czu.cz (email)
  • Vacek, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Kamýcká 129, 16521, Praha 6 – Suchdol, Czech Republic E-mail: vacekz@fld.czu.cz
  • Vacek, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Kamýcká 129, 16521, Praha 6 – Suchdol, Czech Republic E-mail: vacekstanislav@fld.czu.cz
article id 7753, category Research article
Aarne Hovi, Pekka Raitio, Miina Rautiainen. (2017). A spectral analysis of 25 boreal tree species. Silva Fennica vol. 51 no. 4 article id 7753. https://doi.org/10.14214/sf.7753
Keywords: albedo; leaf; needle; reflectance; transmittance; integrating sphere; spectrometer
Highlights: An extensive spectral library containing leaf and needle reflectance and transmittance spectra was collected; The spectra openly available in SPECCHIO Spectral Information System; Effects of tree species, leaf/needle side, canopy position, and needle age on spectra were quantified; Seasonal variations were measured for four species; Spectra analysis highlights the importance of shortwave-infrared region in separating tree species.
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Spectral libraries have a fundamental role in the development of interpretation methods for airborne and satellite-borne remote sensing data. This paper presents to-date the largest spectral measurement campaign of boreal tree species. Reflectance and transmittance spectra of over 600 leaf and needle samples from 25 species were measured in the Helsinki area (Finland) using integrating sphere systems attached to an ASD FieldSpec 4 spectroradiometer. Factors influencing the spectra and red edge inflection point (REIP) were quantified using one-way analysis of variance. Tree species differed most in the shortwave-infrared (1500–2500 nm) and least in the visible (400–700 nm) wavelength region. Species belonging to same genera showed similar spectral characteristics. Upper (adaxial) and lower (abaxial) leaf sides differed most in the visible region. Canopy position (sunlit/shaded) had a minor role in explaining spectral variation. For evergreen conifers, current and previous year needles differed in their spectra, current-year needles resembling those of broadleaved and deciduous conifers. Two broadleaved species were monitored throughout the growing season (May–October), and two conifers were measured twice during summer (June, September). Rapid changes were observed in the spectra in early spring and late autumn, whereas seasonal variations during summer months were relatively small for both broadleaved and coniferous species. Based on our results, shortwave-infrared seems promising in separating tree species, although it is to-date least studied. The spectral library reported here (Version 1.0) is publicly available through the SPECCHIO Spectral Information System.

  • Hovi, Aalto University, Department of Built Environment, P.O. Box 14100, FI-00076 Aalto, Finland E-mail: aarne.hovi@aalto.fi (email)
  • Raitio, Aalto University, Department of Built Environment, P.O. Box 14100, FI-00076 Aalto, Finland E-mail: pekka.raitio@aalto.fi
  • Rautiainen, Aalto University, Department of Built Environment, P.O. Box 14100, FI-00076 Aalto, Finland; Aalto University, Department of Electronics and Nanoengineering, P.O. Box 15500, FI-00076 Aalto, Finland E-mail: miina.a.rautiainen@aalto.fi
article id 7743, category Research article
Sakari Tuominen, Timo Pitkänen, Andras Balazs, Annika Kangas. (2017). Improving Finnish Multi-Source National Forest Inventory by 3D aerial imaging. Silva Fennica vol. 51 no. 4 article id 7743. https://doi.org/10.14214/sf.7743
Keywords: forest inventory; remote sensing; spatial autocorrelation; spatial distribution; aerial imagery; stereo-photogrammetry
Highlights: 3D aerial imaging provides a feasible method for estimating forest variables in the form of thematic maps in large area inventories; Photogrammetric 3D data based on aerial imagery that was originally acquired for orthomosaic production was tested in estimating stand variables; Photogrammetric 3D data highly improved the accuracy of forest estimates compared to traditional 2D remote sensing imagery.
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Optical 2D remote sensing techniques such as aerial photographing and satellite imaging have been used in forest inventory for a long time. During the last 15 years, airborne laser scanning (ALS) has been adopted in many countries for the estimation of forest attributes at stand and sub-stand levels. Compared to optical remote sensing data sources, ALS data are particularly well-suited for the estimation of forest attributes related to the physical dimensions of trees due to its 3D information. Similar to ALS, it is possible to derive a 3D forest canopy model based on aerial imagery using digital aerial photogrammetry. In this study, we compared the accuracy and spatial characteristics of 2D satellite and aerial imagery as well as 3D ALS and photogrammetric remote sensing data in the estimation of forest inventory variables using k-NN imputation and 2469 National Forest Inventory (NFI) sample plots in a study area covering approximately 5800 km2. Both 2D data were very close to each other in terms of accuracy, as were both the 3D materials. On the other hand, the difference between the 2D and 3D materials was very clear. The 3D data produce a map where the hotspots of volume, for instance, are much clearer than with 2D remote sensing imagery. The spatial correlation in the map produced with 2D data shows a lower short-range correlation, but the correlations approach the same level after 200 meters. The difference may be of importance, for instance, when analyzing the efficiency of different sampling designs and when estimating harvesting potential.

  • Tuominen, Natural Resources Institute Finland (Luke), Economics and Society, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: sakari.tuominen@luke.fi (email)
  • Pitkänen, Natural Resources Institute Finland (Luke), Economics and Society, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: timo.p.pitkanen@luke.fi
  • Balazs, Natural Resources Institute Finland (Luke), Economics and Society, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: andras.balazs@luke.fi
  • Kangas, Natural Resources Institute Finland (Luke), Economics and Society, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: Annika.Kangas@luke.fi
article id 7731, category Research article
Anneli Viherä-Aarnio, Pirkko Velling. (2017). Growth, wood density and bark thickness of silver birch originating from the Baltic countries and Finland in two Finnish provenance trials. Silva Fennica vol. 51 no. 4 article id 7731. https://doi.org/10.14214/sf.7731
Keywords: Betula pendula; provenance; bark thickness; frost crack; seed transfer; wood basic density; wood discolouration
Highlights: Baltic origins of silver birch had significantly thicker bark than the Finnish ones; In terms of wood density, no consistent difference was detected between the Baltic and Finnish origins; Incidence of darkened core wood increased with increasing seed origin latitude; Frost cracks were most common in south Latvian origins grown in central Finland.
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Silver birch (Betula pendula Roth) seed origins from the Baltic countries and from Finland were compared in terms of growth, wood density, bark thickness and the incidence of darkened core wood, frost cracks and decay, and the effect of seed origin latitude was examined in two Finnish provenance trials. The material consisted of 21 stand and single tree origins ranging from latitudes 54° to 63°N from the Baltic countries and Finland. The trials, measured at the age of 22 years, were located at Tuusula (60°21´N), southern Finland and at Viitasaari (63°11´N), central Finland. The Baltic origins were superior to the Finnish ones in the southern trial regarding height, whereas in central Finland the Finnish origins grew better. There was no consistent difference between the Baltic and the Finnish group of origins in wood density. Bark thickness decreased with increasing seed origin latitude. The Baltic origins had significantly thicker bark than the Finnish origins. A moderate positive correlation was detected between the seed origin latitude and the incidence of darkened core wood in the southern trial, where the darkened core wood was more common in the Finnish origins than in the Baltic ones. The highest proportion of trees with frost cracks was detected in the south-western Latvian origins growing in central Finland. Seed transfers from the Baltic would have an increasing effect on the bark thickness of birch logs, but no or only minor effects on wood density. Based on our results, there is no reason to recommend the use of non-native Baltic seed origins in Finland instead of the native locally adapted seed sources.

  • Viherä-Aarnio, Natural Resources Institute Finland (Luke), Green technology, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: anneli.vihera-aarnio@luke.fi (email)
  • Velling, E-mail: pike.velling@phnet.fi
article id 7728, category Research article
Liam Donnelly, Sven-Olof Lundqvist, Conor O’Reilly. (2017). Inter- and intra-annual wood property variation in juvenile wood between six Sitka spruce clones. Silva Fennica vol. 51 no. 4 article id 7728. https://doi.org/10.14214/sf.7728
Keywords: productivity; wood properties; Picea sitchensis; clonal forestry; selective breeding; stem height
Highlights: Wood property differences resulted primarily from variation in the proportions of early- and latewood in each annual ring; Width of early- and latewood bands in each ring was found to be a more important determinant of juvenile wood quality than the characteristics of the cells within each band; Wood properties differed greatly between clones, suggesting that there is potential to improve juvenile wood properties through selective breeding.
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Increased growth rates have reduced rotation lengths, increasing the proportion of juvenile wood relative to mature wood, which may negatively affect mechanical performance of sawn timber. However, there is limited information available on the potential impact of breeding for vigour on juvenile wood in Sitka spruce (Picea sitchensis (Bong.) Carrière). In this study, the relationship between vigour (based on total height) and wood properties was investigated in six-year-old Sitka spruce clones grown in two replicated field trials in Ireland. Six clones were evaluated, two clones from each of three vigour (high, intermediate and low) classes. Discs were cut from the base of one ramet per replication for each clone to assess wood quality attributes. Radial tracheid width was significantly and positively correlated with ring width and height, and was negatively correlated with density. The wood of the most vigorous clone had significantly larger ring width with thinner cell walls and wider tracheids than all clones in the two other vigour classes, resulting in lower mean wood density. Latewood properties for all wood attributes measured differed significantly between the two sites. Wood property differences resulted primarily from variation in the proportions of early- and latewood in each annual ring. Additionally, the width of early- and latewood bands in each ring was found to be a more important determinant of juvenile wood quality than the characteristics of the cells within each band. Wood properties differed greatly between clones, suggesting that there is potential to improve juvenile wood properties through selective breeding.

  • Donnelly, UCD Forestry, University College Dublin, Belfield, Dublin 4, Ireland E-mail: liam.donnelly@ucdconnect.ie (email)
  • Lundqvist, Innventia Ab, Drottning Kristinas väg 61, SE-114 86 Stockholm, Sweden E-mail: svenolof.lundqvist@innventia.com
  • O’Reilly, UCD Forestry, University College Dublin, Belfield, Dublin 4, Ireland E-mail: conor.oreilly@ucd.ie
article id 6991, category Research article
Isabel León, Juan José García, Manuel Fernández, Javier Vázquez-Piqué, Raúl Tapias. (2017). Differences in root growth of Quercus ilex and Quercus suber seedlings infected with Phytophthora cinnamomi. Silva Fennica vol. 51 no. 4 article id 6991. https://doi.org/10.14214/sf.6991
Keywords: root rot; holm oak; cork oak; infection; inoculum level; oaks decline
Highlights: Root growth of two Quercus sp. differs significantly after infection with Phytophthora cinnamomi; We observed a marked decrease in the growth of new roots in Quercus ilex with increasing inoculum level; Roots were longer but thinner with a moderate inoculum level in Quercus suber.
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In the southwest of the Iberian Peninsula, Phytophthora cinnamomi Rands is causing irreversible damage to populations of the two most common species of Quercus, the holm oak (Quercus ilex L.) and the cork oak (Quercus suber L.). Although the symptoms are similar in the two species, the mortality rates are different. We found significant differences in the post-infection growth of the root system as a function of tree species, as well as initial plant size, and inoculum level. We observed a marked decrease in the growth of new roots in Q. ilex with increasing inoculum level, while in Q. suber, we found longer but thinner roots with a moderate inoculum level. In both species, we observed a worsening in the water status of the plants from the lowest inoculum level.

  • León, University of Huelva, Agroforestry department, Calle Dr. Cantero Cuadrado, 6, 21004 Huelva, Spain E-mail: isabel.leon@dcaf.uhu.es
  • García, University of Huelva, Agroforestry department, Calle Dr. Cantero Cuadrado, 6, 21004 Huelva, Spain E-mail: juanjose.garcia@dcaf.uhu.es
  • Fernández, University of Huelva, Agroforestry department, Calle Dr. Cantero Cuadrado, 6, 21004 Huelva, Spain E-mail: nonoe@uhu.es
  • Vázquez-Piqué, University of Huelva, Agroforestry department, Calle Dr. Cantero Cuadrado, 6, 21004 Huelva, Spain E-mail: jpique@dcaf.uhu.es
  • Tapias, University of Huelva, Agroforestry department, Calle Dr. Cantero Cuadrado, 6, 21004 Huelva, Spain E-mail: rtapias@uhu.es (email)
article id 6989, category Research article
Åsa Gustafsson. (2017). Assessing work order information quality in harvesting. Silva Fennica vol. 51 no. 4 article id 6989. https://doi.org/10.14214/sf.6989
Keywords: survey study; logging contractor; work orders; work order information components; factor analysis
Highlights: This paper concludes that information quality with regards to accuracy in work order information components pertaining to “Landing – placement and size”, “Cleaning understory trees – not conducted and of low standard”, are lacking and need to be improved in harvesting.
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Being a logging contractor involves several uncertainties, amongst others, information quality in the work order received from customers. The information quality of work orders is of the utmost importance for logging contactors, in order to be able to plan and conduct work properly. The purpose of this paper is three-fold: 1) identifying work order information components in harvesting, 2) identifying work order information quality dimensions in harvesting and 3) assessing work order information quality in harvesting. The paper is based on interviews and a survey. Various interviews took place in Sweden with professionals within the harvesting industry as well as logging contractors, and thereafter a survey was developed. Random selection was conducted and 100 Swedish logging contractors were contacted by telephone in order to answer the survey, with a response rate of 82% from the sample. The paper concludes that the information quality dimension of accuracy concerns the individual work order information components, whereas timeliness is related to receiving the complete work orders. A factor analysis has been conducted with five factors emerging. The assessment of work order information quality in harvesting implies that the potential for improvement exists with regard to increasing the accuracy of the order information for the components of “Cleaning under story trees – not conducted” and “Cleaning under story trees – of low standard” as well as “Landing – size”, and “Landing – placement”. However, their effect on capacity is utilization needs to be explored.

  • Gustafsson, Linnaeus University, School of Business and Economics, Department of Management Accounting and Logistics, 391 82 Kalmar, Sweden E-mail: asa.gustafsson@lnu.se (email)
article id 6977, category Research article
Anett Schibalski, Aleksi Lehtonen, Thomas Hickler, Boris Schröder. (2017). Identifying important topics for model refinement in a widely used process-based model informed by correlative model analyses in a boreal forest. Silva Fennica vol. 51 no. 4 article id 6977. https://doi.org/10.14214/sf.6977
Keywords: Finland; treeline; competition; LPJ-GUESS; statistical modelling; forest ecology
Highlights: Continental-scale model parameterization of widely used LPJ-GUESS experiences problems when applied on the regional level; Competition, disturbances and soil conditions are crucial for explaining treeline position in Finland, besides climatic limitation; Picea abies is overly dominant in LPJ-GUESS model, as key competitive mechanisms are not implemented in sufficient detail.
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Models attempting to predict treeline shifts in changing climates must include the relevant ecological processes in sufficient detail. A previous correlative model study has pointed to nutrients, competition, and temperature as the most important factors shaping the treelines of Pinus sylvestris L., Picea abies (L.) H. Karst. and Betula pubescens Ehrh. in Finnish Lapland. Here, we applied a widely used process-based dynamic vegetation model (LPJ-GUESS) to (i) test its capability to simulate observed spatial and temporal patterns of the main tree species in Finnish Lapland, and (ii) to explore the model representation of important processes in order to guide further model development. A European parameterization of LPJ-GUESS overestimated especially P. abies biomass and the species’ northern range limit. We identified implemented processes to adjust (competition, disturbance) and crucial processes in boreal forests to include (nutrient limitation, forest management) which account for the model’s failure to (edaphically) restrict P. abies in Finnish Lapland and the resulting species imbalance. Key competitive mechanisms are shade and drought tolerance, nutrient limitation, fire resistance, and susceptibility to disturbances (storm, herbivory) which we discussed with respect to boreal ecology and promising model developments to provide a starting point for future model development.

  • Schibalski, Institute of Geoecology, Technische Universität Braunschweig, Langer Kamp 19c, D-38106 Braunschweig, Germany E-mail: a.schibalski@tu-braunschweig.de (email)
  • Lehtonen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: aleksi.lehtonen@luke.fi
  • Hickler, Biodiversity and Climate Research Centre (BiK-F), Senckenberganlage 25, D-60325 Frankfurt am Main, Germany; Department of Physical Geography, Goethe University, Altenhöferallee 1, D-60438 Frankfurt am Main, Germany E-mail: thomas.hickler@senckenberg.de
  • Schröder, Institute of Geoecology, Technische Universität Braunschweig, Langer Kamp 19c, D-38106 Braunschweig, Germany; Berlin-Brandenburg Institute of Advanced Biodiversity Research BBIB, Altensteinstr. 6, D-14195 Berlin, Germany E-mail: boris.schroeder@tu-bs.de
article id 5659, category Research article
Juha Laitila, Anssi Ahtikoski, Jaakko Repola, Johanna Routa. (2017). Pre-feasibility study of supply systems based on artificial drying of delimbed stem forest chips. Silva Fennica vol. 51 no. 4 article id 5659. https://doi.org/10.14214/sf.5659
Keywords: moisture content; heating value; procurement cost; young forests; dry matter loss
Highlights: With artificial drying and quick delivery, avoiding dry material losses of harvested timber, it could be possible to reduce the current costs of the prevailing procurement system based on natural drying of stored timber at roadside landings; The maximum cost for the prospective drying process of fresh chips corresponds to, e.g., organization costs or stumpage price of delimbed stems.
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This study was aimed at determining the maximum cost level of artificial drying required for cost-efficient operation. This was done using a system analysis approach, in which the harvesting potential and procurement cost of alternative fuel chip production systems were compared at the stand and regional level. The accumulation and procurement cost of chipped delimbed stems from young forests were estimated within a 100 km transport distance from a hypothetical end use facility located in northern Finland. Logging and transportation costs, stumpage prices, tied up capital, dry matter losses and moisture content of harvested timber were considered in the study. Moisture content of artificially dried fuel chips made of fresh timber (55%) was set to 20%, 30% and 40% in the comparisons. Moisture content of fuel chips based on natural drying during storing was 40%. Transporting costs were calculated according to new higher permissible dimensions and weight limits for truck-trailers. The procurement cost calculations indicated that with artificial drying and by avoiding dry material losses of timber, it could be possible to reduce current costs of the prevailing procurement system based on natural drying of timber at roadside landings. The maximum cost level of artificial drying ranged between 1.2–3.2 € MWh–1 depending on the supply chain, moisture content and procurement volume of fuel chips. This cost margin corresponds to, e.g., organization, forwarding and transportation costs or stumpage price of delimbed stems.

  • Laitila, Natural Resources Institute Finland (Luke), Bio-based business and industry, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: juha.laitila@luke.fi (email)
  • Ahtikoski, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Paavo Havaksen tie 3, FI-90570 Oulu, Finland E-mail: anssi.ahtikoski@luke.fi
  • Repola, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: jaakko.repola@luke.fi
  • Routa, Natural Resources Institute Finland (Luke), Bio-based business and industry, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: johanna.routa@luke.fi
article id 1760, category Research article
Marek Fajstavr, Kyriaki Giagli, Hanuš Vavrčík, Vladimír Gryc, Josef Urban. (2017). The effect of stem girdling on xylem and phloem formation in Scots pine. Silva Fennica vol. 51 no. 4 article id 1760. https://doi.org/10.14214/sf.1760
Keywords: Pinus sylvestris; radial increment; cambial activity; cell differentiation; phloemogenesis; xylogenesis
Highlights: Stem girdling ceased the cambial activity, below the girdled area, immediately after the removal of the bark strip; Pinus sylvestris survived for up to two years after stem girdling; The girdled trees formed phloem cells above the girdled area but failed to form latewood cells in the next growing season.
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Scots pine (Pinus sylvestris L.) is a resilient, wide spread species. This paper reports on the xylem and phloem cell formation process, before and after, the species was put under artificial stress by stem girdling. Microcore method was applied to a healthy control group and a standing group of girdled trees within an 80-year-old pine forest for two consecutive growing seasons (2013 and 2014). The stem girdling was applied in the middle of the first growing season (July 2013). Cambial activity timings (onset and cessation of cell division), cell formation intensity, cell differentiation, and the dynamics of the annual radial increment in the stem were analyzed. Cambial activity was inhibited and eventually ceased below the stem girdling immediately after the removal of the strip. Therefore, no latewood tracheids were formed. However, above the stem girdling and in the control trees, cell formation and tissue differentiation continued until the end of the growing season, with the girdled trees moving at a less intensive pace but for a longer period of time. During the following growing season (2014), the cambial zone was reactivated only above the stem girdling, not below, and eventually the girdled trees died. In 2014, the onset of the cambial activity was delayed and the division rate of the cells was slower in the girdled trees. Furthermore, the girdled trees formed less phloem cells than the control trees.

  • Fajstavr, Department of Wood Science, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemedelska 3, 613 00 Brno, Czech Republic E-mail: fajstavr.marek@seznam.cz (email)
  • Giagli, Department of Wood Science, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemedelska 3, 613 00 Brno, Czech Republic E-mail: kyriaki.giagli@mendelu.cz
  • Vavrčík, Department of Wood Science, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemedelska 3, 613 00 Brno, Czech Republic E-mail: vavrcik@mendelu.cz
  • Gryc, Department of Wood Science, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemedelska 3, 613 00 Brno, Czech Republic E-mail: gryc@mendelu.cz
  • Urban, Department of Forest Botany, Dendrology and Geobiocenology, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemedelska 3, 613 00 Brno, Czech Republic;  Siberian Federal University, Svobodnyj Prospect 79, Krasnoyarsk, 660041 Krasnoyarsk, Russia E-mail: josef.urban@email.cz
article id 1691, category Research article
Enéas Ricardo Konzen, Raquel Peron, Márcio Akira Ito, Gilvano Ebling Brondani, Siu Mui Tsai. (2017). Molecular identification of bamboo genera and species based on RAPD-RFLP markers. Silva Fennica vol. 51 no. 4 article id 1691. https://doi.org/10.14214/sf.1691
Keywords: molecular markers; molecular screening; restriction enzymes; genetic divergence; Dendrocalamus; Bambusa
Highlights: We investigated the potential of RAPD-RFLP (Random Amplified Polymorphic DNA, Restriction Fragment Length Polymorphism) in detecting genetic relationships among bamboo genera and species; RFLP profiles resolved in acrylamide gels revealed high number of markers, which accurately differentiated species and genera, based on cophenetic correlation coefficients; We recommend RAPD-RFLP for analyses of genetic diversity and divergence among bamboo genera, species and varieties.
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Bamboo species have a very significant ecological and economic impact. Determining morphological and genetic differences among bamboo genera and species are crucial to explore desirable traits for breeding purposes. Several advances have been made in the taxonomy of bamboos by using molecular fingerprinting tools and next generation sequencing technologies. Nevertheless, classical molecular markers such as RAPD (Random Amplified Polymorphic DNA), AFLP (Amplified Fragment Length Polymorphism) and ISSR (Inter Simple Sequence Repeats) also provide an accurate discrimination among genera and species. Moreover, the RAPD-RFLP (Random Amplified Polymorphic DNA, Restriction Fragment Length Polymorphism) method, in which amplification products from RAPD are digested with restriction enzymes, is a reliable, fast and cost-effective method for fingerprinting. RAPD-RFLP has been scarcely used in the literature and no report regarding bamboo taxonomy is available with this method. Here we explored the molecular (RAPD, RAPD-RFLP) variation among genera (Bambusa, Dendrocalamus, Guadua and Phyllostachys) and species of bamboo cultivated in Brazil. Both molecular markers allowed clear distinction among the genera studied. Moreover, high cophenetic correlation values in UPGMA clusters indicated their potential for discriminating bamboo species. The digestion of RAPD products (RFLP) resulted in high number of polymorphic bands and produced very characteristic profiles for each genus with three enzyme combinations (HindIII/HaeIII, HinfI/RsaI, and single digestion with MspI). We recommend RAPD-RFLP as a reproducible and informative method for screening differences among genera, species and varieties of bamboos. Providing a cost-effective and accurate method for species identification and characterization is straightforward for bamboo conservation, management and breeding.

  • Konzen, Cell and Molecular Biology Laboratory, Center for Nuclear Energy in Agriculture (CENA), University of Sao Paulo (USP), Centenário Av., 303, Piracicaba, SP, P.O. Box 96, Brazil; Forest Sciences, Federal University of Lavras (UFLA), Lavras, MG, P.O. Box 3037, Brazil E-mail: erkonzen@gmail.com (email)
  • Peron, Cell and Molecular Biology Laboratory, Center for Nuclear Energy in Agriculture (CENA), University of Sao Paulo (USP), Centenário Av., 303, Piracicaba, SP, P.O. Box 96, Brazil E-mail: rperon@purdue.edu
  • Ito, Embrapa Western Agriculture, Brazilian Agricultural Research Corporation (EMBRAPA), BR 163 Rd., km 253, Dourados, MS, P.O. Box 449, Brazil E-mail: marcio.ito@embrapa.br
  • Brondani, Forest Sciences, Federal University of Lavras (UFLA), Lavras, MG, P.O. Box 3037, Brazil E-mail: gebrondani@gmail.com
  • Tsai, Cell and Molecular Biology Laboratory, Center for Nuclear Energy in Agriculture (CENA), University of Sao Paulo (USP), Centenário Av., 303, Piracicaba, SP, P.O. Box 96, Brazil E-mail: tsai@cena.usp.br
article id 1683, category Research article
Karol Przeździecki, Jarosław Zawadzki, Chris Cieszewski, Pete Bettinger. (2017). Estimation of soil moisture across broad landscapes of Georgia and South Carolina using the triangle method applied to MODIS satellite imagery. Silva Fennica vol. 51 no. 4 article id 1683. https://doi.org/10.14214/sf.1683
Keywords: soil moisture; remotely sensed imagery; satellite observations of forests; triangle method; TVDI; MODIS
Highlights: Temperature vegetation dryness indices were calculated from MODIS satellite imagery to estimate subsurface soil moisture at different depths using the triangle method; Observations were carried out over the vast areas of Georgia and South Carolina, USA, covered with diverse land uses that, included dense forests and agricultural areas; The triangle method may be useful in forestry management applications where the productivity potential of a region and the hydrologic role of forests in that region are of concern.
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We describe here a study based on analysis of vegetation indices and land surface temperatures, which provides relevant information for estimating soil moisture at regional scales. Through an analysis of MODIS satellite imagery and in situ moisture data, the triangle method was used to develop a conceptual land surface temperature−vegetation index model, and spatial temperature-vegetation dryness index (TVDI) values to describe soil moisture relationships for a broad landscape. This study was situated mainly within two states of the southern United States (Georgia and South Carolina). The total study area was about 30 million hectares. The analyses were conducted using information gathered from the 2009 growing season (from the end of March to September). The results of the study showed that soil moisture content was inversely proportional to TVDI, and that TVDI based on the normalized difference vegetation index (NDVI) had a slightly higher correlation with soil moisture than TVDI based on the enhanced vegetation index (EVI).

  • Przeździecki, Warsaw University of Technology, Faculty of Building Services, Hydro and Environmental Engineering, 00-653, Nowowiejska 20, Warszawa, Poland ORCID http://orcid.org/0000-0002-2275-5223 E-mail: karol_przezdziecki@is.pw.edu.pl (email)
  • Zawadzki, Warsaw University of Technology, Faculty of Building Services, Hydro and Environmental Engineering, 00-653, Nowowiejska 20, Warszawa, Poland ORCID http://orcid.org/0000-0003-2842-0018 E-mail: j.j.zawadzki@gmail.com
  • Cieszewski, University of Georgia, Warnell School of Forestry and Natural Resources, 180 E Green St, Athens, GA 30602, USA ORCID http://orcid.org/0000-0003-2842-4406 E-mail: thebiomat@gmail.com
  • Bettinger, University of Georgia, Warnell School of Forestry and Natural Resources, 180 E Green St, Athens, GA 30602, USA ORCID http://orcid.org/0000-0002-5454-3970 E-mail: pbettinger@warnell.uga.edu
article id 7001, category Research article
Fei-Fei Zhai, Jun-Xiang Liu, Zhen-Jian Li, Jin-Mei Mao, Yong-Qiang Qian, Lei Han, Zhen-Yuan Sun. (2017). Assessing genetic diversity and population structure of Salix viminalis across Ergun and West Liao basin. Silva Fennica vol. 51 no. 3 article id 7001. https://doi.org/10.14214/sf.7001
Keywords: genetic variation; microsatellites; geographic barrier; basket willow; common osier
Highlights: High genetic diversity and moderate population differentiation were detected in twelve Salix viminalis populations; Populations from Ergun basin presented higher genetic diversity than populations from West Liao basin; Populations with short distances presented low differentiation.
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Salix viminalis L. is an important shrub that has potential for use as a bioenergy crop, for phytoremediation of heavy metal contaminated soil and sewage sludge treatment. It is mainly distributed in the northeast of China, but the species has not yet been used a resource here. We examined the genetic diversity and population structure of populations from the Ergun basin and West Liao basin using 20 microsatellite markers. A high level of genetic diversity (Na = 16.45, He = 0.742) was detected for S. viminalis, and populations from the Ergun basin exhibited higher genetic diversity and private alleles numbers than the West Liao basin. The 12 populations could be divided into two clusters by both Bayesian analysis and UPGMA clustering which were consistent with the populations derived from the two basins. Moderate population differentiation (FST = 0.076) was shown in S. viminalis, and AMOVA analysis confirmed that most of the genetic variation (86.13%) was attributed to individual differences within populations, while 11.49% was attributed to differences between basins and 2.38% to differences within each basin. Significant correlations of FST/(1–FST) with log (geographic distance) among 12 populations (r = 0.634, p < 0.00) and 10 populations within the Ergun basin (r = 0.482, p = 0.0002) indicated that geographical distance was the principal factor influencing genetic structure. As most of genetic variation exist within populations, so protection measures should be focused on populations with higher genetic diversity and unique alleles, such as Tuli, Mordaga downstream, Zhadun1 and Genhe.

  • Zhai, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China; School of Architectural and Artistic Design, Henan Polytechnic University, Century Avenue, Jiaozuo, Henan, 454000, PR China E-mail: lkyzff@163.com
  • Liu, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China E-mail: liu-jx295@163.com
  • Li, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China E-mail: zhenjianli@163.com
  • Mao, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China; Research Institute of Economic Forest, Xinjiang Academy of Forestry, Anjunanlu, Urumqi, Xinjiang, PR China E-mail: 350512173@qq.com
  • Qian, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China E-mail: qianyq@caf.ac.cn
  • Han, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China E-mail: hdd@caf.ac.cn
  • Sun, State Key Laboratory of Tree Genetics and Breeding; Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Haidian District, Beijing, 10091, PR China E-mail: lkyszy@126.com (email)
article id 6986, category Research article
Mari Tilk, Tea Tullus, Katri Ots. (2017). Effects of environmental factors on the species richness, composition and community horizontal structure of vascular plants in Scots pine forests on fixed sand dunes. Silva Fennica vol. 51 no. 3 article id 6986. https://doi.org/10.14214/sf.6986
Keywords: biodiversity; inland dunes; microhabitats; soil conditions; vegetation–site relationships
Highlights: Factors affecting the species richness, composition and horizontal structure of vascular plants are related to dune topography, resulting in the differentiation of soils and therefore complexes of different microhabitats that are populated by various vascular plant species and causing vegetation zonation.
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Different environmental factors were studied to determine which factors influence the species richness, composition and structure of vascular plants in Pinus sylvestris L. forests in a fixed dune landscape in south-western Estonia. In addition to site topographic factors, different environmental parameters were investigated. Thirty-four vascular plant species were recorded in 232 quadrats. The most abundant species was Vaccinium vitis-idaea L., which was in 82.8% of quadrats, followed by Vaccinium myrtillus L. (74.1%), Melampyrum pratense L. (71.1%) and Deschampsia flexuosa (L.) Trin. (69.8%). The multiple response permutation procedure (MRPP) showed considerable differences in species composition at the bottoms of dunes compared with that on the slopes and at the tops of dunes. Indicator species analysis (ISA) determined species exhibited characteristics specific to zone: V. myrtillus had the highest indicator value at the bottoms of dunes; Calluna vulgaris L., at the tops. Soils were Haplic Podzols, and the presence of humus horizon depended on zone. Soil conditions on the dunes were variable and site specific, in general, soils at the bottoms of the dunes were more acidic and moist compared with those of the slopes and tops of the dunes, and the nutrient content decreased toward the dune tops. According to non-metric multidimensional scaling (NMDS) and linear mixed model analyses, species coverage, composition and richness were controlled by site-specific factors such as absolute height, location and aspect of the quadrat on the dune; soil nitrogen, potassium and phosphorus contents; soil pH and moisture; light conditions; and the thickness of the litter horizon.

  • Tilk, Department of Silviculture, Estonian University of Life Sciences, Kreutzwaldi 5, Tartu, Estonia, 51014; Tallinn Botanic Garden, Kloostrimetsa Road 52, Tallinn, Estonia, 11913 E-mail: Mari.Tilk@botaanikaaed.ee (email)
  • Tullus, Department of Silviculture, Estonian University of Life Sciences, Kreutzwaldi 5, Tartu, Estonia, 51014 E-mail: Tea.Tullus@emu.ee
  • Ots, Department of Silviculture, Estonian University of Life Sciences, Kreutzwaldi 5, Tartu, Estonia, 51014 E-mail: Katri.Ots@emu.ee
article id 5662, category Research article
Samuel Egbäck, Urban Nilsson, Kenneth Nyström, Karl-Anders Högberg, Nils Fahlvik. (2017). Modeling early height growth in trials of genetically improved Norway spruce and Scots pine in southern Sweden. Silva Fennica vol. 51 no. 3 article id 5662. https://doi.org/10.14214/sf.5662
Keywords: Pinus sylvestris; Picea abies; individual tree growth model; genetic component; genetic multiplier; unimproved material; improved material
Highlights: The developed height growth model based on unimproved material predicted the development relatively well for genetically improved Norway spruce; For genetically improved Scots pine, however, the model needed to be modified; By incorporating a genetic component into the Scots pine model, the prediction errors were reduced.
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Genetically improved Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) are used extensively in operational Swedish forestry plantations to increase production. Depending on the genetic status of the plant material, the current estimated genetic gain in growth is in the range 10–20% for these species and this is expected to increase further in the near future. However, growth models derived solely from data relating to genetically improved material in Sweden are still lacking. In this study we investigated whether an individual tree growth model based on data from unimproved material could be used to predict the height increment in young trials of genetically improved Norway spruce and Scots pine. Data from 11 genetic experiments with large genetic variation, ranging from offspring of plus-trees selected in the late 1940s to highly improved clonal materials selected from well performing provenances were used. The data set included initial heights at the age of 7–15 years and 5-year increments for almost 2000 genetic entries and more than 20 000 trees. The evaluation indicated that the model based on unimproved trees predicted height development relatively well for genetically improved Norway spruce and there was no need to incorporate a genetic component. However, for Scots pine, the model needed to be modified. A genetic component was developed based on the genetic difference recorded within each trial, using mixed linear models and methods from quantitative genetics. By incorporating the genetic component, the prediction errors were significantly reduced for Scots pine. This study provides the first step to incorporate genetic gains into Swedish growth models and forest management planning systems.

  • Egbäck, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, 230 53 Alnarp, Sweden E-mail: samuel.egback@slu.se (email)
  • Nilsson, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, 230 53 Alnarp, Sweden E-mail: urban.nilsson@slu.se
  • Nyström, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd, 901 83 Umeå, Sweden E-mail: kenneth.nystrom@slu.se
  • Högberg, Skogforsk, Ekebo, 268 90 Svalöv, Sweden E-mail: karl-anders.hogberg@skogforsk.se
  • Fahlvik, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, 230 53 Alnarp, Sweden E-mail: nils.fahlvik@slu.se
article id 5656, category Research article
Lars-Göran Stener, Johan Westin. (2017). Early growth and phenology of hybrid aspen and poplar in clonal field tests in Scandinavia. Silva Fennica vol. 51 no. 3 article id 5656. https://doi.org/10.14214/sf.5656
Keywords: survival; Populus; Sweden; early test; shoot phenology; Denmark
Highlights: Growth and survival was in general higher for hybrid aspen than for poplar; The poor performance of poplar was likely due to poor climatic adaptation or to high soil acidity; The genetic results indicate good possibilities for effective clonal selection; The results support current recommendations for utilization of selected hybrid aspen and poplar regeneration material in Sweden.
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Results on early survival, growth and shoot phenology of hybrid aspen (Populus tremula L. × P. tremuloides Michx.) and poplar clones (P. trichocarpa Torr. & A. Gray, P. balsamifera L., P. maximowiczii A. Henry and their hybrids) in 13 Scandinavian field trials are presented. The trials were established on forest land (7 sites) or former agricultural land (6 sites) within the latitude range of 56° to 65° N and were assessed 3–4 years after establishment. The main aim was to evaluate phenotypic and genetic differences related to early survival, growth and phenology for hybrid aspen and poplar for different site types and latitudes. Growth and survival was generally higher for hybrid aspen than poplar at all sites. The poor performance of poplar compared to hybrid aspen is likely due to climatic maladaptation or high soil acidity. The early growth performance of the species need to be confirmed at a higher age. The genetic variation and genetic control for growth, phenology and survival was in general intermediate to large indicating good possibilities for effective clonal selection. The genetic site x site correlations (rGE) for growth were for hybrid aspen mostly strong, indicating a weak genotype by environment interaction, while rGE were inconsistent for poplars.The result suggests that southern Sweden can be treated as a single test and utilization zone and in northern Sweden the region along the coast may be another zone. It is too early to make any corresponding conclusions for poplar. In addition, the result backs up the current recommendations for utilization of selected hybrid aspen and poplar regeneration material in Sweden.

  • Stener, The Forestry Research Institute of Sweden, Ekebo 2250, 268 90 Svalöv, Sweden E-mail: lars-goran.stener@skogforsk.se (email)
  • Westin, The Forestry Research Institute of Sweden, Box 3, 918 21 Sävar, Sweden E-mail: johan.westin@skogforsk.se
article id 2017, category Research article
Pentti Niemistö, Soili Kojola, Anssi Ahtikoski, Raija Laiho. (2017). From useless thickets to valuable resource? – Financial performance of downy birch management on drained peatlands. Silva Fennica vol. 51 no. 3 article id 2017. https://doi.org/10.14214/sf.2017
Keywords: forest management; Betula pubescens; thinning; energy wood; pulpwood; profitability; final cutting
Highlights: The most profitable management regimes for pulpwood and energy wood production in dense downy birch stands on drained peatlands include no thinnings, but final cutting at the stand age of 40–45 years as whole-tree harvesting, or as integrated harvesting of pulpwood and delimbed energy wood stems about 10 years later depending on applicable harvesting method; A competitive management regime is early precommercial thinning at 4 m dominant height to a density of 2500 stems per hectare and production of pulpwood with a rotation of 55–65 years. Equal profitability is achieved with or without traditional first thinning, which can thus be included for other reasons, for example to improve regeneration of spruce.
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Downy birch (Betula pubescens Ehrh.) stands on drained peatlands are often considered useless because they typically do not yield good-quality sawn timber. However, covering an area of ca. 0.5 million hectares and with total yields of up to 250 m3 ha–1, downy birch stands on peatlands in Finland have a potential for pulpwood and/or energy wood production. We examined the financial performance of alternative management regimes (with or without thinnings, different thinning intensities, several rotation lengths) combined with alternative harvesting methods (pulpwood, energy wood, or integrated, energy wood being delimbed stems or whole trees). We used data from 19 experimental stands, monitored for 20–30 years. For harvesting removals we considered both actual thinning removals and final-cutting removals with alternative timings that were based on the monitoring data. We assessed the profitability as a combination of the net present value of the birch generation and the bare land value of future generations of Norway spruce (Picea abies (L.) Karst.). The most profitable management was growing without thinnings until whole-tree final cutting at the stand age of 40–45 years with an advanced multi-tree harvesting method. In contrast, the standard method in whole-tree final cutting resulted in the lowest profitability, and an integrated method with the energy wood as delimbed stems was the best of the standard methods. Thinnings were unprofitable especially when aiming to produce energy wood, whereas aiming for pulpwood, light precommercial thinning was competitive. Commercial thinning at the traditional “pulpwood stage” had little effect on profitability. The best stand age for final cutting was 40–65 years – earlier for very dense stands and whole-tree energy wood harvesting with advanced method, later for precommercially thinned stands and pulpwood harvesting.

  • Niemistö, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Kampusranta 9 C, 60320 Seinäjoki, Finland E-mail: pentti.niemisto@luke.fi (email)
  • Kojola, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Latokartanonkaari 9, 00790 Helsinki, Finland E-mail: soili.kojola@luke.fi
  • Ahtikoski, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Paavo Havaksentie 3, 90014 University of Oulu, Finland E-mail: anssi.ahtikoski@luke.fi
  • Laiho, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Latokartanonkaari 9, 00790 Helsinki, Finland E-mail: raija.laiho@luke.fi
article id 1743, category Research article
Gintare Sabalinkiene, Darius Danusevicius, Michael Manton, Gediminas Brazaitis, Kastytis Simkevicius. (2017). Differentiation of European roe deer populations and ecotypes in Lithuania based on DNA markers, cranium and antler morphometry. Silva Fennica vol. 51 no. 3 article id 1743. https://doi.org/10.14214/sf.1743
Keywords: genetic structure; inbreeding; Capreolus capreolus; polymorphism; wild game management
Highlights: Lithuanian roe deer populations are genetically structured into southern and northern groups, most likely affected by a divergent gene flow and Lithuania’s largest rivers slowing down migration; Microsatellite and skull morphology based genetic differentiation between field and forest ecotypes are weak; Geographical location has a significant effect on antler morphometry traits and skull size of male roe deer, the latter increasing northwards.
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The objective of our study was to assess the genetic and morphological differentiation of European roe deer (Capreolus capreolus L.) in Lithuania based on DNA markers, skull and anther morphology. DNA was extracted from 79 culled individuals at 13 locations and genotyped at five nuclear microsatellite loci. Based on culling location, individuals were assigned to either a field (N = 43) or a forest ecotype (N = 36). Skull and antler morphometry was studied on 603 and 292 individuals, respectively. Results showed no significant genetic and skull morphology differentiation between the ecotypes. The forest ecotype tends to exhibit lower genetic diversity compared to the field ecotype, particularly for male individuals. The genetic differentiation of roe deer in Lithuania was significant based on the RST values, but not on the FST values. A STRUCTURE analyses revealed southern and northern genetic clusters, most likely affected by divergent gene flow. The country’s major rivers Nemunas and Neris are likely to increase differentiation between the clusters. ANOVA on skull morphology by gender and age indicated a significant effect of geographical location. Skull size (especially length) is greater in the northern part of the country. We also found significant effects of age, ecotype and geographical location on most of the roe deer male antler morphometric traits.

  • Sabalinkiene, Institute of Forest Biology and Silviculture, Faculty of Forest Sciences and Ecology, Aleksandras Stulginskis University, Studentu street 11, Akademija, Kaunas district, Lithuania E-mail: gintare.sabalinkiene@asu.lt (email)
  • Danusevicius, Institute of Forest Biology and Silviculture, Faculty of Forest Sciences and Ecology, Aleksandras Stulginskis University, Studentu street 11, Akademija, Kaunas district, Lithuania E-mail: darius.danusevicius@asu.lt
  • Manton, Institute of Forest Biology and Silviculture, Faculty of Forest Sciences and Ecology, Aleksandras Stulginskis University, Studentu street 11, Akademija, Kaunas district, Lithuania E-mail: michael.manton@asu.lt
  • Brazaitis, Institute of Forest Biology and Silviculture, Faculty of Forest Sciences and Ecology, Aleksandras Stulginskis University, Studentu street 11, Akademija, Kaunas district, Lithuania E-mail: gediminas.brazaitis@asu.lt
  • Simkevicius, Institute of Forest Biology and Silviculture, Faculty of Forest Sciences and Ecology, Aleksandras Stulginskis University, Studentu street 11, Akademija, Kaunas district, Lithuania E-mail: kastytis.simkevicius@asu.lt
article id 1741, category Research article
Seppo Nevalainen. (2017). Comparison of damage risks in even- and uneven-aged forestry in Finland. Silva Fennica vol. 51 no. 3 article id 1741. https://doi.org/10.14214/sf.1741
Keywords: forest damage; uneven-aged forestry; even-aged forestry
Highlights: Damage risks in two forest management regimes were estimated by means of a literature review and a questionnaire to Finnish forestry experts; Damage risks were usually estimated to be higher in even-aged than in uneven-aged management regimes; In some cases, however, damage risks may be higher in uneven-aged stands (root-rot infected Norway spruce stands and mechanical damage due to repeated thinnings).
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The literature on the most prominent forest damage related to even-aged and uneven-aged forest management regimes was reviewed. A questionnaire to expert researchers was conducted to estimate risks in even-aged and uneven-aged forest management chains in Finland. There are only a few empirical comparisons of damage risks in even- and uneven-aged stands in the literature. The results from the expert survey showed that the damage risks were higher in even-aged management in Norway spruce and Scots pine. However, the variation in the risks between individual chains and between individual causes was high. The highest risks in Scots pine were caused by moose (in even-aged chains) and harvesting damage (in uneven-aged chains). In Norway spruce, root rot caused the highest risks in both even-aged and uneven-aged chains. The higher risks in even-aged forestry are largely due to the many associated practices which favour various types of damage. However, there are some important exceptions: the damage risks may be higher in some uneven-aged stands, especially in Norway spruce stands infected with root rot where the utilization of undergrowth or natural regeneration can be risky. Moreover, the repeated thinnings in uneven-aged stands may lead to increased mechanical damage.

  • Nevalainen, Natural Resources Institute Finland (Luke), Natural Resources and Bioproduction, Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: seppo.nevalainen@luke.fi (email)
article id 1734, category Research article
Jyrki Hytönen, Paula Jylhä, Keith Little. (2017). Positive effects of wood ash fertilization and weed control on the growth of Scots pine on former peat-based agricultural land – a 21-year study. Silva Fennica vol. 51 no. 3 article id 1734. https://doi.org/10.14214/sf.1734
Keywords: Pinus sylvestris; fertilization; vegetation management; shading; agricultural soil; overgrowth; potassium deficiency; production potential
Highlights: Weed control decreased and fertilization increased vegetation height and shading of seedlings; Weed control decreased mortality, but fertilization had no effect; Despite improved foliar K concentration though ash fertilization, all trees in the trial had severe K deficiency after 21 years; Weed control increased growth by 20 m3 ha–1 and fertilization by 35 m3 ha–1 in 21 years.
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The impacts of weed control, ash fertilization and their interaction were tested for the afforestation of former agricultural peat-based soil with Scots pine (Pinus sylvestris L.) in northern Finland in a factorial arrangement of four treatments. Weed control with herbicides was carried out in July 1 and 2 years from planting, and wood ash (5 Mg ha–1) was applied in the spring of the 2nd year. Various vegetation, tree growth and nutrient assessments were made over the 21-year study period. Weed control decreased the weed cover by 36–56 percentage points, vegetation height by 4–26 cm and thus shading of seedlings by vegetation for at least 4 years after planting. For the same period, ash fertilization increased vegetation height by 6–15 cm and shading of seedlings. Weed control reduced seedling mortality by 27 percentage points in 21 years, but ash fertilization had no significant effect. Ash fertilization increased foliar potassium and boron concentrations, but its effect declined, and severe K-deficiency was recorded 21 years after planting. Up to the 9th year, weed control had a greater influence on growth than fertilization. Later the significance of fertilization increased due to an aggravated K-deficiency. Stand volume at year 21 for the untreated control plots was 8 m3 ha–1. Weed control and fertilization increased stand volume by 20 and 35 m3 ha–1, with a combined effect of 55 m3 ha–1. The effects of weed control and fertilization were additive and no significant interactions were found. Due to severe K-deficiencies, re-fertilization of all treatments would be necessary for the continued survival and growth of Scots pine.

  • Hytönen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Teknologiakatu 7, FI-67100 Kokkola, Finland ORCID http://orcid.org/0000-0001-8475-3568 E-mail: jyrki.hytonen@luke.fi (email)
  • Jylhä, Natural Resources Institute Finland (Luke), Green technology, Teknologiakatu 7, FI-67100 Kokkola, Finland E-mail: paula.jylha@luke.fi
  • Little, Nelson Mandela Metropolitan University, George Campus, Western Cape, South Africa E-mail: keith.little@nmmu.ac.za
article id 1666, category Research article
Mareike Wieczorek, Alexei Kolmogorov, Stefan Kruse, Inga Jacobsen, Ingmar Nitze, Anatoly N. Nikolaev, Ingo Heinrich, Luidmila A. Pestryakova, Ulrike Herzschuh. (2017). Disturbance-effects on treeline larch-stands in the lower Kolyma River area (NE Siberia). Silva Fennica vol. 51 no. 3 article id 1666. https://doi.org/10.14214/sf.1666
Keywords: stand structure; fire; treeline; Larix cajanderi; Siberia
Highlights: Disturbances are an important shaping factor of larch stands at lower Kolyma; Youngest larch stands have the highest population densities and highest growth rates; Saplings grow clustered, irrespective of the underlying disturbance regime; Mixed climate-disturbance signals make it difficult to infer future treeline processes.
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Tree stands in the boreal treeline ecotone are, in addition to climate change, impacted by disturbances such as fire, water-related disturbances and logging. We aim to understand how these disturbances affect growth, age structure, and spatial patterns of larch stands in the north-eastern Siberian treeline ecotone (lower Kolyma River region), an insufficiently researched region. Stand structure of Larix cajanderi Mayr was studied at seven sites impacted by disturbances. Maximum tree age ranged from 44 to 300 years. Young to medium-aged stands had, independent of disturbance type, the highest stand densities with over 4000 larch trees per ha. These sites also had the highest growth rates for tree height and stem diameter. Overall lowest stand densities were found in a polygonal field at the northern end of the study area, with larches growing in distinct “tree islands”. At all sites, saplings are significantly clustered. Differences in fire severity led to contrasting stand structures with respect to tree, recruit, and overall stand densities. While a low severity fire resulted in low-density stands with high proportions of small and young larches, high severity fires resulted in high-density stands with high proportions of big trees. At water-disturbed sites, stand structure varied between waterlogged and drained sites and latitude. These mixed effects of climate and disturbance make it difficult to predict future stand characteristics and the treeline position.

  • Wieczorek, Periglacial Research Section, Alfred Wegener Institute Helmholtz Centre for Polar and Marine Research, 14473 Potsdam, Germany; Institute of Earth and Environmental Science, University of Potsdam, 14476 Potsdam, Germany E-mail: mareike.wieczorek@awi.de
  • Kolmogorov, Institute of Natural Sciences, North-Eastern Federal University of Yakutsk, 677000 Yakutsk, Russia E-mail: kilatroooon@gmail.com
  • Kruse, Periglacial Research Section, Alfred Wegener Institute Helmholtz Centre for Polar and Marine Research, 14473 Potsdam, Germany; Institute of Earth and Environmental Science, University of Potsdam, 14476 Potsdam, Germany E-mail: Stefan.Kruse@awi.de
  • Jacobsen, Periglacial Research Section, Alfred Wegener Institute Helmholtz Centre for Polar and Marine Research, 14473 Potsdam, Germany; Institute of Earth and Environmental Science, University of Potsdam, 14476 Potsdam, Germany E-mail: Inga.Jacobsen@awi.de
  • Nitze, Periglacial Research Section, Alfred Wegener Institute Helmholtz Centre for Polar and Marine Research, 14473 Potsdam, Germany; Institute of Geography, University of Potsdam, 14476 Potsdam, Germany E-mail: Ingmar.Nitze@awi.de
  • Nikolaev, Institute of Natural Sciences, North-Eastern Federal University of Yakutsk, 677000 Yakutsk, Russia; Melnikov Permafrost Institute of the Siberian Branch of RAS, 677000 Yakutsk, Russia E-mail: yktnan@rambler.ru
  • Heinrich, GFZ German Research Centre for Geosciences, Telegrafenberg, 14473 Potsdam, Germany E-mail: heinrich@gfz-potsdam.de
  • Pestryakova, Institute of Natural Sciences, North-Eastern Federal University of Yakutsk, 677000 Yakutsk, Russia E-mail: lapest@mail.ru
  • Herzschuh, Periglacial Research Section, Alfred Wegener Institute Helmholtz Centre for Polar and Marine Research, 14473 Potsdam, Germany; Institute of Earth and Environmental Science, University of Potsdam, 14476 Potsdam, Germany E-mail: Ulrike.Herzschuh@awi.de (email)
article id 1631, category Research article
Jonas Koala, Louis Sawadogo, Patrice Savadogo, Ermias Aynekulu, Janne Heiskanen, Mohammed Saïd. (2017). Allometric equations for below-ground biomass of four key woody species in West African savanna-woodlands. Silva Fennica vol. 51 no. 3 article id 1631. https://doi.org/10.14214/sf.1631
Keywords: root; shoot ratio; Burkina Faso; biomass estimation; species-specific equation; root excavation
Highlights: Species-specific equations for belowground biomass (BGB) predicted biomass with less bias than generic equations; All the generic equations underestimated BGB; For accurate estimation of BGB in savanna-woodlands, species-specific equations are needed for more species.
Abstract | Full text in HTML | Full text in PDF | Author Info

Accurate estimates of both above-ground biomass (AGB) and below-ground biomass (BGB) are essential for estimating carbon (C) balances at various geographical scales and formulating effective climate change mitigation programs. However, estimating BGB is challenging, particularly for forest ecosystems, so robust allometric equations are needed. To obtain such equations for savanna-woodlands of the West African north sudanian zone, we selected four common native woody species (Anogeissus leiocarpa (DC.) Guill. & Perr., Detarium microcarpum Guill. & Perr., Piliostigma thonningii (Schumach.) Milne-Redh. and Vitellaria paradoxa C.F. Gaertn.). At two sites in Burkina Faso, we determined the BGB of 30 trees of each of these species by excavation, and measured various above-ground dimensional variables. The root:shoot ratio varied widely among the species, from 0.1 to 3.4. Depending on the species, allometric equations based on stem basal area at 20 cm height, basal area at breast height and tree height explained 50–95% of the variation in BGB. The best generic equation we obtained, based on basal area at 20 cm, explained 60% of the variation in BGB across the species. Three previously published generic allometric equations underestimated BGB by 8 to 63%. The presented equations should significantly improve the accuracy of BGB estimates in savanna-woodlands and help avoid costly needs to excavate root systems.

  • Koala, Centre National de Recherche Scientifique et Technologique (CNRST), Institut de l’Environnement et de Recherches Agricoles (INERA), Département Productions Forestières, 03 BP 7047, Ouagadougou 03, Burkina Faso E-mail: ezeyamb@yahoo.fr (email)
  • Sawadogo, Centre National de Recherche Scientifique et Technologique (CNRST), Institut de l’Environnement et de Recherches Agricoles (INERA), Département Productions Forestières, 03 BP 7047, Ouagadougou 03, Burkina Faso E-mail: sawadogo_ls@hotmail.com
  • Savadogo, World Agroforestry Centre & International Crop Research Institute for the Semi-Arid Tropics (ICRAF-ICRISAT), West and Central Africa Region-Sahel Node, BP 12404, Niamey, Niger E-mail: savadogo.patrice@gmail.com
  • Aynekulu, World Agroforestry Centre (ICRAF), United Nations Avenue, P.O. Box 30677-00100, Nairobi, Kenya E-mail: e.betemariam@cgiar.org
  • Heiskanen, University of Helsinki, Department of Geosciences and Geography, P.O. Box 68, FI-00014 University of Helsinki, Finland E-mail: janne.heiskanen@helsinki.fi
  • Saïd, International Livestock Research Institute (ILRI). P.O. Box 30709, Nairobi, Kenya E-mail: m.said@cgiar.org
article id 1563, category Research article
Kristina Ahnlund Ulvcrona, Dan Bergström, Urban Bergsten. (2017). Stand structure after thinning in 1–2 m wide corridors in young dense stands. Silva Fennica vol. 51 no. 3 article id 1563. https://doi.org/10.14214/sf.1563
Keywords: heterogeneity; conventional thinning; future crop trees; production
Highlights: Boom corridor thinning (BCT) results in more stand structure heterogeneity than conventional thinning or pre-commercial thinning (PCT), maintaining both smaller-diameter trees and deciduous species; Neither dominant height nor number of possible future crop trees is jeopardized, and boom corridor thinning results in higher values of stem volume and biomass; The technique is flexible as various corridor types give similar stand structure results.
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Boom corridor thinning (BCT) has been proposed as a cost-effective technique for biomass thinning (BT) in young dense stands. The objective of this study was to determine how various BCT operations affect stand structure following biomass thinning and to compare the results with conventional selective thinning methods. Two series of field experiments were established; BCT 1-series: Three sites in south of Sweden (9 and 11 m in mean and dominating tree height) with five treatments, including a control, conventional selective thinning and three BCT treatments (1 m and 2 m wide corridors and selective BCT). The second BCT series: Three regions in Sweden (in the north, centre and in the south), with two stand sites in each region with different tree heights (4/9 m and 5/10 m in mean/dominating tree height). Treatments were control, pre-commercial thinning (PCT), conventional selective thinning and BCT (high and low thinning). Following the first biomass thinning, BCT regimes and selective thinning methods resulted in similar stand structures based on the number of possible future crop trees (>80 mm in diameter at breast height). However, BCT maintained a higher diversity of tree sizes as well as more stems per hectare, including deciduous species, than the selective thinning approaches. The stands after BCT should have more vertical complexity, especially when compared to pre-commercial thinning. The structural heterogeneity resulting from BCT may also increase stand biodiversity and ecosystem service values.

  • Ahnlund Ulvcrona, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: kristina.ulvcrona@slu.se (email)
  • Bergström, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: dan.bergstrom@slu.se
  • Bergsten, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: urban.bergsten@slu.se
article id 5643, category Research article
Katarzyna A. Jadwiszczak, Stanisław Kłosowski, Iwona Zalewska, Agata Banaszek, Agnieszka Chrzanowska. (2017). Genetic diversity and sexual reproduction in relict populations of Betula nana. Silva Fennica vol. 51 no. 2 article id 5643. https://doi.org/10.14214/sf.5643
Keywords: flowering; AFLP; dwarf birch; meiotic recombination; seed germination
Highlights: Genetic diversity parameters and meiotic recombination frequencies in the relict populations were comparable to those from widespread localities; Contribution of seeds without ovule was very high; Fully developed seeds germinated better in central populations; Significant differences of groundwater parameters were observed between relict and central populations.
Abstract | Full text in HTML | Full text in PDF | Author Info

In the present study, the impact of geographical isolation and habitat conditions on genetic diversity and sexual reproduction was tested in four relict populations of dwarf birch Betula nana L. in Poland and Belarus. Amplified fragment length polymorphism (AFLP) method revealed that the endangered central European stands were not genetically extirpated compared with the widespread localities from Finland and Russia, which can result from infrequent outcrossing events in long-living clonal populations. However, genetic clustering methods indicated significant differentiation of the Polish populations because of their small sizes and long-term geographical isolation. Considerable numbers of empty seeds were observed in both relict and central locations, although fully developed seeds germinated better in widespread populations. Analysis of groundwater chemical parameters indicated that two relict populations were significantly different from the remaining samples with respect to pH, electrical conductivity and concentrations of phosphorus ions, which can also influence the efficiency of sexual reproduction. In the light of results obtained it seems that endangered B. nana localities are relatively stable.

  • Jadwiszczak, Institute of Biology, University of Białystok, Ciołkowskiego 1J, 15-245 Białystok, Poland ORCID http://orcid.org/0000-0002-9345-8891 E-mail: jadwiszczak2010@gmail.com (email)
  • Kłosowski, Department of Environment Protection and Modelling, The Jan Kochanowski University, Świętokrzyska 15, 25-406 Kielce, Poland E-mail: stanislaw.klosowski@ujk.kielce.pl
  • Zalewska, Faculty of Pharmacy, Medical University of Białystok, Mickiewicza 2a, 15-222 Białystok, Poland E-mail: iwonazalewska1988@gmail.com
  • Banaszek, Institute of Biology, University of Białystok, Ciołkowskiego 1J, 15-245 Białystok, Poland E-mail: banaszek@uwb.edu.pl
  • Chrzanowska, Institute of Biology, University of Białystok, Ciołkowskiego 1J, 15-245 Białystok, Poland E-mail: maga.chrzanowska@gmail.com
article id 2021, category Research article
Jonas Bohlin, Inka Bohlin, Jonas Jonzén, Mats Nilsson. (2017). Mapping forest attributes using data from stereophotogrammetry of aerial images and field data from the national forest inventory. Silva Fennica vol. 51 no. 2 article id 2021. https://doi.org/10.14214/sf.2021
Keywords: airborne laser scanning; National Forest Inventory; photogrammetry; aerial images; forest attribute estimation; image matching; large area
Highlights: Image based forest attribute map generated using NFI plots show similar accuracy as currently used LiDAR based forest attribute map; Also similar accuracies were found for different forest types; Aerial images from leaf-off season is not recommended.
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Exploring the possibility to produce nation-wide forest attribute maps using stereophotogrammetry of aerial images, the national terrain model and data from the National Forest Inventory (NFI). The study areas are four image acquisition blocks in mid- and south Sweden. Regression models were developed and applied to 12.5 m × 12.5 m raster cells for each block and validation was done with an independent dataset of forest stands. Model performance was compared for eight different forest types separately and the accuracies between forest types clearly differs for both image- and LiDAR methods, but between methods the difference in accuracy is small at plot level. At stand level, the root mean square error in percent of the mean (RMSE%) were ranging: from 7.7% to 10.5% for mean height; from 12.0% to 17.8% for mean diameter; from 21.8% to 22.8% for stem volume; and from 17.7% to 21.1% for basal area. This study clearly shows that aerial images from the national image program together with field sample plots from the NFI can be used for large area forest attribute mapping.

  • Bohlin, Department of Forest Resource Management, Swedish University of Agricultural Sciences, S-901 35 Umeå, Sweden ORCID http://orcid.org/0000-0002-3318-5967 E-mail: jonas.bohlin@slu.se (email)
  • Bohlin, Department of Forest Resource Management, Swedish University of Agricultural Sciences, S-901 35 Umeå, Sweden ORCID http://orcid.org/0000-0003-1224-6684 E-mail: inka.bohlin@slu.se
  • Jonzén, Department of Forest Resource Management, Swedish University of Agricultural Sciences, S-901 35 Umeå, Sweden E-mail: jonas.jonzen@slu.se
  • Nilsson, Department of Forest Resource Management, Swedish University of Agricultural Sciences, S-901 35 Umeå, Sweden ORCID http://orcid.org/0000-0001-7394-6305 E-mail: mats.nilsson@slu.se
article id 1781, category Research article
Petr Čermák, Michal Rybníček, Tomáš Žid, Kjell Andreassen, Isabella Børja, Tomáš Kolář. (2017). Impact of climate change on growth dynamics of Norway spruce in south-eastern Norway. Silva Fennica vol. 51 no. 2 article id 1781. https://doi.org/10.14214/sf.1781
Keywords: Picea abies; precipitation; crown condition; decline; tree-ring width; Oslofjord
Highlights: Correlations between tree-ring width and climate parameters showed temporal instability in their relationship during the period 1915–2012; A statistically significant positive correlation of April–May precipitation on tree-ring growth was identified since the mid-1970s; The concomitant temperature increase may have contributed to the changes of growth dynamics.
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The ongoing climate change may have a distinct effect on Norway spruce growth, one of the most important tree species in European forest management. Therefore, the understanding and assessment of climate-growth relationship can help to reveal relevant patterns in temporal variability that may result in lower tree vitality and decline. The main objective of our study was to evaluate the long-term climate-growth variability of Norway spruce in south-eastern Norway, at the northern edge of the temperate zone. We sampled in total 270 dominant and co-dominant trees from 18 plots in south-eastern Norway. We analysed stem cores and evaluated crown condition parameters to assess the retrospective tree growth and vitality. Despite considerable differences in the crown parameters, high similarity among tree-ring width (TRW) series allowed compiling the regional tree-ring width chronology. Correlations between TRW and climate parameters showed temporal instability in their relationship during the period 1915–2012. While we did not detect any significant relationships between TRW and climate parameters in the first half of the study period (1915–1963), a significant correlation between TRW and spring precipitation was observed for the period 1964–2012. This shift appeared concurrent with temperatures reaching above-average values compared to the average of the climate normal period 1961–1990.

  • Čermák, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědělská 3, 613 00 Brno, Czech Republic E-mail: cermacek@mendelu.cz (email)
  • Rybníček, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědělská 3, 613 00 Brno, Czech Republic; Global Change Research Institute, The Czech Academy of Sciences, Bělidla 986/4a, 603 00 Brno, Czech Republic E-mail: michalryb@post.cz
  • Žid, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědělská 3, 613 00 Brno, Czech Republic E-mail: tom.z@centrum.cz
  • Andreassen, Norwegian Institute of Bioeconomy Research (NIBIO), P.O. Box 115, NO-1431 Ås, Norway E-mail: Kjell.Andressen@nibio.no
  • Børja, Norwegian Institute of Bioeconomy Research (NIBIO), P.O. Box 115, NO-1431 Ås, Norway E-mail: Isabella.Borja@nibio.no
  • Kolář, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědělská 3, 613 00 Brno, Czech Republic; Global Change Research Institute, The Czech Academy of Sciences, Bělidla 986/4a, 603 00 Brno, Czech Republic E-mail: koldatom@gmail.com
article id 1721, category Research article
Anna Hebda, Błażej Wójkiewicz, Witold Wachowiak. (2017). Genetic characteristics of Scots pine in Poland and reference populations based on nuclear and chloroplast microsatellite markers. Silva Fennica vol. 51 no. 2 article id 1721. https://doi.org/10.14214/sf.1721
Keywords: Pinus sylvestris; phylogeography; conifers; genetic divergence; population structure; microsatellite markers; neutral variation
Highlights: Similar genetic variation was found between Polish Scots pine populations from a wide variety of habitats based on nSSR and cpSSR markers; Homogeneity was observed in the genetic structures of Polish and Finnish populations from the continuous pine range; Genetic differentiation in microsatellite markers was identified only when populations from the central pine distribution were compared to the marginal stands.
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Polymorphisms at a set of eighteen nuclear (nSSR) and chloroplast (cpSSR) microsatellite loci were investigated in sixteen populations of Scots pine (Pinus sylvestris L.) derived from the provenance trial experiment and representative of the species distribution range and climatic zones in Poland. The patterns of genetic variation were compared to the reference samples from the species distribution in Europe and Asia. A similar level of genetic variation and no evidence of population structure was found among the Polish stands. They showed genetic similarity and homogenous patterns of allelic frequency spectra compared to the Northern European populations. Those populations were genetically divergent compared to the marginal populations from Turkey, Spain and Scotland. The population structure patterns reflect the phylogeography of the species and the divergence of populations that most likely do not share recent history. As the analysed provenance trial populations from Poland are diverged in phenotypic traits but are genetically similar, they could be used to test for selection at genomic regions that influence variation in quantitative traits.

  • Hebda, University of Agriculture in Krakow, Faculty of Forestry, Institute of Forest Ecology and Silviculture, Department of Genetics and Forest Tree Breeding, 29 Listopada 46, 31-425 Kraków, Poland ORCID http://orcid.org/0000-0002-3149-8644 E-mail: ana.hebda@gmail.com (email)
  • Wójkiewicz, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland E-mail: bwojkiew@man.poznan.pl
  • Wachowiak, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland; Institute of Environmental Biology, Faculty of Biology, Adam Mickiewicz University, Umultowska 89, 61-614 Poznań, Poland E-mail: witoldw@man.poznan.pl
article id 1713, category Research article
Lars-Göran Stener, Lars Rytter, Gunnar Jansson. (2017). Effects of pruning on wood properties of planted silver birch in southern Sweden. Silva Fennica vol. 51 no. 2 article id 1713. https://doi.org/10.14214/sf.1713
Keywords: wood defects; discolouration; rot; ingrown bark
Highlights: Pruning silver birch trees increased the production of defect-free wood outside the knots; Most wood defects were found inside the knots; Pruned birch trees provide butt logs with higher value than unpruned trees.
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Pruning was performed at midsummer in two genetically homogenous and managed planted silver birch stands in southern Sweden – one aged 9 and one aged 10 years. Wood defects were analysed 10 years thereafter, using the five uppermost twigs of the stems up to a height of 30 dm. The number of trees examined at each site was around 70, of which half were pruned. The main findings were that: a) compared to unpruned trees, pruned trees produced more defect-free wood outside the knots; b) most wood defects were found inside the knots; and c) wood defects like rot and bark ingrowth were similar for pruned and unpruned trees, while discolouration was marginally higher for pruned trees inside knots but similar outside knots. Overall, the results confirm previous findings that pruned birch trees will provide butt logs with higher value than unpruned trees.

  • Stener, The Forestry Research Institute of Sweden, Ekebo 2250, 268 90 Svalöv, Sweden E-mail: lars-goran.stener@skogforsk.se (email)
  • Rytter, The Forestry Research Institute of Sweden, Ekebo 2250, 268 90 Svalöv, Sweden E-mail: lars.rytter@skogforsk.se
  • Jansson, The Forestry Research Institute of Sweden, Uppsala Science Park, 751 83 Uppsala, Sweden E-mail: gunnar.jansson@skogforsk.se
article id 1693, category Research article
Olalla Díaz-Yáñez, Blas Mola-Yudego, José Ramón González-Olabarria. (2017). What variables make a forest stand vulnerable to browsing damage occurrence? Silva Fennica vol. 51 no. 2 article id 1693. https://doi.org/10.14214/sf.1693
Keywords: moose; forest damage; cervid; machine learning; risk modelling; ungulates
Highlights: Stands more vulnerable to browsing damage are young with lower densities and dominated by birch, pine or mixed species; Stand size could play a role on forest susceptibility to browsing occurrence.
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Ungulate browsing results in important damages on the forests, affecting their structure, composition and development. In the present paper, we examine the occurrence of browsing damage in Norwegian forests, using data provided by the National Forest Inventory along several consecutive measurements (entailing the period 1995–2014). A portfolio of variables describing the stand, site and silvicultural treatments are analyzed using classification trees to retrieve combinations related to browsing damage. Our results indicate that the most vulnerable forest stands are young with densities below 1400 trees ha–1 and dominated by birch, pine or mixed species. In addition, stand diversity and previous treatments (e.g. thinnings) increase the damage occurrence and other variables, like stand size, could play a role on forest susceptibility to browsing occurrence although the latter is based on weaker evidence. The methods and results of our study can be applied to implement management measures aiming at reducing the browsing damages of forests.

  • Díaz-Yáñez, School of Forest Sciences, University of Eastern Finland, PO Box 111, 80101 Joensuu, Finland ORCID http://orcid.org/0000-0003-3829-5759 E-mail: olalla.diaz@gmail.com (email)
  • Mola-Yudego, Norwegian Institute of Bioenergy Research, P.O. Box, 115, 1431 Ås, Norway; School of Forest Sciences, University of Eastern Finland, PO Box 111, 80101 Joensuu, Finland ORCID http://orcid.org/0000-0003-0286-0170 E-mail: blas.mola@uef.fi
  • González-Olabarria, Forest Sciences Centre of Catalonia (CTFC-CEMFOR), Ctra. de St. Llorenç de Morunys, km 2, 25280 Solsona, Spain ORCID http://orcid.org/0000-0002-5040-712X E-mail: jr.gonzalez@ctfc.es
article id 1678, category Research article
Juan F. Jimenez, Pedro Sánchez-Gómez, Jose Luis Cánovas, Isabell Hensen, Miloud Aouissat. (2017). Influence of natural habitat fragmentation on the genetic structure of Canarian populations of Juniperus turbinata. Silva Fennica vol. 51 no. 2 article id 1678. https://doi.org/10.14214/sf.1678
Keywords: genetic diversity; AFLP markers; Canarian juniper; oceanic islands
Highlights: Canarian populations of Juniperus turbinata exhibited levels of genetic diversity similar to those of mainland populations; Despite the historical geologic events occurred in Canary Islands, and the documented decrease of populations since the arrival of settlers, no genetic differentiation between islands has been observed; As it has been observed for other Macaronesian endemics, J. turbinata long dispersal ability seems to be the main force driving the genetic structure of populations.
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Oceanic archipelagos provide an important platform from which to evaluate the effects of isolation and fragmentation on the genetic structure of species. As a result of oceanic isolation, such species usually show lower levels of genetic diversity and higher genetic differentiation than their mainland congeners. However, this is not necessarily the case for long distance dispersal species, whose genetic structure is not strictly defined by population isolation. We assessed the level and distribution of genetic diversity among Canarian populations of Juniperus turbinata in order to evaluate the influence of population isolation on its genetic structure. Using Amplified Fragment Length Polymorphism markers, we analyzed molecular diversity among 175 individuals from five populations occurring across the Canary Island and three Moroccan populations. Principal Coordinate Analysis, neighbor joining clustering, AMOVA and Bayesian-based analysis were applied to examine population structure. Despite the documented habitat loss and decline in Canarian populations, Amplified Fragment Length Polymorphism markers revealed levels of intra-population genetic diversity that were similar to those from mainland populations, and low levels of genetic differentiation. Bayesian analysis of population structure showed three main clusters, one comprising El Hierro population and a few individuals from several islands, a second cluster that grouped the remaining Canarian populations together, and a third cluster grouping Moroccan populations. Our results suggest that the main force driving the genetic structure of Canarian populations of J. turbinata is its capacity for long distance dispersal.

  • Jimenez, Departamento de Biología Vegetal (Botánica), Universidad de Murcia, Campus de Espinardo s/n, E-30100 Murcia, Spain E-mail: fjimenez@um.es (email)
  • Sánchez-Gómez, Departamento de Biología Vegetal (Botánica), Universidad de Murcia, Campus de Espinardo s/n, E-30100 Murcia, Spain ORCID http://orcid.org/0000-0001-6754-1512 E-mail: psgomez@um.es
  • Cánovas, Departamento de Biología Vegetal (Botánica), Universidad de Murcia, Campus de Espinardo s/n, E-30100 Murcia, Spain E-mail: joseluis.canovas@um.es
  • Hensen,  Institut für Biologie, Martin-Luther Universitat, 06099 Halle, Germany E-mail: isabell.hensen@botanik.uni-halle.de
  • Aouissat, Centre Universitaire Salhi Ahmed Naama, BP 66, Naâma, Algérie E-mail: aouissatm@yahoo.fr
article id 1657, category Research article
Razvan Vasile Campu, Arcadie Ciubotaru. (2017). Time consumption and productivity in manual tree felling with a chainsaw – a case study of resinous stands from mountainous areas. Silva Fennica vol. 51 no. 2 article id 1657. https://doi.org/10.14214/sf.1657
Keywords: time study; Husqvarna; work time structure; harvesting systems; resinous temperate forest
Highlights: An important preoccupation in sustainable logging management is represented by the analysis of work time structure and productivity level in manual tree felling with a chainsaw; Sound knowledge of the factors which influence work time allows better planning of harvesting operations so that deadlines could be met and damage to forest ecosystems be minimized.
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The purpose of this research is to establish time consumption and productivity when using Husqvarna 365 chainsaw for resinous tree felling in mountainous regions. The research was conducted in the Romanian Southern Carpathians, in two mixed spruce (Picea abies (L.) H. Karst.) and fir (Abies alba Mill.) tree stands (S1 and S2). Only one team of workers, made up of a feller and an assistant, was used in the felling operation. This was divided into nine specific stages for which work times were measured. Work time structure used here includes WP – workplace time (PW – productive work time; SW – supportive work time, NT – non-work time) and NW – non-workplace time. The results indicated a productivity of 10.138 m3 h–1 (4.55 tree h–1) in S1 and of 11.374 m3 h–1 (4.33 tree h–1) in S2. Work time structure was WP 88.61% (PW 19.59%; SW 33.88%; NT 35.14%) and NW 11.39% in S1 and WP 83.77% (PW 17.66%; SW 30.73%; NT 35.38%) and NW 16.23% in S2. The results obtained showed that the power function best describes the relationship between productivity expressed by tree h–1 and breast height diameter (dbh) (R2 = 0.89 in S1 and R2 = 0.94 in S2). When productivity is expressed by m3 h–1 the results obtained in the case of power, exponential and linear functions are comparable (R2 = 0.65 to 0.67 in S1 and R2 = 0.81 to 0.92 in S2). Productivity is also influenced by stump diameter and the distance between trees. Their influence on productivity was emphasized by linear regression equations.

  • Campu, Transilvania University of Braşov, Faculty of Silviculture and Forest Engineering, Department of Forest Engineering, Forest Management Planning and Terrestrial Measurements, Şirul Beethoven no. 1, 500123, Braşov, Romania E-mail: vasile.campu@unitbv.ro (email)
  • Ciubotaru, Transilvania University of Braşov, Faculty of Silviculture and Forest Engineering, Department of Forest Engineering, Forest Management Planning and Terrestrial Measurements, Şirul Beethoven no. 1, 500123, Braşov, Romania E-mail: ciuboarc@unitbv.ro
article id 1607, category Research article
Yanlin Fu, Juan A. Oliet, Guolei Li, Jiaxi Wang. (2017). Effect of controlled release fertilizer type and rate on mineral nutrients, non-structural carbohydrates, and field performance of Chinese pine container-grown seedlings. Silva Fennica vol. 51 no. 2 article id 1607. https://doi.org/10.14214/sf.1607
Keywords: growth; nursery fertilization; Pinus tabulaeformis; nutrient loading; outplanting performance
Highlights: We demonstrated that Chinese pine container-grown seedling nutrient status and non-structural carbohydrate content were sufficient over a wide range of fertilization rates; Fertilization at 80 mg N seedling–1 was optimal for seedling responses in the nursery and field; Nursery fertilization using controlled release fertilizer (CRF) with a single coating layer yielded better seedling nursery performance than CRF with multiple coatings.
Abstract | Full text in HTML | Full text in PDF | Author Info

Although controlled release fertilizer (CRF) with single and multiple-layer coatings are extensively used in tree seedlings, studies that compare the impact of CRF type and application rate on seedling growth, nutrient storage, and, most importantly, outplanting performance, are lacking. In the current study, container-grown Pinus tabulaeformis Carr. (Chinese pine) seedlings were fertilized with commercial CRF with either one or multiple coating layers with equivalent formulation and longevity, at six rates ranging from 40 to 240 mg N seedling–1. Seedlings were sampled for dry mass, non-structural carbohydrate (NSC) content, and mineral nutrient status at the end of the growing season in the nursery, and subsequently outplanted for one season. Compared to Chinese pine seedlings fertilized with single-layer CRF treatments, seedlings treated with multiple-layer CRF had higher starch concentrations but reduced dry mass and N, P, K concentrations in the nursery, and reduced diameter growth in the field. Fertilization rates of 80 and 120 mg N seedling–1 generally yielded maximal plant dry mass and mineral nutrient content. Field survival peaked at 80 mg N seedling–1. Seedling growth, soluble sugar content, and starch concentration in the nursery and survival in the field consistently decreased at rates of 200 and 240 mg N seedling–1. In our study, optimal nursery and field performance of P. tabulaeformis were observed using single layer CRF at 80 mg N seedling–1 (3.3 g CRF l–1 media).

  • Fu, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing Forestry University; Beijing Laboratory of Urban and Rural Ecological Environment; 35 East Qinghua Road, Haidian District, Beijing 100083, China E-mail: bjfu_fu@163.com
  • Oliet, Department of Natural Systems and Resources, Universidad Politécnica de Madrid, Ciudad Universitaria s/n, 28040 Madrid, Spain E-mail: juan.oliet@upm.es
  • Li, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing Forestry University; Beijing Laboratory of Urban and Rural Ecological Environment; 35 East Qinghua Road, Haidian District, Beijing 100083, China E-mail: glli226@163.com (email)
  • Wang, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing Forestry University; Beijing Laboratory of Urban and Rural Ecological Environment; 35 East Qinghua Road, Haidian District, Beijing 100083, China E-mail: wjx198979@163.com
article id 1599, category Research article
Andrew McEwan, Michal Brink, Raffaele Spinelli. (2017). Factors affecting the productivity and work quality of chain flail delimbing and debarking. Silva Fennica vol. 51 no. 2 article id 1599. https://doi.org/10.14214/sf.1599
Keywords: modeling; plantation; coppice; efficiency; pulpwood; processing
Highlights: Machine productivity averaged 59 m3 ub SMH–1, with a 19% incidence of delay time; Productivity increased 70% if tree volume increased from 0.1 to 0.4 m3 ub; Debarking quality was good for 58% of the trees, medium for 29% and poor for 13%; The more trees in a bunch and the higher BWBS, the lower debarking quality.
Abstract | Full text in HTML | Full text in PDF | Author Info

Chain flail delimbing and debarking may improve value recovery from small tree harvests, without renouncing the benefits of multi-tree processing. The technology is mature and capable of excellent performance, which has been documented in many benchmark studies. This paper offers new insights into the relationship between the performance of chain flail delimbing and debarking and such factors as tree volume, load volume, tree form and bark-wood bond strength (BWBS). The study was conducted in Chile, during the commercial harvesting of a Eucalyptus globulus Labill. plantation. In an observational study, researchers collected production data from over 780 work cycles, and work quality data from over 1000 individual trees. The analysis of these data shows that productivity is affected primarily by load volume. Work quality is affected by BWBS and by the number of trees in a load. Work quality degrades with increasing BWBS and tree number, since more trees tend to shield each other. Tree form has no effect on either productivity or work quality. Regression and probability functions are provided, and can be used for predictive purposes when trying to optimize current operations or to prospect the introduction of chain flail technology to new work environments.

  • McEwan, Postgraduate Forest Programme, Faculty of Natural and Agricultural Sciences, University of Pretoria, Private Bag X20 Hatfield, Pretoria, 0028, South Africa E-mail: Andrew.McEwan@nmmu.ac.za
  • Brink, Postgraduate Forest Programme, Faculty of Natural and Agricultural Sciences, University of Pretoria, Private Bag X20 Hatfield, Pretoria, 0028, South Africa E-mail: michal@cmo.co.za
  • Spinelli, CNR IVALSA, Via Madonna del Piano 10, I-50019 Sesto Fiorentino (FI), Italy ORCID http://orcid.org/0000-0001-9545-1004 E-mail: spinelli@ivalsa.cnr.it (email)
article id 1618, category Research article
Miguel Genin, Mohamed Alifriqui, Abdessamad Fakhech, Mohamed Hafidi, Lahcen Ouahmane, Didier Genin. (2017). Back to forests in pre-Saharan Morocco? When prickly pear cultivation and traditional agropastoralism reduction promote argan tree regeneration. Silva Fennica vol. 51 no. 1B article id 1618. https://doi.org/10.14214/sf.1618
Keywords: arid zones; Argania spinosa; agroforestry parklands; desertification; facilitation factors; land use changes; Opuntia ficus-indica
Highlights: There was a significant positive relationship between the age of implanted prickly pear orchards and natural argan tree regeneration; This relationship is mainly associated with interconnected changes in traditional land uses and the activation of facilitation factors such as an enhancement of the soil’s organic matter and nurse plant phenomena; This example constitutes a remarkable alternative model for thinking about agricultural development while combating desertification.
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In the southwestern pre-Saharan arid zone of Morocco, the endemic argan forest (Argania spinosa) had been almost completely destroyed in the 1960s due to intensive coal mining and mixed cereal-livestock farming. These activities turned out to be unviable and a massive rural exodus occurred in the 1970s. Local populations started to develop maintenance-free prickly pear (Opuntia ficus-indica) cultivation at large scale in order to keep their land ownership rights, while reducing their traditional agropastoral activity. We conducted a survey in order to characterize the relationships between the age of prickly pear orchards and argan tree regeneration. We also explored facilitating factors, such as soil organic matter and mycorrhiza. Results showed a high positive correlation (r2 = 0.75, p < 0.001) between the age of prickly pear orchards and argan tree resprouts, but with differences depending on a continentality gradient. The soil organic matter content also showed highly significant differences (p < 0.001) depending on the age of the prickly pear plantation, while spora density did not show such differences. The recent high economic value attributed to prickly pear fruits, and to both argan and prickly pear seed oil, has given farmers the opportunity to develop a lucrative agricultural activity, while promoting the recovery of native vegetation. This situation constitutes a remarkable example of speculative agricultural development in a very harsh environment, in phase with ecological priorities for combating desertification. It could represent an alternative to the externally-generated projects sustained by high levels of public funding, with ecological, economic and social impacts which are sometimes questionable.

  • Genin, Institut de Recherche pour le Développement (IRD) & Aix-Marseille Université, Laboratoire Population, Environnement, Développement, UMR151 AMU-IRD, Marseille, France E-mail: miguel.genin@gmail.com
  • Alifriqui, Cadi Ayyad University (UCAM), Laboratoire d’Ecologie et Environnement (CNRST, URAC 32), Faculté des Sciences Semlalia, Marrakech, Morocco E-mail: alifriqui@gmail.com
  • Fakhech, Cadi Ayyad University (UCAM), Laboratoire d’Ecologie et Environnement (CNRST, URAC 32), Faculté des Sciences Semlalia, Marrakech, Morocco E-mail: abdessamad.fakhech@edu.uca.ac.ma
  • Hafidi, Cadi Ayyad University (UCAM), Laboratoire d’Ecologie et Environnement (CNRST, URAC 32), Faculté des Sciences Semlalia, Marrakech, Morocco E-mail: hafidi.ucam@gmail.com
  • Ouahmane, Cadi Ayyad University (UCAM), Laboratoire d’Ecologie et Environnement (CNRST, URAC 32), Faculté des Sciences Semlalia, Marrakech, Morocco E-mail: l.ouahmane@gmail.com
  • Genin, Institut de Recherche pour le Développement (IRD) & Aix-Marseille Université, Laboratoire Population, Environnement, Développement, UMR151 AMU-IRD, Marseille, France E-mail: didier.genin@univ-amu.fr (email)
article id 1579, category Research article
Oktay Yildiz, Ernaz Altundağ, Bilal Çeti̇n, Ş Teoman Guner, Murat Sarginci, Bulent Toprak. (2017). Afforestation restoration of saline-sodic soil in the Central Anatolian Region of Turkey using gypsum and sulfur. Silva Fennica vol. 51 no. 1B article id 1579. https://doi.org/10.14214/sf.1579
Keywords: arid zones; Elaeagnus angustifolia; Tamarix smyrnensis; Populus alba; soil amelioration; exchangeable sodium percent
Highlights: Significantly enhanced height and diameter growth of Elaeagnus angustifolia and height growth of Populus alba improved with soil chemical amendments in comparison to control; Infiltration rate on gypsum application sites was 55% higher than on sulfur application sites; Elaeagnus angustifolia survival rates with chemical treatments were 43% greater than controls; Tamarix smyrnensis had the highest mean survival rate of 80%, while Populus alba averaged 36%.
Abstract | Full text in HTML | Full text in PDF | Author Info

A significant amount of land area in the Central Anatolian Region of Turkey has saline-sodic soil properties. The aim of the current study was to use both soil amendment and tree to restore these degraded lands. The primary objective was to ameliorate soils by leaching excess sodium with gypsum and sulfur applications. Following soil treatments, salt cedar (Tamarix smyrnensis Bunge), Russian olive (Elaeagnus angustifolia L.) and silver poplar (Populus alba L.) seedlings were planted on experimental and control sites to evaluate the effects of the treatments on survival and growth of these species. In the fall of 2013, three-year-old seedlings were planted using 1.5 × 1.5 m spacing on each plot. Survival rates were determined and height and diameter were measured at the end of September 2015. Second year infiltration measurements indicated that both chemical treatments had significantly increased the infiltration capacity of the soil (P = 0.0003). Soil infiltration capacity on gypsum treated sites was about 55% higher than on sulfur sites. Following the second growing season, salt cedar had the highest survival rates of 80%. Silver poplar had 36% survival rates across the treatments. Russian olive had 50 cm height growth on both gypsum and sulfur application sites vs. only 25 cm on controls. Diameters of Russian olive on gypsum and sulfur sites were about 9.3 mm vs. 5 mm on the controls. Silver poplars on gypsum treated sites grew 42% taller than controls. Salt cedar had no significant growth responses among treatments. With appropriate soil amendments, especially gypsum, Russian olive gave the best overall two-year results.

  • Yildiz, Düzce University, Faculty of Forestry, 81620 Duzce, Turkey E-mail: oktayyildiz@duzce.edu.tr (email)
  • Altundağ, Düzce University, Faculty of Arts and Sciences, 81620 Duzce, Turkey E-mail: ernazaltundag@duzce.edu.tr
  • Çeti̇n, Düzce University, Faculty of Forestry, 81620 Duzce, Turkey E-mail: bilalcetin@duzce.edu.tr
  • Guner, Eskişehir Soil and Ecological Research Institute, General Directorate of Forestry, 06560 Yenimahalle/Ankara, Turkey E-mail: stguner@ogm.gov.tr
  • Sarginci, Düzce University, Faculty of Forestry, 81620 Duzce, Turkey E-mail: muratsarginci@duzce.edu.tr
  • Toprak, Düzce University, Faculty of Forestry, 81620 Duzce, Turkey E-mail: bulenttoprak@duzce.edu.tr
article id 1553, category Research article
Miguel Angel Salinas-Melgoza, Margaret Skutsch, Jon C. Lovett, Armonia Borrego. (2017). Carbon emissions from dryland shifting cultivation: a case study of Mexican tropical dry forest. Silva Fennica vol. 51 no. 1B article id 1553. https://doi.org/10.14214/sf.1553
Keywords: carbon stocks; slash-and-burn; swidden cultivation; REDD ; land sparing; Borlaug hypothesis
Highlights: Under REDD+, shifting cultivation should be considered degradation rather than deforestation; Carbon stocks in old fallows (>20 years) are higher than those in old growth forests which have never been used for shifting cultivation; Extending length of fallows increases rates of carbon emissions; Shortened fallow cycles result in higher carbon stocks and lower emissions at the landscape level; Cycle lengths could be optimized for carbon sequestration in a land sharing approach.
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The article considers the relation of shifting cultivation to deforestation and degradation, and hence its impacts in terms of carbon emissions and sequestration potential. There is a need to understand these relationships better in the context of international policy on Reduced Emissions from Deforestation and Forest Degradation (REDD+). The article reviews the way in which shifting cultivation has been incorporated in global and national estimations of carbon emissions, and assembles the available information on shifting cultivation in Tropical Dry Forests (TDF) in Mexico, where it is widely practiced. It then takes the case of two villages, Tonaya and El Temazcal, which lie within the basin of the River Ayuquila in Jalisco, Mexico. Field data for the typical carbon stocks and fluxes associated with shifting cultivation are compared with stocks and fluxes associated with more intensive agricultural production in the same dry tropical forest area to highlight the carbon sequestration dynamics associated with the shortening and potential lengthening of the fallow cycles. The biomass density in the shifting cultivation system observed can reach levels similar to that of old growth forests, with old fallows (>20 years) having higher carbon stocks than old growth forests. Per Mg of maize produced, the biomass-related emissions from shifting cultivation in the traditional 12 year cycle are about three times those from permanent cultivation. We did not, however, take into account the additional emissions from inputs that result from the use of fertilizers and pesticides in the case of permanent agriculture. Shortening of the fallow cycle, which is occurring in the study area as a result of government subsidies, results in higher remaining stocks of carbon and lower emissions at the landscape level.

  • Salinas-Melgoza, University of Twente, Drienerlolaan 5, 7522 NB Enschede, the Netherlands ORCID http://orcid.org/0000-0003-3209-1659 E-mail: ma.masm@gmail.com (email)
  • Skutsch, Universidad Nacional Autónoma de México (CIGA-UNAM), Antigua Carretera a Pátzcuaro No. 8701, Col. Ex-Hacienda de San José de la Huerta, Campus Morelia, C.P. 58190, Michoacán, Mexico ORCID http://orcid.org/0000-0001-6120-4945 E-mail: mskutsch@ciga.unam.mx
  • Lovett, University of Leeds, Leeds, LS2 9JT, UK E-mail: j.lovett@leeds.ac.uk
  • Borrego, CONACYT-Centro de Investigaciones en Geografía Ambiental, Antigua Carretera a Pátzcuaro No. 8701, Col. Ex-Hacienda de San José de la Huerta, Campus Morelia, C.P. 58190, Michoacán, México E-mail: aborrego@ciga.unam.mx
article id 1778, category Research article
Adriano Mazziotta, Dmitry Podkopaev, María Triviño, Kaisa Miettinen, Tähti Pohjanmies, Mikko Mönkkönen. (2017). Quantifying and resolving conservation conflicts in forest landscapes via multiobjective optimization. Silva Fennica vol. 51 no. 1 article id 1778. https://doi.org/10.14214/sf.1778
Keywords: biodiversity; ecosystem management; forestry; decision support tools; environmental conflicts; land-use planning; systematic conservation planning
Highlights: We introduce a compatibility index quantifying how targeting a management objective in the forest landscape affects another objective; To resolve conflicts we find compromise solutions minimizing the maximum deterioration among objectives; We apply our approach for a case study of forest management for biodiversity conservation and development; Multiple use management and careful planning can reduce biodiversity conflicts in forest ecosystems.
Abstract | Full text in HTML | Full text in PDF | Author Info

Environmental planning for of the maintenance of different conservation objectives should take into account multiple contrasting criteria based on alternative uses of the landscape. We develop new concepts and approaches to describe and measure conflicts among conservation objectives and for resolving them via multiobjective optimization. To measure conflicts we introduce a compatibility index that quantifies how much targeting a certain conservation objective affects the capacity of the landscape for providing another objective. To resolve such conflicts we find compromise solutions defined in terms of minimax regret, i.e. minimizing the maximum percentage of deterioration among conservation objectives. Finally, we apply our approach for a case study of management for biodiversity conservation and development in a forest landscape. We study conflicts between six different forest species, and we identify management solutions for simultaneously maintaining multiple species’ habitat while obtaining timber harvest revenues. We employ the method for resolving conflicts at a large landscape level across a long 50-years forest planning horizon. Our multiobjective approach can be an instrument for guiding hard choices in the conservation-development nexus with a perspective of developing decision support tools for land use planning. In our case study multiple use management and careful landscape level planning using our approach can reduce conflicts among biodiversity objectives and offer room for synergies in forest ecosystems.

  • Mazziotta, University of Jyväskylä, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland; Center for Macroecology Evolution and Climate, University of Copenhagen, Universitetsparken 15, DK-2100 Copenhagen, Denmark; Stockholm Resilience Centre, Stockholm University, Kräftriket 2b, 11429 Stockholm, Sweden ORCID http://orcid.org/0000-0003-2088-3798 E-mail: a_mazziotta@hotmail.com (email)
  • Podkopaev, University of Jyväskylä, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland; Systems Research Institute, Polish Academy of Sciences, Newelska 6, 01-447 Warsaw, Poland E-mail: dmitry.podkopaev@ibspan.waw.pl
  • Triviño, University of Jyväskylä, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: maria.trivino@jyu.fi
  • Miettinen, University of Jyväskylä, Faculty of Information Technology, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: kaisa.miettinen@jyu.fi
  • Pohjanmies, University of Jyväskylä, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: tahti.t.pohjanmies@jyu.fi
  • Mönkkönen, University of Jyväskylä, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: mikko.monkkonen@jyu.fi
article id 1718, category Research article
Mihails Čugunovs, Eeva-Stiina Tuittila, Lauri Mehtätalo, Laura Pekkola, Ida Sara-Aho, Jari Kouki. (2017). Variability and patterns in forest soil and vegetation characteristics after prescribed burning in clear-cuts and restoration burnings. Silva Fennica vol. 51 no. 1 article id 1718. https://doi.org/10.14214/sf.1718
Keywords: disturbance; restoration; sampling; spatial variability; prescribed fire; SOM stocks
Highlights: Soil parameter variability is similar across sites of different disturbance type; Variability of understory vegetation biomass and cover is higher and more different between sites than soil variability; Sites studied here reflect well the assumed disturbance-type gradient based on PCA; Sampling six forest sites per treatment should provide good statistical power to capture the differences in soil organic matter stocks.
Abstract | Full text in HTML | Full text in PDF | Author Info

Forest ecological restoration by burning is widely applied to promote natural, early-successional sites and increase landscape biodiversity. Burning is also used as a forest management practice to facilitate forest regeneration after clearcutting. Besides the desired goals, restoration burnings also affect soil biogeochemistry, particularly soil organic matter (SOM) and related soil carbon stocks but the long-term effects are poorly understood. However, in order to study these effects, a reliable estimate of spatial variability is first needed for effective sampling. Here we investigate spatial variability of SOM and vegetation features 13 years after burnings and in combination with variable harvest levels. We sampled four experimental sites representing distinct management and restoration treatments with an undisturbed control. While variability of vegetation cover and biomass was generally higher in disturbed sites, soil parameter variability was not different between the four sites. The joint ecological patterns of soil and vegetation parameters across the whole sample continuum support well the prior assumptions on the characteristic disturbance conditions within each of the study sites. We designed and employed statistical simulations as a means to plan prospective sampling. Sampling six forest sites for each treatment type with 30 independent soil cores per site would provide enough statistical power to adequately capture the impacts of burning on SOM based on the data we obtained here and statistical simulations. In conclusion, we argue that an informed design-based approach to documenting the ecosystem effects of forest burnings is worth applying both through obtaining new data and meta-analysing the existing.

  • Čugunovs, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: mihails.cugunovs@uef.fi (email)
  • Tuittila, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland ORCID http://orcid.org/0000-0001-8861-3167 E-mail: eeva-stiina.tuittila@uef.fi
  • Mehtätalo, University of Eastern Finland, School of Computing, Science Park, Länsikatu 15, P.O. Box 111, 80101 Joensuu, Finland ORCID http://orcid.org/0000-0002-8128-0598 E-mail: lauri.mehtatalo@uef.fi
  • Pekkola, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: laura.pekkola@gmail.com
  • Sara-Aho, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: ida.sara-aho@mhy.fi
  • Kouki, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland ORCID http://orcid.org/0000-0003-2624-8592 E-mail: jari.kouki@uef.fi
article id 1714, category Research article
Liam Donnelly, Olga M. Grant, Conor O’Reilly. (2017). Effect of deployment-type on stem growth, biomass partitioning and crown characteristics of juvenile Sitka spruce clones. Silva Fennica vol. 51 no. 1 article id 1714. https://doi.org/10.14214/sf.1714
Keywords: productivity; Picea sitchensis; biomass partitioning; clones; deployment-type
Highlights: Deployment x clone interactions reduced tree height and diameter growth in mixed plots for one clone; Height and diameter heterogeneity was significantly greater in mixed plots; Deployment-type significantly altered relationships between crown variables and competition was more asymmetric in mixed plots compared to monoclonal.
Abstract | Full text in HTML | Full text in PDF | Author Info

Competitive interactions in clonal forestry are not well understood and this needs to be addressed to develop better deployment strategies. Eight juvenile Sitka spruce (Picea sitchensis (Bong.) Carriére) clones were grown in monoclonal and clonal mixtures in a field experiment for three years to assess the effects of genetic diversity on shoot growth, above- and below-ground biomass partitioning and crown characteristics. Shoot elongation was measured throughout the growing season, while diameter was measured twice annually in May and December. After the third year, crown silhouette area was estimated from digitised images for one ramet per plot and ramets were then destructively harvested. Deployment × clone interactions were observed for tree height and diameter with reductions observed in mixed plots. Mixed plots had significantly greater height and diameter heterogeneity and more asymmetrical competition than monoclonal plots. Results from this study demonstrate that stem growth can be significantly altered when clones are planted in multi-clonal mixtures but for most clones, deployment-type will not significantly reduce their productivity.

  • Donnelly, UCD Forestry, University College Dublin, Belfield, Dublin 4, Ireland E-mail: liam.donnelly@ucdconnect.ie (email)
  • Grant, UCD Forestry, University College Dublin, Belfield, Dublin 4, Ireland E-mail: olga.grant@ucd.ie
  • O’Reilly, UCD Forestry, University College Dublin, Belfield, Dublin 4, Ireland E-mail: conor.oreilly@ucd.ie
article id 1709, category Research article
Guoping Chen, Cong Shi, Shanshan Cheng, Tiejian Zhao, Guoquan Liu, Fuchen Shi. (2017). The structure and soil characteristics of a Pinus tabuliformis planted forest after 60 years of natural development in North China. Silva Fennica vol. 51 no. 1 article id 1709. https://doi.org/10.14214/sf.1709
Keywords: species diversity; Chinese pine; plantation forest; mixed species stands; soil chemical properties; soil microbial community
Highlights: Increasing proportions of broadleaf tree species was shown to affect nutrient content of the forest floor and soil, and the soil microbial community in the process of natural development of Pinus tabuliformis planted forest. In this regard, this study can act as a reference for management of the near-natural transformation of P. tabuliformis planted forests and for the choice of the tree species used.
Abstract | Full text in HTML | Full text in PDF | Author Info

This study evaluated the transformation of a Pinus tabuliformis Carrière forest into a near-natural forest after 60 years of natural development. The structure and soil characteristics of P. tabuliformis planted forest, the near-natural forest (coniferous-broadleaved P. tabuliformis mixed forest), and secondary forest (Quercus mongolica Fisch. ex Ledeb. forest) were compared. Tree, shrub and herb species diversity of the mixed and Q. mongolica forests was higher than that of the planted P. tabuliformis forest. Examination of soil characteristics revealed that compared to the pure pine forest, nitrogen (N) and phosphorus (P) concentrations of the mixed and Q. mongolica forests increased in the forest floor and soil, but total carbon (C) concentration decreased in the forest floor, countered by increases in the soil. Furthermore, soil cation exchange capacity (CEC) and pH in the P. tabuliformis forest increased when deciduous broadleaved species were present. Total microbial biomass and bacterial biomass in the soils were greatest in the Q. mongolica forest, followed by the mixed, and then the P. tabuliformis forests. However, fungal biomass did not significantly differ among the three forests. Overall, the findings of this study suggest that different forest types can affect soil microbial biomass and community structure. Meanwhile, the natural development is recommended as a potential management alternative to near-natural transformation of a P. tabuliformis planted forest.

  • Chen, Department of Plant Biology & Ecology, College of Life Sciences, Nankai University, Weijin Road 94, Tianjin 300071, P.R. China E-mail: guopingchern@mail.nankai.edu.cn
  • Shi, Graduate School of Agriculture, Hokkaido University, Sapporo, Hokkaido 060-8689, Japan E-mail: cshi1@for.agr.hokudai.ac.jp
  • Cheng, School of Environment and Energy, Shenzhen Graduate School of Peking University, Shenzhen 518055, China E-mail: 1401213932@sz.pku.edu.cn
  • Zhao, Baxian Mountain National Nature Reserve, Tianjin 301900, China Received 29 September 2016 Revised E-mail: zhaotiejiann456@sina.com
  • Liu, Baxian Mountain National Nature Reserve, Tianjin 301900, China Received 29 September 2016 Revised E-mail: liuguoquan01@163.com
  • Shi, Department of Plant Biology & Ecology, College of Life Sciences, Nankai University, Weijin Road 94, Tianjin 300071, P.R. China E-mail: fcshi@nankai.edu.cn (email)
article id 1705, category Research article
Sulmaz Janfaza, Hamed Yousefzadeh, Seyed Mohammad Hosseini Nasr, Roberto Botta, Asad Asadi Abkenar, Daniela Torello Marinoni. (2017). Genetic diversity of Castanea sativa an endangered species in the Hyrcanian forest. Silva Fennica vol. 51 no. 1 article id 1705. https://doi.org/10.14214/sf.1705
Keywords: population structure; microsatellite markers; sweet chestnut
Highlights: This is the first report of genetic diversity of the few remaining populations of C. sativa in the southern limits of its distribution in Eurasia; Low genetic diversity and high genetic differentiation among small isolated populations of C. sativa with low geographical distance represent serious genetic erosion indicators in the Hyrcanian forest, even hinting at an ongoing extinction vortex.
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Castanea sativa Mill. is one of the most endangered tree species in Iran where it is represented by small fragmented populations in the north of the country. 18 simple sequence repeat (SSR) loci (10 nuclear and 8 chloroplastic) were used to evaluate the genetic diversity and population structure of C. sativa from the Hyrcanian forest. For nuclear SSR, the number of alleles detected per locus ranged from 1 to 5 and observed heterozygosity (HO) was between 0.125 and 1.000. Analysis of molecular variance (AMOVA) indicated a high level of variation within populations (84%) and low levels between populations (16%). Based on structure analysis, the four studied populations were divided into two main clusters that have genetic distance Fst = 0.3. The Shafaroud population was separated in the first cluster, Siyahmazgi, Qalehroudkhan and Veysroud were placed in the second cluster. The UPGMA analysis confirmed the results of Structure analysis, separating the Shafaroud population from the others. The 8 chloroplast SSR loci used to screen the populations showed no polymorphism. In General, low nuclear genetic diversity, no polymorphism in cpDNA and considerable genetic differentiation among populations in short geographical distance represent a serious genetic erosion threat for C. sativa in the Hyrcanian forest, even hinting at an ongoing extinction vortex. Therefore, due to significant decline in genetic diversity, it is essential to introduce constraints protection upon the areas of distribution of all four populations of this species in Iran.

  • Janfaza, Department of Forestry, Sari University of Agriculture Sciences and Natural Resources, Km 9 Darya Road, P.O. Box 578, Mazandaran, Iran E-mail: aristocratka_b@yahoo.com
  • Yousefzadeh, Department of Environment, Faculty of Natural Resources and Marine Science ,Tarbiat Modares University, Noor, Mazandaran, Iran E-mail: h.yousefzadeh@modares.ac.ir (email)
  • Hosseini Nasr, Department of Forestry, Sari University of Agriculture Sciences and Natural Resources, Km 9 Darya Road, P.O. Box 578, Mazandaran, Iran E-mail: s.hosseini@sanru.ac.ir
  • Botta, Dipartimento di Scienze Agrarie Forestali e Alimentari, DISAFA, Università di Torino, Via Verdi 8, 10124 Torino, Italy E-mail: roberto.botta@unito.it
  • Asadi Abkenar, Agricultural Research, Education and Extension Organization (AREEO) of IRAN, Agricultural Biotechnology Research Institute of Iran (RBRII), Branch of North Region, Guilan, Rasht, Iran E-mail: asadiabkenarasad@gmail.com
  • Marinoni, Dipartimento di Scienze Agrarie Forestali e Alimentari, DISAFA, Università di Torino, Via Verdi 8, 10124 Torino, Italy E-mail: daniela.marinoni@unito.it
article id 1704, category Research article
Inger Sundheim Fløistad, Toril Drabløs Eldhuset. (2017). Effect of photoperiod and fertilization on shoot and fine root growth in Picea abies seedlings. Silva Fennica vol. 51 no. 1 article id 1704. https://doi.org/10.14214/sf.1704
Keywords: Norway spruce; nitrogen; nutrients; potassium; root collar diameter; root scanning; short day treatment
Highlights: Effects of photoperiod and fertilization treatment on Norway spruce seedling growth were examined; Short day treatment and ordinary K:N ratio in the fertilization proved the best combination for achieving seedlings with suitable root and shoot properties for field establishment; Increased K:N ratio in the fertilization did not reduce shoot height growth.
Abstract | Full text in HTML | Full text in PDF | Author Info

Picea abies seedlings were given three different fertilization treatments in the nutrient solution by varying the potassium:nitrogen (K:N) ratios (2.5, 3.0 or 3.9 g g–1). All fertilization treatments were combined with short-day (SD) treatment or no such treatment (control). Above- and belowground growth responses in the seedlings were analyzed. The SD treatment resulted in significantly reduced shoot height, compared to untreated control, irrespective of K:N ratio. No combination of photoperiod treatment or fertilization treatment affected the root collar diameter. In the current year root fraction with diameter < 0.5 mm, the highest K:N ratio led to significantly increased root length in control plants. In each 0.1 mm root diameter class up to 0.5 mm, the highest K:N ratio significantly stimulated root growth in control plants, while the effect was less evident for SD plants. SD treatment stimulated length growth in some fine root diameter classes. We conclude that SD treatment is a good and sufficient measure to reduce height growth without compromising fine root growth of P. abies seedlings. Fertilization treatment did not significantly improve aboveground growth in SD treated seedlings, and only limited effects on root growth was seen on control plants.

  • Fløistad, Norwegian Institute of Bioeconomy Research (NIBIO), P.O. Box 115, NO-1431 Ås, Norway E-mail: inger.floistad@nibio.no (email)
  • Eldhuset, Norwegian Institute of Bioeconomy Research (NIBIO), P.O. Box 115, NO-1431 Ås, Norway E-mail: toril.eldhuset@nibio.no
article id 1694, category Research article
Katri Luostarinen, Katja Hakkarainen, Henri Kaksonen. (2017). Wood anatomy of seed and basal bud originated downy birches (Betula pubescens Ehrh.) grown at four different sites. Silva Fennica vol. 51 no. 1 article id 1694. https://doi.org/10.14214/sf.1694
Keywords: fibre; xylem; parenchyma; ray; vessel; sprout; wood structure
Highlights: Young xylem of sprouts did not clearly show more mature characteristics than that of seedlings; Marked differences in xylem structure could be observed between growing sites.
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In trees, xylem must fulfil three important tasks: conducting water to leaves, storing nutrients and water, and supporting the trunk. The origin of the trunk, i.e., seed or basal bud that forms sprouts, and the growth site may affect xylem anatomy, differences of which can affect successful growth of trees. Both seedlings and sprouts of downy birch (Betula pubescens Ehrh.) from four different growth sites with two different soil media, peat and mineral soil, were studied. The diameter of fibres and vessels and the thickness of the double fibre wall were measured, and the number of vessels, rays and axial parenchyma cells was counted. The fibre wall:lumen ratio, vessel percentage area and vessel size:number ratio were calculated. Xylem from sprouts showed only occasionally more mature characteristics than that of seedlings. The number of rays was similar at all four sites, but differences were observed in all other studied characteristics between sites, particularly if soil type was different. The vessel size and number correlated with the number of axial parenchyma cells in juvenile wood, which emphasises the importance of their connections with storage cells particularly at this stage of growth. Good water conductivity was connected with weaker wood, particularly in maturing wood.

  • Luostarinen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: katri.luostarinen@uef.fi (email)
  • Hakkarainen, Natural Resources Institute of Finland (Luke), Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: katjahak@gmail.com
  • Kaksonen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: hkaksone@gmail.com
article id 1692, category Research article
Aleksey Fedorkov, Ludmila Gutiy. (2017). Performance of lodgepole pine and Scots pine in field trials located in north-west Russia. Silva Fennica vol. 51 no. 1 article id 1692. https://doi.org/10.14214/sf.1692
Keywords: Pinus sylvestris; stem defects; mortality; Pinus contorta; stem growth
Highlights: Stem volume was bigger for lodgepole pine in comparison to local Scots pine except for the southernmost origin; The proportion of stems with no defects was lower for all lodgepole pine seed sources than for local Scots pine; Lodgepole pine stem growth traits were significantly related to latitude of seed origin.
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Mortality, stem growth and quality of lodgepole pine (Pinus contorta var. latifolia) originating from the six Swedish seed orchards and local Scots pine (Pinus sylvestris L.) were estimated in four field trials established in the Komi Republic (north-west Russia). A randomized row-plot design with 6–12 replicates of each entry was used. The tree mortality was slightly higher for Scots pine than that for lodgepole pine, except for the lodgepole pine seed sources of the southern origins with lower survival. Scots pine stem quality was better than that of lodgepole pine, but the lodgepole pine stem growth was faster except the seed source of the southernmost origin. The lodgepole pine seed sources of northern origins had better stem growth (height, diameter at breast height and volume), while the effect of latitude on the quality traits was insignificant.

  • Fedorkov, Institute of Biology, Komi Science Center, Russian Academy of Sciences, 28 Kommunisticheskaya st., Syktyvkar 167982, Russia ORCID http://orcid.org/0000-0001-7800-7534 E-mail: fedorkov@ib.komisc.ru (email)
  • Gutiy, Syktyvkar Forest Institute (branch), Saint-Petersburg State Forest Technical University, 39 Lenin st., Syktyvkar 167000, Russia E-mail: lguti@mail.ru
article id 1684, category Research article
Anna Kowalska, Jan Marek Matuszkiewicz, Jerzy Solon, Anna Kozłowska. (2017). Indicators of ancient forests in nutrient-deficient pine habitats. Silva Fennica vol. 51 no. 1 article id 1684. https://doi.org/10.14214/sf.1684
Keywords: forest continuity; central Poland; life-history traits; mixed oak-pine forests; phi coefficient; pine forests
Highlights: Distinct groups of species with a preference for ancient pine and mixed oak-pine forests can be determined; The ancient forest indicator composition in pine habitats differs remarkably from ancient forest indicators in deciduous forests; Dispersal-related traits significantly distinguish ancient forest indicators from other species found in nutrient-poor forest habitats.
Abstract | Full text in HTML | Full text in PDF | Author Info

Pine forests are common in many European regions. Nonetheless, there are only a few studies on regeneration of plant species populations in nutrient-deficient pine habitats. Ancient temperate forests are perceived to be particularly important objects of environmental conservation, due to their ability to sustain a considerable number of rare and vulnerable species. In this paper, we present indicator species of ancient pine and mixed oak-pine forests, together with their trait profiles. Phytosociological relevés were collected from mature stands in the Masuria and Kurpie regions of central Poland. Forest persistence was determined on the basis of historical maps, with the data set divided into three categories. The indicator value of species was evaluated using Tichý and Chytrý’s phi coefficient. Functional response traits of indicator species were identified. Distinct groups of species with a preference for ancient forests can be determined. The dispersal-related traits significantly distinguish ancient forest indicators from other species found in nutrient-poor forest habitats. Since the low potential for long-distance dispersal hinders the establishment of new plant populations in isolated stands, we stress the need to avoid ancient forest clearance and fragmentation of woodland; afforestation should be located in the vicinity of ancient stands. Moreover, as recent forests have turned out to support several rare plant species, to maintain phytodiversity on a landscape level a mixture of ancient and recent forests, both managed and strictly protected, is needed.

  • Kowalska, Institute of Geography and Spatial Organization, Polish Academy of Science, Twarda 51/55, 00-818 Warsaw, Poland E-mail: aniak@twarda.pan.pl (email)
  • Matuszkiewicz, Institute of Geography and Spatial Organization, Polish Academy of Science, Twarda 51/55, 00-818 Warsaw, Poland E-mail: jan.mat@twarda.pan.pl
  • Solon, Institute of Geography and Spatial Organization, Polish Academy of Science, Twarda 51/55, 00-818 Warsaw, Poland E-mail: j.solon@twarda.pan.pl
  • Kozłowska, Institute of Geography and Spatial Organization, Polish Academy of Science, Twarda 51/55, 00-818 Warsaw, Poland E-mail: a.kozl@twarda.pan.pl
article id 1689, category Research article
Juha Laitila, Eeva Lehtonen, Tapio Ranta, Perttu Anttila, Saija Rasi, Antti Asikainen. (2016). Procurement costs of cereal straw and forest chips for biorefining in South-East Finland. Silva Fennica vol. 50 no. 5 article id 1689. https://doi.org/10.14214/sf.1689
Keywords: supply chains; stumps; young stands; logging residues; delimbed stems; agrobiomass
Highlights: Procurement cost at the plant was 59 € dry tonne –1 when the annual procurement volume of biomass was 100 000 tonnes. Of that amount, the proportion of logging residues was 58.4%, stumps 24.3% and delimbed stems 17.3%; Cereal straw represents an important source of biomass in Kouvola but the cost competiveness is poor compared the procurement costs of forest chips due to high baling and transporting costs.
Abstract | Full text in HTML | Full text in PDF | Author Info

In Finland the increasing use of biofuel in transport presupposes a search for new raw material sources for biorefining. The aim of this study was, at the regional level, to compare the procurement costs of logging residues, stumps, delimbed stems and cereal straw for biorefining. The accumulation and procurement costs of forest chips and cereal straw were estimated within a 100-kilometre transporting distance via existing road network from an end-use-facility located in Kouvola in South-East Finland. The analyses were performed as simulated treatments in thinnings of young stands, cereal fields and regeneration fellings using existing productivity and cost functions and yield calculations based on crop statistics, the forest industry stand data and the sample plots data of the National Forest Inventory of Finland. Accumulation of raw material assortments and costs of production stages were defined per dry tonnes. Subsidies and raw material prices were excluded from consideration in the study. The results indicate that recovering logging residues requires lower costs than utilization of stumps, delimbed stems or cereal straw. Cereal straw represents an important source of biomass in Kouvola but the cost competiveness is poor compared the procurement costs of forest chips. When the annual procurement volume of biomass was 50 000 dry tonnes the cost at the plant was 49 € dry tonne –1 and biomass was comprised totally of logging residues. Procurement cost grew to 59 € dry tonne –1 when the annual procurement volume of biomass was doubled to 100 000 dry tonnes. Of that amount, the proportion of logging residues was 58.4%, stumps 24.3% and delimbed stems 17.3%. First cereal straw dry tonnes were delivered to end-use-facility, when procurement cost grew to 60 € dry tonne –1 and annual procurement volume of biomass was 110 000 dry tonnes.

  • Laitila, Natural Resources Institute Finland (Luke), Bio-based Business and Industry, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: juha.laitila@luke.fi (email)
  • Lehtonen, Natural Resources Institute Finland (Luke), Green Technology, Halolantie 31A, FI-71750 Maaninka, Finland E-mail: eeva.lehtonen@luke.fi
  • Ranta, Lappeenranta University of Technology, LUT School of Energy Systems, Laboratory of Bioenergy, Lönnrotinkatu 7, FI-50100 Mikkeli, Finland E-mail: tapio.ranta@luke.fi
  • Anttila, Natural Resources Institute Finland (Luke), Bio-based Business and Industry, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: perttu.anttila@luke.fi
  • Rasi, Natural Resources Institute Finland (Luke), Bio-based Business and Industry, Survontie 9A, FI-40500 Jyväskylä E-mail: saija.rasi@luke.fi
  • Asikainen, Natural Resources Institute Finland (Luke), Bio-based Business and Industry, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: antti.asikainen@luke.fi
article id 1680, category Research article
Liisa Kulmala, Indre Žliobaitė, Eero Nikinmaa, Pekka Nöjd, Pasi Kolari, Kourosh Kabiri Koupaei, Jaakko Hollmén, Harri Mäkinen. (2016). Environmental control of growth variation in a boreal Scots pine stand – a data-driven approach. Silva Fennica vol. 50 no. 5 article id 1680. https://doi.org/10.14214/sf.1680
Keywords: height growth; diameter growth; Gross primary production; turgor pressure; predictive modelling; Least Angle Regression
Highlights: High water potential and carbon gain during bud forming favoured height growth; High water potential during the elongation period favoured height growth; A spring with high carbon gain favoured diameter growth; The obtained regression models had generally low generalization performance.
Abstract | Full text in HTML | Full text in PDF | Author Info

Despite the numerous studies on year-to-year variation of tree growth, the physiological mechanisms controlling annual variation in growth are still not understood in detail. We studied the applicability of data-driven approach i.e. different regression models in analysing high-dimensional data set including continuous and comprehensive measurements over meteorology, ecosystem-scale water and carbon fluxes and the annual variation in the growth of app. 50-year-old Scots pine stand in southern Finland. Even though our dataset covered only 16 years, it is the most extensive collection of interactions between a Scots pine ecosystem and atmosphere. The analysis revealed that height growth was favoured by high water potential of the tree and carbon gain during the bud forming period and high water potential during the elongation period. Diameter growth seemed to be favoured by a winter with high precipitation and deep snow cover and a spring with high carbon gain. The obtained models had low generalization performance and they would require more evaluation and iterative validation to achieve credibility perhaps as a mixture of data-driven and first principle modeling approaches.

  • Kulmala, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: liisa.kulmala@helsinki.fi (email)
  • Žliobaitė, Aalto University, Department of Computer Science and Helsinki Institute for Information Technology, P.O. Box 11000, FI-00076 Aalto, Finland; University of Helsinki, Department of Geosciences and Geography, P.O. Box 64, FI-00014 University of Helsinki, Finland E-mail: zliobaite@gmail.com
  • Nikinmaa, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: eero.nikinmaa@helsinki.fi
  • Nöjd, Natural Resources Institute Finland (Luke), Bio-based business and industry, Tietotie 2, FI-02150 Espoo, Finland E-mail: pekka.nojd@luke.fi
  • Kolari, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland; University of Helsinki, Department of Physics, P.O. Box 64, FI-00014 University of Helsinki, Finland E-mail: pasi.kolari@helsinki.fi
  • Kabiri Koupaei, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: kourosh.kabiri@helsinki.fi
  • Hollmén, Aalto University, Department of Computer Science and Helsinki Institute for Information Technology, P.O. Box 11000, FI-00076 Aalto, Finland; University of Helsinki, Department of Geosciences and Geography, P.O. Box 64, FI-00014 University of Helsinki, Finland E-mail: jaakko.hollmen@aalto.fi
  • Mäkinen, Natural Resources Institute Finland (Luke), Bio-based business and industry, Tietotie 2, FI-02150 Espoo, Finland E-mail: harri.makinen@luke.fi
article id 1672, category Research article
Maiju Peura, María Triviño, Adriano Mazziotta, Dmitry Podkopaev, Artti Juutinen, Mikko Mönkkönen. (2016). Managing boreal forests for the simultaneous production of collectable goods and timber revenues. Silva Fennica vol. 50 no. 5 article id 1672. https://doi.org/10.14214/sf.1672
Keywords: forest management; multifunctional forestry; mushroom; trade-offs; wildberry
Highlights: We found a strong conflict between bilberry production and timber revenues, resulting in large losses of timber revenues when increasing bilberry production; The conflicts between other collectables (cowberry, cep) and timber production were relatively small; With careful forest planning, there is potential to simultaneously produce high levels of collectable goods and timber revenues in the landscape.
Abstract | Full text in HTML | Full text in PDF | Author Info

Timber production is an economically important provisioning ecosystem service in forests, but is often in conflict with the provision of other ecosystem services. In multifunctional forestry, the production of timber and non-timber ecosystem services should coexist in the same landscape. To this end, we explored the capacity of a boreal landscape to simultaneously produce collectable goods bilberry (Vaccimium myrtillus L.), cowberry (Vaccinium vitis-idaea L.) and cep (Boletus edulis Bull.) alongside timber revenues. We also identified optimal forest management plans to achieve this. Furthermore, we analyzed trade-offs between collectable good yields and timber production, as well as between their economic values. We ran forest growth simulations under seven alternative management regimes at a landscape level across 50-year planning horizons. Then, we used multi-objective optimization to explore trade-offs and identify optimal forest management plans. The results showed that the strongest trade-off was between bilberry and timber production, resulting in a large loss in timber revenues for a gain in bilberry production. However, the conflicts between other collectables and timber production were relatively small: it was possible to increase the provision of collectable goods 4–15% with small reductions (35%) from timber revenues. With careful forest planning, there is the potential to simultaneously produce high levels of collectable goods and timber revenues in the landscape.

  • Peura, University of Jyvaskyla, Department of Biological and Environmental Sciences, P.O. Box 35, FI-40014, University of Jyväskylä, Finland E-mail: maiju.h.peura@jyu.fi (email)
  • Triviño, University of Jyvaskyla, Department of Biological and Environmental Sciences, P.O. Box 35, FI-40014, University of Jyväskylä, Finland E-mail: maria.trivino@jyu.fi
  • Mazziotta, University of Jyvaskyla, Department of Biological and Environmental Sciences, P.O. Box 35, FI-40014, University of Jyväskylä, Finland; Center for Macroecology, Evolution and Climate, Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK-2100 Copenhagen, Denmark E-mail: adriano.mazziotta@snm.ku.dk
  • Podkopaev, University of Jyvaskyla, Department of Biological and Environmental Sciences, P.O. Box 35, FI-40014, University of Jyväskylä, Finland; Systems Research Institute, Polish Academy of Sciences, Newelska 6, 01-447 Warsaw, Poland E-mail: dmitry.podkopaev@gmail.com
  • Juutinen, Natural Resources Institute Finland (Luke), Economics and Society, P.O. Box 413, FI-90014 University of Oulu, Finland; University of Oulu, Department of Economics, P.O. Box 4600, FI-90014 University of Oulu, Finland E-mail: artti.juutinen@luke.fi
  • Mönkkönen, University of Jyvaskyla, Department of Biological and Environmental Sciences, P.O. Box 35, FI-40014, University of Jyväskylä, Finland E-mail: mikko.monkkonen@jyu.fi
article id 1615, category Research article
Minna Blomqvist, Päivi Lyytikäinen-Saarenmaa, Tuula Kantola, Maiju Kosunen, Mervi Talvitie, Markus Holopainen. (2016). Impacts of natural enemies and stand characteristics on cocoon mortality of the pine sawfly Diprion pini in a Fennoscandian boreal forest. Silva Fennica vol. 50 no. 5 article id 1615. https://doi.org/10.14214/sf.1615
Keywords: needle loss; Diprionidae; population dynamics; forest floor vegetation; parasitism; predation
Highlights: Annual cocoon mortality caused by natural enemies varied between 66% and 80% during the six-year study period, most of it caused by the family Ichneumonidae; Basal area, and coverage of lichen (Lichenes) and lingonberry (Vaccinium vitis-idaea L.) best explained cocoon parasitism and predation; Combination of suitable stand characteristics, abiotic environmental factors, and incomplete control by natural enemies enabled pest species to extend its gradation phase.
Abstract | Full text in HTML | Full text in PDF | Author Info

We investigated the impact of natural enemies on the cocoon mortality of the common pine sawfly (Diprion pini L.) during a six-year period in eastern Finland. The enemies were classified into parasitoids (insect families Chalcidoidea, Ichneumonidae, and Tachinidae), and predators (birds, small mammals, and insect families Elateridae and Carabidae). The appearance of D. pini was estimated as the intensity of annual defoliation. The impact of stand characteristics on the performance of parasitoids and predators was also investigated. Influence of the natural enemy complex on cocoon mortality of D. pini was nearly stable, but defoliation intensity slowly declined towards the end of the study period. Annual cocoon mortality by natural enemies varied between 66% and 80%. Our results verified that the most significant mortality factors were ichneumonid parasitoids and small mammals. Random Forest classification indicated that stand characteristics, such as basal area, and coverage of lichen (Lichenes) and lingonberry (Vaccinium vitis-idaea L.) affected the performance of parasites and predators. We suggest that a combination of optimal stand characteristics, abiotic environmental factors and mild to moderate control by natural enemies acted as drivers, which drove the pine sawfly population to extended gradation. For future forest health management, detailed information on abiotic and biotic regulating factors, along with long-term monitoring campaigns for conifer sawflies are needed to adapt Fennoscandian forests to altered climatic and silvicultural conditions.

  • Blomqvist, University of Helsinki, Department of Forest Sciences, P.O. Box 27 (Latokartanonkaari 7), FI-00014 University of Helsinki, Finland ORCID http://orcid.org/0000-0003-2328-8839 E-mail: minna.blomqvist@helsinki.fi (email)
  • Lyytikäinen-Saarenmaa, University of Helsinki, Department of Forest Sciences, P.O. Box 27 (Latokartanonkaari 7), FI-00014 University of Helsinki, Finland E-mail: paivi.lyytikainen-saarenmaa@helsinki.fi
  • Kantola, Knowledge Engineering Laboratory, Department of Entomology, Texas A&M University, College Station, TX 77843-2475, USA E-mail: tuula.kantola@helsinki.fi
  • Kosunen, University of Helsinki, Department of Forest Sciences, P.O. Box 27 (Latokartanonkaari 7), FI-00014 University of Helsinki, Finland E-mail: maiju.kosunen@helsinki.fi
  • Talvitie, University of Helsinki, Department of Forest Sciences, P.O. Box 27 (Latokartanonkaari 7), FI-00014 University of Helsinki, Finland E-mail: mervi.talvitie@dnainternet.net
  • Holopainen, University of Helsinki, Department of Forest Sciences, P.O. Box 27 (Latokartanonkaari 7), FI-00014 University of Helsinki, Finland E-mail: markus.holopainen@helsinki.fi
article id 1566, category Research article
Valdir Marcos Stefenon, Jordana Caroline Nagel, Igor Poletto. (2016). Evidences of genetic bottleneck and fitness decline in Luehea divaricata populations from southern Brazil. Silva Fennica vol. 50 no. 5 article id 1566. https://doi.org/10.14214/sf.1566
Keywords: demographic history; effective population size; genetic resources conservation; Pampa biome
Highlights: Signatures of genetic bottlenecks and reduction of populations’ fitness were observed in populations of Luehea divaricata in southern Brazil; Lower levels of observed heterozygosity are correlated with populations’ fitness, decreasing germination capacity and increasing the proportion of anomalous germinated plantlets; Promoting the connection among populations is proposed as a key strategy towards conservation of L. divaricata genetic resources in its southernmost distribution range.
Abstract | Full text in HTML | Full text in PDF | Author Info

Extant populations growing in regions that were refugia during the last glacial period are expected to show higher genetic diversity than populations that moved from these refugia into new areas in higher latitudes. Such new populations likely faced harsher climatic conditions, being established with reduced population size and experiencing the effects of genetic bottlenecks. In this study we employed data from nuclear SSR markers for detecting molecular signatures of genetic bottlenecks, and germination experiments to evaluate reduction of populations’ fitness in natural populations of Luehea divaricata Mart. et Zucc., growing in the southern range of the species distribution (around 30°S latitude). Signatures of genetic bottlenecks and reduction of populations’ fitness were observed in all populations. Lower levels of observed heterozygosity are correlated with populations’ fitness, decreasing germination capacity and increasing the proportion of anomalous germinated plantlets. Promoting the connection among populations is proposed as a key strategy towards conservation of L. divaricata genetic resources in its southernmost distribution range. The offspring from crosses among populations would significantly increase the observed heterozygosity and fitness of multiple populations.

  • Stefenon, Nucleus of Genomics and Molecular Ecology, Interdisciplinary Center of Biotechnological Research, Universidade Federal do Pampa, BR290 km, 97300-000, São Gabriel, RS, Brazil E-mail: valdirstefenon@unipampa.edu.br (email)
  • Nagel, Nucleus of Genomics and Molecular Ecology, Interdisciplinary Center of Biotechnological Research, Universidade Federal do Pampa, BR290 km, 97300-000, São Gabriel, RS, Brazil E-mail: jordana.nagel@yahoo.com.br
  • Poletto, Laboratory of Plant Protection, Universidade Federal do Pampa, BR290 km, 97300-000, São Gabriel, RS, Brazil E-mail: igorpoletto@unipampa.edu.br
article id 1687, category Research article
Hannu Hökkä, Jori Uusitalo, Harri Lindeman, Jari Ala-Ilomäki. (2016). Performance of weather parameters in predicting growing season water table depth variations on drained forested peatlands – a case study from southern Finland. Silva Fennica vol. 50 no. 4 article id 1687. https://doi.org/10.14214/sf.1687
Keywords: peatland; precipitation; bearing capacity; evapotranspiration; mixed linear model; water table depth
Highlights: Four-week precipitation and evapotranspiration explain much of drained peatland water table depth variation during a growing season.
Abstract | Full text in HTML | Full text in PDF | Author Info

The amount of water in peat soil is one factor affecting its bearing capacity, which is a crucial aspect in planning peatland timber harvesting operations. We studied the influence of weather variables on the variation of drained peatland growing season water conditions, here the ground water table depth (WTD). WTD was manually monitored four times in 2014 and three times in 2015 in 10–30 sample plots located in four drained peatland forests in south-western Finland. For each peatland, precipitation and evapotranspiration were calculated from the records of the nearest Finnish Meteorological Institute field stations covering periods from one day to four weeks preceding the WTD monitoring date. A mixed linear model was constructed to investigate the impact of the weather parameters on WTD. Precipitation of the previous four–week period was the most important explanatory variable. The four-week evapotranspiration amount was interacting with the Julian day showing a greater effect in late summer. Other variables influencing WTD were stand volume within the three-metre radius sample plot and distance from nearest ditch. Our results show the potential of weather parameters, specifically that of the previous four-week precipitation and evapotranspiration, for predicting drained peatland water table depth variation and subsequently, the possibility to develop a more general empirical model to assist planning of harvesting operations on drained peatlands.

  • Hökkä, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Paavo Havaksen tie 3, FI-90014 Oulun yliopisto, Finland E-mail: hannu.hokka@luke.fi (email)
  • Uusitalo, Natural Resources Institute Finland (Luke), Green Technology, Kaironiementie 15, FI-39700 Parkano, Finland E-mail: jori.uusitalo@luke.fi
  • Lindeman, Natural Resources Institute Finland (Luke), Green Technology, Kaironiementie 15, FI-39700 Parkano, Finland E-mail: harri.lindeman@luke.fi
  • Ala-Ilomäki, Natural Resources Institute Finland (Luke), Green Technology, Jokiniemenkuja 1, FI-01370 Vantaa, Finland E-mail: jari.ala-ilomaki@luke.fi
article id 1682, category Research article
Jiří Korecký, Yousry A. El-Kassaby. (2016). Pollination dynamics variation in a Douglas-fir seed orchard as revealed by microsatellite analysis. Silva Fennica vol. 50 no. 4 article id 1682. https://doi.org/10.14214/sf.1682
Keywords: selfing; Pseudotsuga menziesii (Mirb.) Franco; genetic quality; pedigree reconstruction; gametic contribution; pollen contamination
Highlights: Important characteristics such as parental reproductive success, pollen contamination, and selfing rate in the second generation Douglas-fir seed orchard have been estimated; Since this research is a part of a multi-year study, outputs were compared to those from two other years; Results are in line and differences in pollination dynamics across years are attributable to the various crop management practices.
Abstract | Full text in HTML | Full text in PDF | Author Info

As part of a multi-year monitoring study of pollination dynamics in a second generation Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) seed orchard, we estimated with the aid of eight microsatellite markers three important reproductive biology characteristics affecting the genetic worth and diversity of seed crops; namely parental reproductive success, pollen contamination, and selfing rate. The obtained results were compared to those from two previous years to gauge appropriate seed crop management practices and ultimately allow approximate generalization of seed crop genetic quality. We determined that 80% of parental gametes were produced by 52% of the parents, 13% of paternal gametes resulted from pollen contamination (i.e., gene flow), and 12% of the seed were the product of selfing. The obtained results were in line with those observed for 2005 and 2009 where 80% of gametes being produced by 37–48% of the parents, 10–18% pollen contamination, and 15–17% selfing rate. The observed reproductive biology parameters differences are attributable to the various crop management practices implemented (i.e., bloom delay and supplemental-mass-pollination) across years and calls for justification due to the observed minimal differences on seed crops genetic quality.

  • Korecký, Department of Genetics and Physiology of Forest Trees, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Praha 6, 165 21, Czech Republic ORCID http://orcid.org/0000-0001-7859-1750 E-mail: korecky@fld.czu.cz (email)
  • El-Kassaby, Department of Forest and Conservation Sciences, Faculty of Forestry, University of British Columbia, Vancouver, BC, V6T 1Z4, Canada ORCID http://orcid.org/0000-0002-4887-8977 E-mail: y.el-kassaby@ubc.ca
article id 1663, category Research article
Marta Kempf, Monika Konnert. (2016). Distribution of genetic diversity in Fagus sylvatica at the north-eastern edge of the natural range. Silva Fennica vol. 50 no. 4 article id 1663. https://doi.org/10.14214/sf.1663
Keywords: Poland; nSSR markers; European beech; limit of natural distribution
Highlights: European beech at the north-eastern edge of its natural range in Poland have a high level of genetic variation, similar to the populations from Central Europe; The differences between the beech provenances from the two centres in Poland, which were previously identified on the basis of pollen analyses and phenotypic traits, have now been genetically confirmed.
Abstract | Full text in HTML | Full text in PDF | Author Info

An understanding of the genetic variation of the beech, especially at the edge of its natural distribution, is essential because of the change in natural distribution of the species resulting from changing climatic conditions. The main aim of the study was to determine the level of genetic diversity of European beech at the north-eastern edge of its natural range. The other aim was to check the genetic variation of beech from the two centres, the north and the south of Poland, which were identified in previous findings based on pollen analyses and phenotypic traits. The research material was the progeny of twelve beech provenances. The genetic structure of the populations was determined by ten highly variable microsatellite DNA loci. The results confirmed the high genetic diversity of beech at the north-eastern edge of its natural distribution, which infers the probability of their good adaptation to the changing climate and an extension of the range. Genetic analyses confirmed the existence of two genetic centres for beech in Poland. The populations from south-eastern Poland had a slightly higher diversity than the populations from the north-western area, which may indicate that the colonisation of Poland occurred by two routes. The results are important for creating the borders of the provenance regions and for limiting the transfer of seeds and seedlings. The choice of forest reproductive material, based on the knowledge of genetic diversity, is very important for the stability of future forests.

  • Kempf, Department of Genetics and Forest Tree Breeding, Faculty of Forestry, University of Agriculture in Krakow, Al. 29-listopada 46, 31–425 Krakow E-mail: m.kempf@ur.krakow.pl (email)
  • Konnert, Bavarian Office for Forest Seeding and Planting, Forstamtsplatz 1, 83317 Teisendorf, Germany E-mail: monika.konnert@asp.bayern.de
article id 1628, category Research article
Jürgen Aosaar, Ülo Mander, Mats Varik, Hardo Becker, Gunnar Morozov, Martin Maddison, Veiko Uri. (2016). Biomass production and nitrogen balance of naturally afforested silver birch (Betula pendula Roth.) stand in Estonia. Silva Fennica vol. 50 no. 4 article id 1628. https://doi.org/10.14214/sf.1628
Keywords: aboveground biomass; birch; afforestation; soil carbon; land use change; agricultural land; nitrogen budget
Highlights: Leafless aboveground biomass of the 17-year-old natural silver birch stand growing in abandoned agricultural land reached 94 Mg ha–1; The largest fluxes in N budget were net nitrogen mineralization and gaseous N2-N emission; Nitrogen leaching was low; Soil N content increased with the stand age, soil C content remained stable; N2O and N2 fluxes in boreal deciduous forest were analysed.
Abstract | Full text in HTML | Full text in PDF | Author Info

Silver birch (Betula pendula Roth.) is one of the main pioneer tree species occupying large areas of abandoned agricultural lands under natural succession in Estonia. We estimated aboveground biomass (AGB) dynamics during 17 growing seasons, and analysed soil nitrogen (N) and carbon (C) dynamics for 10 year period in a silver birch stand growing on former arable land. Main N fluxes were estimated and nitrogen budget for 10-year-old stand was compiled. The leafless AGB and stem mass of the stand at the age of 17-years were 94 and 76 Mg ha–1 respectively. The current annual increment (CAI) of stemwood fluctuated, peaking at 10 Mg ha–1 yr–1 at the age of 15 years; the mean annual increment (MAI) fluctuated at around 4–5 Mg ha–1. The annual leaf mass of the stand stabilised at around 3 Mg ha–1 yr–1. The stand density decreased from 11600 to 2700 trees ha–1 in the 8- and 17-year-old stand, respectively. The largest fluxes in N budget were net nitrogen mineralization and gaseous N2-N emission. The estimated fluxes of N2O and N2 were 0.12 and 83 kg ha–1 yr–1, respectively; N leaching was negligible. Nitrogen retranslocation from senescing leaves was approximately 45 kg ha–1, N was mainly retranslocated into stembark. The N content in the upper 0–10 cm soil layer increased significantly (145 kg ha–1) from 2004 to 2014; soil C content remained stable. Both the woody biomass dynamics and the N cycling of the stand witness the potential for bioenergetics of such ecosystems.

  • Aosaar, Estonian University of Life Sciences, Institute of Forestry and Rural Engineering, Kreutzwaldi 1, 51014 Tartu, Estonia E-mail: jyrgen.aosaar@emu.ee (email)
  • Mander, University of Tartu, Institute of Ecology and Earth Sciences, Ülikooli 18, 50090 Tartu, Estonia E-mail: ulo.mander@ut.ee
  • Varik, Estonian University of Life Sciences, Institute of Forestry and Rural Engineering, Kreutzwaldi 1, 51014 Tartu, Estonia E-mail: mats.varik@emu.ee
  • Becker, Estonian University of Life Sciences, Institute of Forestry and Rural Engineering, Kreutzwaldi 1, 51014 Tartu, Estonia E-mail: hardo.becker@emu.ee
  • Morozov, Estonian University of Life Sciences, Institute of Forestry and Rural Engineering, Kreutzwaldi 1, 51014 Tartu, Estonia E-mail: gunnar.morozov@emu.ee
  • Maddison, University of Tartu, Institute of Ecology and Earth Sciences, Ülikooli 18, 50090 Tartu, Estonia E-mail: martin.maddison@ut.ee
  • Uri, Estonian University of Life Sciences, Institute of Forestry and Rural Engineering, Kreutzwaldi 1, 51014 Tartu, Estonia E-mail: veiko.uri@emu.ee
article id 1622, category Research article
Lingbo Dong, Pete Bettinger, Zhaogang Liu, Huiyan Qin, Yinghui Zhao. (2016). Evaluating the neighborhood, hybrid and reversion search techniques of a simulated annealing algorithm in solving forest spatial harvest scheduling problems. Silva Fennica vol. 50 no. 4 article id 1622. https://doi.org/10.14214/sf.1622
Keywords: spatial harvest scheduling; metaheuristics; combinatorial optimization; neighborhood search; reversion search
Highlights: The performances of neighborhood, hybrid and reversion search strategies of simulated annealing were evaluated when implemented with a forest spatial harvest scheduling problem; The performances of alternative search strategies of simulated annealing were all systematic and clear better than the conventional strategy; The reversion techniques were significant superior to the other search strategies in solving forest spatial harvest scheduling problems.
Abstract | Full text in HTML | Full text in PDF | Author Info

Heuristic techniques have been increasingly used to address the complex forest planning problems over the last few decades. However, heuristics only can provide acceptable solutions to difficult problems, rather than guarantee that the optimal solution will be located. The strategies of neighborhood, hybrid and reversion search processes have been proved to be effective in improving the quality of heuristic results, as suggested recently in the literature. The overall aims of this paper were therefore to systematically evaluate the performances of these enhanced techniques when implemented with a simulated annealing algorithm. Five enhanced techniques were developed using different strategies for generating candidate solutions. These were then compared to the conventional search strategy that employed 1-opt moves (Strategy 1) alone. The five search strategies are classified into three categories: i) neighborhood search techniques that only used the change version of 2-opt moves (Strategy 2); ii) self-hybrid search techniques that oscillate between 1-opt moves and the change version of 2-opt moves (Strategy 3), or the exchange version of 2-opt moves (Strategy 4); iii) reversion search techniques that utilize 1-opt moves and the change version of 2-opt moves (Strategy 5) or the exchange version of 2-opt moves (Strategy 6). We found that the performances of all the enhanced search techniques of simulated annealing were systematic and often clear better than conventional search strategy, however the required computational time was significantly increased. For a minimization planning problem, Strategy 6 produced the lowest mean objective function values, which were less than 1% of the means developed using conventional search strategy. Although Strategy 6 performed very well, the other search strategies should not be neglected because they also have the potential to produce high-quality solutions.

  • Dong, Department of Forest Management, College of Forestry, Northeast Forestry University, Harbin 150040, China E-mail: farrell0503@126.com
  • Bettinger, Warnell School of Forestry and Natural Resources, University of Georgia, Athens 30602, GA, USA E-mail: pbettinger@warnell.uga.edu
  • Liu, Department of Forest Management, College of Forestry, Northeast Forestry University, Harbin 150040, China E-mail: lzg19700602@163.com (email)
  • Qin, Department of Forestry Economic, College of Economic & Management, Northeast Forestry University, Harbin 150040, China E-mail: huiyanqin@hotmail.com
  • Zhao, Department of Forest Management, College of Forestry, Northeast Forestry University, Harbin 150040, China E-mail: zyinghui0925@126.com
article id 1611, category Research article
Ilpo Ervasti. (2016). Wood fiber contents of different materials in the paper industry material chain expressed in roundwood equivalents (RWEs). Silva Fennica vol. 50 no. 4 article id 1611. https://doi.org/10.14214/sf.1611
Keywords: pulpwood; paper production; raw materials; roundwood equivalent (RWE); wood pulp; recycled fiber; recovered paper; material recycling
Highlights: The paper industry material recycling chain consists of different material streams; Consumed paper products form a huge fiber source for industry. However, a reliable measuring system should be developed to quantify these fiber volumes; Converting factors are introduced to quantify the wood fiber contents of different paper industry-related materials of the chain, namely, wood pulp, paper and recovered paper; By using these conversion factors it is possible to convert for example global recovered paper utilization volumes into wood fiber volumes expressed in roundwood equivalents (RWEs).
Abstract | Full text in HTML | Full text in PDF | Author Info

At present, there are no means for reliably comparing the wood fiber contents of different material streams within the paper industry material chain with each other. The aim of this article is to introduce conversion factors that make it possible to quantify the volume of wood expressed in roundwood equivalent (RWE) values for different paper industry-related materials in the material chain. These conversion factors apply to wood pulp, paper, and recovered paper. European data are used in quantifying the paper industry material streams and calculating the RWE conversion factors. The introduced conversion factors can be used to estimate RWE volumes at a global scale. With assumption that paper recycling did not occur and that paper production volume remained unchanged, an additional volume of 666 million m3 RWEs would be required globally per annum to produce 167 million tons of virgin wood pulp to replace 222 million tons of recovered paper utilized by the paper industry in 2010. This volume is approximately the same as 1.6 times the total removal of wood in Europe (EU27), or the total annual removal of wood in the USA, Canada, and Brazil combined.

  • Ervasti, Aalto University, Industrial Engineering and Management, Maarintie 8, 02150 Espoo, Finland E-mail: ilervasti@gmail.com (email)
article id 1588, category Research article
Kouakou Laurent Kouakou, Jonas Patrick Dao, Kouadio Ignace Kouassi, Manehonon Martine Beugré , Mongomaké Koné, Jean-Pierre Baudoin, Irié Arsene Zoro Bi. (2016). Propagation of Garcinia kola (Heckel) by stem and root cuttings. Silva Fennica vol. 50 no. 4 article id 1588. https://doi.org/10.14214/sf.1588
Keywords: coppicing; vegetative propagation; bitter kola; non-mist polypropagators
Highlights: Roots and branches cuttings from mature forest trees are difficult to sprout; The highest mean numbers of roots and buds produced were obtained with softwood cuttings; The aqueous IBA treatment was more effective than the powder in promoting rooting and root development; Non-mist poly-propagator gave the best propagation results; Seedling stockplants cut above node 1 promoted most vigorous shoots production.
Abstract | Full text in HTML | Full text in PDF | Author Info

The availability of appropriate propagation techniques is a major constraint to the domestication of the forest trees widely used by rural communities; such as Garcinia kola (Heckel). This study tested the ability of root and stem cuttings to regenerate vegetatively when treated with indole-3-butyric acid (IBA), and set in non-mist poly-propagator within a shaded nursery. It found that G. kola is amenable to propagation by softwood stem cuttings. Attention was given to the effect of cutting age (softwood, semi hardwood) and IBA application with regard to the sprouting and rooting efficiency. Bud and leaf emergence time were also investigated, as was the coppicing ability of the stump. Results revealed that roots and branches from mature forest trees did not sprout under any culture conditions. The highest percentages of rooting (70–85%) were obtained with softwood cuttings set in non-mist poly-propagator, regardless of hormonal treatment. The mean numbers of buds (2.9 ± 0.4) and roots (2.6 ± 0.1) produced by softwood cuttings was significantly greater than those obtained in the Control. The best average emergence time of buds (25.1 days ± 9.3) and leaves (36.8days ± 8.4) was obtained with cuttings treated with IBA and set in non-mist poly-propagator against, 54.4 ± 12.5 and 72.6 ± 3.4 days respectively for the Control. In general, non-mist poly-propagator gave the best propagation results. When coppiced, the shoots emerging from stumps with one node were the most vigorous.

  • Kouakou, Université Nangui Abrogoua, UFR/SN, Laboratoire de Génomique Fonctionnelle et Amélioration génétique, 02 BP 801 Abidjan, Côte d’Ivoire; Université Nangui Abrogoua, UFR Sciences de la Nature, Laboratoire de Biologie et Amélioration des Productions Végétales, 02 BP 801 Abidjan 02, Côte d’Ivoire E-mail: kk_laurent@yahoo.fr (email)
  • Dao, Université Nangui Abrogoua, UFR/SN, Laboratoire de Génomique Fonctionnelle et Amélioration génétique, 02 BP 801 Abidjan, Côte d’Ivoire E-mail: daojonas@hotmail.fr
  • Kouassi, Université Nangui Abrogoua, UFR/SN, Laboratoire de Génomique Fonctionnelle et Amélioration génétique, 02 BP 801 Abidjan, Côte d’Ivoire E-mail: kouadioignace@yahoo.fr
  • Beugré , Université Nangui Abrogoua, UFR Sciences de la Nature, Laboratoire de Biologie et Amélioration des Productions Végétales, 02 BP 801 Abidjan 02, Côte d’Ivoire E-mail: beugremartine@yahoo.fr
  • Koné,  Université Nangui Abrogoua, UFR Sciences de la Nature, Laboratoire de Biologie et Amélioration des Productions Végétales, 02 BP 801 Abidjan 02, Côte d’Ivoire E-mail: babadaoudi@gmail.com
  • Baudoin, Université de Liège, Gembloux Agro-Bio Tech, Laboratoire Agroécologie tropicale et Horticulture, Passage des Déportés, 2 B 5030 Gembloux, Belgique E-mail: jean-pierre.baudoin@ulg.ac.be
  • Zoro Bi, Université Nangui Abrogoua, UFR/SN, Laboratoire de Génomique Fonctionnelle et Amélioration génétique, 02 BP 801 Abidjan, Côte d’Ivoire E-mail: banhiakalou@yahoo.fr
article id 1567, category Research article
Eetu Kotivuori, Lauri Korhonen, Petteri Packalen. (2016). Nationwide airborne laser scanning based models for volume, biomass and dominant height in Finland. Silva Fennica vol. 50 no. 4 article id 1567. https://doi.org/10.14214/sf.1567
Keywords: forest inventory; LIDAR; regression analysis; remote sensing; calibration; area-based approach; mixed-effect models
Highlights: Pooled data from nine inventory projects in Finland were used to create nationwide laser-based regression models for dominant height, volume and biomass; Volume and biomass models provided regionally different means than real means, but for dominant height the mean difference was small; The accuracy of general volume predictions was nevertheless comparable to relascope-based field inventory by compartments.
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The aim of this study was to examine how well stem volume, above-ground biomass and dominant height can be predicted using nationwide airborne laser scanning (ALS) based regression models. The study material consisted of nine practical ALS inventory projects taken from different parts of Finland. We used field sample plots and airborne laser scanning data to create nationwide and regional models for each response variable. The final models had one or two ALS predictors, which were chosen based on the root mean square error (RMSE), and cross-validated. Finally, we tested how much predictions would improve if the nationwide models were calibrated with a small number of regional sample plots. Although forest structures differ among different parts of Finland, the nationwide volume and biomass models performed quite well (leave-inventory-area-out RMSE 22.3% to 33.8%, mean difference [MD] –13.8% to 18.7%) compared with regional models (leave-plot-out RMSE 20.2% to 26.8%). However, the nationwide dominant height model (RMSE 5.4% to 7.7%, MD –2.0% to 2.8%, with the exception of the Tornio region – RMSE 11.4%, MD –9.1%) performed nearly as well as the regional models (RMSE 5.2% to 6.7%). The results show that the nationwide volume and biomass models provided different means than real means at regional level, because forest structure and ALS device have a considerable effect on the predictions. Large MDs appeared especially in northern Finland. Local calibration decreased the MD and RMSE of volume and biomass models. However, the nationwide dominant height model did not benefit much from calibration.

  • Kotivuori, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: eetu.kotivuori@uef.fi (email)
  • Korhonen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: lauri.korhonen@uef.fi
  • Packalen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: petteri.packalen@uef.fi
article id 1564, category Research article
Stanislav Vacek, Zdeněk Vacek, Lukáš Bílek, Jaroslav Simon, Jiří Remeš, Iva Hůnová, Jan Král, Tereza Putalová, Miroslav Mikeska. (2016). Structure, regeneration and growth of Scots pine (Pinus sylvestris L.) stands with respect to changing climate and environmental pollution. Silva Fennica vol. 50 no. 4 article id 1564. https://doi.org/10.14214/sf.1564
Keywords: Pinus sylvestris; natural regeneration; climate; stand development; production; air-pollution
Highlights: Pine forest stands showed positive development of stand structural characteristics related to their diversity, number of regeneration individuals and growth characteristics; Tree-ring width was positively correlated with precipitation, while it was negatively correlated with temperature in growing seasons; Mean NOx concentrations showed positive effect on radial growth of pine; Serious defoliation was caused by SO2 concentrations and N deposition in combination with extreme climate events.
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Changes in the structure and development of managed Scots pine (Pinus sylvestris L.) stands with respect to changing environmental conditions were set for the period 1979–2015. The study was conducted in conditions of natural pinewoods and pine-oak sites on five permanent research plots (0.25 ha) in Eastern Bohemia, Czech Republic (CR). Studied forest stands showed positive development of stand structural characteristics related to their diversity, number of regeneration individuals and growth characteristics. The standing volume of regularly distributed tree layer in 2015 was in the range of 320–434 m3 ha–1, which indicates an increase by 5.9–20.0% over 10 years. Correlation between pine radial increment and the amount of precipitation was generally the strongest one. Positive statistically significant correlation between diameter increment and temperature was demonstrated only for the average March temperature of the current year. Within the CR, study site can be characterised as a medium polluted area both for sulphur and nitrogen, despite this SO2 concentrations and N deposition in combination with extreme climate events caused severe defoliation in pine stands. Conversely, radial growth was positively significantly correlated with mean NOx concentrations. Drought mainly in combination with even medium environmental pollution can further worsen the health status of pine stands in lowland areas of Central Europe. Thus, formulation of silvicultural techniques able to mitigate the impact of these stress factors is needed.

  • Vacek, Czech University of Life Sciences Prague, Faculty of Forestry and Wood Sciences, Kamýcká 129, 165 21 Prague, Czech Republic E-mail: vacekstanislav@fld.czu.cz
  • Vacek, Czech University of Life Sciences Prague, Faculty of Forestry and Wood Sciences, Kamýcká 129, 165 21 Prague, Czech Republic E-mail: vacekz@fld.czu.cz
  • Bílek, Czech University of Life Sciences Prague, Faculty of Forestry and Wood Sciences, Kamýcká 129, 165 21 Prague, Czech Republic E-mail: bilek@fld.czu.cz
  • Simon, Mendel University in Brno, Faculty of Forestry and Wood Technology, Zemědělská 3, 613 00 Brno, Czech Republic E-mail: jaroslav.simon@mendelu.cz
  • Remeš, Czech University of Life Sciences Prague, Faculty of Forestry and Wood Sciences, Kamýcká 129, 165 21 Prague, Czech Republic E-mail: remesj@email.cz (email)
  • Hůnová, Czech Hydrometeorological Institute, Na Šabatce 17 143 06 Prague, Czech Republic E-mail: hunova@chmi.cz
  • Král, Czech University of Life Sciences Prague, Faculty of Forestry and Wood Sciences, Kamýcká 129, 165 21 Prague, Czech Republic E-mail: kraljan@fld.czu.cz
  • Putalová, Czech University of Life Sciences Prague, Faculty of Forestry and Wood Sciences, Kamýcká 129, 165 21 Prague, Czech Republic E-mail: putalova@fld.czu.cz
  • Mikeska, University of Hradec Králové, Faculty of Science, Rokitanského 62, 500 03 Hradec Králové, Czech Republic E-mail: Mikeska.Miroslav@uhul.cz
article id 1559, category Research article
Karol Bronisz, Mike Strub, Chris Cieszewski, Szymon Bijak, Agnieszka Bronisz, Robert Tomusiak, Rafał Wojtan, Michał Zasada. (2016). Empirical equations for estimating aboveground biomass of Betula pendula growing on former farmland in central Poland. Silva Fennica vol. 50 no. 4 article id 1559. https://doi.org/10.14214/sf.1559
Keywords: silver birch; simplified and expanded models; seemingly unrelated regression
Highlights: We developed equations for aboveground biomass components of young silver birch stands on post-agricultural lands in central Poland for single tree level; Simplified equations were based exclusively on diameter at ground level or breast height, while expanded ones were based on the appropriate diameter and tree height; For large trees, diameter at breast height is a more appropriate explanatory variable than diameter at ground level; Biomass estimations based on models from neighboring countries were consistent with our results.
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We determined empirical models for estimating total aboveground as well as stem, branches, and foliage dry biomass of young (age up to 16 years) silver birch (Betula pendula Roth.) growing on the post-agricultural lands. Two sets of allometric models for trees with a height below or above 1.3 m (small and large trees respectively) were developed. Simplified models were elaborated based exclusively on appropriate tree diameter (diameter at ground level for small trees, diameter at breast height for large trees), while expanded models also included tree height. Total aboveground biomass was estimated as the sum of biomass of all tree components. To assure additivity of the developed equations, the seemingly unrelated regression approach for the final model fitting was used. Expanded models in both tree groups were characterized by a better fit to the data (R2 for total aboveground biomass for small and large trees equaled 0.8768 and 0.9752, respectively). Diameter at breast height appeared to be a better predictor than diameter at ground level – simplified models had better fit for large trees (R2 for total aboveground biomass equals 0.9611) than for small ones (R2 = 0.7516). The developed equations provide biomass predictions consistent with available Latvian, Estonian, Finnish, Swedish, and Norwegian models for silver birch.

  • Bronisz, Laboratory of Dendrometry and Forest Productivity, Faculty of Forestry, Warsaw University of Life Sciences – SGGW, Nowoursynowska 159, 02-787 Warsaw, Poland E-mail: karol.bronisz@wl.sggw.pl (email)
  • Strub, Warnell School of Forestry and Natural Resources, University of Georgia, Athens, GA, 30605, USA E-mail: strub@mcfns.com
  • Cieszewski, Warnell School of Forestry and Natural Resources, University of Georgia, Athens, GA, 30605, USA E-mail: thebiomat@gmail.com
  • Bijak, Laboratory of Dendrometry and Forest Productivity, Faculty of Forestry, Warsaw University of Life Sciences – SGGW, Nowoursynowska 159, 02-787 Warsaw, Poland E-mail: szymon.bijak@wl.sggw.pl
  • Bronisz, Laboratory of Dendrometry and Forest Productivity, Faculty of Forestry, Warsaw University of Life Sciences – SGGW, Nowoursynowska 159, 02-787 Warsaw, Poland E-mail: agnieszka.bronisz@wl.sggw.pl
  • Tomusiak, Laboratory of Dendrometry and Forest Productivity, Faculty of Forestry, Warsaw University of Life Sciences – SGGW, Nowoursynowska 159, 02-787 Warsaw, Poland E-mail: robert.tomusiak@wl.sggw.pl
  • Wojtan, Laboratory of Dendrometry and Forest Productivity, Faculty of Forestry, Warsaw University of Life Sciences – SGGW, Nowoursynowska 159, 02-787 Warsaw, Poland E-mail: rafal.wojtan@wl.sggw.pl
  • Zasada, Laboratory of Dendrometry and Forest Productivity, Faculty of Forestry, Warsaw University of Life Sciences – SGGW, Nowoursynowska 159, 02-787 Warsaw, Poland E-mail: michal.zasada@wl.sggw.pl
article id 1568, category Research article
Jouni Siipilehto, Harri Lindeman, Mikko Vastaranta, Xiaowei Yu, Jori Uusitalo. (2016). Reliability of the predicted stand structure for clear-cut stands using optional methods: airborne laser scanning-based methods, smartphone-based forest inventory application Trestima and pre-harvest measurement tool EMO. Silva Fennica vol. 50 no. 3 article id 1568. https://doi.org/10.14214/sf.1568
Keywords: forest inventory; diameter distribution; Weibull; area-based approach; parameter recovery; k-NN estimation
Highlights: An airborne laser scanning grid-based approach for determining stand structure enabled bi- or multimodal predicted distributions that fitted well to the ground-truth harvester data; EMO and Trestima applications needed stand-specific inventory for sample measurements or sample photos, respectively, and at their best, provided superior accuracy for predicting certain stand characteristics.
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Accurate timber assortment information is required before cuttings to optimize wood allocation and logging activities. Timber assortments can be derived from diameter-height distribution that is most often predicted from the stand characteristics provided by forest inventory. The aim of this study was to assess and compare the accuracy of three different pre-harvest inventory methods in predicting the structure of mainly Scots pine-dominated, clear-cut stands. The investigated methods were an area-based approach (ABA) based on airborne laser scanning data, the smartphone-based forest inventory Trestima app and the more conventional pre-harvest inventory method called EMO. The estimates of diameter-height distributions based on each method were compared to accurate tree taper data measured and registered by the harvester’s measurement systems during the final cut. According to our results, grid-level ABA and Trestima were generally the most accurate methods for predicting diameter-height distribution. ABA provides predictions for systematic 16 m × 16 m grids from which stand-wise characteristics are aggregated. In order to enable multimodal stand-wise distributions, distributions must be predicted for each grid cell and then aggregated for the stand level, instead of predicting a distribution from the aggregated stand-level characteristics. Trestima required a sufficient sample for reliable results. EMO provided accurate results for the dominating Scots pine but, it could not capture minor admixtures. ABA seemed rather trustworthy in predicting stand characteristics and diameter distribution of standing trees prior to harvesting. Therefore, if up-to-date ABA information is available, only limited benefits can be obtained from stand-specific inventory using Trestima or EMO in mature pine or spruce-dominated forests.

  • Siipilehto, Natural Research Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: jouni.siipilehto@luke.fi (email)
  • Lindeman,  Natural Research Institute Finland, Green Technology, Kaironiementie 15, 39700 Parkano E-mail: harri.lindeman@luke.fi
  • Vastaranta, University of Helsinki, Department of Forest Sciences, P.O. Box 62 (Viikinkaari 11), FI-00014 University of Helsinki E-mail: mikko.vastaranta@helsinki.fi
  • Yu, Finnish Geospatial Research Institute (FGI), Department of Remote Sensing and Photogrammetry, National Land Survey of Finland, P.O. Box 15 (Geodeetinrinne 2), FI-02431, Masala, Finland E-mail: xiaowei.yu@maanmittauslaitos.fi
  • Uusitalo,  Natural Research Institute Finland, Green Technology, Kaironiementie 15, 39700 Parkano E-mail: jori.uusitalo@luke.fi
article id 1565, category Research article
Teija Ruuhola, Ari Nikula, Nivala Vesa, Seppo Nevalainen, Juho Matala. (2016). Effects of bedrock and surficial deposit composition on moose damage in young forest stands in Finnish Lapland. Silva Fennica vol. 50 no. 3 article id 1565. https://doi.org/10.14214/sf.1565
Keywords: Pinus sylvestris; plantation; Alces alces; soil; damage risk; topography; geochemistry
Highlights: The effect of bedrock and soil on moose damage in forest plantations were examined; Moose damage were concentrated in nutrient rich bedrock areas; Bedrock of damaged stands contained a higher proportion of mafic and alkaline rocks; Pine-dominated stands on fine grained fertile forest sites had the highest damage risk.
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There is evidence that moose are attracted to fertile growth habitats apparently due to better quality and larger quantities of food. The nutrients in mineral soils originate from the weathering of bedrock and the composition of parental bedrock affects the fertility of produced mineral soil, thus affecting also the import of nutrients into the whole food web. We surveyed the connection between moose damage in forest plantations and the composition of bedrock and surficial deposits in Finnish Lapland. We used a database of compensated moose damage in private forests in years 1997−2010. Undamaged stands in National Forest Inventories (NFI) from years 1986–2008 served as a control data and moose-damaged NFI-stands as a reference data. Bedrock and surficial depositions and the location of studied stands in relation to ancient shorelines were explored by using the digital databases of the Geological Survey of Finland. Moose-damaged stands were concentrated in southwestern and east Lapland in the areas of the Peräpohja Schist Belt and Lapland’s Greenstone Belt that are both composed of nutrient-rich rocks. The bedrock of damaged stands contained a higher proportion of mafic and alkaline rocks than did the control stands. Moose-damaged stands were pine-dominated and grew in more fertile forest sites than did control stands. Part of pine stands probably located in soils formerly occupied by spruce, which may increase the stands’ vulnerability to biotic threats. Especially, there were relatively more moose damage in pine plantations regenerated on fine-grained mineral soils derived from nutrient rich rocks than in less fertile soils.

  • Ruuhola, Natural Resources Institute Finland (Luke), Natural Resources and Bioproduction, Yliopistokatu 6, FI-80101 Joensuu, Finland; University of Eastern Finland, Faculty of Science and Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: teija.ruuhola@uef.fi
  • Nikula, Natural Resources Institute Finland (Luke), Economics and Society, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: ari.nikula@luke.fi (email)
  • Vesa, Natural Resources Institute Finland (Luke), Economics and Society, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: vesa.nivala@luke.fi
  • Nevalainen, Natural Resources Institute Finland (Luke), Natural Resources and Bioproduction, Yliopistokatu 6, FI-80101 Joensuu, Finland E-mail: seppo.nevalainen@luke.fi
  • Matala, Natural Resources Institute Finland (Luke), Natural Resources and Bioproduction, Yliopistokatu 6, FI-80101 Joensuu, Finland E-mail: juho.matala@luke.fi
article id 1562, category Research article
Mats Erik Berlin, Torgny Persson, Gunnar Jansson, Matti Haapanen, Seppo Ruotsalainen, Lars Bärring, Bengt Andersson Gull. (2016). Scots pine transfer effect models for growth and survival in Sweden and Finland. Silva Fennica vol. 50 no. 3 article id 1562. https://doi.org/10.14214/sf.1562
Keywords: Pinus sylvestris; adaptation; tree breeding; reaction patterns; forest regeneration material
Highlights: Scots pine transfer effect models for growth and survival, valid in both Sweden and Finland have been developed; The models use high-resolution gridded climate data and can predict performance in future climatic conditions; The models perform well both for unimproved and genetically improved material and can be used to develop deployment recommendations of contemporary forest regeneration material in Sweden and Finland.
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In this study, we developed models of transfer effects for growth and survival of Scots pine (Pinus sylvestris L.) in Sweden and Finland using a general linear mixed-model approach. For model development, we used 378 provenance and progeny trials with a total of 276 unimproved genetic entries (provenances and stand seed check-lots) distributed over a wide variety of climatic conditions in both countries. In addition, we used 119 progeny trials with 3921 selected genetic entries (open- and control pollinated plus-tree families) for testing model performance. As explanatory variables, both climatic indices derived from high-resolution gridded climate datasets and geographical variables were used. For transfer, latitude (photoperiod) and, for describing the site, temperature sum were found to be main drivers for both survival and growth. In addition, interaction terms (between transfer in latitude and site altitude for survival, and transfer in latitude and temperature sum for growth) entail changed reaction patterns of the models depending on climatic conditions of the growing site. The new models behave in a way that corresponds well to previous studies and recommendations for both countries. The model performance was tested using selected plus-trees from open and control pollinated progeny tests. Results imply that the models are valid for both countries and perform well also for genetically improved material. These models are the first step in developing common deployment recommendations for genetically improved forest regeneration material in both Sweden and Finland.

  • Berlin, Uppsala Science Park, SE-75183 Uppsala, Sweden E-mail: mats.berlin@skogforsk.se (email)
  • Persson, Skogforsk, Box 3, SE-91821 Sävar, Sweden E-mail: torgny.persson@skogforsk.se
  • Jansson, Uppsala Science Park, SE-75183 Uppsala, Sweden E-mail: gunnar.jansson@skogforsk.se
  • Haapanen, Natural Resources Institute Finland (Luke), Green Technology, Box 18, FI-01301 Vantaa, Finland E-mail: matti.haapanen@luke.fi
  • Ruotsalainen, Natural Resources Institute Finland (Luke), Green Technology, Finlandiantie 18, FI-58450 Punkaharju, Finland E-mail: seppo.ruotsalainen@luke.fi
  • Bärring, Rossby Centre, Swedish Meteorological and Hydrological Institute, Folkborgsvägen 17, SE-60176 Norrköping, Sweden E-mail: lars.barring@smhi.se
  • Andersson Gull, Skogforsk, Box 3, SE-91821 Sävar, Sweden E-mail: bengt.andersson@skogforsk.se
article id 1560, category Research article
Marina Gurskaya, Pavel Moiseev, Martin Wilmking. (2016). Does slope exposure affect frost ring formation in Picea obovata growing at treeline in the Southern Urals? Silva Fennica vol. 50 no. 3 article id 1560. https://doi.org/10.14214/sf.1560
Keywords: tree line; topography; frost injury; Siberian spruce
Highlights: Frost ring formation was ubiquitous from continuous forest cover to treeline, but, surprisingly, was not affected by slope exposure; Late frosts in spring were the main cause for frost ring formation; While mainly young trees (< 30 years) recorded frost events, at the climatically harshest site (highest elevation, northeastern exposure), frost events were recorded also in trees older than 70 years.
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Topographic complexity in mountainous ecosystems strongly influences plant growth and as such also wood formation. This wood formation can possibly be used to understand topographic variation of the main climatic drivers, e.g. by modulating frost events. Here we test the influence of different slope exposures on the spatio-temporal distribution of frost rings in Siberian spruce (Picea obovata Ledeb.) in the Southern Urals, Russia. We sampled on two opposite slopes, northeast (NE) and southwest (SW), on three elevation levels from the highest single trees to closed canopy forest and analysed frost ring occurrence and their seasonal timing. Frost ring formation at all exposure-elevation combinations was common and mainly concentrated in the early part of the growing season. The age until trees record frost rings was equally similar (until about 35 years) on both slopes and different elevational levels with the exception of the climatically harshest site, the highest elevation on the NE slope. While we could not deduce a direct, easily identifiable climatic driver for the formation of frost rings, our analysis shows high potential to disentangle the complex interplay between climate, site condition and tree growth in mountainous ecosystems.

  • Gurskaya, Institute of Plant and Animal Ecology (IPAE), Ural Branch of Russian Academy of Sciences , 8 Marta St. 202, 620144 Ekaterinburg, Russia E-mail: mgurskaya@yandex.ru (email)
  • Moiseev, Institute of Plant and Animal Ecology (IPAE), Ural Branch of Russian Academy of Sciences , 8 Marta St. 202, 620144 Ekaterinburg, Russia E-mail: moiseev@ipae.uran.ru
  • Wilmking, Institute of Botany and Landscape Ecology, Soldmanstrasse 15, Ernst-Moritz-Arndt-University Greifswald, 17487 Greifswald, Germany E-mail: wilmking@uni-greifswald.de
article id 1546, category Research article
Jussi Manner, Lauri Palmroth, Tomas Nordfjell, Ola Lindroos. (2016). Load level forwarding work element analysis based on automatic follow-up data. Silva Fennica vol. 50 no. 3 article id 1546. https://doi.org/10.14214/sf.1546
Keywords: harvesting; cut-to-length logging; haulage; data gathering; automatic recording; classification algorithm; Hidden Markov Models
Highlights: Recent developments in on-board technology enables automatic collection of follow-up data on forwarder work; Time consumption per load was more strongly associated with Loading drive distance than with extraction distance, indicating that the relevance of extraction distance as a main indicator of forwarding productivity should be re-considered; Data, within variables, were positively skewed with a few exceptions with normal distributions.
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Recent developments in on-board technology have enabled automatic collection of follow-up data on forwarder work. The objective of this study was to exploit this possibility to obtain highly representative information on time consumption of specific work elements (including overlapping crane work and driving), with one load as unit of observation, for large forwarders in final felling operations. The data used were collected by the John Deere TimberLink system as nine operators forwarded 8868 loads, in total, at sites in mid-Sweden. Load-sizes were not available. For the average and median extraction distances (219 and 174 m, respectively), Loading, Unloading, Driving empty, Driving loaded and Other time effective work (PM) accounted for ca. 45, 19, 8.5, 7.5 and 14% of total forwarding time consumption, respectively. The average and median total time consumptions were 45.8 and 42.1 minutes/load, respectively. The developed models explained large proportions of the variation of time consumption for the work elements Driving empty and Driving loaded, but minor proportions for the work elements Loading and Unloading. Based on the means, the crane was used during 74.8% of Loading PM time, the driving speed was nonzero during 31.9% of the Loading PM time, and Simultaneous crane work and driving occurred during 6.7% of the Loading PM time. Time consumption per load was more strongly associated with Loading drive distance than with extraction distance, indicating that the relevance of extraction distance as a main indicator of forwarding productivity should be re-considered.

  • Manner, The Forestry Research Institute of Sweden (Skogforsk), Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: jussi.manner@skogforsk.se (email)
  • Palmroth, John Deere Forestry, P.O. Box 472, FI-33101 Tampere, Finland E-mail: PalmrothLauri@JohnDeere.com
  • Nordfjell, Swedish University of Agricultural Sciences, Department of Forest Biomaterials and Technology, SE-901 83 Umeå, Sweden E-mail: tomas.nordfjell@slu.se
  • Lindroos, Swedish University of Agricultural Sciences, Department of Forest Biomaterials and Technology, SE-901 83 Umeå, Sweden E-mail: ola.lindroos@slu.se
article id 1516, category Research article
Hamed Yousefzadeh, Abasalt Hosseinzadeh Colagar, Fatemeh Fallah. (2016). Genetic diversity of geographically isolated Iranian populations of Betula pendula Roth: implications for conservation. Silva Fennica vol. 50 no. 3 article id 1516. https://doi.org/10.14214/sf.1516
Keywords: genetic differentiation; habitat isolation; haplotype diversity; genetic drift; long-term isolation
Highlights: The Iranian populations of birches exhibited high levels of genetic diversity, population differentiation, and the presence of unique haplotypes; The high genetic differentiation amongst the populations may contribute to the local geographical structure and poor gene flow amongst individuals; The results can potentially be used to adopt appropriate strategies for the conservation and management of isolated tree populations.
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The effects of long-term habitat fragmentations on genetic and population differentiation of Betula pendula Roth were investigated using chloroplast DNA (cpDNA) variations. Leaf samples were collected from four small remnant populations across the north of Iran. Three pairs of universal primers were used to amplify cpDNA, large single copy regions of trnC-trnD, trnK1-trnK2 and trnD-trnT. A total of 18 of the cpDNA haplotypes in the four populations were identified, however, no clear phylogeographic structuring of haplotypes could be detected. The total genetic diversity (HT) for all populations was high (0.932). Average intra-population genetic diversity was estimated as HS = 0.729 and average differentiation of populations GST = 0.218. Mantel tests of isolation by distance revealed a significant relationship between Wright’s inbreeding coefficient (Fst) and geographical distances for the four populations in Iran (r = 0.77, p < 0.05). The results of the hierarchical analysis of molecular variance (AMOVA) indicated that a 66% variation was partitioned within populations, whilst the variance amongst the four populations was only 34%. We suggest that significant genetic differentiation amongst populations can likely be attributed to reduced gene flow as a result of habitat fragmentation.

  • Yousefzadeh, Department of Forestry, Faculty of Natural Resources, Tarbiat Modares University, Nour, Mazandaran, Iran E-mail: h.yousefzadeh@modares.ac.ir (email)
  • Hosseinzadeh Colagar, Department of Molecular and Cell Biology; Faculty of Basic Sciences, University of Mazandaran, 47416-95447 Babolsar, Iran E-mail: ahcolagar@umz.ac.ir
  • Fallah, Department of Molecular and Cell Biology; Faculty of Basic Sciences, University of Mazandaran, 47416-95447 Babolsar, Iran E-mail: fatemehfalah69@yahoo.com
article id 1514, category Research article
Heidi Hallongren, Ville Kankaanhuhta, Mikael Kukkonen. (2016). Cleaning Scots pine seedling stands with mechanical uprooters – a work quality comparison of two related devices. Silva Fennica vol. 50 no. 3 article id 1514. https://doi.org/10.14214/sf.1514
Keywords: Pinus sylvestris; productivity; early cleaning; mechanized silviculture
Highlights: The productivity of the narrower modified device was significantly better than the wider original device; Work quality did not differ significantly between devices when stand characteristics, regeneration success and pre-existing damage were taken into account; Results indicate that mechanical uprooting devices may be further developed to a cost-effective alternative to motor-manual techniques for the early cleaning of direct seeded commercial Scots pine stands.
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Commercial forests require early cleaning to ensure the unhindered and uniform growth of crop trees. In order to be cost effective, non-crop vegetation should be uprooted to prevent their recovery. Performing this work manually is a labour-intensive task but it can be done mechanically. We evaluated the efficiency of two uprooting devices in direct seeded Scots pine (Pinus sylvestris L.) stands ca. 1 m tall. Productivity and quality of the uprooting work was investigated across eight stands and ca. 160 sample plots in northern Karelia, eastern Finland. Time consumption of the uprooters was analyzed through a linear regression model and the work quality through a multilevel multivariate model in terms of the number of individual Scots pine seedlings, processing units (i.e., a bunch of seedlings to be harvested in the future) and broadleaves. The productivity of the narrower modified device was significantly better in terms of time consumption than the wider original device. Work quality did not differ significantly between devices when stand characteristics, regeneration success and pre-existing damage were taken into account. Results indicate that mechanical uprooting devices may be further developed to a cost-effective alternative to motor-manual techniques for the early cleaning of direct seeded commercial Scots pine stands.

  • Hallongren, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: heidi.hallongren@luke.fi (email)
  • Kankaanhuhta, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: ville.kankaanhuhta@luke.fi
  • Kukkonen, University of Eastern Finland, School of Forest Sciences. P.O. Box 111, FI-80101 Joensuu, Finland E-mail: mikaelkukkonen@hotmail.com
article id 1492, category Research article
Li-Bin Liu, Yang-Yang Wu, Gang Hu, Zhong-Hua Zhang, An-Yun Cheng, Shi-Jie Wang, Jian Ni. (2016). Biomass of karst evergreen and deciduous broad-leaved mixed forest in central Guizhou province, southwestern China: a comprehensive inventory of a 2 ha plot. Silva Fennica vol. 50 no. 3 article id 1492. https://doi.org/10.14214/sf.1492
Keywords: biomass; secondary forest; necromass; karst terrain; carbon inventory
Highlights: Comprehensive inventory of the karst secondary forest based on a 2 ha large plot enhanced the reliability of biomass estimates; The biomass was 158.1 Mg ha−1, and the five dominant tree species accounted for 92.4% of aboveground tree biomass; The estimated necromass of woody debris and litter in the karst secondary forest was 17.6 Mg ha−1.
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The biomass of a secondary evergreen and deciduous broad-leaved mixed forest was comprehensively inventoried in a permanent 2 ha plot in southwestern China. Biomass models, sub-sampling, soil pit method, and published data were utilized to determine the biomass of all components. Results showed that the total biomass of the forest was 158.1 Mg ha−1; the total biomass included the major aboveground (137.7 Mg ha−1) and belowground (20.3 Mg ha−1) biomass components of vascular plants as well as the minor biomass components of bryophytes (0.078 Mg ha−1) and lichens (0.043 Mg ha−1). The necromass was 17.6 Mg ha−1 and included woody debris (9.0 Mg ha−1) and litter (8.6 Mg ha−1). The spatial pattern of the aboveground biomass was determined by the spatial distribution of dominant trees with large diameter, tall height, and dense wood. The belowground biomass differed in terms of root diameter and decreased with increasing soil depth. The belowground biomass in each soil pit in local habitats was not related to the spatial distribution of woody plants and soil pit depth. The karst forest presented lower biomass compared than the nonkarst forests in the subtropical zone. Biomass carbon in the karst terrains would increase substantially if degraded karst vegetation could be successfully restored to the forest. Comprehensive site-based biomass inventory of karst vegetation will contribute not only to provide data for benchmarking global and regional vegetation and carbon models but also for regional carbon inventory and vegetation restoration.

  • Liu, State Key Laboratory of Environmental Geochemistry, Institute of Geochemistry, Chinese Academy of Sciences, Lincheng West Road 99, 550081 Guiyang, China; Puding Karst Ecosystem Research Station, Chinese Academy of Sciences, 562100 Puding, China; University of Chinese Academy of Sciences, 100049 Beijing, China E-mail: liulibin@mail.gyig.ac.cn
  • Wu, State Key Laboratory of Environmental Geochemistry, Institute of Geochemistry, Chinese Academy of Sciences, Lincheng West Road 99, 550081 Guiyang, China; Puding Karst Ecosystem Research Station, Chinese Academy of Sciences, 562100 Puding, China; University of Chinese Academy of Sciences, 100049 Beijing, China E-mail: wuyang2468@hotmail.com
  • Hu, School of Chemistry and Life Science, Guangxi Teachers Education University, 530001 Nanning, China; Key Laboratory of Beibu Gulf Environment Change and Resources Utilization of Ministry of Education, Guangxi Teachers Education University, 530001 Nanning, China E-mail: ahhugang@gmail.com
  • Zhang, School of Chemistry and Life Science, Guangxi Teachers Education University, 530001 Nanning, China; Key Laboratory of Beibu Gulf Environment Change and Resources Utilization of Ministry of Education, Guangxi Teachers Education University, 530001 Nanning, China E-mail: gxtczzh@gmail.com
  • Cheng, State Key Laboratory of Environmental Geochemistry, Institute of Geochemistry, Chinese Academy of Sciences, Lincheng West Road 99, 550081 Guiyang, China; Puding Karst Ecosystem Research Station, Chinese Academy of Sciences, 562100 Puding, China E-mail: chenganyun@vip.skleg.cn
  • Wang, State Key Laboratory of Environmental Geochemistry, Institute of Geochemistry, Chinese Academy of Sciences, Lincheng West Road 99, 550081 Guiyang, China; Puding Karst Ecosystem Research Station, Chinese Academy of Sciences, 562100 Puding, China E-mail: wangshijie@vip.skleg.cn
  • Ni, State Key Laboratory of Environmental Geochemistry, Institute of Geochemistry, Chinese Academy of Sciences, Lincheng West Road 99, 550081 Guiyang, China; Puding Karst Ecosystem Research Station, Chinese Academy of Sciences, 562100 Puding, China E-mail: nijian@vip.skleg.cn (email)
article id 1461, category Research article
Ludmila Novitskaya, Nadezhda Nikolaeva, Natalia Galibina, Tatiana Tarelkina, Ludmila Semenova. (2016). The greatest density of parenchyma inclusions in Karelian birch wood occurs at confluences of phloem flows. Silva Fennica vol. 50 no. 3 article id 1461. https://doi.org/10.14214/sf.1461
Keywords: Betula pendula; patterned wood; parenchyma; branch attachments; forks; sucrose content
Highlights: Dark-colored inclusions creating the figured pattern in Karelian birch wood consist of storage parenchyma cells; Their greatest density is formed above branch attachments and below forks; In these zones, the sucrose content is elevated since photoassimilate flows of the trunk and branches merge into one pathway; A high level of sucrose enhances the differentiation of parenchyma cells.
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The specific pattern of the wood of Karelian birch (Betula pendula Roth var. carelica (Merckl.) Hämet-Ahti), is created mainly by dark-coloured inclusions of parenchyma tissue. Our study revealed that the greatest density of parenchyma inclusions in Karelian birch wood is observed above branch attachments to the trunk and below forks. In the place of branch attachment, phloem flows of photoassimilates (sucrose) from the branch and along the trunk merge into one pathway, causing a rise in sucrose content in tissues there. In the area below the fork, sucrose flows from two (or more) trunk axes are combined. Many studies have demonstrated that elevated sucrose level is associated with the differentiation of parenchyma. We believe that where large phloem fluxes merge a high level of sucrose promotes mass differentiation of parenchyma cells instead of fibers and vessels. As a result, the density of the figured pattern in the wood increases. The obtained data have a practical value and can be used in developing recommendations for Karelian birch cultivation.

  • Novitskaya, Forest Research Institute, Karelian Research Center, Russian Academy of Sciences, Pushkinskaya str. 11, 185910, Petrozavodsk, Russia E-mail: ludnovits@rambler.ru (email)
  • Nikolaeva, Forest Research Institute, Karelian Research Center, Russian Academy of Sciences, Pushkinskaya str. 11, 185910, Petrozavodsk, Russia E-mail: kar-birch@mail.ru
  • Galibina, Forest Research Institute, Karelian Research Center, Russian Academy of Sciences, Pushkinskaya str. 11, 185910, Petrozavodsk, Russia E-mail: galibina@krc.karelia.ru
  • Tarelkina, Forest Research Institute, Karelian Research Center, Russian Academy of Sciences, Pushkinskaya str. 11, 185910, Petrozavodsk, Russia E-mail: karelina.t.v@gmail.com
  • Semenova, Forest Research Institute, Karelian Research Center, Russian Academy of Sciences, Pushkinskaya str. 11, 185910, Petrozavodsk, Russia E-mail: mi7enova@gmail.com
article id 1441, category Research article
Dorota Zawadzka, Stanisław Drozdowski, Grzegorz Zawadzki, Jerzy Zawadzki. (2016). The availability of cavity trees along an age gradient in fresh pine forests. Silva Fennica vol. 50 no. 3 article id 1441. https://doi.org/10.14214/sf.1441
Keywords: Pinus sylvestris; forest management; Poland; Augustów Forest; cavity-nesting birds; woodpeckers
Highlights: The density of cavity trees in pine-dominated, managed forests varied in relation to stand age and was highest in stands older than 130 years of age; Cavities excavated by woodpeckers dominated among all cavities; The number of trees with cavities appears insufficient to ensure the effective protection of bird diversity in managed stands of Augustów Forest.
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Given their importance as a resource for many forest organisms, tree cavities were inventoried in the managed pine forests of north-east Poland, in relation to the: 70–100, 101–130 and >130 year age-classes within the clear-cutting system. The densities at which cavities were present was found to depend on forest age, given that stands 70–100 years old were characterised by an average density of 0.62 trees ha–1, while forests older than 130 years reported 3.28 trees ha–1. Stands aged 70–100 years differed from those aged 130+ in having just 0.27 trees ha–1 of cavity trees, as compared with 2.91 trees ha–1. The total volume of cavity trees in stands up to 100 years old was 0.37 m3 ha–1 on average, as compared with 5.42 m3 ha–1 in stands over 130 years old. The cavities created by woodpeckers constituted 76% of all of those found, and included 53% excavated by great spotted woodpeckers (Dendrocopos major L.) and 23% by black woodpeckers (Dryocopus martius L.) The proportion of cavities excavated by D. major was highest in the youngest age class of stands. There, cavities made by D. martius constituted only 8% of the total, as compared with 31% in the oldest stands. The abundance of cavity trees thus differed along an age gradient, though in any event the availability of cavity trees appears to be too limited to provide for the needs of hole-nesting birds. Forest managers must thus take more account than hitherto of the need to protect cavity trees.

  • Zawadzka, Institute of Forest Science, University of Łódź, Branch in Tomaszów Mazowiecki, Konstytucji 3 Maja 65/67, 97-200 Tomaszów Mazowiecki, Poland E-mail: dorota_zaw@wp.pl
  • Drozdowski, Department of Silviculture, Warsaw University of Life Sciences - SGGW, Nowoursynowska 159, 02-776 Warsaw, Poland E-mail: stanislaw_drozdowski@sggw.pl (email)
  • Zawadzki, Eagle Conservation Committee, Okółek 14, 16-506 Giby, Poland E-mail: grzesiekgfz@op.pl
  • Zawadzki, Eagle Conservation Committee, Okółek 14, 16-506 Giby, Poland E-mail: jerzy_zaw@wp.pl
article id 1313, category Research article
Jonathan Sheppard, Christopher Morhart, Heinrich Spiecker. (2016). Bark surface temperature measurements on the boles of wild cherry (Prunus avium) grown within an agroforestry system. Silva Fennica vol. 50 no. 3 article id 1313. https://doi.org/10.14214/sf.1313
Keywords: valuable timber; sunscald; sunburn; southwest disease; cambium; alley cropping
Highlights: Widely spaced trees within agroforestry systems are at risk of sun induced damages; Bark surface temperature on the south western bole face reached nearly 50 °C in summer and experienced a maximum range of 38 °C within a 24 hour period in spring; Maximum and minimum daily bark surface temperatures are modelled using climatic and solar position data; The application of white paint to stems reduces the bark surface temperature.
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Growing Prunus avium L. within an agroforestry system (AFS) may result in sun damage to cambial tissues on sun-exposed bole faces. There are two periods of risk of damage caused by insolation to exposed tree boles, the summer, when cambial temperatures become too high, or during winter, when the frozen dormant cambium tissue thaws and then rapidly re-freezes, a phenomenon commonly referred to as sunscald or southwest disease. Damage on the south western bole face was observed on a number of P. avium within an AFS. Five trees were sampled to assess the period in time that damage occurred. To retrospectively investigate such damage, bark surface temperature data were collected over a two year period for a further five P. avium and analysed. It was shown that bark surface temperature on the south western bole face reached nearly 50 °C during summer and experienced a maximum range of 38 °C within a 24 hour period in spring. A specially formulated white paint was applied to two trees, thus, testing a method to reduce the risk of sun damage. Two models were constructed to predict maximum and minimum daily bark surface temperature using maximum, minimum and mean daily air temperature, daily sum of sunshine hours, cloud cover, wind speed, relative humidity, maximum solar elevation and height on the tree bole as predictor variables. The damage occurred during winter 2009/2010. The models were used to identify maximum and minimum bark surface temperatures during that winter enabling the identification of possible damage events.

  • Sheppard, Chair of Forest Growth and Dendroecology, Albert-Ludwigs-University Freiburg, Tennenbacher Straße 4, 79106 Freiburg, Germany ORCID http://orcid.org/0000-0002-4959-7069 E-mail: jonathan.sheppard@iww.uni-freiburg.de (email)
  • Morhart, Chair of Forest Growth and Dendroecology, Albert-Ludwigs-University Freiburg, Tennenbacher Straße 4, 79106 Freiburg, Germany ORCID http://orcid.org/0000-0003-1874-5011 E-mail: christopher.morhart@iww.uni-freiburg.de
  • Spiecker, Chair of Forest Growth and Dendroecology, Albert-Ludwigs-University Freiburg, Tennenbacher Straße 4, 79106 Freiburg, Germany E-mail: instww@uni-freiburg.de
article id 1521, category Research article
Kalle Karttunen, Juha Laitila, Tapio Ranta. (2016). First-thinning harvesting alternatives for industrial or energy purposes based on regional Scots pine stand simulations in Finland. Silva Fennica vol. 50 no. 2 article id 1521. https://doi.org/10.14214/sf.1521
Keywords: harvesting; energy wood; stand simulation; stumpage price; small-diameter wood
Highlights: Small-diameter delimbed wood from Scots pine stands delivered directly for energy use was the most cost-efficient option in terms of the total supply-chain cost in comparison with corresponding industrial use or a whole-tree supply chain for energy use; Forest-management and harvesting decisions influenced the removal of forest biomass and stumpage price as well as the total supply-chain costs for forest biomass; The greatest cost-reduction potential (10.0%, 4.00 € m–3) was achieved for the delimbed energy wood’s supply chain in the regional case of South Savo in eastern Finland.
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Combining research into forest management stand conditions and wood supply chain processes has been missing from earlier forestry studies. There is a clear need to develop more cost-efficient small-diameter wood production, harvesting and transportation methods from first thinning, which could be used for either industrial or energy wood purposes. This study considers the total cost for small-diameter wood originating from young Scots pine (Pinus sylvestris L.) dominated stands. Pine pulpwood is the most harvested and most used roundwood assortment, use of which is expected to rise following new pulp-mill investments in Finland. In addition, utilisation of small-diameter trees directly for energy purposes has been increasing steadily in recent years. The aim of the study was to determine the cost-reduction potential of alternative forest management options and supply chains for small diameter-wood in the regional case of South Savo in eastern Finland. The total costs of three distinct scenarios were studied on the basis of forest management, first-thinning harvesting methods, and transportation: 1) industrial wood, 2) delimbed energy wood, and 3) whole trees for energy purposes. The cost-reduction potential for energy-wood supply chains from first thinning was compared to the industrial supply chain. Small-diameter delimbed wood delivered straight for energy purposes was found to be the most cost-efficient as far as the total cost of the supply chain is concerned. More cost-efficient small-diameter wood processes can be found by linking forest stand simulations with supply chain analysis.

  • Karttunen, Lappeenranta University of Technology, LUT School of Energy Systems, Laboratory of Bioenergy, Lönnrotinkatu 7, FI-50100 Mikkeli, Finland E-mail: kalle.karttunen@lut.fi
  • Laitila, Natural Resources Institute Finland (Luke), Bio-based business and industry, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: juha.laitila@luke.fi
  • Ranta, Lappeenranta University of Technology, LUT School of Energy Systems, Laboratory of Bioenergy, Lönnrotinkatu 7, FI-50100 Mikkeli, Finland E-mail: tapio.ranta@lut.fi (email)
article id 1520, category Research article
Tomáš Kolář, Kyriaki Giagli, Miroslav Trnka, Emílie Bednářová, Hanuš Vavrčík, Michal Rybníček. (2016). Response of the leaf phenology and tree-ring width of European beech to climate variability. Silva Fennica vol. 50 no. 2 article id 1520. https://doi.org/10.14214/sf.1520
Keywords: dendroclimatology; Fagus sylvatica; temperature; soil moisture; radial increment
Highlights: The timing of leaf phenological phases in European beech is controlled by temperature; Tree-ring width variations in European beech positively reflect growing season precipitation and soil water availability; The water availability in the top 40 cm of soil layer is more important for European beech growth than that in the deeper layers; Extension of the phenological growing season does not increase tree-ring width.
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Various environmental conditions (heat waves and drought events) strongly affect leaf and xylem phenology. Disentangling the influence of temperature, precipitation and soil moisture content (AWR) on the forest productivity remains an important research area. We analyzed the impact of climate variability on the leaf phenology (10 sample trees) and radial growth (17 sample trees) of European beech (Fagus sylvatica L.). The study was conducted on 130-year-old European beech trees growing in a temperate forest stand in the Czech Republic. Detailed 20-year phenological monitoring was performed at the study site (1992–2011). As expected, leaf phenological events were mainly driven by the growing season temperatures. Leaf unfolding was highly affected positively by spring temperatures and the top-layer (to 40 cm) AWR in March. The correlation of tree-ring width with the interpolated climate data was positive significant for the growing season AWR and precipitation signal. Furthermore, the water availability in the top soil layer was found to be an important predictor of tree growth and extremely low growth occurrence. The extended phenological growing season, which was caused by a temperature increase, was not followed by an increased tree-ring width. The examined relationships point out the significance of the water availability in the top soil layer in European beech stands.

  • Kolář, Department of Wood Science, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědělská 3, 613 00 Brno, Czech Republic; Global Change Research Institute, The Czech Academy of Sciences, Bělidla 986/4a, 603 00 Brno, Czech Republic E-mail: koldatom@gmail.com (email)
  • Giagli, Department of Wood Science, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědělská 3, 613 00 Brno, Czech Republic E-mail: giagli@node.mendelu.cz
  • Trnka, Global Change Research Institute, The Czech Academy of Sciences, Bělidla 986/4a, 603 00 Brno, Czech Republic; Department of Agrosystems and Bioclimatology, Faculty of Agronomy, Mendel University in Brno, Zemědělská 1, 613 00 Brno, Czech Republic E-mail: mirek_trnka@yahoo.com
  • Bednářová, Institute of Forest Ecology, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědelská 3, 61300 Brno, Czech Republic E-mail: bednarov@mendelu.cz
  • Vavrčík, Department of Wood Science, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědělská 3, 613 00 Brno, Czech Republic E-mail: vavrcik@mendelu.cz
  • Rybníček, Department of Wood Science, Faculty of Forestry and Wood Technology, Mendel University in Brno, Zemědělská 3, 613 00 Brno, Czech Republic; Global Change Research Institute, The Czech Academy of Sciences, Bělidla 986/4a, 603 00 Brno, Czech Republic E-mail: michalryb@post.cz
article id 1510, category Research article
Tähti Pohjanmies, Sakina Elshibli, Pertti Pulkkinen, Mari Rusanen, Pekka Vakkari, Helena Korpelainen, Tomas Roslin. (2016). Fragmentation-related patterns of genetic differentiation in pedunculate oak (Quercus robur) at two hierarchical scales. Silva Fennica vol. 50 no. 2 article id 1510. https://doi.org/10.14214/sf.1510
Keywords: marginal populations; Finland; genetic diversity; microsatellites
Highlights: While long-lived, widespread tree species should be resistant to genetic impoverishment, we detected high differentiation among populations and pronounced genetic structure within populations of Quercus robur in Finland; These patterns seem indicative of population processes active at range margins, and of disequilibrium following historic habitat change; Preservation of remaining genetic variation is thus important in the conservation of Q. robur.
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Populations at species’ range margins are expected to show lower genetic diversity than populations at the core of the range. Yet, long-lived, widespread tree species are expected to be resistant to genetic impoverishment, thus showing comparatively high genetic diversity within populations and low differentiation among populations. Here, we study the distribution of genetic variation in the pedunculate oak (Quercus robur L.) at its range margin in Finland at two hierarchical scales using 15 microsatellite loci. At a regional scale, we compared variation within versus among three oak populations. At a landscape scale, we examined genetic structuring within one of these populations, growing on an island of ca 5 km2. As expected, we found the majority of genetic variation in Q. robur to occur within populations. Nonetheless, differentiation among populations was markedly high (FST = 0.12) compared with values reported for populations of Q. robur closer to the core of its range. At the landscape level, some spatial and temporal sub-structuring was observed, likely explained by the history of land-use on the island. Overall, Q. robur fulfils the expectation of the central-marginal hypothesis of high differentiation among marginal populations, but the notable population differentiation has most likely been influenced also by the long, ongoing fragmentation of populations. Finnish oak populations may still be adjusting to the drastic habitat changes of the past centuries. Preservation of genetic variation within the remaining stands is thus an important factor in the conservation of Q. robur at its range margin.

  • Pohjanmies, University of Helsinki, Department of Agricultural Sciences, Spatial Foodweb Ecology Group, P.O. Box 27, FI-00014 University of Helsinki, Finland; University of Jyväskylä, Department of Biological and Environmental Sciences, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: tahti.t.pohjanmies@jyu.fi (email)
  • Elshibli, Natural Resources Institute Finland (Luke), Green technology, P.O. Box 18, FI-01301 Vantaa, Finland; University of Helsinki, Department of Agricultural Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: sakina.elshibli@helsinki.fi
  • Pulkkinen, Natural Resources Institute Finland (Luke), Green technology, Haapastensyrjäntie 34, FI-12600 Läyliäinen, Finland E-mail: pertti.pulkkinen@luke.fi
  • Rusanen, Natural Resources Institute Finland (Luke), Green technology, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: mari.rusanen@luke.fi
  • Vakkari, Natural Resources Institute Finland (Luke), Green technology, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: pekka.vakkari@luke.fi
  • Korpelainen, University of Helsinki, Department of Agricultural Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: helena.korpelainen@helsinki.fi
  • Roslin, University of Helsinki, Department of Agricultural Sciences, Spatial Foodweb Ecology Group, P.O. Box 27, FI-00014 University of Helsinki, Finland; Swedish University of Agricultural Sciences, Department of Ecology, P.O. Box 7044, SE-750 07 Uppsala, Sweden E-mail: tomas.roslin@helsinki.fi
article id 1495, category Research article
Per-Ola Olsson, Tuula Kantola, Päivi Lyytikäinen-Saarenmaa, Anna Maria Jönsson, Lars Eklundh. (2016). Development of a method for monitoring of insect induced forest defoliation – limitation of MODIS data in Fennoscandian forest landscapes. Silva Fennica vol. 50 no. 2 article id 1495. https://doi.org/10.14214/sf.1495
Keywords: remote sensing; insect defoliation detection; coarse-resolution; EVI2; z-score; Sentinel-2
Highlights: We developed and tested a method to monitor insect induced defoliation in forests based on coarse-resolution satellite data (MODIS); MODIS data may fail to detect defoliation in fragmented landscapes, especially if defoliation history is long. More homogenous forests results in higher detection accuracies; The method may be applied to future coarse and medium-resolution satellite data with high temporal resolution.
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We investigated if coarse-resolution satellite data from the MODIS sensor can be used for regional monitoring of insect disturbances in Fennoscandia. A damage detection method based on z-scores of seasonal maximums of the 2-band Enhanced Vegetation Index (EVI2) was developed. Time-series smoothing was applied and Receiver Operating Characteristics graphs were used for optimisation. The method was developed in fragmented and heavily managed forests in eastern Finland dominated by Scots pine (Pinus sylvestris L.) (pinaceae) and with defoliation of European pine sawfly (Neodiprion sertifer Geoffr.) (Hymenoptera: Diprionidae) and common pine sawfly (Diprion pini L.) (Hymenoptera: Diprionidae). The method was also applied to subalpine mountain birch (Betula pubescens ssp. Czerepanovii N.I. Orlova) forests in northern Sweden, infested by autumnal moth (Epirrita autumnata Borkhausen) and winter moth (Operophtera brumata L.). In Finland, detection accuracies were fairly low with 50% of the damaged stands detected, and a misclassification of healthy stands of 22%. In areas with long outbreak histories the method resulted in extensive misclassification. In northern Sweden accuracies were higher, with 75% of the damage detected and a misclassification of healthy samples of 19%. Our results indicate that MODIS data may fail to detect damage in fragmented forests, particularly when the damage history is long. Therefore, regional studies based on these data may underestimate defoliation. However, the method yielded accurate results in homogeneous forest ecosystems and when long-enough periods without damage could be identified. Furthermore, the method is likely to be useful for insect disturbance detection using future medium-resolution data, e.g. from Sentinel‑2.

  • Olsson, Lund University, Department of Physical Geography and Ecosystem Science, Sölvegatan 12, S-223 62 Lund, Sweden E-mail: per-ola.olsson@nateko.lu.se (email)
  • Kantola, Texas A & M University, Knowledge Engineering Laboratory, Department of Entomology, College Station, TX, USA E-mail: tuula.kantola@helsinki.fi
  • Lyytikäinen-Saarenmaa, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: paivi.lyytikainen-saarenmaa@helsinki.fi
  • Jönsson, Lund University, Department of Physical Geography and Ecosystem Science, Sölvegatan 12, S-223 62 Lund, Sweden E-mail: anna_maria.jonsson@nateko.lu.se
  • Eklundh, Lund University, Department of Physical Geography and Ecosystem Science, Sölvegatan 12, S-223 62 Lund, Sweden E-mail: lars.eklundh@nateko.lu.se
article id 1475, category Research article
Jiaxi Wang, Haiqun Yu, Guolei Li, Feng Zhang. (2016). Growth and nutrient dynamics of transplanted Quercus variabilis seedlings as influenced by pre-hardening and fall fertilization. Silva Fennica vol. 50 no. 2 article id 1475. https://doi.org/10.14214/sf.1475
Keywords: nutrient loading; nutrient storage; nursery transplanting; seedling quality
Highlights: High pre-hardening fertilization favored seedling growth and nutrient storage at the rapid growth and hardening phases following transplantation. Overall, high fall fertilization was beneficial only at the hardening phase; The combination of 100 mg N seedling–1 during pre-hardening with 36 mg N seedling–1 during hardening was recommended for satisfactory transplanting performance for Quercus variabilis.
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Stored nutrient reserves are closely correlated with survival and growth of transplanted seedlings. Previous studies have proven that combining pre-hardening fertilization (PF) with fall fertilization (FF) built seedling nutrient reserves more effectively; however, their effect on transplanting performance is poorly documented. We investigated the independent and interacting effects of 2 levels of PF and 4 levels of FF on seedling growth, nutrient acquisition and accumulation during different growth phases 1 year after transplanting of Quercus variabilis Blume in a nursery. High PF benefited nutrient reserves and subsequent transplanted seedling growth and tissue nutrient storage at the end of the rapid growth and hardening phases. Fall fertilization with 36 mg N increased stem dry mass and tissue nutrient content at the end of the hardening phase. At the conclusion of establishment, PF and FF showed a significant interaction for N and K uptake from soil. At the end of the rapid growth and hardening phases, high PF consistently increased nutrient uptake. Enhanced N and K uptake occurred following application of 36 mg N of FF at the end of the hardening phase. Distinct roles for PF and FF on 3 phases of transplanted seedlings demonstrated the necessity to evaluate fertilization in terms of nutrient reserves and subsequent transplanting performance in consecutive phases. Combining 100 mg N seedling–1 during pre-hardening with 36 mg N seedling–1 during fall yielded ideal transplanting performance for Quercus variabilis seedlings.

  • Wang, Key Laboratory for Silviculture and Conservation, Ministry of Education; Beijing Laboratory of Urban and Rural Ecological Environment, Beijing Forestry University, 35 East Qinghua Road, Haidian District, Beijing 100083, China E-mail: wjx198979@163.com
  • Yu, Beijing Forestry Carbon Administration, room 201, No.1 Xiao Huang Zhuang Bei Jie, Dongcheng District, Beijing 100013, China E-mail: yuhq@bfdic.com
  • Li, Key Laboratory for Silviculture and Conservation, Ministry of Education; Beijing Laboratory of Urban and Rural Ecological Environment, Beijing Forestry University, 35 East Qinghua Road, Haidian District, Beijing 100083, China E-mail: glli226@163.com (email)
  • Zhang, Beijing Forestry Carbon Administration, room 201, No.1 Xiao Huang Zhuang Bei Jie, Dongcheng District, Beijing 100013, China E-mail: zhangf@bfdic.com
article id 1474, category Research article
Cedric A. Goussanou, Sabin Guendehou, Achille E. Assogbadjo, Maguette Kaire, Brice Sinsin, Aida Cuni-Sanchez. (2016). Specific and generic stem biomass and volume models of tree species in a West African tropical semi-deciduous forest. Silva Fennica vol. 50 no. 2 article id 1474. https://doi.org/10.14214/sf.1474
Keywords: carbon stock; allometric equations; non-destructive sampling; basic wood density
Highlights: Non-destructive sampling approach applied to derive ground truth observations and generate robust basic wood densities; Species-specific and generic allometric equations; Specific equations have better predictive capabilities than generic models.
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The quantification of the contribution of tropical forests to global carbon stocks and climate change mitigation requires availability of data and tools such as allometric equations. This study made available volume and biomass models for eighteen tree species in a semi-deciduous tropical forest in West Africa. Generic models were also developed for the forest ecosystem, and basic wood density determined for the tree species. Non-destructive sampling approach was carried out on five hundred and one sample trees to analyse stem volume and biomass. From the modelling of volume and biomass as functions of diameter at breast height (Dbh) and stem height, logarithmic models had better predictive capabilities. The model validation showed that in absence of data on height, models using Dbh only as variable was an alternative. The comparison of basic wood densities to data published in literature enabled to conclude that the non-destructive sampling was a good approach to determining reliable basic wood density. The comparative analysis of species-specific models in this study with selected generic models for tropical forests indicated low probability to identify effective generic models with good predictive ability for biomass. Given tree species richness of tropical forests, the study demonstrated the hypothesis that species-specific models are preferred to generic models, and concluded that further research should be oriented towards development of specific models to cover the full range of dominant tree species of African forests.

  • Goussanou, Laboratory of Applied Ecology, Faculty of Agronomic Sciences, University of Abomey-Calavi, 01 BP 526 Cotonou, Benin E-mail: cedricgoussanou@gmail.com (email)
  • Guendehou, Laboratory of Applied Ecology, Faculty of Agronomic Sciences, University of Abomey-Calavi, 01 BP 526 Cotonou, Benin; Benin Centre for Scientific and Technical Research, 03 BP 1665 Cotonou, Benin E-mail: sguendehou@yahoo.fr
  • Assogbadjo, Laboratory of Applied Ecology, Faculty of Agronomic Sciences, University of Abomey-Calavi, 01 BP 526 Cotonou, Benin E-mail: assogbadjo@yahoo.fr
  • Kaire, Centre Régional AGRHYMET, Département Formation et Recherche, BP 11011 Niamey, Niger E-mail: m.kaire@agrhymet.ne
  • Sinsin, Laboratory of Applied Ecology, Faculty of Agronomic Sciences, University of Abomey-Calavi, 01 BP 526 Cotonou, Benin E-mail: bsinsin@gmail.com
  • Cuni-Sanchez, University of Copenhagen, Center for Macroecology, Evolution and Climate, Nørregade 10, P.O. Box 2177, 1017 Copenhagen K, Denmark E-mail: aidacuni@hotmail.com
article id 1462, category Research article
Pekka Punttila, Olli Autio, Janne S. Kotiaho, D. Johan Kotze, Olli J. Loukola, Norbertas Noreika, Anna Vuori, Kari Vepsäläinen. (2016). The effects of drainage and restoration of pine mires on habitat structure, vegetation and ants. Silva Fennica vol. 50 no. 2 article id 1462. https://doi.org/10.14214/sf.1462
Keywords: ecological restoration; ditching; Aichi Biodiversity Target 15; Formicidae; pine bogs and fens; transforming and transformed drained mires; water-table level
Highlights: Mire drainage shifted floristic composition and ant assemblages towards forest communities; Raising the water-table level by ditch filling and the thinning of trees affected mire communities positively already 1–3 years after the start of restoration; The extent of tree cover, the coverage of Sphagnum mosses and the water-table level were major determinants of ant assemblage structure.
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Habitat loss and degradation are the main threats to biodiversity worldwide. For example, nearly 80% of peatlands in southern Finland have been drained. There is thus a need to safeguard the remaining pristine mires and to restore degraded ones. Ants play a pivotal role in many ecosystems and like many keystone plant species, shape ecosystem conditions for other biota. The effects of mire restoration and subsequent vegetation succession on ants, however, are poorly understood. We inventoried tree stands, vegetation, water-table level, and ants (with pitfall traps) in nine mires in southern Finland to explore differences in habitats, vegetation and ant assemblages among pristine, drained (30–40 years ago) and recently restored (1–3 years ago) pine mires. We expected that restoring the water-table level by ditch filling and reconstructing sparse tree stands by cuttings will recover mire vegetation and ants. We found predictable responses in habitat structure, floristic composition and ant assemblage structure both to drainage and restoration. However, for mire-specialist ants the results were variable and longer-term monitoring is needed to confirm the success of restoration since these social insects establish perennial colonies with long colony cycles. We conclude that restoring the water-table level and tree stand structure seem to recover the characteristic vegetation and ant assemblages in the short term. This recovery was likely enhanced because drained mires still had both acrotelm and catotelm, and connectedness was still reasonable for mire organisms to recolonize the restored mires either from local refugia or from populations of nearby mires.

  • Punttila, Finnish Environment Institute, P.O. Box 140, FI-00251 Helsinki, Finland E-mail: pekka.punttila@ymparisto.fi (email)
  • Autio, Centre for Economic Development, Transport and the Environment in South Ostrobothnia, P.O. Box 252, FI-65101 Vaasa, Finland E-mail: olli.autio@ely-keskus.fi
  • Kotiaho, University of Jyväskylä, Department of Biology & Environmental Sciences, P.O. Box 35, FI-40014 Jyväskylä, Finland E-mail: janne.kotiaho@jyu.fi
  • Kotze, University of Helsinki, Department of Environmental Sciences, P.O. Box 65, FI-00014, University of Helsinki, Finland E-mail: johan.kotze@helsinki.fi
  • Loukola, University of Oulu, Department of Biology, P.O. Box 3000, FI-90014 Oulu, Finland E-mail: olli.loukola@gmail.com
  • Noreika, University of Helsinki, Department of Environmental Sciences, P.O. Box 65, FI-00014, University of Helsinki, Finland; University of Helsinki, Department of Biosciences, P.O. Box 65, FI-00014 University of Helsinki, Finland E-mail: norbertas.noreika@gmail.com
  • Vuori, University of Jyväskylä, Department of Biology & Environmental Sciences, P.O. Box 35, FI-40014 Jyväskylä, Finland E-mail: anna@kureniemi.fi
  • Vepsäläinen, University of Helsinki, Department of Biosciences, P.O. Box 65, FI-00014 University of Helsinki, Finland E-mail: kari.vepsalainen@helsinki.fi
article id 1448, category Research article
Andrew McEwan, Natascia Magagnotti, Raffaele Spinelli. (2016). The effects of number of stems per stool on cutting productivity in coppiced Eucalyptus plantations. Silva Fennica vol. 50 no. 2 article id 1448. https://doi.org/10.14214/sf.1448
Keywords: modeling; logging; short rotation forestry; efficiency; operator effect
Highlights: Double- and single stem coppice stools were harvested mechanically; Stem size had the strongest impact on productivity; An experienced operator performed equally well with both stool treatments; Cost was ~10% higher with double stems for the less experienced operator; Operator experience may play a major role when cutting coppice stands.
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A time study was conducted to determine whether stem crowding had any impact on harvester productivity in Eucalyptus grandis stands. This represents an important element when trying to balance the advantages and disadvantages of coppice management in fast growing plantations designated for mechanized harvesting (i.e. machine felling, delimbing, debarking and cross-cutting). The study material consisted of 446 coppice stems, half of which grew as single stems per stool and half as double stems per stool as a result of different coppice reduction strategies. The dataset was balanced and randomized, with both subsets replicating exactly the same stem size distribution and the single and double stems alternating randomly. Harvester productivity ranged between 6 and 50 m3 under bark per productive machine hour, following the variation of tree diameter from 10 to 40 cm at breast height (1.37 m according to South African standards). Regression analysis indicated that both tree size and stem crowding (e.g. one or two stems per stool) had a significant effect on harvester productivity, which increased with stem size and decreased with stem crowding. However, operator experience may overcome the effect of stem crowding, which was not significant when the harvester was manned by a highly experienced operator. In any case, the effect of stem size was much greater than that of stem crowding, which resulted in a cost difference of less than 10%. However, this figure excludes the possible effects of stem crowding on volume recovery and stem development, which should be addressed in the future.

  • McEwan, Nelson Mandela Metropolitan University – George Campus, Saasveld, 6529, George, South Africa E-mail: Andrew.McEwan@nmmu.ac.za
  • Magagnotti, CNR IVALSA, Via Madonna del Piano 10, I-50019 Sesto Fiorentino (FI), Italy E-mail: magagnotti@ivalsa.cnr.it
  • Spinelli, CNR IVALSA, Via Madonna del Piano 10, I-50019 Sesto Fiorentino (FI), Italy E-mail: spinelli@ivalsa.cnr.it (email)
article id 1414, category Research article
Rami Saad, Jörgen Wallerman, Johan Holmgren, Tomas Lämås. (2016). Local pivotal method sampling design combined with micro stands utilizing airborne laser scanning data in a long term forest management planning setting. Silva Fennica vol. 50 no. 2 article id 1414. https://doi.org/10.14214/sf.1414
Keywords: LIDAR; forest management planning; local pivotal method (LPM); segmentation; most similar neighbor (MSN) imputation; suboptimal loss; Heureka; decision support system
Highlights: Most similar neighbor imputation was used to estimate forest variables using airborne laser scanning data as auxiliary data; For selecting field reference plots the local pivotal method (LPM) was compared to systematic sampling design; The LPM sampling design combined with a micro stand approach showed potential for improvement and has the potential to be a competitive method when considering cost efficiency.
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A new sampling design, the local pivotal method (LPM), was combined with the micro stand approach and compared with the traditional systematic sampling design for estimation of forest stand variables. The LPM uses the distance between units in an auxiliary space – in this case airborne laser scanning (ALS) data – to obtain a well-spread sample. Two sets of reference plots were acquired by the two sampling designs and used for imputing data to evaluation plots. The first set of reference plots, acquired by LPM, made up four imputation alternatives (varying number of reference plots) and the second set of reference plots, acquired by systematic sampling design, made up two alternatives (varying plot radius). The forest variables in these alternatives were estimated using the nonparametric method of most similar neighbor imputation, with the ALS data used as auxiliary data. The relative root mean square error (RelRMSE), stem diameter distribution error index and suboptimal loss were calculated for each alternative, but the results showed that neither sampling design, i.e. LPM vs. systematic, offered clear advantages over the other. It is likely that the obtained results were a consequence of the small evaluation dataset used in the study (n = 30). Nevertheless, the LPM sampling design combined with the micro stand approach showed potential for improvement and might be a competitive method when considering the cost efficiency.

  • Saad, Swedish University of Agricultural Sciences (SLU), Department of Forest Resource Management, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: rami.saad@slu.se (email)
  • Wallerman, Swedish University of Agricultural Sciences (SLU), Department of Forest Resource Management, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: jorgen.wallerman@slu.se
  • Holmgren, Swedish University of Agricultural Sciences (SLU), Department of Forest Resource Management, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: johan.holmgren@slu.se
  • Lämås, Swedish University of Agricultural Sciences (SLU), Department of Forest Resource Management, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: tomas.lamas@slu.se
article id 1410, category Research article
Seppo Nevalainen, Juho Matala, Kari T. Korhonen, Antti Ihalainen, Ari Nikula. (2016). Moose damage in National Forest Inventories (1986–2008) in Finland. Silva Fennica vol. 50 no. 2 article id 1410. https://doi.org/10.14214/sf.1410
Keywords: Populus tremula; Pinus sylvestris; regeneration; soil preparation; thinning; Betula spp.; Keywords Alces alces; seedling stands; tree species mixture
Highlights: Almost 100 000 stands were studied; The proportion of damage doubled during the study period; Tree species mixture had a clear effect on the damage frequency; The damage was more common in mineral soils than in peatlands, in artificially than in naturally regenerated stands and in stands that needed thinning or clearing or in which soil preparation was used.
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The occurrence of moose damage was studied using data from three National Forest Inventories (NFIs) accomplished between 1986 and 2008 in Finland. The combined data included a total of 97 390 young stands. The proportion of moose damage increased from 3.6% to 8.6% between the 8th NFI (1986–1994) and the 10th NFI (2004–2008). The majority (75%) of the damage occurred in Scots pine-dominated stands. The proportion of damage was higher in aspen-dominated stands than in stands dominated by any other tree species. The tree species mixture also had a clear effect on the occurrence of damage. Pure Scots pine stands had less damage than mixed Scots pine stands, and moose damage decreased linearly with the increasing proportion of Scots pine. Stands on mineral soil had more frequent moose damage than stands on peatlands. The fertility class of the site had no straightforward effect on the damage frequency. Artificially regenerated stands had more damage than naturally regenerated stands. Accomplished soil preparation measures and the need for thinning or clearing operations increased moose damage. High proportions of moose damage in young stands were found around the country. In the 10th NFI, the largest concentration of damage was found in southwestern Finland. Our study shows the temporal and spatial changes in the occurrence of moose damage and pinpoints some important silvicultural factors affecting the relative risk of young stands over a large geographical area.

  • Nevalainen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: seppo.nevalainen@luke.fi (email)
  • Matala, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: juho.matala@luke.fi
  • Korhonen, Natural Resources Institute Finland (Luke), Economics and society, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: kari.t.korhonen@luke.fi
  • Ihalainen, Natural Resources Institute Finland (Luke), Economics and society, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: antti.ihalainen@luke.fi
  • Nikula, Natural Resources Institute Finland (Luke), Economics and society, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: ari.nikula@luke.fi
article id 1386, category Research article
Håkan Lideskog, Magnus Karlberg. (2016). Simulated continuous mounding improvements through ideal machine vision and control. Silva Fennica vol. 50 no. 2 article id 1386. https://doi.org/10.14214/sf.1386
Keywords: site preparation; silviculture; remote sensing; scarification; clearcut obstacles; mounding simulation; work procedures
Highlights: Different strategies for how to utilise machine vision to streamline the mounding head movements were developed and evaluated; The theoretical minimum rate of encountered obstacles while utilising machine vision in continuous mounding is presented, provided that an optimal continuous mounding has been performed; The needed minimum resolution of a machine vision system at work on a clearcut area was found.
Abstract | Full text in HTML | Full text in PDF | Author Info

To promote the growth and survival of regenerated forests, site preparation prior to tree planting on clearcuts is necessary. This is often performed with scarifiers, either through trenching or mounding. Mounding is generally considered better in a plant survival perspective but is inefficient on obstacle-rich clearcuts. By utilising machine vision through e.g. remote sensing methods, new strategies can enable efficient mound positioning. In this paper, three realistic strategies utilizing ideal clearcut object identification through machine vision have been developed that can be used for more efficient mounding. The results show that mounding efficiency can be significantly improved with a new mound positioning strategy that employs ideal object identification, especially on obstacle-rich clearcuts.

 

 

  • Lideskog, Luleå University of Technology, Department of Engineering Sciences and Mathematics, Division of Product and Production Development, SE-971 87 Luleå, Sweden E-mail: hakan.lideskog@ltu.se (email)
  • Karlberg, Luleå University of Technology, Department of Engineering Sciences and Mathematics, Division of Product and Production Development, SE-971 87 Luleå, Sweden E-mail: magnus.karlberg@ltu.se
article id 1323, category Research article
Tiina Laine, Kalle Kärhä, Antti Hynönen. (2016). A survey of the Finnish mechanized tree-planting industry in 2013 and its success factors. Silva Fennica vol. 50 no. 2 article id 1323. https://doi.org/10.14214/sf.1323
Keywords: regeneration; boreal; forestry; silviculture; mechanization; planting machines; critical success factors (CSFs)
Highlights: In 2013, 31 planting machines were operated by 22 businesses and planted 4.7 million seedlings on 2663 hectares in Finland; Critical success factors included expertise of planting machine operators, high quality planting, adequate amount of work, stoniness, and removal of slash; Growth of the industry will depend on improved cost-efficiency, appropriate worksites, marketing, development of planting machines.
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The aim of the study was to update the information pertaining to mechanized tree-planting activity in Finland in 2013 and its success factors. All businesses providing a mechanized tree-planting service were interviewed and asked to describe their equipment and activities, identify critical success factors (CSFs), and suggest areas for improvement. In 2013, 31 planting machines (18 Bracke P11.a, 11 M-Planter and 2 Risutec) operated by 22 businesses planted approximately 4.7 million seedlings on 2663 hectares. CSFs included expertise of planting machine operators, high quality planting, adequate amount of work, stoniness, and removal of slash. Based on the survey, some recommendations for improving mechanized planting work can be made. Firstly, improving the cost-efficiency of mechanized planting is necessary to enhance businesses’ profitability. Secondly, worksite selection is crucial as stoniness, stumps and slash debris diminish productivity. Lastly, the popularity of mechanized planting in the future will benefit from more marketing. Many businesses were unwilling to increase the area of service, invest in new equipment, or increase the volume of planting work but they believed that mechanized planting will become more popular in the near future.

  • Laine, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: tiina.laine@luke.fi (email)
  • Kärhä, Stora Enso Wood Supply Finland, P.O. Box 309, FI-00101 Helsinki, Finland E-mail: kalle.karha@storaenso.com
  • Hynönen, University of Eastern Finland, Faculty of Science and Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: hynonena@gmail.com
article id 1442, category Research article
Silva Šēnhofa, Mārtiņš Zeps, Roberts Matisons, Jānis Smilga, Dagnija Lazdiņa, Āris Jansons. (2015). Effect of climatic factors on tree-ring width of Populus hybrids in Latvia. Silva Fennica vol. 50 no. 1 article id 1442. https://doi.org/10.14214/sf.1442
Keywords: dendroclimatology; hybrid aspen; hybrid poplar; fast-growing hybrids; weather conditions
Highlights: Hybrid poplar and hybrid aspen were sensitive to temperature in summer and dormant periods, but none of the tested factors were strictly limiting; Hybrid poplar was sensitive to a higher number of climatic factors than hybrid aspen; Temperature showed a negative correlation with tree-ring width.
Abstract | Full text in HTML | Full text in PDF | Author Info

Fast-growing hybrids of Populus L. have an increasing importance as a source of renewable energy and as industrial wood. Nevertheless, the long-term sensitivity of Populus hybrids to weather conditions and hence to possible climatic hazards in Northern Europe have been insufficiently studied, likely due to the limited age of the trees (short rotation). In this study, the climatic sensitivity of ca. 65-year-old hybrid poplars (Populus balsamifera L. × P. laurifolia Ledeb.), growing at two sites in the western part of Latvia, and ca. 55-year-old hybrid aspens (Populus tremuloides Michx. × P. tremula L.), growing in the eastern part of Latvia, have been studied using classical dendrochronological techniques. The high-frequency variation of tree-ring width (TRW) of hybrid poplar from both sites was similar, but it differed from hybrid aspen due to the diverse parental species and geographic location of the stands. Nevertheless, some common tendencies in TRW were observed for both hybrids. Climatic factors influencing TRW were generally similar for both hybrids, but their composition differed. The strength of climate-TRW relationships was similar, but the hybrid poplar was affected by a higher number of climatic factors. Hybrid poplar was sensitive to factors related to water deficit in late summer in the previous and current years. Hybrid aspen was sensitive to conditions in the year of formation of tree-ring. Both hybrids also displayed a reaction to temperature during the dormant period. The observed climate-growth relationships suggest that increasing temperatures might burden the radial growth of the studied hybrids of Populus.

  • Šēnhofa, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: silva.senhofa@gmail.com
  • Zeps, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: martins.zeps@silava.lv
  • Matisons, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: robism@inbox.lv (email)
  • Smilga, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: janis.smilga@silava.lv
  • Lazdiņa, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: dagnija.lazdina@silava.lv
  • Jansons, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: aris.jansons@silava.lv
article id 1428, category Research article
Gernot Erber, Franz Holzleitner, Maximilian Kastner, Karl Stampfer. (2015). Effect of multi-tree handling and tree-size on harvester performance in small-diameter hardwood thinnings. Silva Fennica vol. 50 no. 1 article id 1428. https://doi.org/10.14214/sf.1428
Keywords: productivity; thinning; energy wood; motor-manual; accumulating harvester head; delimbing
Highlights: Harvesting with the accumulating energy wood head EF28 was studied under small tree dimension (8 dm3) in hardwood thinnings; Reasonable productivity was achieved; Maximum achieved cutting diameter in hornbeam stand was 23 cm and 15% lower than in softwood stands; Head has potential under such conditions.
Abstract | Full text in HTML | Full text in PDF | Author Info

Early thinnings are laborious and costly. Thus forest companies are searching for cost and time efficient ways to carry out this task. The study’s purpose was to determine the productivity of the EF28 accumulating energy wood harvesting head in harvesting small-diameter hornbeam (Carpinus betulus L.) undergrowth trees and evaluate the effect of its multi-tree handling (MTH) capacity on time consumption. The harvester was a wheeled, three-axle Komatsu 911. A time study of 7.1 hours on 19 plots, with a total area of 0.76 ha was conducted. On average, the harvested tree volume was 8 dm³ and the stand density was 2666 trees/ha. The productivity was modelled with MTH conduction, mean diameter at breast height and the number of trees handled per cycle as independent variables. On average, MTH took 27% longer per cycle, increased extracted volume per cycle by 33% and consequently increased productivity with 5.0%. In 71.9% of the cycles more than one tree was handled and if so, dimensions were smaller than in single-tree handling (5.8 cm vs. 12.0 cm). Maximum felling diameter of 23 cm was about 15% smaller than in softwood (according to the manufacturer’s specifications) and the driver didn’t exploit the EF28’s theoretical potential in terms of trees handled per cycle. It can be concluded that the head could significantly improve productivity in small-diameter wood procurement.

  • Erber, Addresses University of Natural Resources and Life Sciences Vienna, Department of Forest and Soil Sciences, Institute of Forest Engineering, Peter-Jordan Straße 82/3, A-1190 Vienna, Austria E-mail: gernot.erber@boku.ac.at (email)
  • Holzleitner, Addresses University of Natural Resources and Life Sciences Vienna, Department of Forest and Soil Sciences, Institute of Forest Engineering, Peter-Jordan Straße 82/3, A-1190 Vienna, Austria E-mail: franz.holzleitner@boku.ac.at
  • Kastner, Addresses University of Natural Resources and Life Sciences Vienna, Department of Forest and Soil Sciences, Institute of Forest Engineering, Peter-Jordan Straße 82/3, A-1190 Vienna, Austria E-mail: maximilian.kastner@boku.ac.at
  • Stampfer, Addresses University of Natural Resources and Life Sciences Vienna, Department of Forest and Soil Sciences, Institute of Forest Engineering, Peter-Jordan Straße 82/3, A-1190 Vienna, Austria E-mail: karl.stampfer@boku.ac.at
article id 1413, category Research article
Ilya Potapov, Marko Järvenpää, Markku Åkerblom, Pasi Raumonen, Mikko Kaasalainen. (2015). Data-based stochastic modeling of tree growth and structure formation. Silva Fennica vol. 50 no. 1 article id 1413. https://doi.org/10.14214/sf.1413
Keywords: terrestrial lidar; form diversity; morphological plasticity; stochastic functional-structural plant model; quantitative structure models; data fitting
Highlights: We propose a stochastic version of the tree growth model LIGNUM for producing tree structures consistent with detailed terrestrial laser scanning data, and we provide the proof-of-concept by using model-based simulations and real laser scanning data; Trees produced with the data-based model resemble the trees of the dataset, and are statistically similar but not copies of each other; the number of such synthetic trees is not limited.
Abstract | Full text in HTML | Full text in PDF | Author Info

We introduce a general procedure to match a stochastic functional-structural tree model (here LIGNUM augmented with stochastic rules) with real tree structures depicted by quantitative structure models (QSMs) based on terrestrial laser scanning. The matching is done by iteratively finding the maximum correspondence between the measured tree structure and the stochastic choices of the algorithm. First, we analyze the match to synthetic data (generated by the model itself), where the target values of the parameters to be estimated are known in advance, and show that the algorithm converges properly. We then carry out the procedure on real data obtaining a realistic model. We thus conclude that the proposed stochastic structure model (SSM) approach is a viable solution for formulating realistic plant models based on data and accounting for the stochastic influences.

  • Potapov, Tampere University of Technology, Department of Mathematics, P.O. Box 553, FI-33101 Tampere, Finland E-mail: ilya.potapov@tut.fi (email)
  • Järvenpää, Tampere University of Technology, Department of Mathematics, P.O. Box 553, FI-33101 Tampere, Finland E-mail: marko.jarvenpaa@tut.fi
  • Åkerblom, Tampere University of Technology, Department of Mathematics, P.O. Box 553, FI-33101 Tampere, Finland E-mail: markku.akerblom@tut.fi
  • Raumonen, Tampere University of Technology, Department of Mathematics, P.O. Box 553, FI-33101 Tampere, Finland E-mail: pasi.raumonen@tut.fi
  • Kaasalainen, Tampere University of Technology, Department of Mathematics, P.O. Box 553, FI-33101 Tampere, Finland E-mail: mikko.kaasalainen@tut.fi
article id 1406, category Research article
Tatiana V. Stankova. (2015). A dynamic whole-stand growth model, derived from allometric relationships. Silva Fennica vol. 50 no. 1 article id 1406. https://doi.org/10.14214/sf.1406
Keywords: Scots pine; state-space approach; transition function; algebraic difference approach; growth projection; Base-Height-Invariant dummy variable method; even-aged stands
Highlights: A dynamic whole-stand model was derived from simple allometries and biological rationale; The state-space modelling approach was applied, suggesting several novelties to overcome scarcity of longitudinal data; The model consists of a three-dimensional state vector, defined by dominant stand height, stand density and mean stem volume, and three transition functions; It was tested with data from Scots pine plantations.
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Growth and yield tables are constrained by a standard production regime and the stand-level dynamic models are an attractive alternative for the even-aged monospecific stands. The objective of this study is to derive a parsimonious and biologically sound whole-stand dynamic growth model and to validate its consistency and relevance for prediction of stand growth and yield. The state-space modelling approach was employed, introducing several novelties in comparison with its current usage. The model consists of a three-dimensional state vector, defined by dominant stand height, number of trees per hectare and mean stem volume, and three transition functions. A site index model was incorporated for height growth projection and transition functions for stand density and mean stem volume with respect to dominant height increase were derived from simple allometries and biological rationale. The model was examined with data from 79 permanent sample plots in Scots pine plantations. Nonlinear Seemingly Unrelated Regression was used to account for cross-equation correlations, and the Base-Height-Invariant dummy variable method was employed to estimate dynamic-form equations. Model consistency was validated in terms of accuracy of predictions and applicability to both thinned and unthinned stands. The new dynamic growth model is a parsimonious biometric whole-stand model that simulates the expected stand development for a broad spectrum of potential management alternatives and can be incorporated in a computer program for further use. It may be especially useful for application when a scarcity of longitudinal data prevents the use of more complicated modelling approaches.

  • Stankova, Forest Research Institute of Bulgarian Academy of Sciences, Department of Forest Genetics, Physiology and Plantations, 132 “St. Kliment Ohridski” Blvd., 1756 Sofia, Bulgaria ORCID http://orcid.org/0000-0002-9932-7286 E-mail: tatianastankova@yahoo.com (email)
article id 1393, category Research article
Werner Poschenrieder, Andreas Rais, Jan-Willem G. van de Kuilen, Hans Pretzsch. (2015). Modelling sawn timber volume and strength development at the individual tree level – essential model features by the example of Douglas fir. Silva Fennica vol. 50 no. 1 article id 1393. https://doi.org/10.14214/sf.1393
Keywords: sawing; wood quality modelling; stem structure; grading; sensitivity test; R statistical environment
Highlights: An individual tree timber growth and quality model toolbox was designed; It realistically predicts an increase of bending strength with planting density; Prediction was shown to be based on consideration of essential intrinsic variables; Height‑diameter‑allometry depending on planting density was effective; Consideration of cambial age and knot area ratio was crucial.
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We designed a streamlined timber growth and quality model that aims at the effect of stand management on the efficiency of wood resource use. Applying the R based module toolbox to experimental plots of Douglas fir (Pseudotsuga menziesii [Mirb.] Franco) we analysed essential model features for reflecting the influence of planting density on board strength. The current version realistically predicted a significant increase of centre board bending strength at tree age 40 with initial stand density. Model performance gained clear advantage from a) parameterisation of height to diameter allometry as dependent on planting density b) consideration of cambial age and cross‑sectional knot area in board strength computation. Crown shape was less decisive. The model produced a significant effect of planting density even after a whole rotation period of 70 years as well as a realistic spectrum of board bending strength.

  • Poschenrieder, Technische Universität München, Chair of Forest Growth and Yield Science, Hans Carl von Carlowitz Platz 2, 85354 Freising, Germany ORCID http://orcid.org/0000-0002-9028-8583 E-mail: Werner.Poschenrieder@lrz.tum.de (email)
  • Rais, Technische Universität München, Chair of Forest Growth and Yield Science, Hans Carl von Carlowitz Platz 2, 85354 Freising, Germany; Technische Universität München, Holzforschung München, Winzererstrasse 45, 80797 Munich, Germany E-mail: Andreas.Rais@hfm.tum.de
  • van de Kuilen, Technische Universität München, Holzforschung München, Winzererstrasse 45, 80797 Munich, Germany E-mail: vandekuilen@hfm.tum.de
  • Pretzsch, Technische Universität München, Chair of Forest Growth and Yield Science, Hans Carl von Carlowitz Platz 2, 85354 Freising, Germany E-mail: Hans.Pretzsch@lrz.tum.de
article id 1384, category Research article
Staffan Berg, Erik Valinger, Torgny Lind, Tommi Suominen, Diana Tuomasjukka. (2015). Comparison of co-existing forestry and reindeer husbandry value chains in northern Sweden. Silva Fennica vol. 50 no. 1 article id 1384. https://doi.org/10.14214/sf.1384
Keywords: carbon sequestration; agroforestry; global warming potential; gross value added; employment; ToSIA
Highlights: Forestry adapted to reindeer husbandry results in: potential economic improvement of reindeer husbandry, potential reduced cuttings in forestry and reduced wood flow to industry, reduced gross value added for forest industry and increased carbon storage in standing forest.
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Forestry in Malå, northern Sweden, coexists with other land uses. Reindeer husbandry is in the area for centuries and requires large areas of grazing land. Competing land uses may threaten the Malå Sami village. The aim of the study was to evaluate increased consideration in forest management towards 1) reindeer husbandry, 2) nature and 3) a combination of the two. These scenarios were compared with forest management as it was in 2009. Results indicate that all three scenarios lead to a decrease in annual harvesting volumes of 0.2 to 0.4 million m3. Forest industry dominated the economic viability in the area. Forest management adapted to the needs of reindeer husbandry resulted in less potential for yearly harvest, employment and profits from forest industry. On the other hand, it led to an increase in growing stock and consequently the potential for carbon sequestration over time. Indeed the increased sequestration would compensate for all fossil emissions of carbon from the Forest Wood Chain (FWC). The nature scenario had minor effects on economic result and on the emissions of fossil carbon. The combined scenario gave a reduced economic performance for the FWC. A scenario based on forest management accommodating the needs of reindeer husbandry gave the best economic result for the reindeer chain, due to high survival rate of the reindeer. However the economic importance of reindeer husbandry in the region was small compared to the FWC. Results from scenario analysis could serve as a platform for mutual understanding between stakeholders.

  • Berg, Swedish University of Agricultural Sciences (SLU), Department of Forest Ecology and Management, Skogsmarksgränd, SE-90 183 Umeå, Sweden E-mail: staffan.berg@efi.int (email)
  • Valinger, Swedish University of Agricultural Sciences (SLU), Department of Forest Ecology and Management, Skogsmarksgränd, SE-90 183 Umeå, Sweden E-mail: erik.valinger@slu.se
  • Lind, Swedish University of Agricultural Sciences (SLU), Department of Forest Resource Management, Skogsmarksgränd, SE-90 183 Umeå, Sweden E-mail: torgny.lind@slu.se
  • Suominen, European Forest Institute, Sustainability and Climate Change Research Programme, Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: tommi.suominen@efi.int
  • Tuomasjukka, European Forest Institute, Sustainability and Climate Change Research Programme, Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: diana.tuomasjukka@efi.int
article id 1341, category Research article
Přemysl Humplík, Petr Čermák, Tomáš Žid. (2016). Electrical impedance tomography for decay diagnostics of Norway spruce (Picea abies): possibilities and opportunities. Silva Fennica vol. 50 no. 1 article id 1341. https://doi.org/10.14214/sf.1341
Keywords: sapwood; heartwood; tomogram; decay proportion; impedance dataset
Highlights: Statistical parameters of EIT datasets with values of electrical resistance of heartwood are possible to employ in refining heartwood rot diagnostics; Sapwood proportion is decreasing as the proportion of decay on the radial cut expands; Using EIT datasets and sapwood proportion, trees with rot can be split into two groups as per proportion of decay: [< 35%] and [> 35%].
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The paper aimed at testing the potential of refining tree rot diagnostics carried out by means of electrical impedance tomography (EIT). Examined was the use of EIT datasets with electrical resistance values and sapwood proportion determined on the basis of tomograms. Making use of datasets with resistance values in EIT rot diagnostics is not a default method, although datasets stay unaffected by a fixed colour scale and subsequent subjective evaluation unlike tomograms. Tomography measurement was carried out for 27 individuals of Norway spruce (Picea abies [L.] Karst.) in two stands north-east of Brno, Czech Republic. Once felled down, radial cut-outs were sampled at the measurement site and used for localising rot and determining the extent of the area of decay. The results were subsequently compared with tomograms. EIT datasets containing values of electrical resistance found by measuring were statistically processed and compared with the extent of rot area identified within the cuts. Sapwood proportion values were also detected using the tomograms. The baseline assumption that sapwood proportion decreases as the rot area in the radial cut expands was confirmed. In trees with rot percentage to 35% approximately, sapwood proportion was exceeding 30% except one tree. In trees with rot percentage exceeding 35%, sapwood proportion was below 30%. On the basis of interpreted datasets, the trees can be split into three characteristic groups that correspond to the occurrence, extent and nature of the rot.

  • Humplík, Mendel University in Brno, Faculty of Forestry and Wood Technology, Zemědělská 1665/3, 613 00 Brno, Czech Republic E-mail: premysl.humplik@mendelu.cz (email)
  • Čermák, Mendel University in Brno, Faculty of Forestry and Wood Technology, Zemědělská 1665/3, 613 00 Brno, Czech Republic E-mail: petr.cermak@mendelu.cz
  • Žid, Mendel University in Brno, Faculty of Forestry and Wood Technology, Zemědělská 1665/3, 613 00 Brno, Czech Republic E-mail: tomas.zid@mendelu.cz
article id 1405, category Research article
Lauri Korhonen, Daniela Ali-Sisto, Timo Tokola. (2015). Tropical forest canopy cover estimation using satellite imagery and airborne lidar reference data. Silva Fennica vol. 49 no. 5 article id 1405. https://doi.org/10.14214/sf.1405
Keywords: logistic regression; beta regression; forest area; international forest definition; ALOS AVNIR-2; vegetation index
Highlights: The fusion of airborne lidar data and satellite images enables accurate canopy cover mapping; The zero-and-one inflated beta regression is demonstrated in large area estimation; Forest/non-forest classification should be done directly, for example by using logistic regression.
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The fusion of optical satellite imagery, strips of lidar data and field plots is a promising approach for the inventory of tropical forests. Airborne lidars also enable an accurate direct estimation of the forest canopy cover (CC), and thus a sample of lidar strips can be used as reference data for creating CC maps which are based on satellite images. In this study, our objective was to validate CC maps obtained from an ALOS AVNIR-2 satellite image wall-to-wall, against a lidar-based CC map of a tropical forest area located in Laos. The reference CC values which were needed for model training were obtained from a sample of four lidar strips. Zero-and-one inflated beta regression (ZOINBR) models were applied to link the spectral vegetation indices derived from the ALOS image with the lidar-based CC estimates. In addition, we compared ZOINBR and logistic regression models in the forest area estimation by using >20% CC as a forest definition. Using a total of 409 217 30 × 30 m population units as validation, our model showed a strong correlation between lidar-based CC and spectral satellite features (root mean square error = 12.8%, R2 = 0.82). In the forest area estimation, a direct classification using logistic regression provided better accuracy than the estimation of CC values as an intermediate step (kappa = 0.61 vs. 0.53). It is important to obtain sufficient training data from both ends of the CC range. The forest area estimation should be done before the CC estimation, rather than vice versa.

  • Korhonen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland; (current) University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID http://orcid.org/0000-0002-9352-0114 E-mail: lauri.z.korhonen@helsinki.fi (email)
  • Ali-Sisto, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: dheikkil@student.uef.fi
  • Tokola, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland. E-mail: timo.tokola@uef.fi
article id 1403, category Research article
Kristina Mjöfors, Monika Strömgren, Hans-Örjan Nohrstedt, Annemieke Ingrid Gärdenäs. (2015). Impact of site-preparation on soil-surface CO2 fluxes and litter decomposition in a clear-cut in Sweden. Silva Fennica vol. 49 no. 5 article id 1403. https://doi.org/10.14214/sf.1403
Keywords: boreal forest; soil disturbance; soil respiration; carbon stock; needle decomposition; coarse root decomposition
Highlights: Disturbances of the soil did not lead to higher CO2 emissions from the soil; Heavy mixing of the soil lead to lower CO2 emissions from the soil; Buried needles and coarse roots decomposed faster than those on the surface; Abundance of δ15N decreased in needles and roots after site preparation.
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Boreal forest soil contains significant amounts of organic carbon. Soil disturbance, caused for example by site preparation or stump extraction, may increase decomposition and thus lead to higher CO2 emissions, contributing to global warming. The aim of this study was to quantify responses of soil-surface CO2 fluxes (Rs) and litter (needle and root) decomposition rates following various kinds of soil disturbance commonly caused by mechanical site preparation and stump harvest. For this purpose four treatments were applied in a clear-cut site in central Sweden: i) removal of the humus layer and top 2 cm of mineral soil, ii) placement of a humus layer and 2 cm of mineral soil upside down on top of undisturbed soil, forming a double humus layer buried under mineral soil, iii) heavy mixing of the humus layer and mineral soil, and iv) no disturbance (control). Rs measurements were acquired with a portable respiration system during two growing seasons. To assess the treatments’ effects on litter decomposition rates, needles or coarse roots (Ø = 6 mm) were incubated in litterbags at positions they would be located after the treatments (buried, or on top of the soil). The results indicate that site preparation-simulating treatments have no effect or may significantly reduce, rather than increase, CO2 emissions during the following two years. They also show that buried litter decomposes more rapidly than litter on the surface, but in other respects the treatments have little effect on litter decomposition rates.

  • Mjöfors, Swedish University of Agricultural Sciences (SLU), Department of Soil and Environment, P.O. Box 7014, 150 07 Uppsala, Sweden E-mail: kristina.mjofors@slu.se (email)
  • Strömgren, Swedish University of Agricultural Sciences (SLU), Department of Soil and Environment, P.O. Box 7014, 150 07 Uppsala, Sweden E-mail: Monika.stromgren@slu.se
  • Nohrstedt, Swedish University of Agricultural Sciences (SLU), Department of Soil and Environment, P.O. Box 7014, 150 07 Uppsala, Sweden E-mail: Hans-orjan.nohrstedt@slu.se
  • Gärdenäs, Swedish University of Agricultural Sciences (SLU), Department of Soil and Environment, P.O. Box 7014, 150 07 Uppsala, Sweden E-mail: Annemieke.gardenas@slu.se
article id 1395, category Research article
Joseph Buongiorno. (2015). Income and time dependence of forest product demand elasticities and implications for forecasting. Silva Fennica vol. 49 no. 5 article id 1395. https://doi.org/10.14214/sf.1395
Keywords: modeling; structural change; econometrics; markets
Highlights: Elasticities of demand with gross domestic product and prices were stable over time and income level for sawnwood and particleboard only; Other product elasticities differed with income and time, leading in conjunction with a sector model to higher projected world demand and prices than obtained by ignoring differences between countries and over time.
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In view of improving multi-country forest sector models, this study investigated to what extent the price and income elasticity of demand for forest products had changed in the past two decades, and how much they depended on the countries income level. For each of seven major product groups annual observations were divided between high-income (top 20% in gross domestic product per capita) and low-income, and between recent (2004–2013) and older (1992–2003) observations. The results indicated that for sawnwood and particleboard the data could be pooled across all countries and years. For the other commodity groups (veneer & plywood, fiberboard, newsprint, printing & writing paper, other paper & paperboard), there were statistically significant differences in gross domestic product or price elasticity between high and low-income levels or old and recent observations. Efficient elasticities were obtained by pooling the maximum number of observations while respecting the statistically significant differences. The resulting GDP elasticities were the same, or very close, across income levels for all products. The price elasticities differed by income level only for newsprint and for veneer and plywood. International forest sector projections to 2065 obtained with these elasticities compared with those based on pooling all data across time and income levels gave less than 3% difference for world consumption of sawnwood, particleboard, fiberboard, and newsprint, but 19% higher consumption for veneer and plywood, 31% for printing and writing paper, and 18% for other paper and paperboard. The world price was 1% to 11% higher for end products and 3% to 22% higher for raw materials and intermediate products.

  • Buongiorno, University of Wisconsin-Madison, Department of Forest and Wildlife Ecology, 1630 Linden Drive, Madison, WI 53706, USA E-mail: jbuongio@wisc.edu (email)
article id 1394, category Research article
Sari Karvinen, Tuomas Nummelin. (2015). Finnish wood harvesting contractors’ risks in Russia. Silva Fennica vol. 49 no. 5 article id 1394. https://doi.org/10.14214/sf.1394
Keywords: Russia; contracting; wood harvesting; risk analysis
Highlights: Disagreements on wood measurement and payment delays were the most important economic risks; Dependency on a few clients created risk for unfavourable agreements and work interruptions; Fires in site huts caused the risk of personal injury; Inadequate professional skills were serious economic and work interruption risks; Unhealthy competition, the functioning of the authorities, and infrastructure were important external risk factors.
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Finnish wood harvesting contractors have been working in Russia since the 1990s and new entrepreneurs are still interested in starting operations there, even though Russia is not an easy business environment. This study identifies the most significant risks in contracting in Russia. Risks were identified through expert evaluation and a risk analysis was conducted by using a risk matrix. Possible preventative measures were assessed for the identified risks. Some risks were found to be common in Russia and Finland, for example a limited number of clients, dependency on a few clients, and weak negotiating positions. A stable amount of work, i.e. the availability of stands for harvesting, was also a challenge on the both sides of border. Typical problems in Russia were breaches of contract, especially disagreements on wood measurement and payment delays, potentially causing serious economic losses. Specific to Russia were problems related to machine service and spare parts, as well as security issues. The professional skills of machine operators, as well as changing work motivation were risks in Russia. Cultural differences lead to more challenging supervision and management of staff. Among the external factors, the most challenging in Russia were unhealthy competition in the marketplace and non-transparent and the unpredictable procedures of the authorities. In Russia problems caused by seasonality are amplified by the sparse road network and longer downtime. The revealed specific features of the Russian business environment can help Finnish wood harvesting companies to plan a risk management process for operations in Russia.

  • Karvinen, Natural Resources Institute Finland (Luke), Economics and society, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: sari.karvinen@luke.fi (email)
  • Nummelin, Natural Resources Institute Finland (Luke), Green technology, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: tuomas.nummelin@luke.fi
article id 1382, category Research article
Juha Laitila, Johanna Routa. (2015). Performance of a small and a medium sized professional chippers and the impact of storage time on Scots pine (Pinus sylvestris) stem wood chips characteristics. Silva Fennica vol. 49 no. 5 article id 1382. https://doi.org/10.14214/sf.1382
Keywords: chipping; productivity; quality; fuel; particle size; storage time; sieve; stem wood
Highlights: The storage time of pulpwood had no significant effect on particle size distribution in any chip size classes; The study confirms the knowledge that chipping time consumption is inversely proportional to engine power and grapple load size in feeding; The use of an narrower 80 mm × 80 mm sieve on Scots pine material does not seem to offer any benefit compared to a 100 mm × 100 mm sieve from the perspective of chip quality.
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The primary aim of this study was to clarify the chipping productivity and fuel consumption of tractor-powered and truck-mounted drum chippers when chipping pine pulpwood at a terminal. The secondary aim was to evaluate the impact of wood storage time on the chemical and physical technical specifications of wood chips by chipping pulpwood from eight different storage time groups, using Scots pine (Pinus sylvestris) pulpwood stems logged between 2 and 21 months previously at the terminal with the above-mentioned chippers. Thirdly, the impact of sieve mesh size on the particle size distribution of wood chips from different age groups was compared by using an 80 mm × 80 mm sieve for a tractor-powered chipper and a 100 mm × 100 mm sieve for a truck-mounted chipper. With both chippers, the chipping productivity grew as a function of grapple load weight. The average chipping productivity of the tractor-powered chipper unit was 19 508 kg (dry mass) per effective hour (E0h), and for the truck-mounted chipper the average productivity was 31 184 kg E0h–1. The tractor-powered drum chipper’s fuel consumption was 3.1 litres and for the truck-mounted chipper 3.3 litres per chipped 1000 kg (dry mass). The amount of extractives or volatiles did not demonstrate any statistically significant differences between storage time groups. The particle size distributions with both chippers were quite uniform, and the storage time of pulpwood did not have a significant effect on the particle size distribution in any chip size classes. One reason for this might be that the basic density of chipped wood was homogenous and there was no statistical difference between different storage times. The use of new sharp knives is likely to have affected chip quality, as witnessed by the absence of oversized particles and the moderate presence of fines. The use of narrower 80 mm × 80 mm sieves on Scots pine material does not seem to offer any benefit compared to 100 mm × 100 mm from the chip quality point of view.

  • Laitila, Natural Resources Institute Finland (Luke), Bio-based business and industry, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: juha.laitila@luke.fi (email)
  • Routa, Natural Resources Institute Finland (Luke), Bio-based business and industry, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: johanna.routa@luke.fi
article id 1377, category Research article
Raul Fernandez-Lacruz, Fulvio Di Fulvio, Dimitris Athanassiadis, Dan Bergström, Tomas Nordfjell. (2015). Distribution, characteristics and potential of biomass-dense thinning forests in Sweden. Silva Fennica vol. 49 no. 5 article id 1377. https://doi.org/10.14214/sf.1377
Keywords: bioenergy; GIS; wood fuel; early thinning; small-tree harvesting; biomass potential; biomaterial
Highlights: Biomass-dense thinning forests (BDTF) cover 2.1–9.8 M ha in Sweden, which represents 9–44% of the country’s productive forest land area, depending on the constraints applied; 65% of BDTF area is found in northern Sweden; Analyses revealed a yearly harvesting potential of at least 4.3 M OD t of undelimbed whole trees (3.0 M OD t of delimbed stemwood including tops).
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Understanding the characteristics of unutilized biomass resources, such as small-diameter trees from biomass-dense thinning forests (BDTF) (non-commercially-thinned forests), can provide important information for developing a bio-based economy. The aim of this study was to describe the areal distribution, characteristics (biomass of growing stock, tree height, etc.) and harvesting potential of BDTF in Sweden. A national forest inventory plot dataset was imported into a geographical information system and plots containing BDTF were selected by applying increasingly stringent constraints. Results show that, depending on the constraints applied, BDTF covers 9–44% (2.1–9.8 M ha) of the productive forest land area, and contains 7–34% of the total growing stock (119–564 M OD t), with an average biomass density of 57 OD t ha–1. Of the total BDTF area, 65% is located in northern Sweden and 2% corresponds to set-aside farmlands. Comparisons with a study from 2008 indicate that BDTF area has increased by at least 4% (about 102 000 ha), in line with general trends for Sweden and Europe. Analyses revealed that the technical harvesting potential of delimbed stemwood (over bark, including tops) from BDTF ranges from 3.0 to 6.1 M OD t yr–1 (7.5 to 15.1 M m3 yr–1), while the potential of whole-tree harvesting ranges from 4.3 to 8.7 M OD t yr–1 (10.2 to 20.6 M m3 yr–1) depending on the scenario considered. However, further technological developments of the harvest and supply systems are needed to utilize the full potential of BDTF.

  • Fernandez-Lacruz, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden ORCID http://orcid.org/0000-0001-9284-8911 E-mail: raul.fernandez@slu.se (email)
  • Di Fulvio, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden; International Institute for Applied Systems Analysis (IIASA), Ecosystems Services and Management Program (ESM), Schlossplatz 1, A-2361 Laxenburg, Austria E-mail: Fulvio.di.Fulvio@slu.se
  • Athanassiadis, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: Dimitris.Athanassiadis@slu.se
  • Bergström, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: Dan.Bergstrom@slu.se
  • Nordfjell, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: Tomas.Nordfjell@slu.se
article id 1348, category Research article
Sakari Tuominen, Andras Balazs, Heikki Saari, Ilkka Pölönen, Janne Sarkeala, Risto Viitala. (2015). Unmanned aerial system imagery and photogrammetric canopy height data in area-based estimation of forest variables. Silva Fennica vol. 49 no. 5 article id 1348. https://doi.org/10.14214/sf.1348
Keywords: forest inventory; aerial imagery; unmanned aerial system; UAV; photogrammetric surface model; canopy height model
Highlights: Orthoimage mosaic and 3D canopy height model were derived from UAV-borne colour-infrared digital camera imagery and ALS-based terrain model; Features extracted from orthomosaic and canopy height data were used for estimating forest variables; The accuracy of forest estimates was similar to that of the combination of ALS and digital aerial imagery.
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In this paper we examine the feasibility of data from unmanned aerial vehicle (UAV)-borne aerial imagery in stand-level forest inventory. As airborne sensor platforms, UAVs offer advantages cost and flexibility over traditional manned aircraft in forest remote sensing applications in small areas, but they lack range and endurance in larger areas. On the other hand, advances in the processing of digital stereo photography make it possible to produce three-dimensional (3D) forest canopy data on the basis of images acquired using simple lightweight digital camera sensors. In this study, an aerial image orthomosaic and 3D photogrammetric canopy height data were derived from the images acquired by a UAV-borne camera sensor. Laser-based digital terrain model was applied for estimating ground elevation. Features extracted from orthoimages and 3D canopy height data were used to estimate forest variables of sample plots. K-nearest neighbor method was used in the estimation, and a genetic algorithm was applied for selecting an appropriate set of features for the estimation task. Among the selected features, 3D canopy features were given the greatest weight in the estimation supplemented by textural image features. Spectral aerial photograph features were given very low weight in the selected feature set. The accuracy of the forest estimates based on a combination of photogrammetric 3D data and orthoimagery from UAV-borne aerial imaging was at a similar level to those based on airborne laser scanning data and aerial imagery acquired using purpose-built aerial camera from the same study area.

  • Tuominen, Natural Resources Institute Finland (Luke), Economics and society, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: sakari.tuominen@luke.fi (email)
  • Balazs, Natural Resources Institute Finland (Luke), Economics and society, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: andras.balazs@luke.fi
  • Saari, VTT Technical Research Centre of Finland Ltd, P.O. Box 1000, FI-02044 VTT, Finland E-mail: Heikki.Saari@vtt.fi
  • Pölönen, University of Jyväskylä, Department of Mathematical Information Technology, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: ilkka.polonen@jyu.fi
  • Sarkeala, Mosaicmill Oy, Kultarikontie 1, FI-01300 Vantaa, Finland E-mail: janne.sarkeala@mosaicmill.com
  • Viitala, Häme University of Applied Sciences (HAMK), P.O. Box 230, FI-13101 Hämeenlinna, Finland E-mail: Risto.Viitala@hamk.fi
article id 1347, category Research article
Paulo Borges, Even Bergseng, Tron Eid, Terje Gobakken. (2015). Impact of maximum opening area constraints on profitability and biomass availability in forestry – a large, real world case. Silva Fennica vol. 49 no. 5 article id 1347. https://doi.org/10.14214/sf.1347
Keywords: bioenergy; forest management planning; mixed integer programming; area restriction model; green-up
Highlights: We solved a large and real world near city forestry problem; The inclusion of maximum open area constraints caused 7.0% loss in NPV; Solution value at maximum deviated 0.01% from the true optimum value; The annual energy supply of 20–30 GWh estimated from harvest residues could provide a small, but stable supply of energy to the municipality.
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The nature areas surrounding the capital of Norway (Oslomarka), comprising 1 700 km2 of forest land, are the recreational home turf for a population of 1.2 mill. people. These areas are highly valuable, not only for recreational purposes and biodiversity, but also for commercial activities. To assess the impacts of the challenges that Oslo municipality forest face in their management, we developed four optimization problems with different levels of management constraints. The constraints consider control of harvest level, guarantee of minimum old-growth forest area and maximum open area after final harvest. For the latter, to date, no appropriate analyses quantifying the impact of such a constraint on economy and biomass production have been carried out in Norway. The problem solved is large due to both the number of stands and number of treatment schedules. However, the model applied demonstrated its relevance for solving large problems involving maximum opening areas. The inclusion of maximum open area constraints caused 7.0% loss in NPV compared to the business as usual case with controlled harvest volume and minimum old-growth area. The estimated supply of 20-30 GWh annual energy from harvest residues could provide a small, but stable supply of energy to the municipality.

  • Borges, Department of Ecology and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, 1432 Aas, Norway E-mail: paulo.borges@nmbu.no (email)
  • Bergseng, Department of Ecology and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, 1432 Aas, Norway E-mail: even.bergseng@nmbu.no
  • Eid, Department of Ecology and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, 1432 Aas, Norway E-mail: tron.eid@nmbu.no
  • Gobakken, Department of Ecology and Natural Resource Management, Norwegian University of Life Sciences, P.O. Box 5003, 1432 Aas, Norway E-mail: terje.gobakken@nmbu.no
article id 1342, category Research article
Blas Mola-Yudego, Gianni Picchi, Dominik Röser, Raffaele Spinelli. (2015). Assessing chipper productivity and operator effects in forest biomass operations. Silva Fennica vol. 49 no. 5 article id 1342. https://doi.org/10.14214/sf.1342
Keywords: mixed models; efficiency; comminution; operator effect; raw material supply; wood chippers
Highlights: A model is constructed to assess the productivity in chipping of wood biomass at roadside; The data includes 172 trials and 67 operators in Italy; The operator effect was included in a mixed model approach; The R2 were 0.76 (fixed part) and 0.88 (incl. operator effects).
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The present research focuses on the productivity of energy wood chipping operations at several sites in Italy. The aim was to assess the productivity and specifically the effect attributed to the operator in the chipping of wood biomass. The research included 172 trials involving 67 operators across the country that were analysed using a mixed model approach, in order to assess productivity, and to isolate the operator effect from other potential variables. The model was constructed using different predictors aiming to explain the variability due to the machines and the raw-materials. The final model included the average piece weight of raw material chipped as well as the power of the machine. The coefficients of determination (R2) were 0.76 for the fixed part of the model, and 0.88 when the effects due to the operators were included. The operators’ performance compared to their peers was established, and it was compared to a subjective classification based on the operator’s previous experience. The results of this study can help to the planning and logistics of raw material supply for bioenergy, as well as to a more effective training of future forest operators.

  • Mola-Yudego, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland; NIBIO Norwegian Institute of Bioeconomy Research, P.O. Box 115, 1431 Ås, Norway ORCID http://orcid.org/0000-0003-0286-0170 E-mail: blas.mola@uef.fi (email)
  • Picchi, CNR IVALSA, Via Madonna del Piano 10, I-50019 Sesto Fiorentino, Italy E-mail: picchi@ivalsa.cnr.it
  • Röser, Forest Feedstocks Group, FPInnovations, Vancouver, British Columbia, Canada E-mail: dominik.roser@fpinnovations.ca
  • Spinelli, CNR IVALSA, Via Madonna del Piano 10, I-50019 Sesto Fiorentino, Italy E-mail: spinelli@ivalsa.cnr.it
article id 1340, category Research article
Mostafa Farhadi, Mulualem Tigabu, Per Christer Odén. (2015). Near Infrared Spectroscopy as non-destructive method for sorting viable, petrified and empty seeds of Larix sibirica. Silva Fennica vol. 49 no. 5 article id 1340. https://doi.org/10.14214/sf.1340
Keywords: larch; NIRS; OPLS; precision sowing; seed sorting; seed quality
Highlights: Near Infrared spectroscopy discriminates filled-viable, empty and petrified seeds of Larix sibirica with 98%, 82% and 87% accuracy, respectively based on spectral differences attributed to moisture and storage reserves; The classification accuracy reached 100% when sorting seeds into viable and non-viable class; The results demonstrate that NIR spectroscopy has great potential as non-destructive sorting technique to upgrade seed lot quality.
Abstract | Full text in HTML | Full text in PDF | Author Info

Larix sibirica Ledeb. is one of the promising timber species for planting in the boreal ecosystem; but poor seed lot quality is the major hurdle for production of sufficient quantity of planting stocks. Here, we evaluated the potential of Near Infrared (NIR) Spectroscopy for sorting viable and non-viable seeds, as the conventional sorting technique is inefficient. NIR reflectance spectra were collected from single seeds, and discriminant models were developed with Orthogonal Projections to Latent Structure – Discriminant Analysis (OPLS-DA). The computed model predicted the class membership of filled-viable, empty and petrified seeds in the test set with 98%, 82% and 87% accuracy, respectively. When two-class OPLS-DA model was fitted to discriminate viable and non-viable (empty and petrified seeds combined), the predicted class membership of test set samples was 100% for both classes. The origins of spectral differences between non-viable (petrified and empty) and viable seeds were attributed to differences in seed moisture content and storage reserves. In conclusion, the result provides evidence that NIR spectroscopy is a powerful non-destructive method for sorting non-viable seeds of Larix sibirica; thus efforts should be made to develop on-line sorting system for large-scale seed handling.

  • Farhadi, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: mostafa.farhadi@slu.se
  • Tigabu, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: mulualem.tigabu@slu.se (email)
  • Odén, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: per.oden@slu.se
article id 1334, category Research article
Abolfazl Daneshvar, Mulualem Tigabu, Asaddollah Karimidoost, Per Christer Oden. (2015). Single seed Near Infrared Spectroscopy discriminates viable and non-viable seeds of Juniperus polycarpos. Silva Fennica vol. 49 no. 5 article id 1334. https://doi.org/10.14214/sf.1334
Keywords: NIRS; OPLS; seed sorting; Iran; juniper; near infrared spectroscopy
Highlights: Near Infrared (NIR) Spectroscopy discriminates viable and non-viable (empty, insect-attacked and shriveled) seeds of J. polycarpos with 98% and 100% accuracy, respectively; The origins of spectral differences between non-viable and viable seeds were attributed to differences in seed coat chemical composition and storage reserves; The results demonstrate that NIR spectroscopy has great potential as seed sorting technology to ensure precision sowing.
Abstract | Full text in HTML | Full text in PDF | Author Info

A large quantity of non-viable (empty, insect-attacked and shriveled) seeds of Juniperus polycarpos (K. Koch) is often encountered during seed collection, which should be removed from the seed lots to ensure precision sowing in the nursery or out in the field. The aims of this study were to evaluate different modelling approaches and to examine the sensitivity of the change in detection system (Silicon-detector in the shorter vis-a-vis InGsAs-detector in the longer NIR regions) for discriminating non-viable seeds from viable seeds by Near Infrared (NIR) spectroscopy. NIR reflectance spectra were collected from single seeds, and discriminant models were developed by Partial Least Squares – Discriminant Analysis (PLS-DA) and Orthogonal Projection to Latent Structures – Discriminant Analysis (OPLS-DA) using the entire or selected NIR regions. Both modelling approaches resulted in 98% and 100% classification accuracy for viable and non-viable seeds in the test set, respectively. However, OPLS-DA models were superb in terms of model parsimony and information quality. Modelling in the shorter and longer wavelength region also resulted in similar classification accuracy, suggesting that prediction of class membership is insensitive to change in the detection system. The origins of spectral differences between non-viable and viable seeds were attributed to differences in seed coat chemical composition, mainly terpenoids that were dominant in non-viable seeds and storage reserves in viable seeds. In conclusion, the results demonstrate that NIR spectroscopy has great potential as seed sorting technology to upgrade seed lot quality that ensures precision sowing.

  • Daneshvar, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53, Alnarp, Sweden; (permanent address) Department of Natural Resources, Gonbad Kavous University, Shahid Fallahi Street, P.O. Box 163, Gonbad, Iran E-mail: abolfazl.daneshvar@slu.se
  • Tigabu, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53, Alnarp, Sweden E-mail: mulualem.tigabu@slu.se (email)
  • Karimidoost, Agriculture and Natural Resources Research Center of Golestan Province, Beheshti Ave. P.O. Box 4915677555, Gorgan, Iran E-mail: karimidoost@yahoo.com
  • Oden, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53, Alnarp, Sweden E-mail: per.oden@slu.se
article id 1328, category Research article
Nelly N. Selochnik, Nataliya V. Pashenova, Evgeny Sidorov, Michael J. Wingfield, Riikka Linnakoski. (2015). Ophiostomatoid fungi and their roles in Quercus robur die-back in Tellermann forest, Russia. Silva Fennica vol. 49 no. 5 article id 1328. https://doi.org/10.14214/sf.1328
Keywords: Ophiostomatales; Ophiostoma; oak die-back; pathogenicity
Highlights: Dominant ophiostomatoid fungi associated with Q. robur in the post-outbreak region of oak die-back were investigated; Ophiostoma quercus was the most commonly encountered fungus; This is the first report of O. grandicarpum from Russia; The results of preliminary pathogenicity experiments demonstrate that fungi investigated in this study are unlikely to play causal role in oak die-back
Abstract | Full text in HTML | Full text in PDF | Author Info

Several eastern European countries have reported outbreaks of oak die-back during the 1980’s. Species of Ophiostoma Syd. were isolated from diseased trees and have been suggested to be the possible causal agents of the die-back, but this view have generally not been accepted. In order to monitor the post-outbreak region of oak die-back and to consider the possible role of Ophiostoma spp. in the syndrome, research has been conducted in the Tellerman forest, Voronezh region, Russia between 2005 and 2011. Our study resulted in the isolation of ophiostomatoid fungi from Quercus robur L. trees displaying external signs of desiccation. Fungi were identified based on morphological characteristics and DNA sequence comparisons. Three species of Ophiostoma were identified including O. grandicarpum (Kowalski & Butin) Rulamort, a species closely related to O. abietinum Marm. & Butin, O. fusiforme Aghayeva & M.J. Wingf. and O. lunatum Aghayeva & M.J. Wingf. representing a poorly understood species complex, and most commonly O. quercus (Georgev.) Nannf. Pathogenicity of these fungi was tested using artificial inoculations on Q. robur trees. The fungi were shown to be non-pathogenic and unlikely to play any role in oak die-back. These fungi are most likely only components in a complex of abiotic, biotic and anthropogenic factors that have contributed to a die-back of Quercus spp. in Russia.

  • Selochnik, Forest Science Institute of RAS, Uspenskoe 143030, Moscow Region, Russia E-mail: lenelse@yandex.ru
  • Pashenova, V.N. Sukachev Institute of Forest SB RAS, Krasnoyarsk 660036, Russia E-mail: pasnat@ksc.krasn.ru
  • Sidorov, Department of Forest Protection and Game Management, St. Petersburg State Forest Technical University, St. Petersburg 194021, Russia E-mail: sidorov_evgeny@mail.ru
  • Wingfield, Forestry and Agricultural Biotechnology Institute (FABI), University of Pretoria, 0002 Pretoria, South Africa E-mail: mike.wingfield@up.ac.za
  • Linnakoski, Forestry and Agricultural Biotechnology Institute (FABI), University of Pretoria, 0002 Pretoria, South Africa; Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID http://orcid.org/0000-0002-3294-8088 E-mail: riikka.linnakoski@helsinki.fi (email)
article id 1312, category Research article
Simon Karl Nils Berg, Tomas Nordfjell, Dan Bergström. (2015). Effect of stump size and timing of stump harvesting on ground disturbance and root breakage diameter. Silva Fennica vol. 49 no. 5 article id 1312. https://doi.org/10.14214/sf.1312
Keywords: Picea abies; scarification; Sweden; stump harrow; Ecorex30; site impact; glacial till soil
Highlights: The ground disturbance and root breakage diameter during conventional stump harvesting on mineral soil were quantified; A function for estimating the disturbed area based on stump size was constructed; Many fine roots were found to be harvested; The total ground disturbance at the site after stump harvesting was similar to that caused by soil scarification.
Abstract | Full text in HTML | Full text in PDF | Author Info

Stump wood is a possible alternative to fossil fuel. Its harvesting, however, disturbs the ground and this has not yet been quantified at stump level. Such disturbance is likely to be dependent on stump size, type of soil and timing of stump harvesting. Therefore, we measured ground disturbance and root breakage diameter at two Norway spruce sites with sandy glacial till soil. The sites were harvested with a fork type head, 6 and 18 months after clear cutting. Measurements were made within 2 weeks of harvest. No difference was found between the two sites. The mean area of disturbed ground was 6.06 (std 3.14) m2 per stump and increased exponentially with stump size. A regression function modelling the relationship was constructed. Unexpectedly, many fine roots where extracted in the harvest. The arithmetic and basal area weighted mean root breakage diameter was 4.6 (std 2.2) and 29.5 (std 17.9) mm, respectively. There seems to be a limited increase in root breakage diameter with increased stump size. The small root breakage diameter is associated with reduced fuel quality and greater nutrient removal. It appears that much of the ground disturbance is associated with the creation of ruts rather than stump harvest per se. Stump harvesting disturbs a larger percentage of the area of a harvested site than mounding. Postponing stump harvest by one year did not decrease the ground disturbance or increase the root breakage diameter. To achieve less disturbance and larger root breakage diameter, probably new stump harvesting technology is required.

  • Berg, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: phd.simon.berg@gmail.com (email)
  • Nordfjell, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: tomas.nordfjell@slu.se
  • Bergström, Swedish University of Agricultural Sciences (SLU), Department of Forest Biomaterials and Technology (SBT), Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: dan.bergstrom@slu.se
article id 1310, category Research article
Deliang Lu, Jiaojun Zhu, Yirong Sun, Lile Hu, Guangqi Zhang. (2015). Gap closure process by lateral extension growth of canopy trees and its effect on woody species regeneration in a temperate secondary forest, Northeast China. Silva Fennica vol. 49 no. 5 article id 1310. https://doi.org/10.14214/sf.1310
Keywords: canopy closure; recruitment; canopy opening; opening size; crown expansion; duration; hemispherical photograph
Highlights: Gap closure process by lateral extension growth can be described by quadratic functions; Large gaps (514–621 m2) had higher closure rates but lower closure percentages compared with middle (174–321 m2) and small gaps (68–125 m2); Gaps promoted woody species regeneration in early stage; Large and middle gaps would provide opportunities for filling regeneration, but regeneration in small gaps may eventually fail.
Abstract | Full text in HTML | Full text in PDF | Author Info

Gap formation and its effects on regeneration have been reported as being important in forest development, but seldom studies concentrated on the gap closure process by lateral extension growth of canopy trees surrounding gaps. We monitored the closure process of 12 artificial gaps for 7 years with three size classes: small (from 68 m2 to 125 m2), middle (from 174 m2 to 321 m2), and large (from 514 m2 to 621 m2); and investigated the regeneration twice in a temperate secondary forest, Northeast China. The closure process can be described through quadratic functions, which showed the closure rates slowed down with gap ages. Large gaps had a higher closure rate (39 m2 a–1) than middle gaps (25 m2 a–1) and small gaps (11 m2 a–1). According to the quadratic equations, the lateral growth could last 11, 13 and 16 years for small, middle and large gaps with a remaining size of 12, 69 and 223 m2, respectively. As expected, regeneration exhibited the highest seedling density and volume in large gaps. There was no significant difference in regeneration density between middle gaps, small gaps and forest understory in the final investigation; but the volume of regenerated woody species increased significantly from small gaps to large gaps compared with forest understory. These results may provide references on the choice of appropriate gap sizes to promote the regeneration in temperate secondary forests.

  • Lu, State Key Laboratory of Forest and Soil Ecology, Qingyuan Forest CERN, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110016, China;  University of Chinese Academy of Sciences, Beijing 100049, China E-mail: delianglu14@hotmail.com
  • Zhu, State Key Laboratory of Forest and Soil Ecology, Qingyuan Forest CERN, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110016, China E-mail: jiaojunzhu@iae.ac.cn (email)
  • Sun, State Key Laboratory of Forest and Soil Ecology, Qingyuan Forest CERN, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110016, China E-mail: yirongsun@iae.ac.cn
  • Hu,  Chinese Research Academy of Environmental Sciences, Beijing 100012, China E-mail: lilehu@gmail.com
  • Zhang, State Key Laboratory of Forest and Soil Ecology, Qingyuan Forest CERN, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110016, China;  University of Chinese Academy of Sciences, Beijing 100049, China E-mail: zgq04713@163.com
article id 1408, category Research article
Anna Katharina Franke, Pasi Aatsinki, Ville Hallikainen, Esa Huhta, Mikko Hyppönen, Vesa Juntunen, Kari Mikkola, Seppo Neuvonen, Pasi Rautio. (2015). Quantifying changes of the coniferous forest line in Finnish Lapland during 1983–2009. Silva Fennica vol. 49 no. 4 article id 1408. https://doi.org/10.14214/sf.1408
Keywords: Pinus sylvestris; Picea abies; tree line; forest regeneration; environmental change; stand volume
Highlights: Volume of the growing stock of spruce and pine has increased in forests and in timber lines during the past 26 years; Spruce stem numbers increased on average, while pine stem numbers remained stable and location-specific variation was observed; Presuming that the ongoing trend of increasing temperature will remain, the enhanced forest regeneration and growth may result in extension of forests in the future.
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The boreal timber- and tree-line forests grow in harsh environmental conditions in their outermost distribution limit. Here even small environmental changes may cause dramatic changes in the distribution of tree species. We examined changes of the forest lines of Norway spruce (Picea abies (L.) H. Karst.) and Scots pine (Pinus sylvestris L.) in Finnish Lapland five times during 1983–2009. We monitored the number of stems and the volume of the growing stock in thirteen different locations in forest-line areas. The linear temporal trends and the variations of these response variables were used as indicators of a possible change during the study period. Spruce showed a significant increase both in the volume of the growing stock (up to 40% increase) and in the total stem number (up to 100% increase). A significant increase in the volume of the growing stock was observed in the pine data as well (up to 70% increase), whereas the stem number stagnated or even decreased. The results suggest that spruce needs favourable conditions to have an abundant regeneration, but after the establishment the seedlings seem to be more resistant against biotic and abiotic disturbances than pine seedlings. The increasing stand volume might result in a climate-related northward and upward extension of forests in the future. However, our results show that responses in the boreal forest line are species and location specific and a more favourable climate does not necessarily lead to an advance of the coniferous forest line.

  • Franke, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: anna.franke@fau.de (email)
  • Aatsinki, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: pasi.aatsinki@luke.fi
  • Hallikainen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: ville.hallikainen@luke.fi
  • Huhta, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: esa.huhta@luke.fi
  • Hyppönen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: mikko.hypponen@luke.fi
  • Juntunen, The Sámi Education Institute, Menesjärventie 4, P.O. Box 50, FI-99871 Inari, Finland E-mail: vesa.juntunen@sogsakk.fi
  • Mikkola, Natural Resources Institute Finland (Luke), Economics and society, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: kari.mikkola@luke.fi
  • Neuvonen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 68, FI-80101, Joensuu, Finland E-mail: seppo.neuvonen@luke.fi
  • Rautio, Natural Resources Institute Finland (Luke), Bio-based business and industry, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: pasi.rautio@luke.fi
article id 1391, category Research article
Roberts Matisons, Jānis Jansons, Juris Katrevičs, Āris Jansons. (2015). Relation of tree-ring width and earlywood vessel size of alien Quercus rubra L. with climatic factors in Latvia. Silva Fennica vol. 49 no. 4 article id 1391. https://doi.org/10.14214/sf.1391
Keywords: dendroclimatology; climate-growth relationships; introduced species; earlywood vessel lumen area; wood anatomy; shifting sensitivity
Highlights: Climate-growth relationships of red oak from three sites in Latvia were studied; Tree-ring width was mainly affected by temperature and precipitation in late summer; Vessel size was correlated with temperature parameters in autumn–spring; Sets of climatic factors significant for growth of red oak differed between sites; Changes in climate-growth relationships occurred during 20th century.
Abstract | Full text in HTML | Full text in PDF | Author Info

The effect of climatic factors on wood anatomy of the alien red oak (Quercus rubra L.) growing in three experimental plantations in Latvia was assessed by classical dendrochronological techniques. Two tree-ring proxies – tree-ring width (TRW) and mean area of earlywood vessel lumen (VLA) – were studied on 33 trees. Annual variation of TRW amongst trees was similar (mean r = 0.46), but there was more individuality in VLA (mean r = 0.26); nevertheless, chronologies of both proxies had rather synchronous variation amongst the sites. Annual variation of TRW was affected by factors related to water deficit in late summer, as suggested by the negative effect of temperature and positive effect of precipitation that have intensified during the 20th century, likely due to warming. Although weather conditions during the dormant period did not directly affect TRW, temperature during the autumn-spring period has been the main climatic determinant of VLA likely via influence on overwintering and hence vigour of tree. This suggests that conductive properties of wood and hence the susceptibility to water deficit have been affected by weather conditions before the formation of tree rings. During the 20th century, sensitivity of VLA has shifted from temperature in winter to temperature in autumn likely due to climate change. Still, the positive effect of these factors suggests that warming of climate would increase VLA and hence the risk of embolism and xylem disfunction. Therefore, the importance of availability of water for growth of red oak in Latvia is increasing.

  • Matisons, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: robism@inbox.lv (email)
  • Jansons, Latvian Forest Competence Centre, Dzērbenes str. 27, Riga, Latvia, LV 1006 E-mail: janis.jansons@silava.lv
  • Katrevičs, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: juris.katrevics@silava.lv
  • Jansons, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: aris.jansons@silava.lv
article id 1354, category Research article
Johannes Edvardsson, Anton Hansson. (2015). Multiannual hydrological responses in Scots pine radial growth within raised bogs in southern Sweden. Silva Fennica vol. 49 no. 4 article id 1354. https://doi.org/10.14214/sf.1354
Keywords: Pinus sylvestris; peatland; dendrochronology; precipitation; hydrology; river discharge
Highlights: Annual growth of Scots pine (Pinus sylvestris L.) at Boreal raised bogs was found to reflect a synthesis of climate controlled moisture variability over the preceding one to four year period; Excessive soil moisture is a growth limiting factor for trees at raised bogs; River discharge data reflect hydrological conditions in peat bogs better than precipitation data.
Abstract | Full text in HTML | Full text in PDF | Author Info

To explore interactions between climate and peatland hydrology, Scots pine (Pinus sylvestris L.) growing at four raised bogs in southern Sweden were subject to a dendroclimatological study. Radial tree growth reflecting climate and water table fluctuations over multiannual periods was detected as significant negative correlations between tree-ring width (TRW) chronologies and the preceding one to four years total precipitation or river discharge. Systematically stronger negative correlations were obtained when river discharge instead of precipitation was compared to radial tree growth. This indicates that river discharge reflect moisture variability of peat bogs better than what precipitation data does. Meanwhile, monthly precipitation and radial tree growth did not show any clear correlation, whereas spring and early summer temperatures had a positive influence on the tree growth. Our study shows that growth variability of bog pines in the Boreal zone reflect hydrological responses related to a synthesis of climate controlled moisture variability over several year periods.

  • Edvardsson, Dendrolab.ch, Institute of Geological Sciences, University of Bern, Baltzerstrasse 1+3, CH-3012 Bern, Switzerland & Quaternary Sciences, Department of Geology, Lund University, Sölvegatan 12, SE-223 62 Lund, Sweden ORCID http://orcid.org/0000-0001-5163-1599 E-mail: johannes.edvardsson@dendrolab.ch (email)
  • Hansson, Quaternary Sciences, Department of Geology, Lund University, Sölvegatan 12, SE-223 62 Lund, Sweden E-mail: anton.hansson@geol.lu.se
article id 1326, category Research article
Joanna Bachmatiuk, Jordi Garcia-Gonzalo, Jose Guilherme Borges. (2015). Analysis of the performance of different implementations of a heuristic method to optimize forest harvest scheduling. Silva Fennica vol. 49 no. 4 article id 1326. https://doi.org/10.14214/sf.1326
Keywords: harvest scheduling; simulated annealing; heuristic; cooling schedule; initial temperature
Highlights: The number of treatment schedules available for each stand has an impact on the optimal configuration of opt-moves (i.e. number stands where the treatment schedule is changed in an iteration); Considering a large number of treatment schedules per stand, the one-opt move implementation is preferred, yet when considering a low number of decision choices the two-opt moves option performs better.
Abstract | Full text in HTML | Full text in PDF | Author Info

Finding an optimal solution of forest management scheduling problems with even flow constraints while addressing spatial concerns is not an easy task. Solving these combinatorial problems exactly with mixed-integer programming (MIP) methods may be infeasible or else involve excessive computational costs. This has prompted the use of heuristics. In this paper we analyze the performance of different implementations of the Simulated Annealing (SA) heuristic algorithm for solving three typical harvest scheduling problems. Typically SA consists of searching a better solution by changing one decision choice in each iteration. In forest planning this means that one treatment schedule in a single stand is changed in each iteration (i.e. one-opt move). We present a comparison of the performance of the typical implementation of SA with the new implementation where up to three decision choices are changed simultaneously in each iteration (i.e. treatment schedules are changed in more than one stand). This may allow avoiding local optimal. In addition, the impact of SA - parameters (i.e. cooling schedule and initial temperature) are tested. We compare our heuristic results with a MIP formulation. The study case is tested in a real forest with 1000 stands and a total of 213116 decision choices. The study shows that when the combinatorial problem is very large, changing simultaneously the treatment schedule in more than one stand does not improve the performance of SA. Contrarily, if we reduce the size of the problem (i.e. reduce considerably the number of alternatives per stand) the two-opt moves approach performs better.

  • Bachmatiuk, Centro de Estudos Florestais, Instituto Superior de Agronomia, Universidade de Lisboa, Lisboa, Portugal E-mail: jbachmatiuk@isa.ulisboa.pt (email)
  • Garcia-Gonzalo, Centro de Estudos Florestais, Instituto Superior de Agronomia, Universidade de Lisboa, Lisboa, Portugal E-mail: jordigarcia@isa.ulisboa.pt
  • Borges, Centro de Estudos Florestais, Instituto Superior de Agronomia, Universidade de Lisboa, Lisboa, Portugal E-mail: joseborges@isa.ulisboa.pt
article id 1283, category Research article
Ivana Bjedov, Dragica Obratov–Petković, Danijela Mišić, Branislav Šiler, Jelena M Aleksic. (2015). Genetic patterns in range-edge populations of Vaccinium species from the central Balkans: implications on conservation prospects and sustainable usage. Silva Fennica vol. 49 no. 4 article id 1283. https://doi.org/10.14214/sf.1283
Keywords: marginal populations; genetic structure; Vaccinium sp.; the Balkans; RAPD markers; genetic profiles; interspecific hybridization
Highlights: We studied fragmentary distributed range-edge populations of Vaccinium myrtillus, Vaccinium uliginosum and Vaccinium vitis-idaea from the Balkans using RAPDs; Low genetic diversities and high genetic differentiation were found in all species; The prevalence of clonal individuals was not observed; Past interspecific hybridization among V. vitis-idaea and the other two species was detected; Guidelines for conservation and sustainable usage were provided.
Abstract | Full text in HTML | Full text in PDF | Author Info

Vaccinium myrtillus L., Vaccinium uliginosum L. and Vaccinium vitis-idaea L. are perennial, cold-adapted clonal shrubs distributed throughout Europe, northern Asia and North America. Due to their usage in food (berries) and pharmaceutical industry (berries and leaves), their natural populations are exposed to anthropogenic and other impacts that affect their genetic make-up. We analyzed 14 fragmentary distributed and small-sized peripheral populations of these species from the Balkans, which represents the southeastern-European marginal area of their wide European distributions, using RAPD molecular markers. The contemporary genetic patterns in all three species within the Balkans were generally similar, and in comparison to previous reports on populations of these species found in northward Europe, where they have a more continuous distribution, the levels of genetic diversity were more or less halved, genetic differentiation was several times higher, gene flow exceptionally low, and the expected prevalence of clonal individuals was lacking. The population dynamics of all three species within the Balkans was complex and distinct, and was characterized by a past admixture of individuals from discrete populations of the same species and interspecific hybridisation not only between V. myrtillus and V. vitis-idaea but also between V. uliginosum and V. vitis-idaea, the latter not being reported to date. Conservation measures suitable for preservation of presumably genetically distinct portions of the Balkans’ gene pools of studied species have been suggested, while the utility of interspecific hybrids in breeding programs and/ or in food/pharmaceutical industry is yet to be assessed. 

  • Bjedov, University of Belgrade, Faculty of Forestry, Kneza Višeslava 1, 11000 Belgrade, Serbia E-mail: ivana.bjedov@sfb.bg.ac.rs
  • Obratov–Petković, University of Belgrade, Faculty of Forestry, Kneza Višeslava 1, 11000 Belgrade, Serbia E-mail: dragica.obratov-petkovic@sfb.bg.ac.rs
  • Mišić, University of Belgrade, Institute for Biological Research “Siniša Stanković”, Boulevard Despota Stefana 142, 11000 Belgrade, Serbia E-mail: dmisic@ibiss.bg.ac.rs
  • Šiler, University of Belgrade, Institute for Biological Research “Siniša Stanković”, Boulevard Despota Stefana 142, 11000 Belgrade, Serbia E-mail: branislav.siler@ibiss.bg.ac.rs
  • Aleksic, University of Belgrade, Institute of Molecular Genetics and Genetic Engineering, Vojvode Stepe 444a, P.O. Box 23, 11010 Belgrade, Serbia E-mail: aleksic_jelena@yahoo.com.au (email)
article id 1265, category Research article
Eva Ring, Lars Högbom, Hans-Örjan Nohrstedt, Staffan Jacobson. (2015). Soil and soil-water chemistry below different amounts of logging residues at two harvested forest sites in Sweden. Silva Fennica vol. 49 no. 4 article id 1265. https://doi.org/10.14214/sf.1265
Keywords: clearcutting; final felling; bio fuel; conifer; fuel-adapted felling; nutrient; soil solution
Highlights: Soil-water chemistry, ground vegetation cover and water flux were affected by the amounts of logging residues stored on the ground after harvest; A strong response on soil-water chemistry was recorded at only one of the two sites; At the site showing a weak response, less residue remained after seven years in the treatments giving the most pronounced effects.
Abstract | Full text in HTML | Full text in PDF | Author Info
Logging residues (LR), i.e. tops, branches, and needles, are increasingly being harvested for energy production in Fennoscandia. These residues are temporarily piled on site awaiting transport. This study was undertaken to investigate effects on the soil and soil-water chemistry below different amounts of LR at two recently harvested coniferous sites in Sweden. Seven treatments were included and the studied amounts of LR ranged from no LR left on the ground to four times the estimated LR amount of the harvested stands. Two treatments included eight times the estimated LR amount of the harvested stands but here the LR were removed after 7 or 20 weeks. Soil-water samples were collected during the first six or seven growing seasons. Effects of treatment were detected in the soil water for 11 chemical variables at the northern site, and for the NO3- and Cl- concentrations at the southern site. The strongest response was generally found in the treatment with four times the estimated LR amount, for which the highest concentrations were recorded in most cases. In the first three seasons, the water flux through the LR decreased with an increasing amount of residue. Effects on the exchangeable store of Ca2+ in the mor layer and the upper 20 cm of the mineral soil was detected at both sites. At the northern site, the weight of the remaining LR, ground vegetation and all other material above the mor layer in the treatments with two and four times the estimated LR amount was roughly twice the corresponding weights at the southern site seven years after treatment. Although strong effects on the soil-solution chemistry were detected at one of the study sites, in the treatments corresponding to two and four times the estimated logging residue amount, the effect on the leaching from an entire regeneration area is likely to be relatively small given the percentage of the area hosting these logging residue amounts (ca. 20% after stem-only harvesting and 9% after fuel-adapted felling).
  • Ring, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: eva.ring@skogforsk.se (email)
  • Högbom, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: lars.hogbom@skogforsk.se
  • Nohrstedt, Swedish University of Agricultural Sciences, Department of Soil and Environment, P.O. Box 7014, SE-750 07 Uppsala, Sweden E-mail: hans-orjan.nohrstedt@slu.se
  • Jacobson, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: staffan.jacobson@skogforsk.se
article id 1243, category Research article
Curt Almqvist, Gunnar Jansson. (2015). Effects of pruning and stand density on cone and pollen production in an experimental Pinus sylvestris seed orchard. Silva Fennica vol. 49 no. 4 article id 1243. https://doi.org/10.14214/sf.1243
Keywords: spacing; seed orchard management; crown management; seed production
Highlights: Pollen production of Pinus sylvestris began at the same age for all studied stand density and pruning height combinations but increased more rapidly at higher densities; Treatments with dense spacing increased seed production earlier; Many combinations of stand density and target height gave comparable levels of seed production, yielding a wide range of viable management options.
Abstract | Full text in HTML | Full text in PDF | Author Info

Seed orchards are the link between tree breeding and reforestation. This paper presents data on cone, seed and pollen production and seed quality gathered over 21 years in a Pinus sylvestris (L.) experimental seed orchard containing plots with 14 different combinations of stand density and targeted pruning height. The treatments’ stand densities ranged from 267 to 4000 stems ha-1, and the target graft heights ranged from 2 to 6 meters. Pollen production began at the same orchard age for all studied combinations of stand density and target height but the level of pollen production per hectare increased more rapidly in treatments with higher stand densities. In treatments with dense spacing, cone and seed production initially increased more rapidly than in treatments with wider spacing, thereby providing an earlier return on investment and a shorter seed production lag time. However, the levels of cone and seed production in such treatments over the entire study period were not appreciably different to those achieved in treatments with wider spacing and higher target height. The treatments did not differ substantially with respect to seed quality. These results show that comparable levels of seed production can be obtained with different combinations of stand density and target height, giving seed orchard owners and managers a wide range of viable management options.

  • Almqvist, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: curt.almqvist@skogforsk.se (email)
  • Jansson, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: gunnar.jansson@skogforsk.se
article id 1236, category Research article
Monika Litkowiec, Beata P. Plitta-Michalak, Andrzej Lewandowski, Grzegorz Iszkuło. (2015). Homogenous genetic structure in populations of Taxus baccata with varied proportions of male and female individuals. Silva Fennica vol. 49 no. 4 article id 1236. https://doi.org/10.14214/sf.1236
Keywords: genetic diversity; dioecy; English yew; Poland; sex ratio; nSSR markers
Highlights: Polish populations of Taxus baccata showed a high level of genetic diversity within populations and moderate genetic differentiation between them after nSSR marker testing; No significant differences in the genetic variation between T. baccata male and female individuals were observed, and microsatellite loci neutrality was verified; Determining the sex ratio in T. baccata populations is not essential to develop a clear understanding of genetic differentiation and diversity within and between populations of this species.
Abstract | Full text in HTML | Full text in PDF | Author Info

English yew (Taxus baccata L.) is a strictly outcrossing and dioecious species whose populations are small and isolated. It is known that sex ratios may vary in natural populations due to local environmental conditions or stochastic events. However, unbalanced sex ratios may have negative impacts on genetic diversity through enhanced genetic drift and inbreeding. The present study represents one of the first attempts to compare the genetic variation at microsatellite loci within and between populations with different gender proportions. Our results indicated that there were no significant correlations between sex ratio and the extent of genetic variation in different populations. All populations exhibited high levels of genetic diversity. Additionally, the genetic structure was characterized separately in male and female individuals. Statistical analyses of the set estimators describing the genetic structure of male and female individuals of T. baccata revealed no significant differences between the two groups. Molecular analysis verified that microsatellite nuclear loci neutrality developed for T. baccata, as there were no significant differences in the genetic variation between males and females and no evidence for any outlier loci using coalescent and hierarchical Bayesian simulations. The results demonstrate that ignoring biased sex ratios in T. baccata populations had no effect on the assessment of genetic differentiation and genetic diversity within and between populations of this species. These results are discussed with regards to the practical application of molecular markers in conservation programs.

  • Litkowiec, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland E-mail: mlit@man.poznan.pl (email)
  • Plitta-Michalak, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland E-mail: beata-plitta@wp.pl
  • Lewandowski, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland E-mail: alew@man.poznan.pl
  • Iszkuło, Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland & University of Zielona Góra, Faculty of Biological Sciences, Prof. Z. Szafrana 1, 65-516 Zielona Góra, Poland E-mail: iszkulo@man.poznan.pl
article id 1155, category Research article
Feng Chen, Yujiang Yuan, Wenshou Wei, Tongwen Zhang, Huaming Shang, Shulong Yu. (2015). Divergent response of tree-ring width and maximum latewood density of Abies faxoniana to warming trends at the timberline of the western Qinling Mountains and northeastern Tibetan Plateau, China. Silva Fennica vol. 49 no. 4 article id 1155. https://doi.org/10.14214/sf.1155
Keywords: dendroclimatology; tree rings; faxon fir; climate-growth relationships; warming trend
Highlights: We describe new tree-ring width and maximum latewood density chronologies of Faxon fir; Temperature is the most important limiting factor for the radial growth of Faxon fir; The tree-ring width series captured the warming trends and allowed detecting the recent warming in a long-term context, while the maximum latewood density series showed no upward trend.
Abstract | Full text in HTML | Full text in PDF | Author Info

Tree-ring width (TRW) and maximum latewood density (MXD) data of Faxon fir (Abies faxoniana Rehd. et Wils.) were analyzed for five timberline sites in the western Qinling Mountains and northeastern Tibetan Plateau, to investigate their relationships to climate change, especially twentieth century warming. The cross-correlations among TRW chronologies at the low-frequency band were higher, while the higher correlations among MXD chronologies were found at the high-frequency band. Response analysis showed that the tree-ring formation of fir trees was significantly and positively affected by temperature variations, while it was also negatively affected by precipitation. The TRW series captured the warming trends and allowed detecting the recent warming in a long-term context, while the MXD series showed no upward trend. We also found the temperature sensitivity of the TRW series is unstable over space and time. The divergent response between TRW and MXD might be caused by the seasonal variations of warming trends.

  • Chen, Key Laboratory of Tree-ring Physical and Chemical Research of China Meteorological Administration/Xinjiang Laboratory of Ecology, Institute of Desert Meteorology, China Meteorological Administration, 46 Jianguo Road, Urumqi 830002, China E-mail: feng653@163.com (email)
  • Yuan, Key Laboratory of Tree-ring Physical and Chemical Research of China Meteorological Administration/Xinjiang Laboratory of Ecology, Institute of Desert Meteorology, China Meteorological Administration, 46 Jianguo Road, Urumqi 830002, China E-mail: yuanyuj5502@sina.com
  • Wei, Key Laboratory of Tree-ring Physical and Chemical Research of China Meteorological Administration/Xinjiang Laboratory of Ecology, Institute of Desert Meteorology, China Meteorological Administration, 46 Jianguo Road, Urumqi 830002, China E-mail: cycfqq@sohu.com
  • Zhang, Key Laboratory of Tree-ring Physical and Chemical Research of China Meteorological Administration/Xinjiang Laboratory of Ecology, Institute of Desert Meteorology, China Meteorological Administration, 46 Jianguo Road, Urumqi 830002, China E-mail: tongwenzhang19820705@hotmail.com
  • Shang, Key Laboratory of Tree-ring Physical and Chemical Research of China Meteorological Administration/Xinjiang Laboratory of Ecology, Institute of Desert Meteorology, China Meteorological Administration, 46 Jianguo Road, Urumqi 830002, China E-mail: shang8632@163.com
  • Yu, Key Laboratory of Tree-ring Physical and Chemical Research of China Meteorological Administration/Xinjiang Laboratory of Ecology, Institute of Desert Meteorology, China Meteorological Administration, 46 Jianguo Road, Urumqi 830002, China E-mail: yushl@idm.cn
article id 1337, category Research article
Leszek Bujoczek, Małgorzata Bujoczek, Jan Banaś, Stanisław Zięba. (2015). Spruce regeneration on woody microsites in a subalpine forest in the western Carpathians. Silva Fennica vol. 49 no. 3 article id 1337. https://doi.org/10.14214/sf.1337
Keywords: Picea abies; coarse woody debris; stumps; decomposition; regression model; fallen deadwood
Highlights: The occurrence probability of Picea abies seedlings on fallen deadwood was found to increase with diameter and decay stage of deadwood and with the volume of living trees, and to decrease with the density of living trees, sapling density, and land slope. It was also higher on stumps with greater diameter and in plots with higher sapling density, but decreased with increasing stump height.
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The density of Picea abies [L.] Karst. regeneration on different microsites, the quantity and quality of woody microsites, and seedling occurrence probability on stumps and fallen deadwood were studied in a subalpine forest that has been under protection for approximately 30–40 years (Gorce Mountains in the western Carpathians). Thirty percent of seedlings and 29% of saplings grew on stumps and fallen deadwood, while the remaining regeneration occurred on soil surface and mounds created by uprooted trees. The occurrence probability of Picea seedlings on fallen deadwood increased with deadwood diameter and decay stage and with the volume of living trees, and decreased with increased density of living trees, sapling density, and land slope. Furthermore, seedlings were more likely to grow on stumps with a greater diameter and in plots with higher sapling density, but less likely to grow on higher stumps. Stumps and fallen deadwood covered about 4% of the forest floor, but the material that is most important for promoting regeneration (strongly decomposed logs and those of a diameter exceeding 30 cm) took up only about 22 m2 ha-1. We have concluded that in a subalpine forest that has been protected for 30–40 years regeneration processes take place mostly on soil surface and stumps. The role of fallen deadwood increases over time as a greater number of suitable logs (in terms of size and decay stage) become available.

  • Bujoczek, University of Agriculture in Krakow, E-mail: lbujoczek@gmail.com (email)
  • Bujoczek, University of Agriculture in Krakow, E-mail: bujoczek.m@gmail.com
  • Banaś, University of Agriculture in Krakow, E-mail: rlbanas@cyf-kr.edu.pl
  • Zięba, University of Agriculture in Krakow, E-mail: rlzieba@cyf-kr.edu.pl
article id 1302, category Research article
Nils Fahlvik, Per Magnus Ekö, Nils Petersson. (2015). Effects of precommercial thinning strategies on stand structure and growth in a mixed even-aged stand of Scots pine, Norway spruce and birch in southern Sweden. Silva Fennica vol. 49 no. 3 article id 1302. https://doi.org/10.14214/sf.1302
Keywords: yield; cleaning; heterogeneous forest; dimension distribution; growth simulator
Highlights: Precommercial thinning (PCT) was a useful tool to influence the stand structure in accordance to silvicultural goals; PCT had a great impact on tree species composition; The seemingly great potential to influence the structure of a heterogeneous, mixed stand was restrained by natural settings and unconditional considerations at PCT (e.g. tree vitality, stem quality, regular spacing).
Abstract | Full text in HTML | Full text in PDF | Author Info
Four different management strategies were applied to a young mixed stand of Scots pine (Pinus sylvestris L.), Norway spruce (Pices abies (L.) Karst.) and birch (Betula pendula Roth and Betula pubescens Ehrh.) in southern Sweden. All strategies included an initial precommercial thinning to ca. 2400 stems ha–1. The different aims were: (i) a conifer-dominated stand with focus on high productivity; (ii) a conifer-dominated stand with high quality timber; (iii) keeping a stem-wise species mixture; (iv) a mosaic-wise species mixture. Stem selection according to the different strategies were simulated with a starting point from plots with a 5 m radius. All strategies were applied to all of the plots. A growth simulator was used to simulate the stand development up to final felling. This study illustrates the possibilities for influencing the structure of a mixed stand through precommercial thinning. The study also illustrates the long-term effects on stand structure and volume yield by consequently applying a management strategy from precommercial thinning until final felling. Precommercial thinning was found to be a useful tool to influence the stand structure in accordance to the aims set. However, the opportunities for influencing the stand by precommercial thinning were restricted by natural settings and unconditional considerations (e.g. tree vitality, stem quality, regular spacing), beyond what could be judged from stand average data. The stem volume production during a rotation was 6% lower for (iii) and (iv) compared to (i) and (ii), mainly due to a greater proportion of birch in the former strategies.
  • Fahlvik, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: nils.fahlvik@slu.se (email)
  • Ekö, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: per.magnus.eko@slu.se
  • Petersson, StoraEnso Skog AB, Åsgatan 22, SE-791 80 Falun, Sweden E-mail: nils.petersson@storaenso.com
article id 1301, category Research article
Mikko Moilanen, Jyrki Hytönen, Hannu Hökkä, Anssi Ahtikoski. (2015). Fertilization increased growth of Scots pine and financial performance of forest management in a drained peatland in Finland. Silva Fennica vol. 49 no. 3 article id 1301. https://doi.org/10.14214/sf.1301
Keywords: Pinus sylvestris; stand growth; peatlands; needle nutrients; P deficiency; K deficiency
Highlights: All fertilizers containing phosphorus and potassium improved the P and K status and the stem growth of Scots pine still 26 years from application; Wood ash, containing more nutrients than other fertilizers, gave the strongest stand growth response and the highest net present value; Ash fertilizer treatment outperformed other fertilizer treatments and control in net present value, regardless of the applied discount rate, 3%, 4% or 5%.
Abstract | Full text in HTML | Full text in PDF | Author Info

The long-term effects of fertilization on the needle nutrient concentrations, growth and financial performance of a Scots pine (Pinus sylvestris L.) stand was examined in a thick-peated drained peatland forest located in Central Finland. At the trial establishment in 1985, the trees were suffering from P and K deficiencies, but their N status was good. The fertilizer treatments were Control, PK (rock phosphate + potassium chloride), ApaBio (apatite phosphorus + biotite) and wood ash, applied both with and without N and replicated six times. All treatments containing phosphorus and potassium increased foliar P and K concentrations above the deficiency limits up to the end of the study period of 26 years. The effect of the fertilization on stand volume growth of Scots pine was strong and continued still at the end of the study period. The trees on ApaBio and PK plots grew nearly two-fold and those on Ash plots over two-fold compared with the control plots. In a thinning made at the end of the study period the total logging removal on fertilized plots was 1.5–2.2 times greater and included more saw logs than on the control plots. Ash fertilizer treatment outperformed other fertilizer treatments as well as the control. With a 5% discounted equivalent annual income (EAI) of Ash fertilizer treatment was statistically significantly (p=0.009) almost three times higher than that of control. As a conclusion, fertilization (either using PK fertilizers or Ash) in N-rich drained peatlands is a financially feasible method of management.

  • Moilanen, Natural Resources Institute Finland, Natural resources and bioproduction, Paavo Havaksen tie 3, FI-90014 Oulu, Finland E-mail: mikko.moilanen@luke.fi (email)
  • Hytönen, Natural Resources Institute Finland, Natural resources and bioproduction, Silmäjärventie 2, FI-69100 Kannus, Finland E-mail: jyrki.hytonen@luke.fi
  • Hökkä, Natural Resources Institute Finland, Natural resources and bioproduction, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: hannu.hokka@luke.fi
  • Ahtikoski, Natural Resources Institute Finland, Natural resources and bioproduction, Paavo Havaksen tie 3, FI-90014 Oulu, Finland E-mail: anssi.ahtikoski@luke.fi
article id 1295, category Research article
Jiaxi Wang, Guolei Li, Jeremiah R. Pinto, Jiajia Liu, Wenhui Shi, Yong Liu. (2015). Both nursery and field performance determine suitable nitrogen supply of nursery-grown, exponentially fertilized Chinese pine. Silva Fennica vol. 49 no. 3 article id 1295. https://doi.org/10.14214/sf.1295
Keywords: water stress; exponential fertilization; nursery response; field performance
Highlights: Increasing exponential fertilization rates in the nursery increased seedling biomass, N content, and N concentration for Chinese pine seedlings; Second year seedling survival illustrated a curvilinear response to seedling fertilization rates rather than a linear one; Considering both nursery responses to fertilization and field performance after two years yielded a recommended nitrogen supply rate of 80 mg N seedling–1.
Abstract | Full text in HTML | Full text in PDF | Author Info
Optimum fertilization levels are often determined solely from nursery growth responses. However, it is the performance of the seedling on the outplanting site that is the most important. For Pinus species seedlings, little information is known about the field performance of plants cultured with different nutrient rates, especially with exponential fertilization. In this study, Chinese pine (Pinus tabulaeformis Carr.) seedlings grown in 187 ml containers were fertilized exponentially in 6 treatments ranging from 10 to 120 mg N seedling–1 for 25 weeks before outplanting. Dry mass and N content were measured at planting. Survival and field growth were monitored for two growing seasons. In the nursery, our data showed no difference in dry mass among the 40, 80, 100, and 120 mg N seedling–1 fertilizer treatments; collectively, these treatments were significantly greater than at 10 and 20 mg N seedling–1 treatments. Seedling N content was greatest for the 100 and 120 mg N seedling–1 rates. These data suggested that nursery optimum N fertilization rate was no less than 100 mg N seedling–1. Outplanting height and root-collar diameter growth characteristics were not significantly different after two years, whereas maximum mean survival was best for seedlings nursery-fertilized at 80 mg N seedling–1. In consideration of both nursery and field performance metrics, our data suggest that exponentially fertilizing Chinese pine seedlings at 80 mg N seedling–1 maximizes both nursery biomass accumulation and outplanting survival.
  • Wang, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing Forestry University, 35 East Qinghua Road, Haidian District, Beijing 100083, China E-mail: wjx198979@163.com
  • Li, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing Forestry University, 35 East Qinghua Road, Haidian District, Beijing 100083, China E-mail: glli226@163.com (email)
  • Pinto, US Department of Agriculture, Forest Service, Rocky Mountain Research Station, 1221 South Main Street, Moscow, ID 83843, USA E-mail: jpinto@fs.fed.us
  • Liu, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing Forestry University, 35 East Qinghua Road, Haidian District, Beijing 100083, China E-mail: 1044902638@qq.com
  • Shi, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing Forestry University, 35 East Qinghua Road, Haidian District, Beijing 100083, China E-mail: shiwenhui2008@163.com
  • Liu, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing Forestry University, 35 East Qinghua Road, Haidian District, Beijing 100083, China E-mail: lyong@bjfu.edu.cn
article id 1293, category Research article
Jukka Malinen, Mika Haring, Harri Kilpeläinen, Erkki Verkasalo. (2015). Comparison of alternative roundwood pricing systems – a simulation approach. Silva Fennica vol. 49 no. 3 article id 1293. https://doi.org/10.14214/sf.1293
Keywords: simulation; pricing; roundwood
Highlights: A discrete event simulation model was developed for studying roundwood pricing systems; For a single buyer, pricing based on residual value appraisal produced (RVA) 4.87 per cent higher wood paying capability and 3.70 per cent higher stumpage price than pricing based on average unit prices; As the number of buyers using RVA increases, the competition increased and the advantage decreased.
Abstract | Full text in HTML | Full text in PDF | Author Info

In a closed market, roundwood buyers pricing system affect the roundwood flow from the stands to different roundwood users. If a buyer is capable to discriminate higher value stands from low quality stands better than its competitors, the buyer should be able to buy better raw material. In the study, a discrete event simulation was used to examine the effect of residual value appraisal (RVA) -based pricing of roundwood by log dimensions and grades compared to the traditional pricing based on average unit prices (UP) of roundwood assortments on roundwood flow. The core of the simulation model was the data containing 51 pine dominated stands from southern Finland. Sample trees were theoretically bucked by the bucking simulator in order to estimate the volumes, dimensions and grades of the logs and roundwood assortments. The simulation model of roundwood markets included four roundwood buyers, two corporations and two saw milling enterprises. The main finding of the study was that the buyers who use RVA gains an advantage and receives better quality compared to buyers who use UP. As the number of buyers using RVA increases, the competition increased and the advantage decreased.

  • Malinen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: jukka.malinen@uef.fi (email)
  • Haring, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: mika.haring@gmail.com
  • Kilpeläinen, Natural Resources Institute Finland (Luke), Bio-based business and industry, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: harri.kilpelainen@luke.fi
  • Verkasalo, Natural Resources Institute Finland (Luke), Bio-based business and industry, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: erkki.verkasalo@luke.fi
article id 1267, category Research article
Caroline Mary Adrianne Franklin, Karen A Harper, Liam Kyte Murphy. (2015). Structural dynamics at boreal forest edges created by a spruce budworm outbreak. Silva Fennica vol. 49 no. 3 article id 1267. https://doi.org/10.14214/sf.1267
Keywords: edge influence; balsam fir; insect disturbance; structure; forest influence
Highlights: Insect outbreak edges were 10 m wide with different canopy cover, stem density and tree structural diversity than adjacent ecosystems; Although edge influence on forest structure was weak, forest influence was stronger and extended further, creating an edge zone skewed towards the disturbed area; After thirty years, high-contrast and structurally-diverse transition zones persist on the landscape.
Abstract | Full text in HTML | Full text in PDF | Author Info
Natural disturbances such as insect outbreaks create boundaries that influence vegetation patterns and ecological processes.  To better understand the effects of natural edge creation on relatively intact forests and adjacent disturbed areas, we investigated forest structure on both sides of 30 year-old forest edges created by a spruce budworm (Choristoneura fumiferana Clemens) outbreak in the boreal forest of Cape Breton Highlands National Park, Canada.  Our objectives were: 1) to determine edge influence (compared to interior forest) and forest influence (compared to disturbed areas) on vegetation structure, and 2) to gain insight into the structural development of the edges.  Canopy cover, tree density, radial growth and deadwood were sampled in 5 m x 20 m plots along 120 m transects across six edges.  Randomization tests were used to estimate the magnitude and distance of edge and forest influence.  Narrow transition zones approximately 10 m wide characterized the spruce budworm-induced edges.  Edge influence did not extend into the forest; however, forest influence on structure was detected up to 40 m from the edge into the disturbed area.  We found evidence of the insect outbreak in the form of reduced radial growth during the disturbance across the entire disturbed area-forest gradient, which indicates that spruce budworm activity may not have ceased directly at the edge.  Tree mortality caused by the insect outbreak resulted in snags, many of which have transformed into logs since the outbreak collapsed.  Spruce budworm outbreak-induced forest edges are narrow but dynamic boundaries separating two distinct vegetation communities in the boreal landscape.
  • Franklin, Department of Renewable Resources, University of Alberta, 751 General Services Building, Edmonton, Alberta, T6G 2H1, Canada E-mail: cfrankli@ualberta.ca (email)
  • Harper, School for Resource and Environmental Studies, Dalhousie University, Suite 5010, 6100 University Ave., Halifax, Nova Scotia, B3H 3J5, Canada E-mail: Karen.Harper@dal.ca
  • Murphy, Department of Environmental Science, Saint Mary’s University, 923 Robie St., Halifax, Nova Scotia, B3H 3C3, Canada E-mail: liamkmurphy@gmail.com
article id 1262, category Research article
Āris Jansons, Roberts Matisons, Māra Zadiņa, Linards Sisenis, Jānis Jansons. (2015). The effect of climatic factors on height increment of Scots pine in sites differing by continentality in Latvia. Silva Fennica vol. 49 no. 3 article id 1262. https://doi.org/10.14214/sf.1262
Keywords: Pinus sylvestris; dendroecology; height growth reconstruction; meteorological conditions; Baltic region; increment variation
Highlights: Height increment-climate relationships of Scots pine were assessed using dendrochronological techniques; Annual height increment was significantly affected by climatic factors; In western Latvia, temperature in preceding summer mainly affected height increment; In eastern Latvia height increment was affected by previous autumn temperature; During the 20th century, the effect of climatic factors has altered likely dues to climate change.
Abstract | Full text in HTML | Full text in PDF | Author Info
Height growth of trees is a crucial parameter that influences the composition and productivity of forest stands and quality of timber; however, the relationships between annual height increment (HI) and climatic factors have been poorly studied. In this study, the effect of monthly mean temperature and precipitation sums on the HI of Scots pine in two sites in Latvia have been determined using dendrochronological techniques. Correlation and response function analyses were conducted for entire chronologies of HI and for 50-year intervals within them. Climatic factors significantly affected the HI of Scots pine; however, not only did the suite of significant factors differ between the sites, but the influence of these factors changed during the 20th century. In the site in western Latvia where climate is milder, temperature in the preceding summer was the main climatic determinant of HI. The effect of temperature in the dormant period and spring was significant during the first part of the 20th century, while the effect of temperature in the previous September and November has become significant since the second half of the 20th century. In the site in eastern Latvia where summers are hotter, HI has been affected by both temperature and water deficit related factors in the summer. However, since the later part of the 20th century, the effect of temperature in the previous October has intensified and become the main climatic determinant of HI.
  • Jansons,  LSFRI “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 E-mail: aris.jansons@silava.lv
  • Matisons,  LSFRI “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 E-mail: robism@inbox.lv (email)
  • Zadiņa,  LSFRI “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 E-mail: mara.zadina@silava.lv
  • Sisenis, LUA Forestry Faculty, Akadēmijas str. 11, Jelgava, Latvia E-mail: linards.sisenis@llu.lv
  • Jansons, Forest Competence Centre, Dzērbenes str. 27, Riga, Latvia, LV1006 E-mail: janis.jansons@silava.lv
article id 1226, category Research article
Santiago Pereira, Antonio Prieto, Rafael Calama, Luis Diaz-Balteiro. (2015). Optimal management in Pinus pinea L. stands combining silvicultural schedules for timber and cone production. Silva Fennica vol. 49 no. 3 article id 1226. https://doi.org/10.14214/sf.1226
Keywords: forest management; goal programming; non-timber forest product
Highlights: Three management scenarios are proposed to integrate timber and pine nuts; Different silvicultural regimes for each output are addressed jointly; Goal programming is used in order to solve forest management models; In the mixed scenario, the area allocated to pine nuts should be notably greater.
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This work aimed to tackle a timber harvest scheduling problem by simultaneously integrating into the analysis two forestry products derived from the same species: the timber and the pine nut. For this purpose, three management scenarios were proposed: two in which each of the productions is maximised separately, and a third mixed where, in each management unit, the product to which the silvicultural effort should be devoted is decided. After defining a set of objectives, and optimising the rotation length, a multi-criteria model based on goal programming was considered since no feasible solutions have been obtained when employing linear programming. The results in our case study show how the feasible solutions reached can be more attractive for the manager. Specifically, the area to be devoted to timber and cone/pine-nut production was computed in a scenario where the optimal silviculture (oriented towards timber or pine nuts) in each stand was selected, and it was concluded that the area allocated to pine nuts should be notably greater. This situation is the opposite of the current management.

  • Pereira, Technical University of Madrid, ETS Ingenieros de Montes, Ciudad Universitaria s/n, 28040 Madrid, Spain E-mail: spereirasaez@gmail.com
  • Prieto, Technical University of Madrid, ETS Ingenieros de Montes, Ciudad Universitaria s/n, 28040 Madrid, Spain E-mail: antonio.prieto@upm.es
  • Calama, Dpto. Selvicultura y Gestión Forestal, INIA-CIFOR, Ctra. A Coruña km 7.5, 28040 Madrid, Spain E-mail: rcalama@inia.es
  • Diaz-Balteiro, Technical University of Madrid, ETS Ingenieros de Montes, Ciudad Universitaria s/n, 28040 Madrid, Spain E-mail: luis.diaz.balteiro@upm.es (email)
article id 1056, category Research article
Katri Himanen, Markku Nygren. (2015). Seed soak-sorting prior to sowing affects the size and quality of 1.5-year-old containerized Picea abies seedlings. Silva Fennica vol. 49 no. 3 article id 1056. https://doi.org/10.14214/sf.1056
Keywords: Norway spruce; seed treatment; seed radiography; seedling emergence; seedling morphology
Highlights: After soak-sorting all sunken seeds (bottom fraction) were full and viable, whereas floating seeds contained larvae-filled and immature seed; Seedlings originating from the bottom fraction were greater in height and diameter than control seedlings or those originating from the floating seeds; The proportion of saleable seedlings was four percentage points higher in the bottom fraction than in the other seedlings.
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We studied the effect of soak-sorting Norway spruce (Picea abies (L.) H. Karst.) seeds on emergence, development and quality of container seedlings in two commercial seed lots. The seeds, separated by soaking into bottom and surface fractions, were sown in June, and the seedlings were grown during two growing seasons under typical Finnish nursery conditions. The first summer seedlings were grown in a greenhouse and outdoors for the second, full growing season. All sunken seeds were full and viable according to radiography, whereas the floating seeds contained 2% and 13% larvae-filled and 8% and 11% anatomically immature seeds, depending on the seed lot. Seedlings grown from the bottom fraction seed emerged 2.5–3.5 days earlier than seedlings of storage dry (i.e. control) seed. Height, diameter, and shoot and root dry mass of the seedlings were affected by soaking after both the first and second growing seasons. The largest seedlings originated from the bottom fraction. The proportion of saleable seedlings was four percentage points higher in the bottom fraction than in the other seedlings. The effects of soaking found in this study are more notable than as previously reported for Norway spruce seedlings. This suggests that soaking and soak-sorting may be most useful when the growing conditions are stressful, i.e. when seeds are sown in summer rather than 1-year-old seedling crops sown in spring under the climate conditions typical of Finland.

  • Himanen, Natural Resources Institute Finland (Luke), Green technology, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: katri.himanen@luke.fi (email)
  • Nygren, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: markku.nygren@luke.fi
article id 1280, category Research article
Juha Laitila, Tapio Ranta, Antti Asikainen, Eero Jäppinen, Olli-Jussi Korpinen. (2015). The cost competitiveness of conifer stumps in the procurement of forest chips for fuel in Southern and Northern Finland. Silva Fennica vol. 49 no. 2 article id 1280. https://doi.org/10.14214/sf.1280
Keywords: stumps; quality; delimbed stems; pre-grinding; procurement; heating value
Highlights: Pre-grinding and integrated screening is a way of guaranteeing fuel quality, but, when the stumps’ ash content is six per cent or below, the procurement costs are higher than with grinding of stumps at the plant. Because of high transportation costs, stump harvesting is the most profitable in Southern Finland, where there is greater availability of stumps than in Northern Finland.
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The aim of this study was to evaluate cost competitiveness, at regional level, of various systems for stump transportation and grinding, and to compare the results to the procurement costs of delimbed stems from early thinnings at the stand and regional level. The accumulation and procurement costs of stumps and delimbed stems were estimated within a 100-kilometer radius from two power plants located in Kouvola and in Kajaani. The analyses were performed as simulated treatments in clear cuts and thinnings of young stands, using existing productivity and cost functions, alternative ash percentages for stump wood, and yield calculations based on the forest industry regeneration felling stand data and the sample plots data of the National Forest Inventory of Finland. The results were expressed as Euros per solid cubic meter (€ m–3) and Euros per megawatt hour (€ MWh–1). The results highlight the need to improve stump fuel quality and increase the heating value. The procurement cost of stumps was about 1 € MWh–1 lower in Kouvola compared to Kajaani, when using conceivable ash content of 6% for stumps ground at the plant, and ash content of 1.5% for stumps pre-ground at the roadside landing. The procurement costs of stumps were, on average, 0.55 € MWh–1 lower compared to delimbed stems in Kouvola, and on average 0.6 € MWh–1 higher in Kajaani. Pre-grinding and integrated screening is a feasible way to guarantee the fuel quality expressed as ash content already at roadside landings, but the procurement costs are higher compared to grinding stumps at the plant, when the ash content of ground stumps is 6% or less.
  • Laitila, Natural Resources Institute Finland (Luke), Bio-based Business and Industry, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: juha.laitila@metla.fi (email)
  • Ranta, Lappeenranta University of Technology, Sammonkatu 12, FI-50130 Mikkeli, Finland E-mail: tapio.ranta@lut.fi
  • Asikainen, Natural Resources Institute Finland (Luke), Bio-based Business and Industry, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: antti.asikainen@metla.fi
  • Jäppinen, Lappeenranta University of Technology, Sammonkatu 12, FI-50130 Mikkeli, Finland E-mail: eero.jappinen@lut.fi
  • Korpinen, Lappeenranta University of Technology, Sammonkatu 12, FI-50130 Mikkeli, Finland E-mail: olli-jussi.korpinen@lut.fi
article id 1279, category Research article
Andreas Kreutz, Tuomas Aakala, Russell Grenfell, Timo Kuuluvainen. (2015). Spatial tree community structure in three stands across a forest succession gradient in northern boreal Fennoscandia. Silva Fennica vol. 49 no. 2 article id 1279. https://doi.org/10.14214/sf.1279
Keywords: Picea abies; age structure; spatial structure; natural forest; Betula spp.; point pattern analysis; northern boreal; community composition
Highlights: We studied the tree community spatial structure in three 1.2-ha plots representing naturally developed northern boreal forests of varying ages; Spatial structure showed little differences between the mid-successional, late-successional and old-growth stands; The occurrence of Picea abies relative to Betula spp. indicated a mosaic-like spatial assembly; Mosaics are likely maintained by species-specific replacement, not reciprocal replacement as thought earlier.
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Development of species composition during succession is well studied in natural boreal forests, but empirical assessments of how within-stand spatial structure develops in late-successional stages are few. Here, we quantified spatial patterns in three unmanaged stands consisting of Picea abies (L.) Karst. and Betula pubescens Ehrh. and Betula pendula Roth (hereafter Betula spp.) in northern boreal Fennoscandia. We conducted a comprehensive analysis of small-scale spatial point patterns in three fully mapped 1.2-ha sample plots, representing different forest developmental stages: mid-successional, late-successional and old-growth forest. We used several variants of Ripley’s K-function to analyze the spatial point patterns along the successional gradient. Univariate analyses showed that mature trees of both species were either randomly distributed or clumped. P. abies saplings were clumped, and Betula spp. saplings occurred in a random or clumped manner. In the bivariate analyses, saplings were more likely to be found in the surroundings of mature trees of the same species, but occurred independent of the individuals of other tree species. Mature trees showed interspecific repulsion. Only modest differences occurred in the univariate patterns between the three successional stages, but in the bivariate analyses the most evident patterns, i.e. intraspecific attraction and interspecific repulsion, were stronger in the older successional stages. Overall, the studied stands appear structured as species-specific mosaics. These mosaics, along with mixed species composition, seem to be maintained by species self-replacement, which contrasts with findings from earlier studies.
  • Kreutz, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: andreas.kreutz@wald-rpl.de
  • Aakala, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014 University of Helsinki, Finland ORCID http://orcid.org/0000-0003-0160-6410 E-mail: tuomas.aakala@helsinki.fi (email)
  • Grenfell, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: russell.grenfell@gmail.com
  • Kuuluvainen, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: timo.kuuluvainen@helsinki.fi
article id 1274, category Research article
Juho Hautsalo, Paul Mathieu, Sakina Elshibli, Pekka Vakkari, Juha Raisio, Pertti Pulkkinen. (2015). Variation in height and survival among northern populations of pedunculate oak (Quercus robur L.): results of a 13-year field study. Silva Fennica vol. 49 no. 2 article id 1274. https://doi.org/10.14214/sf.1274
Keywords: adaptation; growth; pedunculate oak; northern stands
Highlights: A height-sum function allowed us to compare the combined growth and survival of northernmost stands of pedunculate oak; Individuals from Turku-Katariinanlaakso performed the best, although other families performed more consistently across trials, which should be considered in future conservation and breeding. Surprisingly, trees planted in a trial location beyond the natural northern limit of pedunculate oak showed the best performance.
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We analysed the adaptive potential of pedunculate oak (Quercus robur L.) in terms of variation in height and survival in five field trials located in southern and central Finland. The trials were established with Finnish native material from six different seed origins. Thirteen years after planting, the number of living trees was counted and measured for height. Analysis of height and survival revealed a significant effect of origin, i.e., a genetic basis to individual tree performance. Two origins from the Turku region (Ruissalo and Katariinanlaakso) performed the best while trees originating from Parainen (Lenholmen) performed the worst. In order to study the effects due to tree origin, a comparison of families (half-sibling trees, i.e. those sharing the same ‘mother’ tree) was made by combining height and survival through a height-sum equation (i.e., the product of mean survival and height of each family in each trial) and used to calculate family- and origin-level ecovalences. Ecovalence is a metric for performance consistency, and indicates how much each variable contributes to the total variation; the higher the value, the lower the consistency of trees across the trials based on their origin or family. Analysis of consistency showed similar results to growth and survival, with Turku families performing the best and families from Parainen performing the worst. Families in the Katariinanlaakso stand (Turku) generally had more stable ecovalence values and more dispersed height-sums, while Ruissalo (Turku) families had higher mean height-sum but higher variability in ecovalence values. These results suggest that seed origins (i.e., genotypes) can be optimized in terms of their suitability for commercial or ecological forest management.
  • Hautsalo,  Natural Resources Institute Finland (Luke), Green technology, Antinniementie 1, FI-41330 Vihtavuori, Finland E-mail: juho.hautsalo@luke.fi (email)
  • Mathieu, Agrocampus Ouest, 35000 Rennes, France E-mail: pm@nn.fr
  • Elshibli, University of Helsinki, Helsinki, Finland E-mail: se@nn.fi
  • Vakkari, Natural Resources Institute Finland (Luke), Vantaa, Finland E-mail: pekka.vakkari@luke.fi
  • Raisio, City of Helsinki, Helsinki, Finland E-mail: jr@nn.fi
  • Pulkkinen, Natural Resources Institute Finland (Luke), Vantaa, Finland E-mail: pertti.pulkkinen@luke.fi
article id 1266, category Research article
Edwyn K. Midmore, Shelagh A. McCartan, Richard L. Jinks, Christine M. Cahalan. (2015). Using thermal time models to predict germination of five provenances of silver birch (Betula pendula Roth) in southern England. Silva Fennica vol. 49 no. 2 article id 1266. https://doi.org/10.14214/sf.1266
Keywords: climate change; seed source; assisted migration; thermal time model
Highlights: Using cumulative germination data, thermal time models were developed for Betula pendula; Models indicated varying degrees of dormancy and pre-chill requirements among provenances; Thermal time parameters were used with climatic data to predict germination times under mild and cold winters in southern England; Predictions suggest that pre-chilled French seeds would germinate about six weeks later than the fastest germinating provenance.
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Climate predictions indicate that growing conditions may become unfavourable for certain tree species in parts of Britain. Guidelines suggest some planting of seed sources from regions between 2° and 5° south of those currently used as part of a climate change adaptation strategy. However, there has been little research on the benefits and risks associated with the use of planting stock from more southerly seed sources. Seeds of five provenances of the ‘relatively’ dormant Betula pendula were germinated over a range of temperatures both with and without a pre-chill. Subsequently, a thermal time model was used to predict the impact of migrating these provenances to southern England. Results identified geographical differences in germination response; those from higher latitude were more sensitive to pre-chill.
  • Midmore, Forest Research Agency, Alice Holt, Surrey. Current: Dolwyddelan, Llandre, Ceredigion, Wales, SY24 5BZ E-mail: emidmore@gmail.com
  • McCartan, Forest Research, Alice Holt, Farnham, Surrey, GU10 4LH, UK E-mail: shelagh.mccartan@forestry.gsi.gov.uk (email)
  • Jinks, Forest Research, Alice Holt Lodge, Farnham, Surrey, GU10 4LH, UK E-mail: richard.jinks@forestry.gsi.gov.uk
  • Cahalan, Bangor University, School of Environment, Natural Resources and Geography, Bangor, Gwynedd, Wales, LL57 2UW E-mail: c.m.cahalan@bangor.ac.uk
article id 1239, category Research article
Tomi Kaakkurivaara, Nuutti Vuorimies, Pauli Kolisoja, Jori Uusitalo. (2015). Applicability of portable tools in assessing the bearing capacity of forest roads. Silva Fennica vol. 49 no. 2 article id 1239. https://doi.org/10.14214/sf.1239
Keywords: stiffness; light weight deflectometer; dynamic cone penetrometer; falling weight deflectometer; elastic modulus
Highlights: The dynamic cone penetrometer (DCP) and light falling weight deflectometer (LFWD) are useful tools for measuring bearing capacity; The measurement results are not same as with the falling weight deflectometer (FWD), but comparable.
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Forest roads provide access to logging sites and enable transportation of timber from forest to mills. Efficient forest management and forest industry are impossible without a proper forest road network. The bearing capacity of forest roads varies significantly by weather conditions and seasons since they are generally made of poor materials and the constructed layers may be mixed with subgrade. A bearing capacity assessment is valuable information when trafficability is uncertain and rutting is obvious. In this study, bearing capacity measurements were carried out using the light falling weight deflectometer (LFWD), the dynamic cone penetrometer (DCP) and the conventional falling weight deflectometer (FWD). The aim was to compare their measurement results in relation to road characteristics and moisture conditions. Data were collected from 35 test road sections in four consecutive springs and during one summer. The test road sections had measurement points both on the wheel path and the centre line. The data show logical correlations between measured quantities, and the study presents reliable regression models between measuring devices. The results indicate that light portable tools, the DCP and the LFWD, can in most cases be used instead of the expensive falling weight deflectometer on forest roads.
  • Kaakkurivaara, Natural Resources Institute Finland, Green technology, Kaironiementie 15, FI-39700 Parkano, Finland E-mail: tomi.kaakkurivaara@gmail.com (email)
  • Vuorimies, Tampere University of Technology, P.O.Box 600, FI-33101 Tampere, Finland E-mail: nuutti.vuorimies@tut.fi
  • Kolisoja, Tampere University of Technology, P.O.Box 600, FI-33101 Tampere, Finland E-mail: pauli.kolisoja@tut.fi
  • Uusitalo, Natural Resources Institute Finland, Green technology, Kaironiementie 15, FI-39700 Parkano, Finland E-mail: jori.uusitalo@luke.fi
article id 1232, category Research article
Pete Bettinger, Mehmet Demirci, Kevin Boston. (2015). Search reversion within s-metaheuristics: impacts illustrated with a forest planning problem. Silva Fennica vol. 49 no. 2 article id 1232. https://doi.org/10.14214/sf.1232
Keywords: forest planning; heuristics; threshold accepting; tabu search; spatial harvest scheduling; adjacency constraints; mixed integer goal programming
Highlights: The interruption of the sequence of events used to explore a solution space and develop a forest plan, and the re-initiation of the search process from a high-quality, known starting point (reversion) seems necessary for some s-metaheuristics; When using a s-metaheuristic, higher quality forest plans may be developed when the reversion interval is around six iterations of the model.
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The use of a reversion technique during the search process of s-metaheuristics has received little attention with respect to forest management and planning problems. Reversion involves the interruption of the sequence of events that are used to explore the solution space and the re-initiation of the search process from a high-quality, known starting point. We explored four reversion rates when applied to three different types of s-metaheuristics that have previously shown promise for the forest planning problem explored, threshold accepting, tabu search, and the raindrop method. For two of the s-metaheuristics, we also explored three types of decision choices, a change to the harvest timing of a single management unit (1-opt move), the swapping of two management unit’s harvest timing (2-opt moves), and the swapping of three management unit’s harvest timing (3-opt moves). One hundred independent forest plans were developed for each of the metaheuristic / reversion rate combinations, all beginning with randomly-generated feasible starting solutions. We found that (a) reversion does improve the quality of the solutions generated, and (b) the rate of reversion is an important factor that can affect solution quality.
  • Bettinger, School of Forestry and Natural Resources, 180 E. Green Street, University of Georgia, Athens, Georgia, USA 30602 E-mail: pbettinger@warnell.uga.edu (email)
  • Demirci, General Directorate of Forestry, Ministry of Forest and Water Affairs, Republic of Turkey E-mail: mehmetdemirci@yahoo.com
  • Boston, Department of Forest Engineering, Resources and Management, College of Forestry, Oregon State University, USA E-mail: Kevin.Boston@oregonstate.edu
article id 1218, category Research article
Mikko Niemi, Mikko Vastaranta, Jussi Peuhkurinen, Markus Holopainen. (2015). Forest inventory attribute prediction using airborne laser scanning in low-productive forestry-drained boreal peatlands. Silva Fennica vol. 49 no. 2 article id 1218. https://doi.org/10.14214/sf.1218
Keywords: remote sensing; forest technology; forest management planning; mapping; k-NN estimation; random forests
Highlights: Following current forest inventory practises, stem volume was predicted in low-productive drained peatlands (LPDPs) with a root mean square error (RMSE) of 13.7 m3 ha–1; When 30 reference plots measured from LPDPs were added to the prediction, RMSE was decreased to 10.0 m3 ha–1; Additional reference plots from LPDPs did not affect the forest inventory attribute predictions in productive forests.
Abstract | Full text in HTML | Full text in PDF | Author Info
Nearly 30% of Finland’s land area is covered by peatlands. In Northern parts of the country there is a significant amount of low-productive drained peatlands (LPDPs) where the average annual stem volume growth is less than 1 m3 ha–1. The re-use of LPDPs has been considered thoroughly since Finnish forest legislation was updated and the forest regeneration prerequisite was removed from LPDPs in January 2014. Currently, forestry is one of the re-use alternatives, thus detailed forest resource information is required for allocating activities. However, current forest inventory practices have not been evaluated for sparse growing stocks (e.g., LPDPs). The purpose of our study was to evaluate the suitability of airborne laser scanning (ALS) for mapping forest inventory attributes in LPDPs. We used ALS data with a density of 0.8 pulses per m2, 558 field-measured reference plots (500 from productive forests and 58 from LPDPs) and k nearest neighbour (k-NN) estimation. Our main aim was to study the sensitivity of predictions to the number of LPDP reference plots used in the k-NN estimation. When the reference data consisted of 500 plots from productive forest stands, the root mean square errors (RMSEs) for the prediction accuracy of Lorey’s height, basal area and stem volume were 1.4 m, 2.7 m2 ha–1 and 13.7 m3 ha–1 in LPDPs, respectively. When 30 additional reference plots were allocated to LPDPs, the respective RMSEs were 1.1 m, 1.7 m2 ha–1 and 10.0 m3 ha–1. Additional reference plot allocation did not affect the predictions in productive forest stands.
  • Niemi, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014, Finland & Centre of Excellence in Laser Scanning Research, Finnish Geospatial Research Institute FGI, Geodeetinrinne 2, FI-02430, Finland E-mail: mikko.t.niemi@helsinki.fi (email)
  • Vastaranta, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014, Finland & Centre of Excellence in Laser Scanning Research, Finnish Geospatial Research Institute FGI, Geodeetinrinne 2, FI-02430, Finland E-mail: mikko.vastaranta@helsinki.fi
  • Peuhkurinen, Arbonaut Oy Ltd., Latokartanontie 7 A, FI-00700, Finland E-mail: jussi.peuhkurinen@arbonaut.com
  • Holopainen, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014, Finland & Centre of Excellence in Laser Scanning Research, Finnish Geospatial Research Institute FGI, Geodeetinrinne 2, FI-02430, Finland E-mail: markus.holopainen@helsinki.fi
article id 1300, category Research article
Jaana Luoranen, Risto Rikala. (2015). Post-planting effects of early-season short-day treatment and summer planting on Norway spruce seedlings. Silva Fennica vol. 49 no. 1 article id 1300. https://doi.org/10.14214/sf.1300
Keywords: Picea abies; height growth; survival; bud burst; short-day treatment; summer planting
Highlights: Summer planting and short-day treatment advanced the bud burst and increased the height of Norway spruce seedlings after planting, compared to autumn and spring planted or untreated seedlings.
Abstract | Full text in HTML | Full text in PDF | Author Info
Effects of short-day (SD) treatment on bud burst, growth and survival of Norway spruce (Picea abies [L.] Karst.) container seedlings after summer planting were studied in an experiment established in Suonenjoki, Central Finland. One-year-old seedlings were SD-treated for three weeks starting on 18 June, 24 June and 8 July 2004 and then planted on 22 July, 5 August, 6 September 2004 and, as a normal spring planting, on 10 May, 2005. Untreated control seedlings were also planted on these dates. Second flush on the planting year and bud burst the following spring was monitored in planted seedlings, whereas seedling height and survival were determined at the end of growing seasons 2004–2006. We observed a non-significant risk of a second flush if seedlings were SD-treated on 18 June. Also, SD-treated seedlings planted in July or August showed advanced bud burst and increased height the following growing season without significant effects on survival, compared to autumn and spring planted seedlings. Planting in July or early August was associated with a significant increase in the incidence of multiple leaders in later years. Based on our results, to begin a three-week SD treatment in late June or early July and then plant seedlings in late July or early August could be a good practice.
  • Luoranen, Natural Resources Institute Finland (Luke), Natural resources and bioproduction, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: jaana.luoranen@luke.fi (email)
  • Rikala, The Finnish Forest Research Institute, Suonenjoki Unit, Suonenjoki, Finland E-mail: rikala@dnainternet.fi
article id 1260, category Research article
Jyrki Hytönen, Anna Saarsalmi. (2015). Biomass production of coppiced grey alder and the effect of fertilization. Silva Fennica vol. 49 no. 1 article id 1260. https://doi.org/10.14214/sf.1260
Keywords: biomass production; coppicing; grey alder; fertilisation
Highlights: Fertilisation (wood ash, N, PK) did not affect grey alder biomass production; Leafless above-ground biomass of 17–20 year old stands was 52–57 Mg ha–1; MAI increased with increase of rotation length to the end of the follow-up period of 17–20 years; Coppicing increased stand density manifold.
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We studied biomass production of two naturally originated grey alder (Alnus incana (L.) Moench) stands having a mixture of birch and willow located in central Finland. One of the stands was growing on a peatland site (Muhos) and the other on a mineral soil site (Juuka). The stands were clear-cut and fertilization experiments were laid out with several treatments. At Muhos, the treatments included nitrogen fertilisation with different amounts of wood ash and an unfertilized control. At Juuka, the treatments included nitrogen fertilisation either with ash or with PK, and ash and PK treatments alone and an unfertilized control. The sprouts at Muhos were grown for 17 years and at Juuka for 20 years. At Juuka the stand was clear-cut second time at the age of 20 years and grown for 8 years. The stands were measured several times and foliar samples were taken twice during the study period. Clear-cutting increased stem number manifold. The stand density of new coppiced forests after the clear-cutting decreased from 67 000–89 000 stems ha–1 at the age of 3–6 years to 10 000–12 000 stems ha–1 at the age of 17–20 years. On neither site fertilization affected biomass production of alders during the study period. Leafless above-ground biomass was 52–57 Mg ha–1 after 17–20 years. Mean annual leafless above-ground biomass production (MAI) increased with increase of rotation time. At the age of 17–20 years the MAI was 2.8–3.0 Mg ha a–1. At Muhos, ash increased foliar P and Ca concentrations, but decreased those of Mn.
  • Hytönen, Natural Resources Institute Finland, Silmäjärventie 2, FI-69100 Kannus, Finland E-mail: jyrki.hytonen@luke.fi (email)
  • Saarsalmi, Natural Resources Institute Finland, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: anna.saarsalmi@luke.fi
article id 1220, category Research article
Elisabeth Düthorn, Lea Schneider, Oliver Konter, Philipp Schön, Mauri Timonen, Jan Esper. (2015). On the hidden significance of differing micro-sites on tree-ring based climate reconstructions. Silva Fennica vol. 49 no. 1 article id 1220. https://doi.org/10.14214/sf.1220
Keywords: Pinus sylvestris; climate change; Picea abies; Finland; temperature reconstruction; RCS detrending
Highlights: Pines and spruces show growth level differences in wet and dry micro-sites with higher growth rates in the dry sites; Spruces show a robust climate-growth relationship with June-July temperatures; Application of collective detrending methods can bias long-term trends in climate reconstructions, if relict and recent samples originate from different micro-sites.
Abstract | Full text in HTML | Full text in PDF | Author Info
Tree-ring chronologies are commonly extended back in time by combining samples from living trees with relict material preserved in man-made structures or natural archives (e.g. lakes). Although spatially close, these natural archives and living-tree-sites often comprise different micro-climates. Inhomogeneous growth conditions among these habitats, which may yield offsets in growth-rates, require caution in data processing. Here we assess species-specific growth dynamics in two micro-habitats and their potential effects on long chronologies by combining tree-ring data from different living-tree-sites with an “artificial” subfossil dataset. Well replicated (n > 80) Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) chronologies from northern Fennoscandia, sampled directly at the lakeshore (wet) and several meters beyond the lakeshore (dry) reveal high coherence of the variance between micro-sites (rspruce = 0.59, rpine = 0.68). Significant differences of the Regional Curves (RC) indicate faster growth of both species at the drier site though. Growth differences are more pronounced between the spruce micro-sites. The combination of recent dry and wet spruce data with artificial relict data results in two long chronologies covering the last 800 years with substantially different trends, although they consist of the same relict material and the micro-site chronologies correlate significantly over the past two centuries. The combination of spruce samples from dry inland micro-sites with subfossil samples originating from the wet lake shore can result in an underestimation of past temperatures prior to the 19th century. Such effects, hidden in the composition of long chronologies (living trees + subfossil samples) can bias long-term trends in climate reconstructions.
  • Düthorn, Department of Geography, Johannes Gutenberg University, Becherweg 21, 55099 Mainz, Germany E-mail: duethorn@uni-mainz.de (email)
  • Schneider, Department of Geography, Johannes Gutenberg University, Becherweg 21, 55099 Mainz, Germany E-mail: l.schneider@geo.uni-mainz.de
  • Konter, Department of Geography, Johannes Gutenberg University, Becherweg 21, 55099 Mainz, Germany E-mail: O.Konter@geo.uni-mainz.de
  • Schön, Department of Geography, Johannes Gutenberg University, Becherweg 21, 55099 Mainz, Germany E-mail: philipp.schoen@gmx.de
  • Timonen, Natural Resources Institute Finland (Luke), Natural resources and bioproduction, FI-96301 Rovaniemi, Finland E-mail: mauri.timonen@metla.fi
  • Esper, Department of Geography, Johannes Gutenberg University, Becherweg 21, 55099 Mainz, Germany E-mail: J.Esper@geo.uni-mainz.de
article id 1214, category Research article
Katri Hamunen, Outi Virkkula, Teppo Hujala, Juha Hiedanpää, Mikko Kurttila. (2015). Enhancing informal interaction and knowledge co-construction among forest owners. Silva Fennica vol. 49 no. 1 article id 1214. https://doi.org/10.14214/sf.1214
Keywords: communities of practice; forest owner clubs; guidance; knowledge sharing; learning communities; peer learning; social network
Highlights: Forest owners’ own communities could complement the present expert-driven forestry extension; Enhancing informal interaction between forest owners calls for sufficiently homogeneous reference groups and also new communication contexts; New purposes of forest ownership and innovative forest management practices are suggested topics for novel owner communities; Forest professionals may initiate these communities, but the continuity depends on the owners themselves.
Abstract | Full text in HTML | Full text in PDF | Author Info
It is a common concern that non-resident private forest owners are less able to make informed decisions regarding their forests. Moreover, the present guidance given by forest professionals is not reaching all owners. In this study, we suggest enhancing knowledge exchange among forest owners by increasing their mutual and informal interaction that could inspire them to co-construct new knowledge. The first objective is to identify present emerging activities that constitute knowledge exchange contexts (communities) for Finnish forest owners. The second objective is to discuss the challenges of current Finnish forest extension and their implications when introducing Communities of Practice as a complementary response to existing, yet insufficient, professional-led extension. Data consist of Finnish forest owners’ and forest professionals’ (n = 43) focus group interviews. The qualitative analysis was theoretically oriented followed by data-driven coding and grouping. According to the results, the role of expert-led encounters is strong in Finland and owners’ opportunities for good mutual communication are rare. Informal communities exist mainly in the countryside among neighbouring owners and within families. To enhance knowledge sharing among owners, one needs to identify innovative topics and activities that would inspire owners to commit to their forest property and perform active silviculture. These communities would operate as creative learning environments allowing participation of different levels. When cultivating forest owners’ communities it is important to consider actors’ roles. Forest owners themselves are responsible for the functions and continuity of these communities, although forest professionals could also initiate new and sufficiently homogenous reference groups for the owners.
  • Hamunen, Natural Resources Institute Finland (Luke), New Business Opportunities, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: katri.hamunen@luke.fi
  • Virkkula, Oulu University of Applied Sciences, Kotkantie 1, FI-90250 Oulu, Finland E-mail: outi.virkkula@oamk.fi
  • Hujala, Natural Resources Institute Finland (Luke), New Business Opportunities, P.O. Box 18, FI-01301 Vantaa, Finland ORCID http://orcid.org/0000-0002-7905-7602 E-mail: teppo.hujala@luke.fi (email)
  • Hiedanpää, Natural Resources Institute Finland (Luke), Economy and Society, Itäinen Pitkäkatu 3, FI-20520 Turku, Finland E-mail: juha.hiedanpaa@luke.fi
  • Kurttila, Natural Resources Institute Finland (Luke), New Business Opportunities, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: mikko.kurttila@luke.fi
article id 1191, category Research article
Tore Skrøppa, Halvor Solheim, Arne Steffenrem. (2015). Genetic variation, inheritance patterns and parent–offspring relationships after artificial inoculations with Heterobasidion parviporum and Ceratocystis polonica in Norway spruce seed orchards and progeny tests. Silva Fennica vol. 49 no. 1 article id 1191. https://doi.org/10.14214/sf.1191
Keywords: blue stain; heritability; root and butt rot; lesion length
Highlights: Genetic variation is demonstrated in response to artificial inoculations with Heterobasidion parviporum and Ceratocystis polonica both between parents and their offspring;Strong relationships are observed between the male parents and their off-spring, less so between the female parents and their offspring.
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Inoculations with the two fungi Heterobasidion parviporum and Ceratocystis polonica were made in two series of progeny tests each containing full-sib families planted at two sites and on grafts of the parents in two seed orchards. Significant variation among families in lesion lengths after inoculation was found for both fungi and a predominantly additive inheritance was indicated. The estimates of narrow sense heritability were 0.13 and 0.22 for H. parviporum and C. polonica, respectively. The estimate of the genetic correlation between the lesion lengths of the two fungi was as low as 0.12. Significant variation in lesion lengths was also found among parental clones, and within ramets of the same clone, in the seed orchards. In one of the series a high positive correlation (r = 0.88) was found between the H. parviporum lesion lengths of the male parents and offspring, but not for the female parents and off-spring. The results confirm earlier conclusions that the genetic variation and heritabilities are large enough for practical breeding for resistance.
  • Skrøppa, Norwegian Forest and Landscape Institute, Box 115, 1431 Ås, Norway E-mail: tore.skroppa@skogoglandskap.no (email)
  • Solheim, Norwegian Forest and Landscape Institute, Box 115, 1431 Ås, Norway E-mail: halvor.solheim@skogoglandskap.no
  • Steffenrem, Norwegian Forest and Landscape Institute, Box 115, 1431 Ås, Norway E-mail: arne.steffenrem@skogoglandskap.no
article id 1141, category Research article
Isabel Miranda, Jorge Gominho, Helena Pereira. (2015). Heartwood, sapwood and bark variation in coppiced Eucalyptus globulus trees in 2nd rotation and comparison with the single-stem 1st rotation. Silva Fennica vol. 49 no. 1 article id 1141. https://doi.org/10.14214/sf.1141
Keywords: stand density; stem analysis; eucalypt silviculture; harvesting cycle; within-tree variation
Highlights: Coppiced E. globulus trees in the 2nd rotation have similar heartwood and sapwood development as single-stem trees in the 1st rotation; The initial tree planting density did not influence heartwood development of coppiced E. globulus trees; Heartwood diameter and height can be modelled with tree diameter and height respectively; Sapwood width is approximately constant within and between coppice and single-stem E. globulus trees.
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Coppiced Eucalyptus globulus trees with 18 years in a 2nd rotation were analysed in relation to heartwood, sapwood and bark content taking into account the effect of the initial planting density by using a spacing trial. A total of 25 stumps, with a variable number of stems per stump from 1 to 3, were analysed. Comparison was made to the previous 1st rotation single stem trees, also harvested at 18 years. In the 2nd rotation, the stump density did not significantly affect stem height and diameter, in opposition to the 1st rotation where spacing significantly impacted on tree dimensions. The effect of the initial planting density is somewhat lost in the coppiced stand in relation with i.e. the number of stems per stump. Heartwood was present in all the coppiced trees up to 49.9% of the total tree height and heartwood volume amounted to 38.9–51.7% of the total tree volume. Within the tree, heartwood content decreased from the base upwards, representing, on average, 54.1% at the base and decreasing to 5.1% at 15.3 m. The sapwood width remained relatively constant with an average radial width of approximately 2 cm. The average stem bark content of coppiced trees was 17.4% of the total stem volume. The comparison of heartwood and sapwood development in the coppiced trees did not show significant differences to the 1st rotation trees, nor did the initial spacing. Heartwood diameter could be modelled using the tree diameter both for 1st and 2nd rotation trees.
  • Miranda, Centro de Estudos Florestais, Instituto Superior de Agronomia, Universidade Lisboa, 1349-017 Lisboa, Portugal E-mail: Imiranda@isa.ulisboa.pt
  • Gominho, Centro de Estudos Florestais, Instituto Superior de Agronomia, Universidade Lisboa, 1349-017 Lisboa, Portugal E-mail: Jgominho@isa.utl.pt (email)
  • Pereira, Centro de Estudos Florestais, Instituto Superior de Agronomia, Universidade Lisboa, 1349-017 Lisboa, Portugal E-mail: Hpereira@isa.utl.pt
article id 1106, category Research article
Jena Ferrarese, David Affleck, Carl Seielstad. (2015). Conifer crown profile models from terrestrial laser scanning. Silva Fennica vol. 49 no. 1 article id 1106. https://doi.org/10.14214/sf.1106
Keywords: prediction; canopy shape; parametric curves; interior Northwest USA
Highlights: Crown models are derived from terrestrial laser data for 3 NW USA conifer species; Crown models require only crown length for implementation; Beta and Weibull curves fit to 95th percentile widths describe crown extent; Crown profile curves are species-specific and not interchangeable; Crown shape is not strongly conditioned by tree size or site.
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Regional crown profile models were derived for three conifer species of the interior northwestern USA from terrestrial laser scans of eighty-six trees across a range of sizes and growing conditions. Equations were developed to predict crown shape from crown length for Pseudotsuga menziesii, Pinus ponderosa, and Abies lasiocarpa from parametric curves applied to crown-length normalized laser point clouds. The 95th width percentile adequately described each crown’s outer limit; alternate width percentiles produced little profile shape variation. For P. menziesii and P. ponderosa, a scaling parameter-modified beta curve gave the most accurate fit (using cross-validated Mean Absolute Error) to aggregated 95th width percentile points. For A. lasiocarpa, beta and Weibull curves (equivalently modified) produced similar results. For all species, modified beta and Weibull curves fit crown points with less error than conic or cylindrical profiles. Crown profile curves were species-specific; interchanging among species increased error significantly. Laser-derived crown base metrics provided objectivity and consistency, but underestimated field-derived base heights through inclusion of dead branches. Profile curve parameters were not correlated with tree or stand characteristics suggesting that crown shape is not strongly conditioned by size and site factors. However, laser sampling necessarily favored more open growing conditions, potentially under-representing variations in crown shape associated with social position. Overall, Terrrestrial Laser Scanning (TLS) lends itself to detailed measurements of external crown architecture with occlusion-imposed limits to characterization of internal features. Yet, the time and cost of collecting and processing individual tree data precludes use of TLS as a common field sampling tool.
  • Ferrarese, College of Forestry and Conservation, The University of Montana, Missoula, MT, USA; (present) Center for the Environmental Management of Military Lands, 1490 Campus Delivery, Colorado State University, Fort Collins, CO 80523, USA E-mail: jena.ferrarese@colostate.edu (email)
  • Affleck, College of Forestry and Conservation, University of Montana, Missoula, MT, USA E-mail: david.affleck@cfc.umt.edu
  • Seielstad, College of Forestry and Conservation, University of Montana, Missoula, MT, USA E-mail: carl.seielstad@firecenter.umt.edu
article id 1219, category Research article
Thomas P. Sullivan, Druscilla S. Sullivan. (2014). Diversifying clearcuts with green-tree retention and woody debris structures: conservation of mammals across forest ecological zones. Silva Fennica vol. 48 no. 5 article id 1219. https://doi.org/10.14214/sf.1219
Keywords: clearcutting; green-tree retention; small mammals; coniferous forests; ecological zones; Myodes gapperi; population dynamics; red-backed voles; woody debris structures
Highlights: Species diversity of small mammals increased with structural complexity left on clearcut sites; Productivity of red-backed vole populations was higher in sites with green-tree retention (GTR) and windrows of woody debris; GTR and windrows may provide additive effect for providing habitat to conserve mammals on clearcuts.
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We tested the hypotheses (H) that on newly clearcut-harvested sites, (H1) abundance and species diversity of the forest-floor small mammal community, and (H2) abundance, reproduction, and recruitment of red-backed voles (Myodes gapperi Vigors), would increase with higher levels of structural retention via green-tree retention (GTR) and woody debris (dispersed and constructed into windrows). Study areas were located in three forest ecological zones in southern British Columbia, Canada. For H1, mean total abundance did generally increase with the gradient of retained habitat structure. Mean species richness and diversity were similar among treatment sites but did show an increasing gradient with structural compexity. For H2, mean abundance, reproduction, and recruitment of M. gapperi were higher in GTR and windrow sites than those without retained structures. There was a positive relationship between mean abundance of M. gapperi and total volume of woody debris across treatments. This study is the first investigation of the responses of forest-floor small mammals to an increasing gradient of retained habitat structure via GTR and woody debris on clearcuts. Our assessment of a combination of these two interventions suggested a potentially strong additive effect that could be cautiously extrapolated across three forest ecological zones. With the advent of low levels of GTR on clearcuts, woody debris structures should help provide some habitat to conserve forest mammals on harvest openings.
  • Sullivan, Department of Forest and Conservation Sciences, Faculty of Forestry, University of BC, 2424 Main Mall, Vancouver, BC, Canada V6T 1Z4 E-mail: tom.sullivan@ubc.ca (email)
  • Sullivan, Applied Mammal Research Institute, 11010 Mitchell Avenue, Summerland, BC, Canada V0H 1Z8 E-mail: dru.sullivan@appliedmammal.com
article id 1217, category Research article
Ulrich J. Wolfsmayr, Peter Rauch. (2014). Primary forest fuel supply chain: assessing barriers and drivers for the modal shift from truck to train. Silva Fennica vol. 48 no. 5 article id 1217. https://doi.org/10.14214/sf.1217
Keywords: biomass; multimodal transport; train transport; quality function deployment; house of quality
Highlights: For combined heat and power plants in Austria procuring forest fuels, the most competitive transport mode is road transport using walking-floor trucks; The main barriers for a modal shift are the plant managers’ negative experiences with the railroad; Rail transport has its benefits, when high volumes are needed and transport distances are long.
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Multimodal primary forest fuel (PFF) transport using the railroad for main haulage has been quite uncommon to present, although it could provide considerable advantages in terms of economical, ecological and social parameters. Accordingly, the objective of this paper is to assess barriers and drivers for the modal shift from truck to train. As methodological tool, we are using the concept of Quality Function Deployment (QFD) with the House of Quality (HoQ) – an approach that has not been used in forest management so far. As the most important barriers for the modal shift from truck to train in PFF transport in Austria, the following were identified: (i) bioenergy plant managers have a negative opinion and negative experience regarding the railroad in terms of high prices, a lot of bureaucracy, etc.; (ii) absence of rail sidings or relatively short rail sidings not suitable for block trains; and (iii) unwillingness to invest in new supply or unloading systems. On the contrary, the most important drivers for a modal shift are: (i) multimodal PFF supply chains using trains can provide high volumes; (ii) increasing catchment areas for larger CHP plants result in increasing transport distances; and (iii) rail transport has less negative environmental and social impact than road transport.
  • Wolfsmayr, University of Natural Resources and Life Sciences, Vienna, Gregor Mendel Straße 33, A-1180 Wien, Austria E-mail: ulrich.wolfsmayr@boku.ac.at (email)
  • Rauch, University of Natural Resources and Life Sciences, Vienna, Gregor Mendel Straße 33, A-1180 Wien, Austria E-mail: peter.rauch@boku.ac.at
article id 1192, category Research article
Hannu Hökkä, Hanna Mäkelä. (2015). Post-harvest height growth of Norway spruce seedlings in northern Finland peatland forest canopy gaps and comparison to partial and complete canopy removals and plantations. Silva Fennica vol. 48 no. 5 article id 1192. https://doi.org/10.14214/sf.1192
Keywords: Picea abies; height growth; peatlands; growth models; advance growth; canopy gap
Highlights: Norway spruce seedlings’ height growth recovered within four years after the cutting of canopy gaps; Growth was linearly related to tree height, being highest for tallest seedlings; Seedlings in the 20 m diameter gap and in the central and northern parts in the 15 m diameter gap showed the best growth; In gaps early height growth was 60% of that in peatland spruce plantations but 2–3 times higher than in uneven-aged cut forests.
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Recent studies have shown the establishment of Norway spruce (Picea abies (L.) Karst.) to be successful in small canopy gaps cut in drained spruce mire stands in northern Finland. The aim of this study was to quantify seedling height growth in gaps and compare it to that observed in other canopy cuttings and plantations. We sampled spruce crop seedlings (maximum density ca. 3000 ha–1) in the spring of 2013 in a field experiment in which canopy gaps of 10, 15 and 20 m in diameter had been cut in winter 2004. The total seedling height in 2013 and the length of annual shoots over the past five years (2012–2008) were recorded in the survey. Seedling height varied from 20 cm to 2.7 m, with an average of 65 cm. The average annual height growth was 7.1 cm. A mixed linear model analysis was carried out to investigate seedling height growth variation. Seedling height was linearly and positively related to growth. Height growth started to increase in the fifth growing season after cutting. Seedling height growth in the 20 m gap was slightly better than in the smaller ones. In the 15 m gap, both the centrally located seedlings and those located at the northern edge grew best. In the 20 m gap, southerly located seedlings grew more slowly than seedlings in all other locations. The average seedling height growth in this study was about 60% of that in peatland plantations, but comparable to that in mineral soil gaps, and 2–3 times higher than in uneven-age cut stands.
  • Hökkä, Finnish Forest Research Institute, Rovaniemi Unit, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: hannu.hokka@metla.fi (email)
  • Mäkelä, Lapland University of Applied Sciences, Jokiväylä 11 C, FI-96300 Rovaniemi, Finland E-mail: hanna.makela@edu.ramk.fi
article id 1190, category Research article
Jaakko Repola, Kristina Ahnlund Ulvcrona. (2014). Modelling biomass of young and dense Scots pine (Pinus sylvestris L.) dominated mixed forests in northern Sweden. Silva Fennica vol. 48 no. 5 article id 1190. https://doi.org/10.14214/sf.1190
Keywords: boreal forests; pine; tree biomass; biomass models; unmanaged stands
Highlights: The biomass allocation to tree components is different in unmanaged and managed young stands; Higher foliage biomass and lower stem and branch biomass were detected in the unmanaged stands; Models for trees from young and dense stands provide better estimates of biomass in such stands than those based on data from managed stands.
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Biomass models for the biomass of above-ground tree components of Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies [L.] Karst.) and birch (Betula pendula Roth and Betula pubescens Ehrh.) in young dense Scots pine dominated forest stands in northern Sweden were constructed. Destructive above-ground biomass sampling was conducted in naturally generated young, dense, Scots pine dominated mixed stands. Three sampling campaigns were undertaken, the first in 1997 and 1998. The second was six years later (2003), and the last 13 years after the first (2010). In total, 280 trees (126 Scots pine, 68 Norway spruce and 86 birches) were sampled from six different stands in northern Sweden. The sampled trees’ diameter at breast height (dbh) was in the range 1–22 cm (Scots pine), 1–21 cm (Norway spruce) and 1–11 cm (birch). Biomass predictions were tested using our models and the widely used biomass models originally constructed for managed stands. The results showed that the biomass allocation to tree components is different in unmanaged and managed young stands; higher foliage biomass and lower stem and branch biomass were detected in the unmanaged stands. The overall conclusion is that the biomass models for managed stands did not produce satisfactory biomass estimates in unthinned, dense, young stands.
  • Repola, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: jaakko.repola@metla.fi (email)
  • Ahnlund Ulvcrona, SLU, Forest Biomaterials and Technology, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: kristina.ulvcrona@slu.se
article id 1140, category Research article
Miguel Fabra Crespo, Olli Saastamoinen, Jukka Matero, Hannes Mäntyranta. (2015). Perceptions and realities: public opinion on forests and forestry in Finland, 1993–2012. Silva Fennica vol. 48 no. 5 article id 1140. https://doi.org/10.14214/sf.1140
Keywords: values; correlation; attitude; reflexive groups; social evolution
Highlights: Analysis of the longest-running survey in the world on public perceptions of forestry; Comparison of perceptions with realities in forestry in Finland; The role of the forests and their management is well regarded by the Finns; More protection of the forests and better performance by the wood industry is demanded.
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The perception of the Finns about forests and forestry has been tracked over a period of more than 15 years. The results of this survey constitute the longest sequence of data of this type at the national level anywhere in the world. The people’s perception of reality represents a factor that influences decisions about policy. For this reason, it deserves monitoring and analysis. Forests in Finland are highly meaningful to the people, who are generally well informed and link their opinions to the facts that they are able to observe. The variability of the responses over the years of the survey is not significant. Silviculture and forest management are perceived as good by most Finns. Finns are aware that more forest grows than is harvested, and they also know that some raw material is still imported. However, they demand that more forest be protected. Finns are aware that their forest industry is not performing well at the international level. They also demand an increased wood supply for building construction. Forest harvesting is viewed as a source of employment and welfare.
  • Fabra Crespo, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: mfabracrespo@yahoo.com (email)
  • Saastamoinen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: olli.saastamoinen@uef.fi
  • Matero, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: materojukka@gmail.com
  • Mäntyranta, Finnish Forest Association, Salomonkatu 17 A, FI-00100 Helsinki, Finland E-mail: hannes.mantyranta@smy.fi
article id 1135, category Research article
Matts Lindbladh, Per-Ola Hedwall, Ida Wallin, Annika M. Felton, Henrik Böhlenius, Adam Felton. (2014). Short-rotation bioenergy stands as an alternative to spruce plantations: implications for bird biodiversity. Silva Fennica vol. 48 no. 5 article id 1135. https://doi.org/10.14214/sf.1135
Keywords: Norway spruce; Picea abies; hybrid aspen; Sweden; Populus × wettsteinii; production forests
Highlights: There is a gap in knowledge regarding the biodiversity implications of replacing production forests with bioenergy stands; We compared the avian biodiversity of early rotation hybrid aspen stands and spruce plantations, the latter being the dominant production forest type in southern Sweden; Our results indicate that young hybrid aspen stands can support relatively diverse and distinctive bird communities.
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Global efforts to decrease dependence on fossil fuels have increased interest in bioenergy production. One source of bioenergy is fast growing deciduous tree species, such as hybrid aspen (Populus × wettsteinii Hämet-Ahti). The majority of research on hybrid aspen which assesses biodiversity implications, has however primarily focused on agricultural lands as the reference condition. This has resulted in a substantial gap in our knowledge regarding the biodiversity implications of replacing production forest types with hybrid aspen, a form of reforestation taking place in northern Europe. In this study we address this knowledge gap by comparing the avian biodiversity of young hybrid aspen and spruce (Picea abies L.) plantations of similar age, the latter being the most prevalent forestry alternative in in southern Sweden. We found that hybrid aspen stands had higher bird species richness and abundance as well as a distinct community composition compared to the spruce stands. We suggest that the most likely driver was the greater structural and tree species complexity in the aspen stands, provided for by the fenced exclusion of ungulates from the regenerating hybrid aspen stands. Our results indicate that at least during early stages of regeneration, and in comparison to the dominating production forest type in the region, hybrid aspen stands can support relatively high levels of bird diversity, and a bird species composition more closely associated with broadleaf habitat types requiring restoration in this region.
  • Lindbladh, Southern Swedish Forest Research Centre, SLU – Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: matts.lindbladh@slu.se (email)
  • Hedwall, Southern Swedish Forest Research Centre, SLU – Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: per-ola.hedwall@slu.se
  • Wallin, Southern Swedish Forest Research Centre, SLU – Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: ida.wallin@slu.se
  • Felton, Southern Swedish Forest Research Centre, SLU – Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: annika.felton@slu.se
  • Böhlenius, Southern Swedish Forest Research Centre, SLU – Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: henrik.bohlenius@slu.se
  • Felton, Southern Swedish Forest Research Centre, SLU – Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: adam.felton@slu.se
article id 1124, category Research article
Āris Jansons, Mārtiņš Zeps, Juris Rieksts-Riekstiņš, Roberts Matisons, Oskars Krišāns. (2014). Height increment of hybrid aspen Populus tremuloides x P. tremula as a function of weather conditions in central part of Latvia. Silva Fennica vol. 48 no. 5 article id 1124. https://doi.org/10.14214/sf.1124
Keywords: height growth; energy wood; tree growth; temperature; intra-annual dynamics; plantation silviculture
Highlights: Intra-annual height growth of hybrid aspen was monitored; Clones with early leaf flushing dates showed faster height growth; Height growth was generally controlled by temperature; Fast-growing hybrids were more robust to weather conditions than slow-growing ones; Potential evapotranspiration (moisture regime) negatively affected height growth of clones with delayed phenology.
Abstract | Full text in HTML | Full text in PDF | Author Info
Height growth of young hybrid aspen (Populus tremula L. × P. tremuloides Michx.) was studied in relation to weather conditions. Height of clones with different leaf flushing phenology (early, intermediate and late) was monitored during the growing periods of 2010 and 2011 in a plantation established on former agricultural land. Mean daily height increment (HI) was calculated. Multiple linear regression was used to determine which weather factors (variables) had significant effect on HI. Mean seasonal height growth (mean seasonal HI) between clones (groups) was compared by ANOVA. In both years, HI was significantly higher for clones with early and intermediate leaf flushing compared to clones with late leaf flushing. The effect of weather factors also differed between clones according to their leaf flushing phenology; it was the weakest for HI of clones with early leaf flushing compared to clones with intermediate and late leaf flushing. Mean temperature was the main factor, which positively affected HI of all clones, suggesting that warmer climate might be beneficial for height growth of young hybrid aspen in Latvia. Nevertheless, significant negative relationship between HI and potential evapotranspiration (PET) was observed for clones with delayed leaf flushing, suggesting negative effect of increasing variability of precipitation on growth. Thus, the differences in height growth intensity might be related to growth sensitivity to weather conditions. On the other hand, such differences in height growth between clones might be caused by competition (i.e. with herbs), as trees with early leaf flushing might conquer more resources and become more robust against the environmental fluctuation.
  • Jansons, LSFRI „SILAVA”, Rigas Str. 111, Salaspils, Latvia, LV2169 E-mail: aris.jansons@silava.lv
  • Zeps, LSFRI „SILAVA”, Rigas Str. 111, Salaspils, Latvia, LV2169 E-mail: martins.zeps@silava.lv
  • Rieksts-Riekstiņš, LSFRI „SILAVA”, Rigas Str. 111, Salaspils, Latvia, LV2169 E-mail: Juris.Riekstins@silava.lv
  • Matisons, LSFRI „SILAVA”, Rigas Str. 111, Salaspils, Latvia, LV2169 E-mail: robism@inbox.lv (email)
  • Krišāns, LSFRI „SILAVA”, Rigas Str. 111, Salaspils, Latvia, LV2169 E-mail: oskars.krisans@silava.lv
article id 1211, category Research article
Xiao Chen, Deborah Page-Dumroese, Ruiheng Lv, Weiwei Wang, Guolei Li, Yong Liu. (2014). Interaction of initial litter quality and thinning intensity on litter decomposition rate, nitrogen accumulation and release in a pine plantation. Silva Fennica vol. 48 no. 4 article id 1211. https://doi.org/10.14214/sf.1211
Keywords: litter quality; litter decomposition; nitrogen cycling; thinning intensity; Pinus tabulaeformis
Highlights: Litter quality and thinning showed an interaction on one year litter decomposition rates, N accumulation, and net N release; N accumulated until the underlying critical acid-unhydrolyzable residue to nitrogen ratio (approximately 57–69) was met; Increased N concentration in litter and thinning intensity induced rapid litter decomposition and N cycling in coniferous plantation with a slow decomposition rate.
Abstract | Full text in HTML | Full text in PDF | Author Info
Thinning alters litter quality and microclimate under forests. Both of these two changes after thinning induce alterations of litter decomposition rates and nutrient cycling. However, a possible interaction between these two changes remains unclear. We placed two types of litter (LN, low N concentration litter; HN, high N concentration litter) in a Chinese pine (Pinus tabulaeformis Carrière) plantation under four thinning treatments to test the impacts of litter quality, thinning or their combination on decomposition rate and N cycling. In our study, N was accumulated to approach an underlying critical acid-unhydrolyzable residue to nitrogen ratio (approximately 57–69) in litter. Moreover, an interaction between litter quality and thinning on decomposition rates, N accumulation and net release did exist. On one hand, one year decomposition rate of LN was elevated after thinning while that of HN remained the same or even lower (under light thinning); N accumulation of LN declined with light thinning and was restored with the increase of thinning intensity whereas that of HN did not decline with thinning and increased under heavy thinning; Net N release from LN was only found in light and heavy thinning while that from HN was found in all treatments, moreover net N release from LN and HN were both elevated under heavy thinning. On the other hand, HN decomposed faster, accumulated less and released more N than LN did under all treatments. Generally, high N concentration in litter and high-intensity thinning can lead to rapid litter decomposition and N cycling in coniferous plantations.
  • Chen, Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing, 100083, China E-mail: chenxiao_0123@126.com
  • Page-Dumroese, U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, 1221 South Main Street, Moscow, ID 83843, USA E-mail: ddumroese@fs.fed.us
  • Lv, College of Plant Science and Technology, Tarim University, Alar Xinjiang, 843300, China E-mail: lvrh514723@126.com
  • Wang, Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing, 100083, China E-mail: fuyuerdejia@126.com
  • Li, Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing, 100083, China E-mail: glli226@163.com
  • Liu, Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing, 100083, China E-mail: lyong@bjfu.edu.cn (email)
article id 1207, category Research article
Olli-Pekka Tikkanen, Irina A. Chernyakova. (2014). Past human population history affects current forest landscape structure of Vodlozero National Park, Northwest Russia. Silva Fennica vol. 48 no. 4 article id 1207. https://doi.org/10.14214/sf.1207
Keywords: boreal forests; forest fires; recovery; forest structure; landscape change
Highlights: In large scale (0–20 km), the proportion of spruce in forest stands was positively affected by distance from old villages. This relationship was non-linear with a threshold distance of 15 km; In small scale (0–5 km), old villages affected tree species composition and age structure of forests. Effect on age structure was the strongest on stands growing on mineral soils.
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The information about location and size of past human settlements can give new insights into the analysis of landscape structures. Vodlozero National Park (NP) is one the largest strictly protected areas in Northwestern Europe. We mapped the location of historic villages, which were abandoned in 1958, and studied the effect of past human activity in the forest landscape in two different scales using forest survey data. We assessed the possible change in age, volume and tree species composition from the edge of open fields up to the distance of 5 km from villages. We made a larger landscape analysis using a grid of forest stands covering the whole northern part of the NP. The past human activity was clearly visible in the present forest landscape. Distance from villages affected age, volume and tree species composition of the forest stands. This effect was the strongest within the first two kilometers from the villages. At the level of whole northern NP, the proportion of spruce markedly increased after approximately 15 km from the nearest old village. The changes in the forests surrounding the villages were most likely the result of the intensive use of wood for different commodities needed in households and farming, in addition to short rotation slash and burn agriculture. If the occurrence of forest fires was more frequent closer to villages than in more remote areas, it can well explain the observed pattern in the abundance of spruce in the larger landscape that is less tolerant to fire than pine.
  • Tikkanen, Department of Biology, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland (Current: School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland) & Interdisciplinary Research and Educational Center of Cross-border Communication CARELICA, Institute of History, Political and Social Sciences, Petrozavodsk State University, 33 Lenin Prospectus, 185910 Petrozavodsk, Republic of Karelia, Russia E-mail: Olli-Pekka.Tikkanen@uef.fi (email)
  • Chernyakova, Interdisciplinary Research and Educational Center of Cross-border Communication CARELICA, Institute of History, Political and Social Sciences, Petrozavodsk State University, 33 Lenin Prospectus, 185910 Petrozavodsk, Republic of Karelia, Russia E-mail: irina.chernyakova@onego.ru
article id 1188, category Research article
Kristina Wallertz, Henrik Nordenhem, Göran Nordlander. (2014). Damage by the pine weevil Hylobius abietis to seedlings of two native and five introduced tree species in Sweden. Silva Fennica vol. 48 no. 4 article id 1188. https://doi.org/10.14214/sf.1188
Keywords: forest regeneration; conifer seedlings; defence; feeding damage; forest plantation; resistance
Highlights: Both native and introduced confer species in Sweden can be highly susceptible to damage by the pine weevil; Douglas fir and Sitka spruce were generally the most damaged among six studied conifer species; The results highlight some of the risks in establishing exotic tree species for forest production.
Abstract | Full text in HTML | Full text in PDF | Author Info
There is increasing interest in using introduced species in Swedish forestry in response to climate change, but it is important to assess their resistance to native pests. Thus, we compared the extent of pine weevil feeding on two dominant native conifers, Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.), the non-host deciduous broadleaf hybrid aspen (Populus × wettsteinii Hämet-Ahti) and four introduced conifers: Douglas fir (Pseudotsuga menziesii (Mirb.) Franco), hybrid larch (Larix × marschlinsii Coaz), Sitka spruce (Picea sitchensis (Bong.) Carriére) and lodgepole pine (Pinus contorta Douglas ex Loudon). The extent of feeding damage on seedlings and its effect on their vitality were examined in a field study in south-central Sweden and a laboratory experiment, which gave largely consistent results. Generally, the species most heavily attacked by the pine weevil, in both experiments, were Douglas fir and Sitka spruce. In the field experiment pine weevils killed or severely damaged significantly higher proportions of Douglas fir and Sitka spruce seedlings (60%) than any other species except Norway spruce (49%). Among conifer seedlings the proportions of killed or severely damaged seedlings were lowest for Scots pine and hybrid larch (27%) and Lodgepole pine (36%). The results indicate that most conifer species planted on young clear-cuttings in Sweden need some kind of pine weevil protection, and the possibility that introducing new tree species might increase damage caused by pests must be considered. For instance, widespread use of hybrid aspen could reduce damage by pine weevils, but increase damage by other, untested pests or pathogens.
  • Wallertz, Unit for Field-based Forest Research, Swedish University of Agricultural Sciences (SLU), Asa Research Station, SE-36030 Lammhult, Sweden E-mail: kristina.wallertz@slu.se (email)
  • Nordenhem, Department of Ecology, Swedish University of Agricultural Sciences (SLU), P.O. Box 7044, SE-75007 Uppsala, Sweden E-mail: henrik.nordenhem@slu.se
  • Nordlander, Department of Ecology, Swedish University of Agricultural Sciences (SLU), P.O. Box 7044, SE-75007 Uppsala, Sweden E-mail: goran.nordlander@slu.se
article id 1176, category Research article
Batoul Al-Hawija, Viktoria Wagner, Monika Partzsch, Isabell Hensen. (2014). Germination differences between natural and afforested populations of Pinus brutia and Cupressus sempervirens. Silva Fennica vol. 48 no. 4 article id 1176. https://doi.org/10.14214/sf.1176
Keywords: nursery; silviculture; drought stress; cold stratification; local adaptation; salt stress; Syria
Highlights: Silvicultural practices of raising and outplanting seedlings yielded contrasting outcomes in our species; Afforested Pinus brutia populations acquired ability to tolerate drought stress at intermediate and hot temperatures compared to natural populations, which may indicate local adaptation; Natural Cupressus sempervirens populations showed higher salt-tolerance than afforested populations; Seed germination was optimal under intermediate temperatures and deionized water for both species.
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In afforestation, silvicultural processes of raising and planting seedlings under certain conditions can yield contrasting outcomes in tree stock performance. Moderate nursery conditions may select against stress tolerance whereas planting seedlings in stressful environments at afforestation sites may select for higher stress tolerance compared to natural populations. We compared germination performance between natural and afforested populations of Pinus brutia Ten. subsp. brutia and Cupressus sempervirens L. var. horizontalis (Mill.) under differing stress treatments. Seeds were collected from both natural stands and from afforested populations outside the natural distribution range, in Syria. Cold, intermediate and hot temperature regimes were simulated (8/4 °C, 20/10 °C and 32/20 °C) along with cold stratification, drought stress (–0.2 and –0.4 MPa), salt stress (50 and 100 mMol l–1), and deionized water (control) conditions. In addition, we tested the effects of seed weight and climatic conditions on seed germination. In general, intermediate temperatures were optimal for both population types. Afforested P. brutia populations outperformed natural ones under drought stress levels at hot and/or intermediate temperatures. Conversely, in C. sempervirens, cold stratification at all temperatures and higher salt stress at intermediate temperatures significantly decreased germination in afforested populations. Seed weight did not significantly affect germination percentages, which were however significantly negatively related to annual precipitation in P. brutia, and to annual temperature in C. sempervirens. We infer that silvicultural processes led to divergent outcomes in our species: local adaptation to drought stress and hot temperatures in afforested P. brutia populations and lower salt-stress tolerance in C. sempervirens.
  • Al-Hawija, Martin-Luther-University Halle-Wittenberg, Institute of Biology/Geobotany and Botanical Garden, Am Kirchtor 1, D-06108 Halle/Saale, Germany E-mail: batoulh@gmail.com (email)
  • Wagner, Department of Botany and Zoology, Masaryk University, Kotlářská 2, CZ-611 37 Brno, Czech Republic E-mail: wagner@sci.muni.cz
  • Partzsch, Martin-Luther-University Halle-Wittenberg, Institute of Biology/Geobotany and Botanical Garden, Am Kirchtor 1, D-06108 Halle/Saale, Germany E-mail: monika.partzsch@botanik.uni-halle.de
  • Hensen, Martin-Luther-University Halle-Wittenberg, Institute of Biology/Geobotany and Botanical Garden, Am Kirchtor 1, D-06108 Halle/Saale, Germany & German Centre for Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig, Deutscher Platz 5e, 04103 Leipzig, Germany E-mail: isabell.hensen@botanik.uni-halle.de
article id 1159, category Research article
Grzegorz Szewczyk, Janusz Michał Sowa, Włodzimierz Grzebieniowski, Mariusz Kormanek, Dariusz Kulak, Arkadiusz Stańczykiewicz. (2014). Sequencing of harvester work during standard cuttings and in areas with windbreaks. Silva Fennica vol. 48 no. 4 article id 1159. https://doi.org/10.14214/sf.1159
Keywords: time consumption; time series; timber harvesting; harvesters; post-disaster stands
Highlights: In standard cutting stands and thinning areas with windbreaks there occurred three-activity operational cycles. In mature stands with windbreaks the occurrence of stable sequences supplemented with five-activity cycles was noted. Consequently, the operational time in post-disaster thinning stands should be increased by 55% whereas in mature stands it should be 30% longer in comparison with standard stands.
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The aim of the study was to characterize repetitive cycles of harvester operation. The study was conducted in thinning, mature and post-disaster pine stands. The sequences of the activities characteristic of harvester operation were described as time series. In order to detect the cyclic variable structure of the analysed time series, the methodology of the single spectrum Fourier analysis was applied. In standard stands, post-disaster late-thinning stands and mature stands, the existence of stable operational cycles with the length of three activities was discovered while in post-disaster mature stands additional five-activity operational phases were noted. Described in this way, the lengths of the operational cycles of harvesters working in post-disaster areas were higher by about 55% and 30% respectively, as compared to standard thinning and mature stands.
  • Szewczyk, University of Agriculture in Krakow, Faculty of Forestry, Institute of Forest Utilization and Forest Technology, Department of Forest and Wood Utilization, Al. 29-Listopada 46, 31-425 Krakow, Poland E-mail: rlszewcz@cyf-kr.edu.pl (email)
  • Sowa, University of Agriculture in Krakow, Faculty of Forestry, Institute of Forest Utilization and Forest Technology, Department of Forest and Wood Utilization, Al. 29-Listopada 46, 31-425 Krakow, Poland E-mail: rlsowa@cyf-kr.edu.pl
  • Grzebieniowski, University of Agriculture in Krakow, Faculty of Forestry, Institute of Forest Utilization and Forest Technology, Department of Forest and Wood Utilization, Al. 29-Listopada 46, 31-425 Krakow, Poland E-mail: wrzoswj@interia.pl
  • Kormanek, University of Agriculture in Krakow, Faculty of Forestry, Institute of Forest Utilization and Forest Technology, Department of Forest Work Mechanisation, Al. 29-Listopada 46, 31-425 Krakow, Poland E-mail: rlkorma@cyf-kr.edu.pl
  • Kulak, University of Agriculture in Krakow, Faculty of Forestry, Institute of Forest Utilization and Forest Technology, Department of Forest and Wood Utilization, Al. 29-Listopada 46, 31-425 Krakow, Poland E-mail: rlkulak@cyf-kr.edu.pl
  • Stańczykiewicz, University of Agriculture in Krakow, Faculty of Forestry, Institute of Forest Utilization and Forest Technology, Department of Forest and Wood Utilization, Al. 29-Listopada 46, 31-425 Krakow, Poland E-mail: rlstancz@cyf-kr.edu.pl
article id 1119, category Research article
Beata Woziwoda, Agnieszka Parzych, Dominik Kopeć. (2014). Species diversity, biomass accumulation and carbon sequestration in the understorey of post-agricultural Scots pine forests. Silva Fennica vol. 48 no. 4 article id 1119. https://doi.org/10.14214/sf.1119
Keywords: biodiversity; Pinus sylvestris plantation; overstorey-understorey interaction; biotic homogenization; Leucobryo-Pinetum community
Highlights: Understorey plant species diversity significantly increases with the age of a Scots pine stand; Biomass of mosses decreases by a quarter, while biomass of herbs increases several times; Total understorey’s carbon stock increases over three times. The highest amount of carbon is accumulated in understorey species like Vaccinium myrtillus and Dicranum polysetum; The growing proportion of vascular plants in the understorey biomass results in an increase in the understorey C/N ratio.
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The purpose of this study was to examine how the age of a stand of post-agricultural Scots pine forests affects the species composition, biomass and the carbon stock of the forest understorey. The community structure and species composition were studied in 75 plots (100 m2 in size), the amount of biomass, organic carbon and total nitrogen were analysed in 75 subplots (1/16 m2 in size). The plots were located in 21 plantations with the stand age of 41–60, 61–80 and over 80-years. Results show that the understorey species diversity increased with the increasing age of Scots pine stands, and the structure and species composition of secondary forests (although managed for timber production) became similar to the fresh pine forest of the European temperate region (Leucobryo-Pinetum community). Despite the increasing species diversity, however, only six understorey vascular and moss species played an important role in the biomass accumulation and C sequestration. Due to the differences in the dominant species composition, the total amount of understorey biomass significantly differed among the forest stands. The mean moss biomass ranged from 3046 kg ha–1 in 41–60-year-old stands, trough 2686 kg ha–1 in 61–80-year-old stands to 2273 kg ha–1 in over 80-year-old stands, and the mean understorey vascular plant biomass amounted to 2 kg ha–1, 1924 kg ha–1 and 3508 kg ha–1, respectively. The concentration of organic C varied considerably between species; it was the highest in Vaccinium myrtillus (50.6%) and in Dicranum polysetum (49.5%). The total mass of C was nearly 800 kg ha–1 in the youngest forests, in the subsequent age series it was two times higher and 3.5 times higher in the oldest ones. Differences in the species composition and in the C/N ratio in different species (generally higher for vascular plants and lower for mosses) were expressed in an increase in the understorey C/N ratio, which was 39.5, 46.6 and 48.6, respectively.
  • Woziwoda, Department of Geobotany and Plant Ecology, Faculty of Biology and Environmental Protection, University of Lodz, Banacha 12/16, 90-237 Lodz, Poland E-mail: woziwoda@biol.uni.lodz.pl (email)
  • Parzych, Environmental Chemistry Research Unit, Institute of Biology and Environmental Protection, Pomeranian University in Słupsk, Arciszewskiego 22b, 76-200 Słupsk, Poland E-mail: parzycha1@op.pl
  • Kopeć, Department of Geobotany and Plant Ecology, Faculty of Biology and Environmental Protection, University of Lodz, Banacha 12/16, 90-237 Lodz, Poland E-mail: domin@biol.uni.lodz.pl
article id 1107, category Research article
Arvo Tullus, Arne Sellin, Priit Kupper, Reimo Lutter, Linnar Pärn, Anna K. Jasinska, Meeli Alber, Maarja Kukk, Tea Tullus, Hardi Tullus, Krista Lõhmus, Anu Sõber. (2014). Increasing air humidity – a climate trend predicted for northern latitudes – alters the chemical composition of stemwood in silver birch and hybrid aspen. Silva Fennica vol. 48 no. 4 article id 1107. https://doi.org/10.14214/sf.1107
Keywords: climate change; Betula; Populus; macronutrients; atmospheric humidity; wood characteristics; structural carbohydrates
Highlights: Hybrid aspen and silver birch trees grew more slowly under increased air humidity conditions and had higher concentrations of N and P and a lower K to N ratio in stemwood; Minor species-specific changes were detected in stemwood concentrations of cellulose and hemicellulose; Density, calorific value and concentrations of lignin and ash in stemwood were not affected by elevated humidity.
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We studied the physicochemical properties of stemwood in saplings of silver birch (Betula pendula Roth) and hybrid aspen (Populus tremula L. × P. tremuloides Michx.), grown for four years under artificially elevated relative air humidity (on average by 7%) in field conditions, using the Free Air Humidity Manipulation (FAHM) research facility in Estonia. Altogether 91 sample trees from three experimental plots with manipulated air humidity and from three control plots were cut in the dormant season and sampled for the analysis of cellulose, hemicellulose, acid detergent lignin, macronutrients (N, P, K), ash content, density, and calorific value of wood. The analysed trees grew significantly more slowly under elevated humidity conditions, with a more pronounced effect on aspens. Significantly higher concentrations of N and P were observed in the stemwood of both aspens and birches grown under elevated humidity. This could be the result of a change in the content of living parenchyma cells and/or enhanced retranslocation of nutrients into wood parenchyma. Additionally, humidification resulted in a significantly higher concentration of cellulose and a lower concentration of hemicellulose in aspen stemwood, and in significantly lower concentrations of cellulose and K in birch stemwood. Elevated humidity did not affect lignin concentration, ash content, basic density and calorific value of stemwood. Results from the FAHM experiment suggest that the increasing air humidity accompanying global warming at northern latitudes will affect the growth and functioning of deciduous trees and forests, with obvious consequences also for forest management and industry.
  • Tullus, Department of Botany, Institute of Ecology and Earth Sciences, Faculty of Science and Technology, University of Tartu, Lai 40, Tartu 51005, Estonia E-mail: arvo.tullus@ut.ee (email)
  • Sellin, Department of Botany, Institute of Ecology and Earth Sciences, Faculty of Science and Technology, University of Tartu, Lai 40, Tartu 51005, Estonia E-mail: arne.sellin@ut.ee
  • Kupper, Department of Botany, Institute of Ecology and Earth Sciences, Faculty of Science and Technology, University of Tartu, Lai 40, Tartu 51005, Estonia E-mail: priit.kupper@ut.ee
  • Lutter, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Kreutzwaldi 5, Tartu 51014, Estonia E-mail: reimo.lutter@emu.ee
  • Pärn, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Kreutzwaldi 5, Tartu 51014, Estonia E-mail: linnar.parn@emu.ee
  • Jasinska, Department of Botany, Institute of Ecology and Earth Sciences, Faculty of Science and Technology, University of Tartu, Lai 40, Tartu 51005, Estonia & Institute of Dendrology, Polish Academy of Sciences, Parkowa 5, 62-035 Kórnik, Poland E-mail: jasiak9@wp.pl
  • Alber, Department of Botany, Institute of Ecology and Earth Sciences, Faculty of Science and Technology, University of Tartu, Lai 40, Tartu 51005, Estonia E-mail: meeli.alber@ut.ee
  • Kukk, Department of Botany, Institute of Ecology and Earth Sciences, Faculty of Science and Technology, University of Tartu, Lai 40, Tartu 51005, Estonia E-mail: maarja.kukk@ut.ee
  • Tullus, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Kreutzwaldi 5, Tartu 51014, Estonia E-mail: tea.tullus@emu.ee
  • Tullus, Institute of Forestry and Rural Engineering, Estonian University of Life Sciences, Kreutzwaldi 5, Tartu 51014, Estonia E-mail: hardi.tullus@emu.ee
  • Lõhmus, Department of Botany, Institute of Ecology and Earth Sciences, Faculty of Science and Technology, University of Tartu, Lai 40, Tartu 51005, Estonia E-mail: krista.lohmus@ut.ee
  • Sõber, Department of Botany, Institute of Ecology and Earth Sciences, Faculty of Science and Technology, University of Tartu, Lai 40, Tartu 51005, Estonia E-mail: anu.sober@ut.ee
article id 1161, category Research article
Tiina Laine, Veli-Matti Saarinen. (2014). Comparative study of the Risutec Automatic Plant Container (APC) and Bracke planting devices. Silva Fennica vol. 48 no. 3 article id 1161. https://doi.org/10.14214/sf.1161
Keywords: productivity; regeneration; cost-efficiency; planting cost; mechanized tree planting
Highlights: As currently designed, the prototype Risutec APC fitted with an automatic feeding system offers no significant advantage over the Bracke planting device in terms of planting productivity or quality; Cost estimates suggest that an idealized automated feeding system could increase productivity and decrease unit costs.
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The productivity of mechanized planting could be increased by minimizing the time spent manually reloading seedling cassettes. This study compared the work-time distribution, productivity and quality of the prototype Risutec APC fitted with an automatic feeding system and the commonly-used and manually-loaded Bracke P11.a. An approach of comparative time study was employed that compared performance of two operators using both machines in four sites where slash and stumps had been removed. Operating costs were estimated and compared for these two machines and an idealized machine with automatic feeding system (referred as AUT). AUT was assumed to be similar to the Bracke planting machine with the only difference being in automatic feeding. Productivity of the Risutec APC (196 seedlings per productive work hour [pl PWh0–1]) was lower than that of Bracke (244 pl PWh0–1), making the unit cost 35.7% higher. A large portion (17.6%) of the productive work time of Risutec APC was interrupted by malfunctions, so it cannot be considered robust and reliable yet. Quality of the planting work was reasonable for both machines. The results suggest that an idealized AUT could increase planting capacity (hectares per year [ha yr–1]) by 15.4% and lower the unit cost (Euro per seedling [€ pl–1]) by 4.7% compared to today’s machines. The importance of an automated feeding system increases with planting efficiency because relatively more time is spent reloading seedlings. Proper automatic feeding system could offer a cost-effective solution and could enhance productivity, but the Risutec APC has yet to meet the technical and economic standards required to be competitive.
  • Laine, Finnish Forest Research Institute (METLA), Suonenjoki Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: tiina.laine@metla.fi (email)
  • Saarinen, Finnish Forest Research Institute (METLA), Suonenjoki Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: veli-matti.saarinen@metla.fi
article id 1153, category Research article
Anu Akujärvi, Ville Hallikainen, Mikko Hyppönen, Eero Mattila, Kari Mikkola, Pasi Rautio. (2014). Effects of reindeer grazing and forestry on ground lichens in Finnish Lapland. Silva Fennica vol. 48 no. 3 article id 1153. https://doi.org/10.14214/sf.1153
Keywords: forest management; timber harvesting; reindeer herding; pastures; lichen cover; lichen biomass
Highlights: Both reindeer grazing and forestry affect the cover and biomass of reindeer lichens; Reindeer grazing has bigger impact than forestry; The lichen cover was about five-fold and the biomass about fifteen-fold in the ungrazed (fenced) sites than in the grazed ones; The decrease of not only the biomass, but also the cover of lichens, is alarming.
Abstract | Full text in HTML | Full text in PDF | Author Info
Reindeer husbandry and forestry are practiced in the same areas in northern Fennoscandia. Reindeer pastures have largely deteriorated. We aimed to quantify the separate and combined effects of reindeer grazing and forestry on the amount of ground lichens. To do this, we mapped and inventoried all larger enclosures (49) in Finnish Lapland where forest management practices were similar in both sides of the fence. The average time since fencing was 43 years. We recorded the cover and estimated dry biomass of ground lichens, as well as parameters describing forest stand characteristics. The effect of reindeer grazing on both the cover and estimated dry biomass of lichens was clear: in the ungrazed (fenced) sites, the lichen cover (35.8%) was on average 5.3-fold and the dry biomass (1929 kg ha–1) 14.8-fold compared with the corresponding estimates in the grazed sites (6.8% and 130 kg ha–1). The effect of forestry on lichens was smaller. In the grazed stands the cover and biomass of lichens were higher in the mature stands compared to the younger stand development classes, whereas in the ungrazed stands there were no significant differences between the development classes. Both reindeer grazing and forestry affect the cover and biomass of ground lichens. The influence of reindeer grazing is, however, much heavier than that of forestry. The decrease of not only the biomass, but also the lichen cover, is alarming. The decrease of lichen cover may hinder the recovery of reindeer pastures, which in the long run endangers the sustainability of reindeer husbandry.
  • Akujärvi, Finnish Environment Institute, Helsinki, Finland E-mail: anu.akujarvi@ymparisto.fi
  • Hallikainen, Finnish Forest Research Institute, Northern Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: ville.hallikainen@metla.fi
  • Hyppönen, Finnish Forest Research Institute, Northern Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: mikko.hypponen@metla.fi (email)
  • Mattila, Finnish Forest Research Institute, Northern Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: eero.mattila@metla.fi
  • Mikkola, Finnish Forest Research Institute, Northern Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: kari.mikkola@metla.fi
  • Rautio, Finnish Forest Research Institute, Northern Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: pasi.rautio@metla.fi
article id 1120, category Research article
Ilari Lehtonen, Petri Hoppula, Pentti Pirinen, Hilppa Gregow. (2014). Modelling crown snow loads in Finland: a comparison of two methods. Silva Fennica vol. 48 no. 3 article id 1120. https://doi.org/10.14214/sf.1120
Keywords: climate; forests; snow damage; rime; dry snow; wet snow; frozen snow
Highlights: A new method to model crown snow loads is presented and compared with a previously published simpler method; The heaviest crown snow loads in Finland are found to typically occur in the eastern parts of the country; The relative importance of different snow load types varies between different regions of Finland.
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The spatial occurrence of heavy crown snow loads in Finland between 1961 and 2010 is studied by using for the first time a model that classifies the snow load into four different types: rime, dry snow, wet snow and frozen snow. In producing this climatology, we used meteorological observations made at 29 locations across Finland. The model performance is evaluated against classified daily images of canopy snow cover and with the help of two short case studies. The results are further compared to those achieved with a simpler method used in previous studies. The heaviest crown snow loads are found to occur typically in eastern Finland. The new method reveals that this holds not only for the total snow loads but also for the different snow load types, although there are certain differences in their geographical occurrence. The greatest benefit achieved with the new method is the inclusion of rime accretion. The forests most prone to heavy riming are those located on tree-covered hills in northern Finland, but as the terrain elevation affects riming efficiency greatly, these small-scale variations in the snow load amounts could not be described in this study in great detail. Moreover, the results are more inaccurate in northern Finland where variations in the terrain elevation are greater than elsewhere. Otherwise, the largest uncertainties in this study are related to wind speed measurements and possibly partly because of that, we were not able to detect any significant trends in the crown snow-load amounts over the study period.
  • Lehtonen, Finnish Meteorological Institute, P.O. Box 503, FI-00101 Helsinki, Finland E-mail: ilari.lehtonen@fmi.fi (email)
  • Hoppula, Finnish Meteorological Institute, P.O. Box 503, FI-00101 Helsinki, Finland E-mail: petri.hoppula@fmi.fi
  • Pirinen, Finnish Meteorological Institute, P.O. Box 503, FI-00101 Helsinki, Finland E-mail: pentti.pirinen@fmi.fi
  • Gregow, Finnish Meteorological Institute, P.O. Box 503, FI-00101 Helsinki, Finland E-mail: hilppa.gregow@fmi.fi
article id 1087, category Research article
Ilkka Korpela, Lauri Mehtätalo, Lauri Markelin, Anne Seppänen, Annika Kangas. (2014). Tree species identification in aerial image data using directional reflectance signatures. Silva Fennica vol. 48 no. 3 article id 1087. https://doi.org/10.14214/sf.1087
Keywords: forestry; reflectance calibration; BRDF; mixed-effects modeling; Monte-Carlo simulation
Highlights: Multispectral reflectance data showed a strong and spectrally correlated tree effect; There was no gain in species classification from using species-specific differences of directional reflectance in real data and only a marginal improvement in simulated data; The directional signatures extracted in multiple images are obscured by the intrinsic within-species variation, correlated observations and inherent reflectance calibration errors.
Abstract | Full text in HTML | Full text in PDF | Author Info
Tree species identification using optical remote sensing is challenging. Modern digital photogrammetric cameras enable radiometrically quantitative remote sensing and the estimation of reflectance images, in which the observations depend largely on the reflectance properties of targets. Previous research has shown that there are species-specific differences in how the brightness observed changes when the viewing direction in an aerial image is altered. We investigated if accounting for such directional signatures enhances species classification, using atmospherically corrected, real and simulated multispectral Leica ADS40 line-camera data. Canopy in direct and diffuse illumination were differentiated and species-specific variance-covariance structures were analyzed in real reflectance data, using mixed-effects modeling. Species classification simulations aimed at elucidating the level of accuracy that can be achieved by using images of different quality, number and view-illumination geometry. In real data, a substantial variance component was explained by tree effect, which demonstrates that observations from a tree correlate between observation geometries as well as spectrally. Near-infrared band showed the strongest tree effect, while the directionality was weak in that band. The gain from directional signatures was insignificant in real data, while simulations showed a potential gain of 1–3 percentage points in species classification accuracy. The quality of reflectance calibration was found to be important as well as the image acquisition geometry. We conclude that increasing the number of image observations cancels out random observation noise and reflectance calibration errors, but fails to eliminate the tree effect and systematic calibration inaccuracy. Directional reflectance constitutes a marginal improvement in tree species classification.
  • Korpela, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014, Finland E-mail: ilkka.korpela@helsinki.fi (email)
  • Mehtätalo, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: lauri.mehtatalo@uef.fi
  • Markelin, Department of Remote Sensing and Photogrammetry, Finnish Geodetic Institute, P.O. Box 15, FI-02431 Masala, Finland E-mail: lauri.markelin@fgi.fi
  • Seppänen, Faculty of Science and Forestry, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: anne.seppanen@arbonaut.com
  • Kangas, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014, Finland E-mail: annika.kangas@helsinki.fi
article id 1054, category Research article
Karin Kolis, Juhana Hiironen, Esa Ärölä, Arvo Vitikainen. (2014). Effects of sale-specific factors on stumpage prices in Finland. Silva Fennica vol. 48 no. 3 article id 1054. https://doi.org/10.14214/sf.1054
Keywords: pulpwood; timber trade; standing timber sales; hedonic price; sawlogs; non-industrial forest owners
Highlights: Data on 4824 individual sales were used to estimate which factors affect stumpage prices; The time of sale, seasonal harvest restrictions, the location and the assortment affected prices; Larger total volumes and shorter forest haulage distances raised unit prices; A higher percentage of the assortment and percentage of sawlogs within the sale corresponded to higher prices.
Abstract | Full text in HTML | Full text in PDF | Author Info
Buyers of standing timber take not only the market situation but also the harvest costs into consideration when making purchase offers. In Finland, 85% of all timber is sold as standing timber, but there is little information for forest owners and third parties regarding how differences in harvest costs are reflected in the stumpage prices. This article analyses the relationship between sale-specific factors and stumpage prices in Finland. Data on 4824 standing timber sales between 2008 and 2012 were gathered from five local Forest Management Associations. Regression analyses were run on the stumpage prices (euros m–3) paid for sawlogs and pulpwood. Seasonal harvest restrictions, the volume of the sale and the timber assortment influenced stumpage prices, as did the presence of forest damages. Prices also differed over time and between locations. Furthermore, the forest haulage distance was statistically significant for pulpwood. The results suggest that the size of the individual sales and the composition of assortments affect the income owners earn from their forest. The results can be used to estimate stumpage prices and the monetary impacts on forest owners of procedures such as forest road network planning and land consolidation, as well as for valuation of forests.
  • Kolis, Aalto University, Department of Real Estate, Planning and Geoinformatics, P.O. Box 12200, FI-00076 Aalto, Finland E-mail: karin.kolis@aalto.fi (email)
  • Hiironen, Aalto University, Department of Real Estate, Planning and Geoinformatics, P.O. Box 12200, FI-00076 Aalto, Finland E-mail: juhana.hiironen@aalto.fi
  • Ärölä, National Land Survey of Finland, Production Support Services, P.O. Box 84, FI-00521 Helsinki, Finland E-mail: esa.arola@nls.fi
  • Vitikainen, Aalto University, Department of Real Estate, Planning and Geoinformatics, P.O. Box 12200, FI-00076 Aalto, Finland E-mail: arvo.vitikainen@aalto.fi
article id 1072, category Research article
Polina Volkova, Alexey Shipunov, Polina Borisova, Reed Moseng, Ranelle Ivens. (2014). In search of hybridity: the case of Karelian spruces. Silva Fennica vol. 48 no. 2 article id 1072. https://doi.org/10.14214/sf.1072
Keywords: cpDNA; Picea; plant mtDNA; introgression; northern taiga
Highlights: Karelian spruces have morphology that is typical for P. obovata and characterized with genetic variation, described previously for P. abies; Karelian spruces evolved due to introgression between P. abies and P. obovata. However, it is unclear whether Karelian spruces could be treated as P. fennica, because unequivocal morphological and genetic characters of the latter are still absent.
Abstract | Full text in HTML | Full text in PDF | Author Info
Distribution ranges of spruces, European Picea abies and Siberian P. obovata intersect in the Eastern Europe and Urals, forming wide zone of introgressive hybridization where species discrimination is difficult. We applied both molecular (mtDNA and cpDNA sequences) and morphological approaches with goals of elucidating the origin of spruces in undisturbed forests of Russian Karelia (considered as part of the abovementioned introgression zone). Karelian spruces have morphology that is typical for P. obovata and characterized with genetic variation, described previously for P. abies. This contradiction between morphology and organellar DNA could be itself an evidence of hybrid origin because morphological data should have a genetic basis. If the genes responsible for the observed morphological differences are nuclear, that explains why we did not see any deviation of Karelian spruces towards P. obovata in organellar markers. In this situation nuclear gene flow from P. obovata should be performed via pollen. Thus, we should expect Karelian spruces to have cpDNA haplotypes (inherited paternally in conifers) typical for P. obovata. However, it is not the case for the majority of plants sampled and requires additional explanation like chloroplast capture in the introgresson zone. In all, we think that Karelian spruces evolved due to introgression between P. abies and P. obovata. However, it is unclear whether Karelian spruces could be treated as P. fennica, because unequivocal morphological and genetic characters of this hybridogenous species are still absent.
  • Volkova, Moscow South-West High School (No. 1543), 26 Bakinskikh komissarov str. 3–5, RU-119571 Moscow, Russia E-mail: avolkov@orc.ru (email)
  • Shipunov, Department of Biology, Minot State University, Minot, North Dakota, USA 58707 E-mail: dactylorhiza@gmail.com
  • Borisova, Biological Department, Moscow State University, Vorobjevy Gory, RU-119899, Moscow, Russia E-mail: salixhastata@ya.ru
  • Moseng, Minot High School, Minot, North Dakota, USA 58701 E-mail: dactylorhiza@gmail.com
  • Ivens, Department of Biology, Minot State University, Minot, North Dakota, USA 58707 E-mail: dactylorhiza@gmail.com
article id 1064, category Research article
Back Tomas Ersson, Urban Bergsten, Ola Lindroos. (2014). Reloading mechanized tree planting devices faster using a seedling tray carousel. Silva Fennica vol. 48 no. 2 article id 1064. https://doi.org/10.14214/sf.1064
Keywords: productivity; cost analysis; seedling feed; silviculture; time study; tree planting machine; containerized seedling
Highlights: Seedling reloading onto the Bracke Planter crane-mounted planting device was twice as fast with the MagMat tray-wise-loaded carousel as today’s seedling-wise-loaded carousel; Tray-wise reloading combined with deplugging seedlings from suitable cultivation trays has the potential to be an efficient and robust way to feed seedlings on any type of tree planting machine.
Abstract | Full text in HTML | Full text in PDF | Author Info
On Nordic clearcuts, today’s tree planting machines produce high-quality but costly regenerations. Much of this high cost is attributable to the planting machines’ low productivity. One promising way of raising productivity is to lessen the time spent manually reloading seedlings onto the carousels of crane-mounted planting devices. Using MagMat, a carousel test-rig designed by engineering students, we studied how much faster tray-wise seedling reloading is on the Bracke Planter compared to reloading with today’s seedling-wise-loaded carousel. The MagMat test-rig held eight Hiko cultivation trays from which seedlings were deplugged individually and dropped into the planting tube. The time study confirmed that seedling reloading was on average twice as fast with MagMat compared to today’s seedling carousel, thereby increasing assumed planting machine productivity by 8–9% depending on the planting device used. MagMat’s cost-efficiency was analysed to be particularly reliant on its added investment cost, mechanical availability and how quickly trays can be switched automatically. Nevertheless, MagMat’s field performance illustrated the overall potential of tray-wise loading compared to piecewise seedling loading for increasing the productivity of crane-mounted planting devices. Also, deplugging proved to be a reliable method of extracting seedlings from the rigid, copper-painted Hiko cultivation trays even when performed at the excavator’s boom-tip during mounding work. We conclude that, rather than piecewise seedling loading, tray-wise loading combined with deplugging seedlings from suitable cultivation trays is a reliable and much more time-efficient method to feed seedlings on probably any type of tree planting machine.
  • Ersson, Department of Forest Biomaterials and Technology, SLU, SE-90183 Umeå, Sweden E-mail: back.tomas.ersson@slu.se (email)
  • Bergsten, Department of Forest Biomaterials and Technology, SLU, SE-90183 Umeå, Sweden E-mail: urban.bergsten@slu.se
  • Lindroos, Department of Forest Biomaterials and Technology, SLU, SE-90183 Umeå, Sweden E-mail: ola.lindroos@slu.se
article id 1019, category Research article
Michael Henke, Stephan Huckemann, Winfried Kurth, Branislav Sloboda. (2014). Reconstructing leaf growth based on non-destructive digitizing and low-parametric shape evolution for plant modelling over a growth cycle. Silva Fennica vol. 48 no. 2 article id 1019. https://doi.org/10.14214/sf.1019
Keywords: growth modelling; non-destructive data acquisition; automated data extraction; image processing tool; leaf shape modelling; reusable modules; Populus x canadensis
Highlights: A complete pipeline for plant organ modelling (at the example of poplar leaves) is presented, from non-destructive data acquisition, over automated data extraction, to growth and shape modelling; Leaf contour models are compared; Resulting “organ” modules are ready for use in FSPMs.
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A simple and efficient photometric methodology is presented, covering all steps from field data acquisition to binarization and allowing for leaf contour modelling. This method comprises the modelling of area and size (correlated and modelled with a Chapman-Richards growth function, using final length as one parameter), and four shape descriptors, from which the entire contour can be reconstructed rather well using a specific spline methodology. As an improvement of this contour modelling method, a set of parameterized polynomials was used. To model the temporal kinetics of the shape, geodesics in shape spaces were employed. Finally it is shown how this methodology is integrated into the 3D modelling platform GroIMP.
  • Henke, Department Ecoinformatics, Biometrics & Forest Growth, University of Göttingen, 37077 Göttingen, Germany E-mail: mhenke@uni-goettingen.de (email)
  • Huckemann, Institute of Mathematical Stochastics, University of Göttingen, 37077 Göttingen, Germany E-mail: huckeman@math.uni-goettingen.de
  • Kurth, Department Ecoinformatics, Biometrics & Forest Growth, University of Göttingen, 37077 Göttingen, Germany E-mail: wk@informatik.uni-goettingen.de
  • Sloboda, Department Ecoinformatics, Biometrics & Forest Growth, University of Göttingen, 37077 Göttingen, Germany E-mail: bslobod@web.de
article id 1013, category Research article
Nils Fahlvik, Björn Elfving, Peder Wikström. (2014). Evaluation of growth functions used in the Swedish Forest Planning System Heureka. Silva Fennica vol. 48 no. 2 article id 1013. https://doi.org/10.14214/sf.1013
Keywords: basal area; simulation; validation; volume; empirical
Highlights: Growth models based on historical growth data gave reliable growth predictions up to the century shift; Detailed single tree growth models had lower precision for estimation of total growth than one single stand-based model; The prediction error was in average about 15% and did not increase with extended prediction period.
Abstract | Full text in HTML | Full text in PDF | Author Info
The performance of growth models implemented in the Swedish Forest Planning System Heureka was evaluated. Four basal area growth models were evaluated by comparing their predictions to data from five-year growth records for 1711 permanent sample plots of the National Forest Inventory (NFI-data). Also, two alternative implementations of Heureka, including a combined stand- and tree-level basal area growth model and a single tree-level model, respectively, were evaluated using data from 57 blocks in a thinning experiment (GG-data) involving Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst) in which the trees were monitored for 30 years after the first thinning. The predicted volume growth was also compared to observed values. Growth models based on data from 1970’s and 1980’s overestimated growth in the NFI test plots from the early 2000’s by about 3%. Stand-level models had larger precision than tree-level models. Basal area growth was underestimated in dense NFI-plots and overestimated in non-thinned GG-plots, illustrating an un-solved modelling problem. Basal area growth was overestimated by 2–5% also in the GG-plots over the whole observation period. Volume growth was however accurately predicted for pine and underestimated by 2% for spruce. The relative prediction error did not increase with increasing length of prediction period. Thinning response models calibrated with GG-data worked well in the total application and produced growth levels for different thinning alternatives in line with observations.
  • Fahlvik, Department of Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: nils.fahlvik@slu.se (email)
  • Elfving, Department of Forest Ecology and Management, Swedish University of Agricultural Sciences, SE-901 83 Umeå, Sweden E-mail: bjorn.elfving@slu.se
  • Wikström, Peder Wikström Skogsanalys AB, c/o Peder Wikström, Huldrans väg 1, SE-907 52 Umeå, Sweden E-mail: peder.wikstrom@slu.se
article id 983, category Research article
Sakari Tuominen, Juho Pitkänen, Andras Balazs, Kari T. Korhonen, Pekka Hyvönen, Eero Muinonen. (2014). NFI plots as complementary reference data in forest inventory based on airborne laser scanning and aerial photography in Finland. Silva Fennica vol. 48 no. 2 article id 983. https://doi.org/10.14214/sf.983
Keywords: airborne laser scanning; National Forest Inventory; aerial imagery; plot sampling
Highlights: Using NFI plots in forest management inventories could provide a way for rationalising forest inventory data acquisition; NFI plots were used as additional reference data in laser scanning and aerial image based forest inventory; NFI plots improved the estimates of some forest variables; There are differences between the two inventory types that cause difficulties in combining the data.
Abstract | Full text in HTML | Full text in PDF | Author Info
In Finland, there are currently two, parallel sample-plot-based forest inventory systems, which differ in their methodologies, sampling designs, and objectives. One is the National Forest Inventory (NFI), aimed at unbiased inventory results at national and regional level. The other is the Forest Centre’s management-oriented forest inventory based on interpretation of airborne laser scanning and aerial images, with the aim of locally accurate stand-level forest estimates. The National Forest Inventory utilises relascope sample plots with systematic cluster sampling. This inventory method is optimised for accuracy of regional volume estimates. In contrast, the management-oriented forest inventory utilises circular sample plots with an allocation system covering certain pre-defined forest classes in the inventory area. This method is optimised to produce reference data for interpretation of the remote-sensing materials in use. In this study, we tested the feasibility of the National Forest Inventory sample plots in provision of additional reference data for the management-oriented inventory. Various combinations of NFI plots and management inventory plots were tested in the interpretation of the laser and aerial-image data. Adding NFI plots in the reference data generally improved the accuracy of the volume estimates by tree species but not the estimates of total volume or stand mean height and diameter. The difference between the plot types in the NFI and management inventories causes difficulties in combination of the two datasets.
  • Tuominen, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: sakari.tuominen@metla.fi (email)
  • Pitkänen, Finnish Forest Research Institute, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: juho.pitkanen@metla.fi
  • Balazs, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: andras.balazs@metla.fi
  • Korhonen, Finnish Forest Research Institute, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: kari.t.korhonen@metla.fi
  • Hyvönen, Finnish Forest Research Institute, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: pekka.hyvonen@metla.fi
  • Muinonen, Finnish Forest Research Institute, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: eero.muinonen@metla.fi
article id 982, category Research article
Karri Uotila, Timo Saksa, Juho Rantala, Nuutti Kiljunen. (2014). Labour consumption models applied to motor-manual pre-commercial thinning in Finland. Silva Fennica vol. 48 no. 2 article id 982. https://doi.org/10.14214/sf.982
Keywords: productivity; pre-commercial thinning; forest vegetation management; early cleaning; release treatment; mixed linear regression
Highlights: When a young stand grows and gets older, the work time needed to make pre-commercial thinning increases. The stands of Norway spruce (Picea abies (L.) Karst.), Scots pine (Pinus sylvestris L.) and hardwoods (Betula spp.) required an additional 8.2%, 5.2%, and 3.3% work-time per year, respectively.
Abstract | Full text in HTML | Full text in PDF | Author Info
Labour models were developed to estimate the time required to Pre-Commercially Thin (PCT) with a clearing saw 4- to 20-year-old stands of the main commercial tree species in Finland. Labour (i.e., work-time consumption) was estimated from the density and stem diameter of the removal of 448 stands via an existing work productivity function. The removal based estimator attained was used as the basis for a priori mixed linear regression models. The main finding was that when a young stand grows and gets older, the work time needed to make a PCT increases. The stands of Norway spruce (Picea abies (L.) Karst.), Scots pine (Pinus sylvestris L.) and hardwoods (Betula spp.) required an additional 8.2%, 5.2%, and 3.3% work-time per year, respectively. Site fertility also played a role in that the most fertile site (mesic OMT) had an estimated labour requirement 114% higher than that for dryish VT. We also note that, per unit area, small stands require less labour than large ones and soil preparation method had a minor effect on the labour estimate. The stands which had previously gone through PCT were separately analysed. In those stands, the only significant variable concerning the labour estimate was age. The a priori models described here can help foresters to develop economic management programmes and issue quotes for forestry services.
  • Uotila, Finnish Forest Research Institute, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: karri.uotila@metla.fi (email)
  • Saksa, Finnish Forest Research Institute, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@metla.fi
  • Rantala, Metsä Group, Lielahdenkatu 10, FI-33400 Tampere, Finland E-mail: juho.rantala@metsagroup.com
  • Kiljunen, Metsähallitus, Asemakatu 7, FI-70107 Kuopio, Finland E-mail: nuutti.kiljunen@metsa.fi
article id 975, category Research article
Renats Trubins, Ola Sallnäs. (2014). Categorical mapping from estimates of continuous forest attributes – classification and accuracy. Silva Fennica vol. 48 no. 2 article id 975. https://doi.org/10.14214/sf.975
Keywords: Sweden; land cover maps; forest type maps; map accuracy assessment; class membership probability; Bayesian network; k-NN estimates
Highlights: The paper presents an approach to classification and accuracy assessment of ad-hoc categorical maps based on existing spatial datasets with estimates of continuous forest variables; Pixel level class membership probabilities are estimated using a Bayesian network model.
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Spatially explicit data on forest attributes is demanded for various research with landscape perspective. Existing datasets with estimates of continuous forest variables are often used as the basis for producing categorical forest type maps. Normally, this type of maps are used without knowing their accuracy. This paper presents a Bayesian network model for estimating pixel level class membership probabilities of thus derived categorical maps. Class membership probabilities can be used as a post-classification measure of map accuracy and in the process of map classification affecting the assignments of class labels. The method is applied in mapping deciduous dominated forests on the basis of the k-NN Sweden 2005 dataset in a study area in southern Sweden. The results indicate rather low accuracy for deciduous class regardless of the map classification method: 0.48 versus 0.50 in the maps classified without and with the use of the class membership probabilities given equal deciduous area. When probability-based classification is applied, the level of accuracy varies with the assumed map class proportions. Thus, when deciduous class area corresponding to the National Forest Inventory estimate was used, the accuracy of only 0.35 was obtained for the deciduous map class.
  • Trubins, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, Box 49, 230 53 Alnarp, Sweden E-mail: renats.trubins@slu.se (email)
  • Sallnäs, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, Box 49, 230 53 Alnarp, Sweden E-mail: ola.sallnas@slu.se
article id 1077, category Research article
Göran Bergqvist, Roger Bergström, Märtha Wallgren. (2014). Recent browsing damage by moose on Scots pine, birch and aspen in young commercial forests – effects of forage availability, moose population density and site productivity. Silva Fennica vol. 48 no. 1 article id 1077. https://doi.org/10.14214/sf.1077
Keywords: Pinus; Betula; Populus; Alces alces
Highlights: Moose damage was most common on aspen and least common on Scots pine; Damage to Scots pine increased when the amount of pine browse decreased, moose index and site index increased and when birch was overtopping pine; Damage to birch increased when moose index increased and damage to aspen increased when the amount of pine and birch browse decreased.
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Moose browsing damage from the winter preceding the study (recent damage) on Scots pine, birch and aspen was examined in relation to forage availability, an index of moose population density and site productivity in young forests in the hemiboreal zone. Recent damage was observed for 4.1 ± 0.54% (mean ± SE; Scots pine), 16.8 ± 1.89% (birch) and 67.6 ± 13.76% (aspen) of the trees. A multiple regression with five independent variables explained 19% (Scots pine) 14% (birch) and 33% (aspen) of the variation in recent damage. Cover of Scots pine browse was the most important variable for predicting damage to Scots pine and accounted for 44% of the explained variation. When birch was overtopping pine there was a significant increase in damage to pine. Moose index was the only significant variable to explain recent damage to birch, and accounted for 64% of the explained variation. For aspen, damage was negatively correlated to coverage of Scots pine and birch browse, each variable accounting for 38% of the explained variation. For Scots pine, increasing the number of pines ha–1 and performing pre-commercial thinning in such a way that pines are not overtopped may be efficient ways of reducing damage proportions, whereas birch needs to be protected from moose (by a reduction of the moose population or otherwise) in order to escape damage. Increased amounts of Scots pine browse and birch browse may also reduce damage levels to aspen, according to this study.
  • Bergqvist, Swedish Association for Hunting and Wildlife Management, Öster Malma, SE-611 91 Nyköping, Sweden E-mail: goran.bergqvist@jagareforbundet.se (email)
  • Bergström, Forestry Research Institute of Sweden, Uppsala Science Park, SE-751 83 Uppsala, Sweden; (present) Gropgränd 2 A, SE-753 10 Uppsala, Sweden E-mail: larsrogerbergstrom@yahoo.com
  • Wallgren, Forestry Research Institute of Sweden, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: martha.wallgren@skogforsk.se
article id 1003, category Research article
Raffaele Spinelli, Carolina Lombardini, Natascia Magagnotti. (2014). The effect of mechanization level and harvesting system on the thinning cost of Mediterranean softwood plantations. Silva Fennica vol. 48 no. 1 article id 1003. https://doi.org/10.14214/sf.1003
Keywords: chipping; biomass; logging; compaction; stand damage
Highlights: Whole-tree harvesting is 40–50% cheaper than cut-to-length harvesting; Mechanization reduced thinning cost by a factor 4; Between 1.5 and 6% of the residual trees were damaged; Mechanized cut-to-length harvesting allows controlled biomass release; Mechanized whole-tree harvesting is the cheapest option for energy chip production.
Abstract | Full text in HTML | Full text in PDF | Author Info
The study compared motor-manual cut-to-length (CTL) harvesting, motor-manual whole-tree (WT) harvesting, mechanized CTL harvesting and mechanized WT harvesting as applied to the production of energy chips from the second thinning of Mediterranean pine plantations in flat terrain. Mechanization increased productivity between 6 and 20 times, depending on process step. It also allowed reducing thinning cost by a factor 4. Shifting from CTL to WT harvesting resulted in a reduction of harvesting cost between 40 and 50%. Fuel consumption was between 40 and 100% higher for CTL harvesting than for WT harvesting. Mechanization entailed a reduction of fuel consumption between 10 and 40%. Stand damage was generally low, between 1.5 and 6%. Mechanized CTL harvesting resulted in the lowest incidence of wounding, and the difference between mechanized CTL and manual WT harvesting was statistically significant. Soil compaction was absent or very small, depending on treatment. Mechanized thinning may produce larger increases of soil bulk density, compared to motor-manual thinning, but the difference is small, although significant. CTL harvesting leaves a larger amount of biomass on the soil, which relieves possible concerns about soil nutrient depletion. On the other hand, heavy residue loads may increase fire risk especially in sensitive Mediterranean environments.
  • Spinelli, CNR IVALSA,Via Madonna del Piano 10, Sesto Fiorentino (FI), Italy E-mail: spinelli@ivalsa.cnr.it (email)
  • Lombardini, CNR IVALSA,Via Madonna del Piano 10, Sesto Fiorentino (FI), Italy E-mail: lombardini@ivalsa.cnr.it
  • Magagnotti, CNR IVALSA,Via Biasi 75, S. Michele all’Adige (TN), Italy E-mail: magagnotti@ivalsa.cnr.it
article id 989, category Research article
Anna Saarsalmi, Pekka Tamminen, Mikko Kukkola. (2014). Effects of long-term fertilisation on soil properties in Scots pine and Norway spruce stands. Silva Fennica vol. 48 no. 1 article id 989. https://doi.org/10.14214/sf.989
Keywords: boreal forest; nitrogen; Pinus sylvestris L.; phosphorus; liming; forest soils; Picea abies (L.) Karst.
Highlights: N fertilisation increased the amount of carbon in the organic layer; N fertilisation decreased the C/N ratio in the surface soil; N addition increased the amount of most nutrients in the organic layer; N fertilisation tended to lower pH, although only slightly.
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The response of surface soil after 45- to 52-years to repeated nitrogen and phosphorus fertilisation was studied. This study included 30 factorial experiments established in young (5- to 30-year-old) stands using plots of 900 m2, on average, and by randomising treatments within each experiment. Total amount of N added varied from 534 to 1908 kg ha–1 and that of P from 69 to 193 kg ha–1, repeated at every second N fertilisation. Liming was performed twice; in total, 6000 kg ha–1 of dolomite was applied. Nitrogen fertilisation increased the mass of the organic layer and the amount of carbon and consequently the amounts of most of the elements in the organic layer. In both the organic layer and the 0–10 cm layer of mineral soil, nitrogen fertilisation decreased the C/N ratio and tended to lower pH, although only slightly. Phosphorus fertilisation increased the amounts of P and Ca. Liming increased the total amounts of most elements in the organic layer, except for C and N. We were able to derive models to describe how changes in the chemical properties of the surface soil depended on doses of elements and on site and stand properties.
  • Saarsalmi, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: anna.saarsalmi@metla.fi (email)
  • Tamminen, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: pekka.tamminen@metla.fi
  • Kukkola, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: mikko.kukkola@metla.fi
article id 980, category Research article
Atte Komonen, Panu Halme, Mari Jäntti, Tuuli Koskela, Janne S. Kotiaho, Tero Toivanen. (2014). Created substrates do not fully mimic natural substrates in restoration: the occurrence of polypores on spruce logs. Silva Fennica vol. 48 no. 1 article id 980. https://doi.org/10.14214/sf.980
Keywords: Norway spruce; boreal forest; ecological restoration; dead wood; protected area management; substrate quality; wood-decaying fungi
Highlights: Polypore communities were more homogeneous among created than among natural logs; The old-growth forest indicator Phellinus ferrugineofuscus occurred frequently on natural logs, but occupied only a few created logs; Results show that created logs do not fully mimic natural logs.
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Many protected areas have been under intensive forest management prior to protection and thus lack natural ecosystem structures and dynamics. Dead wood is a key structure in forests harboring hundreds of threatened species. We investigated the ecological success of dead wood creation as a boreal forest restoration measure. We analysed whether the polypore communities of chain-saw felled and girdled (subsequently fallen) Norway spruce (Picea abies (L.) H. Karst.) logs differ from naturally formed spruce logs of similar decay stage and size. The study was conducted in Leivonmäki National Park in central Finland 8 years after the restoration measures. The average number of polypore species was highest on the chain-saw felled logs and most of the common polypore species were most frequent on this substrate. However, among the natural logs, number of species increased more steeply with increasing number of logs, suggesting greater variation in community composition on this substrate. The old-growth forest indicator Phellinus ferrugineofuscus occurred frequently on natural logs, occupied a few girdled logs but was absent from chain-saw felled logs. Our results show that from the polypore perspective created logs do not fully mimic natural logs, suggesting that creating substrates for species may pose a challenge for restoration.
  • Komonen, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland E-mail: atte.komonen@jyu.fi (email)
  • Halme, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland E-mail: panu.halme@jyu.fi
  • Jäntti, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland E-mail: mari.j.jantti@student.jyu.fi
  • Koskela, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland E-mail: tuuli.e.koskela@student.jyu.fi
  • Kotiaho, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland E-mail: janne.kotiaho@jyu.fi
  • Toivanen, Department of Biological and Environmental Science, P.O. Box 35, FI-40014, University of Jyväskylä, Finland; Current: Birdlife Finland, Annankatu 29 A 16, FI-00100 Helsinki, Finland E-mail: tero.toivanen@birdlife.fi
article id 962, category Research article
Paul A. Klockow, Anthony W. D'Amato, John B. Bradford, Shawn Fraver. (2014). Nutrient concentrations in coarse and fine woody debris of Populus tremuloides Michx.-dominated forests, northern Minnesota, USA. Silva Fennica vol. 48 no. 1 article id 962. https://doi.org/10.14214/sf.962
Keywords: coarse woody debris; fine woody debris; Populus tremuloides; nutrient concentrations; bioenergy feedstock harvesting; Great Lakes
Highlights: We examine effects of size, species, and decay on woody debris nutrient concentrations; Results indicate wide variation in nutrient concentrations across the factors examined; Fine woody debris nutrient concentrations were greater than in coarse woody debris; Coarse woody debris nutrient concentrations generally increased as decay progressed; Results suggest high fine woody debris stocks can represent an important nutrient source.
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Contemporary forest harvesting practices, specifically harvesting woody biomass as a source of bioenergy feedstock, may remove more woody debris from a site than conventional harvesting. Woody debris, particularly smaller diameter woody debris, plays a key role in maintaining ecosystem nutrient stores following disturbance. Understanding nutrient concentrations within woody debris is necessary for assessing the long-term nutrient balance consequences of altered woody debris retention, particularly in forests slated for use as bioenergy feedstocks. Nutrient concentrations in downed woody debris of various sizes, decay classes, and species were characterized within one such forest type, Populus tremuloides Michx.-dominated forests of northern Minnesota, USA. Nutrient concentrations differed significantly between size and decay classes and generally increased as decay progressed. Fine woody debris (≤ 7.5 cm diameter) had higher nutrient concentrations than coarse woody debris (> 7.5 cm diameter) for all nutrients examined except Na and Mn, and nutrient concentrations varied among species. Concentrations of N, Mn, Al, Fe, and Zn in coarse woody debris increased between one and three orders of magnitude, while K decreased by an order of magnitude with progressing decay. The variations in nutrient concentrations observed here underscore the complexity of woody debris nutrient stores in forested ecosystems and suggest that retaining fine woody debris at harvest may provide a potentially important source of nutrients following intensive removals of bioenergy feedstocks.
  • Klockow, Department of Forest Resources, University of Minnesota, St. Paul, MN 55108, USA E-mail: klock039@umn.edu (email)
  • D'Amato, Department of Forest Resources, University of Minnesota, St. Paul, MN 55108, USA E-mail: damato@umn.edu
  • Bradford, US Geological Survey, Southwest Biological Science Center, Flagstaff, AZ 86001, USA E-mail: jbradford@usgs.gov
  • Fraver, School of Forest Resources, University of Maine, Orono, ME 04469, USA E-mail: shawn.fraver@maine.edu
article id 949, category Research article
Ken Olaf Storaunet, Jørund Rolstad, Erlend Rolstad. (2014). Effects of logging on the threatened epiphytic lichen Usnea longissima: an experimental approach. Silva Fennica vol. 48 no. 1 article id 949. https://doi.org/10.14214/sf.949
Keywords: boreal forest; Picea abies; selective logging; Usnea longissima; threatened lichen
Highlights: A re-inventory of the threatened lichen Usnea longissima in ten Norway spruce forest stands where experimental selective loggings had been conducted 5 to 8 years before revealed that the number of lichen thalli had increased with 34%; The number of thalli increased more where the forest was open whether or not the low tree density was caused by the loggings.
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Usnea longissima Ach. is a circumboreal epiphytic lichen draping tree canopies in moist coastal and mountainous forests. It is extinct from many European and North-American localities, presumably due to industrial forestry and air pollution, but still has a stronghold in parts of Scandinavia and U.S. and Canadian Pacific Northwest. In 2005/06 we used a comparative and retrospective approach to evaluate how present and historic tree and stand characteristics influenced the occurrence and abundance of the lichen (Storaunet et al. 2008). In 2012, we re-inventoried ten Norway spruce forest stands with 401 U. longissima-bearing trees and recorded changes in the number of U. longissima thalli. Seven of the stands had been experimentally, selectively logged 5–8 years before, where the lichen-bearing trees had been marked in the field and were avoided during the logging operation. Total number of lichen-bearing trees decreased slightly (2.9%), whereas the number of thalli had increased with 34%. Number of thalli increased more where the forest was open (low basal area, m2ha-1) whether or not the low tree density was caused by the logging events. At high tree densities the change in number of thalli was negligible. We suggest that selective logging, securing lichen-bearing trees, may be a viable management option to keep tree density from becoming too dense, thereby enhancing growth and establishment of U. longissima.
  • Storaunet, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway E-mail: stk@skogoglandskap.no (email)
  • Rolstad, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway E-mail: roj@skogoglandskap.no
  • Rolstad, Skogfaglig Rådgivning, Holmsida 126, NO-1488 Hakadal, Norway E-mail: roe@skogoglandskap.no
article id 1073, category Research article
Pedro Sánchez-Gómez, Juan F. Jiménez, Juan B. Vera, Francisco J. Sánchez-Saorín, Juan F. Martínez, Joseph Buhagiar. (2013). Genetic structure of Tetraclinis articulata, an endangered conifer of the western Mediterranean basin. Silva Fennica vol. 47 no. 5 article id 1073. https://doi.org/10.14214/sf.1073
Keywords: conservation; genetic differentiation; genetic diversity; ISSR markers
Highlights: The employment of ISSR molecular markers has shown moderate genetic diversity and high genetic differentiation in Tetraclinis articulata; Genetic structure of populations seems to be influenced by the anthropogenic use of this species since historical times, or alternatively, by the complex palaeogeographic history of the Mediterranean basin; Results could be used to propose management policies for conservation of populations.
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Tetraclinis articulata (Vahl) Masters is a tree distributed throughout the western Mediterranean basin. It is included in the IUCN (International Union for Conservation of Nature) red list, and protected by law in several of the countries where it grows. In this study we examined the genetic diversity and genetic structure of 14 populations of T. articulata in its whole geographic range using ISSR (inter simple sequence repeat) markers. T. articulata showed moderate genetic diversity at intrapopulation level and high genetic differentiation. The distribution of genetic diversity among populations did not exhibit a linear pattern related to geographic distances, since all analyses (principal coordinate analysis, Unweighted pair group method with arithmetic mean dendrogram and Bayesian structure analysis) revealed that spanish population grouped with Malta and Tunisia populations. Although it is possible that T. articulata earlier was natural in Southeast Spain, results suggest that the current population has been reintroduced into the Iberian Peninsula in historical times, due to its utility in mining and building. In addition, results could be used to propose management guidelines for the conservation of T. articulata.
  • Sánchez-Gómez, Departamento de Biología Vegetal (Botánica), Universidad de Murcia, Campus de Espinardo s/n, E-30100 Murcia, Spain E-mail: psgomez@um.es
  • Jiménez, Departamento de Biología Vegetal (Botánica), Universidad de Murcia, Campus de Espinardo s/n, E-30100 Murcia, Spain E-mail: fjimenez@um.es (email)
  • Vera, Departamento de Biología Vegetal (Botánica), Universidad de Murcia, Campus de Espinardo s/n, E-30100 Murcia, Spain E-mail: jbveraperez@gmail.com
  • Sánchez-Saorín, Dirección General de Medio Ambiente, Consejería de Presidencia de la Región de Murcia, C/ Catedrático Eugenio Úbeda nº3, E-30071 Murcia, Spain E-mail: fjavier.sanchez3@carm.es
  • Martínez, Dirección General de Medio Ambiente, Consejería de Presidencia de la Región de Murcia, C/ Catedrático Eugenio Úbeda nº3, E-30071 Murcia, Spain E-mail: juanf.martinez@carm.es
  • Buhagiar, Argotti Herbarium and Gardens (UOM), Triq Vincenzo Bugeja Floriana VLT 16, Malta E-mail: joseph.buhagiar@um.edu.mt
article id 1071, category Research article
Ursula Kretschmer, Nadeschda Kirchner, Christopher Morhart, Heinrich Spiecker. (2013). A new approach to assessing tree stem quality characteristics using terrestrial laser scans. Silva Fennica vol. 47 no. 5 article id 1071. https://doi.org/10.14214/sf.1071
Keywords: LIDAR; terrestrial laser scanner; tree stem quality assessment; cylinder approximation; bark defects
Highlights: Minimal deviations of the bark surface can be detected and visualized based on terrestrial laser scan data; Additionally the geometrical properties of bark scars and branched knots can be assessed; Two methods using two different approaches are presented: (1) a method using intensity data and (2) a method using bark surface models.
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This paper presents an approach to assess and measure bark characteristics as indicators of wood quality using terrestrial laser scan data. In addition to the detection and measurement by use of the intensity information of the scan data a new approach was established. Bark surface models are calculated for each tree. They offer the representation of the bark as a height model. The reference is the tree stem approximated by a chain of cylinders. Minimal deviations of the bark surface can be detected and visualized and the geometrical properties of bark scars and branched knots can be assessed. Results of the measurement of 18 scars are presented using the two approaches: (1) a method using intensity data or (2) using bark surface models. The selection of the adequate approach depends on the stem characteristics. In a next step, methods for automatic measurement of bark scars will be developed.
  • Kretschmer, Chair of Forest Growth, Albert-Ludwigs-University Freiburg, Tennenbacher Str. 4, 79106 Freiburg, Germany E-mail: ursula.kretschmer@iww.uni-freiburg.de (email)
  • Kirchner, VOLKE Consulting Engineers GmbH, Schätzweg 7-9, 80935 München, Germany E-mail: nadeschda.kirchner@volke.muc.de
  • Morhart, Chair of Forest Growth, Albert-Ludwigs-University Freiburg, Tennenbacher Str. 4, 79106 Freiburg, Germany E-mail: christopher.morhart@iww.uni-freiburg.de
  • Spiecker, Chair of Forest Growth, Albert-Ludwigs-University Freiburg, Tennenbacher Str. 4, 79106 Freiburg, Germany E-mail: instww@uni-freiburg.de
article id 984, category Research article
Christian Rottensteiner, Petros Tsioras, Heinz Neumayer, Karl Stampfer. (2013). Vibration and noise assessment of tractor-trailer and truck-mounted chippers. Silva Fennica vol. 47 no. 5 article id 984. https://doi.org/10.14214/sf.984
Keywords: chippers; noise; whole-body vibration; exposure limit values
Highlights: Truck-mounted chippers were associated with higher vibration values, while tractor-trailer chippers had higher noise level; Chipping hardwood produced higher vibration magnitudes than softwood; Vibration and noise values in most cases did not exceed the exposure limit values set by the European Union.
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During chipping, machine operators are exposed to whole-body vibration and noise bearing a risk to health. Vibration on the operator’s seat and noise inside the chipper cab was measured and analyzed. The factorial design considered two setup variants (tractor-trailer and truck-mounted) of two chipper models from different manufacturers during chipping of softwood and hardwood tree species. Furthermore, exposure to noise was measured during chipping of hardwood. Vibration and noise during chipping, driving between wood piles, and operational delays were measured separately. The results associated truck-mounted chippers with higher vibration values and tractor-trailer chippers with higher noise levels. The highest vibration levels were recorded while driving on the forest road from one log pile to another and the second highest during chipping. On the contrary, the lowest vibration levels were measured during operational delays with the chipper in idling condition. Chipping hardwood produced higher vibration magnitudes than softwood. Exposure to noise was significantly higher during chipping compared to driving and operational delays. Vibration and noise data were combined with time studies data, for the calculation of eight-hour energy equivalent total values, both for vibration and noise. In all cases, the exposure limit values set by the European Union were not exceeded, with the exception of truck-mounted chippers, which are likely to exceed the exposure action value for vibration.
  • Rottensteiner, University of Natural Resources and Life Sciences Vienna, Institute of Forest Engineering, Peter Jordan Straße 82, 1190 Wien, Austria E-mail: christian.rottensteiner@boku.ac.at (email)
  • Tsioras, Aristotle University, P.O. Box 227, GR-541 24 Thessaloniki, Greece E-mail: ptsioras@for.auth.gr
  • Neumayer, Specialist in Occupational Medicine, Wörndlestraße 10, 6020 Innsbruck, Austria E-mail: heinz.neumayer@die-arbeitsmedizin.at
  • Stampfer, University of Natural Resources and Life Sciences Vienna, Institute of Forest Engineering, Peter Jordan Straße 82, 1190 Wien, Austria E-mail: karl.stampfer@boku.ac.at
article id 943, category Research article
Terje Gobakken, Lauri Korhonen, Erik Næsset. (2013). Laser-assisted selection of field plots for an area-based forest inventory. Silva Fennica vol. 47 no. 5 article id 943. https://doi.org/10.14214/sf.943
Keywords: forest inventory; LIDAR; airborne laser scanning; stratified sampling; area-based approach
Highlights: Using laser data as auxiliary information in the selection of field plot locations helps to decrease costs in forest inventories based on airborne laser scanning; Two independent, differently selected sets of field plots were used for model fitting, and third for validation; Using partial instead of ordinary least squares had no major influence on the results; Forty well placed plots produced fairly reliable volume estimates.
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Field measurements conducted on sample plots are a major cost component in airborne laser scanning (ALS) based forest inventories, as field data is needed to obtain reference variables for the statistical models. The ALS data also provides an excellent source of prior information that may be used in the design phase of the field survey to reduce the size of the field data set. In the current study, we acquired two independent modeling data sets: one with ALS-assisted and another with random plot selection. A third data set was used for validation. One canopy height and one canopy density variable were used as a basis for the ALS-assisted selection. Ordinary and partial least squares regressions for stem volume were fitted for four different strata using the two data sets separately. The results show that the ALS-assisted plot selection helped to decrease the root mean square error (RMSE) of the predicted volume. Although the differences in RMSE were relatively small, models based on random plot selection showed larger mean differences from the reference in the independent validation data. Furthermore, a sub-sampling experiment showed that 40 well placed plots should be enough for fairly reliable predictions.
  • Gobakken, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, Ås, Norway E-mail: terje.gobakken@umb.no
  • Korhonen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: lauri.korhonen@uef.fi (email)
  • Næsset, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, Ås, Norway E-mail: erik.naesset@umb.no
article id 937, category Research article
Rene Zamora-Cristales, Kevin Boston, John Sessions, Glen Murphy. (2013). Stochastic simulation and optimization of mobile chipping economics in processing and transport of forest biomass from residues. Silva Fennica vol. 47 no. 5 article id 937. https://doi.org/10.14214/sf.937
Keywords: forest planning; simulation; optimization; economics; decision analysis; forest biomass; renewable energy
Highlights: A stochastic simulation model is proposed to analyze forest biomass operations; The cost of chipper and truck waiting times was estimated in forest biomass recovery operations; The economic effect of truck-machine interactions under uncertainty was analyzed; Road characteristics and processing location have an economic impact in truck and chipper waiting times
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We analyzed the economics of mobile chipping and transport of biomass from forest residues for energy purposes under uncertainty. A discrete-event simulation model was developed and utilized to quantify the impacts of controllable and environmental variables on productivity in order to determine the most cost effective transportation options under steep terrain conditions. Truck-chipper interactions were analyzed to show their effect on truck and chipper standing time. A costing model was developed to account for operating and standing time cost (for the chipper and trucks). The model used information from time studies of each activity in the productive cycle and spatial-temporal information obtained from geographic information system (GIS) devices, and tracking analysis of machine and truck movements. The model was validated in field operations, and proved to be accurate in providing the expected productivity. A cost distribution was elaborated to support operational decisions of forest managers, landowners and risk-averse contractors. Different scenarios were developed to illustrate the economic effects due to changes in road characteristics such as in-highway transport distance, in-forest internal road distance and pile to trailer chipper traveling distances.
  • Zamora-Cristales, Department of Forest Engineering, Resources, and Management, College of Forestry, Oregon State University, 280 Peavy Hall, Corvallis, OR 97331, USA E-mail: rene.zamora@oregonstate.edu (email)
  • Boston, Department of Forest Engineering, Resources, and Management, College of Forestry, Oregon State University, 280 Peavy Hall, Corvallis, OR 97331, USA E-mail: kevin.boston@oregonstate.edu
  • Sessions, Department of Forest Engineering, Resources, and Management, College of Forestry, Oregon State University, 280 Peavy Hall, Corvallis, OR 97331, USA E-mail: john.sessions@oregonstate.edu
  • Murphy, Waiariki Institute of Technology, Rotorua, New Zealand E-mail: glen.murphy@waiariki.ac.nz
article id 1057, category Research article
Jouni Siipilehto, Lauri Mehtätalo. (2013). Parameter recovery vs. parameter prediction for the Weibull distribution validated for Scots pine stands in Finland. Silva Fennica vol. 47 no. 4 article id 1057. https://doi.org/10.14214/sf.1057
Keywords: linear prediction; diameter distribution; Weibull; stand characteristics; parameter recovery
Highlights: A parameter recovery method (PRM) was developed for forest stand inventories and compared with previously developed parameter prediction methods (PPM) in Finland; PRM for the 2-parameter Weibull function provided compatibility for the main stand characteristics: stem number, basal area and one of the four optional mean characteristics; PRM provided comparable and at its best, superior accuracy in volume characteristics compared with PPM.
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The moment-based parameter recovery method (PRM) has not been applied in Finland since the 1930s, even after a continuation of forest stand structure modelling in the 1980s. This paper presents a general overview of PRM and some useful applications. Applied PRM provided compatibility for the included stand characteristics of stem number (N) and basal area (G) with either mean (D), basal area-weighted mean (DG), median (DM) or basal area-median (DGM) diameter at breast height (dbh). A two-parameter Weibull function was used to describe the dbh-frequency distribution of Scots pine stands in Finland. In the validation, PRM was compared with existing parameter prediction models (PPMs). In addition, existing models for stand characteristics were used for the prediction of unknown characteristics. Validation consisted of examining the performance of the predicted distributions with respect to variation in stand density and accuracy of the localised distributions, as well as accuracy in terms of bias and the RMSE in stand characteristics in the independent test data set. The validation data consisted of 467 randomly selected stands from the National Forest Inventory based plots. PRM demonstrated excellent accuracy if G and N were both known. At its best, PRM provided accuracy that was superior to any existing model in Finland – especially in young stands (mean height < 9 m), where the RMSE in total and pulp wood volumes, 3.6 and 5.7%, respectively, was reduced by one-half of the values obtained using the best performing existing PPM (8.7–11.3%). The unweighted Weibull distribution solved by PRM was found to be competitive with weighted existing PPMs for advanced stands. Therefore, using PRM, the need for a basal area weighted distribution proved unnecessary, contrary to common belief. Models for G and N were shown to be unreliable and need to be improved to obtain more reliable distributions using PRM.
  • Siipilehto, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: jouni.siipilehto@metla.fi (email)
  • Mehtätalo, University of Eastern Finland, School of Computing, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: lauri.mehtatalo@uef.fi
article id 1047, category Research article
Kalle Karttunen, Lauri Lättilä, Olli-Jussi Korpinen, Tapio Ranta. (2013). Cost-efficiency of intermodal container supply chain for forest chips. Silva Fennica vol. 47 no. 4 article id 1047. https://doi.org/10.14214/sf.1047
Keywords: trucks; railways; wood fuels; transportation systems
Highlights: The combined availability and simulation study method obtains more realistic results for use in practical decision-making in supply chain management; The total costs of forest chips with intermodal composite container supply chains were lower than traditional options in all scenarios; The most advantageous way to expand the procurement area for forest chips is either to use composite container trucks or start using train transportation instead of trucks for procurement from longer distances.
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Cost-efficient solutions of supply chains for energy wood are required as part of endeavors to reach targets for renewable energy utilization. Long-distance railway transportation is an interesting area of research, especially for high-volume sites where the forest-to-site distance is considerable and rail facilities already exist. The aim of the study was to compare the cost-efficiency of an intermodal container supply chain and traditional multi-modal supply chain with corresponding direct truck logistics for long-distance transportation of forest chips. In the study, site-dependent information for forest biomass transport was integrated into a simulation model to calculate the cost-efficiency of logistic operations related to forest chips transportation in central Finland. The model was tested with several truck and railway transportation scenarios for varying demand of forest chips at the case power plant. The total costs of traditional supply chains were found to be 5–19% more expensive than container supply chain scenarios. The total unit costs of forest chips varied between 15.3 and 20.0 €/MWh depending on the scenario. It is concluded on the basis of the scenario study that intermodal light-structure container logistics and railway transportation could be developed as a viable option for large-scale supply of forest chips.
  • Karttunen, Lappeenranta University of Technology, LUT Savo Sustainable Technologies, Sammonkatu 12, FI-50130 Mikkeli, Finland E-mail: kalle.karttunen@lut.fi (email)
  • Lättilä, Lappeenranta University of Technology, LUT Savo Sustainable Technologies, Sammonkatu 12, FI-50130 Mikkeli, Finland E-mail: lauri.lattila@lut.fi
  • Korpinen, Lappeenranta University of Technology, LUT Savo Sustainable Technologies, Sammonkatu 12, FI-50130 Mikkeli, Finland E-mail: olli-jussi.korpinen@lut.fi
  • Ranta, Lappeenranta University of Technology, LUT Savo Sustainable Technologies, Sammonkatu 12, FI-50130 Mikkeli, Finland E-mail: tapio.ranta@lut.fi
article id 1046, category Research article
Eva-Maria Nordström, Hampus Holmström, Karin Öhman. (2013). Evaluating continuous cover forestry based on the forest owner’s objectives by combining scenario analysis and multiple criteria decision analysis. Silva Fennica vol. 47 no. 4 article id 1046. https://doi.org/10.14214/sf.1046
Keywords: forest planning; decision making; sustainable forest management; uneven-aged forestry
Highlights: Scenario analysis and multiple criteria decision analysis were combined to evaluate alternative forest management strategies for Linköping municipality, Sweden; Continuous cover forestry (CCF) promoted ecological and social objectives better than even-aged forestry but was worse for economic objectives; Ecological and social objectives were important to the municipality and thus, in summary, CCF seemed to be a suitable strategy.
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Forests are increasingly managed both to provide a sustainable yield of timber and for supplying a range of ecosystem services in line with the concept of sustainable forest management. Several incommensurable interests must then be considered, and it is necessary to strike a balance between different objectives. In evaluation of trade-offs to be made, both objective factors and subjective values need to be taken into account. In recent years, continuous cover forestry (CCF) has been put forward as an alternative to even-aged forestry. The aim of this study was to use scenario analysis in combination with multi criteria decision analysis (MCDA) to evaluate whether CCF is a suitable strategy based on the decision makers’ objectives and preferences for sustainable forest management in a specific landscape. This approach was applied to a planning case on the forest estate of the Linköping municipality in southwestern Sweden. The scenario analyses provided insights into relevant quantitative factors, while the MCDA evaluation helped in clarifying the objectives of the forest management and in assessing the relative importance of various objectives. The scenario analyses showed that in this case CCF is a good management strategy in ecological and social terms but yields worse economic outcomes than conventional even-aged forestry. In the Linköping case, there was a relatively strong emphasis on ecological and social aspects and thus, in summary, CCF seemed to be the most suitable option.
  • Nordström, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: eva-maria.nordstrom@slu.se (email)
  • Holmström, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: hampus.holmstrom@slu.se
  • Öhman, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: karin.ohman@slu.se
article id 1043, category Research article
Jorge Martín-García, Luc Barbaro, Julio Javier Diez, Hervé Jactel. (2013). Contribution of poplar plantations to bird conservation in riparian landscapes. Silva Fennica vol. 47 no. 4 article id 1043. https://doi.org/10.14214/sf.1043
Keywords: management; Populus x euramericana; clone I-214; hybrid; native
Highlights: Poplar plantations should not be used as surrogate habitat for native riparian forests with the aim of preserving bird species diversity; Native riparian forests should be preserved or restored as far as possible; Bird communities occurring in poplar plantations can still accommodate rich communities of forest bird species, providing that suitable management is applied at local and landscape levels.
Abstract | Full text in HTML | Full text in PDF | Author Info
In Mediterranean areas, riparian zones are particularly important for maintaining biodiversity. Nevertheless, the native vegetation in these zones has been modified or lost at an alarming rate during the last decades. The main objective of this study was to investigate the influence of poplar plantations on bird diversity in riparian zones, in order to estimate the ecological implications of a substantial expansion of poplar plantations. Breeding birds were sampled by the point-count method in twenty-four poplar plantations of I-214 clone, according to a factorial design combining stand age and understory management. Furthermore, the three native riparian forests remaining in the study area were also surveyed. Explanatory variables included (1) dendrometric, (2) understory and (3) landscape variables within six different radii of circular buffers. The species richness and abundance index were higher in riparian forests than in poplar plantations. Landscape variables (percentage of poplar plantations in the surrounding landscape) strongly influenced bird diversity in poplar plantations. Furthermore, at the local scale, understory cover was also a key factor in shaping bird assemblages. This suggests that poplar plantations should not be used as surrogates for native forests. Nevertheless, poplar plantations can still accommodate rich communities of forest bird species, providing that suitable management is applied at local and landscape levels.
  • Martín-García, Sustainable Forest Management Research Institute, University of Valladolid – INIA, Avenida Madrid, 57, 34004 Palencia, Spain E-mail: jorgemg@pvs.uva.es (email)
  • Barbaro, NRA, UMR 1202 BIOGECO, 69 Route d’Arcachon, F-33612 Cestas cedex, France E-mail: luc@pierroton.inra.fr
  • Diez, Sustainable Forest Management Research Institute, University of Valladolid – INIA, Avenida Madrid, 57, 34004 Palencia, Spain E-mail: jdcasero@pvs.uva.es
  • Jactel, NRA, UMR 1202 BIOGECO, 69 Route d’Arcachon, F-33612 Cestas cedex, France; Université de Bordeaux, UMR 1202, Bordeaux, F-33000 France E-mail: herve.jactel@pierroton.inra.fr
article id 1030, category Research article
Jussi Manner, Tomas Nordfjell, Ola Lindroos. (2013). Effects of the number of assortments and log concentration on time consumption for forwarding. Silva Fennica vol. 47 no. 4 article id 1030. https://doi.org/10.14214/sf.1030
Keywords: productivity; time study; forest haulage; forwarders; cut-to-length logging; log extraction; terrain transport
Highlights: We analysed the effects of total and forwarded log concentrations (m3 (100 m)–1) and the number of loaded assortments on forwarding; The combination of the number of loaded assortments and their abundance (i.e. forwarded log concentration) affected time consumption most; This knowledge enables improved efficiency by optimizing number and assortment proportions in the various loads required to forward a stand.
Abstract | Full text in HTML | Full text in PDF | Author Info
Forwarding has been carried out for 50 years, but much is still unknown about this work. Its complexity comes from both stand features and essential decision-making. Forwarding time consumption is influenced by e.g. log concentrations and number of assortments. Traditionally, focus has been on the total log concentration (TLC), referring to all logs at the harvesting site. However, we focused on forwarded log concentration (FLC), the load-specific log concentration which depends on the assortment distribution at harvesting site and the load-specific number of assortments. To evaluate the effects of TLC, number of assortments in a load and FLC on the loading and unloading times, a standardized field experiment was carried out. Pile and load sizes were constant, while TLC and FLC were manipulated by varying the pile distribution on the test path. For all work elements, the time consumption per m3 was significantly affected by the number of assortments that were loaded, but only the “driving while loading” work element was also significantly influenced by TLC. However, when untangling the intercorrelation between tested factors, it was found that the time consumption for driving while loading significantly decreased as a function of FLC and was unaffected by the number of assortments in a load. That FLC influences the forwarding time consumption highlights the need to study the effects of combining various assortment proportions in a load. Such knowledge will enable analysis of the most efficient number and assortment proportions to combine in the various loads required to forward a given stand.
  • Manner, Swedish University of Agricultural Sciences, Department of Forest Biomaterial and Technology, SE-901 83 Umeå, Sweden E-mail: jussi.manner@slu.se (email)
  • Nordfjell, Swedish University of Agricultural Sciences, Department of Forest Biomaterial and Technology, SE-901 83 Umeå, Sweden E-mail: tomas.nordfjell@slu.se
  • Lindroos, Swedish University of Agricultural Sciences, Department of Forest Biomaterial and Technology, SE-901 83 Umeå, Sweden E-mail: ola.lindroos@slu.se
article id 1022, category Research article
Eero Muinonen, Perttu Anttila, Jaakko Heinonen, Jukka Mustonen. (2013). Estimating the bioenergy potential of forest chips from final fellings in Central Finland based on biomass maps and spatially explicit constraints. Silva Fennica vol. 47 no. 4 article id 1022. https://doi.org/10.14214/sf.1022
Keywords: biomass; stumps; logging residues; remote sensing; forest energy
Abstract | Full text in HTML | Full text in PDF | Author Info
The technical potential of forest chips from final fellings in Central Finland was estimated using a method based on biomass maps derived from a multi-source forest inventory technique. Image segmentation techniques were applied to a satellite image mosaic to detect stand boundaries. The technical potential of forest chips was computed based on primary forestry residues, i.e. logging residues and stumps from final fellings. Harvesting level definitions for final fellings were established using realized statistics for roundwood at the municipality level as well as larger area statistics. The sensitivity of the potential to ecological and technical constraints in the model was also examined. The technical recovery rate of stump harvesting according to biomass harvesting guidelines was evaluated separately. The critical prerequisites for using the advanced, spatially explicit approach to analysing forest energy potentials may lie in the existence of spatially explicit forest inventory data and the biometric models for tree biomass assortments. The method applied was capable of taking into account the constraints that rely upon map data, such the actual forwarding distance or steepness of the slope in the terrain. The calculation results can be used for strategic decision making in the field of forest bioenergy production.
  • Muinonen, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: eero.muinonen@metla.fi (email)
  • Anttila, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: perttu.anttila@metla.fi
  • Heinonen, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jaakko.heinonen@metla.fi
  • Mustonen, Stora Enso, Talvikkitie 40 C, FI-01300 Vantaa, Finland E-mail: jukka.mustonen@storaenso.com
article id 1016, category Research article
Karin Johansson, Eva Ring, Lars Högbom. (2013). Effects of pre-harvest fertilization and subsequent soil scarification on the growth of planted Pinus sylvestris seedlings and ground vegetation after clear-felling. Silva Fennica vol. 47 no. 4 article id 1016. https://doi.org/10.14214/sf.1016
Keywords: nitrogen; establishment; regeneration; Scots pine; disc trenching; carryover; competing vegetation
Highlights: Pre-harvest N fertilization had no significant effect on seedling growth and ground vegetation biomass; Scarification improved seedling survival and growth and reduced the amount of ground vegetation; Without scarification, pre-harvest fertilization increased the amount of damaged seedlings.
Abstract | Full text in HTML | Full text in PDF | Author Info
Fertilization and scarification are both performed to increase tree production at different stages of forest rotation periods. In this study, the effects of previous nitrogen fertilizations and scarification after clear felling on planted Pinus sylvestris L. seedlings and ground vegetation were investigated. Two fertilization experiments established around 1980 were harvested in 2006, after which the plots were scarified by disc trenching and re-planted. The plots had been repeatedly fertilized over a 20-year period before harvesting, with total N doses of 0, 450, 900 or 1800 kg N ha-1. After five growing seasons, the growth, survival and nutrient contents of the seedlings were measured, and ground vegetation was collected to estimate its biomass and nutrient content. Pre-harvest fertilization alone had only minor effects on the results, but scarification increased both the survival and growth of the planted seedlings. However, without scarification, seedling mortality increased with increasing fertilization intensity. The ground vegetation biomass was higher in plots without scarification, but the total biomass of seedlings and ground vegetation was similar in all treatments. Scarification thus favored seedling growth at the expense of ground vegetation. Only a few effects on nutrient content were found, but there were no signs of nutrient imbalance in any of the treatments. At higher levels of fertilization, the K:N ratio in the seedlings decreased while the K content in the ground vegetation increased. Overall, scarification had a greater impact than pre-harvest fertilization on the planted seedlings and the ground vegetation.
  • Johansson, Skogforsk, The Forestry Research Institute of Sweden, Ekebo 2250, SE-268 90 Svalöv, Sweden & Southern Swedish Forest Research Centre, SLU, Box 49, SE-230 53 Alnarp, Sweden E-mail: karin.johansson@skogforsk.se (email)
  • Ring,  Skogforsk, The Forestry Research Institute of Sweden, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: eva.ring@skogforsk.se
  • Högbom,  Skogforsk, The Forestry Research Institute of Sweden, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: lars.hogbom@skogforsk.se
article id 1002, category Research article
Pentti Niemistö. (2013). Effect of growing density on biomass and stem volume growth of downy birch stands on peatland in Western and Northern Finland. Silva Fennica vol. 47 no. 4 article id 1002. https://doi.org/10.14214/sf.1002
Keywords: yield; thinning; silviculture; mortality; pulpwood; stand density; crown; energy-wood
Highlights: The thinning response in young downy birch stands was low and the mortality of merchantable stems was tolerable even in dense unthinned thickets during the rotation of 50 years; The production of above-ground woody biomass and stem volume or even the production of pulpwood increased with stand density, with these values being greatest for unthinned or very lightly thinned plots.
Abstract | Full text in HTML | Full text in PDF | Author Info
The objective of the study was to ascertain the relationship of thinning intensity of downy birch (Betula pubescens Ehrh.) stands with height, crown, and diameter development as well as pulpwood, stem volume, and biomass increment using long-term (20−30 years) field experiments. Diameter growth of birches increased with thinning intensity during the first 15 years from thinning in all development phases, though after that it did so only for the youngest stands. The thinning response was low. Thinning intensity had no influence on increase in height. In terms of stem volume with bark, the mortality in unthinned stands during the study period was 30–45 m3 ha-1. The mean stem number in unthinned birch thickets fell from 25 000 ha-1 at a dominant height of 7 m to 3000 ha-1 at 18 m. The stem volume increment over the first 15 years was highest (5–6 m3 ha-1 a-1) on the very lightly thinned or unthinned plots, but later there was no significant difference between initial thinning intensities. The maximum above-ground leafless biomass (over 100 Mg ha-1) was achieved on very lightly thinned plots. Also, the total production (including thinning removal) of biomass or stem volume or even the production of pulpwood increased with stand density, with these values being greatest for very lightly thinned or unthinned plots. During 50-year rotation, the highest leafless above-ground biomass production was 2.5 Mg ha-1 a-1 as a mean value from the experiments. The highest mean annual production of pulpwood (d > 6.5 cm) was 3.2 m3 ha-1 a-1, and, in practice, no saw timber or veneer timber was produced, because of the small size and low quality of the stems. A thinning in downy birch stands increased slightly the size of stems to be removed in future cuttings, but with exception for very light thinning it decreased the production of biomass and merchantable wood.
  • Niemistö, Finnish Forest Research Institute, Parkano Unit, FI-39700 Parkano, Finland E-mail: pentti.niemisto@metla.fi (email)
article id 938, category Research article
Lars Karlsson, Tommy Mörling, Urban Bergsten. (2013). Influence of silvicultural regimes on the volume and proportion of juvenile and mature wood in boreal Scots pine. Silva Fennica vol. 47 no. 4 article id 938. https://doi.org/10.14214/sf.938
Keywords: fibre length; stand density; juvenile wood
Highlights: Initial stand densities have a large impact on the proportion of mature wood within trees throughout most of their life cycle; The difference between regimes in volume of long fibres in crop trees could be substantial; Long-term silvicultural strategies implemented at juvenile stand ages can be important tools in order to produce wood raw material suited for specific end-uses.
Abstract | Full text in HTML | Full text in PDF | Author Info
Trees from 48 to 78 years old, exposed to three different long-term silvicultural regimes, were examined for transition ages between juvenile (JW), transition (TW) and mature wood (MW), total wood volume and proportions of the same wood types, as defined by fibre length. Twenty one sample trees were collected at sites with similar growing conditions within the same geographical area. Stem discs and fibre samples from breast height (BRH), 20% of tree height, green crown height and 70% of tree height were analysed. Trees growing in an environment with few neighboring trees (Sparse regime) started to produce MW, on average, five years later at BRH and six to nine years later at 20% of total tree height than trees in stands with high stem numbers (Dense regime) and trees growing in stands where the stem number had been heavily reduced after an initial high stand density (Dense/Sparse regime). For all regimes, the greatest mean fibre length was found below the green crown and high initial stem numbers were found to positively influence fibre length. The proportion of MW in the whole stem was 34% at Sparse regime sites, 57–69% at Dense/Sparse sites and 63–64% in sites where there was a Dense regime. The proportion of MW was significantly lower for trees from the Sparse regime on each stem section compared. In conclusion, the results suggest that the initial condition a tree faces affects the stem wood properties and quality output at the end of the rotation period.
  • Karlsson, Department of Forest Biomaterials and Technology, Swedish University of Agricultural Sciences, SE-901 83 Umeå, Sweden E-mail: lars.karlsson@slu.se (email)
  • Mörling, Department of Forest Ecology and Management, Swedish University of Agricultural Sciences, SE-901 83 Umeå, Sweden E-mail: tommy.morling@slu.se
  • Bergsten, Department of Forest Biomaterials and Technology, Swedish University of Agricultural Sciences, SE-901 83 Umeå, Sweden E-mail: urban.bergsten@slu.se
article id 1009, category Research article
Inger Sundheim Fløistad, Aksel Granhus. (2013). Timing and duration of short-day treatment influence morphology and second bud flush in Picea abies seedlings. Silva Fennica vol. 47 no. 3 article id 1009. https://doi.org/10.14214/sf.1009
Keywords: Norway spruce; photoperiod; autumn bud break; root collar diameter; second bud break; sturdyness
Highlights: The duration of short-day treatment, calculated as number of days, influenced the root collar diameter growth more than the timing of the treatment; If short-day treatment starts early in summer, a longer duration of the treatment is recommended to avoid second bud flush.
Abstract | Full text in HTML | Full text in PDF | Author Info
A slower reaction of diameter growth cessation compared to that of height growth in response to short day (SD) treatment is well documented in Picea abies (L.) Karst. seedlings, suggesting that the height/diameter ratio of seedlings could be controlled through appropriate timing and/or duration of SD treatment is forest nurseries. Here, we applied specific combinations of timing (starting date 20 and 27 June, 4 or 11 July) and duration (7, 10, 14 or 17 days) of SD treatment to assess the possibility of obtaining more sturdy seedlings. We observed a rapid and uniform height growth cessation following SD treatment compared with the delayed cessation of diameter growth. Height growth responded significantly only to starting date of SD treatment, resulting in taller seedlings for later starting dates. Diameter growth responded to the duration of SD treatment, with significantly less diameter growth in seedlings exposed to 14 or 17 days of SD treatment than in seedlings exposed to 7 or 10 days of SD treatment. Also starting date influenced diameter growth, resulting in significantly more diameter growth with the earliest starting date compared with the two latest starting dates of the SD treatment.  A second bud flush occurred only in seedlings exposed to SD treatment starting on 20 or 27 of June and only following 7-14 days of duration. This implies a need of longer duration if the SD treatment starts early. This will be at the expense of sustained diameter growth, thus compromising the objective of obtaining more sturdy seedlings.
  • Fløistad, Norwegian Institute for Agricultural and Environmental Research, Høgskolevn 7, N-1430 Ås, Norway & Norwegian Forest and Landscape Institute, P.O. Box 115, N-1431 Ås, Norway E-mail: isf@skogoglandskap.no (email)
  • Granhus, Norwegian Forest and Landscape Institute, P.O. Box 115, N-1431 Ås, Norway E-mail: aksel.granhus@skogoglandskap.no
article id 1006, category Research article
Jörn Rathke, Maria A. Huka, Manfred Gronalt. (2013). The box assignment problem in log yards. Silva Fennica vol. 47 no. 3 article id 1006. https://doi.org/10.14214/sf.1006
Keywords: log yard planning; sawmill; linear optimization
Highlights: Logistic approach for the optimization of log yard in terms of arrangement of storage boxes and ejection boxes reduced transportation time by 16 percent compared with the original solution.
Abstract | Full text in HTML | Full text in PDF | Author Info
This paper presents an optimization approach to minimizing log yard round wood transportation time for a medium sized hardwood sawmill. The log yard, which has to ensure a smooth raw material supply to the entire production process, is the first processing step in a sawmill. The log yard also serves as an internal round wood sorting and storing capacity. Thus, an optimal assignment of ejection boxes, storage boxes and feeding carriages is required to minimize transportation time at a log yard. The main contribution of this paper is to present an integrated approach which simultaneously takes into account log transportation time, storage capacity and yard crane deployment. The approach is based on two steps: a) defining storage spaces per batch and calculating distances and b) determining optimum box assignments in the log yard in order to minimize overall transportation distance. The solution in step b) is compared with the results obtained by random box assignment as well as a spreadsheet based planning method. We have been able to show that our approach is much more flexible and results are more than 16 percent better than the corresponding real life solution.
  • Rathke, University of Natural Resources and Life Sciences, Institute of Production and Logistics, Feistmantelstraße 4, 1180 Vienna, Austria E-mail: joern.rathke@boku.ac.at
  • Huka, University of Natural Resources and Life Sciences, Institute of Production and Logistics, Feistmantelstraße 4, 1180 Vienna, Austria E-mail: maria.huka@boku.ac.at (email)
  • Gronalt, University of Natural Resources and Life Sciences, Institute of Production and Logistics, Feistmantelstraße 4, 1180 Vienna, Austria E-mail: manfred.gronalt@boku.ac.at
article id 1005, category Research article
Marjut Turtiainen, Jari Miina, Kauko Salo, Juha-Pekka Hotanen. (2013). Empirical prediction models for the coverage and yields of cowberry in Finland. Silva Fennica vol. 47 no. 3 article id 1005. https://doi.org/10.14214/sf.1005
Keywords: generalized linear mixed model; abundance; berry yield; Vaccinium vitis-idaea L.
Highlights: The site fertility significantly affected the abundance of cowberry on mineral soils, spruce mires and pine mires; The stand basal area and dominant tree species were among the most important forest structural predictors in the model for the coverage; In the cowberry yield model developed for mineral soil sites, the stand basal area and coverage of cowberry plants were statistically significant predictors.
Abstract | Full text in HTML | Full text in PDF | Author Info
Empirical models for the coverage and berry yield of cowberry (Vaccinium vitis-idaea L.) were developed using generalized linear mixed models (GLMMs). The percentage coverage of cowberry was predicted as a function of site and stand characteristics using data from the Finnish National Forest Inventory (NFI) in 1995. The average annual yield, including the between-year variation in the yield, was predicted as a function of percentage coverage and stand characteristics using permanent experimental plots (MASI) established in different areas of Finland and measured in 2001-2012. The model for cowberry yields (Model 2) was developed for mineral soil forests. The model for the coverage (Model 1) was constructed so that it considers both mineral soil sites and also many other sites where cowberry occurs in the field layer. According to Model 1, the site fertility significantly affected the abundance of cowberry on mineral soils, spruce mires and pine mires. The stand basal area and dominant tree species were among the most important forest structural predictors in Model 1. The site fertility was not a significant predictor in the cowberry yield model. Instead, the stand basal area and coverage of cowberry plants were found to be statistically significant predictors in Model 2. The estimated models were used to predict the cowberry coverage, average annual yield and its 95 % confidence interval along with stand development. The models of this study can be used for multi-objective forest planning purposes.
  • Turtiainen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: marjut.turtiainen@uef.fi (email)
  • Miina, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jari.miina@metla.fi
  • Salo, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: kauko.salo@metla.fi
  • Hotanen, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: juha-pekka.hotanen@metla.fi
article id 1000, category Research article
Sören Wulff, Cornelia Roberge, Anna Hedström Ringvall, Sören Holm, Göran Ståhl. (2013). On the possibility to monitor and assess forest damage within large scale monitoring programmes – a simulation study. Silva Fennica vol. 47 no. 3 article id 1000. https://doi.org/10.14214/sf.1000
Keywords: forest health; forest inventory; environmental monitoring and assessment; forest condition
Abstract | Full text in HTML | Full text in PDF | Author Info
There is a growing demand for information on forest health due to fears that climate change may cause new kinds of damage that have not previously been encountered. In many cases, forest damage monitoring is conducted exclusively within sparse large-scale grids of sample plots and it is doubtful whether these are capable of providing relevant information to support mitigation programmes or other actions required to reduce economic losses due to damage outbreaks. In this study, we used simulated sampling to assess the precision of estimators related to forest state and changes in the damage sustained by trees within an area corresponding to the Swedish region Götaland, assuming a sampling design corresponding to that used in the Swedish National Forest Inventory (NFI) under different damage scenarios. Large and uniformly distributed damage outbreaks were well captured by an NFI-type inventory, but scattered damage outbreaks produced estimates with poor precision. As a consequence, we propose that there might be a need to revise current forest damage monitoring programmes to make them more useful for monitoring the kinds of damage that are likely to arise as a consequence of climate change.
  • Wulff, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: soren.wulff@slu.se (email)
  • Roberge, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: cornelia.roberge@slu.se
  • Ringvall, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: anna.ringvall@slu.se
  • Holm, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: soren.holm@slu.se
  • Ståhl, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: goran.stahl@slu.se
article id 993, category Research article
Jori Uusitalo, Jari Ala-Ilomäki. (2013). The significance of above-ground biomass, moisture content and mechanical properties of peat layer on the bearing capacity of ditched pine bogs. Silva Fennica vol. 47 no. 3 article id 993. https://doi.org/10.14214/sf.993
Keywords: logging; peatland; tree harvesting; trafficability
Abstract | Full text in HTML | Full text in PDF | Author Info
Intensive utilisation of peatland forests calls for logging activities to be increasingly carried out in conditions other than those in harsh winter. The low bearing capacity of peatlands forms a severe obstacle for the prevailing harvesting machinery. The aim of this study was to clarify and quantify the significance of above-ground biomass, brash mat, moisture content and mechanical properties of peat layer on the bearing capacity of pine bogs. In-situ driving tests were conducted in Western Finland on a pine bog covering a large variation of growing stock. Portable tools were tested for measuring strength properties of the top layer of peat. According to the results, shear modulus of top layer of peat, volume of trees and the existence of cutting debris are the most important factors affecting bearing capacity. Spiked shear vane was shown to be a promising tool in predicting the strength properties of peat soil.
  • Uusitalo, Finnish Forest Research Institute, Parkano Unit, Kaironiementie 15, FI-39700 Parkano, Finland E-mail: jori.uusitalo@metla.fi (email)
  • Ala-Ilomäki, Finnish Forest Research Institute, Vantaa Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: jari.ala-ilomaki@metla.fi
article id 973, category Research article
Sergio Rossi, Hubert Morin, François Gionest, Danielle Laprise. (2013). Spatially explicit structure of natural stands dominated by black spruce. Silva Fennica vol. 47 no. 3 article id 973. https://doi.org/10.14214/sf.973
Keywords: boreal forest; Picea mariana; mortality; recruitment; stand dynamics
Abstract | Full text in HTML | Full text in PDF | Author Info
Black spruce [Picea mariana (Mill.) BSP] regeneration emerges in clusters near the pre-existing boles within a few years after the passage of fire. This paper tested the hypothesis that black spruce forests still maintain the spatial structure deriving from postfire stand initiation. Trees and saplings were monitored during 2000-2007 and the horizontal and vertical structure of the stands were investigated on four permanent plots in the boreal forest of Quebec, Canada. Plots showed 1300-2150 trees ha-1, and were composed of trees with homogeneous sizes and a very small proportion of saplings. These characteristics identify single cohorts generated by complete, or almost-complete, stand replacement by fire. Ripley’s L(r) functions showed that the spatial pattern of trees and saplings ranged from random to aggregated, thus demonstrating that the clustering distribution of the individuals in black spruce forests can be maintained even after 80-120 years from stand initiation. These findings could results from incomplete self-thinning or from an environment with heterogeneous distribution of resources. The practices of ecosystem management recently developed in Eastern Canada should take into account both the horizontal and vertical structure to better modulate the competition among individuals during partial harvesting.
  • Rossi, Département des Sciences Fondamentales, Université du Québec à Chicoutimi, 555 boulevard de l’Université, Chicoutimi (QC), G7H2B1, Canada E-mail: sergio.rossi@uqac.ca (email)
  • Morin, Département des Sciences Fondamentales, Université du Québec à Chicoutimi, 555 boulevard de l’Université, Chicoutimi (QC), G7H2B1, Canada E-mail: hubert_morin@uqac.ca
  • Gionest, Département des Sciences Fondamentales, Université du Québec à Chicoutimi, 555 boulevard de l’Université, Chicoutimi (QC), G7H2B1, Canada E-mail: francois_gionest@uqac.ca
  • Laprise, Département des Sciences Fondamentales, Université du Québec à Chicoutimi, 555 boulevard de l’Université, Chicoutimi (QC), G7H2B1, Canada E-mail: danielle_laprise@uqac.ca
article id 972, category Research article
Outi H. Manninen, Rainer Peltola. (2013). Effects of picking methods on the berry production of bilberry (Vaccinium myrtillus), lingonberry (V. vitis-idaea) and crowberry (Empetrum nigrum ssp. hermaphroditum) in Northern Finland. Silva Fennica vol. 47 no. 3 article id 972. https://doi.org/10.14214/sf.972
Keywords: berry yields; commercial picking; wild forest berries
Highlights: Berry production of bilberry, lingonberry and crowberry was studied after picking the berries by plastic hand rake, long-handed metal rake, and powerful picking by long-handed metal rake; Berry production was not affected by the damage caused by any of the picking method; Current commercial picking methods do not endanger the berry production of the berry species at least in short-term.
Abstract | Full text in HTML | Full text in PDF | Author Info
The effect of commercial wild berry picking on berry yields is under a strong public debate in Finland. Especially high concern has been arisen over damages caused by metal rakes used in commercial picking to subsequent berry production. We studied the berry production of bilberry (Vaccinium myrtillus L.), lingonberry (V. vitis-idaea L.) and crowberry (E. nigrum ssp. hermaphroditum (Hagerup) Böcher) after picking the berries by 1) plastic hand rake, 2) long-handed metal rake and 3) powerful picking by long-handed metal rake, in northern Finland during 2010–2012. In the powerful long-handed metal rake treatment the aboveground vegetation was raked twice to the moss layer after berry picking. Biomass, which was removed from the vegetation by rakes was collected and used as a measure of the damage. We assumed that picking by plastic hand rake would result in lowest, long-handed metal rake intermediate and powerful picking by long-handed metal rake highest biomass loss from vegetation. The amount of biomass loss should in turn be reversely reflected into berry production. However, only the powerful picking by long-handed metal rake removed higher amount of biomass than other picking methods in bilberry and lingonberry. In crowberry, the amount of biomass removed by rakes increased from treatment to treatment. Contrary to our assumption, berry production of bilberry, lingonberry and crowberry was not affected by the damage caused by any of the picking method. We conclude that long-handed metal rake used in commercial picking is comparable to hand rake in terms of berry production.
  • Manninen, Agrifood Research Finland MTT, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: outi.h.manninen@mtt.fi (email)
  • Peltola, Agrifood Research Finland MTT, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: rainer.peltola@mtt.fi
article id 970, category Research article
Zhen-Yu Du, Qing-Hua Wang, Shang-Jun Xing, Fang-Chun Liu, Bing-Yao Ma, Hai-Lin Ma, De-Xi Liu. (2013). Fine root distribution, characteristics and rhizosphere soil properties in a mixed stand of Robinia pseudoacacia and Fraxinus velutina in a saline soil. Silva Fennica vol. 47 no. 3 article id 970. https://doi.org/10.14214/sf.970
Keywords: mixed forest; fine root; black locust; velvet ash; rhizhosphere soil
Abstract | Full text in HTML | Full text in PDF | Author Info
The spatial distribution and characteristics of fine roots (< 2 mm in diameter), and rhizosphere soil properties were studied in a mixed planted forest of black locust (Robinia pseudoacacia L.) and velvet ash (Fraxinus velutina Torr.) 27 years after planting in a coastal saline soil of the Yellow River delta, China. The results of fine root analysis showed that the fine roots of both black locust and velvet ash were mainly distributed in the soil layer at 0–20 cm depth and 50–150 cm from trees. The fine root distribution of both species suggests a strategy of avoiding salinity rather than salt –tolerance. The horizontal spread distance of fine roots of velvet ash was evidently longer than that of black locust. The fine root biomass, specific root length, specific root area, specific root volume and root activity were significantly higher for velvet ash in comparison with black locust. The results of soil analysis showed that rhizosphere soil pH of black locust and velvet ash were significantly lower compared with non-rhizosphere soil. The available N content in rhizosphere soil of black locust was higher than that of velvet ash. However, the contents of soluble salt, organic matter, available P and available K in rhizosphere soil of velvet ash were higher than those of black locust. The above results indicated that the differences between black locust and velvet ash in fine root distribution, characteristics and rhizosphere soil properties were the major reasons for that velvet ash showed stronger acclimation responses than black locust to the coastal saline soil.
  • Du, Shandong Academy of Forestry, 42 Wenhua East Road, Jinan 250014, P. R. China E-mail: zydu@qq.com (email)
  • Wang, Shandong Academy of Forestry, Jinan, P. R. China E-mail: wqh0228@163.com
  • Xing, Shandong Academy of Forestry, Jinan, P. R. China E-mail: xingsj-126@126.com
  • Liu, Shandong Academy of Forestry, Jinan, P. R. China E-mail: fchliu@126.com
  • Ma, Shandong Academy of Forestry, Jinan, P. R. China E-mail: mby777@163.com
  • Ma, Shandong Academy of Forestry, Jinan, P. R. China E-mail: mahlin@163.com
  • Liu, Shandong Academy of Forestry, Jinan, P. R. China E-mail: llyldx@163.com
article id 964, category Research article
Liisa Huttunen, Matthew P. Ayres, Pekka Niemelä, Susanne Heiska, Riitta Tegelberg, Matti Rousi, Seppo Kellomäki. (2013). Interactive effects of defoliation and climate change on compensatory growth of silver birch seedlings. Silva Fennica vol. 47 no. 3 article id 964. https://doi.org/10.14214/sf.964
Keywords: root; shoot ratio; biomass; Betula pendula; elevated CO2; elevated temperature; folivory
Highlights: The main components affecting growth compensation in silver birch seedlings are the timing and severity of foliage damage; The ability to compensate growth is also dependent upon the limits of temperature and nutrient availability; The responses of birches imply that folivory does not necessarily lead to reduced net productivity under changing climate
Abstract | Full text in HTML | Full text in PDF | Author Info
Atmospheric warming increases the abundance of insect herbivores and intensifies the risk of defoliation, especially in high latitude forests. At the same time, the effects of increasing temperature and CO2 on plant responses to foliage damage are poorly understood. We examined if previous-year defoliation, varying between 0 and 75% of total leaf area, and different combinations of elevated temperature, CO2 and nutrient availability alter the growth of two-year old silver birch (Betula pendula Roth) seedlings. We measured the greatest height growth in seedlings that were fertilized and defoliated twice at the level of 50% of total leaf area, and subjected to elevated temperature with ambient CO2. The lowest growth was recorded in unfertilized seedlings that were defoliated twice at the level of 25% of total leaf area, and grew under ambient temperature with ambient CO2. The total biomass increased in all seedlings that were fertilized or grew under elevated temperature. The root: shoot ratios were low in defoliated seedlings, or seedlings subjected to fertilization or temperature elevation. Our conclusion is that ability of birches to compensate height growth is highly dependent upon the magnitude and frequency of defoliation on the limits of temperature and nutrient availability. These responses imply that folivory does not necessarily lead to reduced net productivity of trees under changing climate.
  • Huttunen, Section of Ecology, Department of Biology, University of Turku, FI-20014 Turku, Finland E-mail: liisa.huttunen@utu.fi (email)
  • Ayres, Biological Sciences, Dartmouth College, Hanover, NH 03755, USA E-mail: matt.ayres@dartmouth.edu
  • Niemelä, Section of Biodiversity and Environmental Science, Department of Biology, University of Turku, FI-20014 Turku, Finland E-mail: pekka.niemela@utu.fi
  • Heiska, The Finnish Forest Research Institute, Punkaharju Unit, Finlandiantie 18, FI-58450 Punkaharju, Finland E-mail: susanne.heiska@metla.fi
  • Tegelberg, Digitarium - Digitization Centre of the Finnish Museum of Natural History and the University of Eastern Finland, Joensuu Science Park, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: riitta.tegelberg@helsinki.fi
  • Rousi, The Finnish Forest Research Institute, Vantaa Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: matti.rousi@metla.fi
  • Kellomäki, Faculty of Science and Forestry, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: seppo.kellomaki@uef.fi
article id 963, category Research article
Szymon Bijak, Michał Zasada, Agnieszka Bronisz, Karol Bronisz, Maciej Czajkowski, Łukasz Ludwisiak, Robert Tomusiak, Rafał Wojtan. (2013). Estimating coarse roots biomass in young silver birch stands on post-agricultural lands in central Poland. Silva Fennica vol. 47 no. 2 article id 963. https://doi.org/10.14214/sf.963
Keywords: abandoned farmland; belowground biomass; allometric equations; root-to-shoot ratio
Highlights: Age and size of the tree are the most important factors that influence the amount of belowground biomass; Allocation of the biomass to the coarse roots also depends on age and size of the tree
Abstract | Full text in HTML | Full text in PDF | Author Info
Study analyses coarse (d>2 mm of diameter) roots biomass dynamics in young succession stands of silver birch (Betula pendula Roth) growing on abandoned farmlands in central Poland. Research material based on 181 sample trees, which were gathered in 20 pure silver birch stands in 5 locations. The age of the trees varied from 1 to 16 years. Coarse roots biomass of the investigated trees ranged from 0.7 to 4305.5 g/tree (422.6 g/tree on average) showing great variability (coefficient of variation equals 185%). A clear dependence of belowground biomass on the age and size of a tree was observed. Root-to-shoot ratio values vary from 0.1 to 1.0 with evidence of a tendency to decrease with increasing age, diameter at the breast height and height of analysed trees. An allometric equation was elaborated for the estimation of belowground biomass based on height or diameter at breast height of young silver birches. The suitability of this formula should be considered for the estimation of biomass accumulation and carbon sequestration by young successional silver birch stands growing on abandoned agricultural lands.
  • Bijak, Warsaw University of Life Sciences – SGGW, Faculty of Forestry, Laboratory of Dendrometry and Forest Productivity, Nowoursynowska 159, 02-776 Warszawa, Poland E-mail: szymon.bijak@wl.sggw.pl (email)
  • Zasada, Warsaw University of Life Sciences – SGGW, Faculty of Forestry, Laboratory of Dendrometry and Forest Productivity, Nowoursynowska 159, 02-767 Warszawa, Poland E-mail: michal.zasada@wl.sggw.pl
  • Bronisz, Warsaw University of Life Sciences – SGGW, Faculty of Forestry, Laboratory of Dendrometry and Forest Productivity, Warszawa, Poland E-mail: agnieszka.bronisz@wl.sggw.pl
  • Bronisz, Warsaw University of Life Sciences – SGGW, Faculty of Forestry, Laboratory of Dendrometry and Forest Productivity, Warszawa, Poland E-mail: karol.bronisz@wl.sggw.pl
  • Czajkowski, Warsaw University of Life Sciences – SGGW, Faculty of Forestry, Laboratory of Dendrometry and Forest Productivity, Warszawa, Poland E-mail: maciej.czajkowski@wl.sggw.pl
  • Ludwisiak, Warsaw University of Life Sciences – SGGW, Faculty of Forestry, Laboratory of Dendrometry and Forest Productivity, Warszawa, Poland E-mail: lukasz.ludwisiak@wl.sggw.pl
  • Tomusiak, Warsaw University of Life Sciences – SGGW, Faculty of Forestry, Laboratory of Dendrometry and Forest Productivity, Warszawa, Poland E-mail: robert.tomusiak@wl.sggw.pl
  • Wojtan, Warsaw University of Life Sciences – SGGW, Faculty of Forestry, Laboratory of Dendrometry and Forest Productivity, Warszawa, Poland E-mail: rafal.wojtan@wl.sggw.pl
article id 958, category Research article
Back Tomas Ersson, Linus Jundén, Urban Bergsten, Martin Servin. (2013). Simulated productivity of one- and two-armed tree planting machines. Silva Fennica vol. 47 no. 2 article id 958. https://doi.org/10.14214/sf.958
Keywords: discrete-event simulation; mechanized planting; forestry; silviculture; automation; terrain model; root model
Highlights: Using discrete-event simulation and detailed terrain and machine models, the productivities of excavator-based one- and two-armed tree planting machines were simulated; The machines’ arms were equipped with one-and two-headed planting devices; Two planting heads per arm rather than two arms per base machine is better for increasing the productivity of intermittently advancing planting machines on Nordic clearcuts.
Abstract | Full text in HTML | Full text in PDF | Author Info
To increase mechanized planting, planting machine productivity must increase in order to improve cost-efficiency. To determine if excavators with two crane arms could potentially help to increase planting machine productivity under Nordic clearcut conditions, we modelled one-armed and semi-automated two-armed excavators with one- and two-headed planting devices. Using a recently developed tool for discrete-event simulation, these machine models then mounded and planted seedlings on terrain models with moraine soil having various frequencies of obstacles (stumps, roots and stones). Compared to if the two heads were mounted pairwise on only one arm, the results showed that productivity did not increase if two planting heads were attached individually to two separate crane arms. But productivity did increase if the planting machine had four planting heads mounted pairwise on two separate arms. However, despite assuming automated mounding and crane motion between planting spots, the two-armed, four-headed model never achieved high enough productivity levels to make it more cost-efficient than one-armed machines. The simulations illustrate that our terrain models generate realistic root architecture and boulder content distributions in moraine soil, while our machine models functionally describe mechanized planting work. Based on our assumptions, we conclude that further development work on two-armed excavator-based planting machines for Nordic clearcut conditions is not warranted. Our simulations reveal that increasing the number of planting heads per crane arm rather than number of crane arms per base machine offers the greatest potential to raise the productivity of intermittently advancing planting machines.
  • Ersson,  Swedish University of Agricultural Sciences, Department of Forest Biomaterials and Technology, SE-901 83 Umeå, Sweden E-mail: back.tomas.ersson@slu.se (email)
  • Jundén,  UMIT Research Lab, Umeå University, SE-901 87 Umeå, Sweden E-mail: linus.junden@gmail.com
  • Bergsten,  Swedish University of Agricultural Sciences, Department of Forest Biomaterials and Technology, SE-901 83 Umeå, Sweden E-mail: urban.bergsten@slu.se
  • Servin,  UMIT Research Lab, Umeå University, SE-901 87 Umeå, Sweden E-mail: martin.servin@physics.umu.se
article id 956, category Research article
Michał Zasada. (2013). Evaluation of the double normal distribution for tree diameter distribution modeling. Silva Fennica vol. 47 no. 2 article id 956. https://doi.org/10.14214/sf.956
Keywords: diameter distribution; simulated annealing; goodness-of-fit; Weibull distribution; maximum likelihood method; method of moments
Abstract | Full text in HTML | Full text in PDF | Author Info
The double normal distribution consists of two normal distributions truncated at their means and then combined in such a way, that points of truncation now become the overall distribution mode. So far, parameters of the double normal distribution have been estimated exclusively using the method of moments. This study evaluates the maximum likelihood method for the estimation of the double normal distribution parameters in Scots pine stands in Poland, and compares it to the results of the method of moments and the two-parameter Weibull distribution fitted using the maximum likelihood method and the method of moments. Presented results show that it is not recommended to use the maximum likelihood method of parameter fitting with Nelder-Mead and quasi-Newton optimization algorithms for the double normal distribution for small samples. However, it can be used for large samples, giving the fit comparable to the two-parameter Weibull distribution and providing parameters having sound practical and biological meaning. In the case of smaller samples for the double normal distribution it is recommended to apply the maximum likelihood method with the alternative simulated annealing optimization algorithm, use the method of moments or substitute the described distribution with more the flexible and robust Weibull distribution. For the smaller samples, the method of moments was superior to the maximum likelihood method.
  • Zasada, Warsaw University of Life Sciences, Faculty of Forestry, Laboratory of Dendrometry and Forest Productivity, Nowoursynowska 159, 02-776 Warsaw, Poland E-mail: michal.zasada@wl.sggw.pl (email)
article id 954, category Research article
Anna-Maria Eriksson, Jörgen Olsson, Bengt Gunnar Jonsson, Sara Toivanen, Mattias Edman. (2013). Effects of restoration fire on dead wood heterogeneity and availability in three Pinus sylvestris forests in Sweden. Silva Fennica vol. 47 no. 2 article id 954. https://doi.org/10.14214/sf.954
Keywords: Scots pine; CWD; prescribed burning; decay stage; charred wood
Abstract | Full text in HTML | Full text in PDF | Author Info
Restoration fires are increasingly used as a conservation tool in Sweden to recreate forests with characteristics of previous forests that were periodically disturbed by fires and promote fire-dependent species. Restoration fires can result in large inputs of fresh dead wood, but there are risks of losing some of the existing, pre-fire dead wood. To assess these counteracting effects we studied the heterogeneity and availability of dead wood before and after three restoration fires in boreal Scots pine forests. Specifically, we studied volumes of stumps, high stumps, snags and logs. The fires decreased the total volume of pre-fire dead wood (23-41%) and consumed logs in late decay stages (26-54%) to a higher extent than logs in earlier stages. The input of new fresh dead wood after the fires exceeded losses of pre-fire dead wood and resulted in a net increase of dead wood in all three sites. The added dead wood consisted of fresh snags killed by the fires. Fire also affected log characteristics: reducing their vegetation coverage (60-98%), decreasing their ground contact (4-50%) and increasing their surface area of charred wood (>50%). Such changes have important consequences for the micro environmental conditions inside logs, but have been rarely studied in relation to restoration fires. Our results show that restoration fire causes changes in dead wood availability and characteristics of logs. The results imply that ideally stands with low abundance of rare and heavily decayed wood substrates should be burned to optimize dead wood values. Alternatively, management practices should include protection of these substrates during restoration fires.
  • Eriksson, Department of Natural Sciences, Mid Sweden University, SE-851 70, Sundsvall, Sweden E-mail: anna-maria.eriksson@miun.se (email)
  • Olsson, Department of Ecology and Environmental Science, Umeå University, SE-901 87, Umeå, Sweden E-mail: jorgen.m.olsson@slu.se
  • Jonsson, Department of Natural Sciences, Mid Sweden University, SE-851 70, Sundsvall, Sweden E-mail: bengt-gunnar.jonsson@miun.se
  • Toivanen, Department of Natural Sciences, Mid Sweden University, SE-851 70, Sundsvall, Sweden E-mail: sara.toivanen@lansstyrelsen.se
  • Edman, Department of Natural Sciences, Mid Sweden University, SE-851 70, Sundsvall, Sweden E-mail: mattias.edman@miun.se
article id 952, category Research article
Lauri Korhonen, Inka Pippuri, Petteri Packalén, Ville Heikkinen, Matti Maltamo, Juho Heikkilä. (2013). Detection of the need for seedling stand tending using high-resolution remote sensing data. Silva Fennica vol. 47 no. 2 article id 952. https://doi.org/10.14214/sf.952
Keywords: forest management; airborne laser scanning; logistic regression; seedling stand; tending; support vector machine
Abstract | Full text in HTML | Full text in PDF | Author Info
Seedling stands are problematic in airborne laser scanning (ALS) based stand level forest management inventories, as the stem density and species proportions are difficult to estimate accurately using only remotely sensed data. Thus the seedling stands must still be checked in the field, which results in an increase in costs. In this study we tested an approach where ALS data and aerial images are used to directly classify the seedling stands into two categories: those that involve tending within the next five years and those which involve no tending. Standard ALS-based height and density features, together with texture and spectral features calculated from aerial images, were used as inputs to two classifiers: logistic regression and the support vector machine (SVM). The classifiers were trained using 208 seedling plots whose tending need was estimated by a local forestry expert. The classification was validated on 68 separate seedling stands. In the training data, the logistic model’s kappa coefficient was 0.55 and overall accuracy (OA) 77%. The SVM did slightly better with a kappa = 0.71 and an OA = 86%. In the stand level validation data, the performance decreased for both the logistic model (kappa = 0.38, OA = 71%) and the SVM (kappa = 0.37, OA = 72%). Thus our approach cannot totally replace the field checks. However, in considering the stands where the logistic model predictions had high reliability, the number of misclassifications reduced drastically. The SVM however, was not as good at recognizing reliable cases.
  • Korhonen, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: lauri.korhonen@uef.fi (email)
  • Pippuri, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: inka.pippuri@uef.fi
  • Packalén, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: petteri.packalen@uef.fi
  • Heikkinen, School of Computing, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: ville.heikkinen@uef.fi
  • Maltamo, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: matti.maltamo@uef.fi
  • Heikkilä, Finnish Forest Centre, Public Services, Maistraatinportti 4 A, FI-00240 Helsinki, Finland E-mail: juho.heikkila@metsakeskus.fi
article id 935, category Research article
Loice M. A. Omoro, Mike Starr, Petri K. E. Pellikka. (2013). Tree biomass and soil carbon stocks in indigenous forests in comparison to plantations of exotic species in the Taita Hills of Kenya. Silva Fennica vol. 47 no. 2 article id 935. https://doi.org/10.14214/sf.935
Keywords: biomass; carbon stocks; soil carbon; Eastern Arc Montane forest; exotic tree species; indigenous forest
Abstract | Full text in HTML | Full text in PDF | Author Info
Carbon (C) densities of the tree biomass and soil (0–50 cm) in indigenous forest and plantations of eucalyptus, cypress and pine in the Taita Hills, Kenya were determined and compared. The cypress and pine plantations were about 30-years-old and eucalyptus plantations about 50-years-old. Biomass C densities were estimated from breast height diameter and wood density using allometric functions developed for tropical species and an assumed C content of 50%. Belowground biomass C densities were estimated using root:shoot biomass ratios. Soil organic C (SOC) densities were calculated from measured organic carbon contents (0–20 and 20–50 cm layers) and modelled bulk density values. Mean total biomass C and SOC densities for indigenous forest were greater than those of the plantations, and the difference was significant (p < 0.05) in the cases of cypress and pine biomass and pine SOC. The correlation between biomass C and SOC densities was nearly significant in the case of indigenous forest, but negative. Biomass C densities were not significantly correlated with mean annual precipitation, mean annual temperature or potential evapotranspiration, but pine biomass C densities were significantly correlated to actual evapotranspiration. SOC densities were more strongly correlated to mean annual precipitation than biomass C densities, but only significantly so in the case of pine. Neither biomass C nor SOC densities were correlated to plant available water capacity of the soil. Indigenous forest SOC densities were significantly correlated to soil clay contents, but negatively. Indigenous forests sequester more C in biomass and soil than do 30 to 50-year-old plantations of exotics, but it remains unclear if this is an intrinsic difference between indigenous forest and plantations of exotics or because of insufficient time for SOC levels in plantations to recover after clearance of original indigenous forest.
  • Omoro, Viikki Tropical Resources Institute, Department of Forest Sciences, P.O. Box 27 (Latokartanonkaari 7), FI-00014 University of Helsinki, Finland E-mail: loice.omoro@helsinki.fi
  • Starr, Department of Forest Sciences, P. O. Box 27 (Latokartanonkaari 7), FI-00014 University of Helsinki, Finland E-mail: mike.starr@helsinki.fi (email)
  • Pellikka, Department of Geosciences and Geography, P. O. Box 64 (Gustaf Hällströminkatu 2), FI-00014 University of Helsinki, Finland E-mail: petri.pellikka@helsinki.fi
article id 933, category Research article
Per-Ola Hedwall, Harald Grip, Sune Linder, Lars Lövdahl, Urban Nilsson, Johan Bergh. (2013). Effects of clear-cutting and slash removal on soil water chemistry and forest-floor vegetation in a nutrient optimised Norway spruce stand. Silva Fennica vol. 47 no. 2 article id 933. https://doi.org/10.14214/sf.933
Keywords: Picea abies; logging residues; forest fertilisation; forest undergrowth; N-retention; nutrient leakage; whole-tree harvest
Abstract | Full text in HTML | Full text in PDF | Author Info
Fertilisation with nutrient optimisation has in Sweden resulted in large increases in volume growth in young stands of Norway spruce. There are, however, environmental concerns about repeated fertilisation and one is the risk of nutrient leakage to ground water resources and aquatic ecosystems after clear-cutting of such forests. The present study followed soil-water chemistry in optimised fertilised stands after clear-cutting, as well as effects of harvest of slash on nutrient leakage. Parts of a 30-year-old stand of Norway spruce, which had been subject to a nutrient optimisation experiment for 17 years, were clear-cut. A split-plot design with whole-tree harvesting as the sub-plot treatment was applied. Lysimeters were installed and soil-water sampled at nine occasions during the following four years. No significant effects of fertilisation on nitrate leaching were found, while harvest of slash reduced the concentration of Ca, DOC, DON, K, Mg, ammonium and nitrate, as well as pH in the soil solution. While no effects of fertilisation could be seen on the soil water concentration of N, the results indicate an interaction between fertilisation and harvest of slash on the concentration of nitrate in the soil solution. The results indicate that forest-floor vegetation plays an important role in the retention of N after clear-cutting of fertilised forests.
  • Hedwall, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences (SLU), P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: per-ola.hedwall@slu.se (email)
  • Grip, Department of Forest Ecology and Management, SLU, SE-901 83 Umeå, Sweden E-mail: harald@grip2.se
  • Linder, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences (SLU), P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: sune.linder@slu.se
  • Lövdahl, Department of Forest Ecology and Management, SLU, SE-901 83 Umeå, Sweden E-mail: ll@nn.se
  • Nilsson, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences (SLU), P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: urban.nilsson@slu.se
  • Bergh, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences (SLU), P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: johan.bergh@slu.se
article id 932, category Research article
Jun Wang, Le Shi, Shaoyu Song, Ju Tian, Xiangyang Kang. (2013). Tetraploid production through zygotic chromosome doubling in Populus. Silva Fennica vol. 47 no. 2 article id 932. https://doi.org/10.14214/sf.932
Keywords: Populus; colchicine; high temperature; seed hair; tetraploid induction; zygote
Abstract | Full text in HTML | Full text in PDF | Author Info
The most direct approach in breeding triploid Populus is crossing allotetraploids with diploids. However, the lack of allotetraploid Populus restricts application of this approach. In this investigation, zygotic chromosome doubling was induced with colchicine and high temperatures to produce ((Populus pseudo-simonii × P. nigra ‘Zheyin3#’) × (P. × beijingensis)) allotetraploids. We screened 6 and 25 tetraploid individual offspring from the colchicine and high-temperature treatments respectively, indicating that both colchicine and high temperature are effective for tetraploid production by zygotic chromosome doubling of Populus. Developmental characteristics of seed hairs in the ovaries were temporally associated with zygotic development, which was used to successfully guide the colchicine and high-temperature treatments. During certain stages of hair development, the efficiency of tetraploid production was significantly high. However, efficiency of production was not significantly influenced by other factors, i.e. colchicine concentration, temperature or duration of high-temperature treatment. Size and frequency of leaf stomata between tetraploid and diploid plants were significantly different, suggesting that this character can be altered via genomic increase in material. The allotetraploids produced in this investigation, having different genotypes, provide important parental germplasms for further triploid breeding.
  • Wang, National Engineering Laboratory for Tree Breeding; Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants, Ministry of Education; College of Biological Sciences and Technology, Beijing Forestry University, Beijing 100083, P.R. China E-mail: wangjun@bjfu.edu.cn (email)
  • Shi, National Engineering Laboratory for Tree Breeding; Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants, Ministry of Education; College of Biological Sciences and Technology, Beijing Forestry University, Beijing 100083, P.R. China E-mail: 7320932@qq.com
  • Song, National Engineering Laboratory for Tree Breeding; Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants, Ministry of Education; College of Biological Sciences and Technology, Beijing Forestry University, Beijing 100083, P.R. China E-mail: angela-song@foxmail.com
  • Tian, National Engineering Laboratory for Tree Breeding; Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants, Ministry of Education; College of Biological Sciences and Technology, Beijing Forestry University, Beijing 100083, P.R. China E-mail: 1215813245@qq.com
  • Kang, National Engineering Laboratory for Tree Breeding; Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants, Ministry of Education; College of Biological Sciences and Technology, Beijing Forestry University, Beijing 100083, P.R. China E-mail: kangxy@bjfu.edu.cn
article id 930, category Research article
Raffaele Spinelli, Eugenio Cavallo, Lars Eliasson, Alessio Facello. (2013). Comparing the efficiency of drum and disc chippers. Silva Fennica vol. 47 no. 2 article id 930. https://doi.org/10.14214/sf.930
Keywords: productivity; biomass; wood; energy; fuel
Abstract | Full text in HTML | Full text in PDF | Author Info
The study compared the effect of chipper type on productivity, power demand, fuel consumption and product quality. Tests were conducted on two commercial chipper models, a disc and a drum chipper. Both chippers had the same diameter capacity, were applied to the same tractor and fed with the same feedstock types. Fifteen replications were conducted per machine and for each of four different feedstock types, reaching a total of 120 tests. The disc chipper had a higher energy efficiency and used 19% less fuel per unit product, possibly due to its simpler design, integrating comminuting and discharge system in one synergic device. In contrast, the drum chipper was 8% more productive, since it cut with the same energy all along the length of its knives. The drum chipper produced smaller chips, with a higher incidence of fines. Feedstock type had a strong effect on productivity, energy efficiency and product quality. The effect of feedstock type was mainly related to piece size, and may be stronger than the effect of chipper type. Further studies should determine the effect of blade wear on the relative performance of the two chipper types.
  • Spinelli, CNR IVALSA, Via Madonna del Piano 10, Sesto Fiorentino (FI), Italy E-mail: spinelli@ivalsa.cnr.it (email)
  • Cavallo, CNR IVALSA, Via Madonna del Piano 10, Sesto Fiorentino (FI), Italy E-mail: e.cavallo@imamoter.cnr.it
  • Eliasson, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: lars.eliasson@skogforsk.se
  • Facello, CNR IVALSA, Via Madonna del Piano 10, Sesto Fiorentino (FI), Italy E-mail: a.facello@ima.to.cnr.it
article id 928, category Research article
Rune Simonsen. (2013). Optimal regeneration method – Planting vs. natural regeneration of Scots pine in northern Sweden. Silva Fennica vol. 47 no. 2 article id 928. https://doi.org/10.14214/sf.928
Keywords: optimisation; silviculture; genetically improved planting material
Highlights: Two regeneration methods were modelled on stand level and optimised numerically to maximise present value for a range of site indexes and locations; Natural regeneration was optimal in most cases; Planting was optimal for high site indexes, low rate of seedling mortality and for low discount rates; Using genetically improved plant material greatly shifts the preference towards planting
Abstract | Full text in HTML | Full text in PDF | Author Info
In this study the profitability of regenerating Scots pine (Pinus sylvestris L.) was examined for two methods; planting and natural regeneration with seed trees. The methods were modelled on stand level and optimised numerically using nonlinear optimisation. The analysis includes 7 site indexes, 16 to 28 expressed as dominant height in meters at an age of 100 years; and 8 localities in northern Sweden distributed on two latitudes, 60°N and 64°N and four altitudes, 100 to 400 m.a.s.l. Furthermore, two scenarios of genetically improved planting material were examined. The results show that the optimal choice of regeneration method depends on the location, site index and discount rate. Considering the same genetic regeneration material, natural regeneration was the optimal method for most of the evaluated sites. Planting was optimal only for stands of high site index and low rate of seedling mortality, which is associated with localities on low altitudes. The break even site index, where the two methods yielded the same net present value, was 27 on average (25 to 28). The choice between the two regeneration methods was found to be more economically important when the discount rate was low and for low site indexes. The option of using genetically improved plant material shift the preference towards planting. Thus, the two levels of genetic gain of +4% and +10% to maximum mean annual increment resulted in an average break even site index of 25 and 21 respectively.
  • Simonsen, Department of Forest Economics, SLU, SE-901 83 Umeå, Sweden E-mail: rune.simonsen@slu.se (email)
article id 925, category Research article
Steen Magnussen. (2013). An assessment of three variance estimators for the k-nearest neighbour technique. Silva Fennica vol. 47 no. 1 article id 925. https://doi.org/10.14214/sf.925
Keywords: forest inventory; simple random sampling; resampling estimators; bootstrap; jackknife; difference estimator; cluster sampling
Abstract | Full text in HTML | Full text in PDF | Author Info
A jackknife (JK), a bootstrap (BOOT), and an empirical difference estimator (EDE) of totals and variance were assessed in simulated sampling from three artificial but realistic complex multivariate populations (N = 8000 elements) organized in clusters of four elements. Intra-cluster correlations of the target variables (Y) varied from 0.03 to 0.26. Time-saving implementations of JK and BOOT are detailed. In simple random sampling (SRS), bias in totals was ≤ 0.4% for the two largest sample sizes (n = 200, 300), but slightly larger for n = 50, and 100. In cluster sampling (CLU) bias was typically 0.1% higher and more variable. The lowest overall bias was in EDE. In both SRS and CLU, JK estimates of standard error were slightly (3%) too high, while the bootstrap estimates in both SRS and CLU were too low (8%). Estimates of error suggested a trend in EDE toward an overestimation with increasing sample size. Calculated 95% confidence intervals achieved a coverage that in most cases was fairly close (± 2%) to the nominal level. For estimation of a population total the EDE estimator appears to be slightly better than the JK estimator.
  • Magnussen, Canadian Forest Service, Natural Resources Canada, 505 West Burnside Road, Victoria BC V8Z 1M5 Canada E-mail: steen.magnussen@nrcan.gc.ca (email)
article id 904, category Research article
Raul Fernandez-Lacruz, Fulvio Di Fulvio, Dan Bergström. (2013). Productivity and profitability of harvesting power line corridors for bioenergy. Silva Fennica vol. 47 no. 1 article id 904. https://doi.org/10.14214/sf.904
Keywords: accumulating felling head; clearing; maintenance; motor-manual; power grid; wood fuel
Abstract | Full text in HTML | Full text in PDF | Author Info
Trees growing under the wires and around the pylons carrying power lines (PL) represent a significant threat to the power supply because they can cause power outages and damage. The vegetation in these PL corridors is cleared motor-manually on a regular basis, which represents about 50% of the PL maintenance costs. In Sweden, PL corridors account for 140 000 ha of productive forest land, with an estimated bioenergy potential of 3 TWh/year. The aim of this study was to measure the productivity of a harvester (with an accumulating felling head) and a forwarder, performing PL corridor clearing (with the collection of whole trees for energy use) and to calculate how the cost and economic profitability is dependent on tree height, biomass removal, harvested area, forwarding distance and wood fuel price. The study also compared the economic profitability of the mechanized harvesting system with motor-manual clearing. Experimental units were inventoried along a PL corridor in central Sweden and a time study of one harvester and one forwarder (with a single operator per machine), working in those units, was carried out. The results showed that if the tree height was greater than about 6 m, the mechanized harvesting system became a more cost-efficient alternative, when compared to motor-manual clearing, but it was also found that mechanized clearing is not always the most cost-effective option. Nevertheless, mechanization of PL clearing has a huge potential for expansion, requiring further research in the combined management of the PL corridors and side areas.
  • Fernandez-Lacruz, Department of Forest Biomaterials and Technology, Swedish University of Agricultural Sciences, SE-901 83 Umeå, Sweden E-mail: raul.fernandez@slu.se (email)
  • Di Fulvio, Department of Forest Biomaterials and Technology, Swedish University of Agricultural Sciences, SE-901 83 Umeå, Sweden E-mail: fulvio.di.fulvio@slu.se
  • Bergström, Department of Forest Biomaterials and Technology, Swedish University of Agricultural Sciences, SE-901 83 Umeå, Sweden E-mail: dan.bergstrom@slu.se
article id 903, category Research article
Mikko Hyppönen, Ville Hallikainen, Juhani Niemelä, Pasi Rautio. (2013). The contradictory role of understory vegetation on the success of Scots pine regeneration. Silva Fennica vol. 47 no. 1 article id 903. https://doi.org/10.14214/sf.903
Keywords: site preparation; seedling establishment; seedling mortality; field- and ground-layer vegetation; modelling approach; initial growth; prescribed burning
Abstract | Full text in HTML | Full text in PDF | Author Info
In North-East Finland, severe problems have been encountered in the natural regeneration of Scots pine (Pinus sylvestris L.) on sites where regeneration through site preparation usually is quite successful. We hypothesized that in that area understory vegetation, especially heather (Calluna vulgaris), crowberry (Empetrum hermaphroditum), mosses and lichens, could play a key role in this pattern. We found that in general, ground- and field-layer vegetation tends to be in a negative relationship with the establishment, growth and survival of pine seedlings. Some positive relationships were also observed. Lingonberry (Vaccinium vitis-idea) tended to improve seedling height growth. Heather, instead, seemed to have a contradictory role. It was positively related to seedling establishment but negatively to seedling growth. This dual role raises further questions about the primary reasons for the regeneration problems in North-East Finland. All in all, our results suggest that conventional methods of forest regeneration in these kinds of areas are not always effective enough and additional measures are needed. These might include severe prescribed burning along with site preparation in order to decrease the impact of the dominant ground- and field-layer vegetation on the success of Scots pine regeneration.
  • Hyppönen, Finnish Forest Research Institute, Rovaniemi Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: mikko.hypponen@metla.fi (email)
  • Hallikainen, Finnish Forest Research Institute, Rovaniemi Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: ville.hallikainen@metla.fi
  • Niemelä, Finnish Forest Research Institute, Rovaniemi Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: jn@nn.fi
  • Rautio, Finnish Forest Research Institute, Rovaniemi Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: pasi.rautio@metla.fi
article id 902, category Research article
Sakari Tuominen, Reija Haapanen. (2013). Estimation of forest biomass by means of genetic algorithm-based optimization of airborne laser scanning and digital aerial photograph features. Silva Fennica vol. 47 no. 1 article id 902. https://doi.org/10.14214/sf.902
Keywords: forest inventory; airborne laser scanning; biomass modelling
Abstract | Full text in HTML | Full text in PDF | Author Info
Information on forest biomass is required for several purposes, including estimation of forest bioenergy resources and forest carbon stocks. Airborne laser scanning is today considered the most accurate remote sensing method for forest inventory. The three-dimensional nature of laser scanning data enables estimation of the volumes of the tree canopies. The dimensions of the tree canopies show high correlation with the amount of forest biomass. Optical aerial photographs are often used to complement laser data, for improved distinction between tree species. The paper reports on a study testing the estimation of forest biomass variables in two study areas in Southern Finland. The biomass variables were derived on the basis of tree-level field measurements, with biomass models used for pine, spruce, and birch. The sample-plot-level biomass components were derived on the basis of tree-level data and used as reference data for airborne-laser- and aerial‑photograph-based estimation. Results were slightly better for total biomass (RMSE 22.5% and 23.6% for the two study areas) than total volume (RMSE: 23.4% and 26.1%). Species-specific estimation errors were large in general but varied between the study areas, because of differences in their forest structures.
  • Tuominen, Finnish Forest Research Institute, P.O.Box 18, FI-01301 Vantaa, Finland E-mail: sakari.tuominen@metla.fi (email)
  • Haapanen, Finnish Forest Research Institute, P.O.Box 18, FI-01301 Vantaa, Finland E-mail: reija.haapanen@gmail.com
article id 901, category Research article
Luis A. Apiolaza, Rosa M. Alzamora. (2013). Building deployment portfolios for genotypes under performance instability. Silva Fennica vol. 47 no. 1 article id 901. https://doi.org/10.14214/sf.901
Keywords: Pinus radiata; wood quality; breeding objectives; clonal portfolio; deployment portfolio; genotype by environment interaction
Abstract | Full text in HTML | Full text in PDF | Author Info
We used portfolio theory to analyze the tradeoffs between returns and performance instability of deployment units for Pinus radiata D. Don. We considered three groups of 34 trees each grown to produce appearance lumber, structural lumber, or both. Risk was based on the variability of tree returns in scenarios of changing volume, wood stiffness and presence of resin defects due to genotype by environment interaction inducing both changes of scale and differential tree response to environmental scenarios. The return of structural trees was highly variable with a mean of 3.11 NZ $/stem/year, followed by appearance-structural trees (3.48 NZ $/stem/year). In contrast, appearance trees had the lowest returns (1.99 NZ $/stem/year) and variability. The portfolio model selected structural trees in high-risk scenarios, but selection was apportioned between structural and appearance-structural trees as the risk decreased. The model selected only appearance trees for high-risk aversion. The analysis also considered silvicultural regimes, where the appearance-structural regime was selected under high variability. As risk decreased the appearance grades regime was also selected. The structural regime was rarely selected due to the variability of stiffness between trees. Using genotypes improved for stiffness could increase the expected value and reduce variability for structural purposes, making the structural regime more appealing.
  • Apiolaza, School of Forestry, University of Canterbury, Private Bag 4800, 8042 Canterbury, New Zealand E-mail: luis.apiolaza@canterbury.ac.nz (email)
  • Alzamora, Instituto de Manejo Forestal, Universidad Austral, Valdivia, Chile E-mail: ralzamor@uach.cl
article id 899, category Research article
Franz Holzleitner, Christian Kanzian, Norbert Höller. (2013). Monitoring the chipping and transportation of wood fuels with a fleet management system. Silva Fennica vol. 47 no. 1 article id 899. https://doi.org/10.14214/sf.899
Keywords: forest fuel; supply chain; monitoring; logistic; fleet management; wood chips
Abstract | Full text in HTML | Full text in PDF | Author Info
Controlling and organizing the complex forest-to-consumer supply chain of wood fuels is a challenging task, especially for the chipping and transport processes. Truck mounted chippers and transport trailer-trucks must be scheduled to minimize delay to be profitable. Job management within the supply chain, including machine activity based controlling, offers a new way to increase efficiency and productivity. However, detailed data are required to detect and analyze potential gaps and improve forest fuel supply. Generally, data regarding the wood fuel supply chain process are obtained from extensive time studies that are based on a specific process step. Although time studies can detect details during the production of forest fuels, they only describe certain time frames. Long-term data that are recorded during the entire year could encompass seasonal and short term effects. This study aims to monitor the forest fuel supply processes (semi-automated), specifically regarding time and fuel consumption. Large data sets were automatically and efficiently gathered with little effort by drivers and operators. Data were recorded with fleet management equipment for more than 14 months. Vehicle data, including GPS data, were logged at an interval of one minute. Data management was conducted in a pre-configured database that contained pre-defined reports and were run by the Institute of Forest Engineering, Vienna. Work step assignments were implemented with Structured Query Language (SQL)-routines by using the raw machine activities data and GPS. The chipping and transport activities of more than 240 loads were analyzed by focusing on fuel consumption, time needed and traffic. The average distance between chipping sites and plants was approximately 54 kilometers. Fuel consumption from transport reached 50 l/100 km. The chipping unit reached a productivity of 12.8 odt/PSH15 and had a fuel consumption of 58 liters per operating hour.
  • Holzleitner, Institute of Forest Engineering, Department of Forest and Soil Sciences, University of Natural Resources and Life Sciences, Peter Jordanstrasse 82/3, 1190 Vienna, Austria E-mail: franz.holzleitner@boku.ac.at (email)
  • Kanzian, Institute of Forest Engineering, Department of Forest and Soil Sciences, University of Natural Resources and Life Sciences, Peter Jordanstrasse 82/3, 1190 Vienna, Austria E-mail: christian.kanzian@boku.ac.at
  • Höller, Institute of Forest Engineering, Department of Forest and Soil Sciences, University of Natural Resources and Life Sciences, Peter Jordanstrasse 82/3, 1190 Vienna, Austria E-mail: norbert.hoeller@boku.ac.at
article id 894, category Research article
Heimo Karppinen, Mika Korhonen. (2013). Do forest owners share the public’s values? An application of Schwartz’s value theory. Silva Fennica vol. 47 no. 1 article id 894. https://doi.org/10.14214/sf.894
Keywords: demographics; family forest owners; objectives of forest ownership; the public; Schwartz’s value theory
Abstract | Full text in HTML | Full text in PDF | Author Info
The purpose of this study is to investigate the differences in value priorities between Finnish forest owners and the general public. A conclusion is drawn whether and to what extent value changes in society are reflected in forest owners’ values and objectives, and, finally, in their actual forestry behavior. In addition, the study highlights the differences in value priorities among forest owners in various demographic groups. The data set used in this study was based on a nationwide mail survey on Finnish non-industrial private forest owners conducted in 2009 and consisting of 2116 observations of forest owners. Schwartz’s value theory was a good fit for testing the value priorities of forest owners. The three most important values were benevolence, security and conformity, both among the forest owners and the public. Tradition was ranked the fourth most important value by the forest owners, but very low by the public. The forest owners ranked universalism slightly lower than the public in general. This difference was clearly greater when the female forest owners were compared to women in the whole population. The probability of a forest owner belonging to the Softies (high emphasis on universalism and benevolence) increased with age and was higher for the female owners and the owners with recreational or multiple objectives compared to the indifferent owners. The multiobjective owners and recreationists had relatively similar value profiles. The previous literature suggests that multiobjective owners are the most active forest owner group and that recreationists and indifferent owners are the most passive groups in their timber supply behavior. The relationship between values and forestry behavior thus remains ambiguous.
  • Karppinen, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: heimo.karppinen@helsinki.fi (email)
  • Korhonen, Kämnerintie 7e 41, FI-00750 Helsinki, Finland E-mail: mika.korhonen@gmx.com
article id 893, category Research article
Juha Heiskanen, Timo Saksa, Jaana Luoranen. (2013). Soil preparation method affects outplanting success of Norway spruce container seedlings on till soils susceptible to frost heave. Silva Fennica vol. 47 no. 1 article id 893. https://doi.org/10.14214/sf.893
Keywords: Picea abies; pine weevil; vole; mounding; reforestation; glyphosate; plantation establishment; soil water content
Abstract | Full text in HTML | Full text in PDF | Author Info
Soil preparation is a common practice that precedes outplanting of Norway spruce (Picea abies (L.) Karst.) in Finland as it enhances the survival and early growth of seedlings. Mounding in particular has become more common with Norway spruce planting in recent years. However, on fine-grained soils, the postplanting performance of seedlings has been poorer than on coarser soils even with mounding. This study examined the effects of different soil preparation treatments (spot and ditch mounding with varying mound height, inverting, unprepared control with or without a herbicide) on the postplanting performance of Norway spruce container seedlings on till soil susceptible to frost heave in two outplanting forest sites in central Finland. The results indicate higher soil temperature and lower soil water content especially in the highest mounds. Mounds, however, subsided gradually during the study years. Seedling mortality was higher and the proportion of vigorous seedlings was lower in the unprepared treatments, mainly due to increased pine weevil (Hylobius abietis L.) damage. Frost heave was present mainly on ditch mounded and inverted spots. Glyphosate herbicide treatment showed no benefit compared to the untreated control in two years. Consequently, seedling damage and conditions in the planting spots were reflected in seedling growth which was enhanced in the mounded spots. However, varying mound height or thickness of mineral capping showed no clear difference in seedling growth. The results therefore suggest that ditch or spot mounding should be used on frost heave susceptible forest soils to promote plantation establishment. Inverting or having no soil preparation with or without herbicide is not recommended.
  • Heiskanen, Finnish Forest Research Institute, Suonenjoki Unit, Juntintie 154, FI-77600, Suonenjoki, Finland E-mail: juha.heiskanen@metla.fi (email)
  • Saksa, Finnish Forest Research Institute, Suonenjoki Unit, Juntintie 154, FI-77600, Suonenjoki, Finland E-mail: timo.saksa@metla.fi
  • Luoranen, Finnish Forest Research Institute, Suonenjoki Unit, Juntintie 154, FI-77600, Suonenjoki, Finland E-mail: jaana.luoranen@metla.fi
article id 892, category Research article
M. Carmen San José, Laura V. Janeiro, Elena Corredoira. (2013). Micropropagation of threatened black alder. Silva Fennica vol. 47 no. 1 article id 892. https://doi.org/10.14214/sf.892
Keywords: Alnus glutinosa; micropropagation; glucose; mature trees
Abstract | Full text in HTML | Full text in PDF | Author Info
Micropropagation techniques are valuable tools for propagating, conserving and restoring trees. An efficient micropropagation method involving axillary shoot proliferation of material obtained from mature European alder (Alnus glutinosa (L.) Gaertn.) trees was developed. Branch segments from trees aged 20–30 years were forced to flush, and explants derived from new shoots were cultured on Woody Plant Medium supplemented with 8.88 µM benzyladenine and 2.85µM indole-3-acetic-acid. In vitro establishment was achieved in all five genotypes evaluated. Shoot cultures were maintained by sequential subculture of explants on the same medium supplemented with 0.88–0.44 µM benzyladenine and 2.85 µM indole-3-acetic acid. Transfer to fresh medium every 3 weeks during a 9-week multiplication period and the inclusion of 2.28 µM zeatin during the last 3 weeks of culture improved the multiplication rate and shoot quality. Use of 2% glucose as the carbohydrate source produced better results than 3% sucrose for shoot proliferation. In vitro rooting of shoots was achieved with 2% glucose and 0.49 µM indole-3-butyric acid for 7 days, followed by in vitro culture on auxin-free medium for 21 days. Rooted plantlets were acclimatized to the greenhouse and were viable for reintroduction into the natural habitat.
  • San José, Instituto de Investigaciones Agrobiológicas de Galicia (IIAG), CSIC, Apartado 122, 15080 Santiago de Compostela, Spain E-mail: sanjose@iiag.csic.es (email)
  • Janeiro, INLUDES, Diputación Provincial de Lugo, Ronda de la Muralla 140, 27004 Lugo, Spain E-mail: lauravj68@hotmail.com
  • Corredoira, Instituto de Investigaciones Agrobiológicas de Galicia (IIAG), CSIC, Apartado 122, 15080 Santiago de Compostela, Spain E-mail: elenac@iiag.csic.es
article id 924, category Research article
Minna Pulkkinen. (2012). On non-circularity of tree stem cross-sections: effect of diameter selection on cross-section area estimation, Bitterlich sampling and stem volume estimation in Scots pine. Silva Fennica vol. 46 no. 5B article id 924. https://doi.org/10.14214/sf.924
Keywords: basal area; Scots pine; stem volume; forest mensuration; cross-section; non-circularity; Bitterlich sampling
Abstract | View details | Full text in PDF | Author Info
In the common methods of forest mensuration, including stem volume models and Bitterlich sampling, stem cross-sections are assumed to be circular. In nature this assumption is never exactly fulfilled. Errors due to non-circularity have been presumed to be small and unimportant but studied little: theoretical and empirical studies exist on cross-section area estimation, but errors in stem volume estimation have not been investigated at all, and errors in Bitterlich sampling are theoretically known only for stand basal area estimation. In the theoretical part of this study, we developed methods for quantifying the systematic and sampling errors that 22 common ways of selecting diameter within non-circular cross-sections induce (i) in area estimates by the circle area formula, (ii) in stand total estimates by Bitterlich sampling, and (iii) in stem volume estimates by a volume equation, by a cubic-spline-interpolated stem curve, and by a generalised volume estimator. In the empirical part, based on the digital images of 709 discs taken at 6–10 heights in 81 Scots pine stems from different parts of Finland, we investigated the variation in cross-section shape, and demonstrated the magnitude of the errors presented in the theoretical part. We found that non-circularity causes systematic overestimation of area and volume, and inflicts potentially systematic error on stand total estimates by Bitterlich sampling. In our data these effects were small, but the finding is not generalisable due the skewed size distribution and poor geographical representativeness of the data. We recommend using diameter derived from girth for both tree and stand level estimation, as it involves no sampling error and produces clearly the most stable systematic errors.
  • Pulkkinen, E-mail: minna.pulkkinen@iki.fi (email)
article id 922, category Research article
Malin Nilsson, Dianne Staal Westerlund, Olof Wahlberg, Ljusk Ola Eriksson. (2012). Forest planning in a Swedish company – a knowledge management analysis of forest information. Silva Fennica vol. 46 no. 5 article id 922. https://doi.org/10.14214/sf.922
Keywords: forest planning; long-term planning; medium-term planning; knowledge management; knowledge; information
Abstract | View details | Full text in PDF | Author Info
Forest data and forest information are central to forest management planning. The knowledge a large forest-owning company possesses about its forests could potentially be a strategic capability. In this study, the forest-planning process of a large forest company is analyzed in terms of knowledge management (KM). The study was conducted as a case study of Sveaskog – the largest forest-owning company in Sweden. The study focuses on the long-term harvest strategy through medium-term planning until the stands are transferred to the tract bank and ready for operational planning. Interviews with key persons within the organization were conducted to assess how forest knowledge is used in this process. The results are presented for the four knowledge management processes: creation, storage-retrieving, transferring and applying. They show that the planning system relies to a great extent on codified knowledge realized by a push strategy.
  • Nilsson, Swedish University of Agricultural Sciences, Department of Forest Resource Management, SE-901 83 Umeå, Sweden E-mail: malin.nilsson@slu.se (email)
  • Staal Westerlund, Swedish University of Agricultural Sciences, Department of Forest Resource Management, SE-901 83 Umeå, Sweden E-mail: dianne.wasterlund@slu.se
  • Wahlberg, Swedish University of Agricultural Sciences, Department of Forest Resource Management, SE-901 83 Umeå, Sweden E-mail: ow@nn.se
  • Eriksson, Swedish University of Agricultural Sciences, Department of Forest Resource Management, SE-901 83 Umeå, Sweden E-mail: ljusk.ola.eriksson@slu.se
article id 921, category Research article
Katri Luostarinen. (2012). Tracheid wall thickness and lumen diameter in different axial and radial locations in cultivated Larix sibirica trunks. Silva Fennica vol. 46 no. 5 article id 921. https://doi.org/10.14214/sf.921
Keywords: wood; earlywood; latewood; fibre; Siberian larch; xylem
Abstract | View details | Full text in PDF | Author Info
In Larix trunks the properties of wood differ clearly radially, but the axial differences are smaller as well as being less studied. Wood anatomy is in particular poorly studied, even though all other wood properties derive from cell and tissue structure. The aim of this study was to chart variation in tracheid size (double wall thickness (2CWT), diameter of lumen (RD)) within fast grown cultivated Siberian larch (Larix sibirica Ledeb.) trunks. The differences in 2CWT and RD were clear between earlywood (EW) and latewood (LW), 2CWT increasing clearly less in EW than in LW towards the bark, while RD stayed quite stable in LW but in EW increased markedly towards the bark. The difference in 2CWT between EW and LW increased towards the upper trunk. In conclusion, the radial variation in RD and 2CWT was different between the butt and other studied heights. As the difference in 2CWT between EW and LW was smaller at the butt than the upper portion of the trunk, the wood was the most homogenous at the butt.
  • Luostarinen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: katri.luostarinen@uef.fi (email)
article id 920, category Research article
Hannu Hökkä, Jaakko Repola, Mikko Moilanen, Markku Saarinen. (2012). Seedling establishment on small cutting areas with or without site preparation in a drained spruce mire – a case study in northern Finland. Silva Fennica vol. 46 no. 5 article id 920. https://doi.org/10.14214/sf.920
Keywords: Picea abies; site preparation; canopy gaps; natural regeneration; drained peatlands; seedling establishment; clear-cut
Abstract | View details | Full text in PDF | Author Info
A large proportion of drained spruce mire stands is currently approaching regeneration maturity in Finland. We studied the effect of cutting – small canopy openings (78, 177, and 314 m2) and small clear-cuts (0.25–0.37 ha) – with or without site preparation (scalping) on the establishment of natural Norway spruce seedlings in one experimental drained spruce mire stand in northern Finland. The cuttings were made in winter 2004–2005 and site preparation with scalping in early June 2005. The experimental design was composed of four blocks with altogether four clear-cuts and 33 canopy openings. The seedling establishment was surveyed annually (2006, 2008–2010) from five circular sample plots (one 10 m2 and four 5 m2 plots in size) located within the canopy openings and from 18 circular 5 m2 sample plots systematically located in the scalped and untreated halves of the clear-cuts. Site preparation was found unnecessary, because it resulted in a clearly lower number of seedlings in the openings. A slight negative effect was also found in the clear-cuts. In the two years following the cuttings, the number of seedlings increased quickly in the canopy openings, but more gradually in the clear-cut areas. In 2010, on average 15 500 new seedlings were observed in the canopy openings and 6700 in the clear-cut areas, of which 5050 and 1200, respectively, were >0.1 m tall spruces. The proportion of birch increased in the last two years, being ca. 22% in the openings and 45% in the clear-cuts in 2010. The spatial distribution of seedlings was more uneven in the clear-cuts than in the openings, with 41% and 20% of survey plots empty, respectively.
  • Hökkä, Finnish Forest Research Institute, Rovaniemi Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: hannu.hokka@metla.fi (email)
  • Repola, Finnish Forest Research Institute, Rovaniemi Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: jaakko.repola@metla.fi
  • Moilanen, Finnish Forest Research Institute, Oulu Unit, P.O. Box 413, FI-90014 University of Oulu, Finland E-mail: mikko.moilanen@metla.fi
  • Saarinen, Finnish Forest Research Institute, Parkano Unit, Kaironiementie 15, FI-39700 Parkano, Finland E-mail: markku.saarinen@metla.fi
article id 919, category Research article
Karri Uotila, Juho Rantala, Timo Saksa. (2012). Estimating the need for early cleaning in Norway spruce plantations in Finland. Silva Fennica vol. 46 no. 5 article id 919. https://doi.org/10.14214/sf.919
Keywords: cleaning; conifer release; cost-effectiveness; forest vegetation management; early cleaning
Abstract | View details | Full text in PDF | Author Info
Effective management of Norway spruce (Picea abies (L.) Karst.) plantations requires detailed information on stand development, which is costly to measure. However, estimating the need for early stand management from site attributes that persists stabile after ones measured, may provide an inexpensive alternative. This study compared hardwood competition in spruce plantations of varying ages and tested the usability of this information in estimating the need for early cleaning. The data included 197 spruce plantations (4–7 years old) inventoried in southern Finland in 2007. The level (Low, Substantial, High) of need for early cleaning was subjectively determined by contrasting location and size of competing hardwoods to a conifer crop tree. Then the stage of the need for early cleaning was modelled according to site and stand attributes. Nearly 60% of the conifer crop trees in the plantations were subjectively judged to require early cleaning (Substantial 37.2%, High 21.2%), but only 10 per cent of the evaluated area was cleaned. Need for cleaning was intense on peatlands or damp soils, whereas it was mild on unprepared soils or cleaned sites. Traditional site characteristics used in forest management planning can be useful for recognising the peripheral cases, where need for cleaning is probably high or low. However, on a typical mineral soil plantation (uncleaned, soil prepared) the model indicates the differences in the need for early cleaning weakly. The need for early cleaning was already high in 4-year-old plantations, why stand age did not have significant effect on development of the need. Thus, the timing of an operation can not be predicted with the model. Nonetheless, early cleaning very likely opens growth space of crop trees in a 4–7-year-old spruce plantation. Therefore, from an aspect of crop growth, an uncleaned Norway spruce plantation in this age group is quite consistently worth cleaning.
  • Uotila, Finnish Forest Research Institute, Suonenjoki Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: karri.uotila@metla.fi (email)
  • Rantala, Metsä Group, Tampere, Finland E-mail: juho.rantala@metsagroup.com
  • Saksa, Finnish Forest Research Institute, Suonenjoki Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@metla.fi
article id 918, category Research article
Nelson Thiffault, François Hébert, Robert Jobidon. (2012). Planted Picea mariana growth and nutrition as influenced by silviculture x nursery interactions on an ericaceous-dominated site. Silva Fennica vol. 46 no. 5 article id 918. https://doi.org/10.14214/sf.918
Keywords: fertilization; black spruce; scarification; Kalmia angustifolia; Rhododendron groenlandicum; stock type
Abstract | View details | Full text in PDF | Author Info
We aimed at evaluating the interacting effects of silvicultural and nursery practices on planted black spruce (Picea mariana (Mill.) BSP) dimensions, growth, survival and nutrition, 8 years following planting on a carefully logged boreal stand heavily invaded by Kalmia angustifolia L. and Rhododendron groenlandicum (Oeder) Kron & Judd. We also evaluated functional traits related to light and nutrient acquisition and key environmental resource availability to interpret treatment impacts on spruce seedling leaf traits and growth. An experimental black spruce plantation, consisting in a randomized block split-split-split plot design with 13 replicates was established in northeastern Quebec (Canada). Scarification (single-pass, double-pass), fertilization at the time of planting (control; macronutrients only; macro + micronutrients), stock type (container-grown; bare-root), and initial foliar N concentration (4 increasing levels) treatments were applied, and measurements were performed 5 and 8 years following planting. Double-pass scarification significantly increased soil temperature and reduced the competition cover, compared to the single-pass treatment. As a result, double-pass scarification promoted seedling growth over the single-pass treatment, and influenced the expression of other treatment effects. However, the relative gains associated with the second scarification pass have to be balanced against the supplemental investment involved by the treatment before being recommended. Our results point to variable effects of fertilization at planting to stimulate seedling initial growth. In this ecosystem, it appears that the silvicultural gains of this treatment depend on the variable of interest. Bare-root seedlings grew faster than containerized seedlings in the most intense site preparation treatment, but the differences have limited silvicultural impacts.
  • Thiffault, Ministère des Ressources naturelles du Québec, Direction de la recherche forestière, 2700 rue Einstein, Québec, QC, Canada G1P 3W8 E-mail: nelson.thiffault@mrnf.gouv.qc.ca (email)
  • Hébert, Ministère des Ressources naturelles du Québec, Direction de la recherche forestière, 2700 rue Einstein, Québec, QC, Canada G1P 3W8 E-mail: ffrancois.hebert@mrnf.gouv.qc.ca
  • Jobidon, Ministère des Ressources naturelles du Québec, Direction de la recherche forestière, 2700 rue Einstein, Québec, QC, Canada G1P 3W8 E-mail: robert.jobidon@mrnf.gouv.qc.ca
article id 917, category Research article
Anna Gunulf, Rebecka Mc Carthy, Jonas Rönnberg. (2012). Control efficacy of stump treatment and influence of stump height on natural spore infection by Heterobasidion spp. of precommercial thinning stumps of Norway spruce and birch. Silva Fennica vol. 46 no. 5 article id 917. https://doi.org/10.14214/sf.917
Keywords: Picea abies; Betula spp.; precommercial thinning; stump treatment; Heterobasidion spp.; Phlebiopsis gigantea; stump height
Abstract | View details | Full text in PDF | Author Info
An alternative precommercial thinning technique results in trees being cut higher up the stem compared to the normal method using a brush saw. The aims of this study were to investigate if primary infection of Heterobasidion spp. on precommercial thinning stumps of Norway spruce and birch is influenced by stump height and to test the control efficacy of stump treatment with Phlebiopsis gigantea on precommercial thinning stumps of Norway spruce. Small Norway spruce and birch trees were felled on five sites in southern Sweden and their stumps subjected to natural spore infection. For each species, two treatments of stump height were created: 15 and 100 cm. Half of the Norway spruce stumps were treated with P. gigantea. After two months, 896 stumps were sampled and infection by Heterobasidion spp. was quantified. The height of stumps did not significantly influence infection frequency or size of infection on either tree species. Untreated Norway spruce stumps had an infection frequency of 55% while 31% of the treated stumps were infected. The control efficacy of stump treatment in terms of reduced relative infected area was 61–65%. The area occupied by Heterobasidion spp. on birch stumps was generally small, on average 0.4 cm2 per infected stump, although 15% of the stumps were infected. The risk of primary infection in Norway spruce dominated stands should be considered when precommercial thinning is conducted but the control efficacy and economy of stump treatment warrants further investigation before practical recommendations can be made.
  • Gunulf, Swedish University of Agricultural Sciences, Department of Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: anna.gunulf@slu.se (email)
  • Mc Carthy, Skogsforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden E-mail: Rebecka.McCarthy@skogforsk.se
  • Rönnberg, Swedish University of Agricultural Sciences, Department of Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: jonas.ronnberg@slu.se
article id 916, category Research article
M. Carmen San José, Lourdes Romero, Laura V. Janeiro. (2012). Effect of indole-3-butyric acid on root formation in Alnus glutinosa microcuttings. Silva Fennica vol. 46 no. 5 article id 916. https://doi.org/10.14214/sf.916
Keywords: alder; mature trees; histology; rooting
Abstract | View details | Full text in PDF | Author Info
A study of the in vitro rooting process in mature alder (Alnus glutinosa (L.) Gaertn.) shoots is described. Microcuttings from shoots cultured in vitro were transferred to a half-strength Woody Plant Medium containing 0 or 0.1 mg l–1 indole-3-butyric acid (IBA) for 0 to 7 days. The presence of IBA in the medium increased the rooting percentage, number of roots, percentage of lateral roots, and length of the shoots. Histological studies were carried out with shoots treated with 0 or 0.1 mg l–1 IBA for 7 days. According to these criteria, treatment with IBA for 2–3 days proved to be the most successful. In both treatments, substancial reactivation of cell division was observed at the base of the shoots after 1 day. Some cambial zone and adjacent phloem cells became dense cytoplasm, having nuclei with prominent nucleoli. The first cell divisions were also observed at this time. In the treatment with IBA (0.1 mg l–1 for 7 days), meristemoids became individualized, consisting of densely staining cells, by day 3. Identifiable conical shaped root primordia with several cell layers were visible after 4–5 days. Roots with an organized tissue system emerged from the stem after 6 days in the IBA-treated shoots. Meristemoid formation was delayed until the fourth day and root emergence until the eight day in the control treatment (no IBA).
  • San José, Instituto de Investigaciones Agrobiológicas de Galicia, CSIC, Avda de Vigo s/n, Apartado 122, 15780 Santiago de Compostela, Spain E-mail: sanjose@iiag.csic.es (email)
  • Romero, CIFP Politécnico de Santiago, Santiago de Compostela, Spain E-mail: lr@nn.es
  • Janeiro, INLUDES, Diputación Provincial de Lugo, Lugo, Spain E-mail: lauravj68@hotmail.com
article id 913, category Research article
Han Zhang, Joseph Buongiorno. (2012). Markets, government policy, and China's timber supply. Silva Fennica vol. 46 no. 4 article id 913. https://doi.org/10.14214/sf.913
Keywords: market; China; supply; policy; land reform; econometrics
Abstract | View details | Full text in PDF | Author Info
China's domestic demand and exports of wood products are rising rapidly compared to domestic supply. The determinants of timber supply in China were investigated with panel data from 25 provinces from 1999 to 2009. The results indicated that China’s timber supply had responded to both market forces, reflected by timber prices largely determined by world demand and supply, and to government policies expressed by production quotas and the tenure reform on collective forestland. The price elasticity of China’s timber supply was estimated at 0.31±0.12. The inelastic response of production to the quota (0.20±0.02) suggested that government had a limited, though significant, control of timber supply. Other things being equal, the land tenure reform increased timber supply by 18±8 percent, where and when it had been implemented.
  • Zhang, Research Center for Resource Economics and Environment Management, Northwest A&F University. No. 3 Taicheng Road, Yangling, Shaanxi, China 712100 E-mail: hz@nn.fi
  • Buongiorno, Department of Forest and Wildlife Ecology, University of Wisconsin-Madison, Madison, WI 53706, U.S.A. E-mail: jbuongio@wisc.edu (email)
article id 912, category Research article
Heidi Hallongren, Juho Rantala. (2012). Commercialisation and international market potential of Finnish silvicultural machines. Silva Fennica vol. 46 no. 4 article id 912. https://doi.org/10.14214/sf.912
Keywords: mechanised silviculture; silvicultural device; market potential; commercialisation of technology
Abstract | View details | Full text in PDF | Author Info
Recently, the need to mechanise silvicultural operations has increased in the Nordic countries. While several new machine concepts have been developed, the manufacture of silvicultural devices remains at the market introduction stage. Information is required in support of existing and forthcoming Finnish small-scale producers of silvicultural devices, who wish to commercialise and further market their innovations in domestic and export markets. The aim of this study was to identify the opportunities, challenges and market potential of business activities that develop in connection with device or machine production. Small-scale Finnish manufacturers of silvicultural devices, Finnish large-scale harvester manufacturers and international silvicultural experts participated in the study. The results show that participant groups have varying opinions of cooperation methods and export activities, as well as of the export markets with the best potential. According to international silvicultural experts, mechanised planting and pre-commercial thinning have the greatest potential worldwide. However, demand for mechanised pre-commercial thinning and planting has been mainly confined to the Nordic countries. For a foreign firm marketing a new silvicultural machine concept, the most important customers and cooperation partners are locally operating forest firms, machine contractors and research organisations. The results of the study provide a useful overview of the current state of silvicultural device manufacturing in Finland.
  • Hallongren, Finnish Forest Research Institute, Suonenjoki Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: heidi.hallongren@metla.fi (email)
  • Rantala, Metsä Group, Tampere, Finland E-mail: juho.rantala@metsagroup.com
article id 911, category Research article
Marjut Turtiainen, Olli Saastamoinen, Kari Kangas, Matti Vaara. (2012). Picking of wild edible mushrooms in Finland in 1997–1999 and 2011. Silva Fennica vol. 46 no. 4 article id 911. https://doi.org/10.14214/sf.911
Keywords: wild mushrooms; mushroom picking; household use; market use; milk caps; ceps
Abstract | View details | Full text in PDF | Author Info
This study reports on national survey results concerning mushroom picking in Finland during four separate years: 1997–1999 and 2011. The material was collected by mailed questionnaire surveys amongst Finnish households. The sample size varied from 1858 (in 1998) to 6849 households (in 1997) and the response rates varied from 51% (in 2011) to 70% (in 1999). The results indicate that both the rate of participation in mushroom picking and estimates of the quantities collected varied greatly depending on whether the survey was conducted in a favourable or unfavourable year. In 1998, when the mushroom crop was abundant, a total of 47% of all households were engaged in picking and the total harvest was 16.1 million kg. In 1999, when the crop was poor, the estimates were the lowest (23% and 3.3 million kg, respectively) and in a year with a relatively abundant crop (2011), the estimates were 42% and 15.0 million kg, respectively. Mushrooms were collected mainly for home use, which accounted for 85–90% of the total harvest depending on the year. Only a small proportion of all households (0.3–1.3%) were engaged annually in commercial mushroom picking. In 1997–1999, milk caps formed the major part of the total amount picked (i.e. 37–53% depending on the year), whilst in 2011 their share was approximately one fifth of the total harvest. The results also indicate that the proportion of ceps in commercial picking has increased since the 1990s
  • Turtiainen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: marjut.turtiainen@uef.fi (email)
  • Saastamoinen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: olli.saastamoinen@uef.fi
  • Kangas, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: kari.kangas@uef.fi
  • Vaara, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: matti.vaara@uef.fi
article id 910, category Research article
Gernot Erber, Christian Kanzian, Karl Stampfer. (2012). Predicting moisture content in a pine logwood pile for energy purposes. Silva Fennica vol. 46 no. 4 article id 910. https://doi.org/10.14214/sf.910
Keywords: moisture content; modelling; natural wind drying of fuel wood; log pile
Abstract | View details | Full text in PDF | Author Info
Determining the moisture content of naturally dried fuel stock without frequent measuring is a problem still unsolved. Modelling moisture content based on automatically captured meteorological data could provide a solution. An accurate model would allow the drying period and the point of chipping to be optimised. For the experimental study, a metal frame supported by load sensors and loaded with 17 tons of logwood was set up next to a meteorological station. A multiple linear regression model was used to link meteorological and load data to provide a formula for determining the moisture content. The pile dried for a period of 14 months (average temperature of 7.3 °C, a humidity of 81%, and 777 mm of rainfall). The overall moisture content dropped from 50.1% to 32.2%. The regression model, which based on daily means and sums of meteorological parameters, provided a mean deviance from the observed curve of –0.51%±0.71% within the period of investigation. Relative humidity was found to be most important parameter in drying. Increased moisture content resulting from rainfall greater than 30 mm per day reverted back to pre-rainfall values within two to three days, if no other rainfall events followed. Covering the pile would have a positive effect on the drying performance. In terms of economic benefit it could be shown that natural drying is beneficial. Overall this study shows that meteorological data used in site specific drying models can adequately predict the moisture content of naturally dried logwood.
  • Erber, University of Natural Resources and Life Sciences, Peter Jordan-Strasse 82, 1190 Wien, Austria E-mail: gernot.erber@boku.ac.at (email)
  • Kanzian, University of Natural Resources and Life Sciences, Peter Jordan-Strasse 82, 1190 Wien, Austria E-mail: christian.kanzian@boku.ac.at
  • Stampfer, University of Natural Resources and Life Sciences, Peter Jordan-Strasse 82, 1190 Wien, Austria E-mail: karl.stampfer@boku.ac.at
article id 909, category Research article
Veikko Hiltunen, Mikko Kurttila, Jouni Pykäläinen. (2012). Strengthening top-level guidance in geographically hierarchical large scale forest planning: experiences from the Finnish state forests. Silva Fennica vol. 46 no. 4 article id 909. https://doi.org/10.14214/sf.909
Keywords: efficiency; acceptability; hierarchical planning; strategic forest planning
Abstract | View details | Full text in PDF | Author Info
Different planning approaches conclude to different results. The top-down approach allocates resources efficiently from the top-level perspective, while the bottom-up approach provides optimal results for the lower levels. Integrated approach that combines the elements of these two basic approaches provides compromise solutions for decision makers. The aim of this study was to examine potential efficiency improvements in hierarchically structured large scale forest management through increased top-level guidance. The resulting effects on the acceptability of the plans on the lower level were also studied. Large scale planning typically considers forests owned by states, companies and municipalities. In the case study of the Finnish state forests, alternative country level solutions were generated by combining regional forest plans in different ways. The results showed that the currently applied bottom-up approach, which produces regionally optimal management strategies, did not result in the most efficient use of resources on the country level. However, the new country level solutions did not produce huge improvements in the country level objective values compared to the results of the current approach. Furthermore, if country level efficiency improvements were emphasized more, together with wide approval by regional stakeholders and local residents, new kind of interaction and participation between the planning levels and also between the regions would be needed.
  • Hiltunen, Metsähallitus, Viestitie 2, FI-87700 Kajaani, Finland E-mail: veikko.hiltunen@metsa.fi (email)
  • Kurttila, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: mikko.kurttila@metla.fi
  • Pykäläinen, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jouni.pykalainen@metla.fi
article id 908, category Research article
Sattar Ezzati, Akbar Najafi, M. A. Rab, Eric K. Zenner. (2012). Recovery of soil bulk density, porosity and rutting from ground skidding over a 20-year period after timber harvesting in Iran. Silva Fennica vol. 46 no. 4 article id 908. https://doi.org/10.14214/sf.908
Keywords: soil disturbance; timber harvesting; soil conservation; skid trail slope; mountainous forest
Abstract | View details | Full text in PDF | Author Info
Ground-based skidding can have detrimental effects on soil properties trough soil profile disturbance and compaction that can persist for decades. We investigated the recovery of physical properties of disturbed brown soils on four abandoned downhill skid trails in a deciduous mountain forest in northern Iran. The most recent skidding operations had taken place 1–5 yrs, 6–10 yrs, 11–15 yrs, and 16–20 yrs ago, providing a 20-year chronosequence with four 5-year recovery periods. For each recovery period, mean values for soil bulk density (BD), total porosity (TP), macroporosity (MP), soil moisture content (SM), and rut depth (RD) were assessed for three levels of traffic intensity (Primary (PS), Secondary (SS) and Tertiary (TS) skid trails) and two levels of slope gradients (Gentle (G) and Steep (S)) and compared to those in undisturbed (control) areas. Over the 20-year recovery period, PS trails on gentle slopes exhibited mean values that were 35–42% (BD), 3–7% (SM), and 13–19 cm (RD) greater and 18–24% (TP) and 19–28% (MP) lower compared to undisturbed areas; on steep PS trails, values were 40–46% (BD), 2–13% (SM), and 13–21 cm (RD) greater and 23–27% (TP) and 28–35% (MP) lower, respectively. While RD and SM recovered, 20 years was not long enough for the other physical soil properties, particularly on steep slopes. To minimize soil disturbance, skidding should be confined to areas with gentle slopes and alternative harvesting methods such as cable yarding should be used where slope gradients exceed 20%.
  • Ezzati, Department of Forestry and Forest Engineering, Tarbiat Modares University, P.O. Box 64414-356, Iran E-mail: se@nn.ir
  • Najafi, Department of Forestry and Forest Engineering, Tarbiat Modares University, P. Box 64414-356, Iran E-mail: a.najafi@modares.ac.ir (email)
  • Rab, Soil Physics Future Farming Systems Research Division, Department of Primary Industries, Victoria, Australia E-mail: mr@nn.ir
  • Zenner, Department of Ecosystem Science and Management, Pennsylvania State University, University Park, PA, USA E-mail: eric.zenner@psu.edu
article id 907, category Research article
Chunyu Zhang, Yazhou Zhao, Xiuhai Zhao, Klaus von Gadow. (2012). Species-habitat associations in a northern temperate forest in China. Silva Fennica vol. 46 no. 4 article id 907. https://doi.org/10.14214/sf.907
Keywords: species richness; spatial autocorrelation; dispersal limitations; indicator species; topographic differentiation
Abstract | View details | Full text in PDF | Author Info
This contribution identifies species-habitat associations in a temperate forest in north-eastern China, based on the assumption that habitats are spatially autocorrelated and species are spatially aggregated due to limited seed dispersal. The empirical observations were obtained in a large permanent experimental area covering 660 x 320 m. The experimental area was subdivided into four habitat types using multivariate regression tree (MRT) analysis. According to an indicator species analysis, 38 of the 47 studied species were found to be significant indicators of the MRT habitat types. The relationships between species richness and topographic variables were found to be scale-dependent, while the great majority of the species shows distinct habitat-dependence. There are 188 potential species-habitat associations, and 114 of these were significantly positive or negative based on habitat randomization. We identified 139 significant associations using a species randomization. A habitat is not a closed system it may be both, either a sink or a source. Therefore, additional to the randomization, the Poisson Cluster Model (PCM) was applied. PCM considers the spatial autocorrelation of species and habitats, and thus appears to be more realistic than the traditional randomization processes. It identified only 37 associations that were significant. In conclusion, the deviation from the random process, i.e. the high degree of species spatial mingling may be explained by persistent immigration across habitats.
  • Zhang, Key Laboratory for Forest Resources & Ecosystem Processes of Beijing, Beijing Forestry University, Beijing 100083, China E-mail: zcy_0520@163.com (email)
  • Zhao, Department of Landscape Architecture, School of Architecture, Tsinghua University, Beijing 100084, China E-mail: yz@nn.cn
  • Zhao, Key Laboratory for Forest Resources & Ecosystem Processes of Beijing, Beijing Forestry University, Beijing 100083, China E-mail: xz@nn.cn
  • von Gadow, Faculty of Forestry and Forest Ecology, Georg-August-University Göttingen, Büsgenweg 5, D-37077 Göttingen, Germany E-mail: KGadow@gwdg.de
article id 906, category Research article
Eivind Meen, Anders Nielsen, Mikael Ohlson. (2012). Forest stand modelling as a tool to predict performance of the understory herb Cornus suecica. Silva Fennica vol. 46 no. 4 article id 906. https://doi.org/10.14214/sf.906
Keywords: canopy layer; field layer; stand structure dynamics; forest simulator; composite models; Akaike’s information criterion
Abstract | View details | Full text in PDF | Author Info
Forest simulation models have been widely used to predict future stand structure. Generally these models do not include the understory vegetation and its response on stand structure change or other environmental factors. Previous simulation studies have shown that stand structure related variables, e.g. basal area, can explain diversity of the forest floor vegetation in boreal forests. We hypothesise that such variables also can be used to explain the performance of understory species and we conceptualise how plant ecology and forest modelling can be combined to predict the performance of understory plants in Norwegian boreal forests. We predict the performance of an understory plant species (Cornus suecica) over time using simulated values of forest variables as input to models expressing the relationship between forest environment variables and plant performance variables (viz. plant height, plant dry weight, number of flowers, number of branches and number of leaves). We also present relationships between plant performance and explanatory variables commonly used in basic ecological research, variables that currently not are readily compatible with forest simulators (e.g. soil chemical variables).We found basal area of canopy trees being the most important explanatory variable explaining C. suecica performance. The performance variable dry weight was predicted by one single model whereas the other performance variables were best predicted by model averaging. Forest simulations for 150 years showed values of plant performance of C. suecica to be reduced during forest succession.
  • Meen, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, P.O. Box 5003, NO-1432 Ås, Norway E-mail: eivind.meen@umb.no (email)
  • Nielsen, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, P.O. Box 5003, NO-1432 Ås, Norway E-mail: an@nn.no
  • Ohlson, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, P.O. Box 5003, NO-1432 Ås, Norway E-mail: mo@nn.no
article id 905, category Research article
Katarzyna A. Jadwiszczak, Danuta Drzymulska, Agata Banaszek, Piotr Jadwiszczak. (2012). Population history, genetic variation and conservation status of the endangered birch species Betula nana L. in Poland. Silva Fennica vol. 46 no. 4 article id 905. https://doi.org/10.14214/sf.905
Keywords: Betula nana; genetic diversity; microsatellites; conservation genetics; cpDNA; isolation
Abstract | View details | Full text in PDF | Author Info
The effective conservation of species requires data on the levels and distribution of genetic diversity within and among populations. In this study, we estimated the genetic variation in three isolated populations of Betula nana in Poland. An analysis of 11 nuclear microsatellites revealed moderate mean heterozygosities (HO=0.556, HE=0.562), low mean number of alleles per locus (A=4.57) and no inbreeding in the total sample. An M-ratio test indicated that each population had experienced a severe bottleneck in the past. Tests for heterozygosity excess revealed that a significant decrease in the numbers of individuals in two populations had occurred quite recently. The large number of private alleles and very restricted number of migrants between populations (Nm=0.35) strongly suggest that genetic drift and geographic isolation are the primary factors responsible for the reduction of genetic variation in the Polish populations of B. nana. We detected two cpDNA haplotypes in the study populations, which can be explained in terms of either the genetic drift acting on the relict localities or a postglacial recolonisation from distinct refugia. Palynological data indicated that one refugium could be located in the Carpathians and their northern foreland. The primary threat to B. nana in Poland is the overgrowth of its habitats by competing species, which has likely resulted in a lack of generative reproduction in the mountain populations.
  • Jadwiszczak, Institute of Biology, University of Białystok, wierkowa 20B, 15-950 Białystok, Poland E-mail: kszalaj@uwb.edu.pl (email)
  • Drzymulska, Institute of Biology, University of Białystok, wierkowa 20B, 15-950 Białystok, Poland E-mail: dd@nn.pl
  • Banaszek, Institute of Biology, University of Białystok, wierkowa 20B, 15-950 Białystok, Poland E-mail: ab@nn.pl
  • Jadwiszczak, Institute of Biology, University of Białystok, wierkowa 20B, 15-950 Białystok, Poland E-mail: pj@nn.pl
article id 440, category Research article
Chen-Jung Lin, Chih-Hsin Chung, Te-Hsin Yang, Far-Ching Lin. (2012). Detection of electric resistivity tomography and evaluation of the sapwood-heartwood demarcation in three Asia Gymnosperm species. Silva Fennica vol. 46 no. 3 article id 440. https://doi.org/10.14214/sf.440
Keywords: Cryptomeria japonica; Cunninghamia konishii; electrical resistance; nondestructive technique; Taiwania cryptomerioides
Abstract | View details | Full text in PDF | Author Info
The proportions of sapwood and heartwood of trees have significant impacts on various uses. Electric resistivity tomography (ERT) and corresponding electrical resistance (ER) value maps were examined in Japanese cedar (Cryptomeria japonica D. Don), Taiwania (Taiwania cryptomerioides Hayata), and Luanta fir (Cunninghamia konishii Hayata) trees. The position of the sapwood-heartwood demarcation was measured on incremental cores from living trees and the corresponding ER of the sapwood-heartwood boundary was acquired from the ER map. A positive significant relationship was found between the maximum ER plus minimum ER values (ERmax + ERmin) and ER of the sapwood-heartwood demarcation from the tomographic data. The position of the sapwood-heartwood demarcation was determined by corresponding ER, and the critical ER can be established by the ERmax + ERmin value of the tomographic data. The results from this study indicate that ERT technique can be used to determine the position of the sapwood-heartwood boundary and can serve as a methodology in undamaged living trees of Gymnosperm species.
  • Lin, Taiwan Forestry Research Institute, Taipei Taiwan E-mail: d88625002@yahoo.com.tw (email)
  • Chung, Taiwan Forestry Research Institute, Taipei Taiwan E-mail: chc@nn.tw
  • Yang, Taiwan Forestry Research Institute, Taipei Taiwan E-mail: thy@nn.tw
  • Lin, Taiwan Forestry Research Institute, Taipei Taiwan E-mail: fcl@nn.tw
article id 52, category Research article
Mirja Rantala, Teppo Hujala, Mikko Kurttila. (2012). Measuring and monitoring socio-cultural sustainability in the action of forest biodiversity cooperation networks. Silva Fennica vol. 46 no. 3 article id 52. https://doi.org/10.14214/sf.52
Keywords: additive utility function; community adaptation; criteria and indicators; METSO programme
Abstract | View details | Full text in PDF | Author Info
To safeguard overall sustainability in forest resource management, the ecological, economic, social, and cultural dimensions of sustainability should all be considered. However, the socio-cultural impacts are frequently contemplated only weakly in sustainability assessments. Hitherto, attempts to operationalize socio-cultural impacts arising from economic utilization or conservation of forest resources have been perceived as vague when compared to rigorous ecological and economic indicators. One reason is that socio-cultural impacts of forest management on individuals and communities are many and by nature context- and case-specific: they need local definition, which hampers diffusion of good solutions. This study developed a multi-criteria method for measuring and monitoring socio-cultural impacts of forest resource management; the case of cooperation network projects within Forest Biodiversity Programme for Southern Finland (METSO) provided empirical data. Based on a literature review, a set of 10 criteria and 25 indicators was compiled. Cumulative utility scores, presenting networks’ contributions to socio-cultural sustainability, were generated using performance, expert evaluation and weighting data and an additive utility model. The method enables longitudinal monitoring of socio-cultural impacts, which is beneficial because outcomes are different at different time points of projects’ life cycles and some appear with a delay. The method can be used in comparing sub-utility distributions i.e. monitoring units’ performance profiles, providing valuable information for policy-makers. The multi-criteria approach and the list of socio-cultural criteria are internationally transferable to other countries and contexts such as forest bioenergy, nature tourism, watershed management, that call for analysing socio-cultural impacts of forest resource management activity on private lands.
  • Rantala, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: mr@nn.fi
  • Hujala, Finnish Forest Research Institute, Vantaa Unit, Vantaa, Finland E-mail: th@nn.fi
  • Kurttila, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: mikko.kurttila@metla.fi (email)
article id 51, category Research article
Sanna Hautamäki, Antti Mutanen, Jari Viitanen. (2012). Substitution in the Finnish forest industry’s roundwood procurement. Silva Fennica vol. 46 no. 3 article id 51. https://doi.org/10.14214/sf.51
Keywords: roundwood procurement; imports; substitution; translog cost function
Abstract | View details | Full text in PDF | Author Info
In this study, the interaction and substitution between domestic and imported roundwood in the Finnish forest industry’s wood procurement is analysed by timber assortments. The results from the translog cost function approach and quarterly data of the total wood procurement and its components during the euro regime indicate that, to a certain extent, the Finnish forest industry has had the possibility of substituting imported roundwood volumes between countries in the Baltic Sea region. Contrary to earlier studies, also in the case of Russian birch pulpwood, the most important imported timber assortment, the results suggest that Russian birch pulpwood has rather substituted for than complemented the domestic supply in Finland. The increase in roundwood export duties in Russia has had a statistically significant effect on the trade in birch pulpwood and spruce sawlogs. Moreover, the results confirm the earlier findings of a rigid demand for roundwood in Finnish roundwood markets.
  • Hautamäki, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: sh@nn.fi
  • Mutanen, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: antti.mutanen@metla.fi (email)
  • Viitanen, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jv@nn.fi
article id 49, category Research article
Kalle Karttunen, Kari Väätäinen, Antti Asikainen, Tapio Ranta. (2012). The operational efficiency of waterway transport of forest chips on Finland’s Lake Saimaa. Silva Fennica vol. 46 no. 3 article id 49. https://doi.org/10.14214/sf.49
Keywords: barges; discrete-event simulation; forest fuels; logistics; supply chains; tugboats
Abstract | View details | Full text in PDF | Author Info
New and cost-efficient methods for use in supply chains for energy wood should be found, to reach the targets of the renewable energy utilisation set by the European Union. The long-distance waterway transportation of forest fuels should be thoroughly investigated, especially in areas where the transport distance is long and waterways could provide a feasible method of conveying forest fuel. In comparison to transport of forest chips by truck, barge-based waterway transport shows a competitive advantage due to the larger loads and higher bulk density of chips it allows. The cost-efficiency of waterway transportation operations related to forest chips in Finland’s Lake Saimaa region was studied using practical demonstrations and discrete-event simulation. The varying demand for fuel wood in three separate bio-power plants on the Saimaa lakeside (near the cities of Varkaus, Mikkeli, and Savonlinna) was addressed in several barge transportation scenarios. Finally, the economy of barge transportation was compared to the economy of truck transportation as a function of transportation distance and in terms of the annual performance of the transportation methods examined. The waterway supply chain of forest chips was cost-competitive to road transport by truck after 100–150 km. According to the simulation study, the most economical waterway transport options were based on fixed barge system and shift-independent harbor logistics where loading and unloading of barges were carried-out with a wheeled loader and a belt conveyor. Total supply chain costs including the best waterway logistics from road side storage to power plant ranged from 10.75 euros to 11.64 euros/MWh in distances of 100–150 km by waterways. The energy-density of forest chips in the barge load was found to be, on average, 25% higher than that in truck hauling, because of the better compaction of chips. Waterway transport is a viable option for long-distance transportation of forest chips in Eastern Finland.
  • Karttunen, Lappeenranta University of Technology, LUT Savo Sustainable Technologies, Mikkeli, Finland E-mail: kalle.karttunen@lut.fi (email)
  • Väätäinen, The Finnish Forest Research Institute, Joensuu, Finland E-mail: kari.vaatainen@metla.fi
  • Asikainen, The Finnish Forest Research Institute, Joensuu, Finland E-mail: antti.asikainen@metla.fi
  • Ranta, Lappeenranta University of Technology, LUT Savo Sustainable Technologies, Mikkeli, Finland E-mail: tapio.ranta@lut.fi
article id 48, category Research article
Jyrki Hytönen, Lasse Aro. (2012). Biomass and nutrition of naturally regenerated and coppiced birch on cutaway peatland during 37 years. Silva Fennica vol. 46 no. 3 article id 48. https://doi.org/10.14214/sf.48
Keywords: Betula; biomass production; coppicing; fertilization; PK fertilizers; wood ash
Abstract | View details | Full text in PDF | Author Info
Biomass production and nutrient use of birch thickets with a mixture of willow on a cut away peatland in southern Finland over a period of 37 years was studied. Dense, naturally regenerated 16-year-old birch stands were cut down, fertilized with either wood ash (P 108 and K 339 kg ha–1) or PK fertilizer (P 50 and K 95 kg ha–1) or left unfertilized. The biomass production of the coppiced stands and one uncut stand was monitored for a period of 21 years. Soil nutrient and foliar nutrient concentrations were analyzed several times during the study period. Ash fertilization supplied more nutrients than PK fertilization and increased the soil nutrient amounts more. The foliar phosphorus concentration of birch on control plots indicated a severe phosphorus deficiency which was removed by PK and ash fertilization. Fertilization did not increase nutrient concentrations of the stem (wood + bark) nor the amount of nutrients bound in the biomass. Two energy wood rotations (16+21 years) produced 124–158 Mg ha–1 of leafless, above-ground biomass altogether corresponding to 61–78 Mg ha–1 of carbon. The highest biomass yield was achieved with a rotation of 37 years in the uncut stand (211 Mg ha–1). Corresponding values for mean annual increment (MAI) were 3.4–4.3 Mg ha–1 and 5.7 Mg ha–1. This study shows that the length of the rotation for birch in energy wood production should be longer than 21 years. PK and ash fertilization increased the biomass of coppiced 21-year-old birch by 23 Mg ha–1 and 33 Mg ha–1, respectively.
  • Hytönen, Finnish Forest Research Institute, Kannus, Finland E-mail: jyrki.hytonen@metla.fi (email)
  • Aro, Finnish Forest Research Institute, Kannus, Finland E-mail: lasse.aro@metla.fi
article id 47, category Research article
Iulian Dragotescu, Daniel D. Kneeshaw. (2012). A comparison of residual forest following fires and harvesting in boreal forests in Quebec, Canada. Silva Fennica vol. 46 no. 3 article id 47. https://doi.org/10.14214/sf.47
Keywords: forest fires; biodiversity; boreal mixedwood; clearcuts; disturbances; residual forests; tree retention
Abstract | View details | Full text in PDF | Author Info
Residual forests are a key component of post-burned areas creating structure within burns and providing habitat and seed sources. Yet, despite their importance to biodiversity and ecosystem processes there is little information on how similar or different residuals in burned landscape are to harvested landscapes. Our goal was to examine and compare the density, size, shape, and spatial arrangement of residual forest vegetation after fire and clearcutting. We evaluated residual forest in two locations within the boreal mixedwood region of Quebec, Canada using aerial photo interpretation and ArcGIS 9.1 software. We found residual stands to be larger and more abundant in harvested zones relative to sites affected by fire. Differences with respect to shape and spatial arrangement of residual forest were also observed among disturbance types. Factors such as proximity to watercourses, watercourse shape, and physiography affected residual abundance and spatial distribution. Residual forest in harvested zones tended to be more elongated with greater edge due to rules governing forest operations. Despite greater quantity of residual forest in harvested areas than fires, managers should still be prudent as the surrounding forest matrix is reduced in many managed landscapes.
  • Dragotescu, Université du Québec à Montréal, Centre d’Étude de la Forêt (CEF), Montreal, Quebec, Canada E-mail: idragot@hotmail.com (email)
  • Kneeshaw, Université du Québec à Montréal, Centre d’Étude de la Forêt (CEF), Montreal, Quebec, Canada E-mail: ddk@nn.ca
article id 46, category Research article
Matti Rousi, Boy J.H.M. Possen, Risto Hagqvist, Barb R. Thomas. (2012). From the Arctic Circle to the Canadian prairies – a case study of silver birch acclimation capacity. Silva Fennica vol. 46 no. 3 article id 46. https://doi.org/10.14214/sf.46
Keywords: acclimation; Betula papyrifera; Betula pendula; birch adaptability; critical night length; provenance transfers
Abstract | View details | Full text in PDF | Author Info
Earlier provenance research has indicated poor success even in short distance transfers (> 2–3° latitude) of silver birch (Betula pendula Roth) southward from their origin. These results may indicate poor adaptability of silver birch to a warming climate. Some of the scenarios for a warming climate in Finland suggest effective heat sums are likely to double in the north and increase 1.5 fold in the south for the period of 2070–2099. Consequently, the outlook for silver birch appears bleak. To study the acclimation of birch to this projected change we established a provenance trial in northeastern Alberta, Canada, at the temperature area currently predicted for Central Finland (lat. 64–66°N) at the turn of this century (1400 dd). Our 10-year experiment showed that all the Finnish provenances (origins 61–67°N) have acclimated well to the warmer growth conditions experienced in Alberta at 54°N. These results suggest that silver birch has the potential to acclimate to thermal conditions predicted for Finland at the end of the 21st century. Our results also indicate that silver birch has the potential as a plantation species in Canada, where the Finnish birch grew faster in the boreal forest region of Canada than local paper birch (Betula papyrifera Marsh.) provenances.
  • Rousi, The Finnish Forest Research Institute, Finland E-mail: matti.rousi@metla.fi (email)
  • Possen, The Finnish Forest Research Institute, Finland E-mail: bjhmp@nn.fi
  • Hagqvist, The Finnish Forest Research Institute, Finland E-mail: rh@nn.fi
  • Thomas, University of Alberta, Dept of Renewable Resources, Edmonton & Alberta-Pacific Forest Industries Inc., Boyle, Alberta, Canada E-mail: brt@nn.ca
article id 45, category Research article
Guolei Li, Yong Liu, Yan Zhu, Qing Mei Li, R. Karsten Dumroese. (2012). Effect of fall-applied nitrogen on growth, nitrogen storage and frost hardiness of bareroot Larix olgensis seedlings. Silva Fennica vol. 46 no. 3 article id 45. https://doi.org/10.14214/sf.45
Keywords: deciduous trees; fall fertilization; frost hardiness; Olga Bay larch
Abstract | View details | Full text in PDF | Author Info
Nursery response of evergreen trees to fall fertilization has been studied widely, but little attention has been given to deciduous trees. Bareroot Olga Bay larch (Larix olgensis Henry) seedlings were fertilized in the nursery with urea at four rates (0, 30, 60, 90 kg N ha–1), with half of each rate applied on two dates (September 16 and October 1, 2009). The seedlings were excavated for evaluation on October 15. In the unfertilized (control) treatment, root and shoot dry mass increased by 100% and 57% respectively, while N concentration in the roots and shoots increased by 43% and 40% during the 30 day period. This indicated that substantial biomass growth during this period did not lead to internal nutrient dilution. Root dry mass increased when fall fertilization rates were ≥ 60 kg N ha–1. Fall fertilization increased N concentrations in root tissue by 48–73%. Compared with the control, shoot tissues of fall fertilized seedlings had slightly higher N concentration and content and significantly higher frost hardiness.
  • Li, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing 100083, China E-mail: gl@nn.cn
  • Liu, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing 100083, China E-mail: lyong@bjfu.edu.cn (email)
  • Zhu, Key Laboratory for Silviculture and Conservation, Ministry of Education, Beijing 100083, China E-mail: yz@nn.cn
  • Li, Research Institute of Forestry, Chinese Academy of Forestry; Key Laboratory of Forest Silviculture of State Forestry Administration, Beijing 100091, China E-mail: qml@nn.cn
  • Dumroese, US Department of Agriculture, Forest Service, Rocky Mountain Research Station, Moscow, ID, USA E-mail: rkd@nn.us
article id 44, category Research article
Anabel Aparecida de Mello, Leif Nutto, Karla Simone Weber, Carlos Eduardo Sanquetta, Jorge Luis Monteiro de Matos, Gero Becker. (2012). Individual biomass and carbon equations for Mimosa scabrella Benth. (bracatinga) in southern Brazil. Silva Fennica vol. 46 no. 3 article id 44. https://doi.org/10.14214/sf.44
Keywords: carbon offsets; forest inventory; firewood; modeling; tree compartments
Abstract | View details | Full text in PDF | Author Info
Mimosa scabrella Benth. is an important native species of southern Brazil widely used for energy and promising for reforestation carbon offsets. Quantification of biomass and carbon stock is valuable for both purposes. From a forest inventory conducted in southern Brazil, data of M. scabrella were analyzed. Thirty sample trees were felled, excavated and weighed in the field and brought to laboratory for biomass and carbon determination. The total aboveground biomass represented 85% of the tree biomass, while roots corresponded to 15%. Correlation matrix of diameter at 1.3 m height (D), tree height (H) versus total and compartment biomass (P) indicated strong association between tree dimensions and biomasses. Five regression models were tested and equations were fitted to data of five biomass compartments and total tree biomass. The best fitting model for total biomass was P = –0.49361 + 0.034865 x D2H whereas for the partial biomass of the compartments was lnP = β0 + β1 x ln(D) + β2 lnH. Carbon concentration was statistically significantly different in foliage than in other compartments. Three approaches of calculating carbon stocks were evaluated and compared to actual data: 1) Estimated total biomass x weighted mean carbon concentration; 2) Estimated partial (compartment) biomass x compartment average carbon concentration; and 3) Carbon regression equations. No statistical difference was detected among them. It was concluded that biomass equations fitted in this study were accurate and useful for fuelwood and carbon estimations.
  • de Mello, Federal University of Sergipe, Brazil E-mail: aadm@nn.br
  • Nutto, Federal University of Paraná, Brazil E-mail: lnutto.ufpr@gmail.com (email)
  • Weber, Federal University of Paraná E-mail: ksw@nn.br
  • Sanquetta, Carlos Eduardo Sanquetta E-mail: ces@nn.br
  • Monteiro de Matos, Jorge Luis Monteiro de Matos E-mail: jlmdm@nn.br
  • Becker, University of Freiburg, Institute of Forest Utilization and Work Science, Germany E-mail: gb@nn.de
article id 43, category Research article
Anni Markkanen, Panu Halme. (2012). Polypore communities in broadleaved boreal forests. Silva Fennica vol. 46 no. 3 article id 43. https://doi.org/10.14214/sf.43
Keywords: birch; deciduous; slash and burn; species-area relationship; wood-inhabiting fungi
Abstract | View details | Full text in PDF | Author Info
The cover and extent of boreal broadleaved forests have been decreasing due to modern forest management practices and fire suppression. As decomposers of woody material, polypores are ecologically important ecosystem engineers. The ecology and conservation biology of polypores have been studied intensively in boreal coniferous forests. However, only a few studies have focused on the species living on broadleaved trees. To increase knowledge on this species group we conducted polypore surveys in 27 broadleaved forests and 303 forest compartments (539 ha) on the southern boreal zone in Finland and measured dead wood and forest characteristics. We detected altogether 98 polypore species, of which 13 are red-listed in Finland. 60% of the recorded species are primarily associated with broadleaved trees. The number of species in a local community present in a broadleaved forest covered approximately 50 species, of which 30–40 were primarily associated with broadleaved trees. The size of the inventoried area explained 67% of the variation in the species richness, but unlike in previous studies conducted in coniferous forests, dead wood variables as well as forest structure had very limited power in explaining polypore species richness on forest stand level. The compartments occupied by red listed Protomerulius caryae had an especially high volume of living birch, but otherwise the occurrences of red-listed species could not be predicted based on the forest structure.
  • Markkanen, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: anni.e.markkanen@gmail.com (email)
  • Halme, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: ph@nn.fi
article id 42, category Research article
Pablo Martínez-Álvarez, Fernando Manuel Alves-Santos, Julio Javier Diez. (2012). In vitro and in vivo interactions between Trichoderma viride and Fusarium circinatum. Silva Fennica vol. 46 no. 3 article id 42. https://doi.org/10.14214/sf.42
Keywords: pitch canker; antagonism; biocontrol; biological control agents; endophytes; Pinus radiata; Spain
Abstract | View details | Full text in PDF | Author Info
Fusarium circinatum, a fungus that causes pitch canker disease, has been present in Europe since at least 2003, when it was detected in northern Spain and found to be producing severe damage in tree nurseries and pine plantations. In this study, we tested a method of biological control of the disease with Trichoderma viride, a fungal species successfully used against many other pathogens. In vitro and in vivo assays were carried out to test the efficacy of this antagonist in controlling F. circinatum. The T. viride isolate exerted a significant effect on the growth of F. circinatum in the in vitro assay, reducing the length of the pathogen colony by half. However, although we tested three different concentrations of the T. viride spore solution, no clear conclusions were obtained with regard to the effects on the Pinus radiata seedlings. To our knowledge, this is the first study carried out with the aim of using Trichoderma spp. to control pitch canker disease.
  • Martínez-Álvarez, Sustainable Forest Management Research Institute, University of Valladolid – INIA, Avenida de Madrid 44, 34071 Palencia, Spain E-mail: pmtnez@pvs.uva.es (email)
  • Alves-Santos, Sustainable Forest Management Research Institute, University of Valladolid – INIA, Avenida de Madrid 44, 34071 Palencia, Spain E-mail: fmas@nn.es
  • Diez, Sustainable Forest Management Research Institute, University of Valladolid – INIA, Avenida de Madrid 44, 34071 Palencia, Spain E-mail: jjd@nn.es
article id 441, category Research article
Ane Zubizarreta-Gerendiain, Petri Pellikka, Jordi Garcia-Gonzalo, Veli-Pekka Ikonen, Heli Peltola. (2012). Factors affecting wind and snow damage of individual trees in a small management unit in Finland: assessment based on inventoried damage and mechanistic modelling. Silva Fennica vol. 46 no. 2 article id 441. https://doi.org/10.14214/sf.441
Keywords: Pinus sylvestris; Picea abies; Betula spp.; critical wind speed; snow loading; damage risk
Abstract | View details | Full text in PDF | Author Info
In this work, we assessed the factors affecting wind and snow damage of individual trees in a small management unit in western Finland. This was done based on inventoried damage and observed wind speeds and snow loading in storms Pyry and Janika in 2001 and Mielikki in 2002 together with mechanistic model. First, we studied which factors explain the observed damage in individual trees. Secondly, we studied how well the mechanistic model (HWIND) could predict the wind speed needed to uproot individual trees at the margins of permanent upwind edges. We found that Pyry storm caused 70% and Janika and Mielikki 18 and 12% of observed damage. In Janika storm, all trees uprooted. In other storms, both uprooting and stem breakage occurred. Scots pine suffered the most damage. Recently thinned stands on the upwind edges of open areas suffered the most damage. But, damage occurred also on soils with relatively shallow anchorage. HWIND predicted correctly damage for 69% of all uprooted trees. No-uprooting was correctly predicted for 45 and 19% of standing trees (all Scots pines), which were measured within and at the immediate upwind edge of same stands. HWIND model needs further validation at the permanent edges and/or on soils with shallow rooting to improve its prediction accuracy in such conditions.
  • Zubizarreta-Gerendiain, Technical University of Lisbon, School of Agriculture, Forest Research Centre, Lisbon, Portugal E-mail: azg@nn.po
  • Pellikka, University of Helsinki, Dept. of Geosciences and Geography, Helsinki, Finland E-mail: pp@nn.fi
  • Garcia-Gonzalo, Technical University of Lisbon, School of Agriculture, Forest Research Centre, Lisbon, Portugal E-mail: jgg@nn.po
  • Ikonen, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: vpi@nn.fi
  • Peltola, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: heli.peltola@uef.fi (email)
article id 58, category Research article
Ilona Suuriniemi, Jukka Matero, Harri Hänninen, Jussi Uusivuori. (2012). Factors affecting enlargement of family forest holdings. Silva Fennica vol. 46 no. 2 article id 58. https://doi.org/10.14214/sf.58
Keywords: economies of scale; structural change; forest ownership; non-industrial private forest owners; land policy; parcelization; consolidation
Abstract | View details | Full text in PDF | Author Info
This study contributes to the research of enlargement – a counterforce of parcelization – of forest holdings. To help planning policy measures aiming at increased average size of forest holdings, we study the characteristics of family forest owners who acquired additional forestland area during the years 2004–2008. Increases of forestland area due to purchases on the open market, purchases from parents or other relatives, inheritance or gift are studied. Survey data, containing information of 6318 forest owners, are analyzed with logistic regression analysis in order to establish a relationship between the probability of increasing the forestland area and the characteristics of landowners. The results indicate that young male owners, who appreciate economic values of the ownership and are active users of their forest estates, most often expand their forest property. This can be considered as an encouraging result from the point of view of the political objective to boost forest management activity through enlarging family forest holdings.
  • Suuriniemi, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: is@nn.fi
  • Matero, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: jukka.matero@uef.fi (email)
  • Hänninen, Finnish Forest Research Institute, Vantaa Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: hh@nn.fi
  • Uusivuori, Finnish Forest Research Institute, Vantaa Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: ju@nn.fi
article id 57, category Research article
Corrado Costa, Paolo Menesatti, Raffaele Spinelli. (2012). Performance modelling in forest operations through partial least square regression. Silva Fennica vol. 46 no. 2 article id 57. https://doi.org/10.14214/sf.57
Keywords: chipping; productivity; harvesting; skidding
Abstract | View details | Full text in PDF | Author Info
Partial Least Square (PLS) regression is a recent soft-modelling technique that generalizes and combines features from principal component analysis (PCA) and multiple regression. It is particularly useful when predicting one or more dependent variables from a large set of independent variables, often collinear. The authors compared the potential of PLS regression and ordinary linear regression for accurate modelling of forest work, with special reference to wood chipping, wood extraction and the continuous harvesting of short rotation coppice. Compared to linear regression, PLS regression allowed producing models that better fit the original data. What is more, it allowed handling collinear variables, facilitating the extraction of sound models from large amounts of field data obtained from commercial forest operations. On the other hand, PLS regression analysis is not as easy to conduct, and produces models that are less user-friendly. By producing alternative models, PLS regression may provide additional – and not alternative – ways of reading the data. Ideally, a comprehensive data analysis could include both ordinary and PLS regression and proceed from their results in order to get a better understanding of the phenomenon under examination. Furthermore, the computational complexity of PLS regression may stimulate interdisciplinary team-building, to the greater benefit of scientific research within the field of forest operations.
  • Costa, CRA ING, Monterotondo Scalo (Roma), Italy E-mail: cc@nn.it
  • Menesatti, CRA ING, Monterotondo Scalo (Roma), Italy E-mail: pm@nn.it
  • Spinelli, CNR IVALSA, Sesto Fiorentino (FI), Italy E-mail: spinelli@ivalsa.cnr.it (email)
article id 56, category Research article
Johan Holmgren, Andreas Barth, Henrik Larsson, Håkan Olsson. (2012). Prediction of stem attributes by combining airborne laser scanning and measurements from harvesters. Silva Fennica vol. 46 no. 2 article id 56. https://doi.org/10.14214/sf.56
Keywords: ALS data; pre-harvest inventory; tree detection
Abstract | View details | Full text in PDF | Author Info
In this study, a new method was validated for the first time that predicts stem attributes for a forest area without any manual measurements of tree stems by combining harvester measurements and Airborne Laser Scanning (ALS) data. A new algorithm for automatic segmentation of tree crowns from ALS data based on tree crown models was developed. The test site was located in boreal forest (64°06’N, 19°10’E) dominated by Norway spruce (Picea abies) and Scots Pine (Pinus sylvestris).The trees were harvested on field plots, and each harvested tree was linked to the nearest tree crown segment derived from ALS data. In this way, a reference database was created with both stem data from the harvester and ALS derived features for linked tree crowns. To estimate stem attributes for a tree crown segment in parts of the forest where trees not yet have been harvested, tree stems are imputed from the most similar crown segment in the reference database according to features extracted from ALS data. The imputation of harvester data was validated on a sub-stand-level, i.e. 2–4 aggregated 10 m radius plots, and the obtained RMSE of stem volume, mean tree height, mean stem diameter, and stem density (stems per ha) estimates were 11%, 8%, 12%, and 19%, respectively. The imputation of stem data collected by harvesters could in the future be used for bucking simulations of not yet harvested forest stands in order to predict wood assortments.
  • Holmgren, Swedish University of Agricultural Sciences, Forest Resource Management, Umeå, Sweden E-mail: johan.holmgren@slu.se (email)
  • Barth, The Forestry Research Institute of Sweden, Uppsala, Sweden E-mail: ab@nn.se
  • Larsson, Swedish University of Agricultural Sciences, Forest Resource Management, Umeå, Sweden E-mail: hl@nn.se
  • Olsson, Swedish University of Agricultural Sciences, Forest Resource Management, Umeå, Sweden E-mail: ho@nn.se
article id 55, category Research article
Antti Mäkinen, Annika Kangas, Mikko Nurmi. (2012). Using cost-plus-loss analysis to define optimal forest inventory interval and forest inventory accuracy. Silva Fennica vol. 46 no. 2 article id 55. https://doi.org/10.14214/sf.55
Keywords: value of information; prediction error; inventory error
Abstract | View details | Full text in PDF | Author Info
In recent years, optimal inventory accuracy has been analyzed with a cost-plus-loss methodology, where the total costs of inventory include both the measurement costs and the losses from the decisions based on the collected information. Losses occur, when the inaccuracies in the data lead to sub-optimal decisions. In almost all cases, it has been assumed that the accuracy of the once collected data remains the same throughout the planning period, and the period has been from 10 up to 100 years. In reality, the quality of the data deteriorates in time, due to errors in the predicted growth. In this study, we carried out a cost-plus-loss analysis accounting for the errors in (stand-level) growth predictions of basal area and dominant height. In addition, we included the inventory errors of these two variables with several different levels of accuracy, and costs of inventory with several different assumptions of cost structure. Using the methodology presented in this study, we could calculate the optimal inventory interval (life-span of data) minimizing the total costs of inventory and losses through the 30-year planning period. When the inventory costs only to a small extent depended on the accuracy, the optimal inventory period was 5 years and optimal accuracy RMSE 0%. When the costs more and more heavily depended on the accuracy, the optimal interval turned out to be either 10 or 15 years, and the optimal accuracy reduced from RMSE 0% to RMSE 20%. By increasing the accuracy of the growth models, it was possible to reduce the inventory accuracy or lengthen the interval, i.e. obtain clear savings in inventory costs.
  • Mäkinen, Simosol Oy, Rautatietori 4, FI-11130 Riihimäki, Finland E-mail: antti.makinen@simosol.fi (email)
  • Kangas, University of Helsinki, Department of Forest Sciences, Helsinki, Finland E-mail: ak@nn.fi
  • Nurmi, University of Helsinki, Department of Forest Sciences, Helsinki, Finland E-mail: mn@nn.fi
article id 54, category Research article
Urban Nilsson, Björn Elfving, Kjell Karlsson. (2012). Productivity of Norway spruce compared to Scots pine in the interior of northern Sweden. Silva Fennica vol. 46 no. 2 article id 54. https://doi.org/10.14214/sf.54
Keywords: Pinus sylvestris; current annual increment; mean annual increment; Picea abies; site index; yield
Abstract | View details | Full text in PDF | Author Info
Productivity of Norway spruce (Picea abies L. Karst.) and Scots pine (Pinus sylvestris L.) was studied in 12 paired plots in the interior of northern Sweden. Stands were established between 1928 and 1959; yield plots were established between 1974 and 1983 during pre-commercial thinning of the stands. Gross stem-wood production was significantly higher for Scots pine than for Norway spruce, stem-wood production by Norway spruce being 29.4% that of Scots pine. The site index for Norway spruce was lower than for Scots pine at all sites except one; the average difference in site index was 4.8 m. The simulated maximum mean annual increment (MAImax) during the rotation was 19% higher than the MAImax estimated with the site index for Scots pine, whereas simulated MAImax and MAImax estimated from the site index was about the same for Norway spruce. The simulations also indicated that MAI peaked about 50 years later for Norway spruce than for Scots pine. More small trees were included in the diameter distribution of Norway spruce than of Scots pine resulting in a lower stem-wood volume for Norway spruce when stands with the same dominant height were compared. This study shows that the difference in growth and rotation length between Scots pine and Norway spruce has implications when choosing which species to grow in the interior of northern Sweden.
  • Nilsson, Swedish University of Agricultural Sciences (SLU), Southern Swedish Forest Research Centre, Alnarp, Sweden E-mail: urban.nilsson@slu.se (email)
  • Elfving, SLU, Forest Ecology and Management, Umeå, Sweden E-mail: be@nn.se
  • Karlsson, SLU, Unit of Field Based Research, Asa, Sweden E-mail: kk@nn.se
article id 53, category Research article
Saana Kataja-aho, Aino Smolander, Hannu Fritze, Sini Norrgård, Jari Haimi. (2012). Responses of soil carbon and nitrogen transformations to stump removal. Silva Fennica vol. 46 no. 2 article id 53. https://doi.org/10.14214/sf.53
Keywords: biofuel; boreal forest; carbon; nitrogen; nutrient dynamics; stump removal
Abstract | View details | Full text in PDF | Author Info
We studied in central Finland whether stump harvesting after clear felling of coniferous forest poses further short-term changes in soil carbon and nitrogen dynamics when compared to the traditional site preparation method, mounding. Exposed mineral soil patches in Norway spruce (Picea abies) dominated clear-cut stands were sampled 1–5 years after the treatments. The extent of the exposed mineral soil surface was significantly larger at the stump removal sites when compared to the mounding sites. No differences were found in soil pH, organic matter content or total concentration of soil C between the treatments or treatment years. Total concentration of soil N was consistently higher and C:N ratio lower in the stump removal plots than in the mounded plots. Further, both net N mineralisation and nitrification were clearly increased in the stump removal plots one year after the treatments. Soil microbial activity (CO2 production) was higher in the stump removal plots but similar difference was not found in sieved soil samples incubated in the laboratory. Fluxes of other important greenhouse gases (CH4 and N2O) did not seem to be affected by stump removal. The differences between the stump removal and mounding procedures were most obviously attributed to more substantial soil disturbance by stump pulling and/or differences in the microbial communities and quality of soil organic matter in the differently treated soil.
  • Kataja-aho, University of Jyväskylä, Dept. of Biological and Environmental Science, Jyväskylä, Finland E-mail: saana.m.kataja-aho@jyu.fi (email)
  • Smolander, Finnish Forest Research Institute, Vantaa, Finland E-mail: as@nn.fi
  • Fritze, Finnish Forest Research Institute, Vantaa, Finland E-mail: hf@nn.fi
  • Norrgård, University of Jyväskylä, Dept. of Biological and Environmental Science, Jyväskylä, Finland E-mail: sn@nn.fi
  • Haimi, University of Jyväskylä, Dept. of Biological and Environmental Science, Jyväskylä, Finland E-mail: jh@nn.fi
article id 444, category Research article
Sovu, Mulualem Tigabu, Patrice Savadogo, Per Christer Odén. (2012). Facilitation of forest landscape restoration on abandoned swidden fallows in Laos using mixed-species planting and biochar application. Silva Fennica vol. 46 no. 1 article id 444. https://doi.org/10.14214/sf.444
Keywords: black carbon; charcoal; rice husk biochar; secondary forest; shifting cultivation
Abstract | View details | Full text in PDF | Author Info
The cessation of swidden cultivation and the increasing trend of abandonment of swidden fallows have created an opportunity for forest landscape restoration. However, ways need to be found to improve the poor soil fertility at these sites with affordable materials and to generate short-term socio-economic benefits for small-scale swidden fallow holders. This study assessed the feasibility of using mixed-planting of eight native species and application of rice husk biochar as soil amendment measure at a site in Laos. The effect of biochar application was compared against addition of inorganic (NPK) fertilizer and the control. The establishment and growth of the planted seedlings was then monitored for four years. The addition of rice husk biochar and NPK fertilizer did not significantly (p = 0.578) improve the survival rate of planted seedlings, which ranged from 72% to 91% (depending on the species) compared to the control. No significant growth responses to the soil amendments were observed for most of the species during the first year after planting compared to the control. The biochar effect was, however, more evident at the fourth year for diameter (p < 0.01) and height (p < 0.01) of sapling for all species; particularly its effect was more vivid on the diameter of slow-growing species. The results indicate that the species tested in the mixed-planting showed marked growth variation while application of rice husk biochar boosted their growth. Thus, planting mixed-species in swidden fallows has potential to provide continuous supplies of wood from different species to diversify the livelihood of swidden field owners, while maintaining ecosystem services.
  • -, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, Alnarp, Sweden E-mail: s@nn.se
  • Tigabu, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, Alnarp, Sweden E-mail: mulualem.tigabu@slu.se (email)
  • Savadogo, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, Alnarp, Sweden E-mail: ps@nn.se
  • Odén, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, Alnarp, Sweden E-mail: pco@nn.se
article id 70, category Research article
Joonas Järvinen, Jaakko Linnakangas. (2012). Firm capabilities in the Finnish forest cluster: comparisons based on self-organizing map. Silva Fennica vol. 46 no. 1 article id 70. https://doi.org/10.14214/sf.70
Keywords: forest cluster; capability; content analysis; self-organizing map
Abstract | View details | Full text in PDF | Author Info
This paper examines the capability portfolios of Finnish forest cluster firms in 1998–2008. In particular, the focus is on what kind of capabilities the firms in the cluster have developed, whether the firms have developed such capabilities consistently, and whether they have developed similar capability portfolios. Further, a particular focus is on the links between innovativeness and other identified capabilities. We approach the topic with an exploratory quantitative analysis of the annual reports of 11 large firms in the cluster by using computerized content analysis and a self-organizing map as the main research methods. Based on the content analysis and earlier literature, we first identify fifteen capabilities, and then build capability portfolios for the firms on the basis of the results of the self-organizing map. At the firm level, the results reveal both similarities and differences in the capability portfolios of the forest cluster firms. Similarly, we identify both continuity and change in the development of the portfolios. At the cluster level, the focus is on innovativeness capability and its relationship with other capabilities. The results suggest that especially being strategic, market-oriented, and monitoring the external environment are positively related with innovativeness; in contrast, a focus on finance and change seems to suppress innovativeness.
  • Järvinen, Department of Industrial Engineering and Management, Aalto University, P.O.Box 15500, FI-00076 Aalto, Finland E-mail: joonas.jarvinen@aalto.fi (email)
  • Linnakangas, Department of Industrial Engineering and Management, Aalto University, P.O.Box 15500, FI-00076 Aalto, Finland E-mail: jl@nn.fi
article id 69, category Research article
Tarja Wallenius, Risto Laamanen, Jussi Peuhkurinen, Lauri Mehtätalo, Annika Kangas. (2012). Analysing the agreement between an Airborne Laser Scanning based forest inventory and a control inventory – a case study in the state owned forests in Finland. Silva Fennica vol. 46 no. 1 article id 69. https://doi.org/10.14214/sf.69
Keywords: forest inventory; quality assessment; airborne laser scanning
Abstract | View details | Full text in PDF | Author Info
Airborne laser scanning based forest inventories have recently shown to produce accurate results. However, the accuracy varies according to the test area and used methodology and therefore, an unambiguous and practical quality assessment will be needed as a part of each inventory project. In this study, the accuracy of an ALS inventory was evaluated with a field sampling based control inventory. The agreement between the ALS inventory and the control inventory was analysed with four methods: 1) root mean square error (RMSE) and bias, 2) scatter plots with 95% confidence intervals, 3) Bland-Altman plots and 4) tolerance limits within Bland-Altman plots. Each method has its own special features which have to be taken into account when the agreement is analysed. The pre-defined requirements of the ALS inventory were achieved. A simplified control inventory approach with a slightly narrower focus is proposed to be used in the future. The Bland-Altman plots with the tolerance limits are proposed to be used in quality assessments of operational ALS inventories. Further studies to improve the efficiency of quality assessment are needed.
  • Wallenius, Metsähallitus, P.O. Box 94, FI-01301 Vantaa, Finland E-mail: tarja.wallenius@metsa.fi (email)
  • Laamanen, Metsähallitus, P.O. Box 94, FI-01301 Vantaa, Finland E-mail: rl@nn.fi
  • Peuhkurinen, Oy Arbonaut Ltd, Helsinki, Finland E-mail: jp@nn.fi
  • Mehtätalo, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: lm@nn.fi
  • Kangas, University of Helsinki, Department of Forest Sciences, Helsinki, Finland E-mail: ak@nn.fi
article id 68, category Research article
Maria Villikka, Petteri Packalén, Matti Maltamo. (2012). The suitability of leaf-off airborne laser scanning data in an area-based forest inventory of coniferous and deciduous trees. Silva Fennica vol. 46 no. 1 article id 68. https://doi.org/10.14214/sf.68
Keywords: forest inventory; group-specific estimates; leaf-off data; LIDAR
Abstract | View details | Full text in PDF | Author Info
This study examined the suitability of airborne laser scanner (ALS) data collected under leaf-off conditions in a forest inventory, in which deciduous and coniferous trees need to be separated. All analyses were carried out with leaf-on and leaf-off ALS data collected from the same study area. Additionally, aerial photographs were utilized in the Nearest Neighbor (NN) imputations. An area-based approach was used in this study. Regression estimates of plot volume were more accurate in the case of leaf-off than leaf-on data. In addition, regression models were more accurate in coniferous plots than in deciduous plots. The results of applying leaf-on models with leaf-off data, and vice versa, indicate that leaf-on and leaf-off data should not be combined since this causes serious bias. The total volume and volume by coniferous and deciduous trees was estimated by the NN imputation. In terms of total volume, leaf-off data provided more accurate estimates than leaf-on data. In addition, leaf-off data discriminated between coniferous and deciduous trees, even without the use of aerial photographs. Accurate results were also obtained when leaf-off ALS data were used to classify sample plots into deciduous and coniferous dominated plots. The results indicate that the area-based method and ALS data collected under leaf-off conditions are suitable for forest inventory in which deciduous and coniferous trees need to be distinguished.
  • Villikka, University of Easten Finland, Department of Forest Sciences, Joensuu, Finland E-mail: mv@nn.fi
  • Packalén, University of Easten Finland, Department of Forest Sciences, Joensuu, Finland E-mail: petteri.packalen@uef.fi (email)
  • Maltamo, University of Easten Finland, Department of Forest Sciences, Joensuu, Finland E-mail: mm@nn.fi
article id 67, category Research article
Pentti Niemistö, Heikki Korpunen, Ari Laurén, Marika Salomäki, Jori Uusitalo. (2012). Impact and productivity of harvesting while retaining young understorey spruces in final cutting of downy birch (Betula pubescens). Silva Fennica vol. 46 no. 1 article id 67. https://doi.org/10.14214/sf.67
Keywords: Picea abies; harvesting; Betula pubescens; logging; spruce releasing
Abstract | View details | Full text in PDF | Author Info
  • Niemistö, Finnish Forest Research Institute, Parkano & Joensuu, Finland E-mail: pn@nn.fi
  • Korpunen, Finnish Forest Research Institute, Parkano & Joensuu, Finland E-mail: hk@nn.fi
  • Laurén, Finnish Forest Research Institute, Parkano & Joensuu, Finland E-mail: al@nn.fi
  • Salomäki, Finnish Forest Research Institute, Parkano & Joensuu, Finland E-mail: ms@nn.fi
  • Uusitalo, Finnish Forest Research Institute, Parkano & Joensuu, Finland E-mail: jori.uusitalo@metla.fi (email)
article id 66, category Research article
Monika Strömgren, Kristina Mjöfors, Björn Holmström, Achim Grelle. (2012). Soil CO2 flux during the first years after stump harvesting in two Swedish forests. Silva Fennica vol. 46 no. 1 article id 66. https://doi.org/10.14214/sf.66
Keywords: stump removal; Picea abies; C stock; site preparation; soil C; soil disturbance; soil respiration
Abstract | View details | Full text in PDF | Author Info
One way of increasing the supply of renewable energy, thereby decreasing the use of fossil fuels, is to extract the stumps that remain after final stem harvesting. However, little is known about the environmental consequences of stump harvesting, and how ecosystem services, such as carbon sequestration, are affected by the practice. In the present paper, the effects on the soil carbon pool during the first months and years after stump harvesting in former Norway spruce stands are presented. The study was performed at two sites in mid- and southern Sweden. At both sites, the soil CO2 flux was measured on several occasions with a portable respiration system, to compare plots on which stump harvesting had occurred, with reference plots. At one of the sites, CO2 exchange was also followed continuously by means of eddy-covariance measurements before and after stump harvesting. Since there was no vegetation at the beginning of the study, almost all emitted CO2 could be assumed to come from heterotrophic sources, and the soil CO2 flux was measured. This study shows that the effect of stump harvesting on CO2 flux or soil decomposition processes is small or absent compared to site preparation such as mounding in a short-term perspective of months and years. The long-term consequences of stump harvesting are, however, still uncertain.
  • Strömgren, Swedish University of Agricultural Sciences, Department of Soil and Environment, Uppsala, Sweden E-mail: monika.stromgren@slu.se (email)
  • Mjöfors, Swedish University of Agricultural Sciences, Department of Soil and Environment, Uppsala, Sweden E-mail: km@nn.se
  • Holmström, Swedish University of Agricultural Sciences, Department of Soil and Environment, Uppsala, Sweden E-mail: bh@nn.fi
  • Grelle, Swedish University of Agricultural Sciences, Department of Ecology, Uppsala, Sweden E-mail: ag@nn.se
article id 65, category Research article
Kristóf Kelemen, Barbara Mihók, László Gálhidy, Tibor Standovár. (2012). Dynamic response of herbaceous vegetation to gap opening in a Central European beech stand. Silva Fennica vol. 46 no. 1 article id 65. https://doi.org/10.14214/sf.65
Keywords: continuous cover forestry; dispersal; gap colonization; herbs; seed bank type; species richness
Abstract | View details | Full text in PDF | Author Info
Herbaceous ground vegetation in artificially-created gaps was studied in a managed beech (Fagus sylvatica L.) forest over a period of eight years in Northern Hungary, Central Europe. These gaps were being used as an alternative to the regular shelterwood system to create uneven-aged stands. The effects of gap size (15 and 40 m diameter) and canopy openness on herbaceous species colonization and persistence were assessed in a systematic grid of 5 5 m. Overall, herbaceous cover was low before gap creation, increased soon afterwards, and continued to rise over time. The number of herb species increased in the gaps and, to a lesser extent, in adjacent areas under the remaining tree canopy. Colonization of gaps was rapid and there was substantial turnover of species i.e. various species disappeared from the gaps over time whilst others colonized. Species with both long-term persistent seed banks and long distance dispersal abilities were the most successful types colonizing gaps. Six species occurred preferentially in large gaps, while only one species was found to prefer small gaps. Species present before gap creation survived in both gap sizes. Smaller gaps with a diameter of half the height of canopy trees also tended to remain free of common weed species, whereas large cover of Rubus fruticosus L. and Calamagrostis epigejos (L.) Roth could hamper natural regeneration in larger gaps. For the successful regeneration of beech we recommend the use of small gaps complemented by few large gaps.
  • Kelemen, Loránd Eötvös University, Institute of Biology, Department of Plant Systematics, Ecology and Theoretical Biology, Budapest, Hungary E-mail: kk@nn.hu
  • Mihók, Loránd Eötvös University, Institute of Biology, Department of Plant Systematics, Ecology and Theoretical Biology, Budapest, Hungary E-mail: bm@nn.hu
  • Gálhidy, Loránd Eötvös University, Institute of Biology, Department of Plant Systematics, Ecology and Theoretical Biology, Budapest, Hungary E-mail: lg@nn.hu
  • Standovár, Institute of Biology, Department of Plant Systematics, Ecology and Theoretical Biology, Budapest, Hungary E-mail: standy@ludens.elte.hu (email)
article id 64, category Research article
Karin Johansson, Ola Langvall, Johan Bergh. (2012). Optimization of environmental factors affecting initial growth of Norway spruce seedlings. Silva Fennica vol. 46 no. 1 article id 64. https://doi.org/10.14214/sf.64
Keywords: fertilization; establishment; irrigation; regeneration; seedling types; vegetation management
Abstract | View details | Full text in PDF | Author Info
  • Johansson, Skogforsk, Svalöv, Sweden E-mail: karin.johansson@skogforsk.se (email)
  • Langvall, SLU, Asa Forest Research Station, Sweden E-mail: ol@nn.se
  • Bergh, SLU, Southern Swedish Forest Research Centre, Sweden E-mail: jb@nn.se
article id 63, category Research article
Leena Koivuranta, Tarja Latva-Karjanmaa, Pertti Pulkkinen. (2012). The effect of temperature on seed quality and quantity in crosses between European (Populus tremula) and hybrid aspens (P. tremula x P. tremuloides). Silva Fennica vol. 46 no. 1 article id 63. https://doi.org/10.14214/sf.63
Keywords: climate change; gene flow; germination; hybrid vigour
Abstract | View details | Full text in PDF | Author Info
Hybrid aspen (Populus tremula L. Populus tremuloides Michx.) plantations are expanding in Fennoscandia and the Baltic countries; however, the possible effects of plantations on the native European aspen (P. tremula) and the level of gene flow between European and hybrid aspen have not been investigated. We studied seed quantity and quality in intraspecific and interspecific crosses of the European and hybrid aspens over a two year period. In order to study whether elevated temperatures due to climate change would benefit the species differently, we performed the crosses in different temperatures. In both years, interspecific crosses produced more seeds with higher quality than intraspecific crosses. This result was most distinct in crosses between female hybrid aspen and male European aspen. In higher temperatures, relative germination difference between hybrid aspen seeds and seeds from P. tremula P. tremula crosses seems to increase. These results suggest that hybrid aspen may have a significant genetic impact on the European aspen, and this effect may be strengthened by climate warming.
  • Koivuranta, Finnish Forest Research Institute, Haapastensyrjä, Finland E-mail: lk@nn.fi
  • Latva-Karjanmaa, Finnish Forest Research Institute, Haapastensyrjä, Finland E-mail: tlk@nn.fi
  • Pulkkinen, Finnish Forest Research Institute, Haapastensyrjä, Finland E-mail: pertti.pulkkinen@metla.fi (email)
article id 62, category Research article
Nadia Goué, Nathalie Noël-Boizot, Michel Vallance, Elisabeth Magel, Philippe Label. (2012). Microdissection to isolate vascular cambium cells in poplar. Silva Fennica vol. 46 no. 1 article id 62. https://doi.org/10.14214/sf.62
Keywords: fusiform cambial cells; microgenomics; Populus spp.; ray cambial cells
Abstract | View details | Full text in PDF | Author Info
Vascular cambium is the lateral meristem producing xylem cells inwards and phloem cells outwards in plant stem. Thus, in trees, the quality and quantity of wood is a result of highly regulated developmental process depending initially on the vascular cambium cell production. The availability of accurate transcriptomics technologies based on high coverage sequencing raises the level of expectations on tissue sampling to a very high degree. What is the benefit of top-level transcriptomics in wood formation studies if we are using these technologies on raw tissues, mixing cells at the organ level or even higher scale? The presented work describes a nine-step procedure, from standing tree to isolated ray and fusiform cells from cryolyophilized tangential sections of the poplar cambial zone. The aim of this paper is to present a step by step procedure including advices on how to select the optimal tree, how to fell the tree while securing its physiological parameters, how to cryolyophilize and microdissect under binocular, presenting the time schedule of the whole process and RNA analysis.
  • Goué, UMR5546 CNRS-Université Paul Sabatier, Castanet Tolosan, France; (INRA-UAGPF, 2163 Avenue de la Pomme de Pin, 45075 Orléans cedex 2, France) E-mail: goue@lrsv.ups-tlse.fr (email)
  • Noël-Boizot, INRA-UAGPF, 2163 Avenue de la Pomme de Pin, 45075 Orléans cedex 2, France E-mail: nnb@nn.fr
  • Vallance, INRA-UAGPF, 2163 Avenue de la Pomme de Pin, 45075 Orléans cedex 2, France E-mail: mv@nn.fr
  • Magel, Universität Hamburg, Zentrum Holzwirtschaft, Deutschland E-mail: em@nn.de
  • Label, INRA-UAGPF, 2163 Avenue de la Pomme de Pin, 45075 Orléans cedex 2, France E-mail: pl@nn.fr
article id 90, category Research article
Per Angelstam, Kjell Andersson, Robert Axelsson, Marine Elbakidze, Bengt Gunnar Jonsson, Jean-Michel Roberge. (2011). Protecting forest areas for biodiversity in Sweden 1991–2010: the policy implementation process and outcomes on the ground. Silva Fennica vol. 45 no. 5 article id 90. https://doi.org/10.14214/sf.90
Keywords: forest policy; forest protection; restoration ecology; connectivity; green infrastructure; umbrella species; forest disturbance regimes; participation and collaboration
Abstract | View details | Full text in PDF | Author Info
Swedish forest and environmental policies imply that forests should be managed so that all naturally occurring species are maintained in viable populations. This requires maintenance of functional networks of representative natural forest and cultural woodland habitats. We first review the policy implementation process regarding protected areas in Sweden 1991–2010, how ecological knowledge was used to formulate interim short-term and strategic long-term biodiversity conservation goals, and the development of a hierarchical spatial planning approach. Second, we present data about the amount of formally protected and voluntarily set aside forest stands, and evaluate how much remains in terms of additional forest protection, conservation management and habitat restoration to achieve forest and environmental policy objectives in the long-term. Third, a case study in central Sweden was made to estimate the functionality of old Scots pine, Norway spruce and deciduous forest habitats, as well as cultural woodland, in different forest regions. Finally, we assess operational biodiversity conservation planning processes. We conclude that Swedish policy pronouncements capture the contemporary knowledge about biodiversity and conservation planning well. However, the existing area of protected and set-aside forests is presently too small and with too poor connectivity. To bridge this gap, spatial planning, management and restoration of habitat, as well as collaboration among forest and conservation planners need to be improved.
  • Angelstam, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden E-mail: per.angelstam@slu.se (email)
  • Andersson, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden E-mail: ka@nn.se
  • Axelsson, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden E-mail: ra@nn.se
  • Elbakidze, School for Forest Management, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Skinnskatteberg, Sweden E-mail: me@nn.se
  • Jonsson, Dept of Natural Science, Engineering and Mathematics, Mid Sweden University, Sundsvall, Sweden E-mail: bgj@nn.se
  • Roberge, Dept of Wildlife, Fish and Environmental Studies, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Umeå, Sweden E-mail: jmr@nn.se
article id 89, category Research article
Russell Grenfell, Tuomas Aakala, Timo Kuuluvainen. (2011). Microsite occupancy and the spatial structure of understorey regeneration in three late-successional Norway spruce forests in northern Europe. Silva Fennica vol. 45 no. 5 article id 89. https://doi.org/10.14214/sf.89
Keywords: boreal forest; disturbance; microsite; spatial correlation; stand structure; natural forest
Abstract | View details | Full text in PDF | Author Info
We compared microsite occupancy and three spatial structure of regeneration in three primeval late-successional Norway spruce dominated forests. One area lay in the middle boreal zone in Russia (Dvina-Pinega) where larger-scale disturbance from bark beetles and drought had occurred; the other areas lay in the northern boreal zone, one in Finland (Pallas-Ylläs) had encountered only small-scale disturbance, and one in Russia (Kazkim) had been influenced by fire. We mapped all spruce (Picea abies) and birch (Betula pendula and Betula pubescens) trees with diameter at breast height (DBH) ≥ 10 cm on 40 m 400 m plots, and those with DBH < 10 cm on 2 m or 4 m 400 m subplots. On the subplots we also recorded microsite occupancy and estimated microsite availability. At all study areas small seedlings (h < 0.3 m) of both spruce and birch were found largely on disturbance-related microsites. Birch saplings (h ≥ 1.3 m, DBH < 10 cm) disproportionately occupied deadwood-related microsites at Dvina-Pinega. In contrast, spruce saplings at all study areas, and birch saplings at Kazkim and Pallas-Ylläs, showed less, or no, preference. Our results thus confirm the importance of disturbance-related microsites for regeneration establishment, but not necessarily for long-term survival. No spatial segregation between the overstorey (DBH ≥ 10 cm) and saplings (h ≥ 1.3 m, DBH < 10 cm) or seedlings (h < 1.3 m) was found at Pallas-Ylläs or Kazkim, and only three instances of very weak segregation were found at Dvina-Pinega. This suggests that the regeneration gap concept may not be useful for describing the regeneration dynamics of primeval boreal forests.
  • Grenfell, University of Helsinki, Dept of Forest Sciences, Helsinki, Finland E-mail: russell.grenfell@helsinki.fi (email)
  • Aakala, University of Helsinki, Dept of Forest Sciences, Helsinki, Finland E-mail: ta@nn.fi
  • Kuuluvainen, University of Helsinki, Dept of Forest Sciences, Helsinki, Finland E-mail: tk@nn.fi
article id 88, category Research article
Seppo Rouvinen, Jari Kouki. (2011). Tree regeneration in artificial canopy gaps established for restoring natural structural variability in a Scots pine stand. Silva Fennica vol. 45 no. 5 article id 88. https://doi.org/10.14214/sf.88
Keywords: ecosystems; diversity; restoration
Abstract | View details | Full text in PDF | Author Info
In Finland and elsewhere in Europe, many protected forest areas include also stands that were previously managed and that lack several naturally occurring stand characteristics. In these areas, ecosystem restoration can be used to facilitate and accelerate the formation of structural and habitat features resembling those of natural forests. For example, by creating small gaps it could be possible to diversify forest structure and tree species composition and to produce dead wood while still maintaining mostly continous canopy coverage. We examined experimentally the effects of artificial gap formation on post-disturbance tree regeneration in the gaps in a young protected, but formerly commercially managed pine (Pinus sylvestris L.) dominated forest. In the experimental sites, gap size and the portion of girdled trees out of all treated trees (girdled and felled trees combined) in the gaps varied. Natural and artificial (direct seeding of silver birch Betula pendula Roth) tree regeneration and development was monitored both on disturbed (scarified soil patches) and undisturbed forest floor during three growing seasons. Results show that gaps can be valuable in diversifying stand structure but to be successful and rapid, tree regeneration needs disturbed forest floor. Pine regenerated numerously, but birch had clearly lower regeneration, especially on small-sized gaps. In conclusion, increasing tree diversity in young pine-dominated forests seems to be difficult when only small artificial gaps are used. But even small gaps can be used to create and maintain diverse cohort structure of the dominant species and thus they can contribute to restoration goals.
  • Rouvinen, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: seppo.rouvinen@uef.fi (email)
  • Kouki, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: jk@nn.fi
article id 87, category Research article
Radek Bace, Miroslav Svoboda, Pavel Janda. (2011). Density and height structure of seedlings in subalpine spruce forests of Central Europe: logs vs. stumps as a favourable substrate. Silva Fennica vol. 45 no. 5 article id 87. https://doi.org/10.14214/sf.87
Keywords: Norway spruce; Picea abies; coarse woody debris; microsite; natural regeneration
Abstract | View details | Full text in PDF | Author Info
Decaying logs and stumps provide an important seedling substrate in natural subalpine forests. However, only stumps present such a role in managed forests. The aim of this study was to assess the differences in the process of seedling colonization between logs and stumps. The study was carried out in the Czech Republic, in two old-growth subalpine spruce forests located in the Bohemian Forest and Ash Mts., dominated by Athyrium distentifolium Opiz and Vaccinium myrtillus L. undergrowth, respectively. Norway spruce (Picea abies (L.) Karst.) regeneration growing on logs, stumps and non-coarse woody debris (CWD) microsites was surveyed. Regeneration (height 0–2.0 m) densities exceeded 5000 individuals per ha on both sites. The average density of P. abies regeneration per square meter of substrate was 0.3-5.7-19.6 and 0.5-3.8-11.0 on non-CWD microsites, logs and stumps, located in A. distentifolium and V. myrtillus undergrowth, respectively. Stumps and non-CWD microsites dominated by V. myrtillus, supported a higher proportion of taller seedlings per plot compared to the small seedlings growing on logs and non-CWD dominated by A. distentifolium ground-cover. The disproportion in regeneration densities between the stumps and the original logs decreased with increasing stages of decay. The tallest regeneration growing on stumps (root-soil plates) was significantly older than that growing on the logs (stems). Based on these two latter findings, the stumps appeared to provide suitable seedling substrates several years earlier than the logs did. Therefore, we conclude that the stumps play a more important role (relative to their covered area, 21–28 m2 ha–1) in terms of suitable microsites for regeneration, than the logs do.
  • Bace, Czech University of Life Sciences, Faculty of Forestry and Wood Sciences, Prague, Czech Republic E-mail: bace@fld.czu.cz (email)
  • Svoboda, Czech University of Life Sciences, Faculty of Forestry and Wood Sciences, Prague, Czech Republic E-mail: ms@nn.cz
  • Janda, Czech University of Life Sciences, Faculty of Forestry and Wood Sciences, Prague, Czech Republic E-mail: pv@nn.cz
article id 86, category Research article
Mats Jonsell, Jesper Hansson. (2011). Logs and stumps in clearcuts support similar saproxylic beetle diversity: implications for bioenergy harvest. Silva Fennica vol. 45 no. 5 article id 86. https://doi.org/10.14214/sf.86
Keywords: biodiversity; beetles; bioenergy; Coleoptera; logs; insects; stumps
Abstract | View details | Full text in PDF | Author Info
Stumps from clear cuts are increasingly used for bioenergy. Extracting this wood will reduce the habitat available for saproxylic (wood-living) organisms. As little is known about the species assemblages that will be affected, we investigated the diversity of saproxylic beetles in stumps on clear-felled sites and as a reference, we compared it with the diversity in downed logs. Stumps and logs of aspen (Populus tremula L.), birch (Betula pubescens Ehrh. and B. verrucosa Ehrh.[syn. B. pendula Roth]), spruce (Picea abies (L.) Karst.) and pine (Pinus sylvestris L.) were examined in clear cuts of two different ages: one summer old and 4–5 years old. The beetles were sampled by sieving bark (0.25 m2) peeled from the wood. The samples were taken in pairs of one log and one stump situated close together and of the same tree species, age since death and diameter. In total 3348 saproxylic beetles belonging to 124 species were found in 176 samples. The stumps had a similar number of species to the logs both as measured per sample and as an accumulated number. Exceptions were 4–5 years old wood of birch and pine where the number was significantly higher in the stumps. The number of red-listed species was also similar between stumps and logs. Species composition was more different between the stumps and logs of conifers than of deciduous trees. We conclude that clear-felled stumps have a diverse saproxylic insect fauna. This has to be taken into account if large scale extraction of logging stumps is implemented.
  • Jonsell, Swedish University of Agrarian Sciences, Dept of Ecology, Uppsala, Sweden E-mail: mats.jonsell@ekol.slu.se (email)
  • Hansson, Swedish University of Agrarian Sciences, Dept of Ecology, Uppsala, Sweden E-mail: jh@nn.se
article id 85, category Research article
Ville A.O. Selonen, Maija Mussaari, Tero Toivanen, Janne S. Kotiaho. (2011). The conservation potential of brook-side key habitats in managed boreal forests. Silva Fennica vol. 45 no. 5 article id 85. https://doi.org/10.14214/sf.85
Keywords: forest management; saproxylic species; dead wood; vegetation
Abstract | View details | Full text in PDF | Author Info
Today, maintaining biodiversity is included in the targets of boreal forest management. A widespread approach in northern Europe is to identify and preserve woodland key habitats within managed forests. Woodland key habitats are expected to be patches that host populations of threatened and declining species, and the preservation of these patches is assumed to enable the persistence of the focal species in the landscape. In Finland, the criteria for selecting woodland key habitats are defined in the Finnish Forest Act, and the selection has been done by forest practitioners. Our objective was to determine whether the surroundings of boreal brooks and rivulets qualified as key habitats are truly different from brook-side habitats not granted the key habitat status, and whether the brook-side habitats of the two types differ from the forest matrix managed for timber production. We found that the two brook-side habitats were in most aspects rather alike but there was a difference in the composition of ground vegetation assemblages. In contrast, the control forests were distinct from the brook-sides in terms of dead wood, species richness and assemblages of polypores, species richness of epiphytic mosses, and the composition of beetle assemblages. We conclude that brook-sides in general provide an important habitat clearly diverging from the surrounding matrix but that the conservation value of the brook-sides granted the key habitat status may not be substantially larger than that of the brook-sides without the status.
  • Selonen, University of Jyväskylä, Department of Biological and Environmental Sciences, Lahti, Finland E-mail: ville.selonen@juy.fi (email)
  • Mussaari, University of Jyväskylä, Department of Biological and Environmental Sciences, Lahti, Finland E-mail: mm@nn.fi
  • Toivanen, University of Jyväskylä, Department of Biological and Environmental Sciences, Lahti, Finland E-mail: tt@nn.fi
  • Kotiaho, University of Jyväskylä, Department of Biological and Environmental Sciences, Lahti, Finland E-mail: jsk@nn.fi
article id 84, category Research article
Asko Lõhmus, Piret Lõhmus. (2011). Old-forest species: the importance of specific substrata vs. stand continuity in the case of calicioid fungi. Silva Fennica vol. 45 no. 5 article id 84. https://doi.org/10.14214/sf.84
Keywords: forest management; conservation; continuity; habitat; lichen; limiting factor; structural diversity
Abstract | View details | Full text in PDF | Author Info
Appropriate conservation management of old-forest species depends on the causes of their old-forest affinity, which, however, are insufficiently known. Calicioid fungi are often considered old-forest dependent because of their special requirements for microhabitat, microclimate, and stand continuity for at least two tree generations. We demonstrate that, for several methodological or interpretational problems, published studies do not provide unequivocal evidence for such mechanisms and even for old-forest dependency of calicioids in general. We then analyse a large Estonian dataset (ca. 2300 records of 32 species) representing various management types and site types to answer whether old forests have more calicioid species, and any specific species, than could be expected for the substratum availability observed. Although old growth had more species and records than mature managed stands or cutover sites, those substratum types that occurred at roughly similar abundances also hosted comparable numbers of species in different management types. The characteristic substrata adding extra species to old growth were snags and root-plates of treefall mounds; wood surfaces in general comprised more than half of all calicioid records. Although substratum abundance did not fully explain the species-richness contrast between old growth and mature stands, additional evidence suggested that the unexplained variance is rather due to small-scale habitat characteristics than stand-scale continuity or microclimate. Finally, we review the evidence for old-forest affinity of calicioid species and distinguish a set of threatened species. We conclude that the availability of specific substrata is the main limiting factor for calicioid fungi in forests, and its quantitative and stochastic nature explains the large random and region-specific variation in the published lists of ‘old-forest species’.
  • Lõhmus, Department of Zoology, Institute of Ecology and Earth Sciences, University of Tartu, Vanemuise st. 46, EE-51014, Tartu, Estonia E-mail: asko.lohmus@ut.ee (email)
  • Lõhmus, Department of Botany, Institute of Ecology and Earth Sciences, University of Tartu, Tartu, Estonia E-mail: pl@nn.ee
article id 83, category Research article
Inari Ylläsjärvi, Håkan Berglund, Timo Kuuluvainen. (2011). Relationships between wood-inhabiting fungal species richness and habitat variables in old-growth forest stands in the Pallas-Yllästunturi National Park, northern boreal Finland. Silva Fennica vol. 45 no. 5 article id 83. https://doi.org/10.14214/sf.83
Keywords: species-area relationship; species richness; old-growth forest; indicators; northern Fennoscandia; polyporaceae; polypores
Abstract | View details | Full text in PDF | Author Info
Indicators for biodiversity are needed for efficient prioritization of forests selected for conservation. We analyzed the relationships between 86 wood-inhabiting fungal (polypore) species richness and 35 habitat variables in 81 northern boreal old-growth forest stands in Finland. Species richness and the number of red-listed species were analyzed separately using generalized linear models. Most species were infrequent in the studied landscape and no species was encountered in all stands. The species richness increased with 1) the volume of coarse woody debris (CWD), 2) the mean DBH of CWD and 3) the basal area of living trees. The number of red-listed species increased along the same gradients, but the effect of basal area was not significant. Polypore species richness was significantly lower on western slopes than on flat topography. On average, species richness was higher on northern and eastern slopes than on western and southern slopes. The results suggest that a combination of habitat variables used as indicators may be useful in selecting forest stands to be set aside for polypore species conservation.
  • Ylläsjärvi, Rovaniemi University of Applied Sciences, School of Forestry and Rural Industries, Jokiväylä 11 c, FI-96300 Rovaniemi, Finland E-mail: inari.yllasjarvi@ramk.fi (email)
  • Berglund, University of Helsinki, Department of Forest Sciences, Helsinki, Finland E-mail: hb@nn.fi
  • Kuuluvainen, University of Helsinki, Department of Forest Sciences, Helsinki, Finland E-mail: tk@nn.fi
article id 82, category Research article
Nicole J. Fenton, Yves Bergeron. (2011). Dynamic old-growth forests? A case study of boreal black spruce forest bryophytes. Silva Fennica vol. 45 no. 5 article id 82. https://doi.org/10.14214/sf.82
Keywords: disturbances; old-growth forest; succession; boreal; bryophytes; climax; gap dynamics
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Old-growth forests have sparked significant interest over the last twenty years and definitions have evolved from structure based to process based, acknowledging the diversity of forests that could be considered old growth. However studies frequently group all forests over a certain age into a single type, negating the dynamic processes that create old growth. In this study we examine a 2350-year chronosequence in boreal black spruce forests in northwestern Quebec to determine whether continued community change can be observed in the bryophyte layer. Bryophytes dominate the understory of boreal forests and influence ecosystem functioning, particularly in paludified forests where production exceeds decomposition in the organic layer. Community composition and richness changed throughout the chronosequence with no evidence of a steady state associated with an old-growth phase. In contrast the bryophyte community continued to evolve with multiple phases being evident. These results suggest that old-growth forests on the Clay Belt of northwestern Quebec and northeastern Ontario, Canada, should be regarded as part of the continuous gradient in forest development rather than a single state. This complicates conservation of these forests as multiple phases should be considered when planning forest reserves.
  • Fenton, Université du Québec en Abitibi-Témiscamingue, 445 Boulevard de l’Université, Rouyn-Noranda, Québec, Canada J9X 4E5 E-mail: nicole.fenton@uqat.ca (email)
  • Bergeron, Université du Québec en Abitibi-Témiscamingue, 445 Boulevard de l’Université, Rouyn-Noranda, Québec, Canada J9X 4E5 E-mail: yb@nn.ca
article id 81, category Research article
Tuomas Aakala. (2011). Temporal variability of deadwood volume and quality in boreal old-growth forests. Silva Fennica vol. 45 no. 5 article id 81. https://doi.org/10.14214/sf.81
Keywords: forest dynamics; Picea abies; coarse woody debris; range of variability
Abstract | View details | Full text in PDF | Author Info
Reference deadwood volumes from natural forests for forest management and restoration are often derived from one-time measurements or from repeated measurements over short time-scales. Such an approach often assumes an equilibrium state between tree mortality and decomposition, which is questionable in many boreal forest ecosystems due to the occurrence of allogenic disturbances. Using a simulation model based on empirical estimates of tree mortality, disturbance chronologies and models of wood decay class dynamics, this study aimed at characterizing variability in the volume and quality of deadwood for the past 200 years. The variability of deadwood volumes in old-growth forests, arising from differences in disturbance regimes and differing decay rates, was exemplified in two areas of Picea abies-dominated forests in northern Europe. The results imply that with stable deadwood input and slow decay rates the deadwood volume may be in an equilibrium state. On the contrary, if moderate-severity disturbances occur such a state seems improbable. Both study areas displayed continuity in deadwood availability, although temporary paucity in the early decay classes with shortest residence times was also observed. The results stress the importance of understanding the dynamic nature of deadwood in old-growth forests, instead of the traditional view of deadwood as a static ecosystem component.
  • Aakala, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: tuomas.aakala@helsinki.fi (email)
article id 80, category Research article
Mari T. Jönsson, Shawn Fraver, Bengt Gunnar Jonsson. (2011). Spatio-temporal variation of coarse woody debris input in woodland key habitats in central Sweden. Silva Fennica vol. 45 no. 5 article id 80. https://doi.org/10.14214/sf.80
Keywords: boreal forest; Picea abies; dendrochronology; disturbance dynamics; tree mortality; woodland key habitats; windstorms
Abstract | View details | Full text in PDF | Author Info
The persistence of many saproxylic (wood-living) species depends on a readily available supply of coarse woody debris (CWD). Most studies of CWD inputs address stand-level patterns, despite the fact that many saproxylic species depend on landscape-level supplies of CWD. In the present study we used dated CWD inputs (tree mortality events) at each of 14 Norway spruce (Picea abies) dominated woodland key habitat sites to analyze the spatial and temporal patterns of CWD additions between 1950 and 2002 within a small landscape in central Sweden. We found that inputs were episodic within sites, where local windstorms created pulses in CWD input. Pulses occurred simultaneously in many sites, yielding landscape-level synchrony of CWD input. These synchronous pulses, and importantly, the breaks between pulses, may have negative implications for saproxylic species that are dependent on large volume inputs of freshly killed Norway spruce. In addition, the inherent small size and relative isolation of these sites may further increase extinction risks due to stochastic events. However, background CWD input rates occurring between pulses varied substantially among sites, presumably the result of the sites’ varied histories and structural characteristics. This finding suggests that the different sites have varied abilities to provide habitat for saproxylic species during periods with low landscape-level input of CWD.
  • Jönsson, Department of Ecology, SLU, P.O. Box 7044, SE-750 07 Uppsala, Sweden (current); Department of Natural Sciences, Engineering and Mathematics, Mid Sweden University, Sundsvall, Sweden E-mail: mari.jonsson@slu.se (email)
  • Fraver, U.S. Forest Service, Northern Research Station, Grand Rapids, Minnesota, USA (current); Department of Natural Sciences, Engineering and Mathematics, Mid Sweden University, Sundsvall, Sweden E-mail: sf@nn.us
  • Jonsson, Department of Natural Sciences, Engineering and Mathematics, Mid Sweden University, Sundsvall, Sweden E-mail: bgj@nn.se
article id 79, category Research article
Jean-Philippe Légaré, Christian Hébert, Jean-Claude Ruel. (2011). Alternative silvicultural practices in irregular boreal forests: response of beetle assemblages. Silva Fennica vol. 45 no. 5 article id 79. https://doi.org/10.14214/sf.79
Keywords: biodiversity; old-growth forest; irregular stands; selection cutting; coarse woody debris; sustainable management
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In the process of implementing sustainable management in the eastern Canadian boreal forest, we tested two selection cutting methods and compared them with two widely used practices in the boreal forest: clearcutting with protection of the advanced growth and soils and irregular shelterwood cutting leaving small merchantable stems. We used old-growth irregular stands as references in comparing the impact of these silvicultural treatments on the diversity and abundance of beetles. Three groups were targeted: saproxylic flying beetles, epigaeic saproxylic beetles and epigaeic non-saproxylic beetles. A sampling design including 320 pitfall traps and 80 multidirectional flight-interception traps was deployed in 2007. A total of 26 906 beetles was captured including 407 taxa distributed among 52 families. We found that clearcutting with protection of the advanced growth and soils and irregular shelterwood cutting leaving small merchantable stems had a greater impact on beetle communities than both selection cuttings. Canopy opening as well as the presence of snags and downed woody debris appear as important attributes for several saproxylic and non-saproxylic species. Beetle communities in selection cuttings remained more similar to those found in controls; these silvicultural treatments are new tools to implement ecosystemic and sustainable management in irregular boreal forests.
  • Légaré, Université Laval, Faculté de foresterie, de géographie et de géomatique, Pavillon Abitibi-Price, Québec, Canada E-mail: jpl@nn.ca
  • Hébert, Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, 1055 du P.E.P.S., P.O. Box 10380, Stn. Sainte-Foy, Québec (Québec), G1V 4C7, Canada E-mail: christian.hebert@rncan.gc.ca (email)
  • Ruel, Université Laval, Faculté de foresterie, de géographie et de géomatique, Pavillon Abitibi-Price, Québec, Canada E-mail: jcr@nn.ca
article id 78, category Research article
Kris Vandekerkhove, Luc De Keersmaeker, Ruben Walleyn, Frank Köhler, Luc Crevecoeur, Leen Govaere, Arno Thomaes, Kris Verheyen. (2011). Reappearance of old-growth elements in lowland woodlands in northern Belgium: Do the associated species follow? Silva Fennica vol. 45 no. 5 article id 78. https://doi.org/10.14214/sf.78
Keywords: colonisation; saproxylic species; dead wood; very large trees
Abstract | View details | Full text in PDF | Author Info
The forest cover of the western European lowland plain has been very low for centuries. Remaining forests were intensively managed, and old-growth elements like veteran trees and coarse woody debris became virtually absent. Only over the last decades have these old-growth elements progressively redeveloped in parks, lanes and forests, and have now reached their highest level over the last 500–1000 years. Biodiversity associated with these old-growth elements makes up an important part of overall forest biodiversity. The ability of species to recolonise the newly available habitat is strongly determined by limitations in their dispersal and establishment. We analyse the current status and development of old-growth elements in Flanders (northern Belgium) and the process of recolonisation by means of specific cases, focussing on saproxylic fungi and saproxylic beetles. Our results show that ‘hotspots’ of secondary old growth, even isolated small patches, may have more potential for specialised biodiversity than expected, and may provide important new strongholds for recovery and recolonisation of an important share of old-growth related species.
  • Vandekerkhove, INBO, Gaverstraat 4, B-9500 Geraardsbergen, Belgium E-mail: kris.vandekerkhove@inbo.be (email)
  • De Keersmaeker, INBO, Gaverstraat 4, B-9500 Geraardsbergen, Belgium E-mail: ldk@nn.be
  • Walleyn, INBO, Gaverstraat 4, B-9500 Geraardsbergen, Belgium E-mail: rw@nn.be
  • Köhler, Koleopterologisches Forschungsbüro, Bornheim, Germany E-mail: fk@nn.de
  • Crevecoeur, Genk, Belgium E-mail: lc@nn.be
  • Govaere, Agency of Nature and Forests, Brussels, Belgium E-mail: lg@nn.be
  • Thomaes, INBO, Gaverstraat 4, B-9500 Geraardsbergen, Belgium E-mail: at@nn.be
  • Verheyen, Ghent University, Laboratory of Forestry, Gontrode, Belgium E-mail: kv@nn.be
article id 77, category Research article
Annie Claude Bélisle, Sylvie Gauthier, Dominic Cyr, Yves Bergeron, Hubert Morin. (2011). Fire regime and old-growth boreal forests in central Quebec, Canada: an ecosystem management perspective. Silva Fennica vol. 45 no. 5 article id 77. https://doi.org/10.14214/sf.77
Keywords: black spruce-feather moss; fire history; ecosystem management; dendrochronology
Abstract | View details | Full text in PDF | Author Info
Boreal forest management in Eastern Canada has caused depletion and fragmentation of old-growth ecosystems, with growing impacts on the associated biodiversity. To mitigate impacts of management while maintaining timber supplies, ecosystem management aims to narrow the gap between natural and managed landscapes. Our study describes the fire history and associated natural old-growth forest proportions and distribution of a 5000 km2 area located in the black spruce-feather moss forest of central Quebec. We reconstructed a stand-origin map using archival data, aerial photos and dendrochronology. According to survival analysis (Cox hazard model), the mean fire cycle length was 247 years for the 1734–2009 period. Age-class distribution modelling showed that old-growth forests were present on an average of 55% of the landscape over the last 275 years. The mean fire size was 10 113 ha, while most of the burned area was attributable to fires larger than 10 000 ha, leading to old-growth agglomerations of hundreds of square kilometres. In regards to our findings, we propose ecosystem management targets and strategies to preserve forest diversity and resilience.
  • Bélisle, Centre for Forest Research, Université du Québec à Montréal, Montréal, Québec, Canada E-mail: annieclaude_b@hotmail.com (email)
  • Gauthier, Laurentian Forestry Centre, Canadian Forest Service, Natural Resources Canada, Sainte-Foy, Québec, Canada E-mail: sg@nn.ca
  • Cyr, Institut Québécois d’Aménagement de la Fort Feuillue, Université du Québec en Outaouais, Ripon, Québec, Canada E-mail: dc@nn.ca
  • Bergeron, Centre for Forest Research, Université du Québec à Montréal, Montréal, Québec, Canada & NSERC-UQAT-UQAM Industrial Chair in Sustainable Forest Management, Université du Québec en Abitibi-Témiscamingue, Rouyn-Noranda, Québec, Canada E-mail: yb@nn.ca
  • Morin, Département des Sciences Fondamentales, Université du Québec à Chicoutimi, Chicoutimi, Québec, Canada E-mail: hm@nn.can
article id 76, category Research article
Alessandra Bottero, Matteo Garbarino, Vojislav Dukic, Zoran Govedar, Emanuele Lingua, Thomas A. Nagel, Renzo Motta. (2011). Gap-phase dynamics in the old-growth forest of Lom, Bosnia and Herzegovina. Silva Fennica vol. 45 no. 5 article id 76. https://doi.org/10.14214/sf.76
Keywords: disturbance regime; Lom; old-growth forest; canopy gaps; beech-spruce-fir; small-scale
Abstract | View details | Full text in PDF | Author Info
We investigated forest canopy gaps in the mixed beech (Fagus sylvatica L.), silver fir (Abies alba Miller), and Norway spruce (Picea abies (L.) Karst.) old-growth forest of Lom in the Dinaric Mountains of Bosnia and Herzegovina. Gap size, age, gap fraction, gapmaker characteristics and the structure and composition of gapfillers were documented to investigate gap dynamics. The percentages of forest area in canopy and expanded gaps were 19% and 41%, respectively. The median canopy gap size was 77 m2, and ranged from 11 to 708 m2. Although there were many single tree-fall gaps, the majority had multiple gapmakers that were often in different stages of decay, suggesting gap expansion is important at the study site. Of the gapmakers recorded, 14% were uprooted stems, 60% snapped stems, and 26% were standing dead trees. Dendroecological analysis suggests that gap formation varied in time. The density of gapfillers was not correlated to gap size, and the species composition of gapfillers varied between seedling, sapling, and tree life stages. The results suggest that gaps are mainly formed by endogenous senescence of single canopy trees. Exogenous disturbance agents, most likely related to wind and snow, act mainly as secondary agents in breaking weakened trees and in expanding previously established gaps. Although the findings are partially consistent with other studies of gap disturbance processes in similar old-growth forests in central Europe, the observed gap dynamic places the Lom core area at the end of a gradient that ranges from forests controlled by very small-scale processes to those where large, stand replacing disturbances predominate.
  • Bottero, University of Turin, Department Agroselviter, Via Leonardo da Vinci 44, 10095 Grugliasco (TO), Italy E-mail: alessandra.bottero@unito.it (email)
  • Garbarino, University of Turin, Department Agroselviter, Via Leonardo da Vinci 44, 10095 Grugliasco (TO), Italy E-mail: mg@nn.it
  • Dukic, University of Banja Luka, Faculty of Forestry, Banja Luka, Bosnia and Herzegovina E-mail: vd@nn.ba
  • Govedar, University of Banja Luka, Faculty of Forestry, Banja Luka, Bosnia and Herzegovina E-mail: zg@nn.ba
  • Lingua, University of Padua, Department of TeSAF, Legnaro (PD), Italy E-mail: el@nn.it
  • Nagel, University of Ljubljana, Department of Forestry and Renewable Forest Resources, Ljubljana, Slovenia E-mail: tan@nn.si
  • Motta, University of Turin, Department Agroselviter, Via Leonardo da Vinci 44, 10095 Grugliasco (TO), Italy E-mail: rm@nn.it
article id 75, category Research article
Silvia Lamedica, Emanuele Lingua, Ionel Popa, Renzo Motta, Marco Carrer. (2011). Spatial structure in four Norway spruce stands with different management history in the Alps and Carpathians. Silva Fennica vol. 45 no. 5 article id 75. https://doi.org/10.14214/sf.75
Keywords: Picea abies; LISA; old-growth forests; point pattern analyses; random mortality; spatial structure
Abstract | View details | Full text in PDF | Author Info
In Europe most Norway spruce (Picea abies (L.) Karst.) mountain forests have been altered by human activities, leading to a lack of reference condition concerning their original structure. Nonetheless, remnants of Norway spruce primeval forests still exist in the Carpathians. Our objective was to assess the differences in structure between managed and unmanaged stands, concerning diameter distributions, amount of standing deadwood, spatial distributions and spatial structure of trees. We established four permanent plots: one in a virgin forest in the Eastern Carpathians and three in a previously managed forest in the Alps. In each plot, species, DBH, and position of the live and dead standing trees were collected. Spatial distribution and structure of all the trees was analysed through several indices. In the Carpathians forest there are clear signs of natural density-dependent mortality processes whereas in the Alpine plots such dynamics are less evident. In these latter plots, the lower snags volume and the random trees spatial distribution can be considered the legacies of past management. Nonetheless, despite the different history of the four stands, they all seem to converge towards a similar spatial structure with the presence of groups (30–40 m) of trees of similar size.
  • Lamedica, Forest Ecology Research Unit, Dept TeSAF, University of Padova, Legnaro (PD), Italy E-mail: sl@nn.it
  • Lingua, Forest Ecology Research Unit, Dept TeSAF, University of Padova, Legnaro (PD), Italy E-mail: el@nn.it
  • Popa, Forest Research and Management Institute, Research Station for Norway Spruce Silviculture, Câmpulung Moldovenesc, Romania E-mail: ip@nn.ro
  • Motta, Department AGROSELVITER, University of Torino, Grugliasco (TO), Italy E-mail: rm@nn.it
  • Carrer, Forest Ecology Research Unit, Dept TeSAF, University of Padova, Legnaro (PD), Italy E-mail: marco.carrer@unipd.it (email)
article id 447, category Research article
Jim Kiser. (2011). Histochemical and geometric alterations of sapwood in coastal Douglas-fir following mechanical damage during commercial thinning. Silva Fennica vol. 45 no. 4 article id 447. https://doi.org/10.14214/sf.447
Keywords: thinning; sapwood; compartmentalization; damages; Pseudotsuga menziesii
Abstract | View details | Full text in PDF | Author Info
Histochemical and geometric alterations to sapwood in mechanically damaged Douglas-fir (Pseudotsuga menziesii [Mirbel] Franco) trees were quantified 14 years after thinning. Discoloration and decay were measured in felled damaged and undamaged trees. Compartmentalized walls were identified and measured macroscopically. Sapwood to heartwood ratio was measured incrementally along the boles. Results showed a distinct reaction zone forming at the time of injury. Compartmentalized walls 1–3 were less distinct and heavily resinous streaking was evident in extant tissues, particularly in the axial direction. Post-damaged sapwood was burl-like for 4–6 years and tracheids contained resin-filled lumina. Damaged wood volumes were modeled by multiple regression. Wound depth, wound area, and diameter inside bark (DIB) accounted for 73% of the discolored volume (p = 0.02). DIB alone accounted for just over 55% of the response. Post-damaged sapwood averaged 15 mm (SE = 2.3 mm) greater in width on the side opposite the damage along the length of the boards. Wound area explained just over 65% of this response (p = 0.003). Sapwood area was not significantly different between damaged and control trees (p = 0.56). Results indicate that wounded Douglas-fir trees may slow conversion of sapwood to heartwood on the bole side opposite the wound, possibly as a response to maintain sapwood area necessary for physiological maintenance of the existing crown. About 19% of the lower bole volume in damaged trees was affected by discoloration and secondarily by structural changes. Reduction in value of the lower log can be as high as 19% by conventional bucking practices. Alternatives are presented to reduce the value loss to between 2.5% to 3.5%.
  • Kiser, P.O. Box 3729, Pagosa Springs, Colorado, USA E-mail: jim.kiser@parelli.com (email)
article id 103, category Research article
Luis Diaz-Balteiro, Roberto Voces, Carlos Romero. (2011). Making sustainability rankings using compromise programming. An application to European paper industry. Silva Fennica vol. 45 no. 4 article id 103. https://doi.org/10.14214/sf.103
Keywords: indicators; compromise programming; sustainability; paper industry
Abstract | View details | Full text in PDF | Author Info
This paper characterizes the sustainability of the European paper industry. To undertake this task the sustainability of each country is defined by using fourteen indicators of a diverse nature (economic, environmental and social). These indicators are aggregated into a composite or synthetic index with the help of a compromise programming model. In order to associate different weights with each indicator, a survey among international experts has been carried out. In this way, a ranking of seventeen European countries analysed in terms of the sustainability of the European paper industry has been established, where Finland is the most sustainable paper industry in Europe except when the most balanced solution is chosen. Also, the results are robust when different preferential weights are attached. Finally, this methodology can be applied at a more disaggregated level and other indicators can be introduced.
  • Diaz-Balteiro, Research Group “Economics for a Sustainable Environment”, Technical University of Madrid, Ciudad Universitaria s/n, 28040 Madrid, Spain E-mail: luis.diaz.balteiro@upm.es (email)
  • Voces, Research Group “Economics for a Sustainable Environment”, Technical University of Madrid, Ciudad Universitaria s/n, 28040 Madrid, Spain E-mail: rv@nn.es
  • Romero, Research Group “Economics for a Sustainable Environment”, Technical University of Madrid, Ciudad Universitaria s/n, 28040 Madrid, Spain E-mail: cr@nn.es
article id 102, category Research article
Anne Toppinen, Katja Lähtinen, Leena A. Leskinen, Niklas Österman. (2011). Network co-operation as a source of competitiveness in medium-sized Finnish sawmills. Silva Fennica vol. 45 no. 4 article id 102. https://doi.org/10.14214/sf.102
Keywords: collaboration; social capital; strategic resources; competitive advantage; business processes
Abstract | View details | Full text in PDF | Author Info
In the Finnish sawmill industry, inter-firm collaboration has often been brought up as a means of creating a competitive edge in global markets by achieving economies of scale. According to the resource-based view (RBV), a firm can evaluate its current or potential partners by considering firm-level collaboration as a portfolio of complementary strategic resources. The specific focus of the study is on examining the types and forms of sawmill co-operation, how the co-operation emerged and which firm-specific resources are mainly related to co-operation. Based upon this, we can see how the managers of medium-sized sawmills perceive network co-operation as facilitating the achievement of a sustainable competitive advantage. The empirical data for this study were collected by interviewing 16 managers and employees in medium-sized non-integrated sawmills, a joint-venture marketing company and other co-operative partners. The findings of the study show that meaningful and beneficial co-operation partnerships exist in the Finnish sawmilling industry, but the sawmill managers do not perceive this collaboration as a strategic resource. The marketing company was the only firm in this study that relied on its co-operative networks in seeking a sustainable competitive advantage. To make more of co-operative partnerships, the principles of co-operative networking should be understood better in the sawmilling industry in order to know what to expect from co-operation. Furthermore, the managers should have the courage to engage in more extensive co-operation in order for strategic rents to materialize. Since the selection of the right partners is fundamental, further studies could be conducted on the reasons behind failed or terminated co-operative arrangements to gather further empirical knowledge in this subject area.
  • Toppinen, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: anne.toppinen@helsinki.fi (email)
  • Lähtinen, Finnish Environment Institute (SYKE), Joensuu, Finland E-mail: kl@nn.fi
  • Leskinen, Rantalankuja 4, Joensuu, Finland E-mail: lal@nn.fi
  • Österman, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: no@nn.fi
article id 101, category Research article
Risto Laamanen, Annika Kangas. (2011). Large-scale forest owner’s information needs in operational planning of timber harvesting - some practical views in Metsähallitus, Finnish state-owned enterprise. Silva Fennica vol. 45 no. 4 article id 101. https://doi.org/10.14214/sf.101
Keywords: forest inventories; harvest planning; information needs
Abstract | View details | Full text in PDF | Author Info
  • Laamanen, Metsähallitus, Vantaa, Finland E-mail: risto.laamanen@metsa.fi (email)
  • Kangas, Metsähallitus, Vantaa, Finland E-mail: ak@nn.fi
article id 100, category Research article
Annika Kangas, Lauri Mehtätalo, Antti Mäkinen, Kalle Vanhatalo. (2011). Sensitivity of harvest decisions to errors in stand characteristics. Silva Fennica vol. 45 no. 4 article id 100. https://doi.org/10.14214/sf.100
Keywords: forest planning; inventory; measurement errors; decision making; logistic regression; regression tree
Abstract | View details | Full text in PDF | Author Info
In forest planning, the decision maker chooses for each stand a treatment schedule for a predefined planning period. The choice is based either on optimization calculations or on silvicultural guidelines. Schedules for individual stands are obtained using a growth simulator, where measured stand characteristics such as the basal area, mean diameter, site class and mean height are used as input variables. These characteristics include errors, however, which may lead to incorrect decisions. In this study, the aim is to study the sensitivity of harvest decisions to errors in a dataset of 157 stands. Correct schedules according to silvicultural guidelines were first determined using error-free data. Different amounts of errors were then generated to the stand-specific characteristics, and the treatment schedule was selected again using the erroneous data. The decision was defined as correct, if the type of harvest in these two schedules were similar, and if the timings deviated at maximum ±2 for thinning and ±3 years for clear-cut. The dependency of probability of correct decisions on stand characteristics and the degree of errors was then modelled. The proposed model can be used to determine the required level of measurement accuracy for each characteristics in different kinds of stands, with a given accuracy requirement for the timing of treatments. This information can further be utilized in selecting the most appropriate inventory method.
  • Kangas, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: annika.kangas@helsinki.fi (email)
  • Mehtätalo, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: lm@nn.fi
  • Mäkinen, Simosol Oy, Riihimäki, Finland E-mail: am@nn.fi
  • Vanhatalo, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: kv@nn.fi
article id 99, category Research article
Jouni Siipilehto. (2011). Local prediction of stand structure using linear prediction theory in Scots pine-dominated stands in Finland. Silva Fennica vol. 45 no. 4 article id 99. https://doi.org/10.14214/sf.99
Keywords: Pinus sylvestris; linear prediction; diameter distribution; Weibull; Johnson’s SB; height curve; stand characteristics
Abstract | View details | Full text in PDF | Author Info
This study produced a family of models for eight standard stand characteristics, frequency and basal area-based diameter distributions, and a height curve for stands in Finland dominated by Scots pine (Pinus sylvestris L.). The data consisted of 752 National Forest Inventory-based sample plots, measured three times between 1976 and 2001. Of the data, 75% were randomly selected for modelling and 25% left out for model evaluation. Base prediction models were constructed as functions of stand age, location and site providing strongly average expectations. These expectations were then calibrated with the known stand variables using linear prediction theory when estimating the best linear unbiased predictor (BLUP). Three stand variables, typically assessed in Finnish forest management planning fieldwork, were quite effective for calibrating the expectation for the unknown variable. In the case of optional distributions, it was essential to choose the weighting of the diameter distribution model such that the available input variables and the model applied were based on the same scale (e.g. arithmetic stand variables for frequency distribution). Additional input variables generally improved the accuracy of the validated characteristics, but the improvements in the predicted distributions were most noteworthy when the arithmetic mean and basal area-weighted median were simultaneously included in the BLUP estimation. The BLUP method provided a flexible approach for characterising relationships among stand variables, alternative size distributions and the height–diameter curve. Models are intended for practical use in the MOTTI simulator.
  • Siipilehto, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: jouni.siipilehto@metla.fi (email)
article id 98, category Research article
Meeri Pearson, Markku Saarinen, Kari Minkkinen, Niko Silvan, Jukka Laine. (2011). Mounding and scalping prior to reforestation of hydrologically sensitive deep-peated sites: factors behind Scots pine regeneration success. Silva Fennica vol. 45 no. 4 article id 98. https://doi.org/10.14214/sf.98
Keywords: soil preparation; peatland forest regeneration; mound; scalp; Scots pine; OM decomposition; excess moisture
Abstract | View details | Full text in PDF | Author Info
Watering up typically ensues after clearcutting forestry-drained peatland forests. Thus, the effectiveness of maintenance drainage and soil preparation procedures becomes paramount for establishing a new generation of commercial forest. Mounding is the primary method of soil preparation applied in regeneration sites lying on deep peat. As raised planting spots, mounds are resistant to waterlogging and assumed to be beneficial for organic matter (OM) decomposition via, e.g., increased soil aeration and temperature, which would also enhance seedling growth. In recent years, however, less intensive and cheaper alternatives like scalping have been sought with some reported cases of success. Our case study investigated the survival and growth of Scots pine outplants in mounds, scalps, and unprepared microsites along a moisture gradient. After three growing seasons, mounding accelerated neither seedling growth nor OM decomposition relative to the unprepared treatment. Survival in mounds was nonetheless superior overall. Scalps behaved as water collecting depressions leading to a catastrophic regeneration result. Based on our findings, water table level (WTL) overrides other growth-controlling factors in excess moisture conditions. To combat watering up coupled with greater than normal rainfall, we recommend reforestation strategies which provide elevated, prepared planting spots (i.e., mounds) or utilize unprepared, higher microforms.
  • Pearson, Finnish Forest Research Institute, Western Finland Regional Unit, Kaironiementie 15, FI-39700 Parkano, Finland E-mail: meeri.pearson@metla.fi (email)
  • Saarinen, Finnish Forest Research Institute, Western Finland Regional Unit, Kaironiementie 15, FI-39700 Parkano, Finland E-mail: ms@nn.fi
  • Minkkinen, University of Helsinki, Department of Forest Sciences, Helsinki, Finland E-mail: km@nn.fi
  • Silvan, Finnish Forest Research Institute, Western Finland Regional Unit, Kaironiementie 15, FI-39700 Parkano, Finland E-mail: ns@nn.fi
  • Laine, Finnish Forest Research Institute, Western Finland Regional Unit, Kaironiementie 15, FI-39700 Parkano, Finland E-mail: jl@nn.fi
article id 97, category Research article
Hannu Hökkä, Jaakko Repola, Mikko Moilanen, Markku Saarinen. (2011). Seedling survival and establishment in small canopy openings in drained spruce mires in Northern Finland. Silva Fennica vol. 45 no. 4 article id 97. https://doi.org/10.14214/sf.97
Keywords: Picea abies; canopy gaps; natural regeneration; drained peatlands; seedling establishment
Abstract | View details | Full text in PDF | Author Info
A large proportion of drained spruce mire stands is currently approaching regeneration maturity in Finland. Traditional regeneration methods with effective site preparation and planting generally result in satisfactory seedling stands also in spruce mires. However, natural regeneration methods may be more appropriate in protecting watercourses and minimizing regeneration costs. We studied the survival of advance growth and establishment of new seedlings in small canopy openings that were cut at three different diameters in two experimental drained spruce mire stands in Northern Finland (Tervola and Oulu) in 2004. The number of seedlings was repeatedly surveyed from five small circular plots (one 10 m2 and four 5 m2 plots in size) located within the opening. Advance growth which survived the cutting and new seedlings were separated in the surveys. The density of advance growth was on average 9000 ha–1 after cutting, and it decreased by 30% during the five-year monitoring period (2006–2010) due to natural mortality. The number of new seedlings increased rapidly within the three years after cutting the openings. In 2010, 11 000–26 000 new seedlings ha–1 in Tervola and 12 000–16 000 ha–1 in Oulu on average were observed. The size of the opening had no clear effect on the regeneration result. The proportion of birch of the new seedlings increased with time and opening size in Tervola. The results show that Norway spruce regenerates naturally in small canopy openings cut in mature drained spruce mire stands.
  • Hökkä, Finnish Forest Research Institute, Rovaniemi Research Unit, FI-96301 Rovaniemi, Finland E-mail: hannu.hokka@metla.fi (email)
  • Repola, Finnish Forest Research Institute, Rovaniemi Research Unit, FI-96301 Rovaniemi, Finland E-mail: jr@nn.fi
  • Moilanen, Finnish Forest Research Institute, Muhos Research Unit, Muhos, Finland E-mail: mm@nn.fi
  • Saarinen, Finnish Forest Research Institute, Parkano Research Unit, Parkano, Finland E-mail: ms@nn.fi
article id 96, category Research article
Scott R. Abella. (2011). How well do U.S. Forest Service terrestrial ecosystem surveys correspond with measured vegetation properties? Silva Fennica vol. 45 no. 4 article id 96. https://doi.org/10.14214/sf.96
Keywords: ecosystem classification; Pinus ponderosa; potential natural vegetation; reference conditions; understory
Abstract | View details | Full text in PDF | Author Info
Reliable estimates of species composition that forest sites are capable of supporting – specific to ecosystem mapping units across landscapes – are useful for many purposes in forest science and management. Like forestry agencies in numerous countries, the U.S. Forest Service has invested in ecological land classification (termed terrestrial ecosystem survey [TES] in the study region of Arizona) that includes ecosystem-explicit species lists taken to be estimated potential natural vegetation (PNV). Using multivariate community analyses, PNV in the TES was compared to measured species composition on 66 sites representing among the least-disturbed vegetation (considered this study’s measured PNV) spanning 11 ecosystem types on a Pinus ponderosa P. & C. Lawson landscape in northern Arizona, USA. Agreement between the TES PNV and measured species composition was lowest for forbs and shrubs (compared to graminoids), and species composition differed significantly between the TES and this study for at least one plant lifeform in 73% of ecosystems. Reasons for differences between the TES and this study are difficult to resolve, but in some cases appear to result from identification of different species pools in the region. This study suggests that the TES is a useful starting point in understanding vegetation-environment relationships, but further work is needed to refine species lists and more thoroughly account for the influences of fire, grazing, and climate that can influence both PNV and current vegetation. Refining and updating ecosystem-specific species lists may benefit existing forest site classifications and could be planned for when new site classifications are developed, especially with changing climates.
  • Abella, Department of Environmental and Occupational Health, University of Nevada Las Vegas, Las Vegas, Nevada 89154-3064 USA E-mail: scott.abella@unlv.edu (email)
article id 95, category Research article
Venceslas Goudiaby, Suzanne Brais, Yvon Grenier, Frank Berninger. (2011). Thinning effects on jack pine and black spruce photosynthesis in eastern boreal forests of Canada. Silva Fennica vol. 45 no. 4 article id 95. https://doi.org/10.14214/sf.95
Keywords: nitrogen; light; Picea mariana; Pinus banksiana; photosynthesis; thinning
Abstract | View details | Full text in PDF | Author Info
A decrease in the average diameter of commercially harvested tree species in the Eastern boreal forest of Canada has led to a decrease in availability of quality wood for the forest industry. Commercial thinning has been proposed as a means to increase stem diameter growth. However, little is known about physiological responses underlying species responses to thinning. We assessed the effect of canopy opening on the photosynthetic response of mature jack pine (Pinus banksiana Lamb.) and black spruce (Picea mariana (Mill.) BSP) trees. Two years after thinning and for each species, light response curves and the diurnal course of photosynthesis were characterized from measurements taken in a completely randomized block experiment on current-year and one-year-old needles of 12 trees from stands subjected to different levels of canopy opening. Soil water content, air and soil temperatures, and needle N concentration were not affected by thinning for either species. However, light availability increased with basal area removed and could explain the significantly positive relationship between thinning intensity and diurnal course of photosynthesis for one-year-old needles of jack pine. Black spruce photosynthesis did not respond to increases in light. Light-saturated rate of net photosynthesis (Amax), photosynthetic efficiency (α), light compensation point (LCP), and diurnal respiration (Rd) did not vary with thinning for either of the species. Jack pine and black spruce responses to thinning should be interpreted in light of species autecology.
  • Goudiaby, NSERC/UQAT/UQÀM Industrial Chair in Sustainable Forest Management, Université du Québec en Abitibi-Témiscamingue, Québec, Canada E-mail: venceslas.goudiaby@uqat.ca (email)
  • Brais, NSERC/UQAT/UQÀM Industrial Chair in Sustainable Forest Management, Université du Québec en Abitibi-Témiscamingue, Québec, Canada E-mail: sb@nn.ca
  • Grenier, NSERC/UQAT/UQÀM Industrial Chair in Sustainable Forest Management, Université du Québec en Abitibi-Témiscamingue, Québec, Canada E-mail: yg@nn.ca
  • Berninger, University of Helsinki, Faculty of Agriculture and Forestry, Department of Forest Sciences, Finland E-mail: fb@nn.fi
article id 94, category Research article
Youhong Peng, Ke Chen. (2011). Phylogeographic pattern of Populus cathayana in the southeast of Qinghai-Tibetan Plateau of China revealed by cpSSR markers. Silva Fennica vol. 45 no. 4 article id 94. https://doi.org/10.14214/sf.94
Keywords: genetic differentiation; refugia; phylogeography
Abstract | View details | Full text in PDF | Author Info
The vegetation in the Qinghai-Tibetan Plateau is thought to be highly sensitive and more vulnerable to global climate change than that of other areas. The uplift of the plateau as well as the climatic oscillations during glacial periods had a profound impact on plant species distribution and genetic diversity there. In the present study, seven pairs of cpSSR (chloroplast Simple Sequence Repeat) primers were utilized to detect genetic varieties of Populus cathayana Rehd populations from their natural range in the southeastern areas of Qinghai-Tibetan Plateau. A total of 28 alleles and 12 different haplotypes were detected. The proportion of haplotype variation among populations (GST = 0.794, NST = 0.900) indicated high level of genetic differentiation among populations and a significant phylogeographic structure (NST > GST, P < 0.05). This appears to support the hypothesis that these populations were derived from multiple refugia areas during the Quaternary climatic oscillations. Based on the haplotype network and mismatch distribution analyses, we found no evidence of postglacial range recolonization and expansion by P. cathayana in this region. This might be mainly due to the complex topography of the southeastern part of the Qinghai-Tibetan Plateau. The lofty mountain ranges and deep valleys in this region might have prevented long-distance migrations of this species during the climatic amelioration.
  • Peng, Chinese Academy of Sciences, Chengdu Institute of Biology, Chengdu, China E-mail: pengyh@cib.ac.cn (email)
  • Chen, Chinese Academy of Sciences, Chengdu Institute of Biology, Chengdu, China E-mail: kc@nn.cn
article id 93, category Research article
Pertti Pulkkinen, Saila Varis, Raimo Jaatinen, Aulis Leppänen, Anne Pakkanen. (2011). Increasing survival and growth of Scots pine seedlings with selection based on autumn coloration. Silva Fennica vol. 45 no. 4 article id 93. https://doi.org/10.14214/sf.93
Keywords: Pinus sylvestris; adaptation; field trials; harsh conditions; needles; nursery; selection
Abstract | View details | Full text in PDF | Author Info
This study evaluates the possibility of using autumn coloration of young Scots pine (Pinus sylvestris L.) seedlings as an indicator of adaptation to harsh climate conditions. One-year old seedlings from natural stands with different origins and seed orchards were classified as “red/reddish” and “green” based on the needle color after artificially increased night length in nursery and then measured after 14 years in field trials. In almost all the studied groups seedlings classified as “red/reddish” had significantly higher survival rate than seedlings classified as “green”. The survival of “red/reddish” was 14.2% higher than “green” among natural stand seed material and 56.2% among seed orchard material. During the study period the survival difference between “red/reddish” and “green” seedlings tended to increase. The seedling color had limited connection with the height growth, even though the trees classified as “red/reddish” were slightly taller than those classified as “green”. However, the total productivity over all field trials, described here as a heightsum, of “red/reddish” trees was 15% higher than productivity of “green” trees from natural stand material, and 61% higher than those from seed orchard material. It seems that controlled selection based on autumn color can be utilized within seed crops of different types with the aim to increase the adaptability of seed material to different environmental conditions.
  • Pulkkinen, Metla, Haapastensyrjä, Läyliäinen, Finland E-mail: pertti.pulkkinen@metla.fi (email)
  • Varis, Metla, Haapastensyrjä, Läyliäinen, Finland E-mail: sv@nn.fi
  • Jaatinen, Metla, Haapastensyrjä, Läyliäinen, Finland E-mail: rj@nn.fi
  • Leppänen, Metla, Haapastensyrjä, Läyliäinen, Finland E-mail: al@nn.fi
  • Pakkanen, Metla, Haapastensyrjä, Läyliäinen, Finland E-mail: ap@nn.fi
article id 450, category Research article
Miaoer Lu, Pekka Nygren, Jari Perttunen, Stephen G. Pallardy, David R. Larsen. (2011). Application of the functional-structural tree model LIGNUM to growth simulation of short-rotation eastern cottonwood. Silva Fennica vol. 45 no. 3 article id 450. https://doi.org/10.14214/sf.450
Keywords: Populus deltoides; carbon allocation; L-system; photon flux interception; photosynthetic production; voxel model
Abstract | View details | Full text in PDF | Author Info
The functional-structural tree growth model LIGNUM was developed as a general research tool that can be applied to several tree species. The growth simulation of short-rotation eastern cottonwood (Populus deltoides Bartr. ex Marsh.) inherits the basic LIGNUM modeling concepts including modular tree structure, L-system-based description of structural development, and carbon budget. New developments of LIGNUM model in this study were the incorporation of a biochemically-derived photosynthesis submodel; nested time steps for simulating physiological processes, structural development, and annual biomass production; incorporation of field-measured weather data for modeling the response of physiological processes to environmental variation; and application of a Monte-Carlo voxel space submodel for simulating the stochasticity of tree growth and improving computational efficiency. A specific parameter system was applied for modeling P. deltoides growth in the central Missouri, USA, environment. This adaptation of LIGNUM was applied on modeling growth of P. deltoides in a short-rotation agroforestry practice. The simulated height and biomass growth were close to field observations. Visualization of simulation results closely resembled the trees growing in an open site. The simulated response of tree growth to variations in photon flux input was reasonable. The LIGNUM model may be used as a complement to field studies on P. deltoides in short-rotation forestry and agroforestry.
  • Lu, Deparment of Forestry, University of Missouri, Columbia, MO, USA E-mail: ml@nn.us
  • Nygren, The Finnish Society of Forest Science, P.O. 18, FI-01301 Vantaa, Finland E-mail: pekka.nygren@metla.fi (email)
  • Perttunen, Finnish Forest Research Institute, Vantaa, Finland E-mail: jp@nn.fi
  • Pallardy, Deparment of Forestry, University of Missouri, Columbia, MO, USA E-mail: sgp@nn.us
  • Larsen, Deparment of Forestry, University of Missouri, Columbia, MO, USA E-mail: drl@nn.us
article id 449, category Research article
Thomas Wutzler, Ingolf Profft, Martina Mund. (2011). Quantifying tree biomass carbon stocks, their changes and uncertainties using routine stand taxation inventory data. Silva Fennica vol. 45 no. 3 article id 449. https://doi.org/10.14214/sf.449
Keywords: forest management; forest inventory; uncertainty; carbon stocks
Abstract | View details | Full text in PDF | Author Info
For carbon (C) trading or any other verifiable C reports, it would be reasonable to identify and quantify continuous changes in carbon stocks at regional scales without high investments into additional C-specific, time- and labor-intensive inventories. Our study demonstrates the potential of using routine stand taxation data from large scale forestry inventories for verifiable quantification of tree biomass C stocks, C stock change rates, and associated uncertainties. Empirical models, parameters, and equations of uncertainty propagation have been assembled and applied to data from a forest management unit in Central Germany (550 000 ha), using stand taxation inventories collected between 1993 and 2006. The study showed: 1) The use of stand taxation data resulted in a verifiable and sufficiently precise (cv = 7%) quantification of tree biomass carbon stocks and their changes at the level of growth-regions (1700 to 140 000 ha). 2) The forest of the test region accumulated carbon in tree biomass at a mean annual rate of 1.8 (–0.9 to 4.5) tC/ha/yr over the studied period. 3) The taxation inventory data can reveal spatial patterns of rates of C stock changes, specifically low rates of 0.4 tC/ha/yr in the northwest and high rates of 3.0 tC/ha/yr in the south of the study region.
  • Wutzler, Max Planck Institute for Biogeochemistry, Hans-Knöll-Strasse 10, 07745 Jena, Germany E-mail: twutz@bgc-jena.mpg.de (email)
  • Profft, Max Planck Institute for Biogeochemistry, Hans-Knöll-Stra§e 10, 07745 Jena, Germany E-mail: ip@nn.de
  • Mund, Max Planck Institute for Biogeochemistry, Hans-Knöll-Stra§e 10, 07745 Jena, Germany E-mail: mm@nn.de
article id 110, category Research article
Erlend Nybakk, Pablo Crespell, Eric Hansen. (2011). Climate for innovation and innovation strategy as drivers for success in the wood industry: moderation effects of firm size, industry sector, and country of operation. Silva Fennica vol. 45 no. 3 article id 110. https://doi.org/10.14214/sf.110
Keywords: comparative management; innovations; organisational culture; strategy; wood industry; working climate
Abstract | View details | Full text in PDF | Author Info
This study examines the relationships between firm financial performance and a) the climate for innovation and b) innovation strategy in the wood products industry. The focus is on the moderator effects of firm size, country of operation, and industry sector. Using a sample of 460 responses from chief executive officers and top managers of Norwegian and US firms, we conducted a regression analysis to probe for interaction effects. The sample included primary and secondary manufacturers of various sizes. Consistent with previous studies, we found a positive impact for both a climate for innovation and an innovation strategy on firm performance. In terms of moderation, only one interaction was found to be significant, representing a moderator effect of industry size on the climate-performance relationship. Further testing showed that secondary, large manufacturers exhibited a weaker, yet still positive, relationship between climate for innovation and performance. This low level of significant interactions suggests stability of the relationship among the main factors depicted in the model, with important implications for managers and future research. These findings indicate that a positive climate for innovation and a management committed to innovation through an innovation strategy have a positive effect on the bottom line of wood products firms. This effect holds true regardless of industry, size, or country, so most firms can benefit from the implementation of these pro-innovation practices.
  • Nybakk, Norwegian Forest and Landscape Institute, P.O. Box 115, N-1431 Ås, Norway E-mail: nye@skogoglandskap.no (email)
  • Crespell, FPInnovations (Forintek Division), Vancouver, BC, Canada E-mail: pc@nn.fi
  • Hansen, Oregon State University, College of Forestry, OR, USA E-mail: eh@nn.us
article id 109, category Research article
Ann Kristin Raymer, Terje Gobakken, Birger Solberg. (2011). Optimal forest management with carbon benefits included. Silva Fennica vol. 45 no. 3 article id 109. https://doi.org/10.14214/sf.109
Keywords: forest management; Norway spruce; substitution; CO2; greenhouse gas mitigation; optimisation; wood products
Abstract | View details | Full text in PDF | Author Info
In this paper, we analyse how optimal forest management of even aged Norway spruce changes when economic values are placed on carbon fixation, release, and saved greenhouse gas emissions from using wood instead of more energy intensive materials or fossil fuels. The analyses are done for three different site qualities in Norway, assuming present climate and with a range of CO2 prices and real rates of return. Compared to current recommended management, the optimal number of plants per ha and harvest age are considerably higher when carbon benefits are included, and increase with increasing price on CO2. Furthermore, planting becomes more favourable compared to natural regeneration. At the medium site quality, assuming 2% p.a. real rate of return and 20 euros per ton CO2, optimal planting density increases from 1500 per ha to 3000 per ha. Optimal harvest age increases from 90 to 140 years. Including saved greenhouse gas emissions when wood is used instead of more energy intensive materials or fossil fuels, i.e. substitution effects, does not affect optimal planting density much, but implies harvesting up to 20 years earlier. The value of the forest area increases with increasing price on CO2, and most of the income is from carbon. By using the current recommended management in calculations of carbon benefit, our results indicate that the forest’s potential to provide this environmental good is underestimated. The study includes many uncertain factors. Highest uncertainty is related to the accuracy of the forest growth and mortality functions at high stand ages and densities, and that albedo effects and future climate changes are not considered. As such, the results should be viewed as exploratory and not normative.
  • Raymer, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, P.O. Box 5003, N-1432 Ås, Norway E-mail: akr@nn.no
  • Gobakken, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, P.O. Box 5003, N-1432 Ås, Norway E-mail: terje.gobakken@umb.no (email)
  • Solberg, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, P.O. Box 5003, N-1432 Ås, Norway E-mail: bs@nn.no
article id 108, category Research article
Back Tomas Ersson, Urban Bergsten, Ola Lindroos. (2011). The cost-efficiency of seedling packaging specifically designed for tree planting machines. Silva Fennica vol. 45 no. 3 article id 108. https://doi.org/10.14214/sf.108
Keywords: logistics; mechanized planting; containerized seedlings; seedling transport; cost analysis; seedling feed; nursery technology
Abstract | View details | Full text in PDF | Author Info
Today’s crane-mounted planting heads plant seedlings with biologically similar or better results than operational manual planting. However, the total cost of mechanized tree planting in southern Sweden must decrease at least 25% to compete economically with manual planting. Although seedlings packed in machine-specific packaging increase the productivity of planting machines by reducing seedling reloading time, they also increase logistics and investment costs. In this study, we analyzed the total cost of outplanting seedlings with an excavator-mounted Bracke Planter and seedlings packed according to four different concepts: cultivation trays, cardboard boxes, band-mounted seedlings in cardboard boxes and linked pots in container modules. The total cost per planted seedling was calculated for each packaging system as the sum of all costs from nursery to the recovery of empty packaging. The results showed that today’s system of transporting seedlings in cultivation trays is the most cost-efficient of the four alternatives. Machine-specific seedling packaging was 16–23% costlier per planted seedling than cultivation trays when trucking distances were 100 km. Sensitivity analyses indicated that machine-specific seedling packaging increased in cost-efficiency relative to cultivation trays primarily when more planting machines were contracted, but also as planting machine fixed costs and productivity increased. Moreover, the relative cost-efficiency of band-mounted seedlings, but not seedlings in container modules, increased with increasing trucking distance. Thus, we show that investments in machine-specific seedling packaging for today’s planting machines are justified only when the fixed costs, productivity and number of contracted planting machines increase substantially.
  • Ersson, Swedish University of Agricultural Sciences, Department of Forest Resource Management, SE-90183 Umeå, Sweden E-mail: back.tomas.ersson@slu.se (email)
  • Bergsten, Swedish University of Agricultural Sciences, Department of Forest Resource Management, SE-90183 Umeå, Sweden E-mail: ub@nn.se
  • Lindroos, Swedish University of Agricultural Sciences, Department of Forest Resource Management, SE-90183 Umeå, Sweden E-mail: ol@nn.se
article id 107, category Research article
Jaana Luoranen, Risto Rikala, Heikki Smolander. (2011). Machine planting of Norway spruce by Bracke and Ecoplanter: an evaluation of soil preparation, planting method and seedling performance. Silva Fennica vol. 45 no. 3 article id 107. https://doi.org/10.14214/sf.107
Keywords: Picea abies; height growth; machine planting; mechanized forestry; mounding; quality of planting; survival
Abstract | View details | Full text in PDF | Author Info
We evaluated the effects of planting date and planting machine (Bracke: three machines, 69 regeneration areas in three years; Ecoplanter: six areas, two years) on the quality and field performance one and three years after planting of Norway spruce (Picea abies (L.) Karst.) seedlings in central Finland. Both machine types planted on average 1800 seedlings per hectare, and after three years approximately 1600 (Bracke) and 1200 (Ecoplanter) were still alive. This study suggests that planting with a Bracke machine can achieve better regeneration rates than those observed in privately-owned Finnish forests. We characterized the quality of mounding and planting with the Bracke machine as excellent and that of the Ecoplanter as good. The soil preparation method of the Ecoplanter produced humus-rich mounds where seedlings were susceptible to pine weevils and consequently suffered higher mortality. Different machines were used in different regional areas and each machine was operated by different driver/s which may have influenced the results. No negative effects of planting date were observed. Seedling growth decreased if they were tall in relation to their root plug volume, grown too densely in the nursery, and if stored in the field for several months prior to planting. We conclude that mechanized planting is successful when the soil preparation method produces mounds covered by purely mineral soil. Planting from May to the end of September is suitable for seedlings intended for use during this period.
  • Luoranen, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: jaana.luoranen@metla.fi (email)
  • Rikala, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: rr@nn.fi
  • Smolander, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: hs@nn.fi
article id 106, category Research article
Hannu Hökkä, Heli Hyttinen, Hannu Marttila, Juha Jämsen, Bjørn Kløve. (2011). Effect of peak runoff control method on growth of Scots pine stands on drained peatlands in central Finland. Silva Fennica vol. 45 no. 3 article id 106. https://doi.org/10.14214/sf.106
Keywords: peatland; forest drainage; stand growth; runoff control; water quality
Abstract | View details | Full text in PDF | Author Info
In drained peatland forests ditch networks need regular maintenance operations in order to sustain their drainage capacity. These operations however have a significant impact on the quality of the runoff water from the ditched areas. Peak runoff control (PRC) method has been proposed as a possible method to diminish the load to water courses through retention of the runoff temporarily in the ditch network during maximum runoff events using dams with a plastic control pipe. However, blocking water into the ditched area for periods of varying length during the growing season may have a negative impact on the growth of the tree stands. In this study past stand growth was investigated in Central Finland in altogether 10 sample Scots pine thinning stands in which the PRC method has been applied 5 growing seasons earlier. In each stand, a pair of sample plots was established: one plot next to the dam within the influence of periodic flooding and the other one outside the effect of periodic flooding. For determining stand growth, field measurements were made in August 2009. Stand growth near the dam was on average 0.54 m3 ha-1 a-1 lower than farther away from the dam but the analysis of covariance showed that the dam effect was not significant. The results of this study suggest that the PCR method does not decrease Scots pine stand growth during the first five year growth period after ditch cleaning.
  • Hökkä, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: hannu.hokka@metla.fi (email)
  • Hyttinen, Metsänhoitoyhdistys Keski-Suomi, Viitasaari, Finland E-mail: hh@nn.fi
  • Marttila, University of Oulu, Water Resources and Environmental Engineering Lab, Oulu, Finland E-mail: hm@nn.fi
  • Jämsen, Forestry Centre Keski-Suomi, Pihtipudas, Finland E-mail: jj@nn.fi
  • Kløve, University of Oulu, Water Resources and Environmental Engineering Lab, Oulu, Finland E-mail: bk@nn.fi
article id 105, category Research article
Jaana Luoranen, Risto Rikala. (2011). Nutrient loading of Norway spruce seedlings hastens bud burst and enhances root growth after outplanting. Silva Fennica vol. 45 no. 3 article id 105. https://doi.org/10.14214/sf.105
Keywords: Picea abies; late-season fertilization; nitrogen concentration; root growth capacity
Abstract | View details | Full text in PDF | Author Info
We studied the effects of late season nutrient loading (NLOAD) on the timing of bud burst, growth and changes in nitrogen (N) concentrations in the first growing season after seedlings were outplanted. Two-year-old Norway spruce (Picea abies (L.) Karst.) seedlings with three foliar nitrogen concentration levels (NLOAD levels 11.3, 22.5 and 27.5 g N kg-1 for L, M- and H-seedlings, respectively) were examined in the following three experiments: root growth capacity test (RGC), rooting experiment in the field and soil fertility experiment (‘rich’ or ‘poor’ soil) in the field. Bud burst in RGC was monitored daily and foliar N concentration (field experiments), height and root growth (rooting experiment) at monthly intervals. With respect to the RGC test, no differences in root growth were observed among the three NLOAD levels, but buds of H-seedlings burst 2–6 days earlier than others. In the rooting experiment, nutrient loading increased height and root growth but did not affect the timing of height growth. In the soil fertility experiment, foliar N of H- and M-seedlings decreased rapidly, but the decline was slower in rich soil. Current-year needles had more N in seedlings growing in rich soil and the N concentration declined until height growth ceased whereafter it increased until autumn. Improved growth from nutrient loading seems to last only for the first season after planting and the greatest benefits are enjoyed by seedlings planted in poor soils.
  • Luoranen, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600, Suonenjoki, Finland E-mail: jaana.luoranen@metla.fi (email)
  • Rikala, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600, Suonenjoki, Finland E-mail: rr@nn.fi
article id 104, category Research article
Eeva Terhonen, Teresa Marco, Hui Sun, Risto Jalkanen, Risto Kasanen, Martti Vuorinen, Fred Asiegbu. (2011). The effect of latitude, season and needle-age on the mycota of Scots pine (Pinus sylvestris) in Finland. Silva Fennica vol. 45 no. 3 article id 104. https://doi.org/10.14214/sf.104
Keywords: Pinus sylvestris; needles; harsh environment; mycota; needle age; cryptic lifecycle; Hormonema dematioides
Abstract | View details | Full text in PDF | Author Info
The seasonal and latitudinal influences on the diversity and abundance of mycota of Pinus sylvestris needles were investigated. A sample of 1620 needles resulted in a total of 3868 fungal isolates, which were assigned to 68 operational taxonomic units (OTUs). The majority of these OTUs (65%) belong to Ascomycota and only 0.03% was grouped as Basidiomycota. The dominant and most frequently isolated OTU was Hormonema dematioides. Other well-known species with a saprotrophic nutritional mode such as Lophodermium spp. were also observed. The abundance of fungi increased from fall to spring. Frequencies varied significantly in Northern and Southern Finland suggesting that factors associated with latitudinal differences have an impact on the abundance of fungi.
  • Terhonen, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: et@nn.fi (email)
  • Marco, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: tm@nn.fi
  • Sun, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: hs@nn.fi
  • Jalkanen, Finnish Forest Research Institute, Rovaniemi Research Unit, Rovaniemi, Finland E-mail: rj@nn.fi
  • Kasanen, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: rk@nn.fi
  • Vuorinen, Finnish Forest Research Institute, Suonenjoki Research Unit, Suonenjoki, Finland E-mail: mv@nn.fi
  • Asiegbu, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: fa@nn.fi
article id 117, category Research article
Harri Kilpeläinen, Jari Lindblad, Henrik Heräjärvi, Erkki Verkasalo. (2011). Saw log recovery and stem quality of birch from thinnings in southern Finland. Silva Fennica vol. 45 no. 2 article id 117. https://doi.org/10.14214/sf.117
Keywords: Betula pendula; Betula pubescens; thinning; European white birch; silver birch; saw logs; stem quality; stem defects; bucking; recovery
Abstract | View details | Full text in PDF | Author Info
The objective of this study was to examine the timber quality of silver birch (Betula pendula Roth) and European white birch (Betula pubescens Ehrh.) trees in the first and second thinnings in southern parts of Finland, from the viewpoint of sawing of small-diameter, short logs, in particular. The average stem volume of birch was 0.140 m3 in the first thinning stands and 0.206 m3 in the second thinning stands. In planted stands, the trees were larger in the first thinnings but slightly smaller in the second thinnings, compared with naturally regenerated pure birch stands or mixed stands of Norway spruce and birch species. Almost 60% of the harvested and 35% of the remaining stems that could provide saw logs were graded as pulpwood for timber quality due to the occurrence of stem defects. The most common stem defects were multiple crooks and middle crooks. Only minor between-stratum differences were detected in the numbers of defects. Depending on the bucking option, the total percentage of saw and plywood logs from the total birch recovery in the thinning of the sample stands varied between 11.7 and 18.2. The recovery of saw logs was clearly higher in the second thinnings, 12–19%, than in the first thinnings, 8–14%. Of the stand types, saw log recovery was the highest in planted birch stands, 12–19%, but lower in naturally regenerated pure birch stands and mixed stands of Norway spruce and birch. The highest share of saw logs was in the second thinning of planted stands, 17–25%. This study shows that the harvesting recoveries of end-use based timber assortments can be estimated in different kinds of thinning birch stands. Based on tree and log dimensions and stem squality, silver birch firstly from plantations and secondly from mixed stands should be the most interesting source of raw material for the saw milling, furniture and interior product sectors.
  • Kilpeläinen, Finnish Forest Research Institute, Eastern Finland Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: hk@nn.fi
  • Lindblad, Finnish Forest Research Institute, Eastern Finland Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jl@nn.fi
  • Heräjärvi, Finnish Forest Research Institute, Eastern Finland Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: henrik.herajarvi@metla.fi (email)
  • Verkasalo, Finnish Forest Research Institute, Eastern Finland Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: ev@nn.fi
article id 116, category Research article
Eva-Maria Nordström, Ljusk Ola Eriksson, Karin Öhman. (2011). Multiple criteria decision analysis with consideration to place-specific values in participatory forest planning. Silva Fennica vol. 45 no. 2 article id 116. https://doi.org/10.14214/sf.116
Keywords: forest management; decision support; public participation; spatial planning
Abstract | View details | Full text in PDF | Author Info
The combination of multiple criteria decision analysis (MCDA) and participatory planning is an approach that has been applied in complex planning situations where multiple criteria of very different natures are considered, and several stakeholders or social groups are involved. The spatial character of forest planning problems adds further to the complexity, because a large number of forest stands are to be assigned different treatments at different points in time. In addition, experience from participatory forest planning indicates that stakeholders may think about the forest in terms of place-specific values rather than in forest-wide terms. The objective of this study was to present an approach for including place-specific values in MCDA-based participatory forest planning and illustrate the approach by a case study where the objective was to choose a multipurpose forest plan for an area of urban forest in northern Sweden. Stakeholder values were identified in interviews, and maps were used to capture place-specific spatial values. The nonspatial and nonplace-specific spatial values were formulated as criteria and used to build an objective hierarchy describing the decision situation. The place-specific spatial values were included in the creation of a map showing zones of different silvicultural management classes, which was used as the basis for creation of forest plan alternatives in the subsequent process. The approach seemed to work well for capturing place-specific values, and the study indicates that formalized methods for including and evaluating place-specific values in participatory forest planning processes should be developed and tested further.
  • Nordström, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd 1, SE-901 83 Umeå, Sweden E-mail: eva-maria.nordstrom@slu.se (email)
  • Eriksson, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd 1, SE-901 83 Umeå, Sweden E-mail: loe@nn.se
  • Öhman, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Skogsmarksgränd 1, SE-901 83 Umeå, Sweden E-mail: ko@nn.se
article id 115, category Research article
Marjut Turtiainen, Kauko Salo, Olli Saastamoinen. (2011). Variations of yield and utilisation of bilberries (Vaccinium myrtillus L.) and cowberries (V. vitis-idaea L.) in Finland. Silva Fennica vol. 45 no. 2 article id 115. https://doi.org/10.14214/sf.115
Keywords: bilberry; cowberry; total berry yield; yield variation; utilisation of wild berries
Abstract | View details | Full text in PDF | Author Info
So far, only rough estimates for the utilisation rates of wild berries in Finland have been available. One reason for this is that there has been a lack of empirical-knowledge-based studies concerning total yields of wild berries and their yield variations. This study had three aims: 1) total bilberry and cowberry yields of an average crop year were calibrated for different (abundant and poor) crop years using the inventory data on wild berries collected by the Finnish Forest Research Institute (1997–2008); 2) national utilisation rates of bilberries and cowberries were calculated for three different berry years 1997–1999; and 3) regional utilisation rates of these berry species were calculated for the year 1997. According to calculations, annual bilberry yields in Finland vary from 92 to 312 million kg. For cowberry, the range of variation in total berry yields is from 129 to 386 million kg. It was also found that approximately the same proportion of the total yield of bilberries (i.e. 5–6%) was collected between 1997 and 1999. Utilisation rates of cowberries were also quite constant varying from approximately 8% to nearly 10%. In 1997, bilberries and cowberries were utilised most intensively in the eastern parts of the country and in the Oulu-Kainuu region. The results of this present study describe the situation before the phenomenon of foreign pickers. It can be presumed that commercial wild berry picking by migrant collectors has so far affected both national and regional utilisation rates of wild berries.
  • Turtiainen, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: marjut.turtiainen@uef.fi (email)
  • Salo, Finnish Forest Research Institute, Joensuu Research Unit, Joensuu, Finland E-mail: ks@nn.fi
  • Saastamoinen, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: os@nn.fi
article id 114, category Research article
Roy V. Rea. (2011). Impacts of moose (Alces alces) browsing on paper birch (Betula papyrifera) morphology and potential timber quality. Silva Fennica vol. 45 no. 2 article id 114. https://doi.org/10.14214/sf.114
Keywords: deciduous; browse damage; forestry; hardwood; silviculture; ungulate; wood quality
Abstract | View details | Full text in PDF | Author Info
  • Rea, University of Northern British Columbia, 3333 University Way, Prince George, BC, Canada V2N 4Z9 E-mail: reav@unbc.ca (email)
article id 113, category Research article
Jinsong Wang, Chunyu Zhang, Fucai Xia, Xiuhai Zhao, Lianhai Wu, Klaus von Gadow. (2011). Biomass structure and allometry of Abies nephrolepis (Maxim) in Northeast China. Silva Fennica vol. 45 no. 2 article id 113. https://doi.org/10.14214/sf.113
Keywords: biomass structure; aboveground competition; allometry; Abies nephrolepis (Maxim); tree size
Abstract | View details | Full text in PDF | Author Info
Above- and below-ground tree biomass structure and allometric relationships of Abies nephrolepis (Maxim) were assessed in an old secondary forest dominated by A. nephrolepis, Pinus koraiensis, Quercus mongolica, Tilia amurensis, Fraxinus mandshurica and Acer mono in northeast China. Based on the breast-height diameter (D), a total of 21 sample trees were divided into three tree size classes: the small trees (1 cm ≤ D < 10 cm), the medium trees (10 cm ≤ D < 20 cm) and the big trees (D ≥ 20 cm). The greatest amount of live branch biomass was located in the middle and bottom layers of the crown, while the largest foliage biomass was found in the middle layer in each tree size category. The relative contribution of canopy biomass components (live branches and foliage) decreased with increasing tree size, while that of coarse root biomass remained almost constant. The relationship between above- and belowground biomass was linear. D and tree height (H) decreased with increasing competition intensity. The small trees had lower average crown ratio and higher average height-to-diameter ratio than those of the medium and big trees. The big trees had higher average stem to foliage mass ratio than those of the small and medium trees. Crown ratio, height-to-diameter ratio and stem to foliage mass ratio were not correlated with competition intensity in the same tree size class. Root to shoot mass ratio was almost constant among tree sizes. Allometric equations based on D gave higher correlations compared to those with other stem diameters: at tree base, at 30-cm height and at crown base.
  • Wang, Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing 100083, China E-mail: jw@nn.cn
  • Zhang, Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing 100083, China E-mail: cz@nn.cn
  • Xia, Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing 100083, China E-mail: fx@nn.cn
  • Zhao, Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing 100083, China E-mail: zhaoxh@bjfu.edu.cn (email)
  • Wu, Rothamsted Research, Okehampton, Devon, UK E-mail: lw@nn.uk
  • Gadow, Faculty of Forestry and Forest Ecology, Georg-August-University Göttingen, Göttingen, Germany E-mail: kvg@nn.de
article id 112, category Research article
Mike R. Saunders, Shawn Fraver, Robert G. Wagner. (2011). Nutrient concentration of down woody debris in mixedwood forests in central Maine, USA. Silva Fennica vol. 45 no. 2 article id 112. https://doi.org/10.14214/sf.112
Keywords: Acadian Forest; hardwoods; softwoods; carbon-nitrogen ratios; decay classes; disturbance-based silviculture
Abstract | View details | Full text in PDF | Author Info
Both nutrient concentrations and pre- and post-harvest pool sizes were determined across down woody debris decay classes of several hardwood and softwood species in a long-term, natural disturbance based, silvicultural experiment in central Maine. Concentrations of N, P, Ca, Mg, Cu, Fe, and Zn generally increased 2- to 5-fold with increasing decay class. Concentrations of Mn, Al and B did not differ among decay classes, while K decreased by 20–44% from decay class 1 to class 4. C:N-ratios declined with increasing decay class, while N:P-ratios increased from decay class 1 to 2 and then plateaued with further decay. Within decay classes, softwoods generally had lower nutrient concentrations and higher C:N-ratios than hardwoods; N:P-ratios did not differ between hardwoods and softwoods. Although gap harvesting increased the size of the overall down woody debris nutrient pools, mostly through a large pulse of decay class 1 material, harvesting generally reduced the nutrients held in advanced decay classes. Pre-harvest down woody debris pools for N, P, K and Ca were 11.0, 0.6, 2.1 and 21.1 kg ha–1, respectively, while postharvest were 20.0, 1.3, 6.2 and 46.2 kg ha–1, respectively. While the gap-based silvicultural systems sampled in this study doubled the size of the pre-harvest, downed woody debris nutrient pools, the post-harvest pools were estimated to be only 3.2–9.1% of aboveground nutrients.
  • Saunders, Department of Forestry and Natural Resources, Purdue University, 715 State Street, West Lafayette, IN, USA E-mail: msaunder@purdue.edu (email)
  • Fraver, USFS Northern Research Station, Grand Rapids, MN, USA E-mail: sf@nn.us
  • Wagner, School of Forest Resources, University of Maine, Orono, ME, USA E-mail: rgw@nn.us
article id 41, category Research article
Ying Hou, Jintao Qu, Zukui Luo, Chao Zhang, Kaiyun Wang. (2011). Morphological mechanism of growth response in treeline species Minjiang fir to elevated CO2 and temperature. Silva Fennica vol. 45 no. 2 article id 41. https://doi.org/10.14214/sf.41
Keywords: climate change; Abies faxoniana; crown architecture; leaf morphology; response ratio
Abstract | View details | Full text in PDF | Author Info
To test whether and how morphological traits are linked with growth responses of plants to temperature and CO2 is important for understanding the mechanism underlying how plant growth will respond to global warming. In this study, using closed-top chambers to mimic future elevated CO2 and temperature, the growth response, morphological traits of Minjiang fir (Abies faxoniana Rehd.et Wils.) and the relationship of the two were investigated after two years of exposure to the single and combined elevation of CO2 and temperature. The results showed that biomass of Minjiang fir was 21%, 31%, and 35% greater than the control in elevated CO2, elevated temperature and the combination of elevated CO2 and temperature treatments, respectively. Elevated CO2 and temperature significantly affected the morphology of Minjiang fir, and a few morphological traits were highly correlated with growth responses. Larger branch angles at the upper layer, crown volume, and relative crown length contributed to positive growth responses to elevated CO2, while decreased specific leaf area (SLA) constricted any further growth response. Leaf morphological traits were more closely correlated with the response ratio than crown did in the elevated temperature, while in the combination of elevated CO2 and temperature, crown was more correlated with the response ratio than the leaf morphological traits. Thus, our results indicate that morphological traits may contribute differently to growth responses under different experimental conditions.
  • Hou, Department of Life Sciences, Shangqiu Normal University, Shangqiu, China E-mail: yh@nn.cn
  • Qu, Department of Life Sciences, Shangqiu Normal University, Shangqiu, China E-mail: jq@nn.cn
  • Luo, School of Environment and Life Sciences, Kaili University, Kaili, China E-mail: zl@nn.cn
  • Zhang, Shanghai Key Laboratory of Urbanization and Ecological Restoration, East China Normal University, Shanghai, China, and University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: cz@nn.cn
  • Wang, Shanghai Key Laboratory of Urbanization and Ecological Restoration, East China Normal University, Shanghai, China, and University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: kywang@re.ecnu.edu.cn (email)
article id 40, category Research article
Jorge Martín-García, Elba Espiga, Valentín Pando, Julio Javier Diez. (2011). Factors influencing endophytic communities in poplar plantations. Silva Fennica vol. 45 no. 2 article id 40. https://doi.org/10.14214/sf.40
Keywords: endophyte; poplar; management; site quality; foliar nutrients; forest health
Abstract | View details | Full text in PDF | Author Info
The fungal species associated with leaves and twigs from stands of Populus x euramericana in northern Spain were studied with the aim of evaluating the effects of several factors on endophytic communities in these plantations. Endophyte assemblages were analysed in 12 poplar plantations (clone I-214), chosen according to a factorial scheme with two factors: age and site quality. Crown condition, dendrometric variables and foliar nutrients were recorded in each sampled tree to evaluate their effects on endophytic communities. Fungal species richness and relative isolation frequency (RIF) were higher in young stands than in adult stands. Moreover, the age-related differences depended on site quality, with the lowest richness levels observed in adult stands located in poor sites. At stand level, endophyte assemblages varied among stands according to site quality and, to a lesser extent, stand age. On the other hand, crown discoloration, total height and foliar concentrations of iron and zinc may be key indicators of endophytic communities in poplar plantations, at tree level.
  • Martín-García, Sustainable Forest Management Research Institute, University of Valladolid – INIA, Avenida de Madrid 57, 34004 Palencia, Spain, and Forestry Engineering, University of Extremadura, Plasencia, Spain E-mail: jorgemg@pvs.uva.es (email)
  • Espiga, Sustainable Forest Management Research Institute, University of Valladolid – INIA, Palencia, Spain E-mail: ee@nn.es
  • Pando, Statistics and Operations Research Department, University of Valladolid, Spain E-mail: vp@nn.es
  • Diez, Sustainable Forest Management Research Institute, University of Valladolid – INIA, Palencia, Spain E-mail: jjd@nn.es
article id 35, category Research article
Tapio Rantala. (2011). Democratic legitimacy of the forest sector and nature conservation decision-making in Finnish print media discussion. Silva Fennica vol. 45 no. 1 article id 35. https://doi.org/10.14214/sf.35
Keywords: legitimacy; democracy; policy evaluation; forest policy; nature conservation policy; national forest program; nature conservation program
Abstract | View details | Full text in PDF | Author Info
The study explores perceived democratic legitimacy of forest-related decision-making processes in the Finnish print media discourse. The data consists of the readers’ letters in four journals (n = 530), and the comments given during the preparation of the Finnish National Forest Program (n = 140). The objective is to identify the patterns of democratic legitimacy and respective performance evaluations of actual decision-making processes. The patterns can be classified as support for: (A) democracy and other forms of government, (B) different forms of participation, and (C) principles of democracy. The principles can be further classified into 1) core regime, 2) input, 3) throughput, and 4) output principles. Democratic legitimacy was found to be an important source of legitimacy in the public discussion since democratic patterns were found in more than half of the texts. The most common core legitimacy principles included freedom of speech, good national and international standing, forerunnership, and legality at national and international level. The central principles related to input legitimacy included popular sovereignty, a voice for the people, popular participation, openness, presenting alternatives, and urgency. The consensus and majority rules were found to be the most prominent throughput principles. Democratic output legitimacy included accountability, responsibility, cooperation, commitment, responsiveness, the possibility to appeal, credibility, comprehensiveness, and understandability. The findings suggest that among the writers of readers’ letters there is less contestation regarding the principles of democratic legitimacy but there are significant disagreements concerning the performance of decision-making processes. The negative performance evaluations were two times more frequent than the positive evaluations.
  • Rantala, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: tapio.rantala@helsinki.fi (email)
article id 34, category Research article
Natalie Macias, Chris Knowles. (2011). Examining the effect of environmental certification, wood source, and price on architects’ preferences of hardwood flooring. Silva Fennica vol. 45 no. 1 article id 34. https://doi.org/10.14214/sf.34
Keywords: architects; hardwood flooring; conjoint analysis; environmental certification; wood source
Abstract | View details | Full text in PDF | Author Info
This article examines the importance architects place on three factors, environmental certification, wood source, and price, when specifying hardwood flooring. Architects were presented with nine flooring scenarios, in which the three factors were present in varying levels. They were asked to rank the scenarios from the least preferred to the most preferred. Data were obtained from a mail survey of architects in Oregon and Washington, U.S.A. (n = 402). Conjoint analyses determined that architects consider price and wood source to be the most important factors when specifying hardwood flooring. Interestingly, environmental certification was considered the least important factor. The respondents were then separated into three groups for further analysis based on whether they identified themselves as more influenced by environmental factors (biocentric) or human needs (anthropocentric). This analysis showed that the biocentric group favored wood source over price and environmental certification, while the anthropocentric group favored price.
  • Macias, Oregon State University, Department of Wood Science and Engineering, 119 Richardson Hall, Corvallis, OR 97331, USA E-mail: natalie.macias@gmail.com
  • Knowles, Oregon State University, Department of Wood Science and Engineering, 119 Richardson Hall, Corvallis, OR 97331, USA E-mail: chris.knowles@oregonstate.edu (email)
article id 33, category Research article
Raffaele Spinelli, Natascia Magagnotti, Giuseppe Paletto, Christian Preti. (2011). Determining the impact of some wood characteristics on the performance of a mobile chipper. Silva Fennica vol. 45 no. 1 article id 33. https://doi.org/10.14214/sf.33
Keywords: chipping; productivity; biomass; fuel consumption
Abstract | View details | Full text in PDF | Author Info
A study was conducted to determine the effect of some wood characteristics such as species, moisture content and tree part on the performance and product quality offered by a mobile industrial chipper, of the type commonly used for roadside chipping. Two main species, two tree parts and two moisture content levels were combined in a factorial design yielding 8 treatments, each replicated 5 or 6 times. A flow meter was installed on the chipper engine, and all chips produced were weighed and sampled for moisture content and particle size distribution. The results indicated that some wood characteristics such as species and moisture content have a secondary effect on chipper productivity and fuel consumption, which are primarily controlled by piece size. In particular, fuel consumption per unit dry mass seem to be rather constant and in the range of 3.2 l per oven dry ton. Moisture content and tree part may have a significant effect on the particle size distribution of chips. Of course, these results were only verified for the species used in the test and for industrial chippers, and may change if substantially different species or machines are used.
  • Spinelli, CNR IVALSA, Via Madonna del Piano 10, Sesto Fiorentino (FI), Italy E-mail: spinelli@ivalsa.cnr.it (email)
  • Magagnotti, CNR IVALSA, Via Biasi 75, S. Michele all’Adige (TN), Italy E-mail: nm@nn.it
  • Paletto, CNR IMAMOTER, Strada delle Cacce 73, Torino, Italy E-mail: gp@nn.it
  • Preti, CNR IMAMOTER, Strada delle Cacce 73, Torino, Italy E-mail: cp@nn.it
article id 32, category Research article
Susete Marques, Jordi Garcia-Gonzalo, José G. Borges, Brigite Botequim, M. Manuela Oliveira, José Tomé, Margarida Tomé. (2011). Developing post-fire Eucalyptus globulus stand damage and tree mortality models for enhanced forest planning in Portugal. Silva Fennica vol. 45 no. 1 article id 32. https://doi.org/10.14214/sf.32
Keywords: forest fires; forest management; Eucalyptus globulus Labill; damage model; post-fire mortality
Abstract | View details | Full text in PDF | Author Info
Forest and fire management planning activities are carried out mostly independently of each other. This paper discusses research aiming at the development of methods and tools that can be used for enhanced integration of forest and fire management planning activities. Specifically, fire damage models were developed for Eucalyptus globulus Labill stands in Portugal. Models are based on easily measurable forest characteristics so that forest managers may predict post-fire mortality based on forest structure. For this purpose, biometric data and fire-damage descriptors from 2005/2006 National Forest Inventory plots and other sample plots within 2006, 2007 and 2008 fire areas were used. A three-step modelling strategy based on logistic regression methods was used. In the first step, a model was developed to predict whether mortality occurs after a wildfire in a eucalypt stand. In the second step the degree of damage caused by wildfires in stands where mortality occurs is quantified (i.e. percentage of mortality). In the third step this mortality is distributed among trees. Data from over 85 plots and 1648 trees were used for modeling purposes. The damage models show that relative damage increases with stand basal area. Tree level mortality models indicate that trees with high diameters, in dominant positions and located in regular stands are less prone to die when a wildfire occurs.
  • Marques, Technical University of Lisbon, School of Agriculture, Forest Research Center, Tapada da Ajuda, 1349-017 Lisboa, Portugal E-mail: smarques@isa.utl.pt (email)
  • Garcia-Gonzalo, Technical University of Lisbon, School of Agriculture, Forest Research Center, Tapada da Ajuda, 1349-017 Lisboa, Portugal E-mail: jgg@nn.pt
  • Borges, Technical University of Lisbon, School of Agriculture, Forest Research Center, Tapada da Ajuda, 1349-017 Lisboa, Portugal E-mail: jgb@nn.pt
  • Botequim, Technical University of Lisbon, School of Agriculture, Forest Research Center, Tapada da Ajuda, 1349-017 Lisboa, Portugal E-mail: bb@nn.pt
  • Oliveira, Technical University of Lisbon, School of Agriculture, Forest Research Center, Tapada da Ajuda, 1349-017 Lisboa, Portugal E-mail: mmo@nn.pt
  • Tomé, Technical University of Lisbon, School of Agriculture, Forest Research Center, Tapada da Ajuda, 1349-017 Lisboa, Portugal E-mail: jt@nn.pt
  • Tomé, Technical University of Lisbon, School of Agriculture, Forest Research Center, Tapada da Ajuda, 1349-017 Lisboa, Portugal E-mail: mt@nn.pt
article id 31, category Research article
Jeovanna Lowe, David Pothier, Jean-Pierre L. Savard, Ghislain Rompré, Mathieu Bouchard. (2011). Snag characteristics and cavity-nesting birds in the unmanaged post-fire northeastern Canadian boreal forest. Silva Fennica vol. 45 no. 1 article id 31. https://doi.org/10.14214/sf.31
Keywords: snags; cavity nesting birds; boreal forests; old-growth; long term chronosequence; time since fire; mortality rate; basal area
Abstract | View details | Full text in PDF | Author Info
We studied the availability and characteristics of snags and their use by cavity-nesting birds in the northeastern part of the Canadian boreal forest. We built up two long-term (> 200 years) chronosequences following time since the last fire in the unmanaged boreal forest of northeastern Québec, one in the balsam fir-white birch domain (southern region) and one in the spruce-mosses domain (northern region). We then sampled and characterized snags and live trees in 30 stands from each of these two chronosequences. We also looked for nest cavities on all sampled snags, performed bird inventories by point counts, and calculated tree mortality rate from permanent sample plots. Results show that mortality rates follow a U-shaped pattern, with more snags of large diameter (> 20 cm DBH) in young (< 50 years) and in old (> 200 years) forests. In the latter, we also found more nest cavities than in any other age classes. Although abundance of primary cavity nesters (excavating species) did not vary among age classes, secondary cavity nesters (using cavities already available) tend to be more numerous in older forests. Our results highlight the capacity for young and old-growth forests to provide quality habitat for species that are dependent on large snags. Proper forest management should maintain a mosaic of different age forest stands, including snags, to promote biodiversity and provide important resources for resident bird species.
  • Lowe, Centre d’étude de la foret, Département des sciences du bois et de la foret, Pavillon Abitibi-Price, 2405 rue de la Terrasse, Université Laval, Québec, Québec, G1V 0A6, Canada E-mail: jeovannalowe@gmail.com (email)
  • Pothier, Centre d’étude de la foret, Département des sciences du bois et de la foret, Pavillon Abitibi-Price, 2405 rue de la Terrasse, Université Laval, Québec, Québec, G1V 0A6, Canada E-mail: dp@nn.ca
  • Savard, Wildlife Research, Science and Technology, Québec Region, 1141 Route de l’Église, P.O. Box 10100, Québec, Québec, G1V 4H5, Canada E-mail: jpls@nn.ca
  • Rompré, Centre d’étude de la foret, Département des sciences du bois et de la foret, Pavillon Abitibi-Price, 2405 rue de la Terrasse, Université Laval, Québec, Québec, G1V 0A6, Canada & Department of Biology and Health Sciences, 84 West South Street, Wilkes University, PA 18766, USA E-mail: gr@nn.ca
  • Bouchard, Ministère des Ressources naturelles et de la Faune, Direction de l’Environnement et de la Protection des Forets, 880 Chemin Ste-Foy, Quebec, Québec, G1S 4X4, Canada E-mail: mb@nn.ca
article id 30, category Research article
Hilppa Gregow, Heli Peltola, Mikko Laapas, Seppo Saku, Ari Venäläinen. (2011). Combined occurrence of wind, snow loading and soil frost with implications for risks to forestry in Finland under the current and changing climatic conditions. Silva Fennica vol. 45 no. 1 article id 30. https://doi.org/10.14214/sf.30
Keywords: A1B-scenario; climate change; snow loads; soil frost; stem breakage; uprooting; wind
Abstract | View details | Full text in PDF | Author Info
This work focuses on the combined occurrence of wind, snow loading and soil frost with implications for risks to forestry in Finland under the current and changing climatic conditions. For this purpose, we employ meteorological datasets, available for the period of 1971–2009 and global climate model (GCM) simulations for the current climate 1971–2000, and periods 2046–65 and 2081–2100 applying the A1B-climate change scenario. Based on our results, the wind and snow induced risks to Finnish forests are projected to increase in the future although the change in the occurrence of strong winds is small. This is because soil frost depths that support tree anchorage from late autumn to early spring in Finland are projected to nearly disappear in the southern and central parts of the country. Heavy snow loads > 30 kg m–2 are becoming more common in southern and eastern Finland despite that the average cumulative 5-day snow loads decrease in these areas by 18 to 50%, respectively. As a result of the changes in the combined occurrence of wind, snow loading and soil frost, the risk of climatic conditions making conifers liable to uprooting are projected to increase in southern, central and eastern Finland. In the north, the risk of stem breakage is becoming more pronounced under snow loading > 20 kg m–2. Despite some uncertainties related to this work, we assume that the findings can serve as valuable support for the risk assessment of wind and snow induced damages to Finnish forests and for forestry, in general.
  • Gregow, Finnish Meteorological Institute, Helsinki, Finland E-mail: hilppa.gregow@fmi.fi (email)
  • Peltola, University of Eastern Finland, Faculty of Forest Sciences, Joensuu, Finland E-mail: heli.peltola@uef.fi
  • Laapas, Finnish Meteorological Institute, Helsinki, Finland E-mail: ml@nn.fi
  • Saku, Finnish Meteorological Institute, Helsinki, Finland E-mail: ss@nn.fi
  • Venäläinen, Finnish Meteorological Institute, Helsinki, Finland E-mail: av@nn.fi
article id 29, category Research article
Piotr Androsiuk, Roman Zielinski, Kornelia Polok. (2011). B-SAP markers derived from the bacterial KatG gene differentiate populations of Pinus sylvestris and provide new insights into their postglacial history. Silva Fennica vol. 45 no. 1 article id 29. https://doi.org/10.14214/sf.29
Keywords: Pinus sylvestris; KatG gene; molecular markers; population differentiation; migrations
Abstract | View details | Full text in PDF | Author Info
The aim of the studies was to evaluate the efficiency of the KatG gene based B-SAP markers as a tool to distinguish morphologically diversified and geographically distant Scots pine populations and to track the routes of migrations. The 19 populations growing in the IUFRO 1982 provenance experiment and representing the natural distribution of the species in Europe were scored using 103 B-SAP loci. Among them 26% loci were polymorphic. The level of polymorphism was associated with the location of primers on the KatG template. The diversity was low, He = 0.086, and deposited mostly among populations. Seven unique markers were found that identified populations and likely they were associated with morphology. The overall genetic identity was relatively low, I = 0.933 (D = 0.069). The block of six B-SAP markers discriminated populations into two groups in agreement with their geographic origin and thereby further described as the North and the South. The North group was uniform with genetic diversity, He = 0.026 and the overall genetic distance D = 0.022. Presumably, it migrated from refugia in the Alps via France, northern Germany and Denmark, to Scandinavia and Russia. The South group was heterogeneous with He = 0.063 and D = 0.047. This group migrated from the Carpathians via Slovakia to Germany and Poland. The Balkans and Asian refugia did not take part in recolonization of Europe. The block of six B-SAP/KatG markers can be recommended for tracking postglacial history of Scots pine.
  • Androsiuk, University of Warmia and Mazury in Olsztyn, Department of Genetics, Plac Lodzki 3, 10-967 Olsztyn, Poland E-mail: pa@nn.pl
  • Zielinski, University of Warmia and Mazury in Olsztyn, Department of Genetics, Plac Lodzki 3, 10-967 Olsztyn, Poland E-mail: rz@nn.pl
  • Polok, University of Warmia and Mazury in Olsztyn, Department of Genetics, Plac Lodzki 3, 10-967 Olsztyn, Poland E-mail: kpolok@moskit.uwm.edu.pl (email)
article id 28, category Research article
Lu-Min Vaario, Kim Yrjälä, Matti Rousi, Timo Sipilä, Pertti Pulkkinen. (2011). Leaf number indicates salt tolerance of young seedling families of European aspen (Populus tremula L.) growing in different soils. Silva Fennica vol. 45 no. 1 article id 28. https://doi.org/10.14214/sf.28
Keywords: leaf number; Populus tremula; root; shoot ratio; salt tolerance
Abstract | View details | Full text in PDF | Author Info
Soil salinity limits plant productivity and quality. We evaluated the response of 12 aspen (Populus tremula) families to salt stress in two different soils irrigated for 4-weeks with 0, 80 or 160 mM saline solution. Easily measurable characteristics such as shoot height, leaf number, dry mass as well as the distribution of sodium (Na+) ions were measured in 5-month-old aspen seedlings raised in controlled greenhouse conditions on two different soils. Growth among families varied significantly, and the interaction between family and soil type was significant. From 2–5 months, leaf number correlated with that of the first month and salinity tolerance. Sodium ions varied significantly within plants and among families; seedlings that accumulated higher Na+ concentrations in root had more leaves and lower Na+ in shoot. These results suggest that leaf number indicates salt tolerance in young seedlings. Seedling performance was also affected by soil type, especially the root/shoot ratio, suggesting an interaction between salt tolerance and growth medium. This study has identified significant intra-specific variation in salt tolerance of aspen in 160 mM saline and highlighted the potential to select and develop a method for efficient pre-screening of trees to be used in the reclamation of salt-affected land.
  • Vaario, Finnish Forest Research Institute, Vantaa Research Unit, P. O. Box 18, FI-01301 Vantaa, Finland E-mail: lu-min.vaario@metla.fi (email)
  • Yrjälä, MEM group, Department of Biosciences, P.O. Box 56, FI-00014 University of Helsinki, Finland E-mail: ky@nn.fi
  • Rousi, Finnish Forest Research Institute, Vantaa Research Unit, P. O. Box 18, FI-01301 Vantaa, Finland E-mail: matti.rousi@metla.fi
  • Sipilä, Department of Agricultural Sciences, P.O. Box 56, FI-00014 University of Helsinki, Finland E-mail: ts@nn.fi
  • Pulkkinen, Pulkkinen, Finnish Forest Research Institute, Haapastensyrjä Research Unit, Haapastensyrjäntie 34, FI-12600 Läyliäinen, Finland E-mail: pertti.pulkkinen@metla.fi
article id 452, category Research article
Catherine Ky-Dembele, Mulualem Tigabu, Jules Bayala, Patrice Savadogo, Issaka Joseph Boussim, Per Christer Odén. (2010). Clonal propagation of Detarium microcarpum from root cuttings. Silva Fennica vol. 44 no. 5 article id 452. https://doi.org/10.14214/sf.452
Keywords: Burkina Faso; sprouting efficiency; rootling; vegetative propagation
Abstract | View details | Full text in PDF | Author Info
Detarium microcarpum is a valuable tree species for fuelwood, timber, food and medicine in sub-Saharan Africa. However, its population is dwindling due to overexploitation, its seedlings’ low survival rate and slow growth. Vegetative propagation might enhance both survival and growth, but to date a successful clonal method does not exist for D. microcarpum. We conducted four experiments to examine the effects of propagation environment (high versus low humidity), cutting length and diameter, alignment of root segments (horizontal versus vertical) and distance from the root collar of donors on the regeneration ability of root segments collected from field-grown D. microcarpum trees in Burkina Faso. The size of root segments significantly affected their regeneration ability, while alignment had no effect. Sprouting was possible from 10 and 20-cm long segments of 15–60 mm diameter with 7–43% sprouting efficiency and multiple shoots while 5 cm long segments were unsuitable with 0–3% sprouting efficiency. Cuttings maintained at low humidity produced larger diameter sprouts than those in greenhouse. All cuttings showed strong polarity with most of the shoots developing at the proximal end. Rootlings from 20 cm root segments produced more new roots (0.62 ± 0.08 g) than those from 10 cm segments (0.34 ± 0.09 g), but they were similar for sprout and leaf growth. We conclude that lateral roots of field-grown mature trees can be used to produce rootlings in a nursery. Since this study is the first attempt to propagate D. microcarpum from root cuttings, further investigations are required to optimize the technique.
  • Ky-Dembele, Département Productions Forestières, Institut de l’Environnement et de Recherches Agricoles, 03 BP 7047 Ouagadougou 03, Burkina Faso & Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 101, SE-230 53 Alnarp, Sweden (catherine.dembele@ess.slu.se) E-mail: kydembele@hotmail.com (email)
  • Tigabu, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 101, SE-230 53, Alnarp, Sweden E-mail: mt@nn.se
  • Bayala, World Agroforestry Centre, West Africa and Centre Regional Office, Sahel Node, BP E5118 Bamako, Mali E-mail: jb@nn.ml
  • Savadogo, Département Productions Forestières, Institut de l’Environnement et de Recherches Agricoles, 03 BP 7047 Ouagadougou 03, Burkina Faso & Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 101, SE-230 53 Alnarp, Sweden E-mail: ps@nn.bf
  • Boussim, Université de Ouagadougou, Unité de Formation et Recherche en Sciences de la Vie et de la Terre, 03 BP 7021, Ouagadougou 03, Burkina Faso E-mail: ijb@nn.bf
  • Odén, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 101, SE-230 53, Alnarp, Sweden E-mail: pco@nn.se
article id 451, category Research article
Thomas P. Sullivan, Druscilla S. Sullivan, Pontus M. F. Lindgren, Douglas B. Ransome. (2010). Green-tree retention and life after the beetle: stand structure and small mammals 30 years after salvage harvesting. Silva Fennica vol. 44 no. 5 article id 451. https://doi.org/10.14214/sf.451
Keywords: biodiversity; stand structure; Pseudotsuga menziesii; ecological indicators; green-tree retention; small mammals; mountain pine beetle; Pinus contorta; salvage harvest
Abstract | View details | Full text in PDF | Author Info
We report on a retrospective investigation of the impacts of salvage harvesting of lodgepole pine (Pinus contorta var. latifolia Engelm. ex S. Wats.), killed by an outbreak of mountain pine beetle (Dendroctonus ponderosae Hopk.) in the 1970s, with variable retention of Douglas-fir (Pseudotsuga menziesii (Mirbel) Franco). Our inference to biodiversity was coniferous stand structure and four mammal species: the southern red-backed vole (Myodes gapperi Vigors), common shrew (Sorex cinereus Kerr), red squirrel (Tamiasciurus hudsonicus Erxleben) and northern flying squirrel (Glaucomys sabrinus Shaw). We tested hypotheses that, at 30 years after salvage harvest of beetle-killed lodgepole pine trees, (1) abundance and diversity of stand structure, and (2) abundance of mammal species, will increase with higher levels of green-tree retention (GTR). Stand structure attributes and small mammals were sampled during 2005–2008 in young pine stands, with a range of GTR seed-trees (none, dispersed, and aggregated Douglas-fir), and uncut forest in south-central British Columbia, Canada. Diameters and heights of Douglas-fir and lodgepole pine and basal area of total conifers supported hypothesis (1). Mean abundance of the red-backed vole was consistently higher (2.3 to 6.4 times) in the uncut forest than other stands. Overall mean patterns of abundance for common shrews, red squirrels, and northern flying squirrels were similar among treatment stands. Mean abundance of the red-backed vole supported hypothesis (2), but numbers of the other three species did not. There is “life after the beetle” at 30 years after salvage harvesting, and this was enhanced by GTR.
  • Sullivan, Department of Forest Sciences, Faculty of Forestry, University of BC, 2424 Main Mall, Vancouver, BC, Canada V6T 1Z4 E-mail: tom.sullivan@ubc.ca (email)
  • Sullivan, Department of Forest Sciences, Faculty of Forestry, University of BC, 2424 Main Mall, Vancouver, BC, Canada V6T 1Z4 E-mail: dss@nn.ca
  • Lindgren, Applied Mammal Research Institute, 11010 Mitchell Avenue, Summerland, BC, Canada V0H 1Z8 E-mail: pmfl@nn.ca
  • Ransome, Applied Mammal Research Institute, 11010 Mitchell Avenue, Summerland, BC, Canada V0H 1Z8 E-mail: dbr@nn.ca
article id 127, category Research article
Mika Rekola, Annukka Valkeapää, Tapio Rantala. (2010). Nordic forest professionals’ values. Silva Fennica vol. 44 no. 5 article id 127. https://doi.org/10.14214/sf.127
Keywords: forest professionals; values; cognitive mapping; Nordic countries
Abstract | View details | Full text in PDF | Author Info
The present study analyses the values held by forest professionals in three Nordic countries: Finland, Norway, and Sweden. The data is from a large (n = 1113) internet survey that used cognitive mapping as a research tool, which is a novelty in value measurement. The questionnaire is based on the organisational value theory of Schein (1992), supplemented with relevant forest-related and environmental values. The forest-related main value factors were in the following order of importance: Expertise, Private forestry, Forest production, Nature conservation, and Tradition. The measurement included two kinds of cases: action values, referring to present decision-making, and ideal values, referring to decisions concerning future ideals. Most of the values’ scores were similar. Almost all values received higher scores of importance in the ideal cases compared to action cases, a fact that can probably be explained by constraints related to the professionals’ current working environment. Some international differences were also found: Sweden and Norway were closer to each other and both differed from Finland, where private forestry, forest production, and traditions are highly valued. Moreover, respondents working in industry were found to be slightly more production-oriented than other forest professionals. The study also revealed several weaknesses of the cognitive mapping method in measuring values.
  • Rekola, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki E-mail: mika.rekola@helsinki.fi (email)
  • Valkeapää, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki E-mail: av@nn.fi
  • Rantala, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki E-mail: tapio.rantala@helsinki.fi
article id 126, category Research article
Anders Roos, Lotta Woxblom, Denise McCluskey. (2010). The influence of architects and structural engineers on timber in construction – perceptions and roles. Silva Fennica vol. 44 no. 5 article id 126. https://doi.org/10.14214/sf.126
Keywords: timber-frame building; qualitative analysis; specifiers; stakeholders; timber design
Abstract | View details | Full text in PDF | Author Info
This study considers structural engineers’ and architects’ perceptions of structural timber in multi-story construction contexts. Qualitative approaches – interviews and focus groups – were used to investigate attitudes, perceived norms, and perceived factors that hamper or facilitate the prescription of wood use in construction. Wood was perceived as an appropriate building material. Architects’, and even more so engineers’, perceptions of negative aspects of wood focused on decay, instability and sound transmission. Although wood-based construction was seen as a required professional skill, it was not expected to improve one’s professional status. Positive aspects of wood in construction included its strength, environmental friendliness, simple handling and appropriateness for use in conjunction with industrial methods, whereas knowledge gaps and weak support from the wood industry have reduced the use of wood among structural engineers and architects. Both professions perceived their influence on material selection to be weak. They sensed that most of the influence over material selection rested with developers and contractors. The paper contains suggestions on how to make these two professions more influential advocates for wood in construction.
  • Roos, Swedish University of Agricultural Sciences, Department of Forest Products, P.O. Box 7008, SE-750 07 Uppsala, Sweden E-mail: anders.roos@sprod.slu.se (email)
  • Woxblom, Swedish University of Agricultural Sciences, Department of Forest Products, P.O. Box 7008, SE-750 07 Uppsala, Sweden E-mail: lw@nn.se
  • McCluskey, Swedish University of Agricultural Sciences, Department of Forest Products, P.O. Box 7008, SE-750 07 Uppsala, Sweden E-mail: dm@nn.se
article id 125, category Research article
Juho Rantala, Tiina Laine. (2010). Productivity of the M-Planter tree-planting device in practice. Silva Fennica vol. 44 no. 5 article id 125. https://doi.org/10.14214/sf.125
Keywords: silviculture; cost-efficiency; mechanisation; planting; seedling; technology
Abstract | View details | Full text in PDF | Author Info
Need to mechanise tree-planting work have recently increased for many reasons. The newest planting and soil scarification device performing work in Nordic forests is the Finnish M-Planter. This study aims to clarify M-Planter’s productivity in practice and show how various factors affect it. The follow-up data set covers 607 work shifts, of 13 operators with, in total, five M-Planters. The average productivity figures for the operators were 143 and 169 seedlings per effective working hour during the first and second planting season, respectively. Overall, the measured average productivity was 34.2% lower than that observed in an earlier work study of the M-Planter based on an experimental study design. On average, the operators learned to use the combination of the M-Planter and a base machine more efficiently while their experience in using it increased during the follow-up. Increasing number of stones and stumps as well as a thicker humus layer decreased productivity of the M-Planter. The study concludes that utilisation of the full productivity potential of the M-Planter requires not only good operators but also development of the whole planting service supply chain.
  • Rantala, Finnish Forest Research Institute, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: juho.rantala@metla.fi (email)
  • Laine, Finnish Forest Research Institute, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: tl@nn.fi
article id 124, category Research article
Johannes Windisch, Lauri Sikanen, Dominik Röser, David Gritten. (2010). Supply chain management applications for forest fuel procurement – cost or benefit? Silva Fennica vol. 44 no. 5 article id 124. https://doi.org/10.14214/sf.124
Keywords: logistics; forest fuel; supply chain management; cost-benefit analysis
Abstract | View details | Full text in PDF | Author Info
It is commonly agreed that logistics is very demanding in forest fuel business. Even though logistics and supply chain management (SCM) tools already have found their way into forestry business, for example, in roundwood operations, they are not yet very widespread in the field of forest fuel procurement. The present study investigates if modern supply chain management applications are capable of increasing the profitability of forest fuel procurement operations. Since margins are low, decreasing the provision costs could boost wood-based bioenergy business. The study is based on the investigation of two Finnish forest owners associations (FOA) involved in forest fuel procurement using a modern SCM tool. The investigation is done by cost-benefit analysis (CBA) using the net present value (NPV) methodology to determine the profitability. According to the estimates made by the staff, which are based on data such as work time records and delivery notes from before and after introduction of the new system, in both FOAs, the benefits far outweigh the costs over a considered timespan of ten years. However, the amount of the NPV varied significantly. For FOA1, with an annual chip production of 150 000 loose m3, the NPV is 212 739 euro, while for FOA2, with an annual chip production of 37 000 loose m3, the NPV is 969 841 euro. Even if the NPV of FOA2 seems to be very high, the profitability of SCM tools in forest fuel procurement is clearly demonstrated. Additionally, the results indicate that a considerable cost saving potential in forest fuel procurement is attainable through improving work flows and thus reduce the work input.
  • Windisch, Finnish Forest Research Institute, Joensuu Research Unit, Yliopistokatu 6, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: johannes.windisch@metla.fi (email)
  • Sikanen, University of Eastern Finland, School of Forest Science, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: ls@nn.fi
  • Röser, Finnish Forest Research Institute, Joensuu Research Unit, Yliopistokatu 6, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: dr@nn.fi
  • Gritten, University of Eastern Finland, School of Forest Science, Yliopistokatu 7, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: dg@nn.fi
article id 123, category Research article
Staffan Jacobson, Folke Pettersson. (2010). An assessment of different fertilization regimes in three boreal coniferous stands. Silva Fennica vol. 44 no. 5 article id 123. https://doi.org/10.14214/sf.123
Keywords: Pinus sylvestris; boreal forests; Picea abies; biomass yield; carbon sequestration; fertilization regimes; foliar analysis
Abstract | View details | Full text in PDF | Author Info
In 1981–82 three field experiments were established with the aim of elucidating (i) the growth response of middle-aged coniferous stands at different fertilization intensities and, hence, the economic outcomes; and (ii) the need to add nutrients other than nitrogen (N). Nutrient additions were performed at intervals of two, four, six and eight years. The experiments were established on typical podzolized and N-limited mor-humus sites, two in Scots pine (Pinus sylvestris L.) stands and one in a Norway spruce (Picea abies (L.) Karst.) stand, at three different locations in Sweden. The ages of the stands were 65–70 years at the time of establishment. Growth responses were calculated after a 22-year study period. The growth responses were significant in all treatments. The addition of nutrients other than N did not affect stem growth at any of the sites. The growth response tended to increase with decreasing application interval. The results also revealed that the efficiency of fertilization is reduced as the interval between fertilizations is shortened. Accordingly, the growth effect per kg of added N was negatively correlated to fertilization intensity. The least intensive fertilization regime (an eight-year interval) resulted in an average net increase in C sequestration of 35 kg per kg N added. The profitability, in terms of internal rate of return, the present net value at different interest rates and the cost of production, i.e. the cost to produce one extra m3 under the different N regimes, are presented and discussed.
  • Jacobson, Skogforsk, Uppsala Science Park, SE-75183 Uppsala, Sweden E-mail: staffan.jacobson@skogforsk.se (email)
  • Pettersson, Skogforsk, Uppsala Science Park, SE-75183 Uppsala, Sweden E-mail: fp@nn.se
article id 122, category Research article
Benoit Lafleur, Nicole J. Fenton, David Paré, Martin Simard, Yves Bergeron. (2010). Contrasting effects of season and method of harvest on soil properties and the growth of black spruce regeneration in the boreal forested peatlands of eastern Canada. Silva Fennica vol. 44 no. 5 article id 122. https://doi.org/10.14214/sf.122
Keywords: soil disturbance; peatland; Picea mariana; careful logging; clearcutting; paludification; forest productivity
Abstract | View details | Full text in PDF | Author Info
It has been suggested that without sufficient soil disturbance, harvest in boreal forested peatlands may accelerate paludification and reduce forest productivity. The objectives of this study were to compare the effects of harvest methods (clearcutting vs. careful logging) and season (summer vs. winter harvest) on black spruce regeneration and growth in boreal forested peatlands of eastern Canada, and to identify the soil variables that favour tree growth following harvest. Moreover, we sought to determine how stand growth following harvest compared with that observed following fire. The average tree height of summer clearcuts was greater than that of summer carefully logged stands and that of all winter harvested sites. Summer clearcutting also resulted in a higher density of trees > 3 m and > 4 m tall and in a 50% reduction in Rhododendron groenlandicum cover, a species associated with reduced black spruce growth. Height growth of sample trees was related to foliar N and P concentrations, and to soil total N, pH and available Ca and Mg but not to harvest method or season. Our results also indicate that summer clearcutting could produce stand productivity levels comparable to those observed after high-severity soil burns. These results suggest that summer clearcutting could be used to restore forest productivity following harvest in forested peatlands, and offer further support to the idea that sufficient levels of soil disturbance may be required to restore productivity in ecosystems undergoing paludification.
  • Lafleur, NSERC-UQAT-UQAM Industrial Chair in Sustainable Forest Management, Université du Québec en Abitibi-Témiscamingue, 445 boul. de l’Université, Rouyn-Noranda, QC J9X 5E4, Canada E-mail: benoit.lafleur@uqat.ca (email)
  • Fenton, NSERC-UQAT-UQAM Industrial Chair in Sustainable Forest Management, Université du Québec en Abitibi-Témiscamingue, 445 boul. de l’Université, Rouyn-Noranda, QC J9X 5E4, Canada E-mail: njf@nn.ca
  • Paré, Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, 1055 du P.E.P.S., P.O. Box 10380, Stn. Sainte-Foy, QC G1V 4C7, Canada E-mail: dp@nn.ca
  • Simard, Département de Géographie, Université Laval, Pavillon Abitibi-Price, 2405 rue de la Terrasse, Québec, QC G1V 0A6, Canada E-mail: ms@nn.ca
  • Bergeron, NSERC-UQAT-UQAM Industrial Chair in Sustainable Forest Management, Université du Québec en Abitibi-Témiscamingue, 445 boul. de l’Université, Rouyn-Noranda, QC J9X 5E4, Canada E-mail: yb@nn.ca
article id 121, category Research article
Catherine Ky-Dembele, Jules Bayala, Patrice Savadogo, Mulualem Tigabu, Per Christer Odén, Issaka Joseph Boussim. (2010). Comparison of growth responses of Khaya senegalensis seedlings and stecklings to four irrigation regimes. Silva Fennica vol. 44 no. 5 article id 121. https://doi.org/10.14214/sf.121
Keywords: rooted cuttings; water stress; Senegal mahogany
Abstract | View details | Full text in PDF | Author Info
Khaya senegalensis is an important tree species for timber production, native to West Africa, but mahogany shoot borer attacks prevent successful plantations. This research was aimed at comparing the growth of two propagule types, seedlings and stecklings, of Khaya senegalensis subjected to four irrigation regimes, 25, 50, 75 and 100% field capacity in Burkina Faso. The relative growth rate, biomass allocation and intrinsic water use efficiency of the propagules were assessed in a full-factorial pot experiment in block design. Except the relative growth rate of stem basal diameter and specific leaf area, for which mean values were significantly higher for seedlings than stecklings, the two propagule types had similar growth patterns regarding relative growth rates of stem length, leaf, stem, root and the total plant biomass. There was no significant difference between propagule types concerning biomass fraction to total plant biomass of leaf, stem and root, root to stem ratio, leaf area productivity and carbon isotope ratio (δ13C). However, the irrigation regimes significantly affected all parameters. In contrast to 75 and 100% field capacity irrigation regimes, the low water supply of 25 and 50% field capacity resulted in plant stress, which was evident from the significant reduction in plant growth and biomass production and an increase in the root biomass to total plant biomass ratio and δ13C. It can be concluded that seedlings and stecklings have comparable growth patterns, while water stress is a major growth-limiting factor highlighting the need for selecting drought and borer resistant genotypes for successful plantations.
  • Ky-Dembele, Département Productions Forestières, Institut de l’Environnement et de Recherches Agricoles, 03 BP 7047 Ouagadougou 03, Burkina Faso & Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 101, SE-230 53 Alnarp, Sweden (catherine.dembele@slu.se) E-mail: kydembele@hotmail.com (email)
  • Bayala, World Agroforestry Centre, West Africa and Centre Regional Office, Sahel Node, BP E5118 Bamako, Mali E-mail: jb@nn.ml
  • Savadogo, Département Productions Forestières, Institut de l’Environnement et de Recherches Agricoles, 03 BP 7047 Ouagadougou 03, Burkina Faso & Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 101, SE-230 53 Alnarp, Sweden E-mail: ps@nn.bf
  • Tigabu, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 101, SE-230 53, Alnarp, Sweden E-mail: mt@nn.se
  • Odén, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 101, SE-230 53, Alnarp, Sweden E-mail: pco@nn.se
  • Boussim, Université de Ouagadougou, Unité de Formation et Recherche en Sciences de la Vie et de la Terre, 03 BP 7021, Ouagadougou 03, Burkina Faso E-mail: ijb@nn.bf
article id 120, category Research article
Antti J. Lukkarinen, Seppo Ruotsalainen, Teijo Nikkanen, Heli Peltola. (2010). Survival, height growth and damages of Siberian (Larix sibirica Ledeb.) and Dahurian (Larix gmelinii Rupr.) larch provenances in field trials located in southern and northern Finland. Silva Fennica vol. 44 no. 5 article id 120. https://doi.org/10.14214/sf.120
Keywords: height growth; survival; larch; Larix; provenance; seedling damage
Abstract | View details | Full text in PDF | Author Info
The aim of this study was to analyse differences in the survival and height growth of, as well as damages to Siberian (Larix sibirica Ledeb.) and Dahurian (Larix gmelinii Rupr.) larch provenances over four growing seasons in field trials established in 2006 in southern (Punkaharju) and northern Finland (Kivalo). In this context, the study also investigated if the geographical and climatic conditions of the origin of the provenance could explain the differences between the provenances. The study material consisted of 20 Russian Siberian and Dahurian larch provenances and five seed sources from Finland (4) and Russia (1) as comparison lots. It was found that the Finnish seed sources of Siberian larch survived well in both the Kivalo and Punkaharju trials. Five northern latitude Russian provenances, of which one was Dahurian and the remainder were Siberian larches, had the highest survival in Kivalo. However, the differences observed in survival between provenances were only significant (p < 0.05) in Kivalo. Regardless of the trial, the differences, however, in height growth were significant and large between provenances. The southern Dahurian larches had a superior height growth in Punkaharju. The northern Dahurian larch provenance from Magadan (59°50′N, 150°40′E) had the largest height growth in Kivalo, among some northern Siberian larches. Damages were diverse, though Dahurian larches had less mammal damage than the Siberian larches. In general, the differences between provenances were not significant. Latitude and altitude best explained the differences between provenances, but also mean temperature, temperature sum and continentality index affected them (p < 0.05).
  • Lukkarinen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: antti.lukkarinen@pp.inet.fi (email)
  • Ruotsalainen, Finnish Forest Research Institute, FI-58450 Punkaharju, Finland E-mail: sn@nn.fi
  • Nikkanen, Finnish Forest Research Institute, FI-58450 Punkaharju, Finland E-mail: tn@nn.fi
  • Peltola, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: hp@nn.fi
article id 111, category Research article
Ilona Pietilä, Annika Kangas, Antti Mäkinen, Lauri Mehtätalo. (2010). Influence of growth prediction errors on the expected losses from forest decisions. Silva Fennica vol. 44 no. 5 article id 111. https://doi.org/10.14214/sf.111
Keywords: growth prediction; uncertainty; forest information; updating; inoptimality loss
Abstract | View details | Full text in PDF | Author Info
In forest planning, forest inventory information is used for predicting future development of forests under different treatments. Model predictions always include some errors, which can lead to sub-optimal decisions and economic loss. The influence of growth prediction errors on the reliability of projected forest variables and on the treatment propositions have previously been examined in a few studies, but economic losses due to growth prediction errors is an almost unexplored subject. The aim of this study was to examine how the growth prediction errors affected the expected losses caused by incorrect harvest decisions, when the inventory interval increased. The growth models applied in the analysis were stand-level growth models for basal area and dominant height. The focus was entirely on the effects of growth prediction errors, other sources of uncertainty being ignored. The results show that inoptimality losses increased with the inventory interval. Average relative inoptimality loss was 3.3% when the inventory interval was 5 years and 11.6% when it was 60 years. Average absolute inoptimality loss was 230 euro ha–1 when the inventory interval was 5 years and 860 euro ha–1 when it was 60 years. The average inoptimality losses varied between development classes, site classes and main tree species.
  • Pietilä, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ip@nn.fi
  • Kangas, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: annika.kangas@helsinki.fi (email)
  • Mäkinen, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: am@nn.fi
  • Mehtätalo, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: lm@nn.fi
article id 454, category Research article
Paula Jylhä, Olli Dahl, Juha Laitila, Kalle Kärhä. (2010). The effect of supply system on the wood paying capability of a kraft pulp mill using Scots pine harvested from first thinnings. Silva Fennica vol. 44 no. 4 article id 454. https://doi.org/10.14214/sf.454
Keywords: whole-tree harvesting; cut-to-length harvesting; integrated wood harvesting; residual value; whole-tree bundling
Abstract | View details | Full text in PDF | Author Info
The efficiencies of wood supply systems based on cut-to-length (CTL) harvesting, the harvesting of loose whole trees, and whole-tree bundling were compared using the relative wood paying capabilities (WPC) of a kraft pulp mill as decisive criteria. The WPCs from mill to stump were calculated for three first-thinning stands of Scots pine (Pinus sylvestris L.) with mean breast-height diameter of the removal of 6, 8, and 12 cm. Pulp price had a strong effect on the WPC, and the CTL system resulted in the highest WPC per m3 at stump. The savings in procurement costs and gains in energy generation from additional raw material acquired with the harvesting of loose whole trees did not compensate the losses in pulp production. Considering removal per hectare, loose whole trees gave the highest WPCs at stump in the two stands with the smallest trees and the highest proportion of additional raw material. Decrease in pulp price and increase in energy price improved the competitiveness of the whole-tree systems. In the case of whole-tree bundling, savings in transportation costs did not balance the high cutting and compaction costs, and the bundling system was the least competitive alternative.
  • Jylhä, Finnish Forest Research Institute, Kannus, Finland E-mail: paula.jylha@metla.fi (email)
  • Dahl, Aalto University School of Science and Technology, Department of Forest Products Technology, Espoo, Finland E-mail: od@nn.fi
  • Laitila, Finnish Forest Research Institute, Joensuu, Finland E-mail: jl@nn.fi
  • Kärhä, Metsäteho Oy, Helsinki, Finland E-mail: kk@nn.fi
article id 453, category Research article
Eduard Schiessl, Michael Grabner, Gerald Golesch, Thomas Geburek, Silvio Schueler. (2010). Sub-montane Norway spruce as alternative seed source for a changing climate? A genetic and growth analysis at the fringe of its natural range in Austria. Silva Fennica vol. 44 no. 4 article id 453. https://doi.org/10.14214/sf.453
Keywords: Picea abies; climate-growth response; crown type; rear edge; mitochondrial DNA
Abstract | View details | Full text in PDF | Author Info
Insights into the intraspecific variation in climate response of forest trees and the utilization of suitable seed sources are required to maintain forest ecoystems under expected climate change. Individuals of Norway spruce with an anomalous spherical-shaped crown were characterized by genetic (using a mtDNA marker) and dendroclimatic analysis. Such trees occur frequently at the fringe of the spruce distribution in east/south-east Austria. We employed pair-wise comparisons between trees with spherical and “regular” conical crowns on 47 sites. No evidence was found for a different phylogeographic history of spherical and conical spruces, but the high allelic diversity at the nad1 locus highlighted the importance of east/south-east Austria as refugium and migration corridor for Norway spruce. Analysis of mean annual increment revealed a larger amount of earlywood within the sapwood area and fewer negative pointer years for spherical spruces than for conical ones, pointing at a higher vitality and smaller interactions between climate and growth for spherical spruces. Although the results cannot explain the anomalous crown form, they suggest spherical trees to have a higher ability to cope the warm and dry climate of the region than “regular” conical spruces. We discuss the origin of spherical crowns in terms of population history and phenotypic plasticity and speculate on possible effects of crown architecture on canopy-atmosphere exchange.
  • Schiessl, Department of Genetics, Research and Training Centre for Forests, Natural Hazards and Landscape, Hauptstrasse 7, A-1140 Vienna, Austria E-mail: es@nn.at
  • Grabner, Department of Material Sciences and Process Engineering, Institute of Wood Science and Technology, University of Natural Resources and Applied Life Sciences, Peter-Jordan-Strasse 82, A-1190 Vienna, Austria E-mail: mg@nn.at
  • Golesch, Department of Genetics, Research and Training Centre for Forests, Natural Hazards and Landscape, Hauptstrasse 7, A-1140 Vienna, Austria E-mail: gg@nn.at
  • Geburek, Department of Genetics, Research and Training Centre for Forests, Natural Hazards and Landscape, Hauptstrasse 7, A-1140 Vienna, Austria E-mail: tg@nn.at
  • Schueler, Department of Genetics, Research and Training Centre for Forests, Natural Hazards and Landscape, Hauptstrasse 7, A-1140 Vienna, Austria E-mail: silvio.schueler@bfw.gv.at (email)
article id 135, category Research article
Ola Lindroos, Marina Henningsson, Dimitris Athanassiadis, Tomas Nordfjell. (2010). Forces required to vertically uproot tree stumps. Silva Fennica vol. 44 no. 4 article id 135. https://doi.org/10.14214/sf.135
Keywords: forest fuels; Picea abies; bioenergy; stump harvesting; machine development; Betula spp.
Abstract | View details | Full text in PDF | Author Info
Stumpwood attracts renewed interest due to increased use of forest biomass for bioenergy. In Nordic countries stumps are generally uprooted with crawler excavators, which have strong cranes (ca. 400 kNm gross lift torque), but are not designed for moving in forest terrain. Their use is based on practical experience with available and tested machine types rather than thorough examinations of requirements, partly due to limited knowledge of force requirements for uprooting of stumps. Therefore, in this work mean and maximum forces required to vertically uproot stumps of Norway spruce (Picea abies) and birch (Betula spp.) were quantified together with the effects of various soil types and uprooting methods. The used excavator’s crane-mounted uprooting device enabled comparisons between usage of solely crane force, and a method in which preparatory loosening forces were applied prior to crane force. Uprooting stumps in single pieces proved difficult; 61% split unintentionally. Force requirements were similar across tree species, increasing curve-linearly with stump diameter, and stumps uprooted in a single piece required more force than split stumps. Preparatory loosening reduced crane force requirements and, surprisingly, less force was required to uproot stumps from a mesic, till soil than from a moist, finer-textured soil. No stump required more than 60 kN crane force and functions for maximum force requirements indicate that powerful harvesters and forwarders (gross crane lifting capacity of 273 and 155 kNm, respectively) should be able to uproot all stumps with ≤ 61 cm and ≤ 32 cm diameter, respectively, in one piece. Larger stumps could be managed if it is acceptable that stumps are split before uprooting.
  • Lindroos, Department of Forest Resource Management, Swedish University of Agricultural Sciences, SE-901 83 Umeå, Sweden E-mail: ola.lindroos@srh.slu.se (email)
  • Henningsson, Komatsu Forest AB, Box 7124, SE-907 04 Umeå, Sweden E-mail: mh@nn.se
  • Athanassiadis, Department of Forest Resource Management, Swedish University of Agricultural Sciences, SE-901 83 Umeå, Sweden E-mail: da@nn.se
  • Nordfjell, Department of Forest Resource Management, Swedish University of Agricultural Sciences, SE-901 83 Umeå, Sweden E-mail: tn@nn.se
article id 134, category Research article
Dan Bergström, Urban Bergsten, Tomas Nordfjell. (2010). Comparison of boom-corridor thinning and thinning from below harvesting methods in young dense Scots pine stands. Silva Fennica vol. 44 no. 4 article id 134. https://doi.org/10.14214/sf.134
Keywords: bioenergy; comparative time studies; energy wood; geometric thinning; pre-commercial thinnings; systematic thinnings
Abstract | View details | Full text in PDF | Author Info
At present, only a small proportion of the potential extractable bioenergy from young dense forests in Sweden is utilized. The conventional mechanized first thinning systems used in such stands suffer from low productivity, so the operation is only profitable in stands with bigger trees and high standing volumes. Conventional harvesters are used for this operation equipped with accumulating felling heads designed for handling several trees during each crane cycle. In thinning from below the felling and bunching work requires many time-consuming non-linear crane movements to avoid felling or damaging of future crop trees. However, higher productivity can be achieved when trees between strip roads are harvested in about 1 m-wide corridors with a length corresponding to the reach of the crane. We refer to this operation as boom-corridor thinning. The objective of this study was to compare felling and bunching productivity in young dense stands when employing thinning from below or boom-corridor thinning. Experiments were performed using a randomized block design involving between 4400 and 18 600 trees x ha-1 with a corresponding average tree size of 7.2 and 3.2 cm dbh, respectively. Based on the average tree being removed at a dbh of 5.7 cm, the productivity (ODt x PW-hour-1) was significant (almost 16%) higher for the boom-corridor thinning than for thinning from below treatment. At the same time, the time taken for the work element “Crane in-between” (the period between the loaded crane starting to move towards a tree and the felling head rapidly slowing down for positioning) was significantly reduced, by almost 17%. The positive results were achieved even though the operator was new to the method. To achieve a significantly higher efficiency during the felling and bunching operation, development of new harvesting equipment and operating techniques seems crucial.
  • Bergström, Swedish University of Agricultural Sciences, Dept of Forest Resource Management, Section of Planning and Operations Management, Umeå, Sweden E-mail: dan.bergstrom@srh.slu.se (email)
  • Bergsten, Swedish University of Agricultural Sciences, Dept of Forest Resource Management, Section of Planning and Operations Management, Umeå, Sweden E-mail: ub@nn.se
  • Nordfjell, Swedish University of Agricultural Sciences, Dept of Forest Resource Management, Section of Planning and Operations Management, Umeå, Sweden E-mail: tn@nn.se
article id 133, category Research article
Fumiaki Kitahara, Nobuya Mizoue, Shigejiro Yoshida. (2010). Effects of training for inexperienced surveyors on data quality of tree diameter and height measurements. Silva Fennica vol. 44 no. 4 article id 133. https://doi.org/10.14214/sf.133
Keywords: diameter tape; measurement error; National Forest Inventory; training; Vertex III hypsometer
Abstract | View details | Full text in PDF | Author Info
Due to the large number of sample plots and variables to be measured, inexperienced surveyors are expected to take field measurements in National Forest Inventories (NFIs). However, very little information exists on the data quality that can be expected from inexperienced surveyors given different levels of training. We evaluated the quality of data produced by inexperienced undergraduate students when measuring the most fundamental variables: tree diameter using a diameter tape and height using an ultrasonic Vertex III hypsometer. We found that a single training session on how to use the instruments and how to reduce measurement errors was insufficient for inexperienced surveyors to achieve measurement quality objectives (MQOs). Providing a single feedback of control team measurements significantly improved data quality, except in the measurements of tree height of broad-leaved trees, but additional feedback did not contribute to further improvement. We propose that field training courses for inexperienced surveyors incorporate a one-day exercise with feedback instruction.
  • Kitahara, Graduate School of Bioresource and Bioenvironmental Science, Kyushu University, 6-10-1 Hakozaki, Higashiku, Fukuoka 812-8581, Japan E-mail: bunsho@ffpri.affrc.go.jp (email)
  • Mizoue, Faculty of Agriculture, Kyushu University, Fukuoka, Japan E-mail: nm@nn.jp
  • Yoshida, Faculty of Agriculture, Kyushu University, Fukuoka, Japan E-mail: sy@nn.jp
article id 132, category Research article
Mats Berlin, Lars Lönnstedt, Gunnar Jansson, Öje Danell, Tore Ericsson. (2010). Developing a Scots pine breeding objective: a case study involving a Swedish sawmill. Silva Fennica vol. 44 no. 4 article id 132. https://doi.org/10.14214/sf.132
Keywords: Pinus sylvestris; economic weights; bio-economic model; tree breeding; multi-trait selection
Abstract | View details | Full text in PDF | Author Info
The aim of this study was to develop a Scots pine (Pinus sylvestris L.) breeding objective for a vertically integrated sawmill in northern Sweden. The production system was defined as comprising the sawmill and the forests supplying it. Volume per hectare, wood density, survival and straightness were used as objective traits and the related selection criteria were measurements, collected at young tree age, of height, diameter, pilodyn penetration, vitality and straightness. A bio-economic model was used to calculate economic weights for the objective traits identified. We also investigated the efficiency of different selection indices based on these economic weights, in combination with available data on genetic parameters. Furthermore, we studied the effect of different discount rates on the calculated economic weights. The results showed that, compared to the full index (which included all selection criteria), omitting either vitality or straightness had a negligible effect, reducing predicted profit gain per hectare by less than one per cent. Height or diameter each had a greater effect, with a loss of predicted profit gain per hectare of up to 6%. Excluding pilodyn penetration from the selection index caused the largest reduction in predicted profit gain per hectare, amounting to over 10%. However, when both height and diameter were removed the predicted profit gain per hectare dropped to one-third of that based on the full index. Finally, ranking and genetic selection for the developed breeding objective was insensitive to changes in the discount rate.
  • Berlin, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: mats.berlin@skogsforsk.se (email)
  • Lönnstedt, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: ll@nn.se
  • Jansson, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: gj@nn.se
  • Danell, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: od@nn.se
  • Ericsson, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: te@nn.se
article id 131, category Research article
Timo Pukkala, Tatu Hokkanen, Teijo Nikkanen. (2010). Prediction models for the annual seed crop of Norway spruce and Scots pine in Finland. Silva Fennica vol. 44 no. 4 article id 131. https://doi.org/10.14214/sf.131
Keywords: flowering; cone crops; seed years; mixed model
Abstract | View details | Full text in PDF | Author Info
Many studies indicate that the flowering abundance of boreal trees strongly correlates with the weather conditions of the previous summer. This study developed prediction models for the seed crops of Norway spruce and Scots pine using weather variables one and two years prior to flowering year as predictors. Weather data, systematically recorded at many weather stations, were obtained from the Finnish Meteorological Institute. Seed crop monitoring data came from 22 spruce stands and 44 pine stands. In every stand, seed crop has been monitored for many years, the longest continuous period being 45 years. Monthly mean temperatures, monthly rainfalls, and periodical temperature sums were used as predictors in the seed crop models. Generally, both tree species flowered abundantly one year after a warm summer and two years after a cool summer. While the models only explained about 45% of the variation in the annual seed crop, they accurately predicted good and bad seed years: when the models predicted good seed crops the likelihood to have at least a medium seed crop was very high and when the models predicted small seed crops, the likelihood to obtain medium or good seed crop was very low. Therefore, the models reliably predict if a particular year will be a good seed year or a poor seed year. These predictions can be used in forestry practice for proper timing of natural regeneration activities, and when activities in seed orchards are planned.
  • Pukkala, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: timo.pukkala@uef.fi (email)
  • Hokkanen, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: th@nn.fi
  • Nikkanen, Finnish Forest Research Institute, Finlandiantie 18, FI-58450 Punkaharju, Finland E-mail: tn@nn.fi
article id 130, category Research article
Yongqing Yang, Yinan Yao, Xuejiang Zhang. (2010). Comparison of growth and physiological responses to severe drought between two altitudinal Hippophae rhamnoides populations. Silva Fennica vol. 44 no. 4 article id 130. https://doi.org/10.14214/sf.130
Keywords: sea buckthorn (Hippophae rhamnoides L.); physiological response; abscisic acid (ABA); carbon isotope composition
Abstract | View details | Full text in PDF | Author Info
Growth and physiological differences in response to drought were compared between two sea buckthorn (Hippophae rhamnoides L.) populations inhabited in the southeast of the Qinghai-Tibetan Plateau of China. The experimental design included two water regimes (100% and 25% of field capacity) and two populations from the low and high altitude zone. Our experiments were conducted in a naturally lit greenhouse under semi-controlled environmental conditions for a whole growing season in a dry valley (1800 m above the sea level). We found that drought tolerance is highly related to the plant antioxidant capacity and water use efficiency as well as leaf nutrient status in H. rhamnoides. The highland population (HP) experienced a greater inhibition in plant growth and leaf enlargement, lower leaf nitrogen and phosphorus content, lower root nodule biomass and root mass/foliage area ratio, and higher leaf water content loss paralleling with higher enhancement of abscisic acid level in response to drought, as compared with lowland population (LP). Additionally, reduction of leaf lignin content in HP further reduced its drought tolerance. On the contrary, LP showed effective adaptation strategies such as improvement of water economy and maintaining high ascorbic acid content. Therefore, we conclude that LP was more tolerant to drought than HP, and could be selected for reforestation in the dry valleys of upper Minjiang River regions in China.
  • Yang, Xinjiang Institute of Ecology and Geography, Chinese Academy of Sciences, Urumqi, 830011, China & College of Life Sciences, Chongqing Normal University, Chongqing, 400047, China E-mail: yy@nn.cn
  • Yao, Xinjiang Institute of Ecology and Geography, Chinese Academy of Sciences, Urumqi, 830011, China E-mail: yaoya@ms.xjb.ac.cn (email)
  • Zhang, Institute for Plant Protection and Soil sciences, Hubei Academy of Agricultural Sciences, Wuhan, 430064, China E-mail: xz@nn.cn
article id 129, category Research article
Mikko Moilanen, Markku Saarinen, Klaus Silfverberg. (2010). Foliar nitrogen, phosphorus and potassium concentrations of Scots pine in drained mires in Finland. Silva Fennica vol. 44 no. 4 article id 129. https://doi.org/10.14214/sf.129
Keywords: peatland; foliar analysis; drainage; site type; nutrient deficiency
Abstract | View details | Full text in PDF | Author Info
An imbalanced nutrient status in Scots pine stands on drained mires is primarily a consequence of excess nitrogen (N) in relation to mineral nutrients such as phosphorus (P) and potassium (K). In this study, the variation of foliar N, P, and K concentrations relative to some site and environmental characteristics was examined. Foliar nutrient concentrations were determined on needle samples collected from mires representing different drainage ages, site types, geographical locations and annual weather conditions. In the overall data (n = 971 samples in 333 stands) the foliar N concentration varied between 6.7 and 24.2 mg g-1, the P concentration between 0.83 and 2.32 mg g-1, and the K concentration between 2.22 and 6.23 mg g-1. The original (pre-drainage) mire site type proved to be an important factor in explaining the nutrient status of the trees: on originally forested sites, the nutrient balance (N versus K; N versus P) was mostly adequate, whereas on sparsely forested and treeless sites, K deficiency was common. N deficiency was the most common in forested ‘nitrogen-poor’ sites, while P and K deficiencies were more common in originally treeless or sparsely forested ‘nitrogen rich’ sites, where the nutrient imbalance was also the greatest. Over the whole data, 29% of the cases were diagnosed to be N-deficient, 51% P-deficient, and 25% K-deficient. The foliar N concentration increased with increasing temperature sum. The foliar K concentration decreased with increasing depth of the peat layer. On former treeless or sparsely forested sites, foliar K decreased slightly with increasing drainage age. In contrast, on thin-peated sites the foliar P concentration increased with increasing drainage age. The climate conditions (location), the original site type of the mire and peat thickness should be taken into account when planning silvicultural measures on mires drained for forestry.
  • Moilanen, Finnish Forest Research Institute, Muhos and Parkano Research Units, Finland E-mail: mikko.moilanen@metla.fi (email)
  • Saarinen, Finnish Forest Research Institute, Muhos and Parkano Research Units, Finland E-mail: ms@nn.fi
  • Silfverberg, Finnish Forest Research Institute, Muhos and Parkano Research Units, Finland E-mail: ks@nn.fi
article id 128, category Research article
Shelley L. Hunt, Andrew M. Gordon, Dave M. Morris. (2010). Carbon stocks in managed conifer forests in northern Ontario, Canada. Silva Fennica vol. 44 no. 4 article id 128. https://doi.org/10.14214/sf.128
Keywords: forest management; boreal forest; carbon sequestration; carbon stocks; jack pine; black spruce; plantations
Abstract | View details | Full text in PDF | Author Info
Carbon pools and net primary productivity (aboveground) were measured in managed stands of jack pine (Pinus banksiana Lamb.) and black spruce (Picea mariana [Mill.] B.S.P.), ranging in age from 10 to 53 years, in the Lake Nipigon area of northern Ontario. Organic carbon in the forest floor and surface mineral soil (top 15 cm) ranged from 13 to 46 Mg C ha-1 and 10 to 29 Mg C ha-1, respectively. Carbon in aboveground tree biomass ranged from 11 to 74 Mg C ha-1 in crop trees, and 0 to 11 Mg C ha-1 in non-crop trees. Coarse woody debris (downed woody debris and snags) contained between 1 and 17 Mg C ha-1. Understory vegetation rarely represented more than 1% of total ecosystem carbon accumulation, but was responsible for a larger proportion of aboveground net primary productivity (ANPP). Rates of ANPP (expressed as carbon) ranged from 0.8 to 3.5 Mg C ha-1 y-1. Carbon stocks in managed stands were compared with published values from similarly aged fire-origin stands in the North American boreal region. Carbon stocks in our study stands generally exceeded those in unmanaged fire-origin stands of the same age, due to larger tree and forest floor carbon pools.
  • Hunt, University of Guelph, School of Environmental Sciences, 50 Stone Road East, Guelph, Ontario, Canada N1G 2W1 E-mail: shunt@uoguelph.ca (email)
  • Gordon, University of Guelph, School of Environmental Sciences, 50 Stone Road East, Guelph, Ontario, Canada N1G 2W1 E-mail: amg@nn.ca
  • Morris, Ontario Ministry of Natural Resources, Centre for Northern Forest Ecosystem Research, 955 Oliver Rd., Thunder Bay, Ontario, Canada P7B 5E1 E-mail: dmm@nn.ca
article id 146, category Research article
Karri Uotila, Juho Rantala, Timo Saksa, Pertti Harstela. (2010). Effect of soil preparation method on economic result of Norway spruce regeneration chain. Silva Fennica vol. 44 no. 3 article id 146. https://doi.org/10.14214/sf.146
Keywords: Picea abies; soil preparation; cleaning; disc trenching; forestry investment analysis; pre-commercial thinning; regeneration chain; spot mounding; sprouting
Abstract | View details | Full text in PDF | Author Info
Economic result of forest regeneration chains, based either on spot mounding or on disc trenching and planting of Norway spruce (Picea abies [L.] Karst.) seedlings, were clarified and compared to each other. First, effects of soil preparation method on early development of Norway spruce stands were measured from field experiments. Second, the effects of soil preparation method on stand level management programs were modelled. The modelling was based on growth simulation and investment calculations. The soil preparation methods substantially affected early development of a stand. The density of the removed trees in early cleaning was 56% higher on the disc-trenched area compared to the spot-mounded area. The difference was especially high (120%), close by (< 25 cm) the remained spruce seedlings. There was also a difference between the methods in the growth of crop spruces; at biological age of 8 years, the mean height of spruce was 110 cm on the spot-mounded area and 68 cm on the disc-trenched area. The differences led to divergent management programs between the areas. The disc-trenched area needed three young stand management operations whereas two was enough at the spot-mounded area. Although disc trenching is a less expensive method than spot mounding, the total management costs were higher in disc trenching than in spot mounding. Furthermore, incomes from the first commercial thinning were higher when regeneration based on spot mounding. At the interest rate of 3%, the investment in spot mounding had 329 EUR ha–1 higher net present value than the investment in disc trenching.
  • Uotila, Finnish Forest Research Institute, Suonenjoki, Finland E-mail: karri.uotila@metla.fi (email)
  • Rantala, Finnish Forest Research Institute, Suonenjoki, Finland E-mail: jr@nn.fi
  • Saksa, Finnish Forest Research Institute, Suonenjoki, Finland E-mail: ts@nn.fi
  • Harstela, Finnish Forest Research Institute, Suonenjoki, Finland E-mail: ph@nn.fi
article id 145, category Research article
Jianbang Gan, C. T. Smith. (2010). Coupling greenhouse gas credits with biofuel production cost in determining conversion plant size. Silva Fennica vol. 44 no. 3 article id 145. https://doi.org/10.14214/sf.145
Keywords: bioethanol; production cost; carbon balance; feedstock supply radius; life-cycle analysis
Abstract | View details | Full text in PDF | Author Info
Biofuel plant size is one of the key variables in biofuel supply chain analysis as it plays a pivotal role in controlling the efficacy of both feedstock supply and feedstock-to-biofuel conversion. The unit production cost and greenhouse gas (GHG) balance of biofuels vary with plant size. We develop an analytical framework for integrating biofuel production costs and GHG balance derived from life-cycle analysis into supply chain optimization, followed by its application to ethanol production using forest biomass in the southern United States. We derive formulas for determining the optimal biofuel plant size and the corresponding feedstock supply radius based on the minimization of biofuel production costs less GHG benefits. Our results indicate that though biofuel plant size and feedstock supply radius should be augmented by considering GHG benefits, the GHG price will have a more significant impact on net biofuel production costs than on conversion plant size or feedstock supply radius. With a rise in the GHG price the net biofuel production cost tends to increase while the directions of change in plant size and feedstock supply radius are uncertain, depending upon the costs and GHG emissions of biomass transport and feedstock-to-fuel conversion. Combining GHG offset values with biofuel production costs enables us to more holistically examine the biofuel supply chain.
  • Gan, Department of Ecosystem Science and Management, Texas A&M University, Texas, USA E-mail: j-gan@tamu.edu (email)
  • Smith, Faculty of Forestry, University of Toronto, Ontario, Canada E-mail: cts@nn.ca
article id 144, category Research article
Markku Oikari, Kalle Kärhä, Teijo Palander, Heikki Pajuoja, Heikki Ovaskainen. (2010). Analyzing the views of wood harvesting professionals related to the approaches for increasing the cost-efficiency of wood harvesting from young stands. Silva Fennica vol. 44 no. 3 article id 144. https://doi.org/10.14214/sf.144
Keywords: cost-efficiency; energy wood; early thinnings; industrial roundwood; costs; gap analysis; opinion survey
Abstract | View details | Full text in PDF | Author Info
A lot of viable guidelines are currently available for more cost-effective harvesting of energy wood and industrial roundwood (i.e. pulpwood) from young stands. The study ranked the proposed potential approaches for increasing the cost-efficiency of small-diameter (d1.3 < 10 cm) energy wood and industrial roundwood harvesting from early thinnings. Research data, based on a total of 40 personal interviews, was collected in early 2008. The interviewees were divided into four wood harvesting professional groups: 1) Managers in wood procurement organizations, 2) Forest machine contractors, 3) Forest machine manufacturers and vendors, and 4) Wood harvesting researchers. In the opinion of the respondents, there is great potential to increase the cost-efficiency of wood harvesting through improving harvesting conditions (i.e. effective tending of seedling stands, delaying harvesting operations, and pre-clearance of dense undergrowth). The interviewees also underlined that harvesting methods can be rationalized, e.g. multiple-tree handling in industrial roundwood cuttings, crane scale measurement, integrated wood harvesting, and careful selection of stands for harvesting. The strong message given by the interviewees was that the education of forest machine operators must be made more effective in the future. There would be significant possibilities for cost savings in young stands, if methods and techniques with the most potential were utilized completely in wood harvesting.
  • Oikari, Karelwood, Kontiolahti, Finland E-mail: markku.oikari@karelwood.com (email)
  • Kärhä, Metsäteho Oy, Helsinki, Finland E-mail: kk@nn.fi
  • Palander, University of Eastern Finland, Faculty of Science and Forestry, Joensuu, Finland E-mail: tp@nn.fi
  • Pajuoja, Metsäteho Oy, Helsinki, Finland E-mail: hp@nn.fi
  • Ovaskainen, University of Eastern Finland, Faculty of Science and Forestry, Joensuu, Finland E-mail: ho@nn.fi
article id 143, category Research article
Juha Laitila, Jani Heikkilä, Perttu Anttila. (2010). Harvesting alternatives, accumulation and procurement cost of small-diameter thinning wood for fuel in Central Finland. Silva Fennica vol. 44 no. 3 article id 143. https://doi.org/10.14214/sf.143
Keywords: biomass resources; multi-tree-handling; delimbed stemwood; whole trees; Finland; early thinnings; forest chips
Abstract | View details | Full text in PDF | Author Info
This study compared harvesting alternatives, accumulation and procurement costs of small-diameter thinning wood chips for fuel, when trees were harvested either as delimbed stemwood or whole trees. The calculation was made for a hypothetical plant located in Central Finland and the radius of the procurement area was 100 km via the existing road network. Cutting was done with conventional harvester head equipped with multi-tree-handling (MTH) accessories, with the logged trees being chipped at the roadside storage. The cost of delimbed stemwood chips at heating plant was 24% higher compared to the cost of whole tree chips. The availability analysis attested that delimbing lowered the regional cutting removal by 42% compared to the whole tree harvesting, when the minimum accumulation for the fuel fraction at the stand was set at 25 m3/ha. Delimbing diminishes the recovery rate at the site, resulting in a diminishing number of potential recovery sites meeting the threshold volume. However, the study showed that the forest energy potential is increased and procurement costs are reduced, if delimbed stemwood is harvested from stands where the whole tree harvesting is not acceptable due to nutrient loss or for other ecological reasons. Intelligent selection of cutting methods for different stands enables minimization of transport distance and control of procurement cost.
  • Laitila, The Finnish Forest Research Institute, Joensuu, Finland E-mail: juha.laitila@metla.fi (email)
  • Heikkilä, L&T Biowatti Oy, Seinäjoki, Finland E-mail: jh@nn.fi
  • Anttila, The Finnish Forest Research Institute, Joensuu, Finland E-mail: pa@nn.fi
article id 142, category Research article
Helmer Belbo. (2010). Comparison of two working methods for small tree harvesting with a multi tree felling head mounted on farm tractor. Silva Fennica vol. 44 no. 3 article id 142. https://doi.org/10.14214/sf.142
Keywords: Norway spruce; cost functions; forest biomass harvesting; multi-tree handling; young stands
Abstract | View details | Full text in PDF | Author Info
In this study, the efficiency of a small multi-tree felling head, mounted on a farm tractor with a timber trailer was studied, when harvesting small trees for energy in thinnings. Both separate loading and direct loading of the felled trees was studied. Time studies were carried out in a mixed stand of Norway spruce (Picea abies (L.) Karst) and birch (Betula pubescens Ehrh.). The time consumption of the work elements in the different work methods was formulated by regression analysis, where the independent variables were tree size and degree of accumulation. The average size of the harvested trees was 0.035 m3. The time consumption for the harvesting and loading were similar for the two studied methods, 20 minutes per m3 at a tree size of 0.035 m3, but the two methods showed different characteristics for different tree sizes and level of accumulation. The direct loading method had the highest productivity when more than 0.1 m3 were collected in the felling cycle, whereas the separate loading method had the highest productivity when less than 0.05 m3 were collected in the felling cycle. The total effective time consumption for harvesting and forwarding the biomass 300 meters to roadside landing was 27 minutes per m3. The efficiency of the initial felling and collecting of the small trees was the main challenge. Both the harvesting technique and harvesting technology needs further development to provide a feasible production chain for woodfuel from energy thinning.
  • Belbo, Norwegian Forest and Landscape Institute, Box 115, 1431 Ås, Norway E-mail: helmer.belbo@skogoglandskap.no (email)
article id 141, category Research article
Per Kristian Rørstad, Erik Trømborg, Even Bergseng, Birger Solberg. (2010). Combining GIS and forest modelling in estimating regional supply of harvest residues in Norway. Silva Fennica vol. 44 no. 3 article id 141. https://doi.org/10.14214/sf.141
Keywords: bioenergy; forestry; SGIS; supply functions
Abstract | View details | Full text in PDF | Author Info
New and ambitious targets for renewable energy production put attention to increased supply of biomass. Harvest residues are only to a limited extent demanded by the traditional forest industries and represent an unutilized resource for increased production of renewable energy in Norway. The overall objective of this paper is to study how GIS and forest modelling can be combined to improve estimates of the supply of harvest residues, taking different environmental and economic constraints into consideration. The analyses are based on a case study of a forest area of more than 40 000 ha in Southern Norway divided into about 500 private forest properties. The study was carried out by computations of timber harvest using the forestry scenario model SGIS based on extensive forest inventory data at stand level. In the studied area energy utilization of harvest residues is not profitable below an energy price of about EUR 3.2/GJ (NOK 0.10 /kWh) when the distance from roadside to industry is 20 km. Above this level supply increases rapidly over a rather narrow price range and is nearly inelastic above EUR 4.1/GJ (NOK 0.12/kWh). We did not find significant negative shifts in the residues supply caused by changes in location of roundwood harvest over time. Exclusion of collection from stands with a site index (H40) below 14 reduced the potential supply of residues by 16–27%. The optimisation method combined selection of exogenous variables in order to map observed harvesting level and is probably the best approach to map future harvest.
  • Rørstad, Norwegian University of Life Sciences, Dept of Ecology and Natural Resource Management, Ås, Norway E-mail: per.kristian.rorstad@umb.no (email)
  • Trømborg, Norwegian University of Life Sciences, Dept of Ecology and Natural Resource Management, Ås, Norway E-mail: et@nn.no
  • Bergseng, Norwegian University of Life Sciences, Dept of Ecology and Natural Resource Management, Ås, Norway E-mail: eb@nn.no
  • Solberg, Norwegian University of Life Sciences, Dept of Ecology and Natural Resource Management, Ås, Norway E-mail: bs@nn.no
article id 140, category Research article
Jussi Laurila, Risto Lauhanen. (2010). Moisture content of Norway spruce stump wood at clear cutting areas and roadside storage sites. Silva Fennica vol. 44 no. 3 article id 140. https://doi.org/10.14214/sf.140
Keywords: Picea abies; harvesting; bioenergy; moisture content; stump wood
Abstract | View details | Full text in PDF | Author Info
Norway spruce (Picea abies (L.) Karst.) stump wood is a potential source of bioenergy in Finland. The heating value of stump wood depends on, among other things, the moisture, carbon and ash content of the wood. In this study the moisture content of Norway spruce stump wood was examined immediately after harvesting at the clear cutting area and after different drying times at the roadside storage sites. Immediately after stump harvesting the average moisture content (wet basis) was 53%. The stump wood dried fairly fast during spring and summer. One month after stump harvesting, the average moisture content was about 31%. If the stump wood had dried well once, water absorption became very weak and the moisture content increased only slightly in the late autumn. Each spring and summer the moisture content of the stumps was lower than during the previous year. Annually the lowest moisture content was observed at the beginning of July and the highest at both the beginning and the end of the year. The moisture content of stump wood followed an upwards opening parabola over a one year period and was repeated each year. Three years after harvesting the heating value of the stump wood was still 5.241 MWh/ton. Overall, when harvesting took place in the spring or early summer, the stump wood was combustible after a one month drying period immediately after harvesting.
  • Laurila, Seinäjoki University of Applied Sciences, School of Agriculture and Forestry, FI-63700 Ähtäri, Finland E-mail: jussi.laurila@seamk.fi (email)
  • Lauhanen, Seinäjoki University of Applied Sciences, School of Agriculture and Forestry, FI-63700 Ähtäri, Finland E-mail: rl@nn.fi
article id 139, category Research article
Ulf Sikström, Curt Almqvist, Gunnar Jansson. (2010). Growth of Pinus sylvestris after the application of wood ash or P and K fertilizer to a peatland in southern Sweden. Silva Fennica vol. 44 no. 3 article id 139. https://doi.org/10.14214/sf.139
Keywords: fertilization; wood ash; nitrogen; ammonium nitrate; forest yield; needle element concentrations; potassium chloride; raw phosphate
Abstract | View details | Full text in PDF | Author Info
The effects of the application of wood ash and of fertilizer regimes including phosphorus (P) and potassium (K), with and without simultaneous addition of nitrogen (N), were investigated on a stand of Scots pine (Pinus sylvestris L.) saplings growing on a drained oligotrophic peatland site in southern Sweden. A randomized block design was used. Tree growth and concentrations of various elements in the needles were measured. The addition of similar doses of P (approx. 40 kg P ha–1) from different sources resulted in similar growth responses, amounting to 1.6–1.9 m3 ha–1 yr–1 of stem wood over the 26-year study. The P source was either wood ash (2500 kg d.w. ha–1) or PK-fertilizer (raw phosphate and potassium chloride). In response to several treatments there were both increased numbers of trees and increased growth of individual trees. The high PK-dose (40 kg P ha–1 and 80 kg K ha–1) appeared to result in a larger growth increase than the low dose (20 kg P ha–1 and 40 kg K ha–1). The N treatment had no additional effect on growth. In the control plots, tree growth was more or less negligible (0.04 m3 ha–1 yr–1). After almost 26 years, concentrations of P and K in the needles of treated plants were still higher than in the untreated control plants. Nevertheless, in spite of the elevated P concentration, P appears to limit the growth of Scots pine. In conclusion, after sufficient drainage of this type of peatland site, it is possible for a forest stand to develop to the pole stage if wood ash or PK-fertilizer is applied.
  • Sikström, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: ulf.sikstrom@skogforsk.se (email)
  • Almqvist, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: ca@nn.se
  • Jansson, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: gj@nn.se
article id 138, category Research article
Santiago Martín-Alcón, José Ramón González-Olabarría, Lluís Coll. (2010). Wind and snow damage in the Pyrenees pine forests: effect of stand attributes and location. Silva Fennica vol. 44 no. 3 article id 138. https://doi.org/10.14214/sf.138
Keywords: models; stand structure; conifers; Pyrenees; snow damages; stability; wind damages
Abstract | View details | Full text in PDF | Author Info
Wind and snow-induced damage have been analyzed at stand level for three pine forests in the Central-Eastern Pyrenees (Pinus nigra Arn. salzmanii, Pinus sylvestris L. and Pinus uncinata Ram.). Stand-level models have been then developed for the most affected two species, Pinus sylvestris L. and Pinus uncinata Ram., to describe damage severity. The models were based on data from national forest inventory plots. They included variables related to the spatial location and structure of the stands, being validated using a sub-set of the database (25% of the plots randomly selected). Mountain pine forests (Pinus uncinata Ram.) were the most heavily affected by wind and snow disturbances. For both mountain and Scots pine species, topographic exposure and the severity of the local storm regime had an important effect on the degree of damage. Stand’s resistance to wind and snow was found to be dependent on the combined effect of basal area and mean slenderness of the dominant trees. For a given slenderness ratio, damage increased strongly in lower-density stands, particularly in stands with basal areas below 15 m2/ha. Stand structure was particularly important to define the resistance of Scots pine stands, which presented a higher vulnerability to wind and snow under higher degree of even-agedness. The models presented in this study provide empirically-based information that can be used to implement silvicultural practices to minimize the risk of those forests to suffer wind and snow-related damages.
  • Martín-Alcón, Forest Technology Center of Catalonia, Solsona, Lleida, Spain E-mail: santiago.martin@ctfc.es (email)
  • González-Olabarría, Forest Technology Center of Catalonia, Solsona, Lleida, Spain E-mail: jrgo@nn.es
  • Coll, Forest Technology Center of Catalonia, Solsona, Lleida, Spain E-mail: lc@nn.es
article id 137, category Research article
Alberto Bueno, Julio J. Diez, Mercedes M. Fernández. (2010). Ophiostomatoid fungi transported by Ips sexdentatus (Coleoptera; Scolytidae) in Pinus pinaster in NW Spain. Silva Fennica vol. 44 no. 3 article id 137. https://doi.org/10.14214/sf.137
Keywords: Spain; Ips sexdentatus; ophiostomatoid fungi; Pinus pinaster; mycangia
Abstract | View details | Full text in PDF | Author Info
Ips sexdentatus (Coleoptera; Scolytidae) is one of the main vectors of ophiostomatoid blue stain fungi that can cause mortality of healthy conifers. For this reason, our objective was to identify the fungal species carried by this bark beetle in Maritime pine (Pinus pinaster) in north-western Spain. We collected insects from naturally infected pines placed them on malt extract agar (MEA) and left to walk freely on culture plates. Plant tissues (phloem and xylem) from adult pines were cultivated in moist chambers and also on MEA. At the same time, we inoculated pine logs with living insects in the laboratory. Four ophiostomatoid fungi appeared: Ophiostoma ips, Ophiostoma brunneo-ciliatum, Ceratocystiopsis minuta and Ophiostoma sp., as well as Graphium and Sporothrix imperfect stages. Moreover there were seven saprophytic species: Penicillium sp., Trichoderma sp., Verticillium sp., Mucor sp., Aspergillus niger, Gliocladium viride and Scopulariopsis brevicaulis, and the pathogenic Ophiostoma ips. The fructification percentage of the ophiostomatoid species was low, however; its imperfect stage Sporothrix/Hyalorhinocladiella produced high quantity of conidiophores.
  • Bueno, University of Valladolid, Dept of Agroforestry Sciences, Palencia, Spain E-mail: ab@nn.es
  • Diez, University of Valladolid, Dept of Agroforestry Sciences, Palencia, Spain E-mail: jjd@nn.es
  • Fernández, University of Valladolid, Dept of Agroforestry Sciences, Palencia, Spain E-mail: mffernan@agro.uva.es (email)
article id 456, category Research article
Sam B. Coggins, Nicholas C. Coops, Michael A. Wulder. (2010). Improvement of low level bark beetle damage estimates with adaptive cluster sampling. Silva Fennica vol. 44 no. 2 article id 456. https://doi.org/10.14214/sf.456
Keywords: forest inventory; adaptive cluster sampling; mountain pine beetle; object-based classification; high spatial resolution; satellite; digital aerial imagery
Abstract | View details | Full text in PDF | Author Info
Detection of low level infestation in forest stands is of principle importance to determine effective control strategies before the attack spread to large areas. Of particular concern is the ongoing mountain pine beetle, Dendroctonus ponderosae (Hopkins) epidemic, which has caused approximately 14 million hectares of damage to lodgepole pine (Pinus contorta Dougl. ex. Loud var. latifolia Engl.) forests in western Canada. At the stand level attacked trees are often difficult to locate and can remain undetected until the infestation has become established beyond a small number of trees. As such, methods are required to detect and characterise low levels of attack prior to infestation expansion, to inform management, and to aid mitigation activities. In this paper, an adaptive cluster sampling approach was applied to very fine-scale (20 cm) digital aerial imagery to locate mountain pine beetle damaged trees at the leading edge of the current infestation. Results indicated a mean number of 7.36 infested trees per hectare with a variance of 18.34. In contrast a non-adaptive approach estimated the mean number of infested trees in the same area to be 61.56 infested trees per hectare with a variance of 41.43. Using a relative efficiency estimator the adaptive cluster sampling approach was found to be over two times more efficient when compared to the non-adaptive approach.
  • Coggins, Department of Forest Resources Management, University of British Columbia, 2424 Main Mall, Vancouver, B.C., Canada V6T 1Z4 E-mail: scoggins@interchange.ubc.ca (email)
  • Coops, Department of Forest Resources Management, University of British Columbia, 2424 Main Mall, Vancouver, B.C., Canada V6T 1Z4 E-mail: ncc@nn.ca
  • Wulder, Canadian Forest Service (Pacific Forestry Centre), Natural Resources Canada, 506 West Burnside Rd., Victoria, B.C., Canada V8Z 1M5 E-mail: maw@nn.ca
article id 455, category Research article
Seppo Kellomäki, Matti Maajärvi, Harri Strandman, Antti Kilpeläinen, Heli Peltola. (2010). Model computations on the climate change effects on snow cover, soil moisture and soil frost in the boreal conditions over Finland. Silva Fennica vol. 44 no. 2 article id 455. https://doi.org/10.14214/sf.455
Keywords: climate change; soil frost; precipitation; temperature; A2 climate scenario; snow accumulation; soil carrying capacity; soil moisture
Abstract | View details | Full text in PDF | Author Info
This study considered how climate change affects the accumulation of snow, the soil moisture and soil frost at sites without tree cover in boreal conditions in Finland (60°–70°N). An increase of 4.5 °C in annual mean temperature and 20 % in annual precipitation were assumed for Finland by the year 2100 according to A2 emission scenario. Along with climate, the soil type of the permanent inventory plots of the Finnish National Forest Inventory was used. Soil and climate data were combined by using a process-based ecosystem model. Calculations were done for four periods: current climate (1971–2000), near future (2001–2020), mid-term future (2021–2050) and long-term future (2071–2100). According to our simulations, the average monthly duration and depth of snow decreased over the simulation period. However, the increasing precipitation may locally increase the snow depths in the mid-term calculations. In the autumn and winter, the average volumetric soil moisture content slightly increased in southern Finland during the near future, but decreased towards the end of the century, but still remained on a higher level than presently. In northern Finland, the soil moisture in the autumn and winter increased by the end of this century. In the summertime soil moisture decreased slightly regardless of the region. Throughout Finland, the length and the depth of soil frost decreased by the end of the century. In the south, the reduction in the depth was largest in the autumn and spring, while in the mid-winter it remained relatively deep in the middle of the century. In the north, the depth tended to increase during the first two calculation periods, in some areas, even during the third calculation period (2071–2100) due to reduced insulation effects of snow during cold spells. The wintertime increase in soil moisture and reduced soil frost may be reflected to reduced carrying capacity of soil for timber harvesting.
  • Kellomäki, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: seppo.kellomaki@uef.fi (email)
  • Maajärvi, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: mm@nn.fi
  • Strandman, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: hs@nn.fi
  • Kilpeläinen, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: ak@nn.fi
  • Peltola, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: hp@nn.fi
article id 158, category Research article
Ville Hallikainen, Mikko Hyppönen, Leena Pernu, Jouni Puoskari. (2010). Family forest owners’ opinions about forest management in northern Finland. Silva Fennica vol. 44 no. 2 article id 158. https://doi.org/10.14214/sf.158
Keywords: forest management; silviculture; family forest owner; northern Finland; forest owner typology; forest management objective; forest management activity; tourism
Abstract | View details | Full text in PDF | Author Info
  • Hallikainen, Finnish Forest Research Institute, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: ville.hallikainen@metla.fi (email)
  • Hyppönen, Finnish Forest Research Institute, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: mh@nn.fi
  • Pernu, Finnish Forest Research Institute, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: lp@nn.fi
  • Puoskari, Finnish Forest Research Institute, Eteläranta 55, FI-96301 Rovaniemi, Finland E-mail: jp@nn.fi
article id 157, category Research article
Ville Kankaanhuhta, Timo Saksa, Heikki Smolander. (2010). The effect of quality management on forest regeneration activities in privately-owned forests in southern Finland. Silva Fennica vol. 44 no. 2 article id 157. https://doi.org/10.14214/sf.157
Keywords: silvicultural operations; quality control; program evaluation; program effectiveness
Abstract | View details | Full text in PDF | Author Info
This study describes the effect of quality management on forest regeneration activities in privately-owned forests in southern Finland. The study material consists of two mail surveys conducted in 2006 and 2007. The questionnaires were sent to forestry professionals in Forest Owners’ Associations (FOAs), who had either participated (participants) or not (non-participants) in the forest regeneration quality management interventions in years 2000–2006. Quality management interventions of the FOAs included field inventories, feedback meetings, and education sessions about quality techniques. The activities of participant and non-participant FOAs were compared in terms of the available resources, the actions taken, and the aspirations for improvement. In the FOAs, which had participated in quality management, the number of excavator contractors had increased by 16% more than in the non-participant FOAs. The contractors had acquired 23% more soil preparation equipment under the supervision of the participant forestry professionals. The soil preparation method used in conjunction with Norway spruce that had most increased in use was patch mounding. Seedlings that were 1.5 years and older were used by participant forestry professionals in the planting of Norway spruce 11% more than by non-participants. The planting workers had attended 14% more educational sessions, while the use of self-control measurements in soil preparation and planting density had been adopted 10% more frequently under the supervision of the forestry professionals participating in the quality management. A greater interest in obtaining feedback by using quality control inventories was also found among forestry professionals who participated in quality management.
  • Kankaanhuhta, Finnish Forest Research Institute, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: ville.kankaanhuhta@metla.fi (email)
  • Saksa, Finnish Forest Research Institute, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: ts@nn.fi
  • Smolander, Finnish Forest Research Institute, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: hs@nn.fi
article id 156, category Research article
Ilkka Korpela, Hans Ole Ørka, Matti Maltamo, Timo Tokola, Juha Hyyppä. (2010). Tree species classification using airborne LiDAR – effects of stand and tree parameters, downsizing of training set, intensity normalization, and sensor type. Silva Fennica vol. 44 no. 2 article id 156. https://doi.org/10.14214/sf.156
Keywords: airborne laser scanning; ALS; laser; Optech ALTM3100; Leica ALS50-II; canopy; crown modeling; monoplotting; backscatter amplitude; intensity; discriminant analysis
Abstract | View details | Full text in PDF | Author Info
Tree species identification constitutes a bottleneck in remote sensing-based forest inventory. In passive images the differentiating features overlap and bidirectional reflectance hampers analysis. Airborne LiDAR provides radiometric and geometric information. We examined the single-trees-level response of two LiDAR sensors in over 13 000 forest trees in southern Finland. We focused on the commercially important species. Our aims were to 1) explore the relevant LiDAR features and study their dependencies on stand and tree variables, 2) examine two sensors and their fusion, 3) quantify the gain from intensity normalizations, 4) examine the importance of the size of the training set, and 5) determine the effects of stand age and site fertility. A set of 570 semiurban broad-leaved trees and exotic conifers was analyzed to 6) examine the LiDAR signal in the economically less important species. An accuracy of 88 90% was achieved in the classification of Scots pine, Norway spruce, and birch, using intensity variables. Spruce and birch showed the highest levels of confusion. Downsizing the training set from 30% to 2.5% of all trees had only a marginal effect on the performance of classifiers. The intensity features were dependent on the absolute and relative sizes of trees, especially for birch. The results suggest that leaf size, orientation, and foliage density affect the intensity, which is thus not affected by reflectance only. Some of the ecologically important species in Finland may be separable, since they gave rise to high intensity values. Comparison of the sensors implies that performance of the intensity data for species classification varies between sensors for reasons that remained uncertain. Both range and gain receiver normalization improved species classification. Weighting of the intensity values improved the fusion of two LiDAR datasets.
  • Korpela, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ilkka.korpela@helsinki.fi (email)
  • Ørka, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, P.O.Box 5003, NO-1432 Ås, Norway E-mail: hoo@nn.no
  • Maltamo, University of Eastern Finland, School of Forest Science, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: mm@nn.fi
  • Tokola, University of Eastern Finland, School of Forest Science, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: tt@nn.fi
  • Hyyppä, Finnish Geodetic Institute, Department of Photogrammetry and Remote Sensing, P.O.Box 15, FI-02431 Masala, Finland E-mail: jh@nn.fi
article id 155, category Research article
Minna Räty, Annika Kangas. (2010). Segmentation of model localization sub-areas by Getis statistics. Silva Fennica vol. 44 no. 2 article id 155. https://doi.org/10.14214/sf.155
Keywords: eCognition; form height; Getis statistics; image segmentation; local indicators of spatial association
Abstract | View details | Full text in PDF | Author Info
Models for large areas (global models) are often biased in smaller sub-areas, even when the model is unbiased for the whole area. Localization of the global model removes the local bias, but the problem is to find homogenous sub-areas in which to localize the function. In this study, we used the eCognition Professional 4.0 (later versions called Definies Pro) segmentation process to segment the study area into homogeneous sub-areas with respect to residuals of the global model of the form height and/or local Getis statistics calculated for the residuals, i.e., Gi*-indices. The segmentation resulted in four different rasters: 1) residuals of the global model, 2) the local Gi*-index, and 3) residuals and the local Gi*-index weighted by the inverse of the variance, and 4) without weighting. The global model was then localized (re-fitted) for these sub-areas. The number of resulting sub-areas varied from 4 to 366. On average, the root mean squared errors (RMSEs) were 3.6% lower after localization than the global model RMSEs in sub-areas before localization. However, the localization actually increased the RMSE in some sub-areas, indicating the sub-area were not appropriate for local fitting. For 56% of the sub-areas, coordinates and distance from coastline were not statistically significant variables, in other words these areas were spatially homogenous. To compare the segmentations, we calculated an aggregate standard error of the RMSEs of the single sub-areas in the segmentation. The segmentations in which the local index was present had slightly lower standard errors than segmentations based on residuals.
  • Räty, University of Helsinki, Department of Forest Sciences, P.O. Box 27 (Latokartanonkaari 7), FI-00014 University of Helsinki, Finland E-mail: minna.s.raty@helsinki.fi (email)
  • Kangas, University of Helsinki, Department of Forest Sciences, P.O. Box 27 (Latokartanonkaari 7), FI-00014 University of Helsinki, Finland E-mail: ak@nn.fi
article id 154, category Research article
Henrik R. Hallingbäck, Gunnar Jansson, Björn Hannrup, Anders Fries. (2010). Which annual rings to assess grain angles in breeding of Scots pine for improved shape stability of sawn timber? Silva Fennica vol. 44 no. 2 article id 154. https://doi.org/10.14214/sf.154
Keywords: Pinus sylvestris; spiral grain angle; warp; sawn timber; breeding
Abstract | View details | Full text in PDF | Author Info
The shape stability properties of sawn timber could be improved by breeding or grading Scots pine (Pinus sylvestris L.) for reduced grain angles. Currently, only grain angle assessments performed in single annual rings can be considered feasible in forest breeding programmes. The relevance of such methods in assessing shape stability traits was evaluated by taking grain angle measurements beneath the bark in a 36-year-old Scots pine progeny trial. Several grain angle measurements from stem discs were also taken from a sample of 162 trees. Phenotypic correlations were estimated between grain angle and the bow, crook and twist developed in 316 sawn and dried boards. All single annual ring assessments, including measurements taken directly under the bark, were significantly correlated with twist. The highest correlations (0.60–0.70) were observed in annual rings numbered 8–20 and at distances of 30–70 mm from the pith, indicating those parts of logs where grain angles have the largest impact on twist. These results suggests, that grain angles measured beneath the bark are relevant to the twist of sawn small timber, and that any single annual ring could be chosen for the assessment, provided that the tree diameter is within the 60–140 mm range. No appreciable correlations were observed between grain angles and either crook or bow.
  • Hallingbäck, Dept of Plant Biology and Forest Genetics, Swedish University of Agricultural Sciences, Uppsala, Sweden E-mail: henrik.hallingback@vbsg.slu.se (email)
  • Jansson, The Forestry Research Institute of Sweden, Uppsala, Sweden E-mail: gj@nn.se
  • Hannrup, The Forestry Research Institute of Sweden, Uppsala, Sweden E-mail: bh@nn.se
  • Fries, Dept of Forest Genetics and Plant Physiology, Swedish University of Agricultural Sciences, Umeå, Sweden E-mail: af@nn.se
article id 153, category Research article
Dirk Bieker, Steffen Rust. (2010). Non-destructive estimation of sapwood and heartwood width in Scots pine (Pinus sylvestris L.). Silva Fennica vol. 44 no. 2 article id 153. https://doi.org/10.14214/sf.153
Keywords: non-destructive testing; tomography; sapwood; pine
Abstract | View details | Full text in PDF | Author Info
Accurate estimates of the water conducting sapwood area are necessary to scale sapflow measurements to tree and stand level transpiration. We tested a non-destructive method, electric resistivity tomography (ERT), to estimate the area of conductive sapwood in 9 Pinus sylvestris L. trees in lower Saxony, Germany. Tomograms were compared to cross-sections stained with benzidine after harvesting. All tomograms displayed a distinct pattern of low resistivity at the stem perimeter and high resistivity in the stem centre with a steep increase in resistivity in between, assumed to indicate the transition from sapwood to heartwood. The tomograms showed a sapwood width 2 cm smaller than the staining method. This indicates that staining methods overestimate the amount of active sapwood because when heartwood is formed, moisture content decreases before extractive contents reach levels visible by staining. The ERT method is a new powerful method for the non-destructive estimation of sapwood and heartwood width.
  • Bieker, University of Applied Sciences and Arts, Faculty of Resource Management, Büsgenweg 1a, D-37077 Göttingen, Germany E-mail: db@nn.de
  • Rust, University of Applied Sciences and Arts, Faculty of Resource Management, Büsgenweg 1a, D-37077 Göttingen, Germany E-mail: rust@hawk-hhg.de (email)
article id 152, category Research article
Torgny Persson, Bengt Andersson, Tore Ericsson. (2010). Relationship between autumn cold hardiness and field performance in northern Pinus sylvestris. Silva Fennica vol. 44 no. 2 article id 152. https://doi.org/10.14214/sf.152
Keywords: Scots pine; cold hardiness; genetic coefficient of variation; genetic correlation; multivariate analysis; narrow-sense heritability
Abstract | View details | Full text in PDF | Author Info
Results from 3 artificial freezing tests (one-year-old seedlings) and 15 field trials (9- to 21-year old trees) of half-sib offspring from first generation Scots pine (Pinus sylvestris L.) plus-trees were used to estimate the amount of additive genetic variance for autumn cold hardiness and traits assessed in the field, and the genetic correlations between them. Cold hardiness of individual seedlings was scored visually, based on the discoloration of their needles after freezing in a climate chamber. The field traits analyzed were tree vitality, tree height, spike knot frequency, branch diameter, branch angle, stem straightness, and susceptibility to infection by the pathogenic fungi Phacidium infestans L., Gremmeniella abietina (Lagerb.) Morelet, Melampsora pinitorqua (Braun) Rostr. and Lophodermella sulcigena (Rostr.) Höhn. Narrow sense individual heritabilities varied between 0.30 and 0.54 for autumn cold hardiness, 0 and 0.18 for tree vitality, 0.07 and 0.41 for tree height, and 0.01 and 0.26 for the remaining traits. Based on the results of the artificial freeze tests, our estimates of additive genetic correlations indicate that while early selection for cold hardiness can improve seedling survival rates in the field, it may also reduce growth in mild environments. It also has minor effects on quality traits and attack by common fungal diseases. The results indicate that artificial freeze testing is an appropriate method for identifying suitable clones for establishing seed orchards to supply stock for the reforestation of regions with harsh environments.
  • Persson, Forestry Research Institute of Sweden, Sävar, Sweden E-mail: torgny.persson@skogforsk.se (email)
  • Andersson, Forestry Research Institute of Sweden, Sävar, Sweden E-mail: ba@nn.se
  • Ericsson, Forestry Research Institute of Sweden, Sävar, Sweden E-mail: te@nn.se
article id 151, category Research article
Janne Miettinen, Pekka Helle, Ari Nikula, Pekka Niemelä. (2010). Capercaillie (Tetrao urogallus) habitat characteristics in north-boreal Finland. Silva Fennica vol. 44 no. 2 article id 151. https://doi.org/10.14214/sf.151
Keywords: forest management; forestry; grouse; thinnings; landscape composition
Abstract | View details | Full text in PDF | Author Info
This study aimed to identify tools for taking capercaillie habitats into consideration in forest management. This would provide new alternatives for ecologically more sustainable forest management. Capercaillie summer and winter locations, from wildlife monitoring counts (1998–2004) in northern Finland, and reference, non-capercaillie locations were combined with forest planning data, and the area proportions of different landscape classes in an 800-m radius circle surrounding capercaillie and reference locations were compared. Thinning stands (in summer and winter) and spruce mires (in summer) were more abundant in capercaillie habitats than in reference landscapes, whereas e.g. seedling stands, mature stands and waste land areas were less abundant. The relative habitat use was highest in mean tree diameter (DBH) classes from 10.5 to 14.5 cm in summer habitats of adult capercaillie in heath forests, whereas in peatland forests, in brood habitats and in winter habitats it peaked in diameter classes 14.5 to 18.5 cm. The tree layer density was positively associated with the relative habitat use. A trend of lower habitat use was detected in the largest diameters (17–40 cm) in comparison to middle-sized diameters (10–16 cm) in heath forests, but not in peatland forests. Relatively young managed forests (age 30–40 years or more) can form suitable capercaillie habitats in north-boreal forests. However, this suitability is not necessarily permanent. Understorey management, longer rotations and multicohort forest management are suitable tools for capercaillie habitat management, because they can increase the available cover close to the ground, canopy cover, overall forest cover at the landscape scale and bilberry cover.
  • Miettinen, Finnish Game and Fisheries Research Institute, Finland E-mail: janne.miettinen@rktl.fi (email)
  • Helle, Finnish Game and Fisheries Research Institute, Finland E-mail: ph@nn.fi
  • Nikula, Finnish Forest Research Institute, Finland E-mail: an@nn.fi
  • Niemelä, University of Turku, Dept of Biology E-mail: pn@nn.fi
article id 150, category Research article
Sini Eräjää, Panu Halme, Janne S. Kotiaho, Anni Markkanen, Tero Toivanen. (2010). The volume and composition of dead wood on traditional and forest fuel harvested clear-cuts. Silva Fennica vol. 44 no. 2 article id 150. https://doi.org/10.14214/sf.150
Keywords: dead wood; forest fuel harvesting; fine woody debris (FWD); clear-cut
Abstract | View details | Full text in PDF | Author Info
Logging residue and cut stumps are increasingly used as a renewable energy source known as forest fuel. Forest fuel harvesting obviously reduces the volume of dead wood and is likely to alter the dead wood composition, but the magnitude of the change is not known. Such information is important for the evaluation of the effects of forest fuel harvesting on biodiversity because a large proportion of forest dwelling species are directly dependent on dead wood. We measured the volume and characteristics of all dead wood units with a minimum diameter of 2 cm and a minimum length of 20 cm on 10 forest-fuel harvested and 10 traditional (control) clear-cuts. The total volume of dead wood at forest fuel harvested and control clear-cuts was 26.0 and 42.3 m3/ha, respectively. The volumes were much greater than expected suggesting that the volume of dead wood on clear-cuts has been underestimated in previous studies. Forest fuel harvested clear-cuts had 42% less branches and 81% less cut stumps than control clear-cuts but there were no differences in the volume of logs and pieces of logs, snags or roots. The volume of fine woody debris was negatively affected by forest fuel harvesting. We conclude that fine woody debris and cut stumps form a considerable resource on clear-cuts that is reduced by forest fuel harvesting. These components of dead wood have potential to be of importance in managed forests and thus deserve more attention in future biodiversity studies.
  • Eräjää, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: se@nn.fi
  • Halme, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: panu.halme@jyu.fi (email)
  • Kotiaho, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: jsk@nn.fi
  • Markkanen, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: am@nn.fi
  • Toivanen, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: tt@nn.fi
article id 149, category Research article
Juha Kaitera, Heikki Nuorteva. (2010). Effects of Melampyrum extracts on the growth of axenic cultures of Cronartium flaccidum and Peridermium pini. Silva Fennica vol. 44 no. 2 article id 149. https://doi.org/10.14214/sf.149
Keywords: Scots pine; alternate host; anti-fungal phytochemicals; fungal growth regulation; pine stem rust
Abstract | View details | Full text in PDF | Author Info
For 3–6 months, mycelial colonies cultured from 5 isolates of each of two pine stem rusts (Cronartium flaccidum and Peridermium pini) were grown on nutrient-rich agar supplemented with Melampyrum extracts. Non-autoclaved extracts of M. pratense significantly reduced the growth of P. pini. The growth of C. flaccidum isolates was slightly stimulated after the second month of incubation but after that was inhibited during incubation months 4–6. We observed considerable variation in colony growth, a significant component of which was explained by incubation time, isolate, growth medium and their interaction. Rust species (C. flaccidum or P. pini) was not an important factor in growth variation. While sterilized extracts of M. pratense, M. sylvaticum and M. nemorosum did not significantly affect growth, colonies of C. flaccidum were slightly stimulated, whereas colonies of P. pini were slightly inhibited. Generally, isolates of P. pini grew better and showed a slower rate of degeneration than C. flaccidum on all media.
  • Kaitera, Finnish Forest Research Institute, Northern Finland Regional Unit, FI-91500 Muhos, Finland E-mail: juha.kaitera@metla.fi (email)
  • Nuorteva, Finnish Forest Research Institute, Southern Finland Regional Unit, FI-01301 Vantaa, Finland E-mail: hn@nn.fi
article id 459, category Research article
Tomi Tulokas, Jawdat Tannous. (2010). Research method and improvement of log rotation in sawmills. Silva Fennica vol. 44 no. 1 article id 459. https://doi.org/10.14214/sf.459
Keywords: yield; log rotation; sawing; optimizing; precision; standard deviation
Abstract | View details | Full text in PDF | Author Info
  • Tulokas, Centre for Timber Engineering, Edinburgh Napier University, 10 Colinton Road, Edinburgh EH10 5DT, UK E-mail: tomi.tulokas@lut.fi (email)
  • Tannous, School of Engineering and the Built Environment, Edinburgh Napier University, 10 Colinton Road, Edinburgh EH10 5DT, UK E-mail: jt@nn.uk
article id 458, category Research article
Sakari Tuominen, Kalle Eerikäinen, Anett Schibalski, Markus Haakana, Aleksi Lehtonen. (2010). Mapping biomass variables with a multi-source forest inventory technique. Silva Fennica vol. 44 no. 1 article id 458. https://doi.org/10.14214/sf.458
Keywords: National Forest Inventory; remote sensing; biomass models; biomass maps
Abstract | View details | Full text in PDF | Author Info
Map form information on forest biomass is required for estimating bioenergy potentials and monitoring carbon stocks. In Finland, the growing stock of forests is monitored using multi-source forest inventory, where variables are estimated in the form of thematic maps and area statistics by combining information of field measurements, satellite images and other digital map data. In this study, we used the multi-source forest inventory methodology for estimating forest biomass characteristics. The biomass variables were estimated for national forest inventory field plots on the basis of measured tree variables. The plot-level biomass estimates were used as reference data for satellite image interpretation. The estimates produced by satellite image interpretation were tested by cross-validation. The results indicate that the method for producing biomass maps on the basis of biomass models and satellite image interpretation is operationally feasible. Furthermore, the accuracy of the estimates of biomass variables is similar or even higher than that of traditional growing stock volume estimates. The technique presented here can be applied, for example, in estimating biomass resources or in the inventory of greenhouse gases.
  • Tuominen, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: sakari.tuominen@metla.fi (email)
  • Eerikäinen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: ke@nn.fi
  • Schibalski, University of Potsdam, Karl-Liebknecht-Strasse 24–25, 14476 Potsdam, Germany E-mail: as@nn.de
  • Haakana, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: mh@nn.fi
  • Lehtonen, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: al@nn.fi
article id 457, category Research article
Karin Öhman, Ljusk Ola Eriksson. (2010). Aggregating harvest activities in long term forest planning by minimizing harvest area perimeters. Silva Fennica vol. 44 no. 1 article id 457. https://doi.org/10.14214/sf.457
Keywords: optimization; harvest scheduling; mixed integer programming; spatial consideration; trade off
Abstract | View details | Full text in PDF | Author Info
The study presents a new approach for aggregating stands for harvest in strategic forest planning. In fragmented landscapes this could benefit nature conservation as well as reduce costs. The approach is built on the idea of minimizing the outside perimeter of contiguous harvest areas. The formulation allows for the use of exact solution methods such as mixed integer programming. The method was tested in a landscape consisting of 2821 stands. The application showed that large and compact harvest areas were created with limited sacrifice of financial value. The mixed integer programs were in most cases solved within a couple of hours. The method needs to be tested on different landscapes with different degrees of fragmentation. It is also necessary to evaluate the long term consequences of the large clear cuts that appear to be a consequence of this problem formulation.
  • Öhman, SLU, Department of Forest Resource Management, SE-901 83 Umeå, Sweden E-mail: karin.ohman@srh.slu.se (email)
  • Eriksson, SLU, Department of Forest Resource Management, SE-901 83 Umeå, Sweden E-mail: loe@nn.se
article id 167, category Research article
Berit H. Lindstad, Birger Solberg. (2010). Assessing national compliance with international forest policy processes – the role of subjective judgments. Silva Fennica vol. 44 no. 1 article id 167. https://doi.org/10.14214/sf.167
Keywords: sustainable forest management; multiple objectives; implementation; participatory process
Abstract | View details | Full text in PDF | Author Info
Several international policy processes with sustainable forest management (SFM) as a common goal have emerged during the past two decades. Based on an empirical study from Norway, this paper analyses the role of subjective judgments in assessing national compliance with three international forest policy processes, and the implications for determination of the effects of these processes. The Expanded Programme of Work on Forest Biological Diversity, the Ministerial Conference on the Protection of Forests in Europe and the United Nations Forum on Forests, including its predecessors, collectively provide more than 600 recommendations for SFM. While it is nothing new that SFM encompasses value questions, this paper is a systematic review of where in a process of assessing national compliance the role of judgments is most profound. The paper shows that the multiple objectives of the forest recommendations, references to national circumstances and provisions for stakeholder involvement lead to differing opinions about the degree of conformity between international recommendations and national situation, i.e. compliance. These differing opinions mean different prospects for the international processes to have effects, because only implementation, or active responses to international recommendations, constitutes effects. The roles of judgments and values are recommended topics for further investigation. Factors influencing how compliance is assessed, and consequently the degree to which implementation is deemed necessary, require specific attention. Due consideration to substantive and methodological choices in determining national changes and in separation of other sources of influence will provide a better basis for informed discussion of compliance with and effects of international forest-related policy processes.
  • Lindstad, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, Ås, Norway E-mail: berit.lindstad@umb.no (email)
  • Solberg, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, Ås, Norway E-mail: bs@nn.no
article id 166, category Research article
Kim Pingoud, Johanna Pohjola, Lauri Valsta. (2010). Assessing the integrated climatic impacts of forestry and wood products. Silva Fennica vol. 44 no. 1 article id 166. https://doi.org/10.14214/sf.166
Keywords: carbon stocks; managed forests; silvicultural guidelines; harvested wood products; energy and material substitution; displacement of fossil carbon emissions
Abstract | View details | Full text in PDF | Author Info
Managed forests serve as a store of carbon (C) and a renewable source of energy and materials. By using forest products as substitutes for fossil fuels or non-renewable materials, emissions from fossil C sources can be displaced. The efficiency of emissions displacement depends on the product, its lifecycle and the fossil-fuel based reference system that is substituted. Forest management practices have an impact on C stocks in biomass and on the annual supply of products and their mix. There are trade-offs between sequestering C stocks in forests and the climatic benefits obtained by sustainable forest harvesting and using wood products to displace fossil C emissions. This article presents an integrated, steady-state analysis comparing various equilibrium states of managed forests and wood product pools that represent sustainable long-term forestry and wood-use strategies. Two climatic indicators are used: the combined C stock in forests and wood products and the fossil C emissions displaced annually by harvested wood products. The study indicates that long-term strategies could be available that are better according to both indicators than forestry practices based on the existing silvicultural guidelines in Finland. These strategies would involve increasing the basal area and prolonging rotations to produce more sawlogs. Further, the climate benefits appear to be highest in case the sawlog supply is directed to production of long-lived materials substituting for fossil-emission and energy intensive materials and recycled after their useful life to bioenergy.
  • Pingoud, VTT Technical Research Centre of Finland, P.O. Box 1000, FI-02044 VTT, Espoo, Finland E-mail: kim.pingoud@vtt.fi (email)
  • Pohjola, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: jp@nn.fi
  • Valsta, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: lv@nn.fi
article id 165, category Research article
Yrjö Nuutinen, Kari Väätäinen, Antti Asikainen, Robert Prinz, Jaakko Heinonen. (2010). Operational efficiency and damage to sawlogs by feed rollers of the harvester head. Silva Fennica vol. 44 no. 1 article id 165. https://doi.org/10.14214/sf.165
Keywords: productivity; single grip harvester; feed roller; timber damages; work study
Abstract | View details | Full text in PDF | Author Info
In mechanical cutting, deep damage caused by feed rollers can reduce the yield of good quality timber for the sawmill and plywood industries. Additionally the feeding and energy efficiency of feed rollers are important for the profitability of harvester cutting. The objectives of this study were to compare the damages to sawlogs, as well as the time and fuel consumption of stem feeding with six different steel feed rollers during the processing of stems using a single grip harvester. This study tested two rollers with big spikes, two rollers with small spikes, one roller with studs in v-angle and one roller with adaptable steel plates in the ring of the roller. A highly detailed, and accurate processing and fuel consumption projection was recorded using the harvester’s automated data collector on a log and stem level. The roller adaptable plate averaged, for unbarked sawlogs, the lowest damages of 3.7 mm. While the damages of the roller with big spikes were the deepest with an average of 7.8 mm. For medium stems, volume of 0.35 m3, the range of differences between the maximum and minimum effective feeding time per roller was 6–19%, which would increase the effective time consumption of cutting by 1–3%. Corresponding differences in fuel consumption during total stem processing were in the range of 7–15%. According to this study it can be concluded that the traditional rollers with spikes were most effective in processing and fuel consumption, but at the same time they caused the deepest damages to the sawlogs. The roller type with adaptable steel plates was the most effective for small stems, additionally it also caused the least damage to the sawlogs.
  • Nuutinen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: yrjo.nuutinen@metla.fi (email)
  • Väätäinen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: kv@nn.fi
  • Asikainen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: aa@nn.fi
  • Prinz, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: rp@nn.fi
  • Heinonen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jh@nn.fi
article id 164, category Research article
Aki Suvanto, Matti Maltamo. (2010). Using mixed estimation for combining airborne laser scanning data in two different forest areas. Silva Fennica vol. 44 no. 1 article id 164. https://doi.org/10.14214/sf.164
Keywords: airborne laser scanning; area-based method; mixed estimation; regression models
Abstract | View details | Full text in PDF | Author Info
Airborne laser scanning (ALS) data have become the most accurate remote sensing technology for forest inventories. When planning new inventories the costs of fieldwork could be reduced if datasets of old inventory areas are effectively reused in the new area. The aim of this study was to apply mixed estimation using a combination of existing and new field datasets in area-based approach. Additionally, combining datasets with mixed estimation was compared with constructing new local models with smaller datasets. The two forest study areas were in Juuka and Matalansalo, which are located about 120 km apart in eastern Finland. ALS-based regression models were constructed using datasets of Matalansalo (472 reference plots) and Juuka (10–212 reference plots). Models were developed for the basal area median tree diameter and height, mean tree height, stem number, basal area and volume. The work was based on a simulation approach which involved five methods for approximating the regression coefficients. The first method merged the datasets using ordinary least squares (OLS) regression models, whereas the second and third methods combined datasets using mixed estimation on different weighting principles, and the final two estimated local models with predetermined and new independent variables. The results indicate that mixed estimation can improve the accuracy of derived stand variables compared with basic OLS models. Additionally, a sample of 40–50 plots was enough to build local models for basal area and volume and produce at least the equal accuracy of results than any other methods in this study.
  • Suvanto, Blom Kartta Oy, Teollisuuskatu 18, FI-80100 Joensuu, Finland E-mail: aki.suvanto@blomasa.com (email)
  • Maltamo, University of Eastern Finland, School of Forest Sciences, P.O. Box, FI-80101, Joensuu, Finland E-mail: mm@nn.fi
article id 163, category Research article
Blas Mola-Yudego. (2010). Regional potential yields of short rotation willow plantations on agricultural land in Northern Europe. Silva Fennica vol. 44 no. 1 article id 163. https://doi.org/10.14214/sf.163
Keywords: short rotation forestry; willow; production models; regional biomass supply; energy planning
Abstract | View details | Full text in PDF | Author Info
The development of short rotation forestry for bioenergy requires accurate and reliable yield estimates. This paper analyses the current, expected and potential regional productivity of short rotation willow plantations for six countries in Northern Europe. The estimations for present productivity are based on empirical models, using data regarding management, and local productivity based on the regional cereal yields. The estimates of expected yield rely on the current trends of yield increase from commercial willow plantations in the region. The estimates for potential yield are based on climatic restrictions. The results show potential average yields of 9.5, 6.8, 7.9, 9.0, 9.3, and 8.0 odt ha–1 yr–1 for Denmark, Finland, Estonia, Latvia, Lithuania and Sweden, respectively. The results of the study also show that there is a wide regional variation between the different countries. In Denmark, Finland and Sweden there is a convergence between the future forecasts and the climatic potential yields in the areas of high productivity. The Baltic countries seem to present lower estimates of present productivity, reflecting possible socio-economic restrictions, although they show a high biomass potential. The methods presented in this study can be further developed in other areas where willow cultivation is considered, and can serve as a basis for future economic considerations.
  • Mola-Yudego, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: blas.mola@uef.fi (email)
article id 162, category Research article
Ruut Rabinowitsch-Jokinen, Ilkka Vanha-Majamaa. (2010). Immediate effects of logging, mounding and removal of logging residues and stumps on coarse woody debris in managed boreal Norway spruce stands. Silva Fennica vol. 44 no. 1 article id 162. https://doi.org/10.14214/sf.162
Keywords: forest management; soil preparation; bioenergy; log; stump harvesting; CWD; logging residues; snag
Abstract | View details | Full text in PDF | Author Info
Wood fuel production has increased remarkably, but its environmental effects within the forest ecosystem have not yet been studied much. We investigated the immediate effects of two series of forest management treatments, which produce timber and forest chips, on the volume and decay classes of coarse woody debris (CWD). One of the treatment series included logging and residue harvesting (LRH) and mounding (M), while the other series included LRH and mounding combined with stump harvesting (MSH). We hypothesized that, i) LRH reduces CWD, excluding stumps; ii) the more intense the soil preparation treatment is, M vs. MSH, the more CWD is destroyed; iii) both LRH and soil preparation treatments (M and MSH) reduce the occurence of snags, highly decayed CWD and deciduous CWD in particular. Ten sample plots in mature managed Norway spruce (Picea abies (L.) H. Karst.) dominated forests were located in Southern Finland. The total volume of CWD on the sample plots was measured three times: before and after LRH, and after M or MSH. LRH significantly decreased the volume of snags and the combined volume of snags and logs. MSH significantly decreased the total volume of CWD, while M had no significant effect on the volume of CWD. The middle and highly decayed CWD were destroyed most easily in the treatments.
  • Rabinowitsch-Jokinen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301, Vantaa, Finland E-mail: rrj@nn.fi
  • Vanha-Majamaa, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301, Vantaa, Finland E-mail: ilkka.vanha-majamaa@metla.fi (email)
article id 161, category Research article
Mika Nieminen, Erkki Ahti, Harri Koivusalo, Tuija Mattsson, Sakari Sarkkola, Ari Laurén. (2010). Export of suspended solids and dissolved elements from peatland areas after ditch network maintenance in south-central Finland. Silva Fennica vol. 44 no. 1 article id 161. https://doi.org/10.14214/sf.161
Keywords: drained peatlands; forestry; ditching; hydrochemistry; nutrient export; suspended solids
Abstract | View details | Full text in PDF | Author Info
In Finland nearly 6 million hectares of peatlands are drained for forestry purposes. Ditch network maintenance in the drained peatlands, i.e. cleaning old ditches or digging complementary ditches, deteriorates surface water quality by increasing the export of dissolved elements and suspended solids (SS). Effect of ditch network maintenance on the export of SS, dissolved organic carbon (DOC), and dissolved nitrogen (N), phosphorous (P), iron (Fe), aluminum (Al) and manganese (Mn) was studied in nine pairs of treated and control (no maintenance) catchments located in southern and central Finland. In this study we extended the paired catchment approach by combining data from several catchments and identifying the treatment effect on SS and element loads from the entire dataset. Following the method of Laurén et al. (2009) we identified how uncertainty in correlation between treatment and control catchments during pre-treatment period is reflected in the estimated treatment effect on SS and element loads. In the experiment, the export of SS increased significantly for the four year study period following the ditch network maintenance and Al export increased for one year. The export of N, P and Fe was not significantly changed and DOC and Mn export decreased after the ditch maintenance operation.
  • Nieminen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: mika.nieminen@metla.fi (email)
  • Ahti, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: ea@nn.fi
  • Koivusalo, Department of Civil and Environmental Engineering, Aalto University School of Science and Technology, P.O. Box 15200, FI-00076 Aalto, Finland E-mail: hk@nn.fi
  • Mattsson, Finnish Environment Institute, P.O. Box 140, FI-00251 Helsinki, Finland E-mail: tm@nn.fi
  • Sarkkola, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: ss@nn.fi
  • Laurén, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: al@nn.fi
article id 160, category Research article
Fan Yang, Ling-Feng Miao. (2010). Adaptive responses to progressive drought stress in two poplar species originating from different altitudes. Silva Fennica vol. 44 no. 1 article id 160. https://doi.org/10.14214/sf.160
Keywords: antioxidant enzymes; drought stress; free proline; hydrogen peroxide; malondialdehyde; Populus
Abstract | View details | Full text in PDF | Author Info
Cuttings of Populus kangdingensis C. Wang et Tung and Populus cathayana Rehder, originating from high and low altitudes in the eastern Himalaya, respectively, were examined during one growing season in a greenhouse to determine the effects of progressive drought stress. The results manifested that the adaptive responses to progressive drought stress were different in these two species from different altitudes. Significant changes in stem height, leaf development, relative water content (RWC), malondialdehyde (MDA) and hydrogen peroxide (H2O2) appeared earlier in P. cathayana than in P. kangdingensis, whereas changes in soluble protein, soluble sugar, free proline and antioxidant enzymes appeared earlier in P. kangdingensis. In addition, changes in these parameters became more and more significant when the drought stress progressed, especially under severe drought stress in P. cathayana. Plant growth showed significant positive correlations with soluble proteins and sugars, free proline and antioxidants and a significant negative correlation with RWC under water stressed treatment in two poplar species. Compared with P. cathayana, P. kangdingensis was able to maintain a superior height growth and leaf development under drought stress. Also, P. kangdingensis possessed greater increments in soluble protein, soluble sugar, free proline and antioxidant enzymes, but lower increments in MDA and H2O2 than did P. cathayana when the cuttings were exposed to progressive drought stress. Our results suggest that P. kangdingensis originating from the high altitude has a better drought tolerance than does P. cathayana originating from the low altitude. Furthermore, this study manifested that acclimation to drought stress are related the rapidity, severity, duration of the drought event and the altitude of two poplar species.
  • Yang, Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan, Hubei 430074, P. R. China (yangfan@wbgcas.cn) & Chengdu Institute of Biology, Chinese Academy of Sciences, P.O. Box 416, Chengdu, Sichuan 610041, P. R. China E-mail: fanyangmlf6303@163.com (email)
  • Miao, Wuhan Institute of Virology, Chinese Academy of Sciences, Wuhan, Hubei 430071, P. R. China E-mail: lfm@nn.cn
article id 159, category Research article
Johan Stendahl, Maj-Britt Johansson, Erik Eriksson, Åke Nilsson, Ola Langvall. (2010). Soil organic carbon in Swedish spruce and pine forests – differences in stock levels and regional patterns. Silva Fennica vol. 44 no. 1 article id 159. https://doi.org/10.14214/sf.159
Keywords: forest management; Norway spruce; Scots pine; soil carbon; temperature sum; tree species; simulation
Abstract | View details | Full text in PDF | Author Info
The selection of tree species is one factor to consider if we want to mitigate carbon dioxide emissions to the atmosphere through forest management. The objectives of this study were to estimate the differences in soil organic carbon (SOC) stocks under Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) forests and to examine causes of differences in the accumulation of carbon in the forest soil. Large-scale inventory data was used to quantify variations in SOC stock in relation to stand type and the accumulation of carbon for spruce and pine stands was analysed by simulation. Based on field data, the national mean SOC stock was 9.2 kg m–2 in spruce dominated stands and 5.7 kg m–2 in pine dominated stands. For both species, the SOC stock, measured in the field inventory, increased significantly with increasing temperature, although at different rates. The SOC stock was larger for spruce under all temperature conditions, but the difference between species diminished with increasing temperature. The simulations indicated that the build-up of SOC over several rotations was 22% higher in spruce stands than in pine stands under similar environmental conditions. The main difference was found to be the greater input of harvest residues for spruce. Further, the simulations showed that ground vegetation contributed considerably more to the litter production under pine than under spruce. On sites where both Scots pine and Norway spruce are considered suitable, the latter should be selected if the aim of the forest management policy is to maximize the accumulation of SOC in the forest. Further, spruce is more favourable for SOC accumulation in areas with cold temperatures and on sites with low productivity.
  • Stendahl, Department of Soil and Environment, P.O. Box 7001, Swedish University of Agricultural Sciences, SE-75007 Uppsala, Sweden E-mail: johan.stendahl@mark.slu.se (email)
  • Johansson, Department of Soil and Environment, P.O. Box 7001, Swedish University of Agricultural Sciences, SE-75007 Uppsala, Sweden E-mail: mbj@nn.se
  • Eriksson, Department of Energy and Technology, P.O. Box 7061, Swedish University of Agricultural Sciences, SE-75007 Uppsala, Sweden E-mail: ee@nn.se
  • Nilsson, Department of Soil and Environment, P.O. Box 7001, Swedish University of Agricultural Sciences, SE-75007 Uppsala, Sweden E-mail: an@nn.se
  • Langvall, Unit for Field-based Forest Research, Asa Experimental Forest and Research Station, Swedish University of Agricultural Sciences, SE-36030 Lammhult, Sweden E-mail: ol@nn.se
article id 460, category Research article
Hong Ling, Sandhya Samarasinghe, G. Don Kulasiri. (2009). Modelling variability in full-field displacement profiles and Poisson ratio of wood in compression using stochastic neural networks. Silva Fennica vol. 43 no. 5 article id 460. https://doi.org/10.14214/sf.460
Keywords: wood; digital image correlation; displacement profiles; variability; micro structure; stochastic neural networks; Poisson ratio
Abstract | View details | Full text in PDF | Author Info
Vertical and horizontal displacement profiles in compression parallel-to-grain in a 20 x 20 mm area (30 x 21 or 630 points) in the Tangential–Longitudinal (T–L) and Radial Longitudinal (R–L) sections of small wood columns were obtained from digital image correlation applied to simultaneously captured images of the two surfaces. These consisted of 21 displacement realisations of 30 points along the length of the specimen. They revealed considerable local variations. Stochastic neural networks were successfully developed to simulate trends and noise across and along a specimen in both displacements as well as Poisson ratios in T–L and R–L sections for two selected load levels of 20kN and 40kN. These networks specifically embed noise characteristics extracted from data to generate realistic displacement and Poisson ratio realisations with inherent variability. Models were successfully validated using independent data extracted based on bootstrapping method with high accuracy with R2 ranging from 0.79 to 0.91. The models were further validated successfully using a second approach involving Confidence Intervals generated from the data extracted from the models. Models and experimental results revealed that for 20kN load, both vertical and horizontal displacements in T–L section were less heterogeneous across the specimen (smaller vertical shearing and horizontal strain, respectively) than those in the R–L section. For the 40kN load, both displacement profiles in the T–L section were less noisy and more compact than those for the 20kN load indicating less heterogeneity due to compaction of structure. In the R–L section, larger vertical shearing and horizontal strains persisted at 40 kN load. Poisson ratio decreased with load and it was nonlinear in both sections but T–L section showed much less noise across the specimen than the R–L section.
  • Ling, Centre for Advanced Computational Solutions (C-fACS), Lincoln University, Canterbury, New Zealand E-mail: hl@nn.nz
  • Samarasinghe, Centre for Advanced Computational Solutions (C-fACS), Lincoln University, Canterbury, New Zealand E-mail: sandhya.samarasinghe@lincoln.ac.nz (email)
  • Kulasiri, Centre for Advanced Computational Solutions (C-fACS), Lincoln University, Canterbury, New Zealand E-mail: gdk@nn.nz
article id 178, category Research article
Raili Hokajärvi, Teppo Hujala, Leena A. Leskinen, Jukka Tikkanen. (2009). Effectiveness of sermon policy instruments: forest management planning practices applying the activity theory approach. Silva Fennica vol. 43 no. 5 article id 178. https://doi.org/10.14214/sf.178
Keywords: forest policy; forest planning; activity theory; advisory; extension; non-industrial private forests; qualitative study
Abstract | View details | Full text in PDF | Author Info
Recent and ongoing societal changes have brought about a need to foster multiple-use forestry and to strengthen customer orientation in family forestry outreach. The study assesses how forest management planning in family forest holdings could be developed to tackle these challenges. The approach introduces a new way of evaluating the effectiveness of information- and communication-based policy instruments. Here, the cultural-historical activity theory is applied in studying the interwoven practices of present-day planning and the associated advisory services targeted at landowners. The data, comprising semi-structured in-depth interviews with 19 professional planners, were qualitatively examined, and a forest management planning activity model was constructed with the emphasis placed on the inherent contradictions of planning work. As the main contradiction, the forest and the forest owner compete as objects. The aims of making the forest productive and advising the landowners towards an increased activeness motivate forest management planning but the planners feel that they lack the opportunity to respond to the needs of the landowner. A wood-production-emphasizing interpretation of the benefits to the national economy frustrates the policy goal of genuinely promoting the goals of multiple-use forestry. The conclusion drawn is that the actors engaged in forest management planning can reveal the needs for change by discussing their opinions and practical innovations. This can be done with the aid of facilitation by e.g. researchers oriented to developmental work study.
  • Hokajärvi, Oulu University of Applied Sciences, School of Renewable Natural Resources, Metsäkouluntie 4–6, FI-90650 Oulu, Finland E-mail: raili.hokajarvi@oamk.fi (email)
  • Hujala, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: th@nn.fi
  • Leskinen, University of Eastern Finland, Department of Geographical and Historical Studies, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: lal@nn.fi
  • Tikkanen, Oulu University of Applied Sciences, School of Renewable Natural Resources, Metsäkouluntie 4–6, FI-90650 Oulu, Finland E-mail: jt@nn.fi
article id 177, category Research article
Tuomo Nurminen, Heikki Korpunen, Jori Uusitalo. (2009). Applying the activity-based costing to cut-to-length timber harvesting and trucking. Silva Fennica vol. 43 no. 5 article id 177. https://doi.org/10.14214/sf.177
Keywords: logistics; bucking; cutting; forest transport; long-distance transportation; trucking; time consumption; timber assortment; cost driver
Abstract | View details | Full text in PDF | Author Info
The supply chain of the forest industry has increasingly been adjusted to the customer’s needs for precision and quality. This has changed the operative environment both in the forest and on the roads. As the total removal of timber is increasingly divided into more log assortments, the lot size of each assortment decreases and the time consumed in sorting the logs increases. In this respect, the extra assortments have made harvesting work more difficult and affected the productivity of both cutting and forest transport; this has thus increased the harvesting costs. An activity-based cost (ABC) management system is introduced for timber harvesting and long-distance transport, based on the cut-to-length (CTL) method, in which the logistic costs are assigned to timber assortments and lots. Supplying timber is divided into three main processes: cutting, forest transport, and long-distance transportation. An ABC system was formulated separately for each of these main operations. Costs were traced to individual stands and to timber assortment lots from a stand. The cost object of the system is thus a lot of timber that makes up one assortment that has been cut, forwarded, and transported from the forest to the mill. Application of the ABC principle to timber harvesting and trucking was found to be relatively easy. The method developed gives estimates that are realistic to actual figures paid to contractors. The foremost use for this type of costing method should be as a tool to calculate the efficiency of an individual activity or of the whole logistic system.
  • Nurminen, Metsätoimisto Tuomo Nurminen, Joensuuntie 5 B 8, FI-41800 Korpilahti, Finland E-mail: tn@nn.fi (email)
  • Korpunen, Finnish Forest Research Institute, Parkano Research Unit, Kaironiementie 54, FI-39700 Parkano, Finland E-mail: hk@nn.fi
  • Uusitalo, Finnish Forest Research Institute, Parkano Research Unit, Kaironiementie 54, FI-39700 Parkano, Finland E-mail: ju@nn.fi
article id 176, category Research article
María Pasalodos-Tato, Timo Pukkala, Antonio Rigueiro-Rodríguez, Esther Fernández-Nunez, María Rosa Mosquera-Losada. (2009). Optimal management of Pinus radiata silvopastoral systems established on abandoned agricultural land in Galicia (north-western Spain). Silva Fennica vol. 43 no. 5 article id 176. https://doi.org/10.14214/sf.176
Keywords: simulation; economic profitability; optimization; risk reduction; salvage
Abstract | View details | Full text in PDF | Author Info
Timber production has been the main objective in forest production in Galicia for a long time. Nevertheless, factors such as fire risk and the need to obtain non-timber benefits make other production alternatives like silvopastoral systems worth of consideration. Integration of grazing in the production system not only diversifies products and benefits, but also decreases fire risk by enhancing fuel control. Nonetheless, few studies have examined the economic profitability of these systems. This article analyses the economics of silvopastoral systems established on abandoned agricultural soils afforested with Pinus radiata D. Don. Different tree planting densities, discounting rates, grass values and fire risk scenarios were analysed. The technique employed is based on the combination of an optimization algorithm and a simulator of stand growth and grass yield. The most profitable schedules were obtained with initial stand densities of 1500 trees per hectare. However, with high unit values of pasture production (high value of grass), schedules with an initial stand density of 500 trees per hectare were the most profitable. When the risk of fire was included in the analyses, silvopastoral systems were always more profitable than timber production systems. With an assumption that grazing reduces fire risk thinnings should be done earlier and heavier to reduce the expected losses due to fire and to promote grass production. This lengthens the pasture period. In general, rotation lengt
  • Pasalodos-Tato, INIA, Instituto Nacional de Investigación y Tecnología Agraria y Alimentaria. Madrid, Spain E-mail: pasalodos.maria@inia.es (email)
  • Pukkala, University of East Finland, Joensuu, Finland E-mail: tp@nn.fi
  • Rigueiro-Rodríguez, University of Santiago de Compostela, Lugo, Spain E-mail: arr@nn.es
  • Fernández-Nunez, University of Santiago de Compostela, Lugo, Spain E-mail: efn@nn.es
  • Mosquera-Losada, University of Santiago de Compostela, Lugo, Spain E-mail: mrml@nn.es
article id 175, category Research article
Heikki Hänninen, Jaana Luoranen, Risto Rikala, Heikki Smolander. (2009). Late termination of freezer storage increases the risk of autumn frost damage to Norway spruce seedlings. Silva Fennica vol. 43 no. 5 article id 175. https://doi.org/10.14214/sf.175
Keywords: Norway spruce; simulation; autumn frost damage; day degrees; freezer storage; growing season
Abstract | View details | Full text in PDF | Author Info
Over the last few years it has become increasingly common in artificial forest regeneration to extend the planting period by using freezer-stored seedlings for early summer plantings. Developmentally, however, planted freezer-stored seedlings lag behind seedlings planted earlier in the spring. As freezer-stored seedlings also start hardening later, they are more susceptible to early autumn frosts, especially in years when the thermal growing season ends and the first autumn frosts come earlier than usual. By means of computer simulations with a simple temperature sum model and long-term air-temperature data from three locations in Finland, we examined the effect of the freezer-storage termination date on the risk of autumn frost damage to the seedlings. The long-term simulations revealed a drastic effect of year-to-year variation in the thermal conditions during the growing season on the occurrence of autumn frost damage. Such results provide crucial information complementary to those obtained in field experiments, which are always restricted to a relatively short time period. Together with earlier field data, the present results suggest that at an average regeneration site in central Finland, the planting of seedlings whose storage has terminated on 15 June and 22 June involve autumn frost damage every tenth and every fifth year, respectively. The sensitivity analysis revealed that the temperature sum requirement of maturation has a great effect on the risk of autumn frost damage, thus pinpointing the need for experimental studies addressing this ecophysiological trait of the seedlings.
  • Hänninen, Plant Ecophysiology and Climate Change Group (PECC), Department of Biological and Environmental Sciences, Box 65, FI-00014 University of Helsinki, Finland E-mail: heikki.hanninen@helsinki.fi (email)
  • Luoranen, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: jl@nn.fi
  • Rikala, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: rr@nn.fi
  • Smolander, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: hs@nn.fi
article id 174, category Research article
Akihiro Sumida, Taro Nakai, Masahito Yamada, Kiyomi Ono, Shigeru Uemura, Toshihiko Hara. (2009). Ground-based estimation of leaf area index and vertical distribution of leaf area density in a Betula ermanii forest. Silva Fennica vol. 43 no. 5 article id 174. https://doi.org/10.14214/sf.174
Keywords: allometry; leaf area density; LAI; leaf inclination angle; MacArthur–Horn method; pipe model; Betula ermanii
Abstract | View details | Full text in PDF | Author Info
We developed a ground-based method for estimating leaf area index (LAI) and vertical distribution of leaf area density (LAD) for two Betula ermanii plots, combining an allometric method for tree leaf area with the MacArthur–Horn (MH) method using a portable laser rangefinder, including a correction for changes in leaf inclination angle along the vertical gradient measured with a portable digital protractor from a canopy access tower in each plot. Vertical distribution of projected leaf area density obtained by the MH method (LADMH) was transformed to relative distribution for allotting fixed LAI to different heights. Hence, we first developed an allometric method for estimating tree leaf area for LAI determination. Trunk cross-sectional area at branching height (AB) was accurately estimated (r2 = 0.97) from ground-based measurements of tree dimensions. We used this method to apply pipe model allometry between tree leaf area and AB, and estimated LAI (4.56 and 4.57 m2 m–2). We then examined how leaf inclination angle affected estimation of the vertical distribution of actual LAD. Leaf inclination angle measurements revealed that actual LAD in the upper canopy was 1.5–1.8-times higher than LADMH, because of steep leaf inclination, while the correction factor was 1.15–1.25 in the lower canopy. Due to the difference among heights, vertical distribution of LAD estimated with correction for vertical change in leaf inclination was more skewed to the upper canopy than that without correction. We also showed that error in LAD distribution can result if horizontal canopy heterogeneity is neglected when applying the MH method.
  • Sumida, Institute of Low Temperature Science, Hokkaido University, N19W8, Sapporo 060-0819, Japan E-mail: asumida@lowtem.hokudai.ac.jp (email)
  • Nakai, International Arctic Research Center, University of Alaska Fairbanks, 930 Koyukuk Drive, P.O. Box 757340, Fairbanks, Alaska 99775-7340, USA E-mail: tn@nn.jp
  • Yamada, International Meteorological & Oceanographic Consultants Co., Ltd. Kawaguchi-cho 2-6528-87, Choshi, Chiba 288-0001, Japan E-mail: my@nn.jp
  • Ono, Institute of Low Temperature Science, Hokkaido University, N19W8, Sapporo 060-0819, Japan E-mail: ko@nn.jp
  • Uemura, Field Science Center for Northern Biosphere, Hokkaido University, Tokuda 250, Nayoro, Hokkaido 096-0071, Japan E-mail: su@nn.jp
  • Hara, Institute of Low Temperature Science, Hokkaido University, N19W8, Sapporo 060-0819, Japan E-mail: th@nn.jp
article id 173, category Research article
Erik Hellberg, Torbjörn Josefsson, Lars Östlund. (2009). The transformation of a Norway spruce dominated landscape since pre-industrial times in northern Sweden: the influence of modern forest management on forest structure. Silva Fennica vol. 43 no. 5 article id 173. https://doi.org/10.14214/sf.173
Keywords: birch; disturbance dynamics; forest history; deciduous forest; fire
Abstract | View details | Full text in PDF | Author Info
Logging history and the study of reference conditions in Scandinavian boreal forests has tended to focus on Scots pine dominated ecosystems. This paper presents a regional study of pre-industrial forest conditions and examines the effects of the industrial exploitation of ecosystems dominated by Norway spruce in northern Sweden. Historical records covering a period which preceded industrial logging in the study area (1917–1927) were used to obtain quantitative data on forest structure and influence of forest fires. These data were compared with a modern data set (2003) to analyse changes due to the industrial transformation of the forest. The early 20th century landscape was dominated by old, multi-cohorted spruce forests and mixed coniferous forests. It was found that fire affected both the structure and composition of the landscape. In post-burnt areas, even-aged forests dominated by deciduous species were the principal forest type. Between the early and modern data sets, profound changes in tree-species composition and age structure were documented. While the total volume of deciduous species increased substantially, the coverage of forests dominated by deciduous species decreased. There was also a significant increase in pine-dominated forests and in the total volume of pine. The industrial transformation of the studied landscapes has had profound effects on the structure of spruce forests, but much less so on deciduous forests. The study concludes that the present forest structure is a function of past management regimes, and that future transformations of the landscape will continue, thus affecting the natural variability and biodiversity of the forests.
  • Hellberg, Tunstigen 10, SE-831 43 Östersund, Sweden E-mail: eh@nn.se
  • Josefsson, Swedish University of Agricultural Sciences, Department of Forest Ecology and Management, SE-901 83 Umeå, Sweden E-mail: torbjorn.josefsson@svek.slu.se (email)
  • Östlund, Swedish University of Agricultural Sciences, Department of Forest Ecology and Management, SE-901 83 Umeå, Sweden E-mail: lo@nn.se
article id 172, category Research article
Guntis Brumelis, Maris Strazds, Zanna Eglava. (2009). Stand structure and spatial pattern of regeneration of Pinus sylvestris in a natural treed mire in Latvia. Silva Fennica vol. 43 no. 5 article id 172. https://doi.org/10.14214/sf.172
Keywords: Pinus sylvestris; regeneration; natural forest; mire
Abstract | View details | Full text in PDF | Author Info
We examined the regeneration patterns of Pinus sylvestris L. in a natural treed mire in the hemiboreal zone in Latvia. Data on tree stem age and size was collected in 207 fine-scale plots (10 m2) and 4 medium-scale plots (400 m2). Size structure of living and dead trees was also estimated on transects 180-m and 250-m length and 10-m width. In addition, the vegetation was described in 1-m2 plots to determine preferred microsites for P. sylvestris establishment. Pinus sylvestris showed continuous regeneration by an inverse J-shaped age and size structure. Pulses of mortality induced by fire were also evident. Regeneration of P. sylvestris was mostly on Sphagnum magellanicam Brid. hummocks free of competition from Ledum palustre L. and graminoids, which were found in small interspersed patches throughout the mire. The spatial pattern of tree cohorts differed, probably due to changing moisture conditions altering seedbed conditions and by fire-induced mortality in specific areas. Recent rapid invasion by Betula pubescens Ehrh. and Picea abies (L.) Karst. in a post-fire period since 1971 is probably associated with drier conditions.
  • Brumelis, Faculty of Biology, University of Latvia, Kronvalda bulv. 4, Riga LV1586, Latvia E-mail: guntis.brumelis@lu.lv (email)
  • Strazds, Faculty of Biology, University of Latvia, Kronvalda bulv. 4, Riga LV1586, Latvia E-mail: ms@nn.lv
  • Eglava, Faculty of Biology, University of Latvia, Kronvalda bulv. 4, Riga LV1586, Latvia E-mail: ze@nn.lv
article id 171, category Research article
Takuo Nagaike. (2009). Snag abundance and species composition in a managed forest landscape in central Japan composed of Larix kaempferi plantations and secondary broadleaf forests. Silva Fennica vol. 43 no. 5 article id 171. https://doi.org/10.14214/sf.171
Keywords: snags; long-rotation plantation; plantation age; species composition
Abstract | View details | Full text in PDF | Author Info
Larix kaempferi is the main plantation species in the low-snow, cool-temperate zone of Japan. I studied L. kaempferi plantations of various stand ages in central Japan to examine and compare the effect of stand age on the abundance, size, and species composition of snags (standing dead trees) compared to those in secondary broadleaf forests. Plantations that were older than the standard rotation age had more and larger snags than young plantations, and the species diversity of snags was positively correlated with stand age. Because the density of living planted L. kaempferi showed little correlation with snag variables, whereas that of naturally regenerated tree species was positively correlated with snag variables, the density dependence of snag occurrence was stronger in naturally regenerated trees than in planted L. kaempferi. Snag species that were positively correlated with stand age were the main species in secondary broadleaf forests in this area. Basal area, density, and number of species of snags in standard-rotation plantations were significantly lower than in long-rotation plantations and secondary broadleaf forests. Long-rotation plantations are useful for retaining snags compared to standard-rotation plantations.
  • Nagaike, Yamanashi Forest Research Institute, Masuho, Yamanashi 400-0502, Japan E-mail: nagaike-zty@pref.yamanashi.lg.jp (email)
article id 170, category Research article
Ryoichi Doi, Senaratne L. Ranamukhaarachchi. (2009). Community-level physiological profiling in monitoring rehabilitative effects of Acacia auriculiformis plantation on degraded land in Sakaerat, Thailand. Silva Fennica vol. 43 no. 5 article id 170. https://doi.org/10.14214/sf.170
Keywords: multivariate analysis; Acacia auriculiformis; community-level physiological profile; land degradation and rehabilitation; soil bacterial community
Abstract | View details | Full text in PDF | Author Info
This study was conducted to investigate the rehabilitative effects of planting Acacia auriculiformis trees on degraded land by observing variations in soil bacterial community profiles provided by community-level physiological profiling. Soil bacterial and physicochemical comparisons between an original evergreen forest and the Acacia plantation plot, established on an area severely degraded as a result of deforestation, showed that most soil characteristics were rehabilitated 18 to 19 years after the plantation of Acacia according to single variables, Shannon and Simpson diversity indices based on the community-level physiological profiles, principal component analysis and redundancy analysis. However, a more strict statistical comparison, discriminant analysis, completely discriminated between the Acacia plantation and the evergreen forest soils when the community-level physiological profiles were compared. Thus, the Acacia plantation soil was shown to still be in the process to full recovery. Here, we discuss the relevance of planting A. auriculiformis in land rehabilitation schemes in savanna regions.
  • Doi, AFE Building, School of Environment, Resources and Development, Asian Institute of Technology, Pathumthani 12120, Thailand E-mail: roird@aeiou.pt (email)
  • Ranamukhaarachchi, AFE Building, School of Environment, Resources and Development, Asian Institute of Technology, Pathumthani 12120, Thailand E-mail: slr@nn.th
article id 169, category Research article
Susanne von Bargen, Elise Grubits, Risto Jalkanen, Carmen Büttner. (2009). Cherry leaf roll virus – an emerging virus in Finland? Silva Fennica vol. 43 no. 5 article id 169. https://doi.org/10.14214/sf.169
Keywords: Betula pendula; Betula pubescens; Betula nana; IC-RT-PCR; 3' non-coding region; CLRV; phylogenetic relationship
Abstract | View details | Full text in PDF | Author Info
Cherry leaf roll virus, CLRV, is a plant pathogen that infects a variety of deciduous trees and shrubs in temperate regions. Little is known about its occurrence at high latitudes and especially in Finnish birch species. Still, symptoms that seemed to be associated with CLRV such as vein banding, leaf roll and decline have been observed in birch trees throughout the country since the summer of 2002. Six different birch species, subspecies or varieties, i.e. Betula pubescens subsp. pubescens (downy birch), B. pendula (silver birch), B. nana (dwarf birch), B. pubescens var. appressa (Kiilopää birch), B. pubescens subsp. czerepanovii (mountain birch) and B. pendula var. carelica (curly birch) originating from all over Finland were assessed by immunocapture-reverse transcription-polymerase chain reaction (IC-RT-PCR) for CLRV infection. It was shown that CLRV is widely distributed in B. pendula and B. pubescens throughout the country. Furthermore, dwarf birch, mountain birch, Kiilopää birch and curly birch were confirmed to be previously unkown hosts of CLRV. Genetic analysis of virus sequence variants originating from Finnish birch trees revealed atypical phylogenetic relationships. In contrast to CLRV isolates from birches growing in the United Kingdom and Germany which clustered exclusively within group A, Finnish CLRV isolates belonged either to group B, D or E. Thus, virus population structure in Finnish birches seems to be more variable and host plant dependency seems not to apply for Finnish CLRV isolates.
  • Bargen, Humboldt-Universität zu Berlin, Department für Nutzpflanzen- und Tierwissenschaften, Fachgebiet Phytomedizin, Lentzeallee 55/57, 14195 Berlin, Germany E-mail: susanne.von.bargen@agrar.hu-berlin.de (email)
  • Grubits, Humboldt-Universität zu Berlin, Department für Nutzpflanzen- und Tierwissenschaften, Fachgebiet Phytomedizin, Lentzeallee 55/57, 14195 Berlin, Germany E-mail: eg@nn.de
  • Jalkanen, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: risto.jalkanen@metla.fi
  • Büttner, Humboldt-Universität zu Berlin, Department für Nutzpflanzen- und Tierwissenschaften, Fachgebiet Phytomedizin, Lentzeallee 55/57, 14195 Berlin, Germany E-mail: cb@nn.de
article id 168, category Research article
Saila Varis, Anne Pakkanen, Aina Galofré, Pertti Pulkkinen. (2009). The extent of south-north pollen transfer in Finnish Scots pine. Silva Fennica vol. 43 no. 5 article id 168. https://doi.org/10.14214/sf.168
Keywords: Pinus sylvestris; gene flow; adaptation; coniferous phenology; plant population biology; boreal forest dynamics
Abstract | View details | Full text in PDF | Author Info
In order to evaluate the possibility of long distance gene flow in Scots pine (Pinus sylvestris L.), we measured the amount and germinability of airborne pollen and flowering phenology in central, northern, and northernmost Finland during 1997–2000. Totally 2.3% of the detected germinable pollen grains were in the air prior to local pollen shedding. The mean number of germinable pollen grains m–3 air per day was lower prior to local pollen shedding, but in the year 2000 there were more germinable pollen grains in the air of central study site prior to local pollen shedding. Prior to the onset of pollen shedding, 7.5% of female strobili which we observed were receptive. On average female strobili became receptive three days earlier than local pollen shedding started. During the period of pollen shedding in the central study site, we detected germinable airborne pollen in the northern site in years 1997, 1999 and 2000. At the northermost site, we detected germinable airborne pollen during the pollen-shedding period of the northern site in 2000. Our detection of germinable airborne pollen and synchrony of strobili maturation from south to north suggest that populations of Scots pine in central and northern Finland may provide genetic material to populations in northern and northernmost Finland, respectively.
  • Varis, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: saila.varis@metla.fi (email)
  • Pakkanen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: ap@nn.fi
  • Galofré, Passeig de l’estació 21, 5-1, 43800 Valls, Tarragona, Spain E-mail: ag@nn.fi
  • Pulkkinen, Finnish Forest Research Institute, Haapastensyrjä Breeding Station, Karkkilantie 247, FI-12600 Läyliäinen, Finland E-mail: pp@nn.fi
article id 190, category Research article
Katri Luostarinen, Veikko Möttönen. (2009). Effect of felling season, storage and drying on colour of silver birch (Betula pendula) wood from four different growing sites. Silva Fennica vol. 43 no. 4 article id 190. https://doi.org/10.14214/sf.190
Keywords: birch; age; colour darkening; conventional drying; environmental factors; plantation; wood
Abstract | View details | Full text in PDF | Author Info
Darkening of birch wood during artificial drying is a significant problem regarding the use of its timber as raw material by the mechanical wood industry. In the future, an increasing proportion of birch timber will be obtained from plantation forests, which differ from natural forests in many respects. In this investigation sample boards of Betula pendula, both from naturally regenerated stands and plantations, were sawn into the dimensions used as raw material for parquet billets. Growing site, felling season, and storage of logs were taken into account as possible factors affecting wood colour changes during drying. The wood of birches from fertile plantations remained lighter-coloured during conventional drying than the wood of naturally regenerated birches from low- and medium-fertile stands. The reason may be the difference in tree age and growth rate between natural and planted stands. Thus, it could be beneficial to grow birch in fertile stands so that the trees reach log size as young as possible. The results of this study emphasise the good quality of the birch wood from planted stands compared to natural stands with regard to its colour.
  • Luostarinen, Faculty of Forest Sciences, University of Joensuu, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: katri.luostarinen@joensuu.fi (email)
  • Möttönen, Department of Mechanical Engineering, Lappeenranta University of Technology, P.O. Box 20, FI-53851 Lappeenranta, Finland E-mail: vm@nn.fi
article id 189, category Research article
Lars Rytter, Gunnar Jansson. (2009). Influence of pruning on wood characters in hybrid aspen. Silva Fennica vol. 43 no. 4 article id 189. https://doi.org/10.14214/sf.189
Keywords: Populus tremula P. tremuloides; cicatrisation; faultless wood; twig angle; twig diameter
Abstract | View details | Full text in PDF | Author Info
Fast-growing hybrid aspens (Populus tremula L. x P. tremuloides Michx.) are currently of great interest in Sweden since they can produce biomass at high rates and, at the same time, can be used to produce higher value wood products. This study focuses on the effects of pruning hybrid aspen to improve its wood quality. About 50% of the trees in the experimental stand were pruned by removing twigs, at heights up to 4 m, when they were 7–8 years old. Ten years later, 20 pruned and 20 unpruned trees, representing four clones, were randomly selected. Ten knots or twig/stem junctions, respectively, per tree were exposed for inspection using a chain saw and examined. The results revealed that pruned trees cicatrised the knots within about three years and thereafter produced substantial amounts of faultless wood. In contrast, unpruned trees (which had retained almost 80% of their twigs, often as dry twigs with bark pockets) had produced small uneven amounts of quality wood. Removal of twigs with acute angles and/or large diameters resulted in greater colour defects and rot in annual rings outside the pruning position, but the time of cicatrisation was not significantly affected. The results show that pruning can be used to enhance the wood quality of hybrid aspen over a short time period, and that pruning should be performed early during the rotation period when branches are small, in order to minimize discolouration and rot in the new annual rings.
  • Rytter, The Forestry Research Institute of Sweden (Skogforsk), Ekebo 2250, SE-26890 Svalöv E-mail: lars.rytter@skogforsk.se (email)
  • Jansson, The Forestry Research Institute of Sweden (Skogforsk), Uppsala Science Park, SE-751 83 Uppsala E-mail: gj@nn.se
article id 188, category Research article
Mikko Havimo, Juha Rikala, Jari Sirviö, Marketta Sipi. (2009). Tracheid cross-sectional dimensions in Scots pine (Pinus sylvestris) – distributions and comparison with Norway spruce (Picea abies). Silva Fennica vol. 43 no. 4 article id 188. https://doi.org/10.14214/sf.188
Keywords: Norway spruce; Scots pine; cell wall thickness; radial diameter; tangential diameter; distribution
Abstract | View details | Full text in PDF | Author Info
Cell wall thickness and tracheid radial and tangential diameter are important characteristics in papermaking. These fibre cross-sectional dimensions affect paper properties such as light scattering, and tear and tensile indexes. In the authors’ previous article, the mean values and distributions of tracheid cross-sectional dimensions were obtained for Norway spruce (Picea abies). This article characterises the cross-sectional tracheid properties of Scots pine (Pinus sylvestris) using exactly the same methodology as in the previous study on Norway spruce, which enables the comparison between the tree species. The distributions for Scots pine cell wall thickness and tracheid radial diameter were similar: a narrow peak due to earlywood tracheids, and a wide peak due to latewood tracheids. The tangential diameter distributions for Scots pine were very similar in both earlywood and latewood, having one wide peak. Also, the distributions in whole stem, top pulpwood and sawmill chip assortments were quite similar. The differences between Scots pine and Norway spruce tracheid cross-sectional dimensions were fairly marginal. This is at least the case when comparing large tracheid populations, in which differences tend to even out. The situation may be different on a more detailed level of observation, for example, when individual annual rings in the different tree species are compared.
  • Havimo, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: mikko.havimo@helsinki.fi (email)
  • Rikala, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: jr@nn.fi
  • Sirviö, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: js@nn.fi
  • Sipi, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ms@nn.fi
article id 187, category Research article
Tobias Biechele, Leif Nutto, Gero Becker. (2009). Growth strain in Eucalyptus nitens at different stages of development. Silva Fennica vol. 43 no. 4 article id 187. https://doi.org/10.14214/sf.187
Keywords: growth stress; growth strain; Eucalyptus nitens; stages of development; sawnwood
Abstract | View details | Full text in PDF | Author Info
Eucalypts are renowned for their high growth stress levels. These stresses cause splitting, warping and dimensional instability when cutting, processing and drying the wood. In Chile, large Eucalyptus nitens plantations can be found, which, due to these problems, are scarcely utilised for solid wood products (veneer, sawn wood). This study aims to determine the factors influencing growth stress at different stages of tree’s development, and to identify whether the factors influencing growth stress change over time. In five stands of different ages, growth strain, as an indicator of growth stress, was measured at different tree heights with the single hole drilling method. The tree variables DBH, tree height, slenderness (height/diameter ratio) and crown parameters also were measured. A correlation analysis of tree variables and growth strains was undertaken. The results obtained indicate a high variability in growth strain values. It was concluded that growth strain is not correlated with a single growth parameter, but with a combination of factors that variously influence it at different ages and tree heights.
  • Biechele, University of Freiburg, Institute of Forest Utilisation and Work Science, Werthmannstr. 6, DE-79085 Freiburg, Germany E-mail: tobias.biechele@fobawi.uni-freiburg.de (email)
  • Nutto, University of Freiburg, Institute of Forest Utilisation and Work Science, Werthmannstr. 6, DE-79085 Freiburg, Germany E-mail: ln@nn.de
  • Becker, University of Freiburg, Institute of Forest Utilisation and Work Science, Werthmannstr. 6, DE-79085 Freiburg, Germany E-mail: gb@nn.de
article id 186, category Research article
Juho Rantala, Pertti Harstela, Veli-Matti Saarinen, Leo Tervo. (2009). A techno-economic evaluation of Bracke and M-planter tree planting devices. Silva Fennica vol. 43 no. 4 article id 186. https://doi.org/10.14214/sf.186
Keywords: productivity; cost-efficiency; mechanization; planting cost
Abstract | View details | Full text in PDF | Author Info
Techno-economically reasonable mechanization of tree planting has proved to be a difficult task in the Nordic working conditions. Although planting machines and combinations of base machine and planting device have been developed since the 1970s, mechanized planting has not been cost-competitive to manual planting. The aim of this study was to find out work time distributions, productivities, costs and effects of different work difficulty factors on productivities and costs of the state-of-the-art Nordic planting devices, Swedish Bracke and Finnish M-Planter, and to compare the devices with each other. The theory of comparative time studies was the base for the experimental design of this study. In the average working conditions, productivity (E15) of M-Planter (236 seedlings/hour) was 36.0% higher than that of Bracke (174 seedlings/hour). Here, M-Planter performed planting work 23.4% cheaper than Bracke. However, the difference depended greatly on the working conditions; the more stones or stumps the smaller the difference, and the more slash the bigger the difference.
  • Rantala, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: juho.rantala@metla.fi (email)
  • Harstela, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: ph@nn.fi
  • Saarinen, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: vms@nn.fi
  • Tervo, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: lt@nn.fi
article id 185, category Research article
Bianca N. I. Eskelson, Tara M. Barrett, Hailemariam Temesgen. (2009). Imputing mean annual change to estimate current forest attributes. Silva Fennica vol. 43 no. 4 article id 185. https://doi.org/10.14214/sf.185
Keywords: forest inventory and analysis; forest monitoring; national forest inventories; nearest neighbor imputation; Pacific Northwest; paneled inventory data
Abstract | View details | Full text in PDF | Author Info
When a temporal trend in forest conditions is present, standard estimates from paneled forest inventories can be biased. Thus methods that use more recent remote sensing data to improve estimates are desired. Paneled inventory data from national forests in Oregon and Washington, U.S.A., were used to explore three nearest neighbor imputation methods to estimate mean annual change of four forest attributes (basal area/ha, stems/ha, volume/ha, biomass/ha). The randomForest imputation method outperformed the other imputation approaches in terms of root mean square error. The imputed mean annual change was used to project all panels to a common point in time by multiplying the mean annual change with the length of the growth period between measurements and adding the change estimate to the previously observed measurements of the four forest attributes. The resulting estimates of the mean of the forest attributes at the current point in time outperformed the estimates obtained from the national standard estimator.
  • Eskelson, Oregon State University, Department of Forest Engineering, Resources and Management, 204 Peavy Hall, Corvallis, Oregon 97331, USA E-mail: bianca.eskelson@oregonstate.edu (email)
  • Barrett, Oregon State University, Department of Forest Engineering, Resources and Management, 204 Peavy Hall, Corvallis, Oregon 97331, USA E-mail: tmb@nn.us
  • Temesgen, Oregon State University, Department of Forest Engineering, Resources and Management, 204 Peavy Hall, Corvallis, Oregon 97331, USA E-mail: ht@nn.us
article id 184, category Research article
Jaakko Repola. (2009). Biomass equations for Scots pine and Norway spruce in Finland. Silva Fennica vol. 43 no. 4 article id 184. https://doi.org/10.14214/sf.184
Keywords: pine; tree biomass; biomass equations; spruce
Abstract | View details | Full text in PDF | Author Info
In this study, biomass equations for the above- and below-ground tree components of Scots pine (Pinus sylvestris) and Norway spruce (Picea abies [L.] Karst.) were developed. The models were based on 908 pine trees and 613 spruce trees collected in 77 stands located on mineral soil, and represented a wide range of stand and site conditions in Finland. The whole data set consisted of three sub data sets: 33 temporary sample plots, five thinning experiments, and the control plots of 39 fertilization experiments. The biomass equations were estimated for the individual tree components: stem wood, stem bark, living and dead branches, needles, stump, and roots. In the data analysis, a multivariate procedure was applied in order to take into account the statistical dependence among the equations. Three multivariate models for above-ground biomass and one for below-ground biomass were constructed. The multivariate model (1) was mainly based on tree diameter and height, and additional commonly measured tree variables were used in the multivariate models (2) and (3). Despite the unbalanced data in terms of the response variables, the statistical method generated equations that enable more flexible application of the equations, and ensure better biomass additivity compared to the independently estimated equations. The equations provided logical biomass predictions for a number of tree components, and were comparable with other functions used in Finland and Sweden even though the study material was not an objective, representative sample of the tree stands in Finland.
  • Repola, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: jaakko.repola@metla.fi (email)
article id 183, category Research article
Stella Bogino, María José Fernández Nieto, Felipe Bravo. (2009). Climate effect on radial growth of Pinus sylvestris at its southern and western distribution limits. Silva Fennica vol. 43 no. 4 article id 183. https://doi.org/10.14214/sf.183
Keywords: Scots pine; dendroclimatology; tree ring; pulse and interpulse
Abstract | View details | Full text in PDF | Author Info
The associations between tree-ring width and climatic variables, the tree age effect on climate-growth relationship and the drought index impact on radial growth of Scots pine (Pinus sylvestris L.) were studied in Mediterranean environments in the Iberian Peninsula. Dendrochronological techniques were applied to construct six tree-ring width chronologies for P. sylvestris. The association between tree growth and climate was analysed with correlation coefficients and bootstrapped response functions. Drought index (DRI) was used to detect the effects of aridity. Tree-ring width was positively correlated with rainfall in the growing season but this association stated earlier at the lowest altitudinal sites. The effects of temperature varied according to the site. The response to climatic variables depended on the age of the trees: more of the variability was explained by climatic variables in young stands than in old stands. The significant association between radial growth and DRI suggests that drought is a determining factor in the radial growth of P. sylvestris. Climate forecast scenarios show an increase in rainfall irregularity in the Mediterranean region so the differential tree response to rainfall at different elevations can be used to predict tree growth for determining silvicultural treatments.
  • Bogino, Universidad de Valladolid, Departamento de Producción Vegetal y Recursos Forestales, Joint Unit UVa-INIA ‘Sustainable Forest Management’, Avda. de Madrid 44, 34004 Palencia, Spain E-mail: sbogino@fices.unsl.edu.ar (email)
  • Fernández Nieto, Universidad de Valladolid, Departamento de Producción Vegetal y Recursos Forestales, Joint Unit UVa-INIA ‘Sustainable Forest Management’, Avda. de Madrid 44, 34004 Palencia, Spain E-mail: mjfn@nn.es
  • Bravo, Universidad de Valladolid, Departamento de Producción Vegetal y Recursos Forestales, Joint Unit UVa-INIA ‘Sustainable Forest Management’, Avda. de Madrid 44, 34004 Palencia, Spain E-mail: fb@nn.es
article id 182, category Research article
Janne Miettinen, Pekka Helle, Ari Nikula, Pekka Niemelä. (2009). Changes in landscape-scale habitat selection of capercaillie (Tetrao urogallus) in managed north-boreal forest. Silva Fennica vol. 43 no. 4 article id 182. https://doi.org/10.14214/sf.182
Keywords: forestry; landscape composition; capercaillie; wildlife triangle
Abstract | View details | Full text in PDF | Author Info
We studied changes in habitat selection of capercaillie in northern Finnish boreal forests at two spatial scales during two time periods, 1989–1992 and 2000–2003. We studied capercaillie densities and their changes between the study periods in relation to the landscape class proportions within 3-km buffer zones around the wildlife triangle center points. Furthermore, we compared the landscape class proportions in 800-meter buffer zones around capercaillie wildlife triangle count observations and around the counted wildlife triangle transects using t-tests and compositional analysis. At the local population scale (3 km) the change in adult density between the study periods was associated positively with the proportion of young thinning stands in 2003 and reversely with the mature stand (1992 and 2003) and clear-cut (1992) proportions. Capercaillie juvenile and pooled densities during 2000–2003 were positively associated with the advanced thinning stand proportion in 2003. At the capercaillie home range scale (800 m) habitats were rich in mature stands during 1989–1992 in relation to available habitats, but not during 2000–2003 when young thinning stands were more abundant in relation to available habitats. Relatively young managed forests can be suitable for capercaillie, but mature managed forests as capercaillie habitats may have deteriorated between the study periods. Spatial planning may help to form suitable areas that are large enough for the species, but the highest potential may lay in the forest stand scale, where increased cover on the ground could promote the habitat quality.
  • Miettinen, Kankurinhaka 14, FI-90450 Kempele, Finland E-mail: janne.miettinen@rktl.fi (email)
  • Helle, Finnish Game and Fisheries Research Institute, Tutkijantie 2 E, FI-90570 Oulu, Finland E-mail: ph@nn.fi
  • Nikula, Finnish Forest Research Institute, Rovaniemi Research Unit, Eteläranta 55, FI-99600 Rovaniemi, Finland E-mail: an@nn.fi
  • Niemelä, University of Turku, Department of Biology, FI-20014 University of Turku, Finland E-mail: pn@nn.fi
article id 181, category Research article
Jari Miina, Juha-Pekka Hotanen, Kauko Salo. (2009). Modelling the abundance and temporal variation in the production of bilberry (Vaccinium myrtillus L.) in Finnish mineral soil forests. Silva Fennica vol. 43 no. 4 article id 181. https://doi.org/10.14214/sf.181
Keywords: vegetation; generalized linear mixed model; heath forest
Abstract | View details | Full text in PDF | Author Info
  • Miina, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jari.miina@metla.fi (email)
  • Hotanen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jph@nn.fi
  • Salo, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: ks@nn.fi
article id 180, category Research article
Hongzhang Kang, Björn Berg, Chunjiang Liu, Carl J. Westman. (2009). Variation in mass-loss rate of foliar litter in relation to climate and litter quality in Eurasian forests: differences among functional groups of litter. Silva Fennica vol. 43 no. 4 article id 180. https://doi.org/10.14214/sf.180
Keywords: carbon sequestration; climate; Eurasian forests; leaf litter; mass-loss; litter quality
Abstract | View details | Full text in PDF | Author Info
With a data set of litter decomposition collected by means of literature survey, our objectives are 1) to determine the differences in the variation in the first-year mass loss (%) of leaf litter with regard to climate and litter quality among different functional groups of tree species in Eurasian forests, and 2) to determine the difference in effect of mean annual temperature (°C), annual precipitation (dm), as well as concentration of nitrogen (%), and lignin (%) on first-year mass loss over a wide range in climate and litter quality. The main results are as follows. 1) The significant differences between litter types in the relationships between first-year mass loss and climatic factors plus litter quality revealed clearly different decomposition patterns over the continent. Thus, differences were found between coniferous and broadleaf litter, between deciduous broadleaf and evergreen broadleaf as well as between genera and even within a genus, viz. between deciduous and evergreen Quercus. 2) With a change in a relative unit of climate and litter quality variables, there were clear differences in effects of mean annual temperature, annual precipitation, and nitrogen on first-year mass loss for different functional groups of trees. 3) We identified some broadleaf litter species that decomposed to 100% in one year and thus did not contribute to carbon sequestration in a humus layer. Thus, the variation in pattern of foliar litter decomposition with climate and litter quality across functional groups in Eurasian forests showed different decomposition strategies for litter of different groups and genera.
  • Kang, School of Agriculture and Biology, Shanghai Jiao Tong University, Dongchuan Rd. 800, Shanghai 200240, P. R. China E-mail: hk@nn.cn
  • Berg, Department of Forest Ecology, University of Helsinki, Latokartanonkaari 7, FIN-00014 Finland; Dipartimento Biologia Strutturale e Funzionale. Complesso Universitario, Monte S. Angelo, Via Cinthia, IT-80126 Napoli, Italy E-mail: bb@nn.fi
  • Liu, School of Agriculture and Biology, Shanghai Jiao Tong University, Dongchuan Rd. 800, Shanghai 200240, P. R. China; Key Laboratory of Urban Agriculture (South), Ministry of Agriculture, P. R. China, Shanghai Jiao Tong University, 800 Dongchuan Rd., Shanghai 200240, P. R. China E-mail: chjliu@sjtu.edu.cn (email)
  • Westman, Dipartimento Biologia Strutturale e Funzionale. Complesso Universitario, Monte S. Angelo, Via Cinthia, IT-80126 Napoli, Italy E-mail: cjw@nn.it
article id 179, category Research article
Henna Vartiamäki, Jarkko Hantula, Antti Uotila. (2009). Susceptibility of silver birch pruning wounds to infection by white-rot fungus (Chondrostereum purpureum), a potential bioherbicide. Silva Fennica vol. 43 no. 4 article id 179. https://doi.org/10.14214/sf.179
Keywords: birch; vegetation management; biological control; fungal decay; pruning
Abstract | View details | Full text in PDF | Author Info
We artificially inoculated pruning wounds of silver birch (Betula pendula Roth) to study seasonal variation in their vulnerability to infection by the fungal decomposer Chondrostereum purpureum (Pers. ex Fr.) Pouzar. This information is critical to the assessment of incidental infection risks in areas where C. purpureum may be used as a bioherbicide. On seven monthly occasions between April and October 2005, 30 birch trees were pruned to yield a total of 210 experimental trees. On each occasion, 10 trees were inoculated immediately with C. purpureum mycelium, 10 were inoculated with blank inoculum and 10 were only pruned. In the summer of 2007, a survey of 129 experimental trees showed that pruning wounds were most susceptible to infection during May. Treatment with C. purpureum at other times during the growing season also increased the extent of discoloration or decay but the effect was considerably less.
  • Vartiamäki, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: henna.vartiamaki@metla.fi (email)
  • Hantula, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: jh@nn.fi
  • Uotila, University of Helsinki, Hyytiälä Forestry Field Station, Hyytiäläntie 124, FI-35500 Korkeakoski, Finland E-mail: au@nn.fi
article id 204, category Research article
Gaby Deckmyn, Bostjan Mali, Hojka Kraigher, Niko Torelli, Maarten Op de Beeck, Reinhart Ceulemans. (2009). Using the process-based stand model ANAFORE including Bayesian optimisation to predict wood quality and quantity and their uncertainty in Slovenian beech. Silva Fennica vol. 43 no. 3 article id 204. https://doi.org/10.14214/sf.204
Keywords: forest management; wood quality; Bayesian calibration; beech; forest model; mechanistic; red heart
Abstract | View details | Full text in PDF | Author Info
The purpose of this study was to expand an existing semi-mechanistic forest model, ANAFORE (ANAlysing Forest Ecosystems), to allow for the prediction of log quality and the accompanying uncertainty as influenced by climate and management. The forest stand is described as consisting of trees of different cohorts, either of the same or of different species (deciduous or coniferous). In addition to photosynthesis, transpiration, total growth and yield, the model simulates the daily evolution in vessel biomass and radius, parenchyma and branch development. From these data early and latewood biomass, wood tissue composition, knot formation and density are calculated. The new version presented here, includes the description of log quality, including red heart formation of beeches. A Bayesian optimisation routine for the species parameters was added to the stand model. From a given range of input parameters (prior), the model calculates an optimised range for the parameters (posterior) based on given output data, as well as an uncertainty on the predicted values. A case study was performed for Slovenian beech forests to illustrate the main model functioning and more in particular the simulation of the wood quality. The results indicate that the ANAFORE model is a useful tool for analyzing wood quality development and forest ecosystem functioning in response to management, climate and stand characteristics. However, the Bayesian optimization showed that the remaining uncertainty on the input parameters for the chosen stand was very large, due to the large number of input parameters in comparison to the limited stand data.
  • Deckmyn, Research Group Plant and Vegetation Ecology, University of Antwerpen, Universiteitsplein 1, 2610 Antwerpen, Belgium E-mail: gaby.deckmyn@ua.ac.be (email)
  • Mali, Slovenian Forestry Institute, Vecna pot 2, 1000 Ljubljana, Slovenia E-mail: bm@nn.si
  • Kraigher, Slovenian Forestry Institute, Vecna pot 2, 1000 Ljubljana, Slovenia E-mail: hk@nn.si
  • Torelli, Slovenian Forestry Institute, Vecna pot 2, 1000 Ljubljana, Slovenia E-mail: nt@nn.si
  • Op de Beeck, Research Group Plant and Vegetation Ecology, University of Antwerpen, Universiteitsplein 1, 2610 Antwerpen, Belgium E-mail: modb@nn.be
  • Ceulemans, Research Group Plant and Vegetation Ecology, University of Antwerpen, Universiteitsplein 1, 2610 Antwerpen, Belgium E-mail: rc@nn.be
article id 203, category Research article
Matti Maltamo, Jussi Peuhkurinen, Jukka Malinen, Jari Vauhkonen, Petteri Packalén, Timo Tokola. (2009). Predicting tree attributes and quality characteristics of Scots pine using airborne laser scanning data. Silva Fennica vol. 43 no. 3 article id 203. https://doi.org/10.14214/sf.203
Keywords: LIDAR; alpha shape; crown height; height metrics; k-MSN; timber quality
Abstract | View details | Full text in PDF | Author Info
The development of airborne laser scanning (ALS) during last ten years has provided new possibilities for accurate description of the living tree stock. The forest inventory applications of ALS data include both tree and area-based plot level approaches. The main goal of such applications has usually been to estimate accurate information on timber quantities. Prediction of timber quality has not been focused to the same extent. Thus, in this study we consider here the prediction of both basic tree attributes (tree diameter, height and volume) and characteristics describing tree quality more closely (crown height, height of the lowest dead branch and sawlog proportion of tree volume) by means of high resolution ALS data. The tree species considered is Scots pine (Pinus sylvestris), and the field data originate from 14 sample plots located in the Koli National Park in North Karelia, eastern Finland. The material comprises 133 trees, and size and quality variables of these trees were modeled using a large number of potential independent variables calculated from the ALS data. These variables included both individual tree recognition and area-based characteristics. Models for the dependent tree characteristics to be considered were then constructed using either the non-parametric k-MSN method or a parametric set of models constructed simultaneously by the Seemingly Unrelated Regression (SUR) approach. The results indicate that the k-MSN method can provide more accurate tree-level estimates than SUR models. The k-MSN estimates were in fact highly accurate in general, the RMSE being less than 10% except in the case of tree volume and height of the lowest dead branch.
  • Maltamo, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: matti.maltamo@joensuu.fi (email)
  • Peuhkurinen, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: jp@nn.fi
  • Malinen, Finnish Forest Research Institute, Joensuu Research Unit, FI-80101 Joensuu, Finland E-mail: jm@nn.fi
  • Vauhkonen, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: jv@nn.fi
  • Packalén, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: pp@nn.fi
  • Tokola, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: tt@nn.fi
article id 202, category Research article
Tuula Nuutinen, Antti Kilpeläinen, Hannu Hirvelä, Kari Härkönen, Veli-Pekka Ikonen, Reetta Lempinen, Heli Peltola, Lars Wilhelmsson, Seppo Kellomäki. (2009). Future wood and fibre sources – case North Karelia in eastern Finland. Silva Fennica vol. 43 no. 3 article id 202. https://doi.org/10.14214/sf.202
Keywords: National Forest Inventory; fibre length; wood density; proportion of latewood; forest scenario analysis
Abstract | View details | Full text in PDF | Author Info
Information on the potential wood supply is important for the wood industry. In this study, the future development of growing stock, cutting potential and wood properties corresponding to the regional scenario of North Karelian Forest Programme 2006–2010 was analysed. The simulations were performed by employing the Finnish MELA system together with the sample plot and tree data of the 9th Finnish National Forest Inventory (NFI9) as initial data for the simulations. Disc-based models for basic wood density, proportion of latewood and fibre length of Norway spruce and Scots pine in Sweden were calibrated and integrated into the MELA system. The wood properties at breast height of both harvested and standing trees were analysed in different strata (age, site type and cutting method) during the scenario period of 50 years (2002–2052). The average wood properties within the same strata varied only slightly over time. However, the results for different strata differed considerably. In general, wood density, fibre length and proportion of latewood increased, on average, as a function of tree age and along with a decrease in site fertility (excl. wood density and proportion of latewood in harvested Norway spruce in the first case and fibre length in the latter case for both species). For trees less than 80 years, properties in harvested trees were equal to or slightly greater than those of standing trees. The values for clear-cuttings were greater or equal to those of thinnings (excl. wood density and proportion of latewood in Norway spruce). The study demonstrates the value of model-based analyses utilising NFI tree measurements in regions that are considered to be sources of raw material.
  • Nuutinen, European Forest Institute, Torikatu 34, FI-80100 Joensuu, Finland E-mail: tuula.nuutinen@efi.int (email)
  • Kilpeläinen, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: ak@nn.fi
  • Hirvelä, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: hh@nn.fi
  • Härkönen, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: kh@nn.fi
  • Ikonen, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: vpi@nn.fi
  • Lempinen, Finnish Forest Research Institute, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: rl@nn.fi
  • Peltola, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: hp@nn.fi
  • Wilhelmsson, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: lw@nn.fi
  • Kellomäki, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: sk@nn.fi
article id 201, category Research article
Rui Qi, Véronique Letort, Mengzhen Kang, Paul-Henry Cournède, Philippe de Reffye, Thierry Fourcaud. (2009). Application of the GreenLab model to simulate and optimize wood production and tree stability: a theoretical study. Silva Fennica vol. 43 no. 3 article id 201. https://doi.org/10.14214/sf.201
Keywords: wood quality; optimization; biomechanics; FSPM; Particle Swarm Optimization; source-sink dynamics; biomass allocation
Abstract | View details | Full text in PDF | Author Info
The GreenLab model was used to study the interaction between source-sink dynamics at the whole tree level, wood production and distribution within the stem, and tree mechanical stability through simulation and optimization. In this first promising numerical attempt, two GreenLab parameters were considered in order to maximize wood production: the sink strength for cambial growth and a coefficient that determines the way the biomass assigned to cambial growth is allocated to each metamer, through optimization and simulation respectively. The optimization procedure that has been used is based on a heuristic optimization algorithm called Particle Swarm Optimization (PSO). In the first part of the paper, wood production was maximized without considering the effect of wood distribution on tree mechanical stability. Contrary to common idea that increasing sink strength for cambial growth leads to increasing wood production, an optimal value can be found. The optimization results implied that an optimal source and sink balance should be considered to optimize wood production. In a further step, the mechanical stability of trees submitted to their self weight was taken into account based on simplified mechanical assumptions. Simulation results revealed that the allocation of wood at the stem base strongly influenced its global deformation. Such basic mechanical criterion can be an indicator of wood quality if we consider further the active biomechanical processes involved in tree gravitropic responses, e.g. formation of reaction wood.
  • Qi, Ecole Centrale Paris, Laboratory of Applied Mathematics, Grande Voie des Vignes, 92295 Chatenay-Malabry, France; Institute of Automation, Chinese Academy of Sciences, LIAMA/NLPR, P.O.Box 2728, Beijing, China E-mail: qiruitree@gmail.com (email)
  • Letort, Ecole Centrale Paris, Laboratory of Applied Mathematics, Grande Voie des Vignes, 92295 Chatenay-Malabry, France E-mail: vl@nn.fr
  • Kang, Institute of Automation, Chinese Academy of Sciences, LIAMA/NLPR, P.O.Box 2728, Beijing, China E-mail: mk@nn.cn
  • Cournède, Ecole Centrale Paris, Laboratory of Applied Mathematics, Grande Voie des Vignes, 92295 Chatenay-Malabry, France; INRIA saclay Ile-de-France, EPI Digiplant, Parc Orsay Université, 91893 Orsay cedex, France E-mail: phc@nn.fr
  • Reffye, INRIA saclay Ile-de-France, EPI Digiplant, Parc Orsay Université, 91893 Orsay cedex, France; CIRAD, UMR AMAP, Montpellier, F-34000 France E-mail: pdr@nn.fr
  • Fourcaud, CIRAD, UMR AMAP, Montpellier, F-34000 France E-mail: tf@nn.fr
article id 200, category Research article
Peder Gjerdrum, Mauro Bernabei. (2009). Three-dimensional spiral grain pattern in five large Norway spruce stems. Silva Fennica vol. 43 no. 3 article id 200. https://doi.org/10.14214/sf.200
Keywords: grain angle; 3D-model; tree helix; split disk
Abstract | View details | Full text in PDF | Author Info
There is a great deal of interest involved in investigating and understanding grain angle in trees. The objective of the study presented in this paper has been to identify a joint, three-dimensional model for grain angle in stems of Norway spruce (Picea abies (L.) Karst.) Five large spruce trees were sampled. Transverse disks were extracted at regular intervals along the stem, split through the pith with a blunt knife, and observed for grain angle and cambial age along the north radius, setting pith observation to zero angle. The overall finding confirmed grain pattern congruent to distance from pith along the stem, a pattern that varies from tree to tree. Models expressing distance from the pith in cambial age performed slightly better than models in spatial distance. Grain pattern changed slightly along the stem, and this change was found to be consistent for the five stems: the left-handed grain angle in the juvenile wood was more pronounced upwards in the stem, and the angle changed faster towards right-handed spiralling. The model did not account for grain variation related to knots, undulating pith or other irregularities. Model residuals were normally distributed and seemingly homogeneous for all trees and for all height levels. The model is an extension of existing two-dimensional models for stem cross-cuts. After verification in a wider sample, the results should be applicable in tree and wood modelling.
  • Gjerdrum, Norwegian Forest and Landscape Institute, P.O. Box 115, N-1431 Ås, Norway E-mail: peder.gjerdrum@skogoglandskap.no (email)
  • Bernabei, Norwegian Forest and Landscape Institute, P.O. Box 115, N-1431 Ås, Norway E-mail: mb@nn.no
article id 199, category Research article
R. Edward Thomas. (2009). Modeling the relationships among internal defect features and external Appalachian hardwood log defect indicators. Silva Fennica vol. 43 no. 3 article id 199. https://doi.org/10.14214/sf.199
Keywords: modeling; hardwood; log defects; surface scan
Abstract | View details | Full text in PDF | Author Info
As a hardwood tree grows and develops, surface defects such as branch stubs and wounds are overgrown. Evidence of these defects remain on the log surface for decades and in many instances for the life of the tree. As the tree grows the defect is encapsulated or grown over by new wood. During this process the appearance of the defect in the tree’s bark changes. The defect becomes flatter and its dimension changes. This progressional change in appearance is predictable, permitting the size and location of the internal defect to be reliably estimated. This paper concerns the development and analysis of models for the prediction of internal features. With the advent of surface scanning and external detection systems, the prediction of internal features promises to significantly improve the quality, yield, and value of sawn wood products.
  • Thomas, USDA Forest Service, 241 Mercer Springs Road, Princeton, WV 24740, USA E-mail: edthomas@gmail.com (email)
article id 198, category Research article
Henrik Heräjärvi. (2009). Effect of drying technology on aspen wood properties. Silva Fennica vol. 43 no. 3 article id 198. https://doi.org/10.14214/sf.198
Keywords: stability; heat treatment; modification; press drying; strength
Abstract | View details | Full text in PDF | Author Info
  • Heräjärvi, Metla, Joensuu Research Unit, Box 68, FI-80101 Joensuu, Finland E-mail: henrik.herajarvi@metla.fi (email)
article id 197, category Research article
Veli-Pekka Ikonen, Seppo Kellomäki, Heli Peltola. (2009). Sawn timber properties of Scots pine as affected by initial stand density, thinning and pruning: a simulation based approach. Silva Fennica vol. 43 no. 3 article id 197. https://doi.org/10.14214/sf.197
Keywords: three-dimensional modeling; sawing simulator; knots; Pinus sylvestris L.; shading
Abstract | View details | Full text in PDF | Author Info
The aim of this work was to analyze how different management schedules with varying initial stand density, thinning and artificial pruning of branches affect the quality, quantity and value of sawing yield in Scots pine (Pinus sylvestris L.). For this purpose, an integrated model system was employed and further developed to simulate: i) the three dimensional structure of the crown and stem of an average tree grown in a stand related to the changes in the within-stand light conditions as caused by the stand management, and ii) the sawing of logs into pieces and their quality grading based on the size and number of living and dead knots on the surfaces of sawn pieces. To maximize the quality of sawn timber, relatively dense stand is desired in the early phase of the rotation to reduce, especially in the lower part of stem, the growth of branches, and to increase the rate of dying and pruning-off of branches. In the later phase, a relatively sparse stand is desired to increase the self-pruning of branches and the occlusion of knots. However, in any case, artificial pruning is needed to maximize the knot-free zone of the stem. Also the value optimization of individual sawn pieces affects the quality and value of sawn timber. Because, only average tree was simulated, the differences between scenarios for stem volume were small. In the future, further model development is needed to analyze the development of crown and stem properties of trees with different status in a stand.
  • Ikonen, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: veli-pekka.ikonen@joensuu.fi (email)
  • Kellomäki, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: sk@nn.fi
  • Peltola, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: hp@nn.fi
article id 196, category Research article
Aaron R. Weiskittel, Laura S. Kenefic, Robert S. Seymour, Leah M. Phillips. (2009). Long-term effects of precommercial thinning on the stem dimensions, form and branch characteristics of red spruce and balsam fir crop trees in Maine, USA. Silva Fennica vol. 43 no. 3 article id 196. https://doi.org/10.14214/sf.196
Keywords: Acadian Forest; growth and yield; Abies balsamea; Picea rubens; precommercial thinning
Abstract | View details | Full text in PDF | Author Info
The effects of precommercial thinning (PCT) on stem dimensions, form, volume, and branch attributes of red spruce [Picea rubens Sarg.] and balsam fir [Abies balsamea (L.) Mill.] crop trees were assessed 25 years after treatment in an even-aged northern conifer stand. Treatments were a uniform 2.4 x 2.4-m spacing and a control (no PCT). The PCT treatment significantly increased individual tree diameter at breast height (DBH), height growth, crown ratio, and crown width, while it reduced the tree height to DBH ratio. PCT also significantly increased stem taper and consequently, regional volume equations overpredicted observed stem volume by 2 to 15%, particularly for the spaced trees. PCT also increased the number and maximum size of branches on the lower bole. The sizes of knots on half of the sampled spruce crop trees in the spaced plots precluded them from being used as select structural lumber; there were no other effects on log grade. Our findings indicate that PCT can have a long-term influence on the structural attributes of individual trees, and that improved stem-volume prediction equations are needed in the Acadian region of North America.
  • Weiskittel, School of Forest Resources, University of Maine, Orono, ME 04469, USA E-mail: aaron.weiskittel@umit.maine.edu (email)
  • Kenefic, U.S. Forest Service, Northern Research Station, Bradley, ME 04411, USA E-mail: lsk@nn.us
  • Seymour, School of Forest Resources, University of Maine, Orono, ME 04469, USA E-mail: rss@nn.us
  • Phillips, School of Forest Resources, University of Maine, Orono, ME 04469, USA E-mail: lmp@nn.us
article id 195, category Research article
John R. Moore, Andrew J. Lyon, Gregory J. Searles, Leena E. Vihermaa. (2009). The effects of site and stand factors on the tree and wood quality of Sitka spruce growing in the United Kingdom. Silva Fennica vol. 43 no. 3 article id 195. https://doi.org/10.14214/sf.195
Keywords: wood quality; acoustics; timber properties; wood stiffness
Abstract | View details | Full text in PDF | Author Info
The extent and sources of variation in the wood quality of Sitka spruce (Picea sitchensis (Bong.) Carr.) were quantified using data collected from 64 stands in northern Britain. These stands were selected on the basis of elevation, latitude, longitude, yield class, initial spacing and the presence or absence of thinning. Dynamic modulus of elasticity (MOE) was calculated from measurements of stress wave velocity made on standing trees and qualitative descriptions were made of stem form. Dynamic MOE of individual trees ranged from 3.81 kN/mm2 up to 12.29 kN/mm2, with a mean of 7.71 kN/mm2. Approximately 55 percent of the variation in dynamic MOE was due to differences between individual trees within a site, while 35 percent was due to differences between sites. The remaining 10 percent was due to differences between the measurements made on opposite sides of each tree. Variation in dynamic MOE at the site level was significantly influenced by yield class, elevation as well as by a number of the interactions between these factors and latitude, longitude and initial spacing. A multiple regression model incorporating these variables was able to explain 45 percent of the variation in dynamic MOE. Ramicorn branches were the most commonly recorded defect (37.2% of all live trees), followed by stem scarring and basal sweep (6.9% and 6.3%, respectively). Dynamic MOE was not influenced by stem straightness (p = 0.10) which indicates the utility of stress wave velocity measurements for segregating Sitka spruce stands based on potential grade recovery.
  • Moore, Centre for Timber Engineering, Napier University, 10 Colinton Road, Edinburgh EH10 5DT, United Kingdom E-mail: j.moore@napier.ac.uk (email)
  • Lyon, Centre for Timber Engineering, Napier University, 10 Colinton Road, Edinburgh EH10 5DT, United Kingdom E-mail: ajl@nn.uk
  • Searles, Centre for Timber Engineering, Napier University, 10 Colinton Road, Edinburgh EH10 5DT, United Kingdom E-mail: gjs@nn.uk
  • Vihermaa, Department of Analytical and Environmental Chemistry, University of Glasgow, Glasgow, UK E-mail: lev@nn.uk
article id 194, category Research article
Ane Zubizarreta Gerendiain, Heli Peltola, Pertti Pulkkinen. (2009). Growth and wood property traits in narrow crowned Norway spruce (Picea abies f. pendula) clones grown in southern Finland. Silva Fennica vol. 43 no. 3 article id 194. https://doi.org/10.14214/sf.194
Keywords: fibre length; stem volume; height; diameter; wood density; earlywood; latewood
Abstract | View details | Full text in PDF | Author Info
We investigated the growth, yield, wood density traits and fibre properties in 13 narrow crowned Norway spruce (Picea abies f. pendula) clones grown at a spacing of 2 m x 1.5 m (about 3300 seedlings/ha) in a field trial established in 1988 in southern Finland on a forest soil. For comparison, we used 3 normal crowned Norway spruce (Picea abies (L.) Karst.) genetic entries grown as a mixture in the same trial representing southern Finnish breeding regions. We found that wood density traits and fibre properties showed, on average, lower phenotypic variation than growth and yield traits regardless of crown type. Narrow crowned clones also had, on average, lower stem volume and fibre length, but higher overall wood density. More over, the phenotypic correlations between studied growth and wood properties ranged, on average, from moderate (normal crown) to high (narrow crown). These results were opposite to previous findings for narrow and normal crowned genetic entries grown in narrower spacing (1 m x 1 m) in southern Finland. Thus, this indicates lower plasticity of narrow crowned clones to the increasing growing space compared to normal crowned ones, so, they should be grown at denser spacing in order to fully utilise its space efficiency capacity. However, this field trial was established as a mixture of normal and narrow crown trees, so that 90% of genetic entries were narrow crowned ones, and therefore the crown competition would be much higher for normal crowned trees when the whole trial would consist of that entry alone. In the latter case, we could expect significantly lower productivity of normal crowned genetic entries with this spacing.
  • Zubizarreta Gerendiain, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: ane.zubizarreta@joensuu.fi (email)
  • Peltola, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: hp@nn.fi
  • Pulkkinen, Finnish Forest Research Institute, Haapastensyrjä Breeding Station, FI-12600 Läyliäinen, Finland E-mail: pp@nn.fi
article id 193, category Research article
Jaume Gort, Ane Zubizarreta Gerendiain, Heli Peltola, Pertti Pulkkinen, Johanna Routa, Raimo Jaatinen. (2009). Differences in fibre properties in Scots pine (Pinus sylvestris L.) genetic entries grown at different spacing and sites. Silva Fennica vol. 43 no. 3 article id 193. https://doi.org/10.14214/sf.193
Keywords: fibre length; fibre width; genetic entry; growth; wood density; coarseness; spacing
Abstract | View details | Full text in PDF | Author Info
In forest breeding, stem volume growth and sawn timber quality indicators have been used as the most important selection traits for Scots pine, whereas less attention has been given to characteristics such as fibre properties. In the above context, we investigated the differences in fibre properties (i.e. fibre length, fibre width and coarseness) in 20 year old Scots pine (Pinus sylvestris L.) genetic entries as affected by spacing and site, but also the phenotypic correlations between fibre properties, yield and wood density. The study was based on materials harvested from 10 genetic entries grown in a spacing trial (site 1) in central Finland, with a current stand density of 2000 (spacing 1), 2000–2500 (spacing 2) and 4000 trees/ha (spacing 3). In order to study the effects of site, we harvested additional material (4 of 7 genetic entries same as on site 1) from a trial located in southern Finland with a corresponding stand density of 2000 trees/ha (site 2). On site 1, spacing 1 and 3, all average values for analysed fibre properties were similar. In spacing 2 average values were slightly higher. On site 2, the average values for different fibre properties were similar compared to the corresponding spacing 1 on site 1. Spacing affected (p < 0.05) all average fibre properties on site 1; as did also site, when comparing same genetic entries grown on both sites. Regardless of spacing and site, the phenotypic correlations between average fibre length, fibre width and coarseness showed, on average, moderate to strong correlation (p < 0.05). Fibre width showed, in general, low and positive phenotypic correlation with diameter at breast height, stem volume and wood density on site 1. However, as a whole, the ranking of genetic entries changed depending on the trait and spacing considered. Thus, no overall ranking between genetic entries was possible.
  • Gort, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: jaume.gort@joensuu.fi (email)
  • Zubizarreta Gerendiain, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: azg@nn.fi
  • Peltola, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: hp@nn.fi
  • Pulkkinen, Finnish Forest Research Institute, Haapastensyrjä Breeding Station, Karkkilantie E-mail: pp@nn.fi
  • Routa, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: jr@nn.fi
  • Jaatinen, Finnish Forest Research Institute, Haapastensyrjä Breeding Station, Karkkilantie E-mail: rj@nn.fi
article id 192, category Research article
Heli Peltola, Jaume Gort, Pertti Pulkkinen, Ane Zubizarreta Gerendiain, Jouni Karppinen, Veli-Pekka Ikonen. (2009). Differences in growth and wood density traits in Scots pine (Pinus sylvestris L.) genetic entries grown at different spacing and sites. Silva Fennica vol. 43 no. 3 article id 192. https://doi.org/10.14214/sf.192
Keywords: genetic entry; stem volume; height; diameter; wood property traits; phenotypic correlation
Abstract | View details | Full text in PDF | Author Info
In forest breeding, stem volume has typically taken as the most important selection trait, whereas less attention has been given to wood density traits. In this work, we investigated the effects of spacing and genetic entry on the growth, yield and wood density traits in 20 year old Scots pines (Pinus sylvestris L.) based on 10 genetic entries harvested from a spacing trial (stand density range 2000–4000 trees/ha) in central Finland. In order to study also the site effects, we harvested additional material from a trial located in southern Finland (stand density of 2000 trees/ha). Compared to growth and yield properties, wood density traits showed a lower phenotypic variation. Phenotypic correlations among different traits were negative, and mostly moderate to high, suggesting that selection for one trait would simultaneously affect the others. In addition, moderate to strong phenotypic correlations were found among different wood density traits. Stem volume (V) and breast height diameter (DBH) were the largest in widest spacing, whereas in the densest one tree height (H) and latewood percentage were the highest. Genetic entry affected H and wood density traits regardless of spacing. When comparing two sites (with same stand density), genetic entry affected H, whereas site affected DBH and wood density traits. Ranking between genetic entries changed depending on the trait, spacing or site considered. Therefore, no overall ranking was possible. However, we could identify genetic entries having a high V and a relatively high wood density, showing potential for future forest regeneration material.
  • Peltola, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: heli.peltola@joensuu.fi (email)
  • Gort, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: jg@nn.fi
  • Pulkkinen, Finnish Forest Research Institute, Haapastensyrjä Breeding Station, Karkkilantie 247, FI-12600 Läyliäinen, Finland E-mail: pp@nn.fi
  • Zubizarreta Gerendiain, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: azg@nn.fi
  • Karppinen, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: jk@nn.fi
  • Ikonen, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: vpi@nn.fi
article id 191, category Research article
Katri Luostarinen, Noora Huotari, Eila Tillman-Sutela. (2009). Effect of regeneration method on growth, wood density and fibre properties of downy birch (Betula pubescens Ehrh.). Silva Fennica vol. 43 no. 3 article id 191. https://doi.org/10.14214/sf.191
Keywords: coppice; fibre length; fibre width; regeneration methods; wood properties
Abstract | View details | Full text in PDF | Author Info
Short rotation tree stands, established by coppicing, are nowadays used mainly for energy purposes in Fennoscandia, but their usage for pulp raw material may increase in the future. Downy birch (Betula pubescens), which is commonly used for pulp production in boreal zone, has good sprouting capacity. However, it is not known if the fibre properties of sprout-originated downy birches differ from those of seed-originated ones. Therefore, fibre length and width of sprout- and seed-originated downy birches grown on fertile soil were measured at a stand age of 25 years. Additionally annual ring width, stem height and diameter, and wood density were studied to compare the growth and wood properties of sprout- and seed-originated birches. Annual rings were slightly wider in sprout- than in seed-originated birches, whereas no differences were observed in wood density. Fibres, too, were slightly longer and wider in sprout- than in seed-originated trees. Still these minor differences observed here are hardly significant for the industries using birch wood. Consequently downy birch wood from coppiced stands is well suited for pulp. The advantages of coppice, i.e. rapidity and low costs of establishment, productivity, and the ability of downy birch to grow on untypical forest sites, may even increase the importance of the wood coming from coppiced birch stands in the future.
  • Luostarinen, Faculty of Forest Sciences, University of Joensuu, P.O. Box 111, FI-80101, Joensuu, Finland E-mail: katri.luostarinen@joensuu.fi (email)
  • Huotari, Finnish Forest Research Institute, Muhos Research Unit, Kirkkosaarentie, FI-91500 Muhos, Finland E-mail: nh@nn.fi
  • Tillman-Sutela, Finnish Forest Research Institute, Muhos Research Unit, Kirkkosaarentie, FI-91500 Muhos, Finland E-mail: ets@nn.fi
article id 213, category Research article
Bum-Jin Park, Yuko Tsunetsugu, Tamami Kasetani, Takeshi Morikawa, Takahide Kagawa, Yoshifumi Miyazaki. (2009). Physiological effects of forest recreation in a young conifer forest in Hinokage Town, Japan. Silva Fennica vol. 43 no. 2 article id 213. https://doi.org/10.14214/sf.213
Keywords: blood pressure; heart rate variability; pulse rate; relaxation; Shinrin-yoku; therapeutic effects of forest; well-being
Abstract | View details | Full text in PDF | Author Info
It is widely believed that coming into contact with forest environments is somehow beneficial to human well-being and comfort. In Japan, “Shinrin-yoku” (taking in the atmosphere of a forest) has been proposed to be a relaxation activity associated with forest recreation. The purpose of this study was to examine the physiological effects of forest recreation on the autonomic nervous activity. The subjects were twelve male university students (21.8 ± 0.8 years old). On the first day of the experiment, six subjects were sent to a forest area, and the other six to a city area. On the second day, each subject was sent to the area he did not visit on the first day as a cross check. The subjects walked (15 minutes) around their assigned areas before noon, and sat on chairs viewing (15 minutes) the landscapes of their assigned areas in the afternoon. Heart rate variability (HRV), blood pressure, and pulse rate were measured as physiological indices. Measurements were taken at the place of accommodation in the morning, before and after walking, and before and after viewing at their assigned field areas. Pulse rate, diastolic blood pressure and LF/(LF+HF) (LF – low frequency, HF – high frequency) components of HRV were significantly lower in the forest area than in the city area. HF components of HRV tended to be higher in the forest than in the city. In conclusion, the results of the physiological measurements show that forest recreation enabled effective relaxation in people, both of the mind and body.
  • Park, Chiba University, Center for Environment, Health and Field Sciences, Kashiwa-no-ha 6-2-1, 277-0882 Kashiwa, Chiba, Japan E-mail: bjpark@faculty.chiba-u.jp (email)
  • Tsunetsugu, Chiba University, Center for Environment, Health and Field Sciences, Kashiwa-no-ha 6-2-1, 277-0882 Kashiwa, Chiba, Japan E-mail: yt@nn.jp
  • Kasetani, Chiba University, Center for Environment, Health and Field Sciences, Kashiwa-no-ha 6-2-1, 277-0882 Kashiwa, Chiba, Japan E-mail: tk@nn.jp
  • Morikawa, Chiba University, Center for Environment, Health and Field Sciences, Kashiwa-no-ha 6-2-1, 277-0882 Kashiwa, Chiba, Japan E-mail: tm@nn.jp
  • Kagawa, Chiba University, Center for Environment, Health and Field Sciences, Kashiwa-no-ha 6-2-1, 277-0882 Kashiwa, Chiba, Japan E-mail: tk@nn.jp
  • Miyazaki, Chiba University, Center for Environment, Health and Field Sciences, Kashiwa-no-ha 6-2-1, 277-0882 Kashiwa, Chiba, Japan E-mail: ym@nn.jp
article id 212, category Research article
Sandhya Samarasinghe. (2009). Exploration of fracture dynamics properties and predicting fracture toughness of individual wood beams using neural networks. Silva Fennica vol. 43 no. 2 article id 212. https://doi.org/10.14214/sf.212
Keywords: Pinus radiata; wood properties; cracks; initiation; New Zealand; peak stress; speed; video imaging
Abstract | View details | Full text in PDF | Author Info
In this study, the time to crack initiation (Tinit), duration of crack propagation (Tfrac), crack initiation stress, peak stress as well as crack speed and fracture toughness were investigated for three Rates of Loading (ROL) and four sizes of notched wood beams using high-speed video imaging and neural networks. Tinit was consistent for all volumes and the average Tinit was nonlinearly related to volume and ROL. For the smallest ROL, there was a distinct volume effect on Tinit and the effect was negligble at the largest ROL. However, the stress at crack initiation was not consistent. Contrasting these, Tfrac for all volumes appeared to be highly variable but the peak stress carried prior to catastrophic failure was consistent. The crack propagation was a wave phenomenon with positive and negative (crack closure) speeds that varied with the ROL. As accurate estimation of crack initiation load (or stress) and its relationship to peak load (or stress) is important for determining fracture toughness, Artificial Neural Networks (ANN) models were developed for predicting them from volume, Young’s modulus, face and grain angles, density, moisture content and ROL. Models for crack initiation load and peak load showed much higher predictive power than those for the stresses with correlation coefficients of 0.85 and 0.97, respectively, between the actual and predicted loads. Neural networks were also developed for predicting fracture toughness of individual wood specimens and the best model produced a statistically significant correlation of 0.813 between the predicted and actual fracture toughness on a validation dataset. The inputs captured 62% of variability of fracture toughness. Volume and Young’s modulus were the top two contributing variables with others providing lesser contributions.
  • Samarasinghe, Centre for Advanced Computational Solutions (C-fACS), Lincoln University, Canterbury, New Zealand E-mail: sandhya.samarasinghe@lincoln.ac.nz (email)
article id 211, category Research article
Timo Pukkala. (2009). Population-based methods in the optimization of stand management. Silva Fennica vol. 43 no. 2 article id 211. https://doi.org/10.14214/sf.211
Keywords: Particle Swarm Optimization; differential evolution; evolution strategy; Nelder-Mead method; amoeba search; polytope search; evolutionary computing
Abstract | View details | Full text in PDF | Author Info
In Finland, the growth and yield models for tree stands are simulation programs that consist of several sub-models. These models are often non-smooth and non-differentiable. Direct search methods such as the Hooke-Jeeves algorithm (HJ) are suitable tools for optimizing stand management with this kind of complicated models. This study tested a new class of direct search methods, namely population-based methods, in the optimization of stand management. The tested methods were differential evolution, particle swarm optimization, evolution strategy, and the Nelder-Mead method. All these methods operate with a population of solution vectors, which are recombined and mutated to obtain new candidate solutions. The management schedule of 719 stands was optimized with all population-based methods and with the HJ method. The population-based methods were competitive with the HJ method, producing 0.57% to 1.74% higher mean objective function values than HJ. On the average, differential evolution was the best method, followed by particle swarm optimization, evolution strategy, and Nelder-Mead method. However, differences between the methods were small, and each method was the best in several stands. HJ was alone the best method in 7% of stands, and a population based method in 3% (Nelder-Mead) to 29% (differential evolution) of stands. All five methods found the same solution in 18% of stands.
  • Pukkala, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: timo.pukkala@joensuu.fi (email)
article id 210, category Research article
Juha Heiskanen, Markku Lahti, Jaana Luoranen, Risto Rikala. (2009). Nutrient loading has a transitory effect on the nitrogen status and growth of outplanted Norway spruce seedlings. Silva Fennica vol. 43 no. 2 article id 210. https://doi.org/10.14214/sf.210
Keywords: nursery fertilization; plantation performance; soil fertility
Abstract | View details | Full text in PDF | Author Info
In recent years increased fertilization provided to tree seedlings in the nursery in the previous autumn has been introduced in order to promote good outplanting performance. In this paper this nutrient loading has been studied in order to determine how the increased seedling nutrient status with unaffected seedling size affects both the growth and the nutrient concentration, content and uptake of two-year-old Norway spruce container seedlings (Picea abies (L.) Karsten) after outplanting. Seedling development was monitored for three years at two contrasting soil fertility levels on a sandy test field in two planting years and on one natural forest outplanting site in central Finland. Nutrient loading was shown to increase shoot and root growth in a poor fertility soil during the first growing season after planting, while, after the first growing season, nutrient loading was not found to affect seedling performance. However, although nutrient loading cannot compensate for the availability of nutrients to the seedlings from the soil, it may provide an additional input for fast plantation establishment on poorer sites during the first crucial growing season after outplanting.
  • Heiskanen, The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland E-mail: juha.heiskanen@metla.fi (email)
  • Lahti, The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland E-mail: ml@nn.fi
  • Luoranen, The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland E-mail: jl@nn.fi
  • Rikala, The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland E-mail: rr@nn.fi
article id 209, category Research article
Jaana Luoranen, Kyösti Konttinen, Risto Rikala. (2009). Frost hardening and risk of a second flush in Norway spruce seedlings after an early-season short-day treatment. Silva Fennica vol. 43 no. 2 article id 209. https://doi.org/10.14214/sf.209
Keywords: frost hardiness; Picea abies; bud formation; container seedlings; height growth cessation; second flush
Abstract | View details | Full text in PDF | Author Info
There have been years in Finland when container seedlings of Norway spruce (Picea abies (L.) Karsten) planted in the summer have been damaged by early-autumn frosts. For August and September plantings, the seedlings can be hardened by means of short-day (SD) treatment, but little information is available about its usability for earlier plantings. We studied the effects of early-season SD treatment on the frost hardiness and risk of a second flush of Norway spruce seedlings. In three successive years, second-year seedlings were grown in a greenhouse or outdoors in the spring and early summer and then subjected to two or three-week SD treatment beginning on the second, third, or fourth week of June. We monitored the height growth cessation, bud formation, and frost hardiness of the seedlings in the nursery. All SD treatments made the height growth cease, but the risk of a second flush increased if the temperature sum was less than 300 d.d. before the beginning of the SD treatment or more than 450 d.d. between the end of the treatment and mid-August. Clearly, then, SD treatment reduced the risk of a second flush in seedlings that had been grown in a greenhouse in the spring. Early-season SD treatment increased the frost hardiness of both needles and stems for late July to early September in comparison with untreated seedlings. Later in the autumn, however, the differences disappeared. Before recommending the use of early-season SD-treated seedlings for summer planting, the method has to be tested in practical field conditions.
  • Luoranen, The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland E-mail: jaana.luoranen@metla.fi (email)
  • Konttinen, The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland E-mail: kk@nn.fi
  • Rikala, The Finnish Forest Research Institute, Suonenjoki Research Unit, FI-77600 Suonenjoki, Finland E-mail: rr@nn.fi
article id 208, category Research article
Mikko Räisänen, Tapani Repo, Tarja Lehto. (2009). Cold acclimation of Norway spruce roots and shoots after boron fertilization. Silva Fennica vol. 43 no. 2 article id 208. https://doi.org/10.14214/sf.208
Keywords: boron deficiency; dieback; freezing tolerance; mineral nutrition
Abstract | View details | Full text in PDF | Author Info
Boron deficiency, manifested as shoot dieback, is a problem in conifer stands growing on soils with high nitrogen availability in Fennoscandia. Earlier observations on Norway spruce (Picea abies L. Karst.) suggest that freezing tolerance is decreased by boron deficiency. Here, the effect of boron fertilization on cold acclimation of Norway spruce was studied in a young stand with initially low boron status two years after fertilization. Buds, stems, needles and roots were collected at five sampling times during cold acclimation and subsequently exposed to series of freezing temperatures. Lethal temperatures of organs were assessed by electrolyte leakage method (EL) and visual scoring of damage (VS). Freezing tolerance of buds was measured also by differential thermal analysis (DTA). The mean boron (B) concentration in needles was 4 mg kg–1 in unfertilized and 21 mg kg–1 in B-fertilized trees while critical level of B deficiency is considered to be 5 mg kg–1. The risk for increased freezing injuries in the low-B trees was not evident since all trees achieved cold hardiness that would be sufficient in central Finland. At two sampling times out of five, shoots or stem of B-fertilized trees were slightly more freezing tolerant than non-fertilized trees. However, the present study does not give strong evidence for the hypothesis that decreased freezing tolerance in B deficiency would be a triggering factor for leader dieback in Norway spruce at the B levels studied.
  • Räisänen, University of Joensuu, Faculty of Forest Sciences, P.O.Box 111, FI-80101 Joensuu, Finland; (present address), FAForest, FI-83480 Ahonkylä, Finland E-mail: mr@nn.fi (email)
  • Repo, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: tr@nn.fi
  • Lehto, University of Joensuu, Faculty of Forest Sciences, P.O.Box 111, FI-80101 Joensuu, Finland E-mail: tarja.lehto@joensuu.fi
article id 207, category Research article
Mats T. Olsson, Maria Erlandsson, Lars Lundin, Torbjörn Nilsson, Åke Nilsson, Johan Stendahl. (2009). Organic carbon stocks in Swedish Podzol soils in relation to soil hydrology and other site characteristics. Silva Fennica vol. 43 no. 2 article id 207. https://doi.org/10.14214/sf.207
Keywords: temperature sum; soil organic carbon; forest land; soil hydrological class; N deposition; latitude; site capacity; mean annual precipitation; altitude
Abstract | View details | Full text in PDF | Author Info
Site characteristics influence soil organic carbon (SOC) stocks. In Podzols under Swedish forest land, SOC stocks were related to latitude, altitude, soil hydrological class categorized by mean groundwater level, mean annual precipitation, temperature sum during the growing season, total annual nitrogen (N) deposition and site capacity. SOC stocks were determined for the O-horizon and for total soil (O-horizon + mineral soil to a depth of 50 cm). Data from the Swedish National Forest Soil Inventory 1993–2001 were used (1477 field plots). The O-horizon was sampled with a core sampler and carbon (C) stocks were determined. For the mineral soil layers the SOC stock was calculated based on the SOC concentrations, bulk density and content of rock fragments. The results showed that the overall mean SOC stock was 2.8 and 8.2 kg C m–2 for O-horizon and total soil, respectively. Soil hydrological class strongly affected SOC stocks, which increased from on average 6.7 kg C m–2 at dry sites to 9.7 kg C m–2 at slightly moist sites. Corresponding values for the O-horizon were 2.0 to 4.4 kg C m–2. The correlation coefficients for the linear relationship between SOC stock and site characteristics were highest for N deposition, which explained up to 25% of variation, and latitude, which explained up to 20% of variation. Altitude had the lowest degree of explanation.
  • Olsson, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden E-mail: mto@nn.se
  • Erlandsson, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden E-mail: me@nn.se
  • Lundin, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden E-mail: ll@nn.se
  • Nilsson, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden E-mail: torbjorn.nilsson@mark.slu.se (email)
  • Nilsson, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden E-mail: an@nn.se
  • Stendahl, Swedish University of Agricultural Sciences, Dept of Soil and Environment, Uppsala, Sweden E-mail: js@nn.se
article id 206, category Research article
Qiaoying Zhang, Yunchun Zhang, Shaolin Peng, Eshetu Yirdaw, Ning Wu. (2009). Spatial structure of alpine trees in Mountain Baima Xueshan on the southeast Tibetan Plateau. Silva Fennica vol. 43 no. 2 article id 206. https://doi.org/10.14214/sf.206
Keywords: treeline; timberline; Abies forestii var. georgei (Orr) Farjon; Juniperus saltuaria Rehder & E.H. Wilson; spatial point pattern
Abstract | View details | Full text in PDF | Author Info
  • Zhang, State Key Laboratory of Biocontrol, Sun Yat-Sen University, Guangzhou 510275, Guangdong Province, China; Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, Sichuan Province, China E-mail: lsspsl@mail.sysu.edu.cn (email)
  • Zhang, Shandong Institute of Light Industry, Jinan 250353, Shandong Province, China E-mail: yz@nn.cn
  • Peng, State Key Laboratory of Biocontrol, Sun Yat-Sen University, Guangzhou 510275, Guangdong Province, China E-mail: sp@nn.cn
  • Yirdaw, Viikki Tropical Resources Institute (VITRI), P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ey@nn.fi
  • Wu, Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, Sichuan Province, China E-mail: nw@nn.cn
article id 205, category Research article
Pekka Vakkari, Mari Rusanen, Katri Kärkkäinen. (2009). High genetic differentiation in marginal populations of European white elm (Ulmus laevis). Silva Fennica vol. 43 no. 2 article id 205. https://doi.org/10.14214/sf.205
Keywords: marginal populations; genetic differentiation; genetic conservation; Ulmus
Abstract | View details | Full text in PDF | Author Info
Studies on the amount of genetic variation in marginal populations and differentiation between them are essential for assessment of best gene conservation strategies and sampling schemes. Thirteen marginal populations of Ulmus laevis in southern Finland and one in Estonia were investigated for genetic variation in 20 allozyme loci. Population differentiation among Finnish stands was high, Fst = 0.290, and mean genetic diversity low, He = 0.088. The differentiation follows the isolation-by-distance structure within the core of the distribution area (lake Vanajavesi). Fairly high frequency of recurrent genotypes was observed, but this did not have an influence on the genetic parameters. The observed genetic structure is consistent with the central-marginal hypothesis. In the light of the results, the Finnish gene conservation strategy for U. laevis seems to be on a sound basis.
  • Vakkari, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: pekka.vakkari@metla.fi (email)
  • Rusanen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: mr@nn.fi
  • Kärkkäinen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: kk@nn.fi
article id 464, category Research article
Christian Kanzian, Franz Holzleitner, Karl Stampfer, Sarah Ashton. (2009). Regional energy wood logistics – optimizing local fuel supply. Silva Fennica vol. 43 no. 1 article id 464. https://doi.org/10.14214/sf.464
Keywords: logistics; energy wood; transport optimization; GIS
Abstract | View details | Full text in PDF | Author Info
The promotion of electric energy production from solid biomass by the Austrian government has lead to a boom in the construction of new combined heat and power plants. The current total demand for wood chips in the research area for energy purposes is 70 400 m3 of loose volume chips per year. The expected increase in demand due to these new plants is more than 4 times greater than current demand: up to 302 700 m3 of loose volume per year. Even if the energy wood feedstock potential is satisfactory, the design of the supply chain is still unresolved. The aim of this study is to give decision-makers a base for further development. To accomplish this, we designed and tested four different supply scenarios: one for 9 plants and one for 16 plants. The scenarios were developed using a combination of geographic information systems (GIS) and linear programming methods. The results indicate that direct transport of solid fuel wood as round wood and chipping at the plant is the cheapest supply system with a resulting cost of 5.6–6.6 EUR/m3 loose. Using harvesting residues can only be recommended for large plants because of poor fuel quality. In this case, residues would be chipped at or near the landing, piled and transported via self-loading trucks at a cost between 8.4 and 9.1 EUR/m3 loose. In order to meet increasing demand and to ensure a continuous supply, especially during the winter and spring seasons it is necessary to optimize the supply chain by including storage terminals. However, using terminals and increased demand both lead to higher logistical costs. For example, if the total volume is handled via terminals, the average supply costs including storage will increase by 26%. Higher demand increases the costs by 24%.
  • Kanzian, University of Applied Life Sciences Vienna, Institute of Forest Engineering, Peter Jordan Strasse 82, A-1190 Vienna, Austria E-mail: christian.kanzian@boku.ac.at (email)
  • Holzleitner, University of Applied Life Sciences Vienna, Institute of Forest Engineering, Peter Jordan Strasse 82, A-1190 Vienna, Austria E-mail: fh@nn.at
  • Stampfer, University of Applied Life Sciences Vienna, Institute of Forest Engineering, Peter Jordan Strasse 82, A-1190 Vienna, Austria E-mail: ks@nn.at
  • Ashton, Southern Regional Extension Forestry, Forestry Bldg. 4-420, University of Georgia, Athens, GA 30602, USA E-mail: sa@nn.us
article id 222, category Research article
Yi Pan, Yaoqi Zhang, Indrajit Majumdar. (2009). Population, economic welfare and holding size distribution of private forestland in Alabama, USA. Silva Fennica vol. 43 no. 1 article id 222. https://doi.org/10.14214/sf.222
Keywords: consolidation; private forestland owners; parcellation; land tenure
Abstract | View details | Full text in PDF | Author Info
This paper analyzes structural differences in holding size distribution of private forestland at the county level in Alabama, USA. The following categories are used to describe the timberland holding distribution: mean holding size, Gini coefficient of timberland holdings, and share of county timberland in holding sizes of fewer than 80 hectares and more than 800 hectares. Regressions are conducted to explore the socio-economic correlates of forestland holding size distribution. Results indicate that population density, age structure, urban-rural population structure, income structure and land tenure type are strongly associated with the forest holding size distribution.
  • Pan, Auburn University, School of Forestry and Wildlife Sciences, Auburn, AL 36849, USA E-mail: yp@nn.us
  • Zhang, Auburn University, School of Forestry and Wildlife Sciences, Auburn, AL 36849, USA E-mail: yaoqi.zhang@auburn.edu (email)
  • Majumdar, Auburn University, School of Forestry and Wildlife Sciences, Auburn, AL 36849, USA E-mail: im@nn.us
article id 221, category Research article
Glen Murphy, Rod Brownlie, Mark Kimberley, Peter Beets. (2009). Impacts of forest harvesting related soil disturbance on end-of-rotation wood quality and quantity in a New Zealand radiata pine forest. Silva Fennica vol. 43 no. 1 article id 221. https://doi.org/10.14214/sf.221
Keywords: Pinus radiata; harvesting; compaction; wood density; tree growth; stiffness; litter removal; nitrogen deficiency
Abstract | View details | Full text in PDF | Author Info
The long-term effect of soil disturbance (litter removal, topsoil removal and compaction) from forest harvesting on wood quality and quantity of second-rotation Pinus radiata growing on a clay loam soil, was assessed at the end the rotation, 26 years after planting. Relative to Control plots, average tree and stand total volume at rotation end was not significantly affected by litter removal and nil or light compaction, but was significantly reduced by 28% by litter and topsoil removal and moderate subsoil compaction, and further reduced by 38% by heavy compaction. Wood density at breast height in the inner rings of trees in the most disturbed treatments was elevated by up to 30 kg m–3. This occurred because these treatments were more N deficient as reflected by foliar N levels during the first 11 years of growth relative to the Control. However, no treatment differences in wood density were evident in outer rings, and by rotation age overall mean density did not differ significantly between treatments. Neither acoustic velocity of standing trees, nor acoustic velocity of logs, was significantly affected by soil disturbance, indicating that stiffness of lumber cut from trees in the trial was likely to be similar for all treatments. Economic impacts of soil disturbance and compaction on this soil type will therefore result largely from the considerable negative impacts on final tree size, with little or no compensation from improved wood properties.
  • Murphy, Forest Engineering, Resources and Management Department, Oregon State University, Corvallis, Oregon, USA E-mail: glen.murphy@oregonstate.edu (email)
  • Brownlie, Scion Research, Rotorua, New Zealand E-mail: rb@nn.nz
  • Kimberley, Scion Research, Rotorua, New Zealand E-mail: mk@nn.nz
  • Beets, Scion Research, Rotorua, New Zealand E-mail: pb@nn.nz
article id 220, category Research article
Jani Heikkilä, Matti Sirén, Anssi Ahtikoski, Jari Hynynen, Tiina Sauvula, Mika Lehtonen. (2009). Energy wood thinning as a part of stand management of Scots pine and Norway spruce. Silva Fennica vol. 43 no. 1 article id 220. https://doi.org/10.14214/sf.220
Keywords: energy wood thinning; stand management; MOTTI simulator
Abstract | View details | Full text in PDF | Author Info
The effects of combined production of industrial and energy wood on yield and harvesting incomes, as well as the feasibility of energy wood procurement, were studied. Data for 22 Scots pine (Pinus sylvestris L.) and 21 Norway spruce (Picea abies (L.) Karst.) juvenile stands in Central and Southern Finland were used to compare six combined production regimes to conventional industrial wood production. The study was based on simulations made by the MOTTI stand simulator, which produces growth predictions for alternative management regimes under various site and climatic conditions. The combined production regimes included precommercial thinning at 4–8 m dominant height to a density of 3000–4000 stems ha–1 and energy wood harvesting at 8, 10 or 12 m dominant height. Combined production did not decrease the total yield of industrial wood during the rotation period. Differences in the mean annual increment (MAI) were small, and the rotation periods varied only slightly between the alternatives. Combined production regime can be feasible for a forest owner if the price of energy wood is 3–5 EUR m–3 in pine stands, and 8–9 EUR m–3 in spruce stands. Energy wood procurement was not economically viable at the current energy price (12 EUR MWh–1) without state subsidies. Without subsidies a 15 EUR MWh–1 energy price would be needed. Our results imply that the combined production of industrial and energy wood could be a feasible stand management alternative.
  • Heikkilä, L&T Biowatti Oy, P.O. Box 738, FI-60101 Seinäjoki, Finland E-mail: jani.heikkila@biowatti.fi (email)
  • Sirén, Finnish Forest Research Institute, Vantaa Research Unit, P.O.Box 18, FI-01301 Vantaa, Finland E-mail: ms@nn.fi
  • Ahtikoski, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O.Box 16, FI-96301 Rovaniemi, Finland E-mail: aa@nn.fi
  • Hynynen, Finnish Forest Research Institute, Vantaa Research Unit, P.O.Box 18, FI-01301 Vantaa, Finland E-mail: jh@nn.fi
  • Sauvula, Seinäjoki University of Applied Sciences, School of Agriculture and Forestry, Tuomarniementie 55, FI-63700 Ähtäri, Finland E-mail: ts@nn.fi
  • Lehtonen, Finnish Forest Research Institute, Vantaa Research Unit, P.O.Box 18, FI-01301 Vantaa, Finland E-mail: ml@nn.fi
article id 219, category Research article
Arto Haara, Pekka Leskinen. (2009). The assessment of the uncertainty of updated stand-level inventory data. Silva Fennica vol. 43 no. 1 article id 219. https://doi.org/10.14214/sf.219
Keywords: uncertainty; measurement error; simulation; non-parametric methods; observed error; stand-level inventory
Abstract | View details | Full text in PDF | Author Info
Predictions of growth and yield are essential in forest management planning. Growth predictions are usually obtained by applying complex simulation systems, whose accuracy is difficult to assess. Moreover, the computerised updating of old inventory data is increasing in the management of forest planning systems. A common characteristic of prediction models is that the uncertainties involved are usually not considered in the decision-making process. In this paper, two methods for assessing the uncertainty of updated forest inventory data were studied. The considered methods were (i) the models of observed errors and (ii) the k-nearest neighbour method. The derived assessments of uncertainty were compared with the empirical estimates of uncertainty. The practical utilisation of both methods was considered as well. The uncertainty assessments of updated stand-level inventory data using both methods were found to be feasible. The main advantages of the two studied methods include that bias as well as accuracy can be assessed.
  • Haara, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: arto.haara@joensuu.fi (email)
  • Leskinen, Finnish Environment Institute, Research Programme for Production and Consumption, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: pl@nn.fi
article id 218, category Research article
Md. Nurul Islam, Mikko Kurttila, Lauri Mehtätalo, Arto Haara. (2009). Analyzing the effects of inventory errors on holding-level forest plans: the case of measurement error in the basal area of the dominated tree species. Silva Fennica vol. 43 no. 1 article id 218. https://doi.org/10.14214/sf.218
Keywords: inoptimality loss; dominated tree species; erroneous inventory data; forest plan
Abstract | View details | Full text in PDF | Author Info
Accurate inventory data are required for ensuring optimal net return on investment from the forest. Erroneous data can lead to the formulation of a non-optimal plan that can cause inoptimality losses. Little is known of the effect of using erroneous stand inventory data in preparing holding-level forest plans. This study reports on an approach for analyzing such inoptimality losses. Furthermore, inoptimality losses caused by measurement errors in the basal area of the dominated tree species were investigated in a case study. Based on the inventory data including routine measurements by 67 measurers, four measurer groups were created with different measurement error profiles for the basal area of the dominated tree species. This was followed by measurement error simulations for each group and by adding these to the accurate control inventory data to create erroneous data of different error profiles. Three different forest plans were then constructed by using erroneous data of each group. The plans were then analyzed and compared with plans based on correct data. The effect of measurement errors on the net present value from the whole planning period, and on the amount of remaining growing stock at the end of planning period, were analyzed and utilized in calculating the inoptimality losses. It was concluded that even errors involving dominated tree species can cause significant changes in the holding-level forest plans.
  • Islam, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: nurul.islam@joensuu.fi (email)
  • Kurttila, Finnish Forest Research Institute, Joensuu Research Unit, FI-80101 Joensuu, Finland E-mail: mk@nn.fi
  • Mehtätalo, University of Helsinki, Dept. of Forest Resource Management, FI-00014 University of Helsinki, Finland E-mail: lm@nn.fi
  • Haara, University of Joensuu, Faculty of Forest Sciences, FI-80101 Joensuu, Finland E-mail: ah@nn.fi
article id 217, category Research article
Ville Kankaanhuhta, Timo Saksa, Heikki Smolander. (2009). Variation in the results of Norway spruce planting and Scots pine direct seeding in privately-owned forests in southern Finland. Silva Fennica vol. 43 no. 1 article id 217. https://doi.org/10.14214/sf.217
Keywords: Pinus sylvestris; Picea abies; quality control; forest regeneration survey; forest regeneration inventory; quality management
Abstract | View details | Full text in PDF | Author Info
This study describes the variation in the planting results for 3-year-old Norway spruce (Picea abies (L.) Karst.) and 4-year-old Scots pine (Pinus sylvestris L.) using direct seeding in privately-owned regeneration areas in southern Finland. The study material consists of operative forest regeneration quality management inventory areas from the years 2000–2006. The effect of both the regional and the administrative levels as well as ecological factors was modelled on the basis of the hierarchy structure forestry centre, Forest Owners’ Association (= FOA), forestry professional, regeneration area and sample plot. The major part of the variation occurred at the sample plot and regeneration area level. Particular attention was paid to observation of the clustered spatial distribution of Scots pine seedlings. The FOA and forestry professional levels explained 5% of the variation in Norway spruce planting and 11% of the variation in Scots pine direct seeding. Applied forest regeneration operations, site and soil characteristics were included in the fixed effects. In the planting of Norway spruce the most important factor explaining the regeneration result was soil preparation. Mounding produced better results than patching and disc trenching. The site and soil characteristics were other important factors in the operations. The selection of direct seeding of Scots pine on too fertile, fine textured or moist sites yielded poor results.
  • Kankaanhuhta, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: ville.kankaanhuhta@metla.fi (email)
  • Saksa, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: ts@nn.fi
  • Smolander, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: hs@nn.fi
article id 216, category Research article
Juha Siitonen, Jenni Hottola, Auli Immonen. (2009). Differences in stand characteristics between brook-side key habitats and managed forests in southern Finland. Silva Fennica vol. 43 no. 1 article id 216. https://doi.org/10.14214/sf.216
Keywords: dead wood; woodland key habitats; stand structure; CWD; WKH; Forest Act
Abstract | View details | Full text in PDF | Author Info
Preservation of small habitat patches termed as “woodland key habitats” or “especially important habitats” in the Finnish Forest Act has become an integral part of biodiversity-oriented forest management. Forest Act habitats belong to particular habitat types defined in the act, and they are supposed to have natural-like stand characteristics. However, very little is known about the actual stand structure in the designated habitats. Our aim was to compare stand characteristics between brook-side key habitats and comparable managed forests as controls. Seven study areas were selected from four regions across southern Finland. Within each study area ten key habitats and ten controls (140 stands) were randomly selected. Living and dead trees and cut stumps were measured in each stand within a 0.2 ha plot. The average degree of previous cutting was significantly lower whereas the volume of dead wood, volume of deciduous trees, and stand diversity were each significantly higher in key habitats than controls. The average volume of dead wood was 11.7 m3 ha–1 in key habitats and 6.5 m3 ha–1 in controls. However, there was considerable variation among individual stands, and a large part of key habitats could not be distinguished from randomly selected control stands with respect to stand characteristics. The preservation of natural brook channels with their immediate surroundings is undoubtedly important for maintaining aquatic and semiaquatic biodiversity. Nevertheless, when complementing the forest conservation network in the future, main emphasis in selecting potentially valuable stands should be placed on important structural features such as dead wood and old trees.
  • Siitonen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: juha.siitonen@metla.fi (email)
  • Hottola, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: jh@nn.fi
  • Immonen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: ai@nn.fi
article id 215, category Research article
Antti J. Lukkarinen, Seppo Ruotsalainen, Teijo Nikkanen, Heli Peltola. (2009). The growth rhythm and height growth of seedlings of Siberian (Larix sibirica Ledeb.) and Dahurian (Larix gmelinii Rupr.) larch provenances in greenhouse conditions. Silva Fennica vol. 43 no. 1 article id 215. https://doi.org/10.14214/sf.215
Keywords: height growth; larch; Larix; provenance; growth rhythm
Abstract | View details | Full text in PDF | Author Info
The aim of this study was to determine whether the growth rhythm and height growth differ between various larch provenances grown in greenhouse conditions. We also investigated whether the geographic and climatic conditions at the origin of the provenance could explain the possible differences between the provenances. The study material consisted of 16 Russian Siberian (Larix sibirica Ledeb.) and Dahurian larch (Larix gmelinii Rupr.) provenances and four seed sources from Finland as comparison lots. The growth rhythm differences were clearest between the southern and northern provenances; the southern provenances grew for a longer period and the proportion of late summer height growth was larger. Autumn colouration also developed later in these provenances. In the Russian larch material the provenances with a longer growing period had greater height growth. In the whole material the relationship was not so linear due to the deviating behaviour of the comparison material. Several of the Russian larch provenances were taller than the Finnish comparison seed lots, although the difference was not statistically significant. However, the growth of these provenances must be followed for a much longer period in field trials before any conclusions can be drawn about their usability in practical forestry.
  • Lukkarinen, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: antti.lukkarinen@pp.inet.fi (email)
  • Ruotsalainen, Finnish Forest Research Institute, Punkaharju Research Unit, Finlandiantie 18, FI-58450 Punkaharju, Finland E-mail: sr@nn.fi
  • Nikkanen, Finnish Forest Research Institute, Punkaharju Research Unit, Finlandiantie 18, FI-58450 Punkaharju, Finland E-mail: tn@nn.fi
  • Peltola, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: hp@nn.fi
article id 214, category Research article
Michelle de Chantal, Hannu Rita, Urban Bergsten, Mikaell Ottosson Löfvenius, Harald Grip. (2009). Frost heaving of Picea abies seedlings as influenced by soil preparation, planting technique, and location along gap-shelterwood gradients. Silva Fennica vol. 43 no. 1 article id 214. https://doi.org/10.14214/sf.214
Keywords: deep planting; normal planting; mobile container; fixed container; HuMinMix; soil horizon
Abstract | View details | Full text in PDF | Author Info
The effects of soil preparation, planting technique and location along gap-shelterwood gradients (position and orientation) on frost heaving damage to seedlings were studied in Vindeln Experimental Forests, northern Sweden. The forest was harvested in a grid pattern in winter 2004–2005, forming gaps and shelterwood areas of 30 x 40 m each. Gap-shelterwood gradients were delimited in four orientations and subdivided into five positions: 7 m and 15 m into the gap and shelterwood, and at the gap edge. At each position, three replicates of three soil preparations were made: exposed E and B horizons and HuMinMix (milled vegetation and humus layers mixed with surface mineral soil). In early October 2005, one-year-old containerized Picea abies (L.) Karst. seedlings were planted using four techniques: normal and deep planting, and mobile and fixed experimental containers. After one winter, frost heaving damage was highest for seedlings on B horizon combined with the mobile container (51 ± 6%) and normal planting (43±6%). Normal- or deep-planted seedlings in HuMinMix had the least damage (5–6.6 ± 2.5%). Compared to normal planting, deep planting reduced frost heaving damage only on B horizon. When considering the orientation, seedlings in the experimental containers had more or similar frost heaving damage than normal- or deep-planted seedlings. Along the eastern gradient, seedlings incurred more frost heaving damage in the center of the gap than under the canopy.
  • Chantal, University of Helsinki, Dept of Forest Ecology & Dept of Forest Resource Management, Helsinki, Finland E-mail: michelle.dechantal@helsinki.fi (email)
  • Rita, University of Helsinki, Dept of Forest Ecology & Dept of Forest Resource Management, Helsinki, Finland E-mail: hr@nn.fi
  • Bergsten, Swedish University of Agricultural Sciences, Dept of Forest Ecology and Management, Umeå, Sweden E-mail: ub@nn.se
  • Löfvenius, Swedish University of Agricultural Sciences, Dept of Forest Ecology and Management, Umeå, Sweden E-mail: mol@nn.se
  • Grip, Swedish University of Agricultural Sciences, Dept of Forest Ecology and Management, Umeå, Sweden E-mail: hg@nn.se
article id 467, category Research article
Annika Kangas, Ruut Haapakoski, Liisa Tyrväinen. (2008). Integrating place-specific social values into forest planning – Case of UPM-Kymmene forests in Hyrynsalmi, Finland. Silva Fennica vol. 42 no. 5 article id 467. https://doi.org/10.14214/sf.467
Keywords: collaborative planning; experienced values of forests; social value mapping; survey
Abstract | View details | Full text in PDF | Author Info
In participatory forest management planning, the perceived values of local inhabitants concerning the area under planning are collected. The results may, however, depend on the methods used. In this study, values of residents of Hyrynsalmi municipality concerning the nearby forests owned by UPM-Kymmene Ltd. were evaluated with a questionnaire consisting of general value questions and mapping of social values of forests. The data was collected from a public meeting and from a mailed survey from randomly sampled people and from members of municipal council. The aims of the study were to 1) test the social value mapping method in commercial forests in a rural-urban interface as well as to examine the benefits and drawbacks 2) in place-specific and non-specific data collection, and 3) in different data collection methods, from the viewpoint of forest management planning. We noted that while all respondents can claim to represent local values, different data collection methods produced statistically significantly different local values. This needs to be accounted for when planning a participatory process. In operational forest planning, place-specific information is more useful than questions concerning the general values, while the latter may help in defining forest policy goals. The social values mapping method is also relatively easy for the participants. However, in the studied case about one fifth of the area was delineated by the participants per each positive value. The answers were quite scattered, suggesting that most of the area had some social values for local people. This indicates that utilising a social values mapping method in planning needs further development in rural areas, where distinctive patches can not be easily detected.
  • Kangas, University of Helsinki, Department of Forest Management, P.O. Box 27, FI-00014 University of Helsinki E-mail: annika.kangas@helsinki.fi (email)
  • Haapakoski, University of Helsinki, Department of Forest Management, P.O. Box 27, FI-00014 University of Helsinki E-mail: rh@nn.fi
  • Tyrväinen, Finnish Forest Research Institute, P.O. Box 16, FI-96301 Rovaniemi, Finland. E-mail: lt@nn.fi
article id 466, category Research article
Ilkka Korpela, Tuukka Tuomola, Timo Tokola, Bo Dahlin. (2008). Appraisal of seedling stand vegetation with airborne imagery and discrete-return LiDAR – an exploratory analysis. Silva Fennica vol. 42 no. 5 article id 466. https://doi.org/10.14214/sf.466
Keywords: silviculture; remote sensing; forest regeneration; classification; species
Abstract | View details | Full text in PDF | Author Info
The potential for combined use of airborne discrete-return LiDAR and digital imagery in the classification and measurement of common seedling stand vegetation was examined in southern Finland (61°50’N, 24°20’E). Classification was based on spectral and textural image features in addition to geometric and radiometric features of the LiDAR. The accuracy of leaf-on, LiDAR-based terrain elevation models was tested as well as the accuracy of LiDAR in the measurement of vegetation heights. LiDAR-based canopy height and the range-normalized intensity of the LiDAR were strong explanatory variables in vegetation classification. Interspecies variation was observed in the height measurement accuracy of LiDAR for different tree, shrub and low vegetation canopies. Elevation models derived with 1–15 pulses per m2 showed an inherent noise of app. 15–25 cm, which restricts the use of LiDAR in regeneration assessment of very young stands. The spatial pattern of the competing vegetation was reproduced in classification-based raster surfaces, which could be useful in deriving meaningful treatment proposals.
  • Korpela, University of Helsinki, Dept of Forest Management, P.O. Box 27, FI-00014 University of Finland E-mail: ilkka.korpela@helsinki.fi (email)
  • Tuomola, University of Helsinki, Dept of Forest Management, P.O. Box 27, FI-00014 University of Finland E-mail: tt@nn.fi
  • Tokola, University of Helsinki, Dept of Forest Management, P.O. Box 27, FI-00014 University of Finland E-mail: tt@nn.fi
  • Dahlin, University of Helsinki, Dept of Forest Management, P.O. Box 27, FI-00014 University of Finland E-mail: bd@nn.fi
article id 465, category Research article
Ken Olaf Storaunet, Jørund Rolstad, Målfrid Toeneiet, Erlend Rolstad. (2008). Effect of logging on the threatened epiphytic lichen Usnea longissima: a comparative and retrospective approach. Silva Fennica vol. 42 no. 5 article id 465. https://doi.org/10.14214/sf.465
Keywords: Picea abies; forest history; dendroecology; epiphytic lichens; historic logging; selective logging; stand reconstruction; Usnea longissima
Abstract | View details | Full text in PDF | Author Info
  • Storaunet, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway E-mail: ken.storaunet@skogoglandskap.no (email)
  • Rolstad, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway E-mail: jr@nn.no
  • Toeneiet, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway E-mail: mt@nn.no
  • Rolstad, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway E-mail: er@nn.no
article id 227, category Research article
Ola Lindroos. (2008). The effects of increased mechanization on time consumption in small-scale firewood processing. Silva Fennica vol. 42 no. 5 article id 227. https://doi.org/10.14214/sf.227
Keywords: comparative time study; fuelwood; blade saw; wedge splitter; processor; senior worker; operator effects
Abstract | View details | Full text in PDF | Author Info
Firewood, which is mainly processed by the consumer, is still an important source of energy for heating houses in industrialised countries. Possibilities to compare the mechanization’s impact on efficiency of firewood processing are limited, due to variations between working conditions in previous studies. Therefore, the objective was to compare time consumption for two small-scale firewood processing systems with different levels of mechanisation under identical conditions. The systems were tested on two classes of wood: one with a homogeneous and medium-sized diameter of logs and one with a mixture of small and large-diameter-logs. Differences in time consumption were analysed for correlations with physical workloads, deviations to routine operations, operator influences and operator perceptions. Twelve operators (60–79 years old) were studied and they showed large variation in time consumption. However, the within-operator time consumption patterns were consistent. In other words, operators all responded similarly to the different combinations of systems and wood classes, but at different absolute levels. The time required to process a unit volume of wood was 25–33% lower when the more highly mechanised system was used, and the time required was 13–22% lower for the homogeneous wood class. Physical work load, deviations and perceptions of the work varied between operators, but were weakly correlated with time consumption. The results’ implications for analyses of investments in equipment for firewood-processing for self-sufficiency purposes are discussed.
  • Lindroos, Swedish University of Agricultural Sciences, Department of Forest Resource Management, SE-901 83 Umeå, Sweden E-mail: ola.lindroos@srh.slu.se (email)
article id 226, category Research article
Anneli Viherä-Aarnio, Pirkko Velling. (2008). Seed transfers of silver birch (Betula pendula) from the Baltic to Finland – effect on growth and stem quality. Silva Fennica vol. 42 no. 5 article id 226. https://doi.org/10.14214/sf.226
Keywords: Betula pendula; yield; provenance; climatic adaptation; stem defect
Abstract | View details | Full text in PDF | Author Info
Silver birch (Betula pendula Roth) seed origins from the Baltic countries, Finland and Russia were compared for survival, growth and stem quality, and the effect of latitudinal seed transfer distance examined in two provenance trials. The trials were located on moist upland forest soils at Tuusula (60°21’N) in southern Finland and at Viitasaari (63°11’N) in central Finland. The material consisted of 21 stand and single tree origins ranging from latitudes 54° to 63°N. Survival, height, dbh, relative stem taper, stem volume/ha and the proportion of trees with a stem defect (vertical branch or forked stem), were assessed when the trees were 22 years old. Significant differences were detected among the origins regarding all the measured traits in both trials. Southern Finnish origins produced the highest volume per unit area in central Finland, whereas Estonian and north Latvian stand seed origins, as well as the southern Finnish plus tree origins, were the most productive ones in southern Finland. The more southern the origin, the higher was the proportion of trees with a stem defect in both trials. The latitudinal seed transfer distance had a significant but relatively small effect on survival, stem volume/ha and proportion of trees with a stem defect. The proportion of trees with a stem defect increased linearly in relation to the seed transfer distance from the south. The relationship of both survival and stem volume/ha to the seed transfer distance was curvilinear. Volume/ha was increased by transferring seed from ca. 2 degrees of latitude from the south. A longer transfer from the south, as well as transfer from the north, decreased the yield.
  • Viherä-Aarnio, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: anneli.vihera-aarnio@metla.fi (email)
  • Velling, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: pv@nn.fi
article id 225, category Research article
Johan Bergh, Urban Nilsson, Harald Grip, Per-Ola Hedwall, Tomas Lundmark. (2008). Effects of frequency of fertilisation on production, foliar chemistry and nutrient leaching in young Norway spruce stands in Sweden. Silva Fennica vol. 42 no. 5 article id 225. https://doi.org/10.14214/sf.225
Keywords: boreal; sludge; wood-ash; N; P; K
Abstract | View details | Full text in PDF | Author Info
There is a great need to increase the production in Swedish forests to meet future demand from the forest industry and the bio-energy sector. One option to increase the production is to supply nutrients to young stands of Norway spruce. For the practical application it is important to develop and optimise fertilisation regimes in terms of production, economy and leaching of nutrients. The frequency of fertilisation is one important variable in the fertilisation regime, and this study aimed to study effects of different fertilisation frequencies on production and leaching of nitrogen. In 2001, five field experiments were established in southern, central and northern Sweden. Young stands of Norway spruce were fertilised every year, every second year and every third year. In addition, fertilisation with sludge pellets and wood-ash combined with nitrogen was investigated. The current annual increment after five years of treatment was significantly larger in fertilised than in unfertilised treatments. The difference in production between fertilisation every year and every second year was insignificant, while fertilisation every third year resulted in lower production. Sludge pellets and wood-ash fertilisation gave significantly lower production than fertilisation every second year even though approximately the same amount of nitrogen was applied. There was relatively little leaching of nitrate to ground water in all treatments; 0.6–1 kg N ha–1 a–1 from plots with fertilisation every year or every second year; and 2.7 kg N ha–1 a–1 from plots with fertilisation every third year. Most of the leaching was after the first fertilisation, in all treatments at all sites.
  • Bergh, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: johan.bergh@ess.slu.se (email)
  • Nilsson, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: un@nn.se
  • Grip, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: hg@nn.se
  • Hedwall, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: poh@nn.se
  • Lundmark, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: tl@nn.se
article id 224, category Research article
Xiangwen Xiao, Xiao Xu, Fan Yang. (2008). Adaptive responses to progressive drought stress in two Populus cathayana populations. Silva Fennica vol. 42 no. 5 article id 224. https://doi.org/10.14214/sf.224
Keywords: antioxidant enzymes; free proline; malondialdehyde; soluble sugars; drought tolerance; Populus cathayana
Abstract | View details | Full text in PDF | Author Info
The young, vegetatively propagated cuttings of Populus cathayana Rehder were exposed to a progressive drought stress for 12 weeks in a greenhouse to characterize the physiological and biochemical basis of drought adaptation in woody plants. Two contrasting populations were employed in our study, which were from the wet and dry climate regions in western China, respectively. The results showed that the adaptive responses of P. cathayana to drought were affected by drought intensity and poplar genotype (population). The progressive drought stress significantly inhibited plant growth, increased carotenoid contents and, at the same time, accumulated soluble sugars and free proline in the plants of both populations tested. On the other hand, the gradually increasing drought also induced antioxidative systems including the increase of the activities of superoxide dismutase (SOD) and guaiacol peroxidase (POD). Moreover, there were different responses to progressive drought stress between the two contrasting populations. Compared with the wet climate population, the dry climate population had lower shoot height and growth rate, higher free proline content, and more efficient photoprotective system (such as higher carotenoid content and Car/Chl) and antioxidant system (such as higher POD activity), as a result of drought stress. These results suggest that the dry climate population possesses better drought tolerance than the wet climate population. The differences in drought tolerance may be closely related with efficient photoprotective system, accumulation of the osmoprotectant proline as well as the increased capacity of the antioxidative system to scavenge reactive oxygen species, and the consequent suppressed level of lipid peroxidation under drought conditions.
  • Xiao, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O. Box 416, Chengdu 610041, P. R. China; Graduate School of the Chinese Academy of Sciences, Beijing 100039, P. R. China E-mail: xiaoxw@cib.ac.cn (email)
  • Xu, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O. Box 416, Chengdu 610041, P. R. China; Graduate School of the Chinese Academy of Sciences, Beijing 100039, P. R. China E-mail: xx@nn.cn
  • Yang, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O. Box 416, Chengdu 610041, P. R. China; Graduate School of the Chinese Academy of Sciences, Beijing 100039, P. R. China E-mail: fy@nn.cn
article id 239, category Research article
Matti Stendahl, Anders Roos. (2008). Antecedents and barriers to product innovation – a comparison between innovating and non-innovating strategic business units in the wood industry. Silva Fennica vol. 42 no. 4 article id 239. https://doi.org/10.14214/sf.239
Keywords: product development; obstacles for innovation; innovation strategy; organization
Abstract | View details | Full text in PDF | Author Info
Increased competitive pressure from low-cost economies and substituting materials has raised the need for new strategies focusing on product differentiation in the Nordic wood industry. With the aim to identify factors that can facilitate increased product innovation activity, this study compared organizational characteristics and perceived barriers to product development among innovating and non-innovating strategic business units (SBUs) in the Swedish and Finnish wood industry. Multivariate analysis of data from a cross-sectional sample of 110 SBUs suggested that organizational size and educational level among white-collar workers are significant antecedents of product innovation activity. Furthermore, the difficulty of giving practical priority to development work in the everyday stress was identified as the most important perceived barrier to product development among managers in both innovating and non-innovating SBUs. A low competence level among the personnel and a low need to innovate was perceived to be the second most important barriers to product development among managers in, respectively, innovating and non-innovating SBUs. Practitioners who wish to increase product innovation activity in the wood industry are advised to promote an increased educational level in wood industry companies. They are further encouraged to seek ways to reduce the perceived barriers to product innovation identified in this study.
  • Stendahl, Swedish University of Agricultural Sciences, Department of Forest Products, P.O. Box 7008, SE-750 07 Uppsala, Sweden E-mail: matti.stendahl@sprod.slu.se (email)
  • Roos, Swedish University of Agricultural Sciences, Department of Forest Products, P.O. Box 7008, SE-750 07 Uppsala, Sweden E-mail: ar@nn.se
article id 238, category Research article
Mikko Kurttila, Pekka Leskinen, Jouni Pykäläinen, Tiina Ruuskanen. (2008). Forest owners' decision support in voluntary biodiversity-protection projects. Silva Fennica vol. 42 no. 4 article id 238. https://doi.org/10.14214/sf.238
Keywords: biodiversity protection; compensation fee; multi-criteria decision support; multi-objective forest planning
Abstract | View details | Full text in PDF | Author Info
New forest-biodiversity-protection instruments based on temporary protection periods and non-industrial private forest owners’ voluntary participation have been recently introduced and tested in pilot areas located in Southern Finland. Thanks to their several benefits, the use of voluntary instruments is becoming more common in many other countries as well. Voluntary protection here means that forest owners voluntarily set aside tracts of forest to be protected and define their compensation fees. Depending on the objectives of the forest owners, the compensation fee reflects the forest owners’ (positive) attitude towards biodiversity, scenic beauty, recreational values and/or the existence of long-term cutting possibilities. When a forest owner decides to offer part of his/her forest holding to be temporarily protected, the owner faces a new decision problem related to definition of the compensation fee, which should be based on diverse information concerning stand- and holding-level opportunity costs as well as on the biodiversity value of the stand. This article introduces three decision-support elements for assisting forest owners in defining their compensation fees. The first element relates to the assessment of the potential stand-level loss of timber harvesting income that the temporary protection of the stand may cause. The second element sets the holding-level opportunity cost of protection by utilizing the forest owners’ holding level goals, the holdings’ production possibilities and optimization methods. The third element describes the biodiversity value of the stand by means of a multi-criteria expert model. Case study material collected from the area of Central Karelia Herb-rich Forests Network pilot project is used to illustrate the characteristics of the decision-support elements and to point out some development needs for the future use of these elements.
  • Kurttila, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: mikko.kurttila@joensuu.fi (email)
  • Leskinen, Finnish Environment Institute, Research Programme for Production and Consumption, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: pl@nn.fi
  • Pykäläinen, Metsämonex Ltd., Joensuu, Finland E-mail: jp@nn.fi
  • Ruuskanen, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: tr@nn.fi
article id 237, category Research article
Jussi Peuhkurinen, Matti Maltamo, Jukka Malinen. (2008). Estimating species-specific diameter distributions and saw log recoveries of boreal forests from airborne laser scanning data and aerial photographs: a distribution-based approach. Silva Fennica vol. 42 no. 4 article id 237. https://doi.org/10.14214/sf.237
Keywords: diameter distribution; non-parametric estimation; pre-harvest measurement; stem data bank
Abstract | View details | Full text in PDF | Author Info
The low-density airborne laser scanning (ALS) data based estimation methods have been shown to produce accurate estimates of mean forest characteristics and diameter distributions, according to several studies. The used estimation methods have been based on the laser canopy height distribution approach, where various laser pulse height distribution -derived predictors are related to the stand characteristics of interest. This approach requires very delicate selection methods for selecting the suitable predictor variables. In this study, we introduce a new nearest neighbor search method that requires no complicated selection algorithm for choosing the predictor variables and can be utilized in multipurpose situations. The proposed search method is based on Minkowski distances between the distributions extracted from low density ALS data and aerial photographs. Apart from the introduction of a new search method, the aims of this study were: 1) to produce accurate species-specific diameter distributions and 2) to estimate factual saw log recovery, using the estimated height-diameter distributions and a stem data bank. The results indicate that the proposed method is suitable for producing species-specific diameter distributions and volumes at the stand level. However, it is proposed, that the utilization of more extensive and locally emphasized reference data and auxiliary variables could yield more accurate saw log recoveries.
  • Peuhkurinen, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: jp@nn.fi (email)
  • Maltamo, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: mm@nn.fi
  • Malinen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jm@nn.fi
article id 236, category Research article
Jaakko Repola. (2008). Biomass equations for birch in Finland. Silva Fennica vol. 42 no. 4 article id 236. https://doi.org/10.14214/sf.236
Keywords: birch; tree biomass; biomass functions; biomass of tree components
Abstract | View details | Full text in PDF | Author Info
Biomass equations were compiled for the above- and below-ground tree components of birch (Betula pendula Roth and Betula pubescens Ehrh.). The equations were based on 127 sample trees in 24 birch stands located on mineral soil sites. The study material consisted of 20 temporary plots and ten plots from four thinning experiments with different thinning intensities (unthinned, moderately and heavily thinned plots). The equations were estimated for the following individual tree components: stem wood, stem bark, living and dead branches, foliage, stump, and roots. In the data analysis, a multivariate procedure was applied in order to take into account the statistical dependency among the equations. Three multivariate variance component models were constructed for the above-ground biomass components, and one for the below-ground biomass components. The multivariate model (1) was mainly based on tree diameter and height, and in the multivariate models (2) and (3) additional commonly measured tree variables were used. The equations provided logical biomass predictions for a number of tree components, and were comparable with other functions used in Finland and Sweden. The applied statistical method generated equations that gave more reliable biomass estimates than the equations presented earlier. Furthermore, the structure of the multivariate models enables more flexible application of the equations, especially for research purposes.
  • Repola, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: jaakko.repola@metla.fi (email)
article id 235, category Research article
Ari Nikula, Ville Hallikainen, Risto Jalkanen, Mikko Hyppönen, Kari Mäkitalo. (2008). Modelling the factors predisposing Scots pine to moose damage in artificially regenerated sapling stands in Finnish Lapland. Silva Fennica vol. 42 no. 4 article id 235. https://doi.org/10.14214/sf.235
Keywords: Pinus sylvestris; boreal forest; forestry; Alces alces; damage; modelling
Abstract | View details | Full text in PDF | Author Info
Moose (Alces alces) damage in forest plantations have been at a high level in Finland in recent decades. Nowadays, moose is the most severe pest in Scots pine plantations also in Finnish Lapland. So far, despite the high level of damage and different bio-geographical conditions in Northern Finland, most of the moose-damage research has been carried out in Southern Finland. A number of research have also been performed to analyse factors affecting browsing but predictive models are rare. Data from 123 randomly selected and artificially regenerated pine plantations in Northern Finland were used in modelling the risk of moose browsing. The stands had been regenerated during 1984–1995. A total of 508 sample plots (range 2–8 plots per stand) were measured. Hierarchical logistic regression models with a random factor were constructed to predict the probability of leader-shoot browsing of pine on a plot. The number of planted pines and deciduous trees overtopping the pines were the most important predictors increasing the browsing probability. The results support earlier findings that deciduous trees overtopping or reaching the height of the pines should be cleaned from the immediate vicinity of the pines. Seedlings with a height ranging from 75 to 299 centimetres were more susceptible to browsing. Heavy soil scarification, such as ploughing or mounding, increased the browsing probability compared with lighter scarification methods. Soil type did not affect the browsing probability, but paludification decreased it. The within-stand variation in deciduous trees density and height should be taken into account in future moose browsing risk assessments. In Lapland, high moose damage risk areas are characterized by a low elevation and higher temperature sum.
  • Nikula, Finnish Forest Research Institute, Rovaniemi Research Unit, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: ari.nikula@metla.fi (email)
  • Hallikainen, Finnish Forest Research Institute, Rovaniemi Research Unit, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: vh@nn.fi
  • Jalkanen, Finnish Forest Research Institute, Rovaniemi Research Unit, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: rj@nn.fi
  • Hyppönen, Finnish Forest Research Institute, Rovaniemi Research Unit, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: mh@nn.fi
  • Mäkitalo, Finnish Forest Research Institute, Rovaniemi Research Unit, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: km@nn.fi
article id 234, category Research article
Enni Flykt, Sari Timonen, Taina Pennanen. (2008). Variation of ectomycorrhizal colonisation in Norway spruce seedlings in Finnish forest nurseries. Silva Fennica vol. 42 no. 4 article id 234. https://doi.org/10.14214/sf.234
Keywords: Picea abies; nursery; containerised seedlings; ectomycorrhiza; fungal diversity
Abstract | View details | Full text in PDF | Author Info
Ectomycorrhizal (ECM) colonisation patterns and seedling growth of containerised spruce seedlings were studied in five typical Finnish forest nurseries by morphotyping and molecular characterisation. ECM colonisation degree of 1-year-old spruce seedlings was below 20% in all studied Finnish forest nurseries. In 2-year-old spruce seedlings the ECM colonisation degree was ca. 50–60% in three of the nurseries, but negligible in others. The ECM fungal species richness varied from 0.1 to 3.8 types per seedling. Altogether seven ECM morphotypes were distinguished. The clearest factors associated with ECM colonisation patterns were nitrogen and phosphorus fertilisation. Particularly fertilisation in the early stage of seedling development appeared to diminish the degree of colonisation and species richness of ECM fungi. Root/shoot ratio was positively correlated with high colonisation degree and species richness of ECM fungi. Higher fertilisation inputs in these overall fertilisation levels did not increase the size of the seedlings. According to these results moderate fertilisation levels particularly in the beginning of seedling cultivation are critical for generating higher root/shoot ratios and sufficient ECM colonisation degree of the roots.
  • Flykt, University of Helsinki, Department of Applied Chemistry and Microbiology, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ef@nn.fi (email)
  • Timonen, University of Helsinki, Department of Applied Chemistry and Microbiology, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: st@nn.fi
  • Pennanen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: tp@nn.fi
article id 233, category Research article
Raija Laiho, Sakari Sarkkola, Seppo Kaunisto, Jukka Laine, Kari Minkkinen. (2008). Macroscale variation in peat element concentrations in drained boreal peatland forests. Silva Fennica vol. 42 no. 4 article id 233. https://doi.org/10.14214/sf.233
Keywords: drainage; peat soil; nutrient deficiencies; spatial variation; soil nutrients
Abstract | View details | Full text in PDF | Author Info
Information on the variation in soil element concentrations at different spatial scales is needed for, e.g., designing efficient sampling strategies, upscaling the processes related to carbon cycling, and planning land use and management. In spite of intensive land use, such information concerning peat soils is still scarce. We analyzed the variation in peat mineral element concentrations in boreal peatland forests drained 50–60 years earlier. We wanted to quantify the proportions of variation deriving from differences between regions and peatland basins and from within-peatland heterogeneity, and to model the variation using relatively easily measurable site and soil characteristics. We utilized 878 peat samples representing the 0–20 cm layer and collected from 289 sites in 79 peatland basins. The sites represented three different drained peatland forest site types. The two strongest gradients in the element composition captured by principal component analysis were correlated with both the North-South gradient and the site type variation, and the East-West gradient. In general, most of the variation in the element concentrations was contributed by differences among peatland basins, and variation within the floristically determined sites. Most of the element concentrations were best modeled when either the bulk density or the ash content of the peat, or both, were used in addition to site type and geographical location. The explanatory power remained modest for most element concentrations. As for the P concentrations in soil, however, our models provide means for estimating a large part of the variation among drained pine mire sites.
  • Laiho, University of Helsinki, Department of Forest Ecology, FI-00014 University of Helsinki, Finland E-mail: raija.laiho@helsinki.fi (email)
  • Sarkkola, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: ss@nn.fi
  • Kaunisto, Finnish Forest Research Institute, Parkano Research Unit, Kaironiementie 54, FI-39700 Parkano, Finland E-mail: sk@nn.fi
  • Laine, Finnish Forest Research Institute, Parkano Research Unit, Kaironiementie 54, FI-39700 Parkano, Finland E-mail: jl@nn.fi
  • Minkkinen, University of Helsinki, Department of Forest Ecology, FI-00014 University of Helsinki, Finland E-mail: km@nn.fi
article id 232, category Research article
Thomas Wutzler. (2008). Effect of the aggregation of multi-cohort mixed stands on modeling forest ecosystem carbon stocks. Silva Fennica vol. 42 no. 4 article id 232. https://doi.org/10.14214/sf.232
Keywords: stand structure; thinning; inventory; scale; model; stratification; bias; inventorystand structure
Abstract | View details | Full text in PDF | Author Info
Studies of the carbon sink of forest ecosystems often stratify the studied stands by the dominating species and thereby abstract from differences in the mixed-species, multi-cohort structure of many forests. This case study infers whether the aggregation of forestry data introduces a bias in the estimates of carbon stocks and their changes at the scale of individual stands and the scale of a forest district. The empirical TreeGrOSS-C model was applied to 1616 plots of a forest district in Central Germany to simulate carbon dynamics in biomass, woody debris, and soil. In a first approach each stand was explicitly simulated with all cohorts. In three other approaches the forest inventory data were aggregated in several ways, including a stratification of the stands to 110 classes according to the dominating species, age class, and site conditions. A small but significant bias was confirmed. At stand scale the initial ecosystem carbon stocks by the aggregated approach differed from that of the detailed approach by 2.3%, but at the district scale only by 0.05%. The differences in age between interspersed and dominant cohorts as well as differences in litter production were important for the differences in initial carbon stocks. The amounts of wood extracted by thinning operations were important for the differences in the projection of the carbon stocks over 100 years. Because of the smallness of bias, this case study collects evidence that the approaches, that represent stands or stratums by a single cohort, are valid at the scale of a forest district or larger.
  • Wutzler, Max Planck Institute for Biogeochemistry, Hans Knöll Str. 10, DE-07745, Jena, Germany E-mail: thomas.wutzler@bgc-jena.mpg.de (email)
article id 231, category Research article
Hilppa Gregow, Ulla Puranen, Ari Venäläinen, Heli Peltola, Seppo Kellomäki, David Schultz. (2008). Temporal and spatial occurrence of strong winds and large snow load amounts in Finland during 1961-2000. Silva Fennica vol. 42 no. 4 article id 231. https://doi.org/10.14214/sf.231
Keywords: soil frost; coniferous forest; wind climate; precipitation; snow
Abstract | View details | Full text in PDF | Author Info
Information on the temporal and spatial occurrence of strong winds and snow loads on trees is important for the risk management of wind- and snow-induced damage. Meteorological measurements made at 19 locations across Finland during 1961–2000 are used to understand the temporal and spatial occurrence of strong winds and large snow loads. A Kriging interpolation method was used to produce a spatial analysis of wind-speed events above 11 m s–1, 14 m s–1, and greater or equal to 17 m s–1 and snowfall accumulation above 20 kg m–2 and 30 kg m–2. According to the analysis, wind speeds exceeded 14 m s–1 at least 155 times and reached 17 m s–1 only 5 times at inland locations during the 40 years. Large snowfall accumulations were more frequent in the higher-elevation inland areas than along the coast. The snow load on trees exceeded 20 kg m–2 about 65 times a year when averaged over all 40 years, but was as high as 150 times a year during the mild 1990s. The maximum number of heavy snow-load events occurred in 1994 in northern Finland, consistent with a forest inventory by the Finnish Forest Research Institute in 1992–1994. The findings of this study imply that the risk of wind-induced damage is highest in the late autumn when trees do not have the additional support of frozen soil. In contrast, the risk of snow-induced damage is highest at higher-elevations inland, especially in northern Finland.

* Erratum (23 Oct 2012): The authors have requested inclusion of an additional author. Author information should thus be as follows: Hilppa Gregow, Ulla Puranen, Ari Venäläinen, Heli Peltola, Seppo Kellomäki & David Schultz
  • Gregow, Finnish Meteorological Institute, P.O. Box 503, FI-00101 Helsinki, Finland E-mail: hilppa.gregow@fmi.fi (email)
  • Puranen, Finnish Meteorological Institute, P.O. Box 503, FI-00101 Helsinki, Finland E-mail: up@nn.fi
  • Venäläinen, Finnish Meteorological Institute, P.O. Box 503, FI-00101 Helsinki, Finland E-mail: av@nn.fi
  • Peltola, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: heli.peltola@uef.fi
  • Kellomäki, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: seppo.kellomaki@uef.fi
  • Schultz, Finnish Meteorological Institute, P.O. Box 503, FI-00101 Helsinki, Finland E-mail: ds@nn.fi
article id 230, category Research article
Risto Jalkanen, Sheila Hicks, Tarmo Aalto, Hannu Salminen. (2008). Past pollen production reconstructed from needle production in Pinus sylvestris at the northern timberline: a tool for evaluating palaeoclimate reconstructions. Silva Fennica vol. 42 no. 4 article id 230. https://doi.org/10.14214/sf.230
Keywords: Scots pine; age-depth chronology; annual resolution; needle trace method; NTM; pollen accumulation rate; temperature reconstruction; tree line
Abstract | View details | Full text in PDF | Author Info
Annual needle production (PROD) of Scots pine (Pinus sylvestris L.) and pine pollen accumulation rates (PAR) are compared along a 5-site transect from the Arctic Circle to the northern timberline. PROD is calculated using the Needle Trace Method (NTM). PAR is monitored by two series of pollen traps, located in the centres of mires and within forests, respectively. There is a strong year-to-year agreement in PAR and PROD between the sites for the common 19-year period for which both proxies are available. Mean July temperature of the previous year (TJUL–1) correlates statistically significantly with PROD at all five sites and with PAR in the four northernmost sites. There is also a significant relationship between TJUN–1 and PROD at all sites, and TJUN and PAR at the two northernmost sites. PROD and PAR correlate most strongly in the three near tree line sites, where PROD explains up to 51% of the variation in PAR. On the basis of the calibration between PROD, PAR and TJUL–1, PROD and TJUL–1 are used to reconstruct past PAR. That such a reconstruction is realistic is supported by its agreement with the pollen record for 1982–2000 and with records of male flowering for the period 1956–1973. The use of PROD in reconstructing past PAR can help in interpreting the fossil pollen signal in terms of climate rather than vegetation change and in evaluating the high-resolution dating of peat profiles and calculations of the rate of peat accumulation.
  • Jalkanen, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: risto.jalkanen@metla.fi (email)
  • Hicks, Institute of Geosciences, P.O. Box 3000, FI-90014 University of Oulu, Finland E-mail: sh@nn.fi
  • Aalto, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: ta@nn.fi
  • Salminen, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: hs@nn.fi
article id 229, category Research article
Carmen Martín, Teresa Parra, Margarita Clemente-Muñoz, Esteban Hernández-Bermejo. (2008). Genetic diversity and structure of the endangered Betula pendula subsp. fontqueri populations in the south of Spain. Silva Fennica vol. 42 no. 4 article id 229. https://doi.org/10.14214/sf.229
Keywords: Betula pendula; B. pendula subsp. fontqueri; genetic diversity; population; RAPD
Abstract | View details | Full text in PDF | Author Info
Betula pendula subsp. fontqueri, present in the south of Spain, has been considered in danger of extinction and, for this reason, some regional governments in Spain have included their populations in conservation programmes. In order to establish the genetic structure of the Betula pendula subsp. fontqueri populations, a random amplified polymorphic DNA (RAPD) analysis was carried out. Two B. pubescens populations were included in the study as taxonomic controls. B. pendula subsp. fontqueri populations were clearly differentiated through UPGMA, and showed significant pairwise genetic distance (ΦST) values between all pairs of populations obtained by AMOVA. Genetic diversity found between populations was not correlated to geographical distances. The significant differences among populations must be due to progressive isolation of Betula populations along their paleogeographical history, and more recently to the drastic fragmentation and reduction of some of these populations. The results obtained in this work show clear genetic differences which could be considered in the management of conservation strategies for Betula pendula subsp. fontqueri in its Iberian meridional distribution.
  • Martín, Departamento de Biología Vegetal, Escuela Técnica Superior de Ingenieros Agrónomos de Madrid, Universidad Politécnica de Madrid, Ciudad Universitaria s/n, 28040-Madrid, Spain E-mail: mariacarmen.martin@upm.es (email)
  • Parra, Departamento de Ciencias y Recursos Agrícolas y Forestales, Universidad de Córdoba, Avda. Linneo s/n, 14004-Córdoba, Spain E-mail: tp@nn.es
  • Clemente-Muñoz, Departamento de Ciencias y Recursos Agrícolas y Forestales, Universidad de Córdoba, Avda. Linneo s/n, 14004-Córdoba, Spain E-mail: mcm@nn.es
  • Hernández-Bermejo, Departamento de Ciencias y Recursos Agrícolas y Forestales, Universidad de Córdoba, Avda. Linneo s/n, 14004-Córdoba, Spain E-mail: ehb@nn.es
article id 248, category Research article
Tuula Jyske, Harri Mäkinen, Pekka Saranpää. (2008). Wood density within Norway spruce stems. Silva Fennica vol. 42 no. 3 article id 248. https://doi.org/10.14214/sf.248
Keywords: Picea abies; wood density; inter-ring variation; intra-ring variation; maturation
Abstract | View details | Full text in PDF | Author Info
We studied the variation in average wood density of annual rings, earlywood density, and latewood density in addition to ring width and latewood percentage within Norway spruce (Picea abies (L.) Karst.) stems from the pith to the bark, and from the stem base towards the stem apex. Moreover, the variation in wood density within annual rings was studied at the different heights and radial positions in the stem. The material consisted of 85 trees from central and south-eastern Finland. Variation between the annual rings accounted for 11–27% of the total variation in wood density. Only small differences (3–6%) in wood density were found between different heights in the stem. The largest (49–80%) variation in wood density was found within the annual rings. The difference in wood density between earlywood and latewood was smaller in the rings near the pith than in the outer rings. The increasing wood density from the pith outwards was related to increasing latewood density and latewood percentage, whereas the earlywood density increased only slightly from the pith outwards. In a given cambial age (i.e., given rings from the pith), the average wood density of annual rings increased with increasing stem height. In contrast, in the rings formed in the same calendar years (i.e., given rings from the bark), the average wood density of annual rings decreased with increasing stem height. The results of this study verify our knowledge of wood density variation and can further contribute to creating models to predict wood density.
  • Jyske, The Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: tuula.jyske@metla.fi (email)
  • Mäkinen, The Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: hm@nn.fi
  • Saranpää, The Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: ps@nn.fi
article id 247, category Research article
Ane Zubizarreta Gerendiain, Heli Peltola, Pertti Pulkkinen, Veli-Pekka Ikonen, Raimo Jaatinen. (2008). Differences in growth and wood properties between narrow and normal crowned types of Norway spruce grown at narrow spacing in Southern Finland. Silva Fennica vol. 42 no. 3 article id 247. https://doi.org/10.14214/sf.247
Keywords: fibre length; stem volume; height; diameter; wood density; earlywood; latewood
Abstract | View details | Full text in PDF | Author Info
In recent years there has been increased interest in the so called narrow crowned Norway spruce (Picea abies f. pendula), which is a rare mutant of Norway spruce (Picea abies (L.) Karsten), as a suitable wood raw material source for pulp and paper production. This is because it is less sensitive to competition than the normal crowned Norway spruce, and thus, could be more productive especially at dense spacing. In the above context, we investigated how the growth and yield (such as height, diameter, stem volume and ring width) in addition to wood density traits and fibre properties (such as wood density, fibre length and width, cell wall thickness and fibre coarseness) were affected in trees from 9 full-sib families representing narrow crowned Norway spruce grown at narrow spacing of 1 m 1 m in Southern Finland. For comparison, we used normal crowned Norway spruce trees from 6 breeding regions. We found that, compared to growth and yield traits, wood density traits and fibre properties showed, on average, lower phenotypic variations. In addition, these variations were smaller for narrow crowned families than for normal crowned genetic entries. Narrow crowned families also showed, on average, higher growth and yield and fibre length, but lower wood density. Moreover, the phenotypic correlations between growth, yield, wood density traits and fibre properties, ranged, on average, from moderate (narrow crowned) to high (normal crowned). As a whole, the growth and wood properties of narrow crowned families were found to be less sensitive to tree competition than the normal crowned genetic entries used as a comparison.
  • Zubizarreta Gerendiain, University of Joensuu, Faculty of Forest Sciences, Joensuu, Finland E-mail: ane.zubizarreta@joensuu.fi (email)
  • Peltola, University of Joensuu, Faculty of Forest Sciences, Joensuu, Finland E-mail: hp@nn.fi
  • Pulkkinen, Finnish Forest Research Institute, Haapastensyrjä Breeding Station, Läyliäinen, Finland E-mail: pp@nn.fi
  • Ikonen, University of Joensuu, Faculty of Forest Sciences, Joensuu, Finland E-mail: vpi@nn.fi
  • Jaatinen, Finnish Forest Research Institute, Haapastensyrjä Breeding Station, Läyliäinen, Finland E-mail: rj@nn.fi
article id 246, category Research article
Maria Jonsson. (2008). Live-storage of Picea abies for two summers after storm felling in Sweden. Silva Fennica vol. 42 no. 3 article id 246. https://doi.org/10.14214/sf.246
Keywords: Norway spruce; blue stain; MC; storage decay; windthrown
Abstract | View details | Full text in PDF | Author Info
After recent severe storm fellings in Sweden, as harvest, transport, and storage capacities were insufficient, interest in live-storage (leaving windthrown trees in the stand) increased. This study follows windthrown Norway spruce (Picea abies (L.) Karst.) trees during 20 months, i.e. two summers, of live-storage in southern Sweden. Moisture content, blue stain, and storage decay were compared in trees from a site with all trees windthrown and a site with scattered windthrows. After the first summer of live-storage, the quality losses were small. After 20 months, the trees had dried significantly and had numerous infestations of blue stain and storage decay. Trees from the site with scattered windthrows were of better quality compared to trees from the site with all trees windthrown. Live-storage is a suitable method for one year of storage, but the second year losses in wood quality are considerable.
  • Jonsson, Swedish University of Agricultural Sciences, Dept. of Forest Products, Uppsala, Sweden E-mail: maria.jonsson@sprod.slu.se (email)
article id 245, category Research article
Saija Huuskonen, Jari Hynynen, Risto Ojansuu. (2008). Stand characteristics and external quality of young Scots pine stands in Finland. Silva Fennica vol. 42 no. 3 article id 245. https://doi.org/10.14214/sf.245
Keywords: Scots pine; growth and yield; precommercial thinning; external stem quality; young stand management
Abstract | View details | Full text in PDF | Author Info
The effects of silvicultural practices (regeneration method and young stand management) on the stand characteristics of young Scots pine (Pinus sylvestris (L.)) stands were studied. Stand density, mean diameter, crown ratio and external quality of young Scots pine stands were analysed on the basis of extensive inventory data. The study material consisted of 181 stands containing inventory growth plots, representing the most common site types for Scots pine and covering all the important wood production areas in Finland. Intensive management practices, i.e. artificial regeneration and precommercial thinning, clearly enhanced mean diameter development of the stand. The overall stand density of the crop trees was relatively low in the material (1925 trees ha–1). In more than one third of the stands, the stem number of crop trees was below 1500 trees ha–1. Stand density was not affected by forest management, but it was slightly higher in Southern than in Northern Finland. The geographical location, in terms of annual effective temperature sum, affected the average slenderness and crown ratio. At a given mean stand diameter, the dominant height of the stand was lower, and the mean crown ratio was higher, in Northern than in Southern Finland. The average external quality of the Scots pine trees was relatively low. The proportion of trees without any observed defects was 54%. The most common external defects were curved stems (23%) and branchiness (9%). Branchiness was more frequent among the largest trees, while curved stems were more common in smaller trees. Defects were the most frequent in planted stands, and in stands growing on fresh sites. The defects were more frequent in Northern Finland than in Southern Finland. The relatively low stand density and poor external quality of the young stands emphasize the importance of stem quality as a tree selection criterion in commercial thinnings of Scots pine stands, if the goal is to produce high quality timber.
  • Huuskonen, University of Helsinki, Department of Forest Ecology, Helsinki, Finland E-mail: saija.huuskonen@helsinki.fi (email)
  • Hynynen, Finnish Forest Research Institute, Vantaa Research Unit, Vantaa, Finland E-mail: jh@nn.fi
  • Ojansuu, Finnish Forest Research Institute, Vantaa Research Unit, Vantaa, Finland E-mail: ro@nn.fi
article id 244, category Research article
Georg E. Kindermann, Ian McCallum, Steffen Fritz, Michael Obersteiner. (2008). A global forest growing stock, biomass and carbon map based on FAO statistics. Silva Fennica vol. 42 no. 3 article id 244. https://doi.org/10.14214/sf.244
Keywords: biomass map; downscaling; regression analysis
Abstract | View details | Full text in PDF | Author Info
Currently, information on forest biomass is available from a mixture of sources, including in-situ measurements, national forest inventories, administrative-level statistics, model outputs and regional satellite products. These data tend to be regional or national, based on different methodologies and not easily accessible. One of the few maps available is the Global Forest Resources Assessment (FRA) produced by the Food and Agriculture Organization of the United Nations (FAO 2005) which contains aggregated country-level information about the growing stock, biomass and carbon stock in forests for 229 countries and territories. This paper presents a technique to downscale the aggregated results of the FRA2005 from the country level to a half degree global spatial dataset containing forest growing stock; above/below-ground biomass, dead wood and total forest biomass; and above-ground, below-ground, dead wood, litter and soil carbon. In all cases, the number of countries providing data is incomplete. For those countries with missing data, values were estimated using regression equations based on a downscaling model. The downscaling method is derived using a relationship between net primary productivity (NPP) and biomass and the relationship between human impact and biomass assuming a decrease in biomass with an increased level of human activity. The results, presented here, represent one of the first attempts to produce a consistent global spatial database at half degree resolution containing forest growing stock, biomass and carbon stock values. All results from the methodology described in this paper are available online at www.iiasa.ac.at/Research/FOR/.
  • Kindermann, International Institute for Applied Systems Analysis, Laxenburg, Austria E-mail: kinder@iiasa.ac.at (email)
  • McCallum, International Institute for Applied Systems Analysis, Laxenburg, Austria E-mail: im@nn.at
  • Fritz, International Institute for Applied Systems Analysis, Laxenburg, Austria E-mail: sf@nn.at
  • Obersteiner, International Institute for Applied Systems Analysis, Laxenburg, Austria E-mail: mo@nn.at
article id 243, category Research article
Rupert Seidl, Werner Rammer, Petra Lasch, Franz-Werner Badeck, Manfred J. Lexer. (2008). Does conversion of even-aged, secondary coniferous forests affect carbon sequestration? A simulation study under changing environmental conditions. Silva Fennica vol. 42 no. 3 article id 243. https://doi.org/10.14214/sf.243
Keywords: forest management; Norway spruce; climate change mitigation; PICUS; 4C; model comparison
Abstract | View details | Full text in PDF | Author Info
To circumvent problems associated with even-aged, pure coniferous stands propagated outside their natural range alternative management strategies and conversion programs are currently discussed in Central Europe. However, a mainstreaming of such adapted silvicultural systems with climate change mitigation objectives is missing to date. In this study the objective was to assess in situ C storage under conditions of climate change in a secondary Norway spruce (Picea abies (L.) Karst.) forest management unit in Austria. Four management strategies (Norway spruce age class forestry, transition to continuous cover forestry with Norway spruce, conversion to mixed conifer/broadleaved stands, no management) were investigated under current climate and two transient climate change scenarios in a simulation study. By comparing the results of two independent forest ecosystem models (PICUS v1.41, 4C) applied under identical forcings and boundary conditions we aimed at addressing uncertainties in model-based projections. A transition to continuous cover forestry increased C storage in all climate scenarios (+45.4 tC·ha–1 to +74.0 tC·ha–1 over the 100 year analysis period) compared to the approximately balanced C budget under the age class system. For the mixed conifer/broadleaved management variant predictions of the two models diverged significantly (+29.4 tC·ha–1 and –10.6 tC·ha–1 in PICUS and 4C respectively, current climate). With regard to climate change impacts both models agreed on distinct effects on productivity but lower sensitivity of C stocks due to compensation from respiration and adaptive harvest levels. In conclusion, considering the potential effects of silvicultural decisions on C stocks climate change mitigation should be addressed explicitly in programs advocating targeted change in management paradigms.
  • Seidl, Institute of Silviculture, BOKU, Vienna, Austria E-mail: rupert.seidl@boku.ac.at (email)
  • Rammer, Institute of Silviculture, BOKU, Vienna, Austria E-mail: wr@nn.at
  • Lasch, Potsdam Institute for Climate Impact Research e.V., Potsdam, Germany E-mail: pl@nn.de
  • Badeck, Potsdam Institute for Climate Impact Research e.V., Potsdam, Germany E-mail: fwb@nn.de
  • Lexer, Institute of Silviculture, BOKU, Vienna, Austria E-mail: mjl@nn.at
article id 242, category Research article
Jyrki Hytönen, Paula Jylhä. (2008). Fifteen-year response of weed control intensity and seedling type on Norway spruce survival and growth on arable land. Silva Fennica vol. 42 no. 3 article id 242. https://doi.org/10.14214/sf.242
Keywords: Norway spruce; afforestation; frost damage; seedling size; seedling type; vegetation control
Abstract | View details | Full text in PDF | Author Info
The effects of seedling type (2-year-old container seedlings vs. 4-year-old bare-rooted seedlings) and post-planting vegetation control intensity on the growth and survival of Norway spruce (Picea abies (L.) Karst.) seedlings were compared based on 15-year data from a field experiment established on arable land. Vegetation control treatments with terbuthylazine and glyphosate were carried out 1–3 times on successive years, either as overall or spot applications. The highest stand volumes were obtained with the combination of large bare-rooted seedlings and effective vegetation control. Volume of bare-rooted seedlings was greater than that of container seedlings in all treatments (e.g. on the control plots 9.5 m3/ha vs. 4.1 m3/ha). The best results were obtained with the most intensive weed control treatments (spot treatment repeated twice and overall application repeated three times). These treatments increased both bare-rooted and containerised seedlings’ survival by 33–40% units and their height, breast height diameter, and volume by 45–49%, 17–47%, and 249–279%, respectively. In terms of survival, the container seedlings, in due part to their smaller size, benefited from vegetation control more than the bare-rooted seedlings. Successive early summer frosts damaged the seedlings and significantly retarded their growth. The frequency of frost damage was not affected by vegetation control nor was it attributed to seedling type.
  • Hytönen, Finnish Forest Research Institute, Kannus Research Unit, P.O. Box 44, FI-69101, Kannus, Finland E-mail: jyrki.hytonen@metla.fi (email)
  • Jylhä, Finnish Forest Research Institute, Kannus Research Unit, P.O. Box 44, FI-69101, Kannus, Finland E-mail: pj@nn.fi
article id 241, category Research article
Hailemariam Temesgen, Tara M. Barrett, Greg Latta. (2008). Estimating cavity tree abundance using Nearest Neighbor Imputation methods for western Oregon and Washington forests. Silva Fennica vol. 42 no. 3 article id 241. https://doi.org/10.14214/sf.241
Keywords: stand structure; nearest neighbor imputation; snag size; snag frequency; forest landscape modeling
Abstract | View details | Full text in PDF | Author Info
Cavity trees contribute to diverse forest structure and wildlife habitat. For a given stand, the size and density of cavity trees indicate its diversity, complexity, and suitability for wildlife habitat. Size and density of cavity trees vary with stand age, density, and structure. Using Forest Inventory and Analysis (FIA) data collected in western Oregon and western Washington, we applied correlation analysis and graphical approaches to examine relationships between cavity tree abundance and stand characteristics. Cavity tree abundance was negatively correlated with site index and percent composition of conifers, but positively correlated with stand density, quadratic mean diameter, and percent composition of hardwoods. Using FIA data, we examined the performance of Most Similar Neighbor (MSN), k nearest neighbor, and weighted MSN imputation with three variable transformations (regular, square root, and logarithmic) and Classification and Regression Tree with MSN imputation to estimate cavity tree abundance from stand attributes. There was a large reduction in mean root mean square error from 20% to 50% reference sets, but very little reduction in using the 80% reference sets, corresponding to the decreases in mean distances. The MSN imputation using square root transformation provided better estimates of cavity tree abundance for western Oregon and western Washington forests. We found that cavity trees were only 0.25 percent of live trees and 13.8 percent of dead trees in the forests of western Oregon and western Washington, thus rarer and more difficult to predict than many other forest attributes. Potential applications of MSN imputation include selecting and modeling wildlife habitat to support forest planning efforts, regional inventories, and evaluation of different management scenarios.
  • Temesgen, Department of Forest Resources, Oregon State University, Corvallis, OR, USA E-mail: hailemariam.temesgen@oregonstate.edu (email)
  • Barrett, Pacific Northwest Research Station, Anchorage, AK, USA E-mail: tmb@nn.us
  • Latta, Department of Forest Resources, Oregon State University, Corvallis, OR, USA E-mail: gl@nn.us
article id 240, category Research article
Monika Dering, Andrzej Lewandowski, Krzysztof Ufnalski, Aleksandra Kedzierska. (2008). How far to the east was the migration of white oaks from the Iberian refugium? Silva Fennica vol. 42 no. 3 article id 240. https://doi.org/10.14214/sf.240
Keywords: refugia; oaks; postglacial recolonization; Quercus robur; Quercus petrea
Abstract | View details | Full text in PDF | Author Info
The goal of this study was to investigate the postglacial recolonization pathways of the white oaks Quercus robur and Quercus petrea in Poland, and especially to evaluate the impact of Iberian refugium in this part of Europe. Chloroplast DNA polymorphism of 310 individuals older than 200 years was analyzed. Six haplotypes in total were found to differentiate three maternal lineages: the Balkan (haplotypes 4, 5, and 7), the Apennine (haplotypes 1 and 2), and the Iberian (haplotype 12). The most abundant were members of the Balkan (71.5% of all samples) and the Apennine lineage (23.1%), and only 5.4% of individuals were of Iberian origin. The geographic distribution of the three lineages is clearly structured. The northernmost territories of Poland are occupied by Apennine (haplotype 1) and Iberian (haplotype 12) lineages, whereas samples in central and southern Poland represents the Balkan lineage. The population structure might be the result of competitive colonization among lineages after migration from different refugia. It is likely that colonization of northernmost parts of Poland by the Balkan lineage was halted or at least hampered due to the arrival of the Apennine populations. The most significant result of this study concerns the presence and status of the Iberian lineage in Poland, which is most likely of natural origin.
  • Dering, Polish Academy of Sciences, Institute of Dendrology, Parkowa 5, Kornik, Poland E-mail: usnea@o2.pl (email)
  • Lewandowski, Polish Academy of Sciences, Institute of Dendrology, Parkowa 5, Kornik, Poland E-mail: al@nn.pl
  • Ufnalski, Polish Academy of Sciences, Institute of Dendrology, Parkowa 5, Kornik, Poland E-mail: ku@nn.pl
  • Kedzierska, Polish Academy of Sciences, Institute of Dendrology, Parkowa 5, Kornik, Poland E-mail: ak@nn.pl
article id 468, category Research article
Antti Lännenpää, Tuomas Aakala, Heikki Kauhanen, Timo Kuuluvainen. (2008). Tree mortality agents in pristine Norway spruce forests in northern Fennoscandia. Silva Fennica vol. 42 no. 2 article id 468. https://doi.org/10.14214/sf.468
Keywords: Picea abies; northern Fennoscandia; Coniophora; mortality agent; pristine forest
Abstract | View details | Full text in PDF | Author Info
We examined tree mortality agents in pristine old Norway spruce (Picea abies (L.) Karst.) forests in northern Finland and northwestern Russia. The data was collected on nine 40 m   400 m transects. The primary mortality agents of recently dead trees were recorded and their frequencies were calculated. The pattern of tree growth prior to death was studied based on increment core samples and compared with the growth of healthy dominant trees. Of all recently dead trees, 72% could be associated with a primary mortality agent. In both study areas the most common primary mortality agent was a Coniophora (Mérat) DC. -genus fungi, which was found on average in 33% of trees sampled. The fungi Phellinus chrysoloma (Fr.) Don and Onnia leporina (Fr.) H. Jahn as mortality agents were more common in the Finnish area compared to the Russian area. Analysis on the growth patterns indicated weak differences between different pathogens’ influence on prior-to-death growth of trees, so that fungi rotting the whole tree decreased tree growth more rapidly than fungi rotting only the heart wood. The results demonstrated that in old Norway spruce forests of northern Fennoscandia the most common primary tree mortality agents were wood rotting fungi, which weaken the mechanical stability of tree stems until they fall due to snow or wind, which should be considered only as secondary mortality agents. It is evident that tree death in pristine forest typically results from a long-lasting process involving both biotic and abiotic factors.
  • Lännenpää, Department of Forest Ecology, University of Helsinki, Finland E-mail: al@nn.fi
  • Aakala, Department of Forest Ecology, University of Helsinki, Finland E-mail: tuomas.aakala@helsinki.fi (email)
  • Kauhanen, Kolari Research Unit, Finnish Forest Research Institute, Finland E-mail: hk@nn.fi
  • Kuuluvainen, Department of Forest Ecology, University of Helsinki, Finland E-mail: tk@nn.fi
article id 259, category Research article
Jani Laturi, Jarmo Mikkola, Jussi Uusivuori. (2008). Carbon reservoirs in wood products-in-use in Finland: current sinks and scenarios until 2050. Silva Fennica vol. 42 no. 2 article id 259. https://doi.org/10.14214/sf.259
Keywords: climate change; wood products; carbon pool; present value; decay function
Abstract | View details | Full text in PDF | Author Info
This study addresses the question of how much carbon will be sequestered in wood products during the coming decades in Finland. Using sawnwood and other wood material consumption data since the 1950s and inventory data of carbon reservoirs of wood products in the Finnish construction and civil engineering sector, we first derive estimates for the carbon reservoirs in wood products-in-use in that sector. We then extend the estimate to include all wood products-in-use. We find that the carbon pool of wood products in the Finnish construction and civil engineering sector grew by about 12% since an inventory for 2000, and that the overall estimate for carbon reservoirs of Finnish wood products in 2004 was 26.6 million tons of carbon. In building the scenarios until 2050, econometric time series models accounting for the relationship between wood material consumption and the development of GDP were used. The results indicate that the range of carbon reservoirs of wood products in Finland will be 39.6–64.2 million tons of carbon in the year 2050. The impacts of different forms of the decay function on the time-path of a carbon sink and its value in wood products were also studied. When a logistic decay pattern is used, the discounted value of the predicted carbon sink of wood products in Finland is between EUR850 and EUR1380 million – at the price level of EUR15/CO2 ton – as opposed to 440–900 million euros, if a geometric decay pattern is used.
  • Laturi, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: jani.laturi@metla.fi (email)
  • Mikkola, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: jm@nn.fi
  • Uusivuori, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: ju@nn.fi
article id 258, category Research article
Ursula Schatz, Henrik Heräjärvi, Kari Kannisto, Matti Rantatalo. (2008). Influence of saw and secateur pruning on stem discolouration, wound cicatrisation and diameter growth of Betula pendula. Silva Fennica vol. 42 no. 2 article id 258. https://doi.org/10.14214/sf.258
Keywords: Betula pendula; colour defects; pruning saw; secateur; technical quality
Abstract | View details | Full text in PDF | Author Info
The aim of this case study was to compare the impacts of saw and secateur pruning on silver birch (Betula pendula Roth). Data were collected from two saw pruned stands in 2005, and one secateur pruned stand in 2003. All the stands were located in southern Finland. The sample stems were felled, and their butt logs were sawn into flitches, whose knot features and colour defects were measured. In addition, discs were sawn from each stem in order to study the annual ring widths. In this material, pruning with secateurs appeared to cause less colour defects than pruning with a saw. Irrespective of the pruning method used, the colour defects in the stem wood were at their largest in cases where the basal knob or the stem bark appeared to be damaged by pruning. Colour defects spread mainly towards the pith, only in a few cases towards the stem surface. The cicatrisation time of the knots as well as the length of the bark stick remaining inside the stem did not show significant differences between the two pruning methods. Pruning of the lowest living branches appears to have no effect on the diameter growth of silver birch trees.
  • Schatz, Peltotievantie 27, FI-99440 Leppäjärvi, Finland E-mail: us@nn.fin
  • Heräjärvi, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: henrik.herajarvi@metla.fi (email)
  • Kannisto, Finnish Forest Research Institute, Parkano Research Unit, Kaironiementie 54, FI-39700 Parkano, Finland E-mail: kk@nn.fi
  • Rantatalo, Peltotievantie 27, FI-99440 Leppäjärvi, Finland E-mail: mr@nn.fi
article id 257, category Research article
Nadir Ayrilmis. (2008). Effect of compression wood on dimensional stability of medium density fiberboard. Silva Fennica vol. 42 no. 2 article id 257. https://doi.org/10.14214/sf.257
Keywords: compression wood; linear expansion; linear contraction; thickness swelling; thickness shrinkage; medium density fiberboard; dimensional stability
Abstract | View details | Full text in PDF | Author Info
This study evaluated the effect of compression wood on dimensional stability of medium density fiberboards (MDF) manufactured from fiber furnishes of pine (Pinus nigra Arnold var. pallasiana) containing compression wood. Two panel types were manufactured from two different compression wood (CW) portion / normal wood (NW) portions in the furnish, 75/25 and 10/90, respectively. Linear and thickness variations of the panels exposed to various relative humidites at 20 °C, linear expansion/contraction and thickness swelling/shrinkage, were measured according to the procedures defined by DIN EN 318 (2005) standard test method. Panels made from fiber furnish containing 75% the CW had higher linear expansion and linear contraction values with an average value of 0.286% and 0.247% than those of panels made from fiber furnish containing 10% the CW with an average value of 0.184% and 0.152%, respectively. As for thickness swelling and thickness shrinkage properties, panels made from fiber furnish containing 75% the CW had the thickness swelling and thickness shrinkage values with an average of 5.042% and 4.402% while panels made from fiber furnish containing 10% the CW had the values with 3.621% and 2.861%, respectively. Consequently, based on the findings obtained from this study, expansion and swelling properties of the MDF panels were negatively affected by compression wood increase.
  • Ayrilmis, University of Istanbul, Faculty of Forestry, Department of Wood Mechanics and Technology, Bahcekoy, TR-34473 Istanbul, Turkey E-mail: nadiray@istanbul.edu.tr (email)
article id 256, category Research article
Juha Laitila. (2008). Harvesting technology and the cost of fuel chips from early thinnings. Silva Fennica vol. 42 no. 2 article id 256. https://doi.org/10.14214/sf.256
Keywords: productivity; harvesting; logging; energy wood; early thinnings; costs
Abstract | View details | Full text in PDF | Author Info
This study compared and analyzed the procurement cost of whole tree chips when using supply chains based on comminution at the roadside landing or at the terminal. It also identified the bottlenecks of the most common logging systems used in Finland. The study was done by using existing and published productivity parameters and models. The procurement cost calculations were made for a stand where the forwarding distance was 200 metres, removal of whole trees was 60 m per hectare and the area of the stand was 2.0 hectares. The average size of the removed whole trees was 30 litres. The direct transport distance from the stand to the terminal or to the end use facility was 40 km while the secondary distance from the terminal to the end use facility was 10 km. A stumpage price for the harvested raw material was not included in this study. According to the study the cost of whole trees chips were 31.9–41.6 euros/m at the plant, or 14.9–19.4 euros/MWh when the moisture content of chips was estimated to be 40%. The two-machine system was found to be the most cost competitive logging system in pre-commercial thinnings thanks to both efficient cutting and, especially, forwarding work. In the manual worker based logging, the costs of felling bunching were the same as the mechanised system, whereas in forwarding the costs were almost double. Using the harwarder system the logging costs were found to be the highest, but in the larger tree volumes and removals the costs were almost equal to the manual worker based logging. The supply chain based on chipping at the roadside landing was more cost efficient compared to the chipping at the terminal system. The lower comminution cost at the terminal was not enough to cover the higher transportation cost of unprocessed material to the terminal, handling cost of chips at the terminal or the delivery cost to the end use facility.
  • Laitila, Finnish Forest Research Institute, Joensuu Research Unit, Joensuu, Finland E-mail: juha.laitila@metla.fi (email)
article id 255, category Research article
Claude Vidal, Adrian Lanz, Erkki Tomppo, Klemens Schadauer, Thomas Gschwantner, Lucio di Cosmo, Nicolas Robert. (2008). Establishing forest inventory reference definitions for forest and growing stock: a study towards common reporting. Silva Fennica vol. 42 no. 2 article id 255. https://doi.org/10.14214/sf.255
Keywords: national forest inventories; reference definitions; growing stock; harmonisation; analytical decomposition
Abstract | View details | Full text in PDF | Author Info
International agreements such as the Kyoto protocol and Convention on Biological Diversity (1992), as well as, criteria and indicator processes require reports on the status of nations’ forests. Any comparison of the current status and trends of forest resources among nations presumes that the nations’ applied definitions and concepts produce comparable estimates of the status of forests. In spite of this, the FAO has already collected global information for 60 years and made noticeable efforts in creating common definitions, but forest related data are still collected using diverse definitions, even regarding basic concepts such as forest and forest area. A simple consequence is that the cross-countries estimates are not comparable. The reasons behind the differences in the definitions are diverse histories, and sometimes different use of forests. In an ideal case, national forest inventories should fulfil both national and international needs. In addition to the FAO’s Forest Resources Assessment process, other efforts are made to assess the status of forests in European countries, e.g. European Forest Information and Communication System (EFICS). EFICS produced reports about forest inventories but does not suggest any common definition or method to convert estimates from one definition to another one. This article presents principles and methods to create commonly acceptable and adoptable definitions for forest inventories. The principles and methods are demonstrated using two examples: the reference definitions of forest and growing stock. The article is based on the work of COST Action E43 (http://www.metla.fi/eu/cost/e43/).
  • Vidal, Inventaire Forestier National, Château des Barres, Nogent-sur-Vernisson, France E-mail: claude.vidal@ifn.fr (email)
  • Lanz, WSL/FNP, Abteilung Landschaftsinventuren, Birmensdorf, Switzerland E-mail: al@nn.ch
  • Tomppo, Finnish Forest Research Institute, Vantaa Research Unit, Vantaa, Finland E-mail: et@nn.fi
  • Schadauer, Bundesamt und Forschungszentrum für Wald, Wien, Austria E-mail: ks@nn.at
  • Gschwantner, Bundesamt und Forschungszentrum für Wald, Wien, Austria E-mail: tg@nn.at
  • di Cosmo, ISAFA, Villazzano, Italy E-mail: ldc@nn.it
  • Robert, Inventaire Forestier National, Ch‰teau des Barres, Nogent-sur-Vernisson, France E-mail: nr@nn.fr
article id 254, category Research article
Johanna Joensuu, Kari Heliövaara, Eino Savolainen. (2008). Risk of bark beetle (Coleoptera, Scolytidae) damage in a spruce forest restoration area in central Finland. Silva Fennica vol. 42 no. 2 article id 254. https://doi.org/10.14214/sf.254
Keywords: biodiversity; dead wood; damage; forest pests; forest restoration
Abstract | View details | Full text in PDF | Author Info
A beetle inventory using window traps was performed to examine the effect of forest restoration by artificial addition of dead wood on the abundance of beetles and to evaluate the risk of bark beetle damage in a forest restoration area. The number of beetle families was slightly increased, but no consistent differences were found in the abundance of families containing saproxylic Coleoptera between the restoration and control plots. The abundance and species number of bark beetles and longhorn beetles were significantly higher on the restoration plots. Ips typographus and Pityogenes chalcographus increased only slightly in abundance. In the regression models produced, the abundance of bark beetles was best explained by the volume of recently dead wood. However, the bark beetle species whose abundance increased most were secondary and the material also suggests an increase in the abundance of bark beetles’ natural enemies. The risk of bark beetle damage in the area is thus considered insignificant.
  • Joensuu, Dept. of Forest Ecology, University of Helsinki, Finland E-mail: johanna.joensuu@metsanhoitajat.fi (email)
  • Heliövaara, Dept. of Forest Ecology, University of Helsinki, Finland E-mail: kh@nn.fi
  • Savolainen, Kuopio Natural History Museum, Kuopio, Finland E-mail: es@nn.fi
article id 253, category Research article
Riitta Väänänen, Mika Nieminen, Martti Vuollekoski, Hannu Nousiainen, Tapani Sallantaus, Eeva-Stiina Tuittila, Hannu Ilvesniemi. (2008). Retention of phosphorus in peatland buffer zones at six forested catchments in southern Finland. Silva Fennica vol. 42 no. 2 article id 253. https://doi.org/10.14214/sf.253
Keywords: peatland; buffer zone area; phosphorus; retention
Abstract | View details | Full text in PDF | Author Info
Our current knowledge of the P retention efficiency of peatland buffer zone areas used to reduce sediment and nutrient leaching from forestry areas is insufficient. Especially the role of P sorption by soil in buffer zones needs closer examination as there is considerable variation in the efficiency of P retention. Six sites in southern Finland were chosen for the study. The buffer zone areas varied between 0.1–4.9% of the catchment area. A total of 10 kg of solute PO4–P was added to the inflow of the buffer zone areas and the concentrations of PO4–P in inflow and outflow were measured for 2–4 years. P retention characteristics of the surface peat were determined with sorption-desorption isotherms before and after PO4–P addition and the effective buffer zone area over which the added P was spread was determined from soil water samples. P retention in the two largest buffer zone areas was complete (100% retention), and the third largest buffer retained 94%. Retention in the three smallest buffer zones was 24%, 95% and 95% of the added P. As a result of P addition reduction in peat P retention capacity was detected in three out of four cases. The effective buffer zone area varied from 67% to 100% of the total buffer zone area. Factors contributing to efficient P retention were large buffer size and low hydrological load whereas high hydrological load combined with the formation of preferential flow paths, especially during early spring or late autumn was disadvantageous. High P retention capacity in peat contributed to the sustainability of P retention. The study showed that even relatively small buffer zone areas are able to efficiently reduce P load.
  • Väänänen, Department of Forest Ecology, University of Helsinki, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: riitta.vaananen@helsinki.fi (email)
  • Nieminen, Finnish Forest Research Institute, Vantaa Research Unit, Finland E-mail: mn@nn.fi
  • Vuollekoski, Finnish Forest Research Institute, Vantaa Research Unit, Finland E-mail: mv@nn.fi
  • Nousiainen, Finnish Forest Research Institute, Vantaa Research Unit, Finland E-mail: hn@nn.fi
  • Sallantaus, Finnish Environment Institute, Nature Division, Helsinki, Finland E-mail: ts@nn.fi
  • Tuittila, Department of Forest Ecology, University of Helsinki, Finland E-mail: est@nn.fi
  • Ilvesniemi, Finnish Forest Research Institute, Vantaa Research Unit, Finland E-mail: hi@nn.fi
article id 252, category Research article
Tuomo Kalliokoski, Pekka Nygren, Risto Sievänen. (2008). Coarse root architecture of three boreal tree species growing in mixed stands. Silva Fennica vol. 42 no. 2 article id 252. https://doi.org/10.14214/sf.252
Keywords: Pinus sylvestris; Betula pendula; Picea abies; site type; branching pattern; influence area; developmental stage
Abstract | View details | Full text in PDF | Author Info
Root system architecture determines many of the vital functions of a tree, e.g. stability of anchorage and resource uptake. The shoot:root ratio is determined through the allocation of resources. Studies on below-ground architectural elements in boreal mixed forests are relatively scarce despite the fact that knowledge on below-ground interactions and allocation changes in relation to stand developmental stage and soil fertility is needed both in ecological and silvicultural research. In this study, sixty tree root systems of three different tree species, Betula pendula, Picea abies and Pinus sylvestris, were excavated in five mixed forest stands in order to quantify differences between the species and sites in terms of rooting behaviour. Root architecture differed greatly between the species, implying different solutions for the functions of root systems. Half of the P. sylvestris had developed a taproot as a response to anchorage needs, while P. abies correspondingly had pronounced secondary growth of proximal roots. Betula pendula had the most extensive root system, illustrating the greater demand of deciduous trees for water. Betula pendula was also the most sensitive to soil fertility: it favoured exploration on the poorest site, as illustrated by the high total root length, whereas on the most fertile site its strategy was to efficiently exploit soil resources through increased branching intensity. The results obtained in this study provide basic knowledge on the architectural characteristics of boreal tree root systems for use by forestry professionals and modellers.
  • Kalliokoski, The Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: tuomo.kalliokoski@metla.fi (email)
  • Nygren, Department of Forest Ecology, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: pn@nn.fi
  • Sievänen, The Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: rs@nn.fi
article id 251, category Research article
Lina R. Steinke, Andrea C. Premoli, Cintia P. Souto, Mikael Hedrén. (2008). Adaptive and neutral variation of the resprouter Nothofagus antarctica growing in distinct habitats in north-western Patagonia. Silva Fennica vol. 42 no. 2 article id 251. https://doi.org/10.14214/sf.251
Keywords: isozymes; morphological variation; leaf traits; South America; temperate forest
Abstract | View details | Full text in PDF | Author Info
N. antarctica occurs in the widest range of habitat types among all South American Notho–fagus. The aim of this study is to investigate adaptive responses by variation in morphological (tree form and leaf characters), and environmental traits (soils) of the polymorphic N. ant–arctica. Also we analyze the effect of genetic drift and limited gene flow in such predominantly apomict by means of neutral variation (isozymes). We studied four potentially different morphological variants each associated with a separate habitat 1) an arboreal variant growing in optimal environments; 2) a sparsely branched variant of temporarily flooded basins or flats; 3) a dwarf variant growing at high elevation, and 4) a shrub-like variant inhabiting matorral environments. The study was restricted latitudinally to Nahuel Huapi National Park, Argentina. For each habitat type we investigated two sites. Nothofagus antarctica shows locally occurring phenotypes. The forest and the high elevation variants were morphologically distinct from the matorral and the basin types. The latter were undistinguishable except for more profuse branching in the matorral type as a result of sprouting due to recent fires. Isozyme evidence indicates a great deal of within-population genetic diversity which is maintained by outcrossing and significant among-site divergence (FST = 18%) that reflects limited gene flow.The apparent high phenotypic and genetic variability in N. antarctica is due to both plasticity and genotypic effects as a result of stable population structure and long periods of isolation which may be reinforced by selection at diverse biotopes.
  • Steinke, Plant Ecology and Systematics, Lund University, Ecology Department, 223 62 Lund, Sweden E-mail: lks@nn.se
  • Premoli, Universidad Nacional del Comahue, Laboratorio Ecotono – CRUB, Quintral 1250, 8400 Bariloche, Argentina E-mail: apremoli@crub.uncoma.edu.ar (email)
  • Souto, Universidad Nacional del Comahue, Laboratorio Ecotono – CRUB, Quintral 1250, 8400 Bariloche, Argentina E-mail: cps@nn.ar
  • Hedrén, Plant Ecology and Systematics, Lund University, Ecology Department, 223 62 Lund, Sweden E-mail: mh@nn.se
article id 250, category Research article
Saara Lilja-Rothsten, Michelle de Chantal, Chris Peterson, Timo Kuuluvainen, Ilkka Vanha-Majamaa, Pasi Puttonen. (2008). Microsites before and after restoration in managed Picea abies stands in southern Finland: effects of fire and partial cutting with dead wood creation. Silva Fennica vol. 42 no. 2 article id 250. https://doi.org/10.14214/sf.250
Keywords: biodiversity; boreal forest; regeneration; coarse woody debris; disturbance dynamics; CWD; managed forest
Abstract | View details | Full text in PDF | Author Info
Different types of microsites, e.g. CWD (coarse woody debris), mounds, and uprooting pits, are important for tree regeneration and biodiversity. However, microsite diversity is greatly reduced in managed stands. We studied how restoration treatments changed microsite distribution in mature managed Picea abies stands. Four cutting treatments were used: uncut, low-CWD (5 m3 ha–1 of down retention trees, DRT, and 50 m3 ha–1 of standing retention trees), intermediate-CWD (as previous but leaving 30 m3 ha–1 of DRT), and high-CWD (as previous but with 60 m3 ha–1 of DRT). Timber harvested from stands ranged from 108–168 m3 ha–1. Half of the stands were burned, and half remained unburned. Sampling was stratified into upland and paludified biotopes within each stand. The pre-treatment microsite distributions were dominated by level ground in both biotopes; mounds and microsites on or next to CWD or a stump were slightly more abundant in the paludified than in the upland biotopes. Microsites were more diverse after cutting, with and without fire. The cutting treatment increased the relative abundances of microsites on or next to CWD. Fire consumed small diameter dead wood and flattened mounds. Microsites were more diverse in paludified than in upland biotopes. The results demonstrate that microsite diversity can rapidly be restored to structurally impoverished managed Picea stands despite a large portion of wood volume being harvested.
  • Lilja-Rothsten, University of Helsinki, Dept. of Forest Ecology, Finland E-mail: saara.lilja@helsinki.fi (email)
  • Chantal, University of Helsinki, Dept. of Forest Ecology, Finland E-mail: mdc@nn.fi
  • Peterson, Dept. of Plant Biology, University of Georgia, Athens, GA, USA E-mail: cp@nn.us
  • Kuuluvainen, University of Helsinki, Dept. of Forest Ecology, Finland E-mail: tk@nn.fi
  • Vanha-Majamaa, The Finnish Forest Research Institute, Vantaa Unit, Finland E-mail: ivm@nn.fi
  • Puttonen, The Finnish Forest Research Institute, Vantaa Unit, Finland E-mail: pp@nn.fi
article id 267, category Research article
Manfred Gronalt, Peter Rauch. (2008). Vendor managed inventory in wood processing industries – a case study. Silva Fennica vol. 42 no. 1 article id 267. https://doi.org/10.14214/sf.267
Keywords: building materials; service level; solid structure timber; SST; wood construction
Abstract | View details | Full text in PDF | Author Info
Solid structure timber (SST) is an important building material in the wood construction business, in which its production volume is largely related to that respective business. Due to the large variability in the demand and seasonal factors, SST producers’ inventories are likely to be simultaneously overstocked for one type of timber and out of stock of another. An inventory policy that ensures a high service level and relatively low stocks is required. In the present paper, we propose the vendor managed inventory (VMI) approach for controlling the stock of deals that are produced at a sawmill and delivered as raw material for SST-production. We evaluate two VMI implementations against the actual inventory management for three different market scenarios. Furthermore, we layout the necessities for reconfiguring the business processes, and subsequently set up an organisational framework within VMI, which is indeed applicable in this segment of the woodworking industry. In our application background, VMI as an inventory control system is able to reduce the overall raw material stock by more than 37% by simultaneously increasing the SST service level.
  • Gronalt, BOKU – University of Natural Resources and Applied Life Sciences, Feistmantelstr. 4, 1180 Vienna, Austria E-mail: mg@nn.at (email)
  • Rauch, BOKU – University of Natural Resources and Applied Life Sciences, Feistmantelstr. 4, 1180 Vienna, Austria E-mail: peter.rauch@boku.ac.at
article id 266, category Research article
Mikko Havimo, Juha Rikala, Jari Sirviö, Marketta Sipi. (2008). Distributions of tracheid cross-sectional dimensions in different parts of Norway spruce stems. Silva Fennica vol. 42 no. 1 article id 266. https://doi.org/10.14214/sf.266
Keywords: Norway spruce; cell wall thickness; earlywood; latewood; tracheid dimension; tangential width; radial width
Abstract | View details | Full text in PDF | Author Info
Distributions of three cross-sectional dimensions: radial and tangential tracheid width, and cell wall thickness in different timber assortments of Norway spruce were investigated. Wood samples from a mature stand were measured with SilviScan. In the analysis, virtual trees were constructed from measurement data, and divided into three assortments: whole stem, top pulpwood and sawmill chips. Average values and distributions of the properties were calculated for all assortments, and distributions divided into earlywood and latewood across the whole tree assortment. There was considerable variation within latewood in all three cross-sectional dimensions, but variation in earlywood was slight in radial width and cell wall thickness. In earlywood, tangential tracheid width showed considerable internal variation, and the difference between earlywood and latewood in tangential width was small. Within-assortment variation of all three properties was larger than between assortments. We may conclude that only a moderate difference in pulp properties can be achieved by sorting raw material into sawmill chips and top pulpwood. Pulp fractionation into earlywood and latewood seems to be a more efficient method, since it gives classes with small within-class variation and distinct average properties. However, it should be kept in mind that the results are valid only in mature stands, where growth rate variation and juvenile wood content are small.
  • Havimo, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: mikko.havimo@helsinki.fi (email)
  • Rikala, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: jr@nn.fi
  • Sirviö, KCL, P.O. Box 70, FI-02151 Espoo, Finland E-mail: js@nn.fi
  • Sipi, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ms@nn.fi
article id 265, category Research article
Emil Cienciala, Erkki Tomppo, Arnor Snorrason, Mark Broadmeadow, Antoine Colin, Karsten Dunger, Zuzana Exnerova, Bruno Lasserre, Hans Petersson, Tibor Priwitzer, Gerardo Sanchez, Göran Ståhl. (2008). Preparing emission reporting from forests: use of National Forest Inventories in European countries. Silva Fennica vol. 42 no. 1 article id 265. https://doi.org/10.14214/sf.265
Keywords: forest carbon pools; greenhouse gas inventory; Kyoto Protocol; UNFCCC
Abstract | View details | Full text in PDF | Author Info
We examine the current status of greenhouse gas inventories of the sector Land Use, Land-Use Change and Forestry (LULUCF), in European countries, with specific focus on the utilization of National Forest Inventory (NFI) programs. LULUCF inventory is an integral part of the reporting obligations under the United Nations Framework Convention on Climate Change (UNFCCC) and its Kyoto Protocol. The analysis is based on two questionnaires prepared by the COST Action E43 “Harmonisation of National Forest Inventories in Europe”, which were answered by greenhouse gas reporting experts in European countries. The following major conclusions can be drawn from the analysis: 1) definitions used to obtain carbon pool change estimates vary widely among countries and are not directly comparable 2) NFIs play a key role for LULUCF greenhouse gas estimation and reporting under UNFCCC, and provide the fundamental data needed for the estimation of carbon stock changes covering not only living biomass, but increasingly also deadwood, litter and soil compartments. The study highlights the effects of adopting different definitions for two major reporting processes, namely UNFCCC and FAO, and exemplifies the effect of different tree diameter thresholds on carbon stock change estimates for Finland. The results demonstrate that more effort is needed to harmonize forest inventory estimates for the purpose of making the estimates of forest carbon pool changes comparable. This effort should lead to a better utilization of the data from the European NFI programs and improve the European greenhouse gas reporting.
  • Cienciala, Institute of Forest Ecosystem Research (IFER), Areal 1. Jilovske a.s. 1544, 254 01 Jilove u Prahy, Czech Republic E-mail: emil.cienciala@ifer.cz (email)
  • Tomppo, Metla, Finnish Forest Research Institute, Finland E-mail: et@nn.fi
  • Snorrason, Icelandic Forest Research, Iceland E-mail: as@nn.is
  • Broadmeadow, Forestry Commission, Forest Research Alice Holt Logdge, United Kingdom E-mail: mb@nn.uk
  • Colin, French National Forest Inventory, France E-mail: ac@nn.fr
  • Dunger, Federal Research Centre for Forestry and Forest Products, Institute of Forest Ecology and Forest Assessment, Germany E-mail: kd@nn.de
  • Exnerova, Institute of Forest Ecosystem Research, Czech Republic E-mail: ze@nn.cz
  • Lasserre, Department of Environment and Territory Sciences and Technologies, University of Molise, Italy E-mail: bl@nn.it
  • Petersson, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Sweden E-mail: hp@nn.se
  • Priwitzer, National Forest Centre, Forest Research Institute. Slovak Republic E-mail: tb@nn.sk
  • Sanchez, Forest Health Unit, General Directorate for Biodiversity, Environmental Ministry, Spain E-mail: gs@nn.es
  • Ståhl, Swedish University of Agricultural Sciences, Department of Forest Resource Management, Sweden E-mail: gs@nn.se
article id 264, category Research article
Yrjö Nuutinen, Kari Väätäinen, Jaakko Heinonen, Antti Asikainen, Dominik Röser. (2008). The accuracy of manually recorded time study data for harvester operation shown via simulator screen. Silva Fennica vol. 42 no. 1 article id 264. https://doi.org/10.14214/sf.264
Keywords: accuracy of timing; mixed effects models; time studies
Abstract | View details | Full text in PDF | Author Info
The aim of the study was to investigate the effect of work experience on the accuracy and variation of observers recording the operation time of a harvester. A simulated thinning operation using a harvester, shown as video via a television screen in laboratory conditions, was observed by 20 inexperienced students and 10 experienced work study researchers. All the observers timed the different work elements of the harvester work with special fieldwork timers. The duration of different work elements measured by the human observers were compared to the corresponding recordings by the harvester’s automated data collector. Although the inexperienced students made more measurement mistakes than the experienced researchers, the differences in measurement error averages were not statistically significant between the groups. However, the variances of tree specific errors were significantly higher in the measurements done by the students. As inexperienced recorders, the students were not able to properly record short work elements, which lasted a maximum of 4 seconds. Due to systematic measurement errors, there was a large variation in the timing structures of the work elements among all observers. Observers’ skills and experience seems to affect measurement accuracy and thus the derived results, especially in intensive time studies. Therefore, the recorder should receive detailed training and practical experience in timing of different work elements of forest operations. In the future, with the use of automated data collectors time studies with large, detailed and accurate data will be implemented. However, due to the varying timing conditions in the forest, manual data collection is still required because of its greater flexibility.
  • Nuutinen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: yrjo.nuutinen@metla.fi (email)
  • Väätäinen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: kv@nn.fi
  • Heinonen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jh@nn.fi
  • Asikainen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: aa@nn.fi
  • Röser, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: dr@nn.fi
article id 263, category Research article
Aksel Granhus, Dag Fjeld. (2008). Time consumption of planting after partial harvests. Silva Fennica vol. 42 no. 1 article id 263. https://doi.org/10.14214/sf.263
Keywords: planting; work study; mechanical site preparation; partial cutting; scarification
Abstract | View details | Full text in PDF | Author Info
Partial harvesting combined with underplanting may be a means to reduce the risk of regeneration failure when e.g. unfavourable microclimatic conditions or severe damage by bark-feeding insects may be expected after clear-cutting, and to maintain or establish certain stand structures or tree species mixture. In this study, we performed time studies of manual planting with and without prior site preparation (patch scarification, inverting) in partially harvested stands of Norway spruce (Picea abies (L.) Karst.). The harvest treatments included basal area removals of approx. 35, 45, and 55%, and a patch clear-cut treatment that was assumed to provide the same conditions for planting as conventional clear-cutting. Site preparation had a much larger influence on time consumption plant–1 (main time) than the harvest treatment. The lowest time consumption was found with inverting and the highest without site preparation. The time spent on walking between planting spots increased with decreasing harvest intensity, reflecting a lower density of planted seedlings in the partially harvested stands. A corresponding increase in main time per plant only occurred after site preparation, since the time spent on clearing the planting spot (removal of logging residue and humus) on untreated plots was higher at the higher harvest strengths. The variation in time consumption attributed to the six replicate stands was large and mainly due to the difference among stands planted by different workers.
  • Granhus, Norwegian University of Life Sciences, Dept. of Ecology and Natural Resource Management (INA), P.O.Box 5003, NO-1432 Ås, Norway E-mail: aksel.granhus@umb.no (email)
  • Fjeld, Swedish University of Agricultural Sciences, Dept. of Forest Resource Management, SE-901 83 Umeå, Sweden E-mail: df@nn.se
article id 262, category Research article
Julian C. Fox, Huiquan Bi, Peter K. Ades. (2008). Modelling spatial dependence in an irregular natural forest. Silva Fennica vol. 42 no. 1 article id 262. https://doi.org/10.14214/sf.262
Keywords: correlogram; Eucalypt; growth modelling; moving average autoregression; Moran’s I; spatial autocorrelation
Abstract | View details | Full text in PDF | Author Info
The spatial dependence present in a natural stand of Eucalyptus pilularis (Smith) dominated mixed species forest was characterised and modelled. Two wildfires imposed a significant spatial dependence on the post disturbance stand. It was hypothesised that spatial variation in the intensity of the wildfires generated the observed structures. The influence of patch formation, micro-site variability and competitive influences were also noted in the residuals of a distance-dependent individual-tree growth model. A methodology capable of modelling these complicated patterns of observed dependence was sought, and candidates included the spatial interaction, direct specification and Papadakis methods. The spatial interaction method with a moving average autoregression was identified as the most appropriate method for explicitly modelling spatial dependence. Both the direct specification and Papadakis methods failed to capture the influence of competition. This study highlights the possibility that stand disturbances such as natural and artificial fires, insect and fungal attacks, and wind and snow damage are capable of imposing powerful spatial dependencies on the post disturbance stand. These dependencies need to be considered if individual tree growth models are to provide valid predictions in disturbed stands.
  • Fox, School of Forest and Ecosystem Science, University of Melbourne, Burnley Campus, 500 Yarra Blvd, Richmond, Victoria 3121 Australia E-mail: jcfox@unimelb.edu.au (email)
  • Bi, Forest Resources Research, New South Wales Department of Primary Industries, PO Box 100, Beecroft, NSW 2119 Australia E-mail: hb@nn.au
  • Ades, School of Forest and Ecosystem Science, University of Melbourne, Burnley Campus, 500 Yarra Blvd, Richmond, Victoria 3121 Australia E-mail: pka@nn.au
article id 261, category Research article
Miina Rautiainen, Matti Mõttus, Pauline Stenberg, Sanna Ervasti. (2008). Crown envelope shape measurements and models. Silva Fennica vol. 42 no. 1 article id 261. https://doi.org/10.14214/sf.261
Keywords: Norway spruce; Scots pine; crown profile; reflectance model; remote sensing
Abstract | View details | Full text in PDF | Author Info
This paper addresses tree crown envelope shape modeling from the perspective of optical passive remote sensing. The aims are 1) to review the specific requirements of crown shape models and ground measurement techniques in optical remote sensing, and 2) to present preliminary results from empirical, parametric crown shape and volume modeling of Scots pine and Norway spruce applicable in Finland. Results indicated that the basic dimensions (maximum radius, its height and crown length) of tree crowns were better predicted for pines, but the profile shape of the upper part of the crowns varied more than in spruce. Pine crowns were also slightly less concave than spruce crowns. No regularities were observed concerning the lower part of the crowns. The asymmetry of crowns increased as a function of tree age for both species, spruce crowns being more asymmetric than pine crowns. A comparison of measured crown volume with several simple geometrical crown shape envelopes showed that using a cone as a crown shape model for Scots pine and Norway spruce underestimates crown volume most severely. Other crown envelope shape models (e.g. ellipsoids) rendered crown volumes closer to the measured volume and did not differ considerably from each other.
  • Rautiainen, Tartu Observatory, 61602 Tõravere, Estonia, and Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: miina.rautiainen@helsinki.fi (email)
  • Mõttus, Tartu Observatory, 61602 Tõravere, Estonia E-mail: mm@nn.ee
  • Stenberg, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ps@nn.fi
  • Ervasti, City of Vantaa, Land Use and Environment / Green Area Unit, Kielotie 13, FI-01300 Vantaa, Finland E-mail: se@nn.fi
article id 260, category Research article
Panagiotis Michopoulos, George Baloutsos, Anastassios Economou. (2008). Nitrogen cycling in a mature mountainous beech forest. Silva Fennica vol. 42 no. 1 article id 260. https://doi.org/10.14214/sf.260
Keywords: bulk deposition; Fagus sylvatica; litterfall; stemflow; stream water; throughfall
Abstract | View details | Full text in PDF | Author Info
The nitrogen cycle in a mature, mountainous European beech (Fagus sylvatica) forest in Greece was examined for two hydrological years, 2001–2002 and 2002–2003. Bulk deposition was 1383 mm in 2001–2002 and 2392 mm in 2002–2003. Winter temperatures were mild in the first year and low in the second year. Despite these climatological differences, the inorganic N inputs to the forest floor, by means of throughfall and stemflow, were quite similar for the two years, i.e. 12.4 and 14.6 kg ha–1 yr–1. Litterfall production was significantly (p < 0.05) higher in the second year but the N amounts in litterfall did not differ. The ratio of N/P in foliar tissue did not change significantly in 2003 compared with ratio values in the last years. The N amounts used for the annual stem and branch increments are rather high preventing, in this way, some nitrogen from being recycled in the near future. The total soil N content to a depth of 80 cm amounted to more than 5000 kg ha–1, and the C/N ratio in the Oh horizon was approximately 15 but the beech forest did not appear susceptible to N leaching. The concentrations of ammonium and nitrate N in stream water did not reach high values reported in the literature, and did not differ significantly in the two hydrological years. The fluxes of inorganic N in throughfall plus stemflow were higher than those in stream water indicating N retention in soil. Another reason for N retention in the ecosystem is probably the large difference between N requirements and uptake indicating N deficiency. Despite the maturity of the beech trees, the low C/N ratio in the Oh horizon and the relatively high N content in soil, the forest can be considered to be neither saturated nor having reached a N saturation transition stage.
  • Michopoulos, Forest Research Institute of Athens, Terma Alkmanos, Athens 115 28, Greece E-mail: mipa@fria.gr (email)
  • Baloutsos, Forest Research Institute of Athens, Terma Alkmanos, Athens 115 28, Greece E-mail: gb@nn.gr
  • Economou, Forest Research Institute of Athens, Terma Alkmanos, Athens 115 28, Greece E-mail: ae@nn.gr
article id 469, category Research article
Henri Vanhanen, Timo O. Veteli, Sonja Päivinen, Seppo Kellomäki, Pekka Niemelä. (2007). Climate change and range shifts in two insect defoliators: gypsy moth and nun moth – a model study. Silva Fennica vol. 41 no. 4 article id 469. https://doi.org/10.14214/sf.469
Keywords: insects; forest pests; CLIMEX; Lymantria monacha; Lymantria dispar; poleward shift; geographical range
Abstract | View details | Full text in PDF | Author Info
Environmental factors influenced by global climate change determine the distribution ranges of organisms. Especially ectothermic animals are expected to shift their distribution ranges northwards in the next hundred years or so. In this study simulations made with CLIMEX-modelling software were used to predict the future distribution ranges of two Central European serious forest pest species: the nun moth (Lymantria monacha L. (Lepidoptera: Lymantriidae)) and the gypsy moth (L. dispar L). The software calculates an ecoclimatic index based on the life cycle requirements of a species and thus represents the probability of a viable population to exist at a certain location. Three different climate warming scenarios were considered: temperature increase of 1.4, 3.6 and 5.8°C. Simulations generated with the current climate conditions corresponded well to the current distributions of the species. The climate warming scenarios shifted the northern boundary of the distribution for both of these species north by c. a. 500–700 km. Also the southern edge of the ranges retracted northwards by 100–900 km. The results of this study are in agreement with the results of empirical studies on other species. Being serious pest species, these species pose a potential threat to silviculture and therefore, have to be considered in the planning of forest management practices.
  • Vanhanen, Faculty of Forestry, University of Joensuu, P.O.B. 111, FI-80101 Joensuu, Finland E-mail: hv@nn.fi
  • Veteli, Faculty of Biosciences, University of Joensuu, P.O.B. 111, FI-80101, Joensuu, Finland E-mail: timo.veteli@joensuu.fi (email)
  • Päivinen, Faculty of Forestry, University of Joensuu, P.O.B. 111, FI-80101 Joensuu, Finland E-mail: sp@nn.fi
  • Kellomäki, Faculty of Forestry, University of Joensuu, P.O.B. 111, FI-80101 Joensuu, Finland E-mail: sk@nn.fi
  • Niemelä, Faculty of Forestry, University of Joensuu, P.O.B. 111, FI-80101 Joensuu, Finland E-mail: pn@nn.fi
article id 279, category Research article
Gerben Janse. (2007). Characteristics and challenges of forest sector communication in the EU. Silva Fennica vol. 41 no. 4 article id 279. https://doi.org/10.14214/sf.279
Keywords: forest policy; European level; forest sector; communication processes
Abstract | View details | Full text in PDF | Author Info
Fragmentation of forest policy at the EU level and image problems of the forest industry have increased forest sector actors’ interest for cooperation on and coordination of communication activities. Although internal communication in the forest sector at the European level is generally well developed both formally and informally, the desired strengthening of communication with other sectors and the public at large is perceived as difficult. In this pre-study 39 interviews were conducted with EU and UNECE/FAO officials, EU level Forest-based Industries and forestry sector federation representatives, and forest scientists working at the European level. The results indicate that EU level forest sector core actors’ ideas on communication differ. Ideas range from strictly image improvement; being more successful at lobbying with other sectors and high-level policy-makers; up to building long-lasting relations and two-way communication processes with other sectors. This makes it difficult to come to coordinated action among forest sector actors. A more active exchange of information on best practices in forest communication between national as well as European level actors, and increased coordination of communication efforts is desired by industry, forest owners as well as policy-makers. However, clear ideas on how to improve, especially inter-sectoral, communication are largely lacking.
  • Janse, European Forest Institute, Joensuu, Finland E-mail: gerben.janse@efi.int (email)
article id 278, category Research article
Katri Luostarinen. (2007). The effect of annual ring orientation and drying method on deformations, casehardening and colour of silver birch (Betula pendula) boards. Silva Fennica vol. 41 no. 4 article id 278. https://doi.org/10.14214/sf.278
Keywords: wood; timber quality; dimensional stability; distortion; drying schedules; kiln drying; L*a*b* coordinates
Abstract | View details | Full text in PDF | Author Info
Deformations of timber, caused mainly by anisotropic shrinkage, can be partially directed by controlling annual ring orientation through different sawing patterns. Ring orientation also affects the movement of water from within the board to its surface, with rapidity of drying having implications for the wood colour. Here sawn silver birch (Betula pendula Roth) timber was classed into two groups according to ring orientation. Two drying methods were used. The final moisture content was lower and the colour lighter in dried boards with radial than with tangential flats, but deformations were larger in radial than in tangential boards. Both drying and ring orientation affected the final moisture content and moisture gradient of the boards. Very small differences in board sizes or shape had an effect on both colour and deformations. The results support the need for accurate sawing and for classing silver birch timber sawn into parquet billets according to ring orientation in order to optimise the drying quality.
  • Luostarinen, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: katri.luostarinen@joensuu.fi (email)
article id 277, category Research article
Laura Koskela, Bikas K. Sinha, Tapio Nummi. (2007). Some aspects of the sampling distribution of the Apportionment Index and related inference. Silva Fennica vol. 41 no. 4 article id 277. https://doi.org/10.14214/sf.277
Keywords: harvesting; bucking; dissimilarity index; multinomial distribution; overlapping coefficient
Abstract | View details | Full text in PDF | Author Info
As customer-oriented production strategies have gained ground in the sawmill industry, proper measurement of the fit between the log demand and log output distributions has become of crucial importance. The prevailing means of measuring the outcome is the so-called Apportionment Index (AI), which is calculated from the relative proportions of the observed and required distributions. Although some statistical properties of the AI have recently been examined and alternative means of measuring the bucking outcome have been suggested, properties of the sampling distribution of the AI have not yet been widely studied. In this article we examine the asymptotic sampling distribution for the AI by assuming a multinomial distribution for the outcome. Our results are based mainly on large-sample normal approximations. Also some studies of the determination of the number of logs needed to obtain a specified level of accuracy of the AI have been carried out.
  • Koskela, University of Tampere, Dept of Mathematics, Statistics and Philosophy, FI-33014 University of Tampere, Finland E-mail: laura.koskela@uta.fi (email)
  • Sinha, Indian Statistical Institute, Kolkata, India E-mail: bks@nn.in
  • Nummi, University of Tampere, Dept of Mathematics, Statistics and Philosophy, FI-33014 University of Tampere, Finland E-mail: tn@nn.fi
article id 276, category Research article
Jianping Zhu, Pete Bettinger, Rongxia Li. (2007). Additional insight into the performance of a new heuristic for solving spatially constrained forest planning problems. Silva Fennica vol. 41 no. 4 article id 276. https://doi.org/10.14214/sf.276
Keywords: forest planning; harvest scheduling; heuristics; raindrop method
Abstract | View details | Full text in PDF | Author Info
  • Zhu, Warnell School of Forestry and Natural Resources, University of Georgia, Athens, GA 30602, USA E-mail: jz@nn.us
  • Bettinger, Warnell School of Forestry and Natural Resources, University of Georgia, Athens, GA 30602, USA E-mail: pbettinger@warnell.uga.edu (email)
  • Li, Warnell School of Forestry and Natural Resources, University of Georgia, Athens, GA 30602, USA E-mail: rl@nn.us
article id 275, category Research article
Lauri Korhonen, Kari T. Korhonen, Pauline Stenberg, Matti Maltamo, Miina Rautiainen. (2007). Local models for forest canopy cover with beta regression. Silva Fennica vol. 41 no. 4 article id 275. https://doi.org/10.14214/sf.275
Keywords: beta regression; canopy cover; forest canopy
Abstract | View details | Full text in PDF | Author Info
Accurate field measurement of the forest canopy cover is too laborious to be used in extensive forest inventories. A possible alternative to the separate canopy cover measurements is to utilize the correlations between the percent canopy cover and easier-to-measure forest variables, especially the basal area. A fairly new analysis technique, the beta regression, is specially designed for modelling percentages. As an extension to the generalized linear models, the beta regression takes into account the distribution of the model residuals, and uses a logistic link function to ensure logical predictions. In this study, the beta regression method was found to perform well in conifer dominated study area located in central Finland. The same model shape, with basal area, tree height and an additional predictor (Scots pine: site fertility, Norway spruce: percentage of hardwoods) as independent variables, produced good results for both pine and spruce dominated sites. The models had reasonably high pseudo R-squared values (pine: 0.91, spruce: 0.87) and low standard errors (pine: 6.3%, spruce: 5.9%) for the fitting data, and also performed well in a cross validation test. The models were also tested on separate test plots located in a different geographical area, where the prediction errors were slightly larger (pine: 8.8%, spruce: 7.4%). In pine plots, the model fit was further improved by introducing additional predictors such as stand age and density. This improved also the performance of the models in the cross validation test, but weakened the results for the external data set. Our results indicated that the beta regression method offers a noteworthy alternative to separate canopy cover measurements, especially if time is limited and the models can be applied in the same region where the modelling data were collected.
  • Korhonen, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: lauri.korhonen@joensuu.fi (email)
  • Korhonen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: ktk@nn.fi
  • Stenberg, Univ. of Helsinki, Dept of Forest Resource Management, P.O. BOX 27, FI-00014 University of Helsinki, Finland E-mail: ps@nn.fi
  • Maltamo, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: mm@nn.fi
  • Rautiainen, Univ. of Helsinki, Dept of Forest Resource Management, P.O. BOX 27, FI-00014 University of Helsinki, Finland E-mail: mr@nn.fi
article id 274, category Research article
Timo Saksa, Jari Miina. (2007). Cleaning methods in planted Scots pine stands in southern Finland: 4-year results on survival, growth and whipping damage of pines. Silva Fennica vol. 41 no. 4 article id 274. https://doi.org/10.14214/sf.274
Keywords: Pinus sylvestris; birch; competition; tending of sapling stand
Abstract | View details | Full text in PDF | Author Info
The effects of four cleaning treatments on the survival, growth of, and whipping damage to Scots pine (Pinus sylvestris L.) main stems were studied in young planted pine stands in southern Finland. Treatments were: no cleaning, point-cleaning of broadleaves (mainly birch, Betula pendula Roth and B. pubescens Ehrh.) within a radius of 1 m from the pine, topping of competing broadleaves, and total cleaning of broadleaves. A randomised complete block design with three replicates was established in five sapling stands: the mean height of the pines was 1.5 m in the three younger stands (6 or 7 years old), and 3 m in the two older stands (9 years old). Measurements taken four growing seasons later showed that in the younger stands, all cleaning treatments significantly (P < 0.05) decreased the mortality and increased the diameter increment of the pines. The height increment of the pines on point-cleaned and topped plots was significantly greater than on totally cleaned plots. In the older stands, the effects of cleaning treatments on the mortality and increment of pines were non-significant. In the younger and older stands, point-cleaning and total cleaning significantly reduced the whipping damage to pines, whereas the topping of competing broadleaves did not. The preliminary results support the use of point-cleaning in planted Scots pine stands when the mean height of the pines is about 1.5 m.
  • Saksa, Finnish Forest Research Institute, Suonenjoki Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@metla.fi (email)
  • Miina, Finnish Forest Research Institute, Joensuu Unit, P.O.Box 68, FI-80101 Joensuu, Finland E-mail: jm@nn.fi
article id 273, category Research article
Stefan Mattson, Urban Bergsten, Tommy Mörling. (2007). Pinus contorta growth in boreal Sweden as affected by combined lupin treatment and soil scarification. Silva Fennica vol. 41 no. 4 article id 273. https://doi.org/10.14214/sf.273
Keywords: biomass; wood density; cultivation; lodgepole pine; lupinus; nitrogen inputs
Abstract | View details | Full text in PDF | Author Info
Effects of combining lupin (Lupinus nootkatensis L.) establishment and soil scarification on stem volume and stem biomass yield of lodgepole pine (Pinus contorta Dougl. var. latifolia Engelm.) were studied on a poor boreal site in Sweden 18 years after plantation. A field randomized block experiment was established with three different scarification techniques (disc trenching, moulding and ploughing) followed by establishment of lupins by either seeds or roots. There were three blocks without and two blocks with lupins. Overall, on average for the three soil scarification techniques, the lupin treatment significantly increased the volume per hectare by 102%.The lupin treatment significantly increased the stem volume per hectare by 236% for mounding and 139% for disc trenching, whereas the 55% increase for ploughing was not significant. The increase in the total stem biomass yield per tree was more pronounced for larger trees; 46% for average trees and 106% for dominant trees. However, there were no significant differences between scarification techniques for the lupin treatment in total stem biomass yield. Over the 18-year period, the increased growth rate following the lupin treatment resulted in a significantly decreased average stem basic wood density (on average 6%) for the sample trees. Because lupin is a nitrogen-fixing plant species, the large increase in tree growth following the lupin treatment was probably an effect of increased amount of nitrogen in the soil. The results indicate that use of lupin is a possible alternative to increase site productivity of lodgepole pine on poor boreal sites.
  • Mattson, Swedish University of Agricultural Sciences, Dept of Forest Ecology and Management, SE-901 83 Umeå, Sweden E-mail: sm@nn.se
  • Bergsten, Swedish University of Agricultural Sciences, Dept of Forest Ecology and Management, SE-901 83 Umeå, Sweden E-mail: ub@nn.se
  • Mörling, Swedish University of Agricultural Sciences, Dept of Forest Ecology and Management, SE-901 83 Umeå, Sweden E-mail: tommy.morling@ssko.slu.se (email)
article id 272, category Research article
Jonas Rönnberg, Mattias Berglund, Ulf Johansson. (2007). Incidence of butt rot at final felling and at first thinning of the subsequent rotation of Norway spruce stands in south-western Sweden. Silva Fennica vol. 41 no. 4 article id 272. https://doi.org/10.14214/sf.272
Keywords: Picea abies; consecutive rotations; disease transmission; Heterobasidion annosum; Heterobasidion parviporum; stump treatment
Abstract | View details | Full text in PDF | Author Info
The incidence of butt rot in Norway spruce stands at final felling and at first thinning of the subsequent rotation was investigated at 20 sites in south-western Sweden. There was a negative correlation between the incidence at first thinning and the basal area of decayed trees at final felling. Using incidence of decay or basal area of decayed trees to predict disease transfer between rotations is difficult and requires the inclusion of several factors, making predictions uncertain. The level of infection found at final felling in this study varied between 7 and 71.8%, indicating that the risk of spore infection transfer to the next rotation was probably quite low. Long-term experiments are required to reveal the effect of stump treatment on disease transmission between rotations.
  • Rönnberg, SLU, Southern Swedish Forest Research Centre, P.O. Box 49, SE-23053 Alnarp, Sweden E-mail: jonas.ronnberg@ess.slu.se (email)
  • Berglund, SLU, Southern Swedish Forest Research Centre, P.O. Box 49, SE-23053 Alnarp, Sweden E-mail: mb@nn.se
  • Johansson, Tönnersjöheden Experimental Forest, P.O. Box 17, SE-31038 Simlångsdalen, Sweden E-mail: uj@nn.se
article id 271, category Research article
Patrick Insinna, Risto Jalkanen, Bernhard Götz. (2007). Climate impact on 100-year foliage chronologies of Scots pine and Ponderosa pine in the northeast lowlands of Brandenburg, Germany. Silva Fennica vol. 41 no. 4 article id 271. https://doi.org/10.14214/sf.271
Keywords: Scots pine; needle trace method; climate impact; foliage parameters; Ponderosa pine
Abstract | View details | Full text in PDF | Author Info
Due to differences in the high-frequency signal and mean sensitivity of needle parameters in Scots pine and Ponderosa pine revealed in previous investigations, variance caused by climate factors at a dry site in the northeast lowlands of Brandenburg was investigated. Although water is the general limiting factor for both tree species, there are evident differences in the climate-driven impact on individual needle parameters. Autumn precipitation of the previous year was equally important for Scots pine and Ponderosa pine, but summer precipitation was more significant for the needle parameters of Scots pine. In contrast to precipitation, temperature seems to have a minor impact on needle parameters. Although January temperatures are significant predictors for both species, intercorrelations between needle parameters and summer temperatures were found only for Ponderosa pine. Striking correlation was also found between sun activity and needle production in Ponderosa pine, but not Scots pine, indicating possible adaptation to solar radiation.
  • Insinna, Office for Environmental Protection Liechtenstein, Climate Change Division, P.O. Box 684, FL-9490 Vaduz, Liechtenstein E-mail: patrick.insinna@aus.llv.li (email)
  • Jalkanen, Rovaniemi Research Unit, Finnish Forest Research Institute, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: rj@nn.fi
  • Götz, Eberswalde University of Applied Sciences, Department of Forestry, Forest-Botanical Gardens, D-16225 Eberswalde, Germany E-mail: bg@nn.de
article id 471, category Research article
Michael Vohland, Johannes Stoffels, Christina Hau, Gebhard Schüler. (2007). Remote sensing techniques for forest parameter assessment: multispectral classification and linear spectral mixture analysis. Silva Fennica vol. 41 no. 3 article id 471. https://doi.org/10.14214/sf.471
Keywords: Picea abies; remote sensing; stand variables; stem number; multispectral classification; Linear Spectral Mixture Analysis
Abstract | View details | Full text in PDF | Author Info
One of the most common applications of remote sensing in forestry is the production of thematic maps, depicting e.g. tree species or stand age, by means of image classification. Nevertheless, the absolute quantification of stand variables is even more essential for forest inventories. For both issues, satellite data are attractive for their large-area and up-to-date mapping capacities. This study followed two steps, and at first a supervised parametric classification was performed for a German test site based on a radiometrically corrected Landsat-5 TM scene. There, eight forest classes were identified with an overall accuracy of 87.5%. In the following, the study focused on the estimation of one key stand variable, the stem number per hectare (SN), which was carried out for a number of Norway spruce stands that had been clearly identified in the multispectral classification. For the estimation of SN, the approach of Linear Spectral Mixture Analysis (LSMA) was found to be clearly more effective than spectral indices. LSMA is based on the premise that measured reflectances can be linearly modelled from a set of so-called endmember spectra. In this study, the endmember sets were held variable to decompose pixel values to abundances of a vegetation, a background (soil, litter, bark) and a shade fraction. Forest structure determines the visible portions of these fractions, and therefore, a multiple regression using them as predictor variables provided the best SN estimates. LSMA allows a pixel-by-pixel quantification of SN for complete satellite images. This opens the view to exploit these data for an improved calibration of large-scale multi-parameter assessment strategies (e.g. statistical modelling or the kNN method for satellite data interpretation).
  • Vohland, University of Trier, Faculty of Geography and Geosciences, Remote Sensing Department, Trier, Germany E-mail: mv@nn.de (email)
  • Stoffels, University of Trier, Faculty of Geography and Geosciences, Remote Sensing Department, Trier, Germany E-mail: js@nn.de
  • Hau, University of Trier, Faculty of Geography and Geosciences, Remote Sensing Department, Trier, Germany E-mail: ch@nn.de
  • Schüler, Research Institution for Forest Ecology and Forestry (FAWF), Department of Forest Growth and Silviculture, Trippstadt, Germany E-mail: gs@nn.de
article id 286, category Research article
Riitta Hänninen, A. Maarit I. Kallio. (2007). Economic impacts on the forest sector of increasing forest biodiversity conservation in Finland. Silva Fennica vol. 41 no. 3 article id 286. https://doi.org/10.14214/sf.286
Keywords: forest biodiversity conservation; roundwood market; forest industry production
Abstract | View details | Full text in PDF | Author Info
In the next coming years, political decisions will be made upon future actions to safeguard forest biodiversity in Southern Finland. We address the economic consequences on the Finnish forest sector of conserving additional 0.5% to 5% of the old growth forest land in Southern Finland. The impacts on supply, demand and prices of wood and forest industry production are analysed employing a partial equilibrium model of the Finnish forest sector. An increase in conservation raises wood prices and thus the production costs of the forest industry. This makes sawnwood production fall, but does not affect paper and paperboard production. The forest owners’ aggregated wood sales income is unaffected or slightly increased, because an increase in stumpage prices offsets the decrease in the harvests. If conservation increases wood imports, negative effects on forest industry become smaller whereas aggregated forest owners’ income may decline depending on the magnitude of import substitution.
  • Hänninen, Finnish Forest Research Institute, Unioninkatu 40 A, FI-00170 Helsinki, Finland E-mail: riitta.hanninen@metla.fi (email)
  • Kallio, Finnish Forest Research Institute, Unioninkatu 40 A, FI-00170 Helsinki, Finland E-mail: maarit.kallio@metla.fi
article id 285, category Research article
Heli Peltola, Antti Kilpeläinen, Kari Sauvala, Tommi Räisänen, Veli-Pekka Ikonen. (2007). Effects of early thinning regime and tree status on the radial growth and wood density of Scots pine. Silva Fennica vol. 41 no. 3 article id 285. https://doi.org/10.14214/sf.285
Keywords: thinning response; ring width; early wood; late wood; intra-ring wood density; X-ray densitometry
Abstract | View details | Full text in PDF | Author Info
  • Peltola, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: heli.peltola@joensuu.fi (email)
  • Kilpeläinen, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: ak@nn.fi
  • Sauvala, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: ks@nn.fi
  • Räisänen, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: tr@nn.fi
  • Ikonen, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: vpi@nn.fi
article id 284, category Research article
Tuomo Nurminen, Jaakko Heinonen. (2007). Characteristics and time consumption of timber trucking in Finland. Silva Fennica vol. 41 no. 3 article id 284. https://doi.org/10.14214/sf.284
Keywords: logistics; cut-to-length operations; time study; work phases; log-normal distribution
Abstract | View details | Full text in PDF | Author Info
Cost efficiency and flexibility have become increasingly important in the logistics of cut-to-length harvesting operations. At the same time, the operating conditions for long-distance transportation have become more demanding and variable. Since the number of log products has increased and the size of harvesting sites has decreased, loads of timber must increasingly be collected from several log decks, increasing the time consumption and costs of the trucking operation. The objectives of this study were to formulate time-consumption models for typical timber transportation activities in Finland and introduce a statistical procedure for examining the variation in time consumption during the trucking phases. The study used a combination of time studies and follow-up studies based on empirical data for 368 loads (a total volume of nearly 18 000 m3) collected from one wood procurement district in central Finland. The model included the following explanatory factors: driving distance, number of log decks, log product and load volume. Since transportation includes several phases and since many factors affect the work performance, significant variation in the total transportation time was observed. This makes planning and cost accounting more difficult. The models developed in this study are a promising initial tool to support route planning and optimization, and cost and profitability calculations for trucking entrepreneurs and the forest industry.
  • Nurminen, University of Joensuu, Faculty of Forest Sciences; Forest Agency Tuomo Nurminen, Joensuuntie 5 B 8, FI-41800 Korpilahti, Finland E-mail: tn@nn.fi (email)
  • Heinonen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jh@nn.fi
article id 283, category Research article
Ilkka Korpela, Tuukka Tuomola, Esko Välimäki. (2007). Mapping forest plots: an efficient method combining photogrammetry and field triangulation. Silva Fennica vol. 41 no. 3 article id 283. https://doi.org/10.14214/sf.283
Keywords: remote sensing; least squares adjustment; intertree positioning; spatial resection; trilateration
Abstract | View details | Full text in PDF | Author Info
Intra stand spatial information is often collected in ecological investigations, when functioning or interactions in the ecosystem are studied. Local relative accuracy is often given priority in such cases. Forest maps with accurate absolute positions in a global coordinate system are needed in remote-sensing applications for validation and calibration purposes. Establishing the absolute position is particularly difficult under a canopy as is creating undistorted coordinate systems for large plots in the forest. We present a method that can be used for the absolute mapping of point features under a canopy that is efficient for large forest plots. In this method, an undistorted network of control points is established in the forest using photogrammetric observations of treetops. These points are used for the positioning of other points, using redundant observations of interpoint distances and azimuths and a least squares adjustment. The method provides decimetre-level accuracy and only one person is required to conduct the work. An estimate of the positioning accuracy of each point is readily available in the field. We present the method, a simulation study that explores the potential of the method and results from an experiment in a mixed boreal stand in southern Finland.
  • Korpela, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ilkka.korpela@helsinki.fi (email)
  • Tuomola, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: tt@nn.fi
  • Välimäki, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ev@nn.fi
article id 282, category Research article
Annika Kangas, Lauri Mehtätalo, Matti Maltamo. (2007). Modelling percentile based basal area weighted diameter distribution. Silva Fennica vol. 41 no. 3 article id 282. https://doi.org/10.14214/sf.282
Keywords: stand structure; diameter distribution; prediction; interpolation
Abstract | View details | Full text in PDF | Author Info
In percentile method, percentiles of the diameter distribution are predicted with a system of models. The continuous empirical diameter distribution function is then obtained by interpolating between the predicted values of percentiles. In Finland, the distribution is typically modelled as a basal-area weighted distribution, which is transformed to a traditional density function for applications. In earlier studies it has been noted that when calculated from the basal-area weighted diameter distribution, the density function is decreasing in most stands, especially for Norway spruce. This behaviour is not supported by the data. In this paper, we investigate the reasons for the unsatisfactory performance and present possible solutions for the problem. Besides the predicted percentiles, the problems are due to implicit assumptions of diameter distribution in the system. The effect of these assumptions can be somewhat lessened with simple ad-hoc methods, like increasing new percentiles to the system. This approach does not, however, utilize all the available information in the estimation, namely the analytical relationships between basal area, stem number and diameter. Accounting for these, gives further possibilities for improving the results. The results show, however, that in order to achieve further improvements, it would be recommendable to make the implicit assumptions more realistic. Furthermore, height variation within stands seems to have an important contribution to the uncertainty of some forest characteristics, especially in the case of sawnwood volume.
  • Kangas, Department of Forest Resources Management, P.O.Box 27, FI-00014 University of Helsinki, Finland E-mail: ak@nn.fi (email)
  • Mehtätalo, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: lm@nn.fi
  • Maltamo, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: mm@nn.fi
article id 281, category Research article
Florence Renou-Wilson, Edward P. Farrell. (2007). The use of foliage and soil information for managing the nutrition of Sitka and Norway spruce on cutaway peatlands. Silva Fennica vol. 41 no. 3 article id 281. https://doi.org/10.14214/sf.281
Keywords: fertilization; Picea abies; foliar analysis; long-term trend; Picea sitchensis; cutaway peatland
Abstract | View details | Full text in PDF | Author Info
This investigation focuses on the development and nutrient status of the first Sitka spruce and Norway spruce stands established on milled cutaway peatlands in Ireland in the late 1980s and early 1990s. Phosphatic fertilization at planting is critical for the establishment of trees on cutaway peatlands but may not be sufficient to see the stands close canopy. Results from this study indicate the likely demand for P and N fertilizer during the rotation of these plantations. During the ten-year-study period (1994–2004), the nutrient status of both Norway and Sitka spruce stands deteriorated with the passage of time. Twenty-seven out of the twenty-eight examined stands became P deficient before 10 years old and half of the plots were N deficient within 13 to 15 years. Sitka spruce stands became N and P deficient earlier than Norway spruce. Regardless of species, tree stands growing on Sphagnum peat entered the critical N deficiency threshold sooner and were all severely deficient by 2004 compared to 22% of the trees growing on Phragmites peat. The effects of aerial re-fertilization were also site specific and although P deficiency was cured, the trees were likely to suffer from nutrient imbalance (N and Cu especially). These results demand a change of standard fertilization practices, which should be related more specifically to peat type and species requirements. Peat type identification and foliar analysis monitoring should become standard management tools while the long-term continuous monitoring of these new forests would be very valuable throughout their first rotation.
  • Renou-Wilson, University College Dublin, Dublin, Ireland E-mail: frw@nn.ie (email)
  • Farrell, University College Dublin, Dublin, Ireland E-mail: epf@nn.ie
article id 280, category Research article
Mika Nieminen, Mikko Moilanen, Sirpa Piirainen. (2007). Phosphorus allocation in surface soil of two drained peatland forests following wood and peat ash application – why effective adsorption on low sorptive soils? Silva Fennica vol. 41 no. 3 article id 280. https://doi.org/10.14214/sf.280
Keywords: forestry; ash fertilization; peat; aluminium; iron; phosphorus adsorption; phosphorus fractionation
Abstract | View details | Full text in PDF | Author Info
Fertilization of drained peatland forests with wood ash and different commercial phosphorus fertilizers (calcium phosphates) generally give similar stand growth responses. However, it has been shown that use of calcium phosphate fertilizers generally lead to highly increased P release to recipient water courses. On the other hand, no corresponding release has been reported after ash fertilization. Soil samples from the surface peats from a nutrient poor and a fertile drained peatland site were collected before and 1–4 years after application of five different types of ash products. The changes over time in the different chemical forms of P in the substrate were studied. The results indicated that the reason for the low liability to leaching of ash fertilizers was that significant amounts of P are adsorbed by Al and Fe during weathering of the ash fertilizers. However, further studies are needed to clarify whether the adsorption of P occurs with the Al and Fe of the ash or the native Al and Fe compounds present in soil before ash fertilization.
  • Nieminen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: mika.nieminen@metla.fi (email)
  • Moilanen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: mm@nn.fi
  • Piirainen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: sp@nn.fi
article id 303, category Research article
Lars Lönnstedt. (2007). Industrial timberland ownership in the USA: arguments based on case studies. Silva Fennica vol. 41 no. 2 article id 303. https://doi.org/10.14214/sf.303
Keywords: forest management; backward vertical integration; pricing; secured supply; owner structure
Abstract | View details | Full text in PDF | Author Info
The forest product companies’ ownership of timberland is decreasing in the United States as in many other countries. In aggregate the forest product industry owned 26.5 million hectares (11.6% of the U.S. timberland) in 2002 compared with 28.5 million hectares in 1987 (FIA 2006). Reasons for this decrease of timberland ownership are several and complex. This article presents four case studies of U.S.-based forest product companies. The vertical integration theory and empirical studies about timberland ownership give a base for the study. Four hypotheses are formulated on the basis of the literature. The results give support to two of them. An important reason for timberland ownership is a wish to secure deliveries. Market conditions are important for the need of owning timberland. Two of the companies did not own timberland, the main reason being more profitable alternative uses of capital. The ownership structure of the company, tradition, and culture are other important explanations for timberland ownership. This study did not show the advantage of timberland ownership for information and coordination.
  • Lönnstedt, SLU, Department of Forest Products, P.O. Box 7060, SE 750 07 Uppsala, Sweden E-mail: lars.lonnstedt@spm.slu.se (email)
article id 302, category Research article
Gökay Nemli, Turgay Akbulut, Emir Zekoviç. (2007). Effects of some sanding factors on the surface roughness of particleboard. Silva Fennica vol. 41 no. 2 article id 302. https://doi.org/10.14214/sf.302
Keywords: particleboard; sanding; surface roughness; power; grit size; feeding speed
Abstract | View details | Full text in PDF | Author Info
Effects of the grit sizes of the sand belt, feeding speed and the feed power of the heads of the sander on the surface roughness of the particleboard panels were investigated. Two surface roughness parameters, average roughness (Ra) and mean peak-to-valley height (Rz), obtained from board surfaces were used in the analysis. Sanding factors were found to have a significant effect on the surface roughness of the particleboard. Better results were obtained with 40 m/min of feeding speed, 40-60-80-120 of grit sizes, and 67 kW of the feed power of the heads of the sander.
  • Nemli, Karadeniz Technical University, Faculty of Forestry, Trabzon-Türkiye E-mail: nemli@ktu.edu.tr (email)
  • Akbulut, Istanbul University, Faculty of Forestry, Istanbul-Türkiye E-mail: ta@nn.tr
  • Zekoviç, Starwood Forest Product Company, Production Manager, Bursa-Türkiye E-mail: ez@nn.tr
article id 301, category Research article
Sanni Raiskila, Minna Pulkkinen, Tapio Laakso, Kurt Fagerstedt, Mia Löija, Riitta Mahlberg, Leena Paajanen, Anne-Christine Ritschkoff, Pekka Saranpää. (2007). FTIR spectroscopic prediction of Klason and acid soluble lignin variation in Norway spruce cutting clones. Silva Fennica vol. 41 no. 2 article id 301. https://doi.org/10.14214/sf.301
Keywords: Norway spruce; FTIR; lignin; PCR; principal component regression
Abstract | View details | Full text in PDF | Author Info
Our purpose was to develop a FTIR spectroscopic method to be used to determine the lignin content in a large number of samples and to apply this method studying variation in sapwood and heartwood lignin content between three fast-growing cutting clones grown in three sites. Models were estimated with 18 samples and tested with 6 samples for which the Klason lignin + acid soluble lignin content had been determined. Altogether 272 candidate models were built with all-subset regressions from the principal components estimated from differently treated transmission spectra of the samples; the spectra were recorded on KBr pellets of sieved and unsieved unextracted wood powder and subjected to four different preprocessings and two different wavenumber selection schemes. The final model showed an adequate fit in the estimation data (R2 = 0.74) as well as a good prediction performance in the test data (R2P = 0.90). This model was based on the wavenumber range of 1850–500 cm–1 of the line-subtraction-normalised spectra recorded from sieved samples. The model was used to predict lignin content in 64 samples of the same material. One of the clones had a slightly lower sapwood lignin content than the two other clones. The fertile growing site with fast growing trees showed slightly higher sapwood lignin content compared with the other two sites. The model was also used to predict the lignin content in the earlywood of 45 individual annual rings. Variation between individual stems and between annual rings was found to be large. No correlation was found between the lignin content and density of earlywood.
  • Raiskila, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: sr@nn.fi
  • Pulkkinen, Department of Forest Ecology, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: mp@nn.fi
  • Laakso, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: tl@nn.fi
  • Fagerstedt, Department of Biological and Environmental Sciences, Plant Biology, P.O. Box 65, FI-00014 University of Helsinki, Finland E-mail: kf@nn.fi
  • Löija, VTT Building and Transport, P.O. Box 1806, FI-02044 VTT, Finland E-mail: ml@nn.fi
  • Mahlberg, VTT Building and Transport, P.O. Box 1806, FI-02044 VTT, Finland E-mail: rm@nn.fi
  • Paajanen, VTT Building and Transport, P.O. Box 1806, FI-02044 VTT, Finland E-mail: lp@nn.fi
  • Ritschkoff, VTT Building and Transport, P.O. Box 1806, FI-02044 VTT, Finland E-mail: acr@nn.fi
  • Saranpää, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: pekka.saranpaa@metla.fi (email)
article id 300, category Research article
Jouni Siipilehto, Sakari Sarkkola, Lauri Mehtätalo. (2007). Comparing regression estimation techniques when predicting diameter distributions of Scots pine on drained peatlands. Silva Fennica vol. 41 no. 2 article id 300. https://doi.org/10.14214/sf.300
Keywords: Pinus sylvestris; drained peatland; dbh distribution; Johnson’s SB function; regression estimation methods
Abstract | View details | Full text in PDF | Author Info
We compared different statistical methods for fitting linear regression models to a longitudinal data of breast height diameter (dbh) distributions of Scots pine dominated stands on drained peatlands. The parameter prediction methods for two parameters of Johnson’s SB distribution, fitted to basal-area dbh distributions, were: 1) a linear model estimated by ordinary least squares (OLS), 2) a multivariate linear model estimated using the seemingly unrelated regression approach (SUR), 3) a linear mixed-effects model with random intercept (MIX), and 4) a multivariate mixed-effects model (MSUR). The aim was to clarify the effect of taking into account the hierarchy of the data, as well as simultaneous estimation of the correlated dependent variables on the model fit and predictions. Instead of the reliability of the predicted parameters, we focused on the reliability of the models in predicting stand conditions. Predicted distributions were validated in terms of bias, RMSE, and error deviation in the generated quantities of the growing stock. The study material consisted of 112 successively measured stands from 12 experimental areas covering the whole of Finland (total of 608 observations). Two independent test data sets were used for model validation. All the advanced regression techniques were superior to OLS, when exactly the same independent stand variables were included. SUR and MSUR were ranked the overall best and second best, respectively. Their ranking was the same in the modeling data, whereas MSUR was superior in the peatland test data and SUR in the mineral soil test data. The ranking of the models was logical, but may not be widely generalized. The SUR and MSUR models were considered to be relevant tools for practical forest management planning purposes over a variety of site types and stand structures.
  • Siipilehto, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: jouni.siipilehto@metla.fi (email)
  • Sarkkola, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: ss@nn.fi
  • Mehtätalo, University of Joensuu, Faculty of Forestry, P.O. Box 111, 80101 Joensuu, Finland E-mail: lm@nn.fi
article id 299, category Research article
Hongcheng Zeng, Timo Pukkala, Heli Peltola, Seppo Kellomäki. (2007). Application of ant colony optimization for the risk management of wind damage in forest planning. Silva Fennica vol. 41 no. 2 article id 299. https://doi.org/10.14214/sf.299
Keywords: harvest scheduling; heuristics; genetic algorithms; simulated annealing; spatial optimization
Abstract | View details | Full text in PDF | Author Info
Ant colony optimization (ACO) is still quite a new technique and seldom used in the field of forest planning compared to other heuristics such as simulated annealing and genetic algorithms. This work was aimed at evaluating the suitability of ACO for optimizing the clear-cut patterns of a forest landscape when aiming at simultaneously minimizing the risk of wind damage and maintaining sustainable and even flow of periodical harvests. For this purpose, the ACO was first revised and the algorithm was coded using the Visual Basic Application of the ArcGIS software. Thereafter, the performance of the modified ACO was demonstrated in a forest located in central Finland using a 30-year planning period. Its performance was compared to simulated annealing and a genetic algorithm. The revised ACO performed logically since the objective function value was improving and the algorithm was converging during the optimization process. The solutions maintained a quite even periodical harvesting timber while minimizing the risk of wind damage. Implementing the solution would result in smooth landscape in terms of stand height after the 30-year planning period. The algorithm is quite sensitive to the parameters controlling pheromone updating and schedule selecting. It is comparable in solution quality to simulated annealing and genetic algorithms.
  • Zeng, University of Joensuu, Faculty of Forest Sciences, P. O. Box 111, FI-80101 Joensuu, Finland E-mail: hongcheng.zeng@joensuu.fi (email)
  • Pukkala, University of Joensuu, Faculty of Forest Sciences, P. O. Box 111, FI-80101 Joensuu, Finland E-mail: tp@nn.fi
  • Peltola, University of Joensuu, Faculty of Forest Sciences, P. O. Box 111, FI-80101 Joensuu, Finland E-mail: hp@nn.fi
  • Kellomäki, University of Joensuu, Faculty of Forest Sciences, P. O. Box 111, FI-80101 Joensuu, Finland E-mail: sk@nn.fi
article id 298, category Research article
Ulfstand Wennström, Urban Bergsten, Jan-Erik Nilsson. (2007). Seedling establishment and growth after direct seeding with Pinus sylvestris: effects of seed type, seed origin, and seeding year. Silva Fennica vol. 41 no. 2 article id 298. https://doi.org/10.14214/sf.298
Keywords: Scots pine; direct seeding; orchard seed; stand seed
Abstract | View details | Full text in PDF | Author Info
The early effects of seed type, seed origin, and seeding year on seedling emergence, survival, and growth after one to four years was quantified and examined. Two experimental series of Scots pine located at 61°N and 64°N and six orchard seed lots and six stand seed lots of adequate geographical origins in each series were used. Both series were replicated at five sites for up to five years. On average, orchard seed lots had 16% and 12% higher seedling emergence, in relation to sown germinable seeds, than stand seed lots in the northern and southern series. The survival from year 1 to year 4 was also higher for orchard seedlings than for stand seedlings; there was a 77% and 72% survival rate in the northern series and a 58% and 49% survival rate in the southern series for orchard and stand seedlings respectively. On average, after four years orchard seedlings were 26% taller in the northern series and 13% taller in the southern series. The gain in height growth for the orchard seeds was positive at all seeding years, at all sites, and at all seedling ages. If survival was calculated to the height of a four-year-old seedling, the survival of orchard seedlings increased by 3% in the northern and 1% in the southern series as the result of the higher growth of orchard seedlings. Using orchard seeds resulted in 6 percent units higher growth gain when the clear cuts were regenerated with direct seeding than with plants using the same seed material. Changes in the ranking of seed lots and seed types at different sites and seeding years for seedling emergence is an effect of external factors such as grazing and foraging that cannot be related directly to the tested factors.
  • Wennström, Skogforsk, Box 3, SE-918 21 Sävar, Sweden E-mail: ulfstand.wennstrom@skogforsk.se (email)
  • Bergsten, SLU, Dept. of Forest Ecology and Management, SE-901 83 Umeå, Sweden & SLU, Vindeln Experimental Forest, SE-922 91 Vindeln, Sweden E-mail: ub@nn.se
  • Nilsson, SLU, Dept. of Forest Genetics and Plant Physiology E-mail: jen@nn.se
article id 297, category Research article
Antti Marjokorpi, Jukka Salo. (2007). Operational standards and guidelines for biodiversity management in tropical and subtropical forest plantations – How widely do they cover an ecological framework? Silva Fennica vol. 41 no. 2 article id 297. https://doi.org/10.14214/sf.297
Keywords: criteria and indicators; afforestation; conservation; sustainable forest management; reforestation
Abstract | View details | Full text in PDF | Author Info
The development of standards and guidelines to secure sustainable forest management at different geographical scales has expanded greatly during the past fifteen years. Most of these efforts, however, have been formulated for natural forests only; those designed specifically for forest plantations are relatively few. The global forest plantation area is expanding rapidly, with obvious positive and negative impacts on biodiversity. We characterize the key concepts of biodiversity in tropical and subtropical forest plantations and present an analysis of how these elements are covered in the eight principal operational standards and guidelines for sustainable plantation forestry. We also examine the applicability of standards and guidelines in plantations established and managed under different initial settings. The results indicate that the standards and guidelines address certain key elements of biodiversity comprehensively, meanwhile others are ignored or receive only slight attention. There is also substantial variation between the sets in their nature (performance- vs. process-based), scope, congruity in concepts and hierarchy, and specificity. The standards and guidelines seldom take into account the varying initial settings for plantation establishment and the consequent variation in critical factors in biodiversity conservation and management. We recommend that standards and guidelines should be developed so as to pay more attention to the type and operating environment of plantations, to cover all key factors of biodiversity, and to consider building closer relationships between the social and ecological aspects of biodiversity.
  • Marjokorpi, Stora Enso, Wood Supply, Talvikkitie 40 C, FI-01300 Vantaa, Finland E-mail: antti.marjokorpi@storaenso.com (email)
  • Salo, Department of Biology, University of Turku, FI-20140 Turku, Finland E-mail: js@nn.fi
article id 296, category Research article
Samuel Roturier, Sofia Bäcklund, Maria Sundén, Urban Bergsten. (2007). Influence of ground substrate on establishment of reindeer lichen after artificial dispersal. Silva Fennica vol. 41 no. 2 article id 296. https://doi.org/10.14214/sf.296
Keywords: mineral soil; bark; Cladina mitis; image analysis; lichen colonization; moss; thallus; twigs
Abstract | View details | Full text in PDF | Author Info
Methods to improve the recovery of reindeer lichen after soil disturbance or overgrazing are being sought for areas where reindeer are herded. The effects of four substrates – mineral soil, moss, twigs and pine bark – on the establishment of lichen fragments after total removal of the vegetation were thus studied in a middle-aged pine stand and a clear-cut, both located in a lichen-rich pine-heath. Cladina mitis fragments of two sizes were manually dispersed in 1 m2 quadrats and their movements from their respective dispersal points were registered after one year. The natural re-establishment of lichens in the quadrats was monitored over three years by using digital pictures. In the forest stand, no significant differences were detected in either the fragment movement or the lichen establishment between the different substrates, but the fragment size had positive effects on both parameters. In the clear-cut, the moss substrate was the most suitable not only for the artificially dispersed lichens to fasten to, but also for the natural settlement of lichens from the surrounding lichen mat. More lichen thalli fastened to the bark and twigs substrates than to the mineral soil, but the settlement of lichens from the surrounding was greater on bare mineral soil substrate. The results indicate that artificial dispersal of lichen thalli on an appropriate substrate could be a successful strategy for promoting lichen recovery.
  • Roturier, SLU, Vindeln Experimental Forests, Svartberget Fältstation, SE-922 91 Vindeln, Sweden E-mail: samuel.roturier@esf.slu.se (email)
  • Bäcklund, E-mail: sb@nn.se
  • Sundén, E-mail: ms@nn.se
  • Bergsten, SLU, Dept of Forest Ecology, SE-901 83 Umeå, Sweden E-mail: ub@nn.se
article id 295, category Research article
Pekka Kaitaniemi, Janne Riihimäki, Julia Koricheva, Harri Vehviläinen. (2007). Experimental evidence for associational resistance against the European pine sawfly in mixed tree stands. Silva Fennica vol. 41 no. 2 article id 295. https://doi.org/10.14214/sf.295
Keywords: stand structure; natural enemies; species diversity; tri-trophic interactions
Abstract | View details | Full text in PDF | Author Info
This study examined whether the saplings of Scots pine (Pinus sylvestris) exhibit associational resistance against the European pine sawfly Neodiprion sertifer (Hymenoptera, Diprionidae) when grown in a mixture with 50% silver birch (Betula pendula). The number of sawflies on pine trees in pure and mixed stands was manipulated at two experimental sites during two years. Survival of larvae and eggs was monitored, and the numbers of presumed sawfly predators were counted. A lower proportion of sawfly larvae and eggs survived on pines grown in the mixture with birch as compared with pure pine stands. Lower survival of sawfly larvae in the mixed stands was associated with the higher abundance of ants in these stands. The numbers of other sawfly predators (e.g. spiders and predatory heteropterans) differed between the study sites and were negatively associated with the presence of ants, which suggests possible interference between these groups. Although sawfly survival was lower on pines in the mixed stands, providing evidence of associational resistance, a related study shows the same trees had a higher number of ant-tended aphid colonies as compared with pines in the pure stands. Therefore, instead of considering resistance against individual herbivore species, it seems more practical to use associational resistance as a trait representing the resistance of larger systems, such as whole tree stands, against the total damage caused by herbivores in general.
  • Kaitaniemi, Hyytiälä Forestry Field Station, University of Helsinki, Hyytiäläntie 124, FI-35500 Korkeakoski, Finland E-mail: pk@nn.fi (email)
  • Riihimäki, Section of Ecology, Department of Biology, University of Turku, FI-20014 Turku, Finland E-mail: jr@nn.fi
  • Koricheva, School of Biological Sciences, Royal Holloway, University of London, Egham, Surrey, TW20 0EX, UK E-mail: jk@nn.uk
  • Vehviläinen, Section of Ecology, Department of Biology, University of Turku, FI-20014 Turku, Finland E-mail: hv@nn.fi
article id 294, category Research article
Ülle Püttsepp, Krista Lõhmus, Andres Koppel. (2007). Decomposition of fine roots and α-cellulose in a short rotation willow (Salix spp.) plantation on abandoned agricultural land. Silva Fennica vol. 41 no. 2 article id 294. https://doi.org/10.14214/sf.294
Keywords: decomposition; α-cellulose; fine roots; acid detergent lignin; short rotation forest; Salix spp.
Abstract | View details | Full text in PDF | Author Info
Decomposition of fine roots (<1 mm in diameter) of the clones of Salix viminalis, S. dasyclados and α-cellulose sheets (50 x 10 x 1 mm) was studied in a 6-years old Salix spp. plantation established on abandoned agricultural land in Estonia. The substrates were incubated in litterbags (mesh size 0.14 mm) in 5–10 cm topsoil, in non-fertilised plots for one year. Changes in the ash-free weight of the fine roots were best described by negative exponential models (S. viminalis R2 = 0.98, S. dasyclados R2 = 0.96), and by a linear model for α-cellulose (R2 = 0.63). The sheets of α-cellulose decomposed roughly twice as rapidly as the fine roots (S. viminalis k = 0.325, S. dasyclados k = 0.165). The remaining (of the initial) ash-free weights of the fine roots were 73.3 ± 0.8% (mean ± SE) and 85.8 ± 2.2% respectively, and of the α-cellulose 35.9 ± 8.5%, in the end of the one year of decomposition. The amount of acid detergent (AD) lignin in the fine-roots of S. viminalis increased significantly and did not change in S. dasyclados, suggesting higher activity of microbial decomposers in the first substrate. Of the studied quality parameters, the AD lignin was the major factor determining the different rate of decomposition of the fine roots of S. viminalis and S. dasyclados. Nitrogen was recycled in the fine root sub-system in both Salix species. This knowledge can be applied in the management of Salix plantations, aimed at bioenergy production.
  • Püttsepp, Department of Ecology, Swedish University of Agricultural Sciences, P.O. Box 7072, SE-75007 Uppsala, Sweden; Estonian University of Life Sciences, Kreuzwaldi 64, Tartu 51014, Estonia E-mail: ulle.puttsepp@ekol.slu.se (email)
  • Lõhmus, Institute of Geography, University of Tartu, Vanemuise 46, Tartu 51014, Estonia E-mail: kl@nn.ee
  • Koppel, Estonian University of Life Sciences, Kreuzwaldi 64, Tartu 51014, Estonia E-mail: ak@nn.ee
article id 293, category Research article
Conor O'Reilly, Norberto De Atrip. (2007). Seed moisture content during chilling and heat stress effects after chilling on the germination of common alder and downy birch seeds. Silva Fennica vol. 41 no. 2 article id 293. https://doi.org/10.14214/sf.293
Keywords: Betula pubescens; Alnus glutinosa; germination temperature; stress; moist chilling
Abstract | View details | Full text in PDF | Author Info
The effects of seed moisture content (MC) and heat treatment on the germination response of common alder (Alnus glutinosa) and downy birch (Betula pubescens) seeds were examined. Seeds of each species were adjusted to MC of 7% to 50% MC, then chilled for up to 36 weeks, after which they were allowed to germinate at 15°C with 8 hours lighting per day or 20 (dark)/ 30°C (light). Seed lot effects were evident, but treatment effects were consistent in each lot and species. The response to moist chilling treatments was larger at 15°C than at 20/30°C. Chilling had no effect on germination unless seed MC was >15%, but it was low also at 20% MC. The highest germination was achieved following 24–36 weeks chilling at the optimum or target MC (TMC) levels of about 30% in alder and 35% in birch. In a separate experiment, seeds were fully imbibed (FI) (~50% MC; standard method used in operational practice) or adjusted to TMC levels, after which some seeds of each treatment group were chilled to release dormancy. Following this, the seeds were dried back to TMC levels and then subjected to 60°C for up to 4 hours after which they were allowed germinate under the same conditions described above. Heat treatment damaged the prechilled FI seeds, but no damage occurred to the non-chilled seeds. However, heat stress stimulated germination in the non-chilled FI seeds of both species and the TMC seeds of alder.
  • O'Reilly, UCD School of Biology and Environmental Science, UCD College of Life Sciences, University College Dublin, Belfield, Dublin 4, Ireland E-mail: conor.oreilly@ucd.ie (email)
  • De Atrip, UCD School of Biology and Environmental Science, UCD College of Life Sciences, University College Dublin, Belfield, Dublin 4, Ireland E-mail: nda@nn.ie
article id 292, category Research article
Sanna Susiluoto, Frank Berninger. (2007). Interactions between morphological and physiological drought responses in Eucalyptus microtheca. Silva Fennica vol. 41 no. 2 article id 292. https://doi.org/10.14214/sf.292
Keywords: photosynthesis; chlorophyll fluorescence; root/shoot ratio; RuBP; xylem permeability
Abstract | View details | Full text in PDF | Author Info
We studied the response of Eucalyptus microtheca to drought in a greenhouse experiment. As a result of the drought the growth of the seedlings decreased and allocation patterns changed so that allocation to the roots increased. However, changes in photosynthesis and stomatal conductance under drought were rather modest. We showed, using chlorophyll fluorescence and measurements of photosynthesis under high CO2 that the biochemical capacity of photosynthesis increased under drought. The results suggest that changes in root/shoot ratio are the primary reactions that initiate a series of compensatory reactions that mitigate the effects of drought in Eucalyptus microtheca.
  • Susiluoto, University of Helsinki, Department of Forest Ecology, PL 27, 00014 University of Helsinki, Finland E-mail: sannamaija.susiluoto@helsinki.fi (email)
  • Berninger, Departement des Sciences biologiques, Université du Québec à Montréal, CP 8888 Succ Centre Ville, Montreal, Canada E-mail: fb@nn.ca
article id 291, category Research article
Yildiray Lise, Zeki Kaya, Fikret Isik, Rumi Sabuncu, Irfan Kandemir, Sertaç Önde. (2007). The impact of over-exploitation on the genetic structure of Turkish red pine (Pinus brutia Ten.) populations determined by RAPD markers. Silva Fennica vol. 41 no. 2 article id 291. https://doi.org/10.14214/sf.291
Keywords: Pinus brutia Ten.; RAPD-PCR; human over-exploitation; genetic structure; inbreeding
Abstract | View details | Full text in PDF | Author Info
To determine the possible impact of over-exploitation on the genetic structure of Turkish red pine (Pinus brutia Ten.) populations, three natural and three over-exploited (human degraded) populations of the species in the Mediterranean region of Turkey were investigated with Randomly Amplified Polymorphic DNA (RAPD). With the 80 RAPD primers tested, 12 of them yielded 137 polymorphic RAPD fragments. Four of the studied populations maintained unique fragments. The mean proportion of polymorphic fragments for all populations ranged from 89.8 to 98.9% and there were no significant differences between natural (94.8%) vs. over-exploited populations (92.7%). The estimated heterozygosity values suggested that Turkish red pine maintains high levels of genetic diversity (range 0.24–0.28) though studied populations and grouped ones as natural (He = 0.28) vs. over-exploited (0.27) did not differ significantly. The mean FST value indicated that the large portion of the total genetic diversity was within populations (93%), but this value was lower in the natural populations (92%) than in the over-exploited ones (94%). In over-exploited populations, excess of homozygosity was observed (about 6% higher) as compared to natural populations, indicating impacts of inbreeding in P. brutia.
  • Lise, Department of Biological Sciences, Middle East Technical University, 06531, Ankara, Turkey E-mail: yl@nn.tr
  • Kaya, Department of Biological Sciences, Middle East Technical University, 06531, Ankara, Turkey E-mail: kayaz@metu.edu.tr (email)
  • Isik, North Carolina State University, Raleigh, USA E-mail: fi@nn.us
  • Sabuncu, Southwest Anatolia Forest Research Institute, Antalya, Turkey E-mail: rs@nn.tr
  • Kandemir, Department of Biology, Zonguldak Karaelmas University, 67100, Zonguldak, Turkey E-mail: ik@nn.tr
  • Önde, Department of Biological Sciences, Middle East Technical University, 06531, Ankara, Turkey E-mail: so@nn.tr
article id 474, category Research article
Tero Heinonen, Mikko Kurttila, Timo Pukkala. (2007). Possibilities to aggregate raster cells through spatial optimization in forest planning. Silva Fennica vol. 41 no. 1 article id 474. https://doi.org/10.14214/sf.474
Keywords: threshold accepting; spatial optimisation; landscape metrics; old forest
Abstract | View details | Full text in PDF | Author Info
  • Heinonen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: tero.heinonen@joensuu.fi (email)
  • Kurttila, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: mk@nn.fi
  • Pukkala, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: tp@nn.fi
article id 473, category Research article
Ulla Mattila, Tuula Nuutinen. (2007). Assessing the incidence of butt rot in Norway spruce in southern Finland. Silva Fennica vol. 41 no. 1 article id 473. https://doi.org/10.14214/sf.473
Keywords: Norway spruce; logistic regression; Heterobasidion; Butt rot; quality loss
Abstract | View details | Full text in PDF | Author Info
The aim of this study was to analyze the occurrence of butt rot damage to Norway spruce in different parts of southern Finland and to quantify the associated loss of quality. The data used in the study are from the 9th National Forest Inventory and consist of 5998 sample plots and 8007 spruce sample trees of saw-timber size. To predict the probability of damage to stands and trees, logistic regression models were constructed. Separate models were made for the whole study area, for the area where the general risk of Heterobasidion root and butt rot damage is high and for the area where the damage frequency is relatively low. In the high-risk area, the probability of damage decreased with increasing elevation and increased with increasing temperature sum. In addition, damage was more common on fertile sites and less common on peatlands; and thick peat layer decreased the risk of damage. The probability of damage was also higher in stands where special or selective cuttings had been carried out. In the sample tree data, the probability of damage increased slightly with increasing diameter and age of the tree. In the low-risk areas, elevation was the only variable that explained the probability of damage to a spruce tree. Site fertility and previous cuttings (more than ten years ago) explained the probability of damage to stands only weakly. For spruce damaged by butt rot, the saw-timber volume was reduced, on average, by 60% both in the high-risk area and in the low-risk area.
  • Mattila, The Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: um@nn.fi (email)
  • Nuutinen, The Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: tn@nn.fi
article id 309, category Research article
Sandhya Samarasinghe, Don Kulasiri, Tristan Jamieson. (2007). Neural networks for predicting fracture toughness of individual wood samples. Silva Fennica vol. 41 no. 1 article id 309. https://doi.org/10.14214/sf.309
Keywords: Pinus radiata; New Zealand; video imaging; strain energy release rate; Neural Networks; fracture toughness
Abstract | View details | Full text in PDF | Author Info
Strain energy release rate (GIc) of Pinus radiata in the TL opening mode was determined using the compliance crack length relationship. A total of 123 specimens consisting of four sizes of specimen with each size having four different crack lengths were tested. For each specimen, grain and ring angles, density and moisture content were measured. Video imaging, was used to measure crack length during propagation. Since cracks extended in stages, full compliance-crack length relationship was developed for each specimen based on their initial and subsequent crack lengths. No significant differences in GIc, between initial and subsequent crack lengths were found for the smaller specimens by paired sample t-tests, but differences were significant for the largest specimen size. The Average fracture toughness was calculated from GIc and it was 215 kPa.m0.5. Three artificial neural networks were developed to predict the: 1) force required to propagate a crack, 2) crack extension, and 3) fracture toughness of an individual specimen. Each was successful, producing respective R2 of 0.870, 0.865, and 0.621 on validation data. A sensitivity analysis of the networks revealed that the crack length was the most influential with 21% contribution followed by grain angle with 14% contribution for predicting the applied force. This was followed by volume and physical properties. For predicting the crack extension, density had the greatest contribution (20%) followed by previous crack length and force contributing 16% equally. Fracture toughness was dominated by the dimensional parameters of the specimen contributing (42%) followed by anisotropy and physical properties.
  • Samarasinghe, Centre for Advanced Computational Solutions (C-fACS), Lincoln University, New Zealand E-mail: ss@nn.nz (email)
  • Kulasiri, Centre for Advanced Computational Solutions (C-fACS), Lincoln University, New Zealand E-mail: dk@nn.nz
  • Jamieson, Centre for Advanced Computational Solutions (C-fACS), Lincoln University, New Zealand E-mail: tj@nn.nz
article id 308, category Research article
Ville Hallikainen, Mikko Hyppönen, Juha Hyvönen, Juhani Niemelä. (2007). Establishment and height development of harvested and naturally regenerated Scots pine near the timberline in North-East Finnish Lapland. Silva Fennica vol. 41 no. 1 article id 308. https://doi.org/10.14214/sf.308
Keywords: natural regeneration; seedling establishment; Scots pine; timberline; seed-tree method; density model; height development model
Abstract | View details | Full text in PDF | Author Info
Researchers and professionals in practical forestry have faced problems concerning the regeneration success of Scots pine in natural regeneration near the timberline in North-East Lapland. The aim of the study was to analyze the seedling establishment and seedling height development of Scots pine in seed-tree stands in the area. The average number of living pine seedlings in the study stands was about 1000 ha–1, but there was considerable variation between the stands. The seedling density was modelled using a multinomial logistic regression with a random factor. Forest site type and the time since seed-tree cutting were the most significant explanatory variables in the model. The probability of reaching the acceptable seedling density was higher on dry site types than on the more fertile ones. The probability increased with the time elapsed since the regeneration activities. Effective temperature sum and the number of intermediate pines also positively affected the probability, but the presence of residual trees negatively. On northern and eastern slopes the probability was lower than on southern and western ones. Seedling height was modelled using a linear mixed model. The age of a dominant seedling was the most positively effective explanatory variable in the height development model. Other positively affecting significant predictors were time since seed-tree cutting, number of intermediate birches, and distance between a seedling and the nearest seed tree. Degree of paludification had a negative effect. The study suggests that the regeneration of Scots pine in North-East Lapland is a relatively slow process.
  • Hallikainen, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: ville.hallikainen@metla.fi (email)
  • Hyppönen, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: mikko.hypponen@metla.fi
  • Hyvönen, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: juha.hyvonen@metla.fi
  • Niemelä, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: jn@nn.fi
article id 307, category Research article
Jacqueline C. Bolli, Andreas Rigling, Harald Bugmann. (2007). The influence of changes in climate and land-use on regeneration dynamics of Norway spruce at the treeline in the Swiss Alps. Silva Fennica vol. 41 no. 1 article id 307. https://doi.org/10.14214/sf.307
Keywords: tree growth; dendroecology; microsites; subalpine meadow succession; tree invasion
Abstract | View details | Full text in PDF | Author Info
Recent changes of climate and land-use are often regarded to affect the European Alpine region substantially and to trigger an increase in the elevation of the upper treeline. The patterns of tree invasion on a subalpine meadow at 1900 m a.s.l. in Sedrun, Canton Grisons, Switzerland, were studied in order 1) to reconstruct the process of tree establishment and tree–growth dynamics in space and time, and 2) to evaluate the influence of site properties, land-use change and climate on these processes. Dendroecological analysis of 105 Norway spruce combined with an assessment of 48 vegetation plots and 17 soil profiles revealed that the trees were established in one main period (1965–1980s), starting 15 years after the abandonment of the agricultural use of the meadow, and that there is a pronounced environmental gradient along the forest-meadow ecotone. Tree establishment and height growth were favoured close to the former forest edge, but all saplings irrespective of their distance to the forest edge and their age showed increased radial growth since 1990, coinciding with a period of higher summer temperatures in the region. Therefore, we conclude that the observed tree-line dynamics in Sedrun are the result of both land-use and climate change: Tree establishment was triggered by the abandonment of the agricultural use of the meadow, and strongly favoured by particularly good growing conditions in a warm decade, which illustrates the sensitivity of conifers near the alpine tree-line to temperature fluctuations.
  • Bolli, Swiss Federal Research Institute, Zürcherstrasse 111, CH-8903 Birmensdorf, Switzerland E-mail: jacqueline.bolli@wsl.ch (email)
  • Rigling, Swiss Federal Research Institute, Zürcherstrasse 111, CH-8903 Birmensdorf, Switzerland E-mail: ar@nn.ch
  • Bugmann, Swiss Federal Institute of Technology (ETH) Zürich, Universitätstrasse 16, CH-8092 Zürich, Switzerland E-mail: hb@nn.ch
article id 306, category Research article
Niina Tanskanen, Hannu Ilvesniemi. (2007). Spatial distribution of fine roots at ploughed Norway spruce forest sites. Silva Fennica vol. 41 no. 1 article id 306. https://doi.org/10.14214/sf.306
Keywords: Picea abies; forest soil; fine roots; ploughing; spatial distribution; specific root length; understorey roots
Abstract | View details | Full text in PDF | Author Info
We examined the spatial distribution of fine roots at two forest sites that were ploughed 20 (site K1) and 33 years (site K2) before sampling and planted with Norway spruce (Picea abies (L.) Karst.) seedlings. Soil core samples were taken from the tilt and beneath the tilt, the furrow and the intermediate undisturbed soil to a depth of 0.4 m for fine root biomass, length and necromass determinations. Norway spruce fine roots were found throughout the ploughed forest sites. The fine roots were, however, unevenly distributed: the fine root biomass was highest in the tilt (624 and 452 g m–2 at sites K1 and K2, respectively) and lowest in the undisturbed soil at site K1 (79 g m–2) and in the furrow at site K2 (145 g m–2). The estimated average fine root biomass at the ploughed forest sites (268 and 248 g m–2 at sites K1 and K2, respectively) was, however, similar to those presented in other studies concerning sites that had not been ploughed. In the tilt, a substantial proportion of the fine roots was in the inverted mineral soil horizons and in the new organic horizon above the tilt. Consistent with the fine root biomass findings, the Norway spruce necromass was highest in the tilt but the vertical distribution of the dead roots was different: the necromass was highest in the buried OBT horizon. The results of this study suggest that at the ploughed forest sites, a substantial part of Norway spruce nutrient and water uptake occured in the tilt during the first 20 or 33 years after plantation.
  • Tanskanen, Department of Forest Ecology, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: niina.tanskanen@helsinki.fi (email)
  • Ilvesniemi, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: hi@nn.fi
article id 305, category Research article
Anna-Maria Veijalainen, Marja-Liisa Juntunen, Arja Lilja, Helvi Heinonen-Tanski, Leo Tervo. (2007). Forest nursery waste composting in windrows with or without horse manure or urea – the composting process and nutrient leaching. Silva Fennica vol. 41 no. 1 article id 305. https://doi.org/10.14214/sf.305
Keywords: microbial hygiene; lignocellulose; nutrients; organic matter decomposition; temperature; tree seedling waste; waste management
Abstract | View details | Full text in PDF | Author Info
In order to find the best management practices for forest nursery waste composting, organic waste was composted without or with horse manure or urea in six windrows for two years. The windrows were built in four consecutive years during 1999–2002. In 1999, no extra-nutrients were added to the windrow (N99). In 2000, urea fertilizer was used as a nitrogen source (U00). Despite this, the process did not function properly. In 2001, two windrows were built, one (H01) with and the other (N01) without horse manure. Horse manure slightly accelerated the heating process. Consequently, two windrows with more horse manure were built in 2002. One was aerated passively (H02) as earlier windrows, and the other was aerated forcedly (HA02). Horse manure and forced aeration were needed to keep the temperature above 55°C for long enough to ensure microbial hygiene of the material. The degradation of cellulose was greater during the curing stage. Nutrient leaching was low, although the additives increased leaching in conjunction with the inefficient process. The results showed that forest nursery waste alone is ineffective at raising the temperature of the compost, and degrades slowly due to its low nutrient and easily available carbon content. The best management practice for forest nursery waste composting is to use horse manure and aeration to ensure the heating process. Environmental contamination can be avoided by collecting the leachates. Further research is needed to evaluate the usability of the compost.
  • Veijalainen, Finnish Forest Research Institute, FI-77600 Suonenjoki, Finland E-mail: amv@nn.fi (email)
  • Juntunen, Finnish Forest Research Institute, FI-77600 Suonenjoki, Finland E-mail: mlj@nn.fi
  • Lilja, Finnish Forest Research Institute, PO Box 18, FI-01301 Vantaa, Finland E-mail: al@nn.fi
  • Heinonen-Tanski, Univ. of Kuopio, Dept. of Environm. Sc., PO Box 1627, FI-70211 Kuopio, Finland E-mail: hht@nn.fi
  • Tervo, Finnish Forest Research Institute, FI-77600 Suonenjoki, Finland E-mail: lt@nn.fi
article id 304, category Research article
Joanna Kosinska, Andrzej Lewandowski, Wladyslaw Chalupka. (2007). Genetic variability of Scots pine maternal populations and their progenies. Silva Fennica vol. 41 no. 1 article id 304. https://doi.org/10.14214/sf.304
Keywords: Pinus sylvestris L.; genetic variation; allozymes; gene conservation
Abstract | View details | Full text in PDF | Author Info
The genetic variability of Scots pine was investigated in six populations from Poland representing two maternal populations and their natural and artificial progenies. Thirteen enzyme systems controlled by 25 allozyme loci were analyzed using starch gel electrophoresis. Progeny populations maintained a high and similar level of genetic variation to that observed in the maternal populations. As expected, much closer genetic relationships were observed between maternal populations and their respective progeny than between the two maternal populations themselves. Progeny populations had the same major alleles, but differed in the number of rare alleles. Therefore, probably not all rare alleles were transferred from the maternal stands to the progenies. In addition, new rare alleles appeared in the progeny populations, possibly as a result of external pollen flow into the maternal populations.
  • Kosinska, Department of Human Medical Genetics, Medical University of Warsaw, Oczki 1, 02-007 Warsaw, Poland E-mail: jk@nn.pl
  • Lewandowski, Polish Academy of Sciences, Institute of Dendrology, 62-035 Kórnik, Poland E-mail: al@nn.pl (email)
  • Chalupka, Polish Academy of Sciences, Institute of Dendrology, 62-035 Kórnik, Poland E-mail: wc@nn.pl
article id 323, category Research article
Simo Kyllönen, Alfred Colpaert, Hannu Heikkinen, Mikko Jokinen, Jouko Kumpula, Mika Marttunen, Kari Muje, Kaisa Raitio. (2006). Conflict management as a means to the sustainable use of natural resources. Silva Fennica vol. 40 no. 4 article id 323. https://doi.org/10.14214/sf.323
Keywords: sustainability; conflict management; resource management; deliberative participation; frame analysis; assurance game; prisoner’s dilemma
Abstract | View details | Full text in PDF | Author Info
Democratic societies’ emphasis on individual rights and freedoms inevitably opens them up to political disputes. Conflict management should thus be seen as an integral part of democratic institutional design. The evolution and management of policy disputes concerning the use of different natural resources in Finland is analysed by using the theoretical models of frame analysis and strategic interaction. The studied disputes include lake fisheries, watercourse regulation, reindeer herding, and forestry. The institutional design in the case studies varies. Despite the differences, many common features are identified that could explain their successes or difficulties in achieving sustainable and cooperative use of the resources. Among these are problems involving complex and uncertain knowledge, differences in frames held by multiple users of a resource, and distrust between the users and other parties. The analysis concludes with preliminary conclusions on how various disputes related to sustainable resource use could be managed. These include addressing the knowledge and frame problems in order to initiate a learning process; establishing sub-processes in which mutual trust between the parties – including a managing authority or a third party – can emerge; giving explicit roles and a clear division of entitlement to the parties; and providing a credible alternative for co-operation that affects the parties’ payoff assessments during the process. Finally, the conflict management process shouldn’t be regarded as a distinct phase of dispute resolution, but as an essential aspect of ongoing co-management practices of resource use.
  • Kyllönen, Department of Social and Moral Philosophy, P.O. Box 9, FI-00014 University of Helsinki, Finland E-mail: simo.kyllonen@helsinki.fi (email)
  • Colpaert, University of Joensuu, Department of Geography, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: ac@nn.fi
  • Heikkinen, Taida, P.O. Box 1000, FI-90014 University of Oulu, Finland E-mail: hh@nn.fi
  • Jokinen, Finnish Forest Research Institute, Kolari Research Unit, Muoniontie 21 A, FI-95900 Kolari, Finland E-mail: mj@nn.fi
  • Kumpula, Finnish Game and Fisheries Research Institute, Reindeer Research Station, Toivoniementie 246, FI-99910 Kaamanen, Finland E-mail: jk@nn.fi
  • Marttunen, Finnish Environment Institute, P.O. Box 140, FI-00251 Helsinki, Finland E-mail: mk@nn.fi
  • Muje, Department of Biological and Environmental Science, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: km@nn.fi
  • Raitio, Department of Social and Policy, University of Joensuu, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: kr@nn.fi
article id 322, category Research article
Jaakko Repola. (2006). Models for vertical wood density of Scots pine, Norway spruce and birch stems, and their application to determine average wood density. Silva Fennica vol. 40 no. 4 article id 322. https://doi.org/10.14214/sf.322
Keywords: Norway spruce; Scots pine; silver birch; wood density; downy birch; mixed models
Abstract | View details | Full text in PDF | Author Info
The purpose of this study was to investigate the vertical dependence of the basic density of Scots pine, Norway spruce, and birch stems, and how such dependence could be applied for determining the average stem wood density. The study material consisted of 38 Scots pine (Pinus sylvestris), 39 Norway spruce (Picea abies [L.] Karst.) and 15 birch (Betula pendula and Betula pubescens) stands located on mineral soil sites in southern Finland. The stem material mainly represented thinning removal from stands at different stages of development. The linear mixed model technique, with both fixed and random effects, was used to estimate the model. According to the fixed part of the model, wood density was dependent on the vertical location along the stem in all three tree species. Wood density in pine decreased from the butt to the top, and the gradient in wood density was steep at the butt but decreased in the upper part of the stem. The vertical dependence was similar in birch, but the density gradient was much smaller. For spruce the vertical dependence of the basic density was moderate. The model can be calibrated for a tree stem when one or more sample disks are measured at freely selected heights. Using treewise calibrated predictions of the vertical density dependence and measured stem diameters, almost unbiased estimates, and lower prediction errors than with traditional methods, were obtained for the average stem wood density. The advantages of the method were greater for pine with a strong vertical dependence in basic density, than for spruce and birch.
  • Repola, Finnish Forest Research Institute, Rovaniemi Research Unit, Eteläranta 55, FI-96300 Rovaniemi, Finland E-mail: jaakko.repola@metla.fi (email)
article id 321, category Research article
Jori Uusitalo, Anne Puustelli, Veli-Pekka Kivinen, Tapio Nummi, Bikas K. Sinha. (2006). Bayesian estimation of diameter distribution during harvesting. Silva Fennica vol. 40 no. 4 article id 321. https://doi.org/10.14214/sf.321
Keywords: diameter distribution; Bayesian statistics; tree harvesting
Abstract | View details | Full text in PDF | Author Info
This research aims to combine two different data sets with Bayesian statistics in order to predict the diameter distribution of trees at harvest. The parameters of prior distribution are derived from the forest management plans supplemented by additional ocular information. We derive the parameters for the sample data from the first trees harvested, and then create the posterior distribution within the Bayesian framework. We apply the standard normal distribution to construct diameter (dbh) distributions, although many other theoretical distributions have been proved better with dbh data available. The methodology developed is then tested on nine mature spruce (Picea abies) dominated stands, on which the normal distribution seems to work well in mature spruce stands. The tests indicate that prediction of diameter distribution for the whole stand based on the first trees harvested is not wise, since it tends to give inaccurate predictions. Combining the first trees harvested with prior information seems to increase the reliability of predictions.
  • Uusitalo, The Finnish Forest Research Institute, Parkano unit, FI-39700 Parkano, Finland E-mail: jori.uusitalo@metla.fi (email)
  • Puustelli, University of Tampere, Department of Mathematics, Statistics and Philosophy, FI-33014 University of Tampere, Finland E-mail: ap@nn.fi
  • Kivinen, University of Helsinki, Department of Forest Resource Management, Box 27, FI-00014 University of Helsinki, Finland E-mail: vpk@nn.fi
  • Nummi, University of Tampere, Department of Mathematics, Statistics and Philosophy, FI-33014 University of Tampere, Finland E-mail: tn@nn.fi
  • Sinha, Stat-Math Division, Indian Statistical Institute, 203 B.T. Road, Kolkata - 700 108, India E-mail: bks@nn.in
article id 320, category Research article
Saija Huuskonen, Jari Hynynen. (2006). Timing and intensity of precommercial thinning and their effects on the first commercial thinning in Scots pine stands. Silva Fennica vol. 40 no. 4 article id 320. https://doi.org/10.14214/sf.320
Keywords: Pinus sylvestris; Scots pine; growth and yield; precommercial thinning; growth modelling; first commercial thinning; diameter development
Abstract | View details | Full text in PDF | Author Info
The effects of the timing and intensity of precommercial thinning on the stand diameter development and wood production in Scots pine stands was addressed. A model was developed in order to assess the thinning response of the stand diameter development. The effect of precommercial and first commercial thinning on the stand volume and the thinning removal at first commercial thinning were also modelled. The models were developed to be applicable for forest management planning purposes. The results are based on Scots pine (Pinus sylvestris L.) trials (13 experiments and 169 plots) located in Southern and Central Finland. Precommercial thinning considerably enhanced the diameter development. Precommercial thinning (at Hdom 3 m to 2000 trees per hectare) increased the mean diameter by 15% at the first commercial thinning stage (Hdom 14 m) compared to the unthinned stand (3000 trees ha–1). Early and intensive precommercial thinning resulted in the strongest response in diameter development. Wide spacing also enhanced the diameter increment. In naturally regenerated stands the diameter development was ca 13% slower than that in seeded stands. The total volume at the time of first commercial thinning was affected by the timing of thinning and the stand structure. The volume of merchantable thinning removal depended on the timing and intensity of precommercial and first commercial thinnings. Delaying the first commercial thinning from 12 meters (Hdom) to 16 meters increased the volume of thinning removal by ca.70%. The early and light precommercial thinning (Hdom 3 m, to density of 3000 trees per hectare) increased the thinning removal by 40% compared to the late and intensive precommercial thinning (at 7 meters to the density of 2000 trees per hectare).
  • Huuskonen, University of Helsinki, Dept. of Forest Ecology, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: sh@nn.fi (email)
  • Hynynen, The Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: jh@nn.fi
article id 319, category Research article
Ulises Diéguez-Aranda, José Antonio Grandas-Arias, Juan Gabriel Álvarez-González, Klaus von Gadow. (2006). Site quality curves for birch stands in north-western Spain. Silva Fennica vol. 40 no. 4 article id 319. https://doi.org/10.14214/sf.319
Keywords: site quality; Betula pubescens; generalized algebraic difference approach; base-age invariant dynamic equation; Hossfeld model; Galicia
Abstract | View details | Full text in PDF | Author Info
A model for predicting the height growth of even-aged, birch (Betula pubescens Ehrh.) dominated stands in Galicia (north-western Spain) was developed. Data from stem analysis of 214 trees were used for model construction. Two dynamic site equations derived with the generalized algebraic difference approach (GADA) were tested, which combine compatible site index and height models in one common equation. Both equations are base-age invariant and directly estimate height and site index from any height and age. The fittings were done in one stage using the base-age-invariant dummy variables method. A second-order continuous-time autoregressive error structure was used to correct the inherent autocorrelation of the longitudinal data used in this study. Cieszewski’s model best described the data. This model is therefore recommended for height growth prediction and site classification of birch stands in Galicia.
  • Diéguez-Aranda, Departamento de Ingeniería Agroforestal, Universidad de Santiago de Compostela. Escuela Politécnica Superior, Campus universitario, 27002 Lugo, Spain E-mail: udieguez@lugo.usc.es (email)
  • Grandas-Arias, Departamento de Ingeniería Agroforestal, Universidad de Santiago de Compostela. Escuela Politécnica Superior, Campus universitario, 27002 Lugo, Spain E-mail: jaga@nn.es
  • Álvarez-González, Departamento de Ingeniería Agroforestal, Universidad de Santiago de Compostela. Escuela Politécnica Superior, Campus universitario, 27002 Lugo, Spain E-mail: jgag@nn.es
  • Gadow, Institut fur Waldinventur und Waldwachstum, George-Auguts-Universität Göttingen. Büsgenweg 5, D-37077 Göttingen, Germany E-mail: kg@nn.de
article id 318, category Research article
Sofia Backéus, Peder Wikström, Tomas Lämås. (2006). Modeling carbon sequestration and timber production in a regional case study. Silva Fennica vol. 40 no. 4 article id 318. https://doi.org/10.14214/sf.318
Keywords: boreal forest; carbon sequestration; optimization; forest biofuel; forest management planning
Abstract | View details | Full text in PDF | Author Info
Forests make up large ecosystems and by the uptake of carbon dioxide can play an important role in mitigating the greenhouse effect. In this study, mitigation of carbon emissions through carbon uptake and storage in forest biomass and the use of forest biofuel for fossil fuel substitution were considered. The analysis was performed for a 3.2 million hectare region in northern Sweden. The objective was to maximize net present value for harvested timber, biofuel production and carbon sequestration. A carbon price for build-up of carbon storage and for emissions from harvested forest products was introduced to achieve an economic value for carbon sequestration. Forest development was simulated using an optimizing stand-level planning model, and the solution for the whole region was found using linear programming. A range of carbon prices was used to study the effect on harvest levels and carbon sequestration. At a zero carbon price, the mean annual harvest level was 5.4 million m3, the mean annual carbon sequestration in forest biomass was 1.48 million tonnes and the mean annual replacement of carbon from fossil fuel with forest biofuel was 61 000 tonnes. Increasing the carbon price led to decreasing harvest levels of timber and decreasing harvest levels of forest biofuel. Also, thinning activities decreased more than clear-cut activities when the carbon prices increased. The level of carbon sequestration was governed by the harvest level and the site productivity. This led to varying results for different parts of the region.
  • Backéus, SLU, Dept. of Forest Resource Management and Geomatics, SE-901 83 Umeå, Sweden E-mail: sofia.backeus@resgeom.slu.se (email)
  • Wikström, SLU, Dept. of Forest Resource Management and Geomatics, SE-901 83 Umeå, Sweden E-mail: pw@nn.se
  • Lämås, SLU, Dept. of Forest Resource Management and Geomatics, SE-901 83 Umeå, Sweden E-mail: tl@nn.se
article id 317, category Research article
Erik Eriksson, Tord Johansson. (2006). Effects of rotation period on biomass production and atmospheric CO2 emissions from broadleaved stands growing on abandoned farmland. Silva Fennica vol. 40 no. 4 article id 317. https://doi.org/10.14214/sf.317
Keywords: biomass; biofuel; carbon stock; abandoned farmland; atmospheric CO2; broadleaved trees; MAI
Abstract | View details | Full text in PDF | Author Info
The growth rates and carbon stocks of unthinned young and mature stands of broadleaved trees growing on abandoned farmland were determined to assess whether their management regimes should involve short (15-year) or long (45-year) rotations to maximize biomass production and reductions of CO2 emissions. Dry mass production and mean annual increment (MAI) were calculated for 28 young stands and 65 mature stands of European aspen (Populus tremula L.), common alder (Alnus glutinosa (L.) Gaertn.), grey alder (Alnus incana (L.) Moench.), silver birch (Betula pendula Roth) and downy birch (Betula pubescens Ehrh.) ranging in latitude from 57° to 63° N in Sweden. The potential for using biomass from the stands to replace coal as a fuel and to store carbon was then evaluated both in short and long rotation scenarios. The results indicate that long rotations are beneficial if the objective is to maximize the average carbon stock in biomass. If, on the other hand, the intention is to optimize reductions in atmospheric CO2 emissions, rotations should be short for aspen, silver birch and grey alder stands. For downy birch and common alder, the MAI was higher for the mature stands than the young stands, indicating that in these species the mature stands are superior for both storing carbon and replacing fossil fuel. Stands of broadleaved trees grown to produce biofuel on abandoned farmland should be established on fertile soils to promote high MAI. If the MAI is low, the rotation period should be long to maximize the average carbon stock.
  • Eriksson, SLU, Dept of Bioenergy, P.O. Box 7061, SE-750 07 Uppsala, Sweden E-mail: ee@nn.se (email)
  • Johansson, SLU, Dept of Bioenergy, P.O. Box 7061, SE-750 07 Uppsala, Sweden E-mail: tj@nn.se
article id 316, category Research article
Sonja Vospernik. (2006). Probability of bark stripping damage by red deer (Cervus elaphus) in Austria. Silva Fennica vol. 40 no. 4 article id 316. https://doi.org/10.14214/sf.316
Keywords: logistic regression; bark stripping; red deer
Abstract | View details | Full text in PDF | Author Info
Bark stripping by red deer (Cervus elaphus) causes considerable damage to Austrian forests, however, the incidence of bark stripping was never examined from large scale survey data. In this manuscript we present a logistic regression model for bark stripping damage (static model) and a model for recent (5-year period) bark stripping damage to previously undamaged trees (dynamic model) developed from Austrian National Forest Inventory data. Both models showed bark stripping damage to be most frequent in core red deer habitat areas and less frequent in less suitable habitat. Damage was concentrated at elevations of 400–1200 m and in alluvial forests (only static model). Norway spruce (Picea abies), European ash (Fraxinus excelsior), Sweet chestnut (Castanea sativa) and Sorbus spp. had 11–12 times more injuries than all the other species. Red deer preferred the smallest trees with a breast height diameter of 5 cm for bark stripping and damage probability decreased rapidly for trees with a breast height diameter greater than 25 cm. Our static model showed a maximum of bark stripping damage in stands with a mean height of 20 m. In the dynamic model the probability for bark stripping damage decreased with decreasing mean height. Also, in the static model the probability for bark stripping damage increased with increasing spruce proportion and with increasing stand density whereas in the dynamic model the proportion of previous bark stripping damage was a good predictor. Goodness of fit and discrimination of both models were good. In combination with forest growth models, the bark stripping models can be used to predict the risk of damage associated with different forest and habitat management options.
  • Vospernik, University of Natural Resources and Applied Life Sciences, Institute of Forest Growth Research, Department of Forest and Soil Sciences, Peter-Jordan-Stra§e 82, A-1190 Vienna, Austria E-mail: sonja.vospernik@boku.ac.at (email)
article id 315, category Research article
Lauri Korhonen, Kari T. Korhonen, Miina Rautiainen, Pauline Stenberg. (2006). Estimation of forest canopy cover: a comparison of field measurement techniques. Silva Fennica vol. 40 no. 4 article id 315. https://doi.org/10.14214/sf.315
Keywords: canopy cover; forest canopy; canopy closure; Cajanus tube; line intersect sampling; spherical densiometer; digital photographs
Abstract | View details | Full text in PDF | Author Info
Estimation of forest canopy cover has recently been included in many forest inventory programmes. In this study, after discussing how canopy cover is defined, different ground-based canopy cover estimation techniques are compared to determine which would be the most feasible for a large scale forest inventory. Canopy cover was estimated in 19 Scots pine or Norway spruce dominated plots using the Cajanus tube, line intersect sampling, modified spherical densiometer, digital photographs, and ocular estimation. The comparisons were based on the differences in values acquired with selected techniques and control values acquired with the Cajanus tube. The statistical significance of the differences between the techniques was tested with the nonparametric Kruskall-Wallis analysis of variance and multiple comparisons. The results indicate that different techniques yield considerably different canopy cover estimates. In general, labour intensive techniques (the Cajanus tube, line intersect sampling) provide unbiased and more precise estimates, whereas the estimates provided by fast techniques (digital photographs, ocular estimation) have larger variances and may also be seriously biased.
  • Korhonen, University of Joensuu, P.O. Box 68, FI-68101 Joensuu, Finland E-mail: lauri.korhonen@joensuu.fi (email)
  • Korhonen, University of Joensuu, P.O. Box 68, FI-68101 Joensuu, Finland E-mail: ktk@nn.fi
  • Rautiainen, University of Joensuu, P.O. Box 68, FI-68101 Joensuu, Finland E-mail: mr@nn.fi
  • Stenberg, University of Joensuu, P.O. Box 68, FI-68101 Joensuu, Finland E-mail: ps@nn.fi
article id 314, category Research article
Yuhua Wang, Helena Korpelainen, Chunyang Li. (2006). Microsatellite polymorphism in the edaphic spruce, Picea asperata, originating from the mountains of China. Silva Fennica vol. 40 no. 4 article id 314. https://doi.org/10.14214/sf.314
Keywords: genetic diversity; microsatellite polymorphism; micro-geographic differentiation; natural selection; mutation
Abstract | View details | Full text in PDF | Author Info
Microsatellite variation of Picea asperata Mast. originating from the mountains of China was investigated by analyzing variation at seven SSR loci in 250 individuals representing ten populations. A fair degree of genetic diversity and considerable population subdivision occurred with the mean gene diversity (H) of 0.707, and genetic distances among populations varying between 0.121 and 0.224 (FST) and between 0.100 and 0.537 (RST). However, inter-population genetic distances showed no correlation with geographic distances between the population sites. This ruled out a simple isolation by distance model and suggested that migration does not have a great impact. In fact, the amount of gene flow, detected using private alleles, was very low, equaling only 0.753. Allele permutation tests revealed that stepwise-like mutations, coupled with genetic drift, could contribute to population differentiation. Moreover, significant genetic differences between populations were detected at most loci. The results indicate that natural selection, presumably through environmental stress, may be one of the main factors causing micro-geographical differentiation in the genetic structure of P. asperata. Based on SSR genotypes, 70% of the 250 individuals were correctly classified into their sites of origin. This suggests that microsatellites (SSRs) are effective in distinguishing genotypes of P. asperata originating from diverse eco-geographical sites in China.
  • Wang, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O. Box 416, Chengdu 610041, China; Graduate School of the Chinese Academy of Sciences, Beijing 100039, China E-mail: yw@nn.cn
  • Korpelainen, Department of Applied Biology, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: hk@nn.fi
  • Li, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O. Box 416, Chengdu 610041, China E-mail: licy@cib.ac.cn (email)
article id 476, category Research article
Ignazio Monteleone, Diana Ferrazzini, Piero Belletti. (2006). Effectiveness of neutral RAPD markers to detect genetic divergence between the subspecies uncinata and mugo of Pinus mugo Turra. Silva Fennica vol. 40 no. 3 article id 476. https://doi.org/10.14214/sf.476
Keywords: molecular markers; population differentiation; genetic diversity; Pinus mugo; mountain pine; dwarf mountain pine
Abstract | View details | Full text in PDF | Author Info
Fifteen populations of Pinus mugo subsp. mugo (shrub) and Pinus mugo subsp. uncinata (erect), located in the Alps, were investigated through genetic variation scored at 64 polymorphic RAPD loci. In addition, morphological traits of the female cones were analysed. According to AMOVA most of the genetic variation was found within populations (83.39%), and only 1.25% of it between subspecies. Populations differed in terms of their internal genetic variation, with Nei’s gene diversity ranging from 0.227 to 0.397. Morphological data showed differences between subspecies, although none of the populations showed full accordance with expectations. Significant correlation was found between matrices for geographical and morphological distances, while genetic distances were not correlated with any other aspect. The efficacy of morphological and RAPD markers in discriminating between subspecies, and the contribution of the results in relation to the preservation of biodiversity, are discussed.
  • Monteleone, DI.VA.P.R.A., Plant Genetics and Breeding, University of Turin, via Leonardo da Vinci 44, I-10095 Grugliasco, Italy E-mail: im@nn.it
  • Ferrazzini, DI.VA.P.R.A., Plant Genetics and Breeding, University of Turin, via Leonardo da Vinci 44, I-10095 Grugliasco, Italy E-mail: df@nn.it
  • Belletti, DI.VA.P.R.A., Plant Genetics and Breeding, University of Turin, via Leonardo da Vinci 44, I-10095 Grugliasco, Italy E-mail: piero.belletti@unito.it (email)
article id 336, category Research article
Ann Hedlund. (2006). The attractiveness of the work is affected when production of handcrafted log houses moves indoors. Silva Fennica vol. 40 no. 3 article id 336. https://doi.org/10.14214/sf.336
Keywords: log house building; attractive work; indoor; outdoor; work environment
Abstract | View details | Full text in PDF | Author Info
Viewed from a historical perspective, a shift has occurred within the forestry and wood sector towards indoor work. In Sweden, the production of handcrafted log houses has now also begun to move indoors. With a point of departure in development processes within the log house sector involving working indoors, education, work attractiveness, between 2001–2005, the aim of this study was to compare indoor work with outdoor work, based on log house builders’ experience of working on handcrafted log houses. Methods used in the interactive development project involving apprentices, experienced log house builders and researchers, were participation with continuous documentation of experiences and opinions; questions; interviews; and measurement of the work environment. The Attractive Work Model has been used in order to analyse perceptions and values. The changes, 15 out of 22 areas, were perceived both negatively and positively. Therefore, it can not be said that working on traditional, handcrafted log houses becomes more attractive if it is moved indoors. The majority wanted to work both outdoors and indoors, while most of the others only wanted to work outdoors. The results indicate that there is scope for developing more attractive work indoors by utilising experiences from log house builders and closely related activities such as the forestry and wood sector. Changes made within one area of work attractiveness affect other areas. Further research is needed both with regard to comparisons between indoor and outdoor work and regarding the interaction between the areas that are identified in the Attractive Work Model.
  • Hedlund, Dalarna University, SE-781 88 Borlänge, Sweden E-mail: ahd@du.se (email)
article id 335, category Research article
Markus Holopainen, Mervi Talvitie. (2006). Effect of data acquisition accuracy on timing of stand harvests and expected net present value. Silva Fennica vol. 40 no. 3 article id 335. https://doi.org/10.14214/sf.335
Keywords: forest inventory; laser scanning; digital aerial photographs; digital photogrammetry; net present value; expected net present value loss
Abstract | View details | Full text in PDF | Author Info
Modern remote sensing provides cost-efficient spatial digital data that are more accurate than before. However, the influence of increased accuracy and cost-efficiency on simulations of forest management planning has not been evaluated. The aim of the present study was to analyse the effect of data acquisition accuracy on standwise forest inventory by comparing the accuracy and cost of traditional compartmentwise inventory methods with 2D and 3D measurements of digital aerial photographs and airborne laser scanning. Comparison was based on the expected net present value (NPV), i.e. economic losses that consisted of the inventory costs and incorrect timings of treatments. The reference data, totalling 700 ha, were measured from Central Park in the city of Helsinki, Finland. The data were simulated to final cut with a MOTTI simulator, which is a stand-level analysis tool that can be used to assess the effects of alternative forest management practices on growth and timber yield. The results showed that when inventory costs were not considered there were no significant differences between the expected NPV losses in 3D measurements of digital aerial photographs, laser scanning and the compartmentwise method. When inventory costs were taken into account, the compartmentwise method was still the most efficient inventory method in the study area. Forest inventories, however, are usually directed to larger areas when the costs per hectare of remote-sensing methods decrease. As a result of better accuracies, 3D and compartmentwise methods always produce better results than the 2D method when NPV losses are accounted. Simulations of this type are based on the accuracies and costs of the 3D data available today, assuming that the data can be used in tree-level measurements.
  • Holopainen, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: markus.holopainen@helsinki.fi (email)
  • Talvitie, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: mt@nn.fi
article id 334, category Research article
Jouni Siipilehto. (2006). Linear prediction application for modelling the relationships between a large number of stand characteristics of Norway spruce stands. Silva Fennica vol. 40 no. 3 article id 334. https://doi.org/10.14214/sf.334
Keywords: Picea abies; stand characteristics; prediction; calibration estimation; interdependence
Abstract | View details | Full text in PDF | Author Info
The aim was to produce models for a large number of stand characteristics of Norway spruce dominated stands. A total of 227 national forest inventory based permanent stand plots, dominated by Norway spruce (Picea abies), were used in modelling eight stand variables as a function of the stand mean biological age and site characteristics. The basic models were able to characterize the average development of the modelled stand variables, but resulted in a relatively high RMSE. Basal area (G) and stem number (N) were the most inaccurate, having a RMSE of 34–41%, while that of mean diameter and height characteristics varied between 16–20%. The expectations and error variances of the basic models were calibrated with known stand variables using linear prediction theory. The best linear unbiased predictor (BLUP) with a single stand variable used for calibration proved to be ineffective for unknown G and N, but relatively effective for the unknown mean characteristics. However, calibration with one sum and one mean characteristic proved to be effective, and additional calibration variables enhanced the precision only marginally. The BLUP method provided a flexible approach when characterizing the relationships between a large number of stand variables, thus enabling multiple use of these models because they were not fixed to a specific inventory system.
  • Siipilehto, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: jouni.siipilehto@metla.fi (email)
article id 333, category Research article
Lauri Mehtätalo, Matti Maltamo, Annika Kangas. (2006). The use of quantile trees in the prediction of the diameter distribution of a stand. Silva Fennica vol. 40 no. 3 article id 333. https://doi.org/10.14214/sf.333
Keywords: stand structure; inventory; percentile; order statistics
Abstract | View details | Full text in PDF | Author Info
This study deals with the prediction of the basal area diameter distribution of a stand without using a complete sample of diameters from the target stand. Traditionally, this problem has been solved by either the parameter recovery method or the parameter prediction method. This study uses the parameter prediction method and the percentile based diameter distribution with a recent development that makes it possible to improve these predictions by using sample order statistics. A sample order statistic is a tree whose diameter and rank at the plot are known, and is referred to in this paper as a quantile tree. This study tested 13 different strategies for selection of the quantile trees from among the trees of horizontal point sample plots, and compared them with respect to RMSE and the bias of four criterion variables in a dataset of 512 stands. The sample minimum was found to be the most promising alternative with respect to RMSE, even though it introduced a rather large amount of bias in the criterion variables. Other good and less biased alternatives are the second and third smallest trees and the tree closest to the plot centre. The use of minimum is recommended for practical inventories because its rank is probably easiest to determine correctly in the field.
  • Mehtätalo, Yale School of Forestry and Environmental Studies, 205 Prospect Street, New Haven, CT 06511, USA E-mail: lauri.mehtatalo@metla.fi (email)
  • Maltamo, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: mm@nn.fi
  • Kangas, University of Helsinki, Department of Forest Resources Management, P.O.Box 27, FI-00014 University of Helsinki, Finland E-mail: ak@nn.fi
article id 332, category Research article
Marc Palahí, Timo Pukkala, Antoni Trasobares. (2006). Calibrating predicted tree diameter distributions in Catalonia, Spain. Silva Fennica vol. 40 no. 3 article id 332. https://doi.org/10.14214/sf.332
Keywords: calibration estimation; Weibull function; parameter prediction
Abstract | View details | Full text in PDF | Author Info
Several probability density functions have been used in describing the diameter distributions of forest stands. In a case where both the stand basal area and number of stems per hectare are assessed, the fitted or predicted distribution is scaled using only one of these variables, with the result that the distribution often gives incorrect values for the other variable. Using a distribution that provides incorrect values for known characteristics means wasting information. Calibrating the distribution so that it is compatible with the additional information on stand characteristics is a way to avoid such wasting. This study examined the effect of calibration on the accuracy of the predicted diameter distributions of the main tree species of Catalonia. The distributions were calibrated with and without considering the prediction errors of the frequencies of diameter classes. When prediction errors were assumed, the calibration was done with and without making allowance for estimation errors in the stand level calibration variables. Calibrated distributions were more accurate than non-calibrated in terms of sums of different powers of diameters. The set of calibration variables that gave the most accurate results included six stand variables: number of trees per hectare, stand basal area, basal-area-weighted mean diameter, non-weighted mean diameter, median diameter, and basal area median diameter. Of the tested three-variable combinations the best was: number of trees per hectare, stand basal area, and basal-area-weighted mean diameter. Means were more useful calibration variables than medians.
  • Palahí, Centre Tecnológic Forestal de Catalunya. Passeig Lluis Companys, 23, 08010, Barcelona, Spain E-mail: marc.palahi@ctfc.es (email)
  • Pukkala, University of Joensuu, Faculty of Forestry, P.O. Box 111, 80101 Joensuu, Finland E-mail: tp@nn.fi
  • Trasobares, Foreco Technologies, Av. Diagonal 416, Estudio 2, Barcelona 08037, Spain E-mail: at@nn.es
article id 331, category Research article
Jouni Siipilehto. (2006). Height distributions of Scots pine sapling stands affected by retained tree and edge stand competition. Silva Fennica vol. 40 no. 3 article id 331. https://doi.org/10.14214/sf.331
Keywords: Pinus sylvestris; retention; height distribution; Weibull function; percentile prediction; edge effect
Abstract | View details | Full text in PDF | Author Info
The paper focused on the height structure of Scots pine saplings affected by (1) retained solitary pine trees or (2) a pine-dominated edge stand. The study material in (1) and (2) consisted of ten separate regeneration areas in southern Finland. In (1) 2-m radius study plots were located at 1, 3, 6 and 10 m distances from 10 systematically selected, solitary retained trees in each stand. In (2) the study plots were systematically located within 20 m from the edge stand. Competition of the individual trees was modelled using ecological field theory. The 24th and 93rd sample percentiles were used for estimating the height distribution using the two-parameter Weibull function. The models incorporated the effect of varying advanced tree competition on the predicted percentiles. Competition free dominant height was used as a driving variable for the developmental phase. Competition resulted in retarded height development within a radius of about 6 m from the retained tree, while it extended up to roughly half of the dominant height of the edge stand. The height distribution without external competition was relatively symmetrical, but increasing competition resulted in a more peaked and skewed distribution. Slight differences were found between northern sunny and southern shaded stand edges, while the least retarded height occurred at the north-western edge receiving morning sunlight. Kolmogorov-Smirnov goodness-of-fit tests showed acceptable and equal fit for both data sets; 2% and 8% of the distributions did not pass the test at the alpha 0.1 level when the Weibull distribution was estimated with the observed or predicted percentiles, respectively.
  • Siipilehto, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: jouni.siipilehto@metla.fi (email)
article id 330, category Research article
Benoit Lapointe, Robert Bradley, William Parsons, Suzanne Brais. (2006). Nutrient and light availability to white spruce seedlings in partial and clearcut harvested aspen stands. Silva Fennica vol. 40 no. 3 article id 330. https://doi.org/10.14214/sf.330
Keywords: foliar nutrients; DIFN light; Picea glauca; soil mineral nitrogen; vector analysis
Abstract | View details | Full text in PDF | Author Info
  • Lapointe, Université de Sherbrooke, Département de biologie, 2500 boulevard de l’Université, Sherbrooke, QC, Canada J1K 2R1 E-mail: bl@nn.ca
  • Bradley, Université de Sherbrooke, Département de biologie, 2500 boulevard de l’Université, Sherbrooke, QC, Canada J1K 2R1 E-mail: robert.bradley@usherbrooke.ca (email)
  • Parsons, Université de Sherbrooke, Département de biologie, 2500 boulevard de l’Université, Sherbrooke, QC, Canada J1K 2R1 E-mail: wp@nn.ca
  • Brais, UQAT, 445 boulevard Université, Rouyn-Noranda, QC, Canada J9X 5E4 E-mail: sb@nn.ca
article id 329, category Research article
Vesa Juntunen, Seppo Neuvonen. (2006). Natural regeneration of Scots pine and Norway spruce close to the timberline in northern Finland. Silva Fennica vol. 40 no. 3 article id 329. https://doi.org/10.14214/sf.329
Keywords: Norway spruce; regeneration; Scots pine; timberline; tree line; seed maturity; seedling mortality
Abstract | View details | Full text in PDF | Author Info
Two different datasets were analyzed in order to clarify the factors that affect regeneration success of Scots pine (Pinus sylvestris) and Norway spruce (Picea abies) in the climatically extreme areas in northern Finland. First, pine seed maturity and the number of cones in the trees were investigated at five pairs of study sites during the period 1997–2003. Secondly, the rate of seedling establishment and seedling survival of Scots pine and Norway spruce were monitored and compared among three different timberline zones (forest zone, timberline, tree line) in 13 localities during the period 1983–1999. The first study showed that both cone production (bud formation) and seed maturity may be limiting factors for successful reproduction in the climatically marginal habitats. Seed maturity correlated well with the temperature sum of the summer, but variation in the number of cones had a periodic component rather than strictly following the temperature sum of the summer of bud formation. Monitoring surveys since 1983 showed that pine and spruce regenerated more or less regularly in all the zones during 1983–1999. However, seedling mortality of pines was much higher compared to spruce. In general, initially small sized seedlings showed higher mortality compared with larger ones. The results suggest that besides restrictions in reproduction, stand dynamics in the timberline habitats are strongly controlled by seedling mortality due to a variety of causes.
  • Juntunen, Finnish Forest Research Institute, Kolari Research Unit, Muoniontie 21, FI-95900 Kolari, Finland E-mail: vesa.juntunen@metla.fi (email)
  • Neuvonen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: sn@nn.fi
article id 328, category Research article
Pedro J. Aphalo, Markku Lahti, Tarja Lehto, Tapani Repo, Aino Rummukainen, Hannu Mannerkoski, Leena Finér. (2006). Responses of silver birch saplings to low soil temperature. Silva Fennica vol. 40 no. 3 article id 328. https://doi.org/10.14214/sf.328
Keywords: biomass; Betula pendula; photosynthesis; electrical impedance; mineral nutrients; soil temperature; stomatal conductance; water relations
Abstract | View details | Full text in PDF | Author Info
Two-year-old silver birch (Betula pendula) saplings were grown for a third growing season in controlled-environment rooms (dasotrons) at three soil temperatures (5, 10, and 20 °C). All trees grew the first flush of leaves, but the growth of the second flush was almost completely inhibited at the two lower temperatures. The dry weight of the second-flush leaves was 50 times larger at 20 °C than at 5 and 10 °C, with about 100 times more nitrogen. Root growth was less affected than shoot growth. Chlorophyll content, net assimilation rate and stomatal conductance were lower at low soil temperatures. The value of the cytoplasm resistance estimated from the electric impedance spectra was lower at 5 °C than at 10 or 20 °C. Leaf water potential was highest at the lowest soil temperature, and intercellular carbon dioxide concentration was only slightly lower in saplings growing in cooler soil. We conclude that the effect of long-term exposure to cold soil on net assimilation and growth was not caused by stomatal closure alone. It is likely to be additionally mediated by the limited nitrogen acquisition at the low soil temperatures, and perhaps additionally by some other factor. As the growth depression of aboveground parts in response to low soil temperature was more significant in silver birch than what has earlier been found in conifers, the relative changes in air and soil temperature may eventually determine whether birch will become more dominant in boreal forests with climate change.
  • Aphalo, University of Helsinki, Department of Biological and Environmental Sciences E-mail: pja@nn.fi
  • Lahti, The Finnish Forest Research Institute E-mail: ml@nn.fi
  • Lehto, University of Joensuu, Faculty of Forestry, Box 111, FI-80101 Joensuu, Finland E-mail: tarja.lehto@joensuu.fi (email)
  • Repo, The Finnish Forest Research Institute E-mail: tr@nn.fi
  • Rummukainen, University of Joensuu, Faculty of Forestry, Box 111, FI-80101 Joensuu, Finland E-mail: ar@nn.fi
  • Mannerkoski, University of Joensuu, Faculty of Forestry, Box 111, FI-80101 Joensuu, Finland E-mail: hm@nn.fi
  • Finér, The Finnish Forest Research Institute E-mail: lf@nn.fi
article id 327, category Research article
Gong Zhang, Guang-ming Zeng, Yi-min Jiang, Chun-yan Du, Guo-he Huang, Jia-mei Yao, Mei Zeng, Xi-lin Zhang, Wei Tan. (2006). Seasonal dry deposition and canopy leaching of base cations in a subtropical evergreen mixed forest, China. Silva Fennica vol. 40 no. 3 article id 327. https://doi.org/10.14214/sf.327
Keywords: throughfall; bulk precipitation; dry deposition; canopy exchange; base cations; evergreen forest
Abstract | View details | Full text in PDF | Author Info
We evaluated the dry deposition and canopy leaching fluxes of base cations in the growing and the dormant seasons using the Na-ratio method based on the 4-year (2000–2003) monitoring data in Shaoshan subtropical evergreen mixed forest, China. The dry deposition of base cations in the growing seasons was lower than that in the dormant seasons, while the canopy leaching of base cations was higher in the growing seasons than that in the dormant seasons. The precipitation quantity and H+ significantly impacted the canopy leaching processes. The annual canopy leaching of K+, Ca2+ and Mg2+ accounted for 88, 46 and 38% of net throughfall flux, respectively. The canopy retention of proton (H+ and NH4+) is close to the canopy leaching of base cations calibrated by weak acids, indicating that the canopy cations leaching is neutralizing acid precipitation.
  • Zhang, College of Environmental Science and Engineering, Hunan University, Hunan Province, Changsha 410082, P.R. China; Hunan Environmental Protection Bureau, Hunan Province, Changsha, 410076, P.R. China E-mail: gz@nn.cn
  • Zeng, College of Environmental Science and Engineering, Hunan University, Hunan Province, Changsha 410082, P.R. China E-mail: zgming@hnu.cn (email)
  • Jiang, College of Environmental Science and Engineering, Hunan University, Hunan Province, Changsha 410082, P.R. China; Hunan Environmental Protection Bureau, Hunan Province, Changsha, 410076, P.R. China E-mail: ymj@nn.cn
  • Du, College of Environmental Science and Engineering, Hunan University, Hunan Province, Changsha 410082, P.R. China E-mail: cyd@nn.cn
  • Huang, College of Environmental Science and Engineering, Hunan University, Hunan Province, Changsha 410082, P.R. China E-mail: ghh@nn.cn
  • Yao, Xiangya Hospital, Central-south University, Hunan Province, Changsha 410008, P.R. China E-mail: jmy@nn.cn
  • Zeng, College of Environmental Science and Engineering, Hunan University, Hunan Province, Changsha 410082, P.R. China E-mail: mz@nn.cn
  • Zhang, College of Environmental Science and Engineering, Hunan University, Hunan Province, Changsha 410082, P.R. China E-mail: xlz@nn.cn
  • Tan, College of Environmental Science and Engineering, Hunan University, Hunan Province, Changsha 410082, P.R. China E-mail: wt@nn.cn
article id 326, category Research article
Xuejiang Zhang, Helena Korpelainen, Chunyang Li. (2006). Microsatellite variation of Quercus aquifolioides populations at varying altitudes in the Wolong Natural Reserve of China. Silva Fennica vol. 40 no. 3 article id 326. https://doi.org/10.14214/sf.326
Keywords: genetic differentiation; genetic variation; altitudinal gradients; microsatellites; Quercus aquifolioides
Abstract | View details | Full text in PDF | Author Info
Genetic variation and differentiation were investigated among five natural populations of Quercus aquifolioides occurring along an altitudinal gradient that varied from 2000 to 3600 m above sea level in the Wolong Natural Reserve of China, by analyzing variation at six microsatellite loci. The results showed that the populations were characterized by relatively high intra-population variation with the average number of alleles equaling 11.33 per locus and the average expected heterozygosity (HE) being 0.779. The amount of genetic variation varied only little among populations, which suggests that the influence of altitude factors on microsatellite variation is limited. However, there is a significantly positive correlation between altitude and the number of low-frequency alleles (R2 = 0.97, P < 0.01), which indicates that Q. aquifolioides from high altitudes has more unique variation, possibly enabling adaptation to severe conditions. F statistics showed the presence of a slight deficiency of heterozygosity (FIS = 0.136) and a low level of differentiation among populations (FST = 0.066). The result of the cluster analysis demonstrated that the grouping of populations does not correspond to the altitude of the populations. Based on the available data, it is likely that the selective forces related to altitude are not strong enough to significantly differentiate the populations of Q. aquifolioides in terms of microsatellite variation.
  • Zhang, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O. Box 416, Chengdu 610041, P. R. China; Graduate School of the Chinese Academy of Sciences, Beijing 100039, P. R. China E-mail: xz@nn.cn (email)
  • Korpelainen, Department of Applied Biology, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: hk@nn.fi
  • Li, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O. Box 416, Chengdu 610041, P. R. China E-mail: licy@cib.ac.cn
article id 477, category Research article
Pete Bettinger, Jianping Zhu. (2006). A new heuristic method for solving spatially constrained forest planning problems based on mitigation of infeasibilities radiating outward from a forced choice. Silva Fennica vol. 40 no. 2 article id 477. https://doi.org/10.14214/sf.477
Keywords: forest management; integer decision variables; integer programming
Abstract | View details | Full text in PDF | Author Info
A new heuristic method to mitigate infeasibilities when a choice is forced into a solution was developed to solve spatially constrained forest planning problems. One unique aspect of the heuristic is the introduction of unchosen decision choices into a solution regardless of the resulting infeasibilities, which are then mitigated by selecting next-best choices for those spatial units that are affected, but in a radiating manner away from the initial choice. As subsequent changes are made to correct the affected spatial units, more infeasibilities may occur, and these are corrected as well in an outward manner from the initial choice. A single iteration of the model may involve a number of changes to the status of the decision variables, making this an n-opt heuristic process. The second unique aspect of the search process is the periodic reversion of the search to a saved (in computer memory) best solution. Tests have shown that the reversion is needed to ensure better solutions are located. This new heuristic produced solutions to spatial problems that are of equal or comparable in quality to traditional integer programming solutions, and solutions that are better than those produced by two other basic heuristics. Three small hypothetical forest examples illustrate the performance of the heuristic against standard versions of threshold accepting and tabu search. In each of the three examples, the variation in solutions generated from random starting points is smaller with the new heuristic, and the difference in solution values between the new heuristic and the other two heuristics is significant (p<0.05) when using an analysis of variance. However, what remains to be seen is whether the new method can be applied successfully to the broader range of operations research problems in forestry and other fields.
  • Bettinger, Warnell School of Forest Resources, University of Georgia, Athens, GA 30602, USA E-mail: pbettinger@forestry.uga.edu (email)
  • Zhu, Warnell School of Forest Resources, University of Georgia, Athens, GA 30602, USA E-mail: jz@nn.us
article id 347, category Research article
Gordon M. Hickey, John L. Innes. (2006). Monitoring and information reporting through regulation: an inter-jurisdictional comparison of forestry-related hard laws. Silva Fennica vol. 40 no. 2 article id 347. https://doi.org/10.14214/sf.347
Keywords: forest legislation; grounded theory; constant comparison; Europe; North America
Abstract | View details | Full text in PDF | Author Info
In most jurisdictions, the rule of law has been the core instrument used to implement rules, regulations and restrictions relating to forests. The results of this approach have relied on the effectiveness of the system for regulating through monitoring and reporting. Despite the obvious differences in the wider operating environment of forestry internationally, issues related to globalization have increased the need for comparison. The potential impact of certain social, economic and environmental differences on the nature of monitoring and information reporting is, therefore, important to forest policy and management. The analysis presented here considered data associated with forestry-related monitoring and information reporting to provide a comparative description of certain hard-law requirements in a sample of jurisdictions. This was done to shed light on the potential for coordinated monitoring and information reporting objectives to be mandated through inter-jurisdictional hard law. Our research suggests that further comparative analysis of hard law monitoring and information reporting requirements could form a central theme in defining the ‘ground rules’ of a global forest law.
  • Hickey, University of British Columbia, Faculty of Forestry, Department of Forest Resources Management, 2045, 2424 Main Mall, UBC, Vancouver, Canada V6T 1Z4 E-mail: ghickey@interchange.ubc.ca (email)
  • Innes, University of British Columbia, Faculty of Forestry, Department of Forest Resources Management, 2045, 2424 Main Mall, UBC, Vancouver, Canada V6T 1Z4 E-mail: jli@nn.ca
article id 346, category Research article
Tuomo Nurminen, Heikki Korpunen, Jori Uusitalo. (2006). Time consumption analysis of the mechanized cut-to-length harvesting system. Silva Fennica vol. 40 no. 2 article id 346. https://doi.org/10.14214/sf.346
Keywords: cutting; time study; forest haulage; single-grip harvester; forwarder; work phase
Abstract | View details | Full text in PDF | Author Info
The time consumption and productivity of harvesting are dependent on stand conditions, the operators’ skills, working techniques and the characteristics of the forestry machinery. Even if the basic methods and machine types of the cut-to-length harvesting system have not changed significantly in 10 to 15 years, improvements in the operators’ competence, technical solutions in forest machinery and changes in the working environment have undoubtedly taken place. In this study, the objective was to discover the special characteristics in the time consumption of mechanized cutting and forest haulage in Finnish conditions. The empirical time study was conducted with professional operators and medium-sized single-grip harvesters and forwarders in final fellings and thinnings in easy terrain in central Finland. The models for effective time consumption in the work phases and total productivity were formed. Stem size, tree species and bucking affected the cutting, whereas timber density on the strip road, the average driving distance, load capacity, wood assortment and the bunching result of the harvester operator had an effect on the forest haulage performance. The results may be used in simulations, cost calculations and education.
  • Nurminen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: tuomo.nurminen@joensuu.fi (email)
  • Korpunen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: hk@nn.fi
  • Uusitalo, Finnish Forest Research Institute, Parkano Research Unit, Kaironiementie 54, FI-39700 Parkano, Finland E-mail: ju@nn.fi
article id 345, category Research article
Pekka Hyvönen, Perttu Anttila. (2006). Change detection in boreal forests using bi-temporal aerial photographs. Silva Fennica vol. 40 no. 2 article id 345. https://doi.org/10.14214/sf.345
Keywords: forest inventory; discriminant analysis; change detection; aerial photography; continuous updating
Abstract | View details | Full text in PDF | Author Info
Increased need for timely forest information is leading to continuous updating of stand databases. In continuous updating, stand attributes are estimated in the field after an operation and stored in databases. To find the changes caused by operations and forest damage, a semi-automatic method based on bi-temporal aerial photographs was developed. The test data were classified into three classes: No-change (952 stands), Moderate-change (163 stands) and Considerable-change (44 stands). The aerial photographs were acquired in years 2001 and 2004 with almost the same image specifications. Altogether 110 features at stand level were extracted and used in change detection analysis. The test data were classified with stepwise discriminant analysis. The overall accuracy of classification varied between 75.3 and 84.7%. The considerable changes were found without error, whereas the Moderate-change and No-change classes were often confused. However, 84.2% of thinned stands were classified correctly. The best accuracy in classification was obtained by using the histogram and textural features extracted from the original, uncorrected images. Radiometric correction did not improve the accuracy of classification. Soil type, characteristics of the growing stock and the location of a stand in an image were found to affect the change detection. Before the method can be applied operationally, issues related to, e.g., confusion between No-change and Moderate-change must be solved.
  • Hyvönen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: pekka.hyvonen@metla.fi (email)
  • Anttila, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: pa@nn.fi
article id 344, category Research article
Juha Lappi. (2006). Smooth height/age curves from stem analysis with linear programming. Silva Fennica vol. 40 no. 2 article id 344. https://doi.org/10.14214/sf.344
Keywords: height growth; spline; ring counts; second order difference; Issa
Abstract | View details | Full text in PDF | Author Info
Stem analysis data defines a range of possible heights for each age. A smooth stem/age curve is obtained with linear programming (LP) when the sum of the absolute second differences of heights is minimized subject to constraints obtained from the stem analysis. The method is analogous to cubic splines. A LP problem can include additional constraints that are based on the assumption that the crosscut is randomly located within the annual height increment. The method produces smoother height curves than Issa method which is utilizing second order differences of ring counts. It was found using simulated data that the method provides better results than earlier methods for short bolts if height growth is sufficiently regular.
  • Lappi, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: juha.lappi@metla.fi (email)
article id 343, category Research article
Karin Vestlund, Tomas Nordfjell, Lars Eliasson, Anders Karlsson. (2006). A decision support system for selective cleaning. Silva Fennica vol. 40 no. 2 article id 343. https://doi.org/10.14214/sf.343
Keywords: forestry; pre-commercial thinning; automation; practical cleaning
Abstract | View details | Full text in PDF | Author Info
Cleaning (pre-commercial thinning) costs have increased relative to logging and regeneration costs, creating a desire for rationalisation. Cleaning with robots may be a solution, but automating stem selections requires a decision support system (DSS) capable of rendering acceptable results. The aims were to develop a DSS for automation of individual stem selections in practical cleaning, and to test, using simulations, if it renders acceptable results. Data on 17 young forest stands were used to develop a DSS that selects stems by species, position (including distance and density parameters), diameter, and damage. Six simulations were run, following the DSS, with different target settings for density, percentage of deciduous stems and minimum distance between stems. The results depend on the initial state of the stands, but generally met the requested targets in an acceptable way. On average, the density results deviated by –20% to +6% from the target values, the amount of deciduous stems shifted towards the target values, and the proportion of stems with defined damaged decreased from initially 14–90% to 4–13%. The mean diameter at breast height increased and the minimum allowed distance between stems was never violated. The simulation results indicate that the DSS is operational. However, for implementation in robotics a crucial problem is to automatically perceive the selected attributes, so additional simulations with erroneous data were run. Correct measurements of diameters are less crucial than to find the majority of the trees and the majority of trees with damages.
  • Vestlund, Swedish University of Agricultural Sciences, Department of Silviculture, P.O. Box 7060, SE-750 07 Uppsala, Sweden E-mail: karin.vestlund@ssko.slu.se (email)
  • Nordfjell, Swedish University of Agricultural Sciences, Department of Forest Resource Management and Geomatics, SE-901 83 Umeå, Sweden E-mail: tn@nn.se
  • Eliasson, Swedish University of Agricultural Sciences, Department of Silviculture, SE-901 83 Umeå, Sweden E-mail: le@nn.se
  • Karlsson, Swedish University of Agricultural Sciences, Department of Silviculture, SE-901 83 Umeå, Sweden E-mail: ak@nn.se
article id 342, category Research article
Jaana Luoranen, Juha Lappi, Gang Zhang, Heikki Smolander. (2006). Field performance of hybrid aspen clones planted in summer. Silva Fennica vol. 40 no. 2 article id 342. https://doi.org/10.14214/sf.342
Keywords: survival; Populus; growth; hybrid aspen; micropropagation; planting date; root cutting; root egress; survivalgrowth
Abstract | View details | Full text in PDF | Author Info
We investigated the possibility to plant clonal hybrid aspen (Populus tremula x tremuloides) during the summer of propagation when the plants are 20–25 cm tall and only a few months old. In four experiments carried out in years 1998–2001, survival of summer-planted hybrid aspens was at least as high as that of hybrid aspen planted in autumn and spring. In all experiments, compared to planting in September or the following May, height growth was greater with planting in July and early August. Root egress of hybrid aspens planted in July and August was also greater than that of aspens planted in autumn or the following spring. Summer planting was thus possible both with plants produced by micropropagation and with those produced from root cuttings.
  • Luoranen, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: jaana.luoranen@metla.fi (email)
  • Lappi, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: lj@nn.fi
  • Zhang, College of Horticulture, Agricultural University of Hebei, Baoding, Hebei, China E-mail: gz@nn.cn
  • Smolander, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: hs@nn.fi
article id 341, category Research article
Sanni Raiskila, Pekka Saranpää, Kurt Fagerstedt, Tapio Laakso, Mia Löija, Riitta Mahlberg, Leena Paajanen, Anne-Christine Ritschkoff. (2006). Growth rate and wood properties of Norway spruce cutting clones on different sites. Silva Fennica vol. 40 no. 2 article id 341. https://doi.org/10.14214/sf.341
Keywords: Norway spruce; density; growth rate; mechanical strength; modulus of elasticity; modulus of rupture
Abstract | View details | Full text in PDF | Author Info
The effect of growth rate on weight density and strength properties of three Norway spruce cutting clones growing on three different sites in different geographic locations was studied. The purpose was to follow variation in wood physical and mechanical properties and in quality between fast-growing clones grown in environments differing in nutritional and soil properties and climate within the boreal zone. The cloned trees had been selected on grounds of good growth, health and quality. The cuttings were collected from three-year-old seedlings and rooted. The rooted cuttings were planted in the 1970’s and they were on average 26 years old at a time of felling. The variation of weight density was studied within the annual ring and within the stem between the juvenile and mature wood from the pith to the bark with an X-ray densitometric method. The average annual ring width (and latewood proportion, %) varied between the clones from 2.92±1.36 mm (15.34%) to 3.30±1.25 mm (11.80%) and between the sites from 2.76±1.07 mm (14.71%) to 3.70±1.22 mm (13.29%). The mean weight density was 0.461±0.077 g cm–3 and latewood density 0.750±0.125 g cm–3 in this material. The mean modulus of elasticity was 9.88±1.43 GPa, modulus of rupture 67.51±11.50 MPa and weight density of the test samples (ρ12) 414±44 kg m–3 in mature wood. The parameters studied showed clearly that the environment had a large effect while the three clones differed from each other similarly in the different sites, e.g. the fastest growing clone was fastest on all sites.
  • Raiskila, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: sr@nn.fi
  • Saranpää, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: pekka.saranpaa@metla.fi (email)
  • Fagerstedt, Department of Biological and Environmental Sciences, Plant Biology, P.O. Box 65, FI-00014 University of Helsinki, Finland E-mail: kf@nn.fi
  • Laakso, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: tl@nn.fi
  • Löija, VTT Building and Transport, P.O. Box 1806, FI-02044 VTT, Finland E-mail: lm@nn.fi
  • Mahlberg, VTT Building and Transport, P.O. Box 1806, FI-02044 VTT, Finland E-mail: rm@nn.fi
  • Paajanen, VTT Building and Transport, P.O. Box 1806, FI-02044 VTT, Finland E-mail: lp@nn.fi
  • Ritschkoff, VTT Building and Transport, P.O. Box 1806, FI-02044 VTT, Finland E-mail: acr@nn.fi
article id 340, category Research article
Petteri Muukkonen, Raisa Mäkipää, Raija Laiho, Kari Minkkinen, Harri Vasander, Leena Finér. (2006). Relationship between biomass and percentage cover in understorey vegetation of boreal coniferous forests. Silva Fennica vol. 40 no. 2 article id 340. https://doi.org/10.14214/sf.340
Keywords: upland soils; peatlands; biomass models; ground vegetation
Abstract | View details | Full text in PDF | Author Info
In the present study, the aboveground biomass of the understorey vegetation of boreal coniferous forests was modelled according to the percentage cover. A total of 224 observations from 22 stands in upland forests and 195 observations from 14 different studies in peatland forests were utilized for the present analyses. The relationships between biomass and percentage cover can be used in ecosystem and carbon-cycle modelling as a rapid nondestructive method for estimation of the aboveground biomass of lichens, bryophytes, herbs and grasses, and dwarf shrubs in upland forests and bottom and field layers in peatland forests.
  • Muukkonen, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: petteri.muukkonen@metla.fi (email)
  • Mäkipää, Finnish Forest Research Institute, Unioninkatu 40 A, FI-00170 Helsinki, Finland E-mail: rm@nn.fi
  • Laiho, Department of Forest Ecology, P.O. Box 24, FI-00014 University of Helsinki, Finland E-mail: rl@nn.fi
  • Minkkinen, Department of Forest Ecology, P.O. Box 24, FI-00014 University of Helsinki, Finland E-mail: km@nn.fi
  • Vasander, Department of Forest Ecology, P.O. Box 24, FI-00014 University of Helsinki, Finland E-mail: hv@nn.fi
  • Finér, Finnish Forest Research Institute, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: lf@nn.fi
article id 339, category Research article
John W. McCarthy, Gordon Weetman. (2006). Age and size structure of gap-dynamic, old-growth boreal forest stands in Newfoundland. Silva Fennica vol. 40 no. 2 article id 339. https://doi.org/10.14214/sf.339
Keywords: boreal forests; old-growth forests; gap dynamics; age and size structure; rotated sigmoid; Newfoundland
Abstract | View details | Full text in PDF | Author Info
The age and size structure of trees in old Abies-Picea-Betula forests on Newfoundland’s Great Northern Peninsula were examined. It was hypothesized that the size and age structure of both the tree and regeneration “strata” of these stands display the complex structural heterogeneity characteristic of classic, self-regenerating, uneven-aged old-growth stands, and that the development and dynamics of such structures occur over long periods of time. With all tree species combined, dbh (diameter at breast height) and height distributions exhibited a strong reverse-J character, with well-defined, semi-logarithmic rotated sigmoid height and size frequencies. Seedling height and basal diameter frequency distributions were reverse-J in character. Live tree ages for all species, except white birch (Betula papyrifera Marsh), ranged from 25 to 269 years, and were characterized by all-age frequency distributions. Tree age and size were poorly correlated. On average, balsam fir (Abies balsamea (L.) Mill.) required 62 years to reach breast height (1.3 m), with black spruce (Picea mariana (Mill.) B.S.P.) and white spruce (Picea glauca (Moench) Voss) requiring 40 and 48 years, respectively. Total age of dead standing trees ranged from 45 to 232 years. Reverse-J age frequencies characterized the seedling bank, with balsam fir seedlings present in nearly all age classes up to 110, 120 and 85 years in three sample stands. Seedling size (height and basal diameter)-age relationships were characteristic of decades-long suppression. The combination of tree and seedling bank size and age structure provide strong evidence of quasi-equilibrium, small-scale, gap dynamic old-growth boreal forest stands.
  • McCarthy, University of British Columbia, Forest Sciences Department, 2424 Main Mall, Vancouver, B.C., Canada V6T 1Z4 E-mail: jmccarthy@jesuits.ca (email)
  • Weetman, University of British Columbia, Forest Sciences Department, 2424 Main Mall, Vancouver, B.C., Canada V6T 1Z4 E-mail: gw@n.ca
article id 338, category Research article
Andrea Vajda, Ari Venäläinen, Pekka Hänninen, Raimo Sutinen. (2006). Effect of vegetation on snow cover at the northern timberline: a case study in Finnish Lapland. Silva Fennica vol. 40 no. 2 article id 338. https://doi.org/10.14214/sf.338
Keywords: climate; snow; forest-tundra; Lapland; radar; tree-line; wind modelling
Abstract | View details | Full text in PDF | Author Info
The presence of permanent snow cover for 200–220 days of the year has a determining role in the energy, hydrological and ecological processes at the climate-driven spruce (Picea abies) timberline in Lapland. Disturbances, such as forest fires or forest harvesting change the vegetation pattern and influence the spatial variation of snow cover. This variability in altered snow conditions (in subarctic Fennoscandia) is still poorly understood. We studied the influence of vegetation on the small-scale spatial variation of snow cover and wind climate in the Tuntsa area that was disturbed by a widespread forest fire in 1960. Radar was applied to measure snow thickness over two vegetation types, the spruce-dominant fire refuge and post-fire treeless tundra. Wind modelling was used to estimate the spatial variation of wind speed and direction. Due to the altered surface roughness and the increased wind velocity, snow drifting was more vigorous on the open tundra, resulting in a 30-cm thinner snow cover and almost half the water equivalent compared to the forest values. The changes in local climate after the fire, particularly in snow cover, may have played an important role in the poor recovery of vegetation: a substantial area is still unforested 40 years after the fire.
  • Vajda, Finnish Meteorogical Institute, Climate and Global Change, P.O. Box 503, FI-00101 Helsinki, Finland E-mail: andrea.vajda@fmi.fi (email)
  • Venäläinen, Finnish Meteorogical Institute, Climate and Global Change, P.O. Box 503, FI-00101 Helsinki, Finland E-mail: av@nn.fi
  • Hänninen, Geological Survey of Finland, P.O. Box 96, FI-02151 Espoo, Finland E-mail: ph@nn.fi
  • Sutinen, Geological Survey of Finland, P.O. Box 77, FI-96101 Rovaniemi, Finland E-mail: rs@nn.fi
article id 337, category Research article
Hannu Salminen, Risto Jalkanen. (2006). Modelling variation of needle density of Scots pine at high latitudes. Silva Fennica vol. 40 no. 2 article id 337. https://doi.org/10.14214/sf.337
Keywords: Pinus sylvestris; needles; dendroclimatology; NTM; precipitation; temperature; fascicles
Abstract | View details | Full text in PDF | Author Info
The relationship between apical extension and needle density and the effect of temperature and precipitation on needle density was modelled using data gathered from forty-nine felled sample trees in five stands of Scots pine (Pinus sylvestris L.) located along a latitudinal transect from the Arctic Circle up to the northern timberline. The lengths were measured and needle densities assessed from all annual shoots located above 1.3 metres using the Needle Trace Method (NTM), resulting, on average, in 39-year-long chronologies. The mean overall needle density was 7.8 short shoots per shoot centimetre. Needle-density variation in the measured data was mostly due to within-tree differences. Of the total variance, within-tree variation yielded 46%, between-tree 21%, and between-year 27%. The dependence of needle density on annual height growth was studied by fitting a multilevel model with random stand-, tree- and year-intercepts, the independent variables being tree age and height growth. There was a very strong negative correlation between height growth and needle density, and the proportion of between-year variance explained solely by height growth and age was 50%. The stand-wise residual variations and their correlations with the temperature and precipitation time series were further analysed with cross-correlation analysis in order to screen for additional independent variables. The only possible additional independent variable found was the precipitation of April–May (precipitation of May in the two northernmost stands). When it was added to the multi-level model, the proportion of explained between-year needle-density variance was 55%, but the overall fit of the model improved only slightly. The effect of late winter and early spring precipitation indicates the role of snow coverage and snowmelt on the growing conditions in the three southernmost stands. In general, stand-level needle-density variation is mostly due to changes in height growth.
  • Salminen, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: hannu.salminen@metla.fi (email)
  • Jalkanen, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: rj@nn.fi
article id 359, category Research article
Paula Horne. (2006). Forest owners’ acceptance of incentive based policy instruments in forest biodiversity conservation – a choice experiment based approach. Silva Fennica vol. 40 no. 1 article id 359. https://doi.org/10.14214/sf.359
Keywords: forest owners; heterogeneity; incentive based policy mechanism; choice experiment method
Abstract | View details | Full text in PDF | Author Info
Finland has launched a new policy programme (METSO) to enhance conservation of forest biodiversity. In non-industrial private forests, the policy is based on economic incentives and voluntarism on the part of forest owners. While biodiversity conservation is the main target of the policy, social acceptability is considered to be of great importance. This study examined the factors that affect the acceptability of biodiversity conservation contracts among private forest owners, and the amount of compensation needed to keep the forest owners at least as well off as before the contract. Choice experiment method was used to analyse the data that were collected by surveying 3000 Finnish private forest owners. Analysing separately those respondents who were willing to enter into a conservation contract allowed an assessment of the impact of forest owners’ heterogeneity on compensation amount. The results show how the welfare of forest owners shifts when the contract terms are changed. In a base scenario the forest owner was assumed to be the initiator of the contract that would require only small patches of forest to be protected, and would also bind new forest owners over its duration of ten years. For all respondents, the average demand for compensation would be around 224 euros annually. When those always choosing the “no additional conservation” alternative were excluded, the average welfare impact of the base scenario was positive.
  • Horne, Finnish Forest Research Institute, Unioninkatu 40 A, FI-00170 Helsinki, Finland E-mail: paula.horne@metla.fi (email)
article id 358, category Research article
Turgay Akbulut, Nadir Ayrilmis. (2006). Effect of compression wood on surface roughness and surface absorption of medium density fiberboard. Silva Fennica vol. 40 no. 1 article id 358. https://doi.org/10.14214/sf.358
Keywords: compression wood; medium density fiberboard; surface properties; surface analysis
Abstract | View details | Full text in PDF | Author Info
Compression wood is undoubtedly one of the most important raw material variables in wood based panel manufacturing. This study evaluated effect of compression wood on surface roughness and surface absorption (flow distance) of medium density fiberboards (MDF) manufactured from furnishes of pine (Pinus nigra Arnold var. pallasiana) containing compression wood. Panels were manufactured from two different portions of the furnish, one of the portions having a compression wood/normal wood ratio of 75/25, and the other having a ratio of 10/90. Surface absorption and surface roughness were determined according to (EN 382-1) and (ISO 4287), respectively. It was found that panels made from furnish with a 75/25 ratio had a significantly lower surface absorption value (255.78 mm) than panels made from furnish with a 10/90 ratio (317.95 mm). Surface roughness measurements based on three roughness parameters, average roughness (Ra), mean peak-to-valley height (Rz), and maximum peak-to-valley height (Ry) were considered to evaluate the surface characteristics of the panels and supported the above findings as the panels made from furnish with a 75/25 ratio had slightly rougher surface with average values of 4.15 µm (Ra). From the tests performed, we conclude that increasing of the compression wood portion increased the surface roughness and decreased the surface absorption value.
  • Akbulut, Istanbul University, Faculty of Forestry, Bahcekoy, TR-34473 Istanbul, Turkey E-mail: ta@nn.tr
  • Ayrilmis, Istanbul University, Faculty of Forestry, Bahcekoy, TR-34473 Istanbul, Turkey E-mail: nadiray@istanbul.edu.tr (email)
article id 357, category Research article
Elizabeth Dodson Coulter, John Sessions, Michael G. Wing. (2006). Scheduling forest road maintenance using the analytic hierarchy process and heuristics. Silva Fennica vol. 40 no. 1 article id 357. https://doi.org/10.14214/sf.357
Keywords: decision support; simulated annealing; threshold accepting; road environmental impacts; AHP
Abstract | View details | Full text in PDF | Author Info
The management of low-volume roads has transitioned from focusing on maintenance designed to protect a capital investment in road infrastructure to also include environmental effects. In this study, two models using mathematical programming are applied to schedule forest road maintenance and upgrade activities involving non-monetary benefits. Model I uses a linear objective function formulation that maximizes benefit subject to budgetary constraints. Model II uses a non-linear objective function to maximize the sum of benefits divided by the sum of all costs in a period. Because of the non-linearity of the constraints and the requirements that the decision variables be binary, the solutions to both problem formulations are found using two heuristics, simulated annealing and threshold accepting. Simulated annealing was found to produce superior solutions as compared to threshold accepting. The potential benefit for completing a given road maintenance or upgrade project is determined using the Analytic Hierarchy Process (AHP), a multi-criterion decision analysis technique. This measure of benefit is combined with the economic cost of completing a given project to schedule maintenance and upgrade activities for 225 km (140 miles) of road in forested road systems within western Oregon. This combination of heuristics, cost-benefit analysis, environmental impacts, and expert judgment produces a road management schedule that better fits the current road management paradigm.
  • Coulter, College of Forestry and Conservation, University of Montana, 32 Campus Drive, Missoula, MT 59812, USA E-mail: elizabeth.coulter@cfc.umt.edu (email)
  • Sessions, Department of Forest Engineering, College of Forestry, Oregon State University, 204 Peavy Hall, Corvallis, OR 97331-5706, USA E-mail: js@nn.us
  • Wing, Department of Forest Engineering, College of Forestry, Oregon State University, 204 Peavy Hall, Corvallis, OR 97331-5706, USA E-mail: mgw@nn.us
article id 356, category Research article
Hamish D. Marshall, Glen Murphy, Kevin Boston. (2006). Three mathematical models for bucking-to-order. Silva Fennica vol. 40 no. 1 article id 356. https://doi.org/10.14214/sf.356
Keywords: mechanical harvesting/processing; optimal bucking; mixed integer programming; dynamic programming; buck-to-value
Abstract | View details | Full text in PDF | Author Info
The aim of this paper is to investigate different mathematical approaches to buck-to-order log merchandizing. A new bucking-to-order planning model using mixed integer programming was developed to determine the optimal production from a stand given different market constraints and forest inventory data. Three different approaches: market prices, target cutting patterns and adjusted price list were tested for generating cutting instructions to fulfill the plan created by the new planning model. The three approaches were evaluated in four test stands. The market prices approach simply applied the market prices to each stand. The target cutting patterns approach applied the sample cutting patterns generated from the planning model to the stand. The adjusted price list used a dynamic programming algorithm embedded in a search heuristic to adjust both the prices and small end diameters of log products to achieve the production goals of the planning models. The results showed that developing a buck-to-order plan is important in obtaining good order fulfillment. The target cutting patterns and adjusted price list approaches certainly out performed the market prices approach. This paper shows that these two approaches are capable of achieving excellent order fulfillment. Further development and testing is needed to determine which method is the best at generating cutting instructions for buck-to-order merchandizing.
  • Marshall, Ensis Forests, Private Bag 3020, Rotorua, New Zealand E-mail: hamish.marshall@ensisjv.com (email)
  • Murphy, Forest Engineering Department, Oregon State University, Corvallis, Oregon 97331, USA E-mail: gm@nn.us
  • Boston, Forest Engineering Department, Oregon State University, Corvallis, Oregon 97331, USA E-mail: kb@nn.us
article id 355, category Research article
Ilkka Korpela. (2006). Geometrically accurate time series of archived aerial images and airborne lidar data in a forest environment. Silva Fennica vol. 40 no. 1 article id 355. https://doi.org/10.14214/sf.355
Keywords: vegetation; canopy; monitoring; laser scanning; change detection; photogrammetry; 3D; aerial triangulation; direct georeferencing
Abstract | View details | Full text in PDF | Author Info
Reconstructing three-dimensional structural changes in the forest over time is possible using archived aerial photographs and photogrammetric techniques, which have recently been introduced to a larger audience with the advent of digital photogrammetry. This paper explores the feasibility of constructing an accurate time-series of archived aerial photographs spanning 42 years using different types of geometric data and estimation methods for image orientation. A recent airborne laser scanning (lidar) data set was combined with the image block and assessed for geometric match. The results suggest that it is possible to establish the multitemporal geometry of an image block to an accuracy that is better than 0.5 m in 3D and constant over time. Even geodetic ground control points can be omitted from the estimation if the most recent images have accurate direct sensor orientation, which is becoming a standard technique in aerial photography. This greatly reduces the costs and facilitates the work. An accurate multitemporal image block combined with recent lidar scanning for the estimation of topography allows accurate monitoring and retrospective analysis of forest vegetation and management operations.
  • Korpela, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ilkka.korpela@helsinki.fi (email)
article id 354, category Research article
Mervi Talvitie, Olli Leino, Markus Holopainen. (2006). Inventory of sparse forest populations using adaptive cluster sampling. Silva Fennica vol. 40 no. 1 article id 354. https://doi.org/10.14214/sf.354
Keywords: coarse woody debris; adaptive cluster sampling; simple random sampling; drought damage; efficiency
Abstract | View details | Full text in PDF | Author Info
In many studies, adaptive cluster sampling (ACS) proved to be a powerful tool for assessing rare clustered populations that are difficult to estimate by means of conventional sampling methods. During 2002 and 2003, severe drought-caused damage was observed in the park forests of the City of Helsinki, Finland, especially in barren site pine and spruce stands. The aim of the present study was to examine sampling and measurement methods for assessing drought damage by analysing the effectiveness of ACS compared with simple random sampling (SRS). Horvitz-Thompson and Hansen-Hurwitz estimators of the ACS method were used for estimating the population mean and variance of the variable of interest. ACS was considerably more effective than SRS in assessing rare clustered populations such as those resulting from drought damage. The variances in the ACS methods were significantly smaller and the inventory efficiency in the field better than in SRS.
  • Talvitie, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: mervi.talvitie@helsinki.fi (email)
  • Leino, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ol@nn.fi
  • Holopainen, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ah@nn.fi
article id 353, category Research article
Jaakko Repola, Hannu Hökkä, Timo Penttilä. (2006). Thinning intensity and growth of mixed spruce-birch stands on drained peatlands in Finland. Silva Fennica vol. 40 no. 1 article id 353. https://doi.org/10.14214/sf.353
Keywords: Picea abies; Betula pubescens; thinning; forest drainage; growth; spruce swamps
Abstract | View details | Full text in PDF | Author Info
The impacts of thinning at various intensities on the growth and mortality of mixed spruce-birch stands were investigated in thinning experiments on spruce swamps in northern and south-eastern Finland. At the time of establishment, three of the stands had recently reached the first commercial thinning stage and four were more advanced. The monitoring period was mainly 15 years, and the thinning intensity varied from heavy thinning (ca. 46 per cent of the basal area removed) to no thinning. Basal area removals of light and moderate thinning were ca. 22% and 39%, respectively. Unthinned plots had the highest volume increment. Light and moderate thinning slightly decreased the 15-year volume increment by, on an average, 1% and 8%, respectively. Heavy thinning led to a greater reduction (22%) in volume increment. The growth response to thinning intensity was evident as a higher relative volume and mean diameter increment of the living trees with decreasing stand density. Part of the volume increment on the unthinned plots was lost through natural mortality. Even light thinning significantly decreased natural mortality.
  • Repola, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: jaakko.repola@metla.fi (email)
  • Hökkä, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: hh@nn.fi
  • Penttilä, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: tp@nn.fi
article id 352, category Research article
Juha-Pekka Hotanen, Matti Maltamo, Antti Reinikainen. (2006). Canopy stratification in peatland forests in Finland. Silva Fennica vol. 40 no. 1 article id 352. https://doi.org/10.14214/sf.352
Keywords: succession; structural diversity; drainage; site type; canopy layer; crown coverage
Abstract | View details | Full text in PDF | Author Info
  • Hotanen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: juha-pekka.hotanen@metla.fi (email)
  • Maltamo, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: mm@nn.fi
  • Reinikainen, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: ar@nn.fi
article id 351, category Research article
Jiaojun Zhu, Xiufen Li, Zugen Liu, Wei Cao, Yutaka Gonda, Takeshi Matsuzaki. (2006). Factors affecting the snow and wind induced damage of a montane secondary forest in northeastern China. Silva Fennica vol. 40 no. 1 article id 351. https://doi.org/10.14214/sf.351
Keywords: stem breakage; uprooting; stand structure; snow and wind damages; secondary forests; mixed forests
Abstract | View details | Full text in PDF | Author Info
In order to understand the processes of snow and wind induced damage in a natural montane, secondary forest in northeastern China, we examined the impacts of site conditions on the snow and wind damage; analyzed if the dominant tree species differed in their susceptibilities to the damage; and established the relationships between the characteristics of tree and stand and the damage. The results indicated that in regard to the topography factors, slope steepness and soil depth played a relatively important role for the damage. Damage ratios of all types combined were positively related with the composition of dominant tree species. The stand density was also important in determining resistance to the damage, i.e., the densely populated stand exhibited less overall damage ratios; however, the dominant tree species were commonly damaged easily by the snow and wind. Four damage modes found (uprooting, stem breakage, canopy damage and bending) were closely related to the stem taper (p < 0.05), and they could be ranked in following order: bending (92.0 ) > uprooting (85.3) > stem breakage (80.1) > canopy damage (65.0). In regard to differences in tree species’ susceptibilities to the damage, Betula costata exhibited the most uprooting, bending and overall damage ratios; while Quercus mongolica showed the highest breakage (both stem breakage and canopy damage) ratio, and Fraxinus mandshurica exhibited the least damage ratio (overall). The major six tree species could also be divided into two groups according to the overall damage ratios, i.e., more susceptible ones (B. costata, Ulmus laciniata and Q. mongolica), and less susceptible ones (F. mandshurica, Acer mono and Juglans mandshurica) to the snow and wind damage.
  • Zhu, Institute of Applied Ecology, Chinese Academy of Sciences, Wenhua Road 72, Shenyang 110016, China E-mail: zrms29@yahoo.com (email)
  • Li, Institute of Applied Ecology, Chinese Academy of Sciences, Wenhua Road 72, Shenyang 110016, China; Graduate School of Chinese Academy of Sciences, Yuquan Road 19-A, Beijing, 100039, China E-mail: xfl@nn.cn
  • Liu, Institute of Applied Ecology, Chinese Academy of Sciences, Wenhua Road 72, Shenyang 110016, China; Graduate School of Chinese Academy of Sciences, Yuquan Road 19-A, Beijing, 100039, China E-mail: zgl@nn.cn
  • Cao, Institute of Applied Ecology, Chinese Academy of Sciences, Wenhua Road 72, Shenyang 110016, China E-mail: wc@nn.cn
  • Gonda, Faculty of Agriculture, Niigata University, Ikarashi 2-8050, Niigata, 950-2181, Japan E-mail: yg@nn.jp
  • Matsuzaki, Faculty of Agriculture, Niigata University, Ikarashi 2-8050, Niigata, 950-2181, Japan E-mail: tm@nn.jp
article id 350, category Research article
Ehsan Sayyad, Seyed Mohsen Hosseini, Jamshid Mokhtari, Reza Mahdavi, Seyed Gholamali Jalali, Moslem Akbarinia, Masoud Tabari. (2006). Comparison of growth, nutrition and soil properties of pure and mixed stands of Populus deltoides and Alnus subcordata. Silva Fennica vol. 40 no. 1 article id 350. https://doi.org/10.14214/sf.350
Keywords: growth; nitrogen fixing trees (NFT); mixed plantations; nutrition; nutrient return
Abstract | View details | Full text in PDF | Author Info
Concerns about decline in soil fertility and long-term productivity of fast-growing plantations have promoted interest in using nitrogen-fixing trees in mixed species plantations. Populus deltoides and Alnus subcordata were planted in five proportions (100P, 67P:33A, 50P:50A, 33P:67A, 100A) in Noor, Iran. After 7 years, the effects of species interactions on tree growth and nutrient concentration in live and senescent leaves and soil properties were assessed. Diameter at breast height and total height of individual Populus trees were positively affected by the presence of Alnus. Nitrogen concentrations in fully expanded and senescent leaves of Populus were higher in mixed plantations than monoculture plantations. The results of nutrition and nutrient return and growth indicated that mixed plantations of these two species were more productive and sustainable than their monoculture plantations. Within the framework of this experiment, it appeared that production was maximized when these two species were grown together in the relative proportions of 50% Populus and 50% Alnus.
  • Sayyad, Tarbiat Modarres University, Natural Resources Faculty, Emam St. Noor, 46414 Noor, Mazandaran, Iran E-mail: es@nn.ir
  • Hosseini, Tarbiat Modarres University, Natural Resources Faculty, Emam St. Noor, 46414 Noor, Mazandaran, Iran E-mail: hosseini@europe.com (email)
  • Mokhtari, Tarbiat Modarres University, Natural Resources Faculty, Emam St. Noor, 46414 Noor, Mazandaran, Iran E-mail: jm@nn.ir
  • Mahdavi, Tarbiat Modarres University, Natural Resources Faculty, Emam St. Noor, 46414 Noor, Mazandaran, Iran E-mail: rm@nn.ir
  • Jalali, Tarbiat Modarres University, Natural Resources Faculty, Emam St. Noor, 46414 Noor, Mazandaran, Iran E-mail: sgj@nn.ir
  • Akbarinia, Tarbiat Modarres University, Natural Resources Faculty, Emam St. Noor, 46414 Noor, Mazandaran, Iran E-mail: ma@nn.ir
  • Tabari, Tarbiat Modarres University, Natural Resources Faculty, Emam St. Noor, 46414 Noor, Mazandaran, Iran E-mail: mt@nn.ir
article id 349, category Research article
Temel Sariyildiz, J. M. Anderson. (2006). Intra-specific variation in cell wall constituents of needle age classes of Pinus sylvestris in relation to soil fertility status in Southwest England. Silva Fennica vol. 40 no. 1 article id 349. https://doi.org/10.14214/sf.349
Keywords: Pinus sylvestris; nitrogen; soil fertility; lignin; needle age classes; phenylpropanoids; hemicellulose
Abstract | View details | Full text in PDF | Author Info
First, second and third year needles were collected from the same branches of young Scots pine trees growing on soils of two different status (high and low fertility sites) that varied in mineral nutrient concentrations and N mineralisation potential. All needle age classes were analysed for total carbon, acid detergent fibre (ADF), lignin, cellulose, phenylpropanoid derivatives (PPD) of lignin, sugar constituents of non-crystalline, hydrated cellulose and hemicellulose, and nutrient concentrations (N, P, K, Ca, Mg and Mn). Significant intra-specific variation in the litter quality variables in relation to soils of high and low fertility was found in the second and third year needles, whereas there were no differences in the other cell wall constituents and mineral elements of the first year needles. The second and third year needles from the low fertility soil contained higher concentrations of ADF, lignin, cellulose, sugar constituents of non-crystalline, hydrated cellulose and hemicellulose, and phenylpropanoid derivatives (PPD) of lignin, but lower concentrations of N, P and Mg than the same needles from the high fertility and fertilised soils. The results in the present study indicate that under different soil fertilities, needle age classes show significant variations in the cell wall constituents and mineral elements, and suggest that this can result in significant variation in litter quality and decomposition rates.
  • Sariyildiz, Kars Kafkas Üniversitesi, Artvin Orman Fakültesi, 08000 Artvin, Turkey E-mail: t_sariyildiz@yahoo.com (email)
  • Anderson, Department of Biological Sciences, Hatherly Laboratories, University of Exeter, Exeter EX4 4PS, UK E-mail: jma@nn.uk
article id 348, category Research article
Chunyang Li, Xuejiang Zhang, Xingliang Liu, Olavi Luukkanen, Frank Berninger. (2006). Leaf morphological and physiological responses of Quercus aquifolioides along an altitudinal gradient. Silva Fennica vol. 40 no. 1 article id 348. https://doi.org/10.14214/sf.348
Keywords: carbon isotope composition; leaf nitrogen content; specific leaf area; stomata
Abstract | View details | Full text in PDF | Author Info
Quercus aquifolioides Rehder & E.H. Wilson, an evergreen alpine and subalpine shrub species, occupies a wide range of habitats on the eastern slopes of the Himalaya in China. In this study, we measured leaf morphology, nitrogen content and carbon isotope composition (as an indicator of water use efficiency) of Q. aquifolioides along an altitudinal gradient. We found that these leaf morphological and physiological responses to altitudinal gradients were non-linear with increasing altitude. Specific leaf area, stomatal length and index increased with increasing altitude below 2800 m, but decreased with increasing altitude above 2800 m. In contrast, leaf nitrogen content per unit area and carbon isotope composition showed opposite change patterns. Specific leaf area seemed to be the most important parameter that determined the carbon isotope composition along the altitudinal gradient. Our results suggest that near 2800 m in altitude could be the optimum zone for growth and development of Q. aquifolioides, and highlight the importance of the influence of altitude in research on plant physiological ecology.
  • Li, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O. Box 416, Chengdu 610041, P. R. China E-mail: licy@cib.ac.cn (email)
  • Zhang, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O. Box 416, Chengdu 610041, P. R. China; Graduate School of the Chinese Academy of Sciences, Beijing 100039, P.R. China E-mail: xz@nn.cn
  • Liu, Sichuan Academy of Forestry, Chengdu 610081, P. R. China E-mail: xl@nn.cn
  • Luukkanen, Viikki Tropical Resources Institute, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ol@nn.fi
  • Berninger, Département des sciences biologiques, Cp 8888 succ centre ville, Université du Québec à Montréal, Montréal (QC) H3C 3P8, Canada E-mail: fb@nn.ca
article id 478, category Research article
Ronald E. McRoberts, Daniel G. Wendt, Greg C. Liknes. (2005). Stratified estimation of forest inventory variables using spatially summarized stratifications. Silva Fennica vol. 39 no. 4 article id 478. https://doi.org/10.14214/sf.478
Keywords: bias; precision; classified satellite imagery; Internet; variance
Abstract | View details | Full text in PDF | Author Info
Large area natural resource inventory programs typically report estimates for selected geographic areas such as states or provinces, counties, and municipalities. To increase the precision of estimates, inventory programs may use stratified estimation, with classified satellite imagery having been found to be an efficient and effective basis for stratification. For the benefit of users who desire additional analyses, the inventory programs often make data and estimation procedures available via the Internet. For their own analyses, users frequently request access to stratifications used by the inventory programs. When data analysis is via the Internet and stratifications are based on classifications of even medium resolution satellite imagery, the memory requirements for storing the stratifications and the online time for processing them may be excessive. One solution is to summarize the stratifications at coarser spatial scales, thus reducing both storage requirements and processing time. If the bias and loss of precision resulting from using summaries of stratifications is acceptably small, then this approach is viable. Methods were investigated for using summaries of stratifications that do not require storing and processing the entire pixel-level stratifications. Methods that summarized satellite image-based 30 m x 30 m pixel stratifications at spatial scales up to 2400 ha produced stratified estimates of the mean that were generally within 5-percent of estimates for the same areas obtained using the pixel stratifications. In addition, stratified estimates of variances using summarized stratifications realized nearly all the gain in precision that was obtained with the underlying pixel stratifications.
  • McRoberts, North Central Research Station, USDA Forest Service, 1992 Folwell Avenue, Saint Paul, Minnesota, USA 5510 E-mail: rmcroberts@fs.fed.us (email)
  • Wendt, Region 9, USDA Forest Service, 626 East Wisconsin Avenue, Milwaukee, Wisconsin 53202, USA E-mail: dgw@nn.us
  • Liknes, North Central Research Station, USDA Forest Service, 1992 Folwell Avenue, Saint Paul, Minnesota, USA 5510 E-mail: gcl@nn.us
article id 368, category Research article
Eero Muinonen. (2005). Generating a raster map presentation of a forest resource by solving a transportation problem. Silva Fennica vol. 39 no. 4 article id 368. https://doi.org/10.14214/sf.368
Keywords: forest inventory; calibration estimator; transportation problem; nonparametric estimation
Abstract | View details | Full text in PDF | Author Info
Necessary tools for raster map generation, for the approach based on the calibration estimator, were developed and implemented. The allocation of the area weight of each pixel to sample plots was formulated as a transportation problem, using a spectral distance measure as a transportation cost, and solved using the transportation simplex algorithm. Pixel level accuracy was calculated for the methods based on the calibration estimator so that the results could be compared with the results of the nearest neighbour estimation, the reference sample plot method (RSP) at pixel level. Local averaging in a 3 x 3 window was performed for each generated raster map as a postprocessing phase to smooth the map. Test plot results were calculated both for the unfiltered raster map and the filtered raster map. RSP produced the smallest RMSE in the pooled test data. Local averaging with a 3 x 3 filter decreased the pixel level error – and the bias – and the differences between the methods are smaller. Without local averaging, the pixel level errors of the methods based on solving the transportation problem were high. Raster map generation using the methods of this study forms an optional part – followed possibly by the classification of the pixel level results – of the whole computation task, when the area weight computation is based on the calibration estimation. For larger areas than in the present study, such as municipalities, the efficiency of the method based on the transportation model must be improved before it is a usable tool, in practice, for raster map generation. For nearest neighbour methods, the area size is not such a problem, because the inventory area is processed pixel by pixel.
  • Muinonen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: eero.muinonen@metla.fi (email)
article id 367, category Research article
Sakari Tuominen, Markus Haakana. (2005). Landsat TM imagery and high altitude aerial photographs in estimation of forest characteristics. Silva Fennica vol. 39 no. 4 article id 367. https://doi.org/10.14214/sf.367
Keywords: multi-source forest inventory; satellite imagery; high-altitude aerial photographs; image texture
Abstract | View details | Full text in PDF | Author Info
Satellite sensor data have traditionally been used in multi-source forest inventory for estimating forest characteristics. Their advantages generally are large geographic coverage and large spectral range. Another remote sensing data source for forest inventories offering a large geographic coverage is high altitude aerial photography. In high altitude aerial photographs the spectral range is very narrow but the spatial resolution is high. This allows the extraction of texture features for forest inventory purposes. In this study we utilized a Landsat 7 ETM satellite image, a photo mosaic composed of high altitude panchromatic aerial photographs, and a combination of the aforementioned in estimating forest attributes for an area covering approximately 281 000 ha in Forestry Centre Häme-Uusimaa in Southern Finland. Sample plots of 9th National Forest Inventory (NFI9) were used as field data. In the estimation, 6 Landsat 7 ETM image channels were used. For aerial photographs, 4 image channels were composed from the spectral averages and texture features. In combining both data sources, 6 Landsat channels and 3 aerial image texture channels were selected for the analysis. The accuracy of forest estimates based on the Landsat image was better than that of estimates based on high altitude aerial photographs. On the other hand, using the combination of Landsat ETM spectral features and textural features on high altitude aerial photographs improved the estimation accuracy of most forest attributes.
  • Tuominen, Finnish Forest Research Institute, Unioninkatu 40 A, FI-00170 Helsinki, Finland E-mail: sakari.tuominen@metla.fi (email)
  • Haakana, Finnish Forest Research Institute, Unioninkatu 40 A, FI-00170 Helsinki, Finland E-mail: mh@nn.fi
article id 366, category Research article
Lars Eliasson. (2005). Effects of forwarder tyre pressure on rut formation and soil compaction. Silva Fennica vol. 39 no. 4 article id 366. https://doi.org/10.14214/sf.366
Keywords: soil density; cone penetrometer; final felling; till soil
Abstract | View details | Full text in PDF | Author Info
In Swedish forestry, final felling is usually done by a harvester and a forwarder. These machines are heavy and the risk for rutting and soil compaction can be considerable under unfavourable soil conditions. The aim of this study was to evaluate effects of forwarder tyre inflation pressure on rutting and soil compaction after final felling. Three levels of forwarder tyre pressure were studied, 300, 450 and 600 kPa, after 2 and 5 machine passages. The first passage was driven with a 19.7 Mg harvester, and the second to fifth passages with a fully loaded forwarder totalling 37.8 Mg. Rut depths were not significant affected by tyre pressures but increased significantly with the number of machine passages. Soil density was significantly increased by 0.075 Mg m–3 by the harvester passage. Soil density increased significantly with increasing number of forwarder passages, and tyre pressure did not significantly influence this increase but the interaction between number of forwarder passages and tyre pressure was almost significant. Data suggest that density increases occur earlier in the 600 kPa treatment than in the other treatments. Only parts of an area harvested are trafficked in a normal harvesting operation. Outside the research area approximately 12.5 per cent of the area harvested was covered with ruts. On primary strip roads, which are heavily trafficked, soil compaction cannot be avoided by reducing the tyre pressure. On secondary strip roads, not passed more than once by the forwarder, a low forwarder tyre pressure may reduce soil compaction.
  • Eliasson, Swedish University of Agricultural Sciences, Department of Silviculture, SE-901 83 Umeå, Sweden E-mail: lars.eliasson@norraskogsagarna.se (email)
article id 365, category Research article
Raffaele Spinelli, Carla Nati, Natascia Magagnotti. (2005). Harvesting and transport of root biomass from fast-growing poplar plantations. Silva Fennica vol. 39 no. 4 article id 365. https://doi.org/10.14214/sf.365
Keywords: productivity; biomass; logistics; harvesting; root recovery; cost
Abstract | View details | Full text in PDF | Author Info
Recovery of tree root biomass can be attractive, since the stump-root system represents a substantial portion of the tree mass and its removal may prove instrumental to re-cultivation. Most available studies concern Nordic technologies, particularly suited to mature conifer stands. Unlike spruce, plantation poplar develops a deep taproot, whose extraction requires completely different methods. The aim of the study was to investigate poplar root recovery operations in plantations with time studies, and to determine the productivity and delivery costs of the operations. Seven operation systems developed to work with poplar plantations in Italian conditions were studied. Extraction and cleaning units were based on general-purpose prime movers. Under favourable conditions extraction and cleaning units achieved a very high productivity: 150 stumps per hour for the extraction unit and 170 for the cleaning unit. Delivered cost varied widely, ranging from 28 to 66 Euros Mg–1. Transportation was the most expensive single work task. It accounted for about 40% of the total recovery cost. Extraction and cleaning contributed approximately 25% each to the total cost, and loading 9%. Guidelines to recovery system improvement and efficient operation are provided.
  • Spinelli, CNR/IVALSA, Via Madonna del Piano - Palazzo F, I-50019 Sesto Fiorentino (FI), Italy E-mail: spinelli@ivalsa.cnr.it (email)
  • Nati, CNR/IVALSA, Via Madonna del Piano - Palazzo F, I-50019 Sesto Fiorentino (FI), Italy E-mail: cn@nn.it
  • Magagnotti, CNR/IVALSA, Via Madonna del Piano - Palazzo F, I-50019 Sesto Fiorentino (FI), Italy E-mail: nm@nn.it
article id 364, category Research article
Timo Pukkala, Jari Miina. (2005). Optimising the management of a heterogeneous stand. Silva Fennica vol. 39 no. 4 article id 364. https://doi.org/10.14214/sf.364
Keywords: Hooke and Jeeves algorithm; thinning schedules; within-stand variation
Abstract | View details | Full text in PDF | Author Info
The study presents a method for taking the heterogeneity of the stand into account in the optimisation of stand management. Heterogeneity refers to within-stand variation in stand density and/or other characteristics. A set of plots, corresponding to different sub-areas of the stand, represents the stand in calculations. Cuttings and other treatments of the plots are done simultaneously. The method was used to analyse how the optimal management depends on the heterogeneity of a Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) stand. The results supported the hypothesis that the heterogeneity of a stand decreases its optimal prior-thinning density. Also the remaining stand basal areas were lower in heterogeneous stands, especially in spruce. The effect of stand heterogeneity prior to the first commercial thinning still affected the timing of the second thinning, which had to be conducted earlier and at lower prior-thinning basal areas in heterogeneous stands. This happened despite the fact that the first thinning greatly decreased the within-stand variation in stand basal area. In addition, heterogeneity decreased the soil expectation value, net income and timber harvests.
  • Pukkala, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: timo.pukkala@joensuu.fi (email)
  • Miina, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jm@nn.fi
article id 363, category Research article
Karin Vestlund, Tomas Nordfjell, Lars Eliasson. (2005). Comparison of human and computer-based selective cleaning. Silva Fennica vol. 39 no. 4 article id 363. https://doi.org/10.14214/sf.363
Keywords: forestry; decision support; pre-commercial thinning; automation; practical cleaning; training-tool
Abstract | View details | Full text in PDF | Author Info
In silvicultural tending operations like cleaning (pre-commercial thinning), the results are irreversible, so it is important for the decisions to be consistent with the aims for the stand. To enable operational automatic stem selections, a decision support system (DSS) is needed. A previously presented DSS seems to render acceptable cleaning results, but needs further analysis. The aims of the study were to compare the cleaning results of experienced cleaners and DSS simulations when “similar” instructions were given, and to assess the usefulness and robustness of the DSS. Twelve experienced cleaners were engaged to “clean” (mark main stems) six areas; each cleaner “cleaned” two areas. The DSS was used to generate two computer-based cleanings (simulations) of these areas. Four laymen also “cleaned” one of the areas following the DSS. The density results were significantly affected by the areas’ location, whereas the proportions of deciduous stems and damaged stems were significantly affected by both the areas’ location and method, i.e. manual “cleaning” and general or adjusted simulation. The study showed that the DSS can be adjusted so that the results are comparable with the cleaners’ results. Thus, the DSS seems to be useful and flexible. The laymen’s results were close to the results of the “general” simulation, implying that the DSS is robust and could be used as a training tool for inexperienced cleaners. The DSS was also acceptable on a single-tree level, as more than 80% of the main-stems selected in the simulations were also selected by at least one cleaner.
  • Vestlund, Swedish University of Agricultural Sciences, Department of Silviculture, P.O. Box 7060, SE-750 07 Uppsala, Sweden E-mail: karin.vestlund@ssko.slu.se (email)
  • Nordfjell, Swedish University of Agricultural Sciences, Department of Silviculture, SE-901 83 Umeå, Sweden E-mail: tm@nn.se
  • Eliasson, Swedish University of Agricultural Sciences, Department of Silviculture, SE-901 83 Umeå, Sweden E-mail: le@nn.se
article id 362, category Research article
Hannu Salminen, Risto Jalkanen. (2005). Modelling the effect of temperature on height increment of Scots pine at high latitudes. Silva Fennica vol. 39 no. 4 article id 362. https://doi.org/10.14214/sf.362
Keywords: Pinus sylvestris; height growth; dendroclimatology; precipitation; temperature
Abstract | View details | Full text in PDF | Author Info
The effect of temperature and precipitation on the height increment of Pinus sylvestris (L.) was modelled using data gathered from a total of 49 felled sample trees from five stands of Scots pine located along a latitudinal transect from the Arctic Circle up to the northern timberline in Finland. A multilevel mixed effects model and cross-correlation analysis of prewhitened time series was used to analyse the dependence between height increment and monthly meteorological observations. The effect of the mean July temperature of the previous year on height increment proved to be very strong at high latitudes (r > 0.7). The mean November temperature of the year before the previous affected statistically significantly on height increment in the three northernmost stands. There was no correlation between height increment and precipitation in any of the sites. The final height increment model based on all stands included tree age, long-term mean temperature sum of site, and the mean July temperature of the previous year as independent variables. According to the model, one degree’s change in July temperature results on average in 1.8 cm change in the next year’s height increment. There was a modest but significant polynomial age-effect. The proportion of explained variance (at the year level) was 74%. The July temperature dependence on height increment was shown to be very strong, suggesting a high value of height increment in climate modelling at the tree line.
  • Salminen, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: hannu.salminen@metla.fi (email)
  • Jalkanen, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: rj@nn.fi
article id 361, category Research article
Jaana Luoranen, Risto Rikala, Kyösti Konttinen, Heikki Smolander. (2005). Extending the planting period of dormant and growing Norway spruce container seedlings to early summer. Silva Fennica vol. 39 no. 4 article id 361. https://doi.org/10.14214/sf.361
Keywords: Norway spruce; height growth; survival; planting date; carbohydrate; frozen storage; hot-lifting; root growth
Abstract | View details | Full text in PDF | Author Info
In order to make mechanized planting economically viable, the present spring planting period for Norway spruce (Picea abies (L.) Karst.) seedlings in Scandinavia needs to be extended. To evaluate the possibilities to extend the planting period, six field experiments were established in four years during which frozen-stored, dormant seedlings and actively growing seedlings targeted for spring planting were planted regularly from mid-May to mid-July or the end of August. The survival of actively growing seedlings did not differ between planting dates from mid-May to mid-July. For dormant seedlings, however, the later in summer they were planted the lower was the survival. Oversized seedlings grown in the nursery in containers of too small volume, which were usually planted after mid-June, resulted in reduced growth of seedlings after planting. Root egress (growth of roots from root plugs into the surrounding soil) was most rapid in July and early August and slowest in May and September. Results showed that with dormant seedlings the planting period can be extended from May to mid-June without increasing mortality or reducing growth. The planting period for seedlings stored outdoors and those seedlings that were already growing in June for the purpose of spring plantings can be extended even longer, but it must be kept in mind that the risk of mechanical damage and reduced growth increase due to brittleness of the shoot and increased height. Further research is needed to evaluate the risks in practical scale plantings and with seedlings that are specially targeted for planting after mid-June.
  • Luoranen, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: jaana.luoranen@metla.fi (email)
  • Rikala, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: rr@nn.fi
  • Konttinen, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: kk@nn.fi
  • Smolander, Finnish Forest Research Institute, Suonenjoki Research Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: hs@nn.fi
article id 360, category Research article
Timo Kurkela, Tarmo Aalto, Martti Varama, Risto Jalkanen. (2005). Defoliation by the common pine sawfly (Diprion pini) and subsequent growth reduction in Scots pine: a retrospective approach. Silva Fennica vol. 39 no. 4 article id 360. https://doi.org/10.14214/sf.360
Keywords: height growth; NTM; defoliation; insect attack; needle density; needle production; needle retention; radial growth; herbivory
Abstract | View details | Full text in PDF | Author Info
The foliage status in the main stem of Scots pines (Pinus sylvestris) was studied retrospectively using the needle trace method (NTM) on a stand, seriously defoliated by the pine sawfly (Diprion pini) in the 1980s. Needle density increased abruptly in the seasons following the defoliation. The strongest reduction in annual needle production occurred one year later. As a consequence of lower needle production, the annual number of attached needles decreased three to five years after the defoliation. Needle retention and the average age of attached needles tended to increase after defoliation. In analyses of covariance with the NTM variables, needle density (logarithmic transformed values) and average age of attached needles, had the highest, significant, negative relationship with radial and height increments both in the period prior to the defoliation and in the time when the trees were suffering from defoliation. The relationships between height increment and the number of needles and needle loss were positive and significant. Also radial increment had a positive relationship with the number of needles but not with needle loss. Interestingly, an abrupt increase in the needle density gave a good indication of the effects of a sudden defoliation in pines.
  • Kurkela, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: timo.kurkela@metla.fi (email)
  • Aalto, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: ta@nn.fi
  • Varama, Finnish Forest Research Institute, Vantaa Research Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: mv@nn.fi
  • Jalkanen, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: rj@nn.fi
article id 479, category Research article
Ken Olaf Storaunet, Jørund Rolstad, Ivar Gjerde, Vegard S. Gundersen. (2005). Historical logging, productivity, and structural characteristics of boreal coniferous forests in Norway. Silva Fennica vol. 39 no. 3 article id 479. https://doi.org/10.14214/sf.479
Keywords: boreal forest; stand structure; decaying wood; forest history; naturalness; selective logging; dead trees
Abstract | View details | Full text in PDF | Author Info
Conservation of forest biodiversity has brought about an interest in evaluating the naturalness of forests, and to locate and protect semi-natural and old-growth forests in the Fennoscandian countries. However, it is not always clear how natural these forests really are, and how the past management history has affected their present structural composition. We studied the relationships between cut stumps from historical logging activity (50–100 years ago) and forest structural characteristics of today in a total of 385 0.25 ha plots in three boreal coniferous forests which are parts of National Nature Reserves in Norway. We also studied how forest productivity influenced these relationships. In plots with negligible logging impact we found the amount of living trees, dead wood, and size of the oldest trees mainly to increase with increasing productivity, whereas the age of the oldest trees decreased. The amount of deciduous trees was generally low irrespective of productivity. The intensity of logging did not consistently influence most of these forest structural variables, neither at low- nor at high-productive sites. The only consistent relationship in all study areas was a decreasing amount of dead wood with increasing logging intensity at high-productive sites. Also, the decay class distribution of dead wood was more right-skewed (indicating on-going accumulation of dead wood) the more logging had occurred at high-productive sites. Except from the effects on dead wood, previous logging does not show up as a major determinant of other stand structures of today.
  • Storaunet, Norwegian Forest Research Institute, Høgskolevegen 8, NO-1432 Ås, Norway E-mail: ken.storaunet@skogforsk.no (email)
  • Rolstad, Norwegian Forest Research Institute, Høgskolevegen 8, NO-1432 Ås, Norway E-mail: jr@nn.no
  • Gjerde, Norwegian Forest Research Institute, Fanaflaten 4, NO-5244 Fana, Norway E-mail: ig@nn.no
  • Gundersen, Norwegian Forest Research Institute, Fanaflaten 4, NO-5244 Fana, Norway E-mail: vsg@nn.no
article id 377, category Research article
Saara Lilja, Timo Kuuluvainen. (2005). Structure of old Pinus sylvestris dominated forest stands along a geographic and human impact gradient in mid-boreal Fennoscandia. Silva Fennica vol. 39 no. 3 article id 377. https://doi.org/10.14214/sf.377
Keywords: disturbances; restoration; stand structure; managed forest; old growth forest
Abstract | View details | Full text in PDF | Author Info
Stand structural characteristics were examined in old Pinus sylvestris dominated sites in three regions along a broad geographic and human impact gradient in mid-boreal Fennoscandia. The study regions were: 1) Häme in south-western Finland, with a long history of forest utilization, 2) Kuhmo in north-eastern Finland, with a more recent history of intensive forest utilization, and 3) Vienansalo in Russian Karelia, still characterized by a large near-natural forest landscape. Within each region the sampled sites were divided into three human impact classes: 1) near-natural stands, 2) stands selectively logged in the past, and 3) managed stands treated with thinnings. The near-natural and selectively logged stands in Häme and Kuhmo had a significantly higher Picea proportion compared to stands in Vienansalo. In comparison, the proportions of deciduous tree volumes were higher in near-natural stands in Vienansalo compared to near-natural stands in Häme. The pooled tree diameter distributions, both in near-natural and selectively logged stands, were descending whereas managed stands had a bimodal diameter distribution. Structural diversity characteristics such as broken trunks were most common in near-natural stands and in stands selectively logged in the past. The results demonstrate the higher structural complexity of near-natural stands and stands selectively logged in the past compared to managed stands, and highlight that old near-natural stands and stands selectively logged in the past vary widely in their structures. This obviously reflects both their natural variability but also various combinations of pre-industrial land use and human impact on fire disturbance. These factors need to be acknowledged when using “natural” forest structures as a reference in developing strategies for forest management, restoration and nature conservation.
  • Lilja, University of Helsinki, Department of Forest Ecology, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: saara.lilja@helsinki.fi (email)
  • Kuuluvainen, University of Helsinki, Department of Forest Ecology, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: tk@nn.fi
article id 376, category Research article
Nils Lexerød, Trond Eid. (2005). Recruitment models for Norway spruce, Scots pine, birch and other broadleaves in young growth forests in Norway. Silva Fennica vol. 39 no. 3 article id 376. https://doi.org/10.14214/sf.376
Keywords: regeneration; national forest inventories; growth simulators; probability models; conditional models; Norway
Abstract | View details | Full text in PDF | Author Info
The objective of the present study was to develop recruitment models for Norway spruce, Scots pine, birch and other broadleaves in young growth forests in Norway. The models were developed from permanent sample plots established by the National Forest Inventory, and they will be included in a growth simulator that is part of a large-scale forestry scenario model. The modelling was therefore restricted to independent variables directly or indirectly available from inventories for practical forest management planning. A two-stage modelling approach that suited the stochastic nature of recruitment in boreal forests was used. Models predicting the probability of recruitment were estimated in a first stage, and conditional models for the number of recruits were developed in a second. The probability models as well as the conditional models were biologically realistic and logical. The goodness of fit tests revealed that the probability models fitted the data well, while the coefficients of determination for the conditional models were relatively low. No independent test data were available, but comparisons of predicted and observed number of recruits in different sub-groups of the data revealed few large deviations. The high level of large random errors was probably due to the great variability observed in number of recruits rather than inappropriate specifications of the models. Provided the generally high level of uncertainty connected to analysis performed with large-scale forestry scenario models and the stochastic nature of recruitment, the presented models seem to give satisfactory levels of accuracy.
  • Lexerød, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, P.O. Box 5003, NO-1432 Ås, Norway E-mail: nils.lexerod@umb.no (email)
  • Eid, Norwegian University of Life Sciences, Department of Ecology and Natural Resource Management, P.O. Box 5003, NO-1432 Ås, Norway. E-mail nils.lexerod@umb.no E-mail: te@nn.no
article id 375, category Research article
Edgar Víquez, Diego Pérez. (2005). Effect of pruning on tree growth, yield, and wood properties of Tectona grandis plantations in Costa Rica. Silva Fennica vol. 39 no. 3 article id 375. https://doi.org/10.14214/sf.375
Keywords: wood quality; teak; heartwood volume; stem form; knot-free volume
Abstract | View details | Full text in PDF | Author Info
Reduced plantation densities have the effect that obtaining natural pruning and stem straightness are less assured. The physiological process of self-pruning is replaced by manual pruning. Generally, plantations are denser and have more uniform spacing than natural forests. In many, if not most species, natural pruning is never a satisfactory option, even after branch senescence, if production of clear wood is a management objective. Natural pruning can only be considered on a species by species basis. This study reports on the first results of a pruning trial for Tectona grandis L.F. plantations in Costa Rica. The treatments consisted of pruning heights of 3.0, 4.0, and 5.0 meters, and the Control without pruning. Differences among treatments in DBH and total height were significant at 3.2, 5.2, and 6.1 years of age, but not at 7.3 years. Under an intensive pruning regime, a teak tree at rotation (20 years) may yield over 40% of knot-free volume (over 60% of the merchantable tree volume). Current findings open a scope for new management options, aiming at improving stem form and wood quality by means of an intensive pruning regime, without having a detrimental effect on tree growth and stand yield.
  • Víquez, Tropical Agricultural Research and Higher Education Center (CATIE), Apartado 7170 CATIE, Turrialba, Costa Rica E-mail: eviquez@catie.ac.cr (email)
  • Pérez, Ambiente Tierra S.A., Apartado 733-2250, Tres Ríos, Cartago, Costa Rica E-mail: dp@nn.cr
article id 374, category Research article
Jyrki Hytönen, Paula Jylhä. (2005). Effects of competing vegetation and post-planting weed control on the mortality, growth and vole damages to Betula pendula planted on former agricultural land. Silva Fennica vol. 39 no. 3 article id 374. https://doi.org/10.14214/sf.374
Keywords: Betula pendula; vegetation control; herbicides; mulch; cover crop; vole damage; weed cover
Abstract | View details | Full text in PDF | Author Info
Effects of competing vegetation and weed control methods (fibre board mulch, cover crop of clover, various herbicides) on the survival and growth of and vole damage to silver birch (Betula pendula Roth) were analysed based on data from a field experiment established in southern Finland. The cover percentage of competing vegetation and its shading effect were assessed, and seedling size and vitality were recorded several times during the 11-year research period. Mean seedling height and height increment decreased linearly with increasing vegetation cover. Seedling mortality started to significantly increase once the vegetation cover had reached the level of 60–80%. Herbicides significantly retarded increase of weed cover on the initially weedless areas for two to three years, and a cover crop promoted increase in cover percentage. Successful weed control with herbicides significantly increased seedling growth and survival. After 11 years, the average stem volume on the herbicide-treated plots (28.9 m3 ha–1) was 2.5-fold as compared to that of the control plots (11.6 m3 ha–1). Furthermore, seedling mortality on the control plots (21%) was almost 3.5-fold as compared to the seedling mortality on herbicide-treated plots (6%). Having a cover crop proved to be an ineffective weed control method both in terms of seedling growth and survival. The application of mulch had only a slight effect on height increment (0.6 m in 11 years), but on the other hand, it considerably decreased seedling mortality (control: 21%, mulch treatment: 1.5%). These differences were not, however, statistically significant. Small seedling size, high shading class, and high vegetation coverage percentage increased the risk of voles damaging the seedlings.
  • Hytönen, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FI-69101 Kannus, Finland E-mail: jyrki.hytonen@metla.fi (email)
  • Jylhä, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FI-69101 Kannus, Finland E-mail: pj@nn.fi
article id 373, category Research article
Anna Saarsalmi, Pekka Tamminen. (2005). Boron, phosphorus and nitrogen fertilization in Norway spruce stands suffering from growth disturbances. Silva Fennica vol. 39 no. 3 article id 373. https://doi.org/10.14214/sf.373
Keywords: Norway spruce; fertilization; needles; height growth; recovery; boron; growth disorders; soil
Abstract | View details | Full text in PDF | Author Info
Growth disturbance symptoms typical of B deficiency have been reported on Norway spruce (Picea abies (L.) Karst.) trees in many parts of eastern Finland. In order to test the B deficiency hypothesis and explore the possibilities of curing the disturbed trees with B fertilization, three experiments were established in October 1999 in young Norway spruce stands growing on fertile sites in eastern Finland. All the stands contained healthy, slightly and severely damaged trees with growth disturbances typical of B deficiency (B < 5 mg kg–1). 40 healthy, 40 slightly damaged, and 40 severely damaged trees were selected as sample trees in each stand. In May 2000, the trees were fertilized with 2.0 kg B ha–1 as borax (B), 2.0 kg B ha–1 and 40 kg P ha–1 as superphosphate (B+P) or 200 kg N ha–1 as urea (N). The control trees were not fertilized (0). The needle response to B fertilization was rapid, relatively high B concentrations being achieved already after one growing season. Boron fertilization cured the growth disorders and increased height growth within four years, but had no effect on diameter growth. The trees also recovered without B fertilization, but to a lesser extent compared to the B fertilized trees. Compared to the control, boron fertilization increased the height growth in all the disorder classes, i.e. 5, 17 and 19 cm yr–1 for healthy, slightly and severely damaged trees, respectively. As the healthy trees also seemed to benefit from B fertilization, this indicates that B deficiency in fact retards height growth before any disorder symptoms become apparent in individual trees. Compared with B alone, the application of P together with B gave no additional benefit. Nitrogen fertilization alone appeared to have a detrimental effect on height growth in the severely disturbed trees.
  • Saarsalmi, The Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: anna.saarsalmi@metla.fi (email)
  • Tamminen, The Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: pt@nn.fi
article id 372, category Research article
Ulf Sikström. (2005). Pre-harvest soil acidification, liming or N fertilization did not significantly affect the survival and growth of young Norway spruce. Silva Fennica vol. 39 no. 3 article id 372. https://doi.org/10.14214/sf.372
Keywords: Picea abies; regeneration; limestone; sulphur powder; urea
Abstract | View details | Full text in PDF | Author Info
Acidification, liming, and N fertilization affect a number of soil properties. Such changes may have an impact on forest regeneration and yield. The aim of this study was to investigate the survival and growth of Picea abies (L.) Karst. planted on plots that had been acidified (in 12 annual treatments totalling 600 or 1200 kg S ha–1 in the form of elemental sulphur), limed (12 x 500 = 6000 kg lime ha–1 in the form of CaCO3) or N-fertilized (3 x 200 = 600 kg N ha–1 in the form of urea) prior to harvest. Trees growing on plots treated with a combination of the N plus the lower S application were also tested. None of the treatments, applied in three replicate stands, significantly influenced either survival or growth of Picea abies trees during the first 11 growing seasons after planting.
  • Sikström, SkogForsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: ulf.sikstrom@skogforsk.se (email)
article id 371, category Research article
Mika Nieminen, Erkki Ahti, Hannu Nousiainen, Samuli Joensuu, Martti Vuollekoski. (2005). Capacity of riparian buffer zones to reduce sediment concentrations in discharge from peatlands drained for forestry. Silva Fennica vol. 39 no. 3 article id 371. https://doi.org/10.14214/sf.371
Keywords: ditching; suspended solids; peatlands; erosion; riparian buffer zones; water quality protection
Abstract | View details | Full text in PDF | Author Info
In 1995–2001, the efficiency of riparian buffer zone areas to reduce the concentrations of suspended solids in discharge from peatlands drained for forestry purposes was studied at 7 locations in south-central Finland. The two largest buffer zones reduced the concentrations of suspended solids by > 70%. The efficiency of the three medium-sized buffer zones to reduce through-flow sediment concentrations was 50–60%, but no reduction occurred at the smallest two buffer areas. Thus, the capacity of buffer zones to reduce sediment concentrations was strongly related to their size. However, significant correlations were also found between reduction capacity and inflow water sediment concentrations, although the correlations at the two smallest buffer zones were low. The use of buffer zones in reducing sediment load from peatlands drained for forestry purposes is recommended, but relatively large areas for efficient removal capacity are needed.
  • Nieminen, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: mika.nieminen@metla.fi (email)
  • Ahti, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: ea@nn.fi
  • Nousiainen, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: hn@nn.fi
  • Joensuu, Forestry Development Centre Tapio, Soidinkuja 4, FI-00700 Helsinki, Finland E-mail: sj@nn.fi
  • Vuollekoski, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: mv@nn.fi
article id 370, category Research article
Pekka Tamminen, John Derome. (2005). Temporal trends in chemical parameters of upland forest soils in southern Finland. Silva Fennica vol. 39 no. 3 article id 370. https://doi.org/10.14214/sf.370
Keywords: soil chemistry; acidification; time series
Abstract | View details | Full text in PDF | Author Info
Changes in chemical soil properties during periods of 12 to 28 years were studied in 54 stands in southern Finland. Relative slopes (%/year) were calculated for the changes in soil variables in order to utilise all the sampling occasions (2–6) covered by the study period. Only the results of new analyses made on the soil samples could be used owing to unpredictable differences between the results of the original and new analyses. During the study period the acidity (pH, exchangeable acidity) of the organic layer had decreased, and the mineral soil had become more acidic only in terms of increased exchangeable aluminium concentrations. An increasing trend in the amount of soil organic matter best explained the acidity variables: it lowered acidity in the organic layer, but increased it in the mineral soil. Acid ammonium acetate extractable nutrients showed decreasing trends over time, apart from an increasing trend for sulphur in the 0–30 cm mineral soil layer. Total concentrations of most elements in the organic layer, including nitrogen and sulphur, also showed a decreasing trend. Changes in the soil variables could not be firmly connected to deposition, wood production or the amount of nutrients accumulated in woody tissues. However, the decrease in sulphur concentrations in the organic layer was clearly linked with the decrease in sulphur deposition in recent years.
  • Tamminen, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: pekka.tamminen@metla.fi (email)
  • Derome, Finnish Forest Research Institute, Rovaniemi Research Station, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: jd@nn.fi
article id 389, category Research article
Ritva Toivonen, Eric Hansen, Erno Järvinen, Raija-Riitta Enroth. (2005). The competitive position of the Nordic wood industry in Germany – intangible quality dimensions. Silva Fennica vol. 39 no. 2 article id 389. https://doi.org/10.14214/sf.389
Keywords: wood products; quality; the intangible product component; performance
Abstract | View details | Full text in PDF | Author Info
This study examines the importance of various intangible product quality dimensions as perceived by wood-trading retailer and wholesaler companies in Germany. Using perceived importance and perceived performance, the study first examines the dimensionality of intangible product quality and then compares Nordic wood product suppliers with suppliers from other major supply regions. Data was collected from 76 German companies during 2000–2001. Results indicate that intangible product quality can be described in three dimensions, “Behaviour and Image”, “Serviceability and Environment”, and “Reliability”. Results also show that Nordic suppliers do not have a strong competitive position in Germany in terms of intangible product quality dimensions. Thus, Nordic suppliers could improve their competitive position by enhancing their service, logistics and other dimensions of the intangible product offering.
  • Toivonen, Pellervo Economic Research Institute PTT, Eerikinkatu 28 A, FI-00180, Helsinki, Finland E-mail: ritva.toivonen@dnainternet.net (email)
  • Hansen, Oregon State University, Dept. of Wood Science & Engineering, Richardson Hall 108, 97331-5751 Corvallis, OR, USA E-mail: eh@nn.us
  • Järvinen, Pellervo Economic Research Institute PTT, Eerikinkatu 28 A, FI-00180, Helsinki, Finland E-mail: ej@nn.fi
  • Enroth, Finnish Forest Research Institute, Unioninkatu 40 A, FI-00170 Helsinki, Finland E-mail: rre@nn.fi
article id 388, category Research article
Jacob Edlund, Mats Warensjö. (2005). Repeatability in automatic sorting of curved Norway spruce saw logs. Silva Fennica vol. 39 no. 2 article id 388. https://doi.org/10.14214/sf.388
Keywords: Picea abies; compression wood; stem form; bow; scanner; sorting
Abstract | View details | Full text in PDF | Author Info
Sawn wood from curved logs is prone to have cross grain and contain compression wood, both of which affect the dimensional stability. Different types of curvature can, however, have different effects on both the sawing process and board quality, which is why a standard measure of bow height alone is not enough to sort logs or set the log quality. The aim of this study was to evaluate the repeatability when sorting curved saw logs using a 3D log scanner. In the study, 56 logs were categorized into five different curvature types and four different degrees of curvature severity. The logs were run through a Rema 3D log scanner four times, and the external geometry was recorded. From the geometry data, variables describing log shapes were calculated and used to develop models using linear discriminant analysis, which was used to classify the logs according to curvature type. The accuracy and repeatability were evaluated for the classifications with Cohen’s simple Kappa coefficient. The results of this study showed that it is possible to sort logs by curve type using a 3D log scanner, although sorting by curve type was largely dependent on curve severity. The repeatability test determined that the chance of a curved log being graded identically two consecutive times was 0.40, measured as kappa.
  • Edlund, SLU, Department of Forest Products and Markets, P.O. Box 7060, SE-750 07 Uppsala, Sweden E-mail: jacob.edlund@spm.slu.se (email)
  • Warensjö, SLU, Department of Forest Products and Markets, P.O. Box 7060, SE-750 07 Uppsala, Sweden E-mail: mw@nn.se
article id 387, category Research article
Sanna Laukkanen, Teijo Palander, Jyrki Kangas, Annika Kangas. (2005). Evaluation of the multicriteria approval method for timber-harvesting group decision support. Silva Fennica vol. 39 no. 2 article id 387. https://doi.org/10.14214/sf.387
Keywords: group decision support; multicriteria approval; timber-harvesting planning; voting methods
Abstract | View details | Full text in PDF | Author Info
The decision support methods most often used in timber-harvesting planning are based on a single criterion. In this study, a voting-theory-based method called multicriteria approval (MA) is introduced to the group decision support of timber-harvesting. The use of voting methods alleviates the problems caused by the multitude of decision objectives involved in forestry decision-making and by the poor quality of information concerning both the preferences of decision-makers and the evaluation of decision alternatives with respect to the objectives often faced in practical timber-harvesting planning. In the case study, the tactical forest management plan of a forest holding jointly owned by three people was specified at the operative timber-harvesting level. The task was to find the best actual operative alternatives for the harvesting that had been proposed in the tactical plan. These timber-harvesting alternatives were combinations of treatment, timber-harvesting system and the timing of logging. Forest owners established multiple criteria under which the alternatives were evaluated. Two versions of MA were tested, one of them based on individual decision analyses and other one based on a composite analysis. The first was markedly modified from the original MA, combining properties of MA and Borda count voting. The other was an original MA with the order of importance for criteria estimated either using Borda count or cumulative voting. The results of the tested MA versions produced were very similar to each other. MA was found to be a useful tool for the group decision support of timber-harvesting.
  • Laukkanen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: sanna.laukkanen@joensuu.fi (email)
  • Palander, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: tp@nn.fi
  • Kangas, UPM-Kymmene Forest, P.O. Box 32, FI-37601 Valkeakoski, Finland E-mail: jk@nn.fi
  • Kangas, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ak@nn.fi
article id 386, category Research article
Jouni Kalliovirta, Timo Tokola. (2005). Functions for estimating stem diameter and tree age using tree height, crown width and existing stand database information. Silva Fennica vol. 39 no. 2 article id 386. https://doi.org/10.14214/sf.386
Keywords: forest inventory; modeling; crown diameter; stem diameter
Abstract | View details | Full text in PDF | Author Info
The aim was to investigate the relations between diameter at breast height and maximum crown diameter, tree height and other possible independent variables available in stand databases. Altogether 76 models for estimating stem diameter at breast height and 60 models for tree age were formulated using height and maximum crown diameter as independent variables. These types of models can be utilized in modern remote sensing applications where tree crown dimensions and tree height are measured automatically. Data from Finnish national forest inventory sample plots located throughout the country were used to develop the models, and a separate test site was used to evaluate them. The RMSEs of the diameter models for the entire country varied between 7.3% and 14.9% from the mean diameter depending on the combination of independent variables and species. The RMSEs of the age models for entire country ranged from 9.2% to 12.8% from the mean age. The regional models were formulated from a data set in which the country was divided into four geographical areas. These regional models reduced local error and gave better results than the general models. The standard deviation of the dbh estimate for the separate test site was almost 5 cm when maximum crown width alone was the independent variable. The deviation was smallest for birch. When tree height was the only independent variable, the standard deviation was about 3 cm, and when both height and maximum crown width were included it was under 3 cm. In the latter case, the deviation was equally small (11%) for birch and Norway spruce and greatest (13%) for Scots pine.
  • Kalliovirta, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: jk@nn.fi (email)
  • Tokola, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: timo.tokola@helsinki.fi
article id 385, category Research article
Diego Pérez, Markku Kanninen. (2005). Effect of thinning on stem form and wood characteristics of teak (Tectona grandis) in a humid tropical site in Costa Rica. Silva Fennica vol. 39 no. 2 article id 385. https://doi.org/10.14214/sf.385
Keywords: sapwood; stem taper; basic density; heartwood; form factor
Abstract | View details | Full text in PDF | Author Info
The aim of the study was to evaluate the effects of thinning intensity on wood properties, such as heartwood proportion, wood basic density, and stem form of teak (Tectona grandis L.f.). The thinning trial was established on a teak plantation in a humid tropical site in northern Costa Rica. The moderate and heavy thinnings yielded the highest percentage of heartwood volume (25 to 30% of total stem volume). The differences between stem form factors under different treatments were not statistically significant after separating thinning effects from timing effects. Both the highest (> 0.65 g cm–3) and the lowest (< 0.50 g cm–3) wood density values were observed under light thinnings, making it difficult to establish a relationship. Large variations in wood properties found under different thinning regimes suggest that at early stages teak stands can be managed under different thinning programs without negatively affecting the quality of wood under humid tropical conditions.
  • Pérez, Ambiente Tierra S.A., Apartado 733-2250, Tres Ríos, Cartago, Costa Rica E-mail: diegoperez@costarricense.cr (email)
  • Kanninen, Center for International Forestry Research (CIFOR), Bogor, Indonesia E-mail: mk@nn.id
article id 384, category Research article
Jorge Cancino, Joachim Saborowski. (2005). Comparison of randomized branch sampling with and without replacement at the first stage. Silva Fennica vol. 39 no. 2 article id 384. https://doi.org/10.14214/sf.384
Keywords: Randomized Branch Sampling; sampling without replacement; probability proportional to size; unequal selection probabilities; multistage sampling
Abstract | View details | Full text in PDF | Author Info
Randomized Branch Sampling (RBS) is a multistage sampling procedure using natural branching in order to select samples for the estimation of tree characteristics. Usually, sampling units are selected with unequal probabilities. Conventional RBS uses sampling with replacement (SWR) for repeated sampling on the first stage, and the sample size equals 1 on all subsequent stages, thus resulting in n so-called sample paths. When the sampling fraction is large multiple selections of first stage units are likely. Sampling without replacement (SWOR) at the first stage is an alternative that is expected to increase efficiency of the procedure. In this case, the second stage sample size m must be larger than 1 to enable unbiased variance estimation. In the present study, a theoretical and an empirical comparison of the conventional RBS and the SWOR variant was accomplished. Requiring a certain precision of the RBS estimation, the conventional RBS method is mostly more time-consuming than the variant with SWOR at the first stage. Only if m = 1 is chosen as second stage sample size for the SWOR RBS, this is often more time-consuming. In those cases, conventional RBS is up to 5% cheaper. In general, the larger m is, the more expensive is conventional RBS compared with the variant with swor at the first stage. The smaller the ratio of the variance between the primary units to the total variance of the estimate, the larger is the advantage of the SWOR variant. Generally, it can be shown that the gain of efficiency by SWOR is larger in case of weak correlations between auxiliary and target variable.
  • Cancino, Facultad de Ciencias Forestales, Universidad de Concepción, Chile E-mail: jcancino@udec.cl (email)
  • Saborowski, Institut für Forstliche Biometrie und Informatik, Georg-August-Universität Göttingen, Germany E-mail: js@nn.de
article id 383, category Research article
Lena Gustafsson, Leif Appelgren, Anders Nordin. (2005). Biodiversity value of potential forest fertilisation stands, as assessed by red-listed and ‘signal’ bryophytes and lichens. Silva Fennica vol. 39 no. 2 article id 383. https://doi.org/10.14214/sf.383
Keywords: biodiversity; conservation; hemi-boreal forest; Sweden; threatened species
Abstract | View details | Full text in PDF | Author Info
In Sweden ca. 20 000 ha forestland is fertilised each year. By using red-listed and ‘signal’ bryophytes and lichens as indicators, we investigated whether forest stands planned for fertilisation have a biodiversity value, and thus if restrictions due to conservation aspects are motivated. Species occurrences were registered in detailed line-transect analysis, with a record size of 10 x 10 m, in 74 coniferous forest stands with a mean age of 57 years in East-Central Sweden. On the 230 ha totally surveyed, 10 red-listed and 37 signal species were found. The mean number of records ha–1 of red-listed bryophytes and lichens was 0.26 ha–1, which is considerably less than previously found in mature production stands and woodland key habitats. Red-listed species were found in 31% of the stands and signal species in 95%. More than 70% of all records of red-listed species and 30% of the records of the signal species were found in moist micro-sites. If rare bryophytes and lichens are to be preserved in fertilisation stands, improved instructions regarding avoidance of important micro-sites are needed.
  • Gustafsson, Swedish University of Agricultural Sciences, Department of Conservation Biology, Box 7002, SE-750 07 Uppsala, Sweden E-mail: lena.gustafsson@nvb.slu.se (email)
  • Appelgren, Belfragegatan 34H, SE-462 37 Vänersborg, Sweden E-mail: la@nn.se
  • Nordin, Museum of Evolution, Botany, Norbyvägen 16, SE-752 36 Uppsala, Sweden E-mail: an@nn.se
article id 382, category Research article
Steen Magnussen, René I. Alfaro, Paul Boudewyn. (2005). Survival-time analysis of white spruce during spruce budworm defoliation. Silva Fennica vol. 39 no. 2 article id 382. https://doi.org/10.14214/sf.382
Keywords: mortality; hazard rates; defoliation stress index; Cox proportional hazard regression; Choristoneura fumiferana
Abstract | View details | Full text in PDF | Author Info
Mortality and defoliation (DF%) in 987 white spruce (Picea glauca (Moench) Voss) trees were followed from 1992 to 2003 during an outbreak of the spruce budworm Choristoneura fumiferana (Clem.) in 15 white-spruce-dominated uneven-aged stands in the Fort Nelson Forest District near Prince George, British Columbia. Four stands were aerially sprayed with Bacillus thuringiensis (Bt). Defoliation and mortality levels were elevated in non-sprayed stands. The relationship between defoliation and survival-times was captured in a Cox proportional hazard model with a defoliation stress index (DSI), diameter (DBH), crown class (CCL), a random stand effect, Bt-treatment, and number of years of exposure to stand-level defoliation (DYEAR) as predictors. The DSI, optimized for discrimination between survivors and non-survivors, is the discounted sum of five lagged DF% values. Survival probabilities were predicted with a maximum error of 0.02. Hazard rates increased by 0.06 for every one point increase in DSI. CCL and random stand effects were highly significant. Bt-treatment effects were fully captured by DSI, CCL, and DYEAR.
  • Magnussen, Canadian Forest Service, Victoria, BC, Canada. V8Z 1M5 E-mail: smagnussen@pfc.forestry.ca (email)
  • Alfaro, Canadian Forest Service, Victoria, BC, Canada. V8Z 1M5 E-mail: ria@nn.ca
  • Boudewyn, Canadian Forest Service, Victoria, BC, Canada. V8Z 1M5 E-mail: pb@nn.ca
article id 381, category Research article
Jianxun Luo, Yuhua Wang, Helena Korpelainen, Chunyang Li. (2005). Allozyme variation in natural populations of Picea asperata. Silva Fennica vol. 39 no. 2 article id 381. https://doi.org/10.14214/sf.381
Keywords: allozymes; genetic variability; Picea asperata
Abstract | View details | Full text in PDF | Author Info
A survey of allozymic alleles and genetic diversity was conducted for ten natural populations of Picea asperata Mast. originating from the mountains of Southwest China. A total of twenty-seven alleles at seventeen loci were observed. Ten of the loci were found monomorphic. Our results showed that the populations sampled were characterized by low genetic diversity (mean He = 0.096) and a low level of inbreeding (mean Fis = 0.005). The UPGMA tree of genetic relationships indicated that there was significant differentiation among populations. The coefficient of genetic differentiation among populations, based on Fst, equaled 0.311. Such extensive inter-populational differentiation detected in P. asperata could have resulted from allele frequency divergence among populations, particularly, in one population. Introgression from another species, variation in environmental conditions, and differing selection pressures could be some of the factors attributing to significant differences among populations. In addition, our results showed that the geographic and genetic distances were not correlated in the populations of P. asperata. Based on the genetic information obtained, we concluded that monitoring appropriate genetic markers may be an effective means of identifying potential genetic changes occurring during forest tree evolution.
  • Luo, Sichuan Academy of Forestry, Chengdu 610081, P. R. China E-mail: jl@nn.cn
  • Wang, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O. Box 416, Chengdu 610041, P. R. China; Graduate School of the Chinese Academy of Sciences, Beijing 100039, P. R. China E-mail: yw@nn.cn
  • Korpelainen, Department of Applied Biology, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: hk@nn.fi
  • Li, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O. Box 416, Chengdu 610041, P. R. China E-mail: licy@cib.ac.cn (email)
article id 380, category Research article
Tapani Repo, Janne Laukkanen, Raimo Silvennoinen. (2005). Measurement of the tree root growth using electrical impedance spectroscopy. Silva Fennica vol. 39 no. 2 article id 380. https://doi.org/10.14214/sf.380
Keywords: willow; CNLS-curve fitting; displacement method; distributed electric model; hydroponics; impedance analysis
Abstract | View details | Full text in PDF | Author Info
The non-destructive evaluation of plant root growth is a challenge in root research. In the present study we aimed to develop electrical impedance spectroscopy (EIS) for that purpose. Willows (Salix myrsinifolia Salisb.) were grown from cuttings in a hydroponic culture in a growth chamber. Root growth was monitored at regular intervals by a displacement method and compared with the EIS parameters of the plants. To measure its impedance spectrum (IS) (frequency range from 40 Hz to 340 kHz) each plant was set in a measuring cell filled with a solution of the hydroponic culture. The IS was measured using a two-electrode measuring system. A silver needle electrode was connected to the stem immediately above the immersion level and a platinum wire was placed in the solution. The measurements were repeated twice weekly for a root growth period of one month. The IS of the entity consisting of a piece of stem, roots and culture solution were modelled by means of an electric circuit consisting of two ZARC-Cole elements, one constant-phase element, and a resistor. On the plant basis, an increase in root volume by growth correlated with a reduction in the sum of resistances in the ZARC-Cole elements (mean Pearson’s correlation coefficient r = –0.70).
  • Repo, The Finnish Forest Research Institute, Joensuu Research Centre, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: tapani.repo@metla.fi (email)
  • Laukkanen, University of Joensuu, Department of Physics, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: jl@nn.fi
  • Silvennoinen, University of Joensuu, Department of Physics, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: rs@nn.fi
article id 400, category Research article
Antti Mutanen, Anne Toppinen. (2005). Finnish sawlog market under forest taxation reform. Silva Fennica vol. 39 no. 1 article id 400. https://doi.org/10.14214/sf.400
Keywords: forest taxation; reforms; roundwood supply; simultaneous equations
Abstract | View details | Full text in PDF | Author Info
The stepwise transition in forest taxation from site productivity tax to taxation of profits from timber sales was one of the major institutional changes to impact Finnish non-industrial private forest owners in the 1990s. In this study the effect of the forest taxation reform on the aggregate supply of sawlogs was investigated using time series analysis and quarterly data. In particular, we estimated two simultaneous equations systems for the pine and the spruce sawlog markets. According to the results, the forest taxation reform strengthened the supply of spruce and pine sawlogs in the anticipatory stage of the taxation reform in 1992. Also during the fiscal transition period, which started in 1993, the supply effect of the taxation reform has clearly been positive. The strong own-price elasticity of sawlog supply found in this study indicates high sensitivity to actual and expected wood price changes in the determination of sawlog supply in Finland. Furthermore, the results indicate that the theoretical assumption of a competitive market is suitable for the Finnish sawlog market, but that separate analysis of pine and spruce sawlogs provides additional insights into market behaviour.
  • Mutanen, Finnish Forest Research Institute, Joensuu Research Centre, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: antti.mutanen@metla.fi (email)
  • Toppinen, Finnish Forest Research Institute, Joensuu Research Centre, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: at@nn.fi
article id 399, category Research article
Torjus F. Bolkesjø. (2005). Projecting pulpwood prices under different assumptions on future capacities in the pulp and paper industry. Silva Fennica vol. 39 no. 1 article id 399. https://doi.org/10.14214/sf.399
Keywords: paper industry; Norway; capacity; partial equilibrium model; pulpwood; prices
Abstract | View details | Full text in PDF | Author Info
Capacity changes in the pulp and paper industry affect demand for pulpwood and thus pulpwood prices. This paper analyzes the impacts on roundwood prices in Norway of two possible capacity changes (one new machine and one close-down) that currently are high on the agenda in the Norwegian paper industry, and assesses the generality of the results obtained from these case studies. The two cases are implemented exogenously into a regionalized partial equilibrium forest sector model, and the capacity change scenarios are compared with a business as usual scenario assuming no demand shocks. The projected pulpwood prices change significantly in regions near mills where capacity shifts, at least for the close-down case, but only moderately at an aggregated national level. The reduction in prices under the close-down studied is higher than the price increase from the possible capacity increase case. The asymmetric price responses projected for the two case studies are supported by sensitivity analyses on other regions and cases (technologies). For the capacity increase case it is shown that the level of the projected pulpwood price is sensitive to assumptions on base-year prices and transport costs of imported roundwood, but the magnitudes of the price increases projected as a result of increased demand are less affected by these assumptions.
  • Bolkesjø, Norwegian University of Life Sciences, Dept. of Ecology and Natural Resource Management, Høyskoleveien 14, NO-1432 Ås, Norway E-mail: torjus.bolkesjo@umb.no (email)
article id 398, category Research article
Heikki Ovaskainen. (2005). Comparison of harvester work in forest and simulator environments. Silva Fennica vol. 39 no. 1 article id 398. https://doi.org/10.14214/sf.398
Keywords: time study; single-grip harvester; harvester simulator; working technique; PlusCan
Abstract | View details | Full text in PDF | Author Info
Harvester simulators offer a safe and cost-saving method for studying the basics of harvester controls and working technique. Therefore, harvester simulators are increasingly being used in the education of harvester operators. In this study, the objective was to compare harvester work in real and simulator environments, and to determine how a professional harvester operator’s working technique may have changed in the simulator environment. Specific features of the simulator that encumbered operators’ normal work are also presented; and the correspondence of the simulator to reality is evaluated. The work of six professional harvester operators was studied in thinning and in clear cutting stands in both environments: first in the real forest and thereafter on the simulator. The results indicate that the operators’ working technique on the simulator was mainly the same as in the real forest. This means that the same restrictions are valid on the simulator as in the forest. The basic principles of harvesting must be known so that high productivity and good quality can be obtained. However, certain simulator-specific features encumbered the work of harvester operators. Limited visibility to the side increased the need to reverse and the 3D-visualization caused failed catches. Improvements in software would remove some of the defects, e.g. failed felling and cheating in the felling phase. These results also indicate that simulators can be used for research purposes.
  • Ovaskainen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: heikki.ovaskainen@joensuu.fi (email)
article id 397, category Research article
Nuutti Kiljunen. (2005). Pricing the risk of the quality-guarantee in a stand establishment service. Silva Fennica vol. 39 no. 1 article id 397. https://doi.org/10.14214/sf.397
Keywords: quality; forest regeneration; risk; guarantee
Abstract | View details | Full text in PDF | Author Info
A stand-establishment service concept with quality guarantee was analysed. Here, the quality of stand establishment was assessed as the density of good quality seedlings evenly distributed on the plantation three years after planting. The amount of adequate premiums for the guarantee service and the risks accumulating to the service provider were studied. Monte Carlo simulation was used as a tool for analyzing the risks accumulating to the service provider of the stand-establishment in operational environments of different sizes. The premiums calculated to cover the expected amount of claims caused by the plantations not meeting the pre-set criteria were about 4–8% in addition to the approximated costs of stand establishment. The criteria used for determining the success or failure in a stand have a marked effect on the amount of premiums with a reasonable risk of ruin.
  • Kiljunen, Finnish Forest Research Institute, Suonenjoki Research Station, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: nuutti.kiljunen@metla.fi (email)
article id 396, category Research article
Timo Pukkala, Mikko Kurttila. (2005). Examining the performance of six heuristic optimisation techniques in different forest planning problems. Silva Fennica vol. 39 no. 1 article id 396. https://doi.org/10.14214/sf.396
Keywords: genetic algorithms; simulated annealing; ecological planning; habitat suitability index (HSI); Hero; random search; tabu search
Abstract | View details | Full text in PDF | Author Info
The existence of multiple decision-makers and goals, spatial and non-linear forest management objectives and the combinatorial nature of forest planning problems are reasons that support the use of heuristic optimisation algorithms in forest planning instead of the more traditional LP methods. A heuristic is a search algorithm that does not necessarily find the global optimum but it can produce relatively good solutions within reasonable time. The performance of different heuristics may vary depending on the complexity of the planning problem. This study tested six heuristic optimisation techniques in five different, increasingly difficult planning problems. The heuristics were evaluated with respect to the objective function value that the techniques were able to find, and the time they consumed in the optimisation process. The tested optimisation techniques were 1) random ascent (RA), 2) Hero sequential ascent technique (Hero), 3) simulated annealing (SA), 4) a hybrid of SA and Hero (SA+Hero), 5) tabu search (TS) and 6) genetic algorithm (GA). The results, calculated as averages of 100 repeated optimisations, were very similar for all heuristics with respect to the objective function value but the time consumption of the heuristics varied considerably. During the time the slowest techniques (SA or GA) required for convergence, the optimisation could have been repeated about 200 times with the fastest technique (Hero). The SA+Hero and SA techniques found the best solutions for non-spatial planning problems, while GA was the best in the most difficult problems. The results suggest that, especially in spatial planning problems, it is a benefit if the method performs more complicated moves than selecting one of the neighbouring solutions. It may also be beneficial to combine two or more heuristic techniques.
  • Pukkala, University of Joensuu, Faculty of Forestry, P.O. BOX 111, FI-80101 Joensuu, Finland E-mail: timo.pukkala@forest.joensuu.fi (email)
  • Kurttila, Finnish Forest Research Institute, Joensuu Research Centre, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: mk@nn.fi
article id 395, category Research article
Lauri Mehtätalo. (2005). Height-diameter models for Scots pine and birch in Finland. Silva Fennica vol. 39 no. 1 article id 395. https://doi.org/10.14214/sf.395
Keywords: mixed model; longitudinal analysis; random parameter; stand development
Abstract | View details | Full text in PDF | Author Info
Height-Diameter (H-D) models for two shade-intolerant tree species were estimated from longitudinal data. The longitudinal character of the data was taken into account by estimating the models as random effects models using two nested levels: stand and measurement occasion level. The results show that the parameters of the H-D equation develop over time but the development rate varies between stands. Therefore the development of the parameters is not linked to the stand age but to the median diameter of the basal-area weighted diameter distribution (DGM). Models were estimated with different predictor combinations in order to produce appropriate models for different situations. The estimated models can be localized for a new stand using measured heights and diameters, presumably from different points in time, and the H-D curves can be projected into the future.
  • Mehtätalo, Finnish Forest Research Institute, Joensuu Research Centre, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: lauri.mehtatalo@metla.fi (email)
article id 394, category Research article
Rafael Calama, Gregorio Montero. (2005). Multilevel linear mixed model for tree diameter increment in stone pine (Pinus pinea): a calibrating approach. Silva Fennica vol. 39 no. 1 article id 394. https://doi.org/10.14214/sf.394
Keywords: mixed model; diameter increment; calibration; stone pine
Abstract | View details | Full text in PDF | Author Info
Diameter increment for stone pine (Pinus pinea L.) is described using a multilevel linear mixed model, where stochastic variability is broken down among period, plot, tree and within-tree components. Covariates acting at tree and stand level, as breast height diameter, density, dominant height or site index are included in the model as fixed effects in order to explain residual random variability. The effect of competition on diameter increment is expressed by including distance independent competition indices. The entrance of regional effects within the model is tested to determine whether a single model is sufficient to explain stone pine diameter increment in Spain, or if, on the contrary, regional models are needed. Diameter increment model can be calibrated by predicting random components using data from past growth measurements taken in a complementary sample of trees. Calibration is carried out by using the best linear unbiased predictor (BLUP) theory. Both the fixed effects model and the calibrated model mean a substantial improvement when compared with the classical approach, widely used in forest management, of assuming constancy in diameter increment for a short projection period.
  • Calama, CIFOR-INIA, Grupo Selvicultura Mediterranea, Apdo. 8111, 28080 Madrid, Spain E-mail: rcalama@inia.es (email)
  • Montero, CIFOR-INIA, Grupo Selvicultura Mediterranea, Apdo. 8111, 28080 Madrid, Spain E-mail: gm@nn.es
article id 393, category Research article
Seppo Rouvinen, Anne Rautiainen, Jari Kouki. (2005). A relation between historical forest use and current dead woody material in a boreal protected old-growth forest in Finland. Silva Fennica vol. 39 no. 1 article id 393. https://doi.org/10.14214/sf.393
Keywords: Populus tremula; Pinus sylvestris; Picea abies; coarse woody debris; forest utilization; historical ecology; land-use history; nature conservation; spatial pattern analysis
Abstract | View details | Full text in PDF | Author Info
Assessing the human impact on the naturalness and vegetation characteristics of protected areas is one of the key issues when designing forest conservation networks in Fennoscandia. We studied the small-scale, detailed relationship between forest utilization history and the current availability of dead woody material in a protected old-growth forest area in North Karelia, eastern Finland. From the study area of 32.4 ha, all the stumps (diameter ≥ 5 cm and height < 1.3 m, classified as natural, man-made and of undetermined origin) were measured using 25 x 25 m sub-plots. Standing and fallen dead trees (dbh ≥ 5 cm) were measured on 50 x 50 m plots in an area of 7.8 ha. The average number of stumps was 130 per ha, and over half of the stumps were classified as man-made. However, the historical documents since the 1910s showed no logging in the area: some of the largest man-made stumps probably originated from an earlier time, but most of those stumps were made considerably later. The variation in the total number of stumps (per ha) was great (range 0–560/ha, 0–16 m2/ha), with no clear clustering in space. However, clustering of man-made stumps was detected. The average volume of pooled standing and fallen trees was 84 m3/ha, with a range of 37–146 m3/ha. The other noticeable man-made disturbance besides logging was notching of aspens, which has a scatteredly significant influence on the amount of dead trees. In conclusion, the protected old-growth forest was not as a whole in a natural state but showed different degrees of human impact from virtually untouched patches to quite heavily managed patches. The results suggest that the number of man-made stumps may be a relatively quick and easy method of assessing the naturalness of woody biomass structure in the Fennoscandian boreal forests.
  • Rouvinen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: seppo.rouvinen@joensuu.fi (email)
  • Rautiainen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: ar@nn.fi
  • Kouki, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: jk@nn.fi
article id 392, category Research article
Veikko Huhta, Mika Räty. (2005). Soil animal communities of planted birch stands in central Finland. Silva Fennica vol. 39 no. 1 article id 392. https://doi.org/10.14214/sf.392
Keywords: forest management; birch; plantations; soil fauna
Abstract | View details | Full text in PDF | Author Info
The aim of this study was to compare soil animal communities in planted birch (Betula pendula) stands of different origin with each other and with natural forests. Six 30-year-old birch stands were investigated, three planted after clear-cutting of spruce stands, and three planted on cultivated soil. The faunal communities were markedly different in plantations established on spruce forest soil and on arable soil. “Birch after Spruce” communities were relatively similar to those of coniferous forests, though the population densities were generally lower. “Birch after Field” communities were sparse and could be characterised as “impoverished forest communities”, except in Lumbricidae and Enchytraeidae that have affinities with deciduous forests and cultural landscapes. Soil conditions are not sufficient to explain the differences between the forests. Colonisation and transport by man may determine the presence of certain species, especially earthworms. These in turn affect soil properties, and compete with or otherwise have negative effects on other soil fauna. Thus the community differences between different forests are an outcome of several factors: soil characteristics, site history, colonisation ability and interspecific interactions.
  • Huhta, Department of Biological and Environmental Science, University of Jyväskylä, P.O. Box 35, FI-40014 Jyväskylä University, Finland E-mail: v.huhta@pp.inet.fi (email)
  • Räty, Ojalanlenkki 4, FI-80140 Joensuu, Finland E-mail: mr@nn.fi
article id 410, category Research article
Jouni Siipilehto, Juha Siitonen. (2004). Degree of previous cutting in explaining the differences in diameter distributions between mature managed and natural Norway spruce forests. Silva Fennica vol. 38 no. 4 article id 410. https://doi.org/10.14214/sf.410
Keywords: Picea abies; stand structure; naturalness; Johnson’s SB distribution
Abstract | View details | Full text in PDF | Author Info
The degree of naturalness was assessed in 37 mature (stand age 80 198 yrs) Norway spruce dominated stands located in southern Finland by measuring the number (0 610 ha–1) and basal area (0 33 m2 ha–1) of cut stumps. The Johnson’s SB distribution was fitted for living spruce trees to describe the dbh-frequency and basal area-dbh distributions. Regression models were constructed for predicting the parameters of the SB distribution using traditional stand parameters (median diameter, basal area, stem number) and the cut stump variables (number, basal area). Stump variables improved the models and enabled to explain the differences in diameter distributions between stands with varying intensity of past cutting. Model for basal area-dbh distribution was more accurate than dbh-frequency model in terms of regression statistics, but less accurate in terms of generated stand variables. The number and basal area of cut stumps seem to be useful and simple measures of stand naturalness which have potential uses in stand modelling and biodiversity-oriented forestry planning.
  • Siipilehto, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: jouni.siipilehto@metla.fi (email)
  • Siitonen, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: juha.siitonen@metla.fi
article id 409, category Research article
Hubert Sterba. (2004). Equilibrium curves and growth models to deal with forests in transition to uneven-aged structure – application in two sample stands. Silva Fennica vol. 38 no. 4 article id 409. https://doi.org/10.14214/sf.409
Keywords: uneven-aged forests; growth models; stem number distribution; transformation
Abstract | View details | Full text in PDF | Author Info
Stem number distributions in uneven-aged forests are assumed to be stable, if they follow special functions, e.g. de Liocourt’s reverse J-shaped breast height diameter distribution. These distributions therefore are frequently regarded as a target in all-aged forests. Intending to convert an even-aged forest or any other forest, not yet exhibiting this sort of equilibrium, towards a steady state forest, the question rises, how to choose an appropriate equilibrium curve and how to achieve this stem number distribution by an appropriate thinning and harvesting schedule. Two stands are investigated: One dominated by Norway spruce (Picea abies), having developed from a 120 year old even-aged stand 25 years ago, after several “target diameter thinnings”. The other one is a mixed species stand of Norway spruce, white fir (Abies alba), larch (Larix europea), common beech (Fagus silvatica), and Scots pine (Pinus sylvestris), having lost its typical uneven-aged structure 20 years ago. These stands were used, together with the distance independent individual tree growth model PrognAus, to reveal that 1) there are more than only one equilibrium curve per stand, 2) not every hypothesised equilibrium can be reached with any stand, 3) an equilibrium in stem number does not necessarily mean a stable species distribution, and 4) growth models provide an excellent help to decide between several equilibrium curves and harvesting schedules to reach them.
  • Sterba, BOKU – University of Natural Resources an Applied Life Sciences, Vienna, Peter Jordanstrasse 82, A-1190 Vienna, Austria E-mail: hubert.sterba@boku.ac.at (email)
article id 408, category Research article
Sakari Sarkkola, Hannu Hökkä, Timo Penttilä. (2004). Natural development of stand structure in peatland Scots pine following drainage: results based on long-term monitoring of permanent sample plots. Silva Fennica vol. 38 no. 4 article id 408. https://doi.org/10.14214/sf.408
Keywords: Pinus sylvestris; peatland; tree mortality; stand structure; drainage; dbh distribution
Abstract | View details | Full text in PDF | Author Info
We studied the dynamics of stand structure on drained peatland sites in Scots pine dominated stands untreated with thinnings. The data consisted of consecutive stand measurements in 10 permanent sample plots where the monitoring periods varied from 29 to 66 years. We assumed that the stand’s structural development was driven by the natural processes of regeneration, growth, and mortality, all related to inter-tree competition within the stand. The DBH distributions of live and dead trees at different times of post-drainage stand development – smoothed by Weibull function – were analysed to characterise the change in stand structure. The initial uneven-sized structure of the natural, widely-spaced stands became more uneven during the first decades following drainage due to enhanced regeneration. Later, as stand density and mean tree size continuously increased, the DBH distributions approached bell-shaped distributions. Accordingly, the suppressed trees showed their highest mortality rate during the first decades, but the peak of the mortality distribution shifted to larger trees along stand succession. The change in structure was faster in southern Finland than in northern Finland. We assumed the changes in stand dynamics reflected increased inter-tree competition, initiated by enhanced site productivity and increased stand stocking resulting from the ditching operation.
  • Sarkkola, University of Helsinki, Department of Forest Ecology, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: sakari.sarkkola@helsinki.fi (email)
  • Hökkä, Finnish Forest Research Institute, Rovaniemi Research Station, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: hh@nn.fi
  • Penttilä, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: tp@nn.fi
article id 407, category Research article
Soili Kojola, Timo Penttilä, Raija Laiho. (2004). Impacts of different thinning regimes on the yield of uneven-structured Scots pine stands on drained peatland. Silva Fennica vol. 38 no. 4 article id 407. https://doi.org/10.14214/sf.407
Keywords: Pinus sylvestris; stand structure; silviculture; growth and yield; peatland forestry; intermediate cuttings
Abstract | View details | Full text in PDF | Author Info
Drained peatlands in northern Europe comprise more than 10 million ha of forestland and thus constitute a considerable production potential in forestry. Much of this area consists of stands dominated by Scots pine and close to maturity regarding commercial thinning. The trees within these stands typically vary in terms of age, size, and growth rate. The impacts of silvicultural cuttings on these uneven-structured stands are inadequately known. We simulated the impacts of a control regime with no thinnings, and three different thinning regimes, involving different thinning intensities, on the development of fifteen pine-dominated stands in Finland. The simulations started from the first thinnings and were continued until regeneration maturity. The predicted total yields ranged from 244 to 595 m3 ha–1, depending on site and thinning regime. The highest total yields were observed for the control regime in which 18–38% of the yield was, however, predicted to self-thin by the end of the simulation. Thus, the differences in the yields of merchantable wood were fairly small among the compared regimes. However, the regimes involving thinnings generally needed less time than the control regime to reach regeneration maturity. The mean annual increment of total stem volume was at its highest in the control regime. The highest mean annual increment of merchantable wood was obtained in the regime involving two moderate thinnings, but excluding the most low-productive sites where thinnings did not increase the yield of merchantable wood.
  • Kojola, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: soili.kojola@metla.fi (email)
  • Penttilä, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: tp@nn.fi
  • Laiho, University of Helsinki, Department of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: rl@nn.fi
article id 406, category Research article
Bernt-Håvard Øyen, Petter Nilsen. (2004). Growth and recruitment after mountain forest selective cutting in irregular spruce forest. A case study in Northern Norway. Silva Fennica vol. 38 no. 4 article id 406. https://doi.org/10.14214/sf.406
Keywords: Norway spruce; mountain forests; selective cutting; North-Norway
Abstract | View details | Full text in PDF | Author Info
During the last thirty years the interest for the use of selective cutting in the sub-alpine spruce forests of Norway has increased. However, there have been very few investigations on the post harvesting development after such cuttings. Four plots in irregular Norway spruce (Picea abies (L.) Karsten) dominated forests on semi-fertile sites in Northern Norway have been the subjects of a case study. We performed a reconstruction of the stand development by means of biometric assessments and ring widths measurements of all standing trees. Tree ages, stand structure, growth and recruitment were examined. Even though a hypothetical reverse J-curve for the present diameter distribution was identified, the four plots were even-aged. Growth reactions indicate that most of the present sawtimber trees were established after heavy dimension cuttings in the late 19th century. The recruitment situation is characterized as satisfying in one of four plots. The post harvesting mean volume increment on the plots have been about two thirds of the potential yield estimated from site indices and maximum mean annual increment in regular stands. Managing strategies for irregular spruce forest stands are briefly discussed.
  • Øyen, Norwegian Forest Research Institute-Bergen, Fanaflaten 4, N-5244 Fana, Bergen, Norway E-mail: bernt-havard.oyen@skogforsk.no (email)
  • Nilsen, Norwegian Forest Research Institute-Bergen, Fanaflaten 4, N-5244 Fana, Bergen, Norway E-mail: pn@nn.no
article id 405, category Research article
Timo Saksa. (2004). Regeneration process from seed crop to saplings – a case study in uneven-aged Norway spruce-dominated stands in southern Finland. Silva Fennica vol. 38 no. 4 article id 405. https://doi.org/10.14214/sf.405
Keywords: boreal forest; Picea abies; regeneration; uneven-aged
Abstract | View details | Full text in PDF | Author Info
The dynamics of spruce regeneration, from seed crop to saplings, was studied based on five permanent plots in uneven-aged, spruce-dominated, boreal forest stands, cut with single-tree selection in the beginning of the 1990’s. The annual fluctuation of the spruce seed crop was very similar in uneven-aged and even-aged stands. The correlation between seed crop and number of germinants was significant; but stem number, basal area or volume of the stand did not influence on seedling emergence. The effects of good seed crops were seen as peaks or an increase in the number of germinants and smallest seedlings. The mean number of ‘stabilised’ spruce seedlings (height 11 cm to 130 cm) varied from 6000 ha–1 to over 25 000 spruce seedlings ha–1 from one monitoring plot to another. On a monitoring plot the number of ‘stabilised’ spruce seedlings was stable over time. Neither stand basal area nor stand volume influenced the number of ‘stabilised’ spruce seedlings, but the height of these seedlings was higher on subplots with lower stand volume and smaller basal area. In this study the monitoring period, 5–10 years, was too short to obtain reliable figures for ingrowth, i.e. the transition of seedlings to the sapling stage (h > 130 cm). The adjusted mean ingrowth was 26 stems ha–1 year–1.
  • Saksa, Finnish Forest Research Institute, Suonenjoki Research Station, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@metla.fi (email)
article id 404, category Research article
William L. Mason, Colin Edwards, Sophie E. Hale. (2004). Survival and early seedling growth of conifers with different shade tolerance in a Sitka spruce spacing trial and relationship to understorey light climate. Silva Fennica vol. 38 no. 4 article id 404. https://doi.org/10.14214/sf.404
Keywords: seedling growth; light regime; stand density; irregular silviculture
Abstract | View details | Full text in PDF | Author Info
Alternative silvicultural systems to clearfelling are being adopted in Great Britain as a means of increasing the species and structural diversity of conifer plantation forests. One area where knowledge is lacking is the critical level of below-canopy light for survival and growth of young seedlings. This was investigated by planting seedlings of European larch Larix decidua (Mill.), Scots pine Pinus sylvestris L., Sitka spruce Picea sitchensis (Bong.(Carr.)), Douglas fir Pseudotsuga menziesii (Mirb.(Franco.)), and western hemlock Tsuga heterophylla (Raf. (Sarg.)) in a Sitka spruce plantation thinned to 3 different spacings. The incident light intensity beneath the canopy ranged from about 2 to over 60 per cent of full light. Planting in an adjoining open area provided an indication of growth under full light. Growth and survival of these seedlings were followed for 4 growing seasons. The highest seedling survival was found under the widest spacing and declined with closer spacing and lower light intensity. Only Douglas fir and western hemlock seedlings survived at the closest spacing, and in low percentages. The tallest seedlings of each species were found in the open grown conditions but survival was variable due to increased weed competition. Species-specific growth responses showed little difference under high light conditions but performance at low light was generally consistent with shade tolerance rankings in the literature except that Sitka spruce shade tolerance was slightly lower than expected. Minimum light requirements for these species increased from 10 to 30 per cent of full light with decreasing shade tolerance. Other studies of incident light in Sitka spruce plantations indicated that target basal areas in the range 25–30 m2 ha–1 are required if these light conditions are to be met, which suggests an irregular shelterwood system with frequent interventions should be favoured.
  • Mason, Forest Research, Northern Research Station, Roslin, Midlothian, Scotland, UK, EH25 9SY E-mail: bill.mason@forestry.gsi.gov.uk (email)
  • Edwards, Forest Research, Northern Research Station, Roslin, Midlothian, Scotland, UK, EH25 9SY E-mail: ce@nn.uk
  • Hale, Forest Research, Northern Research Station, Roslin, Midlothian, Scotland, UK, EH25 9SY E-mail: seh@nn.uk
article id 420, category Research article
Jukka Matero. (2004). Cost-effective measures for diffuse load abatement in forestry. Silva Fennica vol. 38 no. 3 article id 420. https://doi.org/10.14214/sf.420
Keywords: buffer strips; cost effectiveness; drainage maintenance; nutrient leaching
Abstract | View details | Full text in PDF | Author Info
This paper theoretically and empirically analyzes the design of cost-effective diffuse load abatement in forestry. Harvesting with related forest regeneration and drainage maintenance increases nutrient leaching, while riparian buffer strips and adjustments in drainage maintenance technology can be used to prevent this leaching. By utilizing a two-period model it is shown that cost-efficiency requires the establishment of a buffer strip system and a reduction in both current harvesting, and in drainage maintenance – if practized – relative to the private optimum to reflect their effects on water pollution. A simulation analysis was conducted to assess the magnitudes of the decision variables of the theoretical model, as well as to evaluate alternative technologies for the implementation and use of buffer strips and for the adjustment of drainage maintenance. The results for a representative forest holding in the southern half of Finland show that it is possible to considerably reduce total phosphorus leaching with minor cost.
  • Matero, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: jukka.matero@joensuu.fi (email)
article id 419, category Research article
Tero Heinonen, Timo Pukkala. (2004). A comparison of one- and two-compartment neighbourhoods in heuristic search with spatial forest management goals. Silva Fennica vol. 38 no. 3 article id 419. https://doi.org/10.14214/sf.419
Keywords: simulated annealing; Hero; tabu search; 2-optimal heuristic; spatial optimisation; random ascent
Abstract | View details | Full text in PDF | Author Info
This study presents a comparison of the performance of four heuristic techniques with one- and two-compartment neighbourhoods in harvest scheduling problems including a spatial objective variable. The tested heuristics were random ascent, Hero, simulated annealing and tabu search. All methods seek better solutions by inspecting the neighbourhood solutions, which are combinations that can be obtained by changing the treatment schedule in one (one-compartment neighbourhood) or two (two-compartment neighbourhood) compartments. The methods and neighbourhoods were examined in one artificial and four real landscapes ranging from 700 to 981 ha in size. The landscapes had 608 to 900 stand compartments, and the examined planning problems had 2986 to 4773 binary decision variables. The objective function was a multi-objective utility function. The spatial objective variable was the percentage of compartment boundary that joins two compartments, both of which are to be cut during the same 20-year period. The non-spatial objectives were net incomes of three consecutive 20-year management periods and the remaining growing stock volume at the end of the third 20-year period. In another problem formulation, the total harvest of the first 20-year period was used as an objective variable together with the spatial objective. The results showed that a two-compartment neighbourhood was systematically and often clearly better than a one-compartment neighbourhood. The improvements were greatest with the simplest heuristics, random ascent and Hero. Of the four heuristics, tabu search and simulated annealing proved to be the best methods, but with a two-compartment neighbourhood the differences between methods were negligible.
  • Heinonen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: th@nn.fi
  • Pukkala, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: timo.pukkala@joensuu.fi (email)
article id 418, category Research article
Tron Eid. (2004). Testing a large-scale forestry scenario model by means of successive inventories on a forest property. Silva Fennica vol. 38 no. 3 article id 418. https://doi.org/10.14214/sf.418
Keywords: forest planning; growth; validation; large-scale scenario analyses
Abstract | View details | Full text in PDF | Author Info
Modellers of large-scale forestry scenario models face numerous challenges. Information and sub-models from different disciplines within forestry, along with statistical and mathematical methodology, have to be considered. The individual biological sub-models (i.e. models for recruitment, growth and mortality) applied in large-scale forestry scenario models are in general well documented and extensively evaluated. However, evaluations by means of full-scale comparisons of observed and predicted values for continuous forest areas, where the totality of the large-scale forestry scenario model including interactions between sub-models and other parts of the model, are considered, have rarely been seen. The aim of the present work was to test the totality of the Norwegian large-scale forestry scenario model AVVIRK-2000, and thereby evaluate the applicability of the model for use in management planning. The test was done by means of successive inventories and accurate recordings of treatments over a period of 30 years for a property comprising 78.5 ha forest-land. Seen in the perspective of management planning, the differences between observed and predicted values for potential harvest level, growing stock and growth were small, e.g. a difference between observed growing stock in year 2000 and growing stock in the same year predicted from 1970 of 2.6%. The model may therefore be applied for practical purposes without any fundamental changes or calibrations of the biological model basis. However, the present test should be seen as an example that failed to falsify the model, rather than a final validation. As long as the model is in practical use, further evaluations should continue and subsequent possible calibrations should be performed.
  • Eid, Agricultural University of Norway, Dept of Ecology and Natural Resource Management, P.O. Box 5003 NO-1432 Ås, Norway E-mail: tron.eid@ina.nlh.no (email)
article id 417, category Research article
Jori Uusitalo, Sampsa Kokko, Veli-Pekka Kivinen. (2004). The effect of two bucking methods on Scots pine lumber quality. Silva Fennica vol. 38 no. 3 article id 417. https://doi.org/10.14214/sf.417
Keywords: harvesting; mechanized logging; tree bucking
Abstract | View details | Full text in PDF | Author Info
Modern harvesters are equipped with measurement and bucking optimization systems able not only to continuously measure the length and diameter of the stem but also to predict the profile of the unknown part of a stem and to calculate the optimal cross-cutting points for the whole stem. So far, tree-bucking optimization in the Nordic countries has been efficiently applied only with spruce because the quality of pine and birch varies much more both within a stem and between stems. Since limitations in the measuring equipment mean that the presence and position of grade limits as well as additional defects in the stem will normally have to be detected and estimated manually. Consequently, optimization works inefficiently because the harvester operator is continuously forced to disregard the cutting suggestions supplied by the harvester’s automatic system. This paper presents the outcome of research intended to define how change from the current quality bucking principle to automatic bucking affects lumber quality. The study is based on field experiments and test sawing data on 100 Scots pine (Pinus sylvestris) stems from southwestern Finland in 2001. Automatic bucking does not markedly lower the amount of good-quality lumber compared to quality bucking. Since automatic bucking inevitably leads to log distribution that matches the length requirements of customers better, it may be regarded as appropriate for these harvesting conditions.
  • Uusitalo, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: jori.uusitalo@joensuu.fi (email)
  • Kokko, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: sk@nn.fi
  • Kivinen, University of Helsinki, Department of Forest Resource Management, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: vpk@nn.fi
article id 416, category Research article
Glen Murphy, John G. Firth, Malcolm F. Skinner. (2004). Long-term impacts of forest harvesting related soil disturbance on log product yields and economic potential in a New Zealand forest. Silva Fennica vol. 38 no. 3 article id 416. https://doi.org/10.14214/sf.416
Keywords: Pinus radiata; harvesting; compaction; tree growth; litter removal
Abstract | View details | Full text in PDF | Author Info
The effect of soil disturbance (litter removal, topsoil removal and compaction) from forest harvesting on the productivity, log product yields and economic potential of second-rotation Pinus radiata growing on a clay loam soil, was assessed in a long-term trial 21 years after planting. The results are projected forward to the expected harvest age of 28 years. Relative to control plots, average tree volume at 21 years was reduced by 8% in the plots where the litter had been removed and the topsoil had been compacted, and by up to 42% in the plots where the topsoil had been removed and the subsoil compacted. The “degree of compaction” did not have a significant effect on average tree volume in the plots where litter had been removed but did have a significant effect where the topsoil had been removed. Per tree economic potential was reduced to a greater extent (up to 60% loss in value) than average tree volume was reduced. This was largely due to changes in log product yield distribution. Projecting tree growth forward to the end of the rotation at age 28 indicated that the impacts of soil disturbance on tree growth, economic potential and log product yields are likely to be similar in relative terms to those found at age 21.
  • Murphy, Forest Engineering Department, Oregon State University, Corvallis, OR 97331, USA E-mail: glen.murphy@orst.edu (email)
  • Firth, Forest Research, Sala Street, Rotorua, New Zealand E-mail: jgf@nn.nz
  • Skinner, Forest Research, Sala Street, Rotorua, New Zealand E-mail: mfs@nn.nz
article id 415, category Research article
Sandhya Samarasinghe, Don Kulasiri. (2004). Stress intensity factor of wood from crack-tip displacement fields obtained from digital image processing. Silva Fennica vol. 38 no. 3 article id 415. https://doi.org/10.14214/sf.415
Keywords: Pinus radiata; wood; fracture toughness; stress intensity factor; digital image correlation; orthotropic fracture theory
Abstract | View details | Full text in PDF | Author Info
Stress intensity factor of radiata pine (Pinus radiata) in Tangential-Longitudinal opening mode was determined from crack-tip displacement fields obtained from digital image correlation in conjunction with orthotropic fracture theory. For lower loads, experiments agreed with the linear elastic fracture theory but for higher loads, stress intensity factor and load relationship was nonlinear. For 41% of the specimens tested, tip-displacement based stress intensity factor agreed with that based on the ASTM standard formula for lower loads but deviated for higher loads closer to failure. The tip displacement plots showed that the nonlinear behaviour is due to large displacements which we attributed to large plastic deformations and/or micro-cracking in this region. The other 59% specimens showed a similar trend except that the crack-tip based stress intensity factor was consistently higher than the value obtained from the standard formula. The fracture toughness from tip displacements was larger than the standard values for all specimens and the two were related by a logarithmic function with an R2 of 0.61. The study also established that fracture toughness increases with the angle of inclination of the original crack plane to the Radial Longitudinal plane.
  • Samarasinghe, Lincoln University, P.O. Box 84, Canterbury, New Zealand E-mail: ss@nn.nz
  • Kulasiri, Lincoln University, P.O. Box 84, Canterbury, New Zealand E-mail: kulasird@lincoln.ac.nz (email)
article id 414, category Research article
Markku Larjavaara, Timo Kuuluvainen, Heidi Tanskanen, Ari Venäläinen. (2004). Variation in forest fire ignition probability in Finland. Silva Fennica vol. 38 no. 3 article id 414. https://doi.org/10.14214/sf.414
Keywords: ignition; forest fuel moisture; forest fire; fire regime
Abstract | View details | Full text in PDF | Author Info
We examined climate-caused spatio-temporal variation of forest fire ignition probability in Finland based on empirical ignition experiments and 37 years of meteorological data from 26 meteorological stations scattered across Finland. First, meteorological data was used in order to estimate the variation in forest fuel moisture content with the model of the Finnish forest fire risk index. Second, based on data from empirical ignition experiments, fuel moisture content was linked with forest fire ignition probability. In southern Finland average forest fire ignition probability typically peaks in late May and early June, whereas in the northern part of the country the peak occurs at the end of June. There was a three-fold difference in the average annual ignition probability between the north-eastern part (3%) and south-western part of the country (9%). The observed differences in fire ignition probability suggest that the characteristics of the natural fire regime also vary considerably in the southern versus the northern part of the country.
  • Larjavaara, University of Helsinki, Dept of Forest Ecology, FI-00014 University of Helsinki, Finland E-mail: markku.larjavaara@helsinki.fi (email)
  • Kuuluvainen, University of Helsinki, Dept of Forest Ecology, FI-00014 University of Helsinki, Finland E-mail: tk@nn.fi
  • Tanskanen, University of Helsinki, Dept of Forest Ecology, FI-00014 University of Helsinki, Finland E-mail: ht@nn.fi
  • Venäläinen, University of Helsinki, Dept of Forest Ecology, FI-00014 University of Helsinki, Finland E-mail: av@nn.fi
article id 413, category Research article
Mika Nieminen, Timo Penttilä. (2004). Inorganic and organic phosphorus fractions in peat from drained mires in northern Finland. Silva Fennica vol. 38 no. 3 article id 413. https://doi.org/10.14214/sf.413
Keywords: tree growth; drained peatland forests; phosphorus availability; sequential fractionation; site type classification
Abstract | View details | Full text in PDF | Author Info
Soil samples from 15 eutrophic, 26 herb-rich, 15 tall-sedge, and 11 low-sedge drained peatland sites were analysed for easily soluble and aluminum, iron, and calcium bound phosphorus (P) using the Chang and Jackson sequential fractionation method. Compared to earlier investigations, where only total and easily soluble P contents (e.g. NH4OAc or dilute H2SO4 extractable P) in peat have been analysed, significantly higher differences between sites were observed. The eutrophic sites were characterized by four to six-fold greater Ca-bound organic P and two to three-fold greater Ca-bound inorganic P contents than on the other three site type groups, whereas the average Al-bound inorganic P content of the eutrophic sites was only one-third of that at the other site types. Substantial differences between sites were also observed for Fe-bound inorganic P, i.e. two to four-fold greater Fe-P contents were measured at the herb-rich sites compared with the other three site type groups. The stand volume growth in the 67 studied drained peatland sites correlated significantly with Al-bound organic P and Fe-bound inorganic and organic P. The study showed that a detailed fractionation and discrimination of different forms of soil P is important in increasing the understanding of the relationship between P availability and vegetation community types and stand growth on drained peatlands.
  • Nieminen, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: mika.nieminen@metla.fi (email)
  • Penttilä, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: tp@nn.fi
article id 412, category Research article
Xingliang Liu, Haiyun Xu, Frank Berninger, Olavi Luukkanen, Chunyang Li. (2004). Nutrient distribution in Picea likiangensis trees growing in a plantation in West Sichuan, Southwest China. Silva Fennica vol. 38 no. 3 article id 412. https://doi.org/10.14214/sf.412
Keywords: plantation; nutrient distribution; nutrient translocation; Picea likiangensis
Abstract | View details | Full text in PDF | Author Info
We measured nutrient distribution of Picea likiangensis (Franchet) E. Pritzel var. balfouriana trees growing in a plantation by field investigations, sample tree and plot harvest in West Sichuan, Southwest China. Based on the results in this study, the total biomass of plant compartments in plantation ecosystem was 114 829.1 kg ha–1. Tree, shrub, herb, bryophyte and litter layers accounted for 93.9%, 0.9%, 0.02%, 0.04%, 5.2%, respectively. The total biomass of tree layers was 107 817.1 kg ha–1. Needles, branches, stem wood, stem bark and roots accounted for 13.2%, 19.7%, 42.3%, 10.0% and 14.8%, respectively. The concentration of the nutrients was generally highest in the actively growing parts of the trees (e.g. needles) and lowest in the structural and not actively growing parts (e.g. stem wood). On the other hand, the concentrations of N, P, K and Mg were generally higher in the current year needles and branches than in the older needles and branches. These nutrient concentrations were also higher in the upper stem wood and bark than in the lower stem wood and bark, and in small roots than in large roots, whereas the opposite patterns were observed for the concentration of Ca in these compartments. The results will be helpful in understanding the nutrient behavior in a highly productive forest plantation and thereby providing decisive information for their sustainable management.
  • Liu, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O. Box 416, Chengdu 610041, P. R. China; Sichuan Academy of Forestry, Chengdu 610081, P. R. China E-mail: xl@nn.cn
  • Xu, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O.Box 416, Chengdu 610041, P. R. China E-mail: hx@nn.cn
  • Berninger, Département des sciences biologiques, Cp 8888 succ centre ville, Université du Québec à Montréal, Montréal (QC) H3C 3P8, Canada E-mail: fb@nn.ca
  • Luukkanen, Viikki Tropical Resources Institute, P.O. Box 28, FI-00014 University of Helsinki, Finland E-mail: ol@nn.fi
  • Li, Chengdu Institute of Biology, Chinese Academy of Sciences, P.O.Box 416, Chengdu 610041, P. R. China; Viikki Tropical Resources Institute, P.O. Box 28, FI-00014 University of Helsinki, Finland E-mail: licy@cib.ac.cn (email)
article id 430, category Research article
Mårten Hugosson, Fredrik Ingemarson. (2004). Objectives and motivations of small-scale forest owners; theoretical modelling and qualitative assessment. Silva Fennica vol. 38 no. 2 article id 430. https://doi.org/10.14214/sf.430
Keywords: forest management; small-scale forest owners; professional foresters; forest owners’ objectives; forest owners’ motivations; forest owners’ values
Abstract | View details | Full text in PDF | Author Info
Forest management changes with societal change, and it has been debated if economic development in society places material objectives in a less preferable position: it is assumed this is also the case as regards forest management. The aims of this study were to propose a theoretical model for empirical studies of objectives and motivations within this field and to depict motivations and objectives of small-scale forest owners in Sweden. Comparative literature studies were undertaken and qualitative methodology was used for the empirical studies. Firstly, to depict general trends among forest owners, interviews with professional foresters were conducted. Secondly, forest owners throughout Sweden were interviewed to compare the results of the interviews with the professional foresters on the motivations and objectives of small-scale forest owners. Within the literature, there were no consistent views on the subjective grounds for owning and managing small-scale forest estates. The proposed theoretical model originated from the cultural concept. Sets of interpretive and normative qualities were seen as underlying people’s actions, and such sets were related to basic values. The ‘objectives’ were clustered into groups creating four clusters i.e. ‘motivations’. The four motivations depicted were: Conservation; Utilities; Amenities and Economic Efficiency. The empirical results highlighted that the objectives and motivations of forest-owners covered a broad field and a move towards conservation interests was indicated. The theoretical model presented here is suggested a suitable tool for both depicting the motivations and objectives of forest owners and for making future comparisons.
  • Hugosson, Department of Forest Products and Markets, Swedish University of Agricultural Sciences, P.O. Box 7060, SE-750 07 Uppsala, Sweden E-mail: mh@nn.se
  • Ingemarson, Department of Forest Products and Markets, Swedish University of Agricultural Sciences, P.O. Box 7060, SE-750 07 Uppsala, Sweden E-mail: fredrik.ingemarson@spm.slu.se (email)
article id 429, category Research article
Juho Rantala. (2004). Optimizing the supply chain strategy of a multi-unit Finnish nursery company. Silva Fennica vol. 38 no. 2 article id 429. https://doi.org/10.14214/sf.429
Keywords: economies of scale; supply chain management; optimization; mixed integer programming; seedling production
Abstract | View details | Full text in PDF | Author Info
This paper introduces a capacitated mixed integer programming (CMIP) model for solving an integrated production-distribution system design problem (PDSDP) in the seedling supply chain management (SCM) of a multi-unit Finnish nursery company. The model was originally developed from a strategic perspective in which a company desires to evaluate the expansion or closure of its facilities. Nevertheless, the model is also used for solving operational and tactical level problems by applying applicable constraints. The data were collected from the company studied. The results proved that economies of scale could be exploited in seedling production more than the company does today; Compared to the company’s current supply chain strategy with 5 nursery units producing seedlings, when other supply chain strategies were applied the number of nursery units decreased by 2–4 units, and cost savings in the supply chain varied from 11.3% to 21.3%.
  • Rantala, Finnish Forest Research Institute, Suonenjoki Research Station, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: juho.rantala@metla.fi (email)
article id 428, category Research article
Jonny Andersson, Lars Eliasson. (2004). Effects of three harvesting work methods on Harwarder productivity in final felling. Silva Fennica vol. 38 no. 2 article id 428. https://doi.org/10.14214/sf.428
Keywords: forwarder; harvester; work method
Abstract | View details | Full text in PDF | Author Info
During the last ten years interest in the harwarder has increased, however, studies have concentrated on effects of technical improvements on machine productivity. It has been noted that there is a large potential to increase the productivity through development of suitable work methods. To find efficient work patterns for a harwarder with a turnable loading area, three different harvesting methods were studied in final felling. Three work methods were used. Method 1: the harwarder drove backwards into the stand making a strip road, strip road trees were felled and left on the ground, on the way out of the stand the harwarder cut and processed the trees on both sides of the machine directly into the loading area. Method 2: the harwarder drove forward along the edge of the cut, cutting and processing trees directly into the loading area. Method 3: the harwarder drove forward into the stand and cut and processed strip road trees and trees standing on both sides of the machine directly into the loading area. The most efficient work method was method 2 where the productivity was 13.0 m3 u.b. per E0h (cubic metre under bark per effective hour). The productivities for method 1 and 3 were 12.1 and 11.9 m3 u.b. per E0h, respectively. In addition to work method harwarder productivity was shown to be dependent on load volume, average tree size and hauling distance. The only work elements significantly affected by work methods were processing and movement during processing. The operator had only a few weeks to get used to the machine and even less time to practise on the work methods. Thus, it is probable that the productivity for the studied methods will increase with increasing work experience. Furthermore, as only three work methods were studied, there are still untested work methods. The potential to further improve harwarder work methods is considerable.
  • Andersson, Swedish University of Agricultural Sciences, Department of Silviculture, S-901 83 Umeå, Sweden E-mail: je@nn.se
  • Eliasson, Swedish University of Agricultural Sciences, Department of Silviculture, S-901 83 Umeå, Sweden E-mail: lars.eliasson@ssko.slu.se (email)
article id 427, category Research article
Scott A. Weyenberg, Lee E. Frelich, Peter B. Reich. (2004). Logging versus fire: how does disturbance type influence the abundance of Pinus strobus regeneration? Silva Fennica vol. 38 no. 2 article id 427. https://doi.org/10.14214/sf.427
Keywords: disturbance refugia; interspecific competition; Minnesota; seedling dispersion; seed source; white pine
Abstract | View details | Full text in PDF | Author Info
Eastern white pine (Pinus strobus) has decreased in abundance over the past century throughout the Great Lakes Region of North America, but the relative constraints placed on recruitment under contrasting disturbance regimes are not well understood. The objectives of this study were to determine the extent to which white pine could invade areas recently disturbed by fire or logging (within 10–28 years), and assess the relative limitations placed on recruitment by seed supply, microsite habitat, and competition. We compared white pine regeneration on 61 sites disturbed by fire or logging that were adjacent to intact mature stands that provided a seed source. White pine seedling and sapling densities declined with increasing distance from a seed source, and the rate of decrease was determined by the interaction between seed supply and variation in number and quality of safe sites. For a given combination of seed source and site, white pine seedlings were three times more abundant on burned than logged sites. White pine seedlings grew into the sapling size class more often on burned than logged sites due to lower shrub cover on burned sites. At 25 years after disturbance, regeneration densities of white pine sufficient to achieve eventual future dominance occurred up to 80 m and 20 m from the edge of mature white pine stands after fire and logging, respectively. To attain a similar level of white pine stocking after disturbance, three to four times as many patches of mature white pine need to be left after logging than after fire.
  • Weyenberg, University of Minnesota, Department of Forest Resources, 1530 Cleveland Ave. N., St. Paul, MN 55108, USA E-mail: saw@nn.us
  • Frelich, University of Minnesota, Department of Forest Resources, 1530 Cleveland Ave. N., St. Paul, MN 55108, USA E-mail: freli001@umn.edu (email)
  • Reich, University of Minnesota, Department of Forest Resources, 1530 Cleveland Ave. N., St. Paul, MN 55108, USA E-mail: pbr@nn.us
article id 426, category Research article
Uwe Schmitt, Risto Jalkanen, Dieter Eckstein. (2004). Cambium dynamics of Pinus sylvestris and Betula spp. in the northern boreal forest in Finland. Silva Fennica vol. 38 no. 2 article id 426. https://doi.org/10.14214/sf.426
Keywords: birch; boreal forest; Scots pine; cambium dynamics; wood formation; pinning technique; light microscopy
Abstract | View details | Full text in PDF | Author Info
Wood formation dynamics of pine and birch along a south-north transect in Finnish Lapland were determined by the pinning technique. For all trees at all sites a more or less sigmoid shape of the wood formation intensity is characteristic with a slow beginning, a faster growth in the middle and a decreasing activity towards the end of the vegetation period. Wood formation of pine started at sites 1–3 (southern sites) in the second week of June and at sites 4 and 5 (northern sites) only in the last week of June, whereas wood formation ended within the first half of August. Wood formation of birch started in the second half of June and ended around the beginning of August. First cells were laid down by pine and birch when the temperature sum had reached the level of 85 to 90 degree days and 110 to 120 degree days, respectively. The intensity of wood formation in pine was highest in July, in birch within two weeks in the middle of July. Wood formation in pine lasted for about seven weeks at the southernmost and about six weeks at the northernmost site. In birch, the duration of wood formation was about five weeks at the southernmost site and around three weeks at the other sites.
  • Schmitt, Federal Research Centre for Forestry and Forest Products, Institute for Wood Biology and Wood Protection, and University of Hamburg, Chair for Wood Biology, Leuschnerstr. 91, P. O. Box 800209, D-21002 Hamburg, Germany E-mail: u.schmitt@holz.uni-hamburg.de (email)
  • Jalkanen, Finnish Forest Research Institute, Rovaniemi Research Station, Box 16, FI-96301 Rovaniemi, Finland E-mail: rj@nn.fi
  • Eckstein, Federal Research Centre for Forestry and Forest Products, Institute for Wood Biology and Wood Protection, and University of Hamburg, Chair for Wood Biology, Leuschnerstr. 91, P. O. Box 800209, D-21002 Hamburg, Germany E-mail: de@nn.de
article id 425, category Research article
Petteri Vanninen. (2004). Allocation of above-ground growth in Pinus sylvestris – impacts of tree size and competition. Silva Fennica vol. 38 no. 2 article id 425. https://doi.org/10.14214/sf.425
Keywords: Scots pine; allocation; growth efficiency
Abstract | View details | Full text in PDF | Author Info
The effect of tree age, size and competition on above ground growth allocation was studied with 69 Pinus sylvestris trees. Competition was described by tree-level indicators (needle density, crown ratio and height-diameter ratio). The stem, branch and needle growth were determined by stem and branch radial increments and tree level biomass analysis. Combined growth of compartments was strongly correlated with needle mass. Furthermore, tree age, size and competition indicators affected the allocation of growth among the compartments. The allocation of growth to stem and needle increased with tree age and size while the allocation of growth to branch decreased. The increasing crown ratio increased allocation of growth to branches. The combined growth of the components and separate growth of needles, branches and stem were related to needle mass. However, competition and tree size were significant additional explanatory variables when the stem, branch and needle growth were estimated according to needle mass. The growth efficiency increased with relative tree height and decreased with increasing needle density.
  • Vanninen, University of Helsinki, Department of Forest Ecology; mailing address: SAIMA – Centre for Environmental Sciences, Linnankatu 11, FIN-57130 Savonlinna, Finland E-mail: petteri.vanninen@helsinki.fi (email)
article id 424, category Research article
Mats Warensjö, Göran Rune. (2004). Stem straightness and compression wood in a 22-year-old stand of container-grown Scots pine trees. Silva Fennica vol. 38 no. 2 article id 424. https://doi.org/10.14214/sf.424
Keywords: Pinus sylvestris; stem form; eccentricity; reaction wood
Abstract | View details | Full text in PDF | Author Info
The distribution of compression wood in relation to eccentric growth and development of stem straightness was studied in a 22-year-old Scots pine (Pinus sylvestris L.) stand in central Sweden that was established with container-grown seedlings. Stem straightness was measured on the same 440 trees in 1986 and 1997. The number of stems with straight base sections increased from 60% in 1986 to 89% in 1997. Measurements of 72 sample trees in 2001 showed that 96% of the trees had developed straight stem bases. External geometry data of the logs was obtained with a Rema 3D log scanner. A sub-sample of 16 trees was randomly selected for analysis of compression wood distribution and eccentricity measurements. From each tree, 11 discs were cut at every 60 cm along the stem. All discs, except one, contained compression wood. Compression wood and pith eccentricity was most pronounced near the stem base but not significantly correlated to basal sweep. Severe compression wood content was correlated to pith eccentricity and bow height. In general, correlations were better for the basal sections of the logs. Even though most trees were straight, they contained large amounts of compression wood. It is evident that eccentric growth and compression wood formation play major roles in the development of stem straightness. In several stems, a spiral compression wood distribution pattern was found. Reasons for this are discussed.
  • Warensjö, Department of Forest Products and Markets, Swedish University of Agricultural Sciences, SE-901 83 Umeå, Sweden E-mail: mats.warensjo@spm.slu.se (email)
  • Rune, Department of Forestry and Wood Technology, Dalarna University, SE-776 98 Garpenberg, Sweden E-mail: gr@nn.se
article id 423, category Research article
Olle Rosenberg, Staffan Jacobson. (2004). Effects of repeated slash removal in thinned stands on soil chemistry and understorey vegetation. Silva Fennica vol. 38 no. 2 article id 423. https://doi.org/10.14214/sf.423
Keywords: carbon; nitrogen; thinning; base cations; soil chemistry; understorey vegetation; whole-tree harvesting
Abstract | View details | Full text in PDF | Author Info
The increased interest in harvesting logging residues as a source of bio-energy has led to concerns about the potentially adverse long-term impact of the practice on site productivity. The aim of this study was to examine the effects on soil chemistry (pH, C, N and AL-extractable P, K, Ca and Mg) in three different soil layers (FH, 0–5 cm and 5–10 cm mineral soil) and understorey vegetation after the second removal of logging residues in whole-tree thinned stands. The study was performed at four different sites, established in the period 1984–87, representing a range of different climatic and soil conditions: a very fertile Norway spruce (Picea abies (L.) Karst.) site in south-western Sweden and three Scots pine (Pinus sylvestris L.) sites located in south, south-central and central Sweden. The effects of whole-tree thinning on soil chemistry and understorey vegetation were generally minor and variable. Across all sites the concentrations of Ca and Mg were significantly lower when slash was removed.
  • Rosenberg, Skogforsk – The Forestry Research Institute of Sweden, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: olle.rosenberg@skogforsk.se (email)
  • Jacobson, Skogforsk – The Forestry Research Institute of Sweden, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: sj@nn.se
article id 422, category Research article
Mika Nieminen. (2004). Export of dissolved organic carbon, nitrogen and phosphorus following clear-cutting of three Norway spruce forests growing on drained peatlands in southern Finland. Silva Fennica vol. 38 no. 2 article id 422. https://doi.org/10.14214/sf.422
Keywords: hydrochemistry; forest regeneration; peatland forestry; leaching; nutrient losses
Abstract | View details | Full text in PDF | Author Info
The effect of clear-cutting on the concentrations of dissolved organic carbon (DOC), organic nitrogen (DON), NH4+, NO3–, and P in outflow water from three productive, Norway spruce dominated drained peatlands (RCC, VCC-1, VCC-2) were studied. Changes in runoff and transport loads (concentration x runoff) at two of the catchments during the frost-free period are also presented. Approximately 40% of the area was cut at RCC and VCC-2, and 72% at VCC-1. The volume removed was 250 m3 ha–1 at RCC, 259 m3 ha–1 at VCC-1, and for VCC-2, 317 m3 ha–1. The mean annual increase in outflow concentrations of DOC during the first four years after clear-cutting was 9.0 mg l–1 at RCC, 22.8 mg l–1 at VCC-1 and 8.4 mg l–1 at VCC-2. Corresponding increases in the forms of nitrogen were: 0.23, 0.51 and 0.16 mg DON l–1; 0.06, 0.31 and 0.04 mg NH4+-N l–1; and 0.05, 0.12 and 0.22 mg NO3–-N l–1. Clear-cutting did not significantly (p > 0.05) increase P concentrations. The increase in non-frost season runoff over the first three years after clear-cutting was 107 mm at RCC and 207 mm at VCC-1. The export loads of DOC during the non-frost season increased by 80 kg ha–1 at RCC and by 184 kg ha–1 at VCC-1 over the first three years. Corresponding increases for the other studied solutes were: 1.78 and 3.98 kg DON ha–1; 0.39 and 1.49 kg NH4+-N ha–1; 0.45 and 0.48 kg NO3–-N ha–1, and 0.09 and 0.06 kg P ha–1. The study demonstrated that clear-cutting may significantly increase the export of DOC and different forms of nitrogen from drained productive peatlands while only small increases in phosphorus export may occur.
  • Nieminen, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: mika.nieminen@metla.fi (email)
article id 439, category Research article
Christina Lundgren. (2004). Microfibril angle and density patterns of fertilized and irrigated Norway spruce. Silva Fennica vol. 38 no. 1 article id 439. https://doi.org/10.14214/sf.439
Keywords: ring width; growth rate; juvenile wood; MoE; X-ray diffraction
Abstract | View details | Full text in PDF | Author Info
Two Norway spruce nutrient optimisation trials, one in the north of Sweden and one in the south, were used to study the effects of intensive growth and fertilization on wood density and microfibril angle. Three different treatments and a control were available; daily irrigation, daily liquid fertilization and solid fertilization. The nutrient optimisation was based on foliage analysis and the solid fertilization essentially comprised the same amount of nutrients but was applied annually in solid form. Measurements of density and microfibril angle (MFA) were performed using X-ray diffraction. Growth rate, expressed as a transformation of annual ring width, was very important at the southern site when the effect of cambial maturation had been taken into account. Effects of both fertilization and irrigation remained strong and significant for density, and irrigation was a significant factor explaining MFA. At the northern site distance from pith was the dominant factor but the effect of growth rate was also strong and the treatment effect was significant for both density and MFA. The combination of higher MFA and decrease in density for fertilized trees resulted in a lower calculated strength of the wood. An over 100% increase in ring width only corresponded to approximately a 20% decrease in wood density and the production of wood dry matter was hence increased by treatments.
  • Lundgren, SLU, Dept. of Forest Products and Markets, P.O. Box 7060, SE-750 07 Uppsala, Sweden E-mail: christina.lundgren@spm.slu.se (email)
article id 438, category Research article
Christina Lundgren. (2004). Cell wall thickness and tangential and radial cell diameter of fertilized and irrigated Norway spruce. Silva Fennica vol. 38 no. 1 article id 438. https://doi.org/10.14214/sf.438
Keywords: cell wall thickness; tangential tracheid diameter; radial tracheid diameter; tracheid dimensions; nutrient optimisation; juvenile wood
Abstract | View details | Full text in PDF | Author Info
Two Norway spruce nutrient trials were used to evaluate the effects of fertilization and irrigation on transverse tracheid dimensions. Three different treatments and a control (C) were used; daily irrigation (I), daily liquid fertilization (IL) and an annual solid fertilization (F). The nutrient optimisation was based on foliage analysis and both liquid and solid fertilization essentially comprised the same amount of nutrients but the latter was applied annually in solid form. The cell measurements; cell wall thickness, radial and tangential cell widths, were obtained using image analysis (SilviScan at CSIRO, Melbourne, Australia). Mean annual cell wall thickness was decreased by fertilization (F and IL) on both sites whereas no effect of the irrigation on wall thickness could be detected. Radial cell width was increased by treatment at Flakaliden but at Asa the effect of irrigation and fertilization was reversed when the data structure i.e. development from pith and out and annual ring width was taken into account. Tangential cell width was not significantly affected by treatment at Flakaliden. At Asa fertilization caused a small increase on tangential cell width. Ring width was positively affected by treatment and is an important factor explaining the effects on primarily cell wall thickness and radial cell width.
  • Lundgren, SLU, Dept. of Forest Products and Markets, P.O. Box 7060, SE-750 07 Uppsala, Sweden E-mail: christina.lundgren@spm.slu.se (email)
article id 437, category Research article
Raffaele Spinelli, Philip M. O. Owende, Shane M. Ward, Maximiano Tornero. (2004). Comparison of short-wood forwarding systems used in Iberia. Silva Fennica vol. 38 no. 1 article id 437. https://doi.org/10.14214/sf.437
Keywords: Eucalyptus; forwarding; site disturbance; cut-to-length harvesting
Abstract | View details | Full text in PDF | Author Info
  • Spinelli, CNR - Timber and Tree Institute, Via Madonna del Piano - Pal. F, 50019 Sesto Fiorentino, Italy; University College Dublin, Ireland E-mail: spinelli@ivalsa.cnr.it (email)
  • Owende, Dept. of Agricultural and Food Engineering, University College Dublin, Earlsfort Terrace, Dublin 2, Ireland E-mail: pmoo@nn.ie
  • Ward, Dept. of Agricultural and Food Engineering, University College Dublin, Earlsfort Terrace, Dublin 2, Ireland E-mail: smw@nn.ie
  • Tornero, Servicios Forestales SA, Ctra. SE-184 Km 0.63, E-41970 Santiponce, Sevilla, Spain E-mail: mt@nn.es
article id 436, category Research article
Martti Varmola, Hannu Salminen, Mauri Timonen. (2004). Thinning response and growth trends of seeded Scots pine stands at the arctic timberline. Silva Fennica vol. 38 no. 1 article id 436. https://doi.org/10.14214/sf.436
Keywords: Pinus sylvestris; thinning; growth and yield; northern timberline
Abstract | View details | Full text in PDF | Author Info
Growth patterns and reactions of Scots pine (Pinus sylvestris L.) to thinning in extremely harsh climatic conditions were studied in two seeded Scots pine stands located on the arctic timberline. Coniferous trees usually do not form closed stands at the timberline, but occur only in scattered tree groups. The trial was established in two stands in 1985–1986 when the trees were at an age of 47 and 56 years and an average dominant height of 6.0–6.9 m. The trial was remeasured in 1998. The thinning treatments reduced the stem number for five different levels; final density of 300, 550, 800, 1050, and 1300 stems ha–1 and unthinned. The experiment had a randomised block design with four replications in each stand. The increased growing space provided by thinning accelerated diameter growth after a delay of 2–3 years. The differences between the radial growth of the thinning treatments were very clear during the whole 13- to 14-year observation period. Annual increment of the mean diameter was regularly the higher, the larger the spacing. Dominant diameter was less influenced by treatments. There were no significant differences in dominant height between any of the treatments. Both basal area and volume were regularly the greater the higher the stem number was. Even a relatively light thinning had a distinct positive effect on tree growth, i.e. not carrying out thinning resulted in a production loss of merchantable wood. According to the results, seeded stands on the arctic timberline can grow surprisingly well in favourable conditions and reach a dominant height of 12–14 m in 100 years and a mean annual increment of 1.0–1.5 m3 ha–1 y–1 over a rotation period of 130–160 years. Based on increment figures and thinning reactions, a spacing of ca. 1000 stems ha–1 can be recommended.
  • Varmola, The Finnish Forest Research Institute, Rovaniemi Research Station, P.O.Box 16, FIN-96301 Rovaniemi, Finland E-mail: martti.varmola@metla.fi (email)
  • Salminen, The Finnish Forest Research Institute, Rovaniemi Research Station, P.O.Box 16, FIN-96301 Rovaniemi, Finland E-mail: hs@nn.fi
  • Timonen, The Finnish Forest Research Institute, Rovaniemi Research Station, P.O.Box 16, FIN-96301 Rovaniemi, Finland E-mail: mt@nn.fi
article id 435, category Research article
Rebecca Ralston, Joseph Buongiorno, Jeremy S. Fried. (2004). Potential yield, return, and tree diversity of managed, uneven-aged Douglas-fir stands. Silva Fennica vol. 38 no. 1 article id 435. https://doi.org/10.14214/sf.435
Keywords: diversity; simulation; uneven-aged management; Douglas-fir; WestPro; economics
Abstract | View details | Full text in PDF | Author Info
The effects of different management regimes on uneven-aged Douglas-fir stands in the Pacific Northwest of the United States were predicted with a simulation model. Management alternatives were defined by residual stand structure and cutting cycle. The residual stand structure was set by basal area–diameter-q-ratio (BDq) distributions, diameter-limit cuts (assuming concurrent stand improvement), or the current diameter distribution. Cutting cycles of 10 or 20 years were applied for 200 years. The current diameter distribution was defined as the average of the uneven-aged Douglas-fir stands sampled in the most recent Forest Inventory and Analysis conducted in Oregon and Washington. Simulation results were compared in terms of financial returns, timber productivity, species group diversity (hardwoods vs softwoods), size class diversity, and stand structure. Other things being equal, there was little difference between 10- and 20-year cutting cycles. The highest financial returns were obtained with either a 58.4 cm diameter-limit cut, or a BDq distribution with 8.4 m2 of residual basal area, a 71.1 cm maximum diameter, and a q-ratio of 1.2. Using the current stand state as the residual distribution was the best way to obtain high tree size diversity, and high species group diversity. Several uneven-aged regimes gave net present values comparable to that obtained by converting the initial, uneven-aged stand to an even-aged, commercially thinned, plantation.
  • Ralston, Dept of Forest Ecology and Management, University of Wisconsin, Madison, WI 53706, USA E-mail: rr@nn.us
  • Buongiorno, Dept of Forest Ecology and Management, University of Wisconsin, Madison, WI 53706, USA E-mail: jbuongio@facstaff.wisc.edu (email)
  • Fried, USDA Forest Service, Pacific Northwest Forest Experiment Station, Forest Inventory and Analysis Program, P.O. Box 3890, Portland, OR 97208, USA E-mail: jsf@nn.us
article id 434, category Research article
Igor Drobyshev, Mats Niklasson, Per Angelstam. (2004). Contrasting tree-ring data with fire record in a pine-dominated landscape in the Komi Republic (Eastern European Russia): recovering a common climate signal. Silva Fennica vol. 38 no. 1 article id 434. https://doi.org/10.14214/sf.434
Keywords: boreal; climate; earlywood; latewood; chronology; seasonal dating; weather control; natural disturbance
Abstract | View details | Full text in PDF | Author Info
For the period 1420–1960 we contrasted fire events reconstructed at 14 sites distributed over a 50 km x 50 km area in the central part of the Komi Republic (European Russia) with a set of tree-ring width chronologies of Scots pine (Pinus sylvestris L.), developed for the same area. Our aim was to infer common climatic information contained in tree-ring variables and independently dated fire events with the help of a superposed epoch analysis. The strongest weather–growth link was shown for the latewood width, which was positively correlated with the temperature in April–May and July–August of the current growth season and with previous year precipitation in July–August. Earlywood width was positively affected by previous year precipitation in May and November. The relationship between yearly ring variables and multiple-site fire events was dependent on the seasonal timing of fire events as recorded in the scars. In years with early-season fires (which made up 37% of all fires dated with seasonal resolution) total ring width was significantly narrower. In years with late-season fires (63%) total ring width, earlywood, and latewood width were significantly wider. Years with late-season fires tended to be associated with local highs of the latewood width chronologies over 1400–1960, which implied a link between decadal-scale climate variation and fire regime of the area.
  • Drobyshev, SUFOR Project, Department of Plant Ecology and Systematics, Ecology Building, Sölvegatan 37, Lund University, SE-223 62 Lund, Sweden E-mail: igor.drobyshev@ekol.lu.se (email)
  • Niklasson, SUFOR Project, Southern Swedish Forest Research Centre, SLU, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: mn@nn.se
  • Angelstam, Grimsö Wildlife Research Station, Department of Conservation Biology, Forest Faculty, SLU, SE-730 91 Riddarhyttan, Sweden E-mail: pa@nn.se
article id 433, category Research article
Raija Laiho, Timo Penttilä, Jukka Laine. (2004). Variation in soil nutrient concentrations and bulk density within peatland forest sites. Silva Fennica vol. 38 no. 1 article id 433. https://doi.org/10.14214/sf.433
Keywords: thinning; drainage; peat soil; spatial variation; nutrients; bulk density; sampling; variance components
Abstract | View details | Full text in PDF | Author Info
The within-site variability of soil characteristics on sites with different soil types remains poorly quantified, although this information is crucial for the success of research on soil properties, and especially for monitoring soil properties over time. We used coefficients of variation and multilevel variance component models to examine the within-site variation of soil (0–30 cm) mineral nutrient concentrations (P, K, Ca, Mg, Fe, mg g–1; Mn, Zn, mg g–1) and bulk density (kg m–3) on boreal deep-peat sites. We then evaluated the reliability of the site-level estimates (sample means) obtained using different sampling intensities (numbers of samples per site). Our 11 sites represented a single original site type within the oligotrophic nutrient level. Two of the sites were undrained while the rest had been drained for forestry at different points in time. Overall, P concentrations showed the smallest and Mn concentrations the largest within-site variation. The sampling depth contributed more than 50% of the total variance in all other characteristics except the concentrations of P and Fe, and bulk density. The variance proportions of peatland basin, site (within basin), and sampling location (within site) varied by sampling depth for most soil characteristics. The estimates obtained when using a certain number of samples per site were always more reliable for the 0–30 cm layer’s composite samples than for any single 10-cm layer at any depth sampled. On average, it was found that between 4 (P) and some 200 (Mn) samples per site would be needed for the estimates to have a theoretical 10% maximum deviation.
  • Laiho, Univ. of Helsinki, Dept. of Forest Ecology, Peatland Ecology Group, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: raija.laiho@helsinki.fi (email)
  • Penttilä, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: tp@nn.fi
  • Laine, Univ. of Helsinki, Dept. of Forest Ecology, Peatland Ecology Group, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: jl@nn.fi
article id 432, category Research article
Bo Långström, Claes Hellqvist, Jan Cedervind. (2004). Comparison of methods for estimation of needle losses in Scots pine following defoliation by Bupalus piniaria. Silva Fennica vol. 38 no. 1 article id 432. https://doi.org/10.14214/sf.432
Keywords: Pinus sylvestris; pine looper; undefoliated foliage; needle biomass; CIR-pictures
Abstract | View details | Full text in PDF | Author Info
In 1996, ca. 7000 hectares of pine forests at Hökensås in SW Sweden were defoliated by the pine looper, Bupalus piniara (L.) (Lepidoptera. Geometridae). Following an aerial damage survey using CIR (colour infra red) photography, and estimation of pupal densities in the soil, ca 4000 ha of the most defoliated pine stands were sprayed in early August 1997 with Bacillus thuringiensis var. kurstaki. The control operation was succeessful but probably redundant, as no further defoliation occurred in unsprayed reference areas. In order to assess defoliation levels in different damage classes for later growth loss studies, 47 circular study plots were laid out in pine stands representing different damage and age classes. The remaining foliage was recorded for each tree using the following classes: 0, 10, 30, 50, 70, 90 and 100%. The defoliation levels in 1996 were estimated by disregarding the 1997 needle age class. Thirteen ca. 40-year-old sample trees representing different damage classes were felled, and the remaining foliage of all branches was estimated by needle age class using the above-mentioned scale. One branch in each of the whorls 1996, 1991, 1986 and 1981 was sampled and its needle dry weight was determined. The sample branch data confirmed the field observations that virtually no additional defoliation took place in 1997. The damage classes estimated from the CIR-pictures only agreed with the field damage estimates at the higher end of the damage scale. In contrast, the field estimate correlated well with plot means derived from tree-wise estimates (R2 = 0.93), and with with the calculated needle biomasses per tree (R2 = 0.90). Thus, the field damage classification was supported by the more detailed defoliation estimates, and hence forms a relevant basis for later growth loss studies.
  • Långström, SLU, Dept. of Entomology, P.O.Box 7044, S-750 07 Uppsala, Sweden. Fax +46 18 672 890 E-mail: bo.langstrom@entom.slu.se (email)
  • Hellqvist, SLU, Dept. of Entomology, P.O.Box 7044, S-750 07 Uppsala, Sweden. Fax +46 18 672 890 E-mail: ch@nn.se
  • Cedervind, SLU, Dept. of Entomology, P.O.Box 7044, S-750 07 Uppsala, Sweden. Fax +46 18 672 890 E-mail: jc@nn.se
article id 431, category Research article
Pauline Stenberg, Miina Rautiainen, Terhikki Manninen, Pekka Voipio, Heikki Smolander. (2004). Reduced simple ratio better than NDVI for estimating LAI in Finnish pine and spruce stands. Silva Fennica vol. 38 no. 1 article id 431. https://doi.org/10.14214/sf.431
Keywords: Landsat ETM ; Leaf Area Index; spectral vegetation indices; boreal coniferous forests
Abstract | View details | Full text in PDF | Author Info
Estimation of leaf area index (LAI) using spectral vegetation indices (SVIs) was studied based on data from 683 plots on two Scots pine and Norway spruce dominated sites in Finland. The SVIs studied included the normalised difference vegetation index (NDVI), the simple ratio (SR), and the reduced simple ratio (RSR), and were calculated from Landsat ETM images of the two sites. Regular grids of size 1 km2 with gridpoints placed at 50 m intervals were established at the sites and measurements of LAI using the LAI-2000 instrument were taken at the gridpoints. SVI-LAI relationships were examined at plot scale, where the plots were defined as circular areas of radius 70 m around each gridpoint. Plotwise mean LAI was computed as a weighted average of LAI readings taken around the gridpoints belonging to the plot. Mean LAI for the plots ranged from 0.36 to 3.72 (hemisurface area). All of the studied SVIs showed fair positive correlation with LAI but RSR responded more dynamically to LAI than did SR or NDVI. Especially NDVI showed poor sensitivity to changes in LAI. RSR explained 63% of the variation in LAI when all plots were included (n = 683) and the coefficient of determination rose to 75% when data was restricted to homogeneous plots (n = 381). Maps of estimated LAI using RSR showed good agreement with maps of measured LAI for the two sites.
  • Stenberg, Department of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: pauline.stenberg@helsinki.fi (email)
  • Rautiainen, Department of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: mr@nn.fi
  • Manninen, Finnish Meteorological Institute, Meteorological research, Ozone and UV radiation research, P.O. Box 503, FIN-00101 Helsinki, Finland E-mail: tm@nn.fi
  • Voipio, Finnish Forest Research Institute, Suonenjoki Research Station, FIN-77600 Suonenjoki, Finland E-mail: pv@nn.fi
  • Smolander, Finnish Forest Research Institute, Suonenjoki Research Station, FIN-77600 Suonenjoki, Finland E-mail: hs@nn.fi
article id 488, category Research article
Gürsel Colakoglu, Semra Colak, Ismail Aydin, Umit C. Yildiz, Sibel Yildiz. (2003). Effect of boric acid treatment on mechanical properties of laminated beech veneer lumber. Silva Fennica vol. 37 no. 4 article id 488. https://doi.org/10.14214/sf.488
Keywords: mechanical properties; LVL; boric acid; beech veneer
Abstract | View details | Full text in PDF | Author Info
Laminated veneer lumber (LVL) made from beech wood veneers treated with boric acid by using dipping method was tested for some mechanical properties following different standards. The values for treated LVL varied around the mean values of untreated LVL panels for static bending strength in the grain direction and modulus of elasticity; decreased for compression and splitting strengths, perpendicular to the grain; increased for compression strength parallel to the grain, and for Brinell hardness and pull-out strength of screw, perpendicular to surface.
  • Colakoglu, Karadeniz Technical University, Faculty of Forestry, Forest Industry Engineering Section, 61080 Trabzon, Turkey E-mail: gursel@ktu.edu.tr (email)
  • Colak, Karadeniz Technical University, Faculty of Forestry, Forest Industry Engineering Section, 61080 Trabzon, Turkey E-mail: sc@nn.tr
  • Aydin, Karadeniz Technical University, Faculty of Forestry, Forest Industry Engineering Section, 61080 Trabzon, Turkey E-mail: ia@nn.tr
  • Yildiz, Karadeniz Technical University, Faculty of Forestry, Forest Industry Engineering Section, 61080 Trabzon, Turkey E-mail: ucy@nn.tr
  • Yildiz, Karadeniz Technical University, Faculty of Forestry, Forest Industry Engineering Section, 61080 Trabzon, Turkey E-mail: sy@nn.tr
article id 487, category Research article
Åsa Gustafsson. (2003). Logistic services as a competitive means – segmenting the retail market for softwood lumber. Silva Fennica vol. 37 no. 4 article id 487. https://doi.org/10.14214/sf.487
Keywords: service requirements; retail segments; softwood lumber
Abstract | View details | Full text in PDF | Author Info
Softwood lumber has been considered traditionally as commodity. Subsequently brand names and trademarks were of subordinate value and competition was based on price. Recently, retailers have grown substantially and are forcing their suppliers to improve the production and delivery of products. As retailers are getting more diverse and powerful, suppliers are forced to adapt to the retailers’ service requirements. The new situation also brings opportunities for the sawmills to develop their competitive advantage. The retail industry is continuously changing, and in order for sawmills to develop and offer what retailers are asking for, it is necessary that they understand and interpret retailers’ requirements correctly. One way for sawmills to be successful is to develop accurate service elements and to use the service elements as a segmentation base in order to structure their customer base. This study shows that retailers place considerable emphasis on delivery and value-added logistical services. It generates three hypotheses concerning the following potential retail segments; turnover, category, and customer base.
  • Gustafsson, School of Industrial Engineering (IPS), Växjö University, SE-351 95 Växjö, Sweden E-mail: asa.gustafsson@ips.vxu.se (email)
article id 486, category Research article
Arto Haara. (2003). Comparing simulation methods for modelling the errors of stand inventory data. Silva Fennica vol. 37 no. 4 article id 486. https://doi.org/10.14214/sf.486
Keywords: measurement error; simulation; stand-level inventory; non-parametric estimation; Monte Carlo methods
Abstract | View details | Full text in PDF | Author Info
Forest management planning requires information about the uncertainty inherent in the available data. Inventory data, including simulated errors, are infrequently utilised in forest planning studies for analysing the effects of uncertainty on planning. Usually the errors in the source material are ignored or not taken into account properly. The aim of this study was to compare different methods for generating errors into the stand-level inventory data and to study the effect of erroneous data on the calculation of specieswise and standwise inventory results. The material of the study consisted of 1842 stands located in northern Finland and 41 stands located in eastern Finland. Stand-level ocular inventory and checking inventory were carried out in all study stands by professional surveyors. In simulation experiments the methods considered for error generation were the 1nn-method, the empirical errors method and the Monte Carlo method with log-normal and multivariate log-normal error distributions. The Monte Carlo method with multivariate error distributions was found to be the most flexible simulation method. This method produced the required variation and relations between the errors of the median basal area tree characteristics. However, if the reference data are extensive the 1nn-method, and in certain conditions also the empirical errors method, offer a useful tool for producing error structures which reflect reality.
  • Haara, Finnish Forest Research Institute, Joensuu Research Centre, P.O.Box 68, FIN-80101 Joensuu, Finland E-mail: arto.haara@metla.fi (email)
article id 485, category Research article
Helen Uliczka. (2003). Nature conservation efforts by forest owners – intentions and practice in a Swedish case study. Silva Fennica vol. 37 no. 4 article id 485. https://doi.org/10.14214/sf.485
Keywords: forest policy; boreal forest; indicators; biodiversity assessment; forest conservation; policy implementation
Abstract | View details | Full text in PDF | Author Info
Before a forest operation Swedish forest owners need to fill in a registration form. Since 1994, when a new Swedish Forestry Act came into force, intended nature conservation measures can also be noted on the form. I evaluate 1) if the self-reported nature conservation intentions displayed any trends from 1995 to 2000, and 2) if the intentions were implemented. All forms from these years, in one municipality, were analysed and the stand structure retention was measured on 40 clear-cuts. The intentions, noted as check-marks on the form, showed an increasing trend during these years. However, the increase may be an artefact of changes the form during the time period. The number of check-marks on the forms and the stand structure items actually present on the 40 clear-cuts showed a positive relation. The clear-cuts with ≤ 3 check-marks on the form had lower amounts of the three most common items, than those with ≥ 4 check-marks. To conclude 1) a true increase in the self-reported intentions of the forest owners could not be established; 2) the intentions were generally followed by associated practices on the clear-cuts; 3) the amounts of stand structures retained were probably not enough to reach the biodiversity goal of the Forestry Act. The registration form could be improved to become less open for interpretation and contain quantified recommendations. Self-reported intentions of the forest owners could then possibly be used as indicators of real structural retention, which could facilitate planning and allow for making predictions about the future forests.
  • Uliczka, Swedish University of Agricultural Sciences, Forest Faculty, Department of Conservation Biology, Grimsö Wildlife Research Station, SE-730 91 Riddarhyttan, Sweden E-mail: helen.uliczka@nvb.slu.se (email)
article id 484, category Research article
Pekka E. Kauppi. (2003). New, low estimate for carbon stock in global forest vegetation based on inventory data. Silva Fennica vol. 37 no. 4 article id 484. https://doi.org/10.14214/sf.484
Keywords: carbon stock; global forest; vegetation biomass
Abstract | View details | Full text in PDF | Author Info
Several reports by Intergovernmental Panel on Climate Change (IPCC) have referred to published estimates ranging from 352 to 536 x 109 tons for the global pool of carbon in forest vegetation. However, a rounded estimate of 300 x 109 tons can be derived from the recent Global Forest Resources Assessment 2000 of the FAO, as shown in this paper. By comparing with independent empirical evidence as published in recent scientific literature and by considering sampling principles and the disturbance cycles of large forest regions, it is argued that the new lower estimate is more realistic. A downward correction of the estimate would make an important contribution to balancing the global carbon budget.
  • Kauppi, International Institute for Applied Systems Analysis (IIASA), A-2361 Laxenburg, Austria. Present address: Environmental Science and Policy, Department of Limnology and Environmental Protection, PO Box 27, FIN-00014 University of Helsinki, Finland E-mail: pekka.kauppi@helsinki.fi (email)
article id 483, category Research article
Staffan Jacobson. (2003). Addition of stabilized wood ashes to Swedish coniferous stands on mineral soils - effects on stem growth and needle nutrient concentrations. Silva Fennica vol. 37 no. 4 article id 483. https://doi.org/10.14214/sf.483
Keywords: Pinus sylvestris; Norway spruce; Picea abies; Scots pine; foliar analysis; N fertilization; recycling
Abstract | View details | Full text in PDF | Author Info
Increasing use of forest fuels for energy production is generating increasing quantities of wood ash. A common understanding is that this ash should be spread in forests to counteract soil acidification and potential future nutrient deficiencies, and thus help sustain long-term forest productivity. A series of seven field experiments was established in Sweden in 1988–1995 to study the stem growth and needle nutrient concentrations of 30–60-year-old Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.) stands on mineral soil after additions of wood ash in different doses or a combination of wood ash and N. The results showed that the most pronounced growth responses occurred when N was added, either alone or in combination with wood ash. The stem growth responses to additions of wood ash without N were small and variable, and not statistically significant at any of the studied experimental sites. However, there were indications that the addition of wood ash may increase stem-wood growth on fertile sites and decrease it on less fertile sites. In the short term, the addition of wood ash tended to increase the needle nutrient concentrations of most analyzed elements, except for N, but this could not be correlated to responses in stem growth.
  • Jacobson, The Forestry Research Institute of Sweden, Uppsala Science Park, SE-75183 Uppsala, Sweden E-mail: staffan.jacobson@skogforsk.se (email)
article id 482, category Research article
Yaye Kène Gassama-Dia, Djibril Sané, Mansor N'Doye. (2003). Reproductive biology of Faidherbia albida (Del.) A. Chev. Silva Fennica vol. 37 no. 4 article id 482. https://doi.org/10.14214/sf.482
Keywords: flowering; selfing; Faidherbia albida; fructification; polyad; allogamy
Abstract | View details | Full text in PDF | Author Info
Phenology, flowering and fructification were studied in 5 natural populations of Faidherbia albida in a semi-arid zone in Senegal. In this species, the inflorescence acts as the reproductive unit; the basal flowers, opening first, have a low rate of fertilisation; the maximum rate of fertilisation (65%) was obtained in the apical flowers. Stigmatic receptivity, tested by esterasic reaction, was maximal immediately after anthesis. Stigmata of F. albida can bear simultaneously 2 or 3 polyads. Controlled pollination revealed that allogamy is the dominant reproductive system (ISI = 0.2) in natural populations of F. albida. Intra-specific variability in selfing (ISI ranging from 0 to 0.54) was also observed. Despite of the complete reproductive mechanism during flowering, only a small number of ripe pods (1.25%) is produced, and an average of 70% of the ovules per carpel are fertilized.
  • Gassama-Dia, Université Cheikh Anta Diop de Dakar, Faculté des Sciences et Techniques, Département de biologie végétale, Dakar, Sénégal E-mail: ykdia@ucad.sn (email)
  • Sané, Université Cheikh Anta Diop de Dakar, Faculté des Sciences et Techniques, Département de biologie végétale, Dakar, Sénégal E-mail: ds@nn.sn
  • N'Doye, Université Cheikh Anta Diop de Dakar, Faculté des Sciences et Techniques, Département de biologie végétale, Dakar, Sénégal E-mail: mnd@nn.sn
article id 481, category Research article
K. S. Wang. (2003). Relationship between empty seed and genetic factors in European beech (Fagus sylvatica L.). Silva Fennica vol. 37 no. 4 article id 481. https://doi.org/10.14214/sf.481
Keywords: Fagus sylvatica L.; allozyme; empty seeds; selfing; outcrossing rate; correlation
Abstract | View details | Full text in PDF | Author Info
The relationship between percentage of empty seed (Pes) and genetic factors was explored in an isolated stand of European beech (Fagus sylvatica L.). Nine allozyme loci (GOT-B, IDH-A, LAP-A, MDH-B, MDH-C, MNR-A, 6-PGDH-A, PGI-B and PGM-A) were used to estimate genetic factors. Pes ranged from 4.8% to 40.9% for seed samples of 91 trees within the stand and showed an approximate normal distribution. The average Pes was 21.4% and the repeatability of Pes was 43.4%. The multilocus estimate for outcrossing rate (tm) based on seed samples of 30 trees within the stand was 1.015 (SE = 0.011) and the mean single locus estimate was slightly higher at 1.061 (SE = 0.026). No evidence of biparental inbreeding was found. Weak positive correlation between Pes and maximum selfing rate as well as and significant negative correlation between Pes and multilocus outcrossing rate indicated that self-fertilization may be explained as one of the important causes of empty seeds in beech.
  • Wang, Program in Genetics and Genomic Biology, Research Institute, The Hospital for Sick Children, 555 University Avenue, Toronto, Ontario, M5G 1X8, Canada E-mail: kshengw@yahoo.ca (email)
article id 496, category Research article
Hampus Holmström, Hans Kallur, Göran Ståhl. (2003). Cost-plus-loss analyses of forest inventory strategies based on kNN-assigned reference sample plot data. Silva Fennica vol. 37 no. 3 article id 496. https://doi.org/10.14214/sf.496
Keywords: uncertainty; data acquisition; imputation; forestry planning
Abstract | View details | Full text in PDF | Author Info
The usefulness of kNN (k Nearest Neighbour)-assigned reference sample plot data as a basis for forest management planning was studied. Cost-plus-loss analysis was applied, whereby the inventory cost for a specific method is added to the expected loss due to non-optimal forestry activities caused by erroneous descriptions of the forest state. Four different strategies for data acquisition were evaluated: 1) kNN imputation of sample plots based on traditional stand record information, 2) imputation based on plot-wise aerial photograph interpretation in combination with stand record information, 3) sample plot inventory in the field with 5 plots per stand, and 4) sample plot inventory with 10 plots per stand. Expected losses were derived as mean values of differences between the maximum net present value and the corresponding value obtained when the treatment schedule believed to be optimal (based on data simulated according to method 1–4) was selected. The optimal choice of method was found to depend on factors such as stand maturity, stand area, and time to next treatment (thinning or clearcutting). In general, the field sample plot methods were competitive in large mature stands, especially when the time to the next (optimal) treatment was short. By in each stand (within an estate) employing the method with the lowest cost-plus-loss rather than choosing the method that performed best on average for the entire estate, the total cost for inventory at the estate level could be decreased by 15–50%. However, it was found difficult to identify what method should optimally be employed in a stand based on general stand descriptions.
  • Holmström, Regional Board of Forestry of Västra Götaland, P.O. Box 20008, SE-50420 Borås, Sweden E-mail: hampus.holmstrom@svsvg.svo.se (email)
  • Kallur, ÖKA Skogsplan, Kopparvägen 45 O, SE-90750 Umeå, Sweden E-mail: hk@nn.se
  • Ståhl, Swedish Univ. of Agricultural Sciences, Dept. of Forest Resource Management and Geomatics, SE-90183 Umeå, Sweden E-mail: gs@nn.se
article id 495, category Research article
Christopher Dean. (2003). Calculation of wood volume and stem taper using terrestrial single-image close-range photogrammetry and contemporary software tools. Silva Fennica vol. 37 no. 3 article id 495. https://doi.org/10.14214/sf.495
Keywords: biomass; branches; terrestrial; photography; rectification; volume; taper
Abstract | View details | Full text in PDF | Author Info
A method of estimating trunk and branch volumes of single trees is presented that uses a combination of elementary field measurements, terrestrial photography, image rectification and on-screen digitising using commercial software packages and automated volume calculation. The method is applicable to a variety of different sized trees in situations where the trunks are clearly visible. Results for taper measurement and wood volume calculation are presented for Eucalyptus regnans F. von Muell., Sequoiadendron giganteum (Lindley) Buchholz and Quercus robur L. Branch allometrics are provided for E. regnans. The largest errors arose from field observations. If the trees are asymmetrical in cross-section (e.g. due to irregular buttressing or forked stems), or if there is no vantage point perpendicular to the direction of lean, then photographs from more than one side are recommended. Accuracy and precision of geometric reproduction by the image rectification process, and the volume calculation, were tested using mathematically generated tree components. The errors in the branch volumes of the virtual tree showed complex trends due to interacting factors. Volumes were underestimated by an average 0.5% for stems and 4% for branches. Due to the area deficit resulting from non-circular cross-sections of the buttress, overestimation of stem volumes could be as high as 10% on average for mature trees. However, the area deficit was known for E. regnans and incorporated into the volume calculation. The underestimation of volumes would help counteract over-estimation due to the area deficit. The application of this method to carbon accounting in forests and woodlands is explained.
  • Dean, CRC for Greenhouse Accounting, The Australian National University, GPO Box 475, Canberra, ACT 2601, Australia E-mail: cdean@rsbs.anu.edu.au (email)
article id 494, category Research article
Antero Varhimo, Soili Kojola, Timo Penttilä, Raija Laiho. (2003). Quality and yield of pulpwood in drained peatland forests: pulpwood properties of Scots pine in stands of first commercial thinnings. Silva Fennica vol. 37 no. 3 article id 494. https://doi.org/10.14214/sf.494
Keywords: Pinus sylvestris; peatland; thinning; silviculture; pulpwood; fiber; pulping
Abstract | View details | Full text in PDF | Author Info
The inherent structural dynamics of drained peatland forests may result in a great variation in various wood and fiber properties. We examined variation in fiber and pulp properties i) among stands, ii) among trees within stands, and iii) within trees in young stands dominated by Scots pine (Pinus sylvestris L.). The stands, selected to cover a maximal range of the potential variation, were all at a stage of development where the first commercial thinnings would be feasible. Differences in the processability of the thinning removals were small. In similar kraft cooking conditions, a 5-unit variation in the kappa number of unbleached pulp was observed among the stands. Stand origin had no effect on pulp bleaching. The wood formed prior to drainage had a higher density, shorter fibers, was slightly slower delignified by cooking, and its yield was slightly lower than that of post-drainage wood. These properties, except for high density, are typical for juvenile wood in general, and at stand level they did not correlate with the proportion of pre-drainage wood. When the variation in fiber and pulp properties was broken down into its components, most of it was derived from tree-level in all the cases. On average, the fiber and pulp properties did not differ from those observed for first-thinning pulpwood from upland sites. Consequently, peatland-grown pulpwood may be mixed with other pulpwood in industrial processes. It would probably be best suited as the raw material for pulps with high bonding requirements, e.g. in the top ply of multi-ply board grades or in some specialty grades.
  • Varhimo, KCL, P.O. Box 70, FIN-02151 Espoo, Finland E-mail: antero.varhimo@kcl.fi (email)
  • Kojola, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: sk@nn.fi
  • Penttilä, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: sp@nn.fi
  • Laiho, Department of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: raija.laiho@helsinki.fi
article id 493, category Research article
Göran Rune. (2003). Slits in container wall improve root structure and stem straightness of outplanted Scots pine seedlings. Silva Fennica vol. 37 no. 3 article id 493. https://doi.org/10.14214/sf.493
Keywords: Pinus sylvestris; solidwall container; slitwall container; root morphology; mechanical instability; stem straightness
Abstract | View details | Full text in PDF | Author Info
Root structure and basal sweep were measured on 6-year-old Scots pine (Pinus sylvestris L.) trees at two sites with different soil fertility. Each site was planted with seedlings of identical origin after nursery cultivation in either solidwall container types with vertical ribs or in slitwall container types. Neither container design nor container volume affected tree height or stem diameter on the two sites. The transversal area of lateral roots was larger than the transversal area of bottom roots for the two container types at both sites. The proportion of bottom root transversal area to the total root transversal area was larger in the seedlings growing on the low fertility site than in those growing in the high fertility site for both container types. Seedlings cultivated in slitwall containers had a larger root area in proportion to stem diameter and had less root spiralling compared to the trees cultivated in solidwall containers. At the high fertility site, trees from the slitwall container type had straighter stem bases than seedlings grown in solidwall containers. At the low fertility site, differences in basal sweep formation were small between the container types. Reasons for this are discussed.
  • Rune, Dalarna University, Department of Forestry and Wood Technology, SE-776 98 Garpenberg, Sweden E-mail: gru@du.se (email)
article id 492, category Research article
Seppo Kaunisto, Tytti Sarjala. (2003). Foliar potassium concentrations of bilberry, bog bilberry and downy birch as indicators of potassium nutrition of Scots pine on a drained peatland. Silva Fennica vol. 37 no. 3 article id 492. https://doi.org/10.14214/sf.492
Keywords: Pinus sylvestris; peatland; foliar analysis; dormancy; B. pubescens; deficiency; deficiency limit; needle analysis; potassium; putrescine; V. myrtillus; V. uliginosum
Abstract | View details | Full text in PDF | Author Info
Leaves of bilberry (V. myrtillus), bog bilberry (V. uliginosum) and downy birch (B. pubescens) were collected five times during a growing season from 18 plots in a drainage area, and needles of Scots pine (Pinus sylvestris L.) once during the following winter from the same plots at Parkano, southern Finland. The aim was to study the potassium nutrition of the test plants and relationships between the foliar potassium concentrations of Scots pine and of the test plants. The estimation of the potassium nutrition of test plants was based on the accumulation of putrescine in foliage. Apart from single observations, elevated putrescine concentrations were found when the potassium concentrations were < 5 mg g–1 in bilberry, < 4 mg g–1 in bog bilberry and < 6 mg g–1 in downy birch, and the highest concentrations below the potassium levels of 3.0–3.5 mg g–1. At the concentrations of 2–3 mg g–1 the accumulation increased pronouncedly in bog bilberry but less in downy birch and only slightly in bilberry. The foliar potassium concentrations in test plants correlated closely with the concentrations in pine needles. The concentrations of all species in August were quite stable at the levels of the severe and slight potassium deficiency of pine (3.5 and 4.5 mg g–1 respectively) indicating that August would be suitable for collecting foliage. The concentrations in bog bilberry were very close to the concentrations in pine at the severe and those in bilberry at the slight deficiency level of pine. All test plants could be used for predicting the potassium nutrition of Scots pine, but additional research is needed for the practical application of the method.
  • Kaunisto, Finnish Forest Research Institute, Parkano Research Station, Kaironiementie 54, FIN-39700 Parkano, Finland E-mail: seppo.kaunisto@metla.fi (email)
  • Sarjala, Finnish Forest Research Institute, Parkano Research Station, Kaironiementie 54, FIN-39700 Parkano, Finland E-mail: ts@nn.fi
article id 491, category Research article
Lars Rytter, Lars-Göran Stener. (2003). Clonal variation in nutrient content in woody biomass of hybrid aspen (Populus tremula L. x P. tremuloides Michx.). Silva Fennica vol. 37 no. 3 article id 491. https://doi.org/10.14214/sf.491
Keywords: growth; branches; genetic correlations; heritability; nutrient concentration; nutrient removal; stems
Abstract | View details | Full text in PDF | Author Info
Differences in the nutrient concentrations and nutrient amounts of stems and branches amongst clones of hybrid aspen (Populus tremula L. x P. tremuloides Michx.) were investigated. Seven clones with superior and seven with medium growth rates were selected from a test of 119 clones in southern Sweden. Four trees per clone were randomly identified and harvested in dormant conditions. Sample discs from the stems and branches were collected and analysed for N, K, P, Ca, Mg, and S concentrations, as well as wood density. The analyses revealed significant genetic differences in wood density, K, P, and Mg concentrations in the stems. There were weak (non-significant) and negative genetic correlations between stem volume and concentrations of all the nutrients, except potassium, suggesting that nutrient-efficient clones could be selected without significantly sacrificing genetic gain for growth. In the branches K, Ca, and Mg concentrations differed significantly among clones. After selecting more nutrient efficient clones, the potential savings of nutrients compared with current hybrid aspen material was estimated to be around 5%, which seems fairly low, at least in a short-term perspective. However, the use of clones with different nutrient storage strategies may be regarded as a possible way in the long run to save nutrients in hybrid aspen ecosystems, or of removing them when sludge is applied.
  • Rytter, Forestry Research Institute of Sweden, Skogforsk, Ekebo 2250, SE-26890, Svalöv, Sweden E-mail: lars.rytter@skogforsk.se (email)
  • Stener, Forestry Research Institute of Sweden, Skogforsk, Ekebo 2250, SE-26890, Svalöv, Sweden E-mail: lgs@nn.se
article id 490, category Research article
Veiko Uri, Hardi Tullus, Krista Lõhmus. (2003). Nutrient allocation, accumulation and above-ground biomass in grey alder and hybrid alder plantations. Silva Fennica vol. 37 no. 3 article id 490. https://doi.org/10.14214/sf.490
Keywords: biomass; Alnus incana; Alnus incana x Alnus glutinosa; grey alder; hybrid alder; nutrient allocation; nutrient accumulation
Abstract | View details | Full text in PDF | Author Info
The aim of the present work was to investigate the nutrient (N,P,K) allocation and accumulation in grey alder (Alnus incana (L.) Moench) and hybrid alder (Alnus incana (L.) Moench x Alnus glutinosa (L.) Gaertn.) plantations growing on former agricultural land and to estimate the above-ground biomass production during 4 years after establishment. In August of the 4th year, when leaf mass was at its maximum, the amount of nitrogen accumulated in above-ground biomass of grey alder stand was 142.0 kg ha–1, the amount of phosphorus 16.3 kg ha–1 and the amount of potassium 49.5 kg ha–1. The amount of nitrogen accumulated in a hybrid alder stand totalled 76.8 kg ha–1, that of phosphorus 6.2 kg ha–1 and that of potassium 28.2 kg ha–1. The smaller amounts of N,P and K bound in the hybrid alder plantation are related to the smaller biomass of the stand. Still, the amounts of N,P and K consumed for the production of one ton of biomass were similar in the case of up to 4-year-old grey alder and hybrid alder stands. In the 4th year, the amount of nutrients consumed in one ton of biomass produced were: 16.0 kg N, 1.6 kg P and 5.4 kg K for grey alder and 14.6 kg N, 1.1 kg P and 5.2 kg K for hybrid alder. In the 4th year the total above-ground biomass (dry mass) of grey alder (15750 plants ha–1) amounted to 12.3 t ha–1, current annual increment being 6.7 t ha–1. In hybrid alder stands (6700 plants ha–1), the respective figures were 6.1 t ha–1 and 4.5 t ha–1. Comparison of the production capacity on the basis of mean stem mass in the 4th year revealed that the stem mass of grey alder exceeded that of hybrid alder (0.64 kg and 0.58 kg, respectively). Grey alder outpaced hybrid alder in height growth; in the 4th year after establishment, the mean height of the grey alder stand was 4.6 ± 0.9 m and that of the hybrid alder plantation 3.5 ± 0.9 m.
  • Uri, Institute of Silviculture, Estonian Agricultural University, Kreutzwaldi 5, 51014 Tartu, Estonia E-mail: vuri@eau.ee (email)
  • Tullus, Institute of Silviculture, Estonian Agricultural University, Kreutzwaldi 5, 51014 Tartu, Estonia E-mail: ht@nn.ee
  • Lõhmus, Institute of Geography, University of Tartu, Vanemuise 46, 51014 Tartu, Estonia E-mail: kl@nn.ee
article id 507, category Research article
Tysk Staffan Ericsson, Lars Östlund, Rikard Andersson. (2003). Destroying a path to the past – the loss of culturally scarred trees and change in forest structure along Allmunvägen, in mid-west boreal Sweden. Silva Fennica vol. 37 no. 2 article id 507. https://doi.org/10.14214/sf.507
Keywords: forest history; dendroecology; blazed trees; culturally modified trees; forest trails; tree age structure
Abstract | View details | Full text in PDF | Author Info
The tradition to blaze trees to mark trails and boundaries is very old in northern Scandinavia. The disappearance of culturally modified trees (i.e. trees with trail blazes) and changes in forest structure along a section of an old bridle trail in boreal Sweden was analyzed using historical maps and forest surveys from the period 1876 to the year 2000. Remaining blazed trees were located during a field study and selected scars were dated. In total 104 scarred living and dead trees were found. The scars originated from the early 1500s to the early 1900s. Analysis of the forest surveys showed that the forest along the trail was dominated by older trees, and that the majority of the scarred trees probably were present, throughout the 19th century. By the mid 20th century logging had begun to affect the tree age along the trail and in 1974 no stands older than 180 years were present. A conservative estimate shows that around 90% of the original blazed trees have vanished. The trail was interpreted as have being lined for centuries with scarred trees which gradually have been destroyed during the 20th century. Culturally modified trees constitute an unique source of information for understanding pattern of old trails as well as of past human land use and movement in the landscape prior to the 20th century. This biological archive have to a large extent been destroyed by forestry activities and it is therefore very important to survey, recount and protect the trees that are still present.
  • Ericsson, SLU, Dept. of Forest Vegetation Ecology, S-901 83 Umeå, Sweden E-mail: staffan@delta.se (email)
  • Östlund, SLU, Dept. of Forest Vegetation Ecology, S-901 83 Umeå, Sweden E-mail: lo@nn.se
  • Andersson, SLU, Dept. of Forest Vegetation Ecology, S-901 83 Umeå, Sweden E-mail: ra@nn.se
article id 506, category Research article
Teijo Palander, Katja Turunen, Sanna Laukkanen. (2003). Attitude of Finnish timber buyers towards implementation of a forest computer visualisation. Silva Fennica vol. 37 no. 2 article id 506. https://doi.org/10.14214/sf.506
Keywords: change resistance; competence; forest computer visualisation; timber trade
Abstract | View details | Full text in PDF | Author Info
Timber buyers’ opinions as regards forest computer visualisation are studied. The results indicated that timber buyers are still rather conventional in their computer use since they mainly use only the information system of their own company. The majority of the buyers perceived computer visualisation to be slightly useful for their work, or they had no opinion concerning the usefulness of it. One third of the buyers considered computer visualisation to be a quality factor for timber trade, and the majority were willing to use it if a program is going to adopt by their company. In analysis, different personal characteristics were found for three timber buyer groups: qualified, neutral and reluctant. Qualified buyers were the largest one, about half of the buyers belonging to this group. The greatest barriers to adopting a positive attitude to computer visualisation turned out to be weak computer skills and a general lack of interest in computer use. On the other side, it was found that organisational factors did not influence the buyers’ computer skills or attitudes towards visualisation. The results of this study can be utilised by timber buying organisation in ensuring the successful adoption of a new computer system.
  • Palander, Faculty of Forestry, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: teijo.palander@joensuu.fi (email)
  • Turunen, Faculty of Forestry, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: kt@nn.fi
  • Laukkanen, Faculty of Forestry, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: sl@nn.fi
article id 505, category Research article
Jari Kärnä, Eric Hansen, Heikki Juslin. (2003). Environmental activity and forest certification in marketing of forest products – a case study in Europe. Silva Fennica vol. 37 no. 2 article id 505. https://doi.org/10.14214/sf.505
Keywords: environmental marketing; green marketing; European forest industry
Abstract | View details | Full text in PDF | Author Info
Forest industries and their industrial customers from four European countries were surveyed by interviews to study the environmental emphasis and the role of timber certification in their marketing planning. Most of the Finnish, Swedish, German and British companies have begun to integrate environmental issues in their strategic, structural and functional level marketing decisions. They see forest certification as a necessary tool for marketing forest products. The level of environmental activity (greenness) of the companies was studied by creating a one dimensional factor score rating. The logic of marketing planning was tested by using one functional level marketing tool – forest certification – as an example to examine how well the level of greenness explains the importance of forest certification for the company. The results show that in the surveyed companies the level of greenness has more explanatory power than background factors such as country or industry sector. The integration of environmental issues into marketing planning and the interest in forest certification by these companies can provide meaningful insights for the forest industries worldwide as they confront similar issues.
  • Kärnä, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: jari.karna@metla.fi (email)
  • Hansen, Oregon State University, Department of Wood Science and Engineering, Richardson Hall 108, 97331-5751 Corvallis, OR, USA E-mail: eh@nn.us
  • Juslin, University of Helsinki, Department of Forest Economics, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: hj@nn.fi
article id 504, category Research article
Sylvain Jutras, Hannu Hökkä, Virpi Alenius, Hannu Salminen. (2003). Modeling mortality of individual trees in drained peatland sites in Finland. Silva Fennica vol. 37 no. 2 article id 504. https://doi.org/10.14214/sf.504
Keywords: Pinus sylvestris; Betula pubescens; simulation; peatlands; mortality; generalized linear mixed models; multilevel models
Abstract | View details | Full text in PDF | Author Info
Multilevel logistic regression models were constructed to predict the 5-year mortality of Scots pine (Pinus sylvestris L.) and pubescent birch (Betula pubescens Ehrh.) growing in drained peatland stands in northern and central Finland. Data concerning tree mortality were obtained from two successive measurements of the National Forest Inventory-based permanent sample plot data base covering pure and mixed stands of Scots pine and pubescent birch. In the modeling data, Scots pine showed an average observed mortality of 2.73% compared to 2.98% for pubescent birch. In the model construction, stepwise logistic regression and multilevel models methods were applied, the latter making it possible to address the hierarchical data, thus obtaining unbiased estimates for model parameters. For both species, mortality was explained by tree size, competitive position, stand density, species admixture, and site quality. The expected need for ditch network maintenance or re-paludification did not influence mortality. The multilevel models showed the lowest bias in the modeling data. The models were further validated against independent test data and by embedding them in a stand simulator. In 100-year simulations with different initial stand conditions, the models resulted in a 72% and 66% higher total mortality rate for the stem numbers of pine and birch, respectively, compared to previously used mortality models. The developed models are expected to improve the accuracy of stand forecasts in drained peatland sites.
  • Jutras, Département des sciences du bois et de la forêt, Université Laval, Ste-Foy, Québec, G1K 7P4, Canada E-mail: sj@nn.ca
  • Hökkä, Finnish Forest Research Institute, Rovaniemi Research Station, P.O. Box 16, FIN 96301 Rovaniemi, Finland E-mail: hannu.hokka@metla.fi (email)
  • Alenius, Finnish Forest Research Institute, Rovaniemi Research Station, P.O. Box 16, FIN 96301 Rovaniemi, Finland E-mail: va@nn.fi
  • Salminen, Finnish Forest Research Institute, Rovaniemi Research Station, P.O. Box 16, FIN 96301 Rovaniemi, Finland E-mail: hs@nn.fi
article id 503, category Research article
Jyrki Hytönen. (2003). Effects of wood, peat and coal ash fertilization on Scots pine foliar nutrient concentrations and growth on afforested former agricultural peat soils. Silva Fennica vol. 37 no. 2 article id 503. https://doi.org/10.14214/sf.503
Keywords: wood ash; afforestation; Scots pine; vegetation control; herbicides; coal ash; peat ash; peat soils
Abstract | View details | Full text in PDF | Author Info
The effects of ash and commercial fertilizers on the foliar nutrient concentrations and stand growth of Scots pine were studied in four field experiments established on former cultivated peat soils. The aims were to compare ash types (wood, peat and coal ash), study the effects of ash treatment (pelletization), compare ash fertilization with commercial fertilizers, and to study the interaction between ash fertilization and weed control. Foliar samples were collected 1–3 years and 7–8 years after fertilization. In the unfertilized plots, the foliar nitrogen and phosphorus concentrations were fairly high, while those of potassium were low in all the experiments. The boron levels were low in three out of the four experiments. Application of either loose or pelletized wood ash, as well as of commercial fertilizers, increased foliar potassium and boron concentrations, and thus successfully remedied the existing nutrient imbalances and deficiencies. Since phosphorus deficiencies are rarely encountered on field afforestation sites, poor-quality wood ash with low phosphorus concentration could be used. Peat ash containing phosphorus, but only small amounts of potassium and boron, was not found to be very suitable for soil amelioration in connection with field afforestation. Coal ash, containing only small amounts of potassium, was a good source of boron for pine even when used in small amounts, and thus it can be used in cases where boron deficiencies alone are encountered. Wood ash significantly increased the height growth of Scots pines in two of the experiments, but peat ash and coal ash had no statistically significant effect. Wood ash increased the number of healthy seedlings. Vegetation control decreased seedling mortality by 24%, increased the growth of pine and decreased the proportion of trees damaged by elk and by deciduous trees.
  • Hytönen, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: jyrki.hytonen@metla.fi (email)
article id 502, category Research article
Janne Levula, Hannu Ilvesniemi, Carl Johan Westman. (2003). Relation between soil properties and tree species composition in a Scots pine–Norway spruce stand in southern Finland. Silva Fennica vol. 37 no. 2 article id 502. https://doi.org/10.14214/sf.502
Keywords: Norway spruce; Scots pine; tree species composition; soil texture; soil-water retention; spatial variability
Abstract | View details | Full text in PDF | Author Info
It is commonly known in Finland that Scots pine (Pinus sylvestris L.) is a tree of dry soils and Norway spruce (Picea abies (L) Karst.) is a tree of fresh soils. However, the concepts of dry and fresh soils still lack a precise definition. Consequently, the discussion on which soil/site is a pine or spruce habitat has continued over several decades. Moreover, in forest regeneration, the practice of tree species selection between the pine and the spruce has varied. We investigated the relationship between soil properties and pine–spruce species composition in a mature, naturally regenerated stand in southern Finland. We applied spatial analysis to divide the stand area up into 3–7 classes based on selected soil properties and then investigated the variations in species composition among those classes. The pine–spruce basal area ratio (BA of pines / BA of spruces) increased along with increasing mean particle size and proportion of coarse sand and gravel particle size fraction (0.6–20 mm) of mineral soil, and was lowest in classes, with the highest proportions of fine texture fractions. The results suggest that in southern Finland on sorted soils, pine is more competitive in regeneration and growth than spruce when mean particle size is above 0.44 mm or percentage of coarse sand and gravel is higher than 50%.
  • Levula, University of Helsinki, Department of Forest Ecology, FIN-00014 University of Helsinki, Finland E-mail: janne.levula@helsinki.fi (email)
  • Ilvesniemi, University of Helsinki, Department of Forest Ecology, FIN-00014 University of Helsinki, Finland E-mail: hi@nn.fi
  • Westman, University of Helsinki, Department of Forest Ecology, FIN-00014 University of Helsinki, Finland E-mail: cjw@nn.fi
article id 501, category Research article
Jiaojun Zhu, Yutaka Gonda, Takeshi Matsuzaki, Masashi Yamamoto. (2003). Modeling relative wind speed by optical stratification porosity within the canopy of a coastal protective forest at different stem densities. Silva Fennica vol. 37 no. 2 article id 501. https://doi.org/10.14214/sf.501
Keywords: coastal protective forest; optical porosity; Pinus thunbergii; vertical forest structure; wind speed
Abstract | View details | Full text in PDF | Author Info
Wind speed and optical stratification porosity (OSP) were measured at various heights inside a coastal protective forest thinned to different stem densities to assess whether any characteristics of the wind profile in the coastal protective forest could be predicted from OSP. OSP was defined as vertical distribution of the proportion of sky hemisphere not obscured by tree elements inside a forest stand, and was determined for various heights using hemispherical photographic silhouettes on a computer processing system. The distribution of OSP in the coastal forest follows the Lambert-Beer’s law with an extinction coefficient (v). The relative wind speed within the canopy can be described using an exponential form with an attenuation coefficient (a). Variation in relative wind speed was very closely correlated with the distribution of OSP within the canopy. While below the canopy, i.e., in the trunk space, relative wind speed was little correlated with the distribution of OSP because the distribution of OSP was relatively constant there. Therefore, the linear relationships between relative wind speed and OSP and between the two coefficients v and a were established within the canopy. The results suggest that OSP can be used to predict the wind profile in case of the application within the canopy of the coastal forest.
  • Zhu, Qingyuan Station of Forest Ecology, Institute of Applied Ecology, the Chinese Academy of Sciences, No. 72 Wenhua Road, Shenyang 110016, P.R. China; Faculty of Agriculture, Niigata University, Niigata 950-2181, Japan E-mail: jiaojunzhu@iae.ac.cn (email)
  • Gonda, Faculty of Agriculture, Niigata University, Niigata 950-2181, Japan E-mail: yg@nn.jp
  • Matsuzaki, Faculty of Agriculture, Niigata University, Niigata 950-2181, Japan E-mail: tm@nn.jp
  • Yamamoto, Faculty of Agriculture, Niigata University, Niigata 950-2181, Japan E-mail: my@nn.jp
article id 500, category Research article
Jarkko Hantula, Eeva Vainio. (2003). Specific primers for the differentiation of Heterobasidion annosum (s.str.) and H. parviporum infected stumps in northern Europe. Silva Fennica vol. 37 no. 2 article id 500. https://doi.org/10.14214/sf.500
Keywords: IS group S; IS group P; molecular identification; root and butt rot
Abstract | View details | Full text in PDF | Author Info
Two separate amplification products from random amplified microsatellite fingerprints of Heterobasidion annosum (s.str.) and H. parviporum were converted to specific markers. The markers were tested to be species specific and combined to a single PCR-reaction, which allowed the detection and identification of the two fungi directly from wood samples.
  • Hantula, Finnish Forest Research Institute. Fax +358 9 8570 5531 E-mail: jarkko.hantula@metla.fi (email)
  • Vainio, Finnish Forest Research Institute. Fax +358 9 8570 5531 E-mail: ev@nn.fi
article id 499, category Research article
Mikhail V. Kozlov, Pekka Niemelä. (2003). Drought is more stressful for northern populations of Scots pine than low summer temperatures. Silva Fennica vol. 37 no. 2 article id 499. https://doi.org/10.14214/sf.499
Keywords: Pinus sylvestris; climate change; precipitation; stress; temperature; fluctuating asymmetry; Kola peninsula
Abstract | View details | Full text in PDF | Author Info
Needle fluctuating asymmetry, which is a non-specific stress indicator, was used to evaluate responses of Scots pine (Pinus sylvestris L.) to annual climatic variation in the Kola Peninsula, NW Russia, during 1992–1999. Although the 30 trees surveyed for this study demonstrated individualistic responses to the temperature and precipitation of the growth seasons, at the population level we found no effect of temperature and a significant increase in fluctuating asymmetry with a decline in precipitation during the previous August. This finding suggests that the vitality of Scots pine populations at the northern tree limit is controlled by late summer precipitation rather than by temperatures of the growth season.
  • Kozlov, Section of Ecology, University of Turku, 20014 Turku, Finland E-mail: mikoz@utu.fi (email)
  • Niemelä, Forestry Faculty, University of Joensuu, P.O. Box 111, 80101 Joensuu, Finland E-mail: pn@nn.fi
article id 498, category Research article
Slobodan B. Mickovski, A. Roland Ennos. (2003). Anchorage and asymmetry in the root system of Pinus peuce. Silva Fennica vol. 37 no. 2 article id 498. https://doi.org/10.14214/sf.498
Keywords: Pinus sylvestris; eccentricity; root morphology; anchorage; asymmetry; Pinus peuce; clustering
Abstract | View details | Full text in PDF | Author Info
The relationship between the anchorage mechanics and root architecture of Pinus peuce was investigated by carrying out winching tests and examining excavated root systems of 20 mature trees. The root system was dominated by 6.1±1.3 lateral roots, more than 70% of the lateral root cross sectional area (CSA) being distributed in the uppermost 10 cm of soil. Anchorage strength was related to the size of the tree and CSA. The overturning moment of trees was proportional to the diameter at breast height (DBH) to the power of 1.6. The trees exhibited significant asymmetry in anchorage rigidity, but although there was clustering of lateral roots in a preferred direction the root asymmetry was not significantly correlated with the asymmetry in anchorage rigidity, suggesting that much of the anchorage is provided by tap and sinker roots, rather than the laterals. However, the major laterals showed dorsoventral eccentricity, the more eccentric ones being those that were distributed closer to the soil surface and which pointed perpendicular to the direction of greatest resistance. This suggests that this is a result of thigmomorphogenetic effects. These results are compared with those for the related P. sylvestris and suggest that the assimilation and anchorage characteristics of root systems are controlled independently of each other.
  • Mickovski, School of Biological Sciences, University of Manchester, 3.614 Stopford Building, Manchester M13 9PT, UK E-mail: sbm@nn.uk
  • Ennos, School of Biological Sciences, University of Manchester, 3.614 Stopford Building, Manchester M13 9PT, UK E-mail: roland.ennos@man.ac.uk (email)
article id 516, category Research article
Celeste Lacuna-Richman. (2003). Ethnicity and the utilization of non-wood forest products: findings from three Philippine villages. Silva Fennica vol. 37 no. 1 article id 516. https://doi.org/10.14214/sf.516
Keywords: non-wood forest products (NWFPs); indigenous people; migrants; subsistence; market; Philippines
Abstract | View details | Full text in PDF | Author Info
The utilization and trade of non-wood forest products in three villages in the Philippines were compared in this study. Two villages were situated close to each other on the Island of Palawan. The Tagbanua, an indigenous people, inhabited one village; migrants from the Visayas Region of the Philippines populated the other. The third village is located on the Island of Leyte, in the Visayas Region, populated by native Visayan settlers. There was no significant difference in the number of NWFPs utilized by the indigenous people and the migrants. However, there was a wide disparity in income between the two groups, with migrants earning more, partly due to the marketing of commercial NWFPs. This gap could be decreased by fairer trading practices that are dependent in part on better educational opportunities, land rights, legal assistance and access to markets for the Tagbanua. Specific socioeconomic characteristics, such as the presence of a hunter within the household and size of the family were found to have a positive correlation with the use of NWFPs in some study villages. Income and the food expenditure of the household were inversely related with the use of NWFPs in the native Visayan village.
  • Lacuna-Richman, Faculty of Forestry, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: celeste.lacuna-richman@forest.joensuu.fi (email)
article id 515, category Research article
Juho Rantala, Kari Väätäinen, Nuutti Kiljunen, Pertti Harstela. (2003). Economic evaluation of container seedling packing and disinfection machinery. Silva Fennica vol. 37 no. 1 article id 515. https://doi.org/10.14214/sf.515
Keywords: economies of scale; nursery technology; mechanisation; container seedlings; packing;
Abstract | View details | Full text in PDF | Author Info
Productivity and costs of packing container seedlings were studied in a mechanised line for packing and disinfecting seedling trays. The hypothesis was that adequate cost-efficiency could be achieved when some common principles of mechanisation were applied. Results indicated that the unit costs are lower than those of manual packing, if these principles were applied and the annual number of packed seedlings exceeded 6 million. However, most of the nurseries in Finland are still too small to gain a real advantage from large-scale production.
  • Rantala, Finnish Forest Research Institute, Suonenjoki Research Station, FIN-77600 Suonenjoki, Finland E-mail: juho.rantala@metla.fi (email)
  • Väätäinen, Finnish Forest Research Institute, Joensuu Research Centre, FIN-80101 Joensuu, Finland E-mail: kv@nn.fi
  • Kiljunen, Finnish Forest Research Institute, Suonenjoki Research Station, FIN-77600 Suonenjoki, Finland E-mail: nk@nn.fi
  • Harstela, Finnish Forest Research Institute, Suonenjoki Research Station, FIN-77600 Suonenjoki, Finland E-mail: ph@nn.fi
article id 514, category Research article
Jukka Malinen. (2003). Locally adaptable non-parametric methods for estimating stand characteristics for wood procurement planning. Silva Fennica vol. 37 no. 1 article id 514. https://doi.org/10.14214/sf.514
Keywords: stand characteristics; local non-parametric estimation; MSN method; wood procurement planning
Abstract | View details | Full text in PDF | Author Info
The purpose of this study was to examine the use of the local adaptation of the non-parametric Most Similar Neighbour (MSN) method in estimating stand characteristics for wood procurement planning purposes. Local adaptation was performed in two different ways: 1) by selecting local data from a database with the MSN method and using that data as a database in the basic k-nearest neighbour (k-nn) MSN method, 2) by selecting a combination of neighbours from the neighbourhood where the average of the predictor variables was closest to the target stand predictor variables (Locally Adaptable Neighbourhood (LAN) MSN method). The study data used comprised 209 spruce dominated stands located in central Finland and was collected with harvesters. The accuracy of the methods was analysed by estimating the tree stock characteristics and the log length/diameter distribution produced by a bucking simulation. The local k-nn MSN method was not notably better than the k-nn MSN method, although it produced less biased estimates on the edges of the input space. The LAN MSN method was found to be a more accurate method than the k-nn methods.
  • Malinen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: jukka.malinen@joensuu.fi (email)
article id 513, category Research article
Marjut Ihalainen, Kauko Salo, Timo Pukkala. (2003). Empirical prediction models for Vaccinium myrtillus and V. vitis-idaea berry yields in North Karelia, Finland. Silva Fennica vol. 37 no. 1 article id 513. https://doi.org/10.14214/sf.513
Keywords: mixed models; berry yield index; Vaccinium myrtillus; Vaccinium vitis-idaea
Abstract | View details | Full text in PDF | Author Info
Forest berries and the outdoor experiences related to berry collection are important goods and services provided by Finnish forests. Consequently, there is a need for models which facilitate the prediction of the impacts of alternative forest management options on berry yields. Very few such models are available. In particular, empirical models are lacking. Models used in forest management should express the effect of variables altered in forest management such as stand density and mean tree size. This study developed empirical models for bilberry and cowberry yields in North Karelia. The data consisted of 362 measurements of 40 m2 sample plots. The plots were located in clusters. The same plot was measured over 1 to 4 years. Besides berry yield some site and growing stock characteristics of each plot were measured. A random parameter model was used to express the berry yield as a function of site fertility, growing stock characteristics, and random parameters. The random part of the models accounted for the effect of plot, measurement year, and cluster. The fixed predictors of the model for bilberry were stand age and forest site type. Stand basal area, mean tree diameter and forest site type were used to predict cowberry yields. The most significant random parameter was the plot factor. The fixed model part explained only a few per cent of the variation in berry yields. The signs of regression coefficients were logical and the model predictions correlated rather well with the predictions of earlier models.
  • Ihalainen, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: marjut.ihalainen@joensuu.fi (email)
  • Salo, The Finnish Forest Research Institute, Joensuu Research Centre, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: ks@nn.fi
  • Pukkala, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: tp@nn.fi
article id 512, category Research article
Juha-Pekka Hotanen. (2003). Multidimensional site description of peatlands drained for forestry. Silva Fennica vol. 37 no. 1 article id 512. https://doi.org/10.14214/sf.512
Keywords: site index; vegetation; diversity; classification; macronutrients; ordination; peat properties
Abstract | View details | Full text in PDF | Author Info
  • Hotanen, Finnish Forest Research Institute, Joensuu Research Centre, P.O.Box 68, FIN-80101 Joensuu, Finland E-mail: juha-pekka.hotanen@metla.fi (email)
article id 511, category Research article
Luis Diego Pérez Cordero, Markku Kanninen. (2003). Heartwood, sapwood and bark content, and wood dry density of young and mature teak (Tectona grandis) trees grown in Costa Rica. Silva Fennica vol. 37 no. 1 article id 511. https://doi.org/10.14214/sf.511
Keywords: stand density; cross-sectional area; relative stem curves; stem analysis
Abstract | View details | Full text in PDF | Author Info
The aim of this study was to evaluate the heartwood, sapwood and bark content, and wood dry density in young and mature teak (Tectona grandis) trees. For this, 17 plantations were selected from 11 sites representing different climatic conditions and plantation densities (156 to 1600 trees ha–1, and line planting). From these plantations, a total of 87 trees with ages between 5 and 47 years were felled for stem analysis. The highest heartwood proportion of stem volume (over bark) was 61% and the lowest 0.4%. The sapwood proportion ranged between 24 and 72%, while bark represented from 14 to 37% of the total volume. Heartwood proportion was significantly different (P < 0.05) among climatic zones: ‘wet’ sites producing less heartwood than ‘dry’ sites. Stem diameter (under bark) and heartwood diameter at different stem heights differed among sample trees, even when plotted in relative values to avoid dependency with stem size. Dry density was statistically different between 8-year-old trees or younger and 47-year-old trees, and between line planting trees and 13-year-old trees or younger, but did not differ statistically between line planting trees and mature trees. No significant differences were found between climatic zones or between different stand densities. Dry density values for T. grandis plantations in Costa Rica are similar to those reported elsewhere.
  • Pérez Cordero, Centro Agronómico Tropical de Investigación y Enseñanza (CATIE), Turrialba, Costa Rica E-mail: diegoperez@costarricense.cr (email)
  • Kanninen, Center for International Forestry Research (CIFOR), Bogor, Indonesia E-mail: mk@nn.id
article id 510, category Research article
Helge Dzierzon, Risto Sievänen, Winfried Kurth, Jari Perttunen, Branislav Sloboda. (2003). Enhanced possibilities for analyzing tree structure as provided by an interface between different modelling systems. Silva Fennica vol. 37 no. 1 article id 510. https://doi.org/10.14214/sf.510
Keywords: tree growth; mathematical models; simulation systems; forest growth; analysis tools
Abstract | View details | Full text in PDF | Author Info
In recent years, many different advanced mathematical models and simulation systems for tree and forest growth have been developed. We show a possibility to extend analysis tools for measured and simulated plants using a data interface between the simulation model LIGNUM and the multifunctional software system GROGRA. Both systems were developed by different teams. To demonstrate the enhanced possibilities for analyzing a LIGNUM tree, several examples are given. In these examples three different approaches for analysis are applied to measured and simulated trees: Fractal dimension, deduction of tapering laws, and water potential patterns obtained from simulation of waterflow by the specialized software HYDRA. Conclusions for the interfacing and comparison of different modelling tools are drawn.
  • Dzierzon, Institut für Forstliche Biometrie und Informatik, Universität Göttingen, Büsgenweg 4, D-37077 Göttingen, Germany E-mail: hdzierz@gwdg.de (email)
  • Sievänen, The Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: rs@nn.fi
  • Kurth, Institut für Informatik, Brandenburgische Technische Universität Cottbus, P.O. Box 101344, D-03013 Cottbus, Germany E-mail: wk@nn.de
  • Perttunen, The Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: jp@nn.fi
  • Sloboda, Institut für Forstliche Biometrie und Informatik, Universität Göttingen, Büsgenweg 4, D-37077 Göttingen, Germany E-mail: bs@nn.de
article id 509, category Research article
Michelle de Chantal, Laura Eskola, Hannu Ilvesniemi, Kari Leinonen, Carl Johan Westman. (2003). Early establishment of Pinus sylvestris and Picea abies sown on soil freshly prepared and after stabilisation. Silva Fennica vol. 37 no. 1 article id 509. https://doi.org/10.14214/sf.509
Keywords: Norway spruce; site preparation; Scots pine; direct seeding; soil properties
Abstract | View details | Full text in PDF | Author Info
The aim of this study is to investigate the early establishment of Pinus sylvestris L. (Scots pine) and Picea abies (L.) Karst. (Norway spruce) seedlings on soil freshly prepared and soil left to stabilise for one year after preparation. Three site preparation treatments were studied: exposed C horizon, mound (broken O/E/B horizon piled upside down over undisturbed forest floor), and exposed E/B horizon. The years investigated were different in terms of weather, one being rainy and the other one dry. As such, emergence was very low in the dry year. Content of fine silt particles, bulk density, water retention, air-filled porosity, loss-on-ignition, and near saturated hydraulic conductivity did not differ statistically between fresh and stabilised soil. Nevertheless, early establishment of P. sylvestris seedlings was improved on exposed C and E/B horizon after one year of soil stabilisation. In contrast, early establishment of P. sylvestris on mounds, and that of P. abies on all types of site preparation treatments were not improved by soil stabilisation. In addition, mortality due to frost heaving did not differ significantly between freshly prepared and stabilised soil. Considering the fact that growing season climate had a great influence on the sowing outcome, and that early establishment is also affected by other factors that vary yearly, such as predation, seedbed receptivity, and competition from vegetation, it may not be advantageous to wait for soil to stabilise before regenerating from seeds.
  • Chantal, University of Helsinki, Dept. of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: michelle.dechantal@helsinki.fi (email)
  • Eskola, University of Helsinki, Dept. of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: le@nn.fi
  • Ilvesniemi, University of Helsinki, Dept. of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: hi@nn.fi
  • Leinonen, University of Helsinki, Dept. of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: kl@nn.fi
  • Westman, University of Helsinki, Dept. of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: cjw@nn.fi
article id 508, category Research article
Kai Vellak, Jaanus Paal, Jaan Liira. (2003). Diversity and distribution pattern of bryophytes and vascular plants in a boreal spruce forest. Silva Fennica vol. 37 no. 1 article id 508. https://doi.org/10.14214/sf.508
Keywords: boreal forests; correlation; general linear models; Estonia; field layer; ground layer; spatial patterns
Abstract | View details | Full text in PDF | Author Info
Small scale pattern of bryophyte communities is one of the remarkable sources of diversity in species-poor boreal forests. By means of correlation and general linear model approaches, the relationships between bryophyte vegetation and upper layers, as well as the response of ground and field layer species to several environmental factors, was analyzed in a boreal spruce forest in South-East Estonia. Of the studied factors, the strongest influence on the diversity and spatial distribution of ground and field layer species was found for ‘distance from nearest tree’. Species from different layers react differently to the proximity of trees. Species richness of bryophytes is higher further from trees, whereas more vascular plant species prefer to grow in the vicinity of tree trunks. For bryophyte species richness, the pH of the decay horizon is also important; fewer bryophyte species occur in more acid conditions.
  • Vellak, Institute of Zoology and Botany, Estonian Agricultural University, 181 Riia str., 51014 Tartu, Estonia; Institute of Botany and Ecology, University of Tartu, 40 Lai Str., 51005 Tartu, Estonia E-mail: kvellak@zbi.ee (email)
  • Paal, Institute of Botany and Ecology, University of Tartu, 40 Lai Str., 51005 Tartu, Estonia E-mail: jp@nn.ee
  • Liira, Institute of Botany and Ecology, University of Tartu, 40 Lai Str., 51005 Tartu, Estonia E-mail: jl@nn.ee
article id 526, category Research article
Matleena Kniivilä, Olli Saastamoinen. (2002). The opportunity costs of forest conservation in a local economy. Silva Fennica vol. 36 no. 4 article id 526. https://doi.org/10.14214/sf.526
Keywords: nature conservation; local economy; economic impacts; cost of conservation
Abstract | View details | Full text in PDF | Author Info
Costs generated by nature conservation are repeatedly under discussion. Most often the costs of conservation are estimated as aggregate figures at the national or regional level or alternatively, for a forest owner. In this study they were examined at the local level, in the forestry dependent municipality of Ilomantsi in Eastern Finland. The estimations of lost net revenues (stumpage income less silvicultural costs), wages, entrepreneurial income and profits, employment and value added were based on alternative forest management plans calculated for conservation areas. The annual losses as regards employment during the first decade were estimated to be 5.7–20.4 jobs. Later, the employment effects were estimated to be 2.4–6.3 lost jobs. Although the value added lost during the first decade was estimated to be at maximum only 3.4% of the present total value added of the municipality, the share of the value added of forestry was estimated to be higher than the mere protected forest land share would indicate. The use of conservation areas for forestry would create a moderate increase of employment in forestry, i.e. 3.8–14%, during the first decade, but it would later stabilise at a much lower level. Employment impacts at the municipal level were estimated as very small (at maximum 0.9%), but on the other hand, for some villages even single jobs may matter. The main reasons for the minor impacts were the high mechanisation rate of logging and the major flow of stumpage income outside the locality.
  • Kniivilä, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: matleena.kniivila@joensuu.fi (email)
  • Saastamoinen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: os@nn.fi
article id 525, category Research article
Thomas Knoke. (2002). Value of complete information on red heartwood formation in beech (Fagus sylvatica). Silva Fennica vol. 36 no. 4 article id 525. https://doi.org/10.14214/sf.525
Keywords: value of information; economics of beech; red heartwood; silvicultural treatment
Abstract | View details | Full text in PDF | Author Info
Beech (Fagus sylvatica L.) is the most important deciduous tree species in Germany. The wood of beech shows normally a bright colour (white beech) as long as no coloured heartwood has been formed. The facultative heartwood formation is induced when oxygen enters central parts of older trees, where dead or at least less vital parenchyma exist. The coloured heartwood is usually called ‘red heartwood’. Beech without red heartwood can preferably be found in younger trees which show a high water content even in central parts of the stem. The presence of red heartwood is regarded as a severe reduction of timber quality. Numerous studies have investigated opportunities to derive information on the presence and characteristics of red heartwood of standing beech trees. But until now it has not been tested whether such information could be helpful to improve the economics of beech-silviculture. This paper investigates whether complete information on the heartwood of standing beech could be useful to control the proportion of discoloured timber harvested during one rotation. It is also examined, which kind of information on the heartwood could be used to improve the economic results. To verify this, simulations based on simple algorithms were conducted. The general assumption was made that all information on the heartwood would be available. The results show that information which is restricted on the mere existence of red heartwood is neither suited to significantly reduce the amount of coloured timber nor is it possible to improve economic results based on this information. Only based on information on the recent formation of red heartwood of beech, which is actually still white the amount of discoloured timber can be reduced significantly. Consequently the discounted cash flows can only be substantially improved based on information on an expected formation of recent red heartwood.
  • Knoke, Institute of Silviculture and Forest Management, Technische Universität München, Am Hochanger 13, D-85354 Freising, Germany E-mail: knoke@wbfe.forst.tu-muenchen.de (email)
article id 524, category Research article
Timo Pukkala, Jari Miina, Marc Palahí. (2002). Thinning response and thinning bias in a young Scots pine stand. Silva Fennica vol. 36 no. 4 article id 524. https://doi.org/10.14214/sf.524
Keywords: Pinus sylvestris; growth model; non-spatial model; spatial model
Abstract | View details | Full text in PDF | Author Info
The study analyses the annual post-thinning response and thinning bias of a young Scots pine stand as a function of tree size, competition faced by the tree, and competition that is removed around the tree in the thinning treatment. The thinning response of a tree was defined as the change of tree growth due to a thinning treatment. The thinning bias was defined as the difference between the true growth and model prediction. A distance-dependent (spatial) and a distance-independent (non-spatial) growth model were used in the calculations. The empirical data were measured from a thinning experiment consisting of ten plots, each 40 x 30 m in size, which were thinned to different stand densities. The ten-year post-thinning growth of every remaining tree was measured. The results indicated that the highest thinning response is among medium-sized and co-dominant trees. The thinning response is quite small, and even negative for some trees, for two years after thinning but it becomes clearly positive from the third year onwards. The spatial model underestimated the growth of small trees (which usually face high competition) while the non-spatial model overestimated the growth of trees that are small or face much competition. The spatial model used in this study overemphasized the effect of competition while the non-spatial model underestimated this effect. Both growth models overestimated the growth of trees in heavily thinned places, but this bias disappeared in two years. The negative bias was more pronounced with a spatial growth model because the tendency of the non-spatial model to underestimate the growth of trees facing little competition partly compensated for the negative bias.
  • Pukkala, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: timo.pukkala@joensuu.fi (email)
  • Miina, Finnish Forest Research Institute, Joensuu Research Centre, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: jm@nn.fi
  • Palahí, European Forest Institute, Torikatu 34, FIN-80100 Joensuu, Finland E-mail: mp@nn.fi
article id 523, category Research article
Heli Peltola, Jari Miina, Ismo Rouvinen, Seppo Kellomäki. (2002). Effect of early thinning on the diameter growth distribution along the stem of Scots pine. Silva Fennica vol. 36 no. 4 article id 523. https://doi.org/10.14214/sf.523
Keywords: Scots pine; early thinnings; thinning responses; diameter growth; growth distribution
Abstract | View details | Full text in PDF | Author Info
The absolute and relative effects of the first thinning on the diameter growth distribution along the stems were studied in 98 Scots pines (Pinus sylvestris L.) at heights of 1.3, 4, 6 and 8 m. The data cover one 3-year pre-thinning period and four 3-year post-thinning periods in plots with densities varying from 575 to 3400 stems ha–1. A shift in the point of maximum diameter growth down the bole was found during the first 3 years after thinning, with a shift back up the stem later. The thinning response over the whole 12-year post-thinning period was strongest the nearer the stem base and the heavier the thinning. The largest trees had the highest diameter growth after thinning in absolute terms, and the growth was greater the heavier the thinning. The absolute thinning response over the 12-year post-thinning period was highest in the medium tree size and in the largest trees, especially on the heavily thinned and lightly thinned plots. Whereas in the moderately thinned stand the smaller and larger trees responded more than did those of medium size on average. In relative sense, however, the small trees on heavily or moderately thinned plots responded more rapidly and more strongly than the medium-sized or large trees over the whole stem. The small trees on the lightly thinned plots responded only slightly to thinning. The results suggest that it is possible to affect the uniformity of wood properties (such as ring width) both within and between trees by thinning.
  • Peltola, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: heli.peltola@forest.joensuu.fi (email)
  • Miina, Finnish Forest Research Institute, Joensuu Research Centre, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: jm@nn.fi
  • Rouvinen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: ir@nn.fi
  • Kellomäki, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: sk@nn.fi
article id 522, category Research article
Annika Kangas, Matti Maltamo. (2002). Anticipating the variance of predicted stand volume and timber assortments with respect to stand characteristics and field measurements. Silva Fennica vol. 36 no. 4 article id 522. https://doi.org/10.14214/sf.522
Keywords: diameter distribution; timber assortment; prediction; calibration estimation; volume; accuracy; measurement combination
Abstract | View details | Full text in PDF | Author Info
Several models and/or several variable combinations could be used to predict the diameter distribution of a stand. Typically, a fixed model and a fixed variable combination is used in all conditions. The calibration procedure, however, makes it possible to choose the measurement combination from among many possibilities, although the model used is fixed. In this study, the usefulness of utilizing additional stand characteristics for calibrating the predicted diameter distribution is examined. Nine measurement strategies were tested in predicting the total stand volume, sawlog volume and pulpwood volume. The observed errors of these variables under each strategy were modeled as a function of basal area, basal area median diameter and number of stems. The models were estimated in three steps. First, an Ordinary Least Squares (OLS) model was fitted to the observed errors. Then, a variance function was estimated using the OLS residuals. Finally, a weighted Seemingly Unrelated Regression (SUR) analysis was used to model the observed errors, using the estimated variance functions as weights. The estimated models can be used to anticipate the precision and accuracy of predicted volume characteristics for each stand with different variable combinations and, consequently, to choose the best measurement combination in different stands.
  • Kangas, University of Helsinki, Dept. of Forest Resources Management, P.O. Box 27, 00014 University of Helsinki, Finland E-mail: annika.kangas@helsinki.fi (email)
  • Maltamo, University of Joensuu, Faculty of Forestry, P.O. Box 111, 80101 Joensuu, Finland E-mail: mm@nn.fi
article id 521, category Research article
Gordon Nigh. (2002). Site index conversion equations for mixed trembling aspen and white spruce stands in northern British Columbia. Silva Fennica vol. 36 no. 4 article id 521. https://doi.org/10.14214/sf.521
Keywords: site index; mixed species; trembling aspen; white spruce
Abstract | View details | Full text in PDF | Author Info
White spruce and trembling aspen are two important commercial species in British Columbia. They often grow in association, particularly in the Boreal White and Black Spruce and Sub-Boreal Spruce biogeoclimatic zones. Site index conversion equations are useful for estimating the site index of one species from the site index of another species. This study fills a need for site index conversion equations for mixed spruce/aspen stands. Seventy 0.01 ha study plots were established in mixed spruce/aspen stands. One site tree of each species was selected from each plot. The height and breast height ages of the site trees were measured and the site index was estimated with these data. The correlation between the site index of spruce and aspen was 0.6. Geometric mean regression was used to estimate the parameters of a linear site index conversion equation. The analysis did not reveal any differences in the conversion equations across the three major biogeoclimatic units (BWBSmw1, BWBSmw2, and SBS) that were sampled. Therefore, only one conversion equation is required.
  • Nigh, Ministry of Forests, Research Branch, P.O. Box 9519, Stn. Prov. Govt., Victoria, BC, Canada V8W 9C2 E-mail: gordon.nigh@gems2.gov.bc.ca (email)
article id 520, category Research article
Rüdiger Grote. (2002). Foliage and branch biomass estimation of coniferous and deciduous tree species. Silva Fennica vol. 36 no. 4 article id 520. https://doi.org/10.14214/sf.520
Keywords: Picea abies; Fagus sylvatica; biomass estimation; crown dimensions; allometric equations
Abstract | View details | Full text in PDF | Author Info
Under changing environmental conditions, biomass development on the tree and the stand level may differ from today, regardless if the induced change is due to a shift in the general climate properties or to forest management. Under these conditions, tree biomass can not be derived from tables based on former investigations but has to be defined from particular biomass investigations, which generally calculate tree and stand biomass from sample branches using allometric relationships. Therefore, sample measurements on harvested trees are needed. In this paper, foliage and branch biomass estimation for 6 Norway spruces (Picea abies) and 6 beeches (Fagus sylvatica) harvested in a 56-year-old mixed stand in southern Germany is presented. Different allometric models are investigated to derive branch biomass from branch dimension for both species. The equations that are based on branch length, foliated branch fraction, and branch diameter are used for tree and stand level estimates. However, the variation within the 6 trees of each species was too large for a reliable calculation of stand biomass, especially in case of beech branch wood. Furthermore, the necessity of allometric relations and their applicability in individual-tree models is discussed, and the importance of suitable branch- and tree selection is underlined.
  • Grote, TU München, Chair of Forest Yield Science, Am Hochanger 13, D-85354 Freising, Germany E-mail: ruediger.grote@lrz.tu-muenchen.de (email)
article id 519, category Research article
Magnus Lindén, Gudmund Vollbrecht. (2002). Sensitivity of Picea abies to butt rot in pure stands and in mixed stands with Pinus sylvestris in southern Sweden. Silva Fennica vol. 36 no. 4 article id 519. https://doi.org/10.14214/sf.519
Keywords: Pinus sylvestris; Norway spruce; Picea abies; logistic regression; Heterobasidion annosum; Butt rot; mixed forest
Abstract | View details | Full text in PDF | Author Info
Repeatedly sampled data from permanent experimental plots in southern Sweden were used to model butt rot development in Norway spruce growing in pure stands and in mixed stands with Scots pine. The data come from 29 sites with pure spruce, altogether 100 plots, and from 15 sites of mixed spruce and pine, altogether 22 plots. A logistic model provided the best fit to the data. The study material revealed that in mixed stands the proportion of spruce trees with butt rot is lower than in pure Norway spruce stands. The difference in the incidence of butt rot cannot be explained by silviculture or windthrow since both factors are accounted for in the study. The most significant effect on butt rot development in Norway spruce by an admixture of Scots pine, was found when the Scots pine admixture was 50%. In order to reduce the incidence of butt rot in Norway spruce, the study material indicate that there is little to be gained by increasing the Scots pine admixture to much more than 50%.
  • Lindén, Southern Swedish Forest Research Centre c/o Asa Experimental Forest, SLU, S-360 30 Lammhult, Sweden E-mail: magnus.linden@ess.slu.se (email)
  • Vollbrecht, Southern Swedish Forest Research Centre c/o Asa Experimental Forest, SLU, S-360 30 Lammhult, Sweden E-mail: gv@nn.se
article id 518, category Research article
Mats Hannerz, Curt Almqvist, Roland Hörnfeldt. (2002). Timing of seed dispersal in Pinus sylvestris stands in central Sweden. Silva Fennica vol. 36 no. 4 article id 518. https://doi.org/10.14214/sf.518
Keywords: natural regeneration; Scots pine; seed fall
Abstract | View details | Full text in PDF | Author Info
The objective of the study presented here was to describe the variation in timing of seed dispersal from Scots pine (Pinus sylvestris L.) seed trees in central Sweden. Seeds were collected in traps at two sites, for three years at one, and four years at the other . The traps were emptied from March to August each year at 1–2 week intervals during the main period of seed dispersal. The annual seed fall varied between 200 000 and 1.6 million seeds per hectare. The seed fall started in mid to late April, shortly after the heat sum had started to accumulate. The most intensive seed fall took place in early to mid May. The peak period, when 50% of the total seed dispersal occurred, lasted for 18–28 days at the different sites and years. The variation in timing among years seemed to be mainly due to climatic factors – high temperatures promoted seed dispersal, for instance. The results may be useful for planning the time of scarification to optimise the natural regeneration of Scots pine. The data suggest that scarification in the spring, no later than mid-May, would generally create a good seed-bed for most of the current year’s seeds, whereas scarification in late May or June would bury a large proportion of this cohort.
  • Hannerz, The Forestry Research Institute of Sweden (SkogForsk) Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: mats.hannerz@skogforsk.se (email)
  • Almqvist, The Forestry Research Institute of Sweden (SkogForsk) Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: ca@nn.se
  • Hörnfeldt, Swedish University of Agricultural Sciences, Dept. of Forest Products and Markets, P.O. Box 7060, SE-750 07 Uppsala, Sweden E-mail: rh@nn.se
article id 533, category Research article
Anna Liisa Ruotsalainen, Juha Tuomi, Henry Väre. (2002). A model for optimal mycorrhizal colonization along altitudinal gradients. Silva Fennica vol. 36 no. 3 article id 533. https://doi.org/10.14214/sf.533
Keywords: alpine gradient; mycorrhiza; nutrient uptake kinetics; photosynthetic nutrient use efficiency
Abstract | View details | Full text in PDF | Author Info
Mycorrhizal associations are generally favourable for vascular plants in nutrient-poor conditions. Still, non-mycorrhizal plants are common in high arctic and alpine areas, which are often poor in nitrogen and phosphorus. The relative proportion of mycorrhizal plants has been found to decrease along with increasing altitude, suggesting that the advantage of the mycorrhizal symbiosis may change along an altitudinal gradient. This may be related to the environmental factors that possibly constrain the amount of photosynthesized carbon to be shared with mycorrhizal fungi. We propose a simple optimization model for root colonization by fungal symbionts and analyze the advantages of mycorrhizas in relation to the nutrient use efficiency of photosynthesis (PNUE), the kinetics of nutrient uptake and the soil nutrient levels. Our model suggests that mycorrhizas are not usually favoured at low PNUE values. At low nutrient levels, mycorrhizas may be advantageous if they have a lower threshold concentration of nutrient uptake (xmin) compared to non-mycorrhizal roots. If mycorrhizal roots have a higher maximum capacity of nutrient uptake (Vmax), mycorrhizas can be favourable for the host plant even at relatively low nutrient concentrations and at relatively low PNUE. Consequently, the possible patterns along altitudinal gradients essentially depend on PNUE. If the soil nutrient concentration is constant and PNUE decreases, the advantage of mycorrhizal symbiosis declines independently of the nutrient uptake kinetics. If PNUE remains constant and the soil nutrient concentration decreases along with increasing altitude, the emerging colonization pattern (either increasing, decreasing or intermediate) depends on the nutrient uptake kinetics. Additionally, if both PNUE and the soil nutrient concentration decrease, several patterns may emerge, depending on the nutrient uptake kinetics.
  • Ruotsalainen, Department of Biology, Botanical Museum, Box 3000, FIN-90014 University of Oulu, Finland E-mail: annu.ruotsalainen@oulu.fi (email)
  • Tuomi, Department of Biology, Box 3000, FIN-90014 University of Oulu, Finland E-mail: jt@nn.fi
  • Väre, Botanical Museum, Finnish Museum of Natural History, Box 7, FIN-00014 University of Helsinki, Finland E-mail: hv@nn.fi
article id 532, category Research article
Pedro J. Aphalo, Anna W. Schoettle, Tarja Lehto. (2002). Leaf life span and the mobility of “non-mobile” mineral nutrients – the case of boron in conifers. Silva Fennica vol. 36 no. 3 article id 532. https://doi.org/10.14214/sf.532
Keywords: needle age; conifers; model; boron; leaf life span; nutrient mobility; toxicity
Abstract | View details | Full text in PDF | Author Info
Nutrient conservation is considered important for the adaptation of plants to infertile environments. The importance of leaf life spans in controlling mean residence time of nutrients in plants has usually been analyzed in relation to nutrients that can be retranslocated within the plant. Longer leaf life spans increase the mean residence time of all mineral nutrients, but for non-mobile nutrients long leaf life spans concurrently cause concentrations in tissues to increase with leaf age, and consequently may reduce non-mobile nutrient use efficiency. Here we analyze how the role of leaf life span is related to the mobility of nutrients within the plant. We use optimality concepts to derive testable hypotheses, and preliminarily test them for boron (B), a nutrient for which mobility varies among plant species. We review published and unpublished data and use a simple model to assess the quantitative importance of B retranslocation for the B budget of mature conifer forests and as a mechanism for avoiding toxicity.
  • Aphalo, Faculty of Forestry, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland; Current address Department of Biological and Environmental Science, University of Jyväskylä, P.O. Box 35, FIN-40351 Jyväskylä, Finland. E-mail: pedro.aphalo@jyu.fi (email)
  • Schoettle, Rocky Mountain Research Station, 240 West Prospect Road, Fort Collins, CO 80526, USA E-mail: aws@nn.us
  • Lehto, Faculty of Forestry, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: tl@nn.fi
article id 531, category Research article
Thomas N. Buckley, Jeffrey M. Miller, Graham D. Farquhar. (2002). The mathematics of linked optimisation for water and nitrogen use in a canopy. Silva Fennica vol. 36 no. 3 article id 531. https://doi.org/10.14214/sf.531
Keywords: stomatal conductance; optimality theory; nitrogen allocation; NUE; WUE
Abstract | View details | Full text in PDF | Author Info
We develop, and discuss the implementation of, a mathematical framework for inferring optimal patterns of water and nitrogen use. Our analysis is limited to a time scale of one day and a spatial scale consisting of the green canopy of one plant, and we assume that this canopy has fixed quantities of nitrogen and water available for use in photosynthesis. The efficiencies of water and nitrogen use, and the interactions between the two, are strongly affected by physiological and physical properties that can be modeled in different ways. The thrust of this study is therefore to discuss these properties and how they affect the efficiencies of nitrogen and water use, and to demonstrate, qualitatively, the effects of different model assumptions on inferred optimal strategies. Preliminary simulations suggest that the linked optimisation of nitrogen and water use is particularly sensitive to the level of detail in canopy light penetration models (e.g., whether sunlit and shaded fractions are pooled or considered independently), and to assumptions regarding nitrogen and irradiance gradients within leaves (which determine how whole-leaf potential electron transport rate is calculated from leaf nitrogen content and incident irradiance).
  • Buckley, Environmental Biology Group, Research School of Biological Sciences, The Australian National University, GPO Box 475, Canberra City, ACT 2601, Australia and Cooperative Research Centre for Greenhouse Accounting, RSBS, ANU E-mail: tom_buckley@alumni.jmu.edu (email)
  • Miller, Environmental Biology Group, Research School of Biological Sciences, The Australian National University, GPO Box 475, Canberra City, ACT 2601, Australia E-mail: jmm@nn.au
  • Farquhar, Environmental Biology Group, Research School of Biological Sciences, The Australian National University, GPO Box 475, Canberra City, ACT 2601, Australia and Cooperative Research Centre for Greenhouse Accounting, RSBS, ANU E-mail: gdf@nn.au
article id 530, category Research article
Graham D. Farquhar, Thomas N. Buckley, Jeffrey M. Miller. (2002). Optimal stomatal control in relation to leaf area and nitrogen content. Silva Fennica vol. 36 no. 3 article id 530. https://doi.org/10.14214/sf.530
Keywords: stomatal conductance; optimal leaf area; optimality theory; resource substitution
Abstract | View details | Full text in PDF | Author Info
We introduce the simultaneous optimisation of water-use efficiency and nitrogen-use efficiency of canopy photosynthesis. As a vehicle for this idea we consider the optimal leaf area for a plant in which there is no self-shading among leaves. An emergent result is that canopy assimilation over a day is a scaled sum of daily water use and of photosynthetic nitrogen display. The respective scaling factors are the marginal carbon benefits of extra transpiration and extra such nitrogen, respectively. The simple approach successfully predicts that as available water increases, or evaporative demand decreases, the leaf area should increase, with a concomitant reduction in nitrogen per unit leaf area. The changes in stomatal conductance are therefore less than would occur if leaf area were not to change. As irradiance increases, the modelled leaf area decreases, and nitrogen/leaf area increases. As total available nitrogen increases, leaf area also increases. In all the examples examined, the sharing by leaf area and properties per unit leaf area means that predicted changes in either are less than if predicted in isolation. We suggest that were plant density to be included, it too would further share the response, further diminishing the changes required per unit leaf area.
  • Farquhar, Cooperative Research Centre for Greenhouse Accounting and Environmental Biology Group, Research School of Biological Sciences, Australian National University, ACT 2601, Australia E-mail: farquhar@rsbs.anu.edu.au (email)
  • Buckley, Cooperative Research Centre for Greenhouse Accounting and Environmental Biology Group, Research School of Biological Sciences, Australian National University, ACT 2601, Australia E-mail: tnb@nn.au
  • Miller, Research School of Biological Sciences, Australian National University, ACT 2601, Australia E-mail: jmm@nn.au
article id 529, category Research article
Tuula Aalto, Pertti Hari, Timo Vesala. (2002). Comparison of an optimal stomatal regulation model and a biochemical model in explaining CO2 exchange in field conditions. Silva Fennica vol. 36 no. 3 article id 529. https://doi.org/10.14214/sf.529
Keywords: photosynthesis; Scots pine; optimization; Pinus sylvestris L.; CO2 exchange; biochemical model
Abstract | View details | Full text in PDF | Author Info
Gas exchange of Pinus sylvestris L. was studied in subarctic field conditions. Aspects on optimal control of the gas exchange were examined using approach by Hari et al. (Tree Phys. 2: 169–175, 1986). Biochemical model by Farquhar et al. (Planta 149: 78–90, 1980) was utilized to describe the photosynthetic production rate of needles. The model parameters were determined from field measurements. The results from the optimization approach and biochemical model were compared and their performance was found quite similar in terms of R2 calculated using measured exchange rates (0.89 for optimization model and 0.85 for biochemical model). Minor differences were found in relation to responses to intercellular carbon dioxide concentration and temperature.
  • Aalto, Finnish Meteorological Institute, Air Quality Research, Sahaajankatu 20 E, FIN-00810 Helsinki, Finland E-mail: tuula.aalto@fmi.fi (email)
  • Hari, University of Helsinki, Dept. of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: ph@nn.fi
  • Vesala, University of Helsinki, Dept. of Physics, P.O. Box 64, FIN-00014 University of Helsinki, Finland E-mail: tv@nn.fi
article id 545, category Research article
Pete Bettinger, David Graetz, Kevin Boston, John Sessions, Woodam Chung. (2002). Eight heuristic planning techniques applied to three increasingly difficult wildlife planning problems. Silva Fennica vol. 36 no. 2 article id 545. https://doi.org/10.14214/sf.545
Keywords: forest planning; spatial harvest scheduling; adjacency constraints
Abstract | View details | Full text in PDF | Author Info
As both spatial and temporal characteristics of desired future conditions are becoming important measures of forest plan success, forest plans and forest planning goals are becoming complex. Heuristic techniques are becoming popular for developing alternative forest plans that include spatial constraints. Eight types of heuristic planning techniques were applied to three increasingly difficult forest planning problems where the objective function sought to maximize the amount of land in certain types of wildlife habitat. The goal of this research was to understand the relative challenges and opportunities each technique presents when more complex difficult goals are desired. The eight heuristic techniques were random search, simulated annealing, great deluge, threshold accepting, tabu search with 1-opt moves, tabu search with 1-opt and 2-opt moves, genetic algorithm, and a hybrid tabu search / genetic algorithm search process. While our results should not be viewed as universal truths, we determined that for the problems we examined, there were three classes of techniques: very good (simulated annealing, threshold accepting, great deluge, tabu search with 1-opt and 2-opt moves, and tabu search / genetic algorithm), adequate (tabu search with 1-opt moves, genetic algorithm), and less than adequate (random search). The relative advantages in terms of solution time and complexity of programming code are discussed and should provide planners and researchers a guide to help match the appropriate technique to their planning problem. The hypothetical landscape model used to evaluate the techniques can also be used by others to further compare their techniques to the ones described here.
  • Bettinger, Department of Forest Resources, Oregon State University, Corvallis, OR 97331 E-mail: pete.bettinger@orst.edu (email)
  • Graetz, Department of Forest Resources, Oregon State University, Corvallis, OR 97331 E-mail: dgw@nn.us
  • Boston, Carter Holt Harvey Forest Fibre Solutions, Tokoroa, New Zealand E-mail: kb@nn.nz
  • Sessions, Department of Forest Engineering, Oregon State University, Corvallis, OR 97331 E-mail: js@nn.us
  • Chung, Department of Forest Engineering, Oregon State University, Corvallis, OR 97331 E-mail: wc@nn.us
article id 544, category Research article
Perttu Anttila. (2002). Updating stand level inventory data applying growth models and visual interpretation of aerial photographs. Silva Fennica vol. 36 no. 2 article id 544. https://doi.org/10.14214/sf.544
Keywords: aerial photographs; stand level inventory; MELA; updating of inventory data; visual interpretation
Abstract | View details | Full text in PDF | Author Info
In this study two procedures for updating stand level inventory data were developed and tested. The development of the growing stock of 62 stands over 12 years was simulated in the MELA stand simulator with no prior information of rapid changes, such as clear-cuttings. The acceptability of the simulation was decided standwise with visual interpretation of aerial false-colour photographs. If the simulated data were not accepted, new stand attributes were assessed with photo interpretation in procedure 1. In procedure 2, on the other hand, it was possible to utilise old management proposals. In case a cutting or other operation had been proposed and it looked like the operation had been realised, the interpreters accepted the proposal. Otherwise the last implemented operation and implementation year were interpreted. In case no operation had been carried out during the updating period but the growth model updated data were not acceptable, the same stand characteristics were estimated as in procedure 1. Stands where a proposal had been accepted or an operation interpreted were later updated again in MELA so that the program simulated the operations. The Root Mean Squared Errors of stem volume were 62 and 57 m3 per ha (34 and 30%) with procedures 1 and 2. With procedure 2 the accuracy of updating was comparable with a stand level field inventory carried out in the study area. The productivity of the photo interpretation procedures was 57 and 84 ha per h, respectively, whereas the productivity of a field inventory has been 3.3–5 ha per h.
  • Anttila, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: perttu.anttila@joensuu.fi (email)
article id 543, category Research article
Maarten Nieuwenhuis. (2002). The development and validation of pre-harvest inventory methodologies for timber procurement in Ireland. Silva Fennica vol. 36 no. 2 article id 543. https://doi.org/10.14214/sf.543
Keywords: pre-harvest inventory; simulation; dynamic programming; value maximisation; optimal crosscutting
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This article describes the development and validation of a decision-support system for sawmill wood procurement, dealing specifically with the integration of the pre-harvest inventory procedures, site-specific dbh/height models and a generic taper equation, with a crosscutting simulator. The crosscutting simulation program faithfully mimics the process of cut-to-length harvesting and provides detailed information on the potential volume, logs count and diameter distributions for different log assortment specifications. Four data sets, consisting of a total of 4153 diameter and height measurements, were used in the validation process. The sites included two Sitka spruce clearfells, a Sitka spruce thinning and a Norway spruce clearfell. The evaluation process has shown that the developed decision-support system produced accurate results for a wide range of stand types, as long as sufficient large data sets were used, and that it provides the wood procurement manager of a sawmill with an efficient means of gaining a comprehensive insight into the yield potential of standing timber lots and, as such, represents a valuable aid to timber procurement and production planning.
  • Nieuwenhuis, University College Dublin, Dept. of Forestry, Belfield, Dublin 4, Ireland E-mail: maarten.nieuwenhuis@ucd.ie (email)
article id 542, category Research article
Håkan Lindström. (2002). Intra-tree models of juvenile wood in Norway spruce as an input to simulation software. Silva Fennica vol. 36 no. 2 article id 542. https://doi.org/10.14214/sf.542
Keywords: Picea abies; image analysis; juvenile wood; growth conditions
Abstract | View details | Full text in PDF | Author Info
Juvenile wood found in the first 5–25 growth rings of a conifer has a structure and properties that differ from mature wood. Juvenile wood is therefore said to influence processing and the end-use of sawn products. Consequently, models describing the juvenile wood content, within and between trees, could be useful in improving the utilisation and value of wood as an industrial raw material. The objective of the present study was to develop juvenile wood models, based on Norway spruce trees, which could be used within a model system for conversion simulation studies. Nineteen stands of Norway spruce (Picea abies [L.] Karst.) were selected throughout Sweden. Based on DBH, two small, two medium, and two large diameter timber trees were taken from each stand. DBH varied between 180–470 mm, tree height between 17–34 m, and total age between 51–152 years. Each selected tree was cross-cut into logs; discs were prepared from the large end of each log and from the top end of the top log. Image analysis was used to determine growth ring development on sampled discs. Using tree and growth variables, the juvenile core radius and the logarithmic value of juvenile wood percentage were modelled. The two models had an R2Adj of 0.71 and 0.88 respectively.
  • Lindström, University of Canterbury, School of Forestry, Private Bag 4800, Christchurch, New Zealand E-mail: lindstromhakan@netscape.net (email)
article id 541, category Research article
Azim U. Mallik, F. Wayne Bell, Yanli Gong. (2002). Effectiveness of delayed brush cutting and herbicide treatments for vegetation control in a seven-year-old jack pine plantation in northwestern Ontario, Canada. Silva Fennica vol. 36 no. 2 article id 541. https://doi.org/10.14214/sf.541
Keywords: species richness; Pinus banksiana; species diversity; conifer release treatments; glyphosate; brushsaw; competition index
Abstract | View details | Full text in PDF | Author Info
Efficacy of three conifer release treatments, i) single application of glyphosate (Vision™) herbicide, ii) multiple application of glyphosate herbicide, and iii) motor-manual brush cutting for controlling competing plants, particularly trembling aspen (Populus tremuloides), pin cherry (Prunus pensylvanica), green alder (Alnus viridis spp. crispa), and beaked hazel (Corylus cornuta spp. cornuta), was studied in a seven-year-old jack pine (Pinus banksiana) plantation in northwestern Ontario, Canada. The single and multiple glyphosate applications were equally effective in controlling trembling aspen and pin cherry, causing over 90% stem mortality. The brushsaw treatment caused an initial decrease followed by an increase in stem density of these two species. A high degree of stem thinning by natural mortality in the untreated control plots was observed in trembling aspen (23–46%) and pin cherry (41–69%) over four years. As with trembling aspen and pin cherry, stem density of green alder and beaked hazel initially decreased and then increased following the brushsaw treatment, mainly due to resprouting. Stem mortality in green alder and beaked hazel was 45% and 97%, respectively, two years after the operational glyphosate treatment. Competition index (CI) was low (mean CI = 52, ranging from 18 to 115) in all the plots including the untreated control. There was a significant increase in basal diameter of jack pine in the brushsaw and herbicide-treated plots compared to the control three years after the treatments. Jack pine seedlings in the brushsaw and glyphosate treated plots were taller compared to that of control but differences were not significant. Lower species richness and diversity were recorded in the herbicide-treated plots compared to the brushsaw and control plots in the third growing season following treatment.
  • Mallik, Department of Biology, Lakehead University, Thunder Bay, Ontario, Canada P7B 5E1 E-mail: azim.mallik@lakeheadu.ca (email)
  • Bell, Ontario Forest Research Institute, Ontario Ministry of Natural Resources, Sault Ste. Marie, Ontario, Canada P6A 2E5 E-mail: fwb@nn.ca
  • Gong, Department of Biology, Lakehead University, Thunder Bay, Ontario, Canada P7B 5E1 E-mail: yg@nn.ca
article id 540, category Research article
Finn H. Brække, Nagwa Salih. (2002). Reliability of foliar analyses of Norway spruce stands in a Nordic gradient. Silva Fennica vol. 36 no. 2 article id 540. https://doi.org/10.14214/sf.540
Keywords: Norway spruce; Picea abies; needle analyses; diagnostic methods; nutrient status; imbalanced nutrition; fertilisation; volume growth
Abstract | View details | Full text in PDF | Author Info
Norway spruce stands at eleven sites in Finland, Norway and Sweden have been studied under various climates, atmospheric deposition of N and S and fertilisation regimes. Nitrogen was growth restricting at eight inland sites, while P was growth restricting at three coastal sites. Liming and N fertilisation caused serious B deficiency on some of the inland sites. It is likely that liming affects uptake of B, whereas N fertilisation causes a dilution due to increased growth. Application of S combined with N probably caused K deficiency at one of the sites. The reliability of foliar analyses as a method to diagnose nutrient status and the likely changes after nutrient input to spruce forests in the Nordic countries, are discussed. The CR- and the DOP-method are evaluated for diagnostic purposes. Both methods seem to give reliable conclusions even if the CR-method often produces more specific results. Interpretation based on both current and one year old foliage improved the diagnostic prognoses. The accuracy of diagnosis also relies on knowledge and ability of the interpreter. Based on the results it is reason to be cautious about recommendations of single element fertilisations, e.g. with N alone, because the demand of other elements beyond available pools frequently occurs. Forest trees in the boreal region are probably well adapted to N deficiency, which means that they can handle the physiological consequences rather well, while deficiencies of other elements usually are more detrimental to growth vigour and stress related diseases.
  • Brække, Department of Forest Sciences, Agricultural University of Norway (AUN), P.O. Box 5044, N-1432 Ås, Norway E-mail: finn.braekke@isf.nlh.no (email)
  • Salih, Department of Ecology and Environmental Research, Swedish University of Agricultural Sciences (SLU), P.O. Box 7072, S-750 07 Uppsala, Sweden E-mail: ns@nn.se
article id 539, category Research article
Martti Venäläinen, Seppo Ruotsalainen. (2002). Procedure for managing large-scale progeny test data: a case study of Scots pine in Finland. Silva Fennica vol. 36 no. 2 article id 539. https://doi.org/10.14214/sf.539
Keywords: Pinus sylvestris; selection; progeny testing; performance level; reliability coefficient
Abstract | View details | Full text in PDF | Author Info
Large progeny test networks are typical for conventional forest tree breeding programmes. The individual progeny tests differ with respect to age, composition and ability to screen the breeding values of the parent trees. Several approaches have been introduced to manage the unbalanced and diverse nature of the data generated by progeny tests. This report presents a procedure for ranking breeding material on the basis of ‘messy’ data. Plot means were used as input values and missing plots were estimated from least squares. The differences between test means and variances were standardised by the performance level method. The different precision of the tests was quantified through the reliability coefficient. In order to facilitate the selection of plus trees for different purposes, all the available test results were combined into a single variable that was used for ranking. Three different kinds of ranking variable were calculated and each of them proved to be more useful for the selection of plus trees than an arithmetic or weighted mean. One of them, WMEAN, relied on the reliability and number of the progeny tests, while the others, WCONF0.50 and WCONF0.10, relied on the standard error of the plus tree mean, thus emphasising the precision of the values obtained. The analyses were carried out with SAS® procedures, which require only moderate skills in statistics, programming and data processing technology. The procedure has functioned well throughout an eight-year development phase. Nearly three thousand Scots pine (Pinus sylvestris) plus trees have been ranked for various characters, and the results have been used for roguing the seed orchards, to establish new ones, and to select plus trees for breeding populations.
  • Venäläinen, Finnish Forest Research Institute, Punkaharju Research Station, FIN-58450 Punkaharju, Finland E-mail: martti.venalainen@metla.fi (email)
  • Ruotsalainen, Finnish Forest Research Institute, Punkaharju Research Station, FIN-58450 Punkaharju, Finland E-mail: sr@nn.fi
article id 538, category Research article
Seppo Nevalainen. (2002). The incidence of Gremmeniella abietina in relation to topography in southern Finland. Silva Fennica vol. 36 no. 2 article id 538. https://doi.org/10.14214/sf.538
Keywords: Pinus sylvestris; forest inventory; Gremmeniella abietina; topography; digital elevation models
Abstract | View details | Full text in PDF | Author Info
Field data of the 8th National Forest Inventory (NFI) from southern Finland and digital elevation models (DEMs) were used in this study. Damage due to Gremmeniella abietina increased slightly with an increase in absolute elevation in mineral soils. Severe damage increased almost linearly with an increase in elevation in mineral soil plots. The mean elevation in the tract area (the 7 km x 8 km area surrounding the plot) was more strongly correlated with the disease than the elevation of individual plots. The relative altitude of the plot was important: the disease was most severe in the plots situated lower than the mean elevation of the tract area, especially in the peatland plots. In this group, the damage increased linearly with an increase in absolute elevation. According to detailed DEMs in the most diseased areas, steepness of the slope was negatively correlated with the disease. The aspect of the slope had a weak influence. On mineral soils, the disease was most common in south-facing slopes. The microtopography was not as important for the disease occurrence as the relative elevation of the plot. The disease frequencies were very similarly related to the three most common types of surface features (channels, ridges and planar regions) within the 50-m scale. At the cell size of 100 metres, the disease was more common in channels than in ridges, except in mineral soil plots. Topographic variables only partly explained the regional patterns in the occurrence of this disease. The disease was frequent on upland areas, but, on the other hand, it was also common on lowland areas. The most diseased areas studied in detail differed very much from each other with respect to topography and the disease incidence.
  • Nevalainen, Finnish Forest Research Institute, Joensuu Research Centre, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: seppo.nevalainen@metla.fi (email)
article id 537, category Research article
Timo Kurkela. (2002). Crown condition as an indicator of the incidence of root rot caused by Heterobasidion annosum in Scots pine stands. Silva Fennica vol. 36 no. 2 article id 537. https://doi.org/10.14214/sf.537
Keywords: Pinus sylvestris; Heterobasidion annosum; root rot; symptoms; disease incidence
Abstract | View details | Full text in PDF | Author Info
Trees in three Scots pine stands seriously infected by Heterobasidion annosum were classified according to their crown condition into four classes, from healthy to dead trees. After cutting the stands, the classification was compared with the symptoms of annosum root rot on stump surfaces (pitched area) and with the extension of decay in the roots of excavated stumps. When dead trees were included, the average crown condition on the survey plots correlated with disease incidence. Without dead trees the correlation was not significant. Slightly infected trees could not be distinguished from healthy trees on the basis of crown condition. It was concluded that only the proportion of dead and dying trees in a stand is a reliable indication of the disease incidence for making decisions about the future management.
  • Kurkela, Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: timo.kurkela@metla.fi (email)
article id 568, category Research article
Minna Malmivaara, Irja Löfström, Ilkka Vanha-Majamaa. (2002). Anthropogenic effects on understorey vegetation in Myrtillus type urban forests in southern Finland. Silva Fennica vol. 36 no. 1 article id 568. https://doi.org/10.14214/sf.568
Keywords: fragmentation; GNMDS; trampling; vegetation monitoring; wear
Abstract | View details | Full text in PDF | Author Info
The growth of urban population in Finland has resulted in increased fragmentation of urban forests and consequently increased recreational pressure on these forests. The effects of fragmentation and trampling on the ground and field layer vegetation were studied in mesic Myrtillus type Norway spruce-dominated urban forest stands of varying size in the greater Helsinki area. The number of residents living in the vicinity of the forest stands was an important factor affecting the understorey vegetation in urban forests. The cover of understorey vegetation in urban forests was remarkably lower than in rural areas, especially the ground layer cover, e.g. cover of Pleurozium schreberi, was significantly lower in urban forests than in the reference areas. Thus, the ground layer proved to be most susceptible to trampling. In the field layer, the cover of dwarf shrubs, especially of Vaccinium myrtillus, was lower in deteriorated than in undeteriorated urban forest stands.
  • Malmivaara, Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: minna.malmivaara@metla.fi (email)
  • Löfström, Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: il@nn.fi
  • Vanha-Majamaa, Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: ivm@nn.fi
article id 567, category Research article
Sonia Légaré, Yves Bergeron, David Paré. (2002). Influence of forest composition on understory cover in boreal mixedwood forests of western Quebec. Silva Fennica vol. 36 no. 1 article id 567. https://doi.org/10.14214/sf.567
Keywords: biomass; understory; canopy; composition; nutrient availability
Abstract | View details | Full text in PDF | Author Info
Forest overstory composition influences both light and nutrient availability in the mixed boreal forest. The influence of stand composition on understory cover and biomass was investigated on two soil types (clay and till deposits). Four forest composition types were considered in this study: aspen (Populus tremuloides Michx.), paper birch (Betula papyrifera Marsh.), jack pine (Pinus banksiana Lamb.) and a mixture of balsam-fir (Abies balsamea (L.) Mill.) and white spruce (Picea glauca (Moench) Voss). The cover of all understory species was recorded while the biomass of two important and ubiquitous species was measured: mountain maple (Acer spicatum Lam.) of the shrub layer and large-leaved aster (Aster macrophyllus L.) of the herb layer. Soil analyses were conducted to evaluate the influence of overstory composition on understory biomass through its influences on soil characteristics. Analyses of variance showed a significant effect of forest canopy type on mountain maple biomass, understory cover and shrub cover but not on herb cover and large-leaved aster biomass. Path analysis was performed to explore the relationships between canopy type, nutrient availability and understory biomass. Contrary to what was expected, the variation in plant biomass associated with forest composition was weakly related to soil nutrient availability and more strongly related to stand structural attributes.
  • Légaré, Université du Québec en Abitibi-Témiscamingue, Groupe de recherche en écologie forestière interuniversitaire, 445, boulevard de l'Université, Rouyn-Noranda, QC, Canada J9X 5E4 E-mail: sonia.legare@uqat.uquebec.ca (email)
  • Bergeron, NSERC-UQAT-UQAM, Industrial Chair in sustainable forest management, CP 8888, succ. Centre-Ville, Montréal, QC, Canada H3C 3P8 E-mail: yb@nn.ca
  • Paré, Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, P.O. Box 3800, Sainte-Foy, QC, Canada G1V 4C7 E-mail: dp@nn.ca
article id 566, category Research article
Philip J. Burton. (2002). Effects of clearcut edges on trees in the sub-boreal spruce zone of Northwest-Central British Columbia. Silva Fennica vol. 36 no. 1 article id 566. https://doi.org/10.14214/sf.566
Keywords: tree growth; windthrow; canopy influence; clearcut logging; edge effects; edge orientation; opening influence
Abstract | View details | Full text in PDF | Author Info
Clearcut-forest boundaries were evaluated for their effects on mature and regenerating trees in the northern interior of British Columbia, Canada. Two hundred and eighteen rectangular plots measuring 200 m2 each were arrayed in transects across 12 cutblock edges created 9 to 21 years earlier, with the wall of standing timber facing either north or south. The density of canopy trees on the inner edge was found to be reduced by 19% (on north-facing edges) to 46% (on south-facing edges) from average densities found in forest interiors. This reduction was primarily due to windthrow after logging, which was elevated by 27% (over interior background levels) at north-facing edges, and by 216% at south-facing edges. Of the trees situated within 10 m of south-facing cutblock edges, 11% of the Pinus contorta, 18% of the Abies lasiocarpa, and 42% of the Picea engelmannii x glauca trees have apparently collapsed, primarily those having height-to-dbh ratios greater than 71:1. As a result, irradiance in the forest understory was elevated (over interior levels) at south-facing edges to distances of approximately 65 to 70 m into the forest. Increased irradiance from adjacent cutblocks enhanced the understory growth of Picea for approximately 60 m into the inner edge of forests, 75 m for Abies. Mature Pinus trees on south-facing edges showed an unexplained 48% decrease in radial growth compared to average growth rates in forest interiors, an effect that was detectable up to 45 m into the forest. Elevated densities of conifer seedlings were evident for up to approximately 70 m into clearcuts from north-facing forest edges. Seedling growth in clearcuts was largely unaffected by shade from stand edges. Though the extent of edge effects varies considerably with the statistical techniques used to detect them, it appears that opening effects on trees can extend between 40 and 120 m into this forest type, while canopy effects reach shorter distances into clearcuts.
  • Burton, Symbios Research and Restoration, P.O. Box 3398, 3868 13th Avenue, Smithers, British Columbia, Canada V0J 2N0 E-mail: symbios@bulkley.net (email)
article id 565, category Research article
Sybille Haeussler, Lorne Bedford, Alain Leduc, Yves Bergeron, J. Marty Kranabetter. (2002). Silvicultural disturbance severity and plant communities of the southern Canadian boreal forest. Silva Fennica vol. 36 no. 1 article id 565. https://doi.org/10.14214/sf.565
Keywords: site preparation; species composition; species diversity; ecosystem integrity; degradation; biodiversity indicators; life history strategies
Abstract | View details | Full text in PDF | Author Info
Boreal forest ecosystems are adapted to periodic disturbance, but there is widespread concern that conventional forest practises degrade plant communities. We examined vegetation diversity and composition after clearcut logging, mechanical and chemical site preparation in eight 5- to 12-yr old studies located in southern boreal forests of British Columbia and Quebec, Canada to find useful indicators for monitoring ecosystem integrity and to provide recommendations for the development and testing of new silvicultural approaches. Community-wide and species-specific responses were measured across gradients of disturbance severity and the results were explained in terms of the intermediate disturbance hypothesis and a simple regeneration model based on plant life history strategies. Species richness was 30 to 35% higher 5 to 8 years after clearcut logging than in old forest. Total and vascular species diversity generally peaked on moderately severe site treatments, while non-vascular diversity declined with increasing disturbance severity. On more-or-less mesic sites, there was little evidence of diversity loss within the range of conventional silvicultural disturbances; however, there were important changes in plant community composition. Removing soil organic layers caused a shift from residual and resprouting understory species to ruderal species regenerating from seeds and spores. Severe treatments dramatically increased non-native species invasion. Two important challenges for the proposed natural dynamics-based silviculture will be 1) to find ways of maintaining populations of sensitive non-vascular species and forest mycoheterotrophs, and 2) to create regeneration niches for disturbance-dependent indigenous plants without accelerating non-native species invasion.
  • Haeussler, C2 Site 81 RR#2 Monckton Rd., Smithers, B.C., Canada V0J 2N0 E-mail: skeena@bulkley.net (email)
  • Bedford, B.C. Ministry of Forests, P.O. Box 9513 Stn. Prov. Govt., Victoria, B.C., Canada, V8W 9C2 E-mail: lb@nn.ca
  • Leduc, Groupe de recherche en écologie forestière interuniversitaire, Université du Québec à Montréal, CP 8888, Succursale A, Montréal, Québec, Canada, H3C 3P8 E-mail: al@nn.ca
  • Bergeron, Groupe de recherche en écologie forestière interuniversitaire, Université du Québec à Montréal, CP 8888, Succursale A, Montréal, Québec, Canada, H3C 3P8 E-mail: yb@nn.ca
  • Kranabetter, B.C. Ministry of Forests, Bag 5000, Smithers, B.C., Canada, V0J 2N0 E-mail: jmk@nn.ca
article id 564, category Research article
Brett G. Purdy, S. Ellen Macdonald, Mark R. T. Dale. (2002). The regeneration niche of white spruce following fire in the mixedwood boreal forest. Silva Fennica vol. 36 no. 1 article id 564. https://doi.org/10.14214/sf.564
Keywords: boreal forest; regeneration; natural disturbance; white spruce; mycorrhizae
Abstract | View details | Full text in PDF | Author Info
Early establishment of white spruce (Picea glauca) in mixedwood boreal forest stands following fire was examined at several times-since-fire (1-, 2-, 4-, 6-, 14-years). Abiotic and biotic conditions in the stands were assessed at two scales, tree plot (5 m x 5 m) and microsite (1 m x 1 m), along with presence, density and height of white spruce seedlings. Germination and survival of seed sown 1- and 4-years post fire were quantified. Survival and growth of nursery-grown seedlings, and mycorrhizal colonization, survival and growth of sterile seedlings, planted 1-year post-fire were assessed. At the tree plot scale, presence of white spruce seedlings 1-year post-fire could be reliably predicted by organic layer depth and distance to and strength of seed source. In contrast, none of the biotic or abiotic factors measured was strongly correlated with occurrence or density of white spruce seedlings 6- and 14- years post-fire. At the microsite scale, seedling recruitment immediately post-fire was limited to a distinct subset of available microsites (greater % cover of downed wood and moss, lower % cover of litter and herbaceous vegetation). Likewise, seedling occurrence in older burns was associated with distinct microsite conditions; although this was more likely an effect of white spruce presence, rather than the cause. Less than 3% of seed sown 1 yr post-fire survived to become 1yr old germinants, survival was 41% over the next three years. Availability of suitable regeneration microsites declines rapidly with time-since-fire; less than 0.3% of seed sown 4 yrs post-fire survived one year. High rates of mycorrhizal colonization were found on white spruce seedlings planted 1-year post-fire, including early and late stage fungal species. Microsite characteristics were only weakly correlated with mycorrhizal fungal communities. We propose that immediate post-fire recruitment of white spruce is a key process in mixedwood boreal succession.
  • Purdy, Dept. of Renewable Resources, University of Alberta, Edmonton, AB, Canada T6G 2E3 E-mail: bgp@nn.ca
  • Macdonald, Dept. of Renewable Resources, University of Alberta, Edmonton, AB, Canada T6G 2E3 E-mail: ellen.macdonald@ualberta.ca (email)
  • Dale, Dale, Dept. of Biological Sciences, University of Alberta, Edmonton, AB, Canada T6G 2E9 E-mail: mrtd@nn.ca
article id 563, category Research article
Timo Pakkala, Ilkka Hanski, Erkki Tomppo. (2002). Spatial ecology of the three-toed woodpecker in managed forest landscapes. Silva Fennica vol. 36 no. 1 article id 563. https://doi.org/10.14214/sf.563
Keywords: three-toed woodpecker; territory quality; landscape quality; critical threshold value; metapopulation theory; metapopulation capacity; Finnish National Forest Inventory
Abstract | View details | Full text in PDF | Author Info
The effects of landscape structure and forestry on the abundance and dynamics of boreal forest bird species have been studied widely, but there are relatively few studies in which the spatial structure and quality of the landscape have been related to the spatial ecology of bird species. In this paper, we present methods to measure territory and landscape quality for the three-toed woodpecker (Picoides tridactylus) and similar territorial forest bird species based on data from the Finnish multi-source national forest inventory and metapopulation theory. The three-toed woodpecker was studied with territory mapping within an area of 340 square km in southern Finland in 1987–2000. Altogether 195 breeding territory sites were observed. The spatial occurrence of the territories was aggregated, and the highest densities were observed in spruce-dominated old-growth forest areas. Both territory and landscape quality had significant consequences for the occurrence of the three-toed woodpecker. The spatial patterning and permanence of breeding and non-breeding territories were influenced by a combination of spatial dynamics of the species and the quality of the landscape, the latter being much influenced by forestry. The landscape-level spatial occurrence of the three-toed woodpecker in the study area may represent source-sink dynamics. The results of this paper suggest the presence of threshold values at different spatial scales, which may determine the occurrence of the three-toed woodpecker and similar species in managed forest landscapes.
  • Pakkala, Finnish Museum of Natural History, University of Helsinki, P.O. Box 17, FIN-00014 University of Helsinki, Finland E-mail: timo.pakkala@helsinki.fi (email)
  • Hanski, Department of Ecology and Systematics, University of Helsinki, P.O. Box 47, FIN-00014 University of Helsinki, Finland E-mail: ih@nn.fi
  • Tomppo, Finnish Forest Research Institute, Unioninkatu 40 A, FIN-00170 Helsinki, Finland E-mail: et@nn.fi
article id 562, category Research article
Tero Kokkila, Annikki Mäkelä, Eero Nikinmaa. (2002). A method for generating stand structures using Gibbs marked point process. Silva Fennica vol. 36 no. 1 article id 562. https://doi.org/10.14214/sf.562
Keywords: spatial distribution; stand simulation; Gibbs point process; Markov chain Monte Carlo
Abstract | View details | Full text in PDF | Author Info
Stand growth modelling based on single tree responses to their surroundings requires a description of the spatial structure of a stand. While such detailed information is rarely available from field measurements, a method to create it from more general stand variables is needed. A marked Gibbs point potential theory combined with Markov chain Monte Carlo (MCMC) random process was used to create a spatial configuration for any given number of trees. The trees are considered as charges rejecting each other and building ‘potential energy’. As an analogue of the potential energy in physical systems, the potential of a stand is defined in terms of size-dependent tree-to-tree interactions that can be thought of as related to resource depletion and competition. The idea that bigger trees induce larger potentials brings 3-dimensional effects into the system. Any feasible spatial structure is a state of the system, and the related potential can be calculated. The probability that a certain state occurs is assumed to be a decreasing function of its potential. Because more regular structures have lower potentials, by adjusting the steepness of the probability distribution the spatial structure can be allowed to have a lot of randomness (naturally regenerated stands) or forced to be very regular (planted stands). The MCMC algorithm is a numerical method of finding stand configurations that correspond to the expected level of the potential, given the size distribution of trees and the shape of the probability density function. The method also allows us to take into account spatial variation in the terrain. Some spots can be defined to have lower basic potential than others (ditch, planting furrow, etc.) in order to create areas of higher than average stocking density. A preliminary test of the method was conducted on two measured stands. The results suggest that the method could provide an efficient and flexible means of mimicking variable stand structures.
  • Kokkila, University of Helsinki, Department of Forest Ecology, P.O. Box 27, FIN-00014 Helsingin yliopisto, Finland E-mail: tero.kokkila@helsinki.fi (email)
  • Mäkelä, University of Helsinki, Department of Forest Ecology, P.O. Box 27, FIN-00014 Helsingin yliopisto, Finland E-mail: am@nn.fi
  • Nikinmaa, University of Helsinki, Department of Forest Ecology, P.O. Box 27, FIN-00014 Helsingin yliopisto, Finland E-mail: en@nn.fi
article id 561, category Research article
Karen A. Harper, Yves Bergeron, Sylvie Gauthier, Pierre Drapeau. (2002). Post-fire development of canopy structure and composition in black spruce forests of Abitibi, Québec: a landscape scale study. Silva Fennica vol. 36 no. 1 article id 561. https://doi.org/10.14214/sf.561
Keywords: boreal forest; Picea mariana; windthrow; canopy structural development; spruce budworm outbreaks; stand-level structure
Abstract | View details | Full text in PDF | Author Info
Fire reconstruction and forest inventory maps provided an opportunity to study changes in stand-level characteristics following fire using a data set comprised of all forest stands of fire origin in an area of over 10 000 km2. We assigned the date of the most recent fire occurrence to over 31 000 forest stands in an ecoforestry database. We categorized stands on different substrates into age classes to investigate differences in canopy composition, cover and height, and incidence of secondary disturbance. Stands with over 75% Picea mariana (Mill.) BSP dominated all age classes on organic sites. On other substrates, there was a change in canopy composition from deciduous stands and stands dominated by Pinus banksiana Lamb. to Picea mariana stands after about 100 yr. This transition was later for xeric sites. After a peak in canopy cover and height at about 100 yr, there was a decrease in the area occupied by stands with dense, tall canopies. Structural development was slower on less productive sites. There was little incidence of spruce budworm outbreaks. Partial disturbance by windthrow coincided with canopy break-up at 100 yr, but appeared to have little effect on overall canopy structure in later stages. Structural diversity was independent of compositional diversity; on organic sites, stands with similar composition had different canopy structure. Diversity of stands with different composition and structure was greatest in the first 150 yr following fire. Maintaining stands in different stages of structural development on the landscape would serve to maintain regional biodiversity.
  • Harper, Université de Québec à Montréal, Groupe de recherche en écologie forestière interuniversitaire, CP 8888, succ. A, Montréal, QC, Canada H3C 3P8 E-mail: c1444@er.uqam.ca (email)
  • Bergeron, NSERC-UQAT-UQAM, Industrial Chair in sustainable forest management, CP 8888, succ. Centre-Ville, Montréal, QC, Canada H3C 3P8 E-mail: yb@nn.ca
  • Gauthier, Canadian Forest Service, Laurentian Forestry Centre, P.O. Box 3800, Sainte-Foy, QC, Canada G1V 4C7 E-mail: sg@nn.ca
  • Drapeau, Université de Québec à Montréal, Groupe de recherche en écologie forestière interuniversitaire, CP 8888, succ. A, Montréal, QC, Canada H3C 3P8 E-mail: pd@nn.ca
article id 560, category Research article
Christopher P. Quine, Jonathan W. Humphrey, Karen Purdy, Duncan Ray. (2002). An approach to predicting the potential forest composition and disturbance regime for a highly modified landscape: a pilot study of Strathdon in the Scottish Highlands. Silva Fennica vol. 36 no. 1 article id 560. https://doi.org/10.14214/sf.560
Keywords: restoration; wind disturbance; site suitability; catchment planning
Abstract | View details | Full text in PDF | Author Info
The existing native forests of Scotland are fragmented and highly modified and none are ‘natural’. There is considerable interest in expanding the area of this oceanic boreal forest and restoring forest habitat networks to benefit biodiversity. However, unlike regions with substantial remaining natural forest, it is difficult to provide reference values for forest composition and structure using methods related to historical variability. An alternative approach is to combine models that predict woodland type from knowledge of site conditions, and disturbance regime from knowledge of the disturbance agents (particularly abiotic agents). The applicability of this approach was examined as part of a public participatory planning exercise in a highly managed landscape in Eastern Scotland. Models of site suitability (Ecological Site Classification) and wind disturbance (ForestGALES) were combined to determine potential woodland composition and structure, and derive options for native woodland expansion. The land use of the upper Strathdon catchment is currently dominated by agriculture and planted forests of non-native species, and only small fragments of semi-natural woodland remain (< 0.5% of the land area). Model results indicated that a very substantial proportion of the land area could support woodland (> 90%) but of a restricted range of native woodland types, with Scots pine communities predominant. Structural types likely to be present included wind-induced krummholz (treeline) forest, forest with frequent stand replacement by wind, and also a large area where gap phase (or some other disturbance) would predominate. The merits of the approach are discussed, together with the difficulties of validation, and the implications for the management of existing forests.
  • Quine, Forestry Commission, Northern Research Station, Roslin, Midlothian EH25 9SY, United Kingdom E-mail: chris.quine@forestry.gsi.gov.uk (email)
  • Humphrey, Forestry Commission, Northern Research Station, Roslin, Midlothian EH25 9SY, United Kingdom E-mail: jwh@nn.uk
  • Purdy, Forestry Commission, Northern Research Station, Roslin, Midlothian EH25 9SY, United Kingdom E-mail: kp@nn.uk
  • Ray, Forestry Commission, Northern Research Station, Roslin, Midlothian EH25 9SY, United Kingdom E-mail: dr@nn.uk
article id 559, category Research article
Juho Pennanen. (2002). Forest age distribution under mixed-severity fire regimes – a simulation-based analysis for middle boreal Fennoscandia. Silva Fennica vol. 36 no. 1 article id 559. https://doi.org/10.14214/sf.559
Keywords: modeling; disturbance; age structure; forest fire; landscape
Abstract | View details | Full text in PDF | Author Info
A simulation model was used to study the age structure of unmanaged forest landscapes under different fire regimes. Stand age was defined as the age of the oldest tree cohort in a stand. When most fires are not stand-replacing, the theoretical equilibrium stand age distribution is either bell-shaped or bimodal and dominated by old age-classes. Old-growth forests (oldest cohort > 150 y) dominate the landscape unless fires are both frequent and severe. Simulation results and analytical calculations show that if a regime of frequent fires (about every 50 y) maintains landscapes dominated by old-growth forests, then old-growth dominance persists when the number of fires is decreased, despite the associated increase in fire severity. Simulation results were applied to Pinus sylvestris-dominated landscapes of middle boreal Fennoscandia, which according to empirical results were dominated by old-growth forests when fires were frequent during the 19th century. Since the changes in the fire regime can be plausibly explained by changes in the number of human-caused ignitions, old-growth forests have evidently also dominated the landscapes earlier when fires were less frequent. The simulation model is used to produce plausible age distributions of middle boreal Fennoscandian forest landscapes under different historical fire regimes. In summary, the frequency of large-scale disturbance alone predicts forest landscape dynamics poorly, and the roles played by fire severity and residual stands need to be considered carefully. Maintaining and restoring old-growth structures is essential to regaining the natural variability of Fennoscandian forest landscapes.
  • Pennanen, Department of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: juho.pennanen@helsinki.fi (email)
article id 558, category Research article
Tuomo Wallenius. (2002). Forest age distribution and traces of past fires in a natural boreal landscape dominated by Picea abies. Silva Fennica vol. 36 no. 1 article id 558. https://doi.org/10.14214/sf.558
Keywords: boreal forest; disturbance dynamics; fire refugia; tree age
Abstract | View details | Full text in PDF | Author Info
Forest age distribution and occurrence of traces of past fires was studied in a natural Picea abies -dominated landscape in the Onega peninsula in north-west Russia. Forest age (maximum tree age) was determined and charcoal and fire scars were searched for in 43 randomly located study plots. In 70% of the study plots (30/43) trees older than 200 years existed. The largest 50-year age class consisted of plots with 251–300 year old forests. Traces of fires were found in all types of study plots, in forests on mineral soil as well as on peatlands. However, fire has been a rare disturbance factor, as traces of fires could not be found in 35% of the study plots (15/43). Estimated from the forest age class distribution, the fire rotation time for the whole area has been at least 300 years, but possibly considerably longer. This fire rotation time is much longer than fire history studies (largely based on examination of fire scars) commonly have reported for the average time between successive fires in Fennoscandia and Northwest Russia. The results suggest that the often stated generalisations about the importance and natural frequency of fire disturbance in boreal forests do not apply in landscapes dominated by Picea abies.
  • Wallenius, Department of Ecology and Systematics, University of Helsinki, P.O. Box 56, FIN-00014 University of Helsinki, Finland E-mail: tuomo.wallenius@helsinki.fi (email)
article id 557, category Research article
Tuomo Wallenius, Timo Kuuluvainen, Raimo Heikkilä, Tapio Lindholm. (2002). Spatial tree age structure and fire history in two old-growth forests in eastern Fennoscandia. Silva Fennica vol. 36 no. 1 article id 557. https://doi.org/10.14214/sf.557
Keywords: Pinus sylvestris; Picea abies; disturbance dynamics; spatial autocorrelation; spatial pattern
Abstract | View details | Full text in PDF | Author Info
Two near natural old-growth forests, one dominated by Picea abies and the other by Pinus sylvestris, were studied for their fire history, and spatial patterns of trees and tree ages. The spatial tree age structure and the disturbance history of the forests were examined by drawing age class maps based on mapped and aged trees and by dating fires based on fire scars, and by using spatial analyses at tree scale. The tree age structures of the Picea and Pinus dominated forests were different, mainly due to differences in fire history and sensitivity of the dominant tree species to fire. Fire histories and tree age structures of both sites have probably been affected by human in the ancient past. However, in the Picea dominated site, the fires had been severe, killing most of the trees, whereas in the Pinus dominated site the severity of fires had been more variable, leaving some Pinus and even Picea trees alive. In the Pinus dominated site, the tree age distribution was multimodal, consisting of two Pinus cohorts, which were established after fires and a later Picea regeneration. The Picea dominated site was composed of four patches of different disturbance history. In the oldest patch, the tree age distribution was unimodal, with no distinct cohorts, while a single cohort that regenerated after severe fire disturbances dominated the three other patches. In both sites the overall spatial patterns of living and dead trees were random and the proportion of spatially autocorrelated variance of tree age was low. This means that trees of different age grew more or less mixed in the forest without forming spatially distinct regeneration patches, even in the oldest patch of Picea dominated Liimatanvaara, well over 200 years after a fire. The results show that detail knowledge of disturbance history is essential for understanding the development of tree age structures and their spatial patterns.
  • Wallenius, Department of Ecology and Systematics, University of Helsinki, P.O. Box 65, FIN-00014, Helsinki, Finland E-mail: tuomo.wallenius@helsinki.fi (email)
  • Kuuluvainen, Department of Forest Ecology, University of Helsinki, P.O. Box 27, FIN-00014, Helsinki, Finland E-mail: tk@nn.fi
  • Heikkilä, Research Centre of Friendship Park, Tönölä, FIN-88900 Kuhmo, Finland E-mail: rh@nn.fi
  • Lindholm, Finnish Environment Institute, Nature and Land Use Division, P.O. Box 140, FIN-00251 Helsinki, Finland E-mail: tl@nn.fi
article id 556, category Research article
Timo Kuuluvainen, Juha Mäki, Leena Karjalainen, Hannu Lehtonen. (2002). Tree age distributions in old-growth forest sites in Vienansalo wilderness, eastern Fennoscandia. Silva Fennica vol. 36 no. 1 article id 556. https://doi.org/10.14214/sf.556
Keywords: forest dynamics; boreal forest; regeneration; age structure; fire disturbance
Abstract | View details | Full text in PDF | Author Info
The age and size of trees was sampled and measured on eight sample plots (0.2 ha each) within a Pinus sylvestris -dominated boreal forest landscape in Vienansalo wilderness, Russian Karelia. The fire history of these plots was obtained from a previous dendrochronological study. All the studied sample plots showed a wide and uneven distribution of tree ages, but the shape of the age distributions of trees as well as tree species composition varied substantially. Trees over 250 years of age occurred in every studied plot, despite its small size. This suggests that old Pinus were common and rather evenly distributed in the landscape matrix. The oldest Pinus tree was 525 years of age. The correlations between tree age and size were often weak or even nil. In Pinus the correlation between age and diameter was stronger than that between age and height. In the dominant tree species Pinus and Picea, the largest trees were not the oldest trees. The tree age distributions together with the fire history data indicated that the past fires have not been stand replacing, as many of the older Pinus had survived even several fires. Tree age classes that had regenerated after the last fire were most abundant and dominated by Picea and/or deciduous trees, while the trees established before the last fire were almost exclusively Pinus. The results suggest that periodic occurrence of fire is important for the maintenance of the Pinus-dominated landscape. This is because fire kills most Picea and deciduous trees and at the same time enhances conditions for Pinus regeneration, facilitated by available seed from the continuous presence of old fire-tolerant Pinus trees.
  • Kuuluvainen, Department of Forest Ecology, University of Helsinki, P.O. Box 24, FIN-00014 University of Helsinki, Finland E-mail: timo.kuuluvainen@helsinki.fi (email)
  • Mäki, Department of Forest Ecology, University of Helsinki, P.O. Box 24, FIN-00014 University of Helsinki, Finland E-mail: jm@nn.fi
  • Karjalainen, Department of Forest Ecology, University of Helsinki, P.O. Box 24, FIN-00014 University of Helsinki, Finland E-mail: lk@nn.fi
  • Lehtonen, Faculty of Forestry, University of Joensuu, P.O. Box 24, FIN-80101 Joensuu, Finland E-mail: hl@nn.fi
article id 555, category Research article
Leena Karjalainen, Timo Kuuluvainen. (2002). Amount and diversity of coarse woody debris within a boreal forest landscape dominated by Pinus sylvestris in Vienansalo wilderness, eastern Fennoscandia. Silva Fennica vol. 36 no. 1 article id 555. https://doi.org/10.14214/sf.555
Keywords: forest dynamics; disturbances; dead wood; Scots pine; tree death
Abstract | View details | Full text in PDF | Author Info
The amount, variability, quality and spatial pattern of coarse woody debris (CWD) on mineral soil sites was studied within a natural Pinus sylvestris L. dominated boreal forest landscape in Russian Viena Karelia. Data on the total CWD was collected on 27 sample plots (20 m x 100 m) and data on large CWD was surveyed along four transects (40 m wide and up to 1000 m long). The mean volume of CWD (standing and down combined) was 69.5 m3 ha–1, ranging from 22.2 m3 ha–1 to 158.7 m3 ha–1 from plot to plot. On average, 26.9 m3 ha–1 (39%) of CWD was standing dead wood and 42.7 m3 ha–1 (61%) down dead wood. The CWD displayed a wide range of variation in tree species, tree size, stage of decay, dead tree type and structural characteristics, creating a high diversity of CWD habitats for saproxylic organisms. Large CWD was almost continuously present throughout the landscape and its overall spatial distribution was close to random, although a weak autocorrelation pattern was found at distances less than about 50 m. On small spatial scales total CWD showed wide variation up to a sample area of about 0.1 ha, beyond which the variation stabilized. The fire history variables of the sample plots were not related to the amount of CWD. This and the spatial pattern of CWD suggest that the CWD dynamics in this landscape was not driven by fire, but by more or less random mortality of trees due to autogenic causes of death.
  • Karjalainen, Department of Forest Ecology, P.O. Box 24, FIN-00014 University of Helsinki, Finland E-mail: lk@nn.fi
  • Kuuluvainen, Department of Forest Ecology, P.O. Box 24, FIN-00014 University of Helsinki, Finland E-mail: timo.kuuluvainen@helsinki.fi (email)
article id 554, category Research article
Seppo Rouvinen, Timo Kuuluvainen, Juha Siitonen. (2002). Tree mortality in a Pinus sylvestris dominated boreal forest landscape in Vienansalo wilderness, eastern Fennoscandia. Silva Fennica vol. 36 no. 1 article id 554. https://doi.org/10.14214/sf.554
Keywords: dead wood; disturbance dynamics; spatial pattern analysis; mortality agent; forest structure; temporal variation
Abstract | View details | Full text in PDF | Author Info
Tree mortality and its causes and spatial pattern were examined along four transects (width 40 m, length 2550–3960 m), with a total length of 12 190 m and area of 48.8 ha, in a Pinus sylvestris L. dominated, boreal forest landscape. Tree mortality was determined within a time window of 3 years by identifying those trees (dbh ≥ 10 cm) along the transects that fitted into one of the three categories: 1) current mortality: trees that had died during the year of survey (1998), 2) recent mortality: trees that had died during the year (1997) before the survey year, and 3) predicted mortality: trees that were expected to die during the year (1999) following the survey year. Long-term tree mortality was studied on 10 plots (20 m x 100 m) by dating 87 dead trees using dendrochronological methods. The mean current mortality was 1.4 m3 ha–1 (3.7 trees ha–1). Both the recent and predicted mortalities were also 1.4 m3 ha–1. Mortality was, on the average, higher on peatlands than on mineral soils. The highest mortality was found within an area recently flooded by beavers. Over half of the examined trees (52%) were judged to die without any visible signs of an external abiotic cause. At the landscape scale, tree mortality was continuous although somewhat aggregated in space. Of the 66 dated standing dead Pinus trees, 23 (35%) had died during the 19th century and two during the 18th century, demonstrating that dead Pinus can remain standing for long periods of time before falling. Our results show that autogenic mortality of individual trees or small groups of trees was the predominant mode of disturbance in this Pinus dominated landscape.
  • Rouvinen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: seppo.rouvinen@forest.joensuu.fi (email)
  • Kuuluvainen, University of Helsinki, Department of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: tk@nn.fi
  • Siitonen, Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: juha.siitonen@metla.fi
article id 582, category Research article
Kari Kangas, Pasi Markkanen. (2001). Factors affecting participation in wild berry picking by rural and urban dwellers. Silva Fennica vol. 35 no. 4 article id 582. https://doi.org/10.14214/sf.582
Keywords: non-wood forest products; wild berries; participation; household behaviour
Abstract | View details | Full text in PDF | Author Info
The purpose of this study was to examine the participation of urban and rural dwellers in the activity of berry-picking. The respondents in the study lived in the city of Joensuu and in the municipality of Ilomantsi, in eastern Finland. 68% of Joensuu households compared with 82% of those in Ilomantsi participated in berry-picking. These evident differences in the participation rates may be largely due to the higher costs incurred by urban dwellers in picking, since the probability of participation was not significantly higher for Ilomantsi households compared with those in Joensuu who had access to a summer-cottage which was likely to be located near the berry resources. In both municipalities, the participants were divided into two groups according to the nature of their participation in the activity. The larger group – termed ordinary pickers – were characteristically younger families with children, while the other group, termed active pickers, were distinctly more advanced in age. The quantities picked for home consumption by the groups of pickers in Ilomantsi were twice as large as those picked by the corresponding groups in Joensuu. In Joensuu, households were not significantly involved in commercial picking.
  • Kangas, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: kari.kangas@joensuu.fi (email)
  • Markkanen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: pm@nn.fi
article id 581, category Research article
Henrik Heräjärvi. (2001). Technical properties of mature birch (Betula pendula and B. pubescens) for saw milling in Finland. Silva Fennica vol. 35 no. 4 article id 581. https://doi.org/10.14214/sf.581
Keywords: silver birch; log; white birch; technical properties; saw milling; small-sized log
Abstract | View details | Full text in PDF | Author Info
The purpose of this study was to investigate the variation in selected technical properties of mature (age > 60 years) birch stems in southern and central Finland. Technical properties were defined as the natural external characteristics that cause differences in the usability of a certain section of stem in the mechanical wood industry, saw milling in particular. On mineral soils, birch stems in mixed stands were slightly larger than those in pure birch stands. On peatlands, however, birch stems in pure stands were larger than those in mixed stands. The average stem form of silver birch was straighter than that of white birch. Small-sized log sections of white birch, as well as those of codominant silver birch, typically contain many dead knots. On mineral soils, coniferous admixture had a positive effect on self-pruning of white birch. Self-pruning of silver birch was as good in pure birch stands as in mixed stands of spruce and birch. Occurrence of decay did not differ significantly between the two birch species. Not only silver birch, due to the growth and yield of the stand, but also vigorous and good-quality white birch, because of the possibility to provide high-quality logs, can be maintained profitably as an admixture in coniferous forests until final cutting.
  • Heräjärvi, The Finnish Forest Research Institute, Joensuu Research Centre, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: henrik.herajarvi@metla.fi (email)
article id 580, category Research article
Susanna Sironen, Annika Kangas, Matti Maltamo, Jyrki Kangas. (2001). Estimating individual tree growth with the k-nearest neighbour and k-Most Similar Neighbour methods. Silva Fennica vol. 35 no. 4 article id 580. https://doi.org/10.14214/sf.580
Keywords: pine; spruce; single tree growth models; non-parametric models; local estimates
Abstract | View details | Full text in PDF | Author Info
The purpose of this study was to examine the use of non-parametric methods in estimating tree level growth models. In non-parametric methods the growth of a tree is predicted as a weighted average of the values of neighbouring observations. The selection of the nearest neighbours is based on the differences between tree and stand level characteristics of the target tree and the neighbours. The data for the models were collected from the areas owned by Kuusamo Common Forest in Northeast Finland. The whole data consisted of 4051 tally trees and 1308 Scots pines (Pinus sylvestris L.) and 367 Norway spruces (Picea abies Karst.). Models for 5-year diameter growth and bark thickness at the end of the growing period were constructed with two different non-parametric methods: the k-nearest neighbour regression and k-Most Similar Neighbour method. Diameter at breast height, tree height, mean age of the stand and basal area of the trees larger than the subject tree were found to predict the diameter growth most accurately. The non-parametric methods were compared to traditional regression growth models and were found to be quite competitive and reliable growth estimators.
  • Sironen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: susanna.sironen@forest.joensuu.fi (email)
  • Kangas, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: ak@nn.fi
  • Maltamo, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: mm@nn.fi
  • Kangas, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: jk@nn.fi
article id 579, category Research article
Matti Maltamo, Kalle Eerikäinen. (2001). The Most Similar Neighbour reference in the yield prediction of Pinus kesiya stands in Zambia. Silva Fennica vol. 35 no. 4 article id 579. https://doi.org/10.14214/sf.579
Keywords: stand development; difference equations; non-parametric regression; plantation forests
Abstract | View details | Full text in PDF | Author Info
The aim of the study was to develop a yield prediction model using the non-parametric Most Similar Neighbour (MSN) reference method. The model is constructed on stand level but it contains information also on tree level. A 10-year projection period was used for the analysis of stand growth. First, the canonical correlation matrix was calculated for the whole study material using stand volumes at the beginning and at the end of the growth period as independent variables and stand characteristics as dependent variable. Secondly, similar neighbour estimates were searched from the data categories reclassified according to thinnings. Due to this, it was possible to search for growth and yield series which is as accurate as possible both at the beginning and at the end of the growth period. The reliability of the MSN volume predictions was compared to the volumes predicted with the simultaneous yield model. The MSN approach was observed to be more reliable volume predictor than the traditional stand level yield prediction model both in thinned and unthinned stands.
  • Maltamo, Faculty of Forestry, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: matti.maltamo@forest.joensuu.fi (email)
  • Eerikäinen, Faculty of Forestry, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: ke@nn.fi
article id 578, category Research article
Kevin Boston, Pete Bettinger. (2001). Development of spatially feasible forest plans: a comparison of two modeling approaches. Silva Fennica vol. 35 no. 4 article id 578. https://doi.org/10.14214/sf.578
Keywords: forest planning; heuristics; linear programming; wildlife goals
Abstract | View details | Full text in PDF | Author Info
Spatial goals are becoming more frequent aspects of forest management plans as regulatory and organizational policies change in response to fisheries and wildlife concerns. The combination of green-up constraints (harvesting restrictions that prevent the cutting of adjacent units for a specified period of time) and habitat requirements for red-cockaded woodpeckers (RCW) in the southeastern U.S. suggests that spatially feasible forest plans be developed to guide management activities. We examined two modeling approaches aimed at developing management plans that had both harvest volume goals, RCW habitat, and green-up constraints. The first was a two-stage method that in one stage used linear programming to assign volume goals, and in a second stage used a tabu search – genetic algorithm heuristic technique to minimize the deviations from the volume goals while maximizing the present net revenue and addressing the RCW and green-up constraints. The second approach was a one-stage procedure where the entire management plan was developed with the tabu search – genetic algorithm heuristic technique, thus it did not use the guidance for timber volume levels provided by the LP solution. The goal was to test two modeling approaches to solving a realistic spatial harvest scheduling problem. One is where to volume goals are calculated prior to developing the spatially feasible forest plan, while the other approach simultaneously addresses the volume goals while developing the spatially feasible forest plan. The resulting forest plan from the two-stage approach was superior to that produced from the one-stage approach in terms of net present value. The main point from this analysis is that heuristic techniques may benefit from guidance provided by relaxed LP solutions in their effort to develop efficient forest management plans, particularly when both commodity production and complex spatial wildlife habitat goals are considered. Differences in the production of forest products were apparent between the two modeling approaches, which could have a significant effect on the selection of wood processing equipment and facilities.
  • Boston, Forest Fibre Solutions, Carter Holt Harvey, Tokoroa, New Zealand E-mail: kevin.boston@chh.co.nz (email)
  • Bettinger, Department of Forest Resources, Oregon State University, Corvallis, OR 97331 E-mail: pb@nn.us
article id 577, category Research article
Jouni Siipilehto. (2001). Effect of weed control with fibre mulches and herbicides on the initial development of spruce, birch and aspen seedlings on abandoned farmland. Silva Fennica vol. 35 no. 4 article id 577. https://doi.org/10.14214/sf.577
Keywords: Populus tremula; Betula pendula; Picea abies; glyphosate; dichlobenile; herbicide; mulching; terbuthylazine; weed control
Abstract | View details | Full text in PDF | Author Info
Post-planting weed control methods on abandoned farmland were studied in three field trials in southern Finland using a completely randomized design with four treatments and 30 to 40 replications. Mulches of 60 x 60 cm [sheet mulch – strips of plane waste and plastic fibre, newspaper – waste paper slurry, wood chips, pure wood fibre slurry], herbicides [i.e. glyphosate or terbuthylazine alone or mixed and dichlobenile applied to 1 m2 spots] and hoeing treatments were compared to an untreated control plot. The study material consisted of two-year-old containerized aspen (Populus tremula L.), silver birch (Betula pendula (L.) Roth) and Norway spruce (Picea abies (L.) Karst.) seedlings planted in spring 1996. The ground vegetation was dominated by Elymys repens, Deschampsia cespitosa, Cirsium arvense and Epilobium angustifolium. Monitoring of the trials over a 3-year period showed a moderate effect of weed control, which varied according to the method used and by the crop species. Significant growth responses were found with herbicide in spruce, wood chips in spruce and birch and with sheet mulch in aspen seedlings. Sheet mulch also encouraged vole nesting thus increasing damages. Generally, slurry mulches proved to be insufficiently durable. Mulching had a clear insulating effect, which may increase the risk of winter drought.
  • Siipilehto, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: jouni.siipilehto@metla.fi (email)
article id 576, category Research article
Anneli Viherä-Aarnio, Pirkko Velling. (2001). Micropropagated silver birches (Betula pendula) in the field – performance and clonal differences. Silva Fennica vol. 35 no. 4 article id 576. https://doi.org/10.14214/sf.576
Keywords: micropropagation; herbivory; Betula pendula Roth; clone; clonal variation; field testing
Abstract | View details | Full text in PDF | Author Info
Micropropagated and seed-born silver birches (Betula pendula Roth) were compared for survival, height growth and occurrence of biotic damage (voles, hares, mooses, stem lesions and cankers) in field trials in southern Finland. The material consisted of 11 clones and 10 different lots of seedlings growing in 10 field trials, established in clear-cut forest cultivation areas. The plants were 6–7 years old. The micropropagated and seed-born material types did not significantly differ from each other as regards survival, height growth and frequencies of damage caused by biotic agents. Large and significant differences were, however, detected in survival, height and frequencies of all types of biotic damage between single clones. Careful selection and testing of birch clones in field conditions is recommended before wide-scale commercial micropropagation and practical forest cultivation takes place.
  • Viherä-Aarnio, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box. 18, FIN-01301 Vantaa, Finland E-mail: anneli.vihera-aarnio@metla.fi (email)
  • Velling, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box. 18, FIN-01301 Vantaa, Finland E-mail: pv@nn.fi
article id 575, category Research article
Sylvie Mussche, Roeland Samson, Lieven Nachtergale, An De Schrijver, Raoul Lemeur, Noël Lust. (2001). A comparison of optical and direct methods for monitoring the seasonal dynamics of leaf area index in deciduous forests. Silva Fennica vol. 35 no. 4 article id 575. https://doi.org/10.14214/sf.575
Keywords: Leaf Area Index; mixed deciduous forest; LAI-2000 plant canopy analyser; hemispherical photography; litter trap
Abstract | View details | Full text in PDF | Author Info
During the 1996 growing season the seasonal dynamics of the Leaf Area Index (LAI) were determined by 3 different methods in two forest types: a mixed oak (Quercus robur L.) – beech (Fagus sylvatica L.) stand and an ash dominated (Fraxinus excelsior L.) stand. The results obtained from the two indirect methods, i.e. hemispherical photography and LAI-2000 Plant Canopy Analyser (Li-COR), were compared with the results of the direct measurement of litter fall collected in litter trap systems. In this study the direct method is considered to be the reference, giving the most accurate LAI-values. Both the hemispherical photography and the LAI-2000 PCA introduced an underestimation of LAI when the actual canopy leaf distribution in the crown layer deviates from a random distribution of leaf area in space as is found in the mixed oak/beech stand. However, when the condition of random leaf distribution is nearly fulfilled (ash stand), the LAI-2000 PCA gave LAI-values which were close to the results obtained from the direct method. Regression curves with R2 > 0.93 could be calculated for both indirect methods.
  • Mussche, Laboratory of Forestry, Ghent University, Geraardsbergse Steenweg 267, B-9090 Melle, Belgium E-mail: sm@nn.be
  • Samson, Laboratory of Plant Ecology, Ghent University, Coupure Links 653, B-9000 Gent, Belgium E-mail: rs@nn.be
  • Nachtergale, Laboratory of Forestry, Ghent University, Geraardsbergse Steenweg 267, B-9090 Melle, Belgium E-mail: ln@nn.be
  • De Schrijver, Laboratory of Forestry, Ghent University, Geraardsbergse Steenweg 267, B-9090 Melle, Belgium E-mail: An.Deschrijver@rug.ac.be (email)
  • Lemeur, Laboratory of Plant Ecology, Ghent University, Coupure Links 653, B-9000 Gent, Belgium E-mail: rl@nn.be
  • Lust, Laboratory of Forestry, Ghent University, Geraardsbergse Steenweg 267, B-9090 Melle, Belgium E-mail: nl@nn.be
article id 574, category Research article
Hans W. Linderholm. (2001). Climatic influence on Scots pine growth on dry and wet soils in the central Scandinavian mountains, interpreted from tree-ring width. Silva Fennica vol. 35 no. 4 article id 574. https://doi.org/10.14214/sf.574
Keywords: Scots pine; tree-rings; climatic influence; periodicity
Abstract | View details | Full text in PDF | Author Info
Tree rings are one of the most important proxy data sources for reconstructing past climate variability. In order to understand climate variability, it is necessary to get a spatial and temporal coverage of climate information. Summer temperatures mainly influence tree growth at the altitudinal tree line, while at lower altitudes additional factors affect growth. In addition, the nature of soil where trees grow may affect growth response to climate. To decide climate as well as growth-substrate influences on Scots pine (Pinus sylvestris L.) growing below the tree line, two tree-ring width chronologies, sampled at dry mineral soil and wet peat soil in a mountain valley in the central Scandinavian Mountains, were analysed for climate responses and spectral signals. Temperatures during growth season (May–August) showed the strongest influence on tree growth at both sites. Influence of precipitation in the growing season was low, indicating sufficient amounts of available water during growth. However, at the dry-soil site the influence of late winter/early spring precipitation was significant. Strength of the climate–tree–growth relationship at the dry site was similar to that of trees growing at the present tree line, while weaker at the wet site. Both site chronologies exhibited common spectral peaks at c. 3.5 and 13 years indicating a common growth forcing at those time scales. The wet-site chronology displayed low-frequency variations with a 19-year periodicity, where growth peaks coincided with the lunar tidal maxima indicating a possible influence of lunar forcing. At the dry-site, multi-decadal fluctuations displayed a periodicity of 66 years. Both 13- and 66-year periods can be linked to variations in sea surface temperatures of the North Atlantic Ocean, pointing to a maritime influence, on decadal scales, of pine growth in the area. These results suggest that Scots pine in this environment may be regarded as proxies of North Atlantic Ocean coupled climatic variability.
  • Linderholm, Department of Physical Geography and Quaternary Geology, Stockholm University, 106 91 Stockholm, Sweden E-mail: hasse@natgeo.su.se (email)
article id 590, category Research article
Anna Saarsalmi, Eino Mälkönen, Sirpa Piirainen. (2001). Effects of wood ash fertilization on forest soil chemical properties. Silva Fennica vol. 35 no. 3 article id 590. https://doi.org/10.14214/sf.590
Keywords: Pinus sylvestris; wood ash; Picea abies; nutrients; acidity; neutralization; pH
Abstract | View details | Full text in PDF | Author Info
The effects of wood ash fertilization on soil chemical properties were studied in three young Scots pine (Pinus sylvestris L.) plantations and a Norway spruce (Picea abies (L.) Karst.) plantation with different site fertility in southern Finland. The dose of 3 t ha–1 of loose wood ash was applied to 4 replicate plots (25 x 25 m) at each experiment. Humus layer and mineral soil samples were taken before the treatment and 7 and 16 years after wood ash application. Results showed that neutralization as well as fertilization effects of wood ash on forest soil were of long duration. An ash-induced pH increase of 0.6–1.0 pH units and exchangeable acidity (EA) decrease of 58–83% were detected in the humus layer 16 years after wood ash application. The decrease in acidity was most pronounced on the Calluna site with initially the lowest pH and highest EA. In the mineral soil the increase in pH was observed later than in the humus layer. After 16 years, the mineral soil pH was increased (0.2–0.3 pH units) on the Vaccinium and Myrtillus sites. A corresponding and in most cases a significant increase in the extractable Ca and Mg concentrations was detected in both the humus layer and in the mineral soil. Wood ash significantly increased the effective cation exchange capacity (CECe) and base saturation (BS) but decreased the concentration of exchangeable Al in both soil layers on all the sites. No response of N availability to wood ash application could be found.
  • Saarsalmi, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: anna.saarsalmi@metla.fi (email)
  • Mälkönen, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: em@nn.fi
  • Piirainen, Finnish Forest Research Institute, Joensuu Research Station, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: sp@nn.fi
article id 589, category Research article
Pekka Mäkinen. (2001). Competitive strategies applied by Finnish timber carriers following deregulation. Silva Fennica vol. 35 no. 3 article id 589. https://doi.org/10.14214/sf.589
Keywords: timber carriers; competitive strategy; success factors; deregulation
Abstract | View details | Full text in PDF | Author Info
The present study examines the success of timber carriers and the factors involved in their success immediately following deregulation. In Finland in 1991 the timber trucking sector was deregulated. Means testing was changed to suitability testing, which meant that the Ministry of Transport and Communications, provincial authorities and the trucking association could no longer regulate the entry of new entrepreneurs to the sector. The present research material contains two successful enterprise groups. In the strategically more successful group, good results were obtained with a moderate labour input by the entrepreneurs. The strategic position of this group was considered to be successful because the operating hours of the trucks were fairly high but the work loads imposed on the entrepreneur remained reasonable. The profitability of these enterprises was so good that it was possible to use hired labour to drive the trucks. The work load of close to half of the unsuccessful entrepreneurs had been large or extremely large. In some cases, the obvious reason for failure was their inadequate transportation rates. Others had seemingly satisfactory haulage rates when compared to the average, but still their enterprises performed poorly. In these cases, the explanation lay in the inefficiency of operations or excessive debts, the latter caused, for example, by earlier operations. The results of this study do not support the view that a lot of hard work generally means success in entrepreneurship. The results support the view that both entrepreneurs’ work and management inputs have a significant impact on the success of the enterprise, and that high tariffs alone are not a guarantee of success.
  • Mäkinen, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa E-mail: pekka.makinen@metla.fi (email)
article id 588, category Research article
Marjut Ihalainen, Timo Pukkala. (2001). Modelling cowberry (Vaccinium vitis-idaea) and bilberry (Vaccinium myrtillus) yields from mineral soils and peatlands on the basis of visual field estimates. Silva Fennica vol. 35 no. 3 article id 588. https://doi.org/10.14214/sf.588
Keywords: Vaccinium myrtillus; Vaccinium vitis-idaea; non-wood forest products
Abstract | View details | Full text in PDF | Author Info
This study presents new models for predicting bilberry and cowberry yields from site and stand characteristics. These models enable one to evaluate the future states of forests in terms of berry yields. The modelling data consisted of visual field estimates of site and tree stand characteristics, as well as berry yields from 627 forest stands. Berry yields were estimated using a scale from 0 to 10. Using these data, models were prepared which predict the berry yield scores from those site and stand characteristics which are usually known in forest planning calculations. The model predictions correlated positively and often quite strongly with earlier models. The results were in line with previous studies on the effects of site and tree cover on berry production. According to the models, sites of medium and rather poor fertility produce the highest bilberry yields. Increasing tree height increases, and the basal area of spruce and proportion of deciduous trees decrease, bilberry yield. With mineral soils, cowberry yields are best on poor sites. A high proportion of pine improves cowberry yields. The yields are the highest in open areas and very young stands, on the one hand, and in sparsely populated stands of large and old trees, on the other hand. In pine swamps, the yields are best on rather poor sites. Increasing basal area of deciduous trees decreases cowberry yields.
  • Ihalainen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: marjut.ihalainen@forest.joensuu.fi (email)
  • Pukkala, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: tp@nn.fi
article id 587, category Research article
Erkki Tomppo, Kari T. Korhonen, Juha Heikkinen, Hannu Yli-Kojola. (2001). Multi-source inventory of the forests of the Hebei Forestry Bureau, Heilongjiang, China. Silva Fennica vol. 35 no. 3 article id 587. https://doi.org/10.14214/sf.587
Keywords: models; forest inventory; satellite images; k-nearest neighbour method; China
Abstract | View details | Full text in PDF | Author Info
A multi-source forest inventory method is applied to the estimation of forest resources in the area of the Hebei Forest Bureau in Heilongjiang province in North-East China. A stratified systematic cluster sampling design was utilised in field measurements. The design was constructed on the basis of information from earlier stand-level inventories, aerial orthophotographs, experiences from other sampling inventories and the available budget. Sample tree volumes were estimated by means of existing models. New models were constructed and their parameters estimated for tallied tree volumes and volume increments. The estimates for the area of the Bureau were computed from field measurements, and for the areas of the forest farms estimated from field measurements and satellite images. A k-nearest neighbour method was utilised. This method employing satellite image data makes it possible to estimate all variables, particularly for smaller areas than that possible using field measurements only. The methods presented, or their modifications, could also be applied to the planning and realisation of forest inventories elsewhere in Temperate or Boreal zones. The inventory in question gave an estimate of 114 m3/ha (the multi-source inventory 119 m3/ha) instead of 72 m3/ha as previously estimated from available information. Totally nineteen tree species, genera of species or tree species groups were identified (Appendix 1). The forests were relatively young, 60% of them younger than 40 years and 85% younger than 60 years.
  • Tomppo, Finnish Forest Research Institute, Unioninkatu 40 A, FIN-00170 Helsinki, Finland E-mail: erkki.tomppo@metla.fi (email)
  • Korhonen, Finnish Forest Research Institute, Unioninkatu 40 A, FIN-00170 Helsinki, Finland E-mail: ktk@nn.fi
  • Heikkinen, Finnish Forest Research Institute, Unioninkatu 40 A, FIN-00170 Helsinki, Finland E-mail: jh@nn.fi
  • Yli-Kojola, Finnish Forest Research Institute, Unioninkatu 40 A, FIN-00170 Helsinki, Finland E-mail: hyk@nn.fi
article id 586, category Research article
Tuula Nuutinen, Seppo Kellomäki. (2001). A comparison of three modelling approaches for large-scale forest scenario analysis in Finland. Silva Fennica vol. 35 no. 3 article id 586. https://doi.org/10.14214/sf.586
Keywords: MELA; carbon budget; forestry model; scenario modelling; EFISCEN; SIMA
Abstract | View details | Full text in PDF | Author Info
Forests play an important role in the sequestration of carbon dioxide and the storage of carbon. The potential and efficiency of mitigation options in forestry have been studied using large-scale forestry scenario models. In Finland, three models have been applied in attempts to estimate timber production and related carbon budgets. In this study, these models are compared. The oldest, MELA, was designed in the 1970s for the regional and national analysis of timber production. The European Forest Information Scenario Model, EFISCEN, originally a Swedish area matrix model, was developed in the early 1980s. SIMA, a gap-type ecosystem model, was utilised in the 1990s for regional predictions on how the changing climate may affect forest growth and timber yield in Finland. In EFISCEN, only the development of growing stock is endogeneous because the assumptions on growth, and the removal and rules for felling are given exogeneously. In the SIMA model, the rules for felling are exogeneous but the growth is modelled based on individual trees reacting to their environment. In the MELA model, the management of forests is endogeneous, i.e. the growth, felling regimes and the development of growing stock are the results of the analysis. The MELA approach integrated with a process-based ecosystem model seems most applicable in the analyses of effective mitigation measures compatible with sustainable forestry under a changing climate. When using the scenarios for the estimation of carbon budget, the policy makers should check that the analyses cover the whole area of interest, and that the assumptions on growth and management together with the definitions applied correspond with the forestry conditions in question.
  • Nuutinen, Finnish Forest Research Institute, Joensuu Research Centre, Box 68, FIN-80101 Joensuu, Finland E-mail: tuula.nuutinen@metla.fi (email)
  • Kellomäki, University of Joensuu, Box 111, FIN-80101 Joensuu, Finland E-mail: sk@nn.fi
article id 585, category Research article
Meinrad Rohner, Klaus Böswald. (2001). Forestry development scenarios: timber production, carbon dynamics in tree biomass and forest values in Germany. Silva Fennica vol. 35 no. 3 article id 585. https://doi.org/10.14214/sf.585
Keywords: carbon balance; forestry scenario modeling; forest valuation
Abstract | View details | Full text in PDF | Author Info
The dynamics of the age class structure stands at the center of modeling long-run forestry scenarios. This insight has been applied to the construction of the Forest Development and Carbon Budget Simulation Model (ForCaBSiM), a model which is used for the study of several interrelated questions: the development of timber stocks and the potential level of sustainable harvests, the stocks and fluxes of tree carbon in managed forests, the economy-wide effects of management practices on the value of forest lands and timber stocks. The combined study of these issues allows to assess development scenarios with regard to the productive potential of forestry, the carbon cycle, and forest values. At present, the model is adapted to German data, but it is designed for use with other data sets as well. This paper provides a description of core mechanisms in ForCaBSiM. On this background, the choice and impact of crucial assumptions is examined. Illustrative results are used to demonstrate the use of the model. The paper focuses on the impact of varying rotation ages and the tree species composition. Particular attention is given to the concept of steady states.
  • Rohner, Renewable Resource Modeling, D-63477 Maintal, Germany E-mail: rohner@rrmodeling.de (email)
  • Böswald, Factor Consulting + Management AG, CH-8045 Zurich, Switzerland E-mail: kb@nn.ch
article id 584, category Research article
Ralph Alig, Darius Adams, John Mills, Richard Haynes, Peter Ince, Robert Moulton. (2001). Alternative projections of the impacts of private investment on southern forests: a comparison of two large-scale forest sector models of the United States. Silva Fennica vol. 35 no. 3 article id 584. https://doi.org/10.14214/sf.584
Keywords: forest sector; timber supply; forest resource assessment; plantation area
Abstract | View details | Full text in PDF | Author Info
The TAMM/NAPAP/ATLAS/AREACHANGE (TNAA) system and the Forest and Agriculture Sector Optimization Model (FASOM) are two large-scale forestry sector modeling systems that have been employed to analyze the U.S. forest resource situation. The TNAA system of static, spatial equilibrium models has been applied to make 50-year projections of the U.S. forest sector for more than 20 years. Much of its input on forest management behavior and decisions about use of forestland derives from expert-based systems external to the TNAA system. FASOM, a spatial intertemporal optimization model, directly incorporates decisions on management investment and land use options relative to agricultural alternatives as endogenous model elements. The paper contrasts projections of private forest investment from the TNAA and FASOM models, focusing on the southern United States. Comparison of the TNAA base case and an investment-restricted scenario from FASOM, both of which reflect a continuation of recent behavioral tendencies by nonindustrial private owners, suggests that Southern private timberlands have considerable biological and economic potential for intensified forest management. Unrestricted FASOM projections confirm that added investment could lead to substantially larger timber harvest volumes and lower prices than those projected in the base/restricted cases. But even under the more intensive investment scenarios, naturally regenerated forests would cover three-quarters of the future private timberland base and hardwoods would continue to dominate the inventory structure.
  • Alig, USDA Forest Service, Forestry Sciences Lab, 3200 SW Jefferson Way, Corvallis, Oregon 97331, USA E-mail: ralig@fs.fed.us (email)
  • Adams, College of Forestry, Oregon State University, Corvallis, Oregon 97331, USA E-mail: da@nn.us
  • Mills, USDA Forest Service, Forestry Sciences Lab, 1221 SW Yamhill, Portland, Oregon 97205, USA E-mail: jm@nn.us
  • Haynes, USDA Forest Service, Forestry Sciences Lab, 1221 SW Yamhill, Portland, Oregon 97205, USA E-mail: rh@nn.us
  • Ince, USDA Forest Service, Forest Products Lab, Madison, Wisconsin 53705, USA E-mail: pi@nn.us
  • Moulton, USDA Forest Service (retired), Research Triangle Park, North Carolina 27709, USA E-mail: rm@nn.us
article id 583, category Research article
Hans Fredrik Hoen, Tron Eid, Petter Økseter. (2001). Timber production possibilities and capital yields from the Norwegian forest area. Silva Fennica vol. 35 no. 3 article id 583. https://doi.org/10.14214/sf.583
Keywords: forest management; timber production possibilities; rate of return; capital yield
Abstract | View details | Full text in PDF | Author Info
How intensely should a forest be grown? This is a fundamental question in the process of formulating policy guidelines for the management of a forest area, both at the individual property level as well as at the national level. The question is related to a number of factors; the objective(s) of the forest owner, the productivity of the forestland, the initial growing stock, the accessibility within the forest, assumptions regarding future prices and costs and the required real rate of return. This paper presents an applied analysis with the objective of mapping possible future paths for the growing stock on, and timber harvest from the productive forest area in Norway. The analysis is deterministic. The regeneration strategy is a key factor for the long run development of a forest and is thus given particular attention. The analysis is restricted to deal with timber production only and maximisation of the net present value of the forest area is used as the objective function. The required real rate of return is varied and used as the driving force to find the best (optimal) level of intensity in silvicultural management and thus optimal paths for harvesting and growing stocks.
  • Hoen, Agricultural University of Norway, Dept. of Forest Sciences, P.O. Box 5044, N-1432 Ås, Norway E-mail: hans.hoen@isf.nlh.no (email)
  • Eid, Agricultural University of Norway, Dept. of Forest Sciences, P.O. Box 5044, N-1432 Ås, Norway E-mail: te@nn.no
  • Økseter, Agricultural University of Norway, Dept. of Forest Sciences, P.O. Box 5044, N-1432 Ås, Norway E-mail: po@nn.no
article id 598, category Research article
A. Maarit I. Kallio. (2001). Interdependence of the sawlog, pulpwood and sawmill chip markets: an oligopsony model with an application to Finland. Silva Fennica vol. 35 no. 2 article id 598. https://doi.org/10.14214/sf.598
Keywords: wood market; oligopsony; forest industry
Abstract | View details | Full text in PDF | Author Info
The interdependence of the markets for pulpwood, sawlogs and sawmill chips is analysed using a short-run model, which accommodates the alternative competition structures of wood buyers. We propose that imperfect competition in the pulpwood market tends to make the sawmills owned by the pulp and paper companies larger than the independent ones, even in the absence of transactional economies of integration. The impact of the wood market competition pattern on the profits of the forest owners and forest industry firms depends upon a firm-capacity structure, wood supply elasticities, and business cycles in the output markets. The numerical application of the model to the Finnish softwood market suggests that inflexibility of production capacities tends to make the wood demand rather insensitive with respect to price. Only the large firms, which all produce both pulp and sawnwood, may have oligopsony power under some conditions. Integrated production can increase competition in the sawlog market via the wood chip market.
  • Kallio, Helsinki School of Economics, Department of Economics and Management Science, P.O. Box 1210, FIN-00101 Helsinki E-mail: maarit.kallio@hkkk.fi (email)
article id 597, category Research article
Annika Kangas, Jyrki Kangas, Jouni Pykäläinen. (2001). Outranking methods as tools in strategic natural resources planning. Silva Fennica vol. 35 no. 2 article id 597. https://doi.org/10.14214/sf.597
Keywords: uncertainty; fuzzy relations; multicriteria decision support; multiple-use planning
Abstract | View details | Full text in PDF | Author Info
Two outranking methods, ELECTRE III and PROMETHEE II, commonly used as decision-aid in various environmental problems, and their applications to decision support for natural resources management are presented. These methods represent ‘the European school’ of multi-criteria decision making (MCDM), as opposed to ‘the American school’, represented by, for instance, the AHP method. On the basis of a case study, outranking methods are compared to so far more usually applied techniques based on the ideas of multi attribute utility theory (MAUT). The outranking methods have been recommended for situations where there is a finite number of discrete alternatives to be chosen among. The number of decision criteria and decision makers may be large. An important advantage of outranking methods, when compared to decision support techniques most often applied in today’s natural resources management, is the ability to deal with ordinal and more or less descriptive information on the alternative plans to be evaluated. Furthermore, the uncertainty concerning the values of the criterion variables can be taken into account using fuzzy relations, determined by indifference and preference thresholds. The difficult interpretation of the results, on the other hand, is the main drawback of the outranking methods.
  • Kangas, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: annika.kangas@metla.fi (email)
  • Kangas, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: jk@nn.fi
  • Pykäläinen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: jp@nn.fi
article id 596, category Research article
Sakari Tuominen, Simo Poso. (2001). Improving multi-source forest inventory by weighting auxiliary data sources. Silva Fennica vol. 35 no. 2 article id 596. https://doi.org/10.14214/sf.596
Keywords: forest inventory; remote sensing; two-phase sampling; weighting
Abstract | View details | Full text in PDF | Author Info
A two-phase sampling design has been applied to forest inventory. First, a large number of first phase sample plots were defined with a square grid in a geographic coordinate system for two study areas of about 1800 and 4500 ha. The first phase sample plots were supplied by auxiliary data of Landsat TM and IRS-1C with principal component transformation for stratification and drawing the second phase sample (field sample). Proportional allocation was used to draw the second phase sample. The number of field sample plots in the two study areas was 300 and 380. The local estimates of five continuous forest stand variables, mean diameter, mean height, age, basal area, and stem volume, were calculated for each of the first phase sample plots. This was done separately by using one auxiliary data source at a time together with the field sample information. However, if the first phase sample plot for which the stand variables were to be estimated was also a field sample plot, the information of that field sample plot was eliminated according to the cross validation principle. This was because it was then possible to calculate mean square errors of estimates related to a specific auxiliary data source. The procedure produced as many estimates for each first phase sample plot and forest stand variable as was the number of auxiliary data sources, i.e. seven estimates: These were based on Landsat TM, IRS-1C, digitized aerial photos, ocular stereoscopic interpretation from aerial photographs, data from old forest inventory made by compartments, Landsat TM95–TM89 difference image and IRS96–TM95 difference image. The final estimates were calculated as weighted averages where the weights were inversely proportional to mean square errors. The alternative estimates were calculated by applying simple rules based on knowledge and the outliers were defined. The study shows that this kind of system for finding outliers for elimination and a weighting procedure improves the accuracy of stand variable estimation.
  • Tuominen, Finnish Forest Research Institute, Unioninkatu 40 A, FIN-00170 Helsinki, Finland E-mail: sakari.tuominen@metla.fi (email)
  • Poso, Department of Forest Resource Management, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: sp@nn.fi
article id 595, category Research article
Kenneth Nyström, Göran Ståhl. (2001). Forecasting probability distributions of forest yield allowing for a Bayesian approach to management planning. Silva Fennica vol. 35 no. 2 article id 595. https://doi.org/10.14214/sf.595
Keywords: basal area growth model; mixed-model; uncertainty of predictions; Monte Carlo simulation
Abstract | View details | Full text in PDF | Author Info
Probability distributions of stand basal area were predicted and evaluated in young mixed stands of Scots pine (Pinus sylvestris L.), Norway spruce (Picea abies (L.) Karst.) and birch (Betula pendula Roth and Betula pubescens Ehrh.) in Sweden. Based on an extensive survey of young stands, individual tree basal area growth models were estimated using a mixed model approach to account for dependencies in data and derive the variance/covariance components needed. While most of the stands were reinventoried only once, a subset of the stands was revisited a second time. This subset was used to evaluate the accuracy of the predicted stand basal area distributions. Predicting distributions of forest yield, rather than point estimates, allows for a Bayesian approach to planning and decisions can be made with due regard to the quality of the information.
  • Nyström, SLU, Department of Forest Resource Management and Geomatics, SE-901 83 Umeå, Sweden E-mail: kenneth.nystrom@resgeom.slu.se (email)
  • Ståhl, SLU, Department of Forest Resource Management and Geomatics, SE-901 83 Umeå, Sweden E-mail: gs@nn.se
article id 594, category Research article
Walter Zucchini, Matthias Schmidt, Klaus von Gadow. (2001). A model for the diameter-height distribution in an uneven-aged beech forest and a method to assess the fit of such models. Silva Fennica vol. 35 no. 2 article id 594. https://doi.org/10.14214/sf.594
Keywords: diameter-height distribution; mixture models; bivariate normal distribution; SBB distribution; goodness-of-fit; pseudo-residuals; beech forest
Abstract | View details | Full text in PDF | Author Info
This paper illustrates the application of a mixture model to describe the bivariate diameter-height distribution of trees growing in a pure, uneven-aged beech forest. A mixture of two bivariate normal distributions is considered but the methodology is applicable to mixtures of other distributions. The model was fitted to diameter-height observations for 1242 beech trees in the protected forest Dreyberg (Solling, Germany). A considerable advantage of the model, apart from the fact that it happens to fit this large data set unusually well, is that the individual parameters all have familiar interpretations. The bivariate Johnson SBB distribution was also fitted to the data for the purpose of comparing the fits. A second issue discussed in this paper is concerned with the general question of assessing the fit of models for bivariate data. We show how a device called ‘pseudo-residual’ enables one to investigate the fit of a bivariate model in new ways and in considerable detail. Attractive features of pseudo-residuals include the fact that they are not difficult to interpret; they can be computed using generally available statistical software and, most important of all, they enable one to examine the fit of a model by means of simple graphs.
  • Zucchini, Georg-August-Universität Göttingen, Institute for Statistics and Econometrics E-mail: zucchini@wi-wiss.uni-goettingen.de (email)
  • Schmidt, Forest Research Station of Lower Saxony E-mail: ms@nn.de
  • Gadow, Georg-August-Universität Göttingen, Institute for Forest Management and Yield Sciences E-mail: kvg@nn.de
article id 593, category Research article
Anneli Jalkanen. (2001). The probability of moose damage at the stand level in southern Finland. Silva Fennica vol. 35 no. 2 article id 593. https://doi.org/10.14214/sf.593
Keywords: National Forest Inventory; Alces alces; multilevel logistic regression; risk management
Abstract | View details | Full text in PDF | Author Info
The probability of moose damage was studied in sapling stands and young thinning stands in southern Finland. Data from the eighth National Forest Inventory in 1986–92 were used for modelling. The frequency of damage was highest at the height of two to five meters and at the age of ten to twenty years (at the time of measurement). Moose preferred aspen stands the most and least preferred Norway spruce stands. Scots pine and silver birch were also susceptible to damage. Logistic regression models were developed for predicting the probability that moose damage is the most important damaging agent in a forest stand. The best predictive variables were the age and dominant species of the stand. Variables describing the site were significant as cluster averages, possibly characterizing the area as a food source (fertility and organic soil), as well as the lack of shelter (wall stand). When sample plot, cluster and municipality levels were compared, it was found that most of the unexplained variance was at the cluster level. To improve the model, more information should be obtained from that level. The regression coefficients for aspen as supplementary species, and for pine as dominant species, had significant variance from cluster to cluster (area to area). It was also shown that the occurrence of aspen is closely connected to the occurrence of moose damage in pine sapling stands.
  • Jalkanen, Finnish Forest Research Institute, Unioninkatu 40 A, FIN-00170 Helsinki, Finland E-mail: anneli.jalkanen@metla.fi (email)
article id 592, category Research article
Marja-Liisa Juntunen. (2001). Use of pesticides in Finnish forest nurseries in 1996. Silva Fennica vol. 35 no. 2 article id 592. https://doi.org/10.14214/sf.592
Keywords: herbicides; fungicides; insecticides; seedlings; survey study
Abstract | View details | Full text in PDF | Author Info
In 1996 a questionnaire on seedling production and use of pesticides was sent to 33 forest nurseries in Finland. Twenty-eight nurseries answered the questionnaire; thus the survey covered about 80% of the Finnish production of forest seedlings. According to this study, the Finnish nurseries together are using about 1000 kilograms of pesticides (as active ingredient, a.i.) annually. The most used herbicide was terbutylazine (Gardoprim-Neste®), and half of the total amount of fungicide used was chlorothalonil (Bravo 500®). Three fourths of the insecticide products had permethrin as the active ingredient. The nurseries applied, on average, 1.7 kg pesticides (a.i.)/ha annually, although the amount varied considerably between nurseries. In production of container seedlings the highest mean amounts of pesticides were applied to pine seedlings (9.5 kg/ha) and the lowest to spruce seedlings (0.9 kg/ha). To the fields of bareroot seedlings the nurseries applied, on average, 3.9 kg pesticides (a.i.)/ha. Mean amounts of pesticide (a.i.) per 1000 seedlings grown in containers were almost the same for birch and pine production, 1.6 and 1.7 grams, respectively; for production of spruce seedlings the comparable values were less than 0.5 grams. For production of bareroot seedlings the nurseries used about four times more pesticides than for container seedlings.
  • Juntunen, The Finnish Forest Research Institute, Suonenjoki Research Station, FIN-77600 Suonenjoki, Finland E-mail: marja-liisa.juntunen@metla.fi (email)
article id 591, category Research article
Qibin Yu. (2001). Can physiological and anatomical characters be used for selecting high yielding hybrid aspen clones? Silva Fennica vol. 35 no. 2 article id 591. https://doi.org/10.14214/sf.591
Keywords: photosynthesis; growth; leaf; stomatal characters
Abstract | View details | Full text in PDF | Author Info
Stomatal, CO2 exchange parameters and several leaf and growth traits were recorded in a five-year-old hybrid aspen clone trial. The field trial consisted of four aspen hybrid clones (Populus tremula L. x P. tremuloides Michx.) and one local Populus tremula seedling source. The mean estimated height of hybrid aspen clones was 1.6 times higher than for P. tremula. Similarly, basal diameter was 1.5 times and breast diameter 1.8 times higher in the hybrids. Differences were observed for physiological and growth traits among hybrid clones and P. tremula, but, only stomatal characters of hybrid clones differed significantly from those in P. tremula. Hybrid clones had larger guard cells (22.9 mm) than P. tremula (19.2 mm), whereas P. tremula had a higher stomatal density (211.3/mm2) than the hybrid clones (164.4/mm2). Among four hybrid clones, net photosynthesis was strongly correlated with foliar nitrogen. Height correlated significantly with foliar nitrogen, but negatively with leaf fresh weight, leaf dry weight and stomatal density. The results suggested that yield components could be controlled by many genes, specific to each clone. No single gas exchange or morphological variable can provide a reliable indicator of yield potential.
  • Yu, Department of Plant Biology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: qibin.yu@helsinki.fi (email)
article id 607, category Research article
Jurkka Kuusipalo. (2001). Plastic coating of plywood using extrusion technique. Silva Fennica vol. 35 no. 1 article id 607. https://doi.org/10.14214/sf.607
Keywords: plywood; thermoplastics; polymer; polyethylene; polypropylene; extrusion coating; adhesion
Abstract | View details | Full text in PDF | Author Info
The target was to study and develop industrially suitable methods to apply thermoplastic coatings on plywood. Typically, wear and climatic resistance for wooden sheets, such as chip board and plywood, is achieved by thermosetting resin coating (e.g. phenol-formaldehyde). However, economical and environmental issues are driving towards thermoplastic coatings (e.g. polyethylene) in such products. Several technologies, such as film gluing and laminating, have been introduced, but the costs and processing are limiting factors. This study concentrates on direct coating of plywood with extruded molten plastic film, in other words extrusion coating of plywood. This technique provides fast coating process and cheap raw materials. In the study it was found that extrusion coating of plywood is difficult. The high heat energy of the molten plastic causes moisture evaporation of the top veneer, and the only possible escape direction of the evaporated moisture is through the plastic, which causes bubbling in the plastic coating. Another disadvantage of the plywood is its rigidity, which limits the development of the process. In this work, a new technique was applied for coating the flexible veneers with typical extrusion coating equipment, where the escape of the evaporated moisture was possible through the top veneer to the atmosphere. Another main issue was to reach sufficient adhesion between plastic and plywood. Target was achieved using either monolayer polyethylene (PE-LD) or 2-layer polypropylene/adhesion polymer plastic coatings.
  • Kuusipalo, Tampere University of Technology, Institute of Paper Converting, Korkeakoulunkatu 4 C, P.O. Box 541, FIN-33720 Tampere, Finland E-mail: jurkka@cc.tut.fi (email)
article id 606, category Research article
Pekka Leskinen, Jyrki Kangas. (2001). Modelling future timber price development by using expert judgments and time series analysis. Silva Fennica vol. 35 no. 1 article id 606. https://doi.org/10.14214/sf.606
Keywords: forest planning; uncertainty; expert knowledge; forecasting; price peaks; price trend; price variation
Abstract | View details | Full text in PDF | Author Info
Timber prices belong to the most important variables affecting the optimality of forest management. On the other hand, forecasting of timber prices is very uncertain. One difficulty when using past time series data in forecasting future timber price development is the possibility of changes in the markets and in the society at large. Expert knowledge can be applied in forecasting of timber prices as information additional to that provided by time series modelling. This paper presents an approach utilising both time series data and expert judgments in modelling future timber prices. A time series model is used as the basis for the approach. Parameters describing future timber price trends, variation in future timber prices, and the probabilities of price peaks taking place in the future are estimated with expert judgments as the basis. A case study involving 12 experts was carried out in Finland, and models were estimated for all the six major timber assortments in the country. The model produced can be utilised in the optimisation calculations of forest planning.
  • Leskinen, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: pekka.leskinen@metla.fi (email)
  • Kangas, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: jk@nn.fi
article id 605, category Research article
Paul C. Van Deusen. (2001). Scheduling spatial arrangement and harvest simultaneously. Silva Fennica vol. 35 no. 1 article id 605. https://doi.org/10.14214/sf.605
Keywords: simulated annealing; adjacency constraints; Metropolis algorithm
Abstract | View details | Full text in PDF | Author Info
A method based on the Metropolis algorithm is developed for creating desirable spatial configurations on the landscape while simultaneously dealing with other objectives commonly associated with harvest scheduling. Spatial configurations are loosely specified and stochastically attained, which contrasts with other adjacency constraints based on specific block size limits. This method makes it possible to improve habitat and connectivity, and to create buffer zones as part of the scheduling process. It also works with a mapped set of polygons/forest stands and does not require a gridded system.
  • Van Deusen, NCASIS, Northeast Regional Center, 600 Suffolk Street, Fifth Floor, Lowell, MA 01854, USA E-mail: pvandeusen@ncasi.org (email)
article id 604, category Research article
Dan Glöde, Ulf Sikström. (2001). Two felling methods in final cutting of shelterwood, single-grip harvester productivity and damage to the regeneration. Silva Fennica vol. 35 no. 1 article id 604. https://doi.org/10.14214/sf.604
Keywords: silviculture; time study; cost; forest operations; felling technique; logging-damage
Abstract | View details | Full text in PDF | Author Info
In order to find an efficient and careful way of final-cutting shelterwoods, two felling methods, in a single-grip harvester system, were compared with respect to productivity and damage caused to the regeneration. The shelterwood (140–165 m3/ha) consisted of Norway spruce (Picea abies (L.) Karst.) and Scots pine (Pinus sylvestris L.) and the natural regeneration (9530–11 780 seedlings/ha) mostly of Norway spruce. Treatments were: (i) conventional felling on both sides of the harvester striproad, preferably in blanks of the regeneration; (ii) felling of the trees top-end first into the striproad using a method named “tossing the caber”. Both treatments included forwarding after felling. Conventional felling had a non-significantly higher productivity (27.4 m3/E15–h) and lower cost (25.9 SEK/m3) than tossing the caber (26.1 m3/E15–h and 27.2 SEK/m3). However, tossing the caber was significantly more efficient in the felling and processing of pine trees compared with conventional felling. The mean proportions of the disappeared and damaged seedlings were approximately 40% after both treatments. The logging-related damage to the regeneration decreased with increased distance to the striproad in the tossing the caber treatment but not in conventional felling. The conclusions were that there were no differences between the treatments regarding productivity, cost and total damage to the regeneration in mixed conifer shelterwoods but that tossing the caber could be a more productive method than conventional felling in pine dominated stands. Tossing the caber could also be beneficial at a regeneration height of 2–3 m since at this height the damage to the regeneration seems less than at conventional felling.
  • Glöde, SkogForsk, Uppsala Science Park, S-751 83 Uppsala, Sweden E-mail: dan.glode@skogforsk.se (email)
  • Sikström, SkogForsk, Uppsala Science Park, S-751 83 Uppsala, Sweden E-mail: us@nn.se
article id 603, category Research article
Kari Minkkinen, Jukka Laine, Hannu Hökkä. (2001). Tree stand development and carbon sequestration in drained peatland stands in Finland – a simulation study. Silva Fennica vol. 35 no. 1 article id 603. https://doi.org/10.14214/sf.603
Keywords: carbon; peatland; drainage; growth models
Abstract | View details | Full text in PDF | Author Info
Drained peatland forests form an important timber resource in Finland. They also form a sink for atmospheric carbon (C) because of the increased growth and C sequestration rates following drainage. These rates have, however, been poorly quantified. We simulated the tree stand dynamics for drained peatland stands with and without cuttings over two stand rotations. Simulations were done on four peatland site types and two regions in Finland with different climatic conditions, using recently published peatland tree growth models applied in a stand simulator. We then calculated the amount of C stored in the stands on the basis of previously published tree-level biomass and C content models. Finally, we developed regression models to estimate C stores in the tree stands using stand stem volume as the predictor variable. In the managed stands, the mean growth (annual volume increment) ranged from 2 to 9 m3 ha–1 a–1, depending on the rotation (first/second), site type and region. Total yield during one rotation varied from 250 to 920 m3 ha–1. The maximum stand volumes varied from 220 to 520 m3 ha–1 in the managed stands and from 360 to 770 m3 ha–1 in the unmanaged. By the end of the first post-drainage rotation the total C store in the managed stands had increased by 6–12 kg C m–2 (i.e. 45–140 g C m–2 a–1) compared to that in the undrained situation. Averaged over two rotations, the increase in the total C store was 3–6 kg C m–2. In the corresponding unmanaged stands the C stores increased by 8–15 kg m–2 over the same periods. At stand level, the C stores were almost linearly related to the stem volume and the developed regression equations could explain the variation in the simulated C stores almost entirely.
  • Minkkinen, University of Helsinki, Department of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: kari.minkkinen@helsinki.fi (email)
  • Laine, University of Helsinki, Department of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: jl@nn.fi
  • Hökkä, Finnish Forest Research Institute, Rovaniemi Research Station, P.O. Box 16, FIN-96301 Rovaniemi, Finland E-mail: hh@nn.fi
article id 602, category Research article
Teijo Nikkanen. (2001). Reproductive phenology in a Norway spruce seed orchard. Silva Fennica vol. 35 no. 1 article id 602. https://doi.org/10.14214/sf.602
Keywords: Picea abies; flowering; receptive period; pollination; reproductive synchronisation
Abstract | View details | Full text in PDF | Author Info
Reproductive phenology was studied in a Norway spruce seed orchard, located in southern Finland (62°13’N, 25°24’E), consisting of 67 clones from northern Finland (64°–67°N). Timing of flowering was determined on the basis of data recorded by a pollen catch meter during 1984–1995, and visual observations made on grafts in 1989, 1992, 1993 and 1995. The genetic and environmental factors affecting female and male phenology, and reproductive synchronisation were studied. The between-year variation in the timing of flowering was more than three weeks. However, when it was defined on the basis of the effective temperature sum, the variation was smaller. No phenological reproductive isolation was found between the seed orchard and surrounding natural forests. The duration of the receptive period of the seed orchard varied from 5 to 8 days, and anthesis determined on the basis of airborne pollen from 5 to 10 days. The receptive period started about one day earlier than anthesis, except in one abnormally warm flowering period when female and male flowering started simultaneously. In general, the flowering periods of the different clones overlapped. The clonal differences in the phenology of receptivity were in most cases statistically significant, but in pollen shedding they were not. The broad-sense heritability estimates were higher for female than for male phenology. Environmental factors, conversely, had a stronger effect on male phenology. A wide graft spacing and a graft position that favoured solar radiation on the lower parts of the crown promoted early pollen shedding and, subsequently, better reproductive synchronisation between female and male flowering.
  • Nikkanen, Finnish Forest Research Institute, Punkaharju Research Station, FIN-58451 Punkaharju, Finland E-mail: teijo.nikkanen@metla.fi (email)
article id 601, category Research article
Ricardo Alía, Javier Moro-Serrano, Eduardo Notivol. (2001). Genetic variability of Scots pine (Pinus sylvestris) provenances in Spain: growth traits and survival. Silva Fennica vol. 35 no. 1 article id 601. https://doi.org/10.14214/sf.601
Keywords: Spain; Scots pine; genetic variation; provenances; GE interaction
Abstract | View details | Full text in PDF | Author Info
Plants obtained from seed of 16 Spanish and 6 German provenances of Scots pine (Pinus sylvestris L.) were installed at five trial sites distributed throughout the natural range of the species in Spain. Five years after planting (7 years of age) the experimental material was measured for total height, diameter, number of twigs at the fourth year whorl and survival. The analysis confirmed that the rate of height growth of the Spanish is lower than that of the German provenances, whereas for the other traits the best Spanish compare favourably with the Germans. Provenance by site interaction was very significant (P < 0.01) for most traits. Attempts to model the interaction of Spanish provenances on height by simultaneous introduction of some climatic and geographic covariates on both factors were not successful but a multiplicative model with one bilinear term was enough to provide a sensible explanation of this interaction. Usually, provenances closest to each trial site were found better adapted than more distant ones but some provenances of close origin presented a different behaviour. Processes of adaptation and selection of these ancient populations could be considered as the main factors to cause this interaction.
  • Alía, CIFOR-INIA, Unidad de Mejora Forestal, 28080 Madrid, Spain E-mail: ra@nn.es
  • Moro-Serrano, CIFOR-INIA, Unidad de Mejora Forestal, 28080 Madrid, Spain E-mail: jmoro@inia.es (email)
  • Notivol, Unidad de Recursos Forestales, SIA-DGA, Ca de Montañana 179, 50080 Zaragoza, Spain E-mail: en@nn.es
article id 600, category Research article
Qibin Yu, P. M. A. Tigerstedt, Matti Haapanen. (2001). Growth and phenology of hybrid aspen clones (Populus tremula L. x Populus tremuloides Michx.). Silva Fennica vol. 35 no. 1 article id 600. https://doi.org/10.14214/sf.600
Keywords: phenology; growth rate; hybrid aspen; growth pattern
Abstract | View details | Full text in PDF | Author Info
Height, basal diameter, diameter at breast height, bud burst, and leaf development were recorded in a 5-year-old hybrid aspen clonal trial. The field trial consisted of four aspen hybrid clones (Populus tremula x Populus tremuloides) and one local P. tremula seedling source. Phenological traits were observed in the 3rd year. Growth patterns were recorded during the 3rd and 4th years. Phenological traits were explored in relation to hybrid vigor expressed as growth traits. Differences were observed for phenological and growth traits among hybrid clones and P. tremula. The growth period varied from 143–158 days for the four hybrid clones, and was 112 days for P. tremula. The correlation between growth period and yield was highly significant. The annual growth rate of height for the hybrids was 4.2 cm per 7 days (2.4 for P. tremula) in the 3rd year and 6.4 cm per 7 days (2.9 for P. tremula) in the 4th year. After 5 years, mean estimated stem volume of the hybrids was 3.9 times that of P. tremula. Significant clone by year interaction was observed for height, diameter, and volume growth. The hybrid vigor seems to be mainly attributable to a longer growth period.
  • Yu, Department of Plant Biology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: qibin.yu@helsinki.fi (email)
  • Tigerstedt, Department of Plant Biology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: pmat@nn.fi
  • Haapanen, Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: mh@nn.fi
article id 599, category Research article
Chun-Jiang Liu, Carl J. Westman, Hannu Ilvesniemi. (2001). Matter and nutrient dynamics of pine (Pinus tabulaeformis) and oak (Quercus variabilis) litter in North China. Silva Fennica vol. 35 no. 1 article id 599. https://doi.org/10.14214/sf.599
Keywords: Pinus; Quercus; decomposition; nutrients; fall; forest floor; litter; North China
Abstract | View details | Full text in PDF | Author Info
In the mountainous area of North China, a distinguishing feature of climate is the serious drought of spring and the humidity and high temperature of summer by which the litter production and decomposition of forest litter were strongly characterized. We investigated the dynamic and nutrient characteristics of litter in a 30-year-old mixed stand of Chinese pine (Pinus tabulaeformis Carr.) and deciduous orient oak (Quercus variabilis Bl.) and two comparable pure stands. Oak litterfall peaked in November and pine litterfall in December. The oak stand had the largest annual litterfall (347 g m–2) and the forest floor mass (950 g m–2), the mixed stand the second (236 g m–2 and 634 g m–2), and the pine stand the least (217 g m–2 and 615 g m–2). The nutrient return through litterfall and the storage in forest floor followed corresponding order between three stands. The weight loss of pine and oak foliage litter in first year was 25% and 20%. For senesced pine and oak leaves, the translocation rates of N, P and K were 56–83%. Nutrient concentrations were higher in oak leaf litter than pine needle litter, and the concentration of N and Ca appeared to rise while K concentration decreased in both decomposing litter.
  • Liu, Department of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: cliu@silvia.helsinki.fi (email)
  • Westman, Department of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: cjw@nn.fi
  • Ilvesniemi, Department of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: hi@nn.fi
article id 623, category Research article
Anders Roos, Matti Flinkman, Armas Jäppinen, Mats Warensjö. (2000). Adoption of value-adding processes in Swedish sawmills. Silva Fennica vol. 34 no. 4 article id 623. https://doi.org/10.14214/sf.623
Keywords: sawmilling; production structure; technology adoption
Abstract | View details | Full text in PDF | Author Info
Adding value to lumber by processing it after sawing and standard drying is one means for the sawmilling industry to increase market shares in competition with other materials, e.g. glass, steel, concrete, aluminium, and plastics. In this study the adoption patterns of value-adding processes used in Swedish softwood sawmills were analysed based on production data from 1995. About 90% of the sawmills applied a value-adding process after initial sawing and drying, and 72% of the sawmills applied two or more processes. The total share of processed sawnwood was about 40%. Important dimensions of value-adding processes are: extra drying and production of blanks for doors/windows and for furniture; surface-treatment, mainly planing, which is sometimes associated with preservation and painting; length trimming and pallet production; extra drying and production of edge-glued panels and laminated beams; and stress grading and production of building components. The association of different value-adding dimensions with location, ownership and production characteristics were investigated. The total share of value-added production were higher for private sawmills than for mills owned by forest companies or by forest owners’ associations, and it was higher for mills in southern Sweden than for sawmills in other parts of the country. Value-added share does not clearly correlate with mill size or with the dominating tree species being sawn.
  • Roos, Swedish University of Agricultural Sciences, Department of Forest Management and Products, P.O. Box 7060, SE-750 07 Uppsala, Sweden E-mail: anders.roos@sh.slu.se (email)
  • Flinkman, Swedish University of Agricultural Sciences, Department of Forest Management and Products, P.O. Box 7060, SE-750 07 Uppsala, Sweden E-mail: mf@nn.se
  • Jäppinen, Swedish University of Agricultural Sciences, Department of Forest Management and Products, P.O. Box 7060, SE-750 07 Uppsala, Sweden E-mail: aj@nn.se
  • Warensjö, Swedish University of Agricultural Sciences, Department of Forest Management and Products, P.O. Box 7060, SE-750 07 Uppsala, Sweden E-mail: mw@nn.se
article id 622, category Research article
Håkan Lindström. (2000). Intra-tree models of basic density in Norway spruce as an input to simulation software. Silva Fennica vol. 34 no. 4 article id 622. https://doi.org/10.14214/sf.622
Keywords: Picea abies; wood density; image analysis; basic density; growth conditions
Abstract | View details | Full text in PDF | Author Info
Basic density is said to influence aspects of conversion, properties, and end-use of forest products. Consequently, it is argued that accurate models of basic density variation, within and between trees, could be used to improve the utilisation of wood as an industrial raw material. The objective of the present study was to develop basic density models based on Norway spruce trees, that could be used within a model system for conversion simulation studies. Nineteen stands of Norway spruce (Picea abies (L.) Karst.) were selected throughout Sweden. Based on dbh, two small, two moderate, and two large timber trees were taken from each stand. Dbh varied between 180–470 mm, tree height between 17–34 m, and total age between 51–152 years. Each selected tree was cross-cut into logs; discs were prepared from the butt end of each log and from the top end of the top log. Computed tomography scanning and image analysis were used to determine basic density and growth ring development on sampled discs. Basic density development in 20-mm segments from pith outwards was modelled in models based on ring width, tree and growth condition data. The resulting models had an adjusted R2 of 0.37–0.51 and a RMSE of 37–41 kg/m3.
  • Lindström, University of Canterbury, School of Forestry, Private Bag 4800, Christchurch, New Zealand E-mail: lindstromhakan@netscape.net (email)
article id 621, category Research article
Jyrki Kangas, Pekka Leskinen, Timo Pukkala. (2000). Integrating timber price scenario modeling with tactical management planning of private forestry at forest holding level. Silva Fennica vol. 34 no. 4 article id 621. https://doi.org/10.14214/sf.621
Keywords: decision support; optimization; adaptive behaviour; non-industrial private forestry; tactical planning; timber price modelling
Abstract | View details | Full text in PDF | Author Info
In forest management planning, deterministic timber prices are typically assumed. However, real-life timber prices vary in the course of time, and also price peaks, i.e. exceptionally high timber prices, might occur. If land-owners can utilise the price variation by selling timber with the high prices, they are able to increase their net revenues correspondingly. In this study, an approach is presented to study the timber price variation and its significance in the optimization of forest management. The approach utilizes stochastic timber price scenario modelling, simulation of forest development, and optimization of forest management. The approach is presented and illustrated by means of a case study. It is shown how the degree of uncertainty due to variation in timber prices can be analyzed in tactical forest planning of private forestry, and how the potential benefits of adaptive timber-selling behaviour for a forest landowner can be computed by using the approach. The effects of stochastic timber prices on the choice of forest plan are studied at the forest holding level considering also the spacing and type of cuttings and the optimal cutting order. A forest plan prepared under the assumption of constant timber price very seldom results in optimal forest management. Through studying the effects of stochastic timber prices, forest landowners and other decision makers obtain valuable information about the significance of adaptive timber selling behaviour. The presented methodology can also be used in analysing the land-owners’ economic risks as a function of time-price structure.
  • Kangas, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: jyrki.kangas@metla.fi (email)
  • Leskinen, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: pl@nn.fi
  • Pukkala, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: tp@nn.fi
article id 620, category Research article
Annika Kangas, Matti Maltamo. (2000). Performance of percentile based diameter distribution prediction and Weibull method in independent data sets. Silva Fennica vol. 34 no. 4 article id 620. https://doi.org/10.14214/sf.620
Keywords: stand structure; calibration estimation; Weibull function; diameter distribution prediction; distribution-free method; nearest neighbour method
Abstract | View details | Full text in PDF | Author Info
Diameter distribution is used in most forest management planning packages for predicting stand volume, timber volume and stand growth. The prediction of diameter distribution can be based on parametric distribution functions, distribution-free parametric prediction methods or purely non-parametric methods. In the first case, the distribution is obtained by predicting the parameters of some probability density function. In a distribution-free percentile method, the diameters at certain percentiles of the distribution are predicted with models. In non-parametric methods, the predicted distribution is a linear combination of similar measured stands. In this study, the percentile based diameter distribution is compared to the results obtained with the Weibull method in four independent data sets. In the case of Scots pine, the other methods are also compared to k-nearest neighbour method. The comparison was made with respect to the accuracy of predicted stand volume, saw timber volume and number of stems. The predicted percentile and Weibull distributions were calibrated using number of stems measured from the stand. The information of minimum and maximum diameters were also used, for re-scaling the percentile based distribution or for parameter recovery of Weibull parameters. The accuracy of the predicted stand characteristics were also compared for calibrated distributions. The most reliable results were obtained using the percentile method with the model set including number of stems as a predictor. Calibration improved the results in most cases. However, using the minimum and maximum diameters for parameter recovery proved to be inefficient.
  • Kangas, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: annika.kangas@metla.fi (email)
  • Maltamo, Finnish Forest Research Institute, Joensuu Research Station, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: mm@nn.fi
article id 619, category Research article
Annika Kangas, Matti Maltamo. (2000). Percentile based basal area diameter distribution models for Scots pine, Norway spruce and birch species. Silva Fennica vol. 34 no. 4 article id 619. https://doi.org/10.14214/sf.619
Keywords: stand structure; diameter distribution prediction; distribution-free method; rational spline
Abstract | View details | Full text in PDF | Author Info
Information about diameter distribution is used for predicting stand total volume, timber volume and stand growth for forest management planning. Often, the diameter distribution is obtained by predicting the parameters of some probability density function, using means and sums of tree characters as predictors. However, the results have not always been satisfactory: the predicted distributions practically always have a similar shape. Also, multimodal distributions cannot be obtained. However, diameter distribution can also be predicted using distribution-free methods. In the percentile method, the diameters at certain percentiles of the distribution are predicted with models. The empirical diameter distribution function is then obtained by interpolating between the predicted diameters. In this paper, models for diameters at 12 percentiles of stand basal area are presented for Scots pine, Norway spruce and birch species. Two sets of models are estimated: a set with and one without number of stems as a predictor. Including the number of stems as a predictor improved the volume and saw timber volume estimates for all species, but the improvements were especially high for number of stems estimates obtained from the predicted distribution. The use of number of stems as predictor in models is based on the possibility of including this characteristic to measured stand variables.
  • Kangas, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: annika.kangas@metla.fi (email)
  • Maltamo, Finnish Forest Research Institute, Joensuu Research Station, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: mm@nn.fi
article id 618, category Research article
Steen Magnussen, Paul Boudewyn, Mike Wulder, David Seemann. (2000). Predictions of forest inventory cover type proportions using Landsat TM. Silva Fennica vol. 34 no. 4 article id 618. https://doi.org/10.14214/sf.618
Keywords: neural net; maximum likelihood classification; agreement of predictions
Abstract | View details | Full text in PDF | Author Info
The feasibility of generating via Landsat TM data current estimates of cover type proportions for areas lacking this information in the national forest inventory was explored by a case study in New Brunswick. A recent forest management inventory covering 4196 km2 in south-eastern New Brunswick (the test area) and a coregistered Landsat TM scene was used to develop predictive models of 12 cover type proportions in an adjacent 4525 km2 region (the validation area). Four prediction models were considered, one using a maximum likelihood classifier (MLC), and three using the proportions of 30 TM clusters as predictors. The MLC was superior for non-vegetated cover types while a neural net or a prorating of cluster proportions was chosen for predicting vegetated cover types. Most predictions generated for national inventory photo-plots of 2 x 2 km were closer to the most recent inventory results than estimates extrapolated from the test area. Agreement between predictions and current inventory results varied considerably among cover types with model-based predictions outperforming, on average, the simple spatial extensions by about 14%. In this region, an 11-year-old forest inventory for the validation area provided estimates that in half the cases were closer to current inventory estimates than predictions using the optimal Landsat TM model. A strong temporal correlation of photo-plot-level cover type proportions made old-values more consistent than predictions using the optimal Landsat TM model in all but three cases. Prorating of cluster proportions holds promise for large-scale multi-sensor predictions of forest inventory cover types.
  • Magnussen, Canadian Forest Service, 506 West Burnside Road, Victoria B.C., Canada V8Z 1M5 E-mail: smagnussen@pfc.forestry.ca (email)
  • Boudewyn, Canadian Forest Service, 506 West Burnside Road, Victoria B.C., Canada V8Z 1M5 E-mail: pb@nn.ca
  • Wulder, Canadian Forest Service, 506 West Burnside Road, Victoria B.C., Canada V8Z 1M5 E-mail: mw@nn.ca
  • Seemann, Canadian Forest Service, 506 West Burnside Road, Victoria B.C., Canada V8Z 1M5 E-mail: ds@nn.ca
article id 617, category Research article
Jouni Siipilehto. (2000). A comparison of two parameter prediction methods for stand structure in Finland. Silva Fennica vol. 34 no. 4 article id 617. https://doi.org/10.14214/sf.617
Keywords: Pinus sylvestris; Picea abies; parameter prediction; dbh and height distribution; Johnson’s SBB distribution; Näslund’s height curve
Abstract | View details | Full text in PDF | Author Info
The objective of this paper was to predict a model for describing stand structure of tree heights (h) and diameters at breast height (dbh). The research material consisted of data collected from 64 stands of Norway spruce (Picea abies Karst.) and 91 stands of Scots pine (Pinus sylvestris L.) located in southern Finland. Both stand types contained birch (Betula pendula Roth and B. pubescent Ehrh.) admixtures. The traditional univariate approach (Model I) of using the dbh distribution (Johnson’s SB) together with a height curve (Näslund’s function) was compared against the bivariate approaches, Johnson’s SBB distribution (Model II) and Model Ie. In Model Ie within-dbh-class h-variation was included by transforming a normally distributed homogenous error of linearized Näslund’s function to concern real heights. Basal-area-weighted distributions were estimated using the maximum likelihood (ML) method. Species-specific prediction models were derived using linear regression analysis. The models were compared with Kolmogorov-Smirnov tests for marginal distributions, accuracy of stand variables and the dbh-h relationship of individual trees. The differences in the stand characteristics between the models were marginal. Model I gave a slightly better fit for spruce, but Model II was better for pine stands. The univariate Model I resulted in clearly too narrow marginal h-distribution for pine. It is recommended applying of a constrained ML method for reasonable dbh-h relationship instead of using a pure ML method when fitting the SBB model.
  • Siipilehto, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: jouni.siipilehto@metla.fi (email)
article id 616, category Research article
Markus Lindholm, Hannu Lehtonen, Taneli Kolström, Jouko Meriläinen, Matti Eronen, Mauri Timonen. (2000). Climatic signals extracted from ring-width chronologies of Scots pines from the northern, middle and southern parts of the boreal forest belt in Finland. Silva Fennica vol. 34 no. 4 article id 616. https://doi.org/10.14214/sf.616
Keywords: boreal forest; Scots pine; tree-rings; ring-width chronologies; growth variability; growth responses; spectral analysis
Abstract | View details | Full text in PDF | Author Info
Climatic signals were extracted from ring-width chronologies of Scots pines (Pinus sylvestris L.) from natural stands of the northern, middle, and southern parts of the boreal forest belt in Finland. The strength of the common growth signals (forcing factors) were quantified as a function of time. This was achieved by mean inter-series correlations, calculated over a moving 30-year window, both within and between the regional chronologies. Strong regional signals and also evidence for common forcings were found, especially between northern and central, central and eastern, as well as central/eastern and southern chronologies. Response function analyses revealed that growing season temperatures govern the growth rates of northern pines, while towards south, pine growth becomes less affected by temperatures, and more affected by e.g. precipitation. During some periods, growing conditions seem to have been favorable in the south, while they have been unfavorable in the north (growth inversions). Going from the north to the south, the variability of radial growth clearly decreases, and the variance of ring-width series becomes smaller. Growth variability in the four regions was compared during the common interval of the chronologies, from 1806 to 1991. The spectral densities of the northern, central, eastern and southern chronologies were also compared as functions of frequency, viz. cycles per year. The variance is much greater and there is more periodic behavior in the north than in the south in high, medium, as well as lower frequencies.
  • Lindholm, Saima Centre for Environmental Sciences, University of Joensuu, Linnankatu 11, FIN-57130 Savonlinna, Finland E-mail: ml@nn.fi (email)
  • Lehtonen, Finnish Forest Research Institute, Joensuu Research Station, Box 68, FIN-80101 Joensuu, Finland E-mail: hl@nn.fi
  • Kolström, Finnish Forest Research Institute, Joensuu Research Station, Box 68, FIN-80101 Joensuu, Finland E-mail: tk@nn.fi
  • Meriläinen, Saima Centre for Environmental Sciences, University of Joensuu, Linnankatu 11, FIN-57130 Savonlinna, Finland E-mail: jm@nn.fi
  • Eronen, Department of Geology, Division of Geology and Palaeontology, Box 11, FIN-00014 University of Helsinki, Finland E-mail: me@nn.fi
  • Timonen, Finnish Forest Research Institute, Rovaniemi Research Station, Box 16, FIN-96301 Rovaniemi, Finland E-mail: mt@nn.fi
article id 630, category Research article
S. Samarasinghe, G. D. Kulasiri. (2000). Displacement fields of wood in tension based on image processing: Part 2. Crack-tip displacements in mode-I and mixed-mode fracture. Silva Fennica vol. 34 no. 3 article id 630. https://doi.org/10.14214/sf.630
Keywords: wood; digital image correlation; fracture modes; orthotropic fracture; tip displacement
Abstract | View details | Full text in PDF | Author Info
Near tip displacement fields for tensile loaded cracked rubber and wood with a crack parallel-, perpendicular-to-grain, and a parallel-to-grain crack inclined 30°, 45°, and 60° to the load axis were obtained from digital image correlation (DIC). Theoretical displacements were also obtained for rubber and wood using isotropic and orthotropic fracture theory, respectively. The results showed that DIC can reveal fine details of the nature of displacements and the influences of crack tip in both rubber and wood. Experimental crack tip displacements for wood compare well with theory; particularly, when load is perpendicular-to-grain. Some anomalies were found in the tip displacements in the direction of the tracheids due to the unique nature of their behaviour not accounted for by theory. Mixed-mode crack tip displacement fields for wood clearly showed the increasing influence of crack angle on the displacements, and the displacements perpendicular to crack compared very well with theory. The displacements parallel to crack showed some variations owing to the involvement of tracheids.
  • Samarasinghe, Lincoln University, Appl. Computing, Mathematics and Statistics Group, P.O. Box 84, Canterbury, New Zealand E-mail: ss@nn.nz
  • Kulasiri, Lincoln University, Appl. Computing, Mathematics and Statistics Group, P.O. Box 84, Canterbury, New Zealand E-mail: kulasird@tui.lincoln.ac.nz (email)
article id 629, category Research article
S. Samarasinghe, G. D. Kulasiri. (2000). Displacement fields of wood in tension based on image processing: Part 1. Tension parallel- and perpendicular- to grain and comparisons with isotropic behaviour. Silva Fennica vol. 34 no. 3 article id 629. https://doi.org/10.14214/sf.629
Keywords: wood; deformation profiles; image processing; micro-structure
Abstract | View details | Full text in PDF | Author Info
Displacement fields for tensile loaded rubber and wood in parallel- and perpendicular-to-grain were obtained from digital image correlation. The results showed that the digital image correlation can reveal fine details of the nature of displacements in both rubber and wood. It was found that when load is perpendicular-to-grain, the lignin matrix produces uniform displacement fields similar to that of isotropic rubber. Uniform displacement fields also observed when lignin is involved in contraction due to Poisson effect in parallel-to-grain tension. However, when tracheids carry the load in parallel-to-grain loading, or are compressed in perpendicular-to-grain loading, a complex displacement pattern distorted by internal shear stress and slippage is produced.
  • Samarasinghe, Lincoln University, Appl. Computing, Mathematics and Statistics Group, P.O. Box 84, Canterbury, New Zealand E-mail: ss@nn.nz
  • Kulasiri, Lincoln University, Appl. Computing, Mathematics and Statistics Group, P.O. Box 84, Canterbury, New Zealand E-mail: kulasird@tui.lincoln.ac.nz (email)
article id 628, category Research article
Stefan Daume, Dave Robertson. (2000). A heuristic approach to modelling thinnings. Silva Fennica vol. 34 no. 3 article id 628. https://doi.org/10.14214/sf.628
Keywords: thinning; heuristic model; rule-based; knowledge-based
Abstract | View details | Full text in PDF | Author Info
Thinnings play an important role in guiding forest development and are considered by many to be the most important influence on forests in Central Europe. Due to their importance, thinning models are a major part of any forest growth model for managed forests. Existing thinning model approaches have a number of problems associated with structure and model development that weaken their reliability and accuracy. To overcome some of these problems this paper proposes a heuristic approach to modelling thinnings, where the focus is on distance-dependent, single-tree models. This alternative approach tries to capture the information, strategies and deductive processes likely to be employed by a forester deciding on the removal of individual trees in a stand. Use of heuristics to represent thinning knowledge simplifies the construction and refinement of a thinning model and increases its plausibility. The representation of thinning heuristics in Prolog – a programming language based on formal logic – is a straightforward process without losing expressiveness of the original heuristics. Limited tests of the model implemented in Prolog indicate that the proposed model outperforms its competitors.
  • Daume, The University of Edinburgh, Institute for Representation and Reasoning, Division of Informatics, 80 South Bridge, Edinburgh EH1 1HN, United Kingdom E-mail: stefand@dai.ed.ac.uk (email)
  • Robertson, The University of Edinburgh, Institute for Representation and Reasoning, Division of Informatics, 80 South Bridge, Edinburgh EH1 1HN, United Kingdom E-mail: dr@nn.uk
article id 627, category Research article
Joseph Buongiorno, Audra Kolbe, Mike Vasievich. (2000). Economic and ecological effects of diameter-limit and BDq management regimes: simulation results for northern hardwoods. Silva Fennica vol. 34 no. 3 article id 627. https://doi.org/10.14214/sf.627
Keywords: diversity; hardwoods; simulation; uneven-aged management; economics; mixed-species stands
Abstract | View details | Full text in PDF | Author Info
The long-term financial and ecological effects of diameter-limit regimes and basal-area-diameter-q-ratio (BDq) regimes were compared by simulation in the case of northern hardwood forests. Varying the cutting cycle between 10 and 20 years had little effect on returns or stand structure. A 28-cm diameter-limit cut gave the highest production and financial returns, and the highest species diversity, but considerably lower size diversity. A 38-cm diameter-limit cut and a heavy BDq selection harvest gave high returns, while maintaining high levels of diversity. On lands of equal site quality, Michigan’s stands were more productive than Wisconsin’s. The results suggest that it is possible to manage northern hardwood stands sustainably with diameter-limit cuts, combined with removal of poorly performing understory trees. Adjusting the diameter limit gave rise to stands similar in productivity and structure to those obtained by BDq cutting regimes. Given their simplicity of implementation and monitoring, more attention should be given to diameter-limit cutting regimes, with attendant stand improvement measures, as a practical means for uneven-aged management of northern hardwoods.
  • Buongiorno, Department of Forest Ecology and Management, University of Wisconsin-Madison, 1630 Linden Drive, Madison, WI 53705, USA E-mail: Jbuongio@facstaff.wisc.edu (email)
  • Kolbe, Department of Forest Ecology and Management, University of Wisconsin-Madison, 1630 Linden Drive, Madison, WI 53705, USA E-mail: ak@nn.us
  • Vasievich, USDA Forest Service, North Central Forest Experiment Station, 1407 S. Harrison Road, Suite 220, East Lansing, MI 48823, USA E-mail: mv@nn.us
article id 626, category Research article
Teijo Nikkanen, Seppo Ruotsalainen. (2000). Variation in flowering abundance and its impact on the genetic diversity of the seed crop in a Norway spruce seed orchard. Silva Fennica vol. 34 no. 3 article id 626. https://doi.org/10.14214/sf.626
Keywords: Picea abies; clone; ramet; status number; census number
Abstract | View details | Full text in PDF | Author Info
The variation in flowering abundance was studied in a Norway spruce seed orchard, located in southern Finland (62°13'N, 25°24'E), consisting of 67 clones from northern Finland (64°–67°N). The flowering variation in 1984–1996 was studied at the annual, clonal and graft level. In addition, the genetic diversity of an imaginary seed crop was estimated using a concept of status number. The between-year variation was large in both female and male flowering. Differences in flowering abundance among the clones were large and statistically significant in all the years studied. The average broad-sense heritability values for female and male flowering were 0.37 and 0.38, respectively, but varied considerably from year to year. The correlations between the flowering abundance of the clones in different years were usually positive and significant. However, the correlations for two pairs of successive good flowering years showed that the same clones usually flowered well in the first year in both pairs of years, and the other clones in the second year. The clonal differences in flowering could not be explained by geographic origin, but were more dependent on the graft size. Our results demonstrate that the variation in the ramet number, flowering abundance and pollen contamination must be included when estimating the genetic diversity of the seed crop in a seed orchard. The relative status number of the seed orchard was 84% of the number of clones when the variation in the ramet number was included. The relative status numbers after adjusting for the variation in female and male flowering were on the average 46 and 55%, respectively, and 59% when adjusting for both genders together. Pollen contamination increased the status number considerably.
  • Nikkanen, Finnish Forest Research Institute, Punkaharju Research Station, FIN-58450 Punkaharju, Finland E-mail: teijo.nikkanen@metla.fi (email)
  • Ruotsalainen, Finnish Forest Research Institute, Punkaharju Research Station, FIN-58450 Punkaharju, Finland E-mail: sr@nn.fi
article id 625, category Research article
Hans-Örjan Nohrstedt. (2000). Effects of soil scarification and previous N fertilisation on pools of inorganic N in soil after clear-felling of a Pinus sylvestris (L.) stand. Silva Fennica vol. 34 no. 3 article id 625. https://doi.org/10.14214/sf.625
Keywords: boreal forest; mound; Sweden; furrow; nitrogen mineralisation; podsol
Abstract | View details | Full text in PDF | Author Info
Previous analyses of soil water beneath mounds resulting from scarification have implied that this forestry measure increases leaching of inorganic N. However, more recent soil-water studies have not confirmed this assumption. The soil study presented here examined the pools of inorganic N in different microsites emanating from a simulated disc trenching, i. e. the mound with underlying soil, the furrow bottom and the undisturbed soil. The study was made five years after scarification. The mound itself with underlying soil had a larger pool of inorganic N than the undisturbed soil. This was mainly because of an increase in the embedded humus layer, thus implying a larger net N mineralisation and/or lower losses. However, when pools of inorganic N per hectare were calculated, taking into consideration that a scarified area comprises 25% mounds, 25% furrows and 50% undisturbed soil, there was no increase in pools of inorganic N when compared with an area not subjected to scarification. This observation supports the finding of the more recent soil-water studies mentioned, i. e., that leaching seems not to be influenced by soil scarification. The scarification was made as a split-plot treatment on main-plots in an old experiment with different N doses. Thus, the effect of the previous N fertilisation could also be evaluated. Two N doses were tested beside the unfertilised control: 720N (3 x 240 kg N ha–1 yr–1) and 1800N (3 x 600 kg N ha–1 yr–1). The last fertiliser application was made six years before the clearcutting and 13 years before the soil sampling. The previously fertilised main-plots had larger pools of inorganic N than the control plots.
  • Nohrstedt, The Forestry Research Institute of Sweden, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: hans-orjan.nohrstedt@skogforsk.se (email)
article id 638, category Research article
Gert-Jan Nabuurs, Mart-Jan Schelhaas, Ari Pussinen. (2000). Validation of the European Forest Information Scenario Model (EFISCEN) and a projection of Finnish forests. Silva Fennica vol. 34 no. 2 article id 638. https://doi.org/10.14214/sf.638
Keywords: Finland; EFISCEN; European forests; historic forest inventory data
Abstract | View details | Full text in PDF | Author Info
Large-scale forest scenario models are intensively used to make projections of forest areas of up to hundreds of millions of hectares. Within Europe, such projections have been done for 11 countries at the individual national scale, most often to foresee the long-term implications of the ongoing forest management. However, the validity of the models has rarely been tested. The aim of this study was 1. to validate the European Forest Information SCENario model (EFISCEN) by running it on historic Finnish forest inventory data, 2. to improve the model based on the validation, and 3. to project the Finnish forest development till 2050 with the improved model under alternative scenarios. The results of the validation showed that EFISCEN is capable of making reliable large-scale projections of forest resources for periods up to 50–60 years. Based on the validation, the model was improved concerning simulation of age development, thinning regimes and regrowth after thinning. The projection of the Finnish forests till 2050 with the improved model presented a maximum sustainable felling level of around 70 million m3 per year. That provides an average growing stock of 106 m3 ha–1 in 2050 and a net annual increment of 3.6 m3 ha–1 y–1. If the current trend towards more nature oriented forest management continues and 1.39 million ha of forests have been set aside additionally for nature reserves by 2050, the felling level could meet a realistic demand of 57 million m3 per year in 2050. Under the latter regime the average growing stock will have grown to 160 m3 ha–1 in 2050.
  • Nabuurs, European Forest Institute (EFI), Torikatu 34, FIN-80100 Joensuu, Finland; Wageningen University and Research Center, ALTERRA, P.O. Box 47, NL 6700 AA Wageningen, The Netherlands E-mail: g.j.nabuurs@alterra.wag-ur.nl (email)
  • Schelhaas, European Forest Institute (EFI), Torikatu 34, FIN-80100 Joensuu, Finland; Wageningen University and Research Center, ALTERRA, P.O. Box 47, NL 6700 AA Wageningen, The Netherlands E-mail: mjs@nn.nl
  • Pussinen, European Forest Institute (EFI), Torikatu 34, FIN-80100 Joensuu, Finland; Wageningen University and Research Center, ALTERRA, P.O. Box 47, NL 6700 AA Wageningen, The Netherlands E-mail: ap@nn.nl
article id 637, category Research article
Hirofumi Kuboyama, Hiroyasu Oka. (2000). Climate risks and age-related damage probabilities – effects on the economically optimal rotation length for forest stand management in Japan. Silva Fennica vol. 34 no. 2 article id 637. https://doi.org/10.14214/sf.637
Keywords: simulation; climatic risks; damage probability; age class; optimal rotation period; land expectation value
Abstract | View details | Full text in PDF | Author Info
We estimated the damage probability according to age class and major climatic disasters based on ‘Statistical Yearbook of National Forest Insurance’ from 1960 to 1996. The probability of snow damage is high for young stands, then gradually decreases with age. On the other hand, the risk of wind damage gradually increases with age. Decisions about rotation age should be based on the distribution of damage probability with stand age. Risk of damage has two contradictory effects on the optimal rotation period; one is that the rotation-shortening effect caused by risk of damage around harvest age; another is the rotation-extending effect due to decrease of rent by the risk of damage through the raising period. Change of optimal rotation depends on the relative magnitude of these effects. We examine this by calculating land expectation value (LEV) using a simulation model with the empirical damage probability, price and cost. Change of the optimal rotation period obtained from the national average damage probability is not significant. However, the optimal rotation is shorter in high wind risk areas and is longer in high snow risk areas. It is because the damage probability for a mature stand is high in the case of wind and low in the case of snow. In addition, the extent of decrease in LEV is smaller for wind than for snow. The results of simulation based on empirical data confirm that the optimal rotation period can become either shorter or longer through incorporating risk in decision making, depending on the damage probability distribution with stand age.
  • Kuboyama, Forestry and Forest Products Research Institute Tohoku Center, Nabeyashiki 72, Shimo-Kuriyagawa, Morioka, Iwate, 020-0123 Japan E-mail: kuboyama@ffpri-thk.affrc.go.jp (email)
  • Oka, Forestry and Forest Products Research Institute Tohoku Center, Nabeyashiki 72, Shimo-Kuriyagawa, Morioka, Iwate, 020-0123 Japan E-mail: ho@nn.jp
article id 636, category Research article
Tuula Nuutinen, Hannu Hirvelä, Jari Hynynen, Kari Härkönen, Hannu Hökkä, Kari T. Korhonen, Olli Salminen. (2000). The role of peatlands in Finnish wood production – an analysis based on large-scale forest scenario modelling. Silva Fennica vol. 34 no. 2 article id 636. https://doi.org/10.14214/sf.636
Keywords: peatlands; MELA; wood production; forest scenario modelling
Abstract | View details | Full text in PDF | Author Info
Using the Finnish MELA model, a set of scenarios were produced and used to map the possibilities and risks surrounding the utilisation of peatlands in wood production in Finland. One of the scenarios was an estimate of allowable-cut calculated by maximising the net present value of the future revenues using a four per cent interest rate subject to non-decreasing flow of wood, saw logs and net income over a 50-year period, and net present value after the 50 year period greater or equal than in the beginning. The estimate for maximum regionally sustained removal in 1996–2005 was 68 million m3 per year – approaching 74 million m3 during the next decades. In this scenario, 14 per cent of all cuttings during the period 1996–2005 would be made on peatlands, which comprise ca. 31 per cent of the total area of forestry land. By the year 2025, the proportion of peatland cuttings would increase to over 20 per cent. The increase in future cutting possibilities on peatlands compensated for a temporary decrease in cuttings and growing stock on mineral soils. The allowable-cut effect was especially pronounced in northern Finland, where peatlands play an important role in wood production. In addition, the sensitivity of cutting possibilities for assumptions related to growth and price were analysed. The estimate of maximum sustainable yield as defined here seems to be fairly robust on the whole, except in northern Finland where the cutting scenarios were sensitive to the changes in the price of birch pulpwood. The proportion of peatland stands that are profitable for timber production depends on the interest rate: the higher the rate of interest the less peatland stands are thinned. The effect of cutting profile on future logging conditions and resulting costs were analysed in two forestry centres. If clear cuttings on mineral soils are to be cut first, an increase in future logging costs is inevitable.
  • Nuutinen, Finnish Forest Research Institute, Joensuu Research Station, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: tuula.nuutinen@metla.fi (email)
  • Hirvelä, Finnish Forest Research Institute, Joensuu Research Station, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: hh@nn.fi
  • Hynynen, Finnish Forest Research Institute, Joensuu Research Station, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: jh@nn.fi
  • Härkönen, Finnish Forest Research Institute, Joensuu Research Station, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: kh@nn.fi
  • Hökkä, Finnish Forest Research Institute, Joensuu Research Station, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: hh@nn.fi
  • Korhonen, Finnish Forest Research Institute, Joensuu Research Station, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: ktk@nn.fi
  • Salminen, Finnish Forest Research Institute, Joensuu Research Station, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: os@nn.fi
article id 635, category Research article
Manfred J. Lexer, Karl Hönninger, Helfried Scheifinger, Christoph Matulla, Nikolaus Groll, Helga Kromp-Kolb. (2000). The sensitivity of central European mountain forests to scenarios of climatic change: methodological frame for a large-scale risk assessment. Silva Fennica vol. 34 no. 2 article id 635. https://doi.org/10.14214/sf.635
Keywords: climate change; potential natural vegetation; alpine forests; risk assessment; patch model; multi-attribute decision making
Abstract | View details | Full text in PDF | Author Info
The methodological framework of a large-scale risk assessment for Austrian forests under scenarios of climatic change is presented. A recently developed 3D-patch model is initialized with ground-true soil and vegetation data from sample plots of the Austrian Forest Inventory (AFI). Temperature and precipitation data of the current climate are interpolated from a network of more than 600 weather stations to the sample plots of the AFI. Vegetation development is simulated under current climate (‘control run’) and under climate change scenarios starting from today's forest composition and structure. Similarity of species composition and accumulated biomass between these two runs at various points in time were used as assessment criteria. An additive preference function which is based on Saaty’s AHP is employed to synthesize these criteria to an overall index of the adaptation potential of current forests to a changing climate. The presented methodology is demonstrated for a small sample from the Austrian Forest Inventory. The forest model successfully simulated equilibrium species composition under current climatic conditions spatially explicit in a heterogenous landscape based on ground-true data. At none of the simulated sites an abrupt forest dieback did occur due to climate change impacts. However, substantial changes occured with regard to species composition of the potential natural vegetation (PNV).
  • Lexer, Institute of Silviculture, University of Agricultural Sciences, Peter-Jordanstrasse 70, A-1190 Vienna, Austria E-mail: lexer@edv1.boku.ac.at (email)
  • Hönninger, Institute of Silviculture, University of Agricultural Sciences, Peter-Jordanstrasse 70, A-1190 Vienna, Austria E-mail: kh@nn.at
  • Scheifinger, Institute of Meteorology and Physics, University of Agricultural Sciences, Türkenschanzstrasse 18, A-1180 Vienna, Austria E-mail: hs@nn.at
  • Matulla, Institute of Meteorology and Physics, University of Agricultural Sciences, Türkenschanzstrasse 18, A-1180 Vienna, Austria E-mail: cm@nn.at
  • Groll, Institute of Meteorology and Physics, University of Agricultural Sciences, Türkenschanzstrasse 18, A-1180 Vienna, Austria E-mail: ng@nn.at
  • Kromp-Kolb, Institute of Meteorology and Physics, University of Agricultural Sciences, Türkenschanzstrasse 18, A-1180 Vienna, Austria E-mail: hkk@nn.at
article id 634, category Research article
Marcus Lindner, Petra Lasch, Markus Erhard. (2000). Alternative forest management strategies under climatic change – prospects for gap model applications in risk analyses. Silva Fennica vol. 34 no. 2 article id 634. https://doi.org/10.14214/sf.634
Keywords: climate change; forest management; adaptation and mitigation strategies; forest succession model; FORSKA
Abstract | View details | Full text in PDF | Author Info
The projected global climate change will influence growth and productivity of natural and managed forests. Since the characteristics of the future regional climate are still uncertain and the response of our forests to changes in the atmospheric and climatic conditions may be both positive or negative, decision making in managed forests should consider the new risks and uncertainties arising from climatic change, especially if the rotation periods are long. An extended version of the forest gap model FORSKA was applied to simulate the forest development at 488 forest inventory plots in the federal state of Brandenburg, Germany, under two climate and three management scenarios. The transient growth dynamics from 1990 to 2100 were investigated at four sites in different parts of the state, representing the variability of environmental and forest conditions within Brandenburg. The alternative management strategies led to distinct differences in forest composition after 110 years of simulation. The projected climate change affected both forest productivity and species composition. The impacts of alternative management scenarios are discussed. It is concluded that the extended forest gap model can be a valuable tool to support decision making in forest management under global change.
  • Lindner, Potsdam Institute for Climate Impact Research, P.O. Box 60 12 03, D-14412 Potsdam, Germany E-mail: lindner@pik-potsdam.de (email)
  • Lasch, Potsdam Institute for Climate Impact Research, P.O. Box 60 12 03, D-14412 Potsdam, Germany E-mail: pl@nn.de
  • Erhard, Potsdam Institute for Climate Impact Research, P.O. Box 60 12 03, D-14412 Potsdam, Germany E-mail: me@nn.de
article id 633, category Research article
Tron Eid. (2000). Use of uncertain inventory data in forestry scenario models and consequential incorrect harvest decisions. Silva Fennica vol. 34 no. 2 article id 633. https://doi.org/10.14214/sf.633
Keywords: forest management; uncertain inventory data; final harvest decisions; expected net present value-losses; inventory planning
Abstract | View details | Full text in PDF | Author Info
Uncertainty in long-term timber production analyses usually focus success of regeneration, growth/mortality of trees and future fluctuations of timber prices/harvest costs, while uncertainty related to inventory data is paid less attention. At the same time, evaluations of inventory methods usually stop when the error level is stated, while the uncertainty accompanied by using the data is seldom considered. The present work addresses uncertain inventory data in long-term timber production analyses. Final harvest decisions, i.e. possible outcome intervals with respect to timing and expected net present value-losses due to incorrect timing, were considered. A case study was presented where inventory data errors according to different error levels were generated randomly. The selected error levels were based on observations from practical forest inventories in Norway. The analysis tool was GAYA-JLP. The impact of errors on decisions was derived through repeated computations of management strategies maximising net present value without harvest path constraints. A real rate of discount of 3% and an error level of 15% resulted in expected net present value-losses of 1 NOK ha–1 for basal area, 63 NOK ha–1 for mean height, 210 NOK ha–1 for site quality, 240 NOK ha–1 for stand age, and 499 NOK ha–1 when random errors occurred simultaneously for all these variables. The expected net present value-losses varied considerably. The largest losses appeared for stands with ages around optimal economical rotation ages. The losses were also relatively large for young stands, while they were relatively low for overmature stands. The experiences from the case study along with considerations related to other sources of uncertainty may help us to get a more realistic attitude to the reliability of long-term timber production analyses. The results of the study may also serve as a starting point in a decision oriented inventory planning concept, in which alternatives for inventory design and intensity are based on considerations with respect to inventory costs as well as net present value-losses.
  • Eid, Department of Forest Sciences, Agricultural University of Norway, P.O. Box 5044, N-1432 Ås, Norway E-mail: tron.eid@isf.nlh.no (email)
article id 645, category Research article
Veli-Pekka Heikkinen, Antti Kanto. (2000). Market model with long-term effects – empirical evidence from Finnish forestry returns. Silva Fennica vol. 34 no. 1 article id 645. https://doi.org/10.14214/sf.645
Keywords: forest risk returns; market model; co-integration
Abstract | View details | Full text in PDF | Author Info
The aim of the study is to reformulate the conventional market model by considering also long-run characteristics of forestry returns. Finnish stumpage prices and the stock market index are found co-integrated. Co-integration relationship between timber and stock market indicates that there are factors in timber market like high transaction costs, illiquidity or temporal lack of information, and in the Finnish case, price recommendations that are priced by market. The presence of long-run effects may make the short-term market model mis-specified and it may give misleading and incomplete results concerning the expected risk and return of forestry. In our case, the market model risk beta was only slightly biased. This study indicates that an error-correction model is more appropriate model than the market model.
  • Heikkinen, Helsinki School of Economics, P.O. Box 1210, FIN-00101 Helsinki, Finland E-mail: vheikkin@hkkk.fi (email)
  • Kanto, Helsinki School of Economics, P.O. Box 1210, FIN-00101 Helsinki, Finland E-mail: ak@nn.fi
article id 644, category Research article
Jouni Pykäläinen. (2000). Defining forest owner’s forest-management goals by means of a thematic interview in interactive forest planning. Silva Fennica vol. 34 no. 1 article id 644. https://doi.org/10.14214/sf.644
Keywords: interactive planning; interactive optimization; utility function
Abstract | View details | Full text in PDF | Author Info
Numerical optimization may not be the best way to launch an interactive planning process. The forest owner may not be able to identify precise management goals, as required in numerical optimization, for his forest. The owner may also think that it is impossible to express forest-management goals numerically. Due to these reasons, thematic interview was tested as an introductory method in interactive planning in several actual planning cases. It was observed that these interviews helped the owners in outlining their forest-management goals, and they offered an appropriate framework for defining these goals. It was also noticed that the goals defined in the course of the interviews could be included in the planning model in a way understood and accepted by the owners, and the goals defined could be fulfilled.
  • Pykäläinen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: jouni.pykalainen@joensuu.fi (email)
article id 643, category Research article
Pekka Eskelinen, Harri Eskelinen. (2000). A K-band microwave measuring system for the analysis of tree stems. Silva Fennica vol. 34 no. 1 article id 643. https://doi.org/10.14214/sf.643
Keywords: microwave measurements; wood analysis; knot detection; stem quality evaluation
Abstract | View details | Full text in PDF | Author Info
The internal structure of growing trees and freshly cut logs can be characterized in real time by analysing the transmission and reflection of Ku- or K-band microwave energy injected with a horizontal polarization towards the material. Information about the moisture content, material bends, number and location of knots and sections of spoiled wood e.g. due to insects can be gathered in real time. Most sensitive test parameters are attenuation, group delay and the rotation of a linearly polarized wavefront. A simultaneous recording of reflection reduces errors caused by non-significant surface deformations. The spatial resolution, humidity equalization and noise immunity can be improved by applying a wideband frequency modulation. Commercial building blocks supplemented with a special antenna arrangement give possibilities also for the rough harvester environment.
  • Eskelinen, Lappeenranta University of Technology, Department of Electrical Engineering, P.O. Box 20, FIN-53851 Lappeenranta, Finland E-mail: ari.eskelinen@pp.inet.fi (email)
  • Eskelinen, Lappeenranta University of Technology, Department of Electrical Engineering, P.O. Box 20, FIN-53851 Lappeenranta, Finland E-mail: he@nn.fi
article id 642, category Research article
Jonas Rönnberg. (2000). Logging operation damage to roots of clear-felled Picea abies and subsequent spore infection by Heterobasidion annosum. Silva Fennica vol. 34 no. 1 article id 642. https://doi.org/10.14214/sf.642
Keywords: Norway spruce; logging damage; Butt rot; root rot; clear felling; stump root infection
Abstract | View details | Full text in PDF | Author Info
Two studies were carried out to examine the effects of clear-felling operations on stump roots of Norway spruce (Picea abies (L.) Karst.). In study I, the number of cases and the degree of damage to stump roots of Norway spruce were investigated on three clear-felled sites in northern and southern Sweden respectively. The cutting was done in winter or spring. A mean of 37% of the stumps had signs of root damage caused by clear-felling operations. Study II was carried out on two sites in southern and two sites in northern Sweden. The trees were clear-felled in June or July. The frequency of natural infection by Heterobasidion annosum (Fr.) Bref. through damaged roots was compared to infection through stump surfaces. The total area of damage on roots was 88% of the stump surface area. On average, 54% of the stumps were infected through the stump surface and 19% through locations of root damage. The root infections, however, were generally small in size as compared to stump surface infections. The study shows that damage to roots at clear-felling may be extensive, but this probably is not of great importance for the efficacy of stump treatment against H. annosum.
  • Rönnberg, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: jonas.ronnberg@ess.slu.se (email)
article id 641, category Research article
Juha Kaitera. (2000). Analysis of Cronartium flaccidum lesion development on pole-stage Scots pines. Silva Fennica vol. 34 no. 1 article id 641. https://doi.org/10.14214/sf.641
Keywords: Pinus sylvestris; aecia; Cronartium flaccidum; lesion; resin top
Abstract | View details | Full text in PDF | Author Info
Historical and current lesion development and sporulation of Cronartium flaccidum was investigated in a stand of artificially seeded pole-stage Pinus sylvestris in northern Finland. An average of 6.5 lesions developed per infected tree, most of them occurring on a minority (25%) of the trees. During the monitoring period of five years, fresh aecia appeared mainly in 7–10-year-old shoots, the age of the shoots bearing aecia varying between 3–20 years. Aecia appeared for the first time most frequently in 5–10-year-old shoots. Infection waves occurred, whereas lesions were formed most frequently in shoots formed in various years through the 1980s. After the lesions started to sporulate, sporulation in most lesions that finished sporulating during the monitoring period lasted for 1–2 years. The aecia in between 47% and 59% of the infected shoots developed annually over a longer length in proximal direction than in distal direction next to the previous year’s infection. The aecia-bearing distal part of the shoot was longer in between 19% and 37% of the shoots.
  • Kaitera, Finnish Forest Research Institute, Rovaniemi Research Station, P.O. Box 16, FIN-96301 Rovaniemi, Finland E-mail: juha.kaitera@metla.fi (email)
article id 640, category Research article
Jarkko Koskela. (2000). A process-based growth model for the grass stage pine seedlings. Silva Fennica vol. 34 no. 1 article id 640. https://doi.org/10.14214/sf.640
Keywords: allocation; Monte Carlo simulation; grass stage; Pinus merkusii; dynamic model
Abstract | View details | Full text in PDF | Author Info
A carbon- and nitrogen-balance model, applying pipe model theory and a modification of functional balance as growth-guiding rules, is presented for the grass stage pine seedlings. Three populations of Pinus merkusii Jungh. et de Vriese, originating from northern and northeastern Thailand, were grown under controlled environment for 47 weeks to obtain parameter information, to evaluate the model performance and to investigate genotypic variation in various characteristics among the populations. Monte Carlo simulations were used to evaluate the sensitivity of the model behaviour to varying parameter values and to calibrate the model for each population. With given sets of parameter values, the simulated biomass development fitted rather well the observed one during the experiment. The two most important parameters determining model performance were within-shoot shading and specific nitrogen uptake rate of fine roots. The fit of simulated versus measured fine roots had a major effect on acceptable model performance in Monte Carlo simulations. Significant variation in biomass growth, nitrogen use efficiency, height, stem diameter, total carbon concentrations of stem and fine roots, and total nitrogen concentrations of needles, transport roots and fine roots was found among the populations. The observed genotypic variation in seedling biomass and stem diameter was consistent with the geographical distribution of the populations while the variation in the rest of the measured characteristics was not. It seems that P. merkusii populations in Thailand are adapted to more site specific conditions rather than climatic conditions alone, and that the variation in biomass growth may result from variation in internal carbon and nitrogen dynamics among the populations.
  • Koskela, Department of Forest Ecology/Tropical Silviculture Unit, P.O. Box 28, FIN-00014 University of Helsinki, Finland E-mail: jarkko.koskela@helsinki.fi (email)
article id 652, category Research article
M. J. Youngman, G. D. Kulasiri, I. M. Woodhead, G. D. Buchan. (1999). Use of combined constant rate and diffusion model to simulate kiln-drying of Pinus radiata timber. Silva Fennica vol. 33 no. 4 article id 652. https://doi.org/10.14214/sf.652
Keywords: simulation; timber drying; diffusion; constant drying-rate
Abstract | View details | Full text in PDF | Author Info
This paper presents the use of a combined constant drying-rate and diffusion model to simulate the drying of Pinus radiata timber under kiln-drying conditions. The constant drying-rate and diffusion coefficients of the model, which control the drying rate of individual pieces of timber, were determined from calibrating the model against the experimental drying curves obtained under the kiln-drying conditions. The experimental drying curves were obtained from the gravimetric measurements of the moisture content of timber during kiln drying. Statistical relationships were developed for the constant drying-rate and the diffusion coefficients of the model as functions of kiln temperature and the dry basis density of timber. To determine the effects of variability of timber, a simulation scheme was developed based on the model, the probability distribution of the density of timber, the equations for the constant drying-rate coefficient and the diffusion coefficient. The model and the associated simulation method provides a simple way to estimate the drying time of a stack of timber using parameters determined from experimental results for the specific timber kiln.
  • Youngman, Lincoln University, Appl. Management and Computing Division, P.O. Box 84, Canterbury, New Zealand E-mail: mjy@nn.nz
  • Kulasiri, Lincoln University, Appl. Management and Computing Division, P.O. Box 84, Canterbury, New Zealand E-mail: kulasird@tui.lincoln.ac.nz (email)
  • Woodhead, Lincoln University, Appl. Management and Computing Division, P.O. Box 84, Canterbury, New Zealand E-mail: imw@nn.nz
  • Buchan, Lincoln University, Appl. Management and Computing Division, P.O. Box 84, Canterbury, New Zealand E-mail: gdb@nn.nz
article id 651, category Research article
Annika S. Kangas, Jyrki Kangas. (1999). Optimization bias in forest management planning solutions due to errors in forest variables. Silva Fennica vol. 33 no. 4 article id 651. https://doi.org/10.14214/sf.651
Keywords: forest planning; uncertainty; prediction; decision analysis
Abstract | View details | Full text in PDF | Author Info
The yield of various forest variables is predicted by means of a simulation system to provide information for forest management planning. These predictions contain many kinds of uncertainty, for example, prediction and measurement errors. Inevitably, this has an effect on forest management planning. It is well known that uncertainty in the forest yields causes optimistic bias in the observed values of the objective function. This bias increases with the error variances. The amount of bias, however, also depends on the error structure and the relations between the objective variables. In this paper, the effect of uncertainty in forest yields on optimization is studied by simulation. The effect of two different sources of error, the correlation structure of these errors and relations among the objective variables are considered, as well as the effect of two different optimization approaches. The relations between the objective variables and the error structure had a notable effect on the optimization results.
  • Kangas, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: annika.kangas@metla.fi (email)
  • Kangas, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: jk@nn.fi
article id 650, category Research article
Jouni Siipilehto. (1999). Improving the accuracy of predicted basal-area diameter distribution in advanced stands by determining stem number. Silva Fennica vol. 33 no. 4 article id 650. https://doi.org/10.14214/sf.650
Keywords: dbh distribution; parameter prediction; Johnson’s SB distribution; Weibull distribution
Abstract | View details | Full text in PDF | Author Info
The objective of this paper was to study to what extent the accuracy of predicted basal-area diameter distributions (DDG) could be improved by means of stem number observations in advanced (H > 10 m) stands. In the Finnish forest management planning (FMP) inventory practice, stem number is determined only in young stands; in older stands stand basal area is used. The study material consisted of sixty stands of Norway spruce (Picea abies Karst.) and ninety-one stands of Scots pine (Pinus sylvestris L.) with birch (Betula pendula Roth and B. pubescens Ehrh.) admixtures in southern and eastern Finland. For test data, 167–292 independent, National Forest Inventory-based, permanent sample plots were used. DDGs were estimated with the maximum likelihood method. Species-specific models for predicting the distribution parameters were derived using regression analysis. The two-parameter Weibull distribution was compared to the three-parameter Johnson’s SB distributions in predicting DDGs. The models were based on either predictors that are consistent with current FMP (model G), or assuming an additional stem number observation (model G+N). The predicted distributions were compared in terms of the derived stand variables: stem number, total and timber volumes. The results were similar in modelling and test data sets. Methods, based on the SB distribution obtained with model (G+N), proved to give the most accurate description of the stand structure. Differences were marginal in stand total volumes. However, the error variation in stem number was 20% to 80% lower than when applying model (G). SB and Weibull distributions gave very much the same results if model (G) was applied.
  • Siipilehto, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: jouni.siipilehto@metla.fi (email)
article id 649, category Research article
Tord Johansson. (1999). Biomass production of Norway spruce (Picea abies (L.) Karst.) growing on abandoned farmland. Silva Fennica vol. 33 no. 4 article id 649. https://doi.org/10.14214/sf.649
Keywords: Norway spruce; biomass production; Picea abies; thinning; LAI; abandoned farmland; basic density
Abstract | View details | Full text in PDF | Author Info
Biomass production of forests has been studied for at least a century. Tree biomass is used in Sweden both as industrial raw material and an energy source. Few studies dealing with biomass yield from Norway spruce (Picea abies (L.) Karst.) growing on farmland are published. Practical recommendations are sparsely. The aim of this study was to construct dry weight equations for Norway spruce growing on farmland. Dry weight equations for fractions of Norway spruce trees were made. Biomass production was estimated in 32 stands of Norway spruce growing on abandoned farmland. The stands were located in Sweden at latitudes ranging from 58° to 64° N, and their total age varied from 17 to 54 years. A modified ‘mean tree technique’ was used to estimate biomass production; i.e. the tallest tree was chosen for sampling. The actual mean total dry weight above stump level for the 32 stands was 116 ton ha–1, with a range of 6.0 to 237.4 ton d.w. ha–1. When previous thinning removals were included, the mean biomass value was 127 ton ha–1 (6.0–262.8). In addition to estimating conventional dry weights of trees and tree components, basic density, specific leaf area, total surface area and leaf area index, among other measures, were estimated. Norway spruce biomass yields on plots subjected to different thinning were compared. The total harvested biomass was 75–120 ton d.w. ha–1 in heavy thinnings from below. Stands were thinned four to five times, with the first thinning at 23–27 years and the last at 51–64 years. The harvested biomass obtained in the first thinning was 18–38 ton d.w. ha–1. Total biomass production was 178–305 ton d.w. ha–1. Stands thinned from above supplied 71–130 ton d.w. ha–1 in total and 17–42 ton d.w. ha–1 in the first thinning. The total biomass supply was 221–304 ton d.w. ha–1. Unthinned stands produced a total of 155–245 ton d.w. ha–1.
  • Johansson, Swedish University of Agricultural Sciences, Department of Forest Management and Products, P.O. Box 7060, S-750 07 Uppsala, Sweden E-mail: tord.johansson@sh.slu.se (email)
article id 648, category Research article
Sauli Härkönen. (1999). Forest damage caused by the Canadian beaver (Castor canadensis) in South Savo, Finland. Silva Fennica vol. 33 no. 4 article id 648. https://doi.org/10.14214/sf.648
Keywords: Canadian beaver; Castor canadensis; forest damage; prevention of damage; type of damage
Abstract | View details | Full text in PDF | Author Info
The increasing Canadian beaver (Castor canadensis) population has caused forest damage in Finland. However, the occurrence, extent and importance of the damage have not been comprehensively studied. The field inspection was carried out in all of the beaver damage areas (n = 50) in the Anttola, Juva and Pieksämäki game management units in summer 1998. The characteristics of the damage areas, types of damage and the severity of the damage were examined. The size of the damage areas averaged 2.2 ha. The damage areas occurred on peatland forest more (p < 0.05) than expected. The dominant tree species were commercially valuable trees such as Scots pine (Pinus sylvestris) (33%), Norway spruce (Picea abies) (18%) and birches (Betula spp.) (47%). Birch occurred as dominant tree species more (p < 0.05) than expected. The proportion of older forest development classes was considerable. The most important type of damage was flooding (50%) caused by the damming activity of beavers. The trees were dead or dying in 18% of the damage areas. It was estimated that the ecosystem engineering performed by beavers was of no importance in 28% of the damage areas. Prevention of beaver damage has been carried out in 80% of the damage areas. In the study area, beaver numbers have steadily increased, although the number of beavers taken by hunting and the allowed hunting harvest have sharply increased. It is suggested that the methods used to prevent beaver damage should be improved and a compensation system should be provided by the state for the most damaged areas. Beavers can cause damage to commercial forestry, and this should be taken into account more effectively in the management plans made by the game management districts.
  • Härkönen, South Savo Game Management District, P.O. Box 14, FIN-51901 Juva, Finland E-mail: sauli.harkonen@riistanhoitopiirico.inet.fi (email)
article id 647, category Research article
Tapio Linkosalo. (1999). Regularities and patterns in the spring phenology of some boreal trees. Silva Fennica vol. 33 no. 4 article id 647. https://doi.org/10.14214/sf.647
Keywords: Populus tremula; Pinus sylvestris; flowering; Alnus glutinosa; bud burst; Alnus incana; phenological regulation; phenological time series; Betula sp.
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Phenological time series of flowering and bud burst of Populus tremula (L.) and Betula sp., and the flowering of Pinus sylvestris (L.), Alnus glutinosa (L.) and Alnus incana (L.) were constructed from data collected in Finland during the period 1896–1955. The resulting combined time series were examined with two aims in mind: first, to determine the phenological regularities between different species and, second, to detect patterns of spring advancement over a geographically large area. The results indicate that the geographical pattern of spring advancement is rather uniform from year to year, and between different species. Furthermore, the mechanisms regulating the timing of phenological events in different species seem to function in a similar way, suggesting an unanimous optimal response to climatic conditions.
  • Linkosalo, University of Helsinki, Department of Forest Ecology, Unioninkatu 40 B, P.O. Box 24, FIN-00014 University of Helsinki, Finland E-mail: tapio.linkosalo@helsinki.fi (email)
article id 658, category Research article
Muru Juurola, Pekka Ollonqvist, Heikki Pajuoja, Mikko Toropainen. (1999). Outcomes of forest improvement work in Finland. Silva Fennica vol. 33 no. 3 article id 658. https://doi.org/10.14214/sf.658
Keywords: forest policy; forest improvement work; subsidies; input-output method; allowable cut
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This paper discusses public subsidies aimed at intensifying timber production as an initial part of an evaluation of the profitability of forest investment subsidies in Finland. In many countries there are very few proper ex post evaluations of the forest policy instruments in economic terms. The scarcity of timber among users and their attempts to construct new forest policy are discussed first. The increments in annual growth and growing stock as well as its valuation are then evaluated. The final gross utility increments due to forestry investments are measured through their importance in the forest industry products. The direct and indirect changes in GDP are calculated by using the input-output method.
  • Juurola, The Finnish Forest Research Institute, Helsinki Research Centre, Unioninkatu 40 A, FIN-00170 Helsinki, Finland E-mail: mj@nn.fi
  • Ollonqvist, The Finnish Forest Research Institute, Helsinki Research Centre, Unioninkatu 40 A, FIN-00170 Helsinki, Finland E-mail: po@nn.fi
  • Pajuoja, The Finnish Forest Research Institute, Helsinki Research Centre, Unioninkatu 40 A, FIN-00170 Helsinki, Finland E-mail: heikki.pajuoja@metla.fi (email)
  • Toropainen, The Finnish Forest Research Institute, Helsinki Research Centre, Unioninkatu 40 A, FIN-00170 Helsinki, Finland E-mail: mt@nn.fi
article id 657, category Research article
Paul C. Van Deusen. (1999). Multiple solution harvest scheduling. Silva Fennica vol. 33 no. 3 article id 657. https://doi.org/10.14214/sf.657
Keywords: simulated annealing; Metropolis algorithm; Gibb’s sampler
Abstract | View details | Full text in PDF | Author Info
Application of the Metropolis algorithm for forest harvest scheduling is extended by automating the relative weighting of objective function components. Previous applications of the Metropolis algorithm require the user to specify these weights, which demands substantial trial and error in practice. This modification allows for general incorporation of objective function components that are either periodic or spatial in nature. A generic set of objective function components is developed to facilitate harvest scheduling for a wide range of problems. The resulting algorithm generates multiple feasible solutions rather than a single optimal solution.
  • Van Deusen, Principal Research Scientist, NCASI, Northeast Regional Center, Tufts University, 1 Anderson Hall, Medford, Massachusetts 02155, USA E-mail: pvandeus@tufts.edu (email)
article id 656, category Research article
Klaus Silfverberg, Markus Hartman. (1999). Effects of different phosphorus fertilisers on the nutrient status and growth of Scots pine stands on drained peatlands. Silva Fennica vol. 33 no. 3 article id 656. https://doi.org/10.14214/sf.656
Keywords: needles; Scots pine; stand growth; drainage; phosphorus; peat; fertilisation
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The aim of the study was to compare the effects of phosphorus fertilisers of different solubility and different phosphorus doses. The material was collected from 8 field experiments situated on drained peatlands in southern and central Finland (60°–65°N). The sites were drained, oligotrophic pine fens and pine bogs, which had been fertilised between 1961 and 1977 with different combinations of N, K and P. In 1991–94 stand measurements and foliar and peat sampling were carried out on 162 sample plots. Apatite, rock phosphate and superphosphate affected basal area growth to a rather similar extent. However, apatite slightly surpassed superphosphate and rock phosphate at the end of the study period in two hollow-rich S. fuscum bogs. Higher doses of phosphorus did not significantly increase the basal area growth. The foliar phosphorus concentrations clearly reflected the effect of the P fertilisation. Especially on the pine bogs basic fertilisation with 66 kg P/ha maintained the needle phosphorus concentrations at a satisfactory level for more than 25 years after fertilisation. The amount of phosphorus in the 0–20 cm peat layer was not significantly increased either by basic fertilisation or refertilisation. The phosphorus reserves in the peat in the individual experiments were between 88 and 327 kg/ha. There was a strong correlation between the amounts of phosphorus and iron in the peat. Large amounts of iron in peat may reduce the solubility and availability of phosphorus. According to the foliar phosphorus concentrations in the basic-fertilised plots, the need for refertilisation seems to be unnecessary during the 25-year postfertilisation period at least. None of the basic fertilisation treatments seriously retarded the basal area growth compared to the refertilised treatments. There seems to be a greater shortage of potassium than of phosphorus, because the foliar potassium concentrations and the amounts of potassium in the 0–20 cm peat layer were very low in several of the experiments.
  • Silfverberg, The Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: klaus.silfverberg@metla.fi (email)
  • Hartman, The Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: mh@nn.fi
article id 655, category Research article
Raija-Liisa Petäistö. (1999). Growth phase of bare-root Scots pine seedlings and their susceptibility to Gremmeniella abietina. Silva Fennica vol. 33 no. 3 article id 655. https://doi.org/10.14214/sf.655
Keywords: Pinus; nursery; age; Scleroderris
Abstract | View details | Full text in PDF | Author Info
Bare-root row-sown seedlings of Scots pine (Pinus sylvestris L.) in a forest nursery were inoculated with Gremmeniella abietina conidia at different times during their first and second growing seasons. The following spring, the proportion of diseased seedlings was different in various inoculation time treatments according to the age of the seedlings. The first year seedlings were susceptible to infection until late summer, whereas the second year seedlings were not. It is thought that this difference is due to the different growth rhythms of the first and second year seedlings. The difference in the susceptibility of bare-root seedlings to the disease in various growth corresponded to that reported earlier for container seedlings.
  • Petäistö, Finnish Forest Research Institute, Suonenjoki Research Station, FIN-77600 Suonenjoki, Finland E-mail: raija-liisa.petaisto@metla.fi (email)
article id 654, category Research article
Margus Pensa, Risto Jalkanen. (1999). Needle chronologies on Pinus sylvestris in northern Estonia and southern Finland. Silva Fennica vol. 33 no. 3 article id 654. https://doi.org/10.14214/sf.654
Keywords: needle age; needle trace method; needle retention; crown thinning; needle loss; shedding
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A needle trace method was used to reveal the chronology of needle retention on Scots pine (Pinus sylvestris L.) in southern Finland (two stands) and northern Estonia (two stands). The average annual summer needle retention along the main stem varied from 2.2 to 3.1 in Estonian stands and between 3.4 and 4.2 in Finnish stands during the period 1966–1990. The 23-year-mean needle age was 3.0 and 2.1 years in Finland and Estonia, respectively. In all stands, the mean needle age decreased sharply in 1980s.
  • Pensa, Institute of Ecology, Department of Northeast Estonia, Pargi 15, EE-41537 Jõhvi, Estonia E-mail: margus@ecoviro.johvi.ee (email)
  • Jalkanen, Finnish Forest Research Institute, Rovaniemi Research Station, P.O. Box 16, FIN-96301 Rovaniemi, Finland E-mail: rj@nn.fi
article id 665, category Research article
Kari Kangas. (1999). Trade of main wild berries in Finland. Silva Fennica vol. 33 no. 2 article id 665. https://doi.org/10.14214/sf.665
Keywords: imports; bilberry; non-wood forest products; trade; lingonberry; exports
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The price trends and markets of the main wild berries, bilberry (Vaccinium myrtillus L.) and lingonberry (Vaccinium vitis-idaea L.), were analysed in this study, which covered both domestic use of berries, imports and exports. The periods considered were for bilberries from 1988 to 1997 and for lingonberries from 1979 to 1994. The results indicated that both exports and imports have increased and domestic berries have lost their market share to imports in domestic use. One possible explanation for this trend was found in price development. Both export and import prices have decreased, but export price has still been higher than the import price. Simultaneously the domestic price has decreased the fastest. The formation of the price of lingonberries paid to the pickers in the organised domestic markets was studied with a regression model. The results indicated that domestic price was negatively dependent on the amounts of lingonberries demanded in the domestic markets and positively dependent on the export price. Correlation analysis gave evidence on the same kind of relations concerning bilberries.
  • Kangas, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: kari.kangas@forest.joensuu.fi (email)
article id 664, category Research article
Anssi Niskanen. (1999). The financial and economic profitability of field afforestation in Finland. Silva Fennica vol. 33 no. 2 article id 664. https://doi.org/10.14214/sf.664
Keywords: plantation; efficiency; cost-benefit-analysis; net-present-value
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The aim of the study was to assess the rate of return on invested capital and soil expectation value in field afforestation from the financial (business economic) and economic (national economic) point of views in Finland using 1996 cost and price data. Risks for renewal planting and negative growth impacts of reduction in plantation density were explicitly included in the profitability assessments. Results indicated that due to the subsidies and favorable regulations for obtaining them in 1996, field afforestation was financially profitable for farmers regardless of what species was used for planting. From the national economic point of view, investments in field afforestation provided only substantial return on invested capital, being highest after risk adjustments in Norway spruce (Picea abies) plantations.
  • Niskanen, European Forest Institute, Torikatu 34, FIN-80101 Joensuu, Finland E-mail: anssi.niskanen@efi.fi (email)
article id 663, category Research article
Lennart Moberg. (1999). Variation in knot size of Pinus sylvestris in two initial spacing trials. Silva Fennica vol. 33 no. 2 article id 663. https://doi.org/10.14214/sf.663
Keywords: site quality; silviculture; stand density; azimuth; initial spacing; knot size; within-stand differentiation
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The objective of this study was to investigate the variation in internal knot size of Scots pine (Pinus sylvestris L.) stems sampled from mature permanent plots, and for which the silvicultural history was known. It was based on a sample of mature trees removed from two different spacing trials representing a moderate and high site index. Knot size was measured with non-destructive methods using a CT-scanner and digital image analysis. Initial spacing varied between 0.75 and 3 m on the high site-index trial and between 1.5 and 2.5 m on the moderate site-index trial. Wider initial spacing on the high site index resulted in larger knots near the base of the stem. However, due to successive thinnings which gradually equalised stand density among plots, the difference between most plots was less further up in the stems. The effect of silvicultural regime was much more limited on the lower site index. Within-stand differentiation resulted in a variation of tree diameter (DBH); larger trees had significantly larger knots. Furthermore, knots were larger towards south than towards north in both trials. These results illustrated that, by using non-destructive measurements on trees sampled from permanent research plots, it was possible to simultaneously study the variation of internal knot size at stand (such as site and silviculture effects), within-stand (such as relative tree size) and within-tree levels (such as height and azimuth). However, lack of replication prevented valid statistical inference as to stand-level effects.
  • Moberg, Swedish University of Agricultural Sciences, Department of Forest Management and Products, P.O. Box 7060, S-750 07 Uppsala, Sweden E-mail: lennart.moberg@sh.slu.se (email)
article id 662, category Research article
Lars Björklund. (1999). Identifying heartwood-rich stands or stems of Pinus sylvestris by using inventory data. Silva Fennica vol. 33 no. 2 article id 662. https://doi.org/10.14214/sf.662
Keywords: Pinus sylvestris; sapwood; heartwood; wood utilisation
Abstract | View details | Full text in PDF | Author Info
Variations in heartwood percentage, heartwood radius and sapwood width, within and between stands of Scots pine (Pinus sylvestris L.), were analysed using a database of 198 CT-scanned (computer tomography) stems from 33 research plots (stands) throughout Sweden. Heartwood percentage varied greatly both between individual trees and between stands, and correlated poorly to site, stand and tree variables. This implies that it seems unfeasible to identify heartwood-rich stands or stems, e.g., for production of heartwood products, by using inventory data. Heartwood formation expressed as the number of new heartwood rings formed each year was found to increase with increasing cambial age, from about 0.5 rings per year at a cambial age of 45 years, to about 0.8 rings per year at a cambial age of 115 years.
  • Björklund, Swedish University of Agricultural Sciences, Department of Forest Management and Products, S-750 07 Uppsala, Sweden E-mail: lars.bjorklund@sh.slu.se (email)
article id 661, category Research article
Eira-Maija Savonen, Anna Saarsalmi. (1999). Effects of clone and fertilization on the seed and foliar chemical composition of Scots pine (Pinus sylvestris) grafts. Silva Fennica vol. 33 no. 2 article id 661. https://doi.org/10.14214/sf.661
Keywords: needles; Scots pine; nutrient concentration; seeds; seed orchards
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Effects of clone and fertilization on the seed and foliar nutrient concentrations of Scots pine grafts were investigated in a seed orchard in southern Finland. The seed and foliar samples for chemical analyses were collected during winters 1985–86 and 1988–89 from 39 grafts per clone fertilized in spring 1986. There were 6 clones and 13 treatments for each clone with three replications. The treatments consisted of N, P, K in various combinations, micronutrients, wood ash and grass control. Macro- (N, P, K, Ca, Mg) and micronutrients (Cu, Zn, B) were analysed. There were statistically significant differences between the clones in seed nutrient concentrations. The variation of the K, Mg, Ca, Zn and Cu concentrations between the two study years was considerably larger in the seeds than in the needles. The concentrations of these elements in the seeds were low in the year of an abundant seed crop in spite of fertilization. This had, however, no negative effects on germination of seeds. The proportions of crude fat and crude protein were high in both years (34% and 35% in 1985 ; 33% and 38% in 1988). Fertilization had only minor or no effect at all on the seed chemical composition in the orchard with a satisfactory nutrient status of the soil. Also on the foliar nutrient concentrations the effect of the clone was stronger than that of fertilization. Grafts with large needles produced heavy seeds, which had more storage proteins than the lighter seeds.
  • Savonen, The Finnish Forest Research Institute, Parkano Research Station, Kaironiementie 54, FIN-39700 Parkano, Finland E-mail: eira-maija.savonen@metla.fi (email)
  • Saarsalmi, The Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: as@nn.fi
article id 660, category Research article
Erkki Annila, Bo Långström, Martti Varama, Risto Hiukka, Pekka Niemelä. (1999). Susceptibility of defoliated Scots pine to spontaneous and induced attack by Tomicus piniperda and Tomicus minor. Silva Fennica vol. 33 no. 2 article id 660. https://doi.org/10.14214/sf.660
Keywords: Scots pine; defoliation; Scolytidae; Diprionidae; induced attack; Tomicus piniperda; Tomicus minor; Diprion pini
Abstract | View details | Full text in PDF | Author Info
In 1990–1991, Diprion pini extensively defoliated Scots pine (Pinus sylvestris L.) trees in Lauhanvuori National Park in southwestern Finland. Many trees lost all their foliage, while others had ca. 10% foliage left after the second year of defoliation. Outside the national park, many nearby stands were also heavily defoliated in 1990, but were sprayed with diflubenzuron (Dimilin®) in 1991. This protected the current year needles, corresponding to ca 30% of full foliage. In spring 1992, pine trees with 0, 10, 30 and 100% foliage remaining (10 small and 10 large trees in each category) were baited with pine bolts to induce stem attacks by pine shoot beetles. All baited trees were attacked by Tomicus piniperda and some by T. minor. The attacks failed in all these trees except those that were totally defoliated and some of the small trees with 10% foliage left. Many unbaited trees escaped attack entirely, but only totally defoliated trees were successfully colonized (i.e. produced brood). Attack densities and brood production figures peaked in baited, large and totally defoliated trees. None of the measures (cambial electrical resistance, resin flow, induced lesion length by fungal inoculation, amount of hydrocarbons or phenolic compounds) used to describe tree vigour at the time of attack gave better information than the estimated remaining foliage. We conclude that the risk for beetle-induced mortality following defoliation is a function of remaining needle biomass and beetle pressure. Even at high beetle densities (as was simulated by baiting of trees), trees with 10% of the foliage remaining were able to defend themselves against attacking pine shoot beetles.
  • Annila, Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: erkki.annila@metla.fi (email)
  • Långström, Swedish University of Agricultural Sciences, Dept of Entomology, P. O. Box 7044, S-750 07 Uppsala, Sweden E-mail: bl@nn.fi
  • Varama, Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: mv@nn.fi
  • Hiukka, Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: rh@nn.fi
  • Niemelä, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: pn@nn.fi
article id 671, category Research article
Jouni Vettenranta, Jari Miina. (1999). Optimizing thinnings and rotation of Scots pine and Norway spruce mixtures. Silva Fennica vol. 33 no. 1 article id 671. https://doi.org/10.14214/sf.671
Keywords: mixed forests; optimal treatments; simulation models; Hooke and Jeeves’ method; thinning types
Abstract | View details | Full text in PDF | Author Info
The study describes a simulation-optimization system which uses spatial models for diameter and height growth, crown ratio and tree mortality for Scots pine and Norway spruce mixtures. The optimal one- and two-thinning regimes of six initial stands with varing species composition were solved by using nonlinear optimization. The soil expectation value (SEV) at 3% interest rate was used as a management objective. The regimes are determined by taking into account the stand basal areas before the thinnings, the removal percentages for small, medium-sized and large pines and spruces, and the stand basal area before the final felling. The greatest SEV (8900 FIM ha–1) was attained with the initial stand where the proportion of pines was 65% of the number of the stems. In the two-thinning regime, the first thinning was conducted at the age of 39 years when the stand basal area was 37 m2 ha–1 and the dominant height was about 15 m. After the thinning, the basal area was 27 m2 ha–1. Spruces were thinned from below, but both small and large pines were removed. The second thinning was 8 years later and much heavier: the stand basal area was decreased from 35 m2 ha–1 to 18 m2 ha–1 by removing both small and large pines and spruces. When the optimal two-thinning regime was compared to the regime presented by Forest Centre Tapio, the loss of SEV was about 30% (6070 FIM ha–1) in the case of thinnings from below, and about 20% (7250 FIM ha–1) in the case of thinnings from above.
  • Vettenranta, Kivirinnanpolku 4, FIN-40950 Muurame, Finland E-mail: vettenr@cc.joensuu.fi (email)
  • Miina, Faculty of Forestry, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: jm@nn.fi
article id 670, category Research article
Jouni Vettenranta. (1999). Distance-dependent models for predicting the development of mixed coniferous forests in Finland. Silva Fennica vol. 33 no. 1 article id 670. https://doi.org/10.14214/sf.670
Keywords: Norway spruce; Scots pine; competition; growth models; Monte Carlo simulation; crown models
Abstract | View details | Full text in PDF | Author Info
Distance-dependent growth models and crown models, based on extensive material, were built for Scots pine and Norway spruce growing in a mixed forest. The crown ratio was also used as a predictor in a diameter growth model to better describe the thinning reaction. The effect of crown ratio on the growth dynamics was studied in simulation examples. Monte Carlo simulation was used to correct the bias caused by nonlinear transformations of predictors and response. After thinnings the crown ratio as a predictor was found to be a clear growth-retarding factor. The growth retarding effect was stronger among pines with thinnings from below, whereas the estimated yield of spruces over rotation was slightly greater when the crown ratio was included than without it. With each type of thinning the effect of crown ratio on pine growth was almost the same, but the growth of spruces was clearly delayed when the stand was thinned from above. Simulation examples also showed that it is profitable to raise the proportion of spruces during rotation, since spruces maintain the growth more vigorous at older ages. The total yield during 90 years rotation was about 20% higher if the stand was transformed into a pure spruce stand instead of pine.
  • Vettenranta, Kivirinnanpolku 4, FIN-40950 Muurame, Finland E-mail: vettenr@cc.joensuu.fi (email)
article id 669, category Research article
Simo Poso, Guangxing Wang, Sakari Tuominen. (1999). Weighting alternative estimates when using multi-source auxiliary data for forest inventory. Silva Fennica vol. 33 no. 1 article id 669. https://doi.org/10.14214/sf.669
Keywords: remote sensing; two-phase sampling; forest inventory methods
Abstract | View details | Full text in PDF | Author Info
Five auxiliary data sources (Landsat TM, IRS-IC, digitized aerial photographs, visual photo-interpretation and old forest compartment information) applying three study areas and three estimators, two-phase sampling with stratification, the k nearest neighbors and regression estimator, were examined. Auxiliary data were given for a high number of sample plots, which are here called first phase sample plots. The plots were distributed using a systematic grid over the study areas. Some of the plots were then measured in the field for the necessary ground truth. Each auxiliary data source in combination with field sample information was applied to produce a specific estimator for five forest stand characteristics: mean diameter, mean height, age, basal area, and volume of the growing stock. When five auxiliary data sources were used, each stand characteristic and each first phase sample plot were supplied with five alternative estimates with three alternative estimators. Mean square errors were then calculated for each alternative estimator using the cross validation method. The final estimates were produced by weighting alternative estimates inversely according to the mean square errors related to the corresponding estimator. The result was better than the final estimate of any of the single estimators. The improvement over the best single estimate, as measured in mean square error, was 16.9% on average for all five forest stand characteristics. The improvement was fairly equal for all five forest stand characteristics. Only minor differences among the accuracies of the three alternative estimators were recorded.
  • Poso, Department of Forest Resource Management, P.O. Box 24 (Unioninkatu 40 B), FIN-00014 University of Helsinki, Finland E-mail: simo.poso@helsinki.fi (email)
  • Wang, Department of Forest Resource Management, P.O. Box 24 (Unioninkatu 40 B), FIN-00014 University of Helsinki, Finland E-mail: gw@nn.fi
  • Tuominen, Department of Forest Resource Management, P.O. Box 24 (Unioninkatu 40 B), FIN-00014 University of Helsinki, Finland E-mail: st@nn.fi
article id 668, category Research article
Hannu Hökkä, Timo Penttilä. (1999). Modelling the dynamics of wood productivity on drained peatland sites in Finland. Silva Fennica vol. 33 no. 1 article id 668. https://doi.org/10.14214/sf.668
Keywords: forest drainage; stand development; autoregressive process; site productivity; structural equation models
Abstract | View details | Full text in PDF | Author Info
The dynamics of wood productivity on drained peatland sites was analyzed from the covariance structure generated by stand yield data of repeatedly measured permanent sample plots in 81 Scots pine (Pinus sylvestris L.) or Norway spruce (Picea abies Karst. (L.)) stands with admixtures of birch (Betula pubescens Ehrh.). The site production potential, considered a latent variable, was assumed to follow an autoregressive process over time elapsed since drainage. As a measure of the latent variable, a relative growth rate (RGR) index was determined for all stands at the time of drainage and at four successive measurement time points following drainage (on average 16, 23, 30, and 41 years). The index was calculated as the site index of an upland conifer stand with the ratio of periodic volume growth and standing volume and adjusted by changes in stand stocking and thinning. The observed covariance structure was described by fitting a structural equation model to the data of RGR indices. When only the post-drainage measurement times were included, a quasi-simplex model with equal error variances and equal structural parameters at different measurement times fit the data well indicating a permanent covariance structure among the different measurements. Including the measurement at the time of drainage resulted in a non-permanent structure. The stand parameters at the time of drainage were poorly correlated with post-drainage growth. A considerable increase in the wood productivity of the sites was observed, being greatest during twenty years after drainage and continuing up to 40 years since drainage. This was concluded to be due to changes in site properties rather than stand structure although the effects of the single factors could not be analytically separated from one another. Our modelling approach appeared to improve long-term site productivity estimates based merely on botanical site indices.
  • Hökkä, Finnish Forest Research Institute, Rovaniemi Research Station, PO Box 16, 96301 Rovaniemi, Finland E-mail: hannu.hokka@metla.fi (email)
  • Penttilä, Finnish Forest Research Institute, Rovaniemi Research Station, PO Box 16, 96301 Rovaniemi, Finland E-mail: tp@nn.fi
article id 667, category Research article
Timo Kärki. (1999). Predicting the value of grey alder (Alnus incana) logs based on external quality. Silva Fennica vol. 33 no. 1 article id 667. https://doi.org/10.14214/sf.667
Keywords: alder; log quality; sawn timber quality; log value
Abstract | View details | Full text in PDF | Author Info
The quality of grey alder logs (Alnus incana) was studied by sawing sample logs from two different forests in November 1995–February 1996. For grading of grey alder logs and sawn timber the proposed system of Keinänen and Tahvanainen (1995) plus the reject -grade was used. In general, grey alder logs have knots from the base to the top. All types of knots appear, and the length of the knot-free section is small at the base. In small-dimensioned logs there are fewer knots than in larger logs. Especially in large top logs, there were many more fresh knots than in other types of logs. Evidently, in different types of logs the different grades of sawn timber are located in comparable sections along the length. It also seems that the worse the grade class was, the longer was also the length of the class. The most common reasons for decreasing grade were dry knots and discoloration.
  • Kärki, University of Joensuu, Faculty of Forestry, P.O. BOX 111, FIN-80101 Joensuu, Finland E-mail: timo.karki@forest.joensuu.fi (email)
article id 666, category Research article
Rolf Pape. (1999). Influence of thinning on spiral grain in Norway spruce grown on highly productive sites in southern Sweden. Silva Fennica vol. 33 no. 1 article id 666. https://doi.org/10.14214/sf.666
Keywords: Norway spruce; thinning; grain angle; ring width; growth rate; spiral grain
Abstract | View details | Full text in PDF | Author Info
Grain spirality was investigated in eight stands of Norway spruce (Picea abies (L.) Karst.) subjected to different thinning regimes. The dominating general pattern of spiral grain found in this study was typical for conifers, with a maximum of left-handed spirality close to the pith, which decreased towards the bark and sometimes changed to right-handed spiral grain in the outer growth rings. However, there was a large amount of between-tree variation in spiral grain. The effect of thinning on grain spirality was investigated by relating annual ring width to spiral grain, since thinning affects growth rate. A positive correlation between ring width and grain angle was found, but a considerable number of trees showed no or a negative correlation. A statistically significant effect of ring width was only found in five of the eight stands. Heavy thinnings, removing 60% of the basal area of a stand, considerably increased spiral grain, whereas the effects of light thinnings were inconsistent. These results support the findings of earlier studies indicating that spiral grain formation is under considerable genetic control, while its expression can be changed by silvicultural methods which affect growth rate.
  • Pape, Swedish University of Agricultural Sciences, Department of Forest Yield Research, P.O. Box 7061, S-750 07 Uppsala, Sweden E-mail: rolf.pape@sprod.slu.se (email)
article id 677, category Research article
Pekka Leskinen, Jyrki Kangas. (1998). Analysing uncertainties of interval judgment data in multiple-criteria evaluation of forest plans. Silva Fennica vol. 32 no. 4 article id 677. https://doi.org/10.14214/sf.677
Keywords: forest planning; uncertainty; decision analysis; expert judgment; pairwise comparisons
Abstract | View details | Full text in PDF | Author Info
The use of interval judgments instead of accurate pairwise comparisons has been proposed as a solution to facilitate the analysis of uncertainties in the widely applied pairwise comparisons technique. A method is presented for deriving probability distributions for the pairwise comparisons and for utilizing the distributions in the analysis of uncertainties in the evaluation process. The first step is that the expert or the decision-maker is queried as to the best guess of the priority ratio of the attributes compared. This is followed by an adjusting query concerning the uncertainty in the comparison: what is the probability of the priority ratio being between the best guess ± 1 unit of the pairwise comparison scale? An application of the method is presented in the form of multiple-criteria evaluation of alternative management plans for a forest area.
  • Leskinen, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: pekka.leskinen@metla.fi (email)
  • Kangas, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: jk@nn.fi
article id 676, category Research article
Erik Sundström. (1998). Afforestation of low-productive peatlands in Sweden – a tree species comparison. Silva Fennica vol. 32 no. 4 article id 676. https://doi.org/10.14214/sf.676
Keywords: Pinus sylvestris; fertilization; Picea abies; afforestation; drainage; tree species; climate; Pinus contorta; broadcast fertilization; spot fertilization
Abstract | View details | Full text in PDF | Author Info
In 1970, five low-productive treeless peatlands in Sweden, ranging from latitudes 56°N to 67°N, were drained and fertilized for afforestation. The aim of the study was to determine the effect of four ditch spacings, varying from 7.5 to 60 m, and five NPK-fertilizer combinations, on the survival and growth of planted Scots pine (Pinus sylvestris), lodgepole pine (Pinus contorta), Norway spruce (Picea abies), and silver birch (Betula pendula) seedlings. The assessments were carried out 18–22 years after planting. Neither silver birch, nor Norway spruce was regarded suitable for the site type. The mortality of silver birch was almost complete, and Norway spruce did not grow well in any of the study areas, however, better than Scots pine in the north. Lodgepole pine had better height and diameter growth but also higher mortality rates than Scots pine. In the two northernmost experimental areas no response to fertilization was found. In the other three areas, the response to fertilization did not differ between species. Phosphorus was the most effective of the added fertilizer elements, whereas nitrogen showed no positive effect. Broadcast fertilizer application, with three times higher amount of fertilizer per ha gave the same growth response as spot application.
  • Sundström, Swedish University of Agricultural Sciences, Dept. of Silviculture, S-901 83 Umeå, Sweden E-mail: erik.sundstrom@ssko.slu.se (email)
article id 675, category Research article
Per Linder, Peter Jonsson, Mats Niklasson. (1998). Tree mortality after prescribed burning in an old-growth Scots pine forest in northern Sweden. Silva Fennica vol. 32 no. 4 article id 675. https://doi.org/10.14214/sf.675
Keywords: forest management; coarse woody debris; tree mortality; nature conservation; forest fire; dead trees
Abstract | View details | Full text in PDF | Author Info
Tree mortality and input of dead trees were studied after a prescribed burning in a forest reserve in northern Sweden. The stand was a multi-layered old-growth forest. The overstorey was dominated by Scots pine (Pinus sylvestris L.) and the understorey consisted of mixed Scots pine and Norway spruce (Picea abies L. Karst.). Ground vegetation was dominated by ericaceous dwarf-shrubs and feathermosses. The stand has been affected by six forest fires during the last 500 years. The prescribed burning was a low intensity surface fire that scorched almost 90% of the ground. Tree mortality for smaller pines and spruces (DBH < 10 cm) was over 80% in the burned parts of the reserve. For larger pines, 10–50 cm DBH, mortality showed a decreasing trend with increasing diameter, from 14% in class 10–20 cm DBH to 1.4% in class 40–50 cm DBH. However, pines with DBH ≥ 50 cm had a significantly higher mortality, 20%, since a high proportion of them had open fire scars containing cavities, caused by fungi and insects, which enabled the fire to burn inside the trunks and hollow them out. The fire-induced mortality resulted in a 21 m3 ha–1 input of dead trees, of which 12 m3 ha–1 consisted of trees with DBH ≥ 30 cm. An increased mortality among larger trees after low-intensity fires has not previously been described in Fennoscandian boreal forests, probably owing to a lack of recent fires in old-growth stands. However, since large pines with open fire scars were once a common feature in the natural boreal forest, we suggest that this type of tree mortality should be mimicked in forestry practices aiming to maintain and restore natural forest biodiversity.
  • Linder, Department of Forest Vegetation Ecology, SLU, S-901 83 Umeå, Sweden E-mail: pl@nn.se (email)
  • Jonsson, Department of Forest Vegetation Ecology, SLU, S-901 83 Umeå, Sweden E-mail: pj@nn.se
  • Niklasson, Southern Swedish Forest Research Centre, SLU, Box 49, S-230 53 Alnarp, Sweden E-mail: mn@nn.se
article id 674, category Research article
Tuula Aalto. (1998). Carbon dioxide exchange of Scots pine shoots as estimated by a biochemical model and cuvette field measurements. Silva Fennica vol. 32 no. 4 article id 674. https://doi.org/10.14214/sf.674
Keywords: Pinus sylvestris; photosynthesis; Scots pine; CO2 exchange; cuvette measurements
Abstract | View details | Full text in PDF | Author Info
A biochemical model was used to calculate CO2 fluxes to Scots pine shoots in two boreal measurement stations, Hyytiälä in southern Finland (61°51’N, 24°17’E) and Värriö in northern Finland (67°46’N, 29°35’E). The results of the model were compared with cuvette measurements performed in field conditions. A differential equation for change in gas concentration inside a closed cuvette was constructed and solved in order to obtain conductances and fluxes. The results were generally in a good agreement, the correlation coefficients varied from 0.74 to 0.95. Some discrepancies were also found. The model followed more intensively changes in temperature. This could be seen in northern Finland measurements at low temperatures (< 18 °C). The modelled temperature response indicated low fluxes at low temperatures, but measurements did not show any decrease. The irradiation response was relatively similar in both measuring sites and according to the model. Cuvette measurements showed slightly smaller quantum yields as a result from shading of the needles. The temperature dependences of the biochemical model parameters Jmax and Vc(max) were re-evaluated from the field measurements. The results for Vc(max) agreed well with earlier estimations, while the results for Jmax indicated relatively high values at low temperatures especially in northern Finland. Exponential fitting produced also substomatal concentrations of CO2, which agreed quite well with the model. The daily minimum of substomatal/ambient concentration ratio varied from 0.4 to 0.8.
  • Aalto, Department of Physics, P.O. Box 9, FIN-00014 University of Helsinki, Finland E-mail: tuula.aalto@helsinki.fi (email)
article id 673, category Research article
Lauri Palva, Eero Garam, Sari Palmroth, Raimo Sepponen, Pertti Hari. (1998). Utilizing a multipoint measuring system of photosynthetically active radiation in photosynthetic studies within canopies. Silva Fennica vol. 32 no. 4 article id 673. https://doi.org/10.14214/sf.673
Keywords: carbon dioxide; instrumentation; measurement; photosynthetically active radiation
Abstract | View details | Full text in PDF | Author Info
A novel multipoint measuring system of photosynthetically active radiation (PAR) has been constructed and operated within a Scots pine canopy. A regular grid of 800 measuring points has been incorporated into a cuvette to observe the spatial and temporal distribution of PAR incident on the needles of a twig along with simultaneous measurements of the CO2 exchange in order to determine the dependence of photosynthesis on PAR. It was shown that large errors can result if the photosynthetic rate is estimated using the mean value of PAR instead of the instantaneous values of PAR detected at given points in the region of the needles. The results demonstrate that the obtained regression between the CO2 exchange rate estimated using the multipoint PAR measuring system and the measured CO2 exchange rate is as good within a canopy as in unshaded conditions.
  • Palva, Helsinki University of Technology, Applied Electronics Laboratory, P.O. Box 3000, FIN-02015 HUT, Finland E-mail: lauri.palva@hut.fi (email)
  • Garam, Helsinki University of Technology, Applied Electronics Laboratory, P.O. Box 3000, FIN-02015 HUT, Finland E-mail: eg@nn.fi
  • Palmroth, University of Helsinki, Dept. of Forest Ecology, P.O. Box 24, FIN-00014 University of Helsinki, Finland E-mail: sp@nn.fi
  • Sepponen, Helsinki University of Technology, Applied Electronics Laboratory, P.O. Box 3000, FIN-02015 HUT, Finland E-mail: rs@nn.fi
  • Hari, University of Helsinki, Dept. of Forest Ecology, P.O. Box 24, FIN-00014 University of Helsinki, Finland E-mail: ph@nn.fi
article id 672, category Research article
Rimvydas Vasiliauskas, Jan Stenlid. (1998). Spread of Stereum sanguinolentum vegetative compatibility groups within a stand and within stems of Picea abies. Silva Fennica vol. 32 no. 4 article id 672. https://doi.org/10.14214/sf.672
Keywords: Picea abies; Stereum sanguinolentum; somatic incompatibility; decay; wounds
Abstract | View details | Full text in PDF | Author Info
A total of 57 naturally established Stereum sanguinolentum isolates was obtained from artificially wounded Picea abies stems in a forest area of 2 ha in Lithuania. Somatic incompatibility tests revealed 27 vegetative compatibility groups (VCGs) that contained 1–10 isolates. There was no spatial clustering of S. sanguinolentum VCGs within the forest area. The extent of S. sanguinolentum decay was analysed in 48 P. abies stems, 9–26 cm in diameter at breast height. Within 7 years of wounding, the length of S. sanguinolentum decay column in stems was 107–415 cm (291.5 ± 77.3 cm on average), lateral spread of the fungus at the butt was 38–307 cm2 (142.3 ± 66.8 cm2) and decayed proportion of the stem cross-section at the wound site (the butt) was 3–84% (36.8 ± 19.7%). In average, S. sanguinolentum VCG that infected 10 trees exhibited more slow growth inside the stem than VCGs that infected only one tree, and vertical growth varied to a greater extent within this VCG than among different VCGs. Correlation between stem diameter and vertical spread of S. sanguinolentum was not significant (r = –0.103). Despite uniformity of debarked area on all stems 7 years ago (300 cm2), open wound sizes on individual trees at the time of study were between 97–355 cm2 (215.1 ± 59.2 cm2) indicating large differences in wound healing capacity.
  • Vasiliauskas, Department of Plant Protection, Lithuanian University of Agriculture, LT-4324 Kaunas, Lithuania E-mail: rv@nn.lt (email)
  • Stenlid, Department of Forest Mycology & Pathology, Swedish University of Agricultural Sciences, P.O. Box 7026, S-750 07 Uppsala, Sweden E-mail: js@nn.se
article id 686, category Research article
Jean Yves Fraysse, Loïc Crémière. (1998). Nursery factors influencing containerized Pinus pinaster seedlings’ initial growth. Silva Fennica vol. 32 no. 3 article id 686. https://doi.org/10.14214/sf.686
Keywords: fertilization; nitrogen; nursery; growth; planting date; maritime pine; sowing date
Abstract | View details | Full text in PDF | Author Info
Maritime pine (Pinus pinaster) containerized seedlings were raised outdoors at different fertilizer regimes, sowing date or culture duration to assess nursery factors influencing first year growth in the field. Seedling biomass, and N, P and K content before outplanting were affected by these different factors but the one year field growth was more related to N concentration than with morphological traits. The results are discussed in view to improve the plant stock quality in nursery.
  • Fraysse, AFOCEL, Domaine de Sivaillan, 33480 Moulis-en-Médoc, France E-mail: sudouest@afocel.fr (email)
  • Crémière, AFOCEL, Domaine de Sivaillan, 33480 Moulis-en-Médoc, France E-mail: lc@nn.fr
article id 685, category Research article
S. P. Sah, I. C. Dutta, M. S. Haque. (1998). Nursery and field response of sissoo plants (Dalbergia sissoo) to Rhizobium inoculation. Silva Fennica vol. 32 no. 3 article id 685. https://doi.org/10.14214/sf.685
Keywords: Rhizobium; Dalbergia sissoo; nodules; rehabilitation; soils; nitrogen-fixation
Abstract | View details | Full text in PDF | Author Info
The present research work aims to demonstrate the rehabilitation of a degraded forest land by afforestation of sissoo plants (Dalbergia sissoo Roxb.) inoculated with nitrogen fixing bacteria sp. In this study, effects of the three different Rhizobium isolates from three localities such as i. Kotre-isolate, ii. Pokhara-isolate and iii. Syangaza-isolate were assessed both in nursery and in the field. It was noted that the growth and biomass increment of seedlings in nursery as well as in the field after rhizobial inoculation were significantly high compared to control one. Among these 3-isolates, Kotre-isolate was found to be superior to others. Soil improvement around the root of inoculated seedlings was remarkable high. Nitrogen content of the soil increased in the range of 20–40% compared to control (only 10–20%). Kotre-isolate, in general, caused more soil improvement than the other isolates. The nutrient content in the green foliage, particularly nitrogen, increased in the range of 30–50%, compared to control one. The considerable increase in nutrients content of soil as well as in the foliage indicates the improvement in the quality of site.
  • Sah, Department of Biology, Kathmandu University, Dhulikhel, Nepal E-mail: ssah@wlink.com.np (email)
  • Dutta, Institute of Forestry, Tribhuvan University, Pokhara, Nepal E-mail: icd@nn.np
  • Haque, Institute of Forestry, Tribhuvan University, Pokhara, Nepal E-mail: msh@nn.np
article id 684, category Research article
H. M. McKay. (1998). Root electrolyte leakage and root growth potential as indicators of spruce and larch establishment. Silva Fennica vol. 32 no. 3 article id 684. https://doi.org/10.14214/sf.684
Keywords: root growth potential; functional root
Abstract | View details | Full text in PDF | Author Info
The relationship between the condition of bare-rooted 2-year-old seedlings of Sitka spruce and larch at the time of planting and their survival and growth after 2 years was examined. Data were analysed for 2 experiments using seedlings lifted and stored at +1 °C throughout the winter for planting in April and also for 2 experiments using seedlings planted directly on different dates without cold storage. Electrolyte leakage from the fine roots of spruce was closely correlated to survival following direct planting at different times from September to April and fine root leakage was a more accurate indicator of spruce performance than root growth potential. However the pattern of larch survival of directly planted stock was more closely related to root growth potential than to root leakage. When seedlings were cold-stored, root electrolyte leakage and root growth potential were modified during storage and following cold storage, the performance of both species was more closely related to root electrolyte leakage than root growth potential. These results are interpreted as meaning that successful establishment of bare-rooted seedlings requires a functional nursery root system that is capable of both supplying adequate water for a limited period immediately after transplanting and of producing roots to meet the seedling’s increased water demand later in the growing season.
  • McKay, Forest Commission Research Agency, Northern Research Station, Roslin, Midlothian, EH25 9SY, Scotland E-mail: h.mckay@forestry.gov.uk (email)
article id 683, category Research article
Anders Karlsson, Arne Albrektson, Anders Forsgren, Lennart Svensson. (1998). An analysis of successful natural regeneration of downy and silver birch on abandoned farmland in Sweden. Silva Fennica vol. 32 no. 3 article id 683. https://doi.org/10.14214/sf.683
Keywords: Betula pendula; site preparation; Betula pubescens; afforestation; arable land; site conditions
Abstract | View details | Full text in PDF | Author Info
To improve our understanding of factors influencing the success of natural regeneration with downy birch (Betula pubescens Ehrh.) and silver birch (Betula pendula Roth) on abandoned farmlands, a survey was conducted to analyse the effects of site conditions and site preparation characteristics. The study was based on a sample plot inventory conducted in one northern and one southern district of Sweden, in which 29 successfully established, naturally regenerated stands, about to be cleaned or thinned, were assessed. Radical site preparation increased stand density and uniformity of established regeneration, and gave faster initial development, than establishment without site preparation on former leys or meadows. Large proportions of the total sample area were classified as moist, and soils consisting of sand–fine sand or peat were frequent. The frequency of birch stems was highest in mesic sites, and on soils consisting of sand, sand–fine sand or peat. Distances to seed-trees were generally shorter than 80 m, and downy birch was the dominant species in most stands.
  • Karlsson, Swedish University of Agricultural Sciences, Department of Silviculture, S-901 83 Umeå, Sweden E-mail: anders.karlsson@ssko.slu.se (email)
  • Albrektson, Swedish University of Agricultural Sciences, Department of Silviculture, S-901 83 Umeå, Sweden E-mail: aa@nn.se
  • Forsgren, Swedish University of Agricultural Sciences, Department of Silviculture, S-901 83 Umeå, Sweden E-mail: af@nn.se
  • Svensson, Swedish University of Agricultural Sciences, Department of Silviculture, S-901 83 Umeå, Sweden E-mail: ls@nn.se
article id 682, category Research article
Guangxing Wang, Simo Poso, Mark-Leo Waite, Markus Holopainen. (1998). The use of digitized aerial photographs and local operation for classification of stand development classes. Silva Fennica vol. 32 no. 3 article id 682. https://doi.org/10.14214/sf.682
Keywords: calibration; classification; digitized aerial photographs; plot window location; local operation
Abstract | View details | Full text in PDF | Author Info
The increasing capacity of modern computers has created the opportunity to routinely process the very large data sets derived by digitizing aerial photographs. The very fine resolution of such data sets makes them better suited than satellite imagery for some applications; however, there may be problems in implementation resulting from variation in radial distortion and illumination across an aerial photograph. We investigated the feasibility of using local operators (e.g., non-overlapping moving window means and standard deviations) as auxiliary data for generating stand development classes via three steps: (i) derive 6 local operators intended to represent texture for a 16 by 16 m window corresponding to a forest inventory sampling unit, (ii) apply a calibration process (e.g., accounting for location relative to a photo's principal point and solar position) to these local operators, and (iii) apply the calibrated local operators to classify the forest for stand development. Results indicate that calibrated local operators significantly improve the classification compared to what is possible using uncalibrated local operators and satellite images.
  • Wang, Department of Natural Resources and Environmental Sciences, University of Illinois, Urbana, IL, USA E-mail: wang12@staff2.cso.uiuc.edu (email)
  • Poso, Department of Forest Resource Management, P.O. Box 24, FIN-00014 University of Helsinki, Finland E-mail: sp@nn.fi
  • Waite, Department of Forest Resource Management, P.O. Box 24, FIN-00014 University of Helsinki, Finland E-mail: mlw@nn.fi
  • Holopainen, Department of Forest Resource Management, P.O. Box 24, FIN-00014 University of Helsinki, Finland E-mail: mh@nn.fi
article id 681, category Research article
Virpi Palomäki, Alpo Hassinen, Matti Lemettinen, Timo Oksanen, Heljä-Sisko Helmisaari, Jarmo Holopainen, Seppo Kellomäki, Toini Holopainen. (1998). Open-top chamber fumigation system for exposure of field grown Pinus sylvestris to elevated carbon dioxide and ozone concentration. Silva Fennica vol. 32 no. 3 article id 681. https://doi.org/10.14214/sf.681
Keywords: Pinus sylvestris; CO2; open-top chamber; O3
Abstract | View details | Full text in PDF | Author Info
An open-top chamber fumigation system was built in a young Scots pine stand to study the effects of realistic elevated ozone (O3) and carbon dioxide (CO2) concentrations and their combination on trees in natural conditions. Doubled CO2 concentration compared to present ambient concentration, and O3 concentration between 40 ppb and 70 ppb in the first study year (1994) and doubled O3 concentration in years 1995 and 1996 were the target concentrations in the chambers. The O3 concentration in the chambers was successfully maintained close to the target concentration and differences between chambers were small. The mean CO2 concentration in the CO2 treatment was ca. 100 ppm below the target, but was maintained at this level throughout the growing season. Two degrees higher mean air temperature and slightly lower light intensity compared to open air were measured in the chambers. The operation of the fumigation system was satisfactory during the three study years and repeatability of the gas treatments can be regarded good in this low cost exposure system.
  • Palomäki, University of Kuopio, Department of Ecology and Environmental Science, P.O.Box 1627, FIN-70211 Kuopio, Finland E-mail: virpi.palomaki@uku.fi (email)
  • Hassinen, Mekrijärvi Research Station, FIN-82900 Ilomantsi, Finland E-mail: ah@nn.fi
  • Lemettinen, Mekrijärvi Research Station, FIN-82900 Ilomantsi, Finland E-mail: ml@nn.fi
  • Oksanen, University of Kuopio, Department of Ecology and Environmental Science, P.O.Box 1627, FIN-70211 Kuopio, Finland E-mail: to@nn.fi
  • Helmisaari, Finnish Forest Research Institute, Department of Forest Ecology, P.O.Box 18, FIN-01301 Vantaa, Finland E-mail: hsh@nn.fi
  • Holopainen, Agricultural Research Centre, FIN-31600 Jokioinen, Finland E-mail: jh@nn.fi
  • Kellomäki, University of Joensuu, Faculty of Forestry, P.O.Box 111, FIN-80101 Joensuu, Finland E-mail: sk@nn.fi
  • Holopainen, University of Kuopio, Department of Ecology and Environmental Science, P.O.Box 1627, FIN-70211 Kuopio, Finland E-mail: th@nn.fi
article id 692, category Research article
Eero Mattila. (1998). Use of satellite and field information in a forest damage survey of eastern Finnish Lapland in 1993. Silva Fennica vol. 32 no. 2 article id 692. https://doi.org/10.14214/sf.692
Keywords: National Forest Inventory; remote sensing; forest damage survey
Abstract | View details | Full text in PDF | Author Info
The study area consists of the Finnish part of a Landsat 5 TM image from 1990. Three independent field samples were measured during 1991–93 in the study area. The first sample was used to compile training areas for supervised maximum likelihood classification of the image. Classification accuracy was studied in the second sample. The spectral separability of the forest strata usable in practical forestry was poor. The extent of the damage area was estimated by the principle of stratified sampling. The estimate included considerable bias because the field sample had not been objectively selected from the image classes. The third field sample was measured as part of the National Forest Inventory of Finland. It is wholly objective, and about ten times larger than the two earlier field samples. The poor spectral separability of the forest strata was confirmed by the NFI sample. However, this sample could be used in stratified sampling with little or no bias in the estimation of the damage area estimate. 14 different damage types were separated according to specific damaging agent. A thematic map was produced which presents the spatial distribution of two damage-rich image classes. The study area comprises 18 300 sq.km, of which 38% were damaged. At first sight it would appear that the proportion of damaged forest has tripled in ten years. However, this is not the case because now special attention was paid to forest health in the field work. Despite this, it is possible that some damage caused by unfavourable climatic phenomena in the ’80s was still perceptible in 1993. No damage caused directly by air pollution has yet been verified in the study area.
  • Mattila, Finnish Forest Research Institute, Joensuu Research Station, P.O. Box 68, FIN-80101 Joensuu, Finland E-mail: eero.mattila@metla.fi (email)
article id 691, category Research article
Jacek Oleksyn, Mark G. Tjoelker, Peter B. Reich. (1998). Adaptation to changing environment in Scots pine populations across a latitudinal gradient. Silva Fennica vol. 32 no. 2 article id 691. https://doi.org/10.14214/sf.691
Keywords: Pinus sylvestris; climate change; provenance; growth; temperature; photoperiod
Abstract | View details | Full text in PDF | Author Info
In several growth chamber and field experiments we examined the growth response of Scots pine (Pinus sylvestris L.) populations from a wide latitudinal range to temperature and photoperiod. The duration of the shoot elongation period of one-year-old seedlings was affected by temperature and photoperiod. In contrasting temperatures, 23/20 °C, 20/17 °C, and 17/14 °C (day/night), shoot elongation period for all populations was shortest in the high and longest in the low temperature treatments. The northern populations from 61–57°N ceased height growth earlier than the other populations in the southern 50°N photoperiod. The order of growth cessation among populations at 50°N in the chamber experiment and at 52°N in the field experiment was similar and related to observed population differences in terminal leader growth and total tree height. Since the length of growing season is under strong environmentally-mediated genetic control in Scots pine, potential climatic changes such as increasing temperature will probably alter the length and timing of growth in aboveground tree parts, but likely in the opposite direction (a shorter growing season) than has been often hypothesized (a longer growing season). Tree-ring analyses of a provenance experiment established in 1912 indicate that the main climatic factors that limited ring-width growth in Scots pine were air temperatures in the winter months of December through March. Low winter temperatures were followed by the formation of narrow rings over the next summer. Based on responses to temperature, Scots pine populations from the continuous European range can be divided in several geographic groups along a latitudinal gradient. Our results suggest that in developing new models to predict the response of Scots pine to changing environmental conditions, it is necessary to include intraspecific differentiation in acclimation and adaptation to environmental factors.
  • Oleksyn, Polish Academy of Sciences, Institute of Dendrology, Parkowa 5, PL-62-035 Kórnik, Poland; University of Minnesota, Department of Forest Resources, 1530 Cleveland Ave. N., St. Paul, MN 55108, USA E-mail: oleks001@gold.tc.umn.edu (email)
  • Tjoelker, University of Minnesota, Department of Forest Resources, 1530 Cleveland Ave. N., St. Paul, MN 55108, USA E-mail: mgt@nn.us
  • Reich, University of Minnesota, Department of Forest Resources, 1530 Cleveland Ave. N., St. Paul, MN 55108, USA E-mail: pbr@nn.us
article id 690, category Research article
Bengt Persson. (1998). Will climate change affect the optimal choice of Pinus sylvestris provenances? Silva Fennica vol. 32 no. 2 article id 690. https://doi.org/10.14214/sf.690
Keywords: Pinus sylvestris; yield; survival; temperature sum; climatic warming; provenance transfer
Abstract | View details | Full text in PDF | Author Info
Provenance experiments with Pinus sylvestris (L.) were evaluated in Sweden north of latitude 60°N. Survival and yield were determined as functions of temperature sum of the site and latitudinal origin of the provenance. Altitudinal origin was of negligible importance. The effects of latitudinal transfer were influenced by temperature sum at the growing site. At the harshest situated sites southward transfer longer than 3° was optimal for survival and yield, whereas transfer effects in a mild climate were weak. Climatic warming would reduce demands of hardiness. However, moderate differences in productivity are expected between formerly optimal seed sources and the ones adapted to changed climatic conditions. Since mortality usually was low in plantations older than 20 years or higher than 2 m, established stands are expected to be robust against adverse effects of climate change.
  • Persson, Högskolan Dalarna, S-781 88 Borlänge, Sweden E-mail: bpn@du.se (email)
article id 689, category Research article
Erik W. Andersson, Kostas A. Spanos, Timothy J. Mullin, Dag Lindgren. (1998). Phenotypic selection compared to restricted combined index selection for many generations. Silva Fennica vol. 32 no. 2 article id 689. https://doi.org/10.14214/sf.689
Keywords: genetic diversity; heritability; status number; phenotypic selection; index selection
Abstract | View details | Full text in PDF | Author Info
A breeding population has been subjected to repeated selection and crossing by simulation. Unrestricted phenotypic selection and restricted combined index selection were compared at the same effective number for five generations. Results show that phenotypic selection often achieves the gain and diversity possible to achieve by combined index selection but the relative efficiency is different for different family sizes and heritabilities. When phenotypic selection was compared with restricted combined index method at low heritabilities, both methods performed almost equally in terms of gain at the same effective number in small family sizes, although in large families, phenotypic selection was less efficient. At high heritabilities phenotypic selection was as efficient as combined index selection. Phenotypic selection was more efficient in conserving additive variance than combined index selection over five generations compared at the same gain and effective number. The introduction of a dominance component to the total variance had little effect. An increased breeding population size by a factor of ten resulted in an increased additive gain by app. 15%. The conclusion is that even though combined index selection is superior in identifying and extracting the potential for breeding achievements, it is generally not performing better than mass selection when compared at the same effective population size in small families.
  • Andersson, Department of Forest Genetics and Plant Physiology, Swedish University of Agricultural Sciences, S-901 83, Umeå, Sweden E-mail: erik.andersson@genfys.slu.se (email)
  • Spanos, N.AG.RE.F.-Forest Research Institute, 57006 Vassilika, Thessaloniki, Greece E-mail: kas@nn.gr
  • Mullin, Genesis Forest Science Canada Inc., C.P. 64 Succursale Haute-Ville, Québec, QC G1R 4M8 Canada E-mail: tjm@nn.ca
  • Lindgren, Department of Forest Genetics and Plant Physiology, Swedish University of Agricultural Sciences, S-901 83, Umeå, Sweden E-mail: dl@nn.se
article id 701, category Research article
Antrei Lausti, Markku Penttinen. (1998). The analysis of return and its components of non-industrial private forest ownership by forestry board districts in Finland. Silva Fennica vol. 32 no. 1 article id 701. https://doi.org/10.14214/sf.701
Keywords: non-industrial private forests; risk; return; portfolio management; forestry board district; performance evaluation
Abstract | View details | Full text in PDF | Author Info
Non-industrial private forest ownership returns and risks in Finland are both estimated and disaggregated to local Forestry Board Districts (FBD) level. Additionally, the FBD level return is divided into price change, felling and change in the growing stock components, which are compared with the inflation rate. The results are based on a complete count of the stumpage prices, silvicultural costs and state subsidies as well as the National Forest Inventory (NFI) data. The influence of taxation is discussed as well. Although this database is excellent for economic studies as well, the estimation methodology is vitiated by a host of problems, the resolution of which is the major contribution of this study. The study period is 1972–1996. The results show that there have been fairly large differences in forest ownership returns and prices depending on the Forestry Board District. Results show that the price change component has been larger in Northern Forestry Board Districts, as much as 0.9% above the inflation rate in Lapland FBD, than in Southern Forestry Board Districts, 1.5% less than the inflation rate in southern Helsinki FBD. The net increase, however, has been larger in Southern Forestry Board Districts than in Northern Forestry Board Districts. Using the average net increment in Finland as a comparison base, the net increment in South Karelia exceeded it by 0.6%, but fell below it by 1.8% in Northeastern Finland. Finally, the return over the whole period is compared to the return on private housing and inflation in the case of North Savo. In all, the estimation methodology developed also serves as spin-off product development for the Forest Statistics Information Service (FSIS).
  • Lausti, Helsinki School of Economics and Business Administration, Centre for Doctoral Programme, Runeberginkatu 15 A, FIN-00100 Helsinki, Finland E-mail: al@nn.fi
  • Penttinen, Finnish Forest Research Institute, Helsinki Research Centre, Unioninkatu 40 A, FIN-00170 Helsinki, Finland E-mail: markku.penttinen@metla.fi (email)
article id 700, category Research article
Riitta H. Hänninen. (1998). Exchange rate changes and the Finnish sawnwood demand and price in the UK market. Silva Fennica vol. 32 no. 1 article id 700. https://doi.org/10.14214/sf.700
Keywords: exchange rate; export demand; Johansen’s cointegration method; pass-through; sawnwood price; United Kingdom
Abstract | View details | Full text in PDF | Author Info
This paper examines the long-run influence of exchange rate changes on the Finnish sawnwood price in the United Kingdom (UK) using quarterly data for the period 1978–1994. The degree of influence was measured by a pass-through coefficient (PT) obtained from a markup pricing relation of a system model. The model, which included export demand and price equations, was estimated with the cointegration method of Johansen. The results indicated a large PT, which means that exchange rate changes are reflected almost proportionately in Finnish export price expressed in pounds sterling. Thus, the Finnish price of sawnwood in pounds has lowered as a result of depreciation of the Finnish markka (FIM). This has improved Finnish competitiveness and market share in the UK. Appreciation of the FIM has had the opposite effect. It seems that Finnish exporters have made use of depreciations and devaluations of the FIM to maintain and increase their market shares but not necessarily their markups. For Finland, which is in the process of joining the European economic and monetary union (EMU), knowing the size of the PT is also important in assessing the economic impact of membership.
  • Hänninen, Forest Research Institute, Helsinki Research Centre, Unioninkatu 40 A, 00170 Helsinki, Finland E-mail: riitta.hanninen@metla.fi (email)
article id 699, category Research article
Heimo Karppinen. (1998). Values and objectives of non-industrial private forest owners in Finland. Silva Fennica vol. 32 no. 1 article id 699. https://doi.org/10.14214/sf.699
Keywords: non-industrial private forests; landowner objectives; forest values; owner characteristics; forestry behavior
Abstract | View details | Full text in PDF | Author Info
The purpose of the study was to create an empirical typology of non-industrial private forest owners based on forest values and long-term objectives of forest ownership, to identify these types by owner and holding characteristics, as well as to analyze silvicultural and harvesting behavior in these groups. The analysis was based on survey data on 245 forest owners in southeastern Finland. The results indicated that general forest values and long-term objectives of forest ownership are not strongly correlated. The results further suggested that the sole emphasis on economic benefits of forests does not lead to the most active silvicultural and harvesting behavior. Multiobjective owners, who underlined both monetary and amenity benefits of their forest property, were the most active in their silvicultural and cutting behavior. Non-timber objectives seemed not to exclude wood production: a group called recreationists harvested slightly less than other owners. Recreationists were willing to invest in forestry but were selective with respect to management practices. The results can be used in planning and implementation of public forest policy such as allocation of the resources of forestry extension services. Forest industries should also benefit from a knowledge of the objectives of roundwood sellers.
  • Karppinen, Finnish Forest Research Institute, Helsinki Research Centre, Unioninkatu 40 A, FIN-00170 Helsinki, Finland E-mail: heimo.karppinen@helsinki.fi (email)
article id 698, category Research article
Kalle Kärhä, Sami Oinas. (1998). Satisfaction and company loyalty as expressed by non-industrial private forest owners towards timber procurement organizations in Finland. Silva Fennica vol. 32 no. 1 article id 698. https://doi.org/10.14214/sf.698
Keywords: timber trade; company loyalty; decision support system (DSS); non-industrial private forest (NIPF) owner; satisfaction; timber procurement
Abstract | View details | Full text in PDF | Author Info
The satisfaction and company loyalty as expressed by non-industrial, private forest (NIPF) owners towards timber procurement organizations were clarified via a mail questionnaire. The results denoted that there is a positive correlation between the levels of forest owners’ expectations (EXP) and perceived performance (PERF). In addition, the lower EXP and the higher PERF were, the greater was the overall satisfaction index (SAT). About two thirds of forest owners were characterized by a negative SAT value or then they were dissatisfied with the performance of the procurement organization in their last timber-sales transaction. Furthermore, the results obtained indicated that the SAT index significantly influences the company loyalty expressed by NIPF owners – their willingness to give favourable reports of the company to the others, and their willingness to engage in future timber-sales transactions with the same company. Nonetheless, the SAT index did not affect the market share of a particular company, and this could be partly accounted for by the similar level of performance among the companies and the lack of competition in the timber trade. Finally, the results revealed several dimensions of timber procurement, and some seller segments to which companies could pay more attention to in order to be able to achieve better satisfaction levels and the loyalty of NIPF owners in the future. One approach to successfully addressing these challenges could be the adopting of a satisfaction decision support system (SatDSS).
  • Kärhä, Faculty of Forestry, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: karha@forest.joensuu.fi (email)
  • Oinas, Faculty of Forestry, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: so@nn.fi
article id 697, category Research article
Kari Pasanen. (1998). Integrating variation in tree growth into forest planning. Silva Fennica vol. 32 no. 1 article id 697. https://doi.org/10.14214/sf.697
Keywords: risk; growth index; ARMA models; scenario approach
Abstract | View details | Full text in PDF | Author Info
Forest planning is always influenced by uncertain factors. Variations in growth, outcome of regeneration, timber prices, costs and mortality cannot be avoided, whereas the quality of inventory data and the models used for estimation of the state and development of forests can be improved. Methods have been developed for incorporating risk and attitude toward risk in decision analysis, but there has been a lack of good models for dealing with the various sources of risk. The aim of this study was to estimate stochastic models for the variation in growth of Scots pine (Pinus sylvestris), Norway spruce (Picea abies) and birch (Betula pendula and Betula pubescens). The said models had to be capable of generating growth scenarios, and thus correlations between series had to be taken into account. ARMA models were estimated for mean growth index series from Pohjois-Karjala, eastern Finland. Several ARMA models, some of which had seasonal parameters, were found to be adequate for each series. Non-seasonal AR(1) and seasonal AR(1,1) models were used to produce growth scenarios in the case study, in which variation in growth was integrated into forest planning.
  • Pasanen, University of Joensuu, Faculty of Forestry, P.O. BOX 111, 80101 Joensuu, Finland E-mail: pasanen@gis.joensuu.fi (email)
article id 696, category Research article
Jukka Lippu. (1998). Redistribution of 14C-labelled reserve carbon in Pinus sylvestris seedlings during shoot elongation. Silva Fennica vol. 32 no. 1 article id 696. https://doi.org/10.14214/sf.696
Keywords: Scots pine; root growth; carbon storage; 14C labelling; photosynthates; shoot growth
Abstract | View details | Full text in PDF | Author Info
This study examined the later use of 14C reserves formed in previous autumn in Scots pine (Pinus sylvestris L.) seedlings. The seedlings were allowed to photosynthesise 14CO2 in early September when shoot and needle growth was over. The following spring the seedlings were harvested in five samplings during the shoot growth period. The distribution and concentration of 14C were determined and the results were compared with the growth data. It was observed that reserves were not used markedly for the new growth. Most of the 14C was found in one-year-old needles (30–40%) and in the root system (40–50%) which was due to both their high activity as a storage sink and their large sink size. The high initial 14C-activity in the finest roots decreased indicating respiration of reserves. Only a small percent of the reserve carbon was found in the new shoots which indicated that reserves are of minor importance in building a new shoot. An allocation of about 15% of the autumn storage to the stem suggested that in seedlings the stem is of minor importance as a storage organ.
  • Lippu, Department of Forest Ecology, P.O. Box 24 (Unioninkatu 40 B), FIN-00014 University of Helsinki, Finland E-mail: jukka.lippu@helsinki.fi (email)

Category : Climate resilient and sustainable forest management – Review article

article id 23076, category Climate resilient and sustainable forest management – Review article
Joanne C. White. (2024). Characterizing forest recovery following stand-replacing disturbances in boreal forests: contributions of optical time series and airborne laser scanning data. Silva Fennica vol. 58 no. 2 article id 23076. https://doi.org/10.14214/sf.23076
Keywords: regeneration; fire; time series; Landsat; harvest; forest change
Highlights: Remote sensing contributions to monitoring of post-disturbance forest recovery in the boreal are synthesized; Definitions of forest recovery need to be clear and measurable and will vary by application; Landsat time series represent a significant innovation in recovery assessments, but the boreal biome is underrepresented in this research; Opportunities for future research directions and priorities are highlighted.
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The success and rate of forest regeneration following disturbance has implications for sustainable forest management, climate change mitigation, and biodiversity, among others. Systematic monitoring of forest regeneration over large and often remote areas of the boreal forest is challenging. The use of remotely sensed data to characterize post-disturbance recovery in the boreal forest has been an active research topic for more than 30 years. Innovations in sensors, data policies, curated data archives, and increased computational power have enabled new insights into the characterization of post-disturbance forest recovery, particularly following stand-replacing disturbances. Landsat time series data have emerged as an important data source for post-disturbance forest recovery assessments, with Landsat’s 40-year archive of 30-m resolution data providing consistent observations on an annual time step and enabling retrospective capacity to establish spatially explicit recovery baselines. The application of remote sensing for monitoring post-disturbance forest recovery is a rapidly growing area of research globally; however, despite the large amount of disturbance and the disproportionate effects of climate change in the boreal, the boreal biome is relatively underrepresented in the remote sensing forest recovery literature. Herein, the past and present contributions of optical time series and airborne laser scanning data to the characterization of forest recovery in boreal forests are highlighted, and future research priorities are identified.

  • White, Canadian Forest Service, Pacific Forestry Centre, Natural Resources Canada, 506 West Burnside Road, Victoria, B.C., V8Z 1M5, Canada ORCID https://orcid.org/0000-0003-4674-0373 E-mail: joanne.white@nrcan-rncan.gc.ca (email)
article id 23057, category Climate resilient and sustainable forest management – Review article
Ane Christensen Tange, Hanne K. Sjølie, Gunnar Austrheim. (2024). Effectiveness of conservation measures to support biodiversity in boreal timber-production forests. Silva Fennica vol. 58 no. 2 article id 23057. https://doi.org/10.14214/sf.23057
Keywords: forestry; sustainable forest management; dead wood enhancement; forest certification; green tree retention; riparian buffer zone
Highlights: A systematic review of in-situ conservation measures displays that forest biodiversity levels are largely maintained upon harvest with conservation measures compared to unlogged forest; The type of control impacts the frequency of positive, not significant and negative observations; The relatively few significant results restrain distinct conclusions on the effectiveness of the assessed conservation measures to support biodiversity.
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Large parts of the boreal forest ecosystems have been greatly affected by human use, and the current timber-oriented forest management practice that dominates boreal forests is proven to cause biodiversity and ecosystem services declines. These negative effects are mitigated in various ways, including in-situ measures implemented upon harvest. The measures comprise trade-offs between economic and ecological aims; thus, requiring solid knowledge of their effectiveness. However, comprehensive literature review of the effectiveness of such measures is scarce. We aim to fill part of this void by reviewing the scientific literature that have gauged effects of four in-situ conservation measures: green tree retention (GTR), patch retention (PR), dead wood retention (DW) and riparian buffer zones (RB). Two outcomes were considered, species richness and species abundance across taxa.

From a total of 3012 initial papers, 48 met our inclusion criteria that generated 238 unique results. Results were grouped according to control. 178 studies used mature, unlogged forest as control. Out of those, 68% of the findings were not significant, i.e., suggesting no significant impact of harvest with biodiversity measures on species richness and species abundance compared to no harvest. Eighteen percent of the observations showed negative effects and 14% of the observations showed positive effects compared to no harvest. Sixty studies used harvest with no measures as control, of which 45% showed significant positive effects, meaning that compared to harvest with no measures, harvest with conservation measures has positively effects on species richness and abundance. However, 43% of the studies found no significant effect of the implemented conservation measures compared to harvest with no measures taken.

The relatively few significant results reported restrain distinct conclusions on the effectiveness of the assessed conservation measures, but some degree of conservation measure is likely to have positive effects on biodiversity in timber-production forest. However, the scientific basis does not allow for pointing to threshold levels. Higher transparency of study design and statistical results would allow us to include more studies. There is a clear need for more research of effectiveness of common conservation measures in timber-production forests in order to strengthen the knowledge basis. In particular, there are few studies that employ harvest without any conservation measure as control. This is pivotal knowledge for forest managers as well as for policymakers for preserving biodiversity and the ecosystems in forest.

  • Tange, Inland Norway University of Applied Sciences, Faculty of Applied Ecology, Agricultural Sciences and Biotechnology, Department of Forestry and Wildlife Management, Evenstad, Norway; Glommen Mjøsen Skog SA, Elverum, Norway ORCID https://orcid.org/0009-0001-3145-8159 E-mail: ane.tange@inn.no (email)
  • Sjølie, Inland Norway University of Applied Sciences, Faculty of Applied Ecology, Agricultural Sciences and Biotechnology, Department of Forestry and Wildlife Management, Evenstad, Norway ORCID https://orcid.org/0000-0001-8099-3521 E-mail: hanne.sjolie@inn.no
  • Austrheim, University Museum Norwegian University of Science and Technology, Department of Natural History, Trondheim, Norway ORCID https://orcid.org/0000-0002-3909-6666 E-mail: gunnar.austrheim@ntnu.no

Category : Review article

article id 24021, category Review article
Eva Ring, Märtha Wallgren, Erland Mårald, Per Westerfelt, Line Djupström, Aron Davidsson, Johan Sonesson. (2024). Forest roads in Sweden – infrastructure with multiple uses and diverse impacts. Silva Fennica vol. 58 no. 4 article id 24021. https://doi.org/10.14214/sf.24021
Keywords: insects; forestry; recreation; reindeer; water; plants; wildlife
Highlights: The extensive network of forest roads in Sweden facilitates both forestry transportation and other activities including recreation, berry-picking, hunting, tourism, and firefighting; The multiple uses of roads can lead to conflicts when the interests between stakeholders and forest owners collide; Despite its extent, research into the social, ecological, and environmental consequences of Sweden’s forest road network is limited.
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Since the 1950s, more than 200 000 km of roads have been built in Sweden’s forests, making them easily accessible and open to multiple uses. The aim of this study was to review the impacts of forest roads in Sweden from a broad perspective encompassing social, ecological, and environmental factors. The Swedish case is interesting because it has an extensive network of permanent forest roads which were built primarily for forestry-related transportation but are currently used by many other stakeholders for many different purposes. Forest roads not only facilitate transportation of wood, machinery, personnel, and equipment into and out of the forest but also enable emergency response to wildfires and support berry and mushroom picking, hunting, recreation, tourism, and access to second homes. The roads increase the opportunities for members of the public to experience forests in various ways. Conflicts arise when different interests collide, for example when the interests of the forest owner clash with those of commercial berry-picking companies, tourism entrepreneurs, or reindeer (Rangifer tarandus L.) herding. Forest roads may have ecological impacts such as barrier and disturbance effects, fragmentation or loss of habitats, altering fauna movement patterns, and changing the composition of plant and insect species. The environmental impacts of forest roads relate to, among other things, hydrology, water quality, and erosion. Predicted changes in the climate are likely to place new demands on Swedish forest roads but, despite their extent, this review shows that there is only a small amount of rather fragmented research on their social, ecological, and environmental consequences. Overall, few studies appear to cover both social and ecological/environmental factors and their interactions, either in Sweden or elsewhere. This review provides examples of such interactions in the case of Sweden, and suggests that more research into these and the specific social, ecological, and environmental factors involved is warranted.

  • Ring, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, 751 83 Uppsala, Sweden ORCID https://orcid.org/0000-0002-8962-9811 E-mail: eva.ring@skogforsk.se (email)
  • Wallgren, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, 751 83 Uppsala, Sweden; Swedish University of Agricultural Sciences, Department of Wildlife, Fish, and Environmental Studies, 907 36 Umeå, Sweden ORCID https://orcid.org/0000-0002-3172-4496 E-mail: martha.wallgren@skogforsk.se
  • Mårald, Umeå University, Department of Historical. Philosophical and Religious Studies, 901 87 Umeå, Sweden ORCID https://orcid.org/0000-0002-2291-9910 E-mail: erland.marald@umu.se
  • Westerfelt, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, 751 83 Uppsala, Sweden ORCID https://orcid.org/0000-0003-2040-8305 E-mail: per.westerfelt@skogforsk.se
  • Djupström, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, 751 83 Uppsala, Sweden; Swedish University of Agricultural Sciences, Department of Wildlife, Fish, and Environmental Studies, 907 36 Umeå, Sweden ORCID https://orcid.org/0000-0003-4536-7765 E-mail: line.djupstrom@skogforsk.se
  • Davidsson, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, 751 83 Uppsala, Sweden ORCID https://orcid.org/0009-0007-2736-8656 E-mail: Aron.davidsson@skogforsk.se
  • Sonesson, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, 751 83 Uppsala, Sweden ORCID https://orcid.org/0000-0002-2018-7496 E-mail: Johan.sonesson@skogforsk.se
article id 23056, category Review article
Eva Ring, Ulf Sikström. (2024). Environmental impact of mechanical site preparation on mineral soils in Sweden and Finland — a review. Silva Fennica vol. 58 no. 1 article id 23056. https://doi.org/10.14214/sf.23056
Keywords: carbon; nitrogen; soil disturbance; vegetation; water; chemistry; greenhouse gas
Highlights: Mechanical site preparation is carried out on large areas, but limited research on its environmental impact has been undertaken; It affects nitrogen and carbon cycling over the first few years, and has a minor initial impact on CO2, CH4 and N2O fluxes; It increases tree carbon stores and possibly ecosystem carbon stores; Reducing its soil disturbance intensity is warranted.
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Mechanical site preparation (MSP) is deliberate soil disturbance which is undertaken to improve the conditions for forest regeneration. Disc trenching and mounding are the dominant MSP practices currently used in Sweden and Finland. In this paper, the impacts of MSP on the soil, water quality, greenhouse gas (GHG) emissions and ground vegetation of mineral soil sites in Sweden and Finland are reviewed. The practices considered are patch scarification, mounding, inverting, disc trenching, and ploughing, which together represent a wide range of soil disturbance intensity. The environmental effects of MSP in this region have not been studied extensively. The environmental impact of MSP derives from the process of creating microsites which involves horizontal and/or vertical redistribution of soil and soil mixing. This typically affects decomposition, element circulation and leaching, vegetation coverage and uptake of nutrients and water, and possibly erosion and sediment exports. Following disc trenching or mounding the effects on GHG emissions appear to be minor over the first two years. For a few years after disc trenching concentrations in soil water collected below ridges are higher than that below furrows for some elements (e.g., NO3-, NH4+, Mg2+, and total or dissolved organic C). The physical and chemical effects of ploughing remain detectable for several decades. There is little evidence about how the effects of forestry activities in upland areas on soil-water chemistry are transferred to adjacent surface water bodies, including what role streamside discharge areas play. MSP increases the tree biomass C store and may increase the total ecosystem C store. The impact of MSP on the cover and abundance of ground vegetation species depends on the composition of the original plant community, MSP intensity, and the establishment rate of different species. Species cover generally seems to decline for late succession understory species, while pioneer and ruderal species can benefit from the microsites created. Areas containing lichens which are used for reindeer forage require special consideration. More research is needed on the environmental effects of MSP, particularly regarding its long-term effects. Further efforts should be made to develop efficient site-preparation practices which better balance the disturbance intensity with what is needed for successful regeneration.

  • Ring, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, 751 83, Uppsala, Sweden ORCID https://orcid.org/0000-0002-8962-9811 E-mail: eva.ring@skogforsk.se (email)
  • Sikström, Skogforsk (The Forestry Research Institute of Sweden), Uppsala Science Park, 751 83, Uppsala, Sweden E-mail: ulf.sikstrom@skogforsk.se
article id 22018, category Review article
Felipe de Miguel-Díez, Thomas Purfürst, Mauricio Acuna, Eduardo Tolosana-Esteban, Tobias Cremer. (2023). Estimation of conversion factors for wood stacks in landings and their influencing parameters: a comprehensive literature review for America and Europe. Silva Fennica vol. 57 no. 1 article id 22018. https://doi.org/10.14214/sf.22018
Keywords: log properties; roundwood measurement; roundwood purchase; stack properties; wood supply chain
Highlights: Midpoint diameter, stacking quality, crookedness, log length, delimbing quality and bark thickness strongly influence the conversion factors; In Europe and America, a total of 48 methods were developed to estimate the conversion factors of wood stacks; The conversion factors should be estimated for each wood assortment and tree species.
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In commercial transaction of stacked roundwood, the estimation of the stack net volume plays a key role. One generalized method to determine the net volume is using conversion factors that relate the gross and net volumes. In this literature review the developed methods to estimate the conversion factors as well as their influencing parameters were analyzed based on 153 references from America and Europe. According to the results, 48 different methods (including their variants) for estimating the conversion factors were developed. The newest methods enabled their accurate determination inexpensively, e.g., photo-optical methods or 3D simulation models. The analyzed references revealed that 30 parameters influence the conversion factors. Based on this comprehensive review, each stakeholder involved in the roundwood supply chain can know which method is used for estimating the conversion factors in the analyzed territories and which influencing parameters should be considered when purchasing roundwood in order to accurately assess the solid wood content in the stacks.

  • de Miguel-Díez, Eberswalde University for Sustainable Development, Department of Forest Utilization and Timber Markets, Eberswalde, Germany; University of Freiburg, Chair of Forest Operations, Freiburg, Germany ORCID https://orcid.org/0000-0002-3800-7449 E-mail: felipe.diez@hnee.de (email)
  • Purfürst, University of Freiburg, Chair of Forest Operations, Freiburg, Germany ORCID https://orcid.org/0000-0001-9661-0193 E-mail: thomas.purfuerst@foresteng.uni-freiburg.de
  • Acuna, University of the Sunshine Coast, Forest Research Institute, Sunshine Coast, Queensland, Australia E-mail: macuna@usc.edu.au
  • Tolosana-Esteban, Universidad Politécnica de Madrid, E.T.S.I. Montes, Forestal y del Medio Natural, Madrid, Spain ORCID https://orcid.org/0000-0003-2561-0342 E-mail: eduardo.tolosana@upm.es
  • Cremer, Eberswalde University for Sustainable Development, Department of Forest Utilization and Timber Markets, Eberswalde, Germany ORCID https://orcid.org/0000-0001-7866-944X E-mail: Tobias.Cremer@hnee.de
article id 10709, category Review article
Pavel Škvor, Martin Jankovský, Pavel Natov, Jiří Dvořák. (2023). Evaluation of stress loading for logging truck drivers by monitoring changes in muscle tension during a work shift. Silva Fennica vol. 57 no. 1 article id 10709. https://doi.org/10.14214/sf.10709
Keywords: timber transportation; biofeedback; driver; electromyography; physical load
Highlights: This study compared the workload of drivers when driving different types of vehicles; The combination of a truck and trailer exposes the driver to a significantly greater workload than trucks without trailers; The workload of drivers varied during a work shift i.e. drivers experienced significantly more workload in the middle of their work shifts.
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Our research aimed to quantify and evaluate the stress loading of drivers by monitoring the loading of the radial extensor muscle at the wrist joint (musculus extensor carpi radialis) when they drove different types of timber trucks. We monitored changes in the electric potential of skeletal muscles with electromyographic measurements and measurements of changes of heart rate using the Biofeedback 2000 x-pert device. The drivers were observed throughout their work shifts during normal operation of logging trucks and logging trucks with trailers. As a reference, muscle load was measured when driving a passenger car. We evaluated the normality of the measured data and obtained descriptive statistics from the individual measurements. The differences in stress load associated with driving the different types of vehicles increased whilst driving on lower-class roads. Results showed a high stress load for drivers of loaded vehicles when driving on narrow roads. It was more challenging to control a loaded logging truck with a trailer than driving a logging truck, with the difference in muscular loading reaching 22.5%. Driving a logging truck with a trailer produced 46.5% more muscle loading compared to driving a loaded passenger car. For preventive health and safety reasons, it would be reasonable to alternate between drivers when operating various vehicles, thus minimizing the development of possible health issues.

  • Škvor, Department of Forestry Technologies and Construction, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Kamýcká 129, 165 21 Prague 6 - Suchdol, Czech Republic ORCID https://orcid.org/0000-0001-7540-4761 E-mail: skvorp@fld.czu.cz (email)
  • Jankovský, Department of Forestry Technologies and Construction, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Kamýcká 129, 165 21 Prague 6 - Suchdol, Czech Republic E-mail: jankovskym@fld.czu.cz
  • Natov, Department of Forestry Technologies and Construction, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Kamýcká 129, 165 21 Prague 6 - Suchdol, Czech Republic E-mail: natov@fld.czu.cz
  • Dvořák, Department of Forestry Technologies and Construction, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague, Kamýcká 129, 165 21 Prague 6 - Suchdol, Czech Republic ORCID https://orcid.org/0000-0002-5986-8002 E-mail: dvorakj@fld.czu.cz
article id 10609, category Review article
Jaakko Jussila, Emil Nagy, Katja Lähtinen, Elias Hurmekoski, Liina Häyrinen, Cecilia Mark-Herbert, Anders Roos, Ritva Toivonen, Anne Toppinen. (2022). Wooden multi-storey construction market development – systematic literature review within a global scope with insights on the Nordic region. Silva Fennica vol. 56 no. 1 article id 10609. https://doi.org/10.14214/sf.10609
Keywords: sustainability; construction industry; consumer; forest-wood value chain; municipality; urbanization
Highlights: Enabling factors for WMC market diffusion include benefits from cost-efficiency gains from prefabrication and industrialization and perceived sustainability benefits; Inexperience of using wood, and path dependencies to use concrete and steel in multi-storey building are the key barriers for mainstreaming WMC market development; More research is needed on the development in the wood construction value-chains to challenge the dominant concrete-based construction regime in the housing markets.
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Climate change sets high pressures on the construction industry to decrease greenhouse gas emissions. Due to the carbon storage properties and potential to use renewable resources efficiently, wooden multi-storey construction (WMC) is an interesting alternative for the construction industry to enhance sustainable development combined with the aesthetic and well-being benefits of wood perceived among many consumers. For forest industry firms, industrial wood construction is a possibility to seek for business opportunities and bring socio-economic benefits for local economies. Despite positive drivers, WMC still remains a niche even in the forest-rich countries.The purpose of our study is to add understanding on the WMC market development by conducting a systematic literature analysis on international peer-reviewed studies from the past 20 years. Our special focus is on the role of WMC in the housing markets studied from the perspectives of the demand, supply and local governance factors. As specific aims, we 1) synthesize the key barriers and enabling factors for the WMC market growth; 2) identify the actors addressed in the existing studies connected to the WMC market development, and 3) summarize research methods and analytical approaches used in the previous studies. As a systematic method to make literature searches in Web of Science and Scopus for years 2000–2020, we employed PRISMA guidelines. By using pre-determined keywords, our searches resulted in a sample of 696 articles, of which 42 full articles were after selection procedure included in-depth content analysis. Our results showed cost-efficiency gains from industrialized prefabrication and perceived sustainability benefits by consumers and architects enabled a WMC market diffusion. The lack of experiences on the WMC, and path dependencies to use concrete and steel continue to be key barriers for increased WMC. Although our research scope was the global WMC market development, most of the literature concerned the Nordic region. The key actors covered in the literature were businesses (e.g., contractors, manufacturers and architects) involved in the wood construction value-chains, while residents and actors in the local governance were seldomly addressed. Currently, case studies, the use of qualitative data sets and focus on the Nordic region dominate the literature. This hinders the generalizability of findings in different regional contexts. In the future, more research is needed on how sustainability-driven wood construction value-chains are successfully shaping up in different geographical regions, and how they could challenge the dominant concrete-based construction regime.

  • Jussila, University of Helsinki, Department of Forest Sciences, P.O. Box 4, FI-00014 University of Helsinki, Finland E-mail: jaakko@jussila.fi (email)
  • Nagy, Swedish University of Agricultural Sciences, Department of Forest Economics, P.O. Box 7060, SE-750 07 Uppsala, Sweden E-mail: emil.nagy@slu.se
  • Lähtinen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: katja.lahtinen@luke.fi
  • Hurmekoski, University of Helsinki, Department of Forest Sciences, P.O. Box 4, FI-00014 University of Helsinki, Finland E-mail: elias.hurmekoski@helsinki.fi
  • Häyrinen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: liina.hayrinen@luke.fi
  • Mark-Herbert, Swedish University of Agricultural Sciences, Department of Forest Economics, P.O. Box 7060, SE-750 07 Uppsala, Sweden E-mail: cecilia.mark-herbert@slu.se
  • Roos, Swedish University of Agricultural Sciences, Department of Forest Economics, P.O. Box 7060, SE-750 07 Uppsala, Sweden E-mail: anders.roos@slu.se
  • Toivonen, University of Helsinki, Department of Forest Sciences, P.O. Box 4, FI-00014 University of Helsinki, Finland E-mail: ritva.toivonen@helsinki.fi
  • Toppinen, University of Helsinki, Department of Forest Sciences, P.O. Box 4, FI-00014 University of Helsinki, Finland E-mail: anne.toppinen@helsinki.fi
article id 10516, category Review article
Faujiah N. Ritonga, Jacob N. Ngatia, Run X. Song, Umar Farooq, Sonia Somadona, Andi T. Lestari, Su Chen. (2021). Abiotic stresses induced physiological, biochemical, and molecular changes in Betula platyphylla: a review. Silva Fennica vol. 55 no. 3 article id 10516. https://doi.org/10.14214/sf.10516
Keywords: antioxidant; cis-acting elements; gene; ROS production; transcription factor
Highlights: Abiotic stress influence Betula platyphylla growth, development, and yield production by impairing physiological, biochemical, and molecular functions; Overexpression or RNAi line of transcription factors enhance the abiotic stress tolerance of B. platyphylla; MYB and AP2/ERF are the most frequently transcription factor family that has been explored over the last two decades in B. platyphylla under abiotic stress.
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Abiotic stress is one of the major factors in reducing plant growth, development, and yield production by interfering with various physiological, biochemical, and molecular functions. In particular, abiotic stress such as salt, low temperature, heat, drought, UV-radiation, elevated CO2, ozone, and heavy metals stress is the most frequent study in Betula platyphylla Sukaczev. Betula platyphylla is one of the most valuable tree species in East Asia facing abiotic stress during its life cycle. Using transgenic plants is a powerful tool to increase the B. platyphylla abiotic stress tolerance. Generally, abiotic stress reduces leaves water content, plant height, fresh and dry weight, and enhances shed leaves as well. In the physiological aspect, salt, heavy metal, and osmotic stress disturbs seed germination, stomatal conductance, chlorophyll content, and photosynthesis. In the biochemical aspect, salt, drought, cold, heat, osmotic, UV-B radiation, and heavy metal stress increases the ROS production of B. platyphylla cells, resulting in the enhancement of enzymatic antioxidant (SOD and POD) and non-enzymatic antioxidant (proline and AsA) to reduce the ROS accumulation. Meanwhile, B. platyphylla upregulates various genes, as well as proteins to participate in abiotic stress tolerance. Based on recent studies, several transcription factors contribute to increasing abiotic stress tolerance in B. platyphylla, including BplMYB46, BpMYB102, BpERF13, BpERF2, BpHOX2, BpHMG6, BpHSP9, BpUVR8, BpBZR1, BplERD15, and BpNACs. These transcription factors bind to different cis-acting elements to upregulate abiotic stress-related genes, resulting in the enhancement of salt, drought, cold, heat, osmotic, UV-B radiation, and heavy metal tolerance. These genes along with phytohormones mitigate the abiotic stress. This review also highlights the candidate genes from another Betulacea family member that might be contributing to increasing B. platyphylla abiotic stress tolerance.

  • Ritonga, State Key Laboratory of Tree Genetics and Breeding, Forestry College, Northeast Forestry University, Harbin 150040, China E-mail: ritongafaujiah@ymail.com
  • Ngatia, College of Wildlife and Protected Areas, Northeast Forestry University, Harbin 150040, China E-mail: jacob.ngatia3@gmail.com
  • Song, State Key Laboratory of Tree Genetics and Breeding, Forestry College, Northeast Forestry University, Harbin 150040, China E-mail: 13359850710@163.com
  • Farooq, College of Life Science, Northeast Forestry University, Harbin 150040, China E-mail: uf@nn.ch
  • Somadona, College of Agriculture, Riau University, Pekanbaru 28293, Indonesia E-mail: sonia_hut@yahoo.co.id
  • Lestari, Forestry Major, College of Agriculture, Mataram University, Mataram 83125, Indonesia E-mail: atlestari@unram.ac.id
  • Chen, State Key Laboratory of Tree Genetics and Breeding, Forestry College, Northeast Forestry University, Harbin 150040, China E-mail: chensu@nefu.edu.cn (email)
article id 10381, category Review article
Carl F. Salk. (2020). Interpreting common garden studies to understand cueing mechanisms of spring leafing phenology in temperate and boreal tree species. Silva Fennica vol. 54 no. 5 article id 10381. https://doi.org/10.14214/sf.10381
Keywords: photoperiod; bud break; budburst; chilling; elevation gradients; latitudinal gradients; leaf flush; reciprocal transplant experiments; xylem anatomy
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Trees are particularly susceptible to climate change due to their long lives and slow dispersal. However, trees can adjust the timing of their growing season in response to weather conditions without evolutionary change or long-distance migration. This makes understanding phenological cueing mechanisms a critical task to forecast climate change impacts on forests. Because of slow data accumulation, unconventional and repurposed information is valuable in the study of phenology. Here, I develop and use a framework to interpret what phenological patterns among provenances of a species in a common garden reveal about their leafing cues, and potential climate change responses. Species whose high elevation/latitude provenances leaf first likely have little chilling requirement, or for latitude gradients only, a critical photoperiod cue met relatively early in the season. Species with low latitude/elevation origins leafing first have stronger controls against premature leafing; I argue that these species are likely less phenologically flexible in responding to climate change. Among published studies, the low to high order is predominant among frost-sensitive ring-porous species. Narrow-xylemed species show nearly all possible patterns, sometimes with strong contrasts even within genera for both conifers and angiosperms. Some also show complex patterns, indicating multiple mechanisms at work, and a few are largely undifferentiated across broad latitude gradients, suggesting phenotypic plasticity to a warmer climate. These results provide valuable evidence on which temperate and boreal tree species are most likely to adjust in place to climate change, and provide a framework for interpreting historic or newly-planted common garden studies of phenology.

  • Salk, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, P.O. Box 49, SE- 230 53 Alnarp, Sweden; Faculty of International Studies, Utsunomiya University, 350 Minemachi, Utsunomiya-shi, Tochigi 321-8505 Japan; Institute for Globally Distributed Open Research and Education (IGDORE) E-mail: carl.salk@slu.se (email)
article id 10172, category Review article
Ulf Sikström, Karin Hjelm, Kjersti Holt Hanssen, Timo Saksa, Kristina Wallertz. (2020). Influence of mechanical site preparation on regeneration success of planted conifers in clearcuts in Fennoscandia – a review. Silva Fennica vol. 54 no. 2 article id 10172. https://doi.org/10.14214/sf.10172
Keywords: natural regeneration; regeneration chain; seedling growth; coniferous seedlings; disturbed soil surface; seedling survival
Highlights: Mechanical site preparation (MSP) increases seedling survival rates by 15–20%; Survival rates of 80–90% ca. 10 years after MSP and planting conifers are possible; MSP can increase tree height 10–15 years after planting by 10–25%; The increase in growth rate associated with MSP may be temporary, but the height enhancement probably persists.
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In the Nordic countries Finland, Norway and Sweden, the most common regeneration method is planting after clearcutting and, often, mechanical site preparation (MSP). The main focus of this study is to review quantitative effects that have been reported for the five main MSP methods in terms of survival and growth of manually planted coniferous seedlings of Norway spruce (Picea abies (L.) Karst.), Scots pine (Pinus sylvestris L.) and lodgepole pine (Pinus contorta var. latifolia Engelm.) in clearcuts in these three countries. Meta analyses are used to compare the effects of MSP methods to control areas where there was no MSP and identify any relationships with temperature sum and number of years after planting. In addition, the area of disturbed soil surface and the emergence of naturally regenerated seedlings are evaluated. The MSP methods considered are patch scarification, disc trenching, mounding, soil inversion and ploughing. Studies performed at sites with predominately mineral soils (with an organic topsoil no thicker than 0.30 m), in boreal, nemo-boreal and nemoral vegetation zones in the three Fenno-Scandinavian countries are included in the review. Data from 26 experimental and five survey studies in total were compiled and evaluated. The results show that survival rates of planted conifers at sites where seedlings are not strongly affected by pine weevil (Hylobius abietis L.) are generally 80–90% after MSP, and 15–20 percent units higher than after planting in non-prepared sites. The experimental data indicated that soil inversion and potentially ploughing (few studies) give marginally greater rates than the other methods in this respect. The effects of MSP on survival seem to be independent of the temperature sum. Below 800 degree days, however, the reported survival rates are more variable. MSP generally results in trees 10–25% taller 10–15 years after planting compared to no MSP. The strength of the growth effect appears to be inversely related to the temperature sum. The compiled data may assist in the design, evaluation and comparison of possible regeneration chains, i.e. analyses of the efficiency and cost-effectiveness of multiple combinations of reforestation measures.

  • Sikström, Skogforsk, Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: ulf.sikstrom@skogforsk.se
  • Hjelm, Skogforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden E-mail: karin.hjelm@skogforsk.se (email)
  • Holt Hanssen, Norwegian Institute of Bioeconomy Research (NIBIO), P.O. Box 115, NO-1431 Ås, Norway E-mail: kjersti.hanssen@nibio.no
  • Saksa, Natural Resources Institute Finland (Luke), Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@luke.fi
  • Wallertz, Swedish University of Agricultural Sciences (SLU), Asa Forest Research Station, SE-360 30 Lammhult, Sweden E-mail: kristina.wallertz@slu.se
article id 9984, category Review article
Christoph Kogler, Peter Rauch. (2018). Discrete event simulation of multimodal and unimodal transportation in the wood supply chain: a literature review. Silva Fennica vol. 52 no. 4 article id 9984. https://doi.org/10.14214/sf.9984
Keywords: logistics; supply chain management; forest products industry; decision support systems; validation and verification of simulation models; resilient risk management
Highlights: Focus on discrete event simulation, wood supply chain and multimodal transport; Analyses of 12 review articles and a core of 32 research papers, complemented by 48 related ones; Research focus from unimodal to multimodal transport to build efficient, resilient, green and socially sustainable supply chains; Development of robust risk management considering supply risks, demand risks and external risks is needed.
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This review systematically analyses and classifies research and review papers focusing on discrete event simulation applied to wood transport, and therefore illustrates the development of the research area from 1997 until 2017. Discrete event simulation allows complex supply chain models to be mapped in a straightforward manner to study supply chain dynamics, test alternative strategies, communicate findings and facilitate understanding of various stakeholders. The presented analyses confirm that discrete event simulation is well-suited for analyzing interconnected wood supply chain transportation issues on an operational and tactical level. Transport is the connective link between interrelated system components of the forest products industry. Therefore, a survey on transport logistics allows to analyze the significance of entire supply chain management considerations to improve the overall performance and not only one part in isolation. Thus far, research focuses mainly on biomass, unimodal truck transport and terminal operations. Common shortcomings identified include rough explanations of simulation models and sparse details provided about the verification and validation processes. Research gaps exist concerning simulations of entire, resilient and multimodal wood supply chains as well as supply and demand risks. Further studies should expand upon the few initial attempts to combine various simulation methods with optimization.

  • Kogler, Institute of Production and Logistics, Department of Economics and Social Sciences, University of Natural Resources and Life Sciences, Vienna, Feistmantelstrasse 4, A-1180 Vienna, Austria ORCID https://orcid.org/0000-0001-8811-152X E-mail: christoph.kogler@boku.ac.at (email)
  • Rauch, Institute of Production and Logistics, Department of Economics and Social Sciences, University of Natural Resources and Life Sciences, Vienna, Feistmantelstrasse 4, A-1180 Vienna, Austria ORCID http://orcid.org/0000-0002-5812-4415 E-mail: peter.rauch@boku.ac.at
article id 7760, category Review article
Maria A. Huka, Manfred Gronalt. (2018). Log yard logistics. Silva Fennica vol. 52 no. 4 article id 7760. https://doi.org/10.14214/sf.7760
Keywords: log yard; tactical structural and operational problems; material flow; facility layout planning; storage assignment; vehicle movement; seasonality
Highlights: Characteristics of log yard logistics; Classification into tactical structural and operational problems in the wood industry; Different solution methods such as optimisation, heuristics and simulations and their possible application within the log yard with an overview of existing literature which includes several different case studies with varying emphases, problem analysis and solution methods.
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For sawmills, paper mills, particleboard, oriented strand board (OSB), fiberboard and other wood production factories, the log yard is the first step, where raw materials are sorted and stored before production begins. Due to the size of these production sites great potential exists for the optimisation of internal logistics. In this paper the different planning problems of the log yard are introduced and existing literature examined. Beginning with the tactical problems of structure, such as assessing material flow, planning facility layout and assigning storage areas, it continues with operational problems such as vehicle movement planning within the log yard, empty trip minimisation and the seasonality of raw material availability. Data derived from this study reveals a variety of possible solution methods, the applicability of which depends on the precise nature of the log yard operations. Additionally, several real life examples are provided which illustrate the potential for operational improvement.

  • Huka, University of Natural Resources and Life Sciences, Institute of Production and Logistics, Feistmantelstraße 4, 1180 Vienna, Austria E-mail: maria.huka@boku.ac.at (email)
  • Gronalt, University of Natural Resources and Life Sciences, Institute of Production and Logistics, Feistmantelstraße 4, 1180 Vienna, Austria E-mail: manfred.gronalt@boku.ac.at
article id 1673, category Review article
Eshetu Yirdaw, Mulualem Tigabu, Adrian Monge. (2017). Rehabilitation of degraded dryland ecosystems – review. Silva Fennica vol. 51 no. 1B article id 1673. https://doi.org/10.14214/sf.1673
Keywords: restoration; desertification; land degradation; rangelands; croplands; dry forests; landscapes
Highlights: The prospect of restoring degraded drylands is technically promising; The forest landscape restoration concept can be used as the overarching rehabilitation framework; Development of process-based models that forecast rehabilitation outcomes is needed; Rehabilitation methodologies developed for moist areas are not necessarily suitable for drylands; More data is needed on cost-benefit analysis of rehabilitation interventions.
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Land degradation is widespread and a serious threat affecting the livelihoods of 1.5 billion people worldwide of which one sixth or 250 million people reside in drylands. Globally, it is estimated that 10–20% of drylands are already degraded and about 12 million ha are degraded each year. Driven by unsustainable land use practices, adverse climatic conditions and population increase, land degradation has led to decline in provision of ecosystem services, food insecurity, social and political instability and reduction in the ecosystem’s resilience to natural climate variability. Several global initiatives have been launched to combat land degradation, including rehabilitation of degraded drylands. This review aimed at collating the current state-of-knowledge about rehabilitation of degraded drylands. It was found that the prospect of restoring degraded drylands is technically promising using a suite of passive (e.g. area exclosure, assisted natural regeneration, rotational grazing) and active (e.g. mixed-species planting, framework species, maximum diversity, and use of nurse tree) rehabilitation measures. Advances in soil reclamation using biological, chemical and physical measures have been made. Despite technical advances, the scale of rehabilitation intervention is small and lacks holistic approach. Development of process-based models that forecast outcomes of the various rehabilitation activities will be useful tools for researchers and practitioners. The concept of forest landscape restoration approach, which operates at landscape-level, could also be adopted as the overarching framework for rehabilitation of degraded dryland ecosystems. The review identified a data gap in cost-benefit analysis of rehabilitation interventions. However, the cost of rehabilitation and sustainable management of drylands is opined to be lower than the losses that accrue from inaction, depending on the degree of degradation. Thus, local communities’ participation, incorporation of traditional ecological knowledge, clear division of tasks and benefits, strengthening local institutions are crucial not only for cost-sharing, but also for the long-term success of rehabilitation activities.

  • Yirdaw, Viikki Tropical Resources Institute (VITRI), Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: eshetu.yirdaw@helsinki.fi (email)
  • Tigabu, Sveriges Lantbruks Universitet (SLU), Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, P.O. Box 49, SE-230 53, Alnarp, Sweden E-mail: Mulualem.Tigabu@slu.se
  • Monge, Viikki Tropical Resources Institute (VITRI), Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: adrian.mongemonge@helsinki.fi
article id 1650, category Review article
Uriel Safriel. (2017). Land Degradation Neutrality (LDN) in drylands and beyond – where has it come from and where does it go. Silva Fennica vol. 51 no. 1B article id 1650. https://doi.org/10.14214/sf.1650
Keywords: climate change; desertification; ecosystem services; offsetting mechanism; Rio Conventions
Highlights: LDN, a mechanism for offsetting new losses of land’s productivity by restoring productivity of already degraded lands, would maintain the balance of productive lands; As target of Sustainable Development Goal LDN highlights the significance of land whose biological productivity is critical to human survival; Commissioning UNCCD to oversee the implementation of LDN empowers the UNCCD and its impact on sustainability.
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The paper first reviews the desertification/land degradation syndrome, the shortcomings of attempts to control it and the consequences of this failure, including to climate change and biodiversity. It then examines the experience gained by carbon and biodiversity offsets that helped adapting the offsetting principle to the context of land degradation, by emphasizing the restoration of the many already degraded lands on earth, as major component of the Land Degradation Neutrality (LDN) mechanism. LDN is a new voluntary and aspirational target of a Sustainable Development Goal (SDG) under the UN 2030 Agenda for Sustainable Development, aimed at neutralizing the rate of lands coming under degrading use of their productivity. This by balancing the ongoing added degradation with similar rate of restoring equivalent lands whose productivity had been already degraded. If extensively implemented, LDN would stabilize the global amount of productive land by 2030. This would increase global food security and reduce poverty of land users, thus contributing to global sustainability. This review maintains that the failure of United Nations Convention to Combat Desertification (UNCCD) to reduce desertification triggered the emergence of LDN as a mechanism for addressing land degradation globally, rather than just desertification in the drylands. LDN accepted as target of a Sustainable Development Goal also legitimized UNCCD to lead and oversee the aspired process of achieving land degradation neutral world. This paper reviews the development of the LDN concept expressed in scientific deliberations and political advocacy, throughout the five years from inception in 2011 at the UNCCD Secretariat, to early 2016. It notes the fast and increasing acceptance of LDN, expressed in the initiation of implementation already in April 2015 by an increasing number of countries, and in the growing interest and engagement of scientists and policy-makers. But the paper also express concern regarding potential misuse of the concept.

  • Safriel, Hebrew University of Jerusalem, Givat Ram, Jerusalem 9190401, Israel E-mail: uriel36@gmail.com (email)
article id 1660, category Review article
Lars Rytter, Morten Ingerslev, Antti Kilpeläinen, Piritta Torssonen, Dagnija Lazdina, Magnus Löf, Palle Madsen, Peeter Muiste, Lars-Göran Stener. (2016). Increased forest biomass production in the Nordic and Baltic countries – a review on current and future opportunities. Silva Fennica vol. 50 no. 5 article id 1660. https://doi.org/10.14214/sf.1660
Keywords: fertilization; tree breeding; tree species; coppice; cultivation areas; growth increment; nurse crops
Highlights: Annual growth is 287 million m3 in the forests of the Nordic and Baltic countries; Growth can be increased by new tree species, tree breeding, high-productive management systems, fertilization and afforestation of abandoned agricultural land; We predict a forest growth increment of 50–100% is possible at the stand scale; 65% of annual growth is harvested today.
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The Nordic and Baltic countries are in the frontline of replacing fossil fuel with renewables. An important question is how forest management of the productive parts of this region can support a sustainable development of our societies in reaching low or carbon neutral conditions by 2050. This may involve a 70% increased consumption of biomass and waste to meet the goals. The present review concludes that a 50–100% increase of forest growth at the stand scale, relative to today’s common level of forest productivity, is a realistic estimate within a stand rotation (~70 years). Change of tree species, including the use of non-native species, tree breeding, introduction of high-productive systems with the opportunity to use nurse crops, fertilization and afforestation are powerful elements in an implementation and utilization of the potential. The productive forests of the Nordic and Baltic countries cover in total 63 million hectares, which corresponds to an average 51% land cover. The annual growth is 287 million m3 and the annual average harvest is 189 million m3 (65% of the growth). A short-term increase of wood-based bioenergy by utilizing more of the growth is estimated to be between 236 and 416 TWh depending on legislative and operational restrictions. Balanced priorities of forest functions and management aims such as nature conservation, biodiversity, recreation, game management, ground water protection etc. all need consideration. We believe that these aims may be combined at the landscape level in ways that do not conflict with the goals of reaching higher forest productivity and biomass production.

  • Rytter, The Forestry Research Institute of Sweden (Skogforsk), Ekebo 2250, SE-26890 Svalöv, Sweden E-mail: lars.rytter@skogforsk.se (email)
  • Ingerslev, Copenhagen University, Department of Geosciences and Natural Resource Management, Rolighedsvej 23, DK-1958, Frederiksberg C, Denmark E-mail: moi@ign.ku.dk
  • Kilpeläinen, Finnish Environment Institute, Joensuu Office, P.O. Box 111, FI-80101 Joensuu, Finland; University of Eastern Finland, Faculty of Science and Forestry, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: antti.kilpelainen@ymparisto.fi
  • Torssonen, University of Eastern Finland, Faculty of Science and Forestry, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: Piritta.Torssonen@uef.fi
  • Lazdina, Latvian State Forest Research Institute “Silava”, 111 Riga str, Salaspils, LV 2169 Latvia E-mail: Dagnija.Lazdina@silava.lv
  • Löf, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, Box 49 SE-230 53 Alnarp, Sweden E-mail: magnus.lof@slu.se
  • Madsen, Copenhagen University, Department of Geosciences and Natural Resource Management, Rolighedsvej 23, DK-1958, Frederiksberg C, Denmark E-mail: pam@ign.ku.dk
  • Muiste, Estonian University of Life Sciences, Institute of Forestry and Rural Engineering, Dept. Forest Industry, Kreutzwaldi 5, Tartu 51014, Estonia E-mail: Peeter.Muiste@emu.ee
  • Stener, The Forestry Research Institute of Sweden (Skogforsk), Ekebo 2250, SE-26890 Svalöv, Sweden E-mail: Lars-Goran.Stener@skogforsk.se
article id 1416, category Review article
Ulf Sikström, Hannu Hökkä. (2015). Interactions between soil water conditions and forest stands in boreal forests with implications for ditch network maintenance. Silva Fennica vol. 50 no. 1 article id 1416. https://doi.org/10.14214/sf.1416
Keywords: peatland; forest drainage; conifers; tree growth; drainage intensity; ditch deterioration; forest water use; ground-water table; profitability
Highlights: Ditch network maintenance (DNM) may influence soil water conditions less than initial ditching due to reduced hydraulic conductivity of the peat; Stand stocking and management substantially influence soil-water conditions; DNM can lower the GWL and increase tree growth; DNM growth responses of 0.5–1.8 m3 ha–1 yr–1 during 15–20-years in Scots pine peatland stands reported; Greatest need for DNM in the early phase of a stand rotation; Need for better understanding of the link between soil water and tree growth.
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At sites with either peat or mineral soils in large areas of boreal forests, high soil-water contents hamper tree growth and drainage can significantly increase growth. Hence, areas covering about 15 × 106 ha of northern peatlands and wet mineral soils have been drained for forestry purposes. Usually ditches gradually deteriorate, thus reducing their functionality as drains, and ditch-network maintenance (DNM) might be needed to maintain stand growth rates enabled by the original ditching. This article reviews current knowledge on establishing the need for DNM in boreal forest stands, subsequent growth responses, and the financial outcome of the activity. The issues covered in the review are: (i) ditching, changes in ditches over time and the need for DNM; (ii) interactions between soil water and both stand properties and stand management; (iii) ground-water level (GWL) and tree growth responses to DNM; and (iv) financial viability of DNM. Conclusions about the current understanding of issues related to DNM are drawn and implications for DNM in practice are summarized. Finally, gaps in knowledge are identified and research needs are suggested.

  • Sikström, The Forestry Research Institute of Sweden (Skogforsk), Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: ulf.sikstrom@skogforsk.se (email)
  • Hökkä, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: hannu.hokka@luke.fi
article id 1095, category Review article
Jonas Fridman, Sören Holm, Mats Nilsson, Per Nilsson, Anna Hedström Ringvall, Göran Ståhl. (2014). Adapting National Forest Inventories to changing requirements – the case of the Swedish National Forest Inventory at the turn of the 20th century. Silva Fennica vol. 48 no. 3 article id 1095. https://doi.org/10.14214/sf.1095
Keywords: remote sensing; sampling; sample plot inventory; forest state; ENFIN; harmonization
Highlights: National Forest Inventories supply invaluable long term time series of forest state. Recent developments and international harmonization of modern NFIs widen the scope to even include ecosystem goods, e.g. biodiversity and carbon sequestration. The combination of NFI field data with remote sensing techniques can give good estimates for areas smaller than national and regional level.
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National Forest Inventories (NFIs) are becoming increasingly important worldwide in order to provide information about the multiple functions of forests, e.g. their provision of raw materials to industry, biodiversity and their capacity to store carbon for mitigating climate change. In several countries the history of NFIs is very long. For these countries a specific challenge is to keep the inventories up-to-date without sacrificing the advantages associated with long time series. At the turn of the 20th century European NFIs faced some major challenges. In this article we describe the history and the recent developments of the Swedish NFI as an example from which general observations are made and discussed. The Swedish NFI started in 1923 and has evolved from an inventory with a narrow focus on wood resources to an inventory today which aims to provide information about all major forest ecosystem services. It can be concluded that the traditional approaches of most European NFIs, e.g. to collect data through sample plot field inventories, has proved to be applicable even for a wide range of new information requirements. Specifically, detailed data about land use, trees, vegetation, and soils has found new important uses in connection with biodiversity assessments and the estimation of greenhouse gas emissions. Though time-consuming and difficult, making NFI information comparable across countries through harmonization appears to be a useful approach. The European National Forest Inventory Network (ENFIN) was formed in 2003 and has been successful in pan-European NFI harmonization.
  • Fridman, Swedish University of Agricultural Sciences (SLU), SE-901 83 Umeå, Sweden E-mail: jonas.fridman@slu.se (email)
  • Holm, Swedish University of Agricultural Sciences (SLU), SE-901 83 Umeå, Sweden E-mail: soren.holm@slu.se
  • Nilsson, Swedish University of Agricultural Sciences (SLU), SE-901 83 Umeå, Sweden E-mail: mats.nilsson@slu.se
  • Nilsson, Swedish University of Agricultural Sciences (SLU), SE-901 83 Umeå, Sweden E-mail: per.nilsson@slu.se
  • Ringvall, Swedish University of Agricultural Sciences (SLU), SE-901 83 Umeå, Sweden E-mail: Anna.Ringvall@slu.se
  • Ståhl, Swedish University of Agricultural Sciences (SLU), SE-901 83 Umeå, Sweden E-mail: goran.stahl@slu.se
article id 1008, category Review article
Janusz Szmyt. (2014). Spatial statistics in ecological analysis: from indices to functions. Silva Fennica vol. 48 no. 1 article id 1008. https://doi.org/10.14214/sf.1008
Keywords: spatial analyses; spatial indices; spatial functions; spatial ecology
Highlights: Spatial statistics provides a quantitative description of natural variables distributed in space and time; The objectives of spatial analysis are to detect spatial patterns and to confirm if a pattern found is significant; Spatially explicit indices and functions may be applied depending on the information collected from the field; Development of the specific software supports spatial analyses.
Abstract | Full text in HTML | Full text in PDF | Author Info
This paper presents a review of the most common methods in ecological studies aimed at spatial analysis of population structures (horizontal and vertical), based on point process statistics. Methods based on simple spatially explicit indices as well as more sophisticated methods relying on functions are described in a comprehensible manner. Simple indices revealing the information on spatial structure at the scale of the nearest neighbor can be easily implemented in practical forestry. On the other hand, spatial functions, based on much more detailed data, describe the spatial structure in terms of the spatial relationships between the natural processes and population structures and because of this complexity they are rarely used in forest practice. Including both methods in a single paper is also valuable from the potential reader’s point of view saving their time for searching and choosing the appropriate method to make their spatial analysis. This paper can also serve as an initial guide for young researchers or those who are going to start their studies on spatial aspects of bio-systems. Avoiding the statistical and mathematical details makes this paper understandable for readers who are not statisticians or mathematicians. Readers will find many references related to each method described here, allowing them to find solutions to different problems observed in practice. This paper ends with a list of the most common specific software packages available to support spatial analysis.
  • Szmyt, Department of Silviculture, Faculty of Forestry, Poznań University of Life Sciences, ul. Wojska Polskiego 69, 60-625 Poznań, Poland E-mail: jszmyt@up.poznan.pl (email)
article id 923, category Review article
Katarzyna A. Jadwiszczak. (2012). What can molecular markers tell us about the glacial and postglacial histories of European birches? Silva Fennica vol. 46 no. 5 article id 923. https://doi.org/10.14214/sf.923
Keywords: Betula; cpDNA; glacial refugia; microsatellite; suture zone
Abstract | View details | Full text in PDF | Author Info
The last glaciation was one of the most severe of the Pleistocene epoch. The development of the Scandinavian ice sheet forced many species to reduce their ranges to areas with favourable climatic conditions. Most European species survived the Last Glacial Maximum in refugia in southern parts of Europe. Cold-tolerant species, such as birch trees and shrubs, could also inhabit western, eastern or central Europe. After climate warming, Holocene recolonisation began. This paper presents a comparative analysis of the genetic variation of four European Betula species to reconstruct their glacial and postglacial histories. Two chloroplast DNA haplotypes dominate within the ranges of all birch species, one haplotype is the most common in western and northwestern Europe, the second haplotype occurs mainly in the eastern and southeastern parts of the continent. This finding suggests that birches have recolonised Europe from the western and the eastern refugia, respectively. Most of Europe was likely populated from higher latitude refugia because there was no evidence of isolation by distance and weak genetic structures were detected. Similar patterns of haplotype distributions within Betula ranges indicate that postglacial recolonisation may be disturbed by interspecies hybridisation.
  • Jadwiszczak, Institute of Biology, University of Bialystok, wierkowa 20B, 15-950 Bialystok, Poland E-mail: kszalaj@uwb.edu.pl (email)
article id 446, category Review article
Guntis Brumelis, Bengt Gunnar Jonsson, Jari Kouki, Timo Kuuluvainen, Ekaterina Shorohova. (2011). Forest naturalness in northern Europe: perspectives on processes, structures and species diversity. Silva Fennica vol. 45 no. 5 article id 446. https://doi.org/10.14214/sf.446
Keywords: disturbances; dead wood; natural forest; biological diversity
Abstract | View details | Full text in PDF | Author Info
Saving the remaining natural forests in northern Europe has been one of the main goals to halt the ongoing decline of forest biodiversity. To facilitate the recognition, mapping and efficient conservation of natural forests, there is an urgent need for a general formulation, based on ecological patterns and processes, of the concept of “forest naturalness”. However, complexity, structural idiosyncracy and dynamical features of unmanaged forest ecosystems at various spatio-temporal scales pose major challenges for such a formulation. The definitions hitherto used for the concept of forest naturalness can be fruitfully grouped into three dimensions: 1) structure-based concepts of natural forest, 2) species-based concepts of natural forest and 3) process-based concepts of natural forest. We propose that explicit and simultaneous consideration of all these three dimensions of naturalness can better cope with the natural variability of forest states and also aid in developing strategies for forest conservation and management in different situations. To become operational, criteria and indicators of forest naturalness need to integrate the three dimensions by combining species (e.g. red-listed-, indicator- and umbrella species) with stand and landscape level structural features that are indicative of disturbance and succession processes.
  • Brumelis, Faculty of Biology, University of Latvia, Kronvalda bulv. 4, Riga, LV-1586, Latvia; E-mail: guntis.brumelis@lu.lv (email)
  • Jonsson, Department of Natural Sciences, Engineering and Mathematics, Mid Sweden University, Sundsvall, Sweden E-mail: bgj@nn.se
  • Kouki, School of Forest Sciences, University of Eastern Finland, Joensuu, Joensuu E-mail: jk@nn.fi
  • Kuuluvainen, Department of Forest Sciences, University of Helsinki, Finland E-mail: tk@nn.fi
  • Shorohova, Saint-Petersburg State Forest Academy, Saint-Petersburg, Russia & Finnish Forest Research Institute, Vantaa Research Unit, Vantaa, Finland E-mail: es@nn.ru
article id 74, category Review article
Philip J. Burton, S. Ellen Macdonald. (2011). The restorative imperative: challenges, objectives and approaches to restoring naturalness in forests. Silva Fennica vol. 45 no. 5 article id 74. https://doi.org/10.14214/sf.74
Keywords: afforestation; disturbance regime; ecological restoration; forest rehabilitation; native species; reclamation
Abstract | View details | Full text in PDF | Author Info
Many of the world’s forests are not primeval; forest restoration aims to reverse alterations caused by human use. Forest restoration (including reforestation and forest rehabilitation) is widely researched and practiced around the globe. A review of recent literature reveals some common themes concerning forest restoration motivations and methods. In some parts of the world, forest restoration aims mainly to re-establish trees required for timber or fuelwood; such work emphasizes the propagation, establishment and growth of trees, and equates with the traditional discipline of silviculture. Elsewhere, a recent focus on biocentric values adopts the goal of supporting full complements of indigenous trees and other species. Such ecosystem-based restoration approaches consider natural templates and a wide array of attributes and processes, but there remains an emphasis on trees and plant species composition. Efforts to restore natural processes such as nutrient cycling, succession, and natural disturbances seem limited, except for the use of fire, which has seen widespread adoption in some regions. The inherent challenges in restoring “naturalness” include high temporal and spatial heterogeneity in forest conditions and natural disturbances, the long history of human influence on forests in many regions of the world, and uncertainty about future climate and disturbance regimes. Although fixed templates may be inappropriate, we still have a reasonably clear idea of the incremental steps required to make forests more natural. Because most locations can support many alternative configurations of natural vegetation, the restoration of forest naturalness necessarily involves the setting of priorities and strategic directions in the context of human values and objectives, as informed by our best understanding of ecosystem structure and function now and in the future.
  • Burton, Canadian Forest Service, Natural Resources Canada, 3333 University Way, Prince George, British Columbia, Canada V2N 4Z9 E-mail: Phil.Burton@NRCan-RNCan.gc.ca (email)
  • Macdonald, Department of Renewable Resources, University of Alberta, Edmonton, Alberta, Canada E-mail: sem@nn.ca
article id 73, category Review article
Timo Kuuluvainen, Tuomas Aakala. (2011). Natural forest dynamics in boreal Fennoscandia: a review and classification. Silva Fennica vol. 45 no. 5 article id 73. https://doi.org/10.14214/sf.73
Keywords: disturbance; age structure; stand replacement; cohort; gap; patch; natural disturbance emulation
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The aim here was to review and summarize the findings of scientific studies concerning the types of forest dynamics which occur in natural forests (i.e. forests with negligible human impact) of boreal Fennoscandia. We conducted a systematic search for relevant studies from selected reference databases, using search terms describing the location, structure and processes, and degree of naturalness of the forest. The studies resulting from these searches were supplemented with other known works that were not indexed in the databases. This procedure yielded a total of 43 studies. The studies were grouped into four types of forest dynamics according to the information presented on the characteristics of the native disturbance-succession cycle: 1) even-aged stand dynamics driven by stand-replacing disturbances, 2) cohort dynamics driven by partial disturbances, 3) patch dynamics driven by tree mortality at intermediate scales (> 200 m2) and 4) gap dynamics driven by tree mortality at fine scales (< 200 m2). All four dynamic types were reported from both spruce and pine dominated forests, but their commonness differed. Gap dynamics was most commonly reported in spruce forests, and cohort dynamics in pine forests. The studies reviewed provide the best obtainable overall picture of scientific findings concerning the characteristics and variability of the unmanaged boreal forest dynamics in Fennoscandia. The results demonstrate that the unmanaged Fennoscandian forests are characterized by more diverse and complex dynamics than has traditionally been acknowledged.
  • Kuuluvainen, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: timo.kuuluvainen@helsinki.fi (email)
  • Aakala, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ta@nn.fi
article id 72, category Review article
Ekaterina Shorohova, Daniel Kneeshaw, Timo Kuuluvainen, Sylvie Gauthier. (2011). Variability and dynamics of old-growth forests in the circumboreal zone: implications for conservation, restoration and management. Silva Fennica vol. 45 no. 5 article id 72. https://doi.org/10.14214/sf.72
Keywords: late-successional forest; disturbance regimes; stand dynamic types; cohort dynamics
Abstract | View details | Full text in PDF | Author Info
Due to the unprecedented loss of old-growth forests to harvesting throughout circumboreal regions an understanding of similarities and differences in old-growth dynamics is needed to design effective restoration, management and conservation efforts. This paper reviews concepts, prevalence and variability of old-growth forests across landscapes, and evaluates different stand scale dynamics at the old-growth stage across the circumboreal zone. Old-growth historically dominated many boreal forest landscapes in both Eurasia and North America. Throughout much of North America, and to some extent in western Siberia, the natural prevalence and development of old-growth forests are regulated by the occurrence of stand-replacing fires. In eastern North America and Siberia, insect outbreaks may, however, be more important. Insect outbreaks as well as recurrent non-stand replacing surface fires and windthrows, when occurring at the old-growth stage, often form stands characterized by several tree age-class cohorts. This multi age-class forest development type is common in Europe and eastern Siberia but its prevalence and importance in boreal North-America is not well documented. Similarities in successional dynamics across the circumboreal region are found in the development of mono-dominant even-aged stands, the replacement of shade intolerant tree species by shade tolerant species, as well as in all-aged stands driven by small-scale gap dynamics. The message to land managers is that the focus should not only be on setting aside remaining old-growth forests or in restoring static old-growth attributes, but also in emulating natural disturbances and successional dynamics at landscape and regional scales to maintain natural variability in old-growth attributes through time.
  • Shorohova, Saint-Petersburg State Forest University, Saint-Petersburg, Russia & Finnish Forest Research Institute, Vantaa Research Unit, Vantaa, Finland (ekaterina.shorohova@metla.fi) E-mail: shorohova@ES13334.spb.edu (email)
  • Kneeshaw, Université du Québec à Montréal, Centre d’étude de la forêt, Montreal, Canada E-mail: dk@nn.ca
  • Kuuluvainen, University of Helsinki, Department of Forest Sciences, Helsinki, Finland E-mail: tk@nn.fi
  • Gauthier, Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, Québec, Canada E-mail: sg@nn.fi
article id 38, category Review article
Matieu Henry, Nicolas Picard, Carlo Trotta, Raphaël J. Manlay, Riccardo Valentini, Martial Bernoux, Laurent Saint-André. (2011). Estimating tree biomass of sub-Saharan African forests: a review of available allometric equations. Silva Fennica vol. 45 no. 3B article id 38. https://doi.org/10.14214/sf.38
Keywords: aboveground biomass; databases; inventories; models; Tier method; wood specific gravity
Abstract | View details | Full text in PDF | Author Info
In response to the growing interest in estimating carbon stocks in forests, available allometric equations have been compiled for sub-Saharan Africa. Tree, sprout and stand volume and biomass equations were reviewed. The 850 equations and 125 related references were incorporated into an open-access database on the Carboafrica website (http://www.carboafrica.net). The collected information provides a basic tool for the estimation of biomass and carbon stocks and other purposes, such as bioenergy and fodder supply assessment. A Tier-method approach was developed to illustrate the possible use of the equations. Current available biomass expansion factors that are used to convert a volume to the total aboveground biomass appear to be limited; incomplete species-specific allometric equations are preferred to generalised equations. The analysis of the database highlighted important gaps in available tools to assess forest carbon stocks and changes in these stocks. A quality control assessment revealed that 22% of the equations were misreported and recommendations were proposed to guide further research. Further statistical analyses, such as the Bayesian approach, would help to produce more accurate biomass estimates.
  • Henry, IRD, UMR Eco&Sols, Montpellier SupAgro, 2 place Viala, 34060 Montpellier, France; Di.S.A.F.Ri, Università degli Studi della Tuscia, Viterbo, Italy; and AgroParisTech-ENGREF, GEEFT, Montpellier, France E-mail: henry@unitus.it (email)
  • Picard, CIRAD, Montpellier, France E-mail: np@nn.fr
  • Trotta, Di.S.A.F.Ri, Università degli Studi della Tuscia, Viterbo, Italy E-mail: ct@nn.it
  • Manlay, IRD, UMR Eco&Sols, Montpellier SupAgro, 2 place Viala, 34060 Montpellier, France; and AgroParisTech-ENGREF, GEEFT, Montpellier, France E-mail: rjm@nn.fr
  • Valentini, Di.S.A.F.Ri, Università degli Studi della Tuscia, Viterbo, Italy E-mail: rv@nn.it
  • Bernoux, IRD, UMR Eco&Sols, Montpellier SupAgro, 2 place Viala, 34060 Montpellier, France E-mail: mb@nn.fr
  • Saint-André, CIRAD, Montpellier, France; and INRA, UR1138, Biogeochimie des Ecosystèmes Forestiers, Champenoux, France E-mail: lsa@nn.fr
article id 36, category Review article
Tuomo Wallenius. (2011). Major decline in fires in coniferous forests – reconstructing the phenomenon and seeking for the cause. Silva Fennica vol. 45 no. 1 article id 36. https://doi.org/10.14214/sf.36
Keywords: human influence; fire-history; fire suppression; forest dynamics; Pinus
Abstract | View details | Full text in PDF | Author Info
Steep decline in forest fires about a century ago occurred in coniferous forests over large areas in North America and Fennoscandia. This poorly understood phenomenon has been explained by different factors in different regions. The objective of this study is to evaluate the validity of the four most commonly suggested causes of the decrease in forest fires: fire fighting, over-grazing, climate change and human influence. I compiled the available dendrochronological data and estimated the annually burned proportions of Pinus-dominated forests in four subcontinental regions during the past 500 years. These data were compared to the development of fire suppression, grazing pressure, climate and human livelihoods. The annually burned proportions declined over 90% in all studied regions. In three out of the four regions fires decreased decades before fire suppression began. Available drought data are annually well correlated with fires but could not explain the decrease of the level in annually burned areas. A rapid increase in the number of livestock occurred at the same time with the decrease in fires in the Western US but not in Fennoscandia. Hence, fire suppression in Central Fennoscandia and over-grazing in the Western US may have locally contributed to the reduction of burned areas. More general explanation is offered by human influence hypothesis: the majority of the past forest fires were probably caused by humans and the decrease in the annually burned areas was because of a decrease in human caused fires. This is in accordance with the old written records and forest fire statistics. The decrease in annually burned areas, both in Fennoscandia and the United States coincides with an economic and cultural transition from traditional livelihoods that are associated with high fire use to modern agriculture and forestry.
  • Wallenius, Finnish Forest Research Institute, Vantaa Research Unit, Vantaa, Finland E-mail: tuomo.wallenius@metla.fi (email)
article id 147, category Review article
Arja Lilja, Marja Poteri, Raija-Liisa Petäistö, Risto Rikala, Timo Kurkela, Risto Kasanen. (2010). Fungal diseases in forest nurseries in Finland. Silva Fennica vol. 44 no. 3 article id 147. https://doi.org/10.14214/sf.147
Keywords: damping-off; grey mold; root dieback; needle casts; snowblights; scleroderris canker; Sirococcus; pine twisting rust; stem lesions and top dying; leaf lesions; Venturia; powdery mildews
Abstract | View details | Full text in PDF | Author Info
Norway spruce (Picea abies), Scots pine (Pinus sylvestris) and silver birch (Betula pendula) are the major tree species grown in Finnish forest nurseries where 99% of the seedlings are grown in containers first in plastic-covered greenhouses and later outdoors. The main diseases on conifer seedlings are Scleroderris canker (Gremmeniella abietina), Sirococcus blight and cankers (Sirococcus conigenum), snow blights (Herpotrichia juniperi and Phacidium infestans) and needle casts (Lophodermium seditiosum and Meria laricis). Also grey mould (Botrytis cinerea) and birch rust (Melampsoridium betulinum) are among the diseases to be controlled with fungicides. During last years Scleroderris canker has been a problem on Norway spruce, which has been since 2000 the most common species produced in Finnish nurseries. Root die-back (uninucleate Rhizoctonia sp.) on container-grown spruce and pine was a problem in the 1990s. Today the disease has become less common in modern nurseries due to improvements in hygiene and cultivation practice. Since 1991 stem lesions and top dying caused by Phytophthora cactorum has been a problem on birch. The ongoing climate change has already had effect on rusts and powdery mildews as well as other fungi infecting leaves. All diseases, which gain high precipitation and warm and long autumns. For same reasons winter stored seedlings need sprayings against grey mold. Fungal infections are also possible during short-day (SD) treatment, that is necessary for summer and autumn plantings and a beneficial step prior freezing temperatures outside or in freezer storage. Growers are encouraged to use cultural and integrated pest management techniques such as better nursery hygiene, including removing plant debris in nursery growing areas and hot water washing of containers plus removal of diseased, spore-producing seedlings and trees around the nursery.
  • Lilja, Finnish Forest Research Institute, Vantaa, Finland E-mail: arja.lilja@metla.fi (email)
  • Poteri, Finnish Forest Research Institute, Suonenjoki, Finland E-mail: mp@nn.fi
  • Petäistö, Finnish Forest Research Institute, Suonenjoki, Finland E-mail: rlp@nn.fi
  • Rikala, Finnish Forest Research Institute, Suonenjoki, Finland E-mail: rr@nn.fi
  • Kurkela, Finnish Forest Research Institute, Vantaa, Finland E-mail: tk@nn.fi
  • Kasanen, University of Helsinki, Department of Forest Sciences, Helsinki, Finland E-mail: rk@nn.fi
article id 463, category Review article
Thomas Gschwantner, Klemens Schadauer, Claude Vidal, Adrian Lanz, Erkki Tomppo, Lucio di Cosmo, Nicolas Robert, Daisy Englert Duursma, Mark Lawrence. (2009). Common tree definitions for national forest inventories in Europe. Silva Fennica vol. 43 no. 2 article id 463. https://doi.org/10.14214/sf.463
Keywords: harmonisation; COST Action E43; tree definitions; shrub definitions; tree elements; tree variables; tree characteristics
Abstract | View details | Full text in PDF | Author Info
At the international level, various definitions have been established for the compilation and publication of forest resources assessment results over the last decade. These international definitions frequently rely on terms that are not precisely specified for inventory purposes and do not completely cover the requirements arising from the application of National Forest Inventory (NFI) data. Also, with respect to conventional topics such as forest area and growing stock estimation, several terms and expressions referring to individual trees are not, or are only vaguely, defined until now. Since the individual tree is the basic element of any forest resources assessment, the clarification of tree-related terms is an important part of COST Action E43 to harmonise common reporting of National Forest Inventories. Based on a review of existing definitions and on the requirements for harmonised reporting, common tree-related definitions are established. One objective of this study is to refine and enhance the applicability of available tree and shrub definitions, in particular with regard to the distinction between trees and shrubs. The study also focuses on the parts or “elements” of trees and on the distinction between these elements as they are of particular importance in growing stock and biomass definitions. Furthermore, several definitions for tree characteristics such as “living” and “standing”, as well as tree variables such as height, length, diameter at breast height, and crown projection area are adjusted with respect to NFI purposes. A concluding discussion reflects upon the reviewed, refined and newly established definitions. The definitions presented in this paper provide a firm basis for a common set of harmonised reference definitions developed by COST Action E43 and contribute to the precise and consistent use of terms.
  • Gschwantner, Federal Research and Training Centre for Forests, Natural Hazards and Landscape, Wien, Austria E-mail: thomas.gschwantner@bfw.gv.at (email)
  • Schadauer, Federal Research and Training Centre for Forests, Natural Hazards and Landscape, Wien, Austria E-mail: ks@nn.at
  • Vidal, French National Forest Inventory, Nogent-sur-Vernisson, France E-mail: cv@nn.fr
  • Lanz, Swiss Federal Institute for Forest, Snow and Landscape Research, Birmensdorf, Switzerland E-mail: al@nn.ch
  • Tomppo, Finnish Forest Research Institute, Vantaa, Finland E-mail: et@nn.fi
  • di Cosmo, Agricultural Research Council – Forest Monitoring and Planning Research Unit, Trento, Italy E-mail: ldc@nn.it
  • Robert, French National Forest Inventory, Nogent-sur-Vernisson, France E-mail: nr@nn.fr
  • Englert Duursma, Finnish Forest Research Institute, Vantaa, Finland E-mail: ded@nn.fi
  • Lawrence, Northern Research Station, Roslin, Midlothian, Great Britain E-mail: ml@nn.uk
article id 249, category Review article
Rimvydas Vasaitis, Jan Stenlid, Iben M. Thomsen, Pia Barklund, Anders Dahlberg. (2008). Stump removal to control root rot in forest stands. A literature study. Silva Fennica vol. 42 no. 3 article id 249. https://doi.org/10.14214/sf.249
Keywords: biofuel; stand growth; Armillaria; forest disturbance; Heterobasidion; Phellinus weirii
Abstract | View details | Full text in PDF | Author Info
Tree stumps are expected to be increasingly used for energy production in Fennoscandia, thus environmental consequences of stump removal from forest land must be assessed. Aim of this work was to compile available data on the efficacy of stump removal in eradication of root rot fungi (Heterobasidion, Armillaria, and Phellinus), and to review its potential impacts on establishment and productivity of next forest generation. Site disturbance and some technical and economical aspects are discussed, and needs for future research outlined in northern European context. The review demonstrates that stump removal from clear-felled forest areas in most cases results in, a) reduction of root rot in the next forest generation, b) improved seedling establishment, and c) increased tree growth and stand productivity. Observed disturbances caused to a site by stumping operations are normally acceptable. The available data strongly suggests that possibly many (if achievable, all) rot-containing stumps must be removed during harvesting of stumps. Provided equal availability, the priority should be given for stump removal from root rot-infested forest areas, instead of healthy ones. As most studies were done in North America and Britain, several questions must be yet answered under Fennoscandian conditions: a) if and to which extent the conventional stump removal for biofuel on clear-felled sites could reduce the occurrence of Heterobasidion and Armillaria in the next forest generation, b) what impact is it likely to have on survival of replanted tree seedlings, and c) what consequences will there be for growth and productivity of next forest generation.
  • Vasaitis, Department of Forest Mycology & Pathology, Swedish University of Agricultural Sciences, P.O. Box 7026, SE-75007 Uppsala, Sweden E-mail: rimvys.vasaitis@mykopat.slu.se (email)
  • Stenlid, Department of Forest Mycology & Pathology, Swedish University of Agricultural Sciences, P.O. Box 7026, SE-75007 Uppsala, Sweden E-mail: js@nn.se
  • Thomsen, Forest & Landscape, University of Copenhagen, Hoersholm Kongevej 11, DK-2970 Hoersholm, Denmark E-mail: imt@nn.dk
  • Barklund, Department of Forest Mycology & Pathology, Swedish University of Agricultural Sciences, P.O. Box 7026, SE-75007 Uppsala, Sweden E-mail: pb@nn.se
  • Dahlberg, Swedish Species Information Centre, Swedish University of Agricultural Sciences, P.O. Box 7007, SE-75007 Uppsala, Sweden E-mail: ad@nn.se
article id 313, category Review article
Heikki Hänninen, Koen Kramer. (2007). A framework for modelling the annual cycle of trees in boreal and temperate regions. Silva Fennica vol. 41 no. 1 article id 313. https://doi.org/10.14214/sf.313
Keywords: frost hardiness; climatic adaptation; bud burst; climatic change; dormancy; phenology; photosynthetic capacity
Abstract | View details | Full text in PDF | Author Info
Models of the annual development cycle of trees in boreal and temperate regions were reviewed and classified on the basis of their ecophysiological assumptions. In our classification we discern two main categories of tree development: 1) fixed sequence development, which refers to irreversible ontogenetic development leading to visible phenological events such as bud burst or flowering, and 2) fluctuating development, which refers to reversible physiological phenomena such as the dynamics of frost hardiness during winter. As many of the physiological phenomena are partially reversible, we also describe integrated models, which include aspects of both fixed-sequence and fluctuating development. In our classification we further discern simple E-models, where the environmental response stays constant, and more comprehensive ES-models, where the environmental response changes according to the state of development. On the basis of this model classification, we have developed an operational modelling framework, in which we define an explicit state variable and a corresponding rate variable for each attribute of the annual cycle considered. We introduce a unifying notation, which we also use when presenting a selection of previously published models. To illustrate the various developmental phenomena and their modelling, we have carried out model simulations. Finally, we discuss the ecophysiological interpretation of the model variables, methodological aspects of the empirical development and testing of the models, the introduction of new aspects to the modelling, other closely related models, and applications of the models.
  • Hänninen, Plant Ecophysiology and Climate Change Group (PECC), Department of Biological and Environmental Sciences, Box 65, FI-00014 University of Helsinki, Finland E-mail: heikki.hanninen@helsinki.fi (email)
  • Kramer, Alterra, P.O. Box 47, 6700 AA Wageningen, The Netherlands E-mail: kk@nn.nl
article id 312, category Review article
Katja Lähtinen. (2007). Linking resource-based view with business economics of woodworking industry: earlier findings and future insights. Silva Fennica vol. 41 no. 1 article id 312. https://doi.org/10.14214/sf.312
Keywords: strategy; competitiveness; enterprises; resources; woodworking firms
Abstract | View details | Full text in PDF | Author Info
The business environment and sources of competitiveness in the woodworking industry have changed notably since the 1990s. Wood products are traded globally, and with the increase of trade, abundant forest resources are no longer the main source of sustainable competitiveness. Competition within the woodworking industry is increasing both between European and non-European enterprises, as well as within the EU. Capability to create value-added, making rational strategic choices, and creative usage of intangible and tangible resources have been emphasized as crucial for sustaining woodworking industry competitiveness in higher cost-level countries. Resource-based view (RBV) defines the availability of intangible assets, capabilities and tangible resources, and their heterogeneous combination to form the basis for company success. The objective of this review is to examine the possibilities to employ the RBV to the study of the woodworking industry by combining existing empirical results of the factors of companies’ competitiveness with assessment of the RBV. In the existing literature, strategies implemented in woodworking companies have been approached rather widely, while the role of intangible and tangible resources in building firm-level success has received less focus. In addition, a significant gap exists in linking firms’ financial accounting information with empirical data on their resource-usage and business strategies. In future studies, more information of the effects of these strategic elements on the actual business success of the firms would be needed.
  • Lähtinen, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: katja.lahtinen@metla.fi (email)
article id 369, category Review article
Jaroslaw Zawadzki, Chris J. Cieszewski, Michal Zasada, Roger C. Lowe. (2005). Applying geostatistics for investigations of forest ecosystems using remote sensing imagery. Silva Fennica vol. 39 no. 4 article id 369. https://doi.org/10.14214/sf.369
Keywords: remote sensing; spatial information; semivariance; semivariogram; forest classification
Abstract | View details | Full text in PDF | Author Info
Geostatistically based methods that utilize textural information are frequently used to analyze remote sensing (RS) images. The role of these methods in analyzing forested areas increased rapidly during the last several years following advancements in high-resolution RS technology. The results of numerous applications of geostatistical methods for processing RS forest images are encouraging. This paper summarizes such results. Three closely related topics are reviewed: 1) specific properties of geostatistical measures of spatial variability calculated from digital images of forested areas, 2) determination of biophysical forest parameters using semivariograms and characterization of forest ecosystem structure at the stand level, and 3) forest classification methods based on spatial information.
  • Zawadzki, Environmental Engineering Department, Warsaw Technical University, Ul. Nowowiejska 20, 00-653 Warsaw, Poland E-mail: jaroslaw.zawadzki@is.pw.edu.pl (email)
  • Cieszewski, D. B. Warnell School of Forest Resources, University of Georgia, Athens, GA 30602, USA E-mail: cjc@nn.us
  • Zasada, Department of Forest Productivity, Faculty of Forestry, Warsaw Agricultural University, Poland E-mail: mz@nn.pl
  • Lowe, D. B. Warnell School of Forest Resources, University of Georgia, Athens, GA 30602, USA E-mail: rcl@nn.us
article id 378, category Review article
Yaoqi Zhang, Daowei Zhang, John Schelhas. (2005). Small-scale non-industrial private forest ownership in the United States: rationale and implications for forest management. Silva Fennica vol. 39 no. 3 article id 378. https://doi.org/10.14214/sf.378
Keywords: non-industrial private forest; forest land parcelization; timber supply; transaction costs; economic efficiency; land use change
Abstract | View details | Full text in PDF | Author Info
The transaction cost approach is used to explain why small non-industrial private forest (NIPF) ownerships are increasing in the U.S. We argue that the number of small NIPF owners have increased because: 1) a significant amount of forestland is no longer used economically if primarily for timber production, but rather for non-timber forest products and environmental services (particularly where population density is high), 2) when a person makes frequent use of non-timber products and services, owning forestland is more efficient for them because it saves the transaction costs involved in getting them from the market, 3) forestland parcelization takes place when non-timber value increases faster than timber value, and 4) marginal value for non-timber product is diminishing much faster than that for timber production. The paper also discusses implications of the parcelization of NIPF ownerships on forest management.
  • Zhang, School of Forestry & Wildlife Sciences, Auburn University, AL 36849-5418, USA E-mail: yaoqi.zhang@auburn.edu (email)
  • Zhang, School of Forestry & Wildlife Sciences, Auburn University, AL 36849-5418, USA E-mail: dz@nn.us
  • Schelhas, Southern Research Station, USDA Forest Service, Tuskegee University, AL, USA E-mail: js@nn.us
article id 390, category Review article
Bengt Gunnar Jonsson, Nicholas Kruys, Thomas Ranius. (2005). Ecology of species living on dead wood – lessons for dead wood management. Silva Fennica vol. 39 no. 2 article id 390. https://doi.org/10.14214/sf.390
Keywords: forest management; saproxylic species; coarse woody debris; woodland key habitats; landscape planning; Fennoscandian forests; epixylic species
Abstract | View details | Full text in PDF | Author Info
Dead wood has been identified as a crucial component for forest biodiversity. Recent research has improved our understanding of habitat relations for many species associated with dead wood. However, the consequences for forest management are yet to be explored. In this review we build upon the growing volume of studies on dead wood dependent species, the dynamics of dead wood and ecological theory in order to identify the challenges for forest management at the landscape level. The review has a Fennoscandian focus, but the problems and challenges are similar in many forest ecosystems. We argue that it is necessary to 1) counteract the current shortage in availability of dead wood, 2) concentrate planning at the landscape level in order to minimize isolation and reduce edge effects, 3) create a variety of dead wood types, and 4) utilise available quantitative analytical tools. This calls for new approaches to management that to a large extent includes available knowledge, and to find platforms for planning forested landscapes with diverse holdings.
  • Jonsson, Mid Sweden University, Dept of Natural Sciences, SE-851 70 Sundsvall, Sweden E-mail: bengt-gunnar.jonsson@miun.se (email)
  • Kruys, SLU, Dept of Forest Resource Management and Geomatics, SE-901 83 Umeå, Sweden E-mail: nk@nn.se
  • Ranius, SLU, Dept of Entomology, SE-750 07 Uppsala, Sweden E-mail: tr@nn.se
article id 411, category Review article
Arthur Groot, Sylvie Gauthier, Yves Bergeron. (2004). Stand dynamics modelling approaches for multicohort management of eastern Canadian boreal forests. Silva Fennica vol. 38 no. 4 article id 411. https://doi.org/10.14214/sf.411
Keywords: boreal forests; Canada; dynamic models; multi-cohort forest management
Abstract | View details | Full text in PDF | Author Info
The objective of this paper is to discuss approaches and issues related to modelling stand dynamics for multi-cohort forest management in eastern Canadian boreal forests. In these forests, the interval between wildfires can be rather long, and the development of natural forest stands may include the establishment, growth and mortality of several cohorts of trees. Later cohorts are characterised by increasing structural complexity, including spatial heterogeneity and irregular tree size distribution. A multi-cohort forest management framework has been proposed to maintain this complexity, and associated biodiversity, on the landscape. Multi-cohort forest management planning requires forecasts of the development of stands with complex structure in response to silvicultural treatment and to natural disturbance, but current stand dynamics models in the region are applicable mainly to even-aged mono-specific stands. Possible modelling approaches for complex stands include i) the adaptation of current whole-stand growth and yield models, ii) distance-independent, empirically-derived individual-tree models, such as the USDA Forest Service Forest Vegetation Simulator, and iii) distance-dependent, empirically-derived or process-oriented individual-tree models. We conclude that individual-tree models are needed because observational data for fitting whole-stand models are not available for the full array of silvicultural treatments and natural disturbances encompassed by multi-cohort forest management. Predictive accuracy is a concern with individual-tree models, and the incorporation of coarse-scale constraints into these models is a promising means to control error.
  • Groot, Great Lakes Forestry Centre, Canadian Forest Service, 1219 Queen St. E., Sault Ste. Marie, ON, P6A 2E5, Canada E-mail: agroot@nrcan.gc.ca (email)
  • Gauthier, Laurentian Forestry Centre, Canadian Forest Service, 1055 du P.E.P.S., P.O. Box 3800, Sainte-Foy, QC, G1V 4C7, Canada E-mail: sg@nn.ca
  • Bergeron, University of Quebec at Abitibi-Temiscamingue, 445 Boul de l’Université, Rouyn-Noranda, QC, J9X 5E4, Canada E-mail: yb@nn.ca
article id 489, category Review article
Ralph J. Alig. (2003). U.S. landowner behavior, land use and land cover changes, and climate change mitigation. Silva Fennica vol. 37 no. 4 article id 489. https://doi.org/10.14214/sf.489
Keywords: adaptation; carbon sequestration; forest sector; mitigation
Abstract | View details | Full text in PDF | Author Info
Landowner behavior is a major determinant of land use and land cover changes, an important consideration for policy analysts concerned with global change. Study of landowner behavior aids in designing more effective incentives for inducing land use and land cover changes to help mitigate climate change by reducing net greenhouse gas emissions. Afforestation, deforestation, reforestation, and timber harvest are the most frequent land management practices that influence forest carbon stocks and flux. Research studies provide estimates of how private landowners respond to market signals and government programs and how they alter land management. For example, landowners have tended to retain subsidized afforested stands well beyond program life in the United States, suggesting that similar programs for climate change mitigation could result in high rates of retention. At the same time, policy makers need to be aware that unintended consequences of policies can lead to significantly different outcomes than envisioned, including leakage possibilities.
  • Alig, USDA Forest Service, Forestry Sciences Lab, 3200 SW Jefferson Way, Corvallis, Oregon 97331, USA E-mail: ralig@fs.fed.us (email)
article id 497, category Review article
Oili Kiikkilä. (2003). Heavy-metal pollution and remediation of forest soil around the Harjavalta Cu-Ni smelter, in SW Finland. Silva Fennica vol. 37 no. 3 article id 497. https://doi.org/10.14214/sf.497
Keywords: mulch; copper; forest; ecosystem; Harjavalta; nutrient cycle; remediation; liming
Abstract | View details | Full text in PDF | Author Info
Heavy metals and sulphur have been emitted from the Cu-Ni smelter at Harjavalta since 1945. This article reviews the work that has been published in scientific journals after 1975 concerning heavy metal deposition and the effects of pollution on forest ecosystem around Harjavalta. The pollution has had diverse effects on boreal forest ecosystem, e.g. vegetation, nutrient cycle mediated by microbiota and soil animals, herbivorous insects and pathogens, resistance mechanisms of vegetation, and birds. The deposition of heavy metals has increased up to 30 km distance from the smelter. At 8 km distance the ecosystem began to approximate an undisturbed ecosystem where only slight changes in the understorey vegetation were observed. At 4 km distance the species composition of different ecosystem components (vegetation, insects, birds, soil microbiota) had changed and the growth of trees was retarded. At 0.5–1 km distance, where the nutrient cycling was disturbed and only the most resistant organisms were surviving, the ecosystem had ceased to carry out its essential functions. Remediation through liming or mulching with organic matter, of forest soil has had some positive effects on the ecosystem.
  • Kiikkilä, Vantaa Research Centre, Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: oili.kiikkila@metla.fi (email)
article id 535, category Review article
Thomas J. Givnish. (2002). Adaptive significance of evergreen vs. deciduous leaves: solving the triple paradox. Silva Fennica vol. 36 no. 3 article id 535. https://doi.org/10.14214/sf.535
Keywords: deciduous trees; phenology; evergreens; optimality models; leaf longevity
Abstract | View details | Full text in PDF | Author Info
  • Givnish, Dept of Botany, University of Wisconsin, Madison, WI 53706, USA E-mail: givnish@facstaff.wisc.edu (email)
article id 546, category Review article
Teijo Palander, Mari Toivonen, Sanna Laukkanen. (2002). GroupWare and group decision support systems for wood procurement organisation. A review. Silva Fennica vol. 36 no. 2 article id 546. https://doi.org/10.14214/sf.546
Keywords: collaboration; teamwork-based organisation; group decision-making; communication
Abstract | View details | Full text in PDF | Author Info
Many kinds of decision support systems (DSSs) have been suggested for use of wood procurement organisations, but few meet the real needs of team managers in group decision-making process. Therefore, it has been concluded that the important features of group decision support systems (GDSSs) should be developed for teamwork-based organisations. Electronic meeting systems (EMSs), Computer-aided Visualisations (CAVs) and heuristics as well as other numerical approaches as combined with optimisation seem to be some of the most promising elements of GroupWare, because decisions are made in distributed groups and they deal with human behaviour. Relations between GDSSs and spontaneous decision conferencing (SDC) for modern organisations are also discussed, and suggestions for future research of management approaches are also given.
  • Palander, University of Joensuu, Faculty of Forestry, Box 111, FIN-80101 Joensuu, Finland E-mail: teijo.palander@forest.joensuu.fi (email)
  • Toivonen, University of Joensuu, Faculty of Forestry, Box 111, FIN-80101 Joensuu, Finland E-mail: mt@nn.fi
  • Laukkanen, University of Joensuu, Faculty of Forestry, Box 111, FIN-80101 Joensuu, Finland E-mail: sl@nn.fi
article id 553, category Review article
Yves Bergeron, Alain Leduc, Brian D. Harvey, Sylvie Gauthier. (2002). Natural fire regime: a guide for sustainable management of the Canadian boreal forest. Silva Fennica vol. 36 no. 1 article id 553. https://doi.org/10.14214/sf.553
Keywords: biodiversity; natural disturbance; landscape patterns; coarse filter; harvest pattern; volume retention; historic variability; even-age management
Abstract | View details | Full text in PDF | Author Info
The combination of certain features of fire disturbance, notably fire frequency, size and severity, may be used to characterize the disturbance regime in any region of the boreal forest. As some consequences of fire resemble the effects of industrial forest harvesting, conventional forest management is often considered as a disturbance that has effects similar to those of natural disturbances. Although the analogy between forest management and fire disturbance in boreal ecosystems has some merit, it is important to recognise that it also has its limitations. Short fire cycles generally described for boreal ecosystems do not appear to be universal; rather, important spatial and temporal variations have been observed in Canada. These variations in the fire cycle have an important influence on forest composition and structure at the landscape and regional levels. Size and severity of fires also show a large range of variability. In regions where the natural matrix of the boreal forest remains relatively intact, maintenance of this natural variability should be targeted by forest managers concerned with biodiversity conservation. Current forest management tends to reduce this variability: for example, fully regulated, even-aged management will tend to truncate the natural forest age distribution and eliminate over-mature and old-growth forests from the landscape. We suggest that the development of strategic-level forest management planning approaches and silvicultural techniques designed to maintain a spectrum of forest compositions and structures at different scales in the landscape is one avenue to maintain this variability. Although we use the boreal forest of Quebec for our examples, it is possible to apply the approach to those portions of the boreal forest where the fire regime favours the development of even-aged stands in burns.
  • Bergeron, NSERC-UQAT-UQAM Industrial Chair in Sustainable Forest Management, C.P. 8888, Succursale Centre-ville, Montréal, Québec, Canada H3C 3P8 E-mail: bergeron.yves@uqam.ca (email)
  • Leduc, NSERC-UQAT-UQAM Industrial Chair in Sustainable Forest Management, C.P. 8888, Succursale Centre-ville, Montréal, Québec, Canada H3C 3P8 E-mail: al@nn.ca
  • Harvey, NSERC-UQAT-UQAM Industrial Chair in Sustainable Forest Management, C.P. 8888, Succursale Centre-ville, Montréal, Québec, Canada H3C 3P8 E-mail: bdh@nn.ca
  • Gauthier, NSERC-UQAT-UQAM Industrial Chair in Sustainable Forest Management, C.P. 8888, Succursale Centre-ville, Montréal, Québec, Canada H3C 3P8 E-mail: sg@nn.ca
article id 552, category Review article
Timo Kuuluvainen. (2002). Natural variability of forests as a reference for restoring and managing biological diversity in boreal Fennoscandia. Silva Fennica vol. 36 no. 1 article id 552. https://doi.org/10.14214/sf.552
Keywords: biodiversity; ecosystem management; disturbance dynamics; succession; sustainable forestry; heterogeneity; hierarchy; scaling; spatiotemporal dynamics
Abstract | View details | Full text in PDF | Author Info
In Fennoscandia, use of the natural forest as a reference for restoration and management of forest biodiversity has been widely accepted. However, limited understanding of the structure and dynamics of the natural forest has hampered the applications of the natural variability approach. This is especially the case in areas, where the natural forests have almost totally vanished. This review was motivated by the idea that despite these difficulties the essential features of the natural forest can be reconstructed based on biological archives, historical documents, research done in adjacent natural areas, and modeling. First, a conceptual framework for analyzing the relationship between forest structure, dynamics and biodiversity is presented. Second, the current understanding of the structure and dynamics of natural forests at different spatiotemporal scales in boreal Fennoscandia is reviewed. Third, the implications of this knowledge, and gaps in knowledge, on research and on practical restoration and management methods aimed at forest biodiversity conservation are discussed. In conclusion, naturally dynamic forest landscapes are complex, multiscaled hierarchical systems. Current forest management methods create disturbance and successional dynamics that are strongly scale-limited when compared with the natural forest. To restore some of the essential characteristics of the natural forest’s multiscale heterogeneity, diversification of silvicultural and harvesting treatments, as guided by natural disturbance dynamics, is needed to produce more variation in disturbance severity, quality, extent, and repeatability.
  • Kuuluvainen, Department of Forest Ecology, P.O. Box 24, FIN-00014 University of Helsinki, Finland E-mail: timo.kuuluvainen@helsinki.fi (email)
article id 551, category Review article
Anders Dahlberg. (2002). Effects of fire on ectomycorrhizal fungi in Fennoscandian boreal forests. Silva Fennica vol. 36 no. 1 article id 551. https://doi.org/10.14214/sf.551
Keywords: boreal forests; ectomycorrhizal fungi; fire; population; community
Abstract | View details | Full text in PDF | Author Info
Fire, the primary natural disturbance factor in Fennoscandian boreal forests, is considered to have exerted major selection pressure on most boreal forest organisms. However, recent studies show that few ectomycorrhizal (EM) fungi appear to have evolved post-fire adaptations, no succession of EM fungi following fire is apparent after low intensity fires, and only two EM fungal taxa exclusively fruit at post-fire conditions. In this paper I review the present knowledge of effects of forest fire on EM fungal communities in Fennscandian boreal forests, put into perspective by a comparison from other parts of the world. Characteristically, these boreal forests consist of less than a handful of tree species, e.g. Scots pine and Norway spruce, while the below ground communities of EM fungi is impressively species rich with presently more than 700 known taxa. Commonly, forest fires in Fennoscandia have been of low intensity, with a considerable portion of the trees surviving and the organic humus layer partly escaping combustion. Hence, EM fungi appear to largely have evolved under conditions characterised by a more or less continuous presence of their hosts. In fact, the composition of EM fungi within a forest appear be more variable due to spatial variation than due to wildfire. However, in areas with high intensity burns and high tree mortality, most EM fungi may locally be killed. Thus, the legacy of EM fungi following wildfire depends on the survival of trees, which determine the potential for mycorrhizal growth, and the combustion and heating of the organic soil, which directly correlate to mortality of mycorrhizas. The questions if and to what degree fires may be of significance for yet unidentified spatiotemporal dynamics of EM fungal populations and communities are discussed. Recent experiments indicate a few EM fungi are favoured by high intensity burn conditions whereas others disappear. The consequences of wildfires in temperate conifer forests differ considerably from those in boreal forests. Wildfires in temperate conifer forests are typically high intensity stand-replacing fires that cause a total combustion of organic layers. Subsequently, pre-fire EM fungal communities are largely eradicated and a succession of post-fire EM fungi is initiated.
  • Dahlberg, Department of Forest Mycology and Pathology, Swedish University of Agricultural Sciences, P.O. Box 7026, SE-750 07 Uppsala, Sweden E-mail: anders.dahlberg@artdata.slu.se (email)
article id 550, category Review article
Lars Edenius, Margareta Bergman, Göran Ericsson, Kjell Danell. (2002). The role of moose as a disturbance factor in managed boreal forests. Silva Fennica vol. 36 no. 1 article id 550. https://doi.org/10.14214/sf.550
Keywords: management; boreal forest; disturbance; forestry; Alces alces; monitoring; herbivory; large ungulates; moose
Abstract | View details | Full text in PDF | Author Info
We review the interactions between moose (Alces alces) and native tree species in Fennoscandia. The Fennoscandian boreal forests have been intensively managed for wood production over decades. Moose population density is also relatively high in these northern forests. Forest management affects habitat characteristics and food resources from regeneration to final harvest, with the most significant effects occurring early in the stand development. The plant-animal interactions found in such a situation may be different from what has been observed in natural boreal forests with low densities of moose (e.g. in North America). The strong focus on Scots pine (Pinus sylvestris) in forest regeneration in conjunction with a homogenisation of the landscape structure by clear-cutting has favoured moose. Forest development is controlled by man from regeneration to final harvest, and in relation to human-induced disturbances the disturbance by moose is relatively small, but occurs on different spatial levels. At the landscape level, the most prominent effects of moose seem to be suppression and/or redistribution of preferred browse species. At the forest stand level moose primarily induce spatial heterogeneity by browsing patchily and exploiting existing gaps. At the tree level, moose damage trees and lower timber quality, but also create substrate types (e.g. dead and dying wood) valuable for many organisms. Co-management of moose and forest requires good monitoring programmes for both plants and animals, as well as extensive ecological knowledge on the relations between moose and their food plants on different spatial levels.
  • Edenius, SLU, Department of Animal Ecology, SE-901 83 Umeå, Sweden E-mail: lars.edenius@szooek.slu.se (email)
  • Bergman, SLU, Department of Animal Ecology, SE-901 83 Umeå, Sweden E-mail: mb@nn.se
  • Ericsson, SLU, Department of Animal Ecology, SE-901 83 Umeå, Sweden E-mail: ge@nn.se
  • Danell, SLU, Department of Animal Ecology, SE-901 83 Umeå, Sweden E-mail: kd@nn.se
article id 549, category Review article
Andrei Gromtsev. (2002). Natural disturbance dynamics in the boreal forests of European Russia: a review. Silva Fennica vol. 36 no. 1 article id 549. https://doi.org/10.14214/sf.549
Keywords: boreal forests; disturbance dynamics; European Russia; fires; windthrow
Abstract | View details | Full text in PDF | Author Info
In the European part of the Russian boreal zone the dynamics of pristine forests (taiga) has been studied by several generations of researchers. Many studies have examined the patterns and role of fire, windthrow, insect outbreaks and other natural disturbances. An attempt is made to provide a brief review of these studies. The reviewed studies show that lightning strikes were the only natural source of fires in taiga. The frequency of fires varied in various types of pristine landscape from 1–2 per century to 1–2 per millennium. Fires maintained a dynamic equilibrium between compositionally different forest communities or their certain ratio and areal occurrence. Fires favored the regeneration and recovery of pine forests and prevented the replacement of shade-intolerant species (e.g. pine) by shade-tolerant ones (e.g. spruce). Taiga forests generally displayed a mosaic pattern that varied from pioneer plant communities, growing in open burns, to climax communities that were extremely seldom affected by fire. The reviewed studies suggest that fires were a powerful ecological factor in pristine taiga, being largely responsible for the structure and spontaneous dynamics of forest communities. Windfalls were also common in pristine taiga landscapes and they regulated spontaneous dynamics in a gap-mosaic regime, which is most characteristic of spruce forests.
  • Gromtsev, Forest Research Institute, Karelian Research Centre of the Russian Academy of Sciences, P.O. Box 185610, Petrozavodsk, Pushkin st. 11, Russia E-mail: gromtsev@karelia.ru (email)
article id 548, category Review article
Kevin C. Ryan. (2002). Dynamic interactions between forest structure and fire behavior in boreal ecosystems. Silva Fennica vol. 36 no. 1 article id 548. https://doi.org/10.14214/sf.548
Keywords: boreal forest; disturbance dynamics; stand structure; fire behavior; fire severity
Abstract | View details | Full text in PDF | Author Info
This paper reviews and synthesizes literature on fire as a disturbance factor in boreal forests. Spatial and temporal variation in the biophysical environment, specifically, vegetative structure, terrain, and weather lead to variations in fire behavior. Changes in slope, aspect, elevation, and soil affect site energy and water budgets and the potential plant community. These terrain features also have a major influence on fire-caused disturbance through their role in determining moisture conditions and flammability of fuels on hourly, seasonal, and successional time-scales. On fine time scales (minutes to hours), changes in weather, specifically wind and relative humidity, significantly affect a fire’s intensity and aboveground effects. Normal seasonal changes in dryness and periodic drought influence fire intensity and severity principally by affecting the depth of burn and belowground effects. On decades-long time scales changes in vegetative structure affect the mass of fuel available for burning and therefore the potential energy that can be released during a fire. The severity of fire varies in time and space depending not only on the biophysical environment, but also on the location on the fire’s perimeter (head vs. flank vs. rear). Spatial and temporal variation in severity within a fire can have long-lasting impacts on the structure and species composition of post-fire communities and the potential for future disturbances. Characteristic temperature histories of ground, surface, and crown fires are used to illustrate variations in fire severity. A soil-heating model is used to illustrate the impact of varying depth of burn on the depth at which various fire effects occur in the soil profile. A conceptual model is presented for the effects of fire severity on fire-plant regeneration interactions. The conceptual model can be used by restoration ecologists to evaluate the differential effects of controlled or prescribed fires and wildfires and to plan and implement fire treatments to conserve biodiversity.
  • Ryan, USDA Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory, P.O. Box 8089, Missoula, Montana 59807, USA E-mail: kryan@fs.fed.us (email)
article id 632, category Review article
William F. Hyde, Gunnar Köhlin. (2000). Social forestry reconsidered. Silva Fennica vol. 34 no. 3 article id 632. https://doi.org/10.14214/sf.632
Keywords: deforestation; community forestry; developing countries; forest resources; rural development; welfare
Abstract | View details | Full text in PDF | Author Info
This paper reviews the expectations for forestry’s contribution to rural development – and for its special contributions to the most disadvantaged, to women and the landless users of the forest commons. A growing literature challenges some of these expectations; in particular, certain expectations about cultural differences and physical stocks as explanatory factors for patterns of household behavior. This literature could also be used to support a call for sharper definitions of deforestation, improved indicators of the effects of forest resources on the rural poor, and improved design of forest policy interventions. Our paper reviews the literature, suggests some unifying themes, and identifies the critical issues that remain unanswered. The primary contention arising from this literature is that households follow systematic patterns of economic behavior in their consumption and production of forest resources, and that policy interventions in social forestry should be analyzed with regard to markets, policies, and institutions. Markets for forest resources generally exist in some form – although they may be thin. Successful forestry projects and policies require careful identification of the target populations and careful estimation of market and market-related effects on the household behavior of these populations. Institutional structures that assure secure rights for scarce forest resources are uniquely important in a forest enviornment often characterized by open access resources and weak government administration. Social and community forestry, improved stoves, improved strains of multi-purpose trees, and even private commercial forest operations can all improve local welfare, but only where scarcity is correctly identified and the appropriate institutions are in place. An increasing number of observations of afforestation from developing countries around the world is evidence that forestry activities do satisfy these conditions in selective important cases. The critical point for policy is to identify the characteristics of these successful cases that are predictive of other cases where new forestry activities can be welfare enhancing.
  • Hyde, Centre for International Forestry Research, Bogor, Indonesia E-mail: wfhyde@aol.com (email)
  • Köhlin, Göteborg University, Göteborg, Sweden E-mail: gk@nn.se
article id 639, category Review article
Klaus von Gadow. (2000). Evaluating risk in forest planning models. Silva Fennica vol. 34 no. 2 article id 639. https://doi.org/10.14214/sf.639
Keywords: harvest scheduling; continuous cover forest; activity scenario
Abstract | View details | Full text in PDF | Author Info
The purpose of forest scenario modelling is to evaluate multiple management options and to answer what if questions relating to a particular development path of a given forest. Forest scenario planning can reduce uncertainty in management outcomes by anticipating the future in a systematic way, thus reducing the likelihood of unexpected events. It can also improve the chance that future developments will agree with specified objectives. Numerous techniques have been proposed for generating and evaluating scenarios of forest development. Some of the techniques are limited to applications in simple forest production systems while others are suitable for any type of forest management, including individual tree selection systems. Risk is defined as the expected loss due to a particular hazard for a given area and reference period. An expected loss may be calculated as the product of the damage and its probability. Risk analysis, risk evaluation and risk management are formal procedures for quantifying, evaluating and managing risk within a given hazard domain. Applications of risk analysis in forest scenario planning are rare and greater emphasis needs to be placed on hazard prediction. The aim of this contribution is to discuss some aspects of risk analysis, including examples of specific modelling tools. In a forest planning model risk can be considered in the form of specific constraints limiting the total risk in a given time period. Expected hazards can be used to exclude certain risky alternatives and finally, risk can be calculated and used to reduce the value of an objective function coefficient.
  • Gadow, Georg-August-University Göttingen, Institute for Forest Management, Büsgenweg 5, 37077 Göttingen, Germany E-mail: kgadow@gwdg.de (email)
article id 688, category Review article
John A. Stanturf, Callie J. Schweitzer, Emile S. Gardiner. (1998). Afforestation of marginal agricultural land in the Lower Mississippi River Alluvial Valley, U.S.A. Silva Fennica vol. 32 no. 3 article id 688. https://doi.org/10.14214/sf.688
Keywords: Populus deltoides; bottomland hardwoods; direct-seeding; Liquidambar styraciflua; Fraxinus pennsylvanica; Quercus nuttallii; Quercus nigra
Abstract | View details | Full text in PDF | Author Info
Afforestation of marginal agricultural land in the Lower Mississippi Alluvial Valley (LMAV) relies on native species, planted mostly in single-species plantations. Hard mast species such as oak and pecan are favored for their value to wildlife, especially on public land. Successful afforestation requires an understanding of site variation within floodplains and matching species preferences and tolerances to site characteristics, in particular to inundation regimes. Soil physical conditions, root aeration, nutrient availability, and moisture availability during the growing season also must be considered in matching species to site. Afforestation methods include planting seedlings or cuttings, and direct-seeding. Both methods can be done by hand or by machine. If good quality seedlings are planted properly and well cared for before planting, the chances for successful establishment are high but complete failures do occur. Mortality and poor growth are caused by many factors: extended post-planting drought or flooding; poor planting or seeding practices; poor quality seed or seedlings; animal depredation; or herbicide drift from aerial application to nearby cropland. More species can be planted, even on continuously flooded sites. Direct-seeding, while limited to heavy-seeded species (oaks and hickories), costs less than 50% of planting seedlings. Growth varies considerably by soil type; most bottomland hardwoods grow best on silt loam and less well on clay soils. Up to 200 000 ha of land in the LMAV subject to spring and early summer backwater flooding could be afforested over the next decade.
  • Stanturf, USDA Forest Service, Southern Research Station, P.O. Box 227, Stoneville, MS 38776, USA E-mail: jstantur/srs_stoneville@fs.fed.us (email)
  • Schweitzer, USDA Forest Service, Southern Research Station, P.O. Box 227, Stoneville, MS 38776, USA E-mail: cjs@nn.us
  • Gardiner, USDA Forest Service, Southern Research Station, P.O. Box 227, Stoneville, MS 38776, USA E-mail: esg@nn.us
article id 687, category Review article
Janusz B. Zwolinski, David B. South, E. A. P. Droomer. (1998). Pine mortality after planting on post-agricultural lands in South Africa. Silva Fennica vol. 32 no. 3 article id 687. https://doi.org/10.14214/sf.687
Keywords: afforestation; mortality; Pinus patula; Pythium spp.; scalping
Abstract | View details | Full text in PDF | Author Info
Successful afforestation has been practiced in South Africa for more than a century. Recently, however, problems with afforestation of pines have occurred in the northeastern part of the Eastern Cape Province. Rapid mortality of Pinus patula and P. elliottii have occurred when small container seedlings were planted on old-agricultural soils. Death would often occur within 5 months of planting. Growth of surviving trees was retarded and new needles were chlorotic and stunted. Acceptable survival was obtained when seedlings were planted on virgin grasslands. Apparently, some unseen factor in the post-agricultural soil reduces root growth, increases mortality, and decreases uptake of nutrients. Removal of the infested soil by scalping greatly improves survival and growth as does soil fumigation with methyl bromide.
  • Zwolinski, Northeast Cape Forests, 19 Nassau St., 5470 Ugie, Rep. of South Africa E-mail: janusz.zwolinski@paper.mondi.co.za (email)
  • South, School of Forestry, Auburn University, Auburn, Alabama 36849-5418, USA E-mail: dbs@nn.us
  • Droomer, Northeast Cape Forests, 19 Nassau St., 5470 Ugie, Rep. of South Africa E-mail: eapd@nn.za
article id 695, category Review article
Marja-Leena Päätalo. (1998). Factors influencing occurrence and impacts of fires in northern European forests. Silva Fennica vol. 32 no. 2 article id 695. https://doi.org/10.14214/sf.695
Keywords: tree species; fire; climate; risk assessment; site
Abstract | View details | Full text in PDF | Author Info
The return interval and number of fires vary, depending on the geographical location in interaction with climate, topography and amount of fuel. During recent decades, in northern Europe the number and severity of fires have been insignificant compared with Mediterranean region, in which fire return intervals may be 15–35 years, compared to the average of 60–120 years for boreal forests. This is partly due to the efficient system of fire protection in northern Europe, but is mainly due to the less favourable climate for fire and the smaller human impact on ignition of forest fires. The consequences of fire are related to both site and stand characteristics, site being the most important factor controlling the stability of stands. Dry sites being more flammable and likely to ignite are associated with high risk of fire. In northern Europe, due to the interaction between species and site, the role of species difference in risk of fire damage is not clear. In southern Europe, fire risk cannot be explained by differences between tree species. There, other vegetation (shrubs, etc.) is of major importance for the risk of fire. Management of forests can, to some degree, alter the risk and the occurrence of fire. In northern Europe, logging may have compensated for fire occurrence by decreasing the amount of fuel. In addition, forest roads act as fire-breaks and facilitate fire-fighting. On the contrary, in southern Europe the risk of fire has been found to increase because the traditional forest uses and management have decreased, which increases the accumulation of fuel. However, it is not yet possible to quantify and compare the effect of management in absolute terms. Currently, some tools, such as fire-risk indices, remote sensing and GIS-based techniques, are available for prediction of fire risk in some areas. For example, fire-risk indices are most suitable for areas, like northern Europe, which have a low fire risk. In high-risk areas, such as southern Europe, more sophisticated techniques are needed for assessment of the risk. In the future, assuming global warming at northern latitudes (2 x CO2 climate), the risk of fire damage could also increase in northern Europe. Therefore, to allow the various locational and silvicultural factors to be assessed on the European level, an integrated risk model is needed.
  • Päätalo, Faculty of Forestry, University of Joensuu, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: mlp@nn.fi (email)
article id 694, category Review article
Gösta Eriksson. (1998). Evolutionary forces influencing variation among populations of Pinus sylvestris. Silva Fennica vol. 32 no. 2 article id 694. https://doi.org/10.14214/sf.694
Keywords: Pinus sylvestris; Scots pine; among-population variation; evolutionary forces
Abstract | View details | Full text in PDF | Author Info
The evolutionary forces influencing genetic differentiation among populations are identified. Natural selection, random genetic drift, and mutations promote differentiation while phenotypic plasticity and gene flow delay or prevent differentiation. Evolution is a dynamic force which leads to instability and absence of any perfection in the adaptive process. Natural selection acts mainly on phenotypes and only indirectly on the components of important breeding traits. In the northern part of the distribution of Scots pine there is a large among- and within-population variation in survival. The high among-population variation occurs in spite of an assumed high gene flow. Biomass is a product of many components and it also shows a high population variation. Many markers are neutral and such markers will not reveal adaptive variation.
  • Eriksson, Swedish University of Agricultural Sciences, Department of Forest Genetics, Uppsala, Sweden E-mail: gosta.eriksson@sgen.slu.se (email)
article id 693, category Review article
Erik G. Ståhl. (1998). Changes in wood and stem properties of Pinus sylvestris caused by provenance transfer. Silva Fennica vol. 32 no. 2 article id 693. https://doi.org/10.14214/sf.693
Keywords: Pinus sylvestris; yield; silviculture; provenance; wood properties; growth phenology
Abstract | View details | Full text in PDF | Author Info
Wood properties focused in forest tree breeding should be of economic importance, have a large total variation and a high heritability. The properties of interest are those that influence the strength and durability of sawn products or the amount and properties of pulp produced. The following wood properties are treated: width of the annual ring, juvenile wood, late wood content, heart wood, tracheid dimensions, basic density, stem straightness and branch diameter. The provenance variation in wood properties can be related to differences in growth phenology. In the northern part of distribution P. sylvestris (L.) provenances transferred a few degrees southwards have a high survival and yield but stem wood production is low. Trees from these provenances will be straight and with few spike knots or other injuries. The shoot elongation period will be short and the temperature sum required for wood formation sufficient. Provenances transferred southwards will form thin annual rings, few and thin branches, little early wood, high basic density and slender tracheids with thick cell walls in comparison to local provenances. An example of the effect of alternative transfers on the yield and wood properties is evaluated. In regions with deviating climatic patterns alternative provenance transfer patterns may be better. The objectives of the land owner should influence the provenance choice. The importance of integrating tree improvement with silvicultural management is discussed with reference to spacing.
  • Ståhl, College of Dalarna, CITU Centre for Industrial Technology and Development, S-781 88 Borlänge, Sweden E-mail: esl@du.se (email)

Category : Complex remote sensing-assisted forest surveys - Research article

article id 24061, category Complex remote sensing-assisted forest surveys - Research article
Zsofia Koma, Johannes Breidenbach. (2025). Large-scale validation of forest attribute maps across different spatial resolutions. Silva Fennica vol. 59 no. 2 article id 24061. https://doi.org/10.14214/sf.24061
Keywords: airborne laser scanning; National Forest Inventory; forest resource mapping; resolution dependence
Highlights: The study assesses stand-level uncertainty of biomass, volume, basal area, and Lorey’s height estimates resulting from the prediction of maps across varying spatial resolutions (1–30 m); The changes of RMSE and bias across the different spatial resolutions were generally small (< 5%) for additive forest attributes such as biomass, volume, and basal area; The changes of RMSE and bias were also small for Lorey’s height as a non-additive forest attribute if the resolution difference was less than 2 times of the native resolution; The models fitted at the resolution of the NFI plot size can be used to produce forest attribute maps at 10 m resolution without concerning increases in uncertainty at stand-level.
Abstract | Full text in HTML | Full text in PDF | Author Info

Fine-scale, spatially explicit forest attribute maps are essential for guiding forest management and policy decisions. Such maps, based on the combination of National Forest Inventory (NFI) and remote sensing datasets, have a long tradition in the Nordic countries. Harmonizing the pixel size among national forest attribute maps would considerably improve the utility of the maps for users. However, the maps are often aligned with the NFI plot size, and the influence of creating these maps at different spatial resolutions (i.e. pixel sizes) is little studied. We assess the stand-level uncertainty (RMSE) of biomass, volume, basal area, and Lorey’s height estimates resulting from the aggregation of maps across varying spatial resolutions. Models fit at 16 m native resolution using more than 14 000 NFI plots were applied for predictions at pixels sizes (side lengths) of 1, 5, 10, 16, and 30 m. For independent validation, we used more than 600 field plots – that cover a total area of 24 ha and were clustered within 65 stands across Norway. For all attributes, the lowest RMSEs, ranging from 6.86% for Lorey’s height to 13.86% for volume, were observed for predictions at pixel sizes of 5 m to 16 m. The RMSE changes across resolutions were generally small (< 5%) for biomass, volume, and basal area. For Lorey’s height, changing the spatial resolution resulted in large RMSEs of up to 25%. Overall, our findings suggest that the main forest attributes can be mapped at a finer resolutions without complex adjustments.

  • Koma, Norwegian Institute for Bioeconomy Research (NIBIO), Division of Forest and Forest Resources, Department National Forest Inventory, Høgskoleveien 7, 1433 Ås, Norway ORCID https://orcid.org/0000-0002-0003-8258 E-mail: zsofia.koma@nibio.no (email)
  • Breidenbach, Norwegian Institute for Bioeconomy Research (NIBIO), Division of Forest and Forest Resources, Department National Forest Inventory, Høgskoleveien 7, 1433 Ås, Norway ORCID https://orcid.org/0000-0002-3137-7236 E-mail: johannes.breidenbach@nibio.no

Category : Research note

article id 24062, category Research note
Jussi Manner, Hagos Lundström. (2025). The effect of boom-tip control on harvester time consumption in Picea abies dominated final-felling stands: case study. Silva Fennica vol. 59 no. 1 article id 24062. https://doi.org/10.14214/sf.24062
Keywords: productivity; automation; cut-to-length logging; crane work; Intelligent Boom Control (IBC); Levels of Automation (LoA); operator assistance system
Highlights: Two experienced harvester operators (A and B) participated in the study; Operator A reduced time consumption by 10% with boom-tip control, while operator B neither saved nor lost time; Operator A’s time savings occurred exclusively during the work element felling-processing; Variation in the results between the operators emphasizes the need for further research involving a larger pool of operators.
Abstract | Full text in HTML | Full text in PDF | Author Info

Crane work accounts for the majority of a harvester’s productive working time. Boom-tip control assists operators by coordinating end-effector movements, offering the potential to improve productivity. With boom-tip control, the operator steers the boom-tip directly rather than controlling individual crane joints. Despite being commercially available for several years, research on boom-tip control’s impact on harvester work – particularly compared to forwarder work – is limited. Therefore, this study analysed the effect of boom-tip control on harvester time consumption (s m-3) in final-felling stands, involving two experienced operators (A and B) driving a John Deere 1470G harvester. John Deere’s boom-tip control system, Intelligent Boom Control (IBC), was compared to a conventional boom-control system. Data were analysed separately for each operator. While operator A saved time using IBC, no statistically significant difference between IBC and the conventional boom-control system was observed for operator B. For operator A, IBC reduced total time consumption (s m-3) by approximately 10%. The results indicate a need for further research, involving multiple machine manufacturers, operators, and work environments such as thinning and final-felling stands. Moreover, future studies should preferably utilise automated data recording to generate large follow-up datasets on harvester work.

  • Manner, Skogforsk, Uppsala Science Park, 751 83 Uppsala, Sweden ORCID https://orcid.org/0000-0002-4982-3855 E-mail: jussi.manner@skogforsk.se (email)
  • Lundström, Skogforsk, Uppsala Science Park, 751 83 Uppsala, Sweden E-mail: hagos.lundstrom@skogforsk.se
article id 24027, category Research note
Julian Frey, Zoe Schindler, Patrick McClatchy, Christopher Morhart, Elena Larysch, Thomas Seifert. (2025). The 3D reconstruction of wood and leaves from terrestrial laser scanning – a case study on PAR measurements below a solitary Malus domestica tree. Silva Fennica vol. 59 no. 1 article id 24027. https://doi.org/10.14214/sf.24027
Keywords: terrestrial laser scanning; leaf geometry; LESS; light modelling; Malus domestica; PAR; structural tree model
Highlights: Direct reconstruction of leaf and wood polygons from terrestrial laser scanning data by open source software; Validation of reconstruction based on in-situ PAR measurements and direct comparison to a turbid voxel approach; High correlations between in situ PAR measurements and RTM simulation (r = 0.92).
Abstract | Full text in HTML | Full text in PDF | Author Info

In this paper, we present a new methodology that directly extracts the geometry of woody features (wood and bark) and foliage from 3D data originating from terrestrial laser scans. Our goal was to enhance the precision of radiative transfer models for modelling tree shading by using highly resolved 3D tree models. The approach was tested on a single apple tree (Malus domestica (Suckow) Borkh.) in a peri-urban setting and was validated by utilising an open-source radiative transfer model and comparing the simulation output with in-situ measurements of photosynthetically active radiation (PAR) as well as simulations utilizing turbid voxels of 0.2 m and 1 m edge length. The in-situ measurements of 60 PAR sensors showed a correlation coefficient (r) of 0.92 with the simulated light intensities for the reconstructed polygons which was higher than for the voxel-based approaches (0.2 m:  r = 0.85, 1 m: r = 0.73). We were able to demonstrate that our approach effectively simulates light extinction through the canopy. This innovative method has the potential to easily provide detailed insights into high resolution radiation patterns within forests, which are connected to multiple ecosystem functions like species and habitat diversity.

  • Frey, Chair of Forest Growth and Dendroecology, University of Freiburg, Freiburg im Breisgau, Tennenbacher Str. 4, 79106 Freiburg, Germany ORCID https://orcid.org/0000-0001-7895-702X E-mail: julian.frey@wwd.uni-freiburg.de (email)
  • Schindler, Chair of Forest Growth and Dendroecology, University of Freiburg, Freiburg im Breisgau, Tennenbacher Str. 4, 79106 Freiburg, Germany ORCID https://orcid.org/0000-0003-2972-1920 E-mail: zoe.schindler@wwd.uni-freiburg.de
  • McClatchy, Chair of Forest Growth and Dendroecology, University of Freiburg, Freiburg im Breisgau, Tennenbacher Str. 4, 79106 Freiburg, Germany E-mail: patrickjmcclatchy@gmail.com
  • Morhart, Chair of Forest Growth and Dendroecology, University of Freiburg, Freiburg im Breisgau, Tennenbacher Str. 4, 79106 Freiburg, Germany ORCID https://orcid.org/0000-0003-1874-5011 E-mail: christopher.morhart@wwd.uni-freiburg.de
  • Larysch, Chair of Forest Growth and Dendroecology, University of Freiburg, Freiburg im Breisgau, Tennenbacher Str. 4, 79106 Freiburg, Germany ORCID https://orcid.org/0000-0002-1191-5770 E-mail: elena.larysch@wwd.uni-freiburg.de
  • Seifert, Chair of Forest Growth and Dendroecology, University of Freiburg, Freiburg im Breisgau, Tennenbacher Str. 4, 79106 Freiburg, Germany; Department of Forest and Wood Science, Stellenbosch University, Private Bag X1, Matieland, 7602, Stellenbosch, South Africa ORCID https://orcid.org/0000-0002-9611-6272 E-mail: thomas.seifert@wwd.uni-freiburg.de
article id 24039, category Research note
Jussi Manner, Hagos Lundström. (2024). The effect of forked trees on harvester time consumption in a Pinus contorta final-felling stand. Silva Fennica vol. 58 no. 4 article id 24039. https://doi.org/10.14214/sf.24039
Keywords: productivity; stem quality; stem defect; lodgepole pine; cut-to-length logging; double stem; forking
Highlights: For forked trees, codominant stems double harvester time consumption per tree, while double crowns have only minor impacts on harvester time consumption; Forked trees should be removed during thinning when they can still be time-efficiently multi-tree handled; Dealing with forked trees at a later rotation age, when they are too large for multi-tree handling, becomes excessively time-consuming.
Abstract | Full text in HTML | Full text in PDF | Author Info

The consensus on the factors affecting harvester productivity is generally widely acknowledged in the discipline. However, research results regarding the effect of forking on productivity are diverse. Some studies show that harvester productivity is halved when harvesting double stems compared to single-stem trees, while other studies indicate that forking does not necessarily decrease harvester productivity. These differences in study results can depend on what is considered forking. In our study, the forking occurred above the breast-height level. We defined codominant stems as forked trees too large to be multi-tree handled. In contrast, we defined double crowns as forked trees that could be multi-tree handled. The objective of our study was to analyse how the presence of codominant stems and/or double crowns affects harvester time consumption. The study was conducted in Sweden in 2022, involving two operators and two large harvesters. The 45-year-old Pinus contorta Douglas ex Loudon-dominated stand was clearcut during the study. We found that the presence of codominant stems doubles harvester time consumption per tree, while double crowns had only a minor impact on harvester time consumption. Additionally, total time consumption increased linearly with increasing diameter at breast height. Based on these findings, we recommend that forked trees be removed already during thinning when they can still be time-efficiently multi-tree handled. Dealing with forked trees later during the rotation cycle, when they are too large for multi-tree handling, is excessively time-consuming.

  • Manner, Skogforsk, Uppsala Science Park, 751 83 Uppsala, Sweden ORCID https://orcid.org/0000-0002-4982-3855 E-mail: jussi.manner@skogforsk.se (email)
  • Lundström, Skogforsk, Uppsala Science Park, 751 83 Uppsala, Sweden E-mail: hagos.lundstrom@skogforsk.se
article id 23064, category Research note
Henrik Lindberg, Tuomas Aakala, Ilkka Vanha-Majamaa. (2024). Ignition probability and fuel consumption of boreal ground vegetation fuels – an experimental study in Finland. Silva Fennica vol. 58 no. 3 article id 23064. https://doi.org/10.14214/sf.23064
Keywords: forest fires; ground vegetation; prescribed burning; flammability; fuel moisture content; mass loss rate
Highlights: Ignition probability and mass loss rates clearly differed in experimental burnings among four common circumboreal moss and lichen species; Cladonia rangiferina was the most flammable, Dicranum spp. the least flammable and Pleurozium schreberi and Hylocomium splendens intermediate.
Abstract | Full text in HTML | Full text in PDF | Author Info

In boreal forests fires often ignite and spread within the dominant moss and lichen cover of the ground layer vegetation, which thus greatly influences fire hazard. We used an experimental set-up in greenhouse conditions to study the differences in how (1) fuel moisture and (2) wind velocity influence the ignition probability and fuel consumption among four common circumboreal ground vegetation fuels, Pleurozium schreberi (Willd. ex Brid.) Mitt., Hylocomium splendens Schimp., Dicranum spp. and Cladonia rangiferina (L.) F. H. Wigg. Our results show that the reindeer lichen C. rangiferina was clearly the most flammable species, with high ignition probability even at high moisture contents and low wind velocities. Of the mosses, Dicranum was the least flammable, with low ignition probability and mass loss at low wind velocities regardless of moisture content. P. schreberi and H. splendens behaved somewhat similarly with wind velocities quickly increasing the initially low ignition probability and mass loss observed in the absence of wind. However, especially for mass loss, among-species differences tended to disappear with stronger winds. The observed differences can be explained by the different structures and growth forms of the studied species and open a potential avenue for improving forest fire risk predictions.

  • Lindberg, Häme University of Applied Sciences, School of Bioeconomy, Visamäentie 35 A, P.O. Box 230, FI-13100 Hämeenlinna, Finland ORCID https://orcid.org/0000-0001-9045-2673 E-mail: henrik.lindberg@hamk.fi (email)
  • Aakala, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0003-0160-6410 E-mail: tuomas.aakala@uef.fi
  • Vanha-Majamaa, Natural Resources Institute Finland (Luke), Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: ilkka.vanha-majamaa@luke.fi
article id 23050, category Research note
Anne Carolina, Rita K. Sari, Deded S. Nawawi, Effendi T. Bahtiar, Dai Kusumoto. (2024). Mechanical-chemical induction of balsam from Liquidambar excelsa trees. Silva Fennica vol. 58 no. 2 article id 23050. https://doi.org/10.14214/sf.23050
Keywords: Liquidambar excelsa; balsam; ethylene; methyl jasmonate; sustainable tapping
Highlights: The combination of mechanical and chemical induction on Rasamala branches offers an easy and efficient method for balsam exudation; Balsam exudation is chemically stimulated by methyl jasmonate and ethephon; Stimulant concentration increased the amount of balsam exuded in a dose-dependent manner.
Abstract | Full text in HTML | Full text in PDF | Author Info

Rasamala (Liquidambar excelsa (Noronha) Oken) is an endemic plant in Indonesia. Apart from its use as wood, Rasamala also produces an exudate, known as balsam. Rasamala balsam has the potential to be a substitute for other true balsams derived from Altingiaceae, namely Storax. However, local communities have not used Rasamala balsam to its full potential owing to a lack of knowledge about the tapping method and processing. Therefore, an easy and efficient induction method for plant exudates is required to boost productivity. The use of exogenous hormones as stimulants and less damaging tapping techniques for plant stems requires further investigation. In this study, mechanical and chemical inductions were conducted using 0.1%, 1%, 2%, 5%, and 10% (w/w) methyl jasmonate and ethephon as stimuli. These chemical compounds were applied to young twigs without incision (TW), by incision (TI), to branches perforated with an electric bore (BB), and by incision (BI). After exogenous application for 21 days, Rasamala balsam exuded in all induction techniques, except for the TW treatment. BI treatment showed the highest effective induction, as indicated by the highest balsam exudation. Furthermore, methyl jasmonate was a better chemical stimulant than ethephon. In addition, the induced balsam Rasamala exudate showed a physical characteristic of a clear, thick, sticky colorless to white liquid with a distinctive balsamic odor.

  • Carolina, Department of Forest Products, Faculty of Forestry and Environment, IPB University, Jl. Lingkar Kampus IPB Dramaga, Bogor, 16680, Indonesia ORCID https://orcid.org/0000-0002-2129-8665 E-mail: a_caroline@apps.ipb.ac.id (email)
  • Sari, Department of Forest Products, Faculty of Forestry and Environment, IPB University, Jl. Lingkar Kampus IPB Dramaga, Bogor, 16680, Indonesia ORCID https://orcid.org/0000-0001-5377-1384 E-mail: rita_kartikasari@apps.ipb.ac.id
  • Nawawi, Department of Forest Products, Faculty of Forestry and Environment, IPB University, Jl. Lingkar Kampus IPB Dramaga, Bogor, 16680, Indonesia ORCID https://orcid.org/0000-0001-8367-0349 E-mail: dnawawi@apps.ipb.ac.id
  • Bahtiar, Department of Forest Products, Faculty of Forestry and Environment, IPB University, Jl. Lingkar Kampus IPB Dramaga, Bogor, 16680, Indonesia ORCID https://orcid.org/0000-0003-0003-5855 E-mail: bahtiar_et@apps.ipb.ac.id
  • Kusumoto, The University of Tokyo Chiba Forest, Graduate School of Agricultural and Life Sciences, The University of Tokyo, 770 Amatsu, Kamogawa, Chiba 299-5503, Japan ORCID https://orcid.org/0000-0002-4250-2469 E-mail: kusumoto@uf.a.u-tokyo.ac.jp
article id 23071, category Research note
Mattias Finndin, Per Milberg. (2024). The population development of small trees and shrubs after 100 years of free succession of a wooded meadow in southern Sweden. Silva Fennica vol. 58 no. 1 article id 23071. https://doi.org/10.14214/sf.23071
Keywords: conservation; land-use change; nemoral forest; plants; regrowth; wooded grassland
Highlights: Using a unique map of trees and shrubs from 1937, we estimated the mortality of woody species typical of wooded meadows after management ceased in 1923; Both population size and canopy cover of the studied species had decreased during the past 86 years; On the other hand, several tree and shrub specimens endured for a century, pointing to the slow changes involved as well as the potential for restoration.
Abstract | Full text in HTML | Full text in PDF | Author Info

Wooded meadows are characterised by traditional-historic human use. Deliberate selection of species, pollarding and haymaking has created a complex and biodiverse habitat where small trees and shrubs were prevalent. This study set out to document what happens to such trees and shrubs during succession to forest, the normal fate when wooded meadows are abandoned but also when other open to semi-open patches revert to forest. The study was conducted at a site in southern Sweden where traditional management was abandoned by 1923 when the area was protected for research and allowed to follow natural succession. The current study is a follow-up of a 1937-inventory of small trees and shrubs. The results show a decrease in both population size and canopy cover in the selected species during the past 86 years. Hence, we can expect a loss of these species when wooded meadow are abandoned and left to developed into forests. On the other hand, several tree and shrub specimens endured for a century, pointing to the slow changes involved as well as the potential for restoration.

article id 23026, category Research note
Michael M. Müller, Leena Hamberg, Martti Varama. (2023). Palatability of Norway spruce needles infected with Lophodermium piceae to larvae of two sawfly species. Silva Fennica vol. 57 no. 3 article id 23026. https://doi.org/10.14214/sf.23026
Keywords: antagonism; herbivory; fungi; Neodiprion sertifer; Gilpinia pallida; palatability
Highlights: We describe the first experiment including both endophyte inhabited and endophyte free intact needles of Norway spruce for examining the possible effect of endophytes on the palatability of needles to insect larvae; The extent of needles consumed by neither of two sawfly species, Neodiprion sertifer and Gilpina pallida, differed depending on endophyte presence.
Abstract | Full text in HTML | Full text in PDF | Author Info

Unequivocal evidence on the antagonistic effects of endophytic fungi associated with woody plants against insect herbivores has been documented in only a few cases so far. Experimental evidence of the significance of needle endophytes to coniferous trees has remained scant because it is difficult to obtain trees with needles free of endophytes that could be used as comparable controls for trees infected with endophytes. Previously we reported a new methodology to get Norway spruce (Picea abies (L.)  Karst.) saplings without needle endophytes and to inoculate them with a needle endophyte Lophodermium piceae (Fuckel) Höhn. Here we describe the first trial where spruce saplings with and without needle endophytes were provided as substrate for insect larvae. We transferred larvae of two sawfly species, Neodiprion sertifer Geoffroy and Gilpina pallida Klug, to the seedlings. Even though the main host of these sawfly species is not Norway spruce, but Scots pine (Pinus sylvestris L.), they are also known to occasionally feed on spruce. In this experiment the larvae did not develop to pupae with the provided spruce substrate but consumed measurable amounts of needles. No significant difference was found between the extent of needles consumed by either of the two sawfly species in inoculated and uninoculated saplings.

  • Müller, Natural Resources Institute Finland (Luke), Forest Health and Biodiversity, P.O. Box 2, 00791 Helsinki, Finland ORCID https://orcid.org/0000-0003-3699-2139 E-mail: micms.muller@gmail.com (email)
  • Hamberg, Natural Resources Institute Finland (Luke), Forest Health and Biodiversity, P.O. Box 2, 00791 Helsinki, Finland ORCID https://orcid.org/0000-0002-0009-7768 E-mail: leena.hamberg@luke.fi
  • Varama, Natural Resources Institute Finland (Luke), Forest Health and Biodiversity, P.O. Box 2, 00791 Helsinki, Finland E-mail: martti.varama@pp.inet.fi
article id 23017, category Research note
Yrjö Nuutinen, Jari Miina. (2023). Effect of boom corridor and selective thinning on the post-treatment growth of young Scots pine and birch stands. Silva Fennica vol. 57 no. 2 article id 23017. https://doi.org/10.14214/sf.23017
Keywords: geometrical thinning; systematic thinning; thinning reaction
Highlights: During the 4–5-year post-treatment period, boom corridor thinning did not result in growth and yield losses compared to selective thinning; Within the boom corridor and selective thinning treatments, the increment of trees at the edge of strip roads or corridors was higher than at those trees located in the middle of strip roads and/or corridors.
Abstract | Full text in HTML | Full text in PDF | Author Info

Boom corridor thinning (BCT) is a harvester’s working method, primarily suitable for dense, unmanaged young stands. The method was first studied in Sweden in the early 2000s. In Finland, the idea has been further developed and studied for Finnish forests. The advantage is in the corridor, where the harvester head can move more swiftly, and there is no need to identify trees to grow as much as when using the traditional selective thinning (Sel) method. Moreover, the method can be conducted without cost-intensive pre-clearing of undergrowth, creating post-stands with higher biodiversity. This study is the sequel to a previous study in which experiments on BCT and Sel were established in 2017–2018. The experiments were remeasured 4–5 years after their establishment, and the effect of BCT treatments of Scots pine (Pinus sylvestris L.) and silver birch (Betula pendula Roth) on the post-treatment growth and growth reaction of individual trees within the treatments was compared to traditional Sel. During the post-treatment period, BCT did not result in growth or yield losses compared to Sel. Within the treatments, the increment of trees at the edge of strip roads or corridors was higher than that of trees located in the middle of strip roads and/or corridors. A longer post-treatment period needs to be studied to analyse the effect of BCT on the total yield and especially the yield of saw logs during the rest of the rotation period.

article id 22008, category Research note
Eeva Terhonen. (2023). First report of Diplodia tip blight on Scots pine in Finland. Silva Fennica vol. 56 no. 4 article id 22008. https://doi.org/10.14214/sf.22008
Keywords: Pinus sylvestris; drought; Diplodia sapinea; Sphaeropsis sapinea; emerging fungal disease
Highlights: Diplodia tip blight is a new disease on Scots pine in Finland; Diplodia sapinea can be identified reliably with the presented pipeline.
Abstract | Full text in HTML | Full text in PDF | Author Info

Diplodia sapinea (Fr.) Fuckel causes shoot blight on Scots pine (Pinus sylvestris L.). This fungus has been discovered in Finland as a saprophyte in 2015 on Scots pine cones. The endophytic mode of this fungus was later discovered in healthy Scots pine twigs. In 2021 the disease, Diplodia tip blight was observed on Scots pine in Finland. Currently, the disease symptoms are poorly identified so the role of D. sapinea in disease outbreaks in Finland are easily overlooked. The identification of the fungi is challenging in field conditions and requires targeted identification in laboratory. In this research note I report the first Diplodia tip blight outbreaks observed in Finland, the typical disease symptoms, and methodology for the species identification. Samples were collected from symptomatic trees based on observations made by the citizens. Diplodia sapinea was isolated from defoliated and surface sterilized twigs. The species identification by morphological characters was further confirmed with sequencing of ITS region of rDNA and with species-specific primers. A pathogenicity test confirmed that D. sapinea was the disease agent causing shoot blight. This is the first report of Diplodia tip blight on Scots pine in Finland.

  • Terhonen, Natural Resources Institute Finland (Luke), Forest health and biodiversity, Latokartanonkaari 9, FI‐00790 Helsinki, Finland ORCID 0000-0002-9288-440X E-mail: eeva.terhonen@luke.fi (email)
article id 10765, category Research note
Girmantė Jurkšienė, Virgilijus Baliuckas, Donatas Naugžemys, Donatas Žvingila. (2022). Chloroplast DNA polymorphism and morphometric characteristics of Carpinus betulus in the Lithuania forests. Silva Fennica vol. 56 no. 3 article id 10765. https://doi.org/10.14214/sf.10765
Keywords: polymorphism; european hornbeam; intergenic spacer; involucres; migration refugia
Highlights: A 24 bp deletion was found in the chloroplast DNA region of two populations in the southeastern part of Lithuania; Morphometric differences in hornbeam involucre between the study populations were significant; The existence of two haplotypes of the chloroplast DNA region supports the hypothesis of two migration refugia in Carpinus betulus populations.
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The European hornbeam (Carpinus betulus L.) is a medium-sized deciduous tree that spreads northeast of the middle of Lithuania. Carpinus betulus L. is a native tree in Poland, and its branch is migrated by two Pleistocene refugia. We hypothesised that its branches had spread to Lithuania. In this study, we selected 10 populations of hornbeam that were chosen from their distribution location. We sequenced the chloroplast intergenic spacer psbA-trnH of 70 individuals. We found 24 bp deletion in chloroplast DNA (cpDNA) individuals of two populations in the southeastern part of Lithuania. In the seven forest populations, we examined the morphological variability of hornbeam seed involucres and nuts variations of 30 morphometric characteristics. Initial genetic population studies were conducted over a wider area; when differences were detected, morphological studies were conducted in the contact zone. Morphometric differences between the study populations were significant. The existence of two haplotypes of cpDNA supports the hypothesis of two migration refugia in C. betulus populations. This study contributes to significant novel knowledge about the morphological and cpDNA variability of European hornbeam populations in Lithuania and Europe.

  • Jurkšienė, Institute of forestry, Lithuanian Research Centre for Agriculture and Forestry, Liepų str. 1 Girionys, LT-53101 Kaunas, Lithuania ORCID https://orcid.org/0000-0001-8210-6711 E-mail: girmante.jurksiene@lammc.lt (email)
  • Baliuckas, Institute of forestry, Lithuanian Research Centre for Agriculture and Forestry, Liepų str. 1 Girionys, LT-53101 Kaunas, Lithuania; Faculty of Forest Sciences and Ecology, Agriculture Academy, Vytautas Magnus University, K. Donelaičio g. 58, LT-44248 Kaunas, Lithuania E-mail: virgilijus.baliuckas@lammc.lt
  • Naugžemys, Botanical Garden of Vilnius University, Vilnius University, Kairėnų Str. 43, Vilnius 10239, Lithuania ORCID https://orcid.org/0000-0001-6744-5360 E-mail: genetikas@gmail.com
  • Žvingila, Department of Botany and Genetics, Institute of Biosciences, Life Sciences Center, Vilnius University, Saulėtekio Av. 7, LT-10257 Vilnius, Lithuania ORCID https://orcid.org/0000-0001-7826-1815 E-mail: donatas.zvingila@gf.vu.lt
article id 10561, category Research note
Urszula Zajączkowska, Piotr Dąbrowski, Waldemar Kowalczuk, Grzegorz Tarwacki. (2022). Leaf photosynthetic capacity, trunk wood structure and stem xylem sap flow in 700-years old Quercus robur L.: a pilot study upon oak ‘Bartek’, a natural monument in Poland. Silva Fennica vol. 56 no. 3 article id 10561. https://doi.org/10.14214/sf.10561
Keywords: photosynthetic capacity; ‘Bartek’ oak; tree stem tomography; xylem sap flow
Highlights: Photosynthetic and hydraulic capacity of a 700-year-old Quercus robur is comparable to reference values from the literature measured in younger oak trees.
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Physiological studies of long-lived trees are particularly important at this time, especially in light of the need for trees to adapt to global climate change. The results of the present studies were obtained on an approximately 700-year-old Quercus robur L. – the ‘Bartek’ oak. The tree has to adapt to changing climatic conditions, starting from the transition between the Medieval Warm Period and the Little Ice Age, up to the present time of rapid global climate change. Tomograph imaging showed decay of the tree trunk interior and revealed that undamaged wood forms a thin layer around the trunk perimeter. Two series of experiments were carried out to assess the physiological state of the tree. The first concerned measurements related to photosynthetic capacity: chlorophyll a fluorescence, gas exchange (CO2 assimilation, transpiration), stomatal conductance and leaf water potential. The second series concerned xylem sap flow velocity and anatomical studies of stem wood. Photosynthetic capacity was within the limits reported for young healthy trees. The diurnal pattern of velocity of xylem sap flow was also typical for young vigorous trees and flow velocity correlated positively with solar radiation and negatively with air relative humidity. Anatomical observations of the outermost wood showed relatively narrow annuals rings with large diameter earlywood vessels. The results indicate that the veteran tree does not show signs of water stress probably due to a good balance of water flow and that leaf area of the canopy needs only the current ring of wood to feed transpiration of the canopy.

  • Zajączkowska, Department of Forest Botany, Warsaw University of Life Sciences - SGGW, 159 Nowoursynowska Street, 02-776 Warsaw, Poland ORCID https://orcid.org/0000-0002-7119-7547 E-mail: urszula_zajaczkowska@sggw.edu.pl (email)
  • Dąbrowski, Department of Environmental Development Warsaw University of Life Sciences - SGGW, Warsaw, Poland ORCID https://orcid.org/0000-0002-2867-8839 E-mail: piotr_dabrowski@sggw.edu.pl
  • Kowalczuk, Ekosystem Waldemar Kowalczuk Tomasz Kowalczuk, Otwock, Poland E-mail: ekosystem@ekosystem.waw.pl
  • Tarwacki, Forest Protection Department, Forest Research Institute, Sękocin Las, Poland ORCID https://orcid.org/0000-0001-5979-7788 E-mail: G.Tarwacki@ibles.waw.pl
article id 10683, category Research note
Aarne Hovi, Petr Lukeš, Lucie Homolová, Jussi Juola, Miina Rautiainen. (2022). Small geographical variability observed in Norway spruce needle spectra across Europe. Silva Fennica vol. 56 no. 2 article id 10683. https://doi.org/10.14214/sf.10683
Keywords: albedo; remote sensing; reflectance; transmittance; land surface modeling; leaf optical properties; radiative transfer modeling
Highlights: Spectra of Norway spruce needles were collected from three sites in Europe (49°–62°N); The same acquisition and processing parameters were applied throughout the campaign; Geographical variability in the needle spectra was small; Comparison of the spectra of coniferous needles and broadleaved tree foliage is also presented.
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Foliage spectra form an important input to physically-based forest reflectance models. However, little is known about geographical variability of coniferous needle spectra. In this research note, we present an assessment of the geographical variability of Norway spruce (Picea abies (L.) H. Karst.) needle albedo, reflectance, and transmittance spectra across three study sites covering latitudes of 49–62°N in Europe. All spectra were measured and processed using exactly the same methodology and parameters, which guarantees reliable conclusions about geographical variability. Small geographical variability in Norway spruce needle spectra was observed, when compared to variability observed between previous measurement campaigns (employing slightly varying measurement and processing parameters), or to variability between plant functional types (broadleaved vs. coniferous). Our results suggest that variability of needle spectra is not a major factor introducing geographical variability to forest reflectance. The results also highlight the importance of harmonizing measurement protocols when collecting needle spectral libraries. Furthermore, the data collected for this study can be useful in studies where accurate information on spectral differences between broadleaved and coniferous tree foliage is needed.

  • Hovi, Aalto University, School of Engineering, Department of Built Environment, P.O. Box 14100, FI-00760 Aalto, Finland ORCID https://orcid.org/0000-0002-4384-5279 E-mail: aarne.hovi@aalto.fi (email)
  • Lukeš, Global Change Research Institute of the Czech Academy of Sciences, Bělidla 986/4a, 603 00 Brno, Czech Republic ORCID https://orcid.org/0000-0002-3707-6557 E-mail: lukes.p@czechglobe.cz
  • Homolová, Global Change Research Institute of the Czech Academy of Sciences, Bělidla 986/4a, 603 00 Brno, Czech Republic ORCID https://orcid.org/0000-0001-7455-2834 E-mail: homolova.l@czechglobe.cz
  • Juola, Aalto University, School of Engineering, Department of Built Environment, P.O. Box 14100, FI-00760 Aalto, Finland ORCID https://orcid.org/0000-0002-6050-7247 E-mail: jussi.juola@aalto.fi
  • Rautiainen, Aalto University, School of Engineering, Department of Built Environment, P.O. Box 14100, FI-00760 Aalto, Finland; Aalto University, School of Electrical Engineering, Department of Electronics and Nanoengineering, P.O. Box 15500, FI-00760 Aalto, Finland ORCID https://orcid.org/0000-0002-6568-3258 E-mail: miina.a.rautiainen@aalto.fi
article id 10679, category Research note
Jari Miina, Mikko Kurttila. (2022). A model for the sap yield of birches tapped by citizen scientists. Silva Fennica vol. 56 no. 2 article id 10679. https://doi.org/10.14214/sf.10679
Keywords: Betula spp.; non-timber forest product; linear mixed model; crowdsourcing
Highlights: Tree diameter and mean stand height positively affected the sap yield of birches; The sap yield varied between trees, stands, and years; The sap yield model can be utilised in profitability analyses for sap tapping.
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The sap yield of birches (Betula pendula Roth and B. pubescens Ehrh.) was modelled as a function of tree diameter (girth) at breast height, as well as site and stand characteristics measured and reported by citizen scientists representing mainly non-industrial private forest owners in the South Savo, North Karelia, and Northern Ostrobothnia regions in Finland. Birches (tree species not recorded) growing on both mineral and peatland sites were tapped during the springs of 2019 and 2020. Citizen scientists were mainly voluntary forest owners who received the instructions and equipment (spouts, drop lines and buckets) for collecting sap from three birches of different diameters in the same birch stand. Citizen scientists were instructed to measure and report the sap yield and girth of the trees, as well as stand characteristics from the forest resource data, if available. Based on the linear mixed model fitted to the data, the sap yield increased with the increasing tree diameter and mean stand height, and varied between years, stands, and trees; between-region variation was not significant. In a birch stand, the simulated total sap yield ha–1 was depended on the average tree size and the stem number ha–1 and was at its highest just before the first commercial thinning and again before the second thinning. The sap model can be used to predict the necessary sap yield in profitability analyses for sap tapping.

article id 10600, category Research note
Nea Kuusinen, Aarne Hovi, Miina Rautiainen. (2021). Contribution of woody elements to tree level reflectance in boreal forests. Silva Fennica vol. 55 no. 5 article id 10600. https://doi.org/10.14214/sf.10600
Keywords: reflectance model; bark; hyperspectral; spectral mixture analysis
Highlights: Contribution of woody elements to reflectance of boreal tree species was estimated using spectral mixture analysis and airborne hyperspectral data; Mean woody element contribution varied between 0.14–0.19 (Scots pine), 0.12–0.20 (birches) and 0.09–0.10 (Norway spruce).
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Spectral mixture analysis was used to estimate the contribution of woody elements to tree level reflectance from airborne hyperspectral data in boreal forest stands in Finland. Knowledge of the contribution of woody elements to tree or forest reflectance is important in the context of lea area index (LAI) estimation and, e.g., in the estimation of defoliation due to insect outbreaks, from remote sensing data. Field measurements from four Scots pine (Pinus sylvestris L.), five Norway spruce (Picea abies (L.) Karst.) and four birch (Betula pendula Roth and Betula pubescens Ehrh.) dominated plots, spectral measurements of needles, leaves, bark, and forest floor, airborne hyperspectral as well as airborne laser scanning data were used together with a physically-based forest reflectance model. We compared the results based on simple linear combinations of measured bark and needle/leaf spectra to those obtained by accounting for multiple scattering of radiation within the canopy using a physically-based forest reflectance model. The contribution of forest floor to reflectance was additionally considered. The resulted mean woody element contribution estimates varied from 0.140 to 0.186 for Scots pine, from 0.116 to 0.196 for birches and from 0.090 to 0.095 for Norway spruce, depending on the model used. The contribution of woody elements to tree reflectance had a weak connection to plot level forest variables.

  • Kuusinen, Department of Built Environment, School of Engineering, Aalto University, P.O. Box 14100, FI-00076 Aalto, Finland E-mail: nea.kuusinen@aalto.fi (email)
  • Hovi, Department of Built Environment, School of Engineering, Aalto University, P.O. Box 14100, FI-00076 Aalto, Finland E-mail: aarne.hovi@aalto.fi
  • Rautiainen, Department of Built Environment, School of Engineering, Aalto University, P.O. Box 14100, FI-00076 Aalto, Finland; Department of Electronics and Nanoengineering, School of Electrical Engineering, Aalto University, P.O. Box 15500, FI-00076 Aalto, Finland ORCID https://orcid.org/0000-0002-6568-3258 E-mail: miina.a.rautiainen@aalto.fi
article id 10557, category Research note
Mikko T. Niemi. (2021). Improvements to stream extraction and soil wetness mapping within a forested catchment by increasing airborne LiDAR data density – a case study in Parkano, western Finland. Silva Fennica vol. 55 no. 5 article id 10557. https://doi.org/10.14214/sf.10557
Keywords: remote sensing; interpolation; laser scanning; digital elevation model conditioning; overland flow routing; soil drainage; wetness index
Highlights: Overland flow routing can be improved with high-density airborne LiDAR data; Kriging and inverse-distance weighting outperformed triangulated irregular networks in DEM interpolation; A hybrid breaching-filling workflow performed well for DEM conditioning in the Finnish landscape; Enhanced stream extraction and soil wetness mapping contribute to multi-purpose precision forestry.
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The pulse density of airborne Light Detection and Ranging (LiDAR) is increasing due to technical developments. The trade-offs between pulse density, inventory costs, and forest attribute measurement accuracy are extensively studied, but the possibilities of high-density airborne LiDAR in stream extraction and soil wetness mapping are unknown. This study aimed to refine the best practices for generating a hydrologically conditioned digital elevation model (DEM) from an airborne LiDAR -derived 3D point cloud. Depressionless DEMs were processed using a stepwise breaching-filling method, and the performance of overland flow routing was studied in relation to a pulse density, an interpolation method, and a raster cell size. The study area was situated on a densely ditched forestry site in Parkano municipality, for which LiDAR data with a pulse density of 5 m–2 were available. Stream networks and a topographic wetness index (TWI) were derived from altogether 12 DEM versions. The topological database of Finland was used as a ground reference in comparison, in addition to 40 selected main flow routes within the catchment. The results show improved performance of overland flow modeling due to increased data density. In addition, commonly used triangulated irregular networks were clearly outperformed by universal kriging and inverse-distance weighting in DEM interpolation. However, the TWI proved to be more sensitive to pulse density than an interpolation method. Improved overland flow routing contributes to enhanced forest resource planning at detailed spatial scales.

  • Niemi, Department of Forest Sciences, University of Helsinki, FI-00014 University of Helsinki, Finland E-mail: mikko.t.niemi@helsinki.fi (email)
article id 10533, category Research note
Daniel Schraik, Aarne Hovi, Miina Rautiainen. (2021). Estimating cover fraction from TLS return intensity in coniferous and broadleaved tree shoots. Silva Fennica vol. 55 no. 4 article id 10533. https://doi.org/10.14214/sf.10533
Keywords: terrestrial laser scanning; leaf area; lidar intensity; physically-based; voxel
Highlights: We developed a method to obtain the fraction of TLS pulses’ footprint area covered by a target’s projection area; We tested our method with shoots of Norway spruce, Scots pine and silver birch; We provide a physically-based framework related to unmeasured variables, and provide a robust statistical framework to deal with uncertainty.
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Terrestrial laser scanning (TLS) provides a unique opportunity to study forest canopy structure and its spatial patterns such as foliage quantity and dispersal. Using TLS point clouds for estimating leaf area density with voxel-based methods is biased by the physical dimensions of laser beams, which violates the common assumption of beams being infinitely thin. Real laser beams have a footprint size larger than several millimeters. This leads to difficulties in estimating leaf area density from light detection and ranging (LiDAR) in vegetation, where the target objects can be of similar or even smaller size than the beam footprint. To compensate for this bias, we propose a method to estimate the per-pulse cover fraction, defined as the fraction of laser beams’ footprint area that is covered by vegetation targets, using the LiDAR return intensity and an experimental calibration measurement. We applied this method to a Leica P40 single-return instrument, and report our experimental results. We found that conifer foliage had a lower average per-pulse cover fraction than broadleaved foliage, indicating an increased number of partial hits in conifer foliage. We further discuss limitations of our method that stem from unknown target properties that influence the LiDAR return intensity and highlight potential ways to overcome the limitations and manage the remaining uncertainty. Our method’s output, the per-beam cover fraction, may be useful in a weight function for methods that estimate leaf area density from LiDAR point clouds.

  • Schraik, Aalto University, School of Engineering, Department of Built Environment, P.O. Box 14100, FI-00076 Aalto, Finland; ORCID https://orcid.org/0000-0002-7794-3918 E-mail: daniel.schraik@aalto.fi (email)
  • Hovi, Aalto University, School of Engineering, Department of Built Environment, P.O. Box 14100, FI-00076 Aalto, Finland; ORCID https://orcid.org/0000-0002-4384-5279 E-mail: aarne.hovi@aalto.fi
  • Rautiainen, Aalto University, School of Engineering, Department of Built Environment, P.O. Box 14100, FI-00076 Aalto, Finland; Aalto University, School of Electrical Engineering, Department of Electronics and Nanoengineering, P.O. Box 14100, FI-00076 Aalto, Finland ORCID https://orcid.org/0000-0002-6568-3258 E-mail: miina.a.rautiainen@aalto.fi
article id 10524, category Research note
Arnis Gailis, Pauls Zeltiņš, Roberts Matisons, Andis Purviņš, Juris Augustovs, Valts Vīndedzis, Āris Jansons. (2021). Local adaptation of phenotypic stem traits distinguishes two provenance regions of silver birch in Latvia. Silva Fennica vol. 55 no. 2 article id 10524. https://doi.org/10.14214/sf.10524
Keywords: Betula pendula; population genetics; genetic parameters; climate-smart forestry; phenotypic traits
Highlights: Two provenance subregions in Latvia – coastal and inland – were distinguished; Silver birch populations in inland region possessed better growth, higher heritability, and phenotypic plasticity; Moderate to high heritability for stem quality was estimated in both regions; Silver birch from inland region possesses higher potential for improvement of adaptability.
Abstract | Full text in HTML | Full text in PDF | Author Info

Populations of tree species with a wide geographic range, such as silver birch (Betula pendula Roth), show genetic specialization to native environments, while maintaining high phenotypical plasticity. Accordingly, assessment of local specialization is essential for adaptive management. The aim of the study was to detect geographic patterns of local adaptation of growth and stem quality based on two open-pollinated progeny trials in Latvia testing local material. Two provenance regions differing by continentality were distinguished, which also differed in genetic control of growth traits, likely originating from the post-glacial recolonization of vegetation and subsequent natural adaptation. Heritability of the traits was estimated for each of the distinguished regions, indicating differing patterns of genetic adaptation and potential for future selection. Trees from the more continental inland showed superior growth and possessed higher heritability. The coastal provenance region showed slower growth and intermediate heritability of the respective traits. Moderate to high heritability for stem quality traits was estimated irrespectively of region. Overall, better growth and higher heritability suggests that anthropogenic selection within the best inland provenances may constitute better performing and adaptable breeding population compared to the coastal one. Still, overlapping phenotypical variation and heritability of quality traits implies improved stemwood quality for plywood regardless of the provenance region. High adaptive capacity of silver birch genotypes suggests ability to cope with climatic changes, highlighting its potential for climate-smart forestry.

  • Gailis, Latvian State Forest Research Institute Silava, 111 Rigas street, LV-2169, Salaspils, Latvia E-mail: arnis.gailis@silava.lv
  • Zeltiņš, Latvian State Forest Research Institute Silava, 111 Rigas street, LV-2169, Salaspils, Latvia E-mail: pauls.zeltins@silava.lv (email)
  • Matisons, Latvian State Forest Research Institute Silava, 111 Rigas street, LV-2169, Salaspils, Latvia ORCID https://orcid.org/0000-0002-4042-0689 E-mail: roberts.matisons@silava.lv
  • Purviņš, Latvian State Forest Research Institute Silava, 111 Rigas street, LV-2169, Salaspils, Latvia E-mail: andis.purvins@silava.lv
  • Augustovs, Latvian State Forest Research Institute Silava, 111 Rigas street, LV-2169, Salaspils, Latvia E-mail: juris.augustovs@silava.lv
  • Vīndedzis, Latvian State Forest Research Institute Silava, 111 Rigas street, LV-2169, Salaspils, Latvia E-mail: valts.vindedzis@silava.lv
  • Jansons, Latvian State Forest Research Institute Silava, 111 Rigas street, LV-2169, Salaspils, Latvia ORCID https://orcid.org/0000-0001-7981-4346 E-mail: aris.jansons@silava.lv
article id 10491, category Research note
Atte Komonen, Ilkka Puumala, Gergely Várkonyi, Reijo Penttilä. (2021). Wood-decaying fungi in old-growth boreal forest fragments: extinctions and colonizations over 20 years. Silva Fennica vol. 55 no. 1 article id 10491. https://doi.org/10.14214/sf.10491
Keywords: polypores; fragmentation; extinction debt; habitat loss; spruce forest
Highlights: Rare fungi can persist for decades in isolated old-growth forest fragments; The remaining old-growth forest fragments should be urgently protected, even if they are isolated from larger expanses of natural biotopes.
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According to ecology theory, isolated habitat fragments cannot maintain populations of specialized species. Yet, empirical evidence based on monitoring of the same fragments over time is still limited. We studied the colonization–extinction dynamics of eight wood-decaying fungal species in 16 old-growth forest fragments (<14 ha) over a 20-year period (1997–2017). We observed 19 extinctions and 5 colonizations; yet, the distribution of extinctions and colonizations did not differ from the one expected by chance for any of the species. Twenty-six percent of the extinctions took place in two natural fragments amid large forest–peatland complexes. Phellinus nigrolimitatus (Romell) Bourdot and Galzin decreased and Phellinus ferrugineofuscus (P. Karst.) Bourdot increased in abundance (number of logs occupied). The volume of living spruce trees in the forest fragments correlated positively with the number of logs inhabited in five of the study species. Because fragment characteristics did not affect species turnover, it seems that stochastic processes governed colonizations and extinctions. Although the least abundant species in 1997 had declined, and the most abundant species had become more abundant, it appears that specialized wood-decaying fungi can persist for decades in isolated old-growth forest fragments, if suitable dead wood is continuously available.

  • Komonen, University of Jyväskylä, Department of Biological and Environmental Science, School of Resource Wisdom, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: atte.komonen@jyu.fi (email)
  • Puumala, University of Jyväskylä, Department of Biological and Environmental Science, School of Resource Wisdom, P.O. Box 35, FI-40014 University of Jyväskylä, Finland E-mail: ilkka.puumala1@outlook.com
  • Várkonyi, Finnish Environment Institute, Friendship Park Research Centre, Lentiirantie 342 B, FI-88900 Kuhmo, Finland E-mail: gergely.varkonyi@ymparisto.fi
  • Penttilä, Natural Resources Institute Finland, Natural resources, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: reijo.penttila@luke.fi
article id 10263, category Research note
Paul N. Katjutin, Natalia I. Stavrova, Vadim V. Gorshkov, Andrew Yu. Lyanguzov, Irina Ju. Bakkal, Sergey A. Mikhailov. (2020). Radial growth of trees differing in their vitality in the middle-aged Scots pine forests in the Kola peninsula. Silva Fennica vol. 54 no. 3 article id 10263. https://doi.org/10.14214/sf.10263
Keywords: Pinus sylvestris; northern taiga; radial increment; basal area increment
Highlights: Unmanaged middle-aged boreal Scots pine forests in the Kola peninsula are characterised by the prevalence of moderately and strongly weakened trees (65–70%); Radial increment and basal area increment differ greatly (70–75% and 85–90%, respectively) between Scots pine trees differing in their vitality; The trends of annual ring width in Scots pine trees aged from 15–35 to 65–85 years in green moss and green moss-lichen type pine forests are similar; the dynamics of basal area increment differs substantially in relation to forest site type.
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The research was carried out in unmanaged middle-aged (75–85 years) Northern taiga Scots pine (Pinus sylvestris L.) forests in the Kola peninsula. It was established that forests of green moss-lichen and green moss site types are characterised by a predominance (65–70% by stand volume) of moderately and strongly weakened trees. Trees of differing vitality have significant differences in annual increment. Healthy trees had a radial increment (RI) 70–75% greater than that of dying trees, and a basal area increment (BAI) 85–90% greater. The dynamics of the RI and BAI of Scots pine trees for the 70-year period (from 1945 to 2015) is different. The RI of all individuals in the communities studied decreases consistently. The decrease is expressed more strongly in green moss Scots pine forests (80–95% from 1945 to 2015) compared to green moss-lichen forests (60–80%); it manifests itself more in strongly weakened and dying individuals (75–95%) than in healthy and moderately weakened ones (60–80%). Annual basal area increment in green moss Scots pine forests increases by 45–65% from stand establishment until the trees are 25 to 35 years old and subsequently decreases by 50–80% to 70–80 years of age. In green moss-lichen pine forests the BAI of Scots pine remains rather stable in healthy and moderately weakened trees and decreases in strongly weakened and dying individuals by 45% and 75–80%, respectively throughout the studied period.

  • Katjutin, Komarov Botanical Institute of the Russian Academy of Sciences, Professora Popova str. 2, 197376, Saint-Petersburg, Russia E-mail: paurussia@binran.ru (email)
  • Stavrova, Komarov Botanical Institute of the Russian Academy of Sciences, Professora Popova str. 2, 197376, Saint-Petersburg, Russia E-mail: nstavrova@binran.ru
  • Gorshkov, Komarov Botanical Institute of the Russian Academy of Sciences, Professora Popova str. 2, 197376, Saint-Petersburg, Russia; Saint-Petersburg State Forest Technical University, letter U, 5, Institutsky per., 194021, Saint-Petersburg, Russia E-mail: vgorshkov@binran.ru
  • Lyanguzov, Saint-Petersburg State University, 7/9 Universitetskaya Emb., 199034, Saint-Petersburg, Russia E-mail: andrewlyanguzov@gmail.com
  • Bakkal, Komarov Botanical Institute of the Russian Academy of Sciences, Professora Popova str. 2, 197376, Saint-Petersburg, Russia E-mail: bakkal@binran.ru
  • Mikhailov, Komarov Botanical Institute of the Russian Academy of Sciences, Professora Popova str. 2, 197376, Saint-Petersburg, Russia E-mail: smikhailov@binran.ru
article id 10262, category Research note
Markus Melin, Heli Viiri, Olli-Pekka Tikkanen, Riku Elfving, Seppo Neuvonen. (2020). From a rare inhabitant into a potential pest – status of the nun moth in Finland based on pheromone trapping. Silva Fennica vol. 54 no. 1 article id 10262. https://doi.org/10.14214/sf.10262
Keywords: climate change; forest health; forestry; Lymantria monacha; forest damage; insect; range expansion
Highlights: The nun moth is a significant defoliator of coniferous forests in Central-Europe; In Finland, the populations have grown and expanded northwards; Pheromone trapping confirmed the species’ presence throughout central- and southern Finland; The risk of the nun moth becoming a pest for Finland is real as the area offers endless habitats, and climatic conditions are becoming more favourable; This note describes the results from the first nun moth surveys conducted in 2018 and 2019.
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Forests are affected by climate change in various ways. This includes abiotic factors such as droughts, but also biotic damage by pest insects. There are numerous examples from cases where pest insects have benefitted from longer growing seasons or from warmer summers. Similarly, new pest insects have been able to expand their range due to climatic conditions that have changed from hostile to tolerable. Such seems to be the case with the nun moth (Lymantria monacha), an important defoliator of coniferous trees in Europe. For centuries, the species has had massive outbreaks across Central-Europe, while it has been a rare inhabitant in Northern Europe. Recently, the nun moth population in Finland has not only expanded in range, but also grown more abundant. This research note describes the results from the first years (2018–2019) of a monitoring program that is being conducted with pheromone traps across central and southern Finland. So far, the northernmost individuals were trapped near the 64 N degrees. However, there were more southern locations where no moths were trapped. The species was present in every trapping site below the latitude of 62 N degrees. More importantly, at some sites the abundance of the nun moth suggested that local forest damage may already occur. Given the current climatic scenarios for Fennoscandia, it is likely that the nun moth populations will continue to grow, which is why systematic surveys on their abundance and range expansions will be topical.

  • Melin, Natural Resources Institute Finland, Yliopistokatu 6b, FI-80100 Joensuu, Finland ORCID https://orcid.org/0000-0001-7290-9203 E-mail: markus.melin@luke.fi (email)
  • Viiri, Natural Resources Institute Finland, Yliopistokatu 6b, FI-80100 Joensuu, Finland; UPM-Kymmene Oyj, UPM Forest, Åkerlundinkatu 11 B, FI-33100 Tampere, Finland E-mail: heli.viiri@upm.com
  • Tikkanen, University of Eastern Finland, School of Forest Sciences, Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: olli-pekka.tikkanen@uef.fi
  • Elfving, Natural Resources Institute Finland, Yliopistokatu 6b, FI-80100 Joensuu, Finland; University of Oulu, Department of Biology, Pentti Kaiteran katu 1, FI-90014 Oulu, Finland E-mail: riku.elfving@gmail.com
  • Neuvonen, University of Turku, Biodiversity Unit, Kevo Subarctic Research Institute, FI-20014 Turku, Finland E-mail: seppo.neuvonen@utu.fi
article id 10197, category Research note
Ville Kankare, Ville Luoma, Ninni Saarinen, Jussi Peuhkurinen, Markus Holopainen, Mikko Vastaranta. (2019). Assessing feasibility of the forest trafficability map for avoiding rutting – a case study. Silva Fennica vol. 53 no. 3 article id 10197. https://doi.org/10.14214/sf.10197
Keywords: remote sensing; open data; preharvest information; stand trafficability
Highlights: A static trafficability map was developed to provide information about suitable harvesting season; The majority (91.7%) of the evaluated thinning stands were harvested without causing rutting damage if operations were timed correctly in relation to the static trafficability map information; The static trafficability map provides reliable and slightly conservative estimation of the forest trafficability for supporting forest operations.
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Information on forest trafficability (i.e. carrying capacity of the forest floor) is required before harvesting operations in Southern Boreal forest conditions. It describes the seasons when harvesting operations may take place without causing substantial damage to the forest soil using standard logging machinery. The available trafficability information have been based on subjective observations made during the wood procurement planning. For supporting forest operations, an open access map product has been developed to provide information on trafficability of forests. The forest stands are distributed into classes that characterize different harvesting seasons based on topographic wetness index, amount of vegetation, ground water height and ditch depth. The main goal of this case study was to evaluate the information of the static forest trafficability map in relation to the detected rutting within logging tracks measured in the field. The analysis concentrated on thinning stands since the effect of rutting is significant on the growth of the remaining trees. The results showed that the static trafficability map provided reliable and slightly conservative estimation of the forest trafficability. The majority (91.7%) of the evaluated stands were harvested without causing significant damage if harvesting was timed correctly compared to the trafficability information. However, it should be pointed out that the weather history at small scale, the skills of a driver, and effects of used machinery are not considered in the map product although they can have a considerable impact on the rutting.

  • Kankare, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, Joensuu FI-80101, Finland; Department of Forest Sciences, University of Helsinki, FI-00014 University of Helsinki, Finland ORCID https://orcid.org/0000-0001-6038-1579 E-mail: ville.kankare@uef.fi (email)
  • Luoma, Department of Forest Sciences, University of Helsinki, FI-00014 University of Helsinki, Finland E-mail: ville.luoma@helsinki.fi
  • Saarinen, Department of Forest Sciences, University of Helsinki, FI-00014 University of Helsinki, Finland; School of Forest Sciences, University of Eastern Finland, P.O. Box 111, Joensuu FI-80101, Finland E-mail: ninni.saarinen@helsinki.fi
  • Peuhkurinen, Arbonaut Oy, Malminkaari 13–19, FI-00700 Helsinki, Finland E-mail: jussi.peuhkurinen@arbonaut.com
  • Holopainen, Department of Forest Sciences, University of Helsinki, FI-00014 University of Helsinki, Finland E-mail: markus.holopainen@helsinki.fi
  • Vastaranta, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, Joensuu FI-80101, Finland E-mail: mikko.vastaranta@uef.fi
article id 10195, category Research note
Tiina Laine, Leena Hamberg, Veli-Matti Saarinen, Timo Saksa. (2019). The efficacy of Chondrostereum purpureum against sprouting of deciduous species after mechanized pre-commercial thinning. Silva Fennica vol. 53 no. 3 article id 10195. https://doi.org/10.14214/sf.10195
Keywords: vegetation management; silviculture; mechanization; Chondrostereum purpureum; fungal treatment; stump sprouts
Highlights: Fungal treatments increased stump mortality compared to the control (cutting only); The fungal treatment did not decrease the number of sprouts per stump; Application during mechanized pre-commercial thinning did not yield as high stump mortalities as in earlier treatments performed manually.
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The use of a white-rot fungus, Chondrostereum purpureum (Pers. Ex Fr.) Pouzar, as a biocontrol agent against sprouting has been studied with good results. The aim of the study was to investigate the efficacy of two pre-commercial thinning machines, Tehojätkä and Mense, to spread an inoculum of C. purpureum as a biocontrol agent on freshly cut birch (Betula pendula Roth and B. pubescens Ehrh.), European aspen (Populus tremula L.), rowan (Sorbus aucuparia L.), and goat willow (Salix caprea L.) stumps (the fungal treatment) and compare that to the control (cutting only, done by Tehojätkä). Efficacy was investigated in terms of stump mortality and the number of sprouts per stump. This study was conducted in one stand and sprouting was investigated for three years after treatment. The fungal treatment resulted in higher mortality of stumps (34.0% for Tehojätkä and 41.5% for Mense after three years), compared to the control (13.4%). However, the fungal treatment did not decrease the number of sprouts per stump compared to the control. The low occurrence of basidiomata indicates that the accuracy of the spreading mechanism was not satisfactory, causing low mortality figures for the fungal treatment compared to previous studies. In the future, this mechanized method may provide a promising alternative in sprout control if the spreading mechanisms, the accuracy of the treatment, and consequently the efficacy could be improved.

  • Laine, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: tiina.laine@luke.fi (email)
  • Hamberg, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: leena.hamberg@luke.fi
  • Saarinen, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: mulinvuori@gmail.com
  • Saksa, Natural Resources Institute Finland (Luke), Natural resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: timo.saksa@luke.fi
article id 10012, category Research note
Irving U. Hernández-Gómez, Carlos R. Cerdán, Angélica Navarro-Martínez, Dinora Vázquez-Luna, Samaria Armenta-Montero, Edward A. Ellis. (2019). Assessment of the CLASlite forest monitoring system in detecting disturbance from selective logging in the Selva Maya, Mexico. Silva Fennica vol. 53 no. 1 article id 10012. https://doi.org/10.14214/sf.10012
Keywords: community forestry; forest degradation; tropical forest; Yucatan Peninsula
Highlights: The accuracy of CLASlite to detect forest disturbance from selective logging using Landsat imagery was very low (<19.1%); Selective logging impacts was only detected in one case with the highest logging intensity (7 m3 ha–1); CLASlite shows potential in monitoring forest disturbance from tree biomass impacts greater than 900 m2.
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Detecting and monitoring forest disturbance from selective logging is necessary to develop effective strategies and polices that conserve tropical forests and mitigate climate change. We assessed the potential of using the remote sensing tool, CLASlite forest monitoring system, to detect disturbance from timber harvesting in four community forests (ejidos) of the Selva Maya on the Yucatan Peninsula, Mexico. Selective logging impacts (e.g. felling gaps, skid trails, logging roads and log landings) were mapped using GPS in the 2014 annual cutting areas (ACAs) of each ejido. We processed and analyzed two pre-harvest Landsat images (2001 and 2013) and one post-harvest image (November 2014) with the CLASlite system, producing maps of degraded, deforested and unlogged areas in each ACA. Based on reference points of disturbed (felling and skidding), deforested (log landings and roads) and unlogged areas in each ACA, we applied accuracy assessments which showed very low overall accuracies (<19.1%). Selective logging impacts, mainly from log landings and new logging road construction, were detected in only one ejido which had the highest logging intensity (7 m3 ha–1).

  • Hernández-Gómez, Facultad de Ciencias Agrícolas, Universidad Veracruzana. Circuito Gonzalo Aguirre Beltrán, Isleta, Xalapa, Veracruz. C.P. 91000, Mexico E-mail: urielxal@gmail.com
  • Cerdán, Facultad de Ciencias Agrícolas, Universidad Veracruzana. Circuito Gonzalo Aguirre Beltrán, Isleta, Xalapa, Veracruz. C.P. 91000, Mexico E-mail: ccerdan@uv.mx
  • Navarro-Martínez, El Colegio de la Frontera Sur Av. Centenario km 5.5, Col. Pacto Obrero Campesino s/n. Chetumal, Quintana Roo. C.P. 77014, Mexico E-mail: manavaster@gmail.com
  • Vázquez-Luna, Facultad de Ingeniería en Sistemas de Producción Agropecuaria, Universidad Veracruzana. Carretera Costera del Golfo Km. 220, C. Agrícola y Ganadera Michapan, Acayucan, Veracruz. C.P. 96000, Mexico E-mail: divazquez@uv.mx
  • Armenta-Montero, Centro de Investigaciones Tropicales (CITRO), Universidad Veracruzana. Morelos No. 44 y 46, Zona Centro, Xalapa, Veracruz. C.P. 91000, Mexico E-mail: samaria.am@gmail.com
  • Ellis, Centro de Investigaciones Tropicales (CITRO), Universidad Veracruzana. Morelos No. 44 y 46, Zona Centro, Xalapa, Veracruz. C.P. 91000, Mexico E-mail: eellis@uv.mx (email)
article id 10014, category Research note
Āris Jansons, Roberts Matisons, Virgilijus Baliuckas, Līga Purina, Oskars Krišāns, Jānis Jansons, Imants Baumanis. (2018). Performance variation of lodgepole pine provenances in Latvia. Silva Fennica vol. 52 no. 5 article id 10014. https://doi.org/10.14214/sf.10014
Keywords: provenance trial; Pinus contorta var. latifolia; growth traits; transfer; spike knots
Highlights: Performance of 36 provenances of lodgepole pine in 14 trials was studied; The 29 year survival was ca. 40%; Provenance and provenance × trial interaction affected dimensions of lodgepole pine; Provenances from lower latitudes were the most productive.
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Lodgepole pine (Pinus contorta var. latifolia Engelm.) has been extensively introduced in Scandinavia on less productive sites. Under a changing climate, it also has a high potential in the eastern Baltic region; still, its performance there has scarcely been reported. This study investigated the performance of 36 Canadian provenances in 14 trials in western Latvia. Tree dimensions showed notable provenance and provenance-by-environment variation, implying that local selection by provenance can be applied for improved yield. Southern provenances showed the best height growth, while southwestern (more oceanic) provenances excelled in diameter growth. Most of the quality traits were affected by provenance or provenance-by-environment interaction, yet the variation was lower than for the growth traits.

  • Jansons, Latvian State Forest Research Institute ”Silava”, Department of Forest Tree Breeding, Rigas St.t. 111, Salaspils LV-2169, Latvia E-mail: aris.jansons@silava.lv (email)
  • Matisons, Latvian State Forest Research Institute ”Silava”, Department of Forest Tree Breeding, Rigas St.t. 111, Salaspils LV-2169, Latvia E-mail: robism@inbox.lv
  • Baliuckas, Forest Institute, Lithuanian Centre for Agriculture and Forestry, Department of Forest Tree Genetics and Breeding, Liepu St. 1, Girionys, LT-53101 Kaunas distr., Lithuania E-mail: virgilijus.baliuckas@mi.lt
  • Purina, Latvian State Forest Research Institute ”Silava”, Department of Forest Tree Breeding, Rigas St.t. 111, Salaspils LV-2169, Latvia E-mail: liga.purina@silava.lv
  • Krišāns, Latvian State Forest Research Institute ”Silava”, Department of Forest Tree Breeding, Rigas St.t. 111, Salaspils LV-2169, Latvia E-mail: oskars.krisans@silava.lv
  • Jansons, Latvian State Forest Research Institute ”Silava”, Department of Forest Tree Breeding, Rigas St.t. 111, Salaspils LV-2169, Latvia E-mail: janis.jansons.silava@gmail.com
  • Baumanis, Latvian State Forest Research Institute ”Silava”, Department of Forest Tree Breeding, Rigas St.t. 111, Salaspils LV-2169, Latvia E-mail: imants.baumanis@silava.lv
article id 10009, category Research note
Jānis Donis, Māra Kitenberga, Guntars Šņepsts, Edgars Dubrovskis, Āris Jansons. (2018). Factors affecting windstorm damage at the stand level in hemiboreal forests in Latvia: case study of 2005 winter storm. Silva Fennica vol. 52 no. 4 article id 10009. https://doi.org/10.14214/sf.10009
Keywords: Norway spruce; natural disturbance; windstorm; hemiboreal
Highlights: In hemiboreal forests in Latvia, dominant tree species, admixture of spruce in canopy-layer, mean height, timing of thinnings, upwind forest edges and wind gusts had significant effect on windstorm damage occurrence at stand-level; Stands on peat soils were more damaged than stands on mineral soils; Tree species composition of canopy-layer was not statistically significant in the model.
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In managed European hemiboreal forests, windstorms have a notable ecological and socio-economic impact. In this study, stand properties affecting windstorm damage occurrence at the stand-level were assessed using a Generalized Linear Mixed model. After 2005 windstorm, 5959 stands dominated by birch (Betula spp.), Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.), with mean height > 10 m were inventoried. Windstorm damage was positively associated with spruce and pine-dominated stands, increasing mean height, fresh forest edges, decreasing time since the last thinning and stronger wind gusts. Tree species composition – mixed or monodominant – was not statistically significant in the model; while, the admixture of spruce in the canopy layer was positively associated with higher windstorm damage. Stands on peat soils were more damaged than stands on mineral soils. Birch stands were more damaged than pine stands. This information could be used in forest management planning, selection of silvicultural treatments to increase forest resilience to natural disturbances.

  • Donis, Latvian State Forest Research Institute “Silava”, Rigas 111, Salaspils, Latvia, LV-2169 E-mail: janis.donis@silava.lv (email)
  • Kitenberga, Latvian State Forest Research Institute “Silava”, Rigas 111, Salaspils, Latvia, LV-2169 E-mail: mara.kitenberga@gmail.com
  • Šņepsts, Latvian State Forest Research Institute “Silava”, Rigas 111, Salaspils, Latvia, LV-2169 E-mail: guntars.snepsts@silava.lv
  • Dubrovskis, Latvia University of Life Sciences and Technologies, Liela iela 2, Jelgava, Latvia, LV-3001 E-mail: edgars.dubrovskis@llu.lv
  • Jansons, Latvian State Forest Research Institute “Silava”, Rigas 111, Salaspils, Latvia, LV-2169 E-mail: aris.jansons@silava.lv
article id 9986, category Research note
Ninni Saarinen, Joanne C. White, Michael A. Wulder, Annika Kangas, Sakari Tuominen, Ville Kankare, Markus Holopainen, Juha Hyyppä, Mikko Vastaranta. (2018). Landsat archive holdings for Finland: opportunities for forest monitoring. Silva Fennica vol. 52 no. 3 article id 9986. https://doi.org/10.14214/sf.9986
Keywords: National Forest Inventory; satellite; Landsat time series
Highlights: The 45-year Landsat archive contained 30 076 images for Finland by December 31, 2017; 16.3% of these were acquired within ±30 days of August 1 (northern hemisphere summer), have <70% cloud cover, and a 30 m spatial resolution; Using time series analyses, these data provide unique information that complements other datasets available for forest monitoring and assessment in Finland.
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There is growing interest in the use of Landsat data to enable forest monitoring over large areas. Free and open data access combined with high performance computing have enabled new approaches to Landsat data analysis that use the best observation for any given pixel to generate an annual, cloud-free, gap-free, surface reflectance image composite. Finland has a long history of incorporating Landsat data into its National Forest Inventory to produce forest information in the form of thematic maps and small area statistics on a variety of forest attributes. Herein we explore the spatial and temporal characteristics of the Landsat archive in the context of forest monitoring in Finland. The United States Geological Survey Landsat archive holds a total of 30 076 images (1972–2017) for 66 scenes (each 185 km by 185 km in size) representing the terrestrial area of Finland, of which 93.6% were acquired since 1984 with a spatial resolution of 30 m. Approximately 16.3% of the archived images have desired compositing characteristics (acquired within August 1 ±30 days, <70% cloud cover, 30 m spatial resolution). Data from the Landsat archive can augment forest monitoring efforts in Finland, provide new information for science and applications, and enable retrospective, systematic analyses to characterize the development of Finnish forests over the past three decades. The capacity to monitor trends based upon this multi-decadal record with the addition of new measurements is of benefit to multisource inventories and offers nationally comprehensive spatially-explicit datasets for a wide range of stakeholders and applications.

  • Saarinen, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014 University of Helsinki, Finland; School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0003-2730-8892 E-mail: ninni.saarinen@helsinki.fi (email)
  • White, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014 University of Helsinki, Finland; Canadian Forest Service, (Pacific Forestry Center), Natural Resources Canada, 506 West Burnside Road, Victoria, BC, V8Z 1M5, Canada ORCID http://orcid.org/0000-0003-4674-0373 E-mail: joanne.white@canada.ca
  • Wulder, Canadian Forest Service, (Pacific Forestry Center), Natural Resources Canada, 506 West Burnside Road, Victoria, BC, V8Z 1M5, Canada ORCID https://orcid.org/0000-0002-6942-1896 E-mail: mike.wulder@canada.ca
  • Kangas, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Yliopistokatu 6, FI-80100 Joensuu, Finland E-mail: annika.kangas@luke.fi
  • Tuominen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: sakari.tuominen@luke.fi
  • Kankare, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ville.kankare@helsinki.fi
  • Holopainen, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: markus.holopainen@helsinki.fi
  • Hyyppä, Department of Remote Sensing and Photogrammetry, Finnish Geospatial Research Institute, National Land Survey of Finland, Geodeetinrinne 2, FI-02431 Masala, Finland E-mail: juha.hyyppa@nls.fi
  • Vastaranta, School of Forest Sciences, University of Eastern Finland, P.O. Box 111, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0001-6552-9122 E-mail: mikko.vastaranta@uef.fi
article id 9911, category Research note
Tālis Gaitnieks, Indulis Brauners, Kristīne Kenigsvalde, Astra Zaļuma, Lauma Brūna, Jurģis Jansons, Natālija Burņeviča, Andis Lazdiņš, Rimvydas Vasaitis. (2018). Infection of pre-commercially cut stumps of Picea abies and Pinus sylvestris by Heterobasidion spp. – a comparative study. Silva Fennica vol. 52 no. 1 article id 9911. https://doi.org/10.14214/sf.9911
Keywords: Norway spruce; Scots pine; pre-commercial thinnings; Heterobasidion
Highlights: In pre-commercial thinnings both Heterobasidion infection frequency and the extent of surface colonization correlated positively with stump diameter of both Norway spruce and Scots pine; Spruce stumps were significantly more often subjected to primary infections than pine stumps; The pathogen exhibited more extensive surface colonization of spruce stumps than of pine stumps.
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The aim was to investigate relative susceptibility of stumps of spruce and pine to airborne infections by Heterobasidion following pre-commercial thinnings. The proportions of infected stumps and colonized stump surface areas were analysed in 16 forest stands. In total, 746 spruce and 1063 pine stumps were sampled, and 184 and 105 infected stumps, respectively, were analysed. In conclusion, the present study demonstrated that in the investigated area: i) both Heterobasidion infection frequency and the extent of surface colonization correlated positively with stump diameter of both spruce and pine; ii) spruce stumps were significantly more often subjected to primary infections than pine stumps; iii) the pathogen exhibited more extensive surface colonization of spruce stumps than of pine stumps.

  • Gaitnieks, Latvian State Forest Research Institute “Silava”, 111 Rigas str., Salaspils, LV-2169, Latvia E-mail: talis.gaitnieks@silava.lv (email)
  • Brauners, JSC “Latvia’s State Forests”, 1 Vainodes str., Riga, LV-1004, Latvia E-mail: i.brauners@lvm.lv
  • Kenigsvalde, Latvian State Forest Research Institute “Silava”, 111 Rigas str., Salaspils, LV-2169, Latvia E-mail: kristine.kenigsvalde@silava.lv
  • Zaļuma, Latvian State Forest Research Institute “Silava”, 111 Rigas str., Salaspils, LV-2169, Latvia E-mail: astra.zaluma@silava.lv
  • Brūna, Latvian State Forest Research Institute “Silava”, 111 Rigas str., Salaspils, LV-2169, Latvia E-mail: lauma.bruna@silava.lv
  • Jansons, Latvian State Forest Research Institute “Silava”, 111 Rigas str., Salaspils, LV-2169, Latvia E-mail: jurgis.jansons@silava.lv
  • Burņeviča, Latvian State Forest Research Institute “Silava”, 111 Rigas str., Salaspils, LV-2169, Latvia E-mail: natalija.arhipova@silava.lv
  • Lazdiņš, Latvian State Forest Research Institute “Silava”, 111 Rigas str., Salaspils, LV-2169, Latvia E-mail: andis.lazdins@silava.lv
  • Vasaitis, Swedish University of Agricultural Sciences (SLU), P.O. Box 7026, SE-75007 Uppsala, Sweden E-mail: Rimvys.Vasaitis@slu.se
article id 9901, category Research note
Ilze Matisone, Roberts Matisons, Māris Laiviņš, Tālis Gaitnieks. (2018). Statistics of ash dieback in Latvia. Silva Fennica vol. 52 no. 1 article id 9901. https://doi.org/10.14214/sf.9901
Keywords: mortality rate; National Forest Inventory; Fraxinus excelsior; permanent sampling plots
Highlights: Dynamics of ash dieback during 2005–2015 were summarized; The area of ash dominated stands decreased twofold; The number of mature ash trees and their standing volume decreased by 53.1 and 69.9%, compared to 2005, respectively; The mortality of trees was higher during the first part of the survey, the decrease of standing volume culminated later.
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Dieback of the common ash (Fraxinus excelsior L.) has been spreading throughout Europe since the 1990s, causing severe ecological and economical consequences; however, detailed statistics on its dynamics have been published rarely. This paper presents the dynamics of mature ash-dominated stands in Latvia for the period 2005–2015. Data from the national forest inventory and a permanent sampling plot network were summarised. According to the official statistics, the dieback has caused a twofold decrease in area of the ash stands (from 21 891 to 13 011 ha, which respectively comprised ca. 0.8 to ca. 0.4% of the total forest area). The official statistics on standing volume appeared biased, as they did not account for increased mortality. According to the permanent sampling plots, standing volume and stand density have been affected even more, having decreased by 53.1 and 69.9%, respectively, compared to 2005 (the stand density and standing volume of ash in 2015 was 77 individuals ha–1 and 151 m3 ha–1, respectively). The mortality of the trees has not been stable. Stand density decreased faster during 2005–2009 compared to 2010–2015, with mortality rates of 9.6 and 8.2% year–1, respectively. In contrast, the decrease in standing volume in 2005–2009 was slower than in 2010–2015 (mortality rates were 4.7 and 7.7% year–1, respectively) because trees with smaller dimensions were more susceptible to the dieback. Nevertheless, the observed mortality rates clearly indicate negative prospects for ash stands in Latvia.

  • Matisone, Latvian State Forest Research Institute (LSFRI) Silava, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: ilze.matisone@silava.lv (email)
  • Matisons, Latvian State Forest Research Institute (LSFRI) Silava, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: robism@inbox.lv
  • Laiviņš, Latvian State Forest Research Institute (LSFRI) Silava, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: maris.laivins@silava.lv
  • Gaitnieks, Latvian State Forest Research Institute (LSFRI) Silava, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: talis.gaitnieks@silava.lv
article id 9899, category Research note
Linda Robalte, Diāna Jansone, Didzis Elferts, Roberts Matisons, Āris Jansons. (2018). Bilberry ramet dimensions in relation to stand age in oligotrophic conditions in Latvia. Silva Fennica vol. 52 no. 1 article id 9899. https://doi.org/10.14214/sf.9899
Keywords: structural diversity; Vaccinium myrtillus; dwarf shrubs; projective cover; hemiboreal forests
Highlights: Bilberry ramet dimensions (age, diameter, height) and their structural diversity, as well as cover, increased with stand age; Active rejuvenation of ramets was observed in younger stands; The oldest bilberry ramets (>10 years of age) occurred in stands older than 70 years.
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Dwarf shrub layer is an important component of boreal and hemiboreal forest ecosystems that has received little attention, particularly regarding its structural diversity, which, however, could serve as an additional proxy for habitat quality. Dimensions of bilberry (Vaccinium myrtillus L.) ramets were assessed in two sites in Latvia covered by dry oligotrophic Scots pine (Pinus sylvestris L.) stands 10–230 years of age. In total, 20 sampling plots (10×10 m) with 156 subplots (1×1 m) were sampled and 630 bilberry ramets analysed. The dimensions of ramets (age, diameter, and height) and cover of bilberry increased with stand age. The age of the studied ramets ranged 2–13 years; 5–6 years-old ramets were most frequent in all stands. The skewness of the distribution of the ramet dimensions shifted with stand age, leaning towards the higher values. Lower structural diversity of ramets was observed in stands 50–100 years of age. The highest diversity of ramet age structure occurred in stands younger than 150 years, whereas the oldest and largest ramets mostly occurred in the older stands (>150 years). Considering structural diversity of ramets, recovery of bilberry after stand-replacing disturbance (e.g. clearcut) was a continuous process, similarly to that observed in tree layer.

  • Robalte, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: robalte.l@gmail.com (email)
  • Jansone, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia; University of Latvia, Faculty of Biology, Jelgavas Str. 1, LV 1004, Riga, Latvia E-mail: diana.jansone13@gmail.com
  • Elferts, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia; University of Latvia, Faculty of Biology, Jelgavas Str. 1, LV 1004, Riga, Latvia E-mail: didzis.elferts@lu.lv
  • Matisons, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: roberts.matisons@silava.lv
  • Jansons, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: aris.jansons@silava.lv
article id 7818, category Research note
Roberts Matisons, Guntars Šņepsts, Līga Puriņa, Jānis Donis, Āris Jansons. (2018). Dominant height growth of European beech at the northeasternmost stands in Europe. Silva Fennica vol. 52 no. 1 article id 7818. https://doi.org/10.14214/sf.7818
Keywords: site index; Fagus sylvatica; generalized algebraic difference approach; height models
Highlights: The dominant height growth of the introduced European beech was modelled using the generalised algebraic difference approach; The Chapman-Richards and Sloboda models showed the best fit to the data; Height growth of the second generation trees exceeded the first generation trees; In the western part of Latvia, height growth of beech exceeded that in southern Sweden.
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The height growth of trees influences the productivity of stands and the competitiveness of species, shaping the range of their distribution. Dominant height growth was assessed for European beech (Fagus sylvatica L.), growing outside of its natural distribution range in the western part of Latvia. In 10 neighbouring experimental stands, 20 dominant trees were felled for stem analysis. Height growth was modelled using the generalised algebraic difference approach, applying several non-linear equations and mixed procedures. The Chapman-Richards and Sloboda models showed the best fit to the data. Height growth of the second generation (younger) trees exceeded that of the first generation, and followed curve for a higher site index, suggesting sufficient adaptation and improving conditions. Height growth of the studied beech exceeded predictions for beech in southern Sweden, which is considered to be the northern limit of the species range, yet the growth pattern differed. In Latvia, slower height growth was estimated for site indices < 32 m (in 100 years) during the first 60 years, yet larger maximal height was predicted, suggesting a longer establishment period. Nevertheless, the improving height growth indicated increasing potential for the application of the species in commercial forestry, and an expansion of the species within the region even during the 21th century.

  • Matisons, Latvian State Forest Research Institute “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 E-mail: robism@inbox.lv (email)
  • Šņepsts, Latvian State Forest Research Institute “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 E-mail: guntars.snepsts@silava.lv
  • Puriņa, Latvian State Forest Research Institute “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 E-mail: liga.purina@silava.lv
  • Donis, Latvian State Forest Research Institute “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 E-mail: janis.donis@silava.lv
  • Jansons, Latvian State Forest Research Institute “Silava”, Rīgas str. 111, Salaspils, Latvia, LV2169 E-mail: aris.jansons@silava.lv
article id 7771, category Research note
Māra Kitenberga, Roberts Matisons, Āris Jansons, Jānis Donis. (2018). Teleconnection between the Atlantic sea surface temperature and forest fires in Latvia and Estonia. Silva Fennica vol. 52 no. 1 article id 7771. https://doi.org/10.14214/sf.7771
Keywords: disturbance regime; Atmospheric circulation; weather systems; fire meteorology
Highlights: Forest fire activity in Latvia and Estonia was related to conditions in the Atlantic; Teleconnections differed regionally; Negative correlation between number of fires in Estonia and SST in the North Atlantic was detected; Area of forest fires in Estonia and activity of fires in Latvia were positively correlated with SST in the Baltic, North and Mediterranean Seas in summer.
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Forest fire is one of the natural disturbances, which have important ecological and socioeconomical effect. Although fire activity is driven by weather conditions, during past two centuries forest fires have been strongly anthropogenically controlled. In this study, teleconnection between sea surface temperature (SST) in the Atlantic, which influences climate in Europe, and forest fire activity in Latvia and Estonia was assessed using “Climate explorer” web-tool. Factors affecting number and area of forest fires in Latvia and Estonia differed, suggesting regional specifics. In Estonia, the number of fires correlated with the SST in the North Atlantic in spring and summer, which affects the inflow of cool and dry air masses from the Arctic, hence the aridity and burnability. The area of fires in Estonia and in Latvia was associated with increased SST in Baltic Sea and near the European coast in summer, which likely were consequences of occurrence of warm high-pressure systems in summer, causing hot and dry conditions. Nevertheless, the observed teleconnections could be used to predict activity of forest fires in Latvia and Estonia.

  • Kitenberga, Latvian State Forest Research Institute ‘Silava’, Rigas st. 111, Salaspils, Latvia, LV2169 E-mail: mara.kitenberga@gmail.com (email)
  • Matisons, Latvian State Forest Research Institute ‘Silava’, Rigas st. 111, Salaspils, Latvia, LV2169 E-mail: robism@inbox.lv
  • Jansons, Latvian State Forest Research Institute ‘Silava’, Rigas st. 111, Salaspils, Latvia, LV2169 E-mail: aris.jansons@silava.lv
  • Donis, Latvian State Forest Research Institute ‘Silava’, Rigas st. 111, Salaspils, Latvia, LV2169 E-mail: janis.donis@silava.lv
article id 7798, category Research note
Jānis Liepiņš, Jānis Ivanovs, Andis Lazdiņš, Jurģis Jansons, Kaspars Liepiņš. (2017). Mapping of basic density within European aspen stems in Latvia. Silva Fennica vol. 51 no. 5 article id 7798. https://doi.org/10.14214/sf.7798
Keywords: Populus tremula; wood quality; wood density; bark density; density variations
Highlights: Stem bark is significantly denser than wood and does not follow the same variation patterns along the stem; The main trend in radial variation of wood density was the increase from pith to bark; There is a weak relationship between mean basic density and commonly measured stand and tree parameters.
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The objective of this study was to investigate basic density and its within-stem variation by studying 84 European aspen stems from 28 forest stands in Latvia. The studied forest stands covered all age classes from young stands to matured forests in representative growth conditions of European aspen. The densities of 2722 wood and 1022 bark specimens were measured from the sampled trees. Only the knot-free wood specimens without obvious wood defects were chosen for analyses. A map of basic density summarizing its radial and axial variations was constructed to show species-specific, within-stem variability and the relationships between density and tree and stand variables were examined. Stem wood and bark of the European aspen show different patterns of basic density variation along the tree stem. Wood density increases from pith to bark up to certain dimensions and shows a slight decrease afterwards. The weighted basic density of bark (446 ± 39.6 kg m–3) was higher than stem wood density (393 ± 30.4 kg m–3). Our results suggest that wood and bark density measurements obtained at breast height can be used for reliable estimation of the densities of whole-tree stem components, while tree parameters such as diameter at breast height (DBH), tree height and social status or stand parameters, including number of trees, basal area and age, are weak predictors in this context.

  • Liepiņš, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: janis.liepins@silava.lv (email)
  • Ivanovs, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: janis.ivanovs@silava.lv
  • Lazdiņš, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: andis.lazdins@silava.lv
  • Jansons, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: jurgis.jansons@silava.lv
  • Liepiņš, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: kaspars.liepins@silava.lv
article id 7749, category Research note
Eeva J. Vainio, Sannakajsa M. Velmala, Pertti Salo, Seppo Huhtinen, Michael M. Müller. (2017). Defoliation of Tilia cordata trees associated with Apiognomonia errabunda infection in Finland. Silva Fennica vol. 51 no. 4 article id 7749. https://doi.org/10.14214/sf.7749
Keywords: small-leaved lime; fungal disease; anthracnose; ITS rDNA; direct PCR; Tiliaceae
Highlights: Defoliation of Tilia cordata was investigated by fungal isolation from symptomatic leaf petioles and ITS sequence determination; The disease symptoms were associated with the presence of Apiognomonia errabunda; We report the first nucleotide sequences of A. errabunda from the Nordic countries.
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We investigated the causative agent of a disease outbreak affecting small-leaved limes (Tilia cordata Mill.) and resulting in darkening of the leaf petioles and excessive defoliation during summer 2016 in southern Finland. The fungal species composition of the symptomatic petioles was examined by culture isolation and molecular identification using ITS rDNA sequences, which revealed the most prevalent fungal species present in the petioles as Apiognomonia errabunda (Roberge) Höhn. Based on reviewing curated herbarium specimens deposited at the Universities of Helsinki and Turku, A. errabunda is native and widely distributed in small-leaved limes in Finland, and occasionally infects also other broadleaved trees, including Quercus robur L. and ornamental species of Tilia L. and Fagus L. The ITS sequence analysis conducted during this study revealed minor within-species polymorphisms similar to those observed earlier in the Central European and Russian populations of A. errabunda, and reports the first nucleotide sequences of this species from the Nordic countries.

  • Vainio, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 2, FI-00791 Helsinki, Finland ORCID http://orcid.org/0000-0002-6739-7968 E-mail: eeva.vainio@luke.fi (email)
  • Velmala, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: sannakajsa.velmala@luke.fi
  • Salo, Finnish Museum of Natural History, Botanical Museum, P.O. Box 7, FI-00014 University of Helsinki, Finland E-mail: pertti.salo@helsinki.fi
  • Huhtinen, University of Turku, Herbarium, Biodiversity Unit, FI-20014 Turku, Finland E-mail: sephuh@utu.fi
  • Müller, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 2, FI-00791 Helsinki, Finland E-mail: micms.muller@gmail.com
article id 5644, category Research note
Jiří Korecký, Jan Bílý, Petr Sedlák, Milan Lstibůrek. (2017). Innovative multiplex and its evaluation for effective genotyping of wild cherry. Silva Fennica vol. 51 no. 3 article id 5644. https://doi.org/10.14214/sf.5644
Keywords: genetic structure; microsatellites; Prunus avium; SSRs; pseudo 15-plex; one-run genotyping
Highlights: We present simple and cost-efficient method of SSR genotyping in cherry species; Seven of fifteen previously published primer sequences have been altered in order to produce complementary amplicons in pseudo 15-plex; The effectiveness and discriminatory power of established multiplex were verified by SSRs analysis of 48 wild cherry trees.
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Trees from the family Rosaceae play an important role in forest and agricultural ecosystems. Therefore, they are often an object of interest for both forest and horticultural tree breeders. Here, we present the utilization of an effective microsatellite (SSRs) genotyping method for wild cherry (Prunus avium L.) and verified the discriminatory power of the presented multiplex by genotyping 48 genetically distinctive individuals (plus-trees). Concerned loci were previously proven to be cross-compatible among various cultivars of cherry, hence, the method could have a broader utilization beyond to the field of forestry.
Our technique is based on post-PCR processing of 15 polymorphic SSRs loci amplified in three multiplex reactions with fluorescently labeled primers (6-FAM, VIC, PET and NED). All PCR products could be pooled and analyzed simultaneously (pseudo 15-plex). In order to make this approach feasible, we redefined sequences of several primers. Thus, utilizing modified primers provides non-overlapping amplicons of each fluorescent dye.

  • Korecký, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences, Kamýcká 1176, 165 21, Prague, Czech Republic ORCID http://orcid.org/0000-0001-7859-1750 E-mail: korecky@fld.czu.cz (email)
  • Bílý, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences, Kamýcká 1176, 165 21, Prague, Czech Republic ORCID http://orcid.org/0000-0001-5794-0907 E-mail: bily@fld.czu.cz
  • Sedlák, Faculty of Agrobiology, Food and Natural Resources, Czech University of Life Sciences, Kamýcká 129, 165 21, Prague, Czech Republic ORCID http://orcid.org/0000-0001-8016-8900 E-mail: sedlak@af.czu.cz
  • Lstibůrek, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences, Kamýcká 1176, 165 21, Prague, Czech Republic ORCID http://orcid.org/0000-0002-6304-6669 E-mail: lstiburek@fld.czu.cz
article id 1717, category Research note
Jussi Manner, Olle Gelin, Anders Mörk, Martin Englund. (2017). Forwarder crane’s boom tip control system and beginner-level operators. Silva Fennica vol. 51 no. 2 article id 1717. https://doi.org/10.14214/sf.1717
Keywords: productivity; automation; cut-to-length logging; haulage; crane work; extraction; learning curve
Highlights: Boom tip control (BTC) allows the operator to control boom tip movements directly, instead of controlling each movement separately to achieve the desired boom tip movement; BTC eased boom control, so beginner-level operators using BTC achieved higher productivity than beginner-level operators using a conventional (reference) system; There were no significant differences in the slopes of learning curves between the systems.
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The forwarder loads processed wood and transports it to a landing. Productivity of forwarding could be improved by increasing driving speed, but difficult forest terrain limits this. According to current literature, crane work is the most time-consuming work element of forwarding, so improving crane work productivity is essential for improving forwarding productivity. One way to do this is through automation of recurrent boom movement patterns, or alternatively automation can be used to ease crane work. When using conventional boom control (CBC), the operator manually controls each of the independent boom joint movements and combines them to achieve a desired boom tip movement, but boom tip control (BTC) allows the operator to control boom tip movements directly. The objective of the present study was to examine whether BTC facilitates crane work and affects the slopes of learning curves for beginner-level forwarder operators. The study was carried out using a standardised test routine to evaluate effects of two fixed factors, system (levels: CBC, BTC) and point of time (four levels), on five dependent variables. Four of the five dependent variables measured ease of boom control and the fifth measured crane work productivity. The results showed that there were no significant differences in the slopes of learning curves between the systems but the BTC did increase crane work productivity and made boom control easier.

  • Manner, The Forestry Research Institute of Sweden (Skogforsk), Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID http://orcid.org/0000-0002-4982-3855 E-mail: jussi.manner@skogforsk.se (email)
  • Gelin, The Forestry Research Institute of Sweden (Skogforsk), Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: olle.gelin@skogforsk.se
  • Mörk, The Forestry Research Institute of Sweden (Skogforsk), Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: anders.mork@skogforsk.se
  • Englund, The Forestry Research Institute of Sweden (Skogforsk), Uppsala Science Park, SE-751 83 Uppsala, Sweden E-mail: martin.englund@skogforsk.se
article id 1716, category Research note
Nelson Thiffault, Alain Paquette, Christian Messier. (2017). Early silvicultural guidelines for intensive management of hybrid larch plantations on fertile sub-boreal sites. Silva Fennica vol. 51 no. 2 article id 1716. https://doi.org/10.14214/sf.1716
Keywords: vegetation management; survival; growth; scarification; exotic tree species; intensive silviculture; forest functional zoning (TRIAD)
Highlights: Satisfactory growth can be obtained using a wide range of site preparation intensities; There is a net advantage of performing two motor-manual release treatments over a single release; A second release treatment cannot be replaced by more intensive site preparation; Planting depth had no influence on planted seedling growth after 6 years.
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Use of fast-growing tree plantations on dedicated areas is proposed as a means of reconciling fibre production with conservation objectives. Success of this approach however requires fine-tuning silvicultural scenarios so that survival and growth are optimized while management and environmental costs are minimized. This is particularly challenging for hybrid larch (Larix × marschlinsii Coaz), a shade-intolerant species planted on fertile sites in Quebec (Canada) where legislation prevents the use of chemical herbicides. In this context, multiple motor-manual release treatments are often required, with high impacts on costs and social issues related to the scarcity of a qualified workforce. We established a split-split-plot design on a recently harvested site to assess the main and interaction effects of mechanical site preparation (MSP) intensity (five modalities of trenching or mounding), motor-manual release scenario (one or two treatments) and planting depth (0–3 cm or 3–10 cm) on hybrid larch seedling growth and survival six years after planting. Mechanical site preparation intensity and planting depth did not influence seedling growth after 6 years. The lack of significant interaction between MSP and release scenarios indicates that these operations should be planned independently. A more intensive MSP treatment cannot replace a second motor-manual release on fertile sites, as proposed to reduce costs. Our results also show the significant advantage of performing two motor-manual release treatments two years apart (the first one early in the scenario), over performing a single treatment. Our study provides silvicultural guidelines for the establishment of high-yield exotic larch plantations.

  • Thiffault, Direction de la recherche forestière, Ministère des Forêts, de la Faune et des Parcs du Québec, 2700 rue Einstein, Québec, QC, Canada G1P 3W8; Centre d’étude de la forêt, CP 8888, Succursale Centre-ville, Montréal, QC, Canada H3C 3P8 ORCID http://orcid.org/0000-0003-2017-6890 E-mail: nelson.thiffault@mffp.gouv.qc.ca (email)
  • Paquette, Centre d’étude de la forêt, CP 8888, Succursale Centre-ville, Montréal, QC, Canada H3C 3P8; Université du Québec à Montréal, Département des sciences biologiques, CP 8888, Succursale Centre-ville, Montréal, QC, Canada H3C 3P8 ORCID http://orcid.org/0000-0003-1048-9674 E-mail: alain.paquette@gmail.com
  • Messier, Centre d’étude de la forêt, CP 8888, Succursale Centre-ville, Montréal, QC, Canada H3C 3P8; Université du Québec à Montréal, Département des sciences biologiques, CP 8888, Succursale Centre-ville, Montréal, QC, Canada H3C 3P8; Institut des Sciences de la Forêt tempérée (ISFORT), 58 rue Principale, Ripon, QC, Canada JOV 1V0 E-mail nelson.thiff E-mail: christian.messier@uqo.ca
article id 1674, category Research note
Nevenka Ćelepirović, Monika Karija Vlahović, Aikaterini Dounavi, Mladen Ivanković. (2016). Optimizations of high throughput multiplex polymerase chain reaction with simple sequence repeat markers for genotyping of common walnut populations (Juglans regia L.). Silva Fennica vol. 50 no. 5 article id 1674. https://doi.org/10.14214/sf.1674
Keywords: SSR markers; nuclear microsatellites; Juglans regia L.; multiplex PCR
Highlights: We combined eleven SSR markers in one multiplex PCR to make faster and cost effective amplification of the common walnut DNA from Croatia; Genetic variation of common walnut from Croatia was moderate at analyzed SSR loci; The resultant multiplex PCR could be used for genotyping of common walnut populations.
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Multiplex polymerase chain reaction (PCR) allows amplification of two or more pair of primers in parallel for amplification of multiple target sequences in a single reaction tube. In this study, we combined existing simple sequence repeat (SSR) markers (nuclear microsatellites) in the novel combination of multiplex PCR to study the population genetics of common walnut from Croatia. From twenty one tested SSR markers, eleven produced satisfactory results in one multiplex PCR. Population genetic results achieved from 15 samples of Croatian common walnut showed moderate genetic variability (average value: He 0.473; Ho 0.568). Our multiplex PCR allowed cost effective work concerning chemicals, plastic ware, device, and working time producing optimal results. The optimized multiplex PCR represented the best combination of eleven SSR primers for genotyping common walnut in a single PCR reaction.

  • Ćelepirović, Division of Genetics, Forest Tree Breeding and Seed Science, Croatian Forest Research Institute, Jastrebarsko, Croatia E-mail: celepirovic.nevenka@gmail.com (email)
  • Karija Vlahović, DNA Laboratory, Department of Forensic Medicine&Criminology, School of Medicine, University of Zagreb, Zagreb, Croatia E-mail: monika.karija.vlahovic@mef.hr
  • Dounavi, Department of Forest Protection, Forest Research Insitute of Baden-Württemberg, Wonnhaldesttr. 4, 79100 Freiburg, Germany E-mail: aikaterini.dounavi@forst.bwl.de
  • Ivanković, Division of Genetics, Forest Tree Breeding and Seed Science, Croatian Forest Research Institute, Jastrebarsko, Croatia E-mail: mladeni@sumins.hr
article id 1661, category Research note
Āris Jansons, Linda Robalte, Roberts Čakšs, Roberts Matisons. (2016). Long-term effect of whole tree biomass harvesting on ground cover vegetation in a dry Scots pine stand. Silva Fennica vol. 50 no. 5 article id 1661. https://doi.org/10.14214/sf.1661
Keywords: Pinus sylvestris; soil disturbance; stand replacement; Latvia; biomass removal; soil scarification
Highlights: After 47 years, whole tree harvesting (WTH) increased richness of ground cover species compared to conventionally managed stands; Higher occurrence of the oligotrophic species after WTH suggested reduction of soil nutrient content, hence formation of different plant community; WTH, apparently, facilitated recovery of species typical for later successional stages.
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Long-term (47 years) effect of experimental whole tree harvesting (WTH) with a heavy soil scarification on ground cover vegetation was assessed in a dry nutrient-poor Scots pine (Pinus sylvestris L.) stand in Latvia. Neighbouring conventionally managed young (10 years) and mature (119 years) stands of the same type were used for comparison. Higher species richness was observed in the WTH stand compared to conventionally managed young and mature stands (24, 18 and 16 species, respectively), likely due to the profound disturbance. The Shannon diversity index was higher in the young than in the WTH and mature stands (2.36, 1.77 and 1.63, respectively); still, the composition and structure of ground cover vegetation in WTH was more similar to the mature stand. Nevertheless, the occurrence of oligotrophic species in the WTH stand suggested decreased soil nutrient content and potential development of different plant community. Hence, such method might be considered for restoration of oligotrophic stands. Nevertheless, the period of 47 years appeared sufficient for the ground cover vegetation to recover after the WTH.

  • Jansons, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: aris.jansons@silava.lv
  • Robalte, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: robalte.l@gmail.com (email)
  • Čakšs, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: chakijs95@gmail.com
  • Matisons, Latvian State Forest Research Institute “Silava”, 111 Rigas Str., LV 2169, Salaspils, Latvia E-mail: roberts.matisons@silava.lv
article id 1656, category Research note
Līga Puriņa, Roberts Matisons, Āris Jansons, Silva Šēnhofa. (2016). Survival of European beech in the central part of Latvia 33 years since the plantation. Silva Fennica vol. 50 no. 4 article id 1656. https://doi.org/10.14214/sf.1656
Keywords: Fagus sylvatica; introduction experiment; sapling mortality; northeasternmost beech trial
Highlights: Beech saplings growing in the central part of Latvia had ca. 80% survival during the recent three decades; The dimensions of saplings varied greatly likely due to canopy conditions; Some beech self-regeneration was observed; Mainly saplings had narrow crowns; The distribution of sapling dimensions had the reverse-J shape, suggesting successful development of beech.
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The projections of vegetation zones suggest increasing growth potential of European beech (Fagus sylvatica L.) in Northern Europe. Such changes usually are most apparent in the marginal populations. In this study, survival of young beech growing in an experimental plantation under canopy of a mixed coniferous stand in the central part of Latvia was assessed after 33 years since the establishment. The planting material originated from an older experimental stand in the western part of Latvia. Although, at present, the studied plantation could be considered as the northeasternmost beech stand in Europe, a good survival was observed – ca. 80% of the seedlings have survived, despite several cold spells of ca. –30 °C that occurred during the recent three decades. Additionally, some self-regeneration i.e. branch sprouting was observed. The saplings were rather low, as their mean height was ca. 4 m. Still, some individuals, which were growing under canopy openings, reached considerable dimensions; their height and stem diameter exceeded 10 m and 9 cm, respectively. The distribution of sapling dimensions had the reverse-J shape that is typical for shade tolerant species, indicating normal development of the beech regrowth. The crowns of saplings were narrow and the stems were spindly, suggesting that trees with a good stem quality might be bred. Hence, our results suggest that environmental conditions in the central part of Latvia have been satisfactory for beech, thus encouraging establishment of more extensive trials within the region.

  • Puriņa, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: liga.purina@silava.lv
  • Matisons, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: robism@inbox.lv (email)
  • Jansons, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: aris.jansons@silava.lv
  • Šēnhofa, LSFRI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: silva.senhofa@silava.lv
article id 1573, category Research note
Marjut Turtiainen, Jari Miina, Kauko Salo, Juha-Pekka Hotanen. (2016). Modelling the coverage and annual variation in bilberry yield in Finland. Silva Fennica vol. 50 no. 4 article id 1573. https://doi.org/10.14214/sf.1573
Keywords: abundance; berry yield; generalised linear mixed model; Vaccinium myrtillus L.
Highlights: The highest bilberry coverage was found in mesic heath forests and fell forests; On peatlands the coverage was, on average, lower than on mineral soil sites; The approach introduced in this study to calculating annual berry yield indices is a promising way for estimating total annual bilberry yields over a given period of time.
Abstract | Full text in HTML | Full text in PDF | Author Info

The coverage of bilberry (Vaccinium myrtillus L.) was modelled as a function of site and stand characteristics using the permanent sample plots of the National Forest Inventory (NFI) (Model 1). The sample sites consisted of mineral soil forests as well as fells and peatland sites. Annual variation in the bilberry yield (Model 2) was analysed based on measurements over 2001–2014 in the permanent sample plots (so-called MASI plots) in various areas of Finland. We derived annual bilberry yield indices from the year effects of Model 2 and investigated whether these indices could be used to estimate annual variation in bilberry crops in Finland. The highest bilberry coverage was found in mesic heath forests and fell forests. On peatlands the coverage was, on average, lower than on mineral soil sites; the peatland sites with most bilberry coverage were meso-oligotrophic and oligotrophic spruce mires and oligotrophic pine mires. Our bilberry yield indices showed similar variation to those derived from the mean annual berry yields reported and calculated earlier using the MASI plots; the correlation between the indices was 0.795. This approach to calculating annual berry yield indices is a promising way for estimating total annual bilberry yields over a given period of time. Models 1 and 2 can be used in conjunction with the Miina et al.’s (2009) bilberry yield model when bilberry coverage, average annual yield and annual variation in the yield are to be predicted in forest planning.

  • Turtiainen, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: marjut.turtiainen@uef.fi (email)
  • Miina, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Box 68, FI-80101 Joensuu, Finland E-mail: jari.miina@luke.fi
  • Salo, Natural Resources Institute Finland (Luke), Bio-based Business and Industry, Box 68, FI-80101 Joensuu, Finland E-mail: kauko.salo@luke.fi
  • Hotanen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Box 68, FI-80101 Joensuu, Finland E-mail: juha-pekka.hotanen@luke.fi
article id 1549, category Research note
Francesco Chianucci, Luca Salvati, Tessa Giannini, Ugo Chiavetta, Piermaria Corona, Andrea Cutini. (2016). Long-term response to thinning in a beech (Fagus sylvatica L.) coppice stand under conversion to high forest in Central Italy. Silva Fennica vol. 50 no. 3 article id 1549. https://doi.org/10.14214/sf.1549
Keywords: continuous cover forestry; thinning; Leaf Area Index; coppice management; Plant Canopy Analyzer
Highlights: Canopy recovery after medium-heavy thinning reveals the prompt response of beech to intensive thinning cycles; Active management practices accelerate the transition from coppice to high forest.
Abstract | Full text in HTML | Full text in PDF | Author Info

European beech (Fagus sylvatica L.) forests have a long history of coppicing, but the majority of formerly managed coppices are currently under conversion to high forest. The long time required to achieve conversion requires a long-term perspective to fully understand the implication of the applied conversion practices. In this study, we showed results from a long-term (1992–2014) case-study comparing two management options (natural evolution and periodic thinning) in a beech coppice in conversion to high forest. Leaf area index, litter production, radiation transmittance and growth efficiency taken as relevant stand descriptors, were estimated using both direct and indirect optical methods. Overall, results indicated that beech coppice showed positive and prompt responses to active conversion practices based on periodic medium-heavy thinning. A growth efficiency index showed that tree growth increased as the cutting intensity increased. Results from the case study supported the effectiveness of active conversion management from an economic (timber harvesting) and ecological (higher growth efficiency) point of view.

  • Chianucci, Consiglio per la Ricerca in Agricoltura e l’Analisi dell’Economia Agraria – Forestry Research Centre, viale Santa Margherita 80, 52100 Arezzo, Italy ORCID http://orcid.org/0000-0002-5688-2060 E-mail: fchianucci@gmail.com (email)
  • Salvati, Consiglio per la Ricerca in Agricoltura e l’Analisi dell’Economia Agraria – Research Centre for the Soil-Plant System, via della Navicella 2–4, 00184 Roma, Italy E-mail: bayes00@yahoo.it
  • Giannini, Consiglio per la Ricerca in Agricoltura e l’Analisi dell’Economia Agraria – Forestry Research Centre, viale Santa Margherita 80, 52100 Arezzo, Italy E-mail: tessa.giannini@entecra.it
  • Chiavetta, Consiglio per la Ricerca in Agricoltura e l’Analisi dell’Economia Agraria – Forestry Research Centre, viale Santa Margherita 80, 52100 Arezzo, Italy E-mail: ugo.chiavetta@entecra.it
  • Corona, Consiglio per la Ricerca in Agricoltura e l’Analisi dell’Economia Agraria – Forestry Research Centre, viale Santa Margherita 80, 52100 Arezzo, Italy E-mail: piermaria.corona@unitus.it
  • Cutini, Consiglio per la Ricerca in Agricoltura e l’Analisi dell’Economia Agraria – Forestry Research Centre, viale Santa Margherita 80, 52100 Arezzo, Italy E-mail: andrea.cutini@entecra.it
article id 1544, category Research note
Osmo Mattila, Kaisa Hämäläinen, Liina Häyrinen, Sami Berghäll, Katja Lähtinen, Anne Toppinen. (2016). Strategic business networks in the Finnish wood products industry: a case of two small and medium-sized enterprises. Silva Fennica vol. 50 no. 3 article id 1544. https://doi.org/10.14214/sf.1544
Keywords: competitive advantage; wood products; network-based business models
Highlights: Network-based business models are not yet common in the wood products industry; Further research is suggested on network-based co-operation to vitalize industry.
Abstract | Full text in HTML | Full text in PDF | Author Info

The use of network-based business models has been brought up as a means of creating com-petitive edge in the tightening global competition. In practice, adopting network-based models has not yet become common in the wood products industry. The objective of this study is to gain better understanding of types of network-based business models using a case study of two small and medium-sized wood industry companies in Finland (for a sake of anonymity named as A and B). The network of company A is found to consist of mostly of established actors with a new-in-the-market value creation system, whereas network for company B is more stable and has an established value system aiming at growth and incremental innovations. Both networks had experienced difficulties in finding partners and lacked some strategic resources. Via this example we wish to stimulate further research interest on the sources of network-based competitive advantage in the traditional wood product industry in a need of renewal of business models.

  • Mattila, University of Helsinki, Department of Forest Sciences, FI-00014 Helsingin yliopisto, Finland E-mail: osmo.mattila@helsinki.fi
  • Hämäläinen, Siparila Oy, Varaslahdentie 1, FI-40800 Vaajakoski, Finland E-mail: kaisa.hamalainen@siparila.fi
  • Häyrinen, University of Helsinki, Department of Forest Sciences, FI-00014 Helsingin yliopisto, Finland E-mail: liina.hayrinen@helsinki.fi
  • Berghäll, University of Helsinki, Department of Forest Sciences, FI-00014 Helsingin yliopisto, Finland E-mail: sami.berghall@helsinki.fi
  • Lähtinen, University of Helsinki, Department of Forest Sciences, FI-00014 Helsingin yliopisto, Finland E-mail: katja.lahtinen@helsinki.fi
  • Toppinen, University of Helsinki, Department of Forest Sciences, FI-00014 Helsingin yliopisto, Finland E-mail: anne.toppinen@helsinki.fi (email)
article id 1518, category Research note
Francesco Chianucci. (2015). A note on estimating canopy cover from digital cover and hemispherical photography. Silva Fennica vol. 50 no. 1 article id 1518. https://doi.org/10.14214/sf.1518
Keywords: forest canopy; foliage cover; crown cover; fisheye photography; cover photography
Highlights: Comparison of fisheye (DHP) and cover (DCP) photography for estimating canopy cover; Digital photographic estimates validated against artificial images with known cover; Accuracy of cover estimates from DHP is influenced by mean gap size and actual cover; Accuracy of cover estimates from DCP is not influenced by mean gap size and actual cover.
Abstract | Full text in HTML | Full text in PDF | Author Info

Fast and accurate estimates of canopy cover are central for a wide range of forestry studies. As direct measurements are impractical, indirect optical methods have often been used in forestry to estimate canopy cover. In this paper the accuracy of canopy cover estimates from two widely used canopy photographic methods, hemispherical photography (DHP) and cover photography (DCP) was evaluated. Canopy cover was approximated in DHP as the complement of gap fraction data at narrow viewing zenith angle range (0°–15°), which was comparable with that of DCP. The methodology was tested using artificial images with known canopy cover; this allowed exploring the influence of actual canopy cover and mean gap size on canopy cover estimation from photography. DCP provided robust estimates of canopy cover, whose accuracy was not influenced by variation in actual canopy cover and mean gap size, based on comparison with artificial images; by contrast, the accuracy of cover estimates from DHP was influenced by both actual canopy cover and mean gap size, because of the lower ability of DHP to detect small gaps within crown. The results were replicated in both DHP and DCP images collected in real forest canopies. Finally, the influence of canopy cover on foliage clumping index and leaf area index was evaluated using a theoretical gap fraction model. The main findings indicate that DCP can overcome the limits of indirect techniques for obtaining unbiased and precise estimates of canopy cover, which are comparable to those obtainable from direct, more labour-intensive techniques, being therefore highly suitable for routine monitoring and inventory purposes.

  • Chianucci, Consiglio per la Ricerca in Agricoltura e l’Analisi dell’Economia Agraria – Forestry Research Centre, viale Santa Margherita 80, 52100 Arezzo, Italy ORCID http://orcid.org/0000-0002-5688-2060 E-mail: fchianucci@gmail.com (email)
article id 1496, category Research note
Juha Lappi, Jaana Luoranen. (2016). Using a bivariate generalized linear mixed model to analyze the effect of feeding pressure on pine weevil damage. Silva Fennica vol. 50 no. 1 article id 1496. https://doi.org/10.14214/sf.1496
Keywords: measurement error; best linear predictor; correlated random effects; log-log link
Highlights: Probability of damage of treated seedlings can be predicted from the probability of damage of control seedlings (feeding pressure).
Abstract | Full text in HTML | Full text in PDF | Author Info

The objective of the study is to derive a method by which one can analyze how the probability of damage made by pine weevils on seedlings treated with insecticides depends on the probability of damage on untreated control seedlings, called feeding pressure. Because the probabilities vary from stand to stand and from block to block, the analysis is done using a generalized linear mixed model. The dependency of probability of damage on the feeding pressure cannot be properly analyzed using observed relative frequency of damage of control seedlings as a covariate, but it can be analyzed using a bivariate model. One equation describes damage of control seedlings and another equation damage of treated seedlings. The random stand and block effects of different equations are correlated. For a given probability of stand level control seedling damage, the random stand effect for control seedlings can be computed using a link function, then random stand effects for treated seedlings can be predicted using the best linear predictor from the random effect for control seedlings. Using an inverse link the prediction can again be presented in the probability scale which is of interest to the user. Using these three steps the probability of damage of treated seedlings can be predicted from the control damage probability. The probability of damage of treated seedlings can also be predicted from the observed relative frequency of damaged control seedlings using simulation. The complementary log-log link was used for control seedlings and the log-log link for treated seedlings.

  • Lappi, Natural Resources Institute Finland (Luke), Economics and society, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: juha.lappi@luke.fi (email)
  • Luoranen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: jaana.luoranen@luke.fi
article id 1443, category Research note
Jouni Partanen, Risto Häkkinen, Heikki Hänninen. (2015). Significance of the root connection on the dormancy release and vegetative bud burst of Norway spruce (Picea abies) seedlings in relation to accumulated chilling. Silva Fennica vol. 50 no. 1 article id 1443. https://doi.org/10.14214/sf.1443
Keywords: phenology; bud burst percentage; days to bud burst; dormancy release; forcing
Highlights: Cutting the root connection slightly increased the number of days to bud burst of Norway spruce seedlings under warm conditions but it had no consistent effect on bud burst percentage; Our results obtained with seedlings suggest that using detached tree material in dormancy release experiments may slightly affect the results but it will evidently not lead to drastically erroneous conclusions.
Abstract | Full text in HTML | Full text in PDF | Author Info

The effect of cutting the root connection by detaching the shoot from the root system on dormancy release and vegetative bud burst was examined in 2-year-old seedlings of Norway spruce (Picea abies [L.] Karst.). Seedlings were transferred at 1–4 week intervals between October and January from outdoor conditions to experimental forcing in a heated greenhouse. Before forcing, half of the seedlings were cut above ground line, and the detached shoots were forced with their cut ends placed in water. The intact seedlings were forced with their root system remaining intact in the pots. Vegetative bud burst was observed visually. Cutting the root connection slightly increased days to bud burst in the forcing conditions, however, no consistent effect on bud burst percentage was found. Our preliminary seedling data suggest that using detached tree material in dormancy release experiments may have a small effect on bud burst date but it will evidently not lead to drastically erroneous conclusions. Further studies, using different seed lots, are needed to assess the effect of detaching on the dormancy release and bud burst, especially in adult trees.

  • Partanen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: jouni.partanen@luke.fi (email)
  • Häkkinen, Natural Resources Institute Finland (Luke), Management and Production of Renewable Resources, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: risto.hakkinen@luke.fi
  • Hänninen, University of Helsinki, Department of Biosciences, Viikki Plant Science Centre (ViPS), P.O. Box 65, FI-00014 University of Helsinki, Finland E-mail: heikki.hanninen@helsinki.fi
article id 1346, category Research note
Āris Jansons, Roberts Matisons, Oskars Krišāns, Baiba Džeriņa, Mārtiņš Zeps. (2015). Effect of initial fertilization on 34-year increment and wood properties of Norway spruce in Latvia. Silva Fennica vol. 50 no. 1 article id 1346. https://doi.org/10.14214/sf.1346
Keywords: Picea abies; wood density; proportion of latewood; tree-rings; high-frequency densitometry; legacy effect
Highlights: The initial fertilization increased the productivity of Picea abies increasing stem volume by 17% at the age of 34 years; The tree-ring width was affected for up to15 years; The fertilization did not affect mean tree-ring density, although the latewood density was increased.
Abstract | Full text in HTML | Full text in PDF | Author Info

Initial fertilisation, when the fertilizer is supplied during the plantation, is applied to improve the competitive ability of the seedlings and hence to increase their growth and productivity; however, fertilization could also alter wood properties and timber quality. In this study, the dimensions and tree-ring parameters – width, proportion of latewood, maximum and mean density, mean earlywood and latewood density – of initially fertilized (by 14, 6 and 11 g of N, P and K per seedling, respectively) Norway spruce (Picea abies Karst.) growing in an experimental plantation in Kalsnava, Latvia (temperate climate region) were assessed. The fertilization significantly increased the dimensions of trees in long-term (ca. 17% increase of stemwood volume). The analysis of tree-ring width suggested that the duration of the effect was ca. 15 years. The maximum and latewood density were higher for the fertilized trees only in a few years. The mean and earlywood density of tree-rings were mainly similar for both treatments. Altogether, considering the one-time application of a limited amount of fertilizer, such treatment had notable and lasting effect on Norway spruce.

  • Jansons, LSRFI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: aris.jansons@silava.lv
  • Matisons, LSRFI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: robism@inbox.lv (email)
  • Krišāns, LSRFI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: oskars.krisans@silava.lv
  • Džeriņa, LSRFI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: baiba.dzerina@silava.lv
  • Zeps, LSRFI “Silava”, Rigas str. 111, Salaspils, Latvia, LV2169 E-mail: martins.zeps@silava.lv
article id 1415, category Research note
Zorica Šarac, Tanja Dodoš, Nemanja Rajčević, Srdjan Bojović, Petar Marin, Jelena Aleksić. (2015). Genetic patterns in Pinus nigra from the central Balkans inferred from plastid and mitochondrial data. Silva Fennica vol. 49 no. 5 article id 1415. https://doi.org/10.14214/sf.1415
Keywords: genetic differentiation; genetic diversity; the Balkans; European black pine; chloroplast microsatellites; mitochondrial nad7 intron 1; haplotype network
Highlights: Seven populations of Pinus nigra from the central Balkans, representing four infraspecific taxa, were analyzed with chloroplast microsatellites and a mitochondrial locus; Molecular data failed to support infraspecific circumscriptions; Levels of genetic diversities/differentiation at both genomes were in the range of those reported in western Mediterranean populations of P. nigra; Iberian/African and Balkans’ populations share one mtDNA polymorphism and differ in three mutations.
Abstract | Full text in HTML | Full text in PDF | Author Info

Pinus nigra J.F. Arnold, European black pine, is a typical component of Mediterranean and sub-Mediterranean coniferous forests with highly fragmentary distribution. Western Mediterranean populations of this species have been studied genetically to date, while eastern populations from the central Balkans, which are larger and more abundant, are still genetically understudied. We analyzed seven populations of P. nigra representing all infraspecific taxa recognized within the central Balkans (subspecies nigra with varieties nigra and gocensis Đorđević; and subspecies pallasiana (Lamb.) Holmboe with varieties pallasiana and banatica (Endl.) Georgescu et Ionescu), with three chloroplast microsatellites (cpDNA SSRs) and one mitochondrial (mtDNA) locus. Although our molecular data failed to support circumscription of studied infraspecific taxa, we found that genetic patterns at both genomes are in accordance with those found previously in westward populations of this species, that is – exceptionally high levels of genetic diversity (HT = 0.949) and low genetic differentiation (GST = 0.024) at the cpDNA level, and moderate levels of genetic diversity (HT = 0.357) and genetic differentiation (GST = 0.358) at the mtDNA level. Based on genealogical relations of mtDNA types currently present in Balkans’ and Iberian/African populations, we inferred that the ancestral gene pool of P. nigra already harbored polymorphism at position 328 prior to the divergence to two lineages currently present in westward and eastward parts of the species range distribution. Subsequent occurrence of three mutations, which distinguish these two lineages, suggests their long-term isolation.

  • Šarac, University of Niš, Faculty of Sciences and Mathematics, Department of Biology and Ecology, Višegradska 33, 18000 Niš, Serbia E-mail: saraczorica@gmail.com (email)
  • Dodoš, University of Belgrade, Faculty of Biology, Institute of Botany and Botanical Garden “Jevremovac”, Studentski trg 16, 11000 Belgrade, Serbia E-mail: tanjadodos@bio.bg.ac.rs
  • Rajčević, University of Belgrade, Faculty of Biology, Institute of Botany and Botanical Garden “Jevremovac”, Studentski trg 16, 11000 Belgrade, Serbia E-mail: nemanja@bio.bg.ac.rs
  • Bojović, University of Belgrade, Institute for Biological Research “Siniša Stanković”, Boulevard Despota Stefana 142, 11060 Belgrade, Serbia E-mail: bojovic@ibiss.bg.ac.rs
  • Marin, University of Belgrade, Faculty of Biology, Institute of Botany and Botanical Garden “Jevremovac”, Studentski trg 16, 11000 Belgrade, Serbia E-mail: pdmarin@bio.bg.ac.rs
  • Aleksić, University of Belgrade, Institute of Molecular Genetics and Genetic Engineering, Vojvode Stepe 444a, P.O. Box 23, 11000 Belgrade, Serbia E-mail: aleksic_jelena@yahoo.com.au
article id 1330, category Research note
Hanne K. Sjølie, Hans Asbjørn Kårstad Sørlie, Bjørn Tveite, Birger Solberg. (2015). The performance of two Swedish N fertilization functions evaluated on data from Norwegian fertilization experiments. Silva Fennica vol. 49 no. 4 article id 1330. https://doi.org/10.14214/sf.1330
Keywords: Pinus sylvestris; Picea abies; forest production; forest fertilization; predictive functions
Highlights: The performance of two predictive Swedish fertilization growth response functions was assessed on data from Norwegian fertilization experiments; One function performed well on the full dataset, but overpredicted the growth response in spruce plots and underpredicted in pine plots; The second function performed well in pine stands, but overestimated the growth response in spruce and in total.
Abstract | Full text in HTML | Full text in PDF | Author Info

This study compares the responses of two Swedish 5-year predictive stand-level functions with the observed responses in 721 fertilization experiment plots in Norway fertilized with nitrogen (N). All plots are single-species consisting of Norway spruce (Picea abies (L.) H. Karst.) or Scots pine (Pinus sylvestris L.) fertilized with ammonium nitrate (AN) or urea. The correlations between the observed and the two predicted responses were 0.34–0.40 for all plots taken together. One response function performed well on average, but underestimated the response in pine plots and overestimated the response in spruce plots. The second function overpredicted the response on the full dataset, in spruce plots and old forest, but performed well in pine plots. Both functions overestimated the growth response in high-productive plots. Higher N deposition in Norway than in Sweden may count for parts of the deviations. Testing of fertilization functions on new datasets is rare, but important part of the evaluation of functions. As the functions are not well fit for predicting the growth response in spruce and high-productive plots in our sample, new functions that include N deposition are welcome.

  • Sjølie, Department of Ecology and Natural Resource Management, Norwegian University of Life Sciences, NO-1432 Ås, Norway E-mail: hanne.sjolie@nmbu.no (email)
  • Sørlie, Department of Ecology and Natural Resource Management, Norwegian University of Life Sciences, NO-1432 Ås, Norway E-mail: hans.asbjorn.sorlie@slf.dep.no
  • Tveite, Norwegian Forest and Landscape Institute, P.O. Box 115, NO-1431 Ås, Norway E-mail: bjorn.tveite@skogoglandskap.no
  • Solberg, Department of Ecology and Natural Resource Management, Norwegian University of Life Sciences, NO-1432 Ås, Norway E-mail: birger.solberg@nmbu.no
article id 1320, category Research note
Abbot Okotie Oghenekaro, Geoffrey Daniel, Fred O Asiegbu. (2015). The saprotrophic wood-degrading abilities of Rigidoporus microporus. Silva Fennica vol. 49 no. 4 article id 1320. https://doi.org/10.14214/sf.1320
Keywords: simultaneous decay; wood degradation; white rot; delignification
Highlights: Rigidoporus microporus isolates displayed varying saprotrophic capabilities on wood blocks of Rubber tree (Hevea brasiliensis); Percentage mass loss of (Hevea brasiliensis) wood blocks caused by the pathogenic Rigidoporus microporus was significantly higher than that observed with the endophytic isolate; The endophytic isolate has very poor saprotrophic ability on Hevea brasiliensis wood blocks.
Abstract | Full text in HTML | Full text in PDF | Author Info

Saprotrophic wood-decaying abilities of Rigidoporus microporus (Polyporales, Basidiomycota) syn. Rigidoporus lignosus and the structural alterations induced in wood blocks of Hevea brasiliensis Muell. Arg were studied. Mass loss of wood blocks was analyzed after 3 and 6 months respectively and the patterns of decay by pathogenic and endophytic isolates of this fungus were investigated using light microscopy. Effects of temperature on growth of the isolates on malt extract agar were also investigated. The R. microporus isolated from a non-H. brasiliensis host caused the highest percentage mass loss (27.2% after 6 months), followed by isolates ED310 (21.1%) and M13 (15.7%), both collected from diseased H. brasiliensis plantations. The isolate initially identified as an endophyte showed very low saprotrophic wood decay capability (4.3% after 6 months). The optimal temperature for growth of the isolates was 30 °C; except for the endophytic isolate which showed highest growth at 25 °C. Wood samples degraded by the R. microporus isolates showed simultaneous attack of wood cell walls, typical of white rot fungi. Results of the study indicate variability in the wood degrading abilities of the isolates and the potential differences in their physiology are discussed. Our findings further support the need for a taxonomical revision of the Rigidoporus genus.

  • Oghenekaro, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: abbot.oghenekaro@helsinki.fi (email)
  • Daniel, Department of Forest Products/Wood Science, Swedish University of Agricultural Sciences, P.O. Box 7008, SE-75007 Uppsala, Sweden E-mail: geoffrey.daniel@slu.se
  • Asiegbu, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: fred.asiegbu@helsinki.fi
article id 1275, category Research note
Arshad Ali, Ming-Shan Xu, Yan-Tao Zhao, Qing-Qing Zhang, Liu-Li Zhou, Xiao-Dong Yang, En-Rong Yan. (2015). Allometric biomass equations for shrub and small tree species in subtropical China. Silva Fennica vol. 49 no. 4 article id 1275. https://doi.org/10.14214/sf.1275
Keywords: aboveground biomass; allometric equations; carbon storage; biometric variables; shrubs; subtropical forests
Highlights: Diameter (D) and height (H) are strong predictors in species-specific and multispecies models for the aboveground biomass of subtropical shrubs and small trees; Although wet basic density and crown shape may improve the predictive power of aboveground biomass slightly, the labor intensive measurements for wet basic density and crown shape may be disregarded when a large number of individuals are to be surveyed; Our results extend the generality of D-H models for aboveground biomass for large trees to subtropical shrubs and small trees.
Abstract | Full text in HTML | Full text in PDF | Author Info

Species-specific allometric equations for shrubs and small trees are relatively scarce, thus limiting the precise quantification of aboveground biomass (AGB) in both shrubby vegetation and forests. Fourteen shrub and small tree species in Eastern China were selected to develop species-specific and multispecies allometric biomass equations. Biometric variables, including the diameter of the longest stem (D), height (H), wet basic density (BD), and crown area and shape were measured for each individual plant. We measured the AGB through a non-destructive method, and validated these measurements using the dry mass of the sampled plant components. The AGB was related to biometric variables using regression analysis. The species-specific allometric models, with D and H as predictors (D-H models) accounted for 70% to 99% of the variation in the AGB of shrubs and small trees. A multispecies allometric D-H model accounted for 71% of the variation in the AGB. Although BD, as an additional predictor, improved the fit of most models, the D-H models were adequate for predicting the AGB for shrubs and small trees in subtropical China without BD data.

  • Ali, School of Ecological and Environmental Sciences, East China Normal University, Shanghai 200241, China; Tiantong National Forest Ecosystem Observations and Research Station, Ningbo 315114, Zhejiang, China; Department of Environmental Sciences, Abdul Wali Khan University Mardan, 23200, KPK, Pakistan E-mail: arshadforester@gmail.com
  • Xu, School of Ecological and Environmental Sciences, East China Normal University, Shanghai 200241, China; Tiantong National Forest Ecosystem Observations and Research Station, Ningbo 315114, Zhejiang, China E-mail: yumsh09@lzu.edu.cn
  • Zhao, School of Ecological and Environmental Sciences, East China Normal University, Shanghai 200241, China; Tiantong National Forest Ecosystem Observations and Research Station, Ningbo 315114, Zhejiang, China E-mail: zhaoyantao1991@163.com
  • Zhang, School of Ecological and Environmental Sciences, East China Normal University, Shanghai 200241, China; Tiantong National Forest Ecosystem Observations and Research Station, Ningbo 315114, Zhejiang, China E-mail: qingzq@yeah.net
  • Zhou, School of Ecological and Environmental Sciences, East China Normal University, Shanghai 200241, China; Tiantong National Forest Ecosystem Observations and Research Station, Ningbo 315114, Zhejiang, China E-mail: 792920738@qq.com
  • Yang, School of Ecological and Environmental Sciences, East China Normal University, Shanghai 200241, China; Tiantong National Forest Ecosystem Observations and Research Station, Ningbo 315114, Zhejiang, China E-mail: xjyangxd@sina.com
  • Yan, School of Ecological and Environmental Sciences, East China Normal University, Shanghai 200241, China; Tiantong National Forest Ecosystem Observations and Research Station, Ningbo 315114, Zhejiang, China E-mail: eryan@des.ecnu.edu.cn (email)
article id 1305, category Research note
Israel Sánchez-Osorio, Luis Domínguez, Gloria López-Pantoja, Raúl Tapias. (2015). Antennal response of Prinobius myardi to synthetic tree volatiles. Silva Fennica vol. 49 no. 3 article id 1305. https://doi.org/10.14214/sf.1305
Keywords: Quercus; EAG; electroantennography; wood borers; Cerambycidae; plant volatiles; β-pinene
Highlights: Prinobius myardi is a wood borer considered a major threat for Mediterranean oaks, especially Quercus suber and Q. ilex; We performed electroantennographic bioassays to assess olfactory sensitivity of P. myardi to synthetic plant volatiles; P. myardi exhibits a broad sensitivity to common tree volatiles, including those emitted by oaks (α-pinene and β-pinene) or non-host volatiles (1,8-cineole).
Abstract | Full text in HTML | Full text in PDF | Author Info
Prinobius myardi Mulsant is a wood borer implicated in the decline of Mediterranean oaks, especially Quercus suber L. and Quercus ilex L. Plant volatiles play an important role in plant-insect interactions, and electroantennography (EAG) is an effective tool for exploring the electrophysiological activity of host plant volatiles on insects. To improve our understanding of the olfactory sensitivity of P. myardi, we recorded EAG responses to 20 tree volatiles, and analyzed the dose-dependent response to five doses (10–4:1 to 1:1 v/v) of the three most EAG-active compounds. Antennae of P. myardi responded to 13 chemicals, mainly monoterpenes and green leaf volatiles, with the strongest EAG responses being observed with β-pinene, (+)-α-pinene and 1,8-cineole. Dose–response profiles showed positive dose-dependent responses for all three compounds. Our results suggest a broad sensitivity of P. myardi to common tree volatiles, particularly some host-related compounds and volatiles associated with wounded trees; the olfactory recognition of ratios of these compounds could play a role in host selection by P. myardi.
  • Sánchez-Osorio, Departamento de Ciencias Agroforestales, ETSI La Rábida, University of Huelva, 21819 Palos de la Frontera (Huelva), Spain ORCID http://orcid.org/0000-0002-6852-7699 E-mail: isanchez@uhu.es (email)
  • Domínguez, Departamento de Ciencias Agroforestales, ETSI La Rábida, University of Huelva, 21819 Palos de la Frontera (Huelva), Spain ORCID http://orcid.org/0000-0002-0131-0057 E-mail: luis.dominguez@dcaf.uhu.es
  • López-Pantoja, Departamento de Ciencias Agroforestales, ETSI La Rábida, University of Huelva, 21819 Palos de la Frontera (Huelva), Spain ORCID http://orcid.org/0000-0002-2659-6127 E-mail: pantoja@uhu.es
  • Tapias, Departamento de Ciencias Agroforestales, ETSI La Rábida, University of Huelva, 21819 Palos de la Frontera (Huelva), Spain E-mail: rtapias@uhu.es
article id 1321, category Research note
Sofia Bäcklund, Mari T. Jönsson, Joachim Strengbom, Göran Thor. (2015). Composition of functional groups of ground vegetation differ between planted stands of non-native Pinus contorta and native Pinus sylvestris and Picea abies in northern Sweden. Silva Fennica vol. 49 no. 2 article id 1321. https://doi.org/10.14214/sf.1321
Keywords: boreal forests; bryophytes; managed forests; introduced species; exotics; lichens; vascular plants
Highlights: Differences in ground vegetation patterns can be linked to tree species, forest stand age and differences in canopy cover; Vascular plant cover was higher in stands of P. contorta than in stands of both native tree species; The overall differences and similarities between P. contorta and the two native conifers were not consistent over the different age classes.
Abstract | Full text in HTML | Full text in PDF | Author Info
Intensified forestry increases the interest in replacing native tree species with fast growing non-native species. However, consequences for native biodiversity and ecosystem functioning are poorly understood. We compared cover and composition of major functional groups of ground vegetation between planted stands of non-native Pinus contorta Dougl. var. latifolia Engelm. and native conifers Pinus sylvestris L. and Picea abies (L.) H. Karst. in northern boreal Sweden. We quantified the ground cover of lichens, bryophytes, vascular plants and ground without vegetation (bare ground) in 96 stands covering three different age classes (15, 30 and 85 years old). Our study revealed differences in ground vegetation patterns between non-native and native managed forests, and that these differences are linked to stand age and differences in canopy cover. Total vascular plant cover increased with increasing stand age for all tree species, with P. contorta stands having higher cover than both native conifers. The ground cover of lichens was, although generally low, highest in stands of Pinus sylvestris. P. abies stands had a lower cover of vascular plants, but bare ground was more common compared with P. contorta. Our results suggest that the use of P. contorta as an alternative tree species in Fennoscandian forestry will influence native ground vegetation patterns. This influence is likely to change with time and future research should consider both temporal and landscape-scale effects from shifting tree-species dominance to Pinus contorta and other non-native tree species.
  • Bäcklund, Department of Ecology, Swedish University of Agricultural Sciences, P.O. Box 7044, SE-750 07 Uppsala, Sweden E-mail: sofia.backlund@slu.se (email)
  • Jönsson,  The Swedish Species Information Centre, P.O. Box 7007, SE-750 07 Uppsala, Sweden E-mail: mari.jonsson@slu.se
  • Strengbom, Department of Ecology, Swedish University of Agricultural Sciences, P.O. Box 7044, SE-750 07 Uppsala, Sweden E-mail: joachim.strengbom@slu.se
  • Thor, Department of Ecology, Swedish University of Agricultural Sciences, P.O. Box 7044, SE-750 07 Uppsala, Sweden E-mail: goran.thor@slu.se
article id 1255, category Research note
Āris Jansons, Roberts Matisons, Līga Puriņa, Una Neimane, Jānis Jansons. (2015). Relationships between climatic variables and tree-ring width of European beech and European larch growing outside of their natural distribution area. Silva Fennica vol. 49 no. 1 article id 1255. https://doi.org/10.14214/sf.1255
Keywords: Fagus sylvatica; introduced species; wood increment; climatic limitation; Latvia; Larix decidua
Highlights: In western Latvia, variation of tree-ring width of European beech and European larch within stands was similar; Dry summers and cold winters caused common event years in tree-rings; Moisture availability at the end of summer was apparently the main limiting factor for tree-ring width; Winter and spring temperature did not have significant and lasting effect on variation of tree-ring width.
Abstract | Full text in HTML | Full text in PDF | Author Info
Relationships between climatic variables and tree-ring width (TRW) of dominant European larch (Larix decidua Mill.) and European beech (Fagus sylvatica L.) trees growing outside of their natural distribution area in western Latvia were studied. Chronologies of TRW, which covered the periods 1949–2012 and 1911–2012, were produced for beech and larch, respectively. Common signatures in TRW between both species were observed, but their amplitude differed. Correlation analysis showed that variation of TRW of both species was affected by drought related climatic variables. Tree-ring width of beech was affected by temperature in the previous July and August and the effect of spring and autumn temperature was observed. Since the 1980s, the effect of July precipitation has become significant. Summer precipitation was significant for larch in the mid-part of the previous century; however, temperature in the previous September has become a limiting factor since 1970s. The limiting effect of winter and spring temperature apparently lost its significance around the 1950s.
  • Jansons, LSFRI „SILAVA”, Rigas Str. 111, Salaspils, Latvia, LV2169 E-mail: aris.jansons@silava.lv
  • Matisons, LSFRI „SILAVA”, Rigas Str. 111, Salaspils, Latvia, LV2169 E-mail: robism@inbox.lv (email)
  • Puriņa, LSFRI „SILAVA”, Rigas Str. 111, Salaspils, Latvia, LV2169 E-mail: liga.purina@silava.lv
  • Neimane, LSFRI „SILAVA”, Rigas Str. 111, Salaspils, Latvia, LV2169 E-mail: una.neimane@silava.lv
  • Jansons, Forest Competence Centre, Dzerbenes str. 27, Riga, Latvia, LV1006 E-mail: janis.jansons@mnkc.lv
article id 1114, category Research note
Antonio Villasante, Cristina Fernandez. (2014). Measurement errors in the use of smartphones as low-cost forestry hypsometers. Silva Fennica vol. 48 no. 5 article id 1114. https://doi.org/10.14214/sf.1114
Keywords: forest inventory; calibration; accelerometers; forestry tools; relative error
Highlights: We analysed two smartphones (HTC Desire and Samsung Galaxy Note) to determine the errors in the height measurements; The calibration included with the Android applications is insufficient; After appropriate calibration, the smartphone errors are similar to other forest hypsometers (Blume Leiss and Vertex).
Abstract | Full text in HTML | Full text in PDF | Author Info
Various applications currently available for Android allow the estimation of tree heights by using the 3D accelerometer on smartphones. Some make the estimation using the image on the screen, while in others, by pointing with the edges of the terminal. The present study establishes the measurement errors obtained with HTC Desire and Samsung Galaxy Note compared to those from Blume Leiss and Vertex IV. Six series of 12 measurements each were made with each hypsometer (for heights of 6 m, 8 m, 10 m and 12 m). A Kruskall Wallis test is applied to the relative errors to determine whether there are significant differences between the devices. The results indicate that the errors of the uncalibrated smartphones significantly exceed those of traditional forestry apparatus. However, calibration is a very easy procedure that can be done by means of a linear regression line between real angles (obtained with a Digital Angle Finder or with a series of measurements taken independently of the experiment), and the angles of the accelerometer. With this adjustment, the smartphones achieve adequate quality levels although the bias was not totally eliminated. The relative errors when pointing with the edges of the terminal show no significant differences compared to Blume Leiss. Applications that use the screen image give better results (no significant differences were detected with Vertex). There is currently no application that offers calibration of the linear regression slope, which is an essential requirement for ensuring the accuracy of height measurements obtained with smartphones.
  • Villasante, Agroforestry Departament, Universitat de Lleida, Av. Rovira Roure 177, 25198 Lleida, Spain ORCID http://orcid.org/0000-0002-7549-7424 E-mail: avillasante@eagrof.udl.cat (email)
  • Fernandez, Agroforestry Departament, Universitat de Lleida, Av. Rovira Roure 177, 25198 Lleida, Spain E-mail: cfernandez@eagrof.udl.cat
article id 1145, category Research note
Juha Siitonen. (2014). Ips acuminatus kills pines in southern Finland. Silva Fennica vol. 48 no. 4 article id 1145. https://doi.org/10.14214/sf.1145
Keywords: tree mortality; Scots pine; Ips acuminatus; Phaenops cyanea; drought
Highlights: Recently dead pines colonized by Ips acuminatus were frequently found in southern Finland, in a region where the species was thought to be absent; Colonized trees were typically large (average DBH 30 cm), located at open spots in pine-dominated stands, often forming groups of several trees; The damages may be a consequence of dry and hot summers during the 2000s.
Abstract | Full text in HTML | Full text in PDF | Author Info
Recently dead Scots pines (Pinus sylvestris L.) apparently killed by Ips acuminatus (Gyllenhal) were observed in Sipoo, southern Finland, in summer 2013. This record was unexpected and in contradiction with what is currently known about the distribution and aggressiveness of the species in Finland. The aim of this study was to survey a larger area in Uusimaa region, to find out whether I. acuminatus occurs frequently in recently dead pines, and whether inhabited trees share some common tree- or site-level characteristics. Galleries of I. acuminatus were found in most of the studied trees. A total of 96 inhabited trees were found in 21 separate sites. Colonized pines were typically large (average DBH 30 ± 9 cm) trees located in relatively open pine-dominated heathland stands at half-open, sun-exposed spots. The whole upper part of the trunk with thin bark was usually occupied. Galleries of Tomicus piniperda L. or T. minor Hartig occurred only in few cases in the same trees, indicating that the trees had died later in the summer. Galleries of the jewel beetle Phaenops cyanea F. were found in 13 trees. Trees colonized by I. acuminatus often occurred as small groups, with generally 1­–12 trees (average 3 trees), but in one exceptional group there were no less than 35 trees. It is possible that the hot and dry summers during the 2000s have increased the susceptibility of pines to insect damage, and have contributed to a population growth of I. acuminatus.
  • Siitonen, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: juha.siitonen@metla.fi (email)
article id 1136, category Research note
Håkan Lideskog, Back Tomas Ersson, Urban Bergsten, Magnus Karlberg. (2014). Determining boreal clearcut object properties and characteristics for identification purposes. Silva Fennica vol. 48 no. 3 article id 1136. https://doi.org/10.14214/sf.1136
Keywords: forestry; forest regeneration; clearcut obstacles; terrain description; terrain conditions; site properties
Highlights: We define the quantitative properties and qualitative characteristics of stumps, stones, slash, and roots, the most important objects interacting with machine activities after clearcutting; We develop a flowchart showing how a computer-aided system using clearcut object identification should be executed.
Abstract | Full text in HTML | Full text in PDF | Author Info
After clearcutting, machines traffic the clearcut conducting different silvicultural activities. Many objects on a forest clearcut (slash residues, stones, stumps and roots) may disturb e.g. site preparation and planting. This paper describes properties and characteristics of these objects. A flowchart was developed that describes a possible computer-aided system that identifies the objects, and ultimately, makes a machine avoid or target them. A system for obstacle identification creates conditions for further technical development and (semi)automation of e.g., site preparation, mechanized planting, and stump removal.
  • Lideskog, Division of Product and Production Development, Luleå University of Technology, SE-97187 Luleå, Sweden E-mail: hakan.lideskog@ltu.se (email)
  • Ersson, Department of Forest Biomaterials and Technology, SLU, SE-90183 Umeå, Sweden E-mail: back.tomas.ersson@slu.se
  • Bergsten, Department of Forest Biomaterials and Technology, SLU, SE-90183 Umeå, Sweden E-mail: urban.bergsten@slu.se
  • Karlberg, Division of Product and Production Development, Luleå University of Technology, SE-97187 Luleå, Sweden E-mail: magnus.karlberg@ltu.se
article id 1126, category Research note
Rebecka Mc Carthy, Per Magnus Ekö, Lars Rytter. (2014). Reliability of stump sprouting as a regeneration method for poplars: clonal behavior in survival, sprout straightness and growth. Silva Fennica vol. 48 no. 3 article id 1126. https://doi.org/10.14214/sf.1126
Keywords: Populus spp.; coppice regeneration; vegetative regeneration; short-rotation forestry
Highlights: Clonal differences were found in stump sprout survival; Clonal differences occurred in sprout straightness; It is uncertain if the ability to produce a high number of stump sprouts is beneficial.
Abstract | Full text in HTML | Full text in PDF | Author Info
The interest in poplars (Populus spp.) has increased globally as a result of the current incentive to expand the use of biomass for energy. So far, knowledge about poplar stump sprouting is generally poor. The objective of this study was to investigate poplar stump sprouting as regeneration method in harvested plantations. A 19-year-old poplar clonal test, including 23 different clones, on former arable land was harvested in the winter. After one year, stumps were assessed for sprout survival, sprout straightness, number of sprouts, sprout height and basal diameter. Differences between clones were found for survival and sprout straightness. Clonal differences were also found in mean size of sprouts (basal diameter and height). However, no differences were found between clones when comparing size of the tallest sprout or number of sprouts produced on stumps. This study indicates that stump sprouts from various poplar clones differ in survival, sprout straightness and mean growth, which should be taken into account when planning for the next tree generation. The study needs to be complemented with other studies to better assess sprouting characteristics for different clones.
  • Mc Carthy, Forestry Research Institute of Sweden, Skogforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden & Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: rebecka.mccarthy@skogforsk.se (email)
  • Ekö, Southern Swedish Forest Research Centre, Swedish University of Agricultural Sciences, P.O. Box 49, SE-230 53 Alnarp, Sweden E-mail: Per.Magnus.Eko@slu.se
  • Rytter, Forestry Research Institute of Sweden, Skogforsk, Ekebo 2250, SE-268 90 Svalöv, Sweden E-mail: lars.rytter@skogforsk.se
article id 1125, category Research note
Anssi Krooks, Sanna Kaasalainen, Ville Kankare, Marianna Joensuu, Pasi Raumonen, Mikko Kaasalainen. (2014). Predicting tree structure from tree height using terrestrial laser scanning and quantitative structure models. Silva Fennica vol. 48 no. 2 article id 1125. https://doi.org/10.14214/sf.1125
Keywords: remote sensing; terrestrial lidar; tree modelling; branch size distribution
Highlights: The analysis of tree structure suggests that trees of different height growing in similar conditions have similar branch size distributions; There is potential for using the tree height information in large-scale estimations of forest canopy structure.
Abstract | Full text in HTML | Full text in PDF | Author Info
We apply quantitative structure modelling to produce detailed information on branch-level metrics in trees. Particularly we are interested in the branch size distribution, by which we mean the total volume of branch parts distributed over the diameter classes of the parts. We investigate the possibility of predicting tree branch size distributions for trees in similar growing conditions. The quantitative structure model enables for the first time the comparisons of structure between a large number of trees. We found that the branch size distribution is similar for trees of different height in similar growing conditions. The results suggest that tree height could be used to estimate branch size distribution in areas with similar growing conditions and topography.
  • Krooks, Finnish Geodetic Institute, Geodeetinrinne 2, FI–02431 Masala, Finland E-mail: Anssi.Krooks@fgi.fi
  • Kaasalainen, Finnish Geodetic Institute, Geodeetinrinne 2, FI–02431 Masala, Finland E-mail: Sanna.Kaasalainen@fgi.fi (email)
  • Kankare, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014 Helsinki, Finland E-mail: ville.kankare@helsinki.fi
  • Joensuu, Department of Forest Sciences, University of Helsinki, P.O. Box 27, FI-00014 Helsinki, Finland E-mail: marianna.joensuu@alumni.helsinki.fi
  • Raumonen, Tampere University of Technology, Department of Mathematics, P.O. Box 553, Tampere, FI-33101, Finland E-mail: Pasi.Raumonen@tut.fi
  • Kaasalainen, Tampere University of Technology, Department of Mathematics, P.O. Box 553, Tampere, FI-33101, Finland E-mail: Mikko.Kaasalainen@tut.fi
article id 1053, category Research note
Aleksey Fedorkov. (2014). Vitality and height growth of two Larix species and provenances in a field trial located in north-west Russia. Silva Fennica vol. 48 no. 1 article id 1053. https://doi.org/10.14214/sf.1053
Keywords: tree height; provenance trial; Russian larch; tree condition
Highlights: Differences in tree vitality among provenances were insignificant (p > 0.05) at an early age; The provenance effect was significant (p < 0.05) for total height; These findings were in agreement with those of Lukkarinen et al. (2010) using the same material at the same age for field trial located in Punkaharju (Finland).
Abstract | Full text in HTML | Full text in PDF | Author Info
Vitality and height of Larix species and provenances originating from Russia were estimated in a 5-yr field trial performed in the Komi Republic (north-west Russia) using a fully randomized single-tree plot design with 7–8 blocks. Tree provenance had no significant (p > 0.05) effect on tree vitality, though for Larix sukazcewii originating from the European part of Russia, trees from the northern regions were more vital than those from southern regions. Provenance was a significant (p < 0.05) factor for height, where the average height of 136 cm varied considerably (168 cm for trees from Nizhnij Novgorod and 111 cm for trees from Ufa). There were no significant correlations when vitality and height were compared to geographic and climatic variables for the locations.
  • Fedorkov, Institute of Biology, Komi Science Center, Russian Academy of Sciences, Syktyvkar, 167982, Kommunisticheskaya st., 28, Russia E-mail: fedorkov@ib.komisc.ru (email)
article id 1026, category Research note
Leif Nutto, Ricardo A. Malinovski, Martin Brunsmeier, Felipe Schumacher Sant’Anna. (2013). Ergonomic aspects and productivity of different pruning tools for a first pruning lift of Eucalyptus grandis Hill ex Maiden. Silva Fennica vol. 47 no. 4 article id 1026. https://doi.org/10.14214/sf.1026
Keywords: forest management; wood quality; work science; handsaw; shear
Highlights: Pruning of hardwoods coming from forest plantations is becoming more and more important in Brazil to replace scarce wood from tropical forests; Evaluating productivity of different pruning tools is essential for the economic output of plantations managed for high quality wood; Pruning activities of forest workers can be classified as “hard” or “very hard work”. Depending on the tools used physical long-term damages may be prevented.
Abstract | Full text in HTML | Full text in PDF | Author Info
For the substitution of wood from tropical rainforests pruning of Eucalyptus for producing valuable hardwoods in short rotation plantations has become important. Existing tools and ergonomic aspects of pruning were not yet well analysed under these conditions. The objective of the study is to evaluate the productivity and ergonomics of three different pruning tools in a pruning lift up to 3 m in height. The trees used in the study came from an 18-month-old clonal Eucalyptus grandis stand planted in a 5.0 x 2.8 m spacing. Two manual pruning tools and an electric shear were tested for productivity by using time studies. Ergonomic aspects were evaluated by two test persons using pulse meter equipment. The highest productivity could be shown for the electric shear (236 trees per working day), followed by the manual shear (196 trees/day) and the handsaw (180 trees/day). The heartbeat rate of the two test persons ranged from a level of “very hard work” for the manual tools to “middle hard” and “hard work” for the electric shear. The workload level to achieve the productivity currently reached in practice using purely manual tools is extremely high, exceeding the permanent working capacity of the operators and leading to physical degradation on the long run.
  • Nutto, Department for Forestry, Federal University of Paraná, Av. Pref. Lothário Meissner, 632, Curitiba, PR, Brazil E-mail: lnutto.ufpr@gmail.com (email)
  • Malinovski, Department for Forestry, Laboratory of Forest Operations, Federal University of Paraná, Av. Prefeito Lothario Meissner, 900, Curitiba, PR, Brazil E-mail: ricardo@colheitademadeira.com.br
  • Brunsmeier, Chair of Forest Utilization, University of Freiburg, Werthmannstraße 6, 79085 Freiburg, Germany E-mail: martin.brunsmeier@fobawi.uni-freiburg.de
  • Schumacher Sant’Anna, Department for Forestry, Laboratory of Forest Operations, Federal University of Paraná, Av. Prefeito Lothario Meissner, 900, Curitiba, PR, Brazil E-mail: felipe@colheitademadeira.com.br
article id 1024, category Research note
Petter Nilsen, Line Tau Strand. (2013). Carbon stores and fluxes in even- and uneven-aged Norway spruce stands. Silva Fennica vol. 47 no. 4 article id 1024. https://doi.org/10.14214/sf.1024
Keywords: ground vegetation; C storage; C sequestration; CO2 efflux; litter fall
Highlights: Long term (81 years) C sequestration is slightly higher in an even-aged compared to an uneven-aged spruce stand; The even-aged stand has at 81 years age a slightly lower soil C content than the uneven-aged stand; Present C fluxes indicate that the difference in long term C sequestration will increase in favour the even-aged stand if final felling is postponed.
Abstract | Full text in HTML | Full text in PDF | Author Info
This investigation compares present C stores, fluxes and historic tree C sequestration in an uneven-aged and an even-aged Norway spruce stand under similar high productive soil conditions in south-eastern Norway. A selection cutting system has been performed in the uneven-aged forest stand for 81 years and the even-aged stand was established after clear-cutting 81 years ago. Timber productivity has been measured in the uneven stand for 81 years and in the even-aged stand for 52 years. C storage was determined based on tree measurements, tree biomass functions, soil samples and C analyses from trees and soil. Litter fall was sampled during one year and CO2 efflux from the soil was measured during one growing season. The present tree C storage (including roots) was 210 Mg C ha-1 in the even-aged stand and 76 Mg C ha-1 in the uneven-aged stand, while the corresponding figures for C in the soil was 178 and 199 Mg C ha-1. The long term timber production in the uneven-aged stand was measured to be 95% of the even-aged stand and the difference in net C sequestration was 37 Mg ha-1 in an 81 year period in favour the even-aged stand. The highest present CO2 efflux from soil was measured in the even-aged stand. The total net C sequestered in trees during 81 years minus the present soil C-stock accounts to 16 Mg ha-1 in favour the even-aged system
  • Nilsen, The Research Council of Norway, P.O. Box 2700, St. Hanshaugen, N-0131 Oslo, Norway E-mail: pn@rcn.no (email)
  • Strand, Department of Plant and Environmental Sciences, Norwegian University of Life Sciences, P.O. Box 5003, N-1432 Ås, Norway E-mail: line.strand@umb.no
article id 1017, category Research note
Jeanette Edlund, Urban Bergsten, Hans Arvidsson. (2013). A forest machine bogie with a bearing capacity dependent contact area: acceleration and angular orientation when passing obstacles and drawbar pull force and free rolling resistance on firm ground. Silva Fennica vol. 47 no. 3 article id 1017. https://doi.org/10.14214/sf.1017
Keywords: ground damages; soft ground; traction force
Highlights: The Long Tracked Bogie principle (LTB) has low contact area on firm ground with low load, it increases when higher traction force is needed and on softer soil; Free rolling resistance on firm ground was 60% of the value for a conventional bogie; LTB appears to pass wider ditches/cavities, more smoothly with lower pitch angle, than a conventional bogie.
Abstract | Full text in HTML | Full text in PDF | Author Info
The Long Tracked Bogie with contact area dependent on bearing capacity was compared to a conventional bogie. Two unloaded Vimek 608 forwarders with different bogies and with the traction from the front wheel removed were compared. Two high obstacles, 0.1 and 0.2 m high, respectively and 0.15 m in width, and two deep obstacles/ditches with a depth of 0.2 m and a width 1 and 1.5 m were used for tests. Towing tests on flat ground were done by connecting the machines to each other with a load cell in between.  There were no or small differences in acceleration when passing obstacles between the two types of bogie. LTB passed wider ditches/cavities with lower pitch angles (one bogie/side passing) and 0.2 m obstacles with higher roll angles than a conventional bogie. On firm ground, free rolling resistance of the LTB was about 60% of the resistance of the conventional bogie. The drawbar pull force for the LTB was indicated to be a few percentage units higher than for the conventional bogie when it was driving with a towed machine acting as a braking force. The LTB principle might yield opportunities to improve the way we construct bogies for forest machines. Even if the contact area is low on firm ground when the machine is running with low load, it increases when higher traction force is needed and on softer soil. Further field tests are needed to evaluate the LTB when used on soft ground and with higher load as well.
  • Edlund, Sveaskog AB, SE-941 86 Piteå, Sweden E-mail: jeanette.edlund@sveaskog.se
  • Bergsten, Swedish University of Agricultural Sciences, Department of Silviculture, SE-901 83 Umeå, Sweden E-mail: urban.bergsten@slu.se (email)
  • Arvidsson, SMP Umeå, SE-904 03 Umeå, Sweden E-mail: hans.arvidsson@smp.sp.se
article id 974, category Research note
Kristina Ahnlund Ulvcrona, Lars Karlsson, Ingegerd Backlund, Urban Bergsten. (2013). Comparison of silvicultural regimes of lodgepole pine (Pinus contorta) in Sweden 5 years after precommercial thinning. Silva Fennica vol. 47 no. 3 article id 974. https://doi.org/10.14214/sf.974
Keywords: direct seeding; forest production; corridor thinning; biomass harvest; PCT
Highlights: Management regimes can serve different purposes such as biomass production, pulp and timber production or a combination of those; 30 tons biomass or 38–45 m3 stem volume ha–1 could be derived by schematic corridor thinning (70%) at year 20; Producing large amounts of biomass early in the rotation period does not exclude a conversion into pulp and timber production.
Abstract | Full text in HTML | Full text in PDF | Author Info
Early effects (stem volume, mean diameter at breast height weighted against basal area (Dgv) (Dgv), biomass and damage frequency) of different silvicultural regimes 18-19 years after direct seeding of lodgepole pine in northern Sweden were analysed. A Conventional regime, (i) precommercial thinning (PCT) to 2200 stems ha-1, was compared to: (ii) High biomass production (15 300 stems ha-1, no PCT) with and without corridor thinning at year 20, (iii) production of Large dimension trees (PCT to 1700 stems ha-1), (iv), Combined high biomass production and production of conventional round wood (PCT to 4500 stems ha-1). PCT was done 15 yrs after direct seeding for all PCT treatments. Local biomass functions showed that the regimes aiming at High biomass production displayed ca 144-157% more biomass and 134-143% more stem volume than the Conventional and Large dimension regimes (ca 21 tons and 31 m3 ha-1). Dgv for the 1000 (9.2 cm) and 2000 (8.3 cm) largest trees ha-1 appeared unaffected by regime. By schematic corridor thinning (70% of the total area) at year 20 in the High biomass regime, 27-32 tons of biomass ha-1 and 38-45 m3 ha-1 could be derived while still having a Dgv of the 1000 largest trees ha-1 of about 8 cm. Therefore, this study indicates that it is possible to produce and harvest large amounts of biomass and stem volume early in the rotation period without excluding later pulp and timber production. This initial regime comparison should be continued over time.
  • Ahnlund Ulvcrona, SLU, Forest Biomaterials and Technology, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: kristina.ulvcrona@slu.se (email)
  • Karlsson, SLU, Forest Biomaterials and Technology, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: lars.karlsson@slu.se
  • Backlund, SLU, Forest Biomaterials and Technology, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: ingegerd.backlund@slu.se
  • Bergsten, SLU, Forest Biomaterials and Technology, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: urban.bergsten@slu.se
article id 960, category Research note
Shou-Qin Sun, Liang Peng, Gen-Xu Wang, Yan-Hong Wu, Jun Zhou, Hai-Jian Bing, Dong Yu, Ji Luo. (2013). An improved open-top chamber warming system for global change research. Silva Fennica vol. 47 no. 2 article id 960. https://doi.org/10.14214/sf.960
Keywords: climate change; temperature; open-top chamber; OTC-cable system; forest ecosystem
Abstract | Full text in HTML | Full text in PDF | Author Info
This study is an assessment of an improved temperature warming system developed to enhance global warming research-based forest ecosystem and soil ecophysiological experiments. The architecture couples a standard open-top chamber (OTC) with a heating cable. A 16 m wire cable with an 18 W m-1 and 288 W h-1 power rating was coiled around a polyvinyl chloride (PVC) pipe 2.5 m in length and 3.5 cm in diameter. The pipe was reshaped into a circle and fixed inside the OTC at a height of 15 cm. PVC pipe distance to plants was 10 to 15 cm while distance to OTC inner walls was 15 cm. The cable was constructed from a heating source with an alloy resistance wire, an aluminum foil and copper wire shielded layer, a crosslinking polyethylene inner insulator, a PVC coating, and a tinned copper grounding wire. After the cable is powered up, air and soil inside the OTC-cable system is heated by conductivity. Temperature is manipulated according to the voltage and resistance of the cable. The OTC-cable system was developed to examine plant reaction to an increase in air and soil temperatures by 2.84 °C and 1.83 °C, respectively. Temperature values are adjustable by changing cable and PVC pipe length. It offers a new, affordable, low energy consumption and low running cost method by which to study climate change effects on forest ecosystems. This method is especially useful for application in forest ecosystems of many developing countries or in many remote areas of developed countries where the feasibility in supplying sufficient power from local power grids is questionable.
  • Sun, Key Laboratory of Mountain Surface Processes and Ecological Regulation, Institute of Mountain Hazards and Environment, Chinese Academy of Sciences, No. 9, Block 4, South Renmin Road, Chengdu, China, 610041 E-mail: shouqinsun@imde.ac.cn (email)
  • Peng, Horticulture and Landscape College, Hunan Agricultural University, Furong District, Changsha, China, 410128 E-mail: keith215@126.com
  • Wang, Key Laboratory of Mountain Surface Processes and Ecological Regulation, Institute of Mountain Hazards and Environment, Chinese Academy of Sciences, Chengdu, China E-mail: cookiedot@sina.cn
  • Wu, Key Laboratory of Mountain Surface Processes and Ecological Regulation, Institute of Mountain Hazards and Environment, Chinese Academy of Sciences, Chengdu, China E-mail: free2001@tom.com
  • Zhou, Key Laboratory of Mountain Surface Processes and Ecological Regulation, Institute of Mountain Hazards and Environment, Chinese Academy of Sciences, Chengdu, China E-mail: haitaosun@sohu.com
  • Bing, Key Laboratory of Mountain Surface Processes and Ecological Regulation, Institute of Mountain Hazards and Environment, Chinese Academy of Sciences, Chengdu, China E-mail: 78186181@qq.com
  • Yu, Key Laboratory of Mountain Surface Processes and Ecological Regulation, Institute of Mountain Hazards and Environment, Chinese Academy of Sciences, Chengdu, China E-mail: dongdyu@sohu.com
  • Luo, Key Laboratory of Mountain Surface Processes and Ecological Regulation, Institute of Mountain Hazards and Environment, Chinese Academy of Sciences, Chengdu, China E-mail: 1254157095@qq.com
article id 897, category Research note
Lars Lundqvist, Susanne Spreer, Christer Karlsson. (2013). Volume production in different silvicultural systems for 85 years in a mixed Picea abies–Pinus sylvestris forest in central Sweden. Silva Fennica vol. 47 no. 1 article id 897. https://doi.org/10.14214/sf.897
Keywords: plantation; growth and yield; clear-cutting; silvicultural systems; seed trees; selection system; single-tree selection; diameter limit harvest
Abstract | Full text in HTML | Full text in PDF | Author Info
A long-term comparison of different silvicultural systems was established in 1923 in central Sweden, in an uneven-aged mixed Norway spruce–Scots pine forest (Picea abies (L.) Karst. – Pinus sylvestris L.) with about 85% spruce and 15% pine. The five treatments consisted of two examples of even-aged management 1) clear-cutting followed by planting, and 2) seed tree regeneration, one uneven-aged management 3) selection system, one exploiting treatment 4) diameter limit cut, and 5) one untreated control plot. Each treatment plot was 1 ha, 100 m × 100 m. The plots were measured and managed at irregular intervals, ranging from 7 to 15 years. In 2007–2008 the even-aged treatments and the diameter limit cut were repeated and a new rotation started. Mean annual volume increment during the whole observation period differed widely between the treatments, partly because of differences in species composition over time, with treatment clear-cutting followed by planting at the top, and the control at the bottom. Treatment selection system gave only about 60% of planting, but this was probably largely an effect of too small growing stock during the first roughly 50 years. When the growing stock was increased, periodic annual volume increment increased to about 80% of the mean annual volume increment in the even-aged, planted plot.
  • Lundqvist, Deparment of Forest Ecology and Management, Swedish University of Agricultural Sciences, SE-901 83 Umeå, Sweden E-mail: lars.lundqvist@slu.se (email)
  • Spreer, Sveaskog Förvaltnings AB, Ljusdal, Sweden E-mail: susanne.spreer@sveaskog.se
  • Karlsson, Field Research Unit, Swedish University of Agricultural Sciences, Siljansfors, Sweden E-mail: christer.karlsson@slu.se
article id 915, category Research note
Kalle Kärhä. (2012). Comparison of two stump-lifting heads in final felling Norway spruce stand. Silva Fennica vol. 46 no. 4 article id 915. https://doi.org/10.14214/sf.915
Keywords: productivity; Norway spruce; Picea abies; site preparation; stump harvesting; costs; stump lifting
Abstract | View details | Full text in PDF | Author Info
The use of stump and root wood chips has increased very rapidly in the 21st century in Finland: in the year 2000, the total consumption of stump wood chips for energy generation was 10 GWh, while in 2010 it was approximately 2 TWh. Metsäteho Oy and TTS Research evaluated two stump-lifting devices for the lifting of Norway spruce (Picea abies) stumps. The productivity and costs of stump lifting were determined. There was one base machine with one operator in the time study. When lifting stumps with a diameter of 30 cm, the effective hour productivity of stump lifting was 11.2 m3 solid over bark (sob)/E0 (4.8 tonD/E0) without site preparation using a Väkevä Stump Processor, and when lifting spruce stumps with a diameter of 40 cm, the productivity was 14.9 m3 sob/E0 (6.5 tonD/E0). When the site preparation (mounding) was integrated into lifting work, the stump-lifting productivity decreased 21–27%. The stump-lifting productivity of the other lifting head (Järvinen) was lower than that of the Väkevä Stump Processor. Some development suggestions for the Järvinen lifting head were presented and discussed. The cost calculations showed that stump-lifting costs are extremely high when stump diameter is less than 20 cm. Therefore, the study recommended a change in the current stump-harvesting guidelines of Finland: The study suggested that all the stumps with a diameter less than 20 cm should be left on the harvesting site.
  • Kärhä, Stora Enso Wood Supply Finland, Talvikkitie 40 C, FI-01300 Vantaa, Finland E-mail: kalle.karha@storaenso.com (email)
article id 914, category Research note
Ingegerd Backlund, Urban Bergsten. (2012). Biomass production of dense direct-seeded lodgepole pine (Pinus contorta) at short rotation periods. Silva Fennica vol. 46 no. 4 article id 914. https://doi.org/10.14214/sf.914
Keywords: biomass production; lodgepole pine; direct seeding; Pinus contorta; biorefinery; short rotation
Abstract | View details | Full text in PDF | Author Info
Lodgepole pine (Pinus contorta) is a fast-growing species that is suitable for producing woody biomass in Nordic countries. Direct seeding of this species is cheaper than planting and creates dense, stable stands. The objective of this study was to quantify the stem volume and biomass production of direct seeded lodgepole pine stands grown under different site conditions with different stem densities, at an age that would permit extensive harvesting of biomass. A circle-plot inventory was performed in 16 of the oldest direct seeded lodgepole pine stands in mid-northern Sweden. Stemwood production of almost 200 m3/ha was achieved on average on the best sites, rising to about 300 m3/ha for the best circle-plots within 30 years of direct seeding despite the fact that pre-commercial thinning was made once or twice. This corresponds to 100 and 140 tons of dry weight biomass/ha, respectively. Higher stand stem densities (≥3000 st/ha) yielded more biomass with only slight reductions in diameter at breast height. The development of stem volume with respect to dominant height in direct seeded stands was becoming comparable to that in planted stands with similar spacing. It therefore seems that there is an unutilized potential for cost-effectively growing lodgepole pine in dense stands for biomass production after direct seeding. It may be possible to devise regimes for short(er) rotation forestry that would yield substantial amount of inexpensive biomass for biorefineries within a few decades.
  • Backlund, Department of Forest Ecology and Management, Swedish University of Agricultural Sciences, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: ingegerd.backlund@slu.se (email)
  • Bergsten, Department of Forest Ecology and Management, Swedish University of Agricultural Sciences, Skogsmarksgränd, SE-901 83 Umeå, Sweden E-mail: urban.bergsten@slu.se
article id 442, category Research note
Emil Modig, Bo Magnusson, Erik Valinger, Jonas Cedergren, Lars Lundqvist. (2012). Damage to residual stand caused by mechanized selection harvest in uneven-aged Picea abies dominated stands. Silva Fennica vol. 46 no. 2 article id 442. https://doi.org/10.14214/sf.442
Keywords: selection cutting; logging damage; continuous cover management; residual stand; logging methods
Abstract | View details | Full text in PDF | Author Info
Permanent field plots were established in two uneven-aged Norway spruce (Picea abies (L.) Karst) dominated stands in west-central Sweden. The objective was to quantify level and type of damage caused by harvesting and to quantify the difference between two treatments: T20) only skid road harvest (20 m distance between ca. 4 m wide roads), and T40) skid road harvest (40 m distance between ca. 4 m wide roads) combined with thinning between the roads. In T40, the goal was to harvest approximately the same standing volume as in T20. After harvest, two circular sample plots (radius 18 m, i.e. 1018 m2) were established at random locations within each treated area. All mechanical damage on the stem caused by harvest was measured and registered, including bark stripping larger than 15 cm2, stem broken or split, and tearing of branches causing damage on the stem. About 70–90 per cent of the damaged trees were smaller than 15 cm dbh. Very few trees larger than 25 cm dbh were damaged. In T20, more than 50 per cent of the damaged trees were located less than 5 m from the skid road, compared to less than 25 per cent for T40, in which more than 50 per cent of the damaged trees were located 5–10 m from the skid road. Creating only half the number of skid roads caused no more damage, and was probably more profitable because mean stem volume was about 1.5 times larger than in T20.
  • Modig, Statens fastighetsverk, Jokkmokk, Sweden E-mail: em@nn.se
  • Magnusson, Skogsstyrelsen, Bräcke, Sweden E-mail: bm@nn.se
  • Valinger, Deparment of Forest Ecology and Management, SLU, SE-901 83 Umeå, Sweden E-mail: ev@nn.se
  • Cedergren, Mariehamn, Åland E-mail: jc@nn.se
  • Lundqvist, Deparment of Forest Ecology and Management, SLU, SE-901 83 Umeå, Sweden E-mail: lars.lundqvist@slu.se (email)
article id 61, category Research note
Janine Schweier, Gero Becker. (2012). Harvesting of short rotation coppice – harvesting trials with a cut and storage system in Germany. Silva Fennica vol. 46 no. 2 article id 61. https://doi.org/10.14214/sf.61
Keywords: productivity; harvest operations; marginal sites; SRC; Stemster MK III
Abstract | View details | Full text in PDF | Author Info
Short rotation coppice (SRC) harvesting techniques are available in Germany, but broad experience and knowledge about machine performance and the related effective costs of harvesting operations are still missing. This information is crucial, as harvesting costs strongly influence the economic performance of the overall supply chain. Therefore, it was the aim of this study to collect and analyze productivity data of different harvesting systems for SRC. The combined cut and chip system on the one hand and the cut and storage system on the other hand were studied by literature review. Several studies analyze the combined cut and chip systems and the reported machine productivities showed great variations. The average was 30 green tons per scheduled machine hour (gt smh–1). Few studies are analysing the cut and storage system. They report that machines still are under development and that further research is needed. Therefore, time studies of harvesting operations using the cut and storage system were carried out. Five trials were performed with the harvesting machine “Stemster MK III” developed by Nordic Biomass. The share of productive working time was 85% and the average productivity was 21 gt smh–1. These results were compared with values from the literature. Resulting harvesting costs were calculated per oven dry ton (euros odt–1). The advantages and disadvantages of both harvesting systems are highlighted.
  • Schweier, Albert-Ludwigs-University Freiburg, Institute of Forest Utilisation and Work Science, Freiburg, Germany E-mail: janine.schweier@fobawi.uni-freiburg.de (email)
  • Becker, Albert-Ludwigs-University Freiburg, Institute of Forest Utilisation and Work Science, Freiburg, Germany E-mail: gb@nn.de
article id 60, category Research note
Dominik Röser, Blas Mola-Yudego, Robert Prinz, Beatrice Emer, Lauri Sikanen. (2012). Chipping operations and efficiency in different operational environments. Silva Fennica vol. 46 no. 2 article id 60. https://doi.org/10.14214/sf.60
Keywords: wood-fuel logistics; forest machinery; bioenergy supply
Abstract | View details | Full text in PDF | Author Info
This research analyses the productivity of energy wood chipping operations at several sites in Austria and Finland. The aim of the work is to examine the differences in productivity and the effects of the operational environment for the chipping of bioenergy at the roadside. Furthermore, the study quantifies the effects of different variables such as forest energy assortments, tree species, sieve size and machines on the overall productivity of chipping. The results revealed that there are significant differences in the chipping productivity in Austria and Finland which are largely based on the use of different sieve sizes. Furthermore, the different operational environments in both countries, as well as the characteristics of the raw material also seem to have an effect on productivity. In order to improve the chipping productivity, particularly in Central European conditions, all relevant stakeholders need to work jointly to find solutions that will allow a greater variation of chip size. Furthermore, in the future more consideration has to be given to the close interlinkage between the chipper, crane and grapple. As a result, investments costs can be optimized and operational costs and stress on the machines reduced.
  • Röser, Finnish Forest Research Institute, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: dominik.roser@metla.fi (email)
  • Mola-Yudego, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: bmy@nn.fi
  • Prinz, Finnish Forest Research Institute, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: rp@nn.fi
  • Emer, University of Padova, Department of Land, Agriculture and Forest Systems, Legnaro (PD), Italy E-mail: be@nn.it
  • Sikanen, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: ls@nn.fi
article id 71, category Research note
Jaana Luoranen, Heli Viiri. (2012). Soil preparation reduces pine weevil (Hylobius abietis L.) damage on both peatland and mineral soil sites one year after planting. Silva Fennica vol. 46 no. 1 article id 71. https://doi.org/10.14214/sf.71
Keywords: mineral soil; peatland; pine weevil; soil preparation; vole
Abstract | View details | Full text in PDF | Author Info
We studied pine weevil (Hylobius abietis (L.)) feeding damage to Norway spruce and Scots pine seedlings planted in regeneration areas located on peatlands or on mineral soil sites in Southern and Central Finland. The survey included two planting years and a total of 60 regeneration areas (40 areas on peatlands and 20 on mineral soil sites). Some sites classified as peatland were as transformed or transforming drained peatlands that also contained mineral soil on a prepared surface. The soil preparation method, type of surface material around a seedling, pine weevil, vole-induced or other damage and the health of each seedling were observed in systematically selected circular sample plots. There was slightly more pine weevil damage on peatland than on mineral soil sites. More seedlings were damaged on unprepared peat and humus than on a prepared surface. Seedlings surrounded by a prepared surface had a slightly greater risk of being gnawed by pine weevil when planted on prepared peat compared to planting on prepared mineral soil. Vole damage was observed only in one region during one year. Mounded areas had slightly less vole damage than patched areas. In order to reduce damage caused by pine weevils and voles, it is important to scarify the regeneration area properly before insecticide-treated seedlings are planted. Mounding and patching are recommended so that seedlings can be planted in mineral soil whenever possible, even in the case of peatlands.
  • Luoranen, Finnish Forest Research Institute, Suonenjoki Unit, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: jaana.luoranen@metla.fi (email)
  • Viiri, Finnish Forest Research Institute, Joensuu Unit, Joensuu, Finland E-mail: hv@nn.fi
article id 92, category Research note
Adam Felton, Erik Andersson, David Ventorp, Matts Lindbladh. (2011). A comparison of avian diversity in spruce monocultures and spruce-birch polycultures in southern Sweden. Silva Fennica vol. 45 no. 5 article id 92. https://doi.org/10.14214/sf.92
Keywords: biodiversity; conservation; sustainable forestry
Abstract | View details | Full text in PDF | Author Info
The replacement of some spruce monocultures with stands composed of planted Norway spruce (Picea abies) and naturally regenerated birch (Betula spp.) has a range of potential benefits, but the implications for biodiversity are generally unknown. Here we conduct a paired replicated study in southern Sweden of the avian biodiversity found within Norway spruce monocultures, and within Norway spruce stands possessing approximately 20% birch. Our research leads us to three findings. First, avian diversity was significantly higher in the spruce–birch polycultures. Second, spruce–birch polycultures exclusively attracted broadleaf-associated bird species and retained the majority of conifer-associated bird species found in the spruce monocultures. Third, avian biodiversity within the spruce–birch polycultures did not incorporate threatened taxa. We suggest that in addition to the apparent benefits for stand level diversity, widespread use of spruce–birch polycultures could provide a means of softening the matrix for broadleaved-associated species, while concurrently providing an increased broadleaf base from which future conservation actions could be implemented. Our results are relevant to multi-use forestry, and recent policy initiatives by forest certification agencies which aim to increase broadleaf-associated biodiversity within conifer-dominated production forest landscapes.
  • Felton, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, Alnarp, Sweden E-mail: adam.felton@ess.slu.se (email)
  • Andersson, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, Alnarp, Sweden E-mail: ea@nn.se
  • Ventorp, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, Alnarp, Sweden E-mail: dv@nn.se
  • Lindbladh, Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre, Alnarp, Sweden E-mail: ml@nn.se
article id 91, category Research note
Raisa Mäkipää, Tapio Linkosalo. (2011). A non-destructive field method for measuring wood density of decaying logs. Silva Fennica vol. 45 no. 5 article id 91. https://doi.org/10.14214/sf.91
Keywords: coarse woody debris; carbon stock; decaying wood; wood decomposition; penetrometer; pilodyn
Abstract | View details | Full text in PDF | Author Info
Decaying dead wood density measurements are a useful indicator for multiple purposes, such as for estimating the amount of carbon in dead wood and making predictions of potential diversity of dead wood inhabiting fungi and insects. Currently, qualitative decay phases are used as wood density estimates in many applications, since measuring the density is laborious. A quantitative measure of density would, however, be preferred over the qualitative one. Penetrometers, which are commonly used for measuring the density of standing trees, might also be applicable to dead wood density measurements. We tested the device for making quick, quantitative measurements of decaying logs. The penetrometer measures the depth into which a pre-loaded spring forces a pin in the wood. We tested pins of 5 and 10 mm diameter together with an original 2.5 mm pin and compared the results with gravimetric density measurements of the sample logs. Our results suggest that the standard pin works for less decayed wood, but for more decomposed wood, the thicker 5 mm pin gave more reliable estimates when the penetration measures were converted to densities with a linear regression function (R2 = 0.62, F = 82.9, p = 0.000). The range of wood densities successfully measured with the 5 mm pin was from 180 to 510 kg m–3. With the 10 mm pin, the measuring resolution of denser wood was compromised, while the improvement at the other end of density scale was not large. As a conclusion, the penetrometer seems to be a promising tool for quick density testing of decaying logs in field, but it needs to be modified to use a thicker measuring pin than the standard 2.5 mm pin.
  • Mäkipää, The Finnish Forest Research Institute, Vantaa, Finland E-mail: raisa.makipaa@metla.fi (email)
  • Linkosalo, The Finnish Forest Research Institute, Vantaa, Finland E-mail: ts@nn.fi
article id 118, category Research note
Audrius Menkis, Remigijus Bakys, Vaidotas Lygis, Rimvydas Vasaitis. (2011). Mycorrhization, establishment and growth of outplanted Picea abies seedlings produced under different cultivation systems. Silva Fennica vol. 45 no. 2 article id 118. https://doi.org/10.14214/sf.118
Keywords: forest nursery; cultivation system; ectomycorrhizal fungi; Picea abies seedlings
Abstract | View details | Full text in PDF | Author Info
In forest nurseries seedlings are commonly produced using different cultivation systems which may have a profound effect on the extent of root colonisation by ectomycorrhizal (ECM) fungi and, consequently, on subsequent performance of the outplanted seedlings under field conditions. In this study, we compare effects of bare-root and containerised cultivation systems on mycorrhization, establishment and growth of nursery-produced Picea abies. One hectare experimental plantation was established on poor fertility sandy site. In total, 1250 seedlings of each treatment were planted in rows as ten individual replicates. The results of this study showed that production of P. abies seedlings using containerised system can yield abundant ECM colonisation of seedling roots and significantly improve seedling survival in the field. Some reduction in height increment may occur during the first years as a possible cost for support of ECMs. Study demonstrated that selection of proper cultivation system might result in similar or higher mycorrhization and survival rates of outplanted seedlings than achieved by expensive and laborious artificial mycorrhization.
  • Menkis, Department of Forest Mycology and Pathology, Uppsala BioCenter, Swedish University of Agricultural Sciences, P.O. Box 7026, SE-750 07 Uppsala, Sweden E-mail: audrius.menkis@slu.se (email)
  • Bakys, Department of Forest Mycology and Pathology, Uppsala BioCenter, Swedish University of Agricultural Sciences, P.O. Box 7026, SE-750 07 Uppsala, Sweden E-mail: rb@nn.se
  • Lygis, Laboratory of Phytopathogenic Microorganisms, Institute of Botany at the Nature Research Centre, Vilnius, Lithuania E-mail: vl@nn.lt
  • Vasaitis, Department of Forest Mycology and Pathology, Uppsala BioCenter, Swedish University of Agricultural Sciences, P.O. Box 7026, SE-750 07 Uppsala, Sweden E-mail: rv@nn.se
article id 37, category Research note
D. N. Avtzis, F. A. Aravanopoulos. (2011). Host tree and insect genetic diversity on the borderline of natural distribution: a case study of Picea abies and Pityogenes chalcographus (Coleoptera, Scolytinae) in Greece. Silva Fennica vol. 45 no. 1 article id 37. https://doi.org/10.14214/sf.37
Keywords: bark beetles; genetic bases; marginal populations; northern Greece; Norway spruce
Abstract | View details | Full text in PDF | Author Info
Picea abies (L.) Karst. and Pityogenes chalcographus constitute a commonly observed host tree–insect association in Eurasia, with the natural distribution of the bark beetle overlapping that of Norway spruce. The southernmost borderline of their distributions occurs in the Elatia forest (Mt. Rodopi, Greece), where these interacting organisms may experience severe conditions due to the effects of climate change. In order to assess the dynamics of this host tree–insect association, the genetic diversity of both organisms was studied. In contrast to previous studies, the assessment of molecular diversity was based on the same mitochondrial gene (Cytochrome Oxidase One) sequence for both host and pest. This analysis revealed a remarkably higher genetic diversity of P. chalcographus compared to that of P. abies, something that renders the insect capable not only of adapting to novel environmental conditions, but even of shifting to other host species. On the contrary, P. abies presented a narrow genetic base, a potential drawback at the southern-most region of the species natural distribution. Synthesizing the preliminary outcome for both organisms, it appears that P. chalcographus exhibits an evolutionary advantage over P. abies, something that should be considered when planning conservation strategies for the relict forest of Elatia.
  • Avtzis, Aristotle University of Thessaloniki, Laboratory of Forest Genetics and Tree Breeding, Thessaloniki, Greece E-mail: dimitrios.avtzis@gmail.com (email)
  • Aravanopoulos, Aristotle University of Thessaloniki, Laboratory of Forest Genetics and Tree Breeding, Thessaloniki, Greece E-mail: FAA@nn.fi
article id 136, category Research note
Tomasz D. Mazgajski, Michal Zmihorski, Katarzyna Abramowicz. (2010). Forest habitat loss and fragmentation in Central Poland during the last 100 years. Silva Fennica vol. 44 no. 4 article id 136. https://doi.org/10.14214/sf.136
Keywords: afforestation; fragmentation; landscape history; forest continuity; deforestation; ancient forest
Abstract | View details | Full text in PDF | Author Info
The process of habitat fragmentation consists of two components – habitat loss and fragmentation per se. Both are thought to be among the most important threats to biodiversity. However, the biological consequences of this process such as species occurrence, abundance, or genetic structure of population are driven by current, as well as previous, landscape configurations. Therefore, historical analyses of habitat distribution are of great importance in explaining the current species distribution. In our analysis, we describe the forest fragmentation process for an area of 178 km2 in the northern part of Mazowsze region of central Poland. Topographical maps from the years 1890, 1957 and 1989 were used. Over the 100-year period, forest coverage in this area changed from 17% to 5.6%, the number of patches increased from 19 to 42, while the area of the forest interior decreased from 1933 ha to 371 ha. The two components of fragmentation were clearly separated in time. Habitat loss occurred mainly during the first period (1890–1957) and fragmentation per se in the second (1957–1989). Moreover, we recorded that only 47.7% of all the currently (in 1989) afforested areas constitute sites where forests previously occurred (in 1890 and 1957). For forest dwelling organisms characterized by low dispersal abilities, the effective forest coverage seems to be a half of the real forest area in the studied landscape. New afforestations should be planned especially to increase those patches which contain ancient forest, where various plants and animals sensitive to fragmentation may have survived.
  • Mazgajski, Polish Academy of Sciences, Museum & Institute of Zoology, Wilcza 64, PL 00-679 Warsaw, Poland E-mail: mazgaj@miiz.waw.pl (email)
  • Zmihorski, Polish Academy of Sciences, Museum & Institute of Zoology, Wilcza 64, PL 00-679 Warsaw, Poland E-mail: mz@nn.pl
  • Abramowicz, Department of Ecology, University of Warsaw, Banacha 2, PL 02-097 Warsaw, Poland E-mail: ka@nn.pl
article id 148, category Research note
Ola Lindroos, Magnus Matisons, Petter Johansson, Tomas Nordfjell. (2010). Productivity of a prototype truck-mounted logging residue bundler and a road-side bundling system. Silva Fennica vol. 44 no. 3 article id 148. https://doi.org/10.14214/sf.148
Keywords: bioenergy; compaction; composite residue logs; densification; road-side landing; slash; supply chain
Abstract | View details | Full text in PDF | Author Info
When recovering logging residues (LR) for bioenergy its density should be increased before road transport, otherwise a low proportion of the trucks’ load capacity will be used. One way this can be currently done is to compress LR into bundles that are forwarded to roadside landing. A less well-developed alternative is to forward loose LR and bundle it at landing. In the presented study, a prototype specifically developed for road-side bundling was found to produce larger, heavier bundles than bundling machinery intended for in-field use (mean length, diameter and raw bulk density 4.7 m, 0.8 m and 285 kg m–3, respectively, with 299–445 kg oven dry matter per bundle). The machine was also at least 30% more productive than previously described in-field bundling systems, producing 14–19 bundles per productive work hour (PWh), equivalent to 5.2–7.8 oven-dry tonnes PWh–1. Bundles were estimated to use 67–86% of an LR truck’s 30 tonnes load capacity, similar to proportions used when transporting loose LR. However, a continuous feeding and compressing process would probably almost double productivity, while longer bundles would enable full use of truck load capacity. With such improvements bundling at road-side could provide a viable alternative to current LR-recovering systems.
  • Lindroos, Department of Forest Resource Management, Swedish University of Agricultural Sciences, Umeå, Sweden E-mail: ola.lindroos@srh.slu.se (email)
  • Matisons, Department of Forest Resource Management, Swedish University of Agricultural Sciences, Umeå, Sweden E-mail: mm@nn.se
  • Johansson, Sveaskog Förvaltnings AB, Vindeln, Sweden E-mail: pj@nn.se
  • Nordfjell, Department of Forest Resource Management, Swedish University of Agricultural Sciences, Umeå, Sweden E-mail: tn@nn.se
article id 223, category Research note
Bum-Jin Park, Takeshi Morikawa, Tomohiro Ogata, Kenji Washida, Mario Iwamoto, Hirohiko Nakamura, Yoshifumi Miyazaki. (2009). Physiological effects of ingesting eucalyptus essential oil with milk casein peptide. Silva Fennica vol. 43 no. 1 article id 223. https://doi.org/10.14214/sf.223
Keywords: relaxation; eucalyptus essential oil; POMS; salivary cortisol; type A behavior pattern
Abstract | View details | Full text in PDF | Author Info
This study was conducted to clarify the effect of eucalyptus essential oil mixed with milk casein peptide food for human physiological relaxation. Fifteen male university students (21.2 ± 0.9 yr) participated in study as subjects. The subjects were given one of two types of experimental drink (peptide + eucalyptus flavor (Pep + EF), and peptide + grapefruit·orange flavor (Pep + G·O), each flavor contains natural essential oil). We measured the change in salivary cortisol concentration and POMS scores before and two hours after taking experimental drink. The results of a Type A behavior pattern test were used to classify subjects. The concentration of salivary cortisol decreased significantly two hours after taking Pep + EF. And Type B showed bigger change than Type A. In conclusion, the results show that eucalyptus essential oil has the effect of relaxation, and that the effects on Type A and Type B are different.
  • Park, Chiba University, Center for Environment, Health and Field Sciences, 277-0882 Kashiwa, Chiba, Japan E-mail: bjpark@faculty.chiba-u.jp (email)
  • Morikawa, Chiba University, Center for Environment, Health and Field Sciences, 277-0882 Kashiwa, Chiba, Japan E-mail: tm@nn.jp
  • Ogata, Chiba University, Center for Environment, Health and Field Sciences, 277-0882 Kashiwa, Chiba, Japan E-mail: to@nn.jp
  • Washida, Chiba University, Center for Environment, Health and Field Sciences, 277-0882 Kashiwa, Chiba, Japan E-mail: kw@nn.jp
  • Iwamoto, Chiba University, Center for Environment, Health and Field Sciences, 277-0882 Kashiwa, Chiba, Japan E-mail: mi@nn.jp
  • Nakamura, Chiba University, Center for Environment, Health and Field Sciences, 277-0882 Kashiwa, Chiba, Japan E-mail: hn@nn.jp
  • Miyazaki, Chiba University, Center for Environment, Health and Field Sciences, 277-0882 Kashiwa, Chiba, Japan E-mail: ym@nn.jp
article id 228, category Research note
Aku Riihelä, Terhikki Manninen. (2008). Measuring the vertical albedo profile of a subarctic boreal forest canopy. Silva Fennica vol. 42 no. 5 article id 228. https://doi.org/10.14214/sf.228
Keywords: boreal forest; albedo; validation; canopy transmission
Abstract | View details | Full text in PDF | Author Info
The validation of airborne and satellite-derived albedo measurements suffers from the fact that the surface albedo of forest is difficult to measure in-situ over large areas. The goal of this study is to examine whether or not the estimation of the surface albedo of a forest stand from ground level measurements is possible. In addition, knowledge about the vertical behavior of albedo, and therefore transmitted solar radiation, is important in the modeling of interactions of sunlight with the forest canopy. Such modeling is useful for forest growth estimations among other things. To achieve these goals, an albedometer set-up capable of vertical albedo profiling has been used to obtain data from a boreal forest stand in Northern Finland during periods in summer 2006 and winter 2007. The results show a strong relationship between the data and fitted power-law regression curves. Power-law regression fits best likely because of the radiation transmission characteristics of boreal forest.
  • Riihelä, Finnish Meteorological Institute, P.O. Box 503, FI-00200 Helsinki, Finland E-mail: aku.riihela@fmi.fi (email)
  • Manninen, Finnish Meteorological Institute, P.O. Box 503, FI-00200 Helsinki, Finland E-mail: tm@nn.fi
article id 269, category Research note
Christian Kiffner, Elisabeth Rössiger, Oliver Trisl, Rainer Schulz, Ferdinand Rühe. (2008). Probability of recent bark stripping damage by red deer (Cervus elaphus) on Norway spruce (Picea abies) in a low mountain range in Germany – a preliminary analysis. Silva Fennica vol. 42 no. 1 article id 269. https://doi.org/10.14214/sf.269
Keywords: forest management; logistic regression; wildlife management
Abstract | View details | Full text in PDF | Author Info
Red deer (Cervus elaphus) can cause considerable damage to forest stands by bark stripping. Here, we examined the probability of bark stripping of susceptible Norway spruce (Picea abies) during winter in relation to local environmental characteristics in the western Harz Mountains, Lower Saxony, Germany. We present the results of a multiple logistic regression model for recent bark stripping damage by red deer which we developed from two systematic cluster sampling inventories after two winter periods along with associated meteorological data and records of bagged deer. Our model suggests that the risk of bark stripping increased significantly (P  0.05) with rising slope angle, cumulating snow depth and increasing index values of red deer population density. Spruces growing in closed forest stands were debarked at a higher probability than spruces located close to forest edges. Further on, spruce stands on eastern slopes had a lower probability of bark damage than spruce stands on northern slopes. Other tested variables (altitude, length of daily solar irradiation, duration of snow cover, age of spruce stand within the age range of 16–50 years) had no significant effect on the probability of new bark stripping. We conclude that red deer in the western Harz Mountains seem to use bark as food resource at preferred locations and in times of low food availability. To improve fit and predictive power of bark stripping models we recommend including stand characteristics. We propose to reduce the population size of red deer in order to diminish bark stripping damages to an economically acceptable level.
  • Kiffner, University Göttingen, Büsgen-Institute, Department of Forest Zoology and Forest Protection incl. Wildlife Biology and Game Management, Büsgenweg 3, 37077 Göttingen, Germany E-mail: ckiffne@gwdg.de (email)
  • Rössiger, University Göttingen, Büsgen-Institute, Department of Forest Zoology and Forest Protection incl. Wildlife Biology and Game Management, Büsgenweg 3, 37077 Göttingen, Germany E-mail: er@nn.de
  • Trisl, Planungsbüro Trisl, In der Schleene 7, 36037 Waake, Germany E-mail: ot@nn.de
  • Schulz, University Göttingen, Büsgen-Institute, Department of Ecological Informatics, Biometry and Forest Growth, Büsgenweg 4, 37077 Göttingen, Germany E-mail: rs@nn.de
  • Rühe, University Göttingen, Büsgen-Institute, Department of Forest Zoology and Forest Protection incl. Wildlife Biology and Game Management, Büsgenweg 3, 37077 Göttingen, Germany E-mail: fr@nn.de
article id 268, category Research note
Adam Boratynski, Katarzyna Marcysiak, Amelia Lewandowska, Anna Jasinska, Grzegorz Iszkulo, Jaroslaw Burczyk. (2008). Differences in leaf morphology between Quercus petraea and Q. robur adult and young individuals. Silva Fennica vol. 42 no. 1 article id 268. https://doi.org/10.14214/sf.268
Keywords: morphological diversity; plant variation; biometry; Quercus; leaf polymorphism
Abstract | View details | Full text in PDF | Author Info
The characters of Quercus robur and Q. petraea leaves are of main taxonomic value and the adult trees of both species can be distinguished on them. However, young individuals, mostly seedlings but also saplings, are told to be undistinguishable or only partly distinguishable on the leaf morphology. The aim of the study was to verify this hypothesis on the basis of biometrical analyses of leaf characteristics of adults trees and saplings in two mixed oak woods, one located close to the north-eastern limit, the other about 400 km inside of the Q. petraea range in Poland. The analysis of discriminations and minimum spanning tree on the squares of Mahalanobis distances were analysed to find differences between Q. robur, Q. petraea and intermediate adults and saplings. The differences between saplings of Q. robur and Q. petraea were found lower than between adult trees. Nevertheless, the biometrical analysis confirmed determination of saplings in the field.
  • Boratynski, Polish Academy of Sciences, Institute of Dendrology, 5 Parkowa str., 62-035 Kórnik, Poland E-mail: borata@man.poznan.pl (email)
  • Marcysiak, Kazimierz Wielki University, Institute of Biology and Environment Protection, 12 Ossolinskich str., 85-064 Bydgoszcz, Poland E-mail: km@nn.pl
  • Lewandowska, Kazimierz Wielki University, Institute of Biology and Environment Protection, 12 Ossolinskich str., 85-064 Bydgoszcz, Poland E-mail: al@nn.pl
  • Jasinska, Polish Academy of Sciences, Institute of Dendrology, 5 Parkowa str., 62-035 Kórnik, Poland E-mail: aj@nn.pl
  • Iszkulo, Polish Academy of Sciences, Institute of Dendrology, 5 Parkowa str., 62-035 Kórnik, Poland E-mail: gi@nn.pl
  • Burczyk, Kazimierz Wielki University, Institute of Biology and Environment Protection, 12 Ossolinskich str., 85-064 Bydgoszcz, Poland E-mail: jb@nn.pl
article id 927, category Research note
Risto Jalkanen, Carmen Büttner, Susanne von Bargen. (2007). Cherry leaf roll virus abundant on Betula pubescens in Finland. Silva Fennica vol. 41 no. 4 article id 927. https://doi.org/10.14214/sf.927
Keywords: Betula pendula; Betula nana; Betula pubescens subsp. czerepanovii; virology
Abstract | View details | Full text in PDF | Author Info
Virus-related symptoms such as vein banding, leaf roll, chlorosis and subsequent necrosis on birch leaves were increasingly recorded throughout Finland since 2002. They are widespread in this country and have also been detected in northern Norway and Sweden. Symptomatic foliage has so far been found on Betula pendula, B. pubescens, B. pubescens subsp. czerepanovii, and B. nana. A Cherry leaf roll virus (CLRV) specific IC-RT-PCR was applied to young leaves, buds and catkins of symptomatic shoots of nineteen pubescent and one silver birch trees grown in the centre of Rovaniemi, Finland. CLRV was found in seventeen B. pubescens trees. This is the first time that B. pubescens has been confirmed to be a host species for CLRV in Finland. Nor has CLRV been recorded earlier in northern Finland.
  • Jalkanen, Finnish Forest Research Institute, Rovaniemi Research Unit, P.O. Box 16, FI-96301 Rovaniemi, Finland E-mail: risto.jalkanen@metla.fi (email)
  • Büttner, Humboldt Universität zu Berlin, Institut für Gartenbauwissenschaften, Fachgebiet Phytomedizin, Lentzeallee 55/57, 14195 Berlin, Germany E-mail: Carmen.Buettner@agrar.hu-berlin.de
  • Bargen, Humboldt Universität zu Berlin, Institut für Gartenbauwissenschaften, Fachgebiet Phytomedizin, Lentzeallee 55/57, 14195 Berlin, Germany E-mail: susanne.von.bargen@agrar.hu-berlin.de
article id 470, category Research note
Paula Jylhä, Juha Laitila. (2007). Energy wood and pulpwood harvesting from young stands using a prototype whole-tree bundler. Silva Fennica vol. 41 no. 4 article id 470. https://doi.org/10.14214/sf.470
Keywords: productivity; energy wood; pulpwood; bundling; integrated harvesting; Fixteri
Abstract | View details | Full text in PDF | Author Info
The productivity of cutting and bundling whole trees using the first prototype of a bundle-harvester comprised of a harwarder as the base machine, an accumulating felling head, and a compacting device was studied in three young stands in order to facilitate the further development of the concept. In addition, the removal and its composition were studied as a means of laying the foundations for developing methods for work rating and measurement on delivery. Bundling enables in-depth integration of pulpwood and energy wood procurement. Both energy wood (crown biomass) and pulpwood can be incorporated into the same bundles, and the subsequent separation of these fractions takes place at the debarking phase at the pulpmill. Bundle-harvesting productivities were relatively low (2.8–3.7 m3/E0-h) when compared to current harvesting technology. Improving working techniques, machine structure, and components showed great potential for increasing the efficiency of the concept. The bundles were dimensionally uniform. Their solid volume varied between 0.350 m3 and 0.513 m3, depending on the bundle assortment and stand properties. Integrating energy wood harvesting with pulpwood harvesting increased removal even by 59 per cent.
  • Jylhä, Finnish Forest Research Institute, Kannus Research Unit, P.O. Box 44, FI-69101 Kannus, Finland E-mail: paula.jylha@metla.fi (email)
  • Laitila, Finnish Forest Research Institute, Joensuu Research Unit, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: jl@nn.fi
article id 311, category Research note
Dan Bergström, Urban Bergsten, Tomas Nordfjell, Tomas Lundmark. (2007). Simulation of geometric thinning systems and their time requirements for young forests. Silva Fennica vol. 41 no. 1 article id 311. https://doi.org/10.14214/sf.311
Keywords: bioenergy; forest technology; multi-stem
Abstract | View details | Full text in PDF | Author Info
In Fennoscandia, large areas that have not been subjected to pre-commercial thinning (PCT), and thus support dense stands, are becoming suitable for harvesting biomass. However, efficient systems for harvesting biomass from young stands have not yet been developed. In order to optimise biomass harvesting it is here hypothesized that the handling unit should not be a single tree but a corridor area, i.e., all trees in a specific area should be harvested in the same crane movement cycle. Three types of corridor harvesting approaches (using accumulating felling heads for geometric harvesting in two different patterns) were compared in terms of time required to fell a corridor of standardised size. Corridors are defined as strips of harvested areas between conventional strip-roads. Harvests were simulated in two types of stands, first thinning (FT) and delayed PCT stands, in which the spatial positions of the trees had been mapped. The differences in simulated time consumption per corridor were minor when the only variable changed was the corridor pattern. However, there were ca. 2-fold and 3-fold differences in simulated time consumption per corridor between the harvesting approaches for the FT stand and the PCT-stand, respectively. Furthermore, area handling (felling head accumulating all trees corridor-wise, with no restrictions on the accumulated number of trees except for a certain load limit) was found to give up to 2.4-fold increases in productivity compared to a single-tree (reference) approach for the FT stand. In conclusion, the simulation results clearly show the benefits of applying area-harvesting systems in young, dense stands.
  • Bergström, SLU, Dept. of Forest Resource Management, SE-901 83 Umeå, Sweden E-mail: db@nn.se (email)
  • Bergsten, SLU, Dept. of Forest Ecology and Management, SE-901 83 Umeå, Sweden E-mail: ub@nn.se
  • Nordfjell, SLU, Dept. of Forest Resource Management, SE-901 83 Umeå, Sweden E-mail: tn@nn.se
  • Lundmark, SLU, Vindeln Experimental Forests, Svartberget Field Station, SE-922 91 Vindeln, Sweden E-mail: tl@nn.se
article id 310, category Research note
Timo Tahvanainen, Kalle Kaartinen, Timo Pukkala, Matti Maltamo. (2007). Comparison of approaches to integrate energy wood estimation into the Finnish compartment inventory system. Silva Fennica vol. 41 no. 1 article id 310. https://doi.org/10.14214/sf.310
Keywords: diameter distribution; thinnings; calibration estimation; compartment inventory
Abstract | View details | Full text in PDF | Author Info
The harvesting of energy wood from young stands is increasing as the demand for renewable wood fuel is growing. Energy wood consists of stems, tree tops, branches and needles, depending on the size of the trees and the logging method used. The current forest inventory and planning systems used in private forests in Finland do not produce estimates of energy wood components. In stands typical for energy wood harvesting, a large share of energy wood consists of trees smaller than the minimum size for pulpwood. In this study, energy wood was included into the calculation system of compartment inventory, and a procedure for simulating the thinning treatments in young stands was developed. The results for six inventory alternatives and prediction of energy wood were compared with the use of inventory material from 37 young stands that have plenty of energy wood. The measurement of additional stand characteristics and the use of a calibration estimation method was tested, as well as the use of plot-level inventory data instead of stand level data. The results showed that the measurement of the number of trees per hectare, in addition to stand basal area and mean diameter, improved the energy wood estimates. The additional minimum and maximum diameters improved the precision of the estimates, but did not affect bias. The removal estimates were more precise when plot-level data was used, rather than stand-level data. The removal estimates were higher with plot-level data. The results suggest that, in heterogeneous young stands, plot by plot prediction would give more accurate removal estimates than the calculation of a corresponding prediction at the stand-level.
  • Tahvanainen, Finnish Forest Research Institute, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: timo.tahvanainen@metla.fi (email)
  • Kaartinen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuun yliopisto, Finland E-mail: kk@nn.fi
  • Pukkala, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuun yliopisto, Finland E-mail: tp@nn.fi
  • Maltamo, University of Joensuu, Faculty of Forestry, P.O. Box 111, FI-80101 Joensuun yliopisto, Finland E-mail: mm@nn.fi
article id 325, category Research note
Daniel Eriksson, Henrik Lindberg, Urban Bergsten. (2006). Influence of silvicultural regime on wood structure characteristics and mechanical properties of clear wood in Pinus sylvestris. Silva Fennica vol. 40 no. 4 article id 325. https://doi.org/10.14214/sf.325
Keywords: silvicultural regime; structural characteristics; mechanical properties
Abstract | View details | Full text in PDF | Author Info
The objective of the study presented here was to evaluate the influence of two contrasting silvicultural regimes on the structural characteristics and mechanical properties of different wood tissue types at different heights in Scots pine (Pinus sylvestris L.) trees, and reasons for these differences. Wood samples were taken from two stands (a dense 85-year-old stand established by direct seeding and a 56-year-old widely spaced stand established by planting, designated SDR and PWR, respectively in the boreal zone of Sweden). The wood properties associated with the examined silvicultural regimes differed, in terms of both structural characteristics (with up to fivefold differences between SDR and PWR) and mechanical properties (with up to almost threefold differences between SDR and PWR). Differences between the regimes were highest for stiffness, followed by strength and hardness properties and lowest for relative stiffness after 1000 h of loading (creep) (with higher parameter values for SDR than for PWR in each case). The rankings could be explained by differences among the mechanical properties in their sensitivity to maturation of wood characteristics. In conclusion, silvicultural regimes have great potential to regulate wood structural characteristics and mechanical properties, apparently due to the influences of the green crown and growth rate on the vascular cambium, the strength of which vary throughout the rotation period. A silvicultural regime could therefore be seen as a tool that can be used to select material qualities and to make wood a more homogenous material for engineers.
  • Eriksson, Swedish University of Agricultural Sciences, Vindeln Experimental Forest, Svartberget Fieldstation, SE-922 91 Vindeln, Sweden E-mail: daniel.eriksson@esf.slu.se (email)
  • Lindberg, Luleå University of Technology, Division of Polymer Engineering, SE-971 87 Luleå, Sweden E-mail: hl@nn.se
  • Bergsten, Swedish University of Agricultural Sciences, Department of Silviculture, SE-901 83 Umeå, Sweden E-mail: ub@nn.se
article id 324, category Research note
Matti Katila. (2006). Empirical errors of small area estimates from the multisource National Forest Inventory in Eastern Finland. Silva Fennica vol. 40 no. 4 article id 324. https://doi.org/10.14214/sf.324
Keywords: k-nearest neighbours estimation; Landsat ETM ; multisource forest inventory; synthetic estimation
Abstract | View details | Full text in PDF | Author Info
The precision of multisource national forest inventory (MS-NFI) estimators and simple synthetic estimators based on NFI field data only was assessed employing an independent inventory data set of several small areas in Eastern Finland. There were seven test units of size 100 km2 and three test units of size 1 km2 for which a systematic field sampling was carried out. The ‘improved’ MS-NFI method yielded the most precise estimates for mean volume and mean volume of pine and spruce: relative root mean square errors (RMSE*) were 5%, 12% and 15% for 100 km2 test units and 13%, 27% and 40% for 1 km2 test units respectively. The stratified MS-NFI method was best for broad-leaved volume estimation. Synthetic estimation based on the NFI9 field plots post-stratified with coarse scale forest variable maps from NFI8 resulted in RMSE*s comparable to those of the ordinary MS-NFI in areas of 100 km2 for mean volume and mean volume of pine and spruce. The amount of variation between the field inventory estimates for the test units explained by the MS-NFI estimators remained the same or increased when the size of the area increased from of 1 km2 to 100 km2 and up to 2000 km2. The validation of the largest areas was made against the NFI9 field inventory estimates for groups of municipalities in the study area.
  • Katila, Finnish Forest Research Institute, Unioninkatu 40 A, FI-00170 Helsinki, Finland E-mail: mk@nn.fi (email)
article id 379, category Research note
Daniel Ligné, Lars Eliasson, Tomas Nordfjell. (2005). Time consumption and damage to the remaining stock in mechanised and motor manual pre-commercial thinning. Silva Fennica vol. 39 no. 3 article id 379. https://doi.org/10.14214/sf.379
Keywords: birch; Scots pine; cleaning; time study; brush saw; Husqvarna; Vimek
Abstract | View details | Full text in PDF | Author Info
Selective pre-commercial thinning (PCT) is usually carried out by workers with a brush saw in order to increase the growth of the potential crop trees (main stems) through removal of competing trees. In the last decade relative PCT costs have increased, partly because stands are denser and have higher trees when treated, which has led to new interest for mechanised PCT. The objective was to compare mechanised and motor manual PCT regarding productivity and damage to remaining main stems. Time consumption for, and damage after, mechanised and motor manual PCT were studied on 50 plots per treatment in mixed pine birch stands with an initial stand density exceeding 4500 stems ha–1. In the present study productivity was influenced by stand density, stand height and the quota between height and diameter. Irrespectively of these factors, mechanised PCT was 0.74 hours ha–1 slower than motor manual PCT. Motor manual PCT of the average stand (average height 3.69 m, 10 816 stems ha–1) took 5.06 effective hours. In average 2475 and 2805 main stems ha–1 were left after the mechanised and motor manual treatments, respectively, whereof 1.3 and 2.1% were damaged by the treatments. The results show that efficiency in motor manual PCT has increased in dense and tall stands compared to older studies. Motor manual PCT was more time effective than mechanised PCT, and thereby also even more cost-effective. However, the potential for technical and methodological development of mechanised PCT is probably larger than for motor manual PCT.
  • Ligné, Swedish University of Agricultural Sciences, Department of Silviculture, SE-901 83 Umeå, Sweden E-mail: dl@nn.se
  • Eliasson, Swedish University of Agricultural Sciences, Department of Silviculture, SE-901 83 Umeå, Sweden E-mail: lars.eliasson@norraskogsagarna.se (email)
  • Nordfjell, Swedish University of Agricultural Sciences, Department of Silviculture, SE-901 83 Umeå, Sweden E-mail: tn@nn.se
article id 403, category Research note
Timo Saksa, Juha Heiskanen, Jari Miina, Jaakko Tuomola, Taneli Kolström. (2005). Multilevel modelling of height growth in young Norway spruce plantations in southern Finland. Silva Fennica vol. 39 no. 1 article id 403. https://doi.org/10.14214/sf.403
Keywords: Norway spruce; Picea abies; height growth; mounding; disc trenching; container seedlings; intra-level correlation; variance-component model
Abstract | View details | Full text in PDF | Author Info
Height development of Norway spruce (Picea abies (L.) Karst.) transplants was studied on 22 sites prepared by disc trenching or mounding. At the age of 4–9 years the plantations were surveyed using a multistage sampling design. For every planted spruce on a plot, the past annual height increments were measured as far into the past as possible. Multilevel mixed linear modelling was used to analyse the variation in growth at different levels (year, stand, cluster, plot, tree) and the effects of climatic and site characteristics on height growth. The within-plantation variation in height growth was higher on mounded sites than on disc-trenched sites. The mean temperature and the precipitation sum of the summer months affected height growth positively. Soil characteristics measured from undisturbed soil did not explain the height growth of seedlings on mounded sites, whereas on disc-trenched sites, the depth of the organic layer and the soil temperature had a positive effect and the depth of the eluvial horizon a negative effect. The modelling approach used proved to be a useful method for examining the sources of variation in development of young plantations.
  • Saksa, The Finnish Forest Institute, Suonenjoki Research Station, FI-77600 Suonenjoki, Finland E-mail: ts@nn.fi (email)
  • Heiskanen, The Finnish Forest Institute, Suonenjoki Research Station, FI-77600 Suonenjoki, Finland E-mail: jh@nn.fi
  • Miina, The Finnish Forest Institute, Joensuu Research Centre, P. O. Box 68, FI-80101 Joensuu, Finland E-mail: jm@nn.fi
  • Tuomola, The University of Joensuu, Mekrijärvi Research Station, FI-82900 Ilomantsi, Finland E-mail: jt@nn.fi
  • Kolström, The University of Joensuu, Mekrijärvi Research Station, FI-82900 Ilomantsi, Finland E-mail: tk@nn.fi
article id 402, category Research note
Miina Rautiainen, Pauline Stenberg, Tiit Nilson. (2005). Estimating canopy cover in Scots pine stands. Silva Fennica vol. 39 no. 1 article id 402. https://doi.org/10.14214/sf.402
Keywords: Pinus sylvestris; Cajanus tube; LAI-2000 Plant Canopy Analyzer
Abstract | View details | Full text in PDF | Author Info
The way canopy cover is defined and measured influences the obtained canopy cover percentage. Estimates of canopy cover are needed, for example, in canopy radiation modelling and remote sensing applications and as a tool for political decision-making. In this paper, we demonstrated the use of two methods, the LAI-2000 Plant Canopy Analyzer instrument and the Cajanus tube, in Scots pine stands for canopy cover estimation, and also assessed the number of measurement points required for reliable estimates. The Cajanus tube yielded slightly larger canopy closure values than the LAI-2000 instrument, but the values were nevertheless in good agreement. Both of the methods required approximately 250 measurement points for canopy closure estimates of a stand to become relatively stable. We also present the first measured effective canopy transmittance values for Scots pine stands in Finland and an example of tree pattern mapping with the Cajanus tube.
  • Rautiainen, Department of Forest Ecology, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: miina.rautiainen@helsinki.fi (email)
  • Stenberg, Department of Forest Ecology, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: ps@nn.fi
  • Nilson, Tartu Observatory, EE-61602 Tõravere, Tartumaa, Estonia E-mail: tn@nn.ee
article id 401, category Research note
Kari Taulavuori, Marian Sarala, Jouni Karhu, Erja Taulavuori, Eero Kubin, Kari Laine, Jarmo Poikolainen, Erkki Pesonen. (2005). Elongation of Scots pine seedlings under blue light depletion. Silva Fennica vol. 39 no. 1 article id 401. https://doi.org/10.14214/sf.401
Keywords: Scots pine; blue light; elongation; high latitude
Abstract | View details | Full text in PDF | Author Info
The effect of blue light (400–500 nm) removal on the elongation of Scots pine (Pinus sylvetris L.) seedlings was investigated in a field experiment in Northern Finland during two growing seasons. The seedlings were grown in plexiglass chambers and ambient control plots. The blue wavelengths were removed from the light spectrum by using chambers made of orange plexiglass. The results of nearly daily measurements showed that the elongation of Scots pine seedlings was increased by the removal of blue light only at high latitudes.
  • Taulavuori, Department of Biology, University of Oulu, PO Box 3000, FI-90014, Oulu, Finland E-mail: ktaulavu@cc.oulu.fi (email)
  • Sarala, Department of Biology, University of Oulu, PO Box 3000, FI-90014, Oulu, Finland E-mail: ms@nn.fi
  • Karhu, Muhos Research Station, Finnish Forest Research Institute, Kirkkosaarentie 7, FI-91500 Muhos, Finland E-mail: jk@nn.fi
  • Taulavuori, Department of Biology, University of Oulu, PO Box 3000, FI-90014, Oulu, Finland E-mail: et@nn.fi
  • Kubin, Muhos Research Station, Finnish Forest Research Institute, Kirkkosaarentie 7, FI-91500 Muhos, Finland E-mail: ek@nn.fi
  • Laine, Department of Biology, University of Oulu, PO Box 3000, FI-90014, Oulu, Finland E-mail: kl@nn.fi
  • Poikolainen, Muhos Research Station, Finnish Forest Research Institute, Kirkkosaarentie 7, FI-91500 Muhos, Finland E-mail: jp@nn.fi
  • Pesonen, Vantaa Research Centre, Finnish Forest Research Institute, PO Box 18, FI-01301, Vantaa, Finland E-mail: ep@nn.fi
article id 421, category Research note
Pekka Helenius, Jaana Luoranen, Risto Rikala. (2004). Effect of thawing duration and temperature on field performance of frozen-stored Norway spruce container seedlings. Silva Fennica vol. 38 no. 3 article id 421. https://doi.org/10.14214/sf.421
Keywords: Picea abies; height growth; survival; frozen-planting; storage
Abstract | View details | Full text in PDF | Author Info
Increasing use of frozen storage in nurseries at northern latitudes calls for thawing methods that are safe, economical and easy to apply on a large scale. The easiest and most economical method would be to thaw seedlings in the same boxes they were stored in. However, doing this safely requires more knowledge about how long and at what temperatures seedlings should or can be kept in the boxes without reducing field performance. In this study, 1-yr-old frozen-stored Norway spruce (Picea abies (L.) Karst.) container seedlings were thawed for 4, 8 or 16 days at 4 or 12 °C in cardboard boxes before planting on a reforestation site and on experimental field in mid-June. Some seedlings were also planted on these locations after thawing for only 7 hours at 12 °C in order to separate frozen root plugs. We found some evidence that planting seedlings after short thawing periods (7 hours at 12 °C and 4 days at 4 °C), under which conditions the root plugs remain completely or partly frozen, has a negative effect on field performance of Norway spruce seedlings. Thawing over a 4-8 day period in cardboard boxes at ca. 12 °C appears to ensure complete thawing of the root plugs and unaffected field performance, but is short enough to prevent the growth of mould.
  • Helenius, Finnish Forest Research Institute, Suonenjoki Research Station, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: pekka.helenius@metla.fi (email)
  • Luoranen, Finnish Forest Research Institute, Suonenjoki Research Station, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: jl@nn.fi
  • Rikala, Finnish Forest Research Institute, Suonenjoki Research Station, Juntintie 154, FI-77600 Suonenjoki, Finland E-mail: rr@nn.fi
article id 517, category Research note
Matti Huotari, Mareena Jaskari, Erkki Annila, Vilho Lantto. (2003). Responses of olfactory receptor neurons of the large pine weevil to a possible deterrent Neutroil® and two other chemicals. Silva Fennica vol. 37 no. 1 article id 517. https://doi.org/10.14214/sf.517
Keywords: olfactory receptor neuron; attractant; repellent; Hylobius abietis; action potential; silent period; electroantennogram
Abstract | View details | Full text in PDF | Author Info
Electrophysiological responses of olfactory receptor neurons were measured from the antennal sensilla of large pine weevils (Hylobius abietis L.) at 1 s exposure to Neutroil®-hexane mixture odour as a possible deterrent chemical and, for comparison, to α-pinene, α-pinene-ethanol mixture, and ethanol odours, respectively. Neutroil® is a commercial chemical pulp-mill product which has been studied earlier as a deterrent for large pine weevils with preliminary feed tests. In addition, ethanol, hexane and clean carrier air responses were measured. Odour pulses and clean air, as a zero reference value, were directed at a fixed insect antenna in order to induce olfactory responses. Simultaneous olfactory responses, i.e. Hylobius electroantennograms (EAG) and action potential responses, were recorded and these responses of Hylobius olfactory receptor neurons (ORN), such as action potential rates, silent periods and EAG responses, were analyzed for all simultaneous recordings. The exposures to α-pinene, α-pinene-ethanol mixture, pure ethanol and hexane caused an increase of the action potential rate (up to 70 pulses per second) in the ORNs sensitive to these odours, while the Neutroil®-hexane mixture exposures caused either silent periods with a duration between 0.6 and 1.1 s for 1 s exposure pulses or they had no response at all in the ORNs sensitive to the other odours. Thus, the possible deterrence may be caused by inhibition of some pinene-alcohol ORNs.
  • Huotari, University of Oulu, P.O. Box 4500, FIN-90014 University of Oulu, Finland E-mail: matti.huotari@oulu.fi (email)
  • Jaskari, University of Oulu, P.O. Box 4500, FIN-90014 University of Oulu, Finland E-mail: mj@nn.fi
  • Annila, Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: ea@nn.fi
  • Lantto, University of Oulu, P.O. Box 4500, FIN-90014 University of Oulu, Finland E-mail: vl@nn.fi
article id 534, category Research note
Ilkka Leinonen, Heikki Hänninen. (2002). Adaptation of the timing of bud burst of Norway spruce to temperate and boreal climates. Silva Fennica vol. 36 no. 3 article id 534. https://doi.org/10.14214/sf.534
Keywords: adaptation; modelling; frost damage; bud burst; phenology; stabilizing selection
Abstract | View details | Full text in PDF | Author Info
The adaptation of the annual cycle of development of boreal and temperate trees to climatic conditions has been seen as a result of stabilizing selection caused by two opposite driving forces of natural selection, i.e. the tolerance of unfavorable conditions during the frost exposed season (survival adaptation) and the effective use of growth resources during the growing season (capacity adaptation). In this study, two theories of the effects of climate on the adaptation of the timing of bud burst of trees were evaluated. This was done with computer simulations by applying a temperature sum model for predicting the timing of bud burst of different Norway spruce genotypes on the basis of air temperature data from various climatic conditions. High geographical variation in the temperature response of bud burst, typical for Norway spruce, was included in the theoretical analyses. The average timing of bud burst and the corresponding risk of occurrence of damaging frost during the susceptible period after bud burst were calculated for each genotype in each climate. Two contrasting theories of the stabilizing selection were evaluated, i.e. the overall adaptedness of each genotype was evaluated either 1) by assuming a fixed threshold for the risk of frost damage, or 2) by assuming a tradeoff between the risk of frost damage and the length of the growing season. The tradeoff assumption produced predictions of between provenance variation in bud burst which correspond more closely with empirical observations available in literature, compared to the fixed threshold assumption.
  • Leinonen, University of Oklahoma, Department of Botany and Microbiology, Norman, OK 73019, USA E-mail: leinonen@ou.edu (email)
  • Hänninen, University of Helsinki, Department of Ecology and Systematics, FIN-00014 Helsinki, Finland E-mail: hh@nn.fi
article id 569, category Research note
Heikki Kauhanen. (2002). Occurrence of fires in the eastern Saariselkä area, north-west Russia. Silva Fennica vol. 36 no. 1 article id 569. https://doi.org/10.14214/sf.569
Keywords: fire; anthropogenic fire; burn; Russia; satellite image; vegetation mosaic
Abstract | View details | Full text in PDF | Author Info
The occurrence of fires was studied in the eastern Saariselkä area, North-West Russia, by using satellite images and topographical maps. In total, more than 330 burned areas were pinpointed in the study area of 4770 km2. Old burns were concentrated in the eastern part of the study area, but young burns were more common in the west. Sites affected by fires in the more recent past were much smaller than those burnt over earlier. The abundance of burns along rivers and the border surveillance road provided evidence of human impact. The most significant changes in the landscape were found in the eastern part of the study area, where spruce forests had been replaced by birch woodlands.
  • Kauhanen, Finnish Forest Research Institute, Kolari Research Station, Muoniontie 21 A, FIN-95900 Kolari, Finland E-mail: heikki.kauhanen@metla.fi (email)
article id 609, category Research note
Aleksei Fedorkov. (2001). Climatic adaptation of seed maturity in Scots pine and Norway spruce populations. Silva Fennica vol. 35 no. 1 article id 609. https://doi.org/10.14214/sf.609
Keywords: Norway spruce; Scots pine; climatic adaptation; provenance experiment; seed maturation; X-ray method
Abstract | View details | Full text in PDF | Author Info
Seed maturation of Scots pine and Norway spruce in a provenance experiment at Kortkeros (northern Russia) was examined by the X-ray method. Logarithmic relationships were found between seed anatomy development and long-term average thermal sum. Seed development in the northern populations of Scots pine and Norway spruce was a little faster than in the southern ones.
  • Fedorkov, Russian Academy of Sciences, Komi Science Centre, Institute of Biology, Kommunisticheskya St., 28, 167610 Syktyvkar, Russia E-mail: fedorkov@ib.komisc.ru (email)
article id 608, category Research note
Jouni Partanen, Ilkka Leinonen, Tapani Repo. (2001). Effect of accumulated duration of the light period on bud burst in Norway spruce (Picea abies) of varying ages. Silva Fennica vol. 35 no. 1 article id 608. https://doi.org/10.14214/sf.608
Keywords: Norway spruce; phenology; growth initiation; photoperiod
Abstract | View details | Full text in PDF | Author Info
One-year-old seedlings (two sowing times), two-year-old seedlings and 14- and 18-year-old cuttings of Norway spruce (Picea abies (L.) Karst.) were exposed to shortening photoperiod (initially 16 h), lengthening photoperiod (initially 6 h) and constant short photoperiod (6 h) treatments with uniform temperature conditions in growth chambers. The timing of bud burst was examined. In all plants, shortening photoperiod treatment seemed to promote bud burst compared with other treatments. This effect was clearest in the oldest material. The results suggest that, in addition to temperature sum, the accumulated duration of the light period may promote bud burst of Norway spruce.
  • Partanen, Finnish Forest Research Institute, Punkaharju Research Station, Finlandiantie 18, FIN-58450 Punkaharju, Finland E-mail: jouni.partanen@metla.fi (email)
  • Leinonen, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: il@nn.fi
  • Repo, University of Joensuu, Faculty of Forestry, P.O. Box 111, FIN-80101 Joensuu, Finland E-mail: tr@nn.fi
article id 624, category Research note
Hashim Ali El Atta. (2000). Effect of diet and seed pretreatment on the biology of Bruchidius uberatus (Coleoptera, Bruchidae). Silva Fennica vol. 34 no. 4 article id 624. https://doi.org/10.14214/sf.624
Keywords: Bruchidius uberatus; Bruchidae; Acacia nilotica; seed borer biology
Abstract | View details | Full text in PDF | Author Info
Diet significantly (P < 0.0001) affected fecundity of Bruchidius uberatus Fahraeus. Provision of 1% sugar solution increased fecundity from 15 eggs/female to 47. Furthermore, sugar solution prolonged significantly the oviposition period of B. uberatus from one week to two weeks. Diet also significantly (P < 0.0001) increased adult longevity. Mean adult longevity recorded was 3, 7 and 13 days in control, water and sugar treatments, respectively. Seed pretreatment had a highly significant impact on the various developmental stages of B. uberatus. Maximum egg hatchability occurred in non-husked Acacia nilotica (L.) Willd. ex Del. seeds (83%), moderate in de-husked seeds (74%) and least in seeds presoaked in concentrated sulphuric acid (42%). The frequency of larvae that developed successfully into pupae was greatest in non-husked seeds (72%), nevertheless in de-husked and acid pretreated seeds, absolutely no larvae developed into pupae and hence the adult stage was not reached in these two treatments. Thus, de-husking and acid pretreatment of A. nilotica seeds is highly recommended.
  • El Atta, University of Khartoum, Faculty of Forestry, Department of Forest Conservation and Protection, Shambat, Sudan E-mail: hashimelatta@yahoo.com (email)
article id 631, category Research note
Pekka Ripatti. (2000). Use of log-linear models in forecasting structural changes in Finnish non-industrial private forest ownership. Silva Fennica vol. 34 no. 3 article id 631. https://doi.org/10.14214/sf.631
Keywords: forest policy; forecasting; non-industrial private forestry; log-linear models; ownership changes
Abstract | View details | Full text in PDF | Author Info
This paper presents how log-linear models can be used for modelling and forecasting structural changes of Finnish non-industrial private forest ownership. Two cross-sectional sets of data, which were collected in conjunction with two separate surveys by means of mail questionnaires in 1975 and 1990, were employed. A total of six non-industrial private forest holding and ownership attributes are forecast focusing on the earlier pace of structural change. The results show that the pace of change in the forecast attributes appears to be less than it would be when derived from extrapolation of the earlier trends. The results of the study can be applied to forest policy and forestry extension planning, by providing a more realistic outlook of the future structure of non-industrial private forest ownership.
  • Ripatti, Finnish Forest Research Institute, Helsinki Research Centre, Unioninkatu 40 A, FIN-00170 Helsinki, Finland E-mail: pekka.ripatti@metla.fi (email)
article id 646, category Research note
K. S. Krishna Chaitanya, S. Keshavkant, S. C. Naithani. (2000). Changes in total protein and protease activity in dehydrating recalcitrant sal (Shorea robusta) seeds. Silva Fennica vol. 34 no. 1 article id 646. https://doi.org/10.14214/sf.646
Keywords: recalcitrant; viability; total protein; protease; Shorea robusta
Abstract | View details | Full text in PDF | Author Info
A rapid loss of viability was recorded in sal seeds when dehydrated below 36.7% moisture content, at ambient conditions. Seed becomes non-viable at 6 days after harvest (dah). Gradual decline in total protein content due to corresponding increase in protease activity preceded loss of viability. Almost 43.7% and 52.6% loss in total protein content was observed on 3 dah in embryonic axis and cotyledon respectively of seeds showing 100% viability. No protease activity was detected in the embryonic axis and cotyledon of freshly harvested viable seeds (0 dah). The protease activity was detected after 12 hrs of storage and increased sharply with peak levels on 6 dah (0.71 ± 0.04 / min/mg protein) and 4 dah (0.16 ± 0.01 / min/mg protein) in embryonic axis and cotyledon respectively. Later on the enzyme activity declined sharply in both the tissues. Enhanced protease activity in embryonic axes and cotyledons has been discussed with corresponding decline in total protein during desiccation induced loss of viability in sal seeds.
  • Chaitanya, Pt. Ravishankar Shukla University, Seed Biology Lab, School of Life Sciences, Raipur – 492 010 (M.P.), India E-mail: kskc@nn.in
  • Keshavkant, Pt. Ravishankar Shukla University, Seed Biology Lab, School of Life Sciences, Raipur – 492 010 (M.P.), India E-mail: sk@n.in
  • Naithani, Pt. Ravishankar Shukla University, Seed Biology Lab, School of Life Sciences, Raipur – 492 010 (M.P.), India E-mail: naithani@mantraonline.com (email)
article id 653, category Research note
Desta Fekedulegn, Mairitin P. Mac Siurtain, Jim J. Colbert. (1999). Parameter estimation of nonlinear growth models in forestry. Silva Fennica vol. 33 no. 4 article id 653. https://doi.org/10.14214/sf.653
Keywords: parameter estimation; nonlinear model; partial derivative; Marquardt iterative method; initial value
Abstract | View details | Full text in PDF | Author Info
Partial derivatives of the negative exponential, monomolecular, Mitcherlich, Gompertz, logistic, Chapman-Richards, von Bertalanffy, Weibull and the Richard’s nonlinear growth models are presented. The application of these partial derivatives in estimating the model parameters is illustrated. The parameters are estimated using the Marquardt iterative method of nonlinear regression relating top height to age of Norway spruce (Picea abies L.) from the Bowmont Norway Spruce Thinning Experiment. Formulas that provide good initial values of the parameters are specified. Clear definitions of the parameters of the nonlinear models in the context of the system being modelled are found to be critically important in the process of parameter estimation.
  • Fekedulegn, Department of Statistics, West Virginia University, Eberly College of Arts and Sciences, P.O. Box 6330, Morgantown, WV26506, USA E-mail: fdesta@stat.wvu.edu (email)
  • Mac Siurtain, University College Dublin, Ireland E-mail: mpms@nn.ie
  • Colbert, USDA Forest Service, Northeastern Research Station, Morgantown, West Virginia E-mail: jjc@nn.us
article id 659, category Research note
Anneli Viherä-Aarnio, Pirkko Velling. (1999). Growth and stem quality of mature birches in a combined species and progeny trial. Silva Fennica vol. 33 no. 3 article id 659. https://doi.org/10.14214/sf.659
Keywords: stem quality; growth; Betula pendula Roth; Betula pubescens Ehrh.; Betula papyrifera Marsh.; progeny; timber production
Abstract | View details | Full text in PDF | Author Info
The growth and stem quality of silver birch (Betula pendula), downy birch (B. pubescens) and paper birch (B. papyrifera) were compared in a 32-year-old field trial in southern Finland. The material consisted of different unselected stand origins and progenies of phenotypically selected plus trees of silver and downy birch from southern Finland and differing stand origins of paper birch from the North-West Territories, Canada. Growth, yield and a number of stem quality traits, including taper, sweep, stem defects, heights of different crown limits and length of the veneer timber part of the stem were measured or observed. The native Finnish silver and downy birches were superior to paper birch in terms of both yield and stem quality, silver birch being the best. Progenies of silver birch plus trees were better than the stand origin, indicating that the former are able to reach high quality veneer log size in a shorter time than unselected material. The cultivation of paper birch can not be considered viable in Finland.
  • Viherä-Aarnio, The Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: anneli.vihera-aarnio@metla.fi (email)
  • Velling, The Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: pv@nn.fi
article id 679, category Research note
Hans-Örjan Nohrstedt, Gunnar Börjesson. (1998). Respiration in a forest soil 27 years after fertilization with different doses of urea. Silva Fennica vol. 32 no. 4 article id 679. https://doi.org/10.14214/sf.679
Keywords: mineral soil; Pinus sylvestris L.; carbon mineralization; humus; podzol
Abstract | View details | Full text in PDF | Author Info
A number of previous studies have shown that N fertilization often reduces respiration in forest soils. However, the durability of this effect has not been fully explored. In this study, the response of soil respiration to a single fertilization with urea, applied 27 years earlier, was examined in a field experiment located in a stand of Pinus sylvestris in central Sweden. The doses that had been added were 120, 240 and 600 kg N ha–1. Samples were taken from the humus layer and the upper 7.5 cm of the mineral soil. Sieved samples were incubated in the laboratory. No effect of the previous fertilization on soil respiration was found, thus indicating that the reduction shown in earlier studies is not persistent. There was a tendency that the highest N dose had caused a higher N concentration and a lower C/N-ratio in the humus layer and a higher C concentration in the mineral soil.
  • Nohrstedt, The Forestry Research Institute of Sweden, Uppsala Science Park, S-751 83 Uppsala, Sweden E-mail: hans-orjan.nohrstedt@skogforsk.se (email)
  • Börjesson, Department of Microbiology, Swedish University of Agricultural Sciences, P.O. Box 7025, S-750 07 Uppsala, Sweden E-mail: gb@nn.se
article id 678, category Research note
Ian A. Nalder, Ross W. Wein. (1998). A new forest floor corer for rapid sampling, minimal disturbance and adequate precision. Silva Fennica vol. 32 no. 4 article id 678. https://doi.org/10.14214/sf.678
Keywords: boreal; moss; bulk density; variability; forest floor; corer; western Canada
Abstract | View details | Full text in PDF | Author Info
We describe an effective and inexpensive device for sampling forest floors. It is based on a rechargeable, battery-powered drill that drives a sharpened steel coring tube. The corer is simple to fabricate, is lightweight (3.5 kg) and can be used easily by one person to obtain intact, natural volume cores of the forest floor. It has been used extensively to obtain samples in 114 boreal forest stands of western Canada. We found that coefficients of variation were typically 30% for forest floor organic matter and bulk density, and tended to be higher in Pinus banksiana stands than in Picea glauca and Populus tremuloides. Ten samples per stand gave adequate precision for a study of forest floor dynamics and autocorrelation did not appear to be a problem with five-metre sampling intervals. In addition to sampling forest floors, the corer has proven suitable for sampling moss and lichen layers and mineral soil down to about 20 cm. A similar powered system can also be used for increment boring of trees.
  • Nalder, University of Alberta, Department of Renewable Resources, 442 Earth Sciences Building, Edmonton, Alberta, Canada T6G 2E3 E-mail: inalder@gpu.srv.ualberta.ca (email)
  • Wein, University of Alberta, Department of Renewable Resources, 442 Earth Sciences Building, Edmonton, Alberta, Canada T6G 2E3 E-mail: rww@nn.ca
article id 702, category Research note
Timo Kurkela, Heikki Nuorteva. (1998). Short-needle disease of Scots pine: an abnormal needle length distribution. Silva Fennica vol. 32 no. 1 article id 702. https://doi.org/10.14214/sf.702
Keywords: Pinus sylvestris; Scots pine; needle length; insect feeding; tissue deformation
Abstract | View details | Full text in PDF | Author Info
Short-needle syndrome occurs commonly in southern Finland. The disease is characterized by abnormal length distribution of the needles in shoots. In most cases, affected shoots have needles of normal length as well as very short needles. The short needles are those injured during the needle elongation period; the tissues formed abnormal sclerenchymatic structures and wound periderm. One possible cause could be hemipterous insects feeding on growing needles. Salivary sheaths of such insects were often present in both deformed needle bases and undeformed mature tissues.
  • Kurkela, Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: timo.kurkela@metla.fi (email)
  • Nuorteva, Finnish Forest Research Institute, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: hn@nn.fi

Category : Data note

article id 24066, category Data note
Tuomas Yrttimaa, Lauri Liikonen, Aapo Erkkilä, Mikko Vastaranta. (2025). Terrestrial laser scanning point clouds and tree attributes from 55 sample plots at the Evo test site (spring 2024). Silva Fennica vol. 59 no. 1 article id 24066. https://doi.org/10.14214/sf.24066
Keywords: boreal forest; ground-based LiDAR; LIS TreeAnalyzer; point cloud processing; Riegl VZ-400i; tree characterization; tree reconstruction
Highlights: Terrestrial laser scanning dataset from 55 sample plots (32 × 32 m) representing boreal forests of Southern Finland; Data was acquired in April–May 2024 using a Riegl VZ-400i scanner, providing multiple returns per pulse; Each point is annotated with reflectance, return properties, and tree linkage; Crown and stem diameters were derived for further analysis, advancing tree and forest research applications.
Abstract | Full text in HTML | Full text in PDF | Author Info
Terrestrial laser scanning (TLS) is an active remote sensing technique that digitizes trees and forest stands by capturing range measurements, resulting in detailed point clouds. To support the development of computational methods for tree and forest stand characterization, as well as to facilitate the exploration of tree structures, we collected TLS data from 55 sample plots (32 m × 32 m) and 6320 trees in Evo, Southern Finland. Data acquisition was conducted during April–May 2024 using a Riegl VZ-400i terrestrial laser scanner (Riegl Laser Measurement Systems GmbH, Austria), capable of recording up to eight returns per laser pulse. This dataset includes TLS point clouds in the projected coordinate reference system commonly used in Finland (ETRS-TM35FIN). Each point is annotated with reflectance, return number, return count, and height above ground. Additionally, information linking each return to its originating tree is provided. For individual trees, the point clouds were further processed to derive key attributes such as crown projection area, crown diameter, and stem diameter. In addition, tree species was derived by linking the TLS-based tree measurements with field inventory data. Here, we describe and share these curated TLS data files and related tree measurements, which offer a valuable resource for advancing tree- and forest-related research and applications.
  • Yrttimaa, School of Forest Sciences, University of Eastern Finland, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0003-2648-523X E-mail: tuomas.yrttimaa@uef.fi (email)
  • Liikonen, School of Forest Sciences, University of Eastern Finland, FI-80101 Joensuu, Finland E-mail: lauriliik@uef.fi
  • Erkkilä, School of Forest Sciences, University of Eastern Finland, FI-80101 Joensuu, Finland E-mail: aapo.erkkila@uef.fi
  • Vastaranta, School of Forest Sciences, University of Eastern Finland, FI-80101 Joensuu, Finland ORCID https://orcid.org/0000-0001-6552-9122 E-mail: mikko.vastaranta@uef.fi

Category : Complex remote sensing-assisted forest surveys – Discussion article

article id 24063, category Complex remote sensing-assisted forest surveys – Discussion article
Sara Franceschi, Caterina Pisani, Lorenzo Fattorini, Piermaria Corona. (2025). Statistical considerations for enhanced forest resource mapping. Silva Fennica vol. 59 no. 2 article id 24063. https://doi.org/10.14214/sf.24063
Keywords: design-based inference; consistency; kNN mapping; pseudo-population bootstrap; Random Forest mapping
Abstract | Full text in HTML | Full text in PDF | Author Info

This paper examines forest resource mapping from a statistical perspective, highlighting the opportunity to use a design-based approach to ensure inferential congruency with the estimation of averages and totals of forest attributes. Traditionally, in forest surveys estimates of averages and totals are obtained using design-unbiased estimators, with known variance expressions that can be easily estimated using standard sampling methodologies. The paper emphasizes the prominent role of kNN and Random Forest techniques in forest mapping while addressing the methodological limitations identified over more than thirty years of forest literature in efforts to estimate map precision. The critical importance of design-based map consistency, often overlooked in forest literature, is discussed and clarified, demonstrating that it allows for the development of design-based estimators of map precision through bootstrap resampling from the estimated maps.

  • Franceschi, Department of Economics and Statistics, University of Siena, Via Banchi di Sotto 55, 53100 Siena, Italy ORCID https://orcid.org/0000-0001-6675-4540 E-mail: franceschi2@unisi.it (email)
  • Pisani, Department of Economics and Statistics, University of Siena, Via Banchi di Sotto 55, 53100 Siena, Italy E-mail: caterin.pisani@unisi.it
  • Fattorini, Department of Economics and Statistics, University of Siena, Via Banchi di Sotto 55, 53100 Siena, Italy E-mail: lorenzo.fattorini@unisi.it
  • Corona, CREA, Research Centre for Forestry and Wood, Viale Santa Margherita 80, 52100 Arezzo, Italy E-mail: piermaria.corona@crea.gov.it

Category : Discussion article

article id 25031, category Discussion article
Simon Lebel Desrosiers, Nicolas Bélanger, Evelyne Thiffault, Nelson Thiffault. (2025). Climate change and transformation in forest fire regimes: an opportunity for the implementation of assisted migration of tree species in the Canadian boreal forest? Silva Fennica vol. 59 no. 2 article id 25031. https://doi.org/10.14214/sf.25031
Keywords: fire severity; site suitability; post-fire regeneration; climate change adaptation; forest resilience; reforestation strategies; tree migration
Highlights: Increasing fire activity is reshaping post-disturbance landscapes in boreal forests; Post-fire sites offer new opportunities for introducing climate-resilient tree species; Fire can improve or impair site conditions for forest regeneration; Assisted migration of tree species may enhance reforestation success after severe wildfires; Research on post-fire regeneration of introduced species remains critically limited.
Abstract | Full text in HTML | Full text in PDF | Author Info

Climate change is intensifying fire regimes in boreal forests, leading to ecological disruption and raising concerns about forest resilience and post-disturbance recovery. Altered fire dynamics creates novel opportunities for implementing adaptive silviculture for climate change, including assisted migration, the intentional movement and establishment of tree species or tree populations outside their current range of distribution to better match anticipated future climates. Here, we examine how the increasing frequency, severity, and spatial extent of Canadian boreal wildfires can serve as strategic windows for introducing climate-resilient tree species and genotypes. We review how fire influences the availability and suitability of post-fire sites for assisted migration, highlighting how fire-induced changes in soil abiotic and biotic properties may facilitate or hinder the establishment of relocated tree species. While fire can simplify site preparation, reduce biotic competition, and temporarily enhance soil nutrient availability, it may also degrade soil structure by consuming or altering soil organic matter and increasing soil susceptibility to erosion and disrupt essential mycorrhizal associations. We argue that assisted migration of tree species can be a proactive silvicultural tool when used in areas with regeneration failure or where future climate conditions are likely to exceed the tolerance limits of native species. Whilst scientific evidence remains limited on the regeneration success of migrated species and genotypes in post-fire environments, we argue for an integrated adaptation strategy that combines natural regeneration with targeted assisted migration interventions, guided by local site conditions, genetic considerations, and policy support, to build resilient boreal forests under changing disturbance regimes.

  • Lebel Desrosiers, Laboratoire sur la science des données, Université du Québec (TÉLUQ), 5800, rue Saint-Denis, bureau 1105, Montréal, Québec H2S 3L5, Canada ORCID https://orcid.org/0009-0007-1592-8505 E-mail: simon.lebeldesrosiers@teluq.ca
  • Bélanger, Laboratoire sur la science des données, Université du Québec (TÉLUQ), 5800, rue Saint-Denis, bureau 1105, Montréal, Québec H2S 3L5, Canada E-mail: nicolas.belanger@teluq.ca
  • Thiffault, Centre de recherche sur les matériaux renouvelables, Université Laval, 2425 De la Terrasse St, Québec, Québec G1V 0A6, Canada ORCID https://orcid.org/0000-0001-9586-3834 E-mail: evelyne.thiffault@sbf.ulaval.ca
  • Thiffault, Service canadien des forêts, Ressources naturelles Canada, 1055, rue Du P.E.P.S., C.P. 10380, Québec, Québec G1V 4C7, Canada ORCID https://orcid.org/0000-0003-2017-6890 E-mail: nelson.thiffault@canada.ca (email)
article id 24053, category Discussion article
Francisco X. Aguilar, Marie Lautrup, Dohun Kim, Ane C. Tangen, Aapo Rautiainen, Huntley Brownell, Lucas N. López, Andrew D. H. Stratton, Sebastian Glasenapp, Silvia M. Korth, Hanne K. Sjølie, Jette Bredahl Jacobsen. (2024). Inherently spatial: data and analytical insights for the identification of forest socio-ecological hotspots. Silva Fennica vol. 58 no. 5 article id 24053. https://doi.org/10.14214/sf.24053
Keywords: forest sector; spatial analyses; adaptive interventions; ecological-social systems; robustness; spatial data
Abstract | Full text in HTML | Full text in PDF | Author Info

We draw insights regarding intricacies with spatially explicit data and analyses when studying the vulnerability of forest socio-ecological systems to disruptive abiotic and biotic factors. Common issues associated with data include location precision, spatial delimitation, methodological comparability, and measurement consistency. Spatial data analyses are challenged by issues of interpolation and extrapolation, inferences using data at different spatial scales, and assessment of disruption impacts at detectable spatial scales. The inextricable empirical nature of spatial data and analyses requires carefully conducting and disclosing the sensitivity of findings, and including robustness tests to openly inform decision-makers on issues of uncertainty associated with possible interventions. These considerations might be central to identifying forest socio-ecological hotspots as forest-dominated geographic areas encompassing social and ecological systems vulnerable to disruptions caused by abiotic and biotic factors, but where risks to human wellbeing may be considerably reduced through adaptive interventions.

  • Aguilar, Swedish University of Agricultural Sciences (SLU), Department of Forest Economics, SE-901 83 Umeå, Sweden ORCID https://orcid.org/0000-0003-0226-4467 E-mail: francisco.aguilar@slu.se (email)
  • Lautrup, University of Copenhagen, Department of Food and Resource Economics, Rolighedsvej 23, 1958 Frederiksberg C, Denmark E-mail: ml@ifro.ku.dk
  • Kim, Swedish University of Agricultural Sciences (SLU), Department of Forest Economics, SE-901 83 Umeå, Sweden ORCID https://orcid.org/0000-0002-1919-3346 E-mail: dohun.kim@slu.se
  • Tangen, Inland Norway University of Applied Sciences, Department of Forestry and Wildlife Management, P.O. Box 2400, Koppang, Norway ORCID https://orcid.org/0009-0001-3145-8159 E-mail: ane.tange@inn.no
  • Rautiainen, Natural Resources Institute Finland (Luke), Bioeconomy and environment, Latokartanonkaari 9, FI-00790 Helsinki, Finland E-mail: aapo.rautiainen@luke.fi
  • Brownell, University of Copenhagen, Department of Geosciences and Natural Resource Management, Forest and Bioresources, Rolighedsvej 23, 1958 Frederiksberg C, Denmark ORCID https://orcid.org/0000-0002-3798-8783 E-mail: hb@ign.ku.dk
  • López, Inland Norway University of Applied Sciences, Faculty of Applied Ecology, Agricultural Sciences and Biotechnology, P.O. Box 2400, Koppang, Norway ORCID https://orcid.org/0009-0006-6860-3408 E-mail: lucas.lopez@inn.no
  • Stratton, University of Copenhagen, Department of Food and Resource Economics, Rolighedsvej 23, 1958 Frederiksberg C, Denmark ORCID https://orcid.org/0000-0002-6566-2043 E-mail: adhs@ifro.ku.dk
  • Glasenapp, Johann Heinrich von Thünen Institute, Institute of Forestry, Leuschnerstraße 91, 21031 Hamburg-Bergedorf, Germany E-mail: sebastian.glasenapp@thuenen.de
  • Korth, Universidad Nacional de Misiones Facultad de Ciencias Forestales, Bertoni 124, Eldorado 3380, Argentina ORCID https://orcid.org/0009-0007-3261-8234 E-mail: silviakorth@hotmail.com
  • Sjølie, Inland Norway University of Applied Sciences, Faculty of Applied Ecology, Agricultural Sciences and Biotechnology, P.O. Box 2400, Koppang, Norway ORCID https://orcid.org/0000-0001-8099-3521 E-mail: hanne.sjolie@inn.no
  • Bredahl Jacobsen, University of Copenhagen, Department of Food and Resource Economics, Rolighedsvej 23, 1958 Frederiksberg C, Denmark ORCID https://orcid.org/0000-0002-1313-6228 E-mail: jbj@ifro.ku.dk
article id 24004, category Discussion article
Jussi Manner. (2024). Automatic work-element detection: the missing piece in developing intelligent coaching systems for cut-to-length logging machinery. Silva Fennica vol. 58 no. 1 article id 24004. https://doi.org/10.14214/sf.24004
Keywords: forwarder; harvester; work method; operator effect; assistance system; instructor; trainer
Highlights: Next-generation logging systems will crucially impact the future demand for automatic data gathering and work guidance; Artificial intelligence emerges as a gamechanger, prompting re-evaluation of traditional approaches to automatically gather data, especially for forwarders; Industry-wide, interdisciplinary discussions are vital for charting alternative future paths for automatic data gathering and work guidance.
Abstract | Full text in HTML | Full text in PDF | Author Info

The productivity of cut-to-length machine operators exhibits a significant disparity, with the most productive individuals demonstrating twice the efficiency of their less productive counterparts. This discrepancy is largely attributed to variations in work methods. While supervised training has proven effective in streamlining work methods and enhancing productivity, the availability of forest-machine instructors for supervision is limited. Intelligent coaching systems (ICS) are periodically proposed to address this constraint. ICS are computer-based aids that offer machine operators real-time feedback on their work and guide them on how to rationalize their work. The successful implementation of ICS initially requires the development of systems for automatic work-element detection (AWED). Therefore, this article explores the history, current status, and technological possibilities of AWED. Additionally, key features of ICS are briefly reviewed. Lastly, a broader, interdisciplinary discussion is initiated on how to strategically allocate limited research resources. Questions surrounding the feasible ambition level for ICS and AWED are raised, prompting considerations for the next steps in research and development.

article id 23053, category Discussion article
Back Tomas Ersson, Linnea Hansson, Jussi Manner, Per Sandström, Johan Sonesson. (2023). Forest management in northern Fennoscandia: the need for solutions that mitigate conflicts during forest regeneration and increase the use of continuous cover forestry. Silva Fennica vol. 57 no. 3 article id 23053. https://doi.org/10.14214/sf.23053
Keywords: mechanical site preparation; reforestation; reindeer husbandry; indigenous rights; natural resources conflict; Sámi; Sápmi
Highlights: In Sápmi, increased use of continuous cover forestry (CCF) can reduce the frequent conflicts between forest industry and reindeer herding communities; Nordic forestry needs to develop new technical solutions for gentle and lichen-adapted mechanical site preparation during CCF in Sápmi; Such site preparation technology will promote pioneer tree species’ germination and growth, and increase the forest industry’s acceptance of CCF.
Abstract | Full text in HTML | Full text in PDF | Author Info

Today, conflicts often occur in northern Fennoscandia (also known as Sápmi) between forestry and reindeer husbandry. Continuous cover forestry (CCF) is requested by both reindeer herding communities and the general public and is becoming more common, but the forest industry criticizes CCF for lower wood production. Mechanical site preparation (MSP) increases regeneration success and, thus, increases wood production in CCF. To reduce the conflict between forestry and reindeer husbandry, MSP in Sápmi should destroy as little ground lichen as possible. Today, there are no solutions for gentle and lichen-adapted MSP in CCF. Thus, there is a strong need to develop and test new technical solutions that increase regeneration success in a lichen-adapted way during CCF in Sápmi. We suggest that MSP solutions be developed which are gentle, work selectively and function in shelterwoods, gap cuts, and selection cutting stands. We envision that these solutions could fill the gap between the desired adaptivity on the part of the reindeer herding communities and the desired efficiency on the part of the forest industry. Such MSP technology would contribute to increased acceptance of CCF in the forest industry, higher biodiversity, and considerably reduce the conflict between forestry and reindeer herding communities.

  • Ersson, Swedish University of Agricultural Sciences (SLU), School of Forest Management, SE-739 21 Skinnskatteberg, Sweden ORCID https://orcid.org/0000-0003-2442-7482 E-mail: back.tomas.ersson@slu.se (email)
  • Hansson, The Forestry Research Institute of Sweden (Skogforsk), Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID https://orcid.org/0000-0002-9788-1734 E-mail: linnea.hansson@skogforsk.se
  • Manner, The Forestry Research Institute of Sweden (Skogforsk), Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID https://orcid.org/0000-0002-4982-3855 E-mail: jussi.manner@skogforsk.se
  • Sandström, Swedish University of Agricultural Sciences (SLU), Department of Forest Resource Management, SE-901 83 Umeå, Sweden ORCID https://orcid.org/0000-0003-0977-0071 E-mail: per.sandstrom@slu.se
  • Sonesson, The Forestry Research Institute of Sweden (Skogforsk), Uppsala Science Park, SE-751 83 Uppsala, Sweden ORCID https://orcid.org/0000-0002-2018-7496 E-mail: johan.sonesson@skogforsk.se
article id 23034, category Discussion article
Anu Lähteenmäki-Uutela, Salla Rantala, Brent Swallow, Heidi Lehtiniemi, Tuomas Pohjola, Riikka Paloniemi. (2023). Increasing access to forest data for enhancing forest benefits to all. Silva Fennica vol. 57 no. 3 article id 23034. https://doi.org/10.14214/sf.23034
Keywords: Finland; property rights; indigenous rights; access to data; environmental rights; everyone’s rights; institutions
Highlights: Environmental protection, everyone’s forest use, bioeconomy, and tourism may benefit from increased access to forest data; Environmental rights, everyone’s rights and indigenous rights need to be strong and clear for non-landowners to benefit from forest data; Transparent forest data forms the basis for trade in forest ecosystem services.
Abstract | Full text in HTML | Full text in PDF | Author Info

In this discussion paper, we discuss what benefits Finnish citizens and companies can derive from forest data, and how the benefits of that data depend on rights to forests. Environmental protection, everyone’s forest use, bioeconomy, and tourism may benefit from increased access to forest data. Access to forest data is a democratic right by itself. Forest data allow actors to derive more value from their existing forest rights and may spark demands for clarification or reformulation of forest rights. Transparency of forest data also allows voluntary trade in forest ecosystem services. Increased access to forest data may also contribute to forest-related conflicts, given that various, at times contradictory interests are directed at forests. At best, increased access to forest data and information may support the renewal of forest governance to become more democratic, legitimate, and effective.

  • Lähteenmäki-Uutela, Finnish Environment Institute (Syke), Societal Change, Latokartanonkaari 11, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0003-3230-3869 E-mail: anu.lahteenmaki-uutela@syke.fi (email)
  • Rantala, Finnish Environment Institute (Syke), Societal Change, Latokartanonkaari 11, FI-00790 Helsinki, Finland E-mail: salla.rantala@syke.fi
  • Swallow, University of Alberta, Faculty of Agricultural, Life and Environmental Sciences, 5-67 General Services Building, 9007 - 116 St NW, Edmonton, AB, T6G 2H1, Canada E-mail: bswallow@ualberta.ca
  • Lehtiniemi, Finnish Environment Institute (Syke), Societal Change, Latokartanonkaari 11, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0001-7418-5069 E-mail: heidi.lehtiniemi@syke.fi
  • Pohjola, University of Turku, Development Projects, Pohjoisranta 11 A, FI-28100 Pori, Finland ORCID https://orcid.org/0000-0003-3439-7993 E-mail: tuarpo@utu.fi
  • Paloniemi, Finnish Environment Institute (Syke), Societal Change, Latokartanonkaari 11, FI-00790 Helsinki, Finland ORCID https://orcid.org/0000-0003-2853-535X E-mail: riikka.paloniemi@syke.fi
article id 10350, category Discussion article
Maja Peteh. (2020). A synthesis of the characteristics of current national, regional and international forestry journals. Silva Fennica vol. 54 no. 3 article id 10350. https://doi.org/10.14214/sf.10350
Keywords: forestry; geographical coverage; journal title list; scientific publishing
Highlights: Currently, 451 forestry-related journals are published worldwide (41.7% in Europe, 37.3% in Asia) in 32 different languages (47.9% in English only); 6 categories have been defined for the geographical coverage of a journal (55.5% of which have been defined as national and 18.2% as international); The journals could increase their visibility and value by improving their websites and submitting relevant information to international indexing databases.
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One of the main objectives of the International Union of Forest Research Organization’s WP 9.01.06 (Forest Science Publishing) is to clarify the range of forestry-related journals in the context of the overall objective of supporting authors and readers to deepen their knowledge of forestry. This study extended an earlier list of forestry journals using the ISSN database and the CAB abstracts. The study included 451 journal titles, each categorized by ISSN, publisher, language of publication, website, and geographical coverage (i.e. national, regional, international, or three mixed intermediate categories), as well as information on indexing in Web of Science, Scopus, DOAJ, and SherpaRomeo. The included journals are published in 61 countries and in 32 different languages. Those categorized as international are mostly published in English. 21.7% of the journals are indexed in Web of Science and 34.1% in Scopus. 95.6% of the titles are published online or in the print+online publishing model, but only 57.0% of the titles are published in Open Access, of which only 33.7% are indexed in DOAJ.

  • Peteh, University of Ljubljana, Biotechnical Faculty, Večna pot 83, Ljubljana 1000, Slovenia; Slovenian Forestry Institute, Večna pot 2, Ljubljana 1000, Slovenia ORCID https://orcid.org/0000-0002-9636-1155 E-mail: maja.peteh@gozdis.si (email)
article id 1304, category Discussion article
Annika Kangas, Teppo Hujala. (2015). Challenges in publishing: producing, assuring and communicating quality. Silva Fennica vol. 49 no. 4 article id 1304. https://doi.org/10.14214/sf.1304
Keywords: peer review; open access; altmetrics; citation index
Abstract | Full text in HTML | Full text in PDF | Author Info

This paper is based on a session “How to make forest science available for all? Publishers’, editors’, and authors’ challenges” at the IUFRO XXIV world conference, organized by Pekka Nygren and Eeva Korpilahti from the Finnish Society of Forest Science. The presenters dealt with the topical problems of publishing scientific knowledge from different perspectives. The talks covered the development of journals, publications and submissions, benefits and drawbacks of open access publishing as well as electronic and traditional publishing, and possibilities to promote interesting papers either from the journal’s or from the author’s perspective, and the problems of disseminating the scientific results to the end users. In this paper, a few prevalent viewpoints, inspired by the session, are raised and discussed with some suggestions included.

  • Kangas, Natural Resources Institute Finland (Luke), Economics and society, P.O. Box 68, FI-80101 Joensuu, Finland E-mail: annika.kangas@luke.fi (email)
  • Hujala, Natural Resources Institute Finland (Luke), Bio-based business and industry, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: teppo.hujala@luke.fi
article id 119, category Discussion article
Markku Larjavaara, Helene C. Muller-Landau. (2011). Cross-section mass: an improved basis for woody debris necromass inventory. Silva Fennica vol. 45 no. 2 article id 119. https://doi.org/10.14214/sf.119
Keywords: biomass; coarse woody debris (CWD); deadwood; log; void space; nominal volume and wood density; carbon in woody detritus
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  • Larjavaara, Smithsonian Tropical Research Institute, Unit 9100 Box 0948, DPO AA 34002-9998, USA, and University of Helsinki, Department of Forest Sciences, Finland E-mail: larjavaaram@si.edu (email)
  • Muller-Landau, Smithsonian Tropical Research Institute, Unit 9100 Box 0948, DPO AA 34002-9998, USA E-mail: hcml@nn.us
article id 270, category Discussion article
Seppo Rouvinen, Jari Kouki. (2008). The natural northern European boreal forests: unifying the concepts, terminologies, and their application. Silva Fennica vol. 42 no. 1 article id 270. https://doi.org/10.14214/sf.270
Keywords: human influence; biodiversity; natural ecosystems; natural forests; protection; terminology
Abstract | View details | Full text in PDF | Author Info
Recent emphasis on conserving the biodiversity has stressed the value of natural ecosystems in saving the species from extinction. In the Fennoscandian region the conifer-dominated boreal forests form the largest single ecosystem. The forests have been under varying intensity of human influence for decades or centuries. Recent attempts have tried to seek the last remaining natural forests to be included in the protection programmes. However, due to long and widespread human influence, finding and defining the natural forests has proven to be extremely difficult, not only because they are so rare but also because the concept of natural forest is vague. These difficulties are partly seen through the diverse terminology used. We first review the varying terminology as seen in recent studies. Secondly, we propose the basis for defining the natural forest and show some intriguing and challenging difficulties are involved in the concept. These difficulties are at least partly related to inherent strong and long-term dynamic component in boreal forest ecosystems that is manifested over several temporal and spatial scales. Finally, we outline a more general terminology with associated indicators and measurements that might be used in the classification and terminology. Conceptual clarification is necessary, for example, to compile ecologically justified and representative global, national and regional forest statistics. Many currently applied definitions of “forest” and “natural” that are applied in the context of forest statistics overlook ecologically important components of natural forests, and thus provide quite misleading or inadequate data of existing diversity patterns in these ecosystems.
  • Rouvinen, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: sr@nn.fi
  • Kouki, University of Joensuu, Faculty of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: jari.kouki@joensuu.fi (email)
article id 527, category Discussion article
Jyrki Kangas, Ron Store. (2002). Socioecological landscape planning: an approach to multi-functional forest management. Silva Fennica vol. 36 no. 4 article id 527. https://doi.org/10.14214/sf.527
Keywords: forest planning; landscape ecology; multicriteria analysis; sociocultural sustainability
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  • Kangas, Finnish Forest Research Institute, Joensuu Research Centre, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: jyrki.kangas@metla.fi (email)
  • Store, Finnish Forest Research Institute, Kannus Research Station, P.O. Box 44, FIN-69101 Kannus, Finland E-mail: rs@nn.fi
article id 536, category Discussion article
Ian Cowan. (2002). Fit, fitter, fittest; where does optimisation fit in? Silva Fennica vol. 36 no. 3 article id 536. https://doi.org/10.14214/sf.536
Keywords: survival; optimisation; growth; assimilation; nitrogen-use; water-use
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  • Cowan, Research School of Biological Sciences, Australian National University, Canberra, ACT 0200, Australia E-mail: iancowan@bigpond.com.au (email)
article id 573, category Discussion article
Matti Koivula, Jari Niemelä. (2002). Boreal carabid beetles (Coleoptera, Carabidae) in managed spruce forests – a summary of Finnish case studies. Silva Fennica vol. 36 no. 1 article id 573. https://doi.org/10.14214/sf.573
Keywords: management; logging; succession; boreal; forestry; landscape; Carabidae; clear-cutting; edge; retention felling
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  • Koivula, Department of Ecology and Systematics, Division of Population Biology, P.O. Box 65, FIN-00014 University of Helsinki, Finland E-mail: matti.koivula@helsinki.fi (email)
  • Niemelä, Department of Ecology and Systematics, Division of Population Biology, P.O. Box 65, FIN-00014 University of Helsinki, Finland E-mail: jn@nn.fi
article id 572, category Discussion article
Timo Kuuluvainen, Kaisu Aapala, Petri Ahlroth, Mikko Kuusinen, Tapio Lindholm, Tapani Sallantaus, Juha Siitonen, Harri Tukia. (2002). Principles of ecological restoration of boreal forested ecosystems: Finland as an example. Silva Fennica vol. 36 no. 1 article id 572. https://doi.org/10.14214/sf.572
Keywords: biodiversity; ecosystem management; disturbance dynamics; monitoring; heterogeneity; nature conservation; hierarchy
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  • Kuuluvainen, Department of Forest Ecology, University of Helsinki, P.O.Box 27 FIN-00014, Finland E-mail: timo.kuuluvainen@helsinki.fi (email)
  • Aapala, Finnish Environment Institute, Expert Services Department, Nature Division, P.O. Box 140, FIN-00251 Helsinki E-mail: ka@nn.fi
  • Ahlroth, University Museum, Section of Natural History, P.O. Box 35, FIN-40351, Jyväskylä, Finland E-mail: pa@nn.fi
  • Kuusinen, Ministry of the Environment, Land Use Department, P.O.Box 380, FIN-00131 Helsinki, Finland E-mail: mk@nn.fi
  • Lindholm, Finnish Environment Institute, Expert Services Department, Nature Division, P.O. Box 140, FIN-00251 Helsinki E-mail: tl@nn.fi
  • Sallantaus, Pirkanmaa Regional Environment Centre, P.O. Box 297, FIN-33101 Tampere, Finland E-mail: ts@nn.fi
  • Siitonen, Finnish Forest Research Institute, Vantaa Research Centre, P.O. Box 18, FIN-01301 Vantaa, Finland E-mail: juha.siitonen@metla.fi
  • Tukia, Finnish Environment Institute, Expert Services Department, Nature Division, P.O. Box 140, FIN-00251 Helsinki E-mail: ht@nn.fi
article id 571, category Discussion article
S. C. DeLong. (2002). Using nature’s template to best advantage in the Canadian boreal forest. Silva Fennica vol. 36 no. 1 article id 571. https://doi.org/10.14214/sf.571
Keywords: biodiversity; ecosystem management; sustainable forestry; natural disturbance; British Columbia
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  • DeLong, Ministry of Forests, 1011 4th Ave, Prince George, BC, Canada V2L 3H9 E-mail: craig.delong@gems1.gov.bc.ca (email)
article id 570, category Discussion article
Anke Jentsch, Carl Beierkuhnlein, Peter S. White. (2002). Scale, the dynamic stability of forest ecosystems, and the persistence of biodiversity. Silva Fennica vol. 36 no. 1 article id 570. https://doi.org/10.14214/sf.570
Keywords: biodiversity; disturbance; multi-patch scale; pattern emergence; resilience; qualitative equilibrium
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  • Jentsch, Conservation Biology and Ecological Modelling, UFZ – Centre for Environmental Research, Permoserstr. 15, D-04301 Leipzig, Germany E-mail: jentsch@pro.ufz.de (email)
  • Beierkuhnlein, Department of Landscape Ecology, University of Rostock, Justus-Liebig-Weg 6, D-18051 Rostock, Germany E-mail: cb@nn.de
  • White, Department of Biology, Campus Box 3280, University of North Carolina at Chapel Hill, Chapel Hill, NC 27599-3280, USA E-mail: psw@nn.us
article id 610, category Discussion article
Risto Päivinen, Perttu Anttila. (2001). How reliable is a satellite forest inventory? Silva Fennica vol. 35 no. 1 article id 610. https://doi.org/10.14214/sf.610
Keywords: forest inventory; remote sensing; satellite images
View details | Full text in PDF | Author Info
  • Päivinen, European Forest Institute, Torikatu 34, FIN-80101 Joensuu, Finland E-mail: risto.paivinen@efi.fi (email)
  • Anttila, European Forest Institute, Torikatu 34, FIN-80101 Joensuu, Finland E-mail: pa@nn.fi
article id 680, category Discussion article
Sakari Mykrä, Sami Kurki. (1998). ESC-strategy for rational operationalization of forest biodiversity maintenance in Finland. Silva Fennica vol. 32 no. 4 article id 680. https://doi.org/10.14214/sf.680
Keywords: biodiversity; Finland; ESC
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  • Mykrä, Section of Ecology, Department of Biology, University of Turku, FIN-20014 Turku, Finland E-mail: sakmyk@utu.fi (email)
  • Kurki, Section of Ecology, Department of Biology, University of Turku, FIN-20014 Turku, Finland E-mail: sk@nn.fi

Category : Commentary

article id 6985, category Commentary
Eshetu Yirdaw, Markku Kanninen, Mohamed Elfadl, Daniel Tsegai. (2017). Special issue: Drought and Dryland Management – a commentary. Silva Fennica vol. 51 no. 1B article id 6985. https://doi.org/10.14214/sf.6985
Keywords: restoration; desertification; land degradation; rangelands; croplands; dry forests; landscapes
Full text in HTML | Full text in PDF | Author Info
  • Yirdaw, Viikki Tropical Resources Institute (VITRI), Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: eshetu.yirdaw@helsinki.fi (email)
  • Kanninen, Viikki Tropical Resources Institute (VITRI), Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: markku.kanninen@helsinki.fi
  • Elfadl, Viikki Tropical Resources Institute (VITRI), Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: mohamed.elfadl@helsinki.fi
  • Tsegai, UNCCD Secretariat, Platz der Vereinten Nationen 1, 53113 Bonn, Germany E-mail: dtsegai@unccd.int
article id 445, category Commentary
Bengt Gunnar Jonsson, Jari Kouki, Timo Kuuluvainen. (2011). Northern Primeval Forests – Ecology, Conservation and Management. Silva Fennica vol. 45 no. 5 article id 445. https://doi.org/10.14214/sf.445
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  • Jonsson, Mid Sweden University, Dept. of Natural Sciences, Sundsvall, Sweden E-mail: bengt-gunnar.jonsson@miun.se (email)
  • Kouki, University of Eastern Finland, School of Forest Sciences, Joensuu, Finland E-mail: jari.kouki@uef.fi
  • Kuuluvainen, University of Helsinki, Dept. of Forest Science, Helsinki, Finland E-mail: timo.kuuluvainen@helsinki.fi
article id 448, category Commentary
Pia Katila, Marko Katila. (2011). New paradigm for saving the world’s forests? Silva Fennica vol. 45 no. 4 article id 448. https://doi.org/10.14214/sf.448
Abstract | View details | Full text in PDF | Author Info
Book review: Douglas, J. and Simula, M. 2010. The Future of the World’s Forests. Ideas vs Ideologies. World Forests, volume 7. Springer. 211p.
  • Katila, Finnish Forest Research Institute E-mail: pia.katila@metla.fi (email)
  • Katila, Dasos Capital Oy E-mail: mk@nn.fi
article id 461, category Commentary
Sari Palmroth. (2009). Boreal forest and climate change – from processes and transport to trees, ecosystems and atmosphere. Silva Fennica vol. 43 no. 4 article id 461. https://doi.org/10.14214/sf.461
Keywords: carbon; climate; growth; forests; ecology; book reviews
Abstract | View details | Full text in PDF | Author Info
Book review: Hari, P. & Kulmala, L. (eds.). 2008. Boreal Forest and Climate Change. Advances in Global Change Research 34. Springer. 582 p. ISBN 978-1-4020-8717-2.
  • Palmroth, Duke University, Nicholas School of the Environment, Box 90328 Duke University, Durham, USA E-mail: sari.palmroth@duke.edu (email)
article id 462, category Commentary
Jean-Michel Leban, Heli Peltola, Sanna Leinonen, Christine Todoroki, Sophie D'Amours, Barry Gardiner, Jim Goudie, Eini Lowell, Thomas Seifert, Erkki Verkasalo, Lars Wilhelmsson, Tuula Nuutinen. (2009). Preface. Silva Fennica vol. 43 no. 3 article id 462. https://doi.org/10.14214/sf.462
Abstract | View details | Full text in PDF | Author Info
The 6th IUFRO Workshop on Connection between Forest Resources and Wood Quality: Modelling Approaches and Simulation Software.
  • Leban, E-mail: jml@nn.fr (email)
  • Peltola, University of Eastern Finland, School of Forest Sciences, P.O. Box 111, FI-80101 Joensuu, Finland E-mail: heli.peltola@uef.fi
  • Leinonen, E-mail: sl@nn.fi
  • Todoroki, E-mail: ct@nn.nz
  • D'Amours, E-mail: sda@nn.ca
  • Gardiner, E-mail: bg@nn.uk
  • Goudie, E-mail: jg@nn.ca
  • Lowell, E-mail: el@nn.us
  • Seifert, E-mail: ts@nn.de
  • Verkasalo, Finnish Forest Research Institute, Joensuu Unit, P.O. Box 68, FI-80101, Joensuu, Finland E-mail: erkki.verkasalo@metla.fi
  • Wilhelmsson, E-mail: lw@nn.se
  • Nuutinen, European Forest Institute, Torikatu 34, FI-80100 Joensuu, Finland E-mail: tuula.nuutinen@efi.int
article id 475, category Commentary
Petteri Muukkonen, Raisa Mäkipää. (2006). Biomass equations for European trees: addendum. Silva Fennica vol. 40 no. 4 article id 475. https://doi.org/10.14214/sf.475
Keywords: aboveground biomass; allometry; biomass functions; belowground biomass; dbh; tree diameter; tree height
Abstract | View details | Full text in PDF | Author Info
A review of stem volume and biomass equations for tree species growing in Europe (Zianis et al. 2005) resulted in suggestions for additional equations. The numbers of original equations, compiled from scientific articles were 607 for biomass and 230 for stem volume. On the basis of the suggestions and an updated literature search, some new equations were published after our review, but more equations were also available from earlier literature. In this addendum, an additional 188 biomass equations and 8 volume equations are presented. One new tree species (Pinus cembra) is included in the list of volume equations. Biomass equations for twelve new tree species are presented: Abies alba, Carbinus betulus, Larix decidua, P. cembra, P. nigra, Quercus robur, Salix caprea, S. ‘Aquatica’, S. dasyclados, S. phylicifolias, S. triandra and S. accuparia. The tree-level equations predict stem volume, whole tree biomass or biomass of certain components (e.g., foliage, roots, total above-ground) as a function of diameter or diameter and height of a tree. Biomass and volume equations with other independent variables have also been widely developed but they are excluded from this addendum because the variables selected may reflect locally valid dependencies that cannot be generalized to other geographical regions. Most of the equations presented here are developed for Sweden, Finland and Norway in northern Europe, for Austria in central Europe and for Italy in southern Europe. There are also few equations from Poland and Belgium. Most of the equations deal with above-ground components such as stem, branches and foliage, but some new equations are also available for root biomass. Zianis et al. (2005) and this addendum can be used together as guides to the original publications of these equations. Our updated database of the biomass and volume equations is available also from the website www.metla.fi/hanke/3306/tietokanta.htm.
  • Muukkonen, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: petteri.muukkonen@metla.fi
  • Mäkipää, Finnish Forest Research Institute, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: raisa.makipaa@metla.fi (email)
article id 480, category Commentary
Pasi Puttonen, Sauli Valkonen. (2004). Preface. Silva Fennica vol. 38 no. 4 article id 480. https://doi.org/10.14214/sf.480
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  • Puttonen, University of Helsinki, Department of Forest Sciences, P.O. Box 27, FI-00014 University of Helsinki, Finland E-mail: pasi.puttonen@helsinki.fi (email)
  • Valkonen, Finnish Forest Research Institute, Vantaa Unit, P.O. Box 18, FI-01301 Vantaa, Finland E-mail: sauli.valkonen@metla.fi
article id 528, category Commentary
Annikki Mäkelä, Thomas J. Givnish, Frank Berninger, Thomas N. Buckley, Graham D. Farquhar, Pertti Hari. (2002). Challenges and opportunities of the optimality approach in plant ecology. Silva Fennica vol. 36 no. 3 article id 528. https://doi.org/10.14214/sf.528
Keywords: models; acclimation; adaptation; optimisation; evolution; hypotheses; evaluation
Abstract | View details | Full text in PDF | Author Info
A meeting was held in Hyytiälä, Finland 10–12 April 2000 to assess critically the current challenges and limitations of the optimality approach in plant ecophysiology and botany. This article summarises the general discussions and views of the participants on the use of optimisation models as tools in plant ecophysiological research. A general framework of the evolutionary optimisation problem is sketched with a review of applications, typically involved with balanced regulation between parallel processes. The usefulness and limitations of the approach are discussed in terms of published examples, with special reference to model testing. We conclude that, regardless of inevitable problems of model formulation, wider application of the optimality approach could provide a step forward in plant ecophysiology. A major role of evolutionary theory in this process is simply the formulation of testable hypotheses, the evaluation of which can lead to important advances in our ecophysiological understanding and predictive ability.
  • Mäkelä, University of Helsinki, Dept. of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: annikki.makela@helsinki.fi (email)
  • Givnish, University of Wisconsin, Department of Botany, Madison, WI 53706 USA E-mail: tjg@nn.us
  • Berninger, University of Helsinki, Dept. of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: fb@nn.fi
  • Buckley, Cooperative Research Centre for Greenhouse Accounting and Environmental Biology Group, and Research School of Biological Sciences, Australian National University, ACT 2601, Australia E-mail: tnb@nn.au
  • Farquhar, Cooperative Research Centre for Greenhouse Accounting and Environmental Biology Group, and Research School of Biological Sciences, Australian National University, ACT 2601, Australia E-mail: gdf@nn.au
  • Hari, University of Helsinki, Dept. of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: ph@nn.fi
article id 547, category Commentary
Timo Kuuluvainen. (2002). Introduction. Disturbance dynamics in boreal forests: defining the ecological basis of restoration and management of biodiversity. Silva Fennica vol. 36 no. 1 article id 547. https://doi.org/10.14214/sf.547
Keywords: boreal forests; management; biodiversity; disturbance dynamics; restoration; ecology
View details | Full text in PDF | Author Info
  • Kuuluvainen, Department of Forest Ecology, P.O. Box 27, FIN-00014 University of Helsinki, Finland E-mail: timo.kuuluvainen@helsinki.fi (email)

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